Новости систематики высших растений Том 36 2004 ОТ ...

Новости систематики высших растений 

Том 36 

2004 

ОТ РЕДАКТОРА 

3-6 

О РОДЕ BOTRYCHIUM SWARTZ (OPHIOGLOSSACEAE) В РОССИИ 

Цвелев Н.Н.7-21 

О РОДЕ SELAGINELLA P. BEAUV. S. L. (SELAGINELLACEAE) В РОССИИ 

Цвелев Н.Н.22-27 

НОВЫЕ ВИДЫ РОДОВ FESTUCA L. (РОАСЕАЕ), PHLOMOIDES MOENCH 

(LAMIACEAE), PRIMULA L. (PRIMULACEAE) И ACANTHOLIMON BOISS. 

(LIMONIACEAE) ИЗ КЫРГЫЗСТАНА 

Лазьков Г.А.28-39 

РОД SCIRPUS L. (CYPERACEAE) ВО ФЛОРЕ ЕВРАЗИИ 

Егорова Т.В.40-79 

СИСТЕМА РОДА BOLBOSCHOENUS (ASCHERS.) PALLA (CYPERACEAE) 

Татанов И.В.80-95 

НОВЫЕ ВИДЫ РОДОВ ALLIUM L. (ALLIACEAE) И GLADIOLUS L. 

(IRIDACEAE) С КАВКАЗА 

Михеев А.Д.96-100 

НОВЫЕ КОМБИНАЦИИ В РОДЕ ALLIUM L. (ALLIACEAE) 

Серегин А.П.101-102 

КРИТИЧЕСКИЕ ЗАМЕТКИ О ВИДАХ РОДОВ ORNITHOGALUM L. 

(HYACINTHACEAE) И EREMURUS BIEB. (ASPHODELACEAE) ФЛОРЫ 

КАВКАЗА 

Михеев А.Д.103-105 

РОД HOLOSTEUM L. (CARYОРНYLLACEAE) ВО ФЛОРЕ КАВКАЗА 

Зернов А.С., Соколов И.В.106-111 

ВИДЫ РОДА ROSA L. (ROSACEAE) СЕКЦИИ CINNAMOMEAE DC. ВО 

ФЛОРЕ КАВКАЗА 

Бузунова И.О., Камелин Р.В.112-122 

АСТРАГАЛЫ (ASTRAGALUS L., FABACEAE) СЕКЦИИ XIPHIDIUM 

BUNGE ВО ФЛОРЕ УРАЛА 

Князев М.С., Куликов П.В.123-148 

КОНСПЕКТ КАВКАЗСКИХ АСТРАГАЛОВ (ASTRAGALUS L., FABACEAE) 

СЕКЦИИ ANTHYLLOIDEI DC 

Сытин А.К.149-158 

ОБЗОР ПОДСЕКЦИИ MYRS1NITEAE BOISS. СЕКЦИИ PARALIAS 

DUMORT. РОДА EUPHORBIA L. (EUPHORBIACEAE) 

Гельтман Д.В.159-169 

ДВЕ НОВЫЕ КОМБИНАЦИИ В РОДЕ LEUTEA М. PIMEN. (APIACEAE)

Виноградова В.М.170-177 

О ВИДАХ СЕКЦИИ SIDERITOPSIS BHATTACHARJEE РОДА STACHYS L. 

(LAMIACEAE) 

Крестовская Т.В.178-181 

ОБЗОР РОДА LINARIA MILL. (SCROPHULARIACEAE) ВОСТОЧНОЙ 

ЕВРОПЫ И КАВКАЗА 

Пескова И.М.182-208 

НОВЫЙ ВИД РОДА VERONICA L. (SCROPHULARIACEAE) ИЗ 

ЗАПАДНОЙ МОНГОЛИИ 

Косачёв П.А., Герман Д.А.209-212 

О РОДЕ SPHAERANTHUS L. И СИСТЕМЕ СЛОЖНОЦВЕТНЫХ 

(ASTERACEAE) 

Цвелёв Н.Н.213-222 

НОВЫЕ ТАКСОНЫ И НОВЫЕ КОМБИНАЦИИ В РОДАХ РАСТЕНИЙ 

ФЛОРЫ КАВКАЗА 

Зернов А.С.223-228 

О НОВЫХ И РЕДКИХ ВИДАХ ДЛЯ ФЛОРЫ КЫРГЫЗСТАНА 

Лазьков Г.А.229-233 

ТИПОВЫЕ ОБРАЗЦЫ ТАКСОНОВ СЕМЕЙСТВА ROSACEAE, 

ОПИСАННЫХ ИЗ КИТАЯ, ХРАНЯЩИЕСЯ В ГЕРБАРИИ 

БОТАНИЧЕСКОГО ИНСТИТУТА ИМ. В. Л. КОМАРОВА (LE). I. 

ПОДСЕМЕЙСТВО SPIRAEOIDEAE AGARDH 

Бузунова И.О.234-238 

ТИПИФИКАЦИЯ НАЗВАНИЙ ТАКСОНОВ СЕМЕЙСТВА RUBIACEAE 

БРАЗИЛИИ, СОБРАННЫХ Ф. СЕЛЛО И ОПИСАННЫХ А. ШАМИССО И Д. 

ШЛЕХТЕНДАЛЕМ. I. ПОДСЕМЕЙСТВО RUBIOIDEAE: 

COCCOCYPSELUM Р. BROWNE, FARAMEA AUBL. И RUDGEA SALISB 

Имханицкая Н.Н.239-255 

ТИПОВЫЕ ОБРАЗЦЫ ТАКСОНОВ СЕМЕЙСТВА FABACEAE КИТАЯ, 

ХРАНЯЩИЕСЯ В ГЕРБАРИИ БОТАНИЧЕСКОГО ИНСТИТУТА ИМ. В. Л. 

КОМАРОВА (LE). II 

Новосёлова М.С., Росков Ю.Р.256-262 

ТИПОВЫЕ ОБРАЗЦЫ ТАКСОНОВ РОДОВ ASTRAGALUS L. И 

OXYTROPIS DC. (FABACEAE) КИТАЯ И МОНГОЛИИ, ХРАНЯЩИЕСЯ В 

ГЕРБАРИИ БОТАНИЧЕСКОГО ИНСТИТУТА ИМ. В. Л. КОМАРОВА (LE) 

Новосёлова М.С., Сытин А.К.263-269 

ТИПОВЫЕ ОБРАЗЦЫ НЕКОТОРЫХ ТАКСОНОВ РОДА ABIES MILL. 

(PINACEAE), ХРАНЯЩИЕСЯ В ГЕРБАРИИ БОТАНИЧЕСКОГО 

ИНСТИТУТА ИМ. В. Л. КОМАРОВА (LE) 

Орлова Л.В.270-276 

ХРОНОЛОГИЧЕСКИЙ ПЕРЕЧЕНЬ ДАТ, СОДЕРЖАЩИХСЯ В РАБОТЕ 

«МЕЖДУНАРОДНЫЙ КОДЕКС БОТАНИЧЕСКОЙ НОМЕНКЛАТУРЫ 

(СЕНТ-ЛУИССКИЙ КОДЕКС)» 

Егорова Т.В.277-283

НОМЕНКЛАТУРНЫЕ ЗАМЕТКИ 

284-285 

АЛФАВИТНЫЙ УКАЗАТЕЛЬ НАЗВАНИЙ ТАКСОНОВ 

286-296 

АЛФАВИТНЫЙ УКАЗАТЕЛЬ НОВЫХ НАЗВАНИЙ ТАКСОНОВ 

297-298

OT PEAAKTOPA 

A REDACTORE 

Tpugqarr urecroft ToM exefo4H[ra >, KaK r,r flpeAbrAytque roMa 3Toro vrg1aHyrs, coAepxr.rr orrucaHr4t 

HoBbrx raKcoHoB, Kprlrr,rqecKr,re o6:opu pa3Jrr4qHbrx cr4creMarr,r{ecKr,rx 

rpyrrrr cocyArlcrbrx pacrenufi, 3aMerK]r o6 orAellntrx pacreHnrx, 

coo6ulenur o galr6onee BaxHbrx $;ropucruuecKl4x HaxoArax, a raKxe 

AaHHbre o xpanrr.q[xcs s LE rurroBbrx o6pasuax raKcoHoB prga ceueftcrn 

r,r poAoB. B pas4ene

3ToMy Aatoi.t Aeftcrsuremnoro o6HapoAoBaHr{r Ha3BaHufl raKcoHa cre- 

Ayer crrnrarb Aary ny6narauuu KHrrrrr r,rJrr,r xypHaJra, fAe AaHHbrfi rar

HapoAHbrx corpauleunft nassaHufi uaguefturwx rep6apraen vrnpur (3au- 

KoHHr.rKoBa, 1966 :331 345) u He6oruuruu AonoJrHeHr,reM K HeMy (1974: 

3),

(1996) pa6ore M. 3. KapnuqHuKoBa

H. H.I{neren N. Tzvelev 

o POAE BOTRYCHTUM SWARTZ (OPHTOGLOSSACEAE) 

B POCCNN 

DE GENERE BOTRYCHIUM SWARTZ (OPHIOGLOSSACEAE) 

IN ROSSIA 

OAHo us Apesnefiurux ceMeficrs coBpeMeHHbrx cocyAr4crbrx pacretruit 

- Ophioglossaceae, 3anvrMaer oqest o6oco6leusoe uoJroxeHr.re cpe- 

Ar{ [aloporulrKoB (Polypodiophyta), no:uoxHo Aaxe 3acnyxr,rBar BbrAe- 

JIeHr4r He rolurto B KarrecrBe oco6oro Kiracca - Ophioglossopsida, no u 

B KaqecrBe caMocrorreJrbHoro orAeJra. O.renl r,rH'repecHo, rrro 3Ta fpy[tra 

[oKa3brBaer cyrqecrBeHHoe cxo4slBo c He MeHee opulunzurrHofi rpynuofi 

nrruepurux (ureecrnbrx o'r cpeAHero AeBoHa Ao Harrala xap6oHa) 

ApeBoBr4AHbrx [aroporHr4KoBrlAHbrx -* Kilacca Ar chae opt eridop s ida c 

Tr4rroM Archaeopteris J. F. Dawson, 6nnunrx, oAHaKo, yxe pa3Hoc[opo- 

BbrMr{. Cyar no Jrr{reparypHbrM AaHHbrM (Taxragxru , 1978; Cunrupeacrar, 

1995), BrroirHe BeporrHo, uro'Archaeopteris, lo\o6Ho raKxe pa3- 

HocrropoBoMy poAy Lepidodendrolr Sternb. cpeAr.r rrJrayHoBrlAHbrx (Ly c o - 

podiophyta),6un perynrraroM 3BoJrroq[r4 B oqeHb BbrroAHoM B or].rr4- 

MZUIbHbTX AJIS XU3Hr4 yCJTOBI{flX HarrpaBJleHur,r fpaHAr{3aur{r,r oT KaKr{x-l.o 

repButrHbrx Ophioglossopsida. H. C. CHulrapencral (1995 : 73) nnonHe 

o6ocHoeaHHo c{uraer, tro Archaeopteris 6wn nlrut uplr6lrExasilrefrcr 

K loJloceMeHnuru cneuoil BerBbIo 3BoJIIorIr{H pa3HocuopoBbrx [aflopor- 

HrrKoBttAHbrx, a [peAKoB rlepBbrx roJroceMeHHbrx (cJreAyeT ucKaTb cpe- 

Ar4 KycrapHtrKoBbrx rlJl?r Aaxe rpaB.f,Hr,rcrbrx pacreuufi>. Kpyunue, He 

co6pauuue B copycbr crropaHrnr,r, pacrroJroxeHHbre Ha oco6brx qacrsx 

aaft, xoropue y Ophioglossales npuutro Ha3brBarb cloporfoparvur B or- 

Jruqr4e or Bererarr{nHmx, Sorocr,rHTe:r4pyrcul,rx .racreft - rpo$o$opon, 

He u3orHyTr,re ylnrroo6pa3Ho MoJloAbte Bai4u, [pr,rcyTcTBr{e 6rusxofi no 

crpoeHuro nropuvHoft KcnJreMbl r4 Apyrne npusnaril flBJrflrorcr o6ururr.ul 

Als o6onx KJraccoB. 

Ophiog lossop sida, c oAsoft cropoHbr, rBrrf,rorcq rrpt4Ml4Tr.rBHbtMlr rranoporHl4KoBrrAHbrMr,r 

(rax, oveHt rrp u Mr.trr,rBHbrM rIpIz3HaKoM MoxHo 

crluTaTb rrpr4cyTcTBlre npoBoArulr4x [yrrKoB y ocHoBaHr4s KaxAor-o 

cuopaHrux) r4 B Hacrof,lqee BpeMf, o6H.rHo [oMerlaercr B HaqaJre HX 

crrcreMbr, a c gpyrofi cropoHbr, o6na4aror prAoM rrpoABrrHyrbrx [pr43Haxon, 

c6luxarculux r4x c roJloceMeHHbrMr4 (Pinophyta), qro rro3BoJrrer 

cqr4Tarb r,rx ocrarKoM BbrMeprrJtlx npeAKoB roJroceMeHHrrx (Chau, 1986; 

Kato, 1988; Takabashi, Kato, 1988). flpucyrcrnue y Ophioglossopsidu 

BbrcoKocfleqrlaJrr{3rrporauHofi au4ovrr4ropu3br urt (I{nenen, 2000 : 62) 

TaKXe CKJIOHHbT cql.ITarb [prrMlITrlBHOft OCO6eHHOCrbrc, yHacJreAosauuoft 

rIo3AHee rrepBuqHHMI4 f OrOCeMeHHbrMIt t{ rroKpbtroceMeHHbrMt4 pacre- 

HNAMII.

B oruoureuun cucreMbr Ophioglossopsida toxa eure Her eal'tHol'o MHe- 

Hvtfl, v Bo3MoxHbI aJIbrepHarI,IBHbIe ee BapI,IaHTbL Tax, 3 po4a' ltoLo 

KJracca -- Botrychium Swartz, Helminthostachys Kaulfuss r'r Ophioglossum, 

HacroJrbKo ornlrqalorcfl ApyI'or Apyfa, qro BuoltHe 3acJlyxen- 

Ho BbrAerqnrcs s oco6tre floAceMeficrBa oAHoro ceMeftcrBa Ophioglossaceae 

Agardh (uaupauep, Taxra4xau, 1978) ur'u Aaxe caMocrotreJlbHue 

ceneficrsa (Haupnuep, IIIv36es, 2001), ur Koropblx Botrychiaceae 

to coBoKyrlHocrl,t rIpH3HaKos Hal,I6oree upnMI'ITI4BHo. Pol, Ophioglossum3aMeqarereH 

oqeHb BblcoKI,IMI4 xpoMocoMHbtMl4 rrl'{cJlaMll Ao 

2n = 1320 (

peAcraBJreHbr r,r roJrbKo Bererarr4BHoft qacrnrc naitu -- rpo$oSopou, 

rlro cBr,rAereJrbcrByer o HeKoropofr Hegasucur,rocru rpoQoQopa u crroporfopa 

Apyr or Apyra r4 o Haqzure $yurquouanbHoro pa3AeJreHr{fl lrpe)K- 

Ae eAr,rHoro Surouepa Ha 2 Surouepa. Ognaro [oApoA Osmundopteris, 

Iauerouufi 6olee rpyrrHne oco6u c MHoroKparHo paccerreHuunau rpoQo- 

Qopanau, Kaxercr 6olee npuurrrlrBHbrM, HecMolpr Ha rrptrcyrcrBr,re B 

HeM BUAoB c xopolxo parnnrofi nropuunoft rcnlenrofi, nueroqeft nyun 

(Taxra4xrH, 1978 : 169), .rro He Moxer cqurarbcr rrpr,rMrrrlrBHbrM rrpn- 

3HaKoM. Y cerqnu Lanuginosae Clausen sToro rroApoga, co4epxaulefi 

ruualaficruit B. lanuginosum c 6[usx[Mr,r BtrAaMn, cuopo$opou craHo- 

Br4Tcr oAHa ul sernefi rpoQoSopa, Apyrue BerBfl Koropofo rroxoxlr Ha 

caMocrorreJrtuue rpoQo$opu c AoBoJrbHo AJrr4HHbrMu qepeuxoBr,rAHbt- 

MI{ HO)KKAMI{. 

V 6orrru,lHcrBa BuAoB rroApoAa Sceptridium ocHoBHar nafix, cocr.au- 

JreHHaq rpo$otfoporra u cnopo$opoM, K KoHqy rrera orMr,rpaer, Ho Qop- 

Mrlpyercfl erqe oAHa BrroJtHe caMocrorrejlbHar 3r,IMyroulaa nafrr, upe4- 

CTABJIEHHA' TOJIbKO TPOQOSOPOU,'BJIf,IOIIIIIMCq BTOPbIM, HO BE|CTATNB- 

Huru Surorvrepona. Y 6onlurflHcrsa Apyrlrx rraroporHr4KoBr.rAHbrx qKcro 

BereraruBHblx slrrouepoB cr€uro yBenllr{l,IBarbct tryreM [oJruMeptr3auuH, 

BbrfoAHocrb qero Arr Bo3Hr,rKaroqeft raxuu [yreM

156). Ounaro y B. simplex Her 3l4Mylouux sail rpotfotpopoB, qro 

rro3Borfler ocraBttrb efo B lloApoge Botrychium. 

fra6pra4Hoe rpoucxoxAetue B. matricariifulium u B' simplex xopouo 

corJracyercr c HaruuMt4 Ha6npAeHuqMu B rlpupoAe. Tar, B' simplex 

co6upalca HaMI4 Ha MexAIoHHoM rIoHHxeHlrI4 6lus ycrrr p. Konauru n 

JlenuHrpagcxoft o6lacru u Ha 6eperoBoM BaJly flcroecrol'o o3epa K ce- 

Bepy or Horor.r:l6opcKa, I{ s o6oux MecroHaxoxAeH[tx ge6ottlurrle 

rpyrrrrbr oco6efi B. simplex pacrloJlalar[cb Bo3Jle 6oree usoror{HcJleHur,rx 

oco6efr B. multi/idum. Ha cyxotrl Jlyfy rloJlororo cKJIoHa:tesol'o 6epera 

p. llxopu y x.-A. cr. flyAocrl farqnucroro p-Ha JleHI'rurpa4crofr 

o6racruo rge B. simplex 6ul uafiAeH A. O. Xaape (ycrHoe coo6uresne), 

Bcrpeqalorc x w o6a rlpeAlloJlaraeMblx poAl4TerbcKllx slaAa. Ha 6otturuoil 

noJrqHe B cocHoBoM .uecy 6nns g. E. Kpyrelu Jlyxcroro p-Ha JIeuuHlpa4crofi 

o6[actu, rAe HaMu 6utu HailAenu 6onee AecflTKa xopotuo 

pa3Burbrx oco6efi B. matricariiJblium, n He[ocpeAcrBeHHofi 6nrsocrn or 

Hlrx lrMelorcs xar oco6u B. lunaria, TaK lr oco6u B' multiJidum. 3ru AaH- 

Hbte cBr.rAeTeJIbcTByrcT O BO3MOXHOCTI4 MeXBIIAOnOfi (n AalbHepoAcrBeH- 

Hofi!) ru6puAll3aunl4 u B Hacrotulee BpeMfl. 

Huxe upuroAuM KparKnil o6sop poccuficrux BI'lAoB Botrychium vt' 

Knroq AJrr r4x orlpeAeileHl4r c yqeroM Honefturux pa6or rro ceBepoaMepl4- 

KaHcKr4M BI{AaM 3rofo poAa (Wagner, Wagner, 1993; Stensvold et al', 

2002; Wagner, Grant, 2002). CuHoHllMI{Ka AaHa Jlltlxb cauas ueo6xoAuvraa. 

CyurecrBeHHble rpyAHocrl4 uMelorct rrpn o[peAeJreHun BllAoB lloApo;:a 

Botrvchium, rpo$oQopu roropblx oqeHb Bapbl4pylor B 3aBl{cllMocrr{ 

or BeJlI4qI{Hbr oco6efi. 

l. 

PoD, Botrychiam SwarIz,l802, Journ. Bot. (Goettingen) 1800, 2 : ll0' 

L e c t o t y p u s : B. lunaria (L.) Swartz. 

Tpo$oQop Ha BcerAa galrersofi HoxKe orxoAur n Hl'lxHefi qerBeprr'I o6urefi 

ocn safiu 6rns norepxuocru 3eMJIH ......... 2. 

+ Tpo0o0op cugrrafi vruHa roporxofi HoxKe, orxorllr n nepxneii nofloBnHe 

o6ureft ocn saftfi, AaJIeKo or noBepxHocrx 3eMflfi .. . . . 5. 

2. Pacrenue 3-10 crra nrrc. Tpo{o{op Ha oqeHb xoporxoft (ao 1.5 cna 4r.) nox- 

Ke; nJracrr4HKa ero or rlelr'uoii, oontqno HeMHoro rpexnonacrHofi 4o rpoil- 

.raroft c nepHcropa3AeJIbHbIMI,I cerMeHTaMLI Ha rrepeiloqxax. 3 n uyroullrfi 

rpo$orfop orcyrcrByer . . 9. B. simplex. 

* Pac'reune 7-30 crvr ruc. TpoQoSop Ha ArIIlHHoft (6onee 2 cna rur.) HoxKe; 

rrJracTr4Hxa ero ABaxAbI finr,I TpI,rx,4bI rlepficTopacceqeHHas c nepncropa3- 

AeJrbHbrMr,r rrJrr4 nepr4crotronacrHblMl,I KoHeqHbIMI,I cerMeHTaMI,r. 14ueercr 

srar',ryrcurnfirpo$ooop ......3. 

3. TpoQo$op rouxnft, ne urcncruft, c xopouo 3aMerHbIM xLInKoBaHI{eM; ero 

Konequbre cerMeHTbI ocrpoBarble u o6uqno sy6varue . . . 10. B.lernatum. 

+ Tpo0o0op HeMHoro uxcllcrrtft, c eABa 3aMerHbIM )KI4JIKoBaHuelr... ... 4' 

4. Koneqnrre cerMeHTbr rpor[o$opa ocrpoBarble u o6ntqHo sy6.rarrte. . . . . . 

ll. B. robuslunr. 

l0

* KoneqHrre cerMeHTbr rpo$otfopa rynbre, qenruoxpaftHbre rurn c oAriHor{Hbr- 

Mu rynbrMr.r ay6qaur.r . 12. B. multifidum. 

5 (l). Pacrenus20-40 cM Bbrc. Tpo$oQop cu1nuuir, rpnxAbr nepucropacceqeH- 

Hrrft,uoxerHer.rMerbcnopor[opa ..........6. 

* Pacrenus 3-25 cu enc. Tporfcirfop cugrvufi r4rru Ha xoporxoft HoxKe, oAr{H 

pa3 r.rnn ABaxAbr nepacropacceqennrrfi, BcerAa co cnoporpoporrl . . . . . . 7. 

6. Cnoporlop nauqeruuft; ero 6oxosrre Aonu o6uqno pasxoft aJrr4Hbl, HepeA- 

Ko pa3BerBJreHnue. Hapyxurre 6agan*rue nepbrrxKr,r caMbrx HHxHr,rx nepbeB 

rporlo$opa MeHee qeM n 2 pala Kopoqe caMblx AJrr4HHhrx nepbrueK rroft xe 

cropoHbr. ....1. B.tttttltemoides. 

+ Cnopot[op nr.ruefinufi; ero 6orosrre AoJrn noqrr,r oAunaronoft AJprHr,r, Hepa3BerBJIeHHbre. 

Hapyxurre 6aganrurre nepbn[Kr.t caMbrx HlrxHHx nepbeB 

rpoQoQopa s2-3 paza Kopoqe caMbrx Anr{HHbrx nepbrrueK srofi xe cropo- 

Hbr... ..2.B.strictum. 

7. TpoSoSop B oqepraur.ru rpeyrolrHnfi, cugnuui1 He Airr,rHnee cnoeft urnpn- 

Hbr; nepbr cauofi nlrxnefi ero naprr B or{epraHnn y3Ko[aHrlerHbre t,tJrr,r JIHneftsne, 

MeHee qeM n 1.5 pasa Kopoqe scefi nnacruurr,r rpo$oSopa, y Kpynnrrx 

oco6eft neprrcroJronacrnbre, y MeJIKHX rynoropoAr{arbre Ao noqra r{eJrbnoxpafinrrx 

.. 3. B. lanc'eolatum. 

+ TpoQo0op B oqepraHr.n olrpeyronbHoro Ao rrrnpoKoJlr,rHefinoro, cnrn\uir 

tailu Ha roporxoft HoxKe, noqru BcerAa Alll4HHee cnoefi ruupnubr; fiepr,r cauofi 

nnxsefi ero napbr B oqepraHun or unpoKoJraHrrerHblx Ao nonyJryt{- 

Hbrx, rleJrbHbre, nepucToironacTHbrer{nr,r nepucTopa3AeJrbHble ......... 8. 

8. TpoQorpopu B oqepraHur,r JraHrlerHbre unn uupoKoJrnuefiuue; Bce nepbfl Hx 

qeJIbHbIe, or [oJryJryHHbrx Ao I[lrpoKoKJrr.rHoBtrAHbIx, n repxnefi qacrr4 3aKpyrneHHbre, 

uHorAa ropoAqarbre r.trrn eABa ironacrHbre, xors 6rr Hr.rxHr,re 

urnpe cnoeft AJrr4Hbr 7. B. lunariu. 

+ Tpo0o$opbl B orlepraHr,rur or rpeyroJrbHbrx Ao rrrupoKonaHrlernbrx; nx nepbr 

y Kpy[Hbrx oco6eft nepucropa3AenbHbre r,rJrr,r neprrcrononacrHbre, y Merrnx 

oco6eft or neprrcroJronacrHbrx Ao rreJrbHbrx, Ho 3aMerHo AJrnHHee cBoefi 

unpunlr, n uepxuefi r{acrr,t or y3Ko3aKpyrleHHbrx ao ocrporarrrx . . 9. 

9. Tpot[oSop o6rrqno Ha HoxKe Ao 6 nrnr AJr., peAKo cugar-rtuir: ero nepbt 

o6rrqno B oqepraHlt.r JraHrrerHbre, paccraBJreHHbre, y 6olee KpynHrrx oco- 

6eft nepucropa3AeJlbHbre, qacro ocrpoBarbre, co cla6o ganaerublM xlrJrKo- 

BaHLIeM . . 8. B. matricarii/blium. 

+ Tpo0o0op o6r,rvno cuattui.r; ero nepbq o6rrqno B oqepraHr.ru or npoAorrroBaro-TpeyroJrbHbrx 

4o ruupoxorftlleBlrAHbrx, qacro c6luxennrre, y 6olee 

KpynHbrx oco6eft neprrcropa3AenbHbre r,rJrri nepucroJronacrHbre, qaule rynbre, 

cxopouo3aMerHbrMxlrJrKoBaHueM ........ 10. 

10. Tpo0oSopbr B oqepraHr4l4 JraHrrerHo-rpeyroJrbHbre; nepbr r,rx y KpyrrHbrx 

oco6eft TpeyronbHbre, neplrcropa3AeJrbHbre, rracro ocrpoBarbre, K ocHoBa- 

Hr,rroKJrr,rHoBlrAHocyxeHHbre..... ...... 4. B. alaskense. 

+ TpotDo$opbr B or{epranun qftqeer,rAHbre r{Ju{ npoAonroBaro-rftqeea4Hsre; 

nepbr r.rx o6rrqno sfiqesraAHrre ri rynbre, y nan6onee KpynHbrx oco6eft rraoryr 

6rrrt nephcropa3AeJrr,HbrMr,r, no o6rr.rHo flepncroJroflacrHbre, xors 6lr 

Hr,rxHue y ocHoBaH[r 3aKpyrJreHHbre r{Jrr{ KaK 6u o6py6lenHbre...... ll. 

lt

I l. fleprr o6rrqso npo,4orrroBaro-rftqesl,rAHrte, ropu3oHTaJrbHo pacnoJroxeunrre, 

c 3-5 JronacrtMl4 c xaxAoft cropoHbl, AoBoJrbHo roHKHe I,I noroMy c 

6oaee nrrcrynaoul,IMl,I xHflKaMLt 5. B. crassinert'ium. 

+ fleprr o6rr.rHo sftuesraAHrte, B cropoHy I,I HeMHoro KBepxy HanpaBJIeHHbte, 

o6rrqno c l-3 nonacruwt nfiv AorItMI,r c raxAofi cropoHbl, 6onee rolcro- 

Barbre r{ noroMy c MeHee BblcrynarcIqlrMl,r xI,rrKaMH 6. B. boreule. 

floqpoA l. Osmundopteris (Milde) Clausen, 1938, Mem. Torrey Bot. 

Club, 19, 2:93; Ilne.rr., 1991, s Cocyn. pacr. coB. .{alsH. Bocr. 5 : 2l; 

IITM4Kon, 2001, Turczaninowia 4, l-2: 40. .-- B. sect. Osntundopteris 

Milde, 1867, Filices Eur. : 209; id., 1869, Verh. Zool.-Bot. Ges. Wien l9: 

96. Botrypus Michx. 1803, Fl. Bor. Amer. 2:274, s. str. 

Osmundopteru's (Milde) Small, 1938, Ferns Southeast. Stat. :377,482. 

Typ u s: B. virginianum(L.) Swartz. 

l. B. anthemoides C. Presl, 1847, Abh. Bohm. Ges. Wiss. 5,5 :323. 

B. virginianum var. europaeum Angstr., 1854, Bot. Not. (Lund) 5 6 : 

68. B. virginianum subsp. europaeum (irngstr.) Jav,, 1924, Magyar. Fl. : 

18; Clausen, 1938, l. c. : 101; A. Eo6p., 1974, so {Dr. Eapou. qac'rl4 

CCCP | : 72. -- B. virginianum auct. non (L.) Swartz : OoMKH, 1934, ro 

@r. CCCP I : 100; Acrepon, 1977 , Bol. xypu. 62, 7 : 1332; Kpacuo6opoe, 

1988, so (Dn. Cu6. (Lycopod. --_ Hydroch.) : 52; I-{ner., 1991, qur. 

cou. :21; W. a. F. Wagner, 1993, in Fl. North Amer. 2 :91; Rothm., 

1993, in Fl. Europ., ed.2, I: l0; lUrr.rarou, 1999, flattoporH. Pocc. : 16; 

Jonsell, 2000, in Fl. Nord. I : 36. 

B Poccur,r B rrpeAeJrax recHoil 3oHbI, Ho 3axoArrr n s recocrers (Ao 

orp. Xaprxoea ta 3l,ruesa), ua Kanxa:le I.I3BecreH n.{at'ecraHe, s Cn6nptl 

roJtbKo roxHee 60o c. ur., Ha AanbHeM Bocrore ltuul 6ltus ycrur 

Arrlypa. Bue Poccr,ur B JIecHofi 3oHe EBpotIu, e Asep6aftaxane (3araranlcrufi 

p-H) u Cen. Arraepnxe. Ottucau us AncrpnLr (). 2n = 184. 

Enusrufi BuA -- B. virginianum (L.) Swartz (1802, l. c. : I I l; = Osmunda 

virginiana L. 1153, Sp, Pl. : 1064) r.lrlu noABrrA - - B. virginianum 

subsp. virginianum, orrcaH ra: CIIIA (urrar Bnplxtauur) Id pacrpocrpa- 

HeH s 6onee rerrJrbrx, lpenMyuleffneHuo cy6rpourlqecKllx crpaHax. B Ces. 

Auepuxe oH Bcrpeqaerct B Bocr. L loro-Bocr. Iurarax CIIIA, 3axoAf, B 

KaHa4y Jurub Ha rpafiueu rcfo-BocroKe (Horaa LLIor.uaHAur I{ o-Ba 

flpnuqa S4yapaa), fAe oH Bcrpeqaercf,, B orJluqne o'r B. anthemoides, 

ToJrbKo B JrlrcrBeHHbrx Jrecax (Clausen, 1938). llpunoAu'rct raKx(e AJIfl 

Bocr. u lOro-Bocr. Asu]r. A. E. Eo6pon, rlrrr. coq., ouu6oqHo IIprrBo- 

Anr 3Tor BrrA rroA aJrbrepHaruBHbrM Ha3BaHueu > 

ToJrbKo 4rtx Eapasuu, xorq oH rxnpoKo pacfipocrpaHeH [orrru uo scefi 

necHoff 3oHe CeB. Avrepuru, Bcrpeqarcb 3Aecr' 3Haq[TeJrbHo r{aule, r{eM 

B. virginianuru s. str. flocleAHnfi BlrA orfir{qaercs or B. anthemoides 6o- 

Jree roHKr{MH, 6onee paccraBneHHbrMr{ !r 6onee pacceveHHbrMr{ KoHeqHbr- 

t2

MH AoJrbKaMv rpo$orlopa. Ocu ocHoBHbrx sersefi rpoSoSopa y Het-o 

6olee yrrne u loqrr,r 6ecrpu"rmre. 

2. B. strictum Underw. 1903, Bull. Torrey Bot. Club, 30:52: [nel., 

1991, ur.rr, co.l. :21;LIIMaKon, 1999, qzr. coq. : 15. -.Botrypu,s strictus 

(Underw.) Holub. 1973, Preslia 45,3 :277. 

B Poccua Ha rore.{alruero Bocroxa, BHe ee -- s Bocr. Knrae, Ko_ 

pee u .fluonuu. - Ouucas u:.fluoHuu (). 

flo4poA 2. Botychium. - T y p u s : generis 

lectotypus. 

Cerqnr l. Lanceolata Clausen, 1938, l. c. :89; IIIM4Keg,200l, uur. 

cou.:39, s. str. -- Typu s: B. lanceolatun (S. G. Gmel.) Rupr. 

K arofi ceKqrru, cJleAyfl ee aBTopy, Mhr orHocrrM ToJrbKo B. ranceolatum 

A oqenb 6lu:rnfi K HeMy ceBepoaMepurascrnfi sug B. angustise_ 

gmentum (Rease et Moore) Fern., no oco6euuocrrM uorrKocJro)r(eHr4t 

nafin cxo4Hne c rpeAbrAyuttrM rroApoAoira. A. LI. [Urr4aros (2001) orHocur 

x neft raKxe sr,rArr ,8. boreale u B. matricariifolium. c "", irpy4r,o 

corJIacr4Tbcfl. 

3. B. lanceolatum (5. G. Gmel.) Rupr., 1845, Beitr. pflanzenk. Russ. 

Reich. 3:33, quoad nom.; Angstr., 1854, L c. : 68; (DouuH, 1934, uur. 

cou.:98; A. Eo6p., 1974,gur. cov. :70; Kpacno6opor, l9gg, qr.rr. coq.: 

50; I{nen., 1991, unr. cou. : l9; W. a. F. Wagner, 1993, l. c. :97; Rothm., 

1993, 1. c. : l0; ITIMsKss, 1999, Uu1. coq. : 15. ---, Osmunda lanceolata 

S. G. G5nel., 1768, Novi Comment. Acad. Sci. petropol. 12 : 516, tab. I l, 

hg.2. 

B Poccnl.r cfiopaAuqecKu pacrpocrpaHeH B lecHoft 3oHe or Myprr,raH_ 

cxofi u Jlennurpa4croft o6lacreft go Karra.la'rxu u caxanr,rHa BKJrroqH- 

TeJrbHo; r Apnruxy 3axoAr{'r Ha gyrorcroM n-oBe. BHe poccr,il,r __ s Ces. 

n Cpe4nefi Enpoue, flnonnn u n lecnoft gone Ces. Auepnru. - Oun_ 

caH, rro-BrlAHMoMy, uo Marepuiury Crelnepa ns Bocr. Cu6upn. Bosuox_ 

Ho, oH ecrb B fep6apr,rra Milde. floxa sa rlrrr MoxHo cqr.rrarb pHcyHoK 

r pa6ore S. G. Gmelin. - 2n = 90. VxasaHur 3roro Bulra Nrr. Kulaa lr 

Mouronnu [ptrHaAJrexar cJreAynqeMy Br.rAy. 

Cenqus 2. Botrychium. -Typ u s: generis lectotypus. 

4. B. alaskense W. Wagner et J. Grant, 2002, Amer. Fern Journ. 92, 

2 : 164, frg. l-2. -- B. boreale auct. non Milde : KpacHo6opos, l9gg, 

uur. coq.:50, p. max. p.; [drel., 1991, uur. co.r.: 19; lllrraaxos, 1999, 

rlr,rr. coq. : 15, p. p.-- B. lanceolatunt auct. non (S. G. Gmel.) Angstr. : 

fpy6., 1963, s Pacr. I{eHrp. Aruu l :72; fy6auor, 1996, Koncri. $r. 

Bneurs. MoHr. : ll. 

B Poccul. Bcrpeqaercr B ropHbrx paftonax KrxH. rroJroBnn' Bocr. 

Culupu n na.{zurrueM Bocrore (n Apmr.rre roJrbxo na rlyrorcroM r-oBe), 

nne Poccura - Ha nre Arscru u s Ces.-Bocr. KanaAe, a raKxe s MoH_ 

l3

oJlrr4 (Xaurafi r,r Xanrefi) la Ces.-Bocr. Kurae, rAe rror BrlA upllHl'IMaJIcs. 

Ja B. lanceolatum. --Ouucas us IOxH' Anscxu (

. 

B Poccuu AocroBepHo He H3BecreH, Twr H Sr3eMrurflpbr AaBHux KoJrneKTopoB 

yKa3blBaJlr.rct Nts,

B. boreale, rcarc w 2 npeAbrAyurux B[Aa, [poucxoArrr or 4peruefr nr6pr.r- 

)ln3a1lav B. lunaria x B. lanceolatum, w KapJrr4KoBbre 3K3eMlJrqpbr Bcex 

3 ru6pu4oreHHbrx BrrAoB He BcelAa JrerKo pa3rlrqnMbr, 

7. B. lunaria (L.) Swartz, 1802, l. c. : ll0; (DoMttH, 1934, qu'r. cou. : 

98; A. Bo6p., 1974, uur. cou. : 70; KpacHo6opon, 1988. uur. cou. : 50; 

I-\nen., 1991, qur. co.r. : l8; Rothm., 1993, L c. : l0; W. a. F. Wagner, 

1993, l. c. : 97 LLIMaxos, 1999, uur. coq. : 15. - Osmunda lunaria L. 

1753, Sp. Pl. : 1064.' 

B Pocculr Bcrpeqaercfl troqrlr so scefr lecnoii r4 JrecocrenHofi soHax, 

a raKxe B KoH'ruHeHranrHofi Aprrure l{ Ha o-B Kolryer, a 3a ee rlpe- 

AeJraMr4 

-- rtoqru so scefi BHerpotrurlecxoft Erpasnu u Ces. Auepuxe, 

Brrtroqar 3an. n IOxH. fpeu;rangurc, a raKxe na xpafiueu rcre IOxH. 

Auepuxu, Arcrpaluu u Hosofi 3enauAur,r. -- Onucau us Erpouu (). - 2n = 90 (96). 

Hau6olee uupoKo pac[pocrpaHenHufi BuA poAa, Ho B [pe4eJrax Jrecocrerruofi 

roulr Poccl,ru 6ucrpo BbrMr.rpaer. .fi,onolluo uuoroo6paren, 

14 B ero rrpeAeJrax Mor-yr 6nrl nn4enesu 2 Han6onee 6pocarouluecr n 

rna3a pa3HoBr,rAHocrr.r: var. rhombeum Angstr. (1854, l. c. : 70) c 6olee 

y3Kr4Mr4 (pacruupeuHbrMr4 MeHee qeM Ha 100'), a o6m.rHo r,r 6olee paccraBJreHHbrMu 

rrepbrMr{ uvar. incisum Milde (1858, l. c, : 661) c sy6.ra- 

TbrMLr rrJrrr JrorracrHbrMu rrepbtMl,t, flepraa u3 HI,rx Bcrpeqaercr floql-ra 

rroBcroAy o6uqHo BMecre c var. lunaria u 6rua orrucaHa r.rs CIIIA uoA 

Ha3BaHueM B. onondagen.se Underw. (1903. l. c. : 47), a Bropar 3Haqu- 

TeJrbHo 6olee pe4ra. Ha:arraAe Cer. Arraepuru o6H.reH etrle oArrH oqeHb 

cxoAHuil c var. rhomheum BwA -- B. mindagez.re Victorin (1927, Contr. 

Lab. Bot. Univ. Montreal, ll : 331), nuenuluft 6onee uuor-oqr{cJleHHbre 

u 6olee roHKlrre nepbf,. OH upusuaercf, 3a caMocrorreJnHrrfi nn4 u a 

Hacroruree BpeMr (uanpnuep, Wagner, Wagner, 1993 : 98) 14, Bo3Mox- 

Ho, 3axoAr,rr Ha .{anrnufi Bocror, rAe Mhr He cMouu,t orJll4qfiTb elo or 

B. lunaria var. rhombeum. B oltrlu,rue or B. lunaria, Ars Hero yKa3brBaercs.2n 

= 180, B uocle4Hee BpeMr ul Anrcxu orrncaHbr eu\e 2 suha, 

oqeHb 6ru:rue x B. lunaria: B. yaaxudateil Stensv. et Farrar (2002, 

Amer. Fern Journ. 92,2'. 157), rnonHe cxoAuuft c KpyuHblMlr oco6sN,ru 

B. lunaria, no c 2n = 180, vr B. tunux Stensv. et Farrar (2002, 1. c.:152), 

oqeHb noxoxnfi ua B. mindagense, Ho c 2n = 90, rar y B. lunaria. An- 

Topbr 3Tl4X BUAOB, CaMOCTOSTeJIbHOCTb KOTOpbTX rrpeACTaBJueTCt HaM 

coMHrdrerbHoft, ccn,lrarcTcr Ha anailv3 3H3rrMoB, roropuft rroKa3brBaer 

pa3Jrurrr,rfl MexAy ulluu. Eclu r{x [prr3HaBarb, To r{ poccuftcrue r]orryJlttluu 

B. lunaria s. L rpu4ercx pa3AeJrrdrb Ha HecKoJrbKo Mr4KpoBrrAoB, Ilrr 

r{efo, oAHaKo, ueo6xoAuMo lrpoBeAeHr4e AorroJrHIITeJrbHbIx Kapr4oJrolr4qecKux 

u 6[oxu],nqecKr.rx r4ccJreAonauuft. O parnuuurx oco6eft B. lunaria 

c 2n = 90 n 96 Hn.rero He u3BecrHo. 

Eule oAuu 6lnsrcufi ceBepoaMepnxaHcxufi BLIA -- B. crenulatum 

W. Wagner, orJruqarculuftcs or B. lunaria 6olee ronxuuu (He Mrcr4crbt- 

16

uu) rpotfor[opaMu c paBHoMepHo sy6uaruur{ n paccraBJreHHbtMr4 lrepbflMu, 

Bcrpeqaercr r.i s ftrcnuu:

Mr.{Hr,rpoBaHr,reM feHoB BTopofo lr3 3TI4x BI,IAOB, BCJIeACTBI'Ie qero OH, He 

6es ocuosaHuit, 6:';'n tIoMeuIeH uarvrn (llrenes, 2000 : 156) u rIoApoA 

Sceptridium. OqeHr Bapblrpyer B 3aBI.tcI4MocrI{ or BeJluqnunt oco6n: uau- 

6onee rcpyuHme oco6u HMercr cloxnnft rpo$o$op nl 3 noqut oAKHa- 

KOBbTX [epr4cropa3AeJlbHblx rlepbeB Ha xopouo pa3BI{TbIx qepeuoqKax, 

qro, KaK r4 OrreHb Hr,r3Koe (y nonepxuocrn novnu) flonoxeHl4e rpot[o$opa, 

qBHo cBuAereJlbcrByer o poAcrBe 3'r'oro BuAa c B. multi/idum. Hatt- 

6oree uelxue oco6n llMetoltpexnouacrHvtlt utru Aaxe ueJlbHbrfr rpo$o- 

Qop. Bcrpeuaercr o6rt.rHo Ha BJIaxHbIX uoHtr)KeHl{f,x Me)KAy 6epet'onr,r- 

MU BaJlaMI,I t4JlI4 AIOHaMH, IIO-BIrAI{MOMy, BCefAa BMeCTe C OAHI4M 143 CBOrrx 

poAureneit -- B. multifidum. Hannque y B. simplex AI'IIInoI'tAHol'o 

xpoMocoMHoro tnrcJla 2n = 90, KaK Ir y ero poAureJlbcKlrx BI4AoB, cBI{- 

AereJrbcrByer o roM, .rro cra6l4rl,Izagux rw6puAoB B ?ToM clyqae lIIJIa 

He [yreM aJUlonoJIIrIIJrovtl\t4sat\vrt, a IIyreM l.{HTporpeccl4nHoft ra6pugu- 

3a;4rav (n uegpeullx AorIoJIHI4TeJrb Ho I'o f eHHo ro Marepuafl a B. mul t iJidum 

B reHorr,In ru6puga flpu rIoBTopHbIx cKpeuruBaHl{flx. 

flospoa 3. Sceptridiwr (Lyon) Clausen, 1938, l. c.:24: IIserI., 1991, 

uur. corr.: 19; Ilft',raxos,2001, uur. coq.:40. -- Sceptridium Lyon, 1905, 

Bot. Gaz. 40:457. -- Lectotyp us: S. ohliquum (Muhl exWilld.) 

Lyon (= Botrychium obliquum Muhl. ex Willd.). 

B Poccuu npeAc'raBJleHa JII,IrIrb ceKIIl4t MultiJida Clausen (1938, l. c' : 

26) :roro lroApoAa. 2 Apyt'ue ceKurl{: Elongata Clausen c l'ulloM B. dissectum 

Spreng. u Hiemobotrychium W. Wagner c ranoM B. lunarioicles 

(Michx.) Swartz rpl4ypoqeHbt x Ces. Auepnre. 

10. B. ternstrz (Thunb.) Swartz, 1802, L c. : I I l; Zou a. W. Wagner, 

1888, Amer. Fern Journ. 78,4:134; ll]rvraron, 1998, Turczaninowia 1, 

| :23; oH xe, 1999, uur. coq. : 18. - Osmunda ternata Thunb., 1784, 

Fl. Jap.:329. Sceptridium ternatum (Thunb.) Lyon, 1905, l. c. : 458. 

B Poccun roJIbKo Ha Iore Caxa:ruua u Kypu.tlctcux o-BoB, nue Poccv\kr 

---v f.nouutr, Ha rl-oBe Kopea, n Kurae u Cee. ldtt1uu. OnacaH 

uz f,tosnu (). 

Kpoue orMeqeHHofo A. I4. LLINaaxosltN{ ?K3eMrIJIflpa froro BIIAa c 

lOxH. CaxanuHa, I{Meelct eule co6pauHnft s 1984 r. fK3eMIIJIflp c o-Ba 

KyHaurrap (> LE). 

ll. B. rohusrzrz (Rupr. ex Milde) IJnderw., 1903, L c. ; 51; OouuH, 

1934, uu'r. coq. : 99; KpacHo6opos, 1988, qnr. coq. : 51, p. p.; I{rel., 

1991, qur. co.r.:20; lllv46ss, 1999, uur. coq.:18. B. rutaefblium 

A. Braun ex Doell var. robustunr Rupr. ex Milde, 1858, L c. :763, tab. 

55, fig. 9; Rupr., 1859, l. c.:40. -- B. multifidurz subsp. robustum (Rupr. 

ex Milde) Clausen, 1937, Bull. Torrey Bot. Club 64 :72. '-- Sceptridium 

robustum (Rupr. ex Milde) Lyon, 1905, l. c. : 458. 

B Poccnu -- Ha ,{,alruen Bocroxe K IoI'y or Iroc. KopQ u Ha Ktre 

Bocr. Cu6npu. Bne Poccnu --' n Knrae, Ha rr-Be Kopea, s fluoHwu 14 Ha 

18

ore Anscrn. -- OnucaH c Kauqarru u IOxH. Anscru. L e c t o t y - 

p u s : (LE) - Tzvelev (hoc loco). 

B ra.recrse JIeKTorHrra Mbl rrpeAJralaeM rrpliHrrl rluruponaHutrii 

Bbrrue 3K3eMnJup c Kauqarru, o[peAeJrenHuii Pyupexron u nnaeroulrlfi 

2 u:olexroruna. 3auuuaer upoMexyrorrHoe rronoxeHr,re Morqy B. ternatum 

u B. multiJidum, orJrr,rqarcb or trepBofo rr3 Hux 6olee rolcr.nnan, 

HeMHofo Mrcr{crblMrl rpo$oSoparr,ru c MeHee 3aMerHbrMl,I xHJrKaMr4, a or 

BToporo - ocrpoBarbtMr{ I{ o6tr.rHo ry6.raurlru KoHeqHbrMr4 AoJuMt4 

rrepbeB n repbrrrreK. Mor 6u 6nrt npuHrr 3a rroABtrA B. multi/idunt, 

c KoropblM cBr3aH rlepexoAHblMtr oco6ruu, Ho, BeporrHo, caM rrpoucxo- 

Ar{r or rra6pngu:aquu B. ternatum x B. multiJidum. O.{eHr 6lusor u 

K ceBepoaMepr.rKaHcKoMy B. silaifulium C. Presl. LI. M. KpacHo6opon 

(unr. co.r. : 51, r

Ha3BaH[e B. multiJidum B efo coBpeMeHHoM l4ctloJrb3oBaHnn, oHo AoJIxHo 

6rrrl 3aKoHcepBl{poBaHo c HoBbIM TI{IIoM, uro rpe6yer clleuniurbuoro 

oSoprrlreHuf,. Moxuo roJrbKo noxaJrerb, uro Pyupexr BoccraHo- 

Br4n AIlfl Br,rAa 3To Ha3BaHrre, xors oH trlor 6u [pI{HflTb lurt Helo He BbI- 

3brBaroulee covrnenuft Ha3BaHue B- matricarlae (Schrank) Spreng. 

E. IlIxoAa (Skoda, 1996) c.l-rraer, qro Pyupexr ony6nuronaJl Ha3BaHI4e 

M i I d e J. Die Geftiss-Cryptogamen in Schlesien preussischen und tisterreichischen 

Antheils // Nov. Act. Acad. Caesar. Leopold-Carol. Nat. Curios. 1858. 

Yol.26, pt 2. S. 371-753. 

S t o a a B. Taxonomic comments on the , Vol. 2, with some nomenclatural combinations for Pteridophyta ll 

Preslia. 1997. Vol.68. N 4. P.341-359. 

S tens vo ld M. C., F'arr ar D. R.. Joh n so n - G ro n C. Two new 

species of moon-werts (Botrychium subg. Botrychiunt\ from Alaska // Amer. 

Fern J. 2002. Vol. 92, N 2. P. 150-160. 

Ta kabash i A., Kato M. Developmental anatomyof vascularcambium 

and periderm of Botrypus virginianus and its bearing on the systematic 

position of Ophioglossaceae ll Bot. Mag. Tokyo, 1988. Vol. l0l, N 1064. 

P. 373-385. 

Wagner W. H., Grant J. R. Botrychiumalaskense, a new moonsvort 

from the interior of Alaska // Amer. Fern J. 2002. Vol. 92. N 2. P. 164-170. 

Wagner W. H., W agn er F. S. Evidence for interspecihc hybridization 

in pteridophytes with subterraneam mycoparasitic gametophytes // Proc. 

Roy. Soc. Edinburgh. 1985. Vol. B 86. P. 273-281. 

Wagner W. H.,Wagner F.S. Fam. Ophiogktssoc:eaell l-loraof North 

America north of Mexico. New York, Oxford, 1993. Vol. 2. P. 85-106. 

EoraHrqecruit uncrury'r uu. B. JI. Kouapona PAH 

197376 C.-flerep6ypr, yn. flpo$. flonora,2 

E-mail: tzvelev@DG4972. spb.edu

H. H.llne.nen N. Tzvelev 

O POAE SELAGINELLA P. BEAUV. S. L. (SELAGINELLACEAE) 

B POCCHI{ 

DE GENERE SELAGINELLA P. BEAUV. (SELAGINELLACEAE) 

IN ROSSIA 

Selaginella P. Beauv. s. 1., uo-nu4uMoMy, sBJuercs uau6oree KpyuHblM 

poAoM orAena rrayHoBr,rAHarx (Lycopodiophyta). B Hacroqqee BpeMr oH 

HacqHTHBaer yxe 6olee 750 sr,rAos, Ho Bce HoBbre lr HoBbre Br4Abl froro 

poAa orrrlcbrBaK)rcr n3 rpotlrKoe u cy6rpouuxos o6or4x uolyruapufi. 

B orHoureuun [onoxeHr4x cervreficrsa Selaginellaceae B cvcreue rroKa Her 

eAl,rHor-o MHeHur. Tar, B. P. @rznurr (1978) orHocur ero r xlaccy Iso?'topsida 

B KaqecrBe oco6oro rroprAKa, no y B. Po:rua"nepa (Rothmaler, 

1976 : 77), c[eur{aJrbHo r4HTepecoBaBruerocfl Bblcmr,rMlr c[opoBbrMu pac- 

TeHr,rflMr,r, oH orHeceH K KJraccy Lycopodiopsida sv'ecre c rHtlHqHblMLr 

rrnayHoBbrMn. Ha rratrt B3rJrrA, rrpaBr4JrbHee orHocr4Tb Selaginelloceae x 

caMocrorrerbHoMy KJraccy -- Selaginellopsida A, B. Frank (= ordo 

Selaginellales Wettst. 1903, Handb. Syst. Bot. 2, l:97; t y p u s : Selaginella 

P. Beauv.), HapflI{y c KJlaccaMH Lycopodiopsida w Isodtopsida. 

Pop, SelaginellaP. Beauv. s. l. ou[.raercs oqeHb 3Haqr.rreJ]bHbrM noluuopSuauoM 

B orHorrreHnn AocrarorrHo cyuec'rBeHHbrx rrpr,r3HaroB, 14 

ne cny.rafino yxe co BpeMeHH ero flepBoouucaHvt oH 6rrl pa3AeJreH Ha 

HecKoJrbKo caMocrorrerbHbrx poAoB. flpu ueproouvcatttn STor poa 

BKJrIoqaJr Jrurxb oAr{H BrrA S. spinosa P. Beauv. He3aKoHHoe HoBoe 

Ha3BaHne AJrr rur4poKo pacrpocrpaHeHHoro nuga Se lagine lla se laginoide s 

(L.) Link, roropufi cJreAyer cqurarb HecoMHeHHbrM rr{rroM 3r'oro poAa. 

Buecre c 6nnsrlrlr lanaftcxnu BuAoM S. deflexa Brackenr. elo o6nqso 

orHocrr K rrrrroBoMy uoApoAy poAa - Selaginella, roropufi flli'rflercr. 

Hau6o.rree [pr4Mr,ITr4BHbrM B poAe u rraopQonor-uqecKu oqeHb cr4[bHo or- 

Jru.raercr or Bcex ocraJrbHbrx Br{AoB poAa (Snrtrou2\br v cuopo$ulnou- 

Abr y Hero Bce cuupaJrbHo pacrroJroxeHHbte n cxoAHbre Apyr c ApyfoM, 

uoHoruop$Hr,re). 

Vxe ua ocHoBaHur{ uop$oloruqecKr.rx 14 MrrKpoMopSolror-uuecxux 

rrpr,r3HaKoB, oqeB[AHo, qro oclalbHbre BuAbr poAa AoJr)KHrr 6nrt nn4erleHbr 

B oco6ufr poA, rrpr4opurerHoe Ha3BaHr.{e Koropofo Lycopodi.oi.des 

Boehmer ex Ludwig, rrpt4HuMaJtocb y)r(e prAoM aBropoB (Kuntze, 

I 89 I ; Rothmaler, 1944; Sojik, I 992). Molery.urpno-l-eHeruqecKr4e Hccne- 

AoBaHLu rrocJteAHero BpeMeHr (Korall et al. 1999; Korall, Kenrick, 2002) 

rroATBepAnJru or{eHb 3Haqr4TenbHyrc o6oco6leHHocrb Selaginella s. str. u 

TaKXe rroKa3aJlu Heo6xoAt4Mocrb BbtAeJreHufl ocraJrbHbrx Br4AoB 3Tot.o 

poAa (sensu lato> s oco6trfi po4. flpu 3roM HeKoropue ueygo6crua 

Bbl3brBruro o6vttue HoBbrx rou6nuaquft, oAHaKo 6ourruar qacrb l.rx vxe 

6una cAenasa (Kuntze, 1891). 

22

flaneo6oraunqecKr,re AaHH bre cBr,rAereJrbcrByror o He3aBr{cr4Mofi rs o - 

Jrtoulrlr or,qeJrbHbrx rpy[u Selaginella s. L yxe c HtrxHero rap6oHa. ?ro 

o6croqrerlcrBo raKxe rroAAepxr.rBaer ueo6xo4uMocrb AeneHlrfl Selaginella 

s. 1., no ueHlueft uepe, Ha 2 cauocroflTeJlbHbrx poga (Thomas, 

Quansah, l99l : 59). I,1. Corx (Soj6k, 1992), ne 6es cyuecrreHHbrx ocuoraHuft, 

[peAJroxrrJr BHAerr,rrb s oco6Hft po1 Bryodesna Soj6k eule 

oAHy rpyrry BnAoB (45 maon) Selaginella s. l. c rnuou S. rupe.stris (L) 

Spring, nMerou1yxr clrlpafibHo pacnoJroxeHHbre MoHoMop$Hue Suruorl.4br 

u r,eryccarHo pacloJroxeHHue cuopo$r4jrJror4Abr. Ocra:rlu ue n n4lr 

Selaginella s. l. ou upeAJralaer orHosr,rrb K poAy LycopodioideJ, MHor.oqncreHHbre 

Br4llbt Koroporo (ux 6onee 650) Mor-yr 6url pas4elreHbl Ha 

uoApoAbr u ceKql4u. Xors sr,rAn Bryodesma H rro MoJreKyJrrpHo-r-eHerr4qecKnM 

AaHHblM o6pa:ynr oco6yn rna4y (Korall, Kenrick, 2002 : 510), 

oHr4 Bce xe AaJreKo He raK o6oco6neurr, xax Selaginella s. str., n eule 

HecKoJrbKo KJraA Ha lpeAJralaeMofi gruMu aBTopaMLI rna4ncrnvecroft 

cxeMe Aocrtrraror trorrru rarofi )Ke cre[eHrl o6oco6neuuocrn. flolrouy 

Mbr cqr.rraeM rroKa par-IuoHarbHbrM AeJrurb LycopodioideJ ruurrb Ha [oApoAbr 

rr ceKunlr, xors B 6y4yqerra u lror poA, BeposrHo, 6yAer pa3Ae- 

JleH Ha HecKoJtbKO pOAOB. 

Enle paurrue (Jermy, 1986) poa Selaginella s. l. 6ru pasAelen Ha 5 

rroApoAoB: S e lagine I la, Er ic e t o rum Jermy, Te t r agono s t ac hy s J ermy, S t acltvgynandrun 

(P. Beauv.) Baker u Heterostachys Baker, r,t3 Ko.fopbrx 

IroApoA Selaginella, KaK Mbr yxe orMeqanu, o6oco6neH 3Har{HTenbHo 

6orrure Apyrux u cocraBJlrer pog Selaginella s. str., a u3 ocf.aJrbHbrx 

IIoApoAoB, Koropbre BxoArT y pog Lycopodioides, Tetragonostachys, n 

ocHoBHoM coorBercrByer poAy Bryodesma Sojrik, Ericetorum Bxrroqaer 

scero 3 BHAa u3 Arcrpaluu u lOxnoft ASpurra, Heterostachy.r c Ar4Mop- 

$uurvru rrerbrpexprAHbrMr4 cnoposumrougauu coAepxuT oKoJro 60 nper{MyxIecTBeHHo 

Tpo[uqecKrlrx Br'rAoB, a B [oApoA stachygynandrum sxo- 

Aqr 6onee 600 ocraltubrx BHAoB c uoHouopQHbrMr4 qerbrpexpflAHblMr. 

cnopoSnmotrAaMn. Ha Hau B3rJlxA, n Poccur.r, KaK H Ha reppr.rropr{u 

Ollsruero CCCP, rpeAcraBJreHbr BnAbr AByx rroApoA oz: S tachygynanclrum 

14 Tetragonostachys, u3 Koropbrx rropofi, HeoMorpr ua uonouop$uue 

Su.rrnon4rr, tro coBoKyrrHocrrr rrpr,r3HaKoB rrpeAcraBJlrercr 6oLree snco- 

KO C[erIUtUlu3HpoBaHHbrM. 

Poccr.rficnue BuAbr rroApo ga S ta chygynandrunt, Koropblfi , cornacHo 

HoMeHKJrarypHbrM npaBnlau (ou ruuoaofi), cJreAyer Ha3brBarb Lycopo_ 

dioides 6* urraenu aBTopa, rrpeAcraBneH n Poccuu 3 rpyuuauu BtlAoB, 

KOTOpbre H Ha KJraAucru.recrofi cxeMe IIo MoJreKynrpHo-reHerr.rqecKuM 

AaHHbrM (Korall, Kenrick, 2002 : 510) Soprvtrpyror rpr4 caMocrorrerrb- 

Hbre KJraAbr. Mu upe4nonaraeM AJrr 3Tr4x rpyrr[ paHr cerqufi. I,IcxoAr 

u3 cKa3aHHol'o, HaMr,r Hrixe rrpeAJraraercfl cncleMa poccr,rficrux Br.rAoB 

S_elqsiryelk s. 1., cyxlecrBeHHo orJrr4qaroularcr o.r cHcreMbl, upuurrofr 

C. C. XapreBuqeM (1985) AJrr AaJrbHeBocrorrHbrx BHAoB ?roro poAa 

B ero utrpoKoM rroHuMaHr4r,r. 

23

Cucreuy poga Lycopodioides MbI HaquHaeM c ceKIrI,IH Lycopodioides, 

Br,rAbr Koropoft, HecMorpr Ha cuJrbHo guuopoHue (npucnoco6JleHl4e K 

rracroMy o6I4TaHHIo rlpu HeAocrarre cnera) $ntuon4r,t, LrMeIor AoBoJIb- 

HO [pr4M]rTr,rBHbre ToHKHe H pbIXJIbIe crpo6nlu co crIHpaJIbHo lrJlt{ AByprAHo 

pacflonoxeHHbrMr,l cflopoQunlonAaul,l. BuAbl cJleAylouefr ceKrlr.ru 

.- Stachygynandrum, 6onee KcepoMopOHbI, c oqeHb rycrblMu .re't'btpexrpaHHblMr4 

crpo6I,IJIaMLt, Ho o6blr{Ho co cJlaoo gurrlop$ultun $uLutovrawn. 

Eue 6onee xceporr,ropQHbl BLIAbt ceKITHLI Rosulatae, po3erKoBKA- 

Ho co6paHubre tro6erl4 Koropblx B cyxylo rrorotly 3aBopaqtlBalo'tcfl 

BHyrpb, Qoprvrnpyr 6onee ltnn MeHee urapoo6pasuue oco6u. Cucreuy ra- 

Beplxaer Han6ollee clleq[anl,I3Hponauultfi, a, floxiulyft, u salr6onee rcepoMop$Hbrft 

rroApoA Tetragonostachys c oqeHb y3KI4Mn I,I rycro pacllo- 

JloxeHHbrMrr uonorvropt[HblMu oHJUIoI,IAaMU. 

Cev' Selaginellaceae Reichenb. 1837, Handb. Nat. Pflanzensyst. : 163 

(); 

Mett, 1856, Fil. Hort. Lips. : 16, 122, s. str. 

Pol l. Selaginella P. Beauv., 1804, Magas. Encycl. 9:478; id., 1805, 

Prodr. Aetheogr. : 101. 

T n u : S. spinosa P. Beauv. nom. illeg. = S. selaginoides (L.) Link. 

l. S. selaginoides (L.) Link, 1841, Fil. Sp. : 158; A. Eo6p., 1974,so 

tDn. Eopon. qacrr4 CCCP I : 60; Xapx., 1985, s Cocya. pacr. coB. 

.{anrH. Bocr. l: 51; lUayno, 1988, so (Dt. Cu6., Lycopod. - Hydroch.: 

40. - Lycopodium selaginoidesL.,lT53, Sp. Pl. : 1101. 

B Poccnl,r: cenep Enponeftcrofi Poccl,ttt, Ces. Karras, Cn6wps w 

[atrcuuit Bocrox. - OuncaH H3 EBpoIIbI (). 

Pot2. Lycopodioides Boehmer ex Ludwig, 1760, Defin. Gen. Pl., ed. 

2:485: Adans., 1763, Fam. Pl.2:491: Kuntze, 1891, Rev. Gen. Pl. I : 

824; Rothm., 1944, Feddes Repert. 54 : 69. 

JI e xr or r,r r: L- denticulata(L.) Kuntze. 

floapou l. Lycopodioides. - Stachygynandrum P. Beauv. ex Mirbel, 

1802, in Lam. et Mirbel, Hist. Nat. Yeg.3 :477. Selaginella subgen. 

Stachygynandrum (P. Beauv. ex Mirbel) Baker, 1883, Journ. Bot. London, 

2l :3. 

Tnu: reKTorr4rrpoAa. 

Cexuur l. Lycopodioides. - Diplostachyum P. Beauv., 1804,1. c. : 9: 

481. -- Lycopodioide.s sect. Diplostachyurz (P. Beauv.) Rothm., 1944, 

l. c. : 69. 

Tnn: JreKTorurrpoAa. 

l. L. helveticc (L.) Kuntze, 1891, L c. :824. - Lycopodium helveticunt 

L., 1753,1. c. : 1104. - Selaginella helveticq (L.) Spring, 1838, Flora 

(Regensb.) 21, | :149,214; A. Eo6p., 1974, uur. coq. : 60; Xapr., 1985, 

uur. coir. :52;Ulaynq 1988, ulrr. co.r. : 40. 

24

B Poccr,u,r: Ces. Kasxaa, rcr Bocr. Cuiupn u [atrcunit Bocror. 

Ouucas us lllsefiuapr.ua (). 

Cexqur 2. Stachygynandrum (P. Beauv. ex Mirbel) Tzvel. comb. 

nova. - - StachygynandrumP. Beauv. ex Mirbel, 1802, in Lam. et Mirbel, 

Hist. Nat. Yeg. 3: 477. 

T u u : L. llabellata (L.) Kuntze. 

2. Lycopodioides rossii (Baker) Tzvel. comb. nova. -, Selaginella mongholica 

Rupr. var. rossii Baker, 1883, Journ. Bot. London 2l : 45. 

S. rossii (Baker) Warb., 1900, Monsunia I : 101; Xapr., 1985, urar. 

coq.: 53. 

B Poccuu: ror' .(a.lruero Bocrora. - Ouncau ur Ces.-Bocr. Kuraq 

(). 

3. L. sangainolenta (L.) Kuntze, 1891, l. c.:827..,- L),copodium 

sanguinolentum L., 1753, L c. : I101. -- Selaginella sanguinolenta (L) 

Spring, 1843, Bull. Acad. Brux. 10 : 135; id., 1850, Mem. Acad. Roy. Sci. 

Belg.24:57; Xapr., 1985, uur. cov.:54; IIInyro, 1988, rrnr. co.r. :40. -- 

Stachygynandrum sanguinolentum (L.) P. Beauv. 1805, L c.: I10. 

B Poccuu: Bocr. Cu6npr, !zutt,nuft Boc'i.or. -_- Onncau c n-osa KaM- 

.rarra (

flo.qpo.q 2. Tetragonostachys (Jermy) Tzvel' comb' nova. Seluginella 

subgen. Tetragonostaclqts Jermy,1986, Fern Gaz. 13,2 : ll8. 

Bryodesma Soj6k, 1992, Preslia, 64, 2 : 154' 

T u u : L. rupestris (L.) Kuntze. 

6. Lycopodioides siberica (Milde) Tzvel. comb. nova. '- Selaginella 

rupestris f . siberica Milde, 1867, Fil. Eur. Atlant. : 262. S. siberica 

(Milde) Hieron, 1900, Hedwigia 39 : 290. Bryodesma sibericurtt (Milde) 

Soj6k, 1992, Preslia 64,2:155. Selaginella rupestris auct. non (L.) 

Spring: Xapr. 1985, ul4r. coq.:51; flIayno, 1988, qur. co'r':40. 

B Poccnn: rol Bocr. Cu6upu,,{,anrHrafi Bocrox. -- OrtucaH Kc Bocr' 

Cu6upu u c Aleyrcxnx o-BoB. Jlexrortln: 

(Reed,1965-1966). 

Euurrcuft nnA - L. rupestris (L.) Kuntze, orl4caH llo sK3eMllrtpaM 

Creuepa c KaN,rqarru lt Karlua () u3 Cen. Avrepurn. Eclu trex- 

Torr{rroM cqnrarb ra}rqarcxuft o6paeeq, ro L. siherica 6yler 6olee no- 

3AHHM cLIHoHlrMoM L. rupestris, KaK cql,ITaru MHofue aBTopI'l pocctlilcrnx 

. OAHaro JIeKrorHrtoM L. rupestris o0uquzutrno u:6pau ar- 

3eMrrJrrp Kanrvra (Reed, 1965-66: 200), n rloroMy poccuficrnft BUA cJre- 

Ayer Ha3HBa'rc L. siberica, a L. rupestris s. str. HaAo otrurarb orcyl'crByrolrIHM 

s Poccl4r4 (ou npuypo.reH K BocrotrHuu paftoHau Cen. Arraepraxu), 

K coxa.ueuulo, 3THM xe aBTopoM 3a JIeKTorlttI L. siberica llplrHtr 3K3eMrrnflp 

c Aleyrcrux o-BoB, xorq 3a Hero cJleAoBano 6rt upuHtrb lK3eMtl- 

Jup r.3 flpra6afirzuna () nrlu ng Oxornn 

(

sequences // Internat. J. Plant Sci. (Chicago). 1999. Vol. 160, N 3. p. 5g5_ 

594. 

Ko rall P., Kenreck p. phyrogenetic relationshipsin seragineilaceae 

based on rbc|- sequences // Amer. J. Bot. 2002. Vol. g9-, N 3. p. iOO_Sf z. 

K u n t ze O. Revisio Generarum plantarum. Leipzig, lg9l. pars t.itip. 

R e e d c. F. Index Selaginellarum // Mem. Soc. Bioti. (coimbra). 1965-1966. 

Vol. 18. 287 p. 

R o t h m a I e r w. Pteridophyten-studien I // Feddes Repert. 1944. Bd 54. 

s. 55-82. 

R o t h m a I e r w. Klasse Lycopsidal/ Excursionsflora. Kritischer Band. 

Berlin, 1976. S.74-78. 

S o j 6 k J. Generische Problematik der seragineilaceae ll preslia. 1992. yol. 64. 

N 2. P. l5l-158. 

Thomas 8.A., Qu-ansah N. The palaeobotanical case for dividing 

lelaginella (Selaginellaceae: pteridophyta\ lt Fe.- Gaz. 1991 . Vol. 14, pt 2. 

P.59-65. 

EoraHll.recruft nucruryr nu. B. JI. Kouapona pAH 

197376 C.-flerep6ypr, yl. [IpoS. flouosa,2 

E-mail : tzvelev@DG4972.spb.edu

f. A. Jlasrrcon G. Lazkov 

HOBbIE BTIAbI POAOB FESTUCA L. (POACEAE), 

PHLOMOIDES MOENCH (LAMIACEAE), PRIMULA L. 

(PRIMULACf,AE) I{ ACANTHOLTMON BOTSS. 

(LTMONTACEAE) r43 KbrPfbr3CTAHA 

GENERUM FESTUCA L. (POACEAE), PHLOMOTDES MOENCH 

(LAMTACEAE), PRTMULA L. (PRTMULACEAE) 

YTACANTHoLIMoN BoISS. (LIMoNIACEAE) SPECIES 

NOVAE E KYRGYZSTANIA 

flpra onpe4eJreHr4u MarepuuuroB, co6paHuux Bo BpeMrr 3KcrreAnur4r4 Ha 

reppr,rropr,rr4 Kupln:crana s 2003 r., 6ulu o6Hapyxeuu HoBbte AJrfl HayKH 

BrrAbr r43 pa3Hbrx celleilcre. B crarre Aalorc{ r,rx orrrlcaH}u. Tunonue 

o6pasqbr HoBbrx Br.rAoB rrepeAaHbr n fep6apufi Eorauu.recxofo trHcrr,rryra 

uu. B. JI. Korraapona PAH (LE), ay6nr.rKarbr HaxoArrcr r fep- 

6apwu Enonoro-uoqBeHHoro r,rHcrr4ryra HAH Krrprtr:cxofi Pecuy6nuxu 

(FRU). Heroropue r.r3orr.rrrbr rrepeAaHbr n fep6aprafi Mocxoecroro 

rocyAapcrBeHHofo yHuBepcr.{rera uu. M. B. Jlouosocona (MW). 

Festuca tzveleviana Lazkov sp. nova (subgen. Leucopoa (Griseb.) 

Tzvel.). - Planta perennis rhizomatosa, laxe caespitans. Caules 40 60 cm 

alt., basi nodis crassioribus residuis vaginarum fuscis hbrosis tecti, glabri. 

Folia I 2.5 mm lt., griseo-viridia, plana, margine scabriuscula, versus 

apicem gradatim angustata, convoluta, glabra; ligula 2.5 3 mm lg., apice 

irregulariter lacerata, extus breviter pilosa; auriculae parvae, rotundatae. 

Panicula congesta, ramis scabris 0.3-l cm lg. Spiculae 1.5-2 cm lg., in 

numero 3-5, 4-6-florae, rachillis pubescentibus, pallide virides. Gluma 

inferior 4 mm Ig., lanceolata, uninervis, superior 6 mm lg., oblongoovata, 

trinervis. Lemma 8 9 mm lg., late lanceolatum, indistincte 

ciuinquenerve, basi margine pilis paucis; palea 8 mm lg., apice bifida, 

oblongoJanceolata, margine et in carinis longe ciliat4, apice breviter 

pubescens. Ovarium pubescens. Antherae ca. 5 mm Ig., oblongae, violaceo-flavae. 

(Fig. l). 

T y p u s : Kyrgyzstania,

,'u 

14 

\ , 4 

., ,. 1i 

.i' 

29

npeHHbte, floKpblrble oypbIMI{ BoJroKHr,rcTbrMH ocTaTKaMr{ Jtr{cToBbtx 

BJrafaJrHUL loJlue. JlucroBue [[acrr.IHKtr I 2.5 rraru trrnp., cepoBaro-3e- 

JreHbre, [JrocKr4e, r]o Kparo eABa rlepoxoBaTble, K BepxyuKe [ocTelreHHo 

cyxr,IBalounecr, cBepHyrbre, roJrbre; r3brqoK 2.5 3 narra AJr., Ha Bepxyu- 

Ke HerrpaBuirbHo pa3opBauuuft, cHapyxn KoporKo go;rou{crtrft; ytuKn 

He6onruue, oKpyrJrbre. Melenra cxaras, c BeroqKaMu 0.3 I cM AJr., 

uepoxoBarbruu. Konocru 1.5 2 cM AJr., B qr{cre 3- 5, 4 6-use'rKoBbre, c 

ouyureuHofi ocrrc, 6le4uo-3eJreHbre. Hnxurq KoJrocKoBar ueuryr 4 una 

AJr., JraHuerHar, c o4Hofi xunroft, BepxHr.fl KoJtocKoBafl qeruyfl 6 ruu A.lr,, 

IrpoAoJll'oBaro-gftrlesfiAHax, c 3 xr,rJrKaMI4. Hnxusr rlBerKoBar .reruyr 8 

9 uu gl., urllpoKoJraHrrgrHar, c 5 cna6o 3aMerHbrMr,r xorJrKaMr4, B ocHo- 

BAHL4V rIO KpaIO C HeMHOfOrrUCJreHHblMtr BOIOCKaMU; BepXHflf, qBeTKOBat 

qeruyfl 8 uu gn,, Ha BepxytxKe .q,ay:y6qaras, flpoAonroBaro-JtaHrlerHar, 

rro Kparo u KrlJlflM Airr{HHo pecHrlTqarar, n aepxHeft qacrl,r Koporro o[yrrreHHafl. 

3asq:r o[yrueHHar. flrlnlHunlr oKoJro 5 nau 41., upoAoJrroBarne, 

$uoreroBo-xeJrrbre. (Prc. l). 

T u tt : Kuplr,r:craH,

Pr.rc. 2. Tranoeoit o6paeeu Phlomoides kurpsaica Lazkov 

W 

. 

ll

T y p u s : Kyrgyzstania, 

(LE, isotypi - FRU, MW). 

Affinitas. A P. urodonta (M. Pop.) Adyl., Kamelin et Machmedov 

calyce pilis stellatis differt. 

MnoronerHee rpaBrHucroe pacreHne l'.l.5 rr,r nuc. Kopuenuule.rojrcroe; 

KopHeBaq uefira c MHoroqucJreHHblMlr rxeJlKoBr{cro-uepc.rHcrblMr4 

rroqKaMri. Cre6lu fiprMbre, oKpymo-qerbrpexrpaHHbre, roJ]bre, n nepxueii 

qacrn BerBf,ulrdecq. Ilucrss. [oKpbrrbr peAKr4Mr,t xeJre3ucrbrMr.r BoJrocKa- 

MH; Hr4xHue Jrncrbr 10,12 cu AJr.,8-10 cu urnp., rfiqern4Hlre, Ha tsepxyrxKe 

rynoBaTbre, B ocHoBaHHu cepArleBr4AHbre, IIo Kparo Tyrro_lopoAuaro-sy6varbre, 

qepeuKoBbre c qepeuxona l5 20 cM An,; BepxHue Jrncrbf, 

2 5 cu g1.,0.5 3,5 u"r urup., rfiqear4AHbte uJrrz trrr4poKoJraHrlerHbre, c 

KOpO'IKLIM qepeuIKOM I,IJU,I ltOr{TI4 Ct4l.flqhe, Ha BepxyruKe TynoBaTble hJrtr 

cJrerKa 3aocrpeHHbre, HepaBHo3y6varue. Couserne nernquleecs, c 4.8 

paccraBJreHHhrMlr My'l'oBKaMr.r rIBerKoB; MyroBKH 4, I 0-qnerronue. flpu_ 

UBETHI4KH 0,7-I CU AJI., U]I4JIOBI.IAHbIC, [OKPbITbI 3BE3AqATbIMII BOIIOCKAvru. 

9aureqxa rpy6uaro-KonoKoJrbqaras,, 14-15 MM AJr., 5-6 uu ulrp., 

onyrueHHag 3Be3ArtarbtMr4 BoJrocKaMu; sy6qrr c rJrtrpoKr{M ocHoBaHr4eM, 

r43 Koroporo Br,tcryrlaer urr,rJroBnAHoe ocrpne 3--5 mrra AJL, fipu ocHoBa_ 

uuu sy6qon uMerolcr ue6onrurue yurxu. Beu.rux ? 6enosarHfi,25 27 vrvr 

AJr., CHapyxu rroKpbrT AJTHHHbTMI{ Lt KOpOTKr4Mr,l BOJTOCKaMT{; rpy6r

3 3ara: Nl 915 

PHc. 3. THnosofi o6parcu Plrlomoides 

J-')

pilis rudibus articulatis tectae. Calyx tubulosus, l0 -12 mm lg., 3 4 mm 

It., basi et in parte media glabrus, superne pilis eglandulosis longis et 

glandulosis brevibus tectus, dentibus basi latis, in acuminem subulatum 

2 3 mm lg. pilis raris rudibus articulatis tectum productis, basi auriculis 

minutis praeditis. Corolla ? rosea, l6 17 mm lg., extus pilis stellatis 

obsita, tubo l0 ll mm lg., intus basi anulo pilorum instructo, labio 

superiore 7 mm lg., vix falcato, margine dentato dense barbato-pilosoque, 

labio inferiore 6 mm lg., trilobato, lobo centrale rotundato inaequidentato, 

lobis lateralibus late ovatis. Stamina filamentis superne pilis 

eglandulosis longis, inferne piiis glandulosis brevibus tectis. Pistillum 

stigmate bipartito partibus inaequilongis. Ovarium apice tuberculatum, 

glabrum. (Fig.3). 

T y p u s : Kyrghyzstan, (LE, isotypus -- FRU). 

Affinitas. A P. alaica Adyl., Kamelin et Machmedov calyce subglabro, 

verticillis multifloris et foliis latioribus differt. 

In honorem investigatoris generis Phlomoides cl. T. Adylovii species 

nominatur. 

Muololreruee rpaBflHl,Icroe pacrenue 0.7 I rvr nuc. KopHeaar urefina 

c MHoroqr{cJreHHbrMlr rtoqKaMu. Cre6ru lpocrble, flpflMble, oKpyrJloqerbrpexfpaHHbre, 

roJrbre. Jlncrlg [oKpbrrbl cBepxy orro[bIpeHHbIMH, 

qJreHucTbrMr,r BoJrocKaMn, cHn3y cepble oT 3Be3AqaTbIx BoJlocKoB; HI{x- 

Hue 10-15 cM AJt.,6-ll crr.r urnp., riiqenuAHble, Ha BepxyutKe'ryIIoBa- 

Tbre, B ocHoBaHr.rl,t rny6oro cepArIeBHAHbre, [o Kpalo oKpyrno-sy6varbre, 

qepeurKoBbre c qepeuKoM 10-25 cu An.; BepxHlre 2-5 cu An., 0.5' 

5 cu urup., rfiqeslaAHrte nilrl [rI4poKoJIaHuerHbIe, c KoporKI,IM qepeu- 

KOM I4JILI uOrITI4 Cr44ffl;d.e, Ha BepXyIlIKe TyIIOBaTbIe uJlll CJlefKa 3aOCTpeHHble, 

rro Kpalo otcpyrlto-sy6rrarble. Couserue c 3 paccranJleHHblMlrt 

MyroBKaMu rrBerKoB. Myronru MHoroqBerKoaue (6orrure 50). flpnrrBerHr{Kr,r 

oroJro I cu 41., lnHefiHo-laHqerHble, Ha BepxyrrrKe 3aocrpeHHile, 

roJrue, rro Kparo lroKpbrrbr rpy6uuu qreHl/rcrbrMa BoJIocKaMn. 

9aure.rxa rpy6uarar, 10-12 rr,rvr Nr.,3 4 MM ulup., n nuxHefi H cpeAueft 

qacru roJraf,, r nepxHeft [oKpbrra Ar[HHbIMl4 Hexele3ucrblMll H 

KoporKuMu x(eJre3r,rcrbrMu BonocKaMn, :ly6qrr c uupoKuM ocHoBaHHeM, 

r,r3 Koroporo Bbrcryflaer uLtJroBI4AHoe ocrpr4e 2-3 vrrra AJr., rloKpbrroe 

peAKr,rMH rpy6uuu qreHhcrblMu BoJlocKaMI4, tlprl ocHoBauulr ay6uon 

HMerorcr ne6onrrulre yurKu. Ben.rux ? posonuft, 16 17 rvru AJI., cHapyxn 

rroKpbrr 3Be3ArrarbrMr.r BoJrocKaMra; rpy6xa l0-ll rrarra AJr., rr3Hyrpr,r 

B OCHOBaHLII{ C KOJTbTIOM BOJTOCKOB; BepXHtt ty6a 7 MM AJt., Cnel-Ka 

cep[oBr{AHas, [o Kparo ay6.rarar c lycrofi 6opoAxofi BoJrocKoB: Hl,rx- 

Hrr ry6a 6 uu Al., Tpexnoflacrnar,, c orpyuoft HepaBHo sy6.raroii 

cpe4Hefi n urupororfiqeBnAHbrMn 6oronnuu AoirsMr,r. Hrlru rrr.rnuox 

orryureHbr a aepxuefi qacrr.r AruHHbrMH rrpocrbrMr{, B Hr{xHefi roporrn- 

34

Mr4 xeJre3r{crbrMr,r BoJrocrarun, 6es rp}rAarKoB. flecrnx c ABypa3AeJrb- 

HbrM, HepaBHoJrofracrHbrM pbrJrbueM. 3asrsl Ha BepxylrrKe 6ylop.raral, 

r-oJrafl. (Puc. 3). 

T u u : Knprr'rrcran,

tr4 

iJ 

,r 5,a* 

{ 

JI

T u n : Kuprrr:craH,

Jrr4 c peAKuMtt BoJlocKauu; orlu6 2.5 3 uut AJr., HetBHo KoporKoJlotlacrxnil, 

uypnypHuil. Jlenecrru po3oBbre. (Pnc. 5). 

T u n : Kupru:cras,

T. B. Eropona T. Egorova 

POA SCIRPUS L. (CYPERACEAE) BO OJIOPE EBPA3I4I4 

GENUS SCIRPUS L. (CYPERACEAE) IN FLORA EURASIAE 

B uacroqqeil crarle orpaxeHbr pe3yJrbrarbr Kpr{Tr{qecKoro H3yqeHr4q 

cucreMaruKl,r poAa Scirpus L. na repprETopr{r4 Eopasuu. Zccle4onaHre 

6ruro npe4upr4HtrTo lJraBHbrM o6parorra B cBf,3u c noAt-oloBKofr o6pa6or- 

Kr.r ceM. Cyperaceae AJrf, MHororouuofi $lopucruvecrofi cnogxu , Ho B xoAe pa6oru oKa3aJrocb ueo6xoAraunrM.rlJlfl rroHr,rMa- 

Hr,rr olAeJrbHbrx Br,rAoB Scirpus I,r BbrflBneHrrrr poAcrBeHHbtx cnrseft B poAe 

[3yrrl'ITb raKxe el'o llpeAcralzreleft, Bcrpeqanquxcfl n 3a IIpe/leJIaMH 

Poccun. Kporue roro, 3Tor poA He paccMarpr.rB?urcfl upexAe B [peAeflax 

ucefi Eapasur,r. Mexay reM AaHHar reppr.rropr,rf,, TaK xe, rar

Har Auepuxa, rro Koropofr nnaeercr o6pa6orra Scirpus r He4anuo ouy6- 

Ju4KoBaHHoM 23 rorvre (whittemore, 

Schuyler, 2002), ra.rrrcrcf, ocHoBHofi o6lacrurc pacupocrpaHe- 

Hr{q 3TO[O pO4a. B CBq3r,r c 3TLIM H3yqeHlre eBpa3rrarcKrrx BI4AOB SCirpuS 

I,IMeeT BaXHOe 3HaqeHl,re AJr{ o[peAeneHl,Ir BT4AOBO|O COCTaBa f,.rofo poAa 

B ueJroM r,r ero HaABr,r4orofi crpyKTypbr. 

Por, Scirpus [pr4HflT 3Aecb B y3KoM, 6onee ecrecrBeHHoM o6terue, 6e: 

BKJrK)rreHufl B HeI'o poAoB schoenoplecras (Reichenb.) palla u Isolepis 

R. Br., Koropbre paccMarpuBaJrr.rob B ero cocraBe p. [0. poxeB]rueM Bo 

3apoAbrrx Fimbristylis-rr{fla, BuAaM schaenoplectus (s. lacustrLs, s. mucronatus 

A gp.) u Bolboschoenus -* Schoenoplectus-runa, BuAaM Isolepis 

Cyperus-ruua, Trichophorum u Blysmas -- Carex-rl,rra. CreAyer gaMe_ 

Tl'rrb' qro rr,rrr 3apoAhrua, yKa3aHHHfi SHr.[e rar schoenoplectus-rnu, 

6un nepnonaqaJrbHo o6osHa.reH xar Scirpus-run (Schneid"i, 1932; vun 

der Veken, 1965) ncneacrBue rxnpoxoft s ro BpeMfl rpar(roBKa poAa 

scirpus. Ho nocxo*Ky 3apoAb'u scirpus-rnna 6ufi slisn.rreH y Br{AoB 

Scirpus s. 1., orHocr,rMbrx rerrepb K poAy Schoenoplet:tilJ, lror r.rr, ,upo_ 

Abrilra rroryqun Ha3BaHr4e Schoenoplectus-rut (Lye, l97l; Wilson, lggl; 

Goetghebeur, 1998), BaH Aep BexeHou cpeAu raKcoHoMrzqecxr4x rpytur, 

o'rHocrrMhrx K poAy Scirpus s. 1., 6ruo ycraHoBJreHo 6 rnrron sapoguuefr. 

flo pasgeneMoMy rlu[epoJroraMrl MHeHr,rn 3roro aBropa (Van der Veken, 

1965 : 285), ecrecrBeHHbrfi po4 ulreer roJlbKo oAr,rH rrrfl 3apoAbrrua, Ho 

BMeCTe C TeM AaHHbrfi run Moxer 6nrr csoftcrsen HecKoJrbKI,rM poAaM, 

o6uapyxnnar B HeKoropbrx r.r3 Hr.rx xapaKTepHbre oco6eHnocru. 

Taxuu o6pasou, :u6puonoruqecK'e pa3rvqur, MexAy Scirpus s, str. 

u lpexAe o6neAunqelrbrMn c HtrM poAaMr,r, Hapr.ly c vrop$onortrqecKu_ 

Mr4, rrporrrBopeqar KoHqeflullr trrr{poKoro o6reua poga Scirpus, Vsxar 

KoHuerrqus poga scirpus [oAAepxuBaercr raKxe AaHHbrMr4 suflOreue- 

THqecKoIo aHaJr[3a pe3yJrbraroB ceKBeHnpoBanur lHK y raKcoHoB 

Scirpus s. l. (Muasya et al., 1998, 2000, 2001). 

- K HacrosueMy BpeMeHr.r Br.tAbr, BKJrroqaeMbre flpexAe t pog Scirpus 

s. 1., tpu.rucJreHbt K 6onee qeu 50-rz po4arr.r (Goetghebeur,-199g), pac_ 

rrpocrpaHeHHbrM B pa3Hbrx perlloHax 3enaHoro uapa 14 0.rHocrulr,rMc, K 

pa3HbrM rpra6au. BcneAcrsue SToro qficJro BurAoB, paHee yKa3HBaeMOe 

AJrf, poAa Scirpus - orono 200 (Koyama, l95g) pesxo coKparunocb, l,r 

relepb, corJracHo AaHHbrM oAHnx aBropoB (Wilson, l9gl; Goetghebeur, 

1998), a HeM Hacqr{rhrBaercr oKoJro Z0 nugoa, a Apyrux (Whiitemore, 

Schuyler, 2002) -- orono 35. 

B uocnegHue 20 fler pog Scirpus rroHr,rMaerc, B y3KoM, upuHqroM B 

aausofi crarbe o6reMe, .qoBoJrbHo MHorr{Mt4 aBTopaMr.l (Kit Tan, l9g5; 

Eap6apuv, 1987; Zhan, Yang, 1987; Goetghebiur, l99g; Simpson, 

Koyama, 1998; GudZinskas, 1999; I{nener, 2000; XaHAxrH, 2001; 

Whittemore, Schuyler, 2002; pignotti, 2003 H Ap.). Br*oqael,rHe HepeA_ 

xo s Scirpus pogu Schoenoplectus, Isolepis u Actinoscirpur paccMarpuBa_ 

Iorcr KaK caMocrorrerrrHHe KyrroHeHoM (Kukkonen, 199g, 2001). 3arrae- 

Tr,rM, rrro Ha reppr,IToplrrx, oxBarbrBaeMbrx ero pa6oranau, poA Scirpus 

S. StT' HE BCTPCqACTC'. BUCCTC C TCM P'A ABTOPOB I]POAOJIXACT TIPI{HH- 

l.r.arn Scirpus B TpaAaqr{oHHou, 6olee uJrr4 MeHee r[[poKoM o6lerure 1Ko_ 

xeBHnKoB, 1988,2001; Karthikeyan et al., l9g9; TzuoxuHa, 1990; fy6a_ 

uor, 1996; Oh, Ham, 1998; Gavrilova, Sulcs, 1999; Mosyakin, Fedoron_ 

chuk, 1999; 3epuon, 2002; Khoi,2002; Wadood Khan, iOOZ u' ap.y. 

flpeqlaraeu'fi suxe o6:op eBpa3uarcKrrx Br{AoB scirpus oc"ouu" 

fJraBHbrM O6parou na rep6apnoM Marept{aJre, B TOM qI{CJre u rr,rrroB'M. 

4l

xpaHflxluMcx n fep6apuu EoraHr.r.{ecKoro r.rHcrr4Tyra }rM. B. JI. Korr.rapoaa 

PAH (LE), a raKxe o fep6apnax Myrea EcrecrseHnoft uc'r'opnu 

s r. Crorrorulae (S), EoraHu.{ecroro My3er (Dr,rHcxoro uysel Ec'recrneu- 

Hofi ucropuu Vunnepcurera r', Xe.nrcuuxr,l (H), Oraela raKcoHoMl{rrr 

pacleunii Euonornqecroro $arynrrera VHnsepcurera uu. A. Mnuxe- 

Br4qa B r. flosnasu (POZ) u.{euapraueura EoraHuxu Beurepcroro My- 

3er ecrecrBeuuoii ncropnu B r. Ey4aueure (BP). Eunu r43yqeHbr rnrroaue 

o6pasqbr HeKoropbrx BrrAoB Scirpus, pa3MeqeHHbre Ha se6-cafirax 

Hlro-fr opxcroro EoranrqecKoro caaa (NY, http://www.nybg.org/bsci/ 

vasc/Cyperaceae.html) u EoraHn.recKor-o My3er VHunepcurera s l'. Ocno 

(O, http://www.nhm.uio.no/botanisk/bot-mus/index.html), a rar

3 cexquu rBJlgarcf, MoHorr,rrrHbltuu; 2 cexquu olulcbrBaK)Tct BllepBbre. 

flpu rcex raKcoHax nplrBoAflTcr HoMeHKJrarypHble uHTarbI tr yKa3btBaercr 

rnn, [pr.r BnAax - AaHHbIe o6 grororuu, r'eorpa$uuecKoM pacllpocrpaHeH[Il 

rI xpoMocoMHbrx qucJlax. 

feorpaQr.ruecxoe pacfipocrpaHeHr4e BHAoB rrpnBoAr{rcr no KpynH},rM 

peruoHaM Enpasul,r, B [peAeJrax Koropbrx, KaK npaBuJlo, yKa3blBalol'cq 

rocyAapcrBa. PacnpocrpaHeHne n Poccnn Anf, flonyqeHl{fl qeJlocrHoro 

[peAcraBJreHnr yKa3brBaercr cpa3y no sceft ee reppl4Topnn --- Eepoueftcxofi 

n Asuarcroft uacreir., cornacHo pafiounpoaaHl4lo, nprlHflToMy B 

Mor,rx rpeAHAyulux pa6otax (Eropona, 1999, 2001, 2002). BoraHuKoreorpa$u.recrue 

paftouu Mourolu[ AaHbr no pabore B. I,1. fpy6ona 

(1982). flpunoguulre B crarbe AaHHbre no reorpaSu?r Br{AoB 6a:upyrcrcr 

Ha rl3yqeHHou rep6apHol,r Marepl,taJle, a raKxe Ha cBe.qeHuqx, coAepxarqtaxcr 

o pa6orax, urrrr4pyeMbrx B HoMeHKJrarypHbrx a6sauax. flpn yxa- 

3aHur4 xpoMocoMHbrx quceJr yra3brBalorca pa6oru, B Koropbrx uMeK)T- 

Ct CCI'IJIKI'I Ha OpIIIIIHaJIbHbIe I4CTOTIHUKI. 

Scirpus L., 1753, Sp. Pl. : 47; id., Gen. Pl., ed. 5'. 26, nom. cons.; 

Kunth, 1837, Enum. Pl.2 : 157, p. p.; Steud., 1855, Syn. Pl. Glum. 2 : 

83, p.p.; Benth. et Hook. f., 1883, Gen. Pl. 3:1049, p. p., quoad sect. 

Euscirpus; Poxen., 1935, eo On. CCCP 3:44, p. p., quoad sect. 

Taphrogeton; fpoccr., 1940, @n. Kanx., uzg.2,2: 13;T. Koyama, 1958, 

Journ. Fac. Sci. Univ. Tokyo, sect.3 (Bot.) 7, 6:282, p. p., quoad sect. 

Trichophorurfl et sect. Scirpus; id., 1978, in Fl. Taiwan 5; 203; Ohwi, 

1944, Mem. Coll. Sci. Kyoto Univ. ser. B, 18, l: 95, p.p., quoad subgen. 

Taphrogeton; id., 1965, Fl. Jap. : 200, p. p.;Ts. Tang et F. T. Wang, 1961, 

in Fl. Reip. Pop. Sin. ll : 2, p. p., quoad subgen. Scirpus (excl. sect. 

Bolboschoenas et sect. Actinoscirpu.r); Eropona, 1967, s Pacr. I{eHrp. 

Azuu3: 15, p. p., quoad subgen. Scirpus; oHa xe, 1976a, so @r. Enpon. 

qacrn CCCP 2: 89, p. p., quoad subgen. Scirpus; oHa xe, 19766, s OnpeA. 

pacr. Cpegu. AsuH 5 : 8, p. p.; Kern, 1974, in Steenis, Fl. Males. 

ser. l, 7, 3'.494, p. p., quoad sect. Scirpus et sect. Trichophorum; Y. L. 

Chang et Y. L. Yang, 1976, Fl. Pl. Herb, Chin. Bor,-Or. 1l :2,p.p.; 

quoad subgen. Euscirpus (excl. sect, Bolboschoenas et sect. Dichostylis); 

Kitag., 1979, Neo-Lineam. Fl. Manch. : 153, p. p.; DeFilipps, 1980, in 

Fl, Europ. 5'.277, p. p., quoad sect. Scirpus; fpy6on, 1982, Onpe4. cocyA. 

pacr. MoHr.:51, p. p.; M. Zhanet Y. C. Yang, 1987, Acta Biol. 

Plateau Sin.7: 1l; A. E. KoxesHr,rxos, 1988, n CocyA. pacr. coB. [anlu. 

Bocr. 3: 178, p. p., quoad subgen. Scirpus; Karthikeyan et al., 1989, Fl. 

Ind. Enum. Monocot. :70,p. p.; Tr,rrraoxnua, 1990, so (Dr. Cu6.3; 18, 

p. p.; fy6aHos, 1996, Koucn.

3 : 51, p. p., quoad sect, Sclrpas, Trichophorum et Baeothryon (5. petelotii); 

Whittemore a. Schuyler, 2002, in Fl. North Amer. 23 : 8; Pignotti, 

2003, Webbia, 58, 2:301. -- TrichophorumPers.,1805, Syn. Pl. 1 : 69, 

p. p., excl. T. alpino. - SeidliqOpiz,lS26, Naturalientausch 1l :349. - 

Scirpus sect. Taphrogeton Reichenb., 1830, Fl. Germ. Exc. :79. 

Nemocharis Beurl., 1853, Bot. Not. : 53. Taphrogeton (Reichenb.) 

Montandon, 1856, Fl. Jura Sept. : 346. - Scirpus subgen. Euscirpus Pax, 

1888, in Engl. u. Prantl, Nat. Pflanzenfam. 2,2 : ll2, p. p. ,S. b. 

Cvmelli Meinsh., 1901, Tp. flerep6. 6or. caAa 18,3:249, p. p. ,-S. sect. 

Seidlia (Opiz) C. B. Clarke, 1908, Kew Bull. Add. ser.8: ll3. - O6uee 

couBerr{e o6pa:oraHo o6n.rHo KpyrrHbrM TepMr{HarbHbrM rrln (y nu4on 

HeKoropbrx cexqnfi) Taxxe n (l) 3 4 JrarepaJrbHbrMr,r, 3oHTrlKoBHAHbrMl,r, 

cI4JlbHO BeTBr,rCTblMI{ COUBeTI{'Mr4 -- CJIOXHbTMT4 aHTeJtOAr,rfMlr, cocToqrrIrrMH 

r13 rraprlrraJlbHblx aHTeJroAr{eB; JlaTepalrrHbre corlBeTlrr pacfloiroxeHbr 

Ha AruHHbrx uBeroHocax B rla3yxax cre6nentrx JrurcrbeB. flpra oc- 

HoBaHuu repMr{HarbHolo corrBerr.tr r,rMerorcs (l) 3 4 nuHefrslrx Kporor.qrrx 

Jrrrcra, Huxnnft r43 Kolopbrx qacro npeBbruaer ero. Konocru pacrloJloxeHbr 

Ha KoHrIax Beror{eK corlBernr to | 2 u.rrn co6paulr uo 3 .20 

(25-30) s 6.N{. rrJrorHbre loJloBxr,r. Korocttr,r o6rlqHo MHorotrr4cJreHHbre 

(50 500), AoBorrbHO MerrKrre,2.5-6 (8 l0) MM Arr., 1.3 2.5 (3) vru rurzp., 

6. .r. N4uorouBerKoBbre, siillesr,rAHtre, urnpororftlleBr,rAHbre, f,firturrrt,tqec- 

Kr,re, rrpoAoJrroearo-gfrqeBr4AHble [JrLr JraHr{eroBnAHbre, Kpotoutue qeruyn 

pacnoJrolKeHbl IIo c[Iupanl4, qepHoBaTble, qepHoBaTo-3eleHoBaTbre, l.eM- 

Ho-cepbre rdiu{ KopurrHeBble, trraAKue, or rfrqesnAHnx u uruporoailqe- 

BI{AHbrX AO JraHqeTOBtrAHbrX, Ha BepxyruKe Tyfrbre, SaKpyrJreHHbte l4Jllr 

ocrpbre, c KoporKr4M ocrptreM, oqeHb peAKo KoporKo ocrucrne, (1.2) 

1.5 3 (5) MM AJr.,0.6-1.5 MM rrrr4p. ldnerxn o6oeuolue; oKonorIBerHLrK 

us 6, pexe (0)-4 ruerr,ruoK; ulerr,rHKtr nprMbre vrv v3BuilLrcrue, 6e;mre, 

6e:rosarHe taJruI cBerJro-KopnqHeBble, rJraAKr,re nJrr4 [repoxoBarbte or o6paqeHHbrx 

BBepx r.rJrr4 nnn: :y6qon, y HeKoropbtx Br4AoB (ceruur Pseudotrichophorum) 

eucrynanul[e u3 Kporclqux ,reruyfi r4 rrpuAarcrqr4e KoJroc- 

KaM 14 couBerrrro B qeJroM o6.nux. Tn.ruHox I 3. Crotr- 

6ux c (2) 3 ptrllqertrMn BerBrMr{. flloAn rpexrpaHHbre, pexe c.la6o 

ABoflKoBbrrryKJrbre r4Jtrr trJtocKo-BbrrryKJrbre, (0.6) 0.9-1.3 ( 1.8) Nlrvr Atr., 

0.4--0.6 (-0.9) rrana urup., o6uuHo 6erosarrre, Ha BepxylxKe c KoporKr,rM 

(0.I -0.3 uu)

auoq lIt 

orrpEqEJrEHn.fl BalIoB sclRpus oJlopbr EBpA3a14 

l. O6uree couBerr.re o6pasorano repMr4HaJrbHbru (nepxyurevnurra) coqnerneru n 

l-4 paccranlenHbrMr,r JrarepanbHbrMra (6oxonrrrru) couneruaun, pacnoJroxeHHbrMr,r 

B na3yxax cre6nesHxJrncrbeB ..... 2. 

+ O6rqeecorlBerr4erepMnHanbHoe..... ....... 9. 

2. Kporoqr,rq qeuryn qepHoBarbre r.lJrr{ TeMHo-cepbre; Kporoulne Jrr4crbr nptr oc- 

HoBaHnu r4rn noqrr.r no ecefi Annne qepHbre .... .... 16. S. ntaxinnv,it.zii. 

+ Kpotoulue rreuryu KopnqHegbre, KpacHoBaro- r.rJrn cepoBaro-KopnqHeBbre; 

Kporouue ruc'tbfl npn ocHoBanr.trr 3eJreHble nln 6neAno 6yponaro-xelro- 

Barbre. .. . .. 3. 

3. Korocxr.r pacnoJroxeHbr Ha KoHqax BeroqeK coqBerr,rr no oAHoMy, oqeHb 

peAKo no l-2 (3) .... 4. 

f Korocxr,r co6paurr Ha KoHrrax BeroqeK corrBernr no (2) 5-25 B pbrxJroBarbre 

r,rJu{ orleHb [JroTHbre noJryuapoBr,rAnbte tulu uapoBraAnble foJroBKl,r . . . . 7. 

4. O6uree corrBerl4e cocrouT r,r3 oqeHb MHoroqncJleHHbrx KoJrocKos. Ororo- 

IIBerHbIe xlerr4HKt4 B qnclte 6, cnemo-xopr.rrrHeBble, fi3Br,rJlucrbte, He Bbrcrynarcque 

unu 6. u. BbrcrynaKrulne lr3 Kporotrlnx velryft. Pacreune Jlnosnn 

4. S. u,iclturae. 

+ O6rqee coqBerr4e cocrour r.rg l5-50 KoJrocKoB. OxoroqserHHe uerr4HKr.r B 

quc;rc 2 nln 6, 6elue, nprMbre, Kopoqe Kpolouux .reuryfi, rlJrn oHu orcyrcrByror 

.....5. 

5. Coqserue 3aHnMaer noqru Becb penpo4yKTnBnrrft no6er. Kolocxu He rrponulpmupyollue. 

Kla4onpoQnnau Seprunruue. OxoaoqserHnx uIerHHoK 

Her ... .. ... l. S. paniculatocorymbosus. 

* Coqserue pacnoJroxeHo n eepxuefi noJroBr,rHe penpoAyKTr.rBHoro no6era. 

Konocru B ocHoBaHuu nponn{r.rnr.rpypulrae. KlaAonpoQnmrrr crepr,tJrbHbte. 

Oxoloqrernuxqerr,rHoK2utt*t6 .... 6. 

6. OronoqserHrrx rqerunoK 2.Pac'r.eaue Jlaoca n Brernaua .. 2. ,S. petelotii. 

* OxonouserHbrx rrlerr,rHox 6. Pac'renue lOxnoro Krarar (o-r Xaftnaur) . . . 

3. S. hainanensis. 

7 (3). Konocxra co6panu no (2) 5-10 (13) n puxnosarrre,6. q. norryrrapoBnA- 

Hbre roJtoBKr.r 0.5-l or,r B Ar,raM.; B JrarepanbHbrx corlBerr,ux 2-5 roflosox. 

Kpotoune.reuryu rfiqenuAHbre; nnoAbr l-1.2 rr,rrra gl. Pacreuue Karar, n-osa 

Koper u flnounn ... 7. S. karuisawensis. 

* Korocxlz co6panrr no l0 -25 B oqegb nJrorHbre, 6. .r. uraponnAHbre roJroBrn(0.8) 

l-l.5crraBAr,raM.Pacrenux.flnounu ......... 8. 

8. Jlarepalnnrre coqBerur necyr I, oqeHb peAKo 2 rolonxn rorocxoe. JIuc- 

ToBbre nJracrlrnrn 3-6 MM rur,rp. Kpororqne qeruyu B ocHoBaHr,ra l-1.3 vrrr,r 

rur.rp. flno4u 1.3-1.4 MM An. . . . 6. S. fuirenoides. 

+ JlarepaarHbre coqBernr necyr 6oaee 2 ronosox KoJrocKoB. Jlncrosrre nracrrrHKlr 

zl--8 rtlu nrnp. Kponqt,re qeuryn B ocHoBaHr,rn 0.7 urr,r urnp. flnoArr 

I rvrra gl. . . . 5. S. mitsukurianus. 

9 (l). Prneq 2; nlo4rr nJrocxo-BbrrryKJrbre uJrlr HepaBHo ABorKoBbrnyKJrbre. . . 10. 

+ Prueq 3; nno4rr rpexrpaHHbre 13. 

45

10.Brarannua cre6resHx JrlacrbeB qacro cBerro-nypnypHble. Kpooulne veuryu 

qepHoBarbre. OxoroqgerHrte qerI,IHKH c enur o6paqeHHbIMLI sy6uartrr.r. Pac- 

TeHlren_oBaKauqa.rra. .....22. S.mic:rocarpus. 

Bnaranuura cre6resrrx JItIcrbeB He nypnypHble. KpotoutlIe rreuytr KoptlqHesue. 

OxoroqBerHbre uIerI,IHKLt c BBepx o6paulennrtua sy6qanall. PacteHnq 

Bocr., IOxs. n lOro-Bocr. Asiln . ......... ll. 

I l. Kporoqne qeuryl,t c 5 xurraull. Jlyvu coqner:ns rlepoxoBarble . . 

14. S. cluniunus. 

f Kpoourne qeruylr c I (cpeaneft) xnlrofi. J1yvn coqnerut ruaAKHe HJIu roltb- 

Ko BBepxy uepoxoBarbre ... 12. 

12. Braranr.rqa cre6resrrx JrncrbeB KopuqneBarbre. Jlyvr.r couBernq rnaAKr,te. 

Kolocxlr or npoAoflroBaro- Ao uI.rpoxoqfiqesr.rAHbrx, 2.5-3 rr.ru urnp. Kporculre 

qeuyu oAr.rHaKoBo oKpauenHbre no scefi noBepxHocrn, c l, nepe4xo 

cra6o rrrpaxenHoft, xulxoft ... ... 13. S. ternatanus. 

+ Bnarannqa cre6ressrx JrncrbeB 3eJreHbre. Jlyvr,r coqnernr BBepxy uepoxonarrre. 

KonocKr{ flpoAoJrroBa'ro-xfiuesraAHbre, oKoJro 1.5 rr,rna urup. Kporoqrae 

qetrryr4 c mnpoxofi sereHofi norocofi BAorb xopomo BbrpaxeHHoii cpe,uueii 

xaJrKr{ 15. S. roslhornii. 

l3 (9). Coune"rue ns 4-25 AoBoJrbHo KpynHbrx, 6-8 (10) MM An., 3-3.5 rrarrt urap., 

qepHoBarbrx KoJrocKoB. PacreHue Bocr. Cr.r6r,rpn n fianrnero Bocroxa . . . 

... 16. S. muximou'it':ii. 

* Couserue r43 MHoroqncreHHbrx MeHee KpynHbrx KoJrocKoB 14. 

14. Kporcrque qerrryfi qepHoBarbre HJrH rrepHoBaro-3eJreHoBarbre, peAKo Kopr.rqneerre 

(S. atrovirens). Oxouounernue uIerHHKr,r 6elrre, c nHu: o6pauleHHurrau:y6uaun 

uJrl,r rJraAK?re .. .... ... 15. 

+ Kporoqr.re r{emyr4 KopfiqneBbre. OxoroqserHrre ulerr4gKr.r cBerJro-Kop[qHe- 

Bbre, c BBepx o6parqenuunrlr ry6{auu r.rJrrr uraAKHe ......... 21. 

15. OxonoqserHbre u{err4HKu uraAxlre, yAnuuqroulr4ecr npn co3peBaunu nJroAoB 

r.r BbrcrynaKrrqlre rr3 Kporoqux veuryfi, npr{AaBar KoJrocKaM Br,rA (uepcracrr,rx>. 

Pacrenne Kanra3a 10. S. colclticus. 

t OxoroqserHbre r{err.rHKr,r c ry6uamn, He Bblcrynarcqfie r{3 xpotouqux veruyfi 

16. 

16. Konocxu pacnoJroxeHbr Ha BeroqKax corrBerr,rfl no oAHoMy. 17 . 

* Korocxu cKyrreHbr Ha KoHrrax BeroqeK no (l) 2-25 (30), o6parya nyr{Kr4 r.urn 

rrrapoBnAHbreroJronrr,r ..... 18. 

17. llyuu couBerut rJIaAKr,re. Korocxn 5-8 r,.ttl An. Kporcqne qeuryll Ha BepxyurKe 

rynbre r.rJur 3ar(pyrfiengrte. OxororlBerHbre rqeruHrt,t cnJIbHo t43RuIIucrue. 

Cre6lu r{HorAa npnrlr6arouluecr x cy6crpary n yKopeHrrorqrrecr B 

ysnax . 17. .S. radictuts. 

+ Jlyvn coqBerr,rr n aepxneft qacrr.r ilepoxoBarbre. Kolocru 3.5-5 r,arur .qn. 

Kporculr,re r{euyr.r Ha BepxytrrKe ocrpoBarbre. Oronoqnersrre uleruHKr,r nprurre. 

Cre6lu ne npurn6aroulnecr K cy6crpary r,r He yKopeurrouluecfl . . . . 

18. S. orientalis. 

18. Konocxr.r co6pauu no (l) 2-3 (5) a nyvrr,r. PacteHrq c non3yqilMr.l KopHe- 

Br.nuaMH ... 19. 

46

tI 

19. Konocxr,r o6rrqno pe3Ko cyxeHHble K BepxylxKe, ocrpoBarble I{JrH 3aocrpeHnue. 

Kpotou[e qeuryu ocrpoBarHe. Jlyvlt counerr,rq s HI,IxHefi qacru o6uq- 

Ho Dra.rlKr,re 18. S, orientalis. 

Korocxlr nocreneHHo cyxeHnble K BepxyluKe, Tynble. Kpotoulne qeuyu rynbre, 

3aKpyrJreHnbre nJrL yceqeHHble. Jly.rn counerns no eceft AJII4He tuepoxoBarbre 

... ... 19. S. sy'lvaticus. 

zo. 

Korocxu co6panrr no l0-25 (30) n nnorHrre ruapoBllqHble roroextr. Pacrennr 

6eg non3yrux KopHeBr,rq. . .. . . . 20. 

flnacrnHxn r,r BJraraJrnrqa HuxHLrx JrtrcrbeB y3JroBaro-cerr{arble or 6. u. :a- 

MerHo Bbrcryfiarculnx MeJrKr,rx nonepeqHbtx neperopoAoK. Korocxfi qilue- 

BrrAnbre unn npoAoJrroearo-sfiqesuAnlre, 2-5 MM An. Kponurue qeuyn Kopur{HeBbre. 

fhoAu (0.8) l-1.3 MM Arr. 

...21. S. alrovirens. 

fhacruuxN r.r BJraraJrrrua Hr4xnnx Jr[crbeB o6rtqHo He y3JloBaro-cerqarble. 

Korocru runpororftueBlrAHbre nnu sfiqegfiAnrre, 2-3.5 MM AJr. Kpoorqne 

qerryrl qepHoBarbre. flno.qrt (0.6) 0.7-l.l MM An. ... .. 20. S. hattoriunus. 

2l (14). Hraxnuft xpotoulnfi Jrr,rcr B 3-4 paza npeBbllilaer colrBerr4e. Konocxl.t 

raHrreroBrlAHue. Kporcrqlre qeuryr4 nJrorHble, 6yttlaxuc'rue, HHxHue creplurbHbre. 

OxonoqserHue uIerLrHKI{ nptMble 12. S. lilipes. 

Hnxnnfi xporou{r,rfi irrlcr HeMHoro npeBbruraer corrBerr,re. Korocxn sftqennA- 

Hbre, uupoKoqftuesuAHHe, tuapoBrlAHbre, uHoraa urur,rHApuqecxr.re. Kpotoune 

qeuyn nepenoHqarbre, nce Seprr,urbHbre. OKoroqserHue ulernHKu H3- 

Br{Jrrrcrbre .........2?. 

22. Ilytu coqBernr rJraAKr{e. Korocxr.r uapoBrrAH[,re, MeJrKHe, 2-2.5 uu an., co- 

6panurre Ha Kouqax BeroqeK 6. q. no 5-8 s rorosxn .. 9. ,S. rongdtengensis. 

+ Jlyqn corlBerr,rr uepoxoBarbre. Konocxr.r 6onee xpynnue . . . . . ... . .. 23. 

23. OcHosaHrrr Kpotoqux Jrr.rcrbeB KpacHoBaro-Kopr,rqHeBbre, KopfiqHeBbIe, 

r,rHorAa qepHoBarbre. Konocrn co6pauu no 2-15 B rycrbre noJIy3oHT[rKH, 

uruporoafiuewrgHbre, sftqeprrAnrre, r,rHorAa qr.urfiHApuqecxrae, 3.5-8 MM AJr. 

OxoroqnerrrrreurerriHKr.rrnaAKne ..... ll.S.c'yperinus. 

* OcHosaHnr Kpolourr4x JrncrbeB 3eneHbre aln 6reAno 6yponaro-xenroBarbre. 

Kolocru pacfioJroxeHbr Ha Beror{Kax corrBernr rro oAHoMy r4Jrr4, qaure, co- 

6panu B roM xe co[Bernu no 2-3 (5) n uapounaHHe roroBKr.{, ttrapoBnAno-afiqenuAubre 

HJrr4 qfiqesr.rAurre, 3-3.5 (4) MM An. OrofloqserHrre xrerrrH- 

:i : ::1i:1 r::r: : ::i:::i' :::i:*""::: ::::i :t:il!: i,,nn.",,i, 

Sect. l. Paniculatocorymlosi Kiik., 1930, Acta Horti Gotoburg. 5 : 36 

(

urzp., IrpoAoJrroBaro-sJrJll{rlr[qecKr,Ie, 6 8-qserxoeme. Kporoulr.re qeuryl,t 

KpacHoBarbre nnu Kopr4qHeBarbre, rrocepeAlrne c rufiporof 3ereHofr uonocoft 

u 3 xurKaMu, JlaHueroBrlAHbre, Ha BepxyrlrKe ocrl4crble HJIH ocrpue. 

OrolouBerHbrx u"Ierr4HoK 2 utru wx ner. Prueu 3. flno4u rpexrpaHHbre, 

o6parHorfi qeBrrAHbre, Ha BepxylxKe c qepHbrM rrr,rpaMr.rAaJrb- 

HbrM ocrarKotrr crol6uxa. PacreHus 35-55 crra Bbrc., c roHKr{Mlr croJlo- 

HaMr4 JIo 5 cu rur. nlu 6eg sux. 

T y p u s: S. paniculatocorymbosas Kiik. 

l. S. paniculatocorymhosus Kiik., 1930, Acta Horti Gotoburg, 5 : 35; 

Raymond, 1957, Nat. Canad. 84, 5 : 121 (in annot.); T. Koyama, 1958, 

Journ. Fac. Sci. Univ. Tokyo, sect. 3 (Bot.) 7, 6 : 291 (in annot.); 

Ts. Tang et F. T. Wang, 1961, in Fl. Reip. Pop. Sin. l1 : 15. 

Onucau us loro-3arr. Kuraq (upon. CuuyaHr); typus:

paxeHue r BHAeJra Ha se6-caftre Hrro-fioprccroro Eo'ranu.recroio ca4a. 

A4pec aroro cafira B ceru r,rHTepHer (cu. nuure), rAe pa3Mexleurr u:o6paxeHufl 

xpaHflulnxcr e NY ruuosnx o6parqoB u3 Bcex ceueficrn, nro- 

6esuo npe4ocraBuJr MHe A-p V. Touac (Dr. W. Thomas). B rexcre, corrpoBoxAarcqeu 

nro6paxeHr4e ynoMrHyroro o6pasrla S. petelotii, yxarauo: 

. Ho nonHoe coorBercrBr4e 3TuKerKu AaHHbrM upo- 

ToJrora Aaxrr ocHoBaHr,re paccMarpuBarb sror o6paseu B KaqecrBe rr,nra 

(nnu lerroruna hoc loco, ecJru B AanlHefiueu B Apyrux fep6apuax 

6ygyr o6uapyxeHbr Ay6rnKarH paccMarpuBaeMofo o6parqa). I4urepec- 

Ho orMerr4Tb, qro Ha rr,rrroBoM 3K3eM[Jrrpe ttMeercs rloMerKa

.r 4p.) nrUarvrl,I, o6beAI.{HeHHbIMII MHoft B ceKuI{P Beetleu, a raKxe eAI'{Hc'r- 

BeHHbrM BLTAOM CneAytouleft ceKquu Heterogenes. ouo sarnloqaeTct 

B Hiurtrqn14 y Br,rAOB BCex 3TI,IX Cerquft JraTepaJIbHbIx (tlOruuurO 'IepMI,lHtUIbuoro) 

counerufi. I4ttotu,a JrarepiurbHoe collBerl4e pa3BHTo vr y S. muximowiczii 

(sect. Japonici). Y 6onllrnHcrBa xe BI{AoB Scirpus o6qee couBe- 

Tr,re TOJTbKO TepMuHaJrbHOe. MOI6119 IIpgAISJIOXI{Tb, IITO COIIBSTI,IS, 6JII,I3- 

Koe K orrxcaHHoMy Bblure rl{rty, IIMeJII{ aHuecrpaJlbnnte Qopult Scirpus. 

Sect. 2. Heterogenes L. K. Dai, 1977,in Fl. Hainan 4: 538. ' B o'reHr 

KpaTKOM JIaTUHCKOM A[afHO3e ]TOr CeKIILTn OTMeqeHo' rrTo oHa poAcrBeHHa 

ceKIIr,IH Paniculatocorymbosi Kiik., or roropoil orlur{aercfl IIpucyrcrBr,reM 

B coqBc-rnu crepHJtbHbIX (a ne Seprr'ubHblx) rlta4onpo$ltit- 

JroB r.r [poJrr,rQu"ruapyroqux B ocHoBaHuI4 KoJlocKos. Eue oAHTIM o'rJIlIqlrTeJtbHbtM 

IIpII3HaKOM, CyAfl IIO pI4CyHKy eAI4HCTBeHHOI'O BI4Aa CeKqUn, 

fl BJrflerct HaJlnque 6 oKo.lroqserHblx ulerlllHoK. 

T y p u s: S. hainanensis S. M. Huang. 

3. S. hainunensis S. M. Huang, 1977,in Fl' Hainan 4:538,275, 

fig. I 143. 

Ouncas u:] IOxH. Kurax (o-n XaftnaHb); t y p u s : PE. 

Cupue u 6onorucrHe Mecra. -- IOro-Bocr. A3ufl: Kurafi ([Oxu.: 

o-s XailHaul). - - O6tu. pac[p.: 3HAeMuK. 

Comacso rporoJrory, Bl4A poAcrBen S. paniculatocorymbosus Kiik., 

oT KO'rOpOfO OTJlI,Iqaerc{ [pucyrcTBreM oqeHb KopoTKI{x fJIaAKI',IX OKO- 

JrouBerHbrx ulerr4HoK lI y3KI,IM TIJIOAOM (y s. paniculatocorymbo'trr.t [Ie- 

TT4HOK Her, a trJIoA O6parnorffrlesl4AHlrfr). 

Sect. 3. Beetlea Egorova sect. nova. -- TrichophorumPers., 1805' Syn. 

Pl. I : 69, p. p., quoad T. lineatum. - Scirpus L. sect. Trichophorum 

(Pers.) Darl., 1853, Fl. Cestr., ed. 3 : 340, p. p.; T. Koyama, 1958, Journ. 

Fac. Sci. Univ. Tokyo, sect. 3 (Bot.) 7, 6 : 291, p. p., quoad ser. 

Trichophorum (Pers.) T. Koyama, p.p.; Ts. Tang et F. T' Wang, 1961, 

in Fl. Reip. Pop. Sin. ll : 11, p.p.;A. E. KoNenHnror, 1988, n CocyA. 

pacr. coB. lanrH. Bocr. 3 : 180, p. P.i ou xe, Cutenlte [atIlu. Bocr. 

Pocc. : 58, p. p, -- S. sect. Androcoma (Nees) Benth. et Hook. f. ser. 

Lineatae Beetle, 1944, Amer. Journ. Bot. 31, 5 :264, nom. inval. (sine 

descr. latin.). - S. sect. Lineatus (Beetle) Oteng-Yeboah, 1974, Notes 

Roy. Bot. Gard. Edinb . 33,2: 3l l, comb. inval. S. sect. Androcoma 

auct. non (Nees) Benth. et Hook. f.: Beetle, 1947,in North Amer' Fl. 18' 

8 : 489. - Inflorescentia generalis ex anthelodio terminali et anthelodiis 

partialibus lateralibus I 4 distantibus constans. Spiculae numerosae, 

multiflorae, solitariae, rarius solitariae et binae (ternae) seu numero (5) 10 

25 in capitula plus minusve densa densissimave globosa vel semiglobosa 

congestae, 3.5-8 (10) mm lg., 2-3 (4) mm lt., ellipsoideae, late, oblongoet 

anguste ellipsoideae, rarius ovoideae. Squamae brunneae, rubello- vel 

griseo-brunneae, 1.8-2.5 (3) mm lg., ovatae, oblongo-ovatae vel lanceo- 

50

latae, apice acutae vel breviter mucronatae. Setae hypogynae 6, filiformes, 

pallide brunneae, plicato-flexuosae, leves vel superne denticulis 

parcissimis vix conspicuis, antrorsis, 3-4 plo fructu longiores, ex squamis 

haud exsertae vel plus minusve exsertae. Stigmata 3. Fructus trigoni, l- 

1.4 (1.5) mm lg., 0.6-0.7 mm lt., plerumque elliptici. Plantae 100-150 cm 

alt., dense caespitosae rhizomatibus repentibus destitutae. 

T y p u s: S. lineatus Michx. 

Sectio in memoriam generis Scirpus L. investigatoris americani cl. 

A. A. Beetle nominatur. 

O6uree couBerrre MHofoKoJrocKoBoe, o6paroaaHuoe repMuHaJrbHbrM 

n l- 4 narepanbHbrMlr [apu[iulbHblMll coqBerutMu. KoJrocKn MHororruc- 

JreHHbre, MHoroI{BeTKoBbIe, pac[oJloxeHbt Ha KoHqax BeToqeK couBeTHfl 

rro oAHoMy, peAKo to | 2 (3), nr,r6o cKyqeHbr no (5) 10 25 s pbrxJroBa- 

Tbre r4JrH orreHb [roTHbre uapoBr,rAHbre r{Jrr,r rroJryrxapoBrrAHbre roJroBKr{, 

3.5 8 (10) MM AJr., 2-3 rrarra urup., 3JrJrr4rrcorrAiurbHbre, rrupoKo-, rrpoAoJrro- 

Baro- r,rJrn y3Ko3JrJut[col{AilllbHbre, pexe gftueurcAuue. Kpotou-1ue qerllyu 

Kop[qHeBbre, KpacHoBaro- r{Jrr.r cepoBaro-Kopr{rrHeBbre, 1.8 -2.5 (3) naru gn., 

qfiueBr,rAHbre, [poAoJtroBaro-qftuesraAuble uJtI{ JIaHueroBrrAHbIe, Ha BepxyurKe 

ocrpbre r{Jrrr c KoporKI4M ocrpldeM. OKorouBerHbre uIerllHKH 

B qttcre 6, uutenu4Hble, cBerJro-Kopr,rqHeBHe, r,r3BrrJrncrbre u crryraHHble, 

fJlaAKue r.rJu4 B BepxHefi, qacru c oqeHb peAKuMr{, o6parqeHHuMH BBepx 

sy6qaun, y 34 paza AJrr,rHHee rrJroAa, He Bbrcryrrarcxlr4e r4Ju4 6. M. strc- 

Tyrrarcullze H3 Kporcqux ueuryft. Prrleq 3. fhoAu rpexrpanHble, 1 1.4 

(1.5) uu an.,0.6-0.7 MM ruup.,6. q. unuurr,rqecKr4e. Pacrenraq 100 

150 cu Bbrc., rycroAepHrrcrbre, 6er nomy.rux KopHeBnq. 

Tun:5. lineatus Michx. 

Cerqus Ha3BaHa B rraMflTb A. A. Erarna, lr3BecrHofo aMepKKaHcKoro 

r{ccfieAoBarenr poAa Scirpus L. 

BuAu erofi ceKuur{ -- S. lineatus, S. wichurae, S. /uirenoides w gp. 

o6br.{Ho o6"se4wns;lucl s oAHofi ceKqr{r - Trichophorum (Perc.) Darl., 

BMecre c Br,rAaMu S. cyperinus (L.) Kunth, S. asiaticus Beetle (= S. lushanensis 

Ohwl), S. atrocincta.s Fern. r,r Ap. (T. Koyama, 1958, l. c.; Ts. Tang 

et F. T. Wang, 1961, l. c.; A. E. KoxesHaros, 1988, 2001, uur. coq. r 

Ap.). Eurir (Beetle, 1944,1. c.) cupane,4nnBo ucrrroqw S. lineatus u3 ceKuuu 

Trichophorum, BbrAeJrr,rB ero B pf,A Lineatae Beetle (nom. inval.). 

flocreAuuft Bnrn oruec K ceKrlrru Androcoma (Nees) Benth. et Hook. f., 

trro, oAHaKo, HeJrb3r [pr43Harb [paBHJrbHbrM, TaK KaK S. lineatus He r,rMeer 

6nusrux poAcrBeHHbrx cBr3ef c rr{rroM poga Androcoza Nees n coorBercrBeHHo 

ceKuur4 Androcoma - 6parnllcKnM BLTAoM S. giganteus 

Kunth, 3aueruu, qro poA Androcoma paccMarpuBaercs, B KaqecrBe ca- 

Mocrof, TeJrbHoro HccJIeAoBareJrrMr,r roxHoaMepr{KaHcKHX C y p e r ac e ae 

(Luceflo, Alves, 1997). Cerur4oHHoe Ha3BaHr4e Linestus (Beetle) Oteng- 

Yeboah, l. c., lt;tletcfl HeBaJruAHbIM, B cBr3r4 c qeM HaMll .IIaHo ouncanr.re 

Hosofi ceKrrun. 

5l

OcHosHoe u AoBoJIbHo cyulecrBeHHoe pa3.irfique MexAy rlpeAcraBnreJrrMn 

orrr4cbtBaeMofi ceruuu Beetlea v BL4AAMi1I CeKqI'In Trichophorum 

(reuepr oHa Hocr,tr Ha3BaHLIe Pseudotrichophorum, cn' uuxe) cocrol{r B 

crpoeHr4t4 colrBerut. Y slaAos ueprofi cerullll o6ulee collBerne o6paso- 

BaHO TepMrrHaJrbHblM r.r I 4, pacrroJloxeHHblMlt B na3yxax BepxHl4x cre6- 

JreBbrx Jrr,rcTbeB JIaTepaIbHbIMt4 couBeTuflMLI, TOfAa KaK y BI'IAOB n'rOpOft 

ceKuuH - oAHrrM TepMHHaJIbHnrr,r. .{pyroft saxuHfr rIpI'I3HaK, IIO KOTOpoMy 

orJruqalorct rlpeAcraBl4Teru o6eHx cerqufi 3To coorHoureHlle 

AJrrrHbr oKoJrouBerHblx urer[HoK I{ Kpoloulux ueuryfi. V s}rloa ceKIr}lH 

Beetlea oKoJrouBerHble ureruHKrr He BblcrylrarcT uJIu HeMHoro Bblcry[aror 

r13 Kporoulux ueuryfi u,3a peAKIlM lrcKnloqeHl4eM, He rIplIAaIOT KOJIOC- 

KaM Br{A (ulepcrrrcrblxD. OKoloqBerHble IueruHKI't y BI4AoB ceKqull 

Pseudotrichophorum, KaK rlpaBllJto, 3aMerHo rlpeButuarlr Kpolourue qeuIyH, 

qro rrpflAaer KoJlocKaM Id BceMy couBerl4lo ((uepcrucrblfu> o6ltlx. 

Cnegyer orMerurb, trro aBTopbt o6pa6orru Scirpus no > (Whittemore, Schuyler, 2002) raxxe cqlrraror BHAbr 

poAcrBa S. lineatus c oAsofi cropoHbl, I,I poAcrBa S. cyperinus - c Apyrofi, 

o6oco6JreHHblMrr, o qeM MoxHo cyAI,I'l'b lro pacnoJloxeHun 3Tux 2-x 

rpyrfl BrrAoB B c]IcreMe poAa. BHAbI nepnofi fpy[rtbl rloMeuleHbl B Hauane 

o6pa6orKlr, BTopoft -- B KoHue ee. O6oco6neHHocrb paccMarpl{- 

BaeMbrx rpynu BrIA,oB He [oflytrl4ra n o6pa6orre raKcoHoMnqecKofo BbIpaxeHur, 

[ocKoJrbKy B Heft Her BHyrpl4poAoBblx IloApa3Ae.neuufi. B sacToflulefi 

crarbe 3Tr4 ABe fpyutlbl BHAoB, orHocsull.Iect K ceKuKtM Beetlea 

u Pseudotrichophorum, pacrloJloxeHbl pqAoM, TaK KaK oHI4 6oJlee 6ru3- 

Kn MexAy co6ofr, rIeM c ApyruMu ceKul{sMu Scirpus. 

4. S. wichurae Boeck., 1869-1870, Linnaea 36 :729 (), p. p., 

quoad pl. ex Jap.; Ohwi, 1944, Mem. Coll. Sci. Kyoto Univ., ser. B, 18, 

I : 107; id., 1965, Fl. Jap. : 202, p. p., quoad var. wichurae; Bopoul., 

1966, @.1. con. .{alrH. Bocr. : 80, p. p., quoad pl. kuril.; oH xe, 1982, 

Oupeg. pacr. coB. .{anlu. Bocr. : 99, P. P., quoad pl. kuril.; A. E. KoxeBHr{KoB, 

1988, n Cocya. pacr. coB. .{iurrn. Bocr. 3 : 180, rapra 57. 

S. eriophorura Michx. var. nipponicus Franch. et Sav., 1877, 1878, Enum. 

Pl. Jap. 2: 114,545; Meinsh., 1901, Tp. flerep6. 6or. caAa 18,3:249. 

S. concolor Maxim., 1886, Bull. Acad. Sci. P6tersb.3l, I : 110;id., 1886, 

M6l. Biol. 12 : 556. - S. fuirenoides Maxim. f . minor C. B. Clarke, 1904, 

Bull. Acad. Intern. G6ogr. Bot. 14 : 200, p. p., quoad coll. Faurie 4918 

fide T. Koyama, 1958, Journ. Fac. Sci. Univ. Tokyo, sect. 3 (Bot.) 7, 6 : 

339. S. cyperinus (L.) Kunth var. wichurae (Boeck.) Makino, 1904, 

Bot. Mag. Tokyo 18 : 120. -- S. cyperinus var. concolor (Maxim.) Makino 

f. cylindricus Makino, 1932, Journ. Jap. Bot. 8 : 46. -- S. wichurae f. 

cylindricus (Makino) Nemoto, 1936, Suppl. Fl. Jap. : 1023 Ohwi, 1944, 

L c. : 108. -- S. wichurae f. borealis (Ohwi) Ohwi, 1944, l. c. : 107. 

S. wichurae f . concolor (Maxim.) Ohwi, 1944, l. c. : 107. -- S. borealis (Ohwi) 

T. Koyama, 1957, Journ. Jap. Bot. 32: l5l, quoad nom. - S. lineatus 

52

Michx. subsp. wichurae (Boeck.) T. Koyama var. wichurae (Boeck.) 

T. Koyama, 1958, l. c.:339, p. p., excl. pl. corean. S. lineatus subsp. 

wichurae var, wichurae f . wichurae (Boeck.) T. Koyama, 1958, L c. : 339, 

p. p., excl. pl. corean. --5. lineatus subsp. tt'it'hurae var. wichurae f. 

concolor (Maxim.) T. Koyama et f . cylindrfca.r (Makino) T. Koyama, 

1958, L c. : 341. -- S. lineatu.r Michx. subsp. wichurae var. lushanensi.s 

(Ohwi) T. Koyama, 1958, l. c.:341, quoad pl. 

Onucan r43 BocrorrHoi'tVI::!lu (ropu Kxacra) u.fluonnra (o-a Xoucto, 

I4ororarua) (); s y n t y p i : yrpavenu n fep6apura EeplraH-,[anerraa 

(B) Bo BpeMr Bropofi unponofi sofiHu. 

Eonorucrne Jryra, cbrpble Jleca. --'. Poccrr. ,\anau. Bocmox: Kypul. 

(fOxnue Kypr,rnu). -.- Bocr. A:ur: .fnour,ra (o-na XoxrafiAo, XoHcto, 

Curory ra Krouo). -- O6ur. pac[p.: aHAeMI{K. 

Cncreuaruxa u HoMeHrnarypa S. wichurae AoBoJIbHo cnoxHm. 3ror 

Br{A ocHoBaH, KaK cJreAyer H3 rrporoJlofa, Ha SK3eMrIJItpax, co6paHurrx 

s socroqHoft L4vruu (ropu Kxacn) ra s fltoll,uu (o-e XoHcto, Llorot'avra). 

.{nr r[oHcKlrx pacrenuft Ha3BaHue S. wichurae 6ulo nuepnble lrcrroJrb3oBaHo 

ManuHo (Makino, 1904, l. c,) n paure pa3HoBI,IAHocrLI -- 

S. cyperinus (L.) Kunth var. wichurae (Boeck.) Makino, l. c., a sarela 

Oor,r (Ohwi, 1944,1. c.) a paHre ru4a. flpexge 3rI,I pacreHllfl oroxAecrB- 

JlflJrrrcb c orurcaHHbIMI,I H3 Cee. AuepuKI,I BI,IAaMI,I S. cyperinus (L.) 

Kunth uru c S. eriophorum Michx. (= S. cyperinrzs). Hasnauue S. wi.churae 

ytorpe6flrercfl TaKxe pqAor\4 aBropoB Am o6osHavenut pacleuufr. n 

r,r3 Apyrux perroHoB Asr,rn (Maximowicz, 1886; Korr,rapos, 1901; Poxesuu, 

1935; Raymond, 1957; Kern, 1974; Eropona, Kxoft, 1980; Simpson, 

Koyama, 1998; Kxoil, 2002 u gp.). 

KoqN,ra (T. Koyama, 1958: 344), ocuonblBatcb Ha opHrHHarbHoM 

oilrrcat vr S. wichurae, y6eAr,rrenrHo [oKa3ail, qro oHo ornocnTct K 

rrroHcKHM pacreHr{rM, rrocKoJrbKy orMeqeHHbre B HeM rrpu3Haxr.r (upncyrcrBlre 

B o6qeu corIBerHI,I, troMnMo repMI,IHaJIbHofo 3oHTI,IKoBHAHofo 

coqBerr{r, JrarepaJlbHblx coqsetl4ft, BblxoAtlllfix I,I3 [a3yx cre6lrerrrx nucrbeB, 

pacrroJloxeHr,re KoJrocKoB 6. .r. uo oAHoMy Ha KoHeqnbrx Beroq- 

Kax cor.IBerru, Hal[true rrJroAoB, ua r/, xopo.re Kpoloulrlx .reuryfi) caoftcTBeHHbr 

TOJTbKO 3Tr,rM pacTeHr,rrM, HO OTCyTCTByIOT y BCeX BHAeHHbTX r{M 

o6pasqon r43 Marepr{Konofi qacru Asura (ur I4la1r'ru rr ropHbrx pafiouon 

lOxHoro Knrar). flocneAnue 6rmn uAeurnQuqupoaauu Kosuofi c our4caHHbrM 

us loro-3an. Kurar (npon. IOnrnanr) S. asiaticus Beetle 

(Beetle, 1946). K sroMy BbrBoAy Kosnaa npuueJr Ha ocHoBaHnvr v3yqe- 

HnflTfina 3Toro BnAa, xpaHflrqerocx n NY (nro6paxeHue ero HMeercf, Ha 

ne6-caft re Hrro-fr oprcroro 6oraru{qecKoro cagLa - http://www.nybg. 

org/bsci/vasc/Cyperaceae.html - !). K S. asiaticus (= S. lushanensis) 

rrpr.rHaAlexar pacreHur H r43 Apyfnx peruoHoB Asran (xporrae .fnoHnu), 

53

pnBeAeHHbre uI{TI4poBaHHbIMH Bbrule aBTopaMI{ xar S. wichurae. Ocnos- 

Hbre [pu3HaKI4, o'rruqaloulue S. asiaticus or S. wichurae orcyrcl'Blle 

Jra'repaJlbHblx couseruff I4 HaJluque AJIItHHbtx oKoJIoIIBerHbIx ulgrl4HoK, 

3aMeTHO rrpeBbrilralou-lltx Kpolou-lue qeu]yl4 14 npl.rAaloulux KoJIocKaM u 

BceMy corrBerulo xapaKTepuufr,tr o6lur. Y S. wichurae oKo- 

JroqBeTHbre UIeTUHKI{ He BbICTyIlarcT I{JrU HeMHOIO BbICTyrIarcT lI3 Kporculux 

ueuryfr, BcireAcrBr4e qelo KonocKI{ y 3Toro BHAa He (tuepclncrbleD. 

Koqlaa (T. Koyama, 1958, l. c.) rrpl.rHrn S. wichurae B KaqeclBe roA- 

Br.rAa aMepuKaHcKoro snAa S. lineatus Michx. -' S. lineatas subsp. 

wichurae (Boeck.) T. Koyama, BblAeJluB s uela 3 pa3HoBlaAHocru - var. 

wichurae (Boeck.) T. Koyama, var. asiatfca.s (Beetle) T. Koyama 

(= S. asiaticzs Beetle, = S. lushanensrs Ohwi) n var. lushanensls (Ohwi) 

T. Koyama. B Hacrosqefi crarre 2 uepnue pa3HoBLIAHocrH coorBercroyrcr 

S. wichurae u S. lushanensls (= S. asiaticus);var. lushanerstr e tto- 

HrrMaHrrr4 Kosun - 3'ro S. wichurae (cu. rovrureu'rapuit x S. lushanensi.s). 

Cne4yer paccMorperb Bonpoc o ruuu$urauura S. wichurae. Buure 

6nno ynouxHyro, qro no c[paBeIUIrrBoMy 3aMeqaHulo KoqN,Itt (Koyama, 

1958 : 344), orrncaHr4to S. wichurae orBer{alor trroHcKue pacreHur. Kpo- 

Me rofo, fipu Ha3BaHHr.I pa3HoBrrAHocrl{ var. wichuraeo Kor,ua 

(T. Koyama, 1958, l. c. : 339) rpoul.rrl{poBur:

(Dr. D. A. Simpson) u A-Pa C. Ka

Onucan wz Atuauu (o-n XoHcro) (); I ec t o t y - 

p u s (HonoceroBa, 2001 : 258):

HaAJrelKar K oAHoMy rr roMy xe raxcoHy. flocneAHuft HecoMHeHHo cJre- 

Ayr rrpuHr,rMarb B paHre Brila, T. e. KaK S. karuisawensls. Or S. Jidrenoides, 

pac[pocrpaHeHHoMy roJrbKo n fuonvr4, S. karuisawensls xopouro 

orlnrraercr rro xapaKTepy crpoeHm couBerr.rfl. Y S. fuirenoides xo- 

JrocKrr co6paHu uo l0-25 B oqeHb rrJrorHbre, KpyrrHue (0.8) l-1.5 crvr 

r guau.), uapoBr,rAHbre r{Jur rroJryrxapoBr4AHbte roJroBKr,r, qro rrpuAaer 

pacreHrlro ueo6uqsrfi pw, Scirpus o6nun, rtpnqeM JrarepaJlbHbre (ra:yu- 

Hrre) coqnerur Hecyr l, oueHl pe4rco 2 roJroBKH. B couneruu S. karuisawensis 

roJroBKr,r pbrxJroBarHe, MeHee KpyrrHbre (0.5 I crra n 4uau.), coc- 

Torru[e us (2) 5-13 KoJrocKoB, a JtarepaJrbHbre couBerru o6pa:onaulr 

6..r. 2-5 ToJIoBKaMH KoJrocKoB. Kponae roro, Jryrn couBerr4fl y S. karuisawensis 

roHKue (y S. fuirenoides saueruo ronule), Kporoqne qeuryu 

rftqesr,rAnrre,2-2.2 MM AJr., rrJroAbr l-1.2 rvura AJr.(y S. .fuirenoides xporort.(ne 

qeuyu fipoAoJrroBaro-sftqeelrAHbre Ao JraHue'r'oBlrAuwx, 2 2.5 

(3) rraru AJI., trJroAbr 1.3--1.4 rvrM .qn.). 

Sect. 4. Pseadotrichophorum Egorova, 1998, Hosocrr{ cucr. Bbrctu. 

pacr.3l : 10. - Trichophorum Pers., 1805, Syn. Pl. I :69, p. p., quoad 

T. cyperinum, excl. Z alpino et T. lineato. -- Scirpus L. sect. Trichophorum(Perc.) 

Darl., 1853, Fl. Cestr., ed. 3 : 340,p. p.; Beetle, 1944, Amer. 

Journ. Bot. 3l :264;id., 1947,in North Amer. Fl. 18, 8 :491; Fern., 

1950, in Gray's Man. Bot., ed.8 :2751' T. Koyama, 1958, Journ. Fac. Sci. 

Univ. Tokyo, sect.3 (Bot.) 7,6:291, p.p.; Ts. Tang et F. T. Wang, 

1961, in Fl. Reip. Pop. Sin. ll : ll, p.p.; Y. L. Changet Y. L. Yang, 

1976, in Fl. Pl. Herb. Chin. Bor.-Or. I I : 8 et auct. plur. ,S. sect. 

Sylvaticae C. B. Clarke, 1893, inHook. f., Fl. Brit. Ind.6:661, p. p. et 

excl. typo. -- S. sect. Taphrogeton auct. non Reichenb. : Poxes., 1935. 

no (Dn. CCCP 3 : 44, p. p. --_ Coqnerze repMr,rHuurbHoe, c MHororrr4cJreH- 

HbrMr,r KoJrocKaMtr, pac[oJroxeHHbIMI,I Ha KOHUaX BeTOqeK tlo OAHOMy 

nJrr4 B roJroBr{arbrx rryqKax no 2 5 (8). Konocru 3-6 ( 10) vrrvr ur., 2 3 

(5) uu urnp., sftueuAHbre, ruuporoafiqeBr.rAHbre, uapoBl,rAHbre, peAKo 

3JIJII{rrcor4Atulurue. Kpotoulue r{euyu Kopr,rrrHeBble, pxaBo-Kopr{qHeBbre 

I4Jru qepHoBarne, 1.3-2. 5 rr,ru 4n., rft qean4nue, runpororft ueBr{AHbre, 

rlpoAoJrroBaro-rfiuenltAuble uJur rlpoAoJrfoBaro-SJrnr4rrrl4qecKrde. Oxono- 

IrBerHbte uIeruHKI,t B rrucre 6, cnerno-ropllrrHeBble I,uru 6enosarble, HI,I- 

TeB?IAHbIe, 6. M. ugsl,ull,lcrble I{ crlyraHHble, rJraAKI,Ie l,tJrr,r B BepxHeft ,{acrr{ 

c eAnHr4qHbrMr.r, o6paqeuHuMrr BBepx 3y6qauu, n 4 6 par Arr4HHee 

[JroAa, npu co3peBaHr{r,r rrJroAoB BbIcTy[aK'rIlHe r,r3 KpoKrulux veruyft, 

o6H.{Ho npuglaBar KoJrocKaM

3aHHOe CeKqrroHHoe Ha3BaHI,Ie yxe He MofJlo ooJlbue HclloJlb3oBaTbcfl B 

poge Scirpu.r, BcJreAcrBI4€ r{ero paccMarpuBaeMat rpyutla nuAos 6ltrta 

orrr4caHa MHoIo (Egorova, 1998, l. c.) uol ceKquoHHbIM HasBaHlleM 

Pseudotrichophorum. 

8. S.Iushanensrb Ohwi, 1938, Acta Phytotax. Geobot. (Kyoto) 7 :134; 

Ts. Tang et F. T. Wang, 1961, in Fl. Reip. Pop. Sin. Il: 12 (pro syn. 

S. asiaticus);M.Zhan et Y. C. Yang, 1987, Acta Biol. Plateau Sin. 7: 

12. - S. wichurae Boeck., 1869 1870, Linnaea 36 :129 (), 

p. p., quoad pl. ex Ind. Or. --' S. asisticus Beetle, 1946, Amer' Journ. 

Bot. 33, 8:662; Ts. Tang et F. T. Wang, 1961, l. c.:12, p. p', excl' syn. 

nonn., tab. III, fig. l-5; Y. L. Chang et Y. L. Yang, 1976, in Fl' Pl. 

Herb, Chin. Bor.-Or. ll :9, fig.4; Y.T. Lee,1976, in Fl. Cor.7 :268, 

p. p., excl. syn. nonn.; Kitag., 1979, Neo-Lineam. Fl. Mansh.: 153; 

Y. C. Yang, 1987, in Fl. Xizang. 5 :346; A. E. KoxeeHI4KoB, 1988, s CocyA. 

pacr. con. .{,anrH. Bocr. 3 : 181, rapra 57. S. lineatus Michx. 

subsp. wichurae (Boeck.) T. Koyama var. asiqticus (Beetle) T. Koyama, 

1958, Journ. Fac. Sci. Univ. Tokyo, sect. 3 (Bot.) 7, 6 :342. -' S. lineatus 

subsp. wichurae var. lushanensrs (Ohwi) T. Koyama, 1958, l. c. : 341, 

quoad nom. -- S. wichurae Boeck. var. asiaticus (Beetle) Ohwi, 1965, Fl. 

Iap. : 202. - S. eriophorum avct. non Michx. ; C. B. Clarke, 1893, in 

Hook. f., Fl. Brit. Ind. 6 : 661; id., 1903, Journ. Linn. Soc. London (Bot.) 

36 :249; Meinsh., 1901, Tp. flerep6. 6or. caAa 18,3 : 249; Nakai, 19l l, 

Fl. Kor. 2:293. S. wiclrurae auct. non Boeck. : Maxim., 1886, Bull. 

Acad. Sci. P6tersb. 31, I : 111; Korrr., 1901, Tp. flerep6. 6o'r. cap:a20 

((Dn. MaHr.rx. l) : 339; Poxes., 1935. so (Dn. CCCP 3 : 45; Raymond, 

1957, Nat. Canad.84,5: 115; Bopon., 1966, @r. cos.,{anln. Bocr.: 

80, p. p.; oH xe, 1982, Onpe.q. pacr. cor. ,{anlu. Bocr. : 99, p. p., quoad 

pl. ussur.; Kern, l974, in Steenis, Fl. Males., ser. 1,7, 3:502, Eropoea 

u H. K. Kxofi, 1980, Honocrr{ clrcr. Bblcru. paw. 17 : 55; Karthikeyan 

et al., 1989, Fl. Ind. Enum. Monocot. :71;D. A. Simpson a. T. Koyama, 

1998, in Fl. Thailand,6,4:281; T. B. Lee, 1999, lll. Fl. Kor. : 172; 

N. K. Khoi, 2002, in Fl. Vietn. 3 : 57. S. cyperinus (L.) Kunth var. 

eriophorum (Michx.) Makino, 1904, Bot. Mag. Tokyo l8 : 120, quoad 

pl. S. wichurae var. borealis auct. non Ohwi: Kitag., 1939, Rep. Inst. 

Sci. Res. Manch. 3, App. l(Lineam, Fl. Mansh.) : 123. S. borealis 

(Ohwi) T. Koyama, 1957, Journ. Jap. Bot. 32: l5l, quoad pl. 

Ouncau ras l{eurp. Kuraq (lpon. Ifrrucr,r) (); t y p u s : (KYO; nugela 

nso6paxeHr4e ruua (digital image), pacreqarKa ero r tuuorofi 

KonreKur4l4 n LE!). 

Eolorucrue nyra, 6epera peK, cbrpbre Mecra B recax, B ropax Ha 

Bbrcore l70O 22OO M HaA yp. M. -- Poccufl. .il,anou. Bocm.: Vccyp. 

(rcr). Bocr. Arun: Kurafi (Cen.-3au.: Irpon. faulcy; Cer.-Bocr.; Cen.: 

58

npor. [IIaHlcu; Bocr.; [enrp.: npor. XauaHr, Xy6:il ra I{:laHcn; [Oro- 

3an.: npoa. Cuuyaur, IOHrsanr ra fyfiuxoy; Keichow (T. Koyama, 

1958, l. c.)), n-on Kopea. - IOxu. Asus: I{HAua (Bocr. fnuattau, AccaM, 

ropbr Kxacn), ? Heuan. -- IOro-Bocr. Alus: Knrafr (IOxH.), Blernau, 

I4H4otte:ua (o-a Cyrurarpa). -- O6nr. pacup.: 3HAcMI4K. 

PacreHur irofo BtIAa oAHId aBTopu o6osHaqaJtl,I KaK S. asiatictr,s 

Beetle nrru var. asiaticus (Koyama, 1958; Tang, Wang, 1961; Ohwi, 1965; 

Chang, Yang, 1976;l{ttagawa,1979; Koxensnros, 1988,2001 u Ap.), a 

Apylue -- KaK S. wichurae Boeck., L c. (Maximowicz, 1886; Korvrapon, 

l90l; Poxenraq, 1935; Raymond, 1957; Kern, 1974; Eropoaa, Kxofi, 

1980; Simpson, Koyama, 1998; Kxoft, 2002 u Ap.), upnqeN4 HeKoropbte 

lr3 Hrlx cr{HoHrrMlr3r,rpoBaJrr{ c 3TI,IM Ha3BaHIIeM S. asiaticus. B ronaueHraprlrx 

x S. wichurae (cu. ruure) 6uno [oKa3aHo, qro S. wichurae u 

S. asiaticus -- 3To pa3Hbre BuAbl 14 qro lrporolory S. wichurae coorBercrByK)'r 

pacreHrrt, o6ruee coqBerl,Ie Koropblx cocrol,IT t{3 TepMI4HaJIbHoro 

u JrarepaJrbHbrx (uasyuHux) coqnernfi, KoJrocKlr pac[oJroxeHbl Ha 

KOHTIaX BeTOqeK B OCHOBHOM rro OAHOMy, a OKOJlOqBerHble qerI4HKH He 

rtpeBbrtualor rrJru HeMHoro npeBbruanr Kpotoulue qeuyu n o6ltquo ue 

rrpr4Aaror KoJrocKaM (uepcrr{crbrfi> o6lur. O6pa:r1n oIIucaHHolo rr4lla 

Lr3BecrHbr roJlbxo u3 Anouuu, a B Apyrnx peruoHax A:nlr ncrpeqaK)Tcr 

pacreHl{fl, orJruqalouluecr or flrroHcKlrx lJraBHbrM o6pa:ou repMu- 

HaJlbHbrM corrBerr{eM, KoJIocKaMu, pac[onoxeHHbrMl4 Iro (l) 2-3 (5) u 

oKoJrouBerHbrMu urerr,rnKaMu, 6.u. 3aMerHo rIpeBbIuaIoIqI4MI{ Kpolourl,Ie 

qeuyn, r,pup.aBas. KoJrocKaM rr BceMy corrBerrclo ( suA. Ta- 

Kr4e pacreHus 6rrlu o[r{caHbr ns [Oro-3anaAHoro Kuraq (npon. IOnluaHr), 

xar S. asiaticus Beetle, 1946, l. c. (typus:

Hec x Hefi, KaK cJreAyer u3 rrpr4BeAeHHoro ofilrcaHr,ra (> - ) o6pa:t\vt 

S. wichurae. 

PeffuoHa (Raymond, 1957,1. c.), oruocnnurlrft, noAo6Ho prAy Apyfrrx 

ur4Tl4poBaHHbrx Bbrrxe aBTopoB, paccMaTpr,rBaeMbre 3Aecb pacTeHufl 

x S. wichurae (c aruu Ha3BaH[eM oH crrHoHHMH3r{poBaJr S. asiaticus) 

orrucaJr H3 rleHrpanbHofi .{acru BrerHapra (Annam) pa3HoBrrAHocrb -- 

S. wichurae var. donaicus Raymond (1957, Nat. Canad, 84, 5 : ll7; EropoBa 

r{ H. K. Kxofi, 1980, Horocrlr crrcr. Bbrcur. pacr. l7:55; 

N. K. Khoi, 2002, in Fl. Vietn. 3 : 59. ---- Typus:

vLfl. MexAy ,{ugune H Xo6uc-rasu rI rrcroKlt p, Jle6apge; A4xapna, 

ceB. MaKpocrnoH lll4suercKoro xp. - oKp. c. foraAsee6u u lllyaxencxuft 

p-n - p. T6erra). -- O6q. pacrp.: 3HAeMnK KonxuAu. 

K. P. Kr,ruepu4se (Kimeridze, l. c.) - anrop orrr{caHr.m S. colchicus, 

crrpaBeAJurBo cblnxal ero c ceBepoaMepuKaHcKl{M BI4AoM S. atrocittctus 

Fern. OAnaKo rrprr3HaKu, yKa3aHHbre Klruepugre B KaqecrBe orrurru- 

TeJrbHbrx &rlr S. colchicus, He rBJrrlorcfl, ua uoil B3rJlrA, raxoeHrvru. Aualus 

rep6apHoro Marepuala uo S. colchicus w S. utrocinclas 14 Aela[bHofo 

otrr4caHnfl nocreAHero Br,rAa, [p[BeAeHHolo so (Flora of North 

America> (Whittemore, Schuyler, 2002) nora:aJr, qro S. colchicus tpeznHqafiHo 

6nurox H [paKTHqecKH nAeHTr,rqen S. atrocinctus, olrnqaflcb 

or Hero roJrbKo Solee AlunnbrMr.r u y3KHMrr Kpororru4Mr,r qetuyflMu 

2-25l'al.l 41. (a He 1.3-1.8 vur,r) n 6onee AJurHHbrMr4, cr,rJrbHee rrpeBrttua- 

K)uuMn Kporoque qeuyu oKoJrorIBeTHbrMrr uIeTr{HKaMr,r. 

PaHee unoro 6rmo slrcra3aHo MHeHr4e (Eropooa, 1998) o AoBoJrr,Ho 

6rusrov poAcrBe S. colchicus c BocroqHoa3lrarcKr4Mr,r Br4AaMH - S. asier 

ticus Beetle n S. wichurae Boeck. Cuequallnoe r43yqeHr4e rrocJreAHux rro- 

Ka3aro, ,{ro s AeficrBr4TeJrbHocrr4 S. colchicus ue o6uapyxuBaer c ?Trr- 

MIa B[AaMI4 6nnsxoro poAcrBa, xoporxo orrr4qarcb or Httx 6enosarlrNu.r 

rJraAKr{Mr4 (a ue cnerno-KopuqHeBbtMu, B Bepxueft .racrn c pe4xuuu, o6parreHHbrMrr 

BBepx MenKr{un sy6qaun), uepuoraruuu (a He KopnqHeBbruu) 

rporoull,rMu r{euyrMr.r u qepHbrMr,r flpll ocHoBaHur{ Kporoxlr4ur4 Jrnc- 

TbrMr{. Bce uepevucneHHbre [pr.r3HaKu S. colchicus csofrsrneHHrr u S. atrocinctus. 

Heo6rrqafiHoe cxoAcrBo S. colchicus c S. atrocinctus 14 olcy'tcrBue Ha 

eBpa3l4arcKoM KoHTUHeHTe BrrAoB, 6ltlrsxnx K KaBKa3cKoMy, Moxer Ha- 

Becrr,r Ha MbrcJrb o 3aHoce S. atrocinctus ug Ces. Auepnrclr ua Kanras, 

qro, oAHaKo, rpyAHO AOIIycTLlTb, yrluTblBat [pol,I3pacTauue S. colchicus 

B ropHhrx naecroo6uraHr4rx Ha Bbrcore 800-1500 M HaA yp. ru. l,I:eec'r- 

Ho HecKoJrbKo MecroHaxoxAesnfi BkLAa B 3arI. 3arasxasle (Elopona, 

1998) r.r o4uo ua 3an. Kasxage. flocnegHee 6uro o6HapyxeHo 

A. C.3epHoBbrM B 2001 r. n AnmepoHcKoM p-He KpacuoAapcxoro r(pafl 

(3epnon, 2003). flpou3pacraHue Ha Kasrase (n Kolxl,rge) S. colchicus 

npeAcraBJrrer s 6oaluroft cretreHu ealaAoqHnfi Seuoueu. 

ll. S. cyperinus (L.) Kunth, 1837, Enum. Pl.2: 170; Lenski, 1985, 

Gritting. Flor. Rundbr, 19, I :43; Whittemore a. Schuyler, 2002,inFL. 

North Amer. 23:20. -- Eriophorum cyperinumL., 1762, Sp. Pl., ed.2 : 77. 

Onlrcau ur Cen. Auepuru (). 

Cupue u 6onorlrcrbre Mecra. - 3an. Eopona: fepuaHua (3ena;rx 

HuxHu CaKconux, rauocnoe). - O6q. pacrp.: Cen. Arraepuxa (rcpali- 

Htafi socror KaHa4u, [per,rMyulecrBeHHo BocroqHafl r{acrb CIIIA n Mexcr.rra). 

2n = 66 (Whittemore, Schuyler, 2002). 

O6uapyxen KaK 3aHocHoe pacreHlre Ha ceBepe fepuaHuu (Lenski, 

L c.). 

61

Sect.5. Filipes Ts. Tanget F. T. Wang, 1961, in Fl. Reip. Pop. Sin. 

ll : 221, I 4. -- Coqserr4e repMr,rHiurbHoe, 3oHrr.rKoBrlAHoe, ManoJryqeBoe, 

MHO|OKOJTOCKOBOe, OqeHb peAKO C HeMHOfrrMLr t4Jrld OAHT4M KOJTOCKOM. 

Kpororqr.rft lucr l, n 3-4 paza rpeBbrtuatou1nft coueerue. Ko.rrocxu :raHuerHbre, 

Kopr4rrHeBbre. Kpouqrae qeutyu trJlorHble, 6yuaxncrue, Ha BepxyuKe 

ocrpbre uJrr{ Tyrrbre, Hr,rxHke crepr4JrbHbre. OronoqeerHr,re ure- 

Tr,rHKr.r B rurcre 6, rrpruue, HeMHoro rlpeBbrtuarcuI,te rrrroA. Prrneq 3. 

fho4 rpexrpaHuufi. Pacrenue AepHr.rcroe, 6es nol:yuux KopHeBHr.rl. 

MoHorunsafl ceKrrr{r. 

T y p u s : 5../ilipes C. B. Clarke. 

12. S. ftlfues C. B. Clarke, 1903, Journ. Linn. Soc. London (Bot.) 36 : 

249;Ts. Tang et F. T. Wang, 1961, in Fl. Reip. Pop. Sin. ll : 14, tab. V, 

fig. | 7. 

OuucaH u: Bocr. Kuras (upon. @yu:rH); t y p u s :

Mr4 coqBerr{e. Konocxn co6pauu no (2) 4-10 s l-oJloBrrarble uyuru 7 

10 vrvr s Lvraw.,2.5 6 rvrrra llst., 2-3 MM ruup., uupoKo3JrilurrconAanbHbre 

(>),. rfiueBI{AHbIe I,IJTI{ llpoAoJlroBaro-SJIJIHrlcoI{AaJIb- 

Hbre, Ha BepxylxKe ryrrbre [Jrr,r 3aKpyrJIeHHbIe, oqeHb ryqrble, MHofouBeT- 

KoBbre. Kporculne qetuyn KopnrrHeBarrre, (1.2) 1.5-2 (2.5) MM AJr., rruporoaftqennAHbre 

r4JrH rtorrlh oKpyrJlble, Ha BepxylxKe 3aKpyrJIeHHbIe, c 

oqeHb KoporKr,rM ocrpl{eu, c I -5 xumanau. OronouserHble uIe'ruHKH B 

qrcre 0 -3 (6), 6enorarble, IIpflMbIe, o repxHefi IIoJIoBTHe c o6paqeuun- 

Mr,r BBepx peAKr.rMH ry6qauu, 6. u. panuue IIJIoAy. Puneq 2. fl.uoaut 

rrJrocKo-BbrrryKJrbte nJIn HepaBHo ABorIKoBblrryKJrbre, cxaTbre Aopcl{BeHTpaJrbHo, 

0,7-l urra Al., 0.5 0.7 rrarra urup. Pacreurq 50 200 cM BbIc., fyc- 

ToAepHr,rcrbre, 6e: rolsyr{ux KopHeB[ur. 

T u u : S. ternatanus Reinw. ex Miq. 

BnArr erofi cy6rponn.recxofi n rpo[uqecKofi cexuuu rrpexAe BKJTIoqarrr4cb 

B oAHy ceKrlr4ro BMecre c Br,rAaM[ rtluosofi ceKrruu, poAcrBeHHbrutr, 

S. sylvaticus, uru c BHAaMn poAcrBa S. asiaticus (= S. lushanensis) u 

S. Juirenoides, orHocr4MbrMr,r K ceKrlurr' Trichophorunt (Pers.) Darl. (n Hacroxuleft 

crarbe rrocJreAHr{e 2 sura orHeceHbr K ceKrlr.rf,M Psettdotri- 

'chophorum 

u Beetlea coorBercrBeHHo). Or BHAoB ceKur{r{ Scirpus lpegcraBnreJrr{ 

ceKqr{r4 Ternatani orJrnqalorct KopI4qHeBbrMu (a ue qepHoBa- 

TbrMrr r{Jrr,r 3eneHoBaro-qepnoBarbrMl,r, KpoMe S. atrovirens), oveHt tutor- 

HO pactloJroxeHHbrMr4 KpoloulrrMrr qeuryflMlr, Harr{queM Ha oKoJro(BeT- 

Hbrx ulerr{HKax BBepx (a He nHur) obpauleuuux :y6uou, a raK}I(e 2 pwtluaMr,r 

r4 rrJrocKo-BbrrryKJrbrMr4 lun cna6o ABorKoBbnryKJlt'rMH [JIoaaM14 (a 

ue 3 prulqaMr,r t.r TpexfpaHHbrMrr nJIoAaMIt, 3a ucKflroqeuneu S. microcarpus) 

u apeaJroM (nra4u ceKr1uu Scirpus He Bcrpeqamrcf, B Tporrr4Kax 

u cy6rponurax). Or [peAcraBuTenefi cerqnu Beetlea orucbrBaeMbrfi rarcoH 

orJrr4rraercr o6uruu repMr{HaJrbHbrM coqBerlren (a ue couBerl4eM, 

cocroflurlrM rrs TepMr.rHiurbHofo u flapuui]Jtbnbrx JrarepturbHmx couserr,lf) 

lr, KpoMe rofo, or srofi cexquu, a raK)r(e or ceKunu Pseudotri.chophorunt 

0 3 (6) 6elurvru rrpflMbrMlr oKoJlorrBerHhrMl4 uleruHraun,6. M. paBHblMH 

ruoAy (a ue acel4a 6, ropu.rHenbrMrr, H3Br4JrrcrbrMr4, s 2 4 paza AJrrrH- 

Hee llroAa), 2 (a ue 3) pu.rnuanau !r rrJlocKo-BbrrryKirbrMr4 riJru ABof,KoBbrnyKJrbrMH 

(a ue rpexrpauuurun) luo.rr,aMt,t. 

13. S. ternatanzs Reinw. ex Miq., 1856, Fl. Ind. Batav. 3 : 307; Kiik., 

1930, Acta Horti Gotoburg.5:36; id., 1944, Feddes Repert.53: l0l; 

Ohwi, 1944, Mem. Coll. Sci. Kyoto Univ., ser. 8., 18, I : 108; id., 1965, 

Fl. Jap.:203; Raymond, 1957, Nat. Canad.84,5:l15; T. Koyama, 

1958, Journ. Fac. Sci. Univ. Tokyo, sect.3 (Bot.) 7, 6:348, p. p., quoad 

yar. ternatanus;id.,1978, in Fl. Taiwan 5:203, tab. 1319; Ts. Tang et 

F. T. Wang, 1961, in FI. Reip. Pop. Sin. ll : 13, tab. IV, f,rg. 1,4; Kern, 

1974,in Steenis, Fl. Males., ser. 1,7,3:501; Eropona u H. K. Kxofi, 

1980, Hosocrr4 crrcr. Bbrclu. pacr. 17 : 54; Karthikeyan et a1., 1989, Fl. 

lnd. Enum. Monocot.:71; N. K. Khoi,2002,in Fl. Vietn.3:56. -. 

63

S. chinensis Munro, 1857, in Seem., Bot. Voy. Herald : 428, non Osbeck, 

1757;C. B. Clarke, 1893, in Hook. f., Fl. Brit. Ind,.6:662. 

OuncaH us I,IuAouesuu (MonyxrcKue o-Ba, o-n TepHare) (; t y p u s : U. 

Eonorucrrre Mecra, cblpbre Jreca, cpeAu KycrapHr4KoB, 6epera BoAHbrx 

uoroKoB; B ropax o6H.rHo Ha Bbrcore 800-2000 M HaA yp. M. Bocr'. 

Arnr: .{uoulra (o-n Krouo; apxurrerar Prcruo; o-sa Oracasapa (= o-na 

EoHun)). - IOxn. Acus: I,IHAur (fuualau u ceB.-Bocr. vacrr), lllpr.r- 

Jlasra. - IOro-Bocr. Acuq: Kraraft ([Oxn., BKJtK,qar o-s Tafrsanl). 

Brersau, Manafi:ur, L{uAose:ns (o-na Cyrraarpa, f.sa, Kannuaurau, 

Cynanecu, Molyxrcxue o-Ba, o-s Hosar feuner), @nlnunuuu. -.- 

O6u1. pacnp.: 3HAeMr{K. 

Cornacuo KepHy (Kern, l. c.), eror turpoKo pacrpocrpaHeHHnfi s 

Tporu{qecKofi o6racru, oxBarbrBaeMoft lrsgaHueu , 

Br,rA 3HaquTeJrbHo Bapbr4pyeT rro pa3MepaM KoJtocKoB 14 KpOrcUIr4X qeutyfr., 

a raKxe no qucry oKoJlouBerHbtx qeruHoK (0-3, .qo 6). flo aaH- 

Huu Klapxa (C. B. Clarke, 1893, l. c.), upn o6uraHuu ua 6onorax pacreHue 

.racro o6pasyer croJroHbt, r4, KpoMe roro, Ha3eMHbre, yKopeHrrou{uecs 

uo6eru, uoAo6no S. radicans. 

S. ternatanuJ rptrBoAtrrcfl, vr Arrfl, Fyrana (Noltie, 1994), uo, Bo3Mox- 

Ho, 3To yKa3aHne orHocrlrcr rc S. rosthornir (crra. Hraxe). 

14. S. chunianrs Ts.Tang et F. T. Wang, 1961, in Fl. Reip. Pop. Sin. 

ll :221,13, tab. [V, hg. 5,8. 

Ouucau us IOxH. Knrar (o-n XaftHaub); t y p u s :

npor. Xy6efi; [Oro-3au.: rpoB. CuvyaHr u lOnuraHs), .fnoHur (o-n 

Krocro). - IOxu. Arur: I4H4ua, ?Heual, EyraH. IOro-Bocr. Arnr: 

Knrafi (IOxH.: npon fyarqyH).- O6ur. pacnp.: DHAeMUK. 

A66pernarypa (B>, [pr,lBeAeHHar B rrporoJrore [ocJre yKa3aHr4r Mecra 

c6opa rurroBoro o6pasqa, o3Haqaer (Diels, 1901, L c. : 184), qro o6pa3ubr 

c HoMepaMH 1500-2458 co6lrpalncr c KoHua v^afl, ytru HaqaJra 

I4IOH' AO KOHUA HIOJI'. 

S. kiushuensl's Ohwi, l. c., ouucaHHufi c o-sa Krccro (.fluoHux) tr cqrdraauruirc;r 

ero 3HAeMrrKoM, rro3AHee 6nn oroxAecrBJreH c S. rosthornii 

caMr4M Onra (Ohwi, 1965,1. c.). Koarua (T. Koyama, 1958, l. c. : 349) paccMarprrBrur 

S. kiushuensls n xaqecrse pa3HoBr,rAHocru ^t. ternatsnus-- 

S. ternatanus var. kiushuensrs (Ohwi) T. Koyama, rroMecruB 3ro pacre- 

HLIe B ceKIII{to Scirpus HapflAy c S. sylvaticus, S. radicans n S. atrovirens. 

S. rosthornir, raK xe, xar u Pefiuou4 (Raymond, 1957, L c.), Korua 

(T. Koyama, 1958, l. c.: 291) ouru6o'{Ho cerlrrar BrdAoM, poAcrBeHHbrM 

S. paniculatocorymbosus Kiik. u S. petelotii R. Gross. Bce lru 3 vuga, 

orHeceHHbre PefiuoHAona K ceKrlr.rr4 Paniculatocorymbosi Ktik., Koarvra 

BKnroqu[ z ptg Androcorna (Nees) T. Koyama ceKrrrlr,r Trichophorum 

(Pers.) Darl. C Ha3BaHHbtM prAorr,r oH ctrHoHtrMu3r,rpoBrur ceKutrrc Paniculatocorymbosf. 

Anropn (Tang, Wang, 196l) arlro.rutu 

S. rosthornii, S. ternatanus u S. chunianu.r Ts. Tang et F. T. Wang n 

ceKlulo Trichophorum BMecre c S. karuisawensis Makino u S. asiaticus 

Beetle. flpnHrraa uMr{ ceKqufl Paniculatocorymbosi BKJuoqaer roJrbKo 

S. p anic ulat oc orymbo sus Kiik. 

Bo

xftqenuAnue uJIrI upoAoJlrosaro-rftueBl{AHble. Kpotoulrae qeuryl4 qepHo- 

Barbre rnra reMHo-cepue, 3.5-4 (5) rrlrrl An., 1.5-1.8 MM ullp' OKoloqser- 

Hbre xlerlrHKll B trl,Iclte 6, 6ellte, I,I3Bl,uurcrbte, BBepxy uepoxoBarble or 

BBepx HarrpaBJreHHbrx sy6qos, n 2 3 pasa AJIHHHee rIJroAa. Puleu 3. 

flloAu rpexrpaHHble, npoAoirroaaro-o6paruorfiuesl'rAuHe, (1.3) 1.5 1.8 

(2) rr.rna .qn., 0.8-0.9 MM url{p. Pacreuus ao 50 crra BbIc', fycroAepHucl'ble, 

6e: nonsy.rux KopHeBuu-{; cre6nerutx flucrres 2 3. 

T y p u s: S. maximowiczii C. B. Clarke (= Eriophorum iaponicunt 

Maxim.). 

16. S. maximowiczii C. B. Clarke, 1908, Kew Bull. Add. ser. 8 : 30; 

Ohwi, 1944, Mem. Coll. Sci. Kyoto Univ. ser. B, 18, I : 103; id., 1965, 

Fl. Jap. :202 T. Koyama, 1958, Journ. Fac. Sci. Univ. Tokyo, sect.3 

(Bot.) 7,6:350 et272, fig. l; tOpues, 1965, Erorn. Mocr. o6ur. t'tctt. 

rrpnp., orA. 6Iror.70, I : 132;on xe, 1966, r Aprr. t[1. CCCP 3 : 30; 

Bopour., 1966, Ol. con.,{alru. Bocr.:79;ou xe, 1982, Oupe4. pacr. 

cos. AarrH. Bocr. : 99; Y. L. Chang et Y. L. Yan5,1976, in Fl. Pl. Herb. 

Chin. Bor.-Or. 1l :20; Y.T. Lee, 1976,in Fl. Cor.7:267; A. E. KoxeBHr.rKoB, 

1988, n Cocya. pacr. coB. .{anrH. Bocr. 3 : l8l, xapra 57; 

Tr.rlroxuua, 1990, so @n. Cu6. 3 : 20, rapra 19. -- Eriophorum japonicum 

Maxim., 1886, Bull. Acad. Sci. P6tersb.3l, I : lll; id., 1886, Mel. Biol. 

12:558; IOs., 1935, so (Dn. CCCP 3'.28,ra6r.2, pr.rc. l;Kitag., 1939, 

Rep. Inst. Sci. Res. Manch.3, App. I (Lineam. Fl. Mansh.): l16; id., 

1979, Neo-Lineam. Fl. Mansh. : 149; Ts. Tang et F. T. Wang, 1961, in 

Fl. Reip. Pop. Sin. l1 : 36, tab. XV, fig. 13-18. Scirpus iaponicus 

(Maxim.) Fern., 1905, Rhodora 7 : 130, in annot., non Franch. et Savat., 

1879. - Eriophorum maximowiczt, (C. B. Clarke) Beetle, 1946, Amer. 

Journ. Bot. 33,8 : 663. -- Maximoviczia japonica (Maxim.) A. P. Khokhr., 

1985,

3ee-Eyp. (pegro), Vccyp., Cax. (cea.), Kypnn. (oxu.). - Bocr. Arun: 

Kurafi (Cea.-Bocr.), u-on Kopet, fliottut (o-r XorrafiAo u ceBep o-Ba 

Xoncro). - O6ur. pacrp.: oHAeMnK. -2n= 64 (KoxenHr,rror, 1988). 

flpnregeHuHft sI,IA 6ul ueprouaqaJlbHo orl{caH rcax Eriophorum 

japonicum Maxim., L c., uo 3areM rlepeHeceH B pog Scirpus c o6pasona- 

Hueu r

(a He o6uuuo MHororrllc[eHHrtx) KoJrocKoB, Kpoloque rl{crbf,, qacro pe- 

AyurrpoBaHHble Ao qepHblx qeuryeBl,I,qHblx Braf€LIII{uL IUlacTuHKa Hu)KHefo 

KpoK,Ulero Jrrrrcra (ecnu oHa pasar,rra) o6blquo Kopoqe cousernq (a He 

qacro rpeBbruraer ero),2 3 (a ne 3-10 u 6olee) cre6nesHxJrl{cra, Bepxnuft 

us Koropbrx nHorAa c 6. u. yKopoqeHHoir uxu [oJlHocrblo peAyquporauuofi 

unacrunroft, 6olee KpyrIHbIe, 3.5-4 (5) MM AJI., Kporcque 

qeryr4 (a ue l-2.5, peAKo 3 rvru) u 6olee xpyuHue, 1.5-1.8 (2) vrrvr 4n., 

rrJroAbr (y ocranrHux Br.rAoB Scirpus nJIoAbI o6mqHo 0.6-1.2 MM AJI., LI 

Jurrub r4HorAa, Ha[pr,rMep,y S. microcarpus J. et C. Presl u S. expansus 

Fern., oull Bapbr4pyror B rrpeAeJrax 0.7-1.6 una.qn.). Or scex rpeAcraBl4reteit 

ScirpuJ paccMarpr4saeMHfi Br4A orluqaercx oco6eHHocrrMu reorpaQuuecxoro 

pac[pocrpaHeHrrr u gxororuefi. S. maximowiczii - zro 

eAuHcrBeHHnft n po4e Scirpus BHA, rrporr3pacrarcullrft B BbrcoKoropbflx 

(o6urvHo ua ronrqax) yMepeHHoro rrorca r4 rrpoHlrKaloquft n apruruecKr{e 

uuporu (9yxorxa). 

CusrervraruqecKoe [oJroxesne S. maximowiczii 6uno [peAMeroM cueurraJrbHoro 

o6cyxgeum prAoM aBropoB (Beetle, 1946; Koyama, 1958; 

IOpuea, 1965, 1966; Hosocerona,1994). Croeo6pasue 3roro Br,rAa rocJryxrrJro 

ocHoBaHheM AJU BbrAe[eHr,rr ero B ceKrlr{rc Japonici T. Koyama 

(Koyama, 1958) r,r rroBropHo B ceKrlr,rrc Pseudodriophorum Jurtz. () (IOpqes, 1965). O6e ceKur4r.r ycraHoBneHbr B cocraBe poAa 

Scirpus. He6e:uHrepecHo yrloMflHyrb, qro KosIvIa (Koyama, 1958) cuurar 

S. maximowiczii npororuuov^ Eriophorum. Cexttus. Japonici 6una 

Bo3BeAeHa B paHr rroApoAa B cocraBe poga Eriophorum (Oteng-Yeboah, 

1974),n BcKope n B cocraBe poga Scirpus (Chang, in Chang, Yang, 1976), 

qro rBlrJrocb orpaxeHueM rrpeAcraBJreHr,rft o naop$onourqecrofi o6oco6- 

JreHHocrr4 S. maximowiczii n o6 ero uercuofi pogorofi upuHalure)KHocrr,r. 

A. fI. Xoxpf,KoBhrM (1985) S. maximowiczii 6am BbrAereH B orAerrbuufi 

pog Maximoviczia A.P. Khokhr., o6uapo4onaHHuft uyreu ccburKr{ 

Ha ornrcaHr{e ceKrlurr Japonici T. Koyama. Apryueuror B rroJrb3y caMocrorreJrbHocrn 

3Toro poAa r{M ne 6uno [pr.rBeAeHo. Hasnauue Maxintoviczia 

oxasutocb He3aKoHHbrM KaK 6onee nol4unfi ouonuu (cu. ruure), 

B cBfl3rd c qeu 6uno 3aMeHeHo sa Maximowicziella A. P. Khokhr. (pro 

nom. nov.) (Xoxprrcon, 1989). Caenannaa XoxprKoBbrM xou6uuaqur 

Maximowicziella japonica (Maxim.) A. P. Khokhr., rpeAcraBlrer co6ofi 

nomen invalidum, [ocKoJrbKy rrpu ee o6Hapo4oraHr.ru He 6ul yxaraH 

6asr4oHr4rvr. Anropou, BaJrrrAr{3trpoBaBrrrtrM sro Ha3BaHue, cleAyer cqurarb 

rrpu3HaBrrryrc poA Maximowicziella M. C. Hoeocdnory (Novosselova, 

1994, l. c.; Hoaocenona, 2001, 2004), roropa"r rpr,rBeJra B HoMeHKrarypuoft 

qurare Maximowicziella japonica (Maxim.) A. P. Khokhr. ero 

6asuouuM - Eriophorum japonicum Maxim. 

Ha uoft B3rJuA, rrprrBeAeHHbre Bbr[re uopSolorn.recKr.re [pu3HaKu, 

orlnrrarcqr4e S. maximowiczii or Apyrr,rx fipeAcraBure:neir Scirpus, ue 

r4MeIOT [plrHr.Ir4[r4aJIbHOfO XapaKTepa I1I He MoryT cJryxuTb ocHoBaH[eM 

68

AJrr BbrAeJreHr,rr 3Toro Br,rAa B orAenuuft pog. Cnoeo6pasne S. maximowiczii 

otITr,IMiLrrbHo orpzDKaercr rrpr{ paccMorpeHr{r,r ero B cocraBe orgenrHofi 

ceKrlr,rn B [peAeJrax poga Scirpus, Henrsr corJracnTbcr c [peAcraBJreHr,reM 

prAa aBropor (nanpuuep, Koyama, 1958; Koxenuurcon, 

2001) o roM, rrro S. maximowiczii zalr'uv'aer rrpoMexyrorrHoe [onoxe- 

Hr{e MexAy poAaMr4 Scirpus u Etiophorum. C uocne4HuM poAoM S. maximowiczii 

HMeer roJrbKo HeKoropoe ra6urya.llHoe cxoAcrBo, o .reM 6uno 

cKa3aHo Bbrrue. .fl nonHocrrIo pa3AeJuro MHeHr{e E. A. topqena (1965), 

crrrlraloulero S. maximowiczii gpennuM BbrcoKoropurru (rollqonuu) 

rrporr3BoAHbrM or HeBbrcoKoropHoro rurpo$ulruoro rr4rra Scirpus. 

K erorvry MoxHo 4o6aanrl, qro o ApeBHocrI{, a raKxe u o [pHMr,rrLrB- 

Hocrr4 B[Aa cBr,rAereJrbcrByer Hiuruqr4e y orAeJrbHbtx ero oco6eft nouu- 

MO TepMI,IHaJrbHOfO COIIBeTIT', TaKXe U JlaTepiurbHofo, pacrroJtoxeHHoro 

B rra3yxe BepxHero cre6nesoro Jlrlcra. JlarepanlHtre courernr csoficrBeHHbr 

H HeKorophrM Apyrr{M, rrprrMrrruBHbrM, BHAaM Scirpus (cu. rouueurapufi 

K ceKIIr{u Paniculatocorymbosi), u, HecoMHeHHo, HX r4MeJrr,r aHrlecrpairbHbre 

Qoprvrm Scirpus, y Koropbrx o6ulee cor.lBerr{e 3aHtrMaJro 

Becb perrpoAyrrunHufi no6er. 

Sect. 8. Scirpus; T. Koyama, 1958, Journ. Fac. Sci. Univ. Tokyo, sect. 

3 (Bot.) 7,6 :292, p. p., excl. S. ternatano; Ts. Tang et F.T. Wang, 1961, 

in Fl. Reip. Pop. Sin. ll :9; Kern, 1974,in Steenis, Fl. Males. ser. 1,7, 

3 : 501, quoad nom.; DeFilipp;, 1980, in Fl. Europ.5:277; A. E. KoxeBHuKoB, 

1988, s Cocya, pacr. coB. .{alnu. Bocr. 3: 178. -- Seidlia 

Opiz, 1826, Naturalientausch I | : 349. - Scirpus sect. Taphrogeton 

Reichenb., 1830, Fl. Germ. Exc.:79; Poxen., 1935, so Ol. CCCP 3 : 44, 

p. p.; Beetle,1947, in North Amer. Fl. 18,8 :486; Y. L. Chang et 

Y.L.Yang, 1976,in Fl. Pl. Herb. Chin. Bor.-Or. ll :5. - S. sect. 

Euscirpus Griseb., 1844, Spicil. Fl. Rumel. 2:417, p. p.; Benth. et Hook. 

f., 1883, Gen. Pl. 3 : 1051. - Nemocharus Beurl.. 1853. Bot. Not. : 53. - 

Taphrogeton (Reichenb.) Montandon, 1856, Fl. Jura Sept. : 346. - 

Scirpus Rotte Phyllothryon Ddll, 1857, Fl. Bad. : 305, p. p., excl. 

S. maritimo. * Scirpus a. Phyllantheli Nyman, 1882, Consp. Fl. Eur. : 

763, nom. nud., p. p., excl. S. maritimo. - S. subgen. Euscirpus pax, 

1888, in Engl. u. Prantl, Nat. Pflanzenfam.2,2.. llZ, p.p. --' S. sect. 

Sylvaticae C. B. Clarke, 1893, in Hook. f., Fl. Brit. Ind. 6 : 661, quoad 

nom. - S. subgen. Taphrogeton (Reichenb.) Ohwi, 1944, Mem. Coll. Sci. 

Kyoto LJniv., ser. B, 18, I : 95, p. p. - S. subgen. Scirpus: Eropoua, 

1967, s Pacr. I{enrp. Asuu 3 : 15; oHa xe, 1976, so rDr. Erpou. .racru 

CCCP 2 : 89. - O6uee corlBerue repMuHaJrbHoe, MHoroKoJrocKoBoe, 

c KoirocKaMr,r, pacloJroxeHHbrMr{ Ha BerorrKax corlBerr4r uo l_3 (5) r.rnu 

co6paHnuuu no 8-25 (-30) n rrJrorHbre rxapoBrqHbre r{Jrr{ rroJryuapoBr4.q- 

Hbre roJroBKu; KoJrocKlr 2.5-8 (-10) MM AJr., 1.3,1.1 MM rrrlrp., sfi\esup- 

Hbre r,rJrn naHrreroBtrAHue, Kporc[1ue qeulyfi qepHoBarbre r,uu,r qepHoBa_ 

ro-3eJreHoBarbre, peAKo Kopr4qHeBhre (5. atrovirens), 6. v. rfiqesu.qutre. 

69

1,4-2 vtla AJr., Ha BepxylxKe ryrlble, 3axpyfJleHHbre, ocrpoBarbre UJIII 

c KoporKrrM ocrprreM. OxonoqeerNble uIerLtHKI4 B tlllcrre 6, pexe (0) 3-4, 

6enorarue, c o6paulenublMl4 BHI{3:y6qauu, trpflMble, HeMHoro Kopoqe 

fIJIoAa, 6. u, parnue eMy vJILI HeMHoro AJII4HHee, rla6o u:sl4ilI{crble 

(5. radicans), r 2-3 pa3a ATIHHHee rIJIoAa. Pureu 3, pegxo 2 (5. microcarpus). 

flloAn rpexrpaHHbre, lu6o rlJlocKo-BbrnyKJrbte [JII4 cJla6o ABorxoBbrrryKJrbre, 

(0.7) | 1.2 (1.6) MM AJr. PacreHus c tlor3yql4Mrl KopHe- 

BrruraMrr utu 6ez swx. 

Typus:generistypus. 

17. S. radicans Schkuhr, 1793, Ann. Bot. (Usteri), 2, 4 '. 49, tab. l; 

Poxen., 1935, so (Dn. CCCP 3:45; Ohwi, 1944, Mem. Coll, Sci. Kyoto 

Univ., ser. B, 18, I : 102; id., 1965, Fl. Jap. : 202; T. Koyama, 1958, 

Journ. Fac. Sci. Univ. Tokyo, sect. 3 (Bot.) 7, 6 : 345; Ts. Tang et 

F. T. Wang, 1961, in Fl. Reip. Pop, Sin. ll: l0; Eropooa, 1976, so @r. 

Enpon. .racru CCCP 2:89; Y. L. Chang et Y. L. Yang,1976, in Fl. Pl. 

Herb. Chin. Bor.-Or. 11 : 5; Kitag.,1979, Neo-Lineam. Fl. Mansh. : 155; 

DeFilipps, 1980, in Fl. Europ. 5 : 278; fpy6on, 1982, Oupe4. cocyA. 

pacr. Monr. :51;A. E. Koxensuxoa, 1988, n Cocya.pacr. coB,,{anrH. 

Bocr. 3 : 179, xapra 57; Tranaoxr.ru a, 1990, so Or. Cu6. 3 : 20, xapra 2l ; 

Opnora, 1993, Tp. C.-flerep6. o6q. ecrecrBol4crl. 77, 3 (KoHcu. Sl. 

BonoroAcx. o6n.) : 39; fy6auon, 1996, Koncn. $n. BueurH. MoHl. : 

29.- S. sylvaticus L. var. radicans (Schkuhr) Willd., 1798, Sp. Pl. I : 

308. - Seidlia radicans (Schkuhr) Opiz, Naturalientaush ll: 349. --- 

Nemocharis radicans (Schkuhr) Beurl., 1853, Bot. Not.: 53. - Scirpus 

hokkaidoensis Beetle, 1946, Amer. Journ. Bot. 33, 8 :662. 

Onucan u: fepuauur{ (p. 3rr6a) (); 

I e c t o t y p u s (Pignotti, 2003 : 304): [Icon], Schkuhr, l. c., tab. 1. 

Eepera BoAoeMoB, TpaBrHbre 6orota, KaHaBbr, 6olorncrue lyra, 

cbrpbre Jreca. - 3au. Enpoua: ror Hoprerhn, Lllsequw n cDun:lrrtgnu 

(nes4e oqeHb peAoK), (DpaHrlna, crpaHbr l{enrp. Erportrr, llrzutna (cen.), 

6. IOrocnasru. - Poccus. Eepon. qacmbi Kap.-Myptr.t. (roxH. Kapelna: 

cen. flpula4oxbe - oKp. [oc. Kyprudru, Llurrunaxru n Canrur,r), .{,a.fleu. 

(BonoroAcKar o6n. - Bureropcrufi, 9epenoregxuit u Boloro4crufi 

p-uu), JIaA. -I,fu nu., Bepx.-,{Henp., Bepx. -Bolx., Bolx. -Karvr. (TarapcraH 

- pafiou Ka3aHu, gep. Pyrr

Bocr.), n-on Koper, fl,tolaut (o-ra Xorxaitr,o u XoHcro). - O6ul. 

pacrp.: sHAeMr4K. --2n= 56, 58 (Pignotti, 2003). 

Yra:anHrre Bbrure MecroHaxo)I(AeHut S. radicans a Kapenuu tBJrtK)Tcfl 

caMbrMr4 ceBepHbrMr,r B apeaJre BrrAa Ha reppr,rropllr,r Enpouefrcroft 

Poccull. .{na us nux (Kypxl,t6rn n I4uuulaxru) [puBeAeHbI tro Jrl4reparypHbrM 

AaHHbrM (Paueucraa, 1960; Heikkilii et al., 1999), rperre - oKp. 

uoc. Canuu (; o6pa:eq xpaHhrcr n fep6apur,r Hucruryra 6uonoruu 

Kapenrcrcoro HI{ PAH), Ha ocHoBaur,ru ycrHoro coo6urenns A. B. KpanqeHKo. 

.Ilnx Kanuuuurpa4cxofi o6n. (reppuropnr 6Hsruefi Bocr. flpyccr.ru) 

S. radicans rpnBoAr,urc.f, A6porvrafirorvr (Abromeit et al., 1940). 

B nocneAnnee BpeMr Br,rA 3Aecb He HaxoArrJrr4 (I4. IO. fy6apena, ycrHoe 

coo6ureHue). 

Kosr\aofi (T. Koyama, 1958, l. c.) 6mno orMeqeHo, uro y enpoueficrux 

o6pa:qos S. radicans Jryqu couBerufl coBeptueHHo rJraAKue, B To 

BpeMr KaK y a3uarcKr,rx pacreHuft osu qacro uepoxoBarbre. Llsy.renne 

MaTeprraJra [oKa3aJlo, rtTo y [ocJreAHLIx Jryrilt TaKxe rJIaAKr{e. Boavrox- 

Ho, 3a S. radicsns 6unn npuHrrbr BHeruHe orqacrr,r cxoAHbre c nuu o6pa3qbl 

S. orientalis, y Koroporo Jryqu r repxuefi rroJroBr,rHe BcerAa tuepoxoBarbre 

(cu. rarxe KoMMeHraprrfi x erouy suly). 

17. S. orientalis Ohwi, 1932, Acta Phytotax. Geobot. (Kyoto) | :76; 

id.,1944, Mem. Coll. Sci. Kyoto Univ., ser. B, 18, I : l0l; Kitag.,1939, 

Rep. Inst. Sci. Res. Manch. 3, App. I (Lineam. Fl. Mansh.) : 123; fpy- 

6os, 1955, KoHcu. Qn. MoHr. HapogH. Pecn.: 79; oH xe, 1982, Onpeg. 

cocyA. pacr. Mour. : 5l; Bopour.,1966, @r. cos. ,{anrH. Bocr. : 79; oH xe, 

1982, Onpe.q. pacr. con. .(alrH. Bour.; Erop. , 1967, n Pacr. I-leHrp. A:rara 

3 : 16; Y. L. Chang et Y. L, Yang, 1976, Fl. Pl. Herb. Chin. Bor.-Or. I I : 

8, fig. 3, I -4; M. Zhan a. Y. C. Yang, 1987, Acta Biol. Plateau Sin. 7 : 

12; A. E. KoxesHrron, 1988, n CocyA. pacr. coB. .(anlu. Bocr. 3 : 180, 

xapra 57; Tlur,roxuHa, 1990, so @r. Cu6. 3 :20, rapra 20; fy6anor, 1996, 

KoHcu. Qn. Bueutr. Monr. :29. - S. sylvaticus L. var. maximowiczii 

Regel, 1861, M6m. Acad. Sci. P6tersb. (Sci. Phys.-Math,), s6r. 7,4,4 

(Tent. Fl. Ussur.) : 16l;T. Koyama, 1958, Journ. Fac. Sci. Univ. Tokyo, 

sect. 3 (Bot.) 7, 6:346; Ts. Tang et F. T. Wang, 1961, in Fl. Reip. Pop. 

Sin. 1l :9; Ohwi, 1965, Fl. Jap.:202; Kitag., 1979, Neo-Lineam. Fl. 

Mansh. : 155; T.B. Lee, 1999, Ill. Fl. Kor. : l7l. - S. sylvaticus var. 

subradicans Kiik. exTs.Tang, 1932, Bull. Fan.Inst. Biol.3: 136, 150. 

S. sylvaticus subsp. maximowicnl (Regel) N. R. Cui, 1982, in Clavis Pl. 

Xinjian. l:289. - S. sylvaticus auct. non L. : C. B. Clarke, 1903, Journ. 

Linn. Soc. London (Bot.) 36 : 254; Ktik., 1904, Bot. Jahrb. 34, Beibl., 75 : 

21; Poxen,, 1935, so

MelxosoAtr u 6epera BoAoeMoB, sa6olo.reHHbre Jryra. ,* poccnr. 

3an. Cu6upu: Hpr. (roro-aocr.), Astt; Bocm. Cu6upo: Euuc. (rcro-3arr. ,- 

6acc. uuxn. TeqeHHr AHrapu), JIeH.-Kol. (ror, oveHt pegxo), Anl.- 

Canu., Aayp.; ,\aaau. Bocmox:3ee-Eyp., Vccyp., Cax. - I{enrp. Arur: 

MoHronzs (XeHr,, XaHr., Monr.-!ayp., flpuxuHr., Bocr. Monr.). _- 

Bocr. Asrr: Knrafi (Cen.-3au., BHyrp. Mourolzr, Cen.-Bocr.; Cen.: 

upoa. Xs6sft ulllauscw; Bocr,: npor. IIIaurqyH; ? LleHrp.), n-oa Koper, 

flno:uut (o-na XoxrcaftAo u Xoucro). --- O6u1. pacnp.: 3HAeMrdK. -- 2n - 

60 (Koxeanr.rxos, 1988). 

Onu (Ohwi, 1932, L c.) npn o6uapo4oranuu S. orientalis B KarrecrBe 

HOBO|O Br,rAa [pr{BeJr B ctrHoHnuax s. sylvaticus L. var. maximowiczii 

Regel, L c. flocneAnnfi rarcos 6trn onucas c poccr,rficxoro ,{altHer.o 

Bocroxa (typus: , rIocKoJIbKy HeJIb3t 6ruo so:aecru ptBHoBI,rAsocrr 

3. JL Peren B paHf B:aAav3sa cyutrecrBoBaHbfl,Ha3BaH]/,t s. maximowiczii 

C. B. Clarke 1908 (= Eriophorum japonicum Maxim.), rorAa rr4uou 

.S. orientalis cJreAyer cqurarb qr,rrr.rpoBaHHtrft nnue o6pareq, Ha Ko_ 

ropoM 6ruro ocHosaHo o[ncaHr4e s. sylvaticus var. maximowicziiRegel. 

s. orientalis 6luror x s. sylvaticu.r ll HeKoropbrMu aBTopaMr4 rrpr4Hr.r- 

MaeTct B KaqecTBe efo pa3HoBr,rAHOCTr4 I,IJrI IIOABLIAa, B Orlrtaque or 

s- sylvaticus, KorrocKrr y s. orientalls pacuoloxeHbr Ha BeroqKax couBe- 

Tt4fl. rro oAHoMy, HrItr HApfl,\y c oAr,rHorrHbrMlr KoJrocKaMr4, KOTOpbte [pe_ 

o6la4ator, r{MeK)Tcr [yrrKr4 uz 2_3 KoJrocKoB; KoJrocKI{ o6HqHo losoru- 

Ho pe3Ko cyxeHHble IIo HarIpaBJIeHI'IIo K BepxyIUKe, ocrpoBarble vJru 3aocrpeHHbre, 

KpoIo[IUe qeurylr ocrpoBarbre; ryrru corlBerr4r s Hr.rxnefi qa_ 

cru o6rrqno rJraAKHe. Y s. sylvaticzJ KoJrocKr.r cKyqeHbr no 2-5, Ho He- 

KOTOPbIC I,I3 HI{X MO|YT 6ITrr OAUUOTIHbIMI; KOJIOCKII HA BEPXYIUKC TYrrbre, 

Kporcque qeulyu Tyrrbre, 3aKpyrJreHHble llnu yceqeHHbre; Jryr{fl co_ 

IIBETI,Ifl IIICPOXOBATbIE NO SCCfi AJII{HC. fIO ArrUrrC OKOJIOUBCTHbIX UIETI{. 

HOK paccMarpr{BaeMbre BuA,bI, Ha vrofr nsIJIag, He pa3ll4qarorcf,, Ho no 

MH,eHr,rKr Koslrn (Koyama, 1958), y S. orientalis qerrdHKr{ Kopoqe, qeM 

y s. sylvaticzs. Y o6oux BnAoB ouu o6lrqHo HeMHoro AJrr,rHHee [JroAa, 

XOTS OTAETbHbIC IqCTHHKI,I MOTYT 6UTb TOPOVE. HECUOTPT HA TIPI{BEA,CH- 

Hbre orJrr,rqar, roqHaf, uAeHrlrsurcaqral s. sylvaticus tt s. orientalis ne 

Bcef.{a Bo3MOXHa. 

B 3au. Cu6upu, rAe BrrepBbre 6Hn orlaeqen C. A. Tzuoxuuofi 1qur. 

cov.), S. orientalis oqeHb peAoK. B LE uMerorcfl ero c6opu c Alrar. 

s. orientalis BHerrrHe Moxer 6url cxoAeu c s. radicais, or Koroporo 

orJrrrqaercr ruepoxoBarbruu (a ue rna4ruuu) fiyrraMl{ coqnetur, 6onee 

MeJIKITMTI' 3.5-5 nrrra 4n. (a ne 5-8 uur), KoJrosKaMr4, ocropoBarblMr,r Ha 

BepxyrxKe KporoulrrMlr qeuyrMr,r c 3 xunraun (a ue ryrurr,fl,r r{Jur 3aKpyr- 

JreHHbrMI4' C I XUnXOft) r{ [p{MbrMrr (a He usaranucrbrMr4 oKoJrouBerHbruu 

ulerunrauu). 

72

19. S. sylvaticus L.,1753, Sp. Pl. : 51; Poxen., 1935, no Or. CCCP 

3 : 44, p. p.; fpoccr., 1940, @n. Kam., us1. 2,2 : 14; Eropona, 1967, n 

Pacr. I{eHrp. Asuu 3 : 16; oHa xe, 1976a, so (Dn. Eapou. .racru CCCP 

2:89; oHa xe, 19766, n Oupe4. pacr. Cpe4H. Azuu 5 : l0; ona xe, 1991, 

Hosocua cr.rcr. Bbrcru. pavr. 28:6; DeFilipps, 1980, in Fl. Europ . 5 : 277; 

Kit Tan, 1985, in Davis, Fl. Turk. 9 : 61, p. p., excl. pl. ex As. Or. et 

Amer. Bor.; TnuoxnHa, 1990, so @r. Cu6.3: 21, rapra 22;Xaagxtu, 

2001, so

xopouro cofnacyercr c pegroil BsrpeqaeMocrlrc s. hattorianus s ArtoHun. 

Cornacuo AaHHbrM Kosltrt (T. Koyama, l. c.) ra Onn (Ohwi, l. c.), uocneAuufr 

BlrA, oroxAecrsleHHHil Ha3BaHHbIMlI aBTopaM[ c S. atrovirens 

var. georgianas (Harper) Fern., u:necreH s -gnoHI,II4 ToJtbKo u3 eAI4HI.ttt- 

Hbrx rryHKToB B rrpoB. lwaki, rAe 6unu co6pauu u rllrloBble o6paruu. 

flo Aauurura aMeprlKaHcKnx uccJreAoBarelefi (Schuyler, 1967; Whittemore, 

Schuyler,2002) S. hattorianus u S. georgianus R. M. Harper or- 

JruqaroTcr l1o xapaKTepy oKonouBeTHblx uIerI4HoK 14 oKpacKe KpoIouIHx 

ueuryfi: y [epBofo Br.rAa urerr.rHKu B rtHcJIe (4) 5-6, a nepxnefi qacrl{ c 

nuus o6pauqeHHbrMrr sy6uauu, Kopoqe rrJIoAa urrla paBHbIe eMy, a Kpotou1ue 

qeruyu o6rrqHo qepHoBarbre, y BToporo - ulerItHKI{ orcyrcrByror 

rrJur oHH pyAr4MeHTapHble u rJraAKlle, u3peAKa HaBepxy c sy6qaun u 

rroqru paBHbre ruroAy, Ho B rl.rcrte He 6oree 3, rporoqne xe ueurya (u xo- 

JIOCKI4 n qenOrt{) KOpHrrHeBaTbte, co c.lra6o rupaxeHHofi .repuofr [r4fMeHra:Jleit. 

21. S. atrovirens Willd., 1809, Enum. Pl. Horti Berol. I :79:id.,1812, 

Hortus Berol. 2 : 99, tab. 99; DeFilipps, 1980, in Fl. Europ.5:278; 

Whittemore a. Schuyler, 2002,in Fl. North Amer. 23: 15; Pignotti, 2003, 

Webbia 58, 2 : 307. Seidlia atrovirens Opiz, 1826, Naturalientausch 

ll : 349. 

OnncaH us Ces. Auepnxu (). T y p u s : 

?B. 

Crrpue u 6olorucrne Mecra. -- 3an. Enpona: 3aHocHoe Ha ceBepo- 

BocroKe (DpaHqnn, ceBepe Mranwn (flreuoHr), n fepuaHulr r,r lllueffuapvrr4. 

O6rq. pacnp.: Cer. Auepnra (xpaftuufi roro-BocroK KanaAn u 

rperlMyulecrBeHHo BocrouHlre paftouu CIIIA). 2n = 50 60, 50, 56,62 

(Whittemore, Schuyler, 2002; Pignotti, 2003). 

B pa6ore fluruorru (Pignotti, 2003 : 307) rporlr4rr4poBaH 4pyrofi uc- 

TorrHuK rrepBoorlncanuq S. atrovirens:

pacr. coB. .{alrn. Bocr. : 100; Schuyler,1967, Acad. Nat. Philadelph. 

119,6:315; Eropora, r 1981, Oupe.q. cocyA. pacr. Kauv. o6n.:301; 

A. E. KoxesHuKoB, 1988, s Cocy4. pacr. coB.,{urn. Bocr. 3 : 178, xapra 

57; Whittemore a. Schuyler, 2002, in Fl. North Amer. 23 : 18. 

S.avatshensis Kom., 1914, Feddes Repert. l3 : 163;Poxet., 1935, no @n. 

CCCP 3 :46 (). 

Ouucau H3 roro-3arr. KanaAu (o-n Bauryrep, 3aJluB Hyrra) (; typ u s:PR. 

flec.raHo-r4nlrcrbre 6epera peK, pyqbeB, crapr{rrbt. -, Poccuq. fiamu. 

Bocmorc: u-os KaN4qarKa. - O6u-1. pacnp.: Ces. Auepura (nueaprrnvecrue 

paftonn Kauagu, CIIIA; Mercuxa: Eafia Kulr$opHrla). - 2n = 64, 

66 (Whittemore, Schuyler, 2002). 

S. microcarpr.s - eAflHcrneHnrrft BI4A, He crlurafl 3aHocHblx Ic Harypanr.r3oBaB[rr4xcfl 

, rBJrrrorqufi cr o6uluu 4.nx Eopa:uu (u-or Kau.rarxa) 

u Cee. Arraepuxll. PacreHnl, orll4caHHbre c KaNr.rarrlr xax S. avatshensis 

Kom., L c. (lectotypus:

Pa6ora BblnoJrHeHa upra SrnnaHconofi noggepxre Poccuficxoro SoH- 

4a Syu4aueHraJrbHbrx lrccrreAoBaHnfi (npoexr Nb 03-04-49600) u rpanra 

flpe:n4enra P@ uo rroAAepxre BeAyqrax HayqHbrx urxol (upoerr 

Nb HIIJ-2198.2003.4). 

Jlnreparypa 

E a p 6 ap H q A. W. Scirpus L. // Onpe4eru,rreJrb Bbrcrurrx pacrennft Vxpar.rnn. 

Knes, 1987. C.418. 

B o p o tu !r Jr o s B. H. Onpegennrenr pacrenuft coBercKoro flalrnero Bocrora. 

M.. 1982.672 c. 

f eftaeMaH T. C. OnpegeJrr4reJrb Bbrcrrrrlx pacrenui Monaascroft CCP. 

Kuurnnen. 1986. 637 c. 

f p occ re ft u A. A.

H o s o c d a o s a M. C. Tunosue o6paeurr raKcoHoB ceueficrna Cyperaceae 

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Eoranfiqecruit uucruryr uu. B. JI. Korraapora PAH 

197376, C.-flerep6ypr, yn. IIpoS. flouona,2 

E-mail: egorova@DG4972spb.edu 

tvegorova@mail.ru

LI. B. Taranon I. Tatanov 

cr,rcTEMA poAA BOLBOSCHOENUS (ASCHERS.) PALLA 

(CYPERACEAE) 

SYSTEMA GENERIS BOLBOSCHOENUS (ASCHERS.) PALLA 

(CYPERACEAE) 

PoE Bolboschoenus (Aschers.) Palla -- ruy6uexauluu, orHoctrrcr K 

qr4cny HaLrMeHee l{ccJreAoBaHHbrx B cr4cTeMaTrlqecKoM oTHOlxeHUh pOAOB 

ceNreficrsa Cyperaceae, r,r HHKorAa He u3yrralcf, B Mr,rpoBoM uacura6e. 

Cyqecrryroque perr.roHanbHbre o6pa6orrra poAa B oreqecrBeHHbrx 14 

sapy6exuux u , 6ygyqn orpaHuqeHHbrMr4 B 

ocHoBHoM paMKaMrl r,r:yvaeuofi Teppr,rroprrr4, coAepxar, Ha Hatu B3lruA, 

prA HerrpaBHJrbHbIX rlpeAcraBJreHnfr n orHouteHtrr,r rpaKToBKr4 BtrAoB, r4x 

rnuuQuraqzu n reorpaQ[qecroro pac[pocrpaHeHus. Ocrarorcs Heqc- 

HbrMr{ BrlAoBofr cocras pog.a Bolboschoenus, ero [oJroxeHue B cr4creMe 

ceMeftcrsa r4 poAcrBeHHbre cBr3H. Cyuecrnyer upo6leua caMocroflrerb- 

HOCTLT pOAa, CBr3aHHar, B qacTHocTr4, C OTfpaHUqeHLreM ero oT poAa 

Schoenoplectas (Reichenb.) Palla. .{o Hacroruqefo BpeMeHr.r ue 6una 

pa:pa6oraHa crrcreMa Bolboschoenus. 

Po!, Bolboschoenus 6rrr srrAeneH tr3 poAa Scirpus L. flarla (palla, 

1905) uocpeAcrBoM trpvp:aHws. rpyrne BtrAoB Bolboschoenus Aschers. 

(Ascherson, 1864), onucauuofr 6er o6osua.reHur paHra, poAoBoro cl.a- 

Tyca Ha ocHoBaHur{ aHaroMr{rrecrlrx oco6euHocreft cre6.rrs, HarL4qua ouyrleHHbrx 

Kporoulr4x .reruyfi, Ha BepxytxKe c sneN,rxoft, nl roropoft Bbtxoqur 

ue6onruar ocrb (y Scirpus qeuyu 6el meuxu, ToHxo-ocreBr,rAHo 

3aocrpeHHble I,rnH Tyubre), lucrner 6e: a3uuxa, rJraAxnx pbrJlbueBblx Bernefi 

(y Scirpus oHtr c uarrr4Jrlauu), KpyrrHbrx KoJrocKoB. Ho, Hecuorpa Ha 

r{Meloulnecr uopt}oloruvecKr,re u aHaroMr,rqecKue orrnq[r, TaKcoHoMr,rqecxuil 

craryc poga Bolboschoenus Ao Hacrorrqero BpeMeHr4 rBJrrercfl 

rrpeAMeroM 4ucryccufi MHol'trx ur4rrepoJroroB. 

B oreqecrsenuofi lureparype, HaqrHas c pa6oru B. K. flpo6ona 

(1913) u o6pa6orru P. IO. Poxesrqa (1935) no

1999,2000; Adams, 1994; Browning et al., 1995,1996,1997a, b, 1998a, 

b; Bruhl, 1995; Goetghebeur, 1998; Kukkonen, 1998, 2001; Koyama et 

al., 2000; Hroudov6 et al., 2001; Hayasaka, Ohashi, 2002; Hroudov6, 

2002; Smith,2002; Pignotti, 2003). [o 3roro BrrAbr Bolboschoenus paccMarprdBaJrlr 

B poAe Scirpus B cocraBe ceKr.(rru, HasusaeMofi Bolboschoenus 

(Rouy, l9l2; Beetle,1942,1947; Raymond, 1957; Koyama, 

1958; Tang, Wang, 1961; Chang, Yang, 1976; DeFilipps, 1980; Tucker, 

1987), Maritimi (Chermezon, 1937; Kern, 1974) unu Reigera (Beetle, 

1944), nrnnqairr4 B cocraB rpyrrrbr Phyllantheli (Nyman, 1882; Fernald, 

1950) nn6o BbrAeJrfllrr B rroApoA Bolboschoenas (Ohwi, 1944). 

Prg aaropoa BKJrroqaer por, Bolboschoenus B por, Schoenoplectus (Lye, 

l97la, b, 1995; Haines, Lye, 1983; Strong, 1993, 1994; Karlsson, 1997). 

JIU (Lye, l97la) n pa6ore , 

ocHoBbrBaf,cb Ha AaHHbrx Bau gep BereHa (Van der Veken, 1965), ycra- 

HoBrrBruero, rrro BnAbr Bolboschoenus u Schoenoplectus nvretor oALrH THrr 

3apoAbrrra - Schoenoplectus-rnn (sauernu, uro n pa6ore Ban 4ep Be- 

KeHa oror rrrrr 3apoAbrlla o6osHa.reH xax Scirpus-Trlrr BcJIeAcTBI{e urrporoft 

rpaxroBKr,r poAa Scirpus, Ho ycraHoBJten on 6nn Ha Br4Aax poAa 

Schoenoplectus), npuurl po4 Bolboschoenus B KaqecrBe 6e:panroeofi 

rpyrrrbr B poAe Schoenoplectus, a 3arer (Lye, l97lb) rprrHf,Jr ee KaK ceKuuw 

Bolboschoenus B cocraBe 3roro poAa. flosAHee Xefinc n JIu (Haines, 

Lye, 1983) rroBbrctrJru AaHHyro ceKrlr,rrc Ao paHra rroApoAa. 3aueruu, uro 

rrpeAJro)KeHHbre Ha3BaHHbrMr,r aBTopaMu xou6uHaur4tr B paHre ceKrrr{H H 

uoApoAa He rBJrflIorcfl AefrcrB[TerbHo o6HapogoBaHHbrMtr, TaK KaK rrptr 

Hr.rx He yKa3aHbr 6a:uoHurr,ru. B pa6ore Bulcon (Wilson, l98l) yrcasnnaercq, 

qro xorr 3apoAbll Bolboschoenus vMeer ror xe caunft

foJroBqaroe, cocTotqee Lr3 2-HecKolbKrrx KoJIocKoB, oqeHb peAKo oAHo- 

KOJI OCKO BOC UJIIT 3OHTHKOBI,I.IHOC COI,IBETI,IE. 

flpune4eu eu1e prA AaHHbIx, rlo3BoJrflIoul[x cql,ITarb po!, Bolboschoe- 

NUS CAMOCT Ofl TEJIbHbIM, 

Kosars (Kowal, 1958), nryuunrrnft laop$oJlorl{n uJIoAoB 6olluro- 

fo rIficJIa eBpo[eftcKr,Ix [peAcraBr.{Tereil cyperaceae, rroKa3aJl, rrro BLAI'I 

poga Schoenoplectus (5. tabernaemontsni (C. C. Gmel.) Palla, S. lacusrtis 

(L.) Palla, S. triqueter (L.) Palla) u Bolboschoenus mqritimas (L.) Palla 

pa3Ir4qaroTcf llo cKyJrbrlType rIoBepxHocTI,I rIJIoAoB. 

Hccleqonauur Mapera (Marek, 1958) n Hauu (TaraHon, 2004) nu- 

flBtrrru pa3rvqkrr. MexAy Schoenoplectus u Bolhoschoenus rro aHaroMl,IqecKoMy 

crpoeHuro [epr{Kapfir4fi. Y npeAcraruTeJrefi SchoenoplectuJ KJrer- 

KI,I 3K3OKapUlrr MeJrKr,re, [OqTI4 BCefAa COAepXaT KpeMHe3eMHOe TeJIO, 

Me3OKap[Ufi COCTOTTT rr3 CKJreper4A, [O-pa3HOMy OpueHTrrpoBaHHbrx, 

KlerKr.r gHAoKapIIur roHKocreHHbre. y BIAOB BolboschoenuJ KJTeTKH 3K- 

3oKap[ur KpyrrHble, BbITqHyrbIe B paAu€urbHoM HarrpaBJreHlr[, 6e3 KpeM- 

He3eMHOfO Te[a, pexe MeJrKI,re, coAepxaxll4e efo, cKIepeI4AH Me3oKaprlr4t 

opHeHTupoBaHbI qaqe [apaJrleJrbHo rrpoAoJrbHoft ocH rrJroAa, Krer- 

KU 3HAOKaprrHf, TOJTCTOCTeHHHe. 

CasuleM (Savile, 1979) 6ruo ycraHoBJreHo, rrro KaxAbrfi r,rs pogon 

Scirpus s. str., Bolboschoenus u Schoenoplectus topaxaercfl cBor.rMr.r, po- 

AocueuuQuqHblMu, [apa3urr4qecKr,rMr,r px(aBrrr{HnbrMr,r rpu6auu, rrro lo- 

Bopnr o corrpfl )KeHHofi xoucoprr{BHofi 3BoJrlour{r{ [apa3arr4qecKr4x rpl4- 

6os u coorsercrByloull4x raKcoHoB-xo3fleB la, cJIeAoBareJIbHo, o caMo- 

CTOfl TEJIbHOCTI4 IOCJICAHIIX. 

B nocneAnee BpeMr B rleJrf,x r.r3yqeHr,rq cr4creMarnKr4 u QuJroreHr4u ce- 

Meft crBa Cyperaceae ucloJrb3ynrcr MoreKylspHbre AaHHbre. I,Icc[eAoBa- 

Hr.rrMH prAa aBropon (Muasya et al., 1998, 2000,2001; Young et al., 

2002) uo ceKBeHr{poBaur.to.{HK rroKa3aHo, qro poAbr Bolboschoenus n 

Schoenop leclas o6napyxnBaror cyulecrBeHHbre pa3flr4rruq rto cpaBHr.{BaeMHM 

Kpureprarrra (rbcl- u trnl-F DNA), rrro rrpoAeMoHcrpr{poBauo rro- 

Jro)KeHueM 3TIIX pOAOB B pa3Hbrx qaCTtX KJraAOfpaMMbr. 

Epyl (Bruhl, 1995) B KraAucrr4qecKoM aHarn3e ceMeficrBa Cyperaceae 

(c npuMeHeHr4eM KoMrrbK,repuofi 6a:u,qaHHbrx DELTA), ocHoBaH- 

HoM Ha ucrroJlb3oBaHrar,r uop$onorr4qecKr.rx, aHaroMr{qecrnx, eu6puonorlrqecKr{x, 

Suroxuurauecrux, SuauoJroruqecKr{x, reorpaQnvecKux rr 3KororuqecKrrx 

AaHHbrx, uoKa3aJr, rrro B flpeAerax rpu6m Scirpeae pog 

Bolboschoenns o6oco6neH or poAa Schoenoplectus. 

Pols Bolboschoenus u Schoenoplectus rpr4turbr KaK caMocrorreJrr,- 

HbIe a B cauoft uocneAueft clrcreMe Cyperaceae, [peAJroxeHHoft fere6epou 

(Goetghebeur, 1998), Ho, B orJrr{rrrde or crrcreMbr Epyna (Bruhl, 

1995), ouu orHeceHbr r rpu6e Fuireneae. 

na poga BolboschoerJ B rrenoM yKa3brBaercr pa3Hoe rrHcJto Br,rAoB: 

6-1 5 (Smith, 2002), 7- 8 (Eropona, 197 6), 8-10 (KoxeaHrKoB, 2001), l l 

82

(Smith, Kukkonen, 1999), 16 (Oteng-Yeboah, 1974; Wilson, 1981; 

Goetghebeur, Simpson, 1991; Kukkonen, 2001). 

O Hercuocru rpaHuu MexAy BuAaMLI Bolboschoenils LI Hacrof,TeJrbHoil 

HeooxoAl{Mocr[ B cBr3r.I c gTI{M TaKcoHoMnqecKofi peBI,I3r4v po!.a B IIOJI- 

HoM o6beMe HeoAHoKparHo yKa3bIBaJIocb B nl,Ireparype (Wilson, l98l; 

Goetghebeur, Simpson, l99l; Goetghebeur, 1998; Browning, Gordon- 

Gray, 2000, Smith, 2002). 

Hacroqurafl pa6ora 6rtna uocsrqeHa I,I3yqeHLIIo crlcreMarn xtt B o lbo' 

schoenus B MIdpoBoM MacrrlTa6e c lrelrbKl oupeAeJreHl,tt BI,IAoBofo cocra- 

Ba poAa u parpa6orrr{ efo cr4cTeMbl. 

I,IccreAosanr4e ocHoBaHo Ha I,I3yrIeHuIt o6urupuoro rep6apuoro rraa- 

Tepvara, B ToM qr{cJre r.r rr.rrroBoro, xpaHslqeroca n fep6apr,rrx EoraHI4rrecKoro 

LrHcrrrryra uu. B. JL Kouapona PAH (LE), C.-Ilerep6yprcxofo 

rocyAapcrBeHHofo yHHBepcurera (LECB), flasHoro 6oraunqecroro 

caila vv,. H. B. I{rauraHa PAH (MHA), Mocroscxoro rocyAapcrBeHHoro 

yHuBepcuTera r{M. M. B. JlolaoHocoBa (MW), Mocxoncroro ueAarorr{qecKoro 

focyAapcrBeHHoro yHuBepcrrera (MOSP), I{enrpa.nluoro 

Cu6npcroro 6oraHu.recxoro caAa CO PAH (NS), A;rraficKofo rocyAapcrBeHHoro 

yHuBepcr4rera (EapHayn, SSBG), Llncruryra EoraHltru rau. 

H. f. Xonoluoro (Kuer, KW), JllnoacKoro HalluoHturbHolo yHI,IBepcHrera 

rM. I,I. (DpaHro (LW), IlpupoAoBeAqecKoro My3et (Jlleoa, LWS), 

I4ucruryra 3KoJlorrrr{ Kapuar (Jllnon, LWKS), Hrarurcroro 6oraHu.rec- 

Koro caAa (r. -fllra, rtoc. HuKLITa, YALT), Tanpu.recroro HaIrI{oHaJrb- 

Horo yHr.rBepcr.rrera rau. B. I,I. BepHagcroro (Crana$eporloJlb, SIMF), 

Kpnucroro arpapHoro yHrBepcr{rera (Cur"rQeponolr, CSAU), I,Iucruryra 

EoraHuru AxaAeuI{Lt HayK 9eurcrofi Pecny6luru (flpyrour-rqe, 

PR, PRA), flpaxcnoro yHuBepcr.rrera (PRC), .{euapraueura Eorauurcu 

BeHrepcnoro My3efl ecrecrBeHHoit ucropun (Ey4arterur, BP), EpraraucKoro 

My3ef, ecrecrBeHHoir ucropuu (Jlou4ou, BM), HaquoHaJrbHoro 

My3er ecrecrBeHuofr ucropuu n flapuxe ( P), Huo-fropKcKoro 6oraHlrqecKoro 

ca.qa (NY), ForaHllqecxoro HayqHo-rrccJreAoBareJrbcKoro r,rHc'rrrryra 

B flperopll,r (PRE). Eruu ucnolb3oBaHH Jrr{rrHbre c6opu ll ua6.rrro- 

AeHLrfl B rrpupoAe, cAeJraHHbre s 2001-2004 rr. B HeKoropbrx perr{oHax 

Enpoueilcroft Pocculr (JleuuHrpaAcral, BraguMr,rpcKar, Huxeropo4craa 

o6r., ilyraucKas. v Mop4oacxar pecuy6nurn), a raKxe r Vrpauue 

(Kpuu). 

Huxe npuBoAr4Tcq cucreMa Bolboschoenrrs, rBJrf,Iouraf,cr r{ToroM 

BrrepBbre rrpoBeAeHHoit :r'auu KpuTlrqecKoit penuzuu poAa B MHpoBoM 

uacurra6e. 

Karc 6ruo orMeqeHo Bbrrue, cr{creMa poga Bolboschoenus Ao Hacroqxrero 

BpeMeHu se 6una pa:pa6oraxa. PasAeleuue poAa Ha 2 cew\uvr 

Bolboschoenus u Lentischoenus Oteng-Yeboah, ocyulecrBJreHHoe OreHrou-E6oa 

(Oteng-Yebo ah, 197 4), rBrfl ercr HerrpaBr4JrbH brM, rrocKoJlbKy 

ceKr1rr Lentischoenu,r ocHoBaHa Ha Br.rAe B. paludosus (A. Nelson) 5o6, 

83

6lueropo4crBeHHoM B. maritimus - rr.rfre poga Bolboschoenus, rr Mop- 

Sonoruvecrr{e rr aHaroMr{rrecKue [pu3HaKu, xapaKTepr,r3ylour4e AaHHyIo 

ceKrlrrK), He orJrnqaKrrct or raKoBblx ruuosoft ceKrlrrr4. floeronay ceKur{oHHoe 

Ha3BaHrre Lent is c ho enu.r crrHoHrrM r,r3rrpoBaHo B n3JloxeHHoft uauu 

cr,rcreMe c cerqueft Bolboschoenus. YcranosJreHne ceKrluu Lentischoenus 

6rrno csqsaHo c ouru6oqHbrM rroHr.rMaHrreM B. maritimus, :a xoroptrft 

rlpexAe HeKoropbre aBTopbr, B ToM qrrcJre rr aBTop otrvcarnfl. arofr cerunr4, 

rrprrHlrMann B. glaucus (Lam.) S. G. Smith. BcneAcrsre 3roro, ort{caHr{e 

rrdrroBofi ceruuu Bolboschoen rs, npuBeAeHuoe o pa6ore OreHra- 

E6oa, co4epxr4T npu3Haxn, cnoft crneHHbre rrocJreAHeMy BrrAy. 

flpegloxeunas.Hav.n cucreMa poga Bolboschoenus srrtoqaer 3 cerr1uu, 

5 uoAcerqufi u 14 suros. flouuuo nopQonorurecKrax rrpr43HaKoB, 

nplr paspa6orKe cucreMrr 6unu lrcrroJrb3oBaHbr opr4rnHaJrbHbte H Jrr.rrepaTypHbre 

AaHHbre IIo aHaToMr,rn nepr,rKaplr4r n cKyJrbrrrype rroBepxHocrr{ 

rrroAa (Browning, Gordon-Gray,1992, 1993, 2000; Browning et al., 

1995, 1996, 1997a, b, 1998a, b; Hroudov6 et al., 1997, 1998, 2O0l;' 

Hayasaka, Ohashi, 2002; Eropona, Tarauon, 2002,2003; TaraHoe, 

2004). 

Genus Bolboschoenzs (Aschers.) Palla, 1905, in Hallier u. Brand, 

Koch's Syn. Deutsch. Schweiz. Fl., 3 Aufl., 3 :2531t;Apo6., 1913, Tp. 

Eor. vry:. AraA. Hayrc 11 :86;Poxen., 1935, so @n. CCCP 3:55; Eroposa,1967, 

s Pacr. I{enrp. Aguu 3: 19; ona xe,1976, so @n. Enpon. 

qacrn CCCP 2:93; Oteng-Yeboah,1974, Notes Roy. Bot. Gard. Edinb. 

33,2:315; T. Koyama, 1978, in Fl. Taiwan 5:206; id., 1980, Acta 

Phytotax. Geobot. (Kyoto) 31, 4-6: 139; Wilson, 1981, Telopea 2,2 : 

156; Y. C. Yang et M. Zhan,1987, Acta Biol. Plateau Sin.7: 13; 

A. E. KoxesHuKoB, 1988, s Cocya. pacr. coB. ,{,a.lrnu. Bocr. 3 : 187; Tra- 

Moxr,IHa u Eougapena, 1990, so Or. Cu6.3 :22; Browning a. Gordon- 

Gray, 1992, South. Afr. Journ. Bot. 58, 5 : 380; iid., 1993, l. c. 59, 3 : 

3l l; iid., 2000, L c. 66, | : 6l; Browning et al., 1995, Brittonia 47, 4 : 433; 

iid.,1997, Austral. Syst. Bot. 10, I : 49; Goetgh., 1998, in Kubitzki, Fam. 

a. Gen. Vasc. Pl. 4:165; Kukkonen, 1998, in Rech. f., Fl. Iran. 173:16: 

Hroudov6 et al., 2001, Zprhvy Cesk6 Bot. Spoledn. 36, I : l; Hayasaka 

a. Ohashi, 2002, Journ. Jap. Bot. 77. | :9; Hroudov6,2002, Klid KvEt. 

eesk6 Rep. : 794; Smith ,2002, in Fl. North Amer. 23 : 37. - Scirpus L. 

a. Bolboschoenus Aschers., 1864, Fl. Brand. | : 753. - Reigera Opiz, 

1852, Seznam Rostlin KvEt. eesk6 : 83, nom. nud. - Scirpas Rotte 

Phyllotryon Ddll, 1857, Fl. Bad. : 305, p. p. - S. a. Phyllantheli Nyman, 

1882, Consp. Fl. Eur. 4 : 763, nom. nud., p. p., quoad S. maritimum; 

I Cornacno yKa3anulo na:rurynr,uoii crpaHnqe Storo rona, a raKxe aaHHhrM Cra$ne u 

Kayaxa (Stafleu, Cowan, 1979), pe4axropaMr.r sToro roMa rBnrrorcs E. Hallier r 

A. Brand, a xe E. Hallier a R. Wohlfarth, xar o6uuxo yKa3brBaercr B HoMeHrnaryp- 

H6rx qr{Tarax npn poAe Bolboschoenus u snAe B. maritimus. 

84

Fern., 1950, in Gray's Man. Bot., ed. 8 : 271. - S.b. Cymelfi Meinsh., 

1901, Tp. flerep6.6or. caAa 18,3:249,p. p., quoad S. maritimumet 

S. affinem. -,S. sect. Bolboschoenus (Aschers.) Rouy, 1912, Fl. Fr. 13 : 

370; Raymond, 1957, Nat. Canad.84,5: 118 (cum auct. Aschers.); 

T. Koyama, 1958, Journ. Fac. Sci. Univ. Tokyo, sect. 3 (Bot.) 7, 6 :288; 

Ts. Tang et F. T. Wang, 1961, in Fl. Reip. Pop. Sin. 11 : 2;Y. L. Chang 

et Y. L, Yang, 1976, in Fl. Pl. Herb. Chin. Bor.-Or. 11 : 3. - S. sect. 

Maritimi Cherm., 1937, Fl. Madag. fam.29: 155;Kern, l974,in Steenis, 

Fl. Males. ser. 1, 7,3:499; Eropona n Kxofi, 1980, Hosocrrr ctrcr. 

Bbrcrr. pacr. 17 : 54. - S. sect. Bolboschoenas (Aschers.) Beetle, 1942, 

Amer. Journ. Bot.29, I : 82, comb. superfl.; DeFilipps, 1980, in Fl. 

Europ. 5:278; Tucker, 1987, Journ. Arn. Arbor. 68,4 : 375. - S. sect. 

Reigera (Opiz) Beetle, 1944, Amer. Journ. Bot. 31, 5 :263, comb. inval. - 

S. subgen. Bolboschoenus (Aschers.) Ohwi, 1944, Mem. Coll. Sci. Kyoto 

Univ. ser. B, 18, I : 96. - Schoenoplectus (Reichenb.) Palla sect. 

Bolboschoenas (Aschers.) Lye, 1971, Bot. Not. 124,2 :290, comb. inval. 

(sine bas.). -- Schoenoplectus subgen. Bolboschoenus (Aschers.) Haines et 

Lye, 1983, Sedges a. Rushes East. Afr. : 52, comb. inval. (sine bas.). - 

Konocrclr MHororlBerKoBbre, B uracne (l) 2-50 (y B. nobilis - 100-200, 

uHorAa Ao 450), (0.7) l-4 (6) crra ar., (0.4) 0.5-0.7 (1) crr,r unrp., qftqepr.rA- 

Hbre, [poAonronaro-gftueBl,tAHbre, [poAoJlfoBarbte, pexe yAJtuHeHHorruruH.qpuqecrue, 

co6paHHbte B o6qee repMrlHalbHoe 3oHTrrKoBr4AHoe 

couBerne - aHTeJroAr,rfi, cocrorulee rd3 rleHTpalbHoro, cnArrrero, rryqKa 

KoJrocKoB u nyuefi, Hecyqrrx Ha KoHuax no (l) 2 9 (10) cKyqeHHbrx B 

rryrrKu KoJrocKoB, r.rJrr,r B roJroBqaroe, o6pasoraHuoe oAHr{M ulu 2 3 luyt- 

KaMLr, CIIATTIUMU k[tl4 paC[OJTOXeHHbIMI4 Ha CnJIbHO yKOpOqeHHbIX Jlyqax, 

Couserue oKpyxeHo rrprr ocHoBanuu (1) 24 (5) Kporotqr4Mr.r Jrr{crbflMrr 

c nugeftHuur{ nucroBblMtr rlJlacruHKaulr, HI,IXHI,Ift H3 Koropblx rlpeBbrrrraer 

ero, Kporculr,re qeruyu paclloJloxeHhr rro cfluparru, [eperroHqarbte, 

(4) 5-9 (10) una an., (2.5) 3-4 urvr ruup., orryueHHbre c a6arcuanluofi 

cropoHbr, c nrreurofi Ha BepxyrrrKe, r43 Koropoft errxoAnr KoporKaf, 

ocrr. I{nerru o6oenonue. OroroqserHbre uIerHHKr,r lrrJroBr,rAHbre, c HarrpaBJleHHbrMr.r 

BHr{3 rrrnrrlrKaMH, peAKo riraAKr,re, B qllcfle (0) 1-6, coxpa- 

Hrtoulnecr nJrrr orraAaroque flpu rrJroAax. Trrquuox 3. Crotiux c 2-3 

PbIJIbUCBbIMII BCTB'MI.I. fIIOAU TIJIOCKO-BbITIYKJIbIC, HCPABHO ABO'KOBbIrryKJrbre, 

Bbr[yKno-TpexfpaHHbre, ocTpoTpexlpaHHbre, TynoTpexfpaHHbre, 

cx(aro-TpexfpaHHbre (uuor4a c nua4uHroft ua rpanrx) r,rJrn ABorKoBor- 

Hyrbre, (2) 2.54 (5.5) urra N., (1.2) 1.5-2.5 (2.7) uu runp., Ha Bepxyu- 

Ke c oqeHb KoporKr,rM (HoclrroM) - ocrarKoM cron6ura. Cxynrnrypa 

[oBepxHocru rrJroAa (npu uryvenura Ha C3M) 6yropuarafl, vrwr suqarau; 

sr:oxapnr.rft cocror4T H3 oAHoro pf,Aa KpylHbrx, flprMoyroJrbHblx, Bbrrfl- 

HyTbrx B paAuaJrbHoM HarrpaBJleHr,rr,r KJreToK, He coAepxaqux KpeMHBeM- 

Horo reJra, nn6o uenrux, KBaAparHbIx ltJrr,r flprMoyroJlbHbrx, cJrefKa 

BbrTtHyTbrx B TaHf eHrI[aJrbHoM wrrr4 pa&uailbHo M Ha[paBJIeH 14 11, COAep_ 

85

xarqr{x KpeMHe3eMHoe reJlo; uesorapurafi cocrour us 4-12 (13) paaor 

yAJII4HeHHbIX BepeTeHoBHAHbIX CKJIepehA, opI4eHTI4poBaHHbIX flapiurneJlb- 

Ho rrpoAoJrbHofi ocu tIJIoAa, pexe cKJrepel4Abl' rlpHMbIKaIouHe K SHAo- 

KapIIup, opHeHTIlpoBaHbI rleprleHAl'rKyJrflpHo; auAorapnuil o6pasonaH 

oAHr{M prAoM TOJICTOCTeHHbIX BOJTOKHOO6pa:Hltx KIeTOK, OpI4eHTI',IpO- 

BaHHbrx [eprreHAr{KyJupHo npogollHOft OCn IIJIOAa. celreHHar Koxypa 

clrJlbHo Ae3r4HTefpHpoBaHHaf, cocTofulat I{3 Hapy)I(HOl-o cJlof (ax:orecru), 

npe4craBJreHHof o 4e$opvillpoBaHHbl MI4, pe)Ke coxpaHfl IOUII{MK 

cBoro cTpyKTypy KJrerKaMr{, 11 BHyTpeHHero 6eccrpynrypso|o cltol. 3a- 

-Bolboschoenus-rr.rra 

poAb[rr 

(Goetghebeur, I 986, I 998). Muot'oIeTHI4e 

rpaBbr (30) 50-150 (200) crvr Bbrc., c fopr{3oHraJlbHblMrl ArlrIHHbIMlI Kop- 

*ri""uut, u rny6HenuAHblMll o6paronaHntMu Ao 4 (4'5) cM B AuaM'' 

AaroullrMtr Haqiuto HoBbtM uo6erau. Huxune JlrrcTbt qeulyeBuAHble, Cpea""r"r" 

(cre6nerne) - c BJrarar[uleM rI AnnHnoft unacrunroft 6e: 

,sbt{Ka, BepxHrre (rporcqr,re JrKcTbfl coqserns) - 6es BnaraJll{ula. - 

2n= 26-112. 

T y p u s: Scirpus maritimus L. (= Bolboschoenus maritimus (L') 

Palla). 

PoJI srnrcqaer 14 sraAos, paclpocTpaHeHHblx B yMepeHHblx, CyorporrlrqecKux 

?I orqacrl4 Tpo[nqecKux o6nacrqx o6onx uolyurapufr' 

Sectio l. Browningia Tatanov sect. nova. -' Scirpus L' sect' Maritimi 

cherm., 1937, Fl. tvtadag. fam.29: 155, quoad pl. - Bolboschoenus sect. 

Bolboschoenar sensu Oteng-Yeboah, 1974, Notes Roy. Bot. Gard. Fdinb' 

33,2 :315, 316. - Infloriscentia umbelliformis radiis ordinis primi vel 

primi et secundi, rarius primi, secundi et tertii. Setae hypogynae in fructu 

persistentes. Stigmata 3. Fructus acute vel obtuse trigoni. Exocarpii 

iellulae ad fructus faciem indistinctae. Exocarpii cellulae in fructus 

sectione transversa parvulae quadratae vel rectangulares radiatim vel 

tangenter paulo elongatae. Corpus siliceum in exocarpii cellulis adest. 

T y p u s'. B. glaucus (Lam') S' G. Smith' 

Ai"a geog.aphica. Eurasia (Asia Boreali exclusa), America Borealis 

(Canadae parte boreali exclusa), Africa, Australia' 

Sectio in honorem generis Bolhoschoenus investigatoris cl. J. Browning 

(Pietermaritzburg, Respublica Africa Australis) nominatur' 

Couserue 3oHTHKoBIrAHoe, c JlyqaMu l-2 (3) [opflAKoB' Oronouser- 

Hbre uIeTr4HK14 trpkr trJroAax coxpaHrroquecr. Prneu 3. flloAlt ocTpor{Jrr{ 

TyrroTpexfpaHHbre. Ha nonepxHocTtr rrJroAoB He BIAHbI KJIeTKU 3K- 

3oKap[ur. Ha uouepe.rHoM cpe3e [JroAa KJreTKrr 3K3OKapIIlIfl MenKne' 

KBaApaTHbIe I4nU flpqMoyfoJlbHble' clefKa BbITtHyTble B paAuanbHoM 

IrJtlr TaHTeHII[aIbHoM HaIIpaBJIeHI{n. B xnerrax 3K3oKap[ut I{MeeTCq 

KpeMHe3eMHOe Teno. 

T u u : B. glaucus (Lam.) S. G. Smith' 

PacnpocrpaHeHlle. Enpa:ur (rporrae CenepHofi Arran), Cenepuar Auepraxa 

(sa ucxnroqe*rneu cenepHoft qacru KanaAu), A$prara, Arcrpatrna. 

86

Cerqus Ha3BaHa B qecrb l,IccneAoBarelbHllqbl poga Bolboschoenus 

X. EpoynnHr (flureprr,rapuq6ypr, tOxno-Atppl,IKaHcKat Pecny6nura). 

Subsectio l. Glauci Tatanov subsect. nova. - Inflorescentia radiis 

ordinis primi vel primi et secundi, rarius primi, secundi et tertii. Spiculae 

ovoideae, oblongo-ovoideae, oblongae, cylindricae, interdum elongatocylindricae. 

Squamae nitentes, brunneae, pallide brunneae, ferrugineo- vel 

rubello-fuscae seu pallide flavae. Fructus ellipsoidei vel obovoidei, ad 

3.3 mm lg. Sclereidarum series in numero 7 9 in mesocarpio adsunt. 

T y p u s'. B. glaucus (Lam.) S. G. Smith. 

Area geographica. Europa Australis, Asia Occidentalis, Media, 

Centralis et Australis, Africa. 

Coqnerne c ryqaMl,t l-2 (3) roptAKoB. Kolocxu sfiuesl,IAHue, rpo- 

AOJtfOBaTO-tftUeSnAHrre, [poAoJrfoBarble IIJII{ uI4[I{HApnqecKI,Ie, nHorAa 

yAnuHeHHo-quluHApr,rqecrue. Kpotouue qeuryu 6nectsulne, KopHqHe- 

Bbre, cBerJro-Kopr,rtrHeBble, pxaBo- rIJrI4 KpacHoBaro-6ypue unu 6leAuoxeJrrble. 

flnoAu 3JrJrrr[cor4AaJrbHbre IIJII,I o6paruoaft ueBI4AHbte, Ao 3. 3 rvrrta 

An. B Naesorcaprl,ru 7-9 pa.qon cKJrepelrA. 

T u n : B. glaucus (Lam.) S. G. Smith. 

PacnpocrpaHeHue. IOxHas Erpona, 3ara4Har, Cpe4uar, I{eHrpa"ulnar 

u [OxHat A3ut, A$pr,ma. 

Species: l. B. glaucus (Lam.) S. G. Smith (= B. maritimus auct. 

nonn.). - 2. B. nobilis (Ridl.) Goetgh. et D. A. Simpson. 

Subsectio 2. Fluviatiles Tatanov subsect. nova. - Iflorescentia radiis 

ordinis primi. Spiculae oblongo-ovoideae. Sguamae opacae, fuscobrunneae. 

Fructus oblongo-obovoidei, ultra 3.3 mm lg. Sclereidarum 

series in numero 10-13 in mesocarpio adsunt. 

T y p u s'. B. fluviatlh's (Torr.) Soj6k. 

Area geographica. Eurasia (ab Europa Centrali ad oceanum Pacificum, 

meridiem versus ad Indochinani inclusive), America Borealis, 

Australia. 

Coqserue c nyqaMr,r I nopr4xa. Konocrn npoAoJrroBaro-rftqesuAnue. 

Kporculue rreuyr4 MaroBbre, 6ypo-ropnuHeBbre. fhoArr rrpoAoJrroraro-o6parHorftuesrcAHrre, 

6olee 3.3 rrarvr An. B uesoxapfiuu l0-13 pr- 

AOB CKJreper,rA. 

T u u : B. fluviatilis (Torr.) Soj6k. 

PacnpocrpaHeHne. Espasua (or L{enrpalrnofi Enponu 4o Tuxoro 

oKeaHa, Ha lor Ao HuAornrar BKrlorrurenlHo), Cenepuar Auepnra, 

Arcrpanua. 

Species: 3. B. fluviatifti (Torr.) Soj6k (= Schoenoplectus .fluviatilis 

(Torr.) M. T. Strong.- 4. B. yagara (Ohwi) Y. C. Yang et M. Zhan 

(= B. yagara (Ohwi) A. E. Kozhevnikov, = B. fluviatifts subsp. yagara 

(Ohwi) T. Koyama, = B. maritimus var. desoulavii Drob., = B. desoulavii 

(Drob.) A. E. Kozhevnikov). 

87

Sectio 2. Bolboschoenus. - Reigera Opiz, 1852, Seznam Rostlin Kv6t. 

Cesk6 : 83, nom. nud. - ScipusL. a. PhyllantheliNyman, 1882, Consp. 

Fl. Eur. 4:763, nom. nud., p. p., quoad S. maritimum; Fern., 1950, in 

Gray's Man. Bot., ed. 8 : 27L. - S. sect. Maritimi Cherm., 1937, Fl. 

Madag. fam.29: 155, quoad nom.; Kern, L974,in Steenis, Fl. Males. ser. 

1,7,3:499, quoad nom.; Eropona u Kxofi, 1980, Honocrl,r cr{cr. Bbrcur. 

pacr. 17 : 54, quoad nom. -,S. sect. Reigera (Opiz) Beetle, 1944, Amer. 

Journ. Bot. 31. 5 :263. comb. inval. -- Bolboschoenus sect. Lentischoenus 

Oteng-Yeboah, 1974, Notes Roy. Bot. Gard. Edinb. 33, 2 : 315. 

Inflorescentia umbelliformis radiis ordinis primi vel capitata. Setae 

hypogynae in fructu caducae. Stigmata 2-3 vel in eadem spicula 2 et 3. 

Fructus plano-convexi, inaequaliter biconvexi, trigoni, convexo-trigoni 

vel biconcavi. Exocarpii cellulae polygonales ad fructus faciem distinctae. 

Exocarpii cellulae in fructus sectione transversa magnae, radiatim 

elongatae. Corpus silicium in exocarpii cellulis nullum. 

Typus;generistypus. 

Area geographica. Regio Holarctica, Asia Australis et Austro-Orientalis, 

America Australis (pars borealis), Australia. 

Couserue 3oHTI{KoBI,IAHoe, c [yqaMr4 rrepBofo uoptAKa, r.rJrr.r roJroBrraroe. 

OroilouserHbre r.ueruHKn [pr,r [noAax o[aAarouue. Puneu 2 3 

unu 2 u 3 s oAuol,r r,r roM xe KoJrocKe. flloArr rlJlocKo-BbrrryKfibre, HepaBHo 

ABOfl KOBbrlryKJrbre, Tpexf paHHbre, BbrrryKJro -TpexrpaHHbr e v1il4. 

ABoqKoBorHyrue. Ha [oBepxHocrr{ rrJroAoB orqerJttrBo BtrAHbr rroJrr{fo- 

HaJrbHbre KirerKr,r 3K3oKaprrufl. Ha uonepeqHoM cpe3e rrJroAa KJlerK[ 3K- 

3oKap[[r KpyrrHbre, BbrrrHyrbre B paAr,raJrbHoM HarrpaBJrenuu. Kpeuue- 

3eMHOe TeJtO B KJIeTKaX 3K3OKapU[f, OTCyTCTByeT. 

Tun:Tr.rrrpoAa. 

PacupocrpaHeHr{e. folapxruxa, IOxHax u lOro-Bocro.{Haq A:rr, 

IOxsar Auepuxa (cerep), Ancrpalua. 

Kouueurapun, KacaloulI,tecr cHHoHtrM v3a:1'rru ceKur{oHHoro Ha3BaHr{q 

Lentischoenus Oteng-Yeboah, l. c., rp[BeAeHbt HaMr,r ouure (crp. 83). 

Subsectio 3. Bolboschoenus. - lnflorescentia umbelliformis vel 

capitata. Spiculae oblongo-ovoideae vel oblongae. Scuamae brunneae, 

rubello-brunneae, rarius luteolo-albidae. Stigmata 2 3 vel in eadem 

spicula 2 et 3. Fructus plano-convexi, inaequaliter biconvexi, convexotrigoni 

vel trigoni. Pericarpii crassitudo in partibus omnibus fructus 

sectionis transversae aequalis. 

Typus:generistypus. 

Area geographica. Eurasia (ab Europa Atlantica et Asia Occidentali 

orientem versus ad lacum Baical), America Borealis, America Australis 

septentrio, Africa Borealis. 

Coqserue 3oHTHKoBr{AHoe uJlr4 roroBr{aroe. Konocru [poAoJrroBarorfi 

uesuAuue r,rJrr,r upoAonroBarhre. Kporoulrae qe[yu KopuqHeBbre, Kpac- 

HoBaro-Kopr,rqHeBbre, pexe xeJrrosaro-6erosarue. Ptrneq 2 3 ulu 2 u 

88

3 n oAsou Lr roM xe KoJrocKe. flloAu rrJrocKo-BbrrryKJrbre, HepaBHo ABorKoBbrrryKJrble, 

BblrryKJrorpexfpaHHbre HJrLI TpexrpasHne, TonuII,rHa rrepl,IKap[ut 

tlo BceMy IIepI.IMeTpy llofiepeqHoro cpe3a IIJIOAa OALIHaKoBat. 

T u n: rr,r[ poAa. 

Pacnpocrpaneuue. Erpasux (or AuaHurqecxoft Enponu u 3auaAsoft 

Asrrra Ha BocroK Ao 03. Eaftraa), CenepHar u loxsar (cenep) Auepuxa, 

Cenepuar AQpnxa. 

Species: 5. B. maritimas (L.) Palla (= B. compactus (Hoffm.) Drob., 

= Schoenoplectus maritimus (L.) Lye). -- 6. B. schmidil (Raymond) 

Holub. - 7. B. paludosus (A. Nelson) 5o6 (- B. maritimus subsp. 

paludosus (A. Nelson) T. Koyama). -- 8. B. robustus (Pursh) Soj6k 

(= Schoenoplectus robustus (Pursh) M.T. Strong). 

Subsectio 4. Popoviorum Tatanov subsect. nova. - lnflorescentia 

capitata. Spiculae dolioliformes, densissimi. Squamae luteolo-albidae. 

Stigmata 2. Fructus inaequaliter biconvexi vel plano-convexi. Pericarpii 

crassitudo in partibus omnibus fructus sectionis transversae aequalis. - 

Subsectio monotypica. 

T y p u s: B. popovii Egorova. 

Area geographica. Europa Orientalis (pars austro-orientalis), Caucasus 

(pars orientalis), Asia Media et Centralis. 

Coqnerue roJroBqaroe, Konocru 6ouoHronu4Hbre, oqenb rycrbre. 

Kporoulne qeuryr4 xerroBaro-6eflosarne. Prrleu 2. flnoau HepaBHo 

ABotKoBbrrryKJlbte r,rJrr{ rIJrocKo-BbIrryKJrHe, Tolrquga flepr4Kaprrnfl tlo 

BceMy flep?rMerpy flotrepeqHoro cpe:a [JroAa oAuHaKoBar. - MoHoruu- 

Hafl [oAceKr.([g. 

T u n : B. popoviiEgorova. 

PacnpocrpaHeHue. Ioro-socror Bocro.{Hoft Erponu, Kanrar (rocroqHar 

vacrr), Cpe4nxr u I{enrpanlnas, Agus. 

Species: 9. B. popovii Egorova (= B. alfinis (Roth) Drob., p. p., excl. 

tvpo). 

Subsectio 5. Affines Tatanov subsect. nova. - Inflorescentia capitata, 

rarissime umbelliformis. Spiculae oblongo-ovoideae. Squamae brunneae, 

pallide brunneae vel flaventes purpureo-striatae. Stigmata 2. Fructus 

biconcavi cum cavitate in latere abaxiali et adaxiali vel leviter biconvexi. 

Pericarpii crassitudo in sectione transversa inaequalis: haec in angulis 

fructus multo major quam in lateribus. 

T y p u s : B. affinis (Roth) Drob. 

Area geographica. Eurasia (ab Europa Centrali ad oceanum 

Pacificum et a czukotka Kamtschatkaque ad Indochinam inclusive, 

aeque ac Asia Media, Centralis et Australis), Australia. 

Coqnerue foJloBqaroe, orleHb peAro 3oHTr4KoBtrAHoe. Konocrr4 upo- 

AoJlroBaro-rfiqenlr4nne. Kporcque qeuryu Kop[qHeBbre, cBerJro-Kopr,rq- 

HeBbre rdJrr4 xeJrroBarbre c [ypflypHbrMr,r rurpr,rxaMr,r. prmeq 2. flnoArr 

89

ABorKoBorHyrble, Ha o6eux cropoHax c BuaAuHKoit, uttu crrerKa ABot- 

KoBbrrryKJrbre. Toruruna fiepuKaprllrr B pa3Hblx qacrtx flo[epeqHoro cpe- 

3a [JroAa HeoAuHaKoBaq, B yrlax [JroAa oHa 3Haql4TenlHo 6olr'ue' qeM 

Ha cropoHax. 

T u u : B. ffinis (Roth) Drob. 

PacupocrpaHeHue. Enpasnr (or l{enrpanlnoft Erponu Ao Tuxoro 

oKeaHa u or 9yrorxu n KarvrqarKu Ao I'luAorurar BKJrK)qurerbHo, a 

TaK)r(e Cpe4Hrr, I{eurpzutruaq n [OxHas Asur), Ancrpalur. 

Species: 10. B. aJfinis (Roth) Drob. (= B. strobilirr.rs (Roxb.) 

V. Krecz., = B. grandispicus (Steud.) Lewej. et Lobin). - 11. B- planiculmis 

(F. Schmidt) Egorova (= B. caldwellii (Y. J' Cook) Soj6k, 

= B. koshewnikowii (Litv. ex Zinger) A. E. Kozhevnikov). 

Sectio 3. Mediani Tatanov sect. nova. lnflorescentia umbelliformis 

radiis ordinis primi. Setae hypogynae in fructu caducae, rarius 

persistentes. Stigmata 3 vel in eadem spicula 2 et 3. Fructus compressotrigoni 

saepe cum cavitate in faciebus vel plano-convexi. Exocarpii 

cellulae polygonales ad fructus faciem solum in areis nonnullis distinctae. 

Exocarpii cellulae in fructus sectione transversa magnae, radiatim 

elongatae. Corpus siliceum in exocarpii cellulis saepius nullum. - Sectio 

species hybridogenas includens. 

T y p u s: B. medianas (V. J. Cook) Soj6k. 

Area geographica. Europa, Asia Orientalis (Oriens Extremus rossicus, 

Chinae pars orientalis), Asia Austro-Orientalis, America Borealis, 

Australia. 

Coqserue 3oHTr4KoBIdAHoe, c ryqaM[ I nopagxa. OrotrouserHute 

ulerr,rHKl,r rrprr uJroAax ollaAalorrue, pexe coxpaHtlouuecq. PltJteu 3 nttu 

2 u 3 s oAHoM 14 ToM xe KoJIocKe. flloArt cxaro-TpexrpaHHble, qacro c 

nuaAuuroft Ha rpaHtx I,IJIH IIJIocKo-BbIuyKnbIe. ICrerru 3K3oKaplllrt BI4A- 

Hbr roJrbKo Ha HeKoropbrx yqacrKax IIoBepxHocrI4 ruloAa. Ha uouepeu- 

HOM Cpe3e [noAa KJIeTKI4 3K3OKaplUS KpyrlHble, BblTflHyTbIe B paAnaJlb- 

HoM HarrpaBileHr4v. B rlerxax 3K3oKaprII4t KpeMHe3eMHoe reJlo qaue 

orcyrcrByer. -- Ceruuq, BKJIIoqaIourar ru6pn4oreHHble BI4AbL 

T u rr : B. medianus (V. J. Cook) Soj6k . 

PacnpocrpaHeHue. Enpona, Bocroqnas (,{anrHuft Bocror Poccuu, 

BocrorrHbre paftouu Klrrar) n lOro-BocroqHat Arua, Cenepuar Auepnra, 

Aactparux. 

Species: 12. B. medianus (Y. J. Cook) Sojak. - 13. B. novae-angliae 

(Britt.) S. G. Smith (= Schoenoplectus novae-angliae (Britt.) M. T. Strong). 

14. B. laticarpas Marhold, Hroudova, et al. 

B :axno.{eHne xoqy Bbrpa3nrb caMyrc ncKpeHHrcIo 6larogapuocrr 

MoeMy HayqHoMy pyKoBoAureJrp T. B. Eroporofi 3a ueHHbIe coBerbl no 

BorrpocaM HoMeHKraTypbI r4 c[cTeMaTIrKrr, Bo3HLIKIJIIM npn parpa6orxe 

cr4creMbr poga Bolhoschoenus, a raxxe 3a IIepeBoA Ha JlaruHcKl4ft ssrtr 

90

ouucasnft BbrAereHHbrx 3Aecb raKcoHor. -f, upuruareneH T. B. lllyntrnuofi 

(Missouri Botanical Garden, St Louis, USA) ra rrc6esHo oKa3aH- 

Hyro rroMorqb B [peAocraBJleHlrl4 HeKoropblx peAKux illrreparypHblx I4c- 

TOqHUKOB. 

Pa6ora BbrrroJIHeHa upu SunaHcosoft uoAAepxre Poccuficxoro $ouga 

SyH4arvreHraJrbHbrx nccileAoBaHl4ft (npoerr Nb 03-04-49600) r,r rpaura 

flpe:u4eura PO no rloAAepxKe BeAyuII4x HayqHblx urron (npoerr 

Nb Hru-2198.2003.4). 

Jlnreparypa 

.Il p o 6 o n B. K. K ctrcreMarrKe poga Bolboschoenus Palla (Scirpus L. ex 

parte) n ero pacrpocrpaHenrll n Cn6r.rpn // Tp. Bor. nryr. Axa4. nayrc. 1913. 

Bun. ll. C.86-96. 

E r o p o n a T. B. Bolboschoenus Palla ll Pactesus l-(eurpanrnofi AsHu. Jl., 

1967. Bun. 3. C.19-22. 

Eropora T. B. Kny6HeKaMbIIIr - Bolboschoenus(Aschers.) Palla ll @ttopa 

Enponeftcxofi qacru CCCP. il., 1976a. T.2. C.93-96. 

Eropona T. B. Poa Bolboschoenus (Aschers.) Palla- Kny6nerarrarrrrt // 

OnpeAenlrrenr pac'renufi Cpeauefi AsnH. Tauxesr, 19766. T. 5. C. 1720. 

Eropora T.8., TaraHoB A. B. Bolboschoenus glaucus (Lam.) 

S. G. Smith (Cyperaceae) - nosHfi Br r\ p.:l,fl. Qnopu Kasxasa // Hosocra clrcreMarr.rKr.r 

Bbrcrul4x pacrenufi. Cll6., 2002. T. 34. C. 34-43. 

Eropona T. B., TaraHoB 14. B. O cucreMarl{qecKoM noJroxeHHLI 

Bolboschoenus planiculmis u Bolboschoenus koshewnikowii (Cyperaceae) ll 

Eor. xypH. 2003. T. 88, I& 4. C.133-144. 

K o x e B H r.r K o B A. E. Kny6ueKaMbllrr - Bolboschoenas (Aschers.\ Palla ll 

CocyAracrrre pacreHr.lt coBercKoro .Ilanrnero Bocroxa. Jl., 1988. T. 3. 

c. 187-190. 

KoxeBHHKoB A. E. Csrrenue (cenreficrno Cyperaceae Juss.),!,anrHero 

Bocroxa Poccnu (conpeueuurrft raxconouuqecxufi cocraB n ocHoBHbre 3a- 

KoHoMepHocru ero Qoprur.rpooaunr). BnaArsocror , 2001 . 27 5 c. 

Poxesrq P. IO. Kly6neKaMbrru - BolboscltoenusPallall @rcpa CCCP. 

n.. 1935. T. 3. C. 55-57. 

T a r a H o s I,l. B. Poa Bolboschoenus (Cyperaceae) Poccur,r n conpeAenbHblx 

rocyAapcrB // Eoraun.recrue [ccJreAoBaHns s A3lrarcxofi Pocculr: naarep. Xl 

cbe3Aa Pyccroro 6or. o6ut. (18-22 anrycra 2003 r., Honocn6r.rpcr-Eapnayn). 

Eapnayn, 2003. T. l. C.293-294. 

T a r a n o n l,l. B. CparnureJrbHar Kapnorornr nugon Bolboschoenus (Cyperaceae) 

B cBq3u c cr4creMarr.rKoft poaa ll Eor. xypn. 2004. T. 89, I1! 8. 

c. 1225-1248. 

Tzuoxr.rHa C. A., EoHAapeBa H. B. Bolboschoenus (Aschers.) 

Palfa - Kny6uexaunur ll Qnopa Cn6upn. Honocn6npcr, 1990. T. 3. C.22-23. 

I n e n € n H. H. Onpeaerrnrerrb pacrennfi Cenepo-3anaqHoft Poccnu. CI]6., 

2000.781 c. 

A d a m s C. D. Bolboscltoenus (Asch.) Palla / G. Davidse et al. (eds.). Flora 

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9l

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2002

A..[. Mnxeen 

HOBbIE BII.IbI POAOB ALLIUM L. (ALLIACEAE) 

LI GLADIOLUS L. (IRIDACEAE) C KABKA3A 

A. Mikheev 

GENERUM ALLIUM L. (ALLIACEAE) ET GLADIOLUS L. 

(IRIDACEAE) SPECIES NOVAE E CAUCASO 

B Hacroqueft crarte orrlrcblBalorcfl ABa HoBblx BvLLa v3 poAoB Allium 

L. vt Gladiolus L. c reppnroplll4 Kanrasa. 

l. Hosnft BrA poAa Allium L. 

B 1991 r. Bo BpeMt rIoe3AKn uo 3anaAsovry Kanrcasy Ha orporax 

Cranncroro xpe6ra 6nua r. flce6afi (KpacHo4apcxr'rfi xpaft) sanan 6rtnn 

co6pauu o6pasqu JlyKa u3 ceKurrl{ Oreiprason F. Herm. \ogpoga Rhizi' 

rideum G. Don f. ex K. Koch. BseurHe or BuAeHHbIx MHoroKparHo pacrennfi 

Allium globosum Bieb. ex Redoute u A. saxutile Bieb. ouu orlturralorcfl 

6olee rpenrvv'l1, vI ToJIcrbIMI4, rollqeo6pa3Ho I'I3BI{TbIMI4 B cra- 

4UU 6yrOUn3auHu, cI43bIMI4, KaK lI JILICTb', CTpeJIKaMI4 I4 MeJIKHMI',I pO3O- 

BbtMt4 rIBerKaMI4 C XeTTbIMI,I nbIJlbHHKaMI,I. B3flTbIe HaMU JIyKOBI{IIbI 

6nnu srrcaxeHbl Ha yqacrKe 3roloro-6oraHnqecKofi craHI{I4I4 6oraHuqecKoro 

LrHcnryra uu. B. JI. Kouapona PAH n flaruropcxe, rAe HerlpepbrBHo 

KJroHr,rpylorcq Ao cHx uop. Bce orMeqeHHble upr'r c6ope Marepuana 

[pu3HaKr4 coxpaHl{Jlr{cb B Kynbrype Ao Hacrotqero BpeMeHrr' Eltlo 

orreBr4AHo, trro Artfl CenepHoro Kasrasa (u ncero Kanra:a) ero HosHfi 

TaKCOH. c rlenlrc ycraHoBllTb, fBrterc{ JII4 oH HOBbIM AJlt HayKU, MbI 

rrpocMoTpeJru rep6apHufi ltarepual c 6oJlee 3arla,[,H],lx reppuroprafi . 

I,Is Kpnrvra A. A, BseaeHcKr4M (1935) 6rur ouucaH A. mqrschallianum 

Vved., xapaKTepl43ytouqufrcr, B orJlfiqfie or A. globosum u A. saxatile, 

6onee ueKuMu pa3MepaMlr, xeJlroBarblM orreHKoM JII,IcrotrKoB oKoJrouBerHr{Ka 

u xeJITbrMH rIbrJrbHLIKaMu. flo nocleAHeMy rlpn3Haxy uce6aficKafl 

rrorryJrf,rrug c6Jll{xaercg c A. msrschallianum, orJll{qatcb or Hero 

6onee rpy6uMH, ToJIcTbIMI{ I4 BbIcoK}rMrI crpeJlKaMu, KoJlblleo6pa:uo 

rr3Br4TbrMn n $asy 6yrouu:aqnu, u apealloM. B creuqx rora Poccnu o6u- 

Taer raKxe A. savranicurz Bess., 6nusxuft r A. marschallianum I{ He Bceuu 

6oraHuraMu rrprr3Hanaeurril. 

Tarcuur o6pa3oM, rlprr3HaKu uce6aficroft norlyJrturll4 ryKa sBrflIorcf, 

flpoMexyrorrHbrMu MexAy A. globosum u A. saxatile, c oAHofi cropoHbt, 

u A. mar schallianum, c .qpyrofr . Buurecra:asHoe [o3BoJlter paccMarpu- 

Barb ee KaK oco6brfi TaKcoH B paHfe BuAa, 

Allium psebaicum Mikheev sp. nova (sect. Oreiprqson F. Herm.). - 

Plantae bulbis oblongo-obovatis, aggregatis, tunicis brunneis, coriaceis. 

Scapi laevi, aeque ac folia cum pruina glauca, alabastris formandis 

'circinatis, sub anthesin erecti, ad tln-tlupartem foliati, foliis lineari- 

96

filiformibus, teretibus, sulcatis. Spatha partis binis basi subdilatatis, 

rostratis, rostro partis longioris umbellam superante. Umbella globosa, 

pedicellis 3-7 mm lg., flore aequilongi vel sesquilongioribus. Perigonium 

roseum, globoso-campanulatum, phyllis parvis, concavis, ovatis vel late 

ovatis, acuminatis, deflexis, exteris 3 mm lg., 2 mm lt., internis 4 mm 

lg., 3 mm lt., nervo medio prominente laete coloratis. Filamenta rosea, 

5-6 mm lg., perigonio sesqui- vel subduplo-longioribus, antheris 

flavis. 

T y p u s: Caucasus Borealis. Prov. Krasnodar, haud procul ab urb. 

Psebaji, l0 VIII 1991, A. Mikheev (LE!). 

Affinitas. Species nostra A. globoso Bieb. ex Redoute et A. saxatili 

Bieb. affinis est, sed antheris flavis, perigonii phyllis minoribus, ovatis vel 

orbiculare-ovatis (nec ovatolanceolatis), roseis coloratis differt. A specie 

A. marschqllianumYved,. pedicellis longioribus, floribus roseis (nec roseis 

aliquantulum flavescentia), area geographica (Caucasus Borealis, nec 

Tauria) dignoscitur. A speciebus omnibus scapis magis crassioribus et 

rudibus, aeque ac folia cum pruina glauca tectis, alabastris formandis 

circinatis. sub anthesin erectis differt. 

Area geographica. Caucasus Borealis, jugum Skalisty, in systemate 

fluminis Laba. 

Habitat in declivibus siccis fruticosis. 

Jlyxoruqu rrpoAoJrroBaro-xfiqenu4ubte, cKyrreHHbre, qeuytr Koxl4c- 

Tbre, Kopr4qHero-6ypue. Crpelrn nprMbre, B rrepr4oA 6yroHu:aqnn KoJrbqeo6pasuo 

l,I3orHyrble, rJtaAKI,Ie, KaK IE Jllrcrbr, c oIt3bIM H:uIeroM, B HI{xuefi 

verrepru o6lucrnenHbre. Jlucrbr ;rnneftHo-Hr,rreBr,rAHble, BarbKoBarrre, 

6oposgqarbre. 9exol H3 AByx vacreft, B ocHoBaHr,r[ HecKoJrbro pacurnpeHHbrx, 

c Hocr,rKaMI,I, Hocr,rK 6onee ArurnHoft qacru [peBbrrlraer 3oHrur. 

3oHrux urapoBr.rAHbrfi; qaeroHoxxu 3-7 MM AJr., paBHbre rro AJrr4He 

uBerKaM ulv s 1.5 pa3a r,rx [penbruraK,urue. OKoloqBerHr.rK posonufi, 

uapoBl,IAHo-KoJroKoJrbqarufi, nuclorrKil ero MeJrKHe, rorHyrue, afiqe- 

Br4AHbre nJu4 uupoKoafiqeau4Hue c ororHyrbrM 3aocrpeHueM, Hapyx- 

rrr,rHoqHbre HuTr4 po3oBbre, 5-6 uu.4rr., BABoe IUrnHHee oKoJrouBeTHrrKa. 

flunruuru xeJrrHe. 

T u n : Ceoepuufi Kaaral, Kpacno4apcrufr rpaft, B oKpecrHocrrx 

r. flce6aft, l0 VUI 1991, A. !. Mnxeen (LE!). 

PoAcrso. Haur uoeufi sr,rA 6rusor x Allium globosum Bieo. ex 

Redoute u A. saxatile Bieb., uo orluqaercr or Hrrx xeJlrbrMr4 [brJ]bHrrrauu, 

6olee MeJrKHMrlr rfiqeoz4Hrruu r4Jll,r oKpyrJro-gfrqesr4AHnuu (a He 

sftuesr4AHo-nanqerHuuu) nl4croqKaMr,r oKonorlBerHr4Ka, po3oBofi r,tx oxpacrofi; 

or A. marschallianum ornnqaercq 6onee gnuuubrMr,r rrBeroHox- 

KAMI4' PO3OBTTUU, 6CT XCJITOTO OTTCHKA, JII4CTOqKAMU OKOJIOIIBCTHI4KA 14 

reorpaQu.recKr4M pacupocrpaHeHrreM (Ceaepnufi Kaaxas, a ue Kptrna); 

OT BCCX BI,IAOB - 6OICE TPY6UUU }I XCCTKHMI,T CTPCJIKAMI4, IIOKPbITbIMI,I, 

7 3arar Ns 915 97

KaK ll Jrlrcrbt, cLI3t'IM HareroM, B fleplroA 6yroHnsaquu KoJlblreo6pa3Ho 

n3o f HyTbrMH, [pl4 I{BeTeHI{}t flptMocTofl quMl't. 

feor-paSn.recKoe paclpoc.TpaHeHl,Ie. CenepHuft Kalras, Cra'rtncurfi 

xpe6er, s 6accefiHe peru JIa6a. 

O6r,rraer Ha cyxltx, IloKpblrbtx KycrapHI{KaMu, cKJloHax. 

2. Honufi BrA poAa Gludiolus L. 

B 1989 r. Bo BpeMfl oxcrypcrafi ro ceBepHbIM cKJIoHaM Mypry:lcroro 

xpe6ra r ApueHuu Ha 3aKycrapeHHblx Jlyrax 6JII{3 r. Eepu (l44xeraucxnfi 

paftoH) rrau o6parnlltt BHITMaHI{e Ha AoBoJIbHo o6I'IlbHo qnaryqufi 

Br,rA rilrraxHl4Ka, Koropbfit npttu'.nu cHaqala sa Gladiolus communis L. 

Co6pauuue rcny6nelyrcosnult 6urI,I BblcaxeHbt Ha lKctlepl4MeHriulbHoM 

yqacrKe 3roloro-6oraHl,IqecKoft craHIILtu EoraHuqecroro I{HcrI'ITyra I4M 

B. JI. Korvrapona PAH r r. flaruropcre. Bolee BHHMareJlbHoe H3yqeHue 

BbrpaureHHblx pacreHl4fr rIoKa3aJIo, qro lrx nplI3HaKI't He yKJIaAbIBaH)'l'ca 

B paMKrr rlprr3HaKoB G. communis, o6urarculet'o n Enpotte (Hamilton, 

1980). Ha Kasra:e urupoKo pacrpocrpaHeH A,pyroft BHA I13 31'oro poAcrsa 

-- G. caucasicus Herb., xoroprtfi o6ut'rHo trpl4Hl{Miulcfl 3a G. commu' 

nis. G. caucasicus HaM orleHb xopotxo 3HaKoM rlo MHoroq[cJIeHHbIM lloe3AKaM 

nO cenepuouy Kanrasy, a raKxe llo Ha6JuoAeHI',ItM B sKclrepu- 

MeHTe Ha yqacrKax ynovxHyroft BbIIle craHquu. floerouy 6luo oqenuA- 

Ho, qro Haur c6op u3 I,1A)(eBaHcKoro paftona Apueuun He rIpKHaAle)I(I4r 

K 3TOMy nU4y. BCe CKa3aHHOe BbIIXe [pUBeJIO HaC r HeO6xoAuMocrH 

o[ucaHl,rs HoBofo BuAa. 

Gladiolus murgusicus Mikheev sp. nova. -- Plantae bulbis 2-2,5 cm 

diam., tunicis coriaceis, solutes bulbillis accessoriis, numerosis cingens, 

caulibus erectis, 60 80 cm altis, basi vaginis 2 supreme I vestitis 3foliatis, 

foliis inaequinervis, 2 3.5 cm lt., apice obtusatis. lnflorescentia 

laxiuscula, unilateralis, 8-10 cm lg,, 6-8-florae, bracteis ovato-lanceolatis 

vel oblongis. Perigonum 38-45 mm lg., laete roseum, perigonii tubo 

incurvo, laciniis campanulatim subconniventibus lobo medio superiore 

orbiculare-obovato, 25 mm lg., 19 mm lt., lateralibus superiore orbiculare-rhombeis 

22 mm Ig., 17 mm lt., lateralibus inferiore obovate-cuneatis, 

18 mm lg.,12 mm lt., medio inferiore orbiculare-rhombeis 20 mm lg., 

15 mm lt. Filamenta 20 mm lg., antherislO mm lg., stylo 37 mm lg., 

stigmatibus a basi ad apicem sensim dilatatis, stigmatis ramis ad 5 mm 

lg. Capsula obovata, l5- l8 mm lg., seminibus ferrugineis, angularibus 

vix compressis et alatis. 

T y p u s: Armenia. In pratis fruticosis in adjacentibus urb. Berd, 20 V 

1989, A. Mikheev (LE!). 

Affinitas. Nostra species G. communi L. affinis, sed foliis latioribus 

(2 3.5 cm, non 0.5-1.5), apice obtusatis, inflorescentia nunquam ramosa, 

pauciflora (6-8, nec l0 l2-flora), perigonii tubo valde incurvo et area 

98

geographica differt; a G. caucasico Herb. inflorescentia laxior, perigonio 

patenti, floribus laete roseis (non atropurpureis) discrepat, 

Area geographica. Adhuc tantum in adjacentibus urb. Berd (Armenia) 

notum est. 

Habitat in declivibus fruticosis montanis. 

PacreHue c rly6ueryroBurraMu, rloKpbrrbtMu Koxr{crblMl{ pacuetl- 

JreHHbrMH qeuyrMl,t 11 MHofortI4cJleHHbrMH oKpyxarcuuMr4 r4x,q,eTKaMr4. 

Cre6ell aucoxnil, 60- 80 crra Bbrc., c 2 veuryeau4HbtMr,r Jrr4crbflMr.r B oc- 

HoBaHr,ru u I nepxunu; nucrbfl c pa3BurbrMtr JttrcroBbrMr,r [[acrr{HKaMrl 

B qucJre 3, 2 3.5 cM rul4p., c xr,rJrKaMr4, B cepeAr{He rrJracrr4HKtr cABr4HyrbrMr.r 

K o4Hofr cTopoHe, Ha BepxytxKe TyrroBaTbre. coqnerue oAHocTopoHHee, 

8 l0 crra Arr., c 6 8 uBerKaMu. flpnqnernuru rffuenuAHo-JraHuel'- 

Hsre. OxoloqBerHnK 40 45 uvr AJr., flpKo-pososnfi, c corHyroft rpy6roff, 

AoJrrl elo KonoKoJrbqaro rrepeKpblBalouluecg, orru6 uepxHer.o cpeAHet o 

Ju,rcroqKa 25 narrl 4n., 19 MM rrrup., BepxHnx 6orosnx 22tnnr 41., l7 nrra 

rrrup., Hrl)KHr{x 6or

K y n pr ur'o,B a f. Jl. O6sop,nI{AoB poga .Alliwn(Alliacvae'S Kasraim'/l.Fotr 

xypH. 2001. T. 86, I&,4. C. I t9-132r ,. :,,, 

H a m i l t,o n A. 'P. Glcdiolus L, ll Floyg Europaea. Carnbridge,.1980, ,Yol., 5. 

P. l0l-102. . . .. . 

EoraHu.recK[ft r,rHcruTyr l{rvi. B. JI. KorrrapoBa PAH" 

Sroloto-6oran[qecKat crdHunt 

: 

'r 

357506 llrruropcK, re/r[aitc (87933) 21762 

E-mail: stbot@yarldex.ru 

i: ' : 

! ,

A. fI. CepernH A. Seregin 

HOBbIE KOMFI4HAUI,II,I B POAE ALLIUM L. (ALLIACEAE) 

COMBINATIONES NOVAE IN GENERE ALLIUM L. 

(ALLIACEAE) 

Jlerovr 2003 r. B cBr3n c peBu3ueft poga Allium L. uu roceruJrr{ ABa 

rpyuHefiurnx fep6apna Beluro6puraHun: fep6apraft KopoJreBcKoro 

6orasuqecroro caAa s Knrc (K) n fep6apr,rfi Epr,rrancKoro My3er ecrecrseHHofi 

r{cropr4rr (BM). LlsyueHr4e cpeArr3eMHoMopcKrlx Br4AoB Allium, 

xopolIIo [peAcTaBJreHHbrx B 3THx KoJUIeKuI,Iflx, IIO3BOJIIIJIO HaM [peArIOxI,ITb 

HecKoJIbKo HoBbrx rovrbuuauufi H HatsaHrrfi. B 6olluruucrBe cryqACB, 

3TO BO3BCACHUC B PAHT BTTAA HCKOTOPbIX TIOABI,IAOB, IIPCACTABJIf,- 

Ioultax H3oJrltpoBaHHbre rroflynf,rlr{u c xopouo BbrAepxaHuuuu uopSo- 

JrofurrecKuMr,r rrpu3HaKann. fIoAsuAbI, otr[caHHbre rro [prr3HaKaM, He 

flBJrflroulr{Mct, Ha HaIIt B3fJIg.q, cyqecTBeHHbrMr{ AJrfl cr,rcTeMaT[Kr4 JryKoB, 

rrepeBeAeHbr B paHf pa3HoBrrAHocTn. 

,{na rynruroupyeMbrx BuAa -- A. kurrat (xyppar) u A. ascalonicum L.o 

1762, non L., 1756 (uranor), Mbr upr.rHr4MaeM B KaqecrBe pa3HoBr.rAHocreir 

A. porrum (nyr-nopeft) u A. cepa (nyx peuuaruft) coornercrBeHHo. 

Ocnosauux Alx uoAo6Horo perueHfir AeralbHo paccMorpeHrr E. Msrrro 

(Mathew, 1996). 

AlliumL. 

Sect. Allium. 

Allium aegyptiacum (Tiickholm et Drar) Seregin comb. et stat. nov. - 

A. curtum Boiss. et Gaill. subsp. aegyptiacum Tiickholm et Drar, 1954, 

Fl. Egypt 3:69. 

A. durandoi (Batt. et Trabut) Seregin comb. et stat. nov. - A. sphaerocephalon 

L. var. durandoi Batt. et Trabut, 1884, Fl. Alger. Monocot. : 

155. - A. sphaerocephalon subsp. durandoi (Batt. et Trabut) Duyfies, 

1976, Revis. Allium Afr. : 50. 

A. junceum Smith var. tridentatum (Kollmann, N. Ozhatay et Koyuncu) 

Seregin comb. et stat, nov. - A. junceum Smith subsp. tridentatum 

Kollmann, N. Ozhatay et Koyuncu, 1983, Notes Roy. Bot. Gard. Edinb. 

41,2 :248. 

A. parviflorum (Kollmann) Seregin comb. et stat. nov. A. proponticumstearn 

et N. Ozhatay var. parviflorumKollmann, 1983, Notes Roy. 

Bot. Gard. Edinb. 41, 2 : 264. 

A. phanerantherum Boiss. et Hausskn. var. deciduum (Kollmann et 

Koyuncu) Seregin comb. et stat. nov. - A. phanerantherum Boiss. et 

Hausskn. subsp. deciduum Kollmann et Koyuncu, 1983, Notes Roy. Bot. 

Gard. Edinb. 41,2 :263. 

l0l

A. porrum L. var. kurrat (Schweinf' ex K. Krause) Seregin comb' et 

stat. nov. - A. kurrat Schweinf. ex K. Krause, 1926, Notizbl. Bot. Gart. 

Berlin 9:524. 

A. sanctum Seregin nom' nov. -- A. curtum Boiss' et Gaill' subsp' 

palaestinum Feinbrun, 1943, Pal. Journ. Bot', Jer' ser' 3: 14, non 

A. palaestinun Kollmann. 

A. sanctum Seregin var. negevense (Kollmann) Seregin comb' nova' 

A. curtum Boiss. et Gaill. var, negevense Kollmann, 1986, in Feinbrun, 

Fl.Palaest.4:397. 

A. vanense (Kollmann et Koyuncu) Seregin comb. et stat' nov' 

A. stearnianum Koyuncu, N. Ozhatay et Kollmann subsp. vanense 

Kollmann et Koyuncu, 1983, Notes Roy' Bot. Gard. Edinb ' 41,2 :267 ' 

Sect. Codonoprasurz Reichenb. 

A. villosiusculum Seregin nom. nov. A. ./lavum L' var' pilosum 

Kollmann et Koyuncu, 1983, Notes Roy' Bot. Gard' Edinb ' 41,2 : 247, 

non A. pilosum Sibth. et Smith, 1809. 

A. staticiforme Smith subsp. flexuosum (d'Urv.) Seregin comb. et stat. 

nov. A. flexuosum d'Urv', 1822,}dtm. Soc. Linn' Paris I : 294' 

A. staticiforme f , flexuosum (d'urv.) zahar.,1975, Biol. Gallo-Hellen. 6: 46. 

Sect. Br ev is p at h a Y alsecchi. 

A. haemostictum (Stearn) Seregin comb. et stat. nov. A. callimischon 

Link subsp. haemostictum Stearn, 1978' Ann' Mus' Goulandr' 4 : 154' 

SecI. Cepa (Mill.) Prokh. 

A. cepa L. var. schallot Seregin nom. nov. -- A. ascalonicum L', 1762, 

Sp. Pl., ed.2:429, nonL.,1756, Fl. Palaest. : 17. 

Anrop 6narogapur Kopolercrufr 6oraHuqecxwit ca[, s KIIo (Berrraxo6pnraunr) 

u SuHancoBylo troAAepx(y yqacrl4r n MexgyuapoAnblx 

*ypiu^ rep6apHoro Aera, [poBoAl4Brxrrxcfl 3Aecb B KIoHe - u'ole 2003 r'., 

qro Aalo Bo3MOXHOCTb BbIrloJIHIlrb Hacroqulyrc pa6ory' 

JlureparyPa 

M a t h e w B. A review of Allium section Allium. Kew, 1996. 176 p' 

Mocroecxufr rocyAapcrBennufi yHnoepcuTer 

urra. M. B. JlouottocoBa, reorpatbuqecKl4ft $axylrrer, 

xaQegpa 6r.roreorpaQuv,, reP6aPwir 

Mocxra, 119992, BoPo6reru foPu 

E-mail: allium@hotbox.ru

A. [,. Mnxeen A. Mikheev 

KPLITI4IIECKLIE 3AMETKI4 O BI,IAAX POAOB 

ORNITHOGALUM L. (HYACINTHACEAE) 14 EREMURUS BIEB. 

(ASPHODELACEAE) OJrOPhr KABKA3A 

NOTAE CRITICAE DE SPECIEBUS Gf,Nf,RUM 

ORNITHOGALUM L. (HYACINTHACEAE) ET EREMURUS 

BIEB. (ASPHODELACEAE) FLORAE CAUCASI 

B Hacroxqefi craue Aalorcr Kpr.rrr{qecKr,re 3aMeqaHur rro AByM Brr- 

AaM poAoB Ornithogalum L. u Eremuras Bieb. Snopu Kanxaga. !oxa- 

3brBaercr, uro Ornithogalum magnun Krasch. et Schischk. roxAecrBeH 

O. arcuatum Stev. flpranoAflTcr AaHHbre B rroJrb3y caMocrorreJlbHocrr,r 

HeAaBHo orrr4caHHoro su4a Eremurus zangezurica.s Mikheev. 

1. O nuaax Ornithogalum mo,gnum Krasch. et Schischk. 

u O. arcustum Stev. 

B 1935 l'. H. M. KpaureHuunuKoB H E. K. lIIuuKt4H Bo

O. magnum. KpoMe flxrnroptx, [prrBoAtrrcfl TaKXe gtt lpytnn 

tr{. M. KpaureHr.rHHr,rKoBbrM (1935), A. A. fpoccrefiuou (1949) r.r A. C. [IIxuru 

(1969). B fep6apll,r Eorasuqecxoro HHcrr4ryra uM. B. JI. Korraapona 

PAH (LE) r4Merorcr o6pa:qu rrr,rrleMrerrHnna u3 Bopxoruu n Carypaun 

(fpy:ur) r43 rroApoAa Beryllis, orpeAeJreHHue H. {. Ar.auoaoft rar 

O. magnum. Ecnn upuurrb Harrry rorrKy 3peHufl, qro rr{noBofi uarepuan 

\o O. magnum orHocltrcr x O. arcuatum, To raKoe olpeAeleHl4e cJreAyer 

crrr.rrarb HeBepHbrM. Vnouruyrue o6pasqu lrs fpy:ur,r Aonxsrr o6o- 

3Haqarbcr rax O. arcuetumo rrocKoJrbKy Ha3BaH[e O. magnum qBrtre.rct 

cLIHOHI,IMOM o. ArCUAtUm. 

He6eruHrepecHo raKx(e orMerrrrb, qro H. ,{. Arauoaa (1977) cr{[TaeT 

coMHKTeJrbHbrM rrporlr3pacTaHlre o. magnum u flpe4ranxa3be, xoTf oH 

u o[ucaH u3 3Toro peruoHa. Kaxux-lu6o raKcoHoMr.{qecKr.rx BbtBoAoB B 

orHolreHnu rrocJreAHero Blaqa v O. arcuatum oHa He AeJraer. 

2. K noupocy o suAonoil caMocroflTeJrbnocrrr 

Eremurus zangezuricus Mikheev 

B 1996 r. HaMr{ 6un onucau us 3aHrerypa (Aprraenua) nw,g Eremurus 

zangezuricus Mikheev (Mnxeea, 1996), xoporuo naop$onorr4qecKlr u 6pro- 

JroruqecKu o6oco6leHnuft or E. spectablis Bieb. flosftree6*nv rrprlBe- 

AeHbr H 6uoxlrl,ru.recKr,Ie [pr,r3HaKn, orfi[qaroulue Harrr Br{A or uocneAHero 

(Mnxeen, CuupHora, 2000; Mraxees ra Ap., 2003). OAuaxo so (Onope 

Apnaeunn K. f. TaMaHqu (2001) orHecJra Ha3BaHr4e E. zangezuricus 

B craHoHr,rMbr x E. spectab/ls. OcuosaHrleM AJrr cr4HoHr{Mr,r3arlur{ rBJuerct, 

tlo MHeHr{ro TalaaHqu (TaruauaH, 2001 : 386) ro, qro (...r43yr{eHHbre 

...3K3eM[Jrflprr, co6paHuue r.r3 pa3Hbrx Mecr B oxpecrHocrf,x fopnca, ue 

BbrAeJrrJrr.rcb rro yKa3aHHbrM aBTopoM rrpll3HaKaM, 6Onluruncrso r43 Ko- 

Topbrx yKraAbrBarorcq B [peAenbr napua6enluocrrl Bcex apurHcxux o6pa3rloB 

E. spectabilly. Mu He MoxeM cornacnrbcr c roqrofi 3peHr{t 

TauaHrH n rro-[pexHeMy cr{r,rraeM, qro Bce apMrHcrue o6parqu ornoctrcr 

K E. zangezuricus, a He r E. spectabilis n'lro r ApnaeHuu rrocJreAuufi, 

nug He rlpol,r3pacraer. Eo.nee roro, [peAcraBfirercq oqeBuAHbtM, rrro 

r 3aranragle noo6qe He Bcrper{aercs. E. spectabilis, Ho fipor{3pacraror 

Apyfr,re Br,rAbr poAa Eremurus, Bo3MoxHo 6onee 6lurr?re K rrepeAHe- r4 

cpeAHea3r{arcKr.rM BuAaM. Taros E. zangezuricltJ, a raKxe E. azerbaidshanicus 

Charkev. C. C. Xapreeuu (1964 :47) ornevaer:

MexAy co6ofi rro Hiuruql,rrc xeJrroBaro-6elonaroro oKoJrorrBerHr,rKa I,r 

uo.rarroo6pa3Horo (a He xnocroau4uoro) couBernfl B cra1L4r4 6yroHra- 

3a\uu. floerouy MbI fioJrHocrblo corracHbr c [pr,rBeAeHHbrMl{ Bbr[re cJronavru 

XapreBr.rqa o neo 6xoAl,rrr,rocrH MoHorpaQuuecrcoro uccrreAoBa Hr,rt 

3aKaBKa3cKtrx BtrAoB Eremurus, KaK B rrpupoAe, raK r,r B 3Kc[epr4MeHTe. 

BrrssneHr.re llrav'u E. zangezuricus B ropax 3anrerypa [oATBepxAaer MHenue 

Xapretuqa o Bo3MoxHocrtl HaxoxAeHuf, n Apureuuu HoBbrx AJrt 

HayKr,r Br,rAoB sTofo poAa. 

Pa6ora BbrroJrHeHa upu $nnanconofi nog4epxxe poccuficroro 

QoH- 

4a SyH4arraeHrulrrbHbrx HccJreAoBaH r.r fi (npoerr Nb 02-04-49 7 8 8). 

Jlureparypa 

A r a n o s a H. A.Por ornithogalum L. -opunroranroM, HJrn nrnqeuleunur// 

,l,enoparnnHrre rpaBrHr.rcrbre pacreHr,rr Arrq orKpbrroro fpynTa CCCP. jf., 

1977 . T. 2. C. t76-184. 

f poccneftu A. A. Onpe4eru.rrenbpacrenrfi Kasxasa. M., 1949.74gc. 

KpaureHr,rHHr,rKon tl. M. Pog nraqeuleqHrrK- OrnithogalumL. ll 

@aopa CCCP. Jl., 1935. T. 4. C. 379-394. 

M n x e e n A. [. Pe.qx[e H ucqe3aroque BHAbr {noprr CCCP, coxpaHfleMbre 

n flepranrcxov ap6operynre // Eron. fn. 6or. ca.qa. 1984. Brrn. 134. C.43- 

47. 

MHxees A.{.HonufiBr,rApoAa Eremurus cKanxasa ll Bor.xypg. 1996. 

T. 81, Nb 5. C. 79-81 

Mt.txees A.A., CMr.rpHoBa H. 14. OAonorHr,rreJrbHorra 6noxfiruuqecKoM 

npr,r3HaKe B xapaKTepr,rcrr,rKe vnga Eremurus zangezuricus (Asphodela_ 

ceae) ll 6or. xypH. 2000. T. 85, J\b 8. C.92*94. 

Muxe es A.A., CMr,rpHosa H. I4., lllep6yxnn B. [. Co4epxa- 

Hr4e r.r MoHocaxap[4urrft cocraB BoAopacrBopr.rMbrx floJrtrcaxapr.rAoB xopaefi 

HeKoropbrx nngon Ere.muras Bieb., fiHTpoAyrltrpoBaHHbrx r flrrraropcr // 

Pacr. pecypcu. 2003. T. 39, nun. 2. C.69-72. 

T a u a s x n K. f. Poa Eremurus Bieb., opeuypyc // @nopa Apuerutr. Liechtenstein, 

2001. T. 10. C. 38,$-386. 

XapxeBuq C. C. O HeKoropbrx llrleftaux Qnopu Kanrasa // Honocrrr 

cncreMaruKr.r Bbrcrutrx pacrennft. 1966. T. l3l. C. 4147. 

IrI x !r s s A. c. ornithogalum L. // onpe4ernrerrb pacreHuir lpytuu. T6unqcu, 

1969. T . 2. C. 317-31 8. 

EoraHu.{ecrufi.uucruryr uu. B. JL Kouapona pAH 

Sronoro-6oraHr,rqecKar craHuLtt 

357 506 fkruropcr, ren/Qarc (87 933)217 62 

E-mail: stbot@yandex.ru

A. C.3epnon, 

LI. B. Corco;ron 

POA HOLOSTEUM L. (CARYOPHYLLACEAE) 

BO OJIOPE KABKA3A 

GENUS HOLOSTEUM L. (CARYOPHYLLACEAE) 

IN FLORA CAUCASI 

L. Zetnov, 

l. Sokolov 

flpooo4a peBr{3r,rrc Snopu Cerepo-3anaAHor-o Kasxa:a, Mbl croJIK- 

HyJrucb c rpyAHocrsMrr B r.rAeHTl{Quraqnu BTIAoB Holosteum L. us po4clsa 

H. umbellatum L. Pacresur rrofi nonnuopSnofi fpytrllbl uMetor 

x(ere3r4croe orryueHrre rto6efoB u MenKI,Ie [pI4LIBerHuKu 0.5 2 MM AJI. 

Pa4 anropoa (Fenzl, 1842; Ilftvranbtay3eH, 1895; lllnauH-lypni,tcxnfr, 

1928; fpoccrefrrra, 1949; KapxrnH, 1952; Kocenxo, 1970; fanyurro, 1980) 

Ars Kasra:a yKa3blBaer 4 sw\a r43 lrofr rpyIIIIbI: H. umbellstunt, H. glutinosum 

(Bieb.) Fisch. et C. A. Mey,, fI. liniflorum Stev. ex Fisch. et 

C. A. Mey. u H. polygamunt K. Koch. EoruuruHcrBo xe I4ccJIeAoBareJIeii 

npr{BoAtrr tuutr H. umhellatum v, H. glutinosum, cBot\fl, H. polygamunt 

B Cr4HOHr4Mbr K rrOCJreAHeMy BLU\y, a H. liniflOrum ro K H. glutinosum, to 

t< H. umbellatum. Cw'raercr, qro H. umbellatum vueer 3 5 ruqunox; 

JrerrecTKr,r paBHbre r{Jlr,r HeMHofo Anr4HHee qarlleqKl4, IlpI{ ocHoBaHVv pes- 

Hrrrqarbre; r'oJlbre rrpr4uBeTHnKr{, HMerculue no Kpalo y3Kylo IUIeHqaryIo 

xaftrray; roJrbre coqJreHeHtlr uBeroHoxer; ropo6o.rxy 4 6 rrau ArI., a 

H. glutinosum -- l0 TbrrruHoK; re[ecrKr4 s 1.5-2 pa3a AnuHHee qarueq- 

Kr4, rrprl ocHoBaHur4 6e: pecHnveK; xene3r4cro o[yueHHble nprl]BerHH- 

Krr c []r4poKofi flneH.raroft raftuoft I1o Kparo; xene3Hcro o[yueHHbre coqJreHeHns 

qBeroHo)r(eK; ropo6o.rry 6.5 9 urr,r An. Ho pa3rpaHlrqeHue 

H. umbellatum u H. glutinosum rro 3Tl{M rrpH3HaKaM rpeAcraBJlqe]' 

cepbe3Hbre 3arpyAHeHnx. Ilorrorrly Volrepc (Walters, 1964) ro 4la Enpourr [prrBoAl4T H. umbellatum KaK ttotruuop$ultft 

raxcoH c AByM{ rroABraAaMrr -- - subsp. umbellatum r.r subsp. glutinosunt 

(Bieb.) Nyman. Taroe xe rroHrrMaHne BVua orpaxeHo n pa6orr A. JI. TaxraAxuHa 

ra As. A. (De4opona (1972). KooAe (Coode, 1967), npuzuauan 

norur,roptfu3u H. umbellatum, [pr4Hr4Maer B ero cocraBe 3 pa3HoBnAHocrr4- 

var. umbellatum, var. glutinosum (Bieb.) Gay u var. tenerrinum 

(Boiss.) Gay. Bo 2-M usraLruLa

urvr. M. B. Jlououocona (MW), EoraHuqecxolo l,rHcrr{ryra vM. B. JI. Ko- 

MapoBa PAH (LE), BcepoccuilcKoro HayqHo-r4ccJreAoBareJrbcKor.o r.rHcrnryra 

JreKapcrBeHHbrx u apoMaruqecKr,rx pacrennft (MOSM), flanHoro 

6oraHuqecKolo caAa PAH (MHA), Ky6aHcroro focyAapcrBeHHor.o 

arpapHot'o yHuBepcr4rera (KBAI), Mocroscxoro ueAarolnqecKor.o [ocyAapcrBeHHoro 

yHr{Bepcnrera (MOSP) u Pocroncroro-na-floHy focy- 

AapcrBeHHoI'o yHnBepcr4Tera (RV), Mbl npr{trrJlt4 K 3aKJrIoqeHuK), qro trrupr4Ha 

flreHqarofi oxpauHbr IIpt{uBerHr4KoB, a raKxe orryrxeHr{e ocHoBa- 

Hrrf, JrerrecrKoB He r{Meror raKcoHoMr,rrrecKofo 3HaqeHr4f,. Cpe4uee xe 3HaqeHue 

A[r4Hbr ropo6o.rxu yBeJruquBaercr u Enpasur c ceBepo-3arra1a Ha 

Ioro-Bocror, 14 3To I{3MeHeHI4e sBHo HocIlT xruuanrgufi xapaxrep. Tax, 

n 3ana4Hofi Eepone cpeAHrs AJrr,rHa ropo6o,rxu 5.41 rvru (otrrn6xa cpeguero 

apuQuernqecroro (m") = 0.39), n Enponeftcxoit qactu 6usruero 

CCCP (6er Kpurr.ra) 5.67 narra (m, = 0.23), n Kpurury 5.65 rrarr,r (m, = 0.2), 

Ha Kasrase 6 urra (m" = 0.26) u n CpegHefi Azuu 6.07 uu (m" = 0.68). 

flpoqeur oco6efi c [e[ecrKaMu 6olnuefi AJrr4Hbr, rreM qaueruc.l.HK]r, orryrrreHHbrM 

ocHoBaHueM uBeroHoxeK u l0 rnqraHKaMr4 yBeJrr4q[Baercq B 

ToM xe HarrpaBreHr4l,r, flpusuaru 3Ttr rlMeror AoBoJtbHo 6onturofr ro:Q- 

Ququeur KoppeJrrur4u -- 6onee 0.8, c nucoxr,rM ypoBHeM HaAexHocru. 

Ha rnugatrluuft xaparrep [3MeHqr4Bocrr{ KocBeHHo yKa3brBaer. n 

BblcKa3brBaHne E. K. Lllnurrusa (1947) o roM, rrro rraBHbrfi apean 

H. glutinosum Haxo1r4Tcfl B HH3oBbrx Bolru, Cpe4Heft Azuu u 3arasra- 

3be, a B Kpulay 3To peAKoe pacrgHr{e, B orflHrrue or H. umbellatunt. 

Kluna flBuercr, noxanyfi, cauofi cnoxHoft curyaqnefi Arfl HoMeH- 

KJlarypHbrx peueuufi. 34ecl 4ouycrr,rMbt pa3Hbre rrollxoAbr. MoxHo upr,r- 

Hrrb c?rMrrarpuqecKoe [por,r3pacrasue H. glutinosunt u H. untbellatum. 

Moxuo, yqr4TblBar, qro Ha Kasra:e upeo6ragarcr oco6n c [pr43HaKavu 

H. glutinosum, rrpuHqrb Alrr aroft Teppr,rroptir4 Jrnub oAr4H TaKcoH 

IloABlrAoBofo paHr.a. Ho, Ha Hatu B3fJrrA, 60nee [paruJrbHblM f,BJlserct 

[pu3HaHue uoluuopSuocru H. umbellatum, c BbtAerreHueM B ero cocraBe 

AByx pa3HoBngHocreft: var. umbellatum v var. glutinosum. 

H. glutinosum oluapogonaH (Fischer, Meyer, Ave-Lallemant, 1840) 

KaK rrpflMa.s 3aMeHa orrr4caHHoro pauee H. liniflorum (Fischer, Meyer, 

Trautvetter, 1837). Hasnasue H. liniflorum Stev. ex Fisch. et C. A. Mey., 

1837, npu ouy6nnronasuu 6rmo He3aKoHHbrM KaK r43Jrr{lxHee HaHMeHo- 

BaHUe, TaK KaK B KaqecrBe cr.rHoHlrMa K HeMy 6ul npune4eu Arenaria glutinosa 

Bieb.3arueuue H. liniflorum, nom. illeg. ua H. glutinosum., aBTopbr 

co3Aiunr HoByro rou6uuaquro (cu.: fpeftrep r{ Ap., 2001, vrarnx33.2). 

Tar

ltefi>1, r.r rro3ToMy cJreAyer paccMorperb o6a AuafHo3a, corJracHo Koroprrvr: 

^F1. glutinosum: (crasoe; BepxHrle rlrcrbfl. rroqrr.r sftqeslrAHrre, B ocHoBaHHr4 

cpocrxfiecfl; JrerrecrKn rrpoAoJrroBarbre, rroqru uelrHorpafrHbre, rro AJln- 

He r{ rrrr4pr4He eABa rrpeBbrrlalor qarueJr[crrrKu). H. liniflorum: (npuqaernuqKlr eABa 

orafi unesurre, rIBerKr.{ Aecrrr{Tbrrr[uronue). Taruu o6pa:ou, creAyer 

rlpl43Harb roucnequSuqHocrb H. liniflorum u H. glutinosum. HaEo o'r- 

Merr{Tb, .tTo otll{caHlrt o6oux BI{AoB 3aKaHqI,IBaIorc{ 3HaqKoM >. As- 

TOpbI, OqeBHAHO, CqHTaJlU, qTO 3Tl4 pacTeHut I4MeK)T pa3AeIbHOrIOirbIe 

uBerKrd, rrro He coorBercrByer AeftcrBr{TeJrbHocrr4. 3ra ourn6ra, Bepof,rHo, 

ro6yAura Koxa (Koch, 1841 : 708) orucarb H. polygamum v oco- 

6o yrasarr: (qserKra B 3onrrlKax, uHoro6pa.ruue, o6oerrorbre, 

Aecrrr,rrbrquHKoBble, ropo6o.ma c 8 ulu 10 npruuuu sy6vuxaruu), 

(neuecrru MeJrbqe, 

IrBerKu MHoro6parrHrre). {Dpara AI{arHo3a

ueHr,rr uo6eros 14 KpyrrHblMrr rrpuuBerHuKauu, 4 5 MM AJr. 3ror sna xopouro 

o6oco6reH u BbrAepxaH B cBor{x rrpr{3HaKax. 

flo naruuu AaHHbrM, Ha KasKase pog Holosteum rpe1craBJreH ABy- 

Mfl BnAaMU, OALTH l{3 KOTOpbtx I{MeeT ABe pa3HOBI,IAHOCTII. Blt4eleuue 

JreKTOTtrrroB rrpotr3BeAeHo coBMecrHo c f. A. Jlasrrosnr4, [oAforoBr{Brrrr,rM 

pyrorrucr no Caryophyllaceae Anr

Ha Kasxa:e Bcrpeqaercr n CranpouoJlbcKoM xpae, {arecraHe, Bocro.rHofi 

fpyrr.u,r, ApueHnn r,r Asep6aft4xaHe. -- O6u1. pacup.: Malar 

Azun. 

B:axlrcqesne BbtpaxaeM HcKpeHHrorc 6lar.o4apHocrr, H. H. I,Iuxauuqroft 

3a [oMoulb B r]poqreHr.ru anrorpaQa @. P. foreHaKepa Ha rr4noBoM 

o6pa:qe H. marginatunt nT. B. Eroponofi 3a KoHcyJrbl.aqau rro 

HOMeHKJraTypHbrM BorrpocaM. 

Pa6ora BburoJrHeHa upu SuHaHcupoBaHnrl lpanroM flpesu4eura p@ 

Anfl rroAAepxrrr BeAyurux HayqHbrx urxon (Ns HLlI2125.2003.4). 

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Br4Ahr POAA ROSA L. (ROSACEAE) CEKIIIII4 

CINNAMOMEAE DC. BO OJIOPE KABKA3A 

l. Buzunova, 

R. Kamelin 

GENERIS ROSA L. (ROSACEAE) SECTIONIS CINNAMOMEAE 

DC. SPECIES IN FLORA CAUCASI 

B crarre Aaercr o6sop KaBKa3cKl4x BHAoB po4a Rosa L., ttpnuaA.rexaulux 

K cerrrr{u Cinnamomeae DC. ?ra ceKqr4r ua Kasra:e rtpeAcraBneHa 

cpaBHu'reJlbHo He6o;rlruuu q[cJroM TaKcoHoB. flo Harunna AaHHbrM, 

oHa BKnroqaer 8 sr.rAos u 3 HorosuAa. flo cyMMe uopQoloruqecKnx 

rrprr3HaKoB eugn rrofi ceKrrnu MoxHo o6reAuuuru B HecKoJrbKo lpytrn, 

oAHaKo cpeArl Hrrx HeJrb3r uailrn seruft upe4xoauft Tr4rr, rr3 Koroporo 

yAaJrocb 6n arrsecru pflA TaKcoHos c 6oaee clequaJru3r4poBaHHbrMr4 

upr,r3HaKaMr4. O,{esraAuo, qro 3Trr rpyrtrrbr o6oco6unracb B npoqecce 3Bonloulru{ 

He3aBucr.rMo u HeKoropbre u3 Hr,rx BHe [peAeJros Kanxa:a, a [po- 

Hr4Kirr4 TyAa B pe3yJrbrare HeoAHoKparHoro r{3MeHeHr{f, KJrr4Marn,{ecrofr 

o6craHosru pa3HbrMr,r rryrrMr,r r,r B pa3Hoe BpeMr. 

Hau6oree 4perHeft HaM npeAcraBJlrercr rpy[ua Br.rAoB poAcrBa 

R. oxyodon Boiss. s. l. Oua BKJrK)r{aer rpn BuAa R. oxyodon Boiss., 

R. prokhanovii Galushko u R. sosnovskyana Tamamsch. Vxe HeoAHoKparHo 

orMeqaJracb 6rtnsocrl R. oxyodon erpouefrcroMy Br.rAy R. pendulina 

L., rpor43pacraroqeMy r A-rluax u Ha Kapnarax H TaK)r(e upr4Halr,nexaqeMy 

K ceKrrr4rl Cinnamomeqe. Vlx o6'ae$r4HflK)T cJreAylorque [pr4- 

3HaK?I: o6rr.rHo KyBrul4Hqaraa Qoprraa rrJloAoB, rroHr{Karoulux Ha AIHH- 

Hbrx xeJre3r4cTo-qeTl{HtrcTbrx uBeToHo)KKax, AJll4HHbIe qa[IeJrhcTlrKr4 c 

lo[aroqKoBr4AHbrM pacurrpeHr4eM Ha Ko Hue, 3JrJlr,r rrrr4qecKar tpopua rua - 

crorrKoB rr KpacHoBalo-Snoreronafl oKpacKa uo6eros. Kax u R. pendulina, 

R. oxyodon 3Haqr{reJrbHo Bapbrlpyer rro cBor.rM uopQoloruuecKrrM 

Ilprr3HaKaM, Ho BbrAeJreHr,re B rrpeAerax 3Tofo BlrAa MeJlKux Br4AoB 14 pa3- 

HoBlrAHocreft rro.rrr4 He rrpeArrpr{Hr,rMaJrocb, B orJrr4rrrie o'r R. pendulina. 

3ro o6ycnoBJreHo He roJIbKo ueurueft r43yqeHHocrblo rrrr4rroBHr,rKoB 

Kanraaa uo cpaBHeHr.rro c eopoueftcx[Mr,r, Ho H orreBr,rAHofi ruergocrrrc 

TaKr4x rroubrroK. tleu 6olnure MarepuaJra to R, oxyodon [ouaAaer B 

rlone 3peH]rr nccneAoBareJu, TeM 6olrure pa3MbrBarcTcr rpaur4ubr Mex- 

Ay HaMeqeHHbrMrr orAeJrbHocrtMl{, xarne 6rt [prr3HaKrr se 6unu uolroxeHbr 

B ocHoBy 3Toro pa3AeJreHr.u. 

R. pendulina u R. oxyodon orHocrrcr K rrr.rKny Pendulinae Juz. (incl. 

cycl. Oxyodontae Galushko ). .{pyr or Apyra oHr,r orruqarorcr, rroMuuo 

reorpa$nqecxofr [oKanu3auul4, orcyrcrBneM utnuoB n nepxHefr uacrl4 

reHeparr,rBHbrx no6eron y R. pendulina, 'rorga xax y R. oxyodon 

rulrrrbr BcerAa rrpncyrcrByror r,r r4Meror pa:uoo6parHyro Qopuy: upflMbre 

IXI4JrOBr4AHbre fOprl3OHTarbHO O'fKJrOHeHHbre r4Jr[ HarrpaBJreHHbte BBepx, 

tt2

CJICTKA CEPTIOBI,IAHO I43OI'HYTbIC; I4HOTAA KPYTIHbIC ITITTIbI TIEPCMCXAIOTC' 

HrnoBuAHbrMu trrunuKaMn. 3y6uu Kpar n[cror{ror y R. oxyodon o6u.r_ 

Ho upocrbre, Ly R.pendulina cJroxHbre. Ecnu y R. oxyodon KpynHbre, 

HarrpaBJreHHbre BBepx urufibr Bcrpeqarorcs ua no6ere oAHoBpeMeHHO 

c loplr3oHTalbHo oTcTorull,tMu r,IJIu cepIIoBlIAHO l43OfHyTbrMI4, 1.O y 

R. prokhanovt uprEcyrcrByrcT ToJrbKo lrrr4rrbr rrepBoro rr.r[a, xorr orry.'e- 

HHe UBeTOHOXeK n Jlr4cToqKoB y HeIo TaKXe BapbupyeT, KaK u y 

R. oxyodon. Tperrafi nug eroft rpyflrrbr, R. sosnovskyana, orilytqaerc, or 

ABYX UPCAbIAYUIUX BI4AOB IXUTIAMLI BBCPX HATIPABJICHHbIMI,I, rrO O6Ir.tHO 

He crpy[m'rpoBaHHbrMr{ rIapaMI4 y ocHoBaHHr Jrr.rcrbeB ra 60Kosblx no6efoB' 

a paccesHHbtMa uo cte6ln, I,I fycro xeJre3I,ICTO-IqerI{HI{CTbIMI4 qBe- 

TOHOXKaMU H [JrOAaMLr. BO3UOXIO, HecMorps Ha Hzuu{que qubuorpafi- 

HbIX rraurerltcrl{KoB, npu 4anlnefrIueM u3yqeHHlr Ir o[peAelte:f|?']y- XpOMOcoMHoro 

rrncra 3Tor nu4 6y4er orHeceH K ceKrrllr{ Caninae DC., xar:ro 

rlMeno Mecro c R. glauca Pourr. Hanuque crpy[[LrpoBaHHbrx B ocHoBa- 

Hr.Ilr JrHcTbeB u 60rosmx uo6eron, a He paccerHHbrx fro cre6flro KpynHbu 

urnnoB ( uo He MeJrKr4x, rcar y R. acicularis Lindl. u R. oxyacantha Bieb.) 

rBrrerc.rr 6onee unu MeHee BbrAepxaHHbtM rrprr3HaKoM y [pegcrarurelefi 

ceKur'ru Cinnamomeae. Ha cxoAcrso R. sosnovskyana c HeKoropbrMr,r BH- 

AaMr{ ceKrlLtu caninae BuepBbre ooparula BHr{MaHue aBTop oru]caHr4t 

3Toro raKcosa C. f. Tauauurrn. 

flpn:uaru - BBepx Ha[paBJreHHbre trrurrbr u KyBtur{Hrrara_r Qopua 

ruroAoB Bcrpeqalorct AoBoJrbHO qacro y HeKoropbrx BocroqHoa3uarcKrlx 

Br{AoB, Ha[puMep, y xurafrcrux po3 ceKrlr,il,r Cinnqmomeae: R. webbiana 

Wall. ex Royle, R. giraldii Cr6p,, R. forestiana Bouleng. n 4p. B cny- 

.rae c R. pendulina u R. oxyodo,? Mbr r4MseM eule oAuH rrpnMep Bo3Mox_ 

HOCTr{ AByx uyrefi pasaurraa @nop u cnarapulux r{x Br,rAoB: l) anroxrouuhlfi, 

xor4a B ocHoBe BocrorrHoa3llarcKrrx, KaBKa3cKr4x n enpoueficrux 

Br.rAoB Jrexr4T oAHH ApeBHnft, ruuporo paclpocrpaHeHHufr nu4 (c apea_ 

JIOM TIOqTII TAKIIM XC O6UTUPHUM, KAK Y COBPEMCHHOIO BI4AA R. SPiNOsissima 

L. ), roropufr co rpenaeHerra rpanc$opMrrpyercr B pa3Hbrx qac- 

T'X APC'UIA B COOTBCTCTBI4Id C MCHflIOIqYIMVICfl, 

YCJIOBI4'MII CYXIECTBOBAHI4-fl 

c o6pasosaHr4eM cepr{r4 BrrAoB rroJrr4TorrHoro rrpoucxoxA eH:zt, u 2) uurpaquounHfi' 

flpu Koroporr,r o6u1ue [pr{3HaKrr 6uru npurueceHbr c rrpo- 

ABI4XCHI{CM HA KASTAS I,I AUUICC r ENPONY BI,IAOB, I{MCUIII4X BOCTOT{HOruraficxne 

KopHr4. flpu rr,fl.rrpaqr4oHHoM rryrr,r pa3BHTuq Br4AoB poAa 

Rosa Aonxnrr 6unn 6rr coxpaHnrbcr BHAbr-Bexu srofi Murparrr4rr, oco- 

6euHo a Malofr Asuu, rAe Ao cux rrop [pou3pacraror BHAbr ApeBHux 

rpyrlrr [ru[oBHrrKoB: crrHcrrrJrburre R. phoenicia Boiss., R. sempervirens 

L. n 6eloqaerKoBbre r{3 poAcrBa R. beggerana Schrenk, ,ro ,rpa4"ru"r_ 

TeJrH CeKrrl[r cinnamomeae raM orcyrcrByrcT. B nolwy aBToXTOHHOIO 

pa3B'dTldl BTTAOB roBoprrr z Qarr, Ha xoropufi o6parula BHUMaH?re 

I,I. fI. Mau4euona (1969 : 50) -

uroro Kaercasa. HNrelou1laecs B nl,ITeparype yKa3aHI4t Ha IIpoLI3pafl'aHI'Ie 

ero Ha Malou Kasrase He noATBepAnIrvtcb,. . r'ep6aputte o6pasqlt' Ha 

ocHoBe Koropbrx fror BKA yKa3blBaJlcq A"rtq Matrol-o Kasxasa , 6unn HerrpaBr{JrbHo 

o[peAeJIeHbI>. C nocneAHI{M yrBepxAeHueM I]oJIHoc'I'bIo corJracurbcfl 

HeJIb3f,, TaK KaK nlraeruult,tftcs n fep6apun Eotaunqecroro 

r.rHcrr4ryra (LE) o6pa:eq us Barypuanra (Ma.nufr Kanras) orHocl4lcfl K 

R. oxyodon, Ho HyxHo ITpI,IHI{Marb Bo BHI'IMaHI{e cKopocrb, c xoropofr 

paclpocrpaHttorct po3bl B HacTOtXIee BpeMq flpu HerIocpeAcrBeHHoM 

yr{acrur.r qeloBeKa. E6lruraq )Ke qacrb apeara R. oxyodon, geftcrnurenl- 

Ho, pac[oraraercf, s o6racru Eolluroro Kanrasa, 

K quxny Pendulinae, rto HaIxeMy MHeHnIo, orHocl4Tcfl eure oAI{H BI4]r, 

ouucansuil c,Il,oueuxoro Kptxa O. H. ly6osnr (1966 : 169) R. donetzica 

Dubovik. HssecrHo HecKonbKo ero MecroHaxoxAeunfi s AoHeuxoii, 

Pocroscrofr u AcrpaxaHcxofr o6lacrsx. Kar ?I Apyrl{e Bu.ubl frofo ql{Kta, 

R. donetzica uueer KpyrIHbIe rlptMble url'IIIbI It KyBuluHr{aryro Sop- 

My rlJroAoB, Ho flBfif,ercfl KapnHKoBbIM Kycrapu[Kol'r srtcorofi 40 80 cu' 

CneAypulaq rpy[fia BI{AoB - 1To BlrAbl poAcrna R. maialis Herrm. 

s. l. JlureparypHl,Ie AaHHbIe o roM, ecrb na Ha Kasxase rulrflr{Hat 

R. majalis, [poruBopeqnnn. 9eu AerilrbHee n3yqanacb Qnopa Karxa3a, 

TeM MeHee cKJroHHbI 6nnu uccleAoBareJll'I orHocl{Tb safrAeuHlte nuu 

pacreHnfl K 3roMy rr.r4y. B pa6ore A. O.

na BHr.rMaHHe O. H. fiy6onnr (in sched.), orpeAerl,tB HX KaK R. pratorum 

Sukacz. fIoA raruu xe Ha3BaHrIeM oHll BoIuJuI a o6pa6orry po4a Rosa 

ro > (EysyHona, 2001). R. pratorum qau\e 

Bcero oroxAecrBflslor c R. glabrifolia, t

suA 6rusox R. gorenkensis Bess., oco6euuo ero [peAcraBHTeJLflM I'I3 Bocro.ruoft, 

Ka3axcraHcKofi qacru apeara, Bosuoxuo, oH tBJrterct pe3yJlb- 

TaroM LlHTporpeccurHofi ru6pNAn3a\r4v auga, 6nusroro R' maialis ua 

BrrA rr3 rroAceKlrun Vestitue Christ ceKltrI{u Caninae DC., xar u R' gorenkensis. 

C cenepHnx cKJIoHoB I-l,eurpatuHoro KaBKa3a A' lI' fanyuKo orll4ciul 

eure ABa ru4a, 6.uu:rux R. majalis -- R. balcarica Galushko u R. obtegens 

Galushko, BbrAenlrB Hx B orAerbnltil quxn Balcaricae Galushko. O6a 

oHn oTJlr4qarcTct IIII{pOKO3IJII4ITUqeCKIIMII AO OKpyfJIblX II4CTOqKaMU H 

KpynHbrMlr IrpLIuBerHLlKaMU, rlotrrl4 cKpblBaIouII4MlI KOpOTKI4e qBero- 

HoxKrr. Y R. balcarica segovra neo6u'tuas AJII BI4AoB ceKql{u Soprvra 

111urroB: KprorrKoBuAHo tl3orHyrbte c HalrpaBJleHuofr BHIr3 rlapaJuleJrbHo 

Cre6lrO aepxHefr qaCTbIO. EuIe raxne-lU6O COnpeueHHble uIuIlOBHl{KIl C 

TaKr{Mkt rurruaMl4 HeIr3BecrHbI. ITlnporoelll4tlTl4qecKar go oxpyrlofr 

Sopua rucrotrKoB AoBoJIbHo IuupoKo rlpeAcraBleHa y KaBKa3cKI{x Bl4- 

AoB ceKur4u Caninae. BHe KasKa:a sTor rIpH3HaK AoMI{HlIpyer y a3ua'rcKrrx 

ruurroBHrlKoB ceKuun Cinnamomeqe. 

Huxe uu [pI{BoAItM o6:op BI,IAoB ceKuul4 Cinnamomeae poga Rosa 

Qnopu Kanra:a t{ KJrrcq AJII I{x o[peAereHl4a. PacupocrpaHeHl4e BUAoB 

Ha reppnropur4 Kasrala Aaerct corrracHo pafiouuponaHlllo' [pl{HtroMy 

n pa6ore IO. JI. MeHuunoro (1991), a o6ulee pacrpocrpaHeHue - IIo 

paftoHau, [pnHtrbtM B r431a*;z,la

Jlncro.rxn rto KpasM c npocrbrMr,r 3y6qauu, r.rHorAa orAeJrbHbre sy6qrr c 4onoJrHrlTeJrbHbrnrr.r 

sasy6pnnraMn, c npocrbrMu BoJrocKaMu unn 6es Hr.rx, xe- 

[e3r4crbre BoJrocKr{ HHorAa r,rMercTcr Ha paxuce u cpeAneft xHJrKe Ha Hr,rxneftcroponerr,rcrorrKoB 

.....2. 

2. Jlncroqru errrrunrr.rqecKr.re, 2 (3)-5 (6) cru rur., l-Z (2.5) cM rrrup., c ryroBa- 

To 3aocrpenHoft nepxyurxofi u urnpororirlrHoBltAnblM ocHoBaHr.teM . . . . 3. 

Jlucroqxu or rrlrpoKo3JlflnnrHrrecKr.rx Ao oKpyulbrx, c orpyrnoft l,rJrr,r ryno- 

Baro 3aocrpeunofi nepxyumoft u oxpyrnbrM ocHoBaHr{eM ... .. -...... 7. 

3. lllnnsr a nepxHeft qacrr.r reneparr{Bnbrx no6eros o6uqno MeJrKue, zl-5 nanr 

AJL, CIerKa CepnOBr,rAHO r,rJrr,r KpIOTTKOBT.TAHO r,r3orHyTbre, oTAeJrbHble turnbr 

uoryr 6rrrr nprMbrMr,r, ropr,r3oHTanbHo orcrof,rrlt.rMu wtur cJrerKa orrlroHrrculr,rMr,rcr 

nrepx. fho4rr o6rrqno SJrJrr.r[co]rAanbHbre, pe)Ke rrrapoBuAHbre 

HJrH KyBrrrr,rHrlaTble; IIBeTOHOXKU, ilInaHTnn U qaIuen4CIttKI,I Ha CrrHHKe rOnbre,pexexerre3r{cTo-rqeTl,rul,tcTbre 

.......... 4. 

LIInnlr r nepxneft qacrr,r reHeparnBublx no6eros KpynHbre, Ha ueHTpiurbHoM 

no6ere l-1.2 ctr,r An., nptMbre, ropu3oHTaJrbHo orcrorullre rinr.t Ha[paBJreH- 

Hbre Koco BBepx, r.rJrr,r cepnoBrlAno tnorHyrbre. flnoAsr o6uqno KyBtuHHqa- 

Tbre, pexe SJrJr[ncol{AanbHbre; rIBeToHoxKr,r, rI,ItIaHTr.rrl u qaureJrncTr,rKr,r Ha 

cnlrnKe6..r.xeresficro-rrrerr,rHl,rcrble.... .... 5. 

4. 

Kpar lucro.rrcoo my6oxo-ocrponunbqarbre; Jrr,rcroqKlr c o6eux cropoH ro- 

Jrbre, pexe cHr43y onyueHHble flpocrbrMrr BoJIocKaMu. {reroHoxKr,r, runaH- 

TI.II,I H qAITCJII4CTI,IKII HA CNI'HKC TIACTO XEN$ICTO-I.IIETI,IHI,ICTbIC , . 

...... 5. R. glabrifulia. 

Kpar nncrovKoB c ryfloBaro-3aocrpefi HbrMH, rrpr,rireraroq]rMu sy6qaM]r; 

JlncroqKr,r c o6enx cropou r.rrrr4 ToJrbKo c snxneft noBepxHocra onymeHHbre 

npocrblMr.I BoJIocKaMr,r. [neroHoxxn, fltIIaHTI{u 14 qaueJll,rcrl,rKh Ha cnuHre 

o6H.rxo roJrbre . 4. R. majalis. 

5. lUunrr n nepxneft qacrr,r reHeparr,rBnbrx no6eros cepnoBuAHo H3orHyrHe, 

pexe ropr.r3oHTaJrbHo oTcToflulne, HeMHoro BBepx nJII,I BHr,r3 oTKJroHeHHbre, 

I,rHor,qa nepeMexarcqnecr MeJrKI,rMr.r n nroBnAHbrMr.r firtrnr.rKaMr.r. fllo4u rynrrrr{Hqarbre, 

Kpynnbre, Ao 3 crvr gn., o6uvno xeJre3r.lcro-rqerl,IHtrcrbre, noHl,t- 

Karcrque Ha AJrr,rHHbrx 6. q. xeneauc'ro-qerr.tHl,rcrbrx rlBeroHoxKax. . . . . . . 

. l. R. oxyodon. 

lllnnu n nepxueft qacrr,r reHeparlrBnbrx no6eros erepx o6paqeHHbre, c Htr3- 

6erarculnrr,r no creonro ocHoBaHr{eM. flno4rr o6rrqno uapoBnAHbre t{Jnr 3Jrnnncdugalrnble, 

pexe KyBrrrr.rHqarbre, Ha Koporxux rlBeroHoxKax, paBHbrx 

r{M no Airr,rHe nnu 6onee KoporKr,rx, He noHnKaKrurve .. .. ... . . 6. 

6. lllnnrr r nepxHefi qacrr.r reHeparr,rBnbrx no6eros pacnoJraraKrrcr napaMn y 

ocHoBaHr.rr JrncrbeB n 6orconrx no6eros. flno4u ilapoBr,r4Hbte, pexe an.[tlncouAaJlbHbre, 

roJlbre, Ha roJrbrx I4II,I HefycTo xeJle3l,rcTo-rqeTnHl,rcTbrx uBeroHoxKax 

... 2. R. prokhanovii. 

IIIunrr n nepxnefi qacrr,r reHeparr.rBHblx no6eros o6uqso paccerHHbre no 

no6ery, pexe napHbre y ocHoBaHHr Jrr{crbeB. fhoAu 3nJruncor,rAaJrbHbre, 

pexe KyBI[[rHqaTbre, rycTo xeJre3ncTo-rrleTnHr{cTble, Ha xeJIe3[cTo_rIIeTr{HH_ 

crbrx rlBeroHoxKax. . 3. R. sosnovskyana. 

lt7

7 (2). lllunu KpynHbre, 8-10 tuu Ar., KpIoqKoBLIAHbIe, KoHIIbI nx u3orHyrbt noqrr,r 

noA [prMbrM yrnoM. [aeroxoxxtt no AJIHHe paBHbIe nJIoAaM r,rlu npe- 

BbrrrraK)T ux 6olee.relr n 2 para, o6uvno rorble. Jlucroqxu c o6enx cropoHroJrbre 

..... T.R.bulcaricu. 

f Lllunu MeJlKue, 4-5 rrav AJL, cJIerKa cepfloBttAHo tr3orHyrble. IlneroHoxxu 

no aJIl,tHe paBHbre rrJIoAaM I,uI,I KOpOqe UX, rOJIbIe LIJIU paCCetHHO )(eJIe3UC- 

To-urernnr,rcrue. Jlucroqrlr cBepxy roJrble, cHl,t3y onyueHHble npocrblM14 

BoJrocKaMr,r 8. R. oblegens. 

L R. oxyodoz Boiss., 1872, Fl. Or.2:674. -- R. haemathodes Boiss., 

op.cit. : 684. - R. oxyodon Boiss. var. pleiantha Boiss., op. cit. : 674. 

Onucan c Bocr. Kasxasa. T u u :

Mr4 UBeTOHOXKaMI{, HO C pa3pexeHHblM otlyueHl{eM, KpyrIHblMI',I XeJIe3ncrbrMr{ 

BoJrocKaMlr Ha HlIxHefr cropoHe [I4croqKoB, qro l-oBopur o Ilx 

ru6pltAnsaulll4 c BI{AoM n3 [oAceKIII4u Rubiginosae ceKr\vr4 Caninae- 

.[arecraHcrue o6pa3ubr R. oxyodon uMexrr f,BHbIe cJleAbl rn6pu4uraqnn 

c BrrAoM, 6nn:ruu R. gadzhievii Chrshan. et Iskend. I43 ceKul4lr Canilwe 

cyAfl rro xapaKTepHblM, yAJIuHeHHO-3JIJII'IIITuqeCXOfi $OpUu' fycro xeJle- 

3r{cTo-ulernHr4srbtM 14 qacro co6paHuutr,t tto 3 nnoAaru, a raK)I(e IIO llpeo6raAaHulo 

AJIIIHHbIX, ToHKItx, ceprIoBI,IAHo l,I3ofHy'tblx tuHrloB. 

2. R. prokhanavii Galushko, 1959, Eor. uar. (JleHraurpa4) 19 : 205' 

pr.rc. l; fa:ryurrco, 1967, !ep.I{ Kycr. Ces. Kasr. :277; ou xe' 1980, On. 

Cen. Kasr. 2: ll0. 

OuucaH c I{eHrp. Kasxa:a. T u n :

O roru, rITo rcHocrr4 B rroHrrMaHr,Iu R. roopae Ao cux nop Her, roBopHT 

ra ror Qaxr, qro I,I. fl. ManAenoBa orHoctrr R. roopae K BtrAaM ceKur,rn 

Caninae, a ocriulbHbre aBTopbr - K ceKrlrlr,r Cinnamomeae. B ra6laue, 

cocraBneHHofi glr oupe4eJreHnfl KpbrMcKl4x ta KaBKa3cKr,Ix rrr4rroBHnKoB, 

A. A. JloHaqescrufi (1912:105) noMeulaer R. roopae cpeAr,r Br4AoB, r.rMercIuux 

(qarrreJrrdcrttKn rleJrbHbre r,rJru c HeMHoroqucJreHHbltuu yrxuun (lu- 

Hefruo-surenu4nunau) lepbrurraMrrD, orJruqa{ ero or Apyrrlx BtrAoB c 

rreJrbHbrMu qatueJrr,rsrr,rKaMn (tunflaMr,r KpyuHbrMH naepx o6pauleHHbrMr{, 

rrBerKaMu 6olrrueft qacrbK) B MHororrBerKoBbtx couBerr,rflx)). guclo 

uBerKoB yKa3brBaercfl or 3 Ao 12, gtrwrua uBeroHoxeK l0-15 uu, couBe- 

TI{e fycro ulerrrHl,Icroe. Taroro runa couBerur, LJrfl. BtrAoB ceKr.It{t,r 

Cinnamomeae o6n.rno He xapaxTepHbr. 

4. R, majalis Herrm., 1762, Diss. Rosa : 8; falyuro, 1980,

B rpynry R. majalis Herrm. s. 1., u AaHHHe o6 ux pacnpocrpaueHuu 6y- 

Ayr 3aBr{cerr or nrr6opa JreKTort{ua 3Toro Br4Aa, 

6. R. adenophyllc Galushko, 1961, Eor. uar. (Jleuuurpaq) 2l :208, 

pr,rc. l; fzuryurro, 1967, [ep. n Kycr. Cee. Kasr. :279; ou xe, 1980, (Dl. 

Ces. Kaer. 2:109. 

Onucan c l{eHrp. Kasraga. T n u : (LE!). 

Ha rpaoruucrbrx r4 KaMeHIdcrbIx cKJroHax B cpeAHeM fopHoM rrof,ce. 

-- llenmp. Kaex. .' Manr. - O6q. pacrp.: 3HAeMr4K. 

flpuue.ranue. BraA lr3BecreH ToJlbKo rro rr.rrroBoMy 3K3eMrrJrrpy. 

7. R. balcarica Galushko, 1960, Eor. uar. (Jleuranrpa4),20 :205, 

puc. l; falymxo, 1967, [ep. r,r Kycr. Ces. Kanr. : 280; on xe, 1980, @1. 

Ces. Kasx. 2:109. 

Ouucan c l{eHrp. Kasrcasa. T n u ;

f a n y ur r o A. I,L

M. C. Knrsen, 

II. B. Kvmucon 

M. Kniasev, 

P. Kulikov 

ACTPATAJIbI (ASTRAGALUS L.o FABACEAE) CEKUI4I,I 

XIPHIDIUM BUNGE BO OJIOPE YPAJIA 

GENERIS ASTRAGALUS L. (FABACEAE) SPECIES 

E SECTIONE XIPHIDIUM BUNGE IN FLORA URALENSI 

flareapxruuecKat cerulrt Xiphidium Bunge tBJltercfl oAuofi I{3 HaH- 

6onee rpyuubrx [o qllcrly BTIAoB B poAe Astragalus L. Apeau ceKuHIl 

oxBarbrBaer flaBHblM o6parou apttAHble BHyrpnKoHT[HeHraJIbHbre pafroHu 

Erparurl c rleHrpoM pa:Hoo6pasl4s B topax CpeaHeft A':un. Bo 

$lope 6brsurero CCCP lra ceKulrfl rBJlserctt nan6o.lree uuoro.lncleuHofi 

B poAe r{ HacqurbrBaer.6onee 100 nraAos (foHuapor, flouos, 1946; 

Yakovlev et al., 1996). Bo (rlope Poccnu, reppuropltt rcoropofi BKJIIorraer 

rtr.rtrlb nepn$epuvecrue, Hau6olee 6eAHrte BuAaMI,I qacru apeaJla 

ceKr.IHu Xiphidium, oHa rlpeAcrasreHa 3l BuAoM (nrlru.rar onucrtsaeMltfr 

Hraxe). Cpe.qu perrroHoB Poccuu uau6onuuee BuAoBoe 6orarcrso acrparanoB 

ceKquu Xiphidium orMeqaercfl ua Vpzure (upeuuyulecrBeHHo 

loxnou), ao $nope Koropofo [peAcraBneHa rtotrrlr rIoJIoBI4Ha BHAoB 

ceKulrn, r43BecrHbrx Ha reppuropwu Poccun. 

Hosne AaHHbre o BI4AoBoM cocraBe I{ pacllpocrpaHeHnr acrpalaJloB 

ceKuun Xiphidium ua Vpaite, flonyqeHHble B rIocJIeAHI,Ie roAbl, cyqe- 

CTBeHHO yTOqHflIOT I{ AOIIOJTHflIOT CBeAeHHt, COAepXaqI4eCt He rOJIbKo 

no

urne Qropr,rcrr,rqecKr.re n raKcoHoMr,rqecKue pa6orn, trMeroulne orHoue- 

HHe K ypanbcKoMy pernouy), ocHoBHbre cuHoHr.rMbr, cBeAeHur o rlruo- 

BOM MaTepr{aJre, KpaTKaf, xapaKTepr4cTnKa paclpocTpaHeHr,r{ B paccMaTpL4BaeMOM 

peruoHe n 3a ero [peAeJraMr,r, KapTbr pacnpocTpaHeHr4r; 

s Heo6xoAulabrx cJryqaf,x AaHbr KoMMeHTapr,rH. PacnpocrpaHeHr4e Br{AoB 

yKa3aHo rro urrporHbrM ceKTopau Vpalrcrofr ropHofi crpaHbt tr rro 

o6lacrsru u pecuy6rnrau Poccuu, BxoAqrquM B cocl'aB Vpalrcroro 

perr,roHa. M rr orragaJrr4cb or r,rcrroJrb3oBa nut pait ouwpoBaHr4q rto 

* Mnoronernae no6eru AoBoJIgHo cr4JrbHo oApeBecHeBarourue, 6olee Anunsue, 

6. M. BBepx HanpaBJreHHbre. Xors 6u nexoroplre Jrr.rcrbr c 6olee urupoKr,rMr,r 

Jrr,rcroqxaMu. Eo6rr 2-3.5 cM An.,2.5-3.5 MM rrrnp. .......... 7. 

6. @:l^ar 23-27 MM AJr. 9aureqra I l-13 r',rrrr Al. Eo6u 3-5 cu AJr., nprMbre HJrR 

eABar.r3orHyrbre ... ......5. A.pseudotataricus. 

+ Onar 18-20 (22) uu Ar. 9aureqxa 8-10 MM Arr. Eo6u 2-3.5 cM Arr., HecKorrb- 

Ko r{3orHyrbre, pexenprMbre..... ..... 6. A. ucrainicus. 

7. [rerxu 6eroearHe; nJracrnHKa JroAoqKu n 1.5 pasa Kopoqe nororra. Jluc- 

ToqKI,r B ,{Iacre 4-5 nap. Pacrenux, o6urarorqne Ha cKaJrbnbrx obnaxenusx 

yJrbrpaocHoBHbrx nopoA (pexe ua nrnecrnrxax) H B KaMeHr.rcrbrx crenrx 

. .. . 14. A. storozhevae. 

+ IIBerKr,r nypnypHbre; [JracrriHKa JroAoqKr.r 6oree .{e}r e 2 paza Kopoqe Hororxa. 

JlncroqKr,r B qucre 5-7 nap. Pacrenur, o6urarculue na velonux o6- 

HaxeHutrx, pexe Ha rrecKax . . . . . 13. A. pseudomacropus. 

8 (4). tlaurevxh oflyureHbr He BnoJrHe np]rxarbrMr.r, HeMHoro r,r3orHyrbrMr,t BoJrocraun. 

JlncroqKlr B r{r{cJre 4-5 nap. I{nerru fiyptrypnbre. Eo6u 3-4 cM An., 

2-2.5 uu rrr4p., nprMbre, npnxaroofiy[reHHbre .. 2. A. tenrirensis. 

* 9aureqxu onyuleHbr rBcrBeHHo ItplrxarblMl,I nprMbrMlr BoJrocKaMI,r. Jlucro.r- 

Kr{ B qlrcrre (5) 6-9 nap. I{nerrra 6enonarue, oqeHb peAKo nypnypHne. Eo6u 

2.5-3 cu gn., 3-3.5 MM rrlrp., r.rHorAa HeMHoro H3orHyrbre, onyureubr npnxaTblMr,r 

r4 KopoTKrrMlr noJryoTTonbrpeHHblMlr BoJrocKaMrr ..... 9. 

9. Har.r6onee pa3Br.rrbre rr.rcrbr c 5-6 (7) napaMr,r JrncroqKoB. Eo6u 6. q. HecxoJrbKo 

r{3oruyrbre, ony[reHbr roJrbKo 6enrtult, orHocr,rreJrbno AJrnHHblMl,r, 

HIaBHbTM o6paeou noJryorronbrpeHHbrMlr BoJrocKaMH. 9aue.rra rpy6varaa, 

6enoonyrueunar, c npl{Mecblo qepnbrx BoJtocKoB, pacnoJroxeHHbrx npenMyulecrBeHHo 

na sy6qax u nnxnefi cropoHe. Eyronrr n qBerKr,r 6erosarrre 

4. A. austrouralensis. 

t Hau6onee pa3Bnrbre rrr.rcrbr c (6) 7-9 napauu rrr,rcroqKoB. Eo6rr 6. q. npr- 

Mbre, o[y[reHbr KopoTKlrMr., npeuMyrrlecTBeHHo npuxaTbrMr.r H HeMHofo oT- 

KJroHeHHbrMrr 6enrrun BoJtocKaMr,r, BcerAa c npr,rMecblo nptrxarbrx qepHbrx 

BoJrocKoB. 9aureqxa rpy6varo-ronoKoJlbqarar, 6eloonyurennar, c npnMecbro 

qepnblx BoJlocKoB, pacnoJroxenubrx paBHoMepno no ncefi fioBepxHocrlr. 

Eyronrr xeJrroBarbre r.rJrr{ opaHxeBbre; rIBerKH 6euosarrre (n rep6apuu 

xenreloque), oqeHb peAKo flypuypHbre 3. A. karelinianus. 

l0 (3). Courernr rurorHbre, rorroBqarbre, npu ruroAax He yAnr4Hflrour,recq . . . 11. 

t Coqserfiq pbrxJlbre, yxe B HaqaJre rrBereHl,tt KoJrocoBHAHbIe trnn npoAonro- 

Baro-roJloBqarbre, npu nJroAax yAJrnHrloqlrecfl .... ........ 13. 

11. MnorolerHr,re oApeBecHeBarcrrlne no6erlr pacnpocreprbre r4Jrn HeMHoro BocxoArrrlue. 

Eo6u urnnonnAnbre, 3-4 cu w., 2-2.5 MM rrrnp., np[xaroony- 

-f 

IrreHHbIe ......2.A.temirensis. 

Mnoronernze no6ern 6. na. npauocrorque. Bo6rr npoaonroBarbre, l-2 cu 

ur.,3-4.5 MM rrr[p., noJryorronbrpeHHo onyruenHbre .. ...... 12. 

12. ,[|,ononruo BbrcoKr4e (.qo 70 cr\a arrc.) rycrapruK?r c noA3eMnnun no6eraun 

(o6pasyrcr unorocre6enrHbre Kyprr4Hbr), qneryulr,re c HaqaJra Maf, rro rlroHb. 

l-(nerxr.r 23-28 urvt gl., o6uvno po3oBo-uyprypHbre, recrpbre (c xpxo ox- 

t25

paureHnbrM Snarou n 6erosarrrNrfi xpulrrlrra), pe4xo 6enrre. Eo6rt o6rtqso 

cjrerKa r.{3orHyTble, AJII,THHO nOnyOTTOnbIpeHHO O[ylIeHHbIe . . 

.... 8. A. oropolitaruts. 

+ O6uqno MeHee BbrcoKr.re KycrapHLIKH, 6el noAseMHrtx no6eron, c oAI,IHoq- 

HbrMr,r Ha3eMHurun no6erauu, uBeryurne c KoHua Mat no urcnr. I-[eerru l8- 

20 rvur,r an., paBHoMepHo reMHo-nypnypnrre. 6o6u noqrl{ nptMbte, 6onee 

KoporKo onyrueHHble ....7. A. conrutus. 

13. Jlracr, r43 na3yxl4 Koroporo BbIxoAI,IT suxuufi uBeroHoc, cvtr,s\uia [rJILt c qeperuKoM 

rrlesee I cm gr. KycrapHl4Kl4 Ltnlt nonyKycTapHl,lKl'r c cqnbHo oApe- 

BecHeBarouruMu, 6. trl. npsMocrotqnMll MHorolrerszuu noOeraun . ... 14. 

+ Jlncr, r.r3 [a3yxr,I Koroporo BbIxoAHT HtIxsuft lrBeroHoc, c veperuronr 6olee 

1.5 cu al. flonyxycrapufiqKl{ c pacnpocreprblMvt vtrv BocxoAtulI,rMLI, cna- 

6o oApenecueBaIouII4MI,I MHororerH[Mu no6eraun 15. 

14. Jlucro.{xq sau6onee pa3Burblx JlrrcrbeB 2-4 cu AJL, He 6onee qeu n2paza 

Kopoqe paxr{ca, HepeAKo paBHbI eMy LIJII,I HeMHoro AJII,IHHee. Llaerxu 22-26 

MM AJr., paBHoMepHo pacstaBJlenHble tro oAHoMy Ha oct{ couBerllr. 9aureqra 

o6uqno 3erleHat, otrytueHHat (rar n 6o6rr) npel,rMyu]ecrBeHHo pbtxno npuxarbrMrr 

BoJrocKaMn. Eo6rr 3-4 MM llrnp. KycrapHnx, pacryu{r.rft na o6naxeHurx 

MeJra H n3BesrHtKa . . . 9. A. brachylttbus. 

* Jft{croqru 6oree uenrne, y uan6onee pa3Brlrblx JIItcrbeB ao 1.5 (2) cM An., 

6onee qeu y 2 pata Kopoqe paxfica. [serru I 8*22 uu An., B rycroM KonocoBriAHoM 

couBerr.ru, qacrb I,I3 HLIx HepeAKo c6ltlxeua Ha ocn couBerllt no 

2-3. 9aureqxa po3oBarat, onyureHnat (xax ra 6o6rr) npel,IMyu{ecrBeHno no- 

JryorronbrpeHHblMl4 BoJlocraura. 6o6rt 2.5 -3 uu urr.rp. llonyxycrap HI4K, 

rcrpevarouluftcr o6rtqHo Ha fiecKax ll. A. 't'arius. 

15. [aerxu 6erorarrre, peqxo 6re4uo-po3oBble; npuIIBerHI'IKIt JIaHL\erHbIe, ocrpue. 

6o6rr fioqrr.r nprMbre, orronblpeHno 6enoonyrueHHble. Onyruenue va- 

[reqKu c ne6onsnroft npflMecbn npHxarbtx qepHbtx BoJIocKoB 

15. A. pallescens. 

+ l-[nerxn nypnypHbre, c 6erosarsrMlt KpblJrbtMl{, pegxo 6elute; npnqBerHuxu 

rft uesr4AHrre, Tynbre. 6o6u uronHyrble, orronblpenuo 6enoony[reHHble, 

BcerAa c npr.rMecblo npnxarbrx qepHbrx BoJIocKoB. B onyurennu qaureqKu 

o6nlsnrrirepHbreBoJrocKrr .....12.A.macropus. 

Ser. l. Arbusculi Vass., 1970, Honocru crrcr. Bblctu. pacr.6: 148; id., 

1946, so

upr4Hrr K. H. HrournHofi) n o6Hapyxunaer 6oJtee orAaJIeHHoe poAcl'Bo 

c 3arraAHoKa3axcraHcKo-IrpuypaJlbcKnu A. temir ensrs M. Pop. Haxo4ra 

BHAa co croJrb IoxHbtMH poAcrBeHHbIMu cBq3f,Ml{ Ha flonrpHolr Vpane 

cBfl3brBaercf ero aBTopaML4 c Hatl,Jl4tg'veM B rIo3AHeM rIJIefrCTOIleHe IIyTI4 

rrpoHr{KHoBeHur rcepo$HTHblx 3[eMeHToB I'I3 rlpuaJlTafr cxoro IIeHTpa 

Qnoporeuesa B ceBepHble pafiouu Ypanlcrofr ropnofi crpaHbl ueper 3arraAHocr46r4pcKyrc 

H[3MeHHOCTb. o.reSI,IAHO, 3TOT rlyrb mOr QyUnqu o Ht'tpoBarb 

roJrbKo r laHquratprHo-KJr[Marl4qecKrrx ycroBuqx Kpl4oapnAHblx 

nHTepBaJroB rlo3AHero nneilcroqena, 3HarrI,ITerbHo o'rJluqarcu{Hxca or 

coBpeMeHHbrx (Kauelun, IOpuea, 1982). 

K coxanesulo, ToqHoe Mecro rtporl3pacraHllfl Ha flolapHona Vpale 

3TO|O 3aMerrareJlbHoro IIo xapaKTepy pac[pocrpaHeHilfl Bl'rAa, 3Haqu- 

TeJrbHo r,t3oJrrdpoBaHHoro oT Bcero apeara ceKul,tll, Ao CI'IX flop ocraercr 

HeBbrrcHeHHbIM, HecMorpt Ha HeoAHoKparHbIe tIoIIbITKl4 o6uapyxurs 

ero rroBTopHo. PeKu, creKaoulue c [epeBa[a Xofina KaK B 3anaAHoM 

(6accefrH p. fle.ropu), raK I,I B BocroqHoM (6accefru p. O6u) HarlpaBre- 

Htrflx, HocflT oAI4HaKoBbIe Ha3BaHI{f,, u Ha rar

cr yKa3aHr{fl Ha HaxoxAesue A. stenoceras s Axro6raHcroil, KycraHaficrofi 

u Vpa.irlcxoft o6t. Ka:axcrana (faualoHona, OuuoH, 196l; yakovlev 

et al., 1996) ureayer ornoctrrb x A. temirensls. flpn rpocMorpe 3K3eMrrrupoB 

r,r: fep6apnx flasHoro 6orauuqecroro caAa PAH (MHA), na 

KOTopbIx OCHoBbtBiUrHCb yKa3aHr.Ur o HaxoAKax A. karelinianas M. Pop. n 

Cpe4Herra flosonxre (Yakovlev et al., 1996), Hanan 6ulo ycraHoBJleHo, 

rrro oHr,r raKxe orHocf,Tcf, x A. temirensis. 

Ser. 2. Macrotropidi Vass., 1970, Hosocru cncr. Bbrctx. pacr.6:149 

id,.,1946, no Or. CCCP 12:686, descr. ross. 

3. A. kurelinianus M. Pop., 1946, so Or. CCCP 12 : 695; Chater, 

1968, in Fl. Europ. 2 : 122: Bacunrena, 1987, so (D.r. Enporr. .racru 

CCCP 6 :72; Yakovl. et al.,1996, Legum. North. Euras. :256, p. p. 

A. rnacrolobz.r Bieb. var. B. nigropilosus Bunge, 1847, Arb. Naturf. 

Vereins Riga, l, 2:239; id., 1852, Beitr. Kenntn. Fl. Russl. Stepp. Centr.- 

As. ; 261 (seors. impr.); id., 1854, M6m. Sav. Etr. P6tersb. 7 : 261: id.. 

1869, M6m. Acad. Sci. P6tersb. (Sci. Phys. Math.), s6r. 7, 15, I (Astrag. 

Geront.2):222, p. p., quoadpl. ural. -- A. ortholoDasBunge, 1869,1.c.: 

223,p. p., quoad pl. ural.; Kpun., 1933, (Dn.3an, Cu6. 7: 1701, p. p., 

quoad pl. ural. ?A. virescens Kar. ex Ledeb., 1842, Fl. Ross. l: 630, 

non Ait., 1789. -- A. mauolobas auct. non Bieb. : Korsh., 1898, 3an. 

Axa4. Hayr. @us.-Naar. orA. cep.8 (Tent. Fl. Ross. Or.): ll7. (pxc. l). 

JIe rrornn (Podlech, 1998): (P, isolectotypi LE!, P, K). (TuuoM BrAa rBJlrerct 

rurr pa3HoBlrAHocrtr A. macrolobas Bieb. var. nigropilosrzs Bunge). 

IOxnufr Vpal: Pecny6luxa Eauxoprocran (3r.uar,rpcroe ruraro, 6accefru 

p. Eon. lIx rpr{roKa p. Carnaapu; cpeAHee reqeHr4e p. Eenofi; 

Earuxnpcxoe flpe4ypanbe - 03. Acluxyll; Barurupcxoe 3aypalrc , 

Vqanuucxufi, A6sennnoacrcufi vi Eafiuarcxufr p-Hbl), gels6nHcxa.fl 06r. 

(xp. Br,luuentre fopu K ropa Ero:uucxax 6ru: r. Kururmua; os. Eon. 

Eyro4arc 6rus r. BepxueypalbcKa; ropa gexa ua neno6epexle p. Vpal), 

Opeu6yprcxar o6n. (6nlr: g. AHgpeeaxa Caparraurcxoro p-Ha). 

(Puc.4). - O6ur. pacnp.: 3HAeMuK lOxnoro Vpa.ua. 

VrasaHua 3Toro Br4Aa 4lr Vnlauoscrofi IE Caparoncrofi o6"rr. poccuvr 

u Y patrcrofr (3aua4Ho-Kasaxclascxoft) o6n. Kasaxcraua (yakovlev 

et al., 1996) ornocrrcfl K A. temirensls. Kar rroKa3aJro u3yqeHue 

uMeBrur{xcf, B HarueM paclopr)r(eHuu o6pa:qon, co6pauulrx H. lI. Ha_ 

yMeHKo Ha reppr.rropuu Kypraucrofr o6l. (orp. c. Vcrt-yftcroe) u co_ 

npegellHofr vacrn Kycranaficxofr o6n. Ka:axcraHa (A. JlroruHr

=l 

0rlur$, 

il 

I 

,rV I 

I6 

A" 

! 

I26 

\t\ 

V I 

2a 26 

:" d Ie 

f;1 

lww, l,, 

2a2z2dv 

Pnc. l. Astragalus austrouralensrs Kulikov (1, Ia-le), 

A. karelinianus M. Pop. (2a-2e): 

/ - o6rqafi sug; |a, 2a * $tar; 16, 26 - xpuno; 1a, 2e -- noAo,rxai lz, 2z qaureqKa 

B pa3BepHyroM sw\e; Id, 2d -- .raueqxa H npnqBerrurri Ie, 2e --- 6o6. Macrura6nar 

nnnefixa I cu. 

\ 

t29

BoAfl ToiIbKo pyccKoe orrucaHlle AaHHofo raKcoHa, ccbulaercfl Ha Aeficr- 

Br{TerrbHo o6uapo4onaHHoe Ha3BaHr.re pa3HoBI{AHocrIr A. macrolohus 

Bieb. var. nigropilosus Bunge, xparxuft AHafHo3 xoropofi Ha [arrHcKoM 

fl3brKe uMeercr n pa6orax A. A. EyHre (Bunge, 1847 : 239; id., 1852 : 

261):

B pa6ore I4. A. fy6auoBa c coaBropaura (fy6aHoB rr Ap., 1998), coaepxaurefi 

cIIr4coK TaKcoHoB, orrucaHHbrx f. C. KapennHbrM, B KaqecrBe 

JreKTorlr[a A. virescens [pr,IBe.qeH BHueyfloMtuyrufi ruuosofi fK3eMrItrtp 

A. karelinianus (), Ho KoJrneKropoM yKaraH 

Kapenlrn, xorg 3TlIKerKa 3Toro 3x3eMrrJrrpa He ocraBJrf,er couueHuft 

B roM, rrro pacreHne co6pauo Jleuanou. Bo K. @. JIe- 

.qe6ypa (Ledebour, 1842) A. virescens rrpr,rBeAeH B rpuMeqaHlrr,r, reKcr Koroporo 

He coAepxur xarux-nu6o cseAeHr,rft o uop$oloruqecKux rrpu- 

3HaKax 3lofo Br,rAa utl,u o6 orrlrqurx ero or Apyrr{x Br{AoB; yKa3aHo 

Jrr,rurb, rrro oH Lr3BecreH rro He[onHoMy 3K3eMrrJrflpy 6es rrJroAoB (), co6paHHorvry n OpeH6yprcnoft 

ry6epHura (), u upe4uonaraercfl 

ero 6rusocrr r a-nraftcxouy A. pycnolobu.r Bunge. Tuuosofi Nrarepr{an 

no A. virescens a fep6apr.ru LE, rae xpaHr4Tcr ocHoBHar .{acrl c6opor 

f. C. Kapeluua, He o6uapyxeH (nosnaoxHo, oH HaxoAl4Tcf, B eu{e He 

paeo6pauuoft qacru KolJreKrll.rn); us yuonaxuyrofr Bbrure pa6oru 

14. A. fy6aHoBa c coaBTopaMn cJreAyer, qro sToro Marepr4aJra Her 14 B 

fep6aprau MW. Buagy orcyrcrBr.rf, cneAeuuff o uop$ouoruqecrr4x rrpu- 

3HaKax B lporoJrore A. virescens Kar. ex Ledeb., oro Ha3BaHue rBJrflercr 

nomen nudum, a Boflpoc o6 ugeuu.rHocr]r SToro pacreHr.rfi c A. karelinianus 

rlJrr,r KaKrrM-Jrn6o 4pynana rcx(HoypiurbcKraM acrpafaJroM ocraercf, 

oTKpbrTbrM. 

Bo

flypnypuoqBerKoBar Qoprraa A. karelinianus HaMelrailact M. f. flouo- 

BbrM K orrr4caHr4ro xax A. karelinianus var. violascens M. Pop. in sched., 

Ho raK u ne 6tua ouy6uuroaaHa, Huxe Mbr uptrBoAr,ru onucaune a'rofr 

pa3HoBr4AHOCTr4, 

A. karelinianus var. violaceus Knjasev et Kulikov var, nova. Ab 

A. kareliniano var. karelinrano floribus purpureis, in sicco violascentibus 

(nec albis, in sicco flavescentibus) differt. 

T y p u s :

0qr 

ffi 2d 2e 

\/l/ 

t/-( 

lill 

lirrl 

ryt 

2a 26 2e 2e 

/ t 

\ l 

V 

r'!t!fl 

ll# 

I 

I 

// 

/l 

/,'N 

lt;i'iil 

{;iil 

L ' i I 

t., I 

\\,,t 

v 

td Ie 

ntr \ tilil 

V I[U, 

r a 2 

r.ffi 

llm 

36 3e 3e 

ffi 3d 

Puc.2. Astragalus pseudomacropus Knjasev et Kulikov sp. nova (1, Ia-le), 

A. varius S. G. Gmel. (2a-2e), A. macropus Bunge (3a-3e): 

/ - obqufi smx la,2a,3a-Qnar;16,26,36.- xpruro; Ia, 2e,3e -- no1otxa; Iz,2z, 

.Ja - qaureqxa u npnuBerHrK; Id, 2d, 3d - qaueqKa B pa3BeptyroM auae; Ie, 2e, 3e 6o6. 

Macrura6sar nnxeftra I cu. 

133

HoHrlMoM A. or tholobus. flo-an4uuoMy, [poucxo)r(AeHr.re roxHoypanbcKI4x 

3H.qeMrrxos A. austrouralensis u A. kqrelinianus cBr3aHo c Mr4fparlt4flMt 

Ha Vpan lerpoSuruo-creflHbrx 3JreMeHToB ti3 npuaJrrafrcror.o 

qeHTpa au4oo6paso Batkrfl.. 

5. A. pseudotataricus Boriss., 1946, so

(rcr u nro-nocr.), Poccur (toro-sana4 Erponefrcrofr .racrn, CpeAHee 

flononxle: Cauapcraa o6n.; uuerorct yKa3aHI4a Attr Ces. Kaexa:a: 

Cranpouo.rrlcrufi rpafi ). 

Onracauue sHAa bltrro ony6nurconaHo ABaxAbI c nHTepBaJIoM B HecroJrbKo 

MecrrreB B KoHue 1946 t'. u s 1947 r. Ha JraruHcKoM r3brKe, Ho 

Tr4uoM BHAa B srnx ny6nuraqr{rx yKa3brBaJrucb pa3Hbre rep6apHue ax- 

3eMrrJrrpbr: n nepaoft -- Bbr[rerrpuneAensrrft o6paseq, a no nropoft 

ulo 3Tr,rx MecroHaxox\eunfi u pacfloroxeHue r{x rro orHoueHHro K CO- 

BpeMeHHbrM aAMr4Hr,rcrparr,rBHbrM rpaHr4uaM 3arpyAHr,rrelluo (; >;

Puc. 3. PacnpocrpaueHue Ha Ypate Astragalus cornutus Pall. (l), A. temirensis 

M. Pop. (2) u A. storozltevae Knjasev (3). 

Hecuorpr Ha yKinaHr4e no , 'rnrr 3Toro BLIAa orcyrcrByer n LE, u MecroHaxox- 

AeHr4e ero BbucHr4Tb HaM He yAaJrocb. Bo3lroxHo, oH xpaHuTcfl n JIou- 

AoHe, rAe HaxoAKTcr 6onluras .racrr rep6apur fl. C, flanraca. 

B upeqelax paccMarpuBaeMoro perrroHa oco6urfr [Hrepec upeAcraB- 

Jrrlor crrJrbHo o6oco6neuuHe cpeAHeypilrrbcKue MegroHaxoxAeuna, o6pa- 

3ynuue n3oJrr{poBaHHuft ceaepuufi QparrvreHT apeaila, oqeBr4AHo, pe- 

JTT4KTOBO|O XapaKTepa. 

8. A. oropolitazrs Knjasev et Kulikov,2002, Eor. xypH. 87,2 : l3l. 

T u u : Poccnl,

cKJroH, KaMeHr.rcrafl crerrb Ha u3BecrHflKax,14V 1999, qB., fI. B. Kyluroa> 

(LE, r43orr,rrr - SVER). 

lOxHufi Vpa.rr: OpeH6yplcrar o6n. (ueHrpaJrbHbre p-Hbr, rnaBHbrM 

o6pa:ou MexAypeqbe pp. Vpana u Caruapn H3BecrHrKoBbre yBaJrbr 

s Eersencrou u CapaxraucKoM p-Hax, Kynau4urcrnfi ra fy6epLruucruii 

MeJrKocouoqHr4Kr4 B KyaaHAnxcxon, fafrcrou u Hoaorpouuxonr p-uax); 

Pecrry6nuxaEamroprocrau(3naH.rypuucrufi p-n oAHoMecroHaxox- 

AeHI{e Ha rpaHr{qe c Opeu6yprcxofi o6n.). (Puc. 5). O6ur. pacnp.: 

Bocr. Ka:axc'rau (orp. r. Vcrt-KaueHoropcra), A;rrafi. 

Apeiur Bl,u'Aa Ha IOxHoru Vpale 3aHHMaer orHocureJrbHo ue6oluuyru 

Teppu'ropr4rc B ueHrpaJrbHoi4 \ac'ru OpeH6yprcxoft o6n., rAe oH [or{TH 

rronHocrbrc 3aMerqaer 6nnsrufr A. cornutus. Pacnpocrpanenue A. oropolitanus 

ua Anrae HyxAaercr B n3yqeHuu. lult,rourrusunft arraftcro- 

IoxHoypaJrbcr

* r. 1 Fl': 

-\-*ill 

{ 

'-t 

A 

a l 

l 2 

. A 3 

Puc. 4. PacnpocrpaHeHr4e na Vpane Astagalus karelinianus M. Pop. (l), 

A. varius S. G. Gmel. (2), A. brachylobus Fisch. ex DC. (3) u A. austrouralensis 

Kulikov (4). 

BAoJrb p, Vpan n Ha Vpalo-To6olrcnoM BoAopa3Aene, o6pasyrcxlux 

n3oJurpoBaHuuft SparueHT apeaJra, 3Haqr{reJrbHo yAareHunfi K ceBepo- 

BocroKy or ocHoBHofi o6lacrn pacrpocrpaHeHr{r nu4a (puc. 4). 

Ha foxuoru Vpane, KaK r4 rro BceMy apei]Jry, A. brachylobus o6na[aet 

BecbMa 3HaqlrreJrbHoft K:Naen.rusocrblo rr, BepoflTHo, rrpellcraBnfler co- 

6ofi conoryuHocrb eue HeAocraroqHo H3yqeHHbrx JroKaJrbHbrx pac. O4ua 

us Har46onee cyxlecrBeHHo orJu4qaroulr4xcr pac 3Tofo KoMnlerca 6rrla 

HeAaBHo ouucaHa xax A. aktiubensis Sytin (Clrrran, 2000). Apeal rro-' 

ro Br{Aa oxBarbrBaer MeJroBbre ropu llo4ypiurbcKolo rrJraro B Arrrc6r,rgcrofi 

o6n. Kasaxcrana, B qacrHocru, n 6acceftue p. Vnn. Ha xpaftHena 

rore OpeH6yprcrofi o6n. (Bepxne.re6eHcrr,re u Tpouqxue MeJloBbre ropbr 

s Colr-Hrerlr

cTcq raKxe parHoo6parHbre rrpoMe)KyroqHbre Soprrau MexAy AByMr Ba- 

AaMr4, qro, Bo3MoxHo, yKa3brBaer Ha HaJr[rr[e Ha rore OpeH6yprcrofi 

o6r. soHrr r,rHTpolpeccusHofi ru1puguzagtrr{ MexAy A. aktiubensis tr 

A. brachylobus. 

Ser. 4. Mugodsharici Vass,, 1970, Hosocru cr{cr. Bbrclu. pacr. 6 : 149; 

id., 1946,.so tDr. CCCP 12:710, descr. ross. 

10. A. mugosaricus Bunge, 1847, Arb. Naturf. Vereins Riga l, 2 : 

236; id., 1852, Beitr. Kenntn. Fl. Russl. Stepp. Centr.-As. : 259 (seors. 

impr,); id., 1854, M6m. Sav. E,tr. Petersb. T :259; id., 1868, M6m. 

Acad. Sci. P6tersb. (Sci. Phys. Math.), s6r.7, ll, 16 (Astrag. Geront. l) 

:124 (); id,, 1869, L c, : 15, I (Astrag. Geront. 2): 

216; foHv. ra M. flon., 1946, so @r. CCCP 12 : 710; faMarcuoea r 

@ucloHo 1961, so

Tr.ru: [Ic.], S. G. Gmelin, 1770, Reise Russl. I : ll0, tab. XVII. 

Onucan r,r3 oKp. (c. Kocrenru n Bopouexcrofi o61., Ha rrpanou 

6epery,{oHa) (Curnn, 1999). 

IOxnuft Vpan: Pecny6lura Eaumoprocrau (e4uHuunbre HaxoAKH Ha 

rpafr Heu K)re, Bo3MoxHo, 3aHocHbre Mr,rrporrorryJrrrlrdr4 ; Eaurxupcxoe 

flpe4ypa.rne - os. Acnurynr), 9elr6uHcxas o6n. (xpafiuuff rcr Krasr,urrcxufi 

u EpeguHcrufi p-Hrr), OpeH6yprcrat o6l. (Puc. 4). - O6u. 

pacfip.: Eonrapua, Beurpnr, Pyurruua, Mon4anur, Vxparana (rporrae ce- 

Bepa u 3axapnarr,a), Poccur (ror Erpoueficxofr qacrvr; Ces. Kasxas: 

Cranponolrcxufi rpaft, .(arecraH), Ces. n l{eurp. KasaxcraH (Arrro- 

6uHcrar, Arupaycraa, Ypanrcrcax, KycraHaftcraa,,{xerra3r'aHcKafl , 

Typraftcrar u flauo4apcrar o6n.). 

B paccuarpuBaeMoM perr4oHe A. varius orHoclrreJrbHo o6rr.reH rolr- 

Ko B roxHbrx pafioHax OpeH6yprcxoft o6n., rAe qBrrflercs [per,rMyulecrBeHHo 

ucauuo$urou. B 6olee ceBepHbrx paftouax, rAe coo6uecrea 

[ecqaHbrx creneft He I,IMeIor rurlpoKoro pac[pocrpaHeHr4r, BlrA claHo- 

BI,TTCf, OqCHb PEAKI,IM I,I, BO3MOXHO, B PfiA€ CTYqACB lBJIflETCfl,3AHOCHbIM 

(r uacrHocru, orMerreHo [polrcpacraHr,re ero tro o6oql4Halu ruoccefiurrx 

Aopor r,r xeJre3HoAopoxHbtM uacunru). 

12. A. macropns Bunge, 1847, Arb, Naturf. Vereins Riga, l, 2:238: 

id., 1852, Beitr. Kenntn. Fl. Russl. Stepp. Centr.-As. : 261 (seors. impr.); 

id., 1854, M6m. Sav. Etr. P6tersb.7:261; id., 1868, Mdm. Acad. Sci. 

P6tersb. (Sci. Phys. Math.), s6r.7, 11, 16 (Astrag. Geront. 1): 125; id., 

1869,1. c.: 15, I (Astrag. Geront. 2):217; O. n E.@es.r., 1893, Mar. Qn. 

YQnu. ry6. : 98; Korsh., 1898, 3au. Ara4. nayx. (Dug.-nar. or4. cep. 8 

(Tent. Fl. Ross. Or.) : I l6; Kpun., 1933,

IOxnou Vpane. Apezur ero rtorrrlr ueJrr{KoM HaxoAtrrcfl B rrpeAeJ]ax cre[- 

Hoft gonn n rr4urb He3HaqHTeJrbHo 3axoAr4T s HaH6oree loxHbre paffourr 

Jrecocrerru. OAsaro s nxuofi qacrr,r crelHofi soHlr (r ror.y or 50" c. ur,, 

rJraBHbrM o6parou Ha reppr4Topuu Ces. Ka:laxcraua) A. macropus zaveulaercr 

6nu:ruu BItAoM A. pseudomacropus sp. nova, ourrcblBaeMbrM 

Huxe. Tunonoft uarepuat to A. macropus He o6HapyxeH HaMu n fep- 

6apuu LE (necrraorpr Ha yKa3aHue Bo ), Ho rrpaBuJrbHocl.b rrpuMeHe- 

Hr,rr AaHHofo Ha3BaHHr nMeHHO K 3TOMy BIlAy, trMpOKO paC[pOCl'paHeH_ 

HOMy B Creuuofi soue loxsoro vpala, JrefKo ycraHaBJrr{Baercr rro rrpq- 

BOAI'MbIM A. A. EYUTC XAPAKTEPHbIM IIPTT3HAKAM, OTII,IqAIOUIIIM PACCMATpl4BaeMoe 

pacreHr4e or 6nusxnx BrrAoB (A. pseudomacropus, A. storozhevae 

w A. varius): KoporKr4M rfiuesr.rArro-npoAoJrroBarbrM [prruBerHr{KaM, 

3Harrul'erbHo rrpeBbrurarcq[M roA,r4rrHble uo6eru rrBe.roHocaM, yKopoqeH_ 

HbIM peAKr,rM corlBernrM r,r orrorrbrpeHHoMy .repuo-6elouy orryrueHnro 

qarueqKr,r (Bunge, 1847,1852, 1868, 1869). Y A. ntocroprls AoBorrbHo peryJrflpHo 

Bcrpeqarorcq 6eroqserxosrre aJrl6uHocbr, npou3pac.ranurue 

o6u,{Ho orAeJrbHbrMrr oco6ql4u cpeAn pafl.eHzil c o6nI.troft (nypuypnofr) 

oxpacrofi qBerKoB. Or A. pallescens 6enognerKoBa{ Soprraa A. macropus 

orJr u q aercr orrylreH r{eM r{aruerlKu r,r Qoprr,rofi trplr uBe.rH HKo B. 

13. A. pseudomacropus Knjasev et Kulikov sp. nova. -- A. macropus 

auct. non Bunge: falrarouosa z (Dncnn, 1961, so

T y p u s :

cxal o6n., Hoooy:excrufi p-u, 40 xrr,r Ha roro-Ioro-Bocrox or r. Hoeoy:eacxa, oxp. rt.6a- 

6oruxun,40'25'N, 48"41'8,27 Vl 1993, A. K. Cxoopuor u ap. (MHA). Oucxar o6,'.: 

Pyccxono.nrHcxui't p-u, oKp. coBxo3a Ala6aruncrolo, Tl{rqaKoBo-KoBbrJrbHar crerr, 7 

vlll 1949' E. B. Baaqarypora (SSBG); flaraorpa4cxr.ii p-n, oxp. c. flaurorpalcxor.o, 

crenb ru[qaKoBo-KoBbrrbHar, 7 Vlll 1949, ona xe (SSBG). Karaxcran. Ax.rrc6uucxal 

ry6epHnr, Moxayper{be Yran-3rra6a, nofiua Vuna, Z Vl 1926, O. 3. KxoppraHr.; 6acc. 

p. Xo6qu, B AoltlHe p. Tamau, orolo KaparaHAbr-car, 6 Vll 1926, M. M. l4Lrtun, 

M. H.Anparuvrax; lrgis in deserto Kirgisico, 18 V 1881, A. Regel; Typr-ar-rcxar o6'. u 

ye3a, srenHbre ryra u cKrroHbr x p. qynax-caro, l6 vl lg0g, H. M. Kparuenfinnuxoo; l_le- 

Jrr4HorpaacKar o6Lr., Kyprynl.u,xnucxr,tii p-H, n l6 KM K ceBepo-Bocl.o(y or c. Apururu, 

s 2 rM or 6epera o:. Ba6afi,2l V 1973, O. Jl. Jloneluyc; AxuoLrnscrar o6l. u ye:14, 

5 uepcr or o:. ,{eurur, l9 vI 1914, B. ceuenon; In collibus arenosi ad fl. Taldv (ad oed. 

m' Ulutau), 28 v 1842, A. Schrenk; In deserris arenosis ad fl. lschim, zz vttt t'g+r, 

A. Schrenk; CeMnuanarnHcrar o61., flanno4apcxnit yesa, 6Lrn: p. rle4upr,u, ll VI 1914. 

c. E Kyveposcral; 3anaqxo-Ka:axcrancxar o61., o6pun ,reno'o 6eper.a Eo"ruuro'o 

Vsena y n. flopr-Apr.yp,9 Vl 1955, B. l,l. fpy6oe, H. flro6apcxnfi (nce o6pa:rqu LE). 

Po4crso. O'r A. varius S. G. Gmel. u A. macroprz.r Bunge orruqaercs 

6oree peAKr4Mr.r rr MaJror{BerKoBbrMr,r corrBerr4qMr.r, loAoqrofi c rrJracruHroft 

6onee qeM n 2 .(a rue n 1.5) para Kopoqe HororKa, qarueqKaMu 

n 6o6aruu, o[yrueHHbrMu rrpr4xarbrMr,r (a He 6. u. orrourrpeHHbrMu) Bo- 

JIOCKaMT4; I(poMe rol.o, or uepBofo Br,r.qa olrtrqaercr 60nee AJrr4HHbrMt4 

uBeroHocaMvr, a or BToporo -- 6otree AJTHHHbTMI4 JlanuerHbrMll, ocrpo_ 

BarbrMu (a He ryuurr,ru qfiqeenAHmrrau) npuqnerHuKaMtr, rpy6rofi rrauler{- 

Kr4 I]pqMo (a He xoco) cpesanuoft, 6olee roporKr,rMu qepeurann, 6onee 

oApeBecHeBaIourI'{MlI, Koco BBepx Ha[paBJIeHHbrMrr MHoroJre'rHnrrau uo6eraun, 

6eloaarbrMlr (a He 6e"uonaro-3eJreHoBarlruu) or 6olee rvcror.o 

ofly[reHnr foAnqHbrMr.r uo6erauu. BosN{oxHo, Hau60nee 6Jrusox i,s. t*stanaicus 

M. Pop.r, Ho orJruqaercr or Hefo 6o.lee xoporKuMr,r qepeuKa_ 

rrau, 6o;ree uupoKr4Mr4 14 MHororrapHbrMlr JrncroqKaMrl, 6oree srlCoxnMu 

uo6eraun, 6onee AnuHubrMr4, ocrpbrMr,r, JraHuerHrruu (a ue xfiqenn4Huuu) 

upuqaerH[KaMu. 

pxul! Vpan: Open6yprcxar o6n. (Vpano-I,Inercroe MexAyperrbe, 

rcr Coru-I{reuKoro p-ua). (Puc. 5). - O6u. pacrp.: poccur 

1rcib-noc_ 

ror Erpoueficrofi qacrni caparoncxax o6n.; ror'3an. cu6npvr: oncrar 

o6n.); aona creuefi lr rroJryrrycrbrHr cer. KasaxcraHa (Kaparan4r.rHcKaq, 

,{xerra:raucrar, Typraftcxal, Tan4'-KypraHcKas H Axrrc6nHcraq 

o6;r., ror Kyclauaficxofi o61., :anag Vpaltcxofi o6.rr.). 

A. pseudomacropus He Bcrpeqaercr ceBepHee rx[porHor.o yrracrKa re_ 

rreHrir p. Ypan (51"30' c. u.), a K rcry or 50" c. Ix. [paKTzqecKr.r rroJl_ 

Hocrbro 3aMeu{aer 6nuzxuit B:aL A. mecropus, c KoropbrM Ao cr4x rrop 

rrocrorHHo cMetuuB€urcr (rar, n LE 60ntruag rracrb Marepuana uz Ces. 

KasaxcraHa, otrpeAeneHHoro KaK A. macropus, n gefictsureJrbHocru o.r- 

HocHTcr x A. pseudomacropus). Hau6olee cxoAHbr c A. macropu.r MeHee 

I 

K coxrutenuto' cpaaHur' c 3Tr.rM B,naoM BHoBb onucbrBaenuft r. pseudontacropus ne y\a- 

Jrocb' rocxonbxy a LE xe o6napyxulocb H]r rula, Hn xorb cxonlxo-xr.r6yai bnrcroro 

no o6nnxy x A. kustanaicus o6pasqa, no)ToMy nprBoAuMbre o.rJrlli{Hr lrux B'aoB oc- 

HoBaHrr ToJrbKo Ha rrepBoont,rcauHn A. k*slanaicus. 

t44

Pr,rc. 5. PacnpocrpaHenfie Ha Ypane Astragalus macropus Bunge (l), 

A. oropolitanus Knjasev et Kulikov (2), A. pseudomacropus Knjasev et Kulikov 

sp. nova (3) u A. pallescens Bieb. (4). 

pa3Burbre gK3eMIlJItpbr A. pseudomacropus, rorAa KaK 6o.rree uoulHlte 

HarroMr4Halor A. varius. flo gro[oruqecKl4M oco6euuocrflM BHoBb ollllcunaeultft 

BuA TaKxe 6olee 6ln:ox x l. varius, rIeM K A. macropus. 

O6H.rHo A. pseudomacropus [pol43pacraer Ha necKax, Ho B sauaAHoft 

qacru apeala (a vacrHoc'rr4, Ha rore Opeu6yprcxoft o6n.) ou ut6elaer 

rlecqaHblx yqacrKoB I,I AocrarorrHo [ocrotHHo Bcrpeqaercfl JIHlub Ha 

o6saxesurx MeJra r{ MepreJlt. A. varius B 3ToM pafroHe, Hao6opor, us- 

6eraer MeJroBbrx 14 r43BecrHrKosnx o6Haxesuit u [pol43pacraer rlpelrMyurecrBeHHo 

Ha [ecKax (onuaro n CpegHeru u HuxHeu flosorxle oH 

Bcrpeqaerc.fl TaKXe Ha Iraenax). 

14. A. storozhevae Knjasev , 2002, Eor. xypn., 87 , 4 : 149 . 

T n n : Poccnl,

lOxuufi Vpan: OpeH6yprcraa o6l. (KynauAnrcxufr r.r fy6epnuucrufr 

uenrocouo.IHHKH, r{3Bec'rHrKoBbre yBaJrbr s FerseBcxovr n capaxrarucKoM 

p-Hax), Pecuy6.rrura Eaurroprocran (xpaftHuil u. 3nauuy_ 

puucrufi p-H, Ha r{3BecrHrKoBbrx yBaJrax BAoJrb r.paHtrrln c Open6ypr.c_ 

xoft o6n.). - O6q. pacrp.: cenep Axrrc6uscrofr o6n. KasaxcraHa. 

(Puc. 3). 3nAeuux ropuofi qacru loxnoro Vpala u flo/lypa.llcror.o 

IIJIATO. 

Bu4 ovenr, 6nw:or K BocroqHoerpoueiicrouy A. pallescens Bieb. r.r 

3aMerrlaer efo B |opHofi. \acru loxHoro vpara. creAennr o HaxoAKax 

ua l0xuopr Vpane A. albicaulis DC. (Yakovlev et al., 1996; pxluuuua, 

1998), rax 6ruro ycranoBJreHo HaMr{ rrpu rrpocMorpe coorBercrByrou.tux 

rep6apuux 3K3eM[[spoB, raKxe orHocrrcfl x A. storozhevqe. 

15. A. pallescens Bieb., 1819, Fl. Taur.-Cauc. 3 :4g9; DC., 1g25, 

Prodr. 2:2941' Ledeb., 1842, Fl. Ross. I ..626;lJJvawr-., 

1g95,

oco6u c [poMexyroqHHMu rIpI'I3HaKaMn' flo scefi BLIAI{MocrI'I, 3TI4 pe- 

JrI,IKTOBbre 1rOIIyJrtUuH A. palleScenS B HaCTOtu1ee BpeMt g SHa'IUte;1rHofi 

crerreHl,I rloABeprJptcb l,IHTporpecclrsHoft ru6puAu:aqnu c 6nusxutu uu- 

Aovr r4. macropus. OAHaxo 3TI{ HaxoAKI{ AeJIaIor BecbMa BepotrHblM Hailwrue 

B coceAHHx paftonax sanaAHoft .racrlr EauxoprocTaHa rrorryJrfluui,r 

A. pallescens s. str.n He 3arpoHyrblx ru6pnAnsaqueff. 

Taruu o6parovr, ceKIIHfl Xiphidium poga Astragalus npe4crauleHa Bo 

$nope Vpara l5 stlAaNar, I{3 Koropblx | (A. igosclince) pacupocrpaHeH 

Ha flonapuou Ypane, | (A. cornutus) - na CpegHeu u lOxuou n 13 - 

roJrbKo Ha IOxHovr. Bo $nopax o6lacreft n pecuy6nur Vpalrcroro pernoHa 

ceKlrr4fl Xiphidium [peAcraBreHa cJreAyrcquM trlaclroM suAos: Pecny6lura 

Kovut - 1, fleprucrar o6n. - 1, CnepgloncKafl 06lr. -- 0, t{els6lrHcrar 

o6r. -'6, Pecny6ntlrca BanroprocraH -- 7, Open6yprcxar 

o6n. - 12. 4 supla (A. igoschinae, A. karelinianus, A. austrouralensi.s u 

A. storozhevae) rsnrnorcf, 3HAeMuKaun Vpana u flpuypanla (nepeuft 

BlrA - florupuolo, ocrtulbHble rpu --- IOxnoro). Eue 2 B:zp.a (4. oropolitanus 

n A. mugosaricus) uzvecrHbl Ha reppl{Topulr Poccull roJrbKo Ha 

IOxnou Vpane. OrsocnreltHo BblcoKoe BHAoBoe 6orarc'rno ceKultrl 

Xiphidium ro Srope Ypara cBfl3aHo, c oAnofi cropoHbl, c HaJIt{qHeM 

pflAa Br.rAoB, HaxoAtuluxcq Ha BocroqHoM (nocrovHoenponefrcrue 

A. pseudotataricus, A. ucrainicus u A. pallescens) vt ceBepHoM (ceneporypaHcK?Ie 

A. tem.irensis, A. mugosaricus u A. pseudomacropus' a raKxe BocrorrHoeBponeficro-raraxcrancrufi 

A. brachylobus) npeAelax pac[poc- 

TpaHenur, c 4pyrofi, co cuequtfutecKnMl4 qepraMfi tfloporeHera Vpanlcroft 

rOpuofi cTpaHbI, O6ycnoauarullMg B93HI{KHOBeHI{g pqAa gHAeMI4tI- 

Hbrx u pe,rnrKToBbIx (AnrtnurrusHltft anraftcro-roxuoypallcxufi 

A. o r o p o I i t anas) ane.ueuro B, tIpLIAaIouIux oflpeAeJleuHyrc cauo6blTHocrb 

Qnope AaHHoro perfioHa. 

Pa6ora BhIrIoJlHeHa fipu uoAAepxxe Poccuficroro SoHga tfyH4alvreurarbHbrx 

LIccJreAoBaHHfi (npoerr Nb 02-04-49 I 6 I ). 

Jlureparypa 

EopncoBa A. l. AstragaluspseudotataricasBoriss. ll (D.rcpa CCCP. M.; 

n., t946. T. 12. C. 70t. 

B a c u n b rr e H K o LI. T. Knroq Anq ofipeAereHl,Ifl Bl'tAoB ceKIrI'IrI Xiphidiunt 

Burge n pa3AerreglIe Ha BI4AoBbIe pngr ll (Dnopa CCCP. M.; JI., 1946. 

-f ' 12. 

c.646-738. 

f auaroHoBa A. ll., @nclon B. B. Iloapoa Cercidothrix Bungepo4a 

acrparaJr - Astragalus L. ll @nopa KasaxcraHa. Annra-Ara, 1961. T. 5. 

c. 184-299. 

f onqapoB H. O., flonos M. f. Ceruns Xiphidium Bungepo4a 

Astragalus L. //

f opu a K o B c r u fi fl. JI. Acrpariur KapeJrr{HcKnit- Astragaluskarelinianus 

M. Pop. // Kpacrar KHr{ra Cpeqnero Ypana (CrepanoBcrar n flepucrar 

o6nacnr): Pe4rae r,r HaxoArul[ecr noA yrposoft ncqe3HoBeHr,rr BrrAbr pacrennfi 

u xnrornux / flo4 pea. B.H. Eonruaxora, fl.JI. fopvaxoscxoro. E,rarepr.rn6ypr, 

1996. C. 134. 

f opua Ko Bcx ufi fl. n., LLIypoBa E. A. peArne H ncqe3attlulnepacreuur 

Vpala u flpuypanra. M., 1982. 208 c. 

f y 6 a n o n I,I. A., E a ra a ca po Ba T. B., E a I a HA r,r H a T. ll. HayqHoe 

HacreArre BbrAaro[Irrxcr pyccKHx $lopucron f. C. Kapelr.rHa n 

14. fl. Knplraona. M., 1998. 95 c. 

KaMer qH P.8., IOpqeB E. A. HoBrrfi augAstragalasigoschinae(Faba 

ceae) c flonapnoro Ypata ll Eor. xypn. 1982. T. 67, Ns 9. C. 1285-12g9. 

HayrrreHKo H. 14. l,lurepeqrue $nopr.rcrr.rrrecKr,re HaxoAKn r Kypraucxoii 

o6racrn ll Beqn. JI|Y. Cep. 3. 1991. Bsrn. 3. J& 17. C.4249. 

P fl,6 n H u H a 3. H. Koncnerr $noplr Opeu6yprcxofi o6.nacru. ErcarepuH- 

6ypr, 1998. 164 c. 

C rr r u H A. K. K cucrenarnrce Astragalus brachylctbus u Astragalus varius 

(Fabaceae) // 6or. xypH. 1999. T. 84, Ne 12. C. ll7-124. 

C n r u n A. K. Hontrfi Br,ra acrpararra (Astragalus L., Fabaceae) n:] 3anaAsoro 

Kagaxcrana // Hosocrn cr{creMarrrKr4 Bbrcrunx pacreurafi. Cn6., 2000. 

T. 32. C. 92-96. 

l-( n e n € o H. H. 3naxn CCCP. L, 1976.788 c. 

B u n g e A. Alexandri Lehmanni Reliquiae botanicae, sive enumeratio 

plantarum in itinere per deserta Asiae mediae ab A. Lehmannis lg39 ad lg42 

collectarum ll Arbeit. Naturf. Vereins Riga. 1847. Bd 1, H. Z. p. 115-256. 

B u n g e A. Beitrag zur Kenntnis der Flora Russlands und der Steppen 

Central-Asiens, i. e. Alexandri Lehmanni Reliquiae botanicae, sive enumeratio 

plantarum ab A. Lehmannis collectarum (seors. impr.). St. p6tersburg, 

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B u n g e A. Generis Astragali species Gerontogeae. pars 2. Specierum enumeratio 

ll M6m. Acad. Sci. P6tersb. (Sci. Phys. Math.). 56r. 7. 1869. T. 15. N l. 

254 p. 

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1968. Vol. 2. P. 108-124. 

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herbaria of Paris (P) and Geneva (G) // Sendtnera. 1998. Bd 5. S. 24j_263. 

P o d I e c h D., Sytin A. K. Typification of Russian and some other species 

of Astragalus. I // Sendtnera. 1996. Bd 3. S. 149-176. 

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Eurasia: A checklist. Kew, 1996.724 p. 

Eorauuqecxufi, calVpallcroro orAeneunr PAH 

620026, ErarepaH6ypr, alt 315 

E-mail: tilia@r66.ru

A. K. CrtrnH 

A. Sytin 

KOHCIIEKT KABKA3CKLIX ACTPA|AJIOB (ASTRAGALUS L., 

FABACEAE) CEKUIIN ANTHYLLOIDEI DC. 

CONSPECTUS GENERIS ASTRAGALUS L. (FABACEAE) 

SPECIERUM CAUCASICARUM E SECTIONE 

ANTHYLLOTDEI DC. 

B nacroxulefi crarre Aaerc{ o6sop KaBKa3cKI,Ix BI,tAoB Astragalus L., 

BKnroqaB[rr4xcr [pexAe B ceKulrrc Halicacabus Bunge., Anq roropoft, 

oAHaKo, rpnopurerHblM fl BJrterct Ha3BaHI,Ie Ant hy lloidei DC. flpr,ruo4urcfl 

Krtroq AJrr oflpeAefleHl{fl BI,IAoB, flocrotxy BI,IAbI, rlpeAcraBJrtloullre 

3Ty CeKur,rrc Ua Kanxa3e, f,BJlflIoTcfl peAKIrMlr pacTeHt{tMr'I H HyxAaIoTcr 

B oxpaHe, B crarbe llepeqflcfleHbl r,(3yqeHHble rep6apuue o6pasqu. 

O6cyx4anrcr poAcrBeHHbIe cBt3l,t BI,IAoB ceKuI4Ir Anthylloidei u ee tto- 

JroxeHr4e cpegu 6nusrnx cerqufr. 

Cerqus Anthylloidei DC. 1= sect. Hqlicacabas Bunge) orHoclrrcfl K 

[oApoAy Astragalus poga Astragalus L. u o6teAunrer po3eroqHbre nru 

[onypo3eToqHbre rpaBtHI,IcTbIe MHOIOJIeTHUKIT C OApeBeCHeBaU.UI{M KayAeKcoM, 

otryueHHbre [pocrblMl4 BoJIocKaMI,t. Jlncrlq HeflapHorlepHcrble, 

JrlrcToqKu oTAeJrf,rcTcr oT ocn B rIepI{oA oTIlBeTaHufl I{ HaqaJra [noAo- 

HorrreHrrr pacreHnfi . Jluruu$uqnpyrcu[ect rKaHI4 rIpI,IAanr )I(ecrKocrb 

qepeuKaM u cnoco6creylor MHoronerHeMy coxpaHeHl4lo I4x tIpI{ ocHoBa- 

Hr,ur uo6eron. Cnoeo6pasuoft oco6eHHocrblo couserufi qBflterct HaJluque 

lapHbrx [pnIIBerH[ttKoB y ocHoBaHu.fl rraureqrn. Vame'Ira B Haqa- 

Jre rIBeTeHr4r rpy6uaraa, K KoHIIy - KyBIIrLlHqarat, rIpH TIJIOAOHOIIIeHnH, 

B3AyBaflCb, CTaHOBI,rrCr UapOnU4UOfi . 3arrro.Iaq ue6olrruoft, oAHofHe3- 

Annfi vranocervrxssuft 6o6, oHa cuoco6crnyer aHeMoxopHoMy pac[pocrpaHeHr,rKr 

ceMeHu. 3auaAuoa:ltarcKat ceKIItIt Anthylloidei o6leArHqer 

oKoJro 10 ar.{Aos, fipuypoqeHHux x QpuraHo[AHbIM coo6qecreaNa u 

apr4AHbrM peAKoJrecbflM, o6naAarculrlx y3KHM apeaJroM, a I4HorAa flBJIfl- 

Iouruxct .rpe:anuafr uo peAKHMr,t. 

BcneA sa A. floArexorra (Podlech, 1990), MbI IrcrIoJIb3yeM IIpIlopI,Irer- 

Hoe Ha3BaHr4e ceKqr,ru Anthylloidei DC., xorq sra rpyrr[a B [epBoHaqzutb- 

HoM rroHrrMaHl,ur BecbMa [cKyccrBeHHa u o6r'e4l,rHqer BI,IAbr, B Hacroturee 

BpeMfl orHoclrMbre K pa3HbrM rroApoAaM. Tar, o4nH I{3 rIpeAcraBJIeH- 

Hbrx B neft suAos A. calycinus Bieb., nnHe BKlK,qaIor B ceKIrHIo 

Laguropsis Bunge uoApoAa Cercidothrix Bunge. A. A. Eynle (Bunge, 

1868) orHocr.rr K ceKur4r4 Halicacabus 9 ruqon, cpeAlr Hrrx pe4.rafturue 

BrrAbr - A. physaloides DC. - pacreHr4e, co6pannoe Jrumb oAHax(Abl, 

Bo3MoxHo n Cupuu, a raKxe A. wagneri Bunge, o6urarcqufi Ha rope 

Ghodeh Dag 6nur c. 6a3eraH s 3auaAHola Arep6aft4xaue (I4paH/. CnopaAuqecKu 

pac[pocrpaHeuHrrfi na Anaronnftcnou [JrocKoropbe n Typ- 

\wu A. zederbaueri Stadlmann 6nurropogcrBeH orrucaHHoMy r43 Cupuu 

t49

A. surugensis Boiss. et Hausskn. A. khoshjailensrs Sirj. et Rech. f. ncrpeqaercr 

B ropHo-cre[HoM rrorce K)xHoro cKJroHa xp. 3trt6ypc. B flarrarapo-Arae 

ceKur,rn upeAcraBnfler A. w,illisii M. Pop. ,- sHAeMHqHlrii nuA 

xpe6ra Kyrnraur u rop Karra-Kyp.ryx, a B 3a[aAHoil. \ac1y fuccapcroro 

xpe6ra - A. pseudanthylloides Gontsch., o6r.rrarcutufi s lroJtoce 

B3ar.IMOrrpoHI4KHOBeH[r pacTuTeJrbHocru ooerraeponofi nonynyorbrHr4 ]r 

ApeBecHo-KycrapHuKoBoft Qoprraaqlrt.r cpeAHero r.{ HrlxHet.o ropHbrx lroacon 

(fou.rapor, l94l : 102). 

H. O. fosqapoa, o6parr4B BHHMaHr4e Ha orrepraHl.re Slrara, rrpeArro- 

Jroxr4Jl, qro rrpocrar K qelocrHas pou6onuquar Qopua er.o rrpeo6pa:yercs 

B 6oree cnoxHyro rurapoBrdAHyro, au$SepeHrlt4pyrcb Ha rroJroc- 

Ky 14 nJracrl{HKy, [pu 3'roM B ocHoBaHr,ul rroclreAgefi Mol.yr HaJrr.tqecrBo- 

Barb nilv orcylcrBoBarb yrnoBarbre yuKr.l. YsnAes o[peAeJreHHylo ]Bo- 

Jrroulr oHHyro HaupaBJreHHocrb B pa:Hoo6pas vn i-rLrx uop$ololnuecxux 

napuaqui.t, o H Bocuotr b3o Barcr ero AJu BbrAeJreHur BHyrpr4ceKu tr oH Hbrx 

rpyrrr, orrr{caB B ceKr.lr4r{ Hslicacabus p.r^gu Angusfa/l Gontsch., Subauriculati 

Gontsch. u Densiflorl Gontsch. o o6pa6o.me poga Astragalus Lut 

Tbr, TtroreK)rune K 6uorraopQolorl4qecKoMy rr,rrry ((TpalaKaHToBbrx acrparaJroB)). 

BuAur cexunu MegalocystrJ cQopMr4poBaHbr cyxr.rM u xapKHM 

cy6rpouuuecKraM KJrHMarou l4paHcxofo Haropbfl, B [peAeJrax Koroporo 

oHrr n pac[pocrpaHeHbr. E4r,lHcrneHnbrM npeAcraBr4TeJleM arofi cerur4u 

Ha Kanrase sBJrflelcfl Jtumb cBoeo6pasuufi A. szovitsii Fisch. et 

C. A. Mey. - 3HAeMr4rrHuft sraA Cpegneapaxcuucxofi rornonuHtr. K 

Heuy 6nu:ox A. ebenoides Boiss., [pou3pacrarcqufi Ha 3acoJreHHblx ltor{Bax 

uorblHHbrx nycrbrHt c Hulthemia persica (Michx. ex Juss.) Bornm., 

a rarxe Ha aHTpolroleHHbrx Aepr.rBarax arux coo6qecrB, r{3MeHeHHbrx 

3eMneAeJrr4eM or Bocror*roft qacru ApuaHcxolo Halopbq (llpaucrufi 

Asep6afigxau) 4o :ana4Hbrx cKJroHor Hpancroro Haropbr (cenepo-eanaAsufi 

3arpoc n Eaxrnrpcxne ropur). EyHre orHocl4:r A. ebenoides x 

ceKrlr,rr4 Halicacabus (1868), Ho ero BKJrroqeHr4e B ceKr(uro Megalocystis 

Bunge (Tietz, Zarre, 1994), rue A. ebenoides 3aHHMaer flpoMexyrorrHoe 

noJro)r(eHne MexAy ueHrpaJrbHoKo[erAalo-xopacaHcK trM A. r adde i Bazll. 

tt A. szovitsii, upe4craanrercr 6onee [paBIlJlbHbtM perreHrreM, [o3BoJlf,- 

IourHM rcHee oqeprnrb KaK rraop$oloruuecKrle, TaK H reorpa$uuecrne 

fpaHr,rrlbt cerqufi. 

C ro.rxu 3peHur $v roreorpa$rauecroft upr4HaAJrexHocrl4 ceK rrr4 ro 

Anthylloidei uoxuo paccMarpl.rBarb KaK arporrareHcKyro rpytruy 

(r currcre A, JI. TaxraAxrHa, 1972), coc'raBJrqrouyrc yMepeHHo .repMo- 

QznrHyro Sparuuro Snopu apuguo{r u cy6apra4Hoit oltacrefi Haropuff 

flepe4uefi Arrzu Aprvreno-I4paHcxofi upoanuquu. B srofi cBs3u H3ornrpoBaHHoe 

[oJroxeH]re A. daghest anicus 4onolufler HeMHot-oqncneHntrfi 

crrrrcor cJrytraeB arpolareHo-AarecraHcKofr 4utrrouxquu (n Hr4M orHocr4tct 

vr apeast Artemisia incana (L.) Druce), Bo3MoxHo, cBr.rAereJr[,crByer 

o ApeBHr{x MHrparIHoHHbrx cBr3rx MexAy Q.rropaurE .{arecraucxofo oKpyra 

Kanra:crofi nponuHrlun n Arponareucrofi rroAnpoBrrHuuu Apue- 

Ho-tr4paHcrofi nponuHqun. 

Astragalus L. sect. Anthylloidei DC., 1825, Prodr. 2 : 300; Podlech, 

1990, Mitt. Bot. Staatssamm. Mtinchen2g :464; Podlech a. Zarre,200l, 

in Rech. f. , Fl. Iran. 175 :97,p. p., excl. sect. Megalocystis. L sect. 

Halicqcabus Bunge, 1868, M6m. Acad. Sci. P6tersb. (Sci. Phys. Math.), 

s6r. 7, ll,16:74; id., 1869, l. c. 15, | :123: loH.r., 1946, so (Dn. CCCp 

12 : 413; fpoccr., 1952, @n. KanK., nzg. 2, 5 : 295; MaH4eu., 1962, so 

@r. Aprra. 4:175; Chamberlain a. Matthews,1970, in Davis, Fl. Turk. 

3: 178. 

T u n : A. anthylloides Lam. 

l. 

KJTIOq IIJI' ONPETEJIEHNfl RABKA3CKAX BI4,TOB 

,rlncrrq (3) 4-14 cu An. flo6eru ceAoBaro-cepsre, 6apxarncro-onyureHHbre 

.... 2. 

Jlucrrs l+25 (40) crr,r An. flo6eru cu3o-3eJreHbte, npr4)r(aro-onyueHHHe . . , 

J. 

l5l

2. Jlucrrs 4-5 cu AJr., noqr[ crrArque Hrru c qepe[rKarrrn 0.5 cM An.; mtcrorr- 

Kr,r npoAonroBaTo-3JrJrr{nTl,rqecKr,re, Ha BepxyulKe BbreMqaTble, BABOe CJTOXeH- 

Hbre, B qr{cJre (6) 8-10 nap. Counerne 2-5-qgerxoBoe; rlBeroHoc 5-7 cru an. 

9aureqxa l8-20 rrana An., onyureHHas 6elrruu BonocKaMlr (ranorAa c npr.rMecblo 

r{epHblx nonocr

,+'4t? 

, 

t*&V.a*r*"r,{,; ,*,/ 

4o dj' t/"1!' 

:::.. : .::, 

t4t11t,,14Vtg1,, ,. tV'a. A"; l; 

Prac. L @ororpaQnx rexroruna Astragalus mesites Boiss. et Buhse 

153

Ouucan r.r: Asep6aftAxaHa (HaxuuenaHcxar AP) (AoluHa p. Arrnu- 

4xauafr, MexAy cc. Xanara u Kaeauun). Lectotypus (Podlech, 

Sytin, 1996 : 164 ):

dl,;., 

$" 

:i 

&**-.,,-. ffi 

ffi' 

Pnc. 2.

TIarffi" 

156 

M tu 

.epffi'"'' 

, ?.:lF. 

'{j 

.b4lri 

, l d # { 

8pr?rrah rrlrttl$l*r r"j$# 

itnE:Sf. 

1 v"W 

- 

. 

.Siiift 

:sts+# 

..i#,$ffi 

Pnc. 3. @ororpaQnr ronorr,rna Asnagalus karakuschensis Gontsch.

IOxn. 3axasxasle:Apuenua. -* O6rq. pacrp.: Ces.-socr. Typurar 

(ApurHcroe naropre). 

Pearr,rfi su!, A. kaghysmani Bcrperraercr Ha rJrr{Htrcro-ule6Hucrux u 

rurrcoHocHbrx cy6crparax Apaparcrofi rornosnHu n KapccKoM rrJroc- 

Koropbe o Typqnra. Erusox x A. halicacabus Lam., [o-Br.rArrMoMy, Ao- 

BoJrbHo rur{poKo pacnpocrpaHeHHoMy B npeAeJrax ApurHcxoro Haropbfl, 

TorAa KaK MaJroa3r4arcxuit A. anthylloides Lam. eAsa 3axoAtrr B 3arraA- 

HyIo qacrb groft o6racru. HagsaHHHe BHAbr cocraBJlflrcT qenr 6lu:ropoAcTBeHHbrx 

BHKap14aHToB. 

I4ry.rernue o6parqu (specimina examinata): Apmentn: Apmuxcr

*Ou"ro" xxura Apurrucxoft CCP. I4c.reraroulne n peAKr.re suArr pacreHlrfi / 

3. |I. fa6pr.rexrH r.r Ap. Epenan, 1988. 284 c. 

T a x r a A)K A n A. JI. Eorannxo-reorpatfu.recxuft ovepr Apaparcxoft romosun;l 

ll A. JI. TaxraAxrs, As. A. @eaopor. (Dnopa Epeoana. Il., 1972. 

c.7-37. 

P o d I e c h D. Die Typifizierung der altweltlichen Sektionen der Gattung 

Astragalus L. // Mitt. Bot. Staatssamml. Miinchen. 1990. Bd 29. S. 461-494. 

Po dlech D., Sy t i n A. Typificationof Russian and someotherspecies of 

Astragalus L. // Sendtnera. 1996. Bd 3. S. 149-176. 

Podlech D., Zarre Sh. M. Sect. AntlrylloideiDC. ll Flora iranica. Graz, 

2001. lF 175. (Papilionaceae lY, Astragalus ll\. P.97-126. 

Tietz S.,Za rre Sh. M. Revision vonAstragalus L., sect. Megalocystis 

Bunge (Fabaceae) ll Sendtnera. 1994. Bd 2. S. 287-303. 

EoraHu.recxufi nncruryr una. B. JI. Korr,rapora PAH 

197376, C.-flerep6ypr, yl. rrpoQ. flonona, 2 

sytin@DG4972spb.edu

A. B. feruruaH D. Geltman 

oE3Op IOACEKTIUI,I MYRSTNTTEAE BOISS. CEKIIIil4 

PARALIAS DUMORT. PONA EUPHORBIA L. 

(EUPHORBIACEAE) 

GENERIS EUPHORBIA L. (EUPHORBIACEAE) SECTIONIS 

PARALIAS DUMORT. SUBSECTIONIS MYRSINITEAE BOISS. 

REVISIO 

Cerqur Paralias Dumort. -- oAHa ur xpynuefiurnx cerqufr rroApoAa 

Esula Dumort. po4a Euphorbia L., tK.nrouarouar, ilo MoeMy MHeHhro, 

rrrru rrorceruuft. 

floAcerquq Myrsiniteae Boiss. qnuercr, flox(aJryfr, Han6olee ecrecrseHHofi 

rpyuuofi ceKrrurl. OHa 6ula onucaHa Eyacre (Boissier, 1862) 

rar B rpegerrax ceKr{rlu Tithymalus (Scop.) Boiss. Kar 

6nuo uora:aHo K. C. Eafirosuvr (1994), Eyacre s srofi pa6ore o6o:uaqafl 

rroAceKuuh 3HaKoM $ (o verra HMeercr ero yKa3aHue). flo:rouy uox- 

Ho cqr,rrarb, qro rroAceKrlus.6ura ycraHoBJreHa vMeHHo e rrofi pa6ore, 

a He B 6onee nor4neft ny6nuraqun (Pax, 1890--91), KaK 3ro rrprrHr4Manocb 

B pr4e pa6or, B ToM rrucrte t.l Morrx (ferlrvran, 1996). 

Kparrnft o6:lop rroAceKrlr{r{ 6ur Aau B. fI. MaleesHru (1930) B cBq- 

3r,r c paccMoTpeHKeM pacflpocTpaHeHr{s BxoArulux B Hee Br,rAoB, HX [pOrrcxo)KAeHlu 

n Sunoreuun. fl.V. flpoxauoa (1949,1964) BKJrroqrr AaH- 

HyIo rroAceKur.rro B ceKq[ro Paralias (cuHonnu ceKrlr{r Murtekias (Raf .) 

Prokh.) poga Euphorhiu, xors, pauee (flpoxaHoB, 1931) npr.r4asaJr eil 

paHr roApoAa B upeAeJrax poAa Tithymalus Scop. 

B page pa6or (Smith, Tutin, 1968; Benedi et al., 1997) xa raKcoHo- 

Mr4r{ecKar fpyflrra paccMaTpr{B€Liracb B paH|e ceKur4l4, oAHaKo f cril,rTaK) 

qenecoo6parHbrM rrpr.lAepxnBarbcr ToqKr4 3peHuq flpoxauona 

(1949, 1964), yqr,rrbrBar o6qee pasHoo6pasue ceKqr{r4 Paralias. Taror-o 

xe rroAxoAa K cxcreMarr,rre srofi rpyrrrrbr rrpr,rgepxuBanca u pa4rlu$- 

Cvtar (Radcliffe-Smith, 1982) n o6pa6orxe poAa a,nn Typuar.r -- reppu- 

TopHI4, oqeHb BaxHofr A.ru lroHt4MaHr4r cl.IcreMarrKt{ rroft rpyltrrsr. 

.{aHnuft o6:op nogroroBJreH B ocHoBHoM rro MalrcpntrJrarvr fep6apur 

Eoranuqecxoro rrHcruryra uu. B. Jl. Korraapoaa PAH (LE) c rrpuBneqe- 

HrreM AaHHbtx prAa Apyrr,rx fep6apuen. flpu xapaxreprcrrlKe pactrpocl'paHeHlu 

trcrroJrb3oBaHbr KaK AaHHbre rep6apHux rolnerquft, TaK 14 

o6pa6orru poAa B pr4e (oco6euHo Hayek, 1927; Rechinger, 

Schiman-Czeika, 1964; Smitlr, Tutin, 1968; Janrconr.rf, Hraxonutl, 1972; 

Zohary, 1972; Ky:uauov, 1979; Radcliffe-Smith, 1980, 1982; pignatti, 

1982; Zohary et al., 1983; Med-Checklist, 1986; Benedi et al., 1997 

r4 Ap.). 

floAceruur Aocrarorrno ecrecrBeHHo AeJu4Tcfl Ha 3 pxga, BbrAeJIeHnux 

flpoxaHonrru (1949, 1964) ua MarepuaJre c reppr{ropr{u 6lrsruero 

159

CCCP, oAHaKo BIIoJIHe xopotuo orpaxarcqux o6ulee parHoo6paute u 

poAcTBeHHbIe cBf,3tt Bcex ee BI4AOB. 

Euphorbia L. 

Sect. Paralias Dumort. subsect. Myrsiniteae Boiss., 1862, in DC., 

Prodr. 16,2 '. 173; Pax, 1891, in Engl. u. Prantl, Nat. Pflanzenfam. 3, 

5: 1ll. - Euphorbict L. sect. Myrsiniteae (Boiss.) Lojac., 1904, Fl' 

Sicula, 2,2 : 345 (

+ flnr,rua cre6resHx irr,rcrbeB npeBbrrrraer rrrnp]rHy s 2.54 pasa. Jlyvefi eepxyuerrHoro 

couBerur 3-7 (10). Cre6enr 2-5 nana B ocHoBaHr,nr. ?u,q,euux 

o-na Kuup 2. E. veneris. 

6 (2). Cre6leBbre nr4crbr o6parnorfiqeBuAno-Jrorrarqarbre, pacu[peHHbre y cauofi 

nepxyuru, n nepxueft qacru c ry6qarun HenpaBr4nbHofi $oprvmr oxono 

0.2-0.5 MM An. . . 7. E. spinidens. 

* Cre6nenue Jlr.tcrbt SJrJrr,rlrnqecKl4e, npoAoJrroBaro-?rfin[TnirecKr4e, Jlonarrraro-Srlunrr.rqecKne, 

r4HorAa (y E. anacampsercrs) ponr6ra.recxr,t-gJrJIr,rnrr{qec- 

Kue Hlrr,r floqrr.r oKpyrrbre, pacrur4peHHbre y cepeAr.rHbr unu n nepxuefr rpe- 

TH Jrucra, Ho He y Bepxyrrrrn, n nepxnefi qacrn c :y6qarr,rr,r ao 0.2 nana Ail. r4rn 

noqrr.rqenbHoxpafiuue. .....7. 

7. Bepxyurevnoe o6ulee couserue uz 2-6 (7) nyveft. . . . . . 8. 

+ Bepxyureqnoe o6ulee cor{Berr4e r.ra (7) 8-20 lyvefi. . . . 12. 

8. Jlr,rcroqxn o6epro.rrnrpeyroJrbHo-noqKoBr{AHbre ..... 9. 

* Jlucro.rxn o6eprovxn noqrfi oKpyrrbre r,urr.t cepArleBnAHbre. Su4ervrnxn Ea- 

JreapcKl,rxo-BoBuJrr4o-naKpnr. .... ll. 

9. Jlyvu nepxyureqHoro o6qero couBerr,rr 2-3 rrarrl ronul. Cre6leBbre Jrr.rcrbt 

oKpynro-3Jrnr{nrr,rqecKHe, poru6uuecrn-SJUrr,rrrrr,rqecrrre Ao nor{Ta oKpyrnbrx 

. . . 9. E. unecampseros. 

+ Jlyqr.r BepxyrueqHoro o6ulero coqBerr,rr orono I MM ToJrul. Cre6resbre 

JIr{CTbt npoAo'IroBaTo-3flIunTI4r{ecKl,te HJIl,t JIOnaTqaTO-3JIII,InTr,rqeCKAe . . . 

... 10. 

10. Cerraena MoprquHr,rcrbre, c yceqeHHo-xoHr.rqecKnM, Ty[brM rrpuAarKoM . . . . 

* c.*.ru ;;*;;r;;,. ,", ."uo*r., ;;;;;tilr;-;;;";.:" I K;;#'{:" 

.. 5. E. armena. 

I I (8). Jlncme o6pa'uroafiueBlrAHo-Jro[arqarne. Ceuena rnaAKr,re, 2.5-3 naru ar., 

c yceqenuo-(onrirrecKuM, AoBoJrbHo 3aMerHbrM rrpnAarKoM. 3ugerunr o-na 

Kprr . ll. E. rechingeri. 

* Jlncrus floqrtt oKpyrJlble nJII{ [[poKononarqarbre. Ceuena riraAKt{e HJIr.r 

cJrerKa MopulnHr,rcrbre, 2-2.5 MM .(n., c noqrr,r He3aMerHbrM npr.rAarKoM. 

3nAerunx Eaaeapcrnx o-BoB . 10. E. fontquerfuru. 

l2 (7). CerueHa c 3aocrpeHHo-KoHl{qecKr,rM npnAarr(oM, nraAKr.re HJrr,r Mopuln- 

Hr.rcrbre . ... 6. E. marsc:halliana. 

* Ceuena c yceqeHHo-KoHrrqecxr.rM r,uru noqru oKpyrnbrM, TynblM npr,rAarKoM, 

MOpUr,rHtICTbte nJrr4 ToqeqHbre, r{3peAxa nor{Tu rnaAKr,re.... ... ... ... 13. 

13. Cemena .repreo6pasno-MoptqrrHr.rcrbre, c 3aogrpeHHo-KoHl.rqecKnM npuAar- 

KoM. . . 

g. E. myrsinites. 

* CeMeHa r.tlaAKrie, I4HorAa HecKoJrbKo tMr{aro-Mopqr,rHl4crbre, c noqru orpyr- 

JrbrM, Ha Bepxyruxe BorHyrbrM npHAarKoM . 12. E. corsicu. 

14 (l). Cre6neBHe Jrncrbr 6. u. qelrHorpafinue, BepxyrueqHoe o6u1ee coqrerne 

rs 5-8 nyueft . ... 13. E. denticulata. 

+ Cre6aesrIe irncrbr rrlelxory6varbre, Bepxyuequoe o6u1ee coqBerr,re lrs 8-17 

ryveii . 14. E. craspedia. 

ll 3amg Ns 915 l6l

Ser. L Biglandalosae Prokh., 1964, Hosoclr4 ctlcr. Bbtcru. pacr. 1964 : 

232. Cre6resue Jrr{crbq lnnefiurre, pexe truuefiHo-o6parnorfruealu{- 

Hbre, IUrr4HHo 3aoc'rpeHHbre, AJrnHa l{x npeBbruaer ut{pr4Hy s (2.5) 3 5 

pa:1. HerrapsuHn 6. M. ueubHoKpaftHbre, c 2 poxronuAHbrMn trpr{AarKa- 

MI,r, pacrur4peHHbrMrr Ha BepxytlKe. 

T y p u s : E. biglandulosrz Desf. (= E. rigida Bieb.). 

l. E. rigida Bieb., I 1808, Fl. Taur.-Cauc. | :375. E. pungens Banks 

et Sol., 1794, in Russel, Aleppo, ed. 2, 2 : 268, nom illeg., non Larn. 

1788. E. biglandulosa Desf., VlIl 1808, Ann. Mus. Paris 12: l14. 

Tithymalus rigidus (Bieb.) Klotzsch et Garcke, 1859 (publ. 1860), Abh. 

Akad. Wiss. Berlin | :94. -^ Euphorbia phlomos Candargy, 1897, Bull. 

Soc. Bot. Fr.44: 156. 

Onucau us Kpuua (). L e c t o t y p u s (Geltman. hic designatus; 

Byalt, in sched.):

3. E. oxyphyllc Boiss., 1866, in DC', Prodr. 15,2 : 1268, Benedi y 

Vicens, 1998, Ann. Jard. Bot. Madrid 56, | : 173. E- broterl Daveau, 

1885, Bol. Soc. Brot.3:33. -- Tithymalusbroteri (Daveau) Soj6k' 1972' 

eas. Nar. Mus. (Praha) 140 : 170. 

OnucaH 143 VIcnaH:z^V (>). Lectotypus (Burdet etal., 1984):

subsp. armena (Prokh.) Oudejans, 1992 (publ. lg93), Collect. Bot. 

(Barcelona) 21 : 186. 

Onucau us Apueuau. T y p u s : ((3aKaBKa3be, ?.ln{naAruu, 25 IV 

1910, A. A. fpoccreftv> (LE!). 

B peqxonecbqx, KaMeHuc'rbrx u rJrr4Hllcrbrx crerrrx 14 rroJlyrrycrbt- 

Hqx. -- Apueuua (roxn.), A:ep6afr.qxau (Haxu.renaur), Typuna (Kapc), 

Hpan (or

Mecn BTopofo noAsHla) I{3BecTeH, [o-BlrAuMoMy, ToJlbKo s Tarlru]e, a 

Ha ocraJrbHoi4 qacrv apeaila Bcrpeqalorcf, pacreHr4t, orHocfluruecf, K 

o6ouvt noABuAaM, xorfl B orAeJrbHbrx Mecrax pacreHur oAHoro Lr3 Hr.{x 

Moryr rrr,IcreHHo fipeo6raAarb. 

7. E, spiniders Bornm. ex Prokh., 1933, Cucr. o6sop uorovaen Cp. 

Azwu :208. - Tithymalus spinidens Prokh., 1933, l. c, : 208, nom. alt. 

OrrucaH us TyprueHucraHa. T y p u s : (CeB-3au. cKJroH xp. Kylra- 

TaHr, rrporr4B KrnrrJr. KyruraHr, 3oHa apqeBoro lreca, AHo r4 creHbr yulensn,12 

VI 1936, C. A. Hencxufi> (LE!). 

Ha srrxoAax rr{rrcoHocHbrx [opoA, B TpeuluHax cKaJr. -- Typxrraenucran 

(nocr.), Vr6exucran (Cypxangapbr,rHckafl o6n.), I,IpaH, A$raHzcrau 

(cen.). 

Ha reppuropuu Cpe4uefi A:uu apeal Bur,a rpeAcrarneu 4nyua Spar- 

MeHTaMrr: BocrorrHbrM (Eaaxur) u 3arIaAHbrM (ropHue cucreMbr flauupo- 

Araq: fuccapcrufi xpe6er, Kyrnraur), K [epBoMy rrprrMbrKaer eAHHcrBeHHoe 

r{3BecrHoe MecroHaxoxAeHr4e B l{pane (rrpor. XopacaH). flpoxaHoB 

(1933) [peA[oJraraJr, qro 3HaqHTeJ]bHar qacrb aperura Blr't\a, cBfl.- 

3brBarcurafl gru ,qBa $parvreura, HaxoAHTcr B MaJror4ccJregoranur,rx pafroHax 

ceBepHoro A$raHlrcraHa; TeM He MeHee 3a 70 ner, rrpo[reAr.uux co 

BpeMeH[ orrncaHr{r E. spinidens, ou 6lur o6napyxeu B ]Toft crpane rolb- 

Ko B rrpoB. @apna6 (orp. Mefirvreue). BoauoxHo, apeaJr 3roro BuAa, KaK 

u 6onruruHcrBa Apyrnx npeAcraBr{TeJrefr rtoAceKunn, ]rMeer penurroerrfr 

xapaKTep H cocror{T H3 HecKoJrbKr,rx r43oJlttpoBaHHbtx $parueuron. 

8. E. myrsinites L, 1753, Sp. Pl. : 461. -- Tithymalus myrsinites (L.) 

J. Hill, 1768, Hort. Kew : 172. 4. -- E. curttJblia Chaub., 1838, Nouv. 

Fl. Pelop. : 30. - E. pontica Prokh., 1949, so (Dr. CCCP 14 : 740, 408. 

Orucau u: IOxnoft Erponu (). T y p u s : 

Herb. Linn. 630168 (LINN!). 

Ha xauesncrbrx Mecrax, ropHblx creurx u nacr6uqax, pa3pexeHHbtx 

Jrecax. I4r'atut (xpoue cer.), Xopaarul, EocHru n fepueronuua, Cep- 

6r,rr n tlepHoropuq, Pyurruua, At6auux, Maxe4oulra, Eolrapua, fpequr, 

Typqm (Enpou. qacrb, Ces. u 3an. Auarolt-rr), Yrpauua (Kpuru), 

fpysnr, llpau (cen.); 3aHeceH B HeKoropbre crpaHbr I{eHrpalluofr Enpouu 

(9eucxaa Pecuy6nura), 4oaonlHo ycuetrruo HarypaJru3oBaJrcr B 

px4e pafioHoa Cenepuofi Auepuxl,r. 

I{enrporr,r coBpeMeHHoro apeana 3roro, HecoMHeHHo, 6onee runpoKo 

pac[pocrpaHeHHoro B [pournoM BnAa Mo)KHo cqr,rrarb Ealrraucxrail 

rr-oB, c HHM HerrocpeAcrBeHHo cBt3aH yqacroK Ha ceBepe u ranaAe AHa- 

ToJrllr,r. OcranrHtte qacru apeara s roft ulu uttoitt Mepe u3oJrnpoBaHbt, 

ttro trpuBoAr{T r{HofAa K HeKoropblu uopSonoluqecKr,rM o'utr4qnrM pacreurzfi 

u3 3Tr4x $parueuron, Koropbre, oAHaKo, HeAocraroqHbr AJtr Bbt- 

AeJreHlrq rarcoHoB rroABr{AoBoro u reM 6olee nn4oaoro paHra. 

Han6onee lr3oJrr,tpoBaHuufi SparnaeHT apeana HaxoAr.rrcq r :aua4Hofr 

tacru I4paucKor-o Haropbr, xorr He Bce o6parun, qr{Tr{poBauHbre KaK 

165

E. myrsinites Bo (Flora lranica> (Rechinger, Schiman-Czeika, 1964), 

B AeficTBHTeJrbHocru orHocsrcr K lroMy nugy. Tax, yKa3aHr4e Als KorrerAal'a, 

ocHoBaHHoe Ha o6pasuax, co6paHHbrx B Be|erarr{BHoM cocror- 

Hur.r, (Ashkhabad, Suluklu (Saratowka), ad ftnes Persiae, VII 1900, N 931, 

P. Sintenis (LE)) npuHaAJrexur x E. monostyla. VIz rpocMorpeHHofo 

MHOK) r4paHCKOr-O Marepr.{aJra K E. m1'75i77ite.r MO)KHO Ol'HeCl'r{ TOJTbKO 

o6pa:rr,rr r{3 ceBepHbrx pafiouon crpaHbr: npouaHrlnff lurqH (prov. Ghilan, 

ad jug. Amad-choni supra p. Kagostan,25 VII 1902, N 460, Th. Alexeenko 

(LE)) u Maseugepau (distr. Kudjur, in monte Ulodi, K.-H. et 

F. Rechinger, 9 VIII 1948, N 6492. (W); Khalak (zwischen Kandavan- 

Pall und Pol-e-Zanguleh, des Elburs-Gebirges, 27 Vl 

1991, Weiner (W)), a raK)Ke oKpec'rHoc'refi Terepaua (entre Karadj et 

Gach-i-Sar, l6 V 1956, F. Schmidt, N 5690 (W)). 

Xorr npusHaK Mopqr.iHrrclocrn ceMrH y 3Toro Br.rAa B ueJroM oqeHb 

BblAep)KaH, Ha rore lpyurr,r Bcrpeqalorcfl.pacl'err4fl. c fJraAKnMrl LrJrr,r uoqrn 

TJIaAKHMLT ceMeHaMH (Agxapnr, oxp. c. Maxyuqern, cKJronbr yuerra 

A4xapr4cqKalr{,24M941 , A. [unrpueea (LE); Transcaucasia 

occidentalis, Likani, 15 IV 1899, G. Radde (LE) u Ap.), Koropbre Mor- 

Jur Bo3HrrKHyrb B rrponrJrou 6.u arogapa HHTpolpeccnnuoft na6pu4 v3ar\wvr 

c E. marschalliana nnu E. armena; c E. myrsinites 3'tw pacreHufl c6luxaer 

$opua KapyHKyJrbr. K coxanenun, HeAocraroK Marepniura co 3peJrbr- 

MH CCMCHAMI4 HC IO3BOJI'CT TOIIHO BbI'CHI4Tb XAPAKTCP B3AUMOOTHOUICuufi 

lrux Br4AoB s 3axasxalle v.I4paue, qro 6rlro 6sr xe;rarenluo. 

9. E. anacampseras Boiss., 1846, Diagn. Pl. Or. Nov. ser. 1, 5 : 55. 

Tithymalus anacampseros (Boiss,) Klotzsch et Garcke, 1859 (publ. 1860), 

Abh. Akad. Wiss. Berlin I :86. 

Orrncau ra:] Marofi A:r.ru (). Lecto ty p u s (Geltman, hic 

designatus): (G-BOISS!, 

iso. - LE!). 

Ha raueuucrbrx Mecrax, B lopHbrx crelux, pa3pexeHHbrx rrecax. 

Typunr (aH4euux I{eurp. n loro-3a1. AHaro.nuu). 

Pacreunr, orrr4caHHbre rar E. anacampseros vat. tmolea Khan, ros- 

MOXHO, 3aCJTyXUBaIOT paHfa TTOABKAa Vnr4 Aa)Ke B'jAA. 

10. E. fontqueriana Greuter, 1965, Candollea 20 : 172. E. myrsinites 

L. subsp. litardierei Font Quer et Garcias Font, 1949, Buttl. Inst. 

Catalana Hist. Nat. 37 : 55. Tithymalus Jbntquerianus (Greuter) Soj6k, 

1972, Cas. Nar. Mus. (Praha) 140 : 172. 

Onucas c Eareapcrr4x ocrpoBoe. T y p u s : (G!). 

Ha u:secrusKoBbrx KaMeHr{crbrx cKJroHax. 3u.u.eunr o-na MartopKa. 

t66

ll. E. rechingeri Greuter, 1965, Candollea20 : 172. ' Tithymalus 

rechingeri (Greuter) Soj6k, 1972, eas. Nar. Mus. (Praha) 140 : 176. 

Euphorbia myrsinites L. subsp. rechingeri (Greuter) Ald6n, 1986, Mount. 

Fl.Greecel:570. 

OrrncaH c o-Ba Kpur. T y p u s : (Ep. Sfakia: Hugel zwischen den 

Alpen Mavro Korfali und Eklisrradiam, N-Fuss des Berges P6chnes. 1800 

1900 m. 3 VIII 1962, N 4849, W. Greuten (G!). 

Ha raueHucrbrx n3BecrHtKoBblx I'I AoJIoMI{ToBbIx cKroHax Ha Bblcore 

1800-2300 u HaA yp. Mopr. - SHAeuur saraAHofi qacrl4 o-Ba Kpur 

(ropuuft MaccnB Jle$ra-Opn). 

12. E. corsica Req., 1825, Ann. Sci. Nat., ser l, 5 : 384. -- Tithymalus 

corsicus (Req.) Soj6k, 1972,eas. Nar. Mus. (Praha) 140: 17l. 

Ouncan c o-Ba KopcI.IKa ((a Campolite en Corse>>). T y p u s : (?AV). 

Ha cyxux KaMeHI4crbIx yqacrKax n cy6alluttffcKoM lrorce Ha Bblcore 

1500-1800 ru uaA yp. Mopt. 3nAeuux ueHrpaJllHoil qacru o-sa 

Kopcnxa. 

Ser. 3. Denticulatae Prokh., 1964, Hosocrtt cl4cr. Bbtctr. pacr. 1964 : 

232. Herrapuuru c HepoBHbIM, 3y6qarblM KpaeM 14 MHoroqucJleHHbl- 

Mr4 rrpr4AaTKaMu, He paclIrupeHHblMu Ha BepxyuKe. cre6lenrte flI',tcTbq 

3JrJrL4nTlrqecKHe uJllE oKpyfno-3JIJu{[THqeCKUe, 

pr.{Hy B 1.2-1.5 pasa. 

T y p u s: E. denticulataLam. 

AJII,IHa HX llpeBbllilaer ullt- 

13. E. dcnticulutaLam.,lT8S, Encycl. Meth. Bot. 2 : 435. - E. rotundata 

Hochst, 1845, in Lorent, Wahderungen : 344. = E. cilicica Boiss., 

1859, Diagn. Pl. Or. Nov. ser. 2,4:88. -' ? E. pectinata Albov, 1894, 

Bull. Herb. Boiss. 2 : 640. 

Ouucan r.{3 Marofi Arun (). T y p u s : I4rlrIIocrpaqrlf, 

Aubriet (P), ony6rraxoBaHa s Desf. 1808, Ann. Mus. Hist. Nat. 

Paris 12 : l15, tabl. 15 (Radcliffe-Smith, 1982). 

Ha rauenncrblx yqacrKax, B crerlflx, pa3pexeHHblx necax, 800 

3000 rra HaA yp. Mopfl. --- Typuna (nHyrpeuuu Asarolua u xpe6er AHrlrrarp), 

HpaH (upon. KypAlrcrau, Xauagau, Eaxrapan, Jlypncrau), 

Hpax (cea. ropHbre pailouu). 

E. pectinata Albov, L c., ouracaHHnft u: orp. AprnuHa, flpoxaHoB 

(1949) orHoorJr K crrHoHnMau E. denticulata. Cylx rlo reKcry rlepBoollu- 

caij.vfl,, c 3Tl4M MOXHO COfnaCHTbCq, XOT' AJIfl rIOnHOfi yBepeHHOCTU He- 

o6xoAr,rMo r43yqeHl,Ie rl{rroBblx o6pasqon E. pec:thtata, Koropble, cKopee 

Bcero, He 6brrn [pocMorpeHbr fIpoxaHoBbIM. 

14. E. uaspedra Boiss., 1846, Diagn. Pl. Or. Nov. ser. 1,7 : 95. 

Tithymalus craspedius (Boiss.) Klotzsch et Garcke, 1859 (publ. 1860), 

Abh. Akad. Wiss. Berlin I : 87. 

OnzcaH nr Manofi Asr.la (

, _ -Kauesucrbte yqacrKrd B 3apocJrrx uz euercus aegilops u e. hrantii, Tg0 

1000 rvr uaa yp. Mopfl. -- Typuur (nnlafreru lmpOaruf, u Map4nu), 

Hpan (Eaxrapau, JlypucraH), I4par (cen. ropuue r4 rrpeAropHbre p_uu). 

Kparxo Kacaflcb SuloreHun :rofi uo4cexq[r{, MoxHo col)racr4rbc, c 

Maneenuu (I930), qro rpyIrttbr poAcrBa, coorBercrByroqre rrpr4HrrbrM 

n uacroaulefr cTaTbe psAaM, CsOprranponannab r.r pacceJrunr4cb AocTaToq_ 

HO AABHO, TO-BHAIIMOMY, B IUIITOqCHE. B rO ,*C BPEMS BPflT, TIW MOXHO 

couracarbcq c efo MHeHneu o6 0ruocureJrbHo HeAaBHeM (VerneprnvHou) 

pacceJreHur.r u auQ0epeH\Lra\nL4 BnroB ptga Myrsiniteae n, " "u"rno".l", 

upo'cxoxAe''un E. corsica HeuocpeAcrBeHHo or E. myrsinites. BoLree 

o6ocHosaHa roqKa 3p€Hr,rx fpefrrepa (Greuter, l96g), coi'acHo roropori 

(ocrpoBHbre)) BrlAbl (E. corsica, E. fbntqueriana, E. rechingeri) rpeAcras_ 

lxror co60ft oco6yro QuloreHeruqecKyro Be.ltsb, BeAyrqyro HaqaJro Herroc- 

PCACTBCHHO OT AHUECTPAJIbHO'O KOMIIJIEKCA IIOT,CEKIIUI{, TIPUqEM B I'OPA' 

'aKaq BerBb Aara Haqa,.o E. marschailiana, E. myrsinites u E. anac:ampseros. 

B ro xe BpeMf HeJrb3q cofJracr'rbcfl c cy)r(Ae'r{eM fpefirepa o rrpoucxoxAeHl4r,r 

E. rigida or (ocrpoBHbrx)) Bl,rAoB, yr{r4TblBa, AoBoJrbgo cy_ 

ulecrBeHHbre uopsololuvecK[e orJil4r{r{fl r{ xapaKTep reorpaQuqecKor-o 

pac[pocTpaHeHnq 3rofo Br{Aa. 

Cxopee Bcero, auQQepeHqraarrrrf, KaK ocHoBHbrx r-pyrr[ poAcrBa, TaK 

I,I BI1AOB B HX rrpeAeJrax [poH3otlJra eule B KOHqe .rpeT[qHofo 

I|ep]roAa. 

B veraepru'rHoM rrepr4oAe rrpoucxoArrrr r,r3MeHeHur B pacrrpoc.r.paHeHa' 

Br'rAoB' fipuqeM B .'HOBHOM B CTopoHy sparueuraq vL4 pa*ee 60.uee o6rur'rpHbrx 

apeilroB. Eonee o60cnoBaHHbre $uroreuernqecKue uocrpoeHu, 

BO3MOXHbI TOJIbKO NOCNC IPIIBJICqEHI4' AOTIONHHTEJIbHbIX MCTOAOB I,IC- 

cJleAoBaHHt, B lrepBylo oqepeAb, MOleKyJIrpHbrx, AJrf, KOTOpbrx IJaHHAfl, 

CPABHHTCJIbHO HEMHOTOqHCJIEHHAfl U CCTCCTBCHHAf, TIOACCKqI{ f, flBTflE'ICfl 

BecbMa rrHTepecHbrM o6terrou. 

Pa6ora BbrrroJrHeHa flpu rroAAepxxe poccuftcroro 

Sou4a Syu4aneu_ 

ranbHbrx r,lccleAoBaHHfi (upoerr 0l-04-49427) u rpaHra flpesnAeura p@ 

rro rroAAepxKe BeAyulrx HayqHbrx urxon (npoerr Nb HIII 219g.2003.4). 

Jltlreparypa 

E a ft x o s K. c. flo.noxenae ca6npcxrx B,.A.B B cl.rcreMe pora Euprutrbia L. 

ll Bron. MOI4fl. Ors.6aon. lgg4.T.99, nrrn. 6.C. 122_12g. 

f enrrMaH [. B. Ceu. Eupltorbiac.eae _ Moro.rafrsse ll

flpoxaHoB A.V.Pop, uonoqaft - EuphorbiaL. ll @rcpa CCCP. M.; JL, 

1949. T. t4. C. 304-495. 

fl p oxa H o B .f. 14. KoncneKrcr.rcreMbrMoroqaeB CCCP. fio6anneuur lr ua- 

MeHeHr,n // Hosocrn cncreMarr,rKu Bbrctrrr4x pacrenufi. Il., 1964. C.226-237. 

{.e p n r K o B c x a s E. f. Xopacan u Ceucran (6oranr,rrco-arponouuvecxl.rfi 

o.repr Bocrowrofi flepcun) ll Tp. npuxl. 6or., reHer. H @rreKrr., l93l . T . 23. 

c.3-271. 

Benedi C., M olero J., Simon J., Vicens J. Euphorbial. // Flora 

lberica. Madrid, 1997. Vol. 8. P. 210-285. 

B o i s s i e r E. Eupfutrbiaceae - Eupltorbieae ll CandolleA. P. de. Prodromus 

systematis naturalis regni vegetabilis. Parisiis, 1862. Pars 15, sect. 2. P.3- 

188. 

Burdet H. M., Chapin A., J acquemoud F. Types nomenclaturaux 

des taxa ib6riques d6crits par Boissier ou Reuter. Vl. Euphorbiac6es a 

Guttifdres. Candollea. 1984. Vol. 39, N 2. P.771-789. 

G r e u t e r W. Une espdce m6connue end6mique des iles Baldares: Euphorbiu 

fontqueriana // Collect. Bot. 1968. Vol. 7, fasc. 1, N 26. P. 531-537. 

H a y e k A. Prodromus florae peninsulae Balcanicae. Dahlem b. Berlin, 1927. 

Bd l. 1193 S. (Regnum Veg. Beihefte, Bd 30 (1). 

Janxorn! M. M., Huxoau!B.

B. M. BnHorpa4ona V. Vinogradova 

ABE HOBbIE KOMEI{HAUIIIi B pO.rIE LEUTEA M. prMEN. 

(APIACEAE) 

COMBINATIONES DUAE NOVAE IN GENERE LEUTEA 

M. PTMEN. (APTACEAE) 

B nacroquefi craue ocyrqecrBneHo [epeHeceHue AByx Br4AoB poAa 

Peucedanunr L. P. turcomenicurn Schischk. u P. sclerophyllum Boiss. 

et Hausskn., n pog Leutea M. Pimen. 

Pog Leutea 6nl onucau so (Flora iranica> (Pimenov, 1987) n coc.rane 

6 au4on, BbrAeneHHbrx r.r3 poAoB Ferula L. v. Peucedanunt L. Ha ocuo- 

BaHHr4 Qoprum JrucrbeB u oco6enHocrefr crpoeuar uJroAa. flsrl n: l,urx 

BrrAoB 3HAeMuKr,l I4pana, a oAnH Leutea petiolaris (DC.) M. pimen., 

c KoropbrM M. f. llraueHoB oroxecrsJrser P. turcomanicun?, 3axo- 

Ar4T, fro ero MHeHnro, e TypxrueHucraH. P. turcomanicum,flezycloBHo, 

cJreAyer BK[Ioqarb s pog Leutea, oAHaKo, rax 6yAer [ora3aHo Hr4xe, Mbr 

He MoxeM colJrac['rbcq c reM, r{To Ha3BaHue P. turcomanicum orseceso 

M. f. fllrueHoBbrM B criHoHr.lMbl L. petiolaris. 

klpaucrcuit 6orauux H. Akhani (University of Tehran) rocJre rroceurenux 

fep6apvrs, LE s 2003 r. B KaqecrBe > nu4a Leutea 

petiolaris (DC.) M. Pimen. nn6pan 3K3eMrrnrp Peucedanum turcomanicaru 

Schischk., co6pauHnfr F. K. IlluurraHtru s1923 r. n TyprueHucra- 

He. HeuparuJrbHo cAenanHr,rfi ru6op rtrrroBoro o6parqa L. petiolaris 

no6y4ul Hac H3yql{Tb BlrAbr poAa Leutea M. Pimen. n HX TaKcoHoMrrqecKne 

cBr3r4 c P. turconnnicum. 

Butt, P. turcomonicum Schischk. 6lur uepnouaqiurbHo orrlrcaH E. fI. KopoBrrHbrM 

(1924) c rop Eonlurue Salxaun (ror-o-raua4Hufr TypxueuucraH) 

no4 Ha3BaHKeM P. involucratumKorov. OAuaro, KaK BbucHl4Jrocb 

rro3AHee, Ha3BaHr,re P. involucratum Korov. f,BJrreror 6onee nogrunu 

oMoHrrMoM rro orHorrreHurc x P. involucratum W. D. J. Koch (Koch, 

1824). B cBq3r,r c trHu E. K. Illuruxnu (1950) lrpeAnox(r{Jt Ha3BaHHe 

P. turcomanicurn Schischk. B KaqecrBe nomen novum Nrs p. involucratum 

Korov. 

B fep6apurz LE xpauurcs ruuosofi o6paserl vuga peucedanurn 

turc'omanicum Schischk. Collacno (n 

AaHHoM cJryqae, P. turcomanicum), tutufiaqr{pyercf, THnoM Ha3BaHr4r, 

Koropoe oHo 3aMeHrer (r. e. P. irlolucratumKorov.). paHee (Buuolpaaosa, 

2001) naHoro 6nl o6osHa,reH 'rr{rr 

Ha3BaHLrs. P. involucratunt. xo-rfl, 

cJreAoBaJro 6tr nH6parr JreKTorr4rr, rrocKoJtbKy ABa rurroBbtx o6parqa 

3TO|O Br'rAa, co6pauune aBTopoM efo orrHcaHur B Iopax Bolrrullre 6allxaHbr, 

uMeror pa3Hbre HoMepa. Bu6pauuuft 3Aecb JIeKTOTUIJ p. involucratum 

upLtBo!'r,rrcfl HaMu Hnxe. 

170

*S,r# 

t\y"-' 

'"'u 1l 

Prac. I . (Dororpa$nr raeoruna Leutea petiolaris {DC.) M. Pimen. 

t7l

Ferula petiolaris DC. (1830, Prodr. 4 : 173), Ha Ko.l.opoM ocHoBaH 

L. petiolaris (DC.) M. Pimen., orrucaH utVlpaua (. B fep6apnu 

LE unoro 6un o6Hapyxeu r{3orr4rr Ferula petiolaris (isotypus:

Pnc.2.

Leutea turcomanica (Schischk.) V. Vinogr. comb. nova. -- Peuced(tnunt 

turcomanicun Schischk., 1950, eo

t 

'd ', 

:',3 

': 

il 

..t 

,'.i ! 

; 

. $ 

-l# 

.aaaa:::;::1* 

. 

1 t,l 

ffi E 

E. 

! 

'r{ 

ltt',X 

--: 

i:. :" 

t : 

;!. 

Pr.rc. 3. @ororpaQnr nexroruna Leutea lurcomanica (Schischk.) V. Vinogr 

1"15

1,1w" tt'd.* w#{t* 

4,' t ** n"'vt&* I'r ' d-{f' 

Puc. 4. (DororpaQr,ra c'Hrnna peucedanum sclerophyllum Boiss. et Hausskn. 

176

1015. - Leutea petiolaris (DC.) M. Pirnen., 1987, in Rech' f., Fl. Iran. 

162:449, p. p., quoad syn. P. sclerophyllum. 

Ouucan ns klpaua (

T. B. KpecroBcKaf T. Krestovskaja 

O BIIAAX CEKIII{U SIDERITOPSIS BHATTACHARJEE POAA 

STACHYS L. (LAMIACEAE) 

DE SPECIEBUS SECTIONIS SIDERITOPSIS BHATTACHARJEE 

GENERIS STACHYS L. (LAMIACEAE) 

B upoqecce H3yqeHnq BuAoB Stachys L. c reppr,rropuu [Or.o-3arraguofi 

Arnu Mbr BbrqcHr,rlu, qro pacreHur, rlpr4BeAeHHbte no 

(Bhattacharjee, 1982) xax Stachys woronowii (Schischk. ex Grossh.) 

R. Mill ex Bhattacharjee us ceKuur{ Sideritopsis Bhattacharjee, uo HeKo- 

TOpbrM rrpr,r3HaKaM 3HaquTelbHo oTJrnqaroTcfl oT Bcex Apyfr4x Br.rAoB 3Toro 

poAa. Bo-nepnsrx, r4x rbrqnHKtr noqrr,r ueJrHKoM cKpbtrbr a rpy6re 

BeHquKa, pexe cJrerKa BbrAarcTcr !r3 Hee, a Bo-BTopbrx, rx cror6un uueeT 

HepaBHbre Jro[acTr.r ,-- yceqeHHyK] BepxHrorc 14 cJrer.Ka pacunpeHHyro 

I4 yKopoqeHHylo HlrxHrcro. V noAaBrflIoulero 6olturnscrBa BnAoB 

Stachys rblrrr,rHKr4 He 3arJrlorreHtr n rpy6xy BeHquKa, a pacrroJlaralorct 

rroA ero repxuefi ry6ofr. JlnItrl y S. iherica Bieb., S. sparsipilosa Bhattacharjee 

et Hub.-Mor. r4 HeKoropbrx ApylHX Br{AoB Stachys rbrr{rHKu 

cJIerKa Bbrcryflaror ras rpy6rra BeHr{r{Ka. MosoruuHnfi poa Phlomidoschema 

(Benth.) Vved., TbtqHHKr,r Koropolo 3aKJrroqeHbr e rpy6xy BsHrrr{- 

Ka, Mr,r paccMarpr,rBaeM B paHre caMocrotrTeJrbHor-o poAa. tlro racaercr 

rronacreii cron6ura, To Bce yugl't' Stachys r/rMeror cron6ux roJrbKo c 

oAr,rHaKoBbrMH AoJtflMlt. Buecre c reM, yra3aHHbre BbtIIIe [pr43HaKr4 xapaKrepHbr 

AJrr rrpeAcrasr,rrefleft poga Sideritis L. 

B csoe BpeMfl Eyacle (Boissier, 1859) onr.rcail pacreHufl n: Typqnr,r 

B KaqecrBe HoBolo BuAa poAa Sideritis -- S. balansce Boiss. flo rouu- 

JreKcy rrpr,r3HaKoB -- l0-xlurosar qatlreqKa c rpeMr BbrcoKo cpocullrMucr 

BepxHr,rMu zy6uauw, BeHqr4K c y:rofi rpy6xofi, rrprMocrosqeri u noqrr.r 

rrJrocKofr nepxHefi ry6oft, a raKxe uo o6uleuy ra6arycy fror Bl.rA 

BrrorHe coorBercrBye'r poAy Sideritis (upusHaxu (rrJroAtrKoB)) (:perraoo) 

r4 nbrJrbueBbrx 3epeH y oAHonerHr4x Sideritis n Stachys oveHu oquoo6pas- 

Hbr rr He Mofyr 6nrt ucuoltr,3oBaHbr B KarrecrBe AuarHocrr4qecrux). Tuqr,rHKra 

Sideritis balansae, TaK )Ke, xar u y Stachys woronowii cJrer.Ka Bbr- 

Aanrcr ra: rpy6rcu BeHqtlKa.3aueruu, rrro rto AaHHoMy upusHaxy S. balansae 

orJrr,rqaercr or 6o;u,uruscrBa Br4AoB poga Sideritl.s, orHocauler-ocr 

rc rpu6e Marrubieae Endl., rlanuoft orrtn.rr4renurofr oco6euuocruro 

xoropoft flBJrrercr HaJruque rbrqr.rHoK, crpbrrblx n rpy6xe BeHquKa. OA- 

HaKo H y HeKoropbtx Apyrr4x BlrAoB Sideritis, Harrpr4Mep y S. ntacrostachyos 

Poir. ns ceKur4r.r Marrubiastrunt Benth., pacrrpocrpaHeHHofi Ha 

Kanapcrux o-Bax, TbrrrHHKr4 TaKxe cJrerKa BbrAarol.cfl ra: rpy6ru BeHquxa. 

K rouy xe, BeHqrlK S. balansae fipeBbruraer o6ulyro AJnlHy qa[req- 

K!r, a He cKpbrr ee :y6qanar.r, KaK y ocraJrbHbrx oAHoJrerHr,rx BrraoB 

sideritis n3 ceKrrr{r4 Hesiodia (Moench) Benth. Bo:vroxuo. rocneaHee 

t78

o6crorreJrbcrBo Aaer rIoBoA AJtt ttepecMorpa cl{creMbl poga Sideritis, 

rrro rpe6yer 6ollee AeranbHblx 3HaHI4ft no cucreuarHKe 3Toro poAa. 

I,IsyueHue pacre*Vir, orHeceHHblx B yrloMflHyl'ofr srrrue 

K Stachys woronowii, tto3BoJIHJlo HaM cAeJlarb BbIBoA, trro B Aefrcr- 

BUTeJrbHOCTr.r OHrr rrpnHaAJlexaT K poly sideritls, a llueHuo r s. beilansae. 

flpunoguu ero crrHoHI4MttKy, a raKxe cBeAeHuq o rutre, 3KoJIorHr{ u reorpa$u.recroM 

pacnpocrpaHeHulr. 

Sideritis balansae Boiss., 1859, Diagn. Pl. Nov. Or. s6r. 2, 4 : 35: id., 

1879, Fl. Or. 4 : 707; Briquet, 1896, in Engler u. Prantl, Nat. Pflanzenfam. 

4, 3a : 231 IOs., 1954, so @n. CCCP 20 : 272; Rech. f., 1982, Fl. 

Iran. 150 : 106; Meur.rqr., 1987, Bo Or. Aprvr. 8 : 34. - S. woronow'ii 

Schischk. ex Grossh., 1932, n fpoccr., @n. Kasr. 3 : 290. - Stachys 

pseudosiderifi.s Bhattacharjee et Hub.-Mor., 1974, Notes Roy. Bot. Gard. 

Edinb. 33,2:291; Bhattacharjee, 1980, Notes Roy. Bot. Gard. Edinb. 

38, I : 90. - Stachys woronowii (Schischk. ex Grossh.) R. Mill ex 

Bhattacharjee, 1982, in Davis, Fl. Turk. 7 :253: Heller et Heyn, 1986, 

Consp, Fl. Or. 3 : 138. 

Onncan u: Typuuu (>); t y p u s : G -- Boiss. 

Ha cyxnx rpaBgHl{crbrx u ure6Hlrcrbrx cKJIoHax, B 3apocJrtx MaKBHca. 

- KaBKa3: IO)KH. 3aKasrasl,e (AprrleHur, A:ep6afigxatl), [Oro-3au. 

AsNr (cer.-sau. I,lpau, Typunr, Cuput. Jlnnau). 

K coxaneuurc, HaM He yAzurocb o3HaKoMllTbcfl c rl.ttroBblu o6pa:uou 

3roro BuAa, Ho 3K3eMrIJItp, npolrl4rl.rpoBaHHblft Eyacte (Boissier, 1879) 

so (Flora Orientalis> (Gumush-Khane, Bourgeau), 6un H3yqeH HaMrr B 

fep6aprara tr{epycalurucKoro yHl,IBepclrrera (HUJ), raM xe 6btrl 

r,rccJreAoBaH px4 o6pa:qoe nr Typunu u Cupuu. flocrolrry n fep6aprara 

LE upe4craBreHbr Marepualbl ToJIbKo I.r3 [OxH. 3arasragtr (Aprr.reHur, 

A:ep6aftgxaH), a us 3an. A:utE oHrI IroJIHocrbIo orcyrcrByK,T, cqlrraeM 

qenecoo6pa3HbrM rlpounrrlpoBarb r4ecr rep6apuue o6paruH [r3 llocJre- 

AHero peruoHa, [3yqeHHble HaMH s HUJ. 

llccfleaosauuHe 3K3eMn[tpbr (specimina examinata): Typunr: > (HUJ!). Capur:

Bot. Gard. Edinb. 38, I : 90, excl. sp. Stachys obscura Boiss. et Balansa 

et S. zoharyana Eig. 

T y p u s: Stachys woronowii (Schischk. ex Grossh.) R. Mill ex 

Bhattacharjee (= Sideritis balansae Boiss.). 

,{na 4pyrux nuga Stachys obscurq Boiss. et Balansa u S. zohar},ona 

Eig., npex4e BxoAr,rBurue B cocraB ceKul{u Sideritopsis, MbI o6beAHHteM 

B HoByrc ceKunro - Pseudosideritopsis, HMeIouIyrc poAcrBo c ceKUHefi 

Campanistrurm (Habrl.) Reichenb. Or uocneAuefi oHa orlur{aerct AJIHH- 

HbrMr.r [puuBerHuqKaMr4, 6. u. goyry6urrun qa[teqKaMH H AJII,IHHbIMI{ 

UBCTOHO}I(KAMU. 

Stachys L. sect. Pseudosideritopsis Krestovsk. sect. nova, Stachys 

sect. Sideritopsr^r Bhattacharjee, 1980, Notes Roy. Bot. Gard. Edinb. 38, 

I : 90, p. p., excl. typo, quoad S. obscuram et S. zoharyanam; ead.,1982, 

in Davis, Fl. Turk. 7 :253, p.p.- Herbae annuae. Caules floriferi erecti, 

a basi ramosi, dense patente pilosi vel hispidi. Folia caulina mediana 

oblongo-ovata vel ovata, serrata vel crenato-serrata, basi cordata vel 

attenuata, petiolata vel subsessilia. Verticillastres 4" 6-flori. Bracteolae 

lineares vel lanceolatae, herbaceae, spinescentes, tubo calycis aequales. 

Pedicelli 2-5 mm lg. Calyx subbilabiatus, campanulatus, prominente 

nervatus; calycis os cum annulo pilorum densorum provisum. Corollae 

tubus in calyce subinclusus. 

T y p u s: S. obscura Boiss. et Balansa. 

OAHonerunru. I{nerouocnue cre6lu IrpflMble, or ocHoBaHuq BerBl{- 

cTble, fycTo oTTorIbIpeHHo BoJIocHcTblg I4JlIl UIeTLIHIIc'to BoJlocplcTble. 

Cpegurae cre6neBbre Jrucrbt rlpoAorroBaro-f,ftIreB[AHble l{illr sfrqenuA- 

Hbre, [nJrbqaTble I1IJln f opoAqaTo-[HJlbilaTble, B ocHoBaHuI{ cepAIIeBHA- 

Hbre r{JII4 OTTrHyTbre, qepeu]qaTbre HJII4 tlor{T}I cHAtqIIe. flapultarlulte 

cor.lBeruq 4--6-qserxonue. flpnqnerHl4qKu lusefiHlte I,IJU{ JIaHueroBI{A- 

Hbre, TpaBtHncrbre, cJrerKa KoJlIoqI4e, paBHbre rpy6xe qaureqKl4. I{nero- 

HoxK?r 2-5 rrarr,r rur. tlaureqra cJrefKa 4ryry6aa, KororoJrbqarat, c BblpaxeHHbrMr{ 

xr4rraMu, B ycrbe c KoJrbuoM fycrbrx BoJlocKoB, Tpy6xa nenqltKa 

He BbtcTy[aeT I43 rrau]eqKu. 

T n n : S. obscura Boiss. et Balansa. 

PacnpocrpaHeHne: 3an. A:rar. 

I{r'I IO q ln n O II P E!, EJI E H U fl I A/IO B 

l. Jllrcrss nunrvaro-ly6qarrre.{aure.{Ka rrlerr.rHrrcro-Boiroc}rcrar, l0-10.5 rurrr 

Ar. Beuqux 14-15 rr,tt"r Ar. Pacrenus 40-45 crr,r Bbrc. ....... l. S. obscura. 

* llttc'rrs MenKoropoArrarrre. 9aureqKa rycro AJrrlHHoBoJrocucras, I l*14 rrru 

An. BeH.rr.rr 18-19 rrau An. PacreHr..rs 25*35 crrl Bblc. . .. .. 2. S. zoharyana. 

l. S. ohscura Boiss. et Balansa,1859, Diagn. Pl. Nov. Or. slr.2, 4 : 

40; Mouterde, 1879, Nouv. Fl. Liban Syrie, 3, 2 : 153; Bhattacharjee, 

1982, in Davis, Fl. Turk. 7 : 254; Heller et Heyn, 1986, Consp. Fl. Or. 

180

3 : 136. - S. arqbica Hornem. var' minor Boiss., 1879, Fl. Or. 4 : 747 ' 

onucan r.r3 Typur,rH (); t y P u's : G - Boiss' 

Ha raNaeHucrblx cyxnx cKrroHax' -t' [Oro-3au. Arus (toxH., tot-orocr. 

Typuufl, CuPL4fl, JIranaH). 

2. S. zoharyanaEig.,1948, Palaest. Journ. Bot' Jerusalem ser.,4: 172, 

fig. 1; Feinbrun, 1978, Fl. Palaest. 3:132; ead', 1991, Analitic Fl' Eretz- 

Israel : 569; Heller et Heyn, 1986, Consp. Fl' Or. 3 : 138. 

Ouncan uzVlzpautx: hoIotypus:

I4. M. flecnona [. Pesskova 

OE3OP POAA LTNARTA MILL. (SCROPHULARIACEAE) 

BOCTOTIHOIZ EBPOIbI I{ KABKA3A 

CONSPECTUS GENERIS LINARIA MILL. 

(SCROPHULARIACEAE) EUROPAE ORIENTALIS 

ET CAUCASI 

B cucreuaruKe poAa Linaria Mill. (nluruxa), uecuorpfl Ha MHoloqucJreHHbre 

uccJreAoBaulrfl, ocraercf, AoBonbHo MHofo upo6.rreu. Asropbr 

3Haqr4TeJIbHo pacxoAflTcfl B IloHnMaHHr4 OTAeJtbHblX Bt4AOts U HaABLT- 

AoBbrx raKcoHoB. ,{;u reppnropr.rn CCCP JL A. Kyrrpnsnonoil (1950, 

1955) npunoAuJlocb 59 nu4on rbHqHoK, r.rs Hr,rx 38 glr Enporrefrcxoil 

.racru Lr 20 --- Als, Kasxasa. O6pa6orra poga Linaria ao (utrue ), cAeJraHnas 

Jl. VI. VlsatuHoft (1981) 6oree 20 rrer Ha3aA, coAepxur 20 nu4on. 

B uocle4uee BpeMfl glr Kanxala M. E. KauanaAse (1988, 1989) 6nno 

yKa3aHo 23 suta, tO. JL Menuurun (1998, 1999) - 25 era4oe. B mraporoft 

Snope Hacqr,rrbrBaercr oKoJro 150 nuqoa rbHqHoK (Sutton, 1988). 

BaxHoe Mecro B crlcreMarr{Ke po4a Linaria yAeJrrercr KaprroJroruqec- 

KrrM rrpn3Haravr. flepanfi uonorpaS rpu6n Antirrhineqe Chav. (oua 

BKrloqae'r u pog Linanc) lllanau (Chavannes, 1833) nuepntre o6pa'na"rr 

BHr4MaHr4e Ha CBr3b UOptpOnOlUlr CeMrH 1,1 BCefO pacTeHut, oAHaKo efo 

oilucaHnr, ceMf,H KacarcTct roJtr,Ko sHerunefr uopQotrourn u ucltonb3y- 

IOTCT BO BHyTpT4pOAOBOM AeJleHr4H AJI' pa3fpaHuqeHtrf KpyrrHbrx fpyrlu, 

BbrAeJleHHbrx [o rrpu3HaKaM Bere'raruBHl,rx opraHor. PoA Linaria l].fa- 

BaH pa3Aenun Ha ceKr-tr4r{ Cymbalaria, Elatinoides (= Kickxitt), Linaria.strum 

(= Linaria), Chaenorhirutm. ?ru cerur{r{ coorBercrByro'r orAerrbHbtM 

poAaM B r4x coBpeMeHHOM rrOHUMaHnr4. 

CoepenaeHurre cncreMa'rr{qecKr{e pa6oru uo rpu6e Antirrhineae Chav. 

se o6xoArrca 6es npuueHeHnfl cKaHlrpyrcuelo 3JreKrpoHHoro Mr,rKpocKona 

(C3M) Arr KaprroJrofr4rrecKux r{ccJreAoBaHr4fi. floly.reHHbre AaHHble 

6unu o6o6uleHbr B uouorpaQuu Carrona (Sutton, 1988). Pe:lylb'r.arbr 

r4ccJleAoBaHlrr ceMrH KaBKa3cKrlx BHAoB orpaxeHbl n pa6orax M. E. KarraHaA3e 

(1988, 1989). Hcnaucrue au4u 6nlu r43yqeHbr Xyauov c coaBropaMr,r 

(Juan et al., 1999), a raKxe Cerappa u Marey (Segarra, Mateu. 

200 I ), xoropbre HccreAoBaJrH yJrbrpacryJrbrrrypy lt aHaroMr4K) ceueHuoli 

Koxypbr. OAuaro ?'rr.rMr{ aBTopaMr,r c rroMourbrc C3M 6lura uccneAoBaHbl 

He Bce BHAbr, rrpon3pacraloulr4e Ha paccMarpluaeuofr HaMr{ TepptlTopt4r4. 

Kporrae roro, Mbr pacxoAuMcfl c yKa3aHHblMu aBTopaMu B rroHr4MaHuu 

fPAHUq OTACJIbHbIX TAKCOHOB I,I B BOI]POCAX BHYTPI4POAOBO|O ACJIEHI4'. 

Hauu 6luro [peAlplrHrro KoMrri]eKcHoe HccJreAoBaHLre ynbrpacKyJrburypbr 

rroBepxHocrrl ceuennofr Koxypbr rrptr roMoulu C3M (20 un4oo) r,r 

ee aHaroMuqecKofo crpoeHrlfl (24 r'n4a, u3 Kotopbrx 23 ncrpeuarcrcr Ha 

182

eppuropul{ Bocroquofi Erpouu ta Karrara). flonyveuuue AaHHbte lto- 

3BoJrHJrH BbItBr4Tb HoBbre raKcoHoMlrqecKu 3Har{LIMbIe rIpK3HaKIl. Ceueaa 

L. volgensis Rakov et Tzvel. u L. sabulosa Czern. ex Klok, [3yrreHbr 

HaMr4 BrrepBbre. 

I4cuorrgosaH?re KapfioJroruqecKux 14 MaKpoMopQoloruvecrrix l]pr4- 

3HaKoB no3BoJII4Jlo HaM rrpeAJroxuru Hosnfi BapnaHT cncreMbl poAa 

Linaria [pr4MeHureJ]bHo K Br.raaM Snopn Bocroquofi Enporru (n rpanuuax 

EBporeficrofi qacrn 6Hstuero CCCP) u Kaexa:a. Harua cr4creMa 

rrpeAcTaBJlfler co6ofr AorroJrHeHHyK) H r,r3MeHeHHya Bepcr4lo ceKqHoHHoro 

AeneHr4r poya Linaria, [peAJroxeHHoro EeHrauou n BerrurreftHou 

(Bentham, 1846; Wettstein, 1891) r rprrHnMaeMoro MHorr4Mu aBropaMl4. 

TaxcoHottrltqecKn 3Haq[MbIMI,r ceKIII{oHHblMrI IIpr,I3HaKaMu flBJrqlo'rcr: 

xr43HeHHa.fl QOprraa, ocooeHHocrH Ju,rcropacrroJroxeHr4fl r4 crpoeH[s counerufi, 

pa3Mepbr BeHqr,rKa, uop$onorur c€MrH, a raKxe prA apyrprx Kaprtortofr4qecKl4x 

AaHHbIx, Koropbre BKJrroqeHbr HaMu B onucaHuq cexuuft. 

Hauru Kap[ororuqecKr{e r{ccJreAo BaHr,u corJracylorcr c [peAJrox(eHurrn 

JI. A. Kynpuxuonofi (1950) pa3AereHr4eM ceKqr.rr4 Linaria Ha noAceKrlrru 

Linariq u Laeves; rrocJreAHflt rroAceKuufl BaJruAr43r,rpoBaHa HaMrr 

[ocpeAcTBoM rrpnBeAeHl.rt JlaTr,rHcKoro AuafHo3a. 

OnncaHa HoBar ceKur.u . Cretaceae Pesskova, BKrroqanqas.2 slr- 

Aa--- L. cretacea Fisch. ex Spreng. (Enponeficras,Poccufl, u VrpanHa) r.r 

L. ma c r op hy I I a Kuprian. (Cenepo-3anaAH ufi Karaxcrau). 

.II;rr nsyueHHofi 'reppr,rropr{u HaMr.r upr4BoAflTcq 24 sn&a, o6leAnueHsue 

n 8 cexunfi, H3 Hr,rx Ha Kasra:e flpou3pacraror 19 nu4on, s Bocroq- 

Hofi Enpoue -- 13. 34 BHAoBbre HalrMeHoBaHur cBeAeHbr B cr4HoHr4Mbr. 

PrA suAos -- L. genistifolia (L.) Mill., f. grandiflora Desf ., L. odora 

(Bieb.) Fisch., L. macroura (Bieb.) Bieb., n L. kurdica Boiss. et Hohen., 

nprrHrrbr HaMrr B rur.rpoKoM cMbrcJre. L. acutilobu Fisch. ex Reichenb. paccMarpr.rBaercr 

3Aecb B KarrecrBe pa3HoBraAHocrn L. vulgaris Mill. BsraeileHue 

L. canadensls (L.) Dum.-Cours. B pog Nuttallanthus D. A. Sutton, 

IIO HaIULIM AaHHbIM, HeOIIpaBAaHHO. 

Clegyer 3aMer[Tb, rrro AereHHe poga Linaria Ha 2 ceKuHH c KpbrJrarrrrrau 

($ Lycorrhinum) r,r 6ecrpulurrau ($ Leontoruhinurz) ceueHarur4, BocxoArrrlee 

r .(rouoprre (Dumortier, 1827) u Byacre (Boissier, 1879) 

rpyuubr $ Oblongae w $ Discoideae coorBercrBeHHo, He ot'pax(aer Bcelo 

uuoroo6pasnr ceMrH JlbHtHoK, Ho, TeM He MeHee, [clroJtb3ye'rcfl Aocra- 

'rorrHo rxr{poKo. JL M. Kervrylapua-Hara4:e (1967) Bo3BeJra lpyuuu Eyacbe 

B paHr cexqufi. B:apy6exHofi lnreparype (Chater et al.,1972; 

Davis, 1978) ncuolb3yercfl Kareropr4r (r-pytrtta>). V 6o.rrurlrucrBa BHAoB 

c ceMeHaMH, Ha3brBaeMbrMr,r

Ceuena Br4AoB poga Linaria qepHbre uJrrl qepHo-KoplrqHeBbre, AHcKo- 

BI{AHbre, c ruHpoKr.{M KpbrJroM (Auarrrerp ceMeHr,r 2.5 3.5 nana), 6lecuulne, 

lrnn 6es rBHo BbrpaxeHHoro KpbrJra (

Boiss., 1879, Fl. Or.4:370, p. p. -- L. sect. Discoideae (Boiss.) Grossh. 

ex Kem.-Nath., 1967, B fpoccr., @1. Kanr., utg. 2, 7 : 476, p. p' - L. 

grex Discoideae (Boiss.) P. H. Davis, 1978, Fl. Turk. 6 : 666, p' p 

Typ u s: generis lectotypus. 

MuoloneruuKtl c oqepeAHbIMH JtuHeft HbIMLI I{JIII HureBUAHbIMvr Irklc- 

TbTIMLI; y HeKOTOpblx BlIJloB ocb couBeTllfl, uBeToHoxKlr u Bel'erarl4BHble 

opfaHH c onylxeHlreM [r3 xeJre3I{cTbIX t4JlI4 tIpOCTbIX BOJIOCKOB. qaUreq- 

Ka rracro [ileHrraraf,, xoTt 6bI I{3HyrpI,I C XeJle3I4CTbIM OIlyIUeHUeU. BeHqlrr 

xerrmfi r,IJu4 Ju,IJroBbtfi. 

CeueHa 2.5-3.5 MM B ALIaM., AI,IcKoBhAHbIe, c unpoKnM llepe[oHqa- 

TbrM KOJrbUeBbrM KpbIJIOM, ITpOXOAfluILIM qepe3 

Qyunrynyc (urupraHa 

Kpr'ura 6. m. panHa Al4aMerpy reJla ceMeHn). Telo ceMeHI'I l]otlKoBuAHoe, 

rrraAKoe unn 6yropuaroe. CreHKI.r KJreroK 3K3o'recrbl, oco6eHHo Hapyx- 

Hbre [epI4KJruHaJIbHbIe, 3aMeTHO yroJIqeHHbIe. Bropuvuafl cKyJlbllTypa 

la[KJuro3Hat, IlHorAa c reHAeHlruefi r o6pasonaHl'IK) cerKI'I. Snuryrnrynrpuuft 

BocK rlpeAcraBJreH orAeJIbHbIMI,I, AoBoJIbHo MeJlKnMK, IlJtoc- 

K[{MH fpaHynauu. 3HLocIIepM rnaArnfi . 

JI. A. Kynpl,tf,HoBa (1950, 1955) pa3Aerll{rla ceKql{rc Linaria Ha rIoAceKuI,tI{ 

Tuberculatae Kuprian, (= subsect. Linaria) u Laeves Kuprian. uo 

HaJrrrqulo I,tJIn OTCyTCTBIAIo 6ytOpKoB Ha AI4CKe CeMeHn, qro, KaK 6bIJIO 

ycraHoBireHo HaMI4, KoppeJrllpyer c [pI43HaKaM[ BTOpLrrrHoii cnytrlrtrypbr 

KJreroK 3K3OTeCTbL 

Subsectio l. Linaria. - - Linarit sect. Linaria subsect. Tuherculatae 

Kuprian., 1950, Tp. 6or. uncr. AH CCCP, cep. l, 9 :42; ead., 1955, so 

@r. CCCP 22:193. 

T y p u s : generis lectotypus. 

Beuqur xeJlrblft. CeueHa c 6yropvaroft nepnu'troft cryllurypoii; 

KpbrJro c saMerHofi BbreMKoft B o6Jlacrl4 SyHurynyca, B'l'opl{qHafl cKyJIbrrTypa 

cerqaTafl. 

l. L. vulgarlls Mill., 1768, Gard. Dict., ed. 8, no 1; Chav., 1833, 

Monogr. Antirrh. : l3l; Benth., 1846, in DC., Prodr. l0 : 273; Boiss., 

1879, Fl. Or.,4 : 372;Kyrryutu., 1950, Tp. Eor. uucr. AH CCCP, cep. 

l,9 : 46; oHa xe, 1955, so

Iurua, Eelapycl, Vrpauua, MorAosa. Ka:axcraH (:an.). Kanxar: fpysnr, 

Apueu O6q. pacup.: CxauA., Ar.n. u [enrp. Enpoua, Cen. 

A:nr, 3arr. Cu6upa, Cea. Aueprara (:auocuoe). -- 2n = 12. 

CarroH (Sutton, 1988) orvre.raer, r{ro orrncaHue Mntlepa (Miller, 

1768) ouroraHo Ha xuBbrx pacreHr,rrx EoraHuqecroro caAa gencn, u oH 

He AaBaJr ccbrJroK Ha JurHHeeBcrne o6pa:qu Antirrhinum linaria; B KaqecrBe 

rlrra Carron yKa3brBaer o6pareu u: rcp6apur Mulnepa (, 

sine leg., Herb. Mill, -.* BM) 

Ha6nrcAaercfl H3MeHqr4Bocrb rro pa3MepaM u $opue Jrncra v ryclo- 

Te xeJre3r,rcToro o[yu]eHr4r Bet'eTaTr{BHbrx 14 r'eHepaTr4BHbtx opf aHoB. 

,{,onr oco6efi c olyueHr4eM couBsrtrr u cre6trq H er-o rycrora KrtrHzurbHo 

Hapacralor K Ioro-Bocroxy. t{acrr pacreuuft r,rMeer Hr{TeBI,tAHbre Jtucl'bq. 

JL A. KynprlsHoBa (1955) yKa3btBaJra, rrro HrlreBr.rAune nucrtq csofrcrBeHHbr 

pacreHr,rqM L. vulgaris, rrpor{3pacraloulr{M Ha 6oponux rrecKax. 

b) var. acutiloba (Fisch.) Chav., 1833, Monogr. Antirrh. : 132. 

L. acutiloba Fisch. ex Reichenb., 1827,Ic. Pl. Crit. 5,14,t.6ll; Kyupraru., 

1955, so Or. CCCP 22:201; Llvauwua, 1981, so Ol. Erpon. uacru 

CCCP 5 :234. -- L. vulgaris f.latiJblia (Kryl.) Kryl., 1907, (Dn. A.,rlr., 

4:926. --L. vulgarisvar.latifolia Kryl., 1939, (Dl.3au. Cra6., l0:418. 

Ouncau us fiaypuu rro pacreHnrM, BbrpaxleHHbrM r.r3 ceMrH, rrpncnaH- 

HbIx aBropy oflucaHuq BnAa (D. E. @ltueporra (?LE). 

Jlyra. -- Bocr. Enpoua: Poccur (MypuaHcxar n ApxaufeJrbcKaq 

o6r., Pecny6.rrnra Koun). O6q. pacrp.: 3au. u Bocr. Cr.r6npr. 

2n = 12. 

Ornlr.raercq o'r L. vulgaris var. vulgari.s 6olee Kpy[HbrMr4 14 rxr4poKtruu 

(ao 12 nau) Jr[crbrM]r, orcy'r'crBr,reM ofiytueHuq, Hepa3BerBJreHHbrM 

couBerueM u 6onee KpyrrHbrM BeHqtlKoM (15-20 MM AJr., He crrurar ruflopqa), 

no ceMeHa r4.qeHTurrHbl ceMeHaM ruuosofi pa3HoBlrAHocru. B o6;racrr,r 

pac[pocrpaHeHr,rr L. vulgaris var. acutiloba Bcrpeqaorcq 14 Tuuuir- 

Hbre 3K3eM[ttpst L. vulgaris; qacro o6e pa3HoBrrAHocrr.r lrpor,r3pacraror 

COBMCCTHO. 

2. L. hiebersteinii Bess., 1822, Enum. Pl. Volhyn. : 25; Chav., 1833, 

Monogr. Antirrh. : 155; Benth., 1846, in DC., Prodr. l0:273; KyrrprarH,, 

1950, Tp. 6or. uHcr. AH CCCP, cep. l, 9 :44; ona xe, 1955, Bo (Dl. 

CCCP 22 : 197: Kena.-Har., 1967, s fpoccr., (Dlr. Kasx., uzg. 2, 7 : 477; 

Chater et al., 1972, in Fl. Europ.3:232; klsa*uua,lg8l, so

Ivanina, 1981, so @n. Enpon. .racru CCCP 5 : 233. - L. biebersteinii 

subsp. maeotica (Klok.) lvanina, 1981, l. c.:233. --- L. vulgaris auct. non 

Mill.:llluarnr., 1886, (Dl. [Oro-3an. Pocc. :429,p.p. --,L. italicaauct. 

non Trev. : CraHKoB u Tarres, 1949, OnpeA. pacr. Enpon. qaclr4 

CCCP: 798. 

OuucaH c Vrpauuu (). 

CrenHne r4 cJra6o3aAepHoBaHHbre [ecqaHbre, KaMeHnorbre u MeJroBbte 

rroqBbr. Bocr. Erpona: Poccur (Bonx.-Kana.: rcr; Bonx.-[ou.: tor; 

HnxH.-r{ou.; 3arolx. : rcr; HuxH.-Bonx.), VxpauHa, KasaxcraH (rarr.). 

KasICa3: Poccus (,{arecraH). - O6q. pacrp.: IOxH. Erporn. - 2n= 12. 

Tuuonofi ?K3eM[Jrflp L. biebersteinii nueer rycroolyuenuue cre6elr, 

u Ju.{cTbq, Ho qaxle BcTpeqanTcr pacTeHHfl c AoBotrbHo peAKr,rM olryrxeuneu 

(oHo r,rMeerct roJrbKo Ha cre6le), pexe -- [oJrHocrblo roJrbre, c 

eAHHr4r{HbrMrr BoJrocKaMI4 B y3rrax; Ha6lroAarorcr Bce npoMexyrorrHbre 

cogrof,Hr{r. lonr oco6efi c rycrnvr orryueHtreM KJtr{HaJlbHo Hapacrael'K 

rcry. flpurrleqareJrbHo, qro l. biebersteinii o6br.rHo r4Meer roJroe couBe- 

TVe, TorAa KaK AJrfl JrbHrHoK Han6oree xapaKTepHo Huuurqne B corrBel'r{tr 

xe[e3r4crbrx BoJrocKoB H o'rcyrcrBue olryrueHlrt ua cre6:re u Jlncrbrx. 

VsxorncrHrre pacreHm L. biebersteinlf npnuuualucb B oreqec'rBeH- 

Hux

Ouucau usApueuuu, Typus:

subsp. pycnophylla (Boiss. et Bal.) P. H. Davis u subsp. eriocalyx (Boiss.) 

P. H. Davis. HasnanHufi anrop yKa3blBaer, qro qacrb sK3eMIlJrtpoB 

L. kurdica r{Meer ceMeHa c TJIaAKI4M AHcKoM (subsp. aucheri, subsp. 

eriocalyx), oAHaKo Mbr He BlaAeITLT pacreuufi c rraAKI4MH ceMeHaMI4. 

5. L. lineolata Boiss., 1853, Diagn. Pl. Or. Nov. ser, l,12: 42; id., 1879, 

Fl. Or. 4 :379; fpoccr., @n. Kasr, 1932,3 :372;KyrryurH., 1950, Tp. Eor. 

uncr. AH CCCP, cep. l, 9 :44; Kaparuu, 1957, so @.rr, Arep6. 7 :4491, 

Kenr.-Har., 1967, s fpoccr., (Dn. Karx., v3A. 2,7 : 483; MeHr.rux., 1998, 

Eor. xypu. 83,12'.129. *-? L. elymaitica (Boiss.) Kuprian., 1955, so @r. 

CCCP 22 :217. - ? L. lineolata p. elymaitica Boiss., 1879, L c. : 379. 

Ouucau uz Llpaua. Typus: (G), 

Tpananucrrre crJroHbr B cpeAHeM ropHoM rrof,ce. -- Kanxar: Asep- 

6afiAxaH. -- O6ur. pacilp.: 3an, Asuq, 

Bosuoxno, L. lineolata roHcuequt[uqHa c l. kurdica (Sutton, 1988). 

rc. n. MeHurlxnft (1998) yKa3aJr, qro y 3roro BHAa qaureqKa rro KpaK) 

xeJre3l4crztfl, y ocHoBaHHg MeJlKo o[yueHHat, r43Hyrpr.r xene3ucro-BoJrocwcrafl, 

J[rcrbr cBepHyrbl a rpy6xy, HI{reBI,IAHbre. L. lineolata IIpeAcraBileHa 

B [3yqeHHbrx lep6apHux KoJIJIeKuutx HeMHoI'orrHc[eHHbIMn )K3eMrrJrf,paMu. 

Kpoue roro, Mbr He Br{AeJru ceMflH 3Toro su1a. !1. A. Kyupr.rrHoBa 

(1955) orMerr4Jla, uro y L. lineolata ceMeHa Ha ArrcKe 6yropvarue, 

a y L. elymaitica -- rJraAK[e, r4 3Tor Br{A orHocrrrcr K rloAceKuuu Laeves. 

6. L. pyramidalt's (Vent.) F. G. Dietr,, 1818, Nachtr. Vollst. Lexic. 4: 

417:.Chav., Monogr, Antirrh. 1833 : 125; Benth., 1846, in DC., Prodr. 

l0:272; Boiss., 1879, Fl. Or., 4 : 370; fpoccr., 1949, Onpe4. pacr. 

Kanx.: 305; Keu.-Har., 1967, r fpoccr., @n. Kam., vrsr,.2,7 :476; 

P. H, Davis, 1978, Fl. Turk. 6 : 668; fa6prasnsH, 1986, so On. Apvr. 8 : 

314; D. A. Sutton, 1988, Revis. Antirrh. : 297; Ranaual:e, 1988, Eor. 

xypH. 73, l0 : 1478; MeHuqr,, 1998, Eor. xypH. 83, 12 : 130. 

Antirrhinumpyramidatum Vent., 1797,in Lam., Encycl. M6th. Bot.4: 

360. -- L. lenkoranica Kuprian,, 1950, Tp. Eor. uHcr. AH CCCP, cep. 

l, 9 : 67; ead., 1955, no (Dr. CCCP 22 : 194. -- L. kopetdaghensis 

Kuprian., 1950, l. c. 9, I '. 66; ead., 1955, l. c. : 194. - L. pyramidalis 

subsp. lenkoranica (Kuprian.) D. A, Sutton, 1988, L c. : 297; Menuqx., 

1998, urar. co,{. : l3l. -- L. pyramidalis subsp. kopetdaghensrs (Kuprian.) 

D, A. Sutton. 1988. l. c. :299. 

OnucaH r,rs Typrlran. T y p u s : (P, 

Tourn. 967). 

lopHue crerrHbre cKJroHbr, go 2600 M. HaA yp. M. - Kasrar: Apnaeuur, 

Asep6afiAxan. -- O6q, pacnp.: 3an. u CpegH. (Typrrvreur,rcraH) 

Azus,. 

IO, JL MeHnqxraft (1998) paccMarprrBat L. pyramidalis s rrogceKurdll 

Pyramidatae (Vent.) F. G. Dietr. runosofi ceKur4r4, Koropaq BKJtrorraer 

t89

aKxe L. conJbrtiflorc Benth. u L. grffithii Boiss. 3ru elr4u xapaKrepr4- 

3ylorcf, vaurevxoft o rycro orryueunoft AJu,rHHbrMK HexeJre3ucrbrMl4 BorocKaMld. 

Teu ue MeHee, rro Kap[oJrol'uqecKuM rIp[3HaKaM pe3K[rx ol.Ju{- 

.rufi uexgy 3Tr,IMr4 Br,rAaMn I{ ocrilJtbHbrMkl B:ap:avlr' tIoAceKrIHr4 Linaria ue 

ua6nroAaercs. Mtr crreAyeM M. E. Kauanaase (1988) n urnporoii rpaxrosxe 

l. pyramidulis. 

Subsectio 2. Laeves Kuprian. ex Pesskova subsect. nova; Kuprian., 

1950, Tp. Eor. nscr. AH CCCP, cep. l, 9:45, sine descr. latin.; ead., 

1955, eo (Dn. CCCP 22:203, sine descr. latin. 

Corolla lutea vel violacea. Folia filiformia. Semina disci laevia. 

Sculptura secundaria papillosa. 

T y p u s : L. odora (Bieb.) Fisch. 

BeHqux xelruff unu $nolerostrfi. Jlncrbr Hr.rreBr,rAHue. CeueHa sa 

Ar,rcKe fJraAKue; BTopr4t{Han cKyJrbrrTypa [alhJrJto3Haf, . 

T u u : L. odora (Bieb.) Fisch. 

7. L. odora (Bieb.) Fisch., 1812, in Cat. Jard. Gorenk.:25; Chav., 

1833, Monogr. Antirrh. : 136; Benth., 1846, in DC., Prodr. l0:274; 

Boiss., 1879, Fl. Or., 4 : 373; KyupuaH., 1955, so Ol. CCCP 22 :208; 

Keu.-Har., 1967, s fpoccr'., (Dl. Kaor., wsr,.2,7 : 479; Chater et al., 

1972,in Fl, Europ. 3:232, P. H. Davis, 1978, Fl. Turk. 6:670;I[sauusa, 

1981, so (Dn. Enpou. .racru CCCP 5 :234; Karrauaqse, 1988, 6or. 

xypu. 73, l0 :1479; MeHnr1r., 1998, Eor. x(ypH. 83, 12: 128. - 

Antirrhinum odorum Bieb., 1808. Fl. Taur.-Cauc.2:414. ? L. altaica 

Fisch. ex Kuprian., 1936, Tp. 6or. ancr. AH CCCP, cep. l, 2:301. -- 

L. uralensis Kotov, 1946, 6or. xypu. AH yPCP, 3=4 :26, Chater et al., 

1972, l. c. : 233,in obs. - L. dulcis Klok. 1949, Eor. xypu. 34, | : 7l; 

Kyupuau., 1955, unr. coq. : 209; Korot, 1960, so (Dn. VPCP 9:445, 

Chater et al., 1972,l. c. : 232, in obs. -- L. odora subsp. dulcis (Klok.) 

Ivanina, 1981, so (Dr. Enpon. qacru CCCP 5:234. . . L. loeselii auct. 

non Schweigg. : Spreng., 1825, Syst. Yeg.2: 797; Bunge, 1830, in Ledeb., 

Fl. Alt. 2:448. 

Ouucan Hs Poccuu (Bonrorpa4crar o6lacrt). Lectotyp us 

(?Kyrrpraruoea, 1955 : 209; Davis,1978:669): (ex arena mobilis agri 

Sareptani, I 807, [Bieberstein]> (?LE). 

Creuu, rpupycnoBbre rrecKu. -- Bocr. Enpoua: Poccux (Bolx.-[oH., 

Hnxn.-{oH., Huxu.-Bonx. Yrpauua, Ka:axcraH (sau.). Kanrcal: poccnq 

(flpe4ranrarle). - O6ur. pacup.: 3au. Cra6r,rpr. - 2n = 12. 

Hecuorpa Ha 6onturyrc n3MeHqnBocra L. odora, BbrAeJreHr{e r.{3 Hero 

L. dulcis Klok. ra L. uralensis Kotov, rro HarueMy MHeHHro, HeorrpaBAaH- 

Ho, IlocKoJrbKy cyxlecrBeHHblx orJrr4qnft rraex4y yKpar.rHcKr4Mr4 14 loxHoyparbcKr4Mtr 

pacreHraqMr,r He BbuBJreHo. L. odora r4Meer recHoe poAc.rBo 

c L. volgensis u L. loeselii; Bce 3'r[1 BKAbI xapaKrepr43yrorcr cHJrr,Ho pa3- 

BETBJIEHHbIMN NO6CTAUZ, IUHCft UrrU L4 I4JIU HATEBI,IAHbIMII II,ICTb'MII 14 

HeMHororIBerKoBbrMr{ corlBertrflMl4; BeHqr4K 6- l0 rrarra 4I. (6es uuopua); 

190

ceMeHa c ur4poKr4M KpbrJroM u lauernoii 'r'peyloJrbHoii eurelaxofi a o6- 

Jracrn SyHuxylyca; rpaHrrrrbr KJreroK crrepMoAepMu yr.rry6neuHbte, c 

oqeHb'IoHKI,IMI OTJIOXeHKflMH KyTHKyJII,L 

8. L. loeselii Schweigg., 1812, in Konig. Arch. I : 228; KyrrprarH., 

1955, so @n. CCCP 22 : 2091' Chater et al, 1972, in Fl, Europ. 3 : 233; 

I4yauwa, 1981, so On. Enpon. qac'rr.r CCCP 5 : 234. - L. odoro Chav., 

1833, Monogr. Antirrh. : 136. 

OuncaH u3 BocrorrHoft flpyccrau (HuHe KaluuuurpaAcras o6Lr. Poccran). 

Typus: (?B). 

flprarvropcrue ArouHbre rrecKr{. -_- Bocr. Enpona: Poccm (KaluuuurpaAcKat 

o6n.), 3croHlar., JlarBr4fl,, Jlursa. -- O6ur. pacup.: 3HAeMr.rK. 

Apean grofo BI{Aa I.{3oJu4poBaH or ocrtutbHblx, cre[Hbrx, un4oa :roff 

rpy[trbr. VxasaHraa L. loeselii Alr AoHa u Kpluraa (: Spreng., 1825, Syst. Yeg.2:797) u Arraq (Bunge, 1830, in 

Ledeb., Fl. Alt. 2: 448) orHocflrcfl x L. odora. 

9. L. volgensis Rakov et Tzvel., 1993, Hosocrvt cucr. Bbrctu. pacr. 29 '. 

l19; Paros, 1995, no @.rr. lenrp. Pocc. : 88. 

Orucau us Poccr,ru (Vnrrnoncrar o6nacrt). Typ us:

Bonx.), VrpanHa (flpravepH.); Kasnar: Poccur (flpe4raerasrc), Asep- 

6afiaxau. - O6ur. pacrp.: 3at, Cu6. (roxH.), Ka:axcrau. - 2n = 12. 

V rroro sr4Aa Ha6lrcAaerct BHyrpl{lolyrqq[oHHrtfr nolunopQtl:u 

rro oKpacKe BsHqI{Ka (Hannuue HJrl'l orcyrcraue Quoneronrrx xunox). 

BrrAenesue L. incompletc Kuprian., l. c. He lrpegcraBJltercfl HaM npa- 

BrrJrbHbtM, rrocKoJlbKy pacTeHlrt c HeAopa3BlITbIMll qBe'I'KaMU BCTpeqaIOTcr 

rro BceMy apeaJly L. macroura 14 He nMelor reot'paQnuecxoft ttpnypoqeHHocru 

K ero Bocrotruofr qacrl4. 9efirep c coaB'l'opaMn (Chater et al., 

I972) nucra3anv npeAlloJtoxeHI4e o ru6pt'tgHotvt llpol4cxoxAeHnu 

L. incompleta (L. macroura x L. vulgaris). L. debilis Kuprian., l. c. xapaKTeph3yercq 

c6nuxeHubrMtl Jltrcrbfltuu u cre6neM, BerBtuII4Mct o'r oc- 

HoBaHI{f,. llo uaureuy MHeHI4Io, 3To pacreHue cKopee 3Korol'nqecKar 

$oprvra L. macroura, qeM caMoctoqremHltft sIaA. Tfiuosoft o6paseu, 

L. baxanensls Galushko, L c. (

HaAse, 1988, Eor. xypH. 73, l0 :1479; Menuqr., 1998, Eor. xypn. 83, 

12:129. - L. rupestns C. A. Mey., 1831, Verzeichn. Pfl. Cauc. Casp. : 

I l0; non Guss., 1828; Benth. , 1846, in DC., Prodr. l0 :282. 

OurEcancKaerara ([apurrcroeyqerle). Ty p u s:

KpyrrHbrMr{ crpyKTypr{poBaHHbrMH rpaHyJraMn, pacrroJloxeHHbrM14 Ha 

cocoqxax KJreroK rro cropoHarra rpe6uefi n n 6opor4ax Me)I(Ay HrrMI4. 

?ugocuepn rnaAruaft. 

13. L. genistifulia (L.) Mill., 1768, Gard. Dict. ed. 8 : no 14; Chav., 

1833, Monogr. Antirrh. : 127; Benth., 1846, in DC., Prodr. l0 : 275; Kynpr,rrn., 

1955, so cDr. CCCP 22:188; Kerra.-Har., 1967, s fpoccr., @n. 

Kanr., uzg. 2, 7 : 480; Chater et al., 1972, in Fl. Europ. 3 : 230, p. p.; 

P. H. Davis, 1978, Fl. Turk.6:659;Illyavuna, 1981, so (Dr. Eepon. vacru 

CCCP 5:232; D. A. Sutton, 1988, Revis. Antirrh. :313; Karraua,qse, 

1988, For. xypu. 73, l0 : 1476, p. p.; Menrzqr. , 1999, For. xypn. 84, 

| : 126, p. p.; 3epuon, 2000, Pacr. Ces.-3ar. 3axaer. : 100. 

Antirrhinum genisti.folium L., 1753, Sp. Pl. : 616. L. svspirensis 

K. Koch, 1849, Linnaea22:717; KyupurH., 1955, urlr. coq.: 190; l{ua- 

HnHa, 1981, unr. corr. :232;KatauaA:|e, 1988, rrr4'r. coq. :1477; MeHuqr., 

1999, qrr. co\.:127. L. petraea Stev., 1857, Bull. Soc. Nat. Moscou 

30:437. L. euxina Velen., 1866, Abh. Boehm. Ges. Wiss. 7, | :34; 

Kynpuau., 1955, urar. co.r. : 193. - L. ponticc Kuprian., 1950, Tp. Eor. 

nscr. AH CCCP, cep. 1,9:64; ead., 1955, l. c.: 189; Kerra.-Har,, 1967, 

rlr,rr. coq. : 482:' }Isaillsa, 1981, \ur. co,t. : 232. - L. scenoreina Juz., 

1951, Eor, uar. (JleHuHrpaa) 14:30. -- L. adzharica Kem.-Nath., 1963. 

3au. cucr. reorp. pacr. (T6unucu) 23: 40; ead., 1967, s fpoccr., tDn. 

KaeK., trzg.2,7 :481. - L. iberica Kem.-Nath.,1963, l. c. : 38; ead., 

1967,1. c. : 481. - L. imeretica Kem.-Nath., 1963, L c. :37; ead.,1967, 

L c.:481. L. caucasigenc Kem.-Nath., 1963.1. c.:39; ead.,1967,l.c.: 

482. - L. kontschaveli Kem.-Nath., 1963, L c.,23: 38; ead., 1967, l. c. : 

482. - L. genistiJbha subsp. euxina (Velen.) D. A. Sutton, 1988, Revis. 

Antirrh. : 317. 

OuucaH usPoccrau. Lectotypus (Davis, 1978): Herb. Linn. 

N 767140 (LINN). 

Creuu, [ecqaHble, KaMeHI,Icl'bIe I4 MeJroBbre IIoqBbr, B HK)|tHeM llotce 

rop. - Bocr. Enpona: Poccr,rs (Bonx.-,{oH.; 3arorx.; HuxH.-Bo.nx.), 

Eelapycr, Vxpauua, Mongoaa, KasaxcraH (sau.). Kanrar: Poccl,rs 

(flpe4ranxasre), fpy:r,rx, ApueHux, Asep6afigxaH. - - O6u. pacup.: 

CrasA. (ror), I{eurp. u [OxH. Enporra, 3au. Cn6nprn 3au. ra Cpegn. 

Arur. - 2n= 12. 

Brrnr,qrr ua o6neu uolr,uuopQHoro BHAa L. genistilolia 3Haqr.r'reJrbHo 

pa3Ju.rqarorcr. 9eftrep c coaBTopaMu (Chater et al., 1972) paccMa rpuBator 

L. genistifolia n trrupoKoM cMbrcJre r,r BKJrKlqaK)T n nee L. dalmatica 

(L.) Mill. u L. soJiana Velen. .{rnr.rc (Davis, 1978) upr.rsnaer caMocrof,- 

TelbHocrb Ha3BaHHbrx BrrAoB r{ BbrAeJrser uz L. genistifuli.a 6 cta6o orrpaHnqeHHbrx 

rroABuAoB, [per{My[IecrBeHHo nr Typuun, rpe6yrcu1r.rx 

AzurrHefiurero u3yrreHur. Carros (Sutton, 1988) rarxe pa3AeJrrer L. genistiJblia 

Ha [oABrrAbr. B ore.recrseHHofi nnTeparype [pr{Hrro B ocHoBHoM 

y3Koe rroHr4Mauue L. genistiJblia (Kynplrauoaa, 1950, 1955; Koron, 196l; 

t94

Keuylrpux-Haragse, 1967;l4salutLtta, l98l). B urupororu cMblcJre 3ror 

BrlA rrpr{Hflr M. E. KananaAre (1988) r.r oco6eHuo IO. JL Meuuqrcurvr 

(1999), qbfl ToqKa 3peHuf uavr uau6onee 6nusxa. Mu c.IuraeM Bo3Mox- 

HbrM BKJrro{r{rb B crrHoHuvu L. genistiJblia lr lpl{3HaBaervrHfi 3runau asropaMr4 

L. syspirensis. 

L. genistiJblia xapaKreprcyercs 6omruofr HsraeHqnsocrrro uopSo:tofr{qecKr4x 

rrprr3HaKoB Be|eraTI{BHbIx opfaHOB (pa3Mepbl I{ pa3BerBJIeH- 

Hocrb pacreHufl, pa3Mepu n $opua JrncrbeB, crefleHb pa3Burl,Ifl BocKo- 

Bol'o Huera). feorpar[raqecrofr [puypoqeHHocrr4 pacreHlrft c yra:laHun- 

Mn rlpl43HaKawr4 Hauvr He 6nuo BbrrBJreHo. 

Carrou (Sutton, 1988) orrraeuaer raKxe nonuuop$r,rsu ruop$oloruqecKux 

rrpH3HaKoB ceMflH, q'ro coB[aAaer I{ c HaltruMn AaHHbrMn. CpeAn 

r4ccireAoBaHHux pacreHnil L. genistiJblia 3 o6pa:qa 3acJryxuBarcr oco6oro 

BH?rMaHlrx. Vnrrpacryilbrrrypa ux ceMf,H H aHaroMnqecKoe crpoeHne 

ceuesuofi Koxypbr oAr{HaKoBa 14 coorBercrByer Apyrr4M ceMeHaM pacrenufi 

AauHolo BrrAa. V cervrxu A,ayx o6pasuon ( - LE!;

Ouzcan us Apueuuu. Typus:

16. L. sabulosa Czern. ex Klok., 1949, Eor. xypH. 34, | : 69; Kynpurn., 

1950, Tp. 6or. uHcr. AH CCCP, cep. 1, 9:42; oHa xe, 1955, eo (Dr. 

CCCP 22: 190: Koros. 1960, no

ocrrlrrbHbrx BrrAoB ceKrrr4u Speciosae: rroflylyHHaq, a He rer-parApl{qecl(afl 

$opnaa ceMeHr4 n 6o;ree recHo pac[oJroxeHHble ocrpble Kyrl4KyrspHble 

6yropru, o6pa:lyrcqne rhflr,rrrHylo AJrfl J]bHrHoK cerqarylo BTopI,It{HyIo 

cKynbrrrypy. flounuo 3Toro, L. cretacea u L. macrophylla, s orrluqne or 

Apyrux BrrAoB c 6ecrpunuMr4 ceMeHaMn (nrltouar v Bvl\br ceKlrnu 

Speciosae), xapaKTepu3yrorcq Mo3r'oBrrAHofi uepnn,ruofi cryllrurypofi 

cevreuuoil Koxypbr, orcyrcrBr,reM cocoqKoBuAubrx KJreroK r4 orrlrKyrrrKy- 

JrflpHoro BocKa. HasuaHHne BuAbr r4MeK)T, KpoMe roro, BeHr{r4K c roJrcrbrM 

H3orHyrbrM ru[oprreM H rBJrflrorcr o6rrlrarHbrMu KaJrbueQurauu. 

Bce aru oco6euuocru [o3BolqK)r BblAeJrr{rb L. cretacea w L. macrophylla 

B orAenbHylo ceKurllo Cretaceae, 

lJ. L. cretacea Fisch. ex Spreng., 1825, Syst. Veg.2:791;Chav., 1833, 

Monogr. Antirrh. : 122; Benth.,1846, in DC., Prodr. l0:285; Kyupnru., 

1955, no On. CCCP 22 : 221; Chater et al., 1972, in Fl. Europ. 3 : 

229; I4sauuua, 1981, no (Dr. Enpon. qacru CCCP 5 : 236; D. A. Sutton, 

1988, Revis, Antirrh. :325. - L. menisperma Klok., 1949, 6or. xypH. 

34, | :70. - L. creticolc Kuprian., 1950, Tp. Eor. uucr. AH CCCP, cep. 

1,9 :63; ead., 1955, so @r. CCCP 22:221; D. A. Sutton, 1988, Revis. 

Antirrh. : 325. L. cretacea var. creticola (Kuprian.) Kotov, 1960, no 

Or. YPCP 9 : 447: Vlsanuua, 1981, qnr. co-r. : 236. 

OnucanHsPoccun. Typ us: (LE?, photo LE!). 

MenosHe u MepreJlr{crlte o6naxeHnq, ocburn MeJroBbrx cKnoHoB u 

KapbepoB. Bocr. Enpouar Poccus (Bo.ux.-!oH.; Huxu.-{oH.: ceB.nocr.; 

3anolx.; HuxH.-Bonx.: cea.), Vxpauua, Kasaxcrau (:an.). 

O6n1. pacup.: 3au, Cu6wpr (Bepx.-To6.). 

YrasaHr.rs 4lu Caparogcroil o6nacru (I{nauuna, 1981 : 236) He uo4rBepxAeHbr 

rep6apuuruu Marepr{araMu (Pagurnu a, 2002). 

M. B. KnoroB onr4caJr c Bonrn L. menispermc Klok., l. c., roropar, 

rro ero MHeHuIo, orluqaercr or L. cretacea HecroJrbKo 6onee KpyuHbr- 

Mlr rroJryJryHHbrMrr ceMeHarvru. JI. A. KyupuauoBa cBeJra 3To Ha3BaHrre B 

crrHoHr,rMbr L. cretacea 14 orrlrcaJra c BepxHero lona L. creticola Kuprian., 

l. c., Koropar xapaKTepr{3yercq, rraBHbrrrl o6parou, 6onee vrenrr.r- 

MI{ n JraHIIerHbrMn, a He urltpoKoqftqesuAulrN,rv, JrvLcrbflMyt 3apy6exHue 

aBTopbr nu6o upu:uaBarrr4 caMocroflTerbHocrb 3Toro Br{Aa (Sutton, 

1988), rn6o oro)KAecrBJrrrr4 ero c L. cretacea (Chater et al., 1972). [lo 

Harlr4M AaHHbrM, BbrAereHne L. creticola HeorrpaBAaHHo, TaK KaK pa3Mep 

u Qoprr,ra Jrncra noABepx(eHbl 3Haqr.rrgJrbHofi ugNaenqusocrrl Aaxe Ha oA- 

HoM pacreHuu. Carvtre KpyrrHbre r{ rxlrpoKr4e Jlr4crbfl paclonoxeHbr B Bepx- 

Hefi qacru cte6la, rroA corrBerr4eM. 

JI. A. Kyupr4sHoBa (1950) orncaJra raKxe poAcrBeHHbrft sraA ur Cerepo-3anagHoro 

KaraxcraHa - L. macrophylla Kuprian, xoropnfi 

orrr4qaercr or L. cretacec 6onrurnulr pa3MepaMr,r pacreHr.{q, 6olee rpyu- 

Hnvru (Ao 20 rr,rrta gn.) rncruur{, MeHbrrrtrMr.{ pa3MepaM[ BeHqr4Ka, y3KtrM 

KOJrbrIeBbrM KpHnoM ceMeHr.r. 

198

B. H. Paanru:na (2002) orMerraer, qro rto crpoeHl4lo ceMtH L' cre' 

tacea LoBoillHo 6nusra K cpeAl{3eMuouopcrofi L' flava (Poir.) Desf. 

oAsaxo uocleAnufi BUA - OAHOJIeTHT.TK n rrMeeT Apyfylo 3KoJrofI4IO 

(npnuopcrr,re flecKl,t no6epexnft Kopcnru u Caphnruuu). Ceuxu aroro 

BUAA MbI HC BI,IACJIH. 

Sectio 4. Diffusae (Benth.) Wettst', 1891, in Engler u. Prantl, Nat' 

Pflanzenfam. 4,3b: 59; Kyupr'raH., 1955, so @r. CCCP 22 :220; l4sa- 

Hlrsa, 1981, no (Dl. Enporr.'racrll CCCP 5:236; D. A. Sutton' 1988' 

Revis. Antirrh. : 335; Menuur., 1999, Eor. xypu. 84, I : 130. Linaria 

$. DiJJusae Benth., 1846, in DC., Prodr. l0:284. 

L e c t o t y p u s (Viano, 1978b : 223): L. reilexa (L.) Chaz' 

OAHOnerunrri u MHOIOJIeTHnKH, C IIpII[OAHIIMaIOqIIMIaCfl Cre6lsnlt. 

Jlucrlx MyroBqarble, gftqeslaAHo-JlaHuerHble, cyxeHHble B oqeHb Koporxufi 

.reperuOn, I-lnerxu B pbIXJIbIX KHCTTX llJItI paCIIOJrOXeHbl llo OAHo- 

My B rra3yxax Ju.IcrbeB. t{aureqra TpaBflHncraq. BeH'rIar xenrnfi, t'olry- 

6oit utm 6enuft. CeueHa 6ecrpulue, 0.9-l'5 MM AJl.' flepBllqHafl cKyJIbflrypa 

rpe6ueeu4nar. 

Carron (Sutton, 1988) o6rcAnHl{n ceKurltt DifJusae u Minutdlorae, n 

eMy rrocleAosar [O. JL MeHuqxuft (1999). 

18. L. reflexa (L.) Chaz., 1790, Suppl. Dict. Jard' 2:40; Chav', 1833, 

Monogr. Antirrh. : l19; Benth.,1846, in DC', Prodr. l0 :284, Boiss., 

1879, Fl. Or. 4 : 380, KyupraxH., 1955, ao Or. CCCP 22:220; Ker'l.-Har., 

1967, s fpoccr., @n. Kasrc., uzg.2, 1 :483; Chater et al., in Fl. Europ. 

1972, 3 : 228, D. A. Sutton, 1988, Revis. Antirrh' : 336; KanaHaA3e' 

1988, Eor, xypH. 73, l0 : 1481; Menuqx., 1999, Eor' xypu. 84, 1 : 130. - 

Antirrhinum reJlexum L., 1762, Sp. Pl. : 857. 

Ouucan us Arxupa. L e ct o t yp u s (Wijnands, 1983 : 189): 

Herb. Linn. N 767154

19. L. albifrons (Sibth. et Smith) Steud., 182l-1824, Nomencl. Bot. ; 

482; Chav.,l833, Monogr. Antirrh. : 156; Benth.,1846, in DC., Prodr. 

l0 : 280; Bois.s., 1879, Fl. Or.4 :382; KynpraaH., 1955, so Ol. CCCP 22: 

225; Kerr,r.-Har., 1967, n fpoccr., On. Kanr., vrzg. 2, 7 : 484; Kanauagse, 

1988, Eor. xypu. 73, l0: l48l; MeHuqx., 1999, 6or. xypu. 84, I : 

130. Antirrhinum albiJrons Sibth. et Smith, 1826, Ic. Fl. Gr. : tab. 

588. - Linaria minutiflora C. A. Mey., 1831, Verzeichn. Pfl. Cauc. 

Casp. : 109; Chav., 1833, l. c. : 155; Benth., 1846, in DC., Prodr. l0 : 280. 

Onracau c o-Ba PoAoc e 3refrcrou uope. 

Kasras; Asep6afiDKaH (Bocr. 3arcasragle: lllupraHcxnfi Qnopucruuecxufi 

pafiou). - 06111. pacrp.: IOxu. Enpoua, 3an. A:us, Ces. At[puxa. 

Buq npune4eH AJtfl Kasrara Ha ocHoBaHnr,r Jrr,rreparypHbrx AaHHbrx. 

Sectio 5. Versicolores (Benth.) Wettst., 1891, in Engler u. Prantl, Nat. 

Pflanzenfam. 4,3b: 59; Kyrpraau., 1955, no @r. CCCP 22:217; Koroe, 

1960, so

My MHeHr4IO, LBTAeTCfl KOHBepfeHTHbrM, B aHarOM[rreCKOM CTpOeHr{u 14 

ynbTpacKyJrb[Type ceMrH BbrflBJrrK)Tcr cyulecTBeHHbre oTJrHrrHr. 

Eesravr (Bentham, 1846) nulenufl B cocraBe rpyrrrrbr Versicolores 6o- 

Jree MeJIKI{e noApiBAeJreHrrf,, Koropbre 6mnn nonuureHbr Ao paHra ceKu}rr4 

Buauo (Viano, 1978a,1978b). Carron (Sutton 1988) paccuarpr,rBaer HX 

B paHre fioAceKuun. Hauru KaprroJrorr{qecKr4e l4ccJreAoBaHut, BbrqBuBttltre 

3HaqurerbHbre pa3Jruql4q Mex4y H3yrreHHbrMr4 BuAaMlr ceKrll{tr Ver,sicolores 

s. 1., noqrnepAuJru fereporeHHoc'rb. flpe4loxeHHoe Ha3BaHHbrMH 

aBTopaMu AeneHHe ceKrlnr.r Versicolores lpeAcraBJrflercq o6ocHoraHHbrM, 

HO, TeM He MeHee, Mbl He peuaeMcfl efo [pr4HrTb, [ocKoJrbKy HaMr{ r13yrleHa 

roirbKo Marat qacrb BI,IAoB ceKql{r4 Versicolores s. 1., roropue rrpor,r3pacrarcT 

n BocroqHoft Enpone u Ha Kasxase, a L. canadensis (cu. 

nuxe) He paccMarpr4Baercfl r pa6orax Bnauo (Viano, 1978a, 1978b). 

20. L. canadensr's (L.) Dum.-Courset., 1802, Bot. Cult, 2:96;' Chav., 

1833, Monogr. Antirrh. : 149; Benth., 1846, in DC., Prodr. l0 : 278; KyupvrnH., 

1955, so @r. CCCP 22 : 218;, Chater et al., 1972, in Fl. Europ. 

3 : 23I; I4nayura, 1981, so (Dn. Erpon. qacrlr CCCP 5 : 235. 

Antirrhinum canadense L., 1753, Sp. Pl. : 618. - Nuttallanthus canadensis 

(L.) D.A. Sutton, 1988, Revis. Antirrh. : 457. 

OuucaH us Cen. Auepuxu. Lect o typ u s (Pennell, l9l9 : 151 

152): Herb. Linn. N 767169

Ouucas u3 ceB.-Bocr. Typurau (6unru' Typeuraa ApueHraa). 

Lectotypus (Viano, 1978b):

et DC., Fl. Fr., ed. 3, 3 : 586; Benth., 1846, in DC., Prodr. l0 :278. 

L. repens B. monspessulana (L.) Dumort., 1827, Fl. Belg. : 34. 

Onucau us 3an. Enpouu. Lectotypus (Davis, 1979:664): 

Herb. Linn. N 767114 (repens) Alstroemer 123 (LINN). 

Copuue Mecra, BAoJIb Aopot. -- Bocr. Enpona: ?crouur (pe.qxo), 

Jlarnur, Jlursa, VxpanHa (Aoneqr: EoraHu.Iecrvrit cag, sauocuoe). - 

O6u1. pacnp.: Arrr., I{eH'rp. u [Oxn. Erpoua. 

CeueHa c rpe6HenuqHofi nonepxHocrblo, sHAoclepM llaArufi. 

Sectio 6. Arvenses (Benth.) Wettst., 1891, in Engler u. Prantl, Nat. 

Pflanzenfam.4, 3b : 59; KynpmH,, 1950, Tp. Eor. uncr. AH CCCP, cep. 

1, 9 : 53; oHa xe, 1955, so @n. CCCP 22:223;l4nauuna,l98l, so (DrI. 

Enporr. qacru CCCP 5 :236.'- Linaria sect. Linariastrum Chav. $ 4. 

Arvenses Benth., 1846, in DC., Prodr. l0 :279. -- .L. sect. Linariastrum 

Chav., 1833, Monogr. Antirrh. : ll4, p. p. 

T yp u s: L. arvensis (L.) Desf. 

OAuonerHnru c [pqMocrotrll{Ml4 cre6;tqMn 10-20 cu BbIc. 14 tntneit- 

HbrMrr JlucrbflMH, pacrroJloxeHHblMl,I no 2-3. Couserue fycroe, xeJle3[tcro 

ofiy[reHHoe, ttpl{ rlroAax cI4JlbHo yAJll{Hf,Ioueecr. t{aueqra IUIeHqarax. 

BeH.rar uelruft, 6enrrft, roly6ofi, xelrrtft. Kopo6ouxa 5-6 runa 

B Ar{aM., rxapoBl{AHafl. 

Ceueua 2 2,5 uu B Auav,,, AI{cKoBI4AHbre, c IIIHpoKI,IM repe[oHr{arblM 

KorbueBbrM KpbIJroM, rlpoxoAturl4M uepes QyHraxynyc. Tero ceMeHu oKpyrro-[orrKoBuAHoe. 

Knerrn SK3orecrbr KpbrJra paAI,IaJrbHo Bbl'tfl H yrble, 

pa3Mep ux yBefl[rruBaercfl K nepuQepuu, orAeflbHble KJIerKu o6paryror 

cocoqKu, HarrpaBJreHHne r nepn$epuu. KlerxIE oK3orecrbl TeJIa ceMeHlr 

6orree unn MeHee u3oAnaMerpuqecKne. flepnuuuafl cKyJrbrrrypa 6yrop- 

Yarat. 3HAocnepu uaAxnfr. 

IO. JI. MeHuqrufi (1999) yKa3aJr, .rro EeHrara (Bentham, 1846) pas- 

AeJruJr rpylnrt Arvenses Benth. u Supinae Benth., BKflIorrarculue oAHoJrel'- 

Hrre BuAbr c ArrcKoBl4AHbIMr,r ceMeHaMH, rro xapaKTepy pocra (cre6lr.r 

npflMocrorqne y neprofi rpy[rrbr 14 rlpr4rroAHrlMalourr4ecfl ''- y nropofr) 

r{ pa3MepaM rrBerKoB (y nepnoft rpytrrlbl 4 -9 rrau AJI., y Bropoii - 6olee 

l0 uu). ?rz pasluur{r B AarbHeftureu 6rrnu upr{Hrrbr KaK ceKrrr4oHHbre 

MHorrrMr4 rrccJreAoBarenruu (Wettstein, 1891; KynpurHora, 1950, 1955; 

Vald6s, 1970, \nr. no Sutton, 1988; I,Inanrana, l98l; KananaA:e, 1988). 

OAHaxo rlo 3TrrM rrpH3HaKaM Ha3BaHHbre rpy[rlbr, [o3AHee Bo3BeAeHHbre 

B paHr ceKrlr4r{, [paKTr,rqecKn Hepa3JrurruNarr. BsnAy f,Toro Carron 

(Sutton 1988) o6re.qartntr vrx rroA Ha3BaHr4eu Supinae. Mr,r He peuaeMct 

3ToI'o cAeJraTb, trocKoJrbKy He u3yrrarr4 KaprrololuqecKr.r Br.tAr,t ceKrlr4r,r 

Supinae (Benth.) Wettst. s. str. 

fleprouauanbHo B ceKur{to Arvenses Bxrroqaracb L. pelisseriana (L.) 

Mill., anocleAcrBrrr BbrAeJreHHar BalrAecolr (Vald6s, 1970, qur. no 

Sutton, 1988) n ceKrlr4ro Pelisserisnae Vald6s tro KaproJrorr4qecxuM rrpu- 

3HAKAM. 

203

24. L. simplex (Willd.) DC., 1815, in Lam. et DC., Fl. Fr., ed. 3 : 588; 

Benth., 1846, in DC., Prodr. 10 : 280; Kyupnru., 1955, so Qn CCCP 22: 

223;Koros,1960, so (Dr. VPCP 9 :450; Keu.-Har., 1967, s fpoccr., (Dl. 

Kasr., usq,. 2, 7 : 479; Chater et al., 1972, in Fl. Europ. 3 : 236; 

P. H. Davis, 1978, Fl. Turk., 6:672;I4varuuna,l98l, so (Dr. Erpon. .racrra 

CCCP 5:236; Sutton, 1988, Revis. Antirrh.:405; Kanauaa:e, 1988, 

Eor. xypn. 73, l0:1481; MeHuur., 1999, Eor. xypH. 84, I : 129. 

Antirrhinum simplex Willd., 1800, Sp. Pl. 3, I : 243. - L. arvensis var. 

simplex (Willd.) Chav. 1833, Monogr. Antirrh. : I57. * L. arvensis subsp. 

simplex (Willd.) Lange, 1870, in Willk. et Lange, Prodr. Fl. Hisp. 2 : 570. 

Onucas ur 3au. Erponu. Lectotypus (Valdds 1970:91): 

Typ us: L. pelisseriana (L.) Mill. 

BanrAec (Valdes, 1970, urar. no Sutton, 1988) suaeruJt 3ro'r BIIA B 

or4eJlbHyro ceKrrnlo Ha ocHoBaHulr KaprloJlof LIqecKI4x npu3HaKo B (ce nae- 

Ha c rpeyroJtbHbrM Ar{cKoM, KpbIJro 6axpouuaroe, [poxoAI{T B rlJlocKocrr4, 

rrepneH4nrylrpuoft py6urary). Branlrou u JIaHre (Willkomm, 

Lange, 1870) errlenlru L. pelisseriana B orAeJrbHylo rpyn[y ro HaJrI{uurc 

6axpoMqaroro Kpbrna. flocne4yrcrque aBTopbr paccMarpuBaJlu 

L. pelisseriana B cocraBe ceKuul{ Arvenses. 

26. L. pelisseriana (L.) Mill., 1768, Gard. Dict. ed. 8, no ll; Chav., 

1833, Monogr. Antirrh. : 156; Benth.,1846, in DC., Prodr. l0 : 279; KyrrpkrflH.,1955, 

so @r. CCCP 22 :224; Kena.-Har., 1967, s fpoccr., tDr. 

I(anr., uzr. 2, 7 : 479; Chater et al., 1972, in Fl. Europ. 3 : 232: 

P. H. Davis, 1978, Fl. Turk., 6: 672; D. A. Sutton, 1988, Revis. Antirrh. : 

422;Menuux., 1999, Eor. xypn. 84, I : 130. - Antirrhinumpelisserianum 

L.,1762, Sp. Pl. 2 : 885. 

Orncan us (DpaHrlun. L e c t o t y p u s (Davis, 1978): Herb. Linn. 

N 767122 (LINN). 

Kasras (? I{enrp. 3arcanrare). -- O6ur. pacrp.: I-leurp. u IOxn. 

Enpona, 3au. Asur. -- 2n = 24. 

Vrasusaercq gnr I{eHrp. 3axasrasrs (Kynpmnona, IIHr. co.r.; Keuylapux-HaraA3e, 

qrar. co.r.). rc. n. Mesnurufi (1999) orMerl{Jr, qro 

3Tu yKa3aHufl coMHr{TeJrbuu. L. pelisseriana [peAcraBneHa B r{3yqeHHblx 

uauu rep6apHbrx KoJrJreKrlr4flx HeMHororrucJIeHHbIMla QparueHrapuurnalt 

c6opaulr XIX s. PacreHns He r{Meror rrJroAoB, H rorrHo ycraHoBl4'r'b Ltx 

Br,rAoByro rrpHHaAJrexHocTb He [peAcraBJrtrercf, Bo3MoxHbrM. 

Sectio 7. Macrocentrum D. A. Sutton, 1980, Journ. Linn. Soc. London 

(Bot.) 8l : 171; id., 1988, Revis. Antirrh. :452. -- Linaria sect. Dillusae 

subsect. Macrocentrum (D. A. Sutton) Menits., 1999, Eor. xypn. 84, I : 

131. --.L. sect. Versicolores (Benth.) Wettst., 1891, in Engler u. Prantl, 

Nat. Pflanzenfam. 4, 3b : 59, p.p.; Kyupr,rau., 1955, so (Dn. CCCP 22: 

217;VInauuua, 1981, no (Dl. Eapou. .racrr,r CCCP 5 :235. L. sect. Linariastrum 

Chav. $ 3 Versicolores Benth. 1846, in DC., Prodr. l0 :275, p. p. 

Typ us: L. armeniacaChav. 

OAuoneruue loJrbre pacreHur l0-55 crra Bbrc., c nuueftuuuu vrrrr4 HLr- 

TeBLrAHbrMrr oqepeAHbrMI,I Jlr{crbtMrr. 9aurenucruru y:ronraueftHbre, cpocuruect 

roJIbKo B ocHoBaHrru. Beuqux 4-8 errvr AJr.; 3eB orrpmruil, runoperl 

roHrufi, 3HaqrrreJrbHo AJnrHHee BeHrrr,rKa (.qo l5 vru), culrHo usor-- 

Hyrnft. 

Ceuena 6ecxprrlrre, fipl{3MaruqecKr{e, rroqKoBHAHo-Terpa]Apr,rqecrne, 

c 5-8 KpyrrHbrMn ro[epeqHbrMr,r BolaHaMr{. Py6ur,rx cy66a:alrHurft, 

Ha AopcaJrbuou rpe6He. BropuuHafl cKynbrrrypa ocrpo6yropqaraq. 

Knerru 3K3orecrbr l-2-cocoqxonue (c ne6onluoft ueHrpaJrbHoft u enlpaxeHuofr 

MaprrlHiurbHofi uanu.nnarvru). 3ur,myruxynpHuft BocK rrpeA- 

205

cTaBJreH AOBOJTbHO KpyrrHbrMu, He CTpyKrypHpoBaHHbtMH, fpaHyJraMtr. 

SH4ocuepu pyMHHr,rpoBanuuft . 

Hamn Kap[olor-I4rlecKue r4ccJreAoBanl{r cBuAereJrbc'rBylor B troJlb3y 

o6oco6neHsocrr.r Br.rAoB ceKr1utl M ac rocent rum. Py vruuuposaHHtril rH- 

AocrrepM Bcrper{aercr raKxe y L. canadensis u L. incarnata., uo, no nceii 

BnAr,rMocrr4, uo 3ToMy rrpl43HaKy rrMee'l' Mecro KoHBepreHuufl. Harsau- 

Hbre B?rAbr olJlr{qatorcfl or BrlAoB ceKuul,r Macrocentrutn oco6euHocrrMr,r 

ana'rourr4 r.r yxbrpacKyrrbrrrypbt ceueHuofi Koxypbr. A;u slrAos cexur4r4 

M ac r oc entrutn xapaKTepHbr orKpbrrbrft ses seH.rraKa, AJrr.rHHbrfi nsoruyrlrfr 

uuopeq H [pr4AarKH Ha rbrqnHoqHbrx Htll'f,x. 

27. L. armeninca Chav., 1833, Monogr. Antirrh, : 147; Benth., 1846, 

inDC.,Prodr. l0:277; Boiss., 1879,FI. Or.4:381; Kynpnxu, l955,ro 

Ka n a H aA3e M.6. Kpu:ruxo-cuureMarl{qecKoelr3yqeHueKaBKa3cKux npe.qcrasrErenefi 

poga Linaria: Anroper!. [nc. ... raHA. 6uon. nayr. T6nnuclr, 

1989. 17 c. 

Ke u y ilapu a - H araA3e Il.M. LinariaL. ll A. A. fpoccreftr'l.

Segarra J. G., Mateu l. Seed morphology of Linaria species from 

Eastem spain: identification and taxonomic implications ll Bot. l. Linn. Soc. 

2001. Vol. 135, N 4. P. 375-389. 

S u t t o n D. A. A revision of the tribe Antirrhineae. London. oxford. l9gg. 

575 p. 

v i a n o J. Les linaires a graines apteres du bassin mediterraneen occidental. 

l. Linaria sect. Versicolores ll Candollea.1978a. Vol. 33, N l. p. 43_gg. 

v i a n o J. Les linaires a graines apteres du bassin mediterraneen occidental. 

2_, Linaria sect. Elegantes, Bipunctatae, Diffusae, Speciosae, Repentes ll 

Candollea.1978a. Vol.33, N 2. p.209-267. 

w e t t s t e i n R. Antirrhinoideae - Antirrhineael/ Engler A., prantl K. Die 

nattirlichen Pflanzenfamilien. Leipzig, lg9l. Teil 4, Abteil. 3b. S. 56_62. 

Willkomm H. M., Lange J. M. C. prodromus florae hispanicae. 

Stuttgartiae, 1870. Vol. 2. 680 p. 

Mocroscrufi reAaroruqecruft r-ocygapcrseHHHft yHr{Bepcrlrer, 

Euonoro-xuuuqecruft Qaryllrer, raSe4pa 6orauuru 

129164, Mocxna, yn. Ku6alr.rnva, 6, xopn. 5

fl. A. Kocaq6n, 

[. A. fepuan 

P. Kossaczev, 

D. German 

HOBbIII BI,I,( POAA VERONICA L. (SCROPHULARIACEAE) 

lI3 3ATTAAHOIZ MOHI OJIIIII 

GENERIS VERONICA L. (SCROPHULARIACEAE) 

SPECIES NOVA E MONGOLIA OCCIDENTALI 

flpu orcue4uIIHoHHoM o6cne.[osaH[14 AxyHrapcKoro MaKpocKroHa 

Mouronlcroro Alras JIeroM 2003 r. oAHItM H3 aBTopoB coo6uleHtu 

6rrnlr co6panu o6pasubl BepoHI,IKH I'13 ceKIII4H Pseudolysirttachium 

W. D. J. Koch (n rapy6exHoft Jlureparype oHa o6blqHo paccMarpl'IBaercr 

B paHre poAa), uoptfonorn.recKl,I pe3Ko orJruqatoullect or [poqux 

BLTAoB ceKuun, ceofrcrseHnnx AaHHoMy peruoHy AsuH. Huxe [pHBoAt4Tcr 

orrr{caHl,Ie 3Toro pacTeHut. 

I,I: ugsecrHrrx K HacrotuleMy BpeMeHu n Alraftcxofi ropuoft crpaHe 

14 sn.[os H 3 noAeuAoB Ha3BaHHofi ceKuuu 4 su4a u I noAnuA tBJlflIorcfl 

3HAeMrrKaMr4 aroft reppraroplr,:,z,, a pacflpocrpaHeHl,te eure AByx BI4AoB 14 

oAHoro rroABr4Aa eABa BbIxoAl{T 3a ee flpeAeJtbt. flpu arou Bce floKiulb- 

Ho paclpocrpaHeHHble raKcoHbl, B ToM rtl,tcJre ottl{cbtBaeMbtfi, 3a ncrnroqeHneM 

eALrHcrBeHHoro TIoABHAa (V. spicata subsp. kamelinii), l{Melor 

ru6pu4oreHHoe llpoucxoxAeHue. OuucaHne HoBoro BuAa etqe 6onee 

uoAqepKrlBaer orMeqeHHyn paHee (Koca.rdr, 2002,2003) polt Alraficrofi 

ropHoft crpaHbl KaK MoulHoro oqara ru6pu4oreuHoro nugoo6pa- 

3oBaHHfl B ceKIII{u Pseudolysimachium W. D. J. Koch. 

Veronica (sect. Pseudolysimachium) smirnovii Kossaczev et D. German 

sp. nova. - Herbae perennes, l5-32 cm alt., pilis brevibus patentibus 

glandulosis dense pubescentes. Rhizomata brevia, adscendentia; caules 

tenui, simplices, solitarii vel pauci, erecti vel adscendentes, basi lignosi. 

Folia numero 16-22, integra, ab ellipticis ad lineari-lanceolata, margine 

indistincte dentato-serrata, rarius integerrima, cuneata, apice acuminata 

vel obtusiuscula, in axillis ramulis abbreviatis sterilibus praedita; folia 

basalia et media ad 45 mm lg. et 8 mm lt., opposita, elliptica, oblongoelliptica 

vel anguste-obovata, petiolata, petiolis ad l5 mm lg.; folia 

superiora - minora et angustiora, alterna, sessilia. Racemi terminales, 

solitarii, spicati, densi, obtusiusculi. Pedicelli tenui, l-1.5 (2) mm lg.; 

bracteae lineari-lanceolatae, pedicellis 2-2.5-plo longiores. Calyx 2 2.5 

(3) mm lg., in tl., vel 2/, in lacinias ovato-lanceolatas, acutatas dissectus, 

pilis brevibus glandulosis obtecti. Corolla atro-caerulea, 6.5-7.5 mm lg.; 

tubus parce curvatus, circa 3 mm lg.; petala 4.5 6.5 mm lg., anguste 

ovatolanceolata, acutiuscula. Antherae atro-caeruleae; stamina corollae 

aequilonga. Capsulae (immaturae) glandulosae; stylus 5.5-6 mm lg. (Vide 

fie.). 

l4 3ara: Np 915 209

T y p u s : Mongolia, Khobdo ditio, Altai Mongolicus, cursus 

superior fluminis Baian-gol, jugi Arschantyn-Nuruu declive generale 

boreale a haud procul a trajectu Baian-Hutel ,46"22'lat. bor., 9lol0' 

long. orient., alt. 1900 2100 s. m., in declivibus lapidosis, cum specie 

Juniperus sabina, et rupibus. 14 VII 2003, D. German (SSBG, isotypus 

LE). 

Aff'rnitas. Species inter V. porphyriana Pavl. et V. laeta Kar. et Kir. 

intermedia est. A primo pilis abbreviatis, caulibus tenuioribus, foliis 

angustioribus et numerosis, in parte caulis superne alternis, praesentia in 

axillis foliorum ramulis abbreviatis sterilibus, pedicellis longioribus; ab 

altero - indumento glanduloso, foliis in parte caulis inferne oppositis, 

capsulis pilosis differt. 

Species in honorem investigatoris florae Altaicae cl. S. Smirnovii 

nominatur. 

Msororeruue rpaBflHr,rcrbre pacleHnr l5'-32 cu Bbtc., rycro orlyueH- 

Hbre KoporKr4Mrl orcrorrql4Ml,r xeJre3r4crbrMu BolocKaMn. Kopneeuula 

KoporKr{e, BocxoAflurfle; cre6lu roHKrie, [pocrbre, oA[Hoquble rtJrr4 B 

ql4cJre HecKoJIbKI4x, [psMocroxrtwe vJwr BocxoAtulIle, r unxHefi uact'u 

oApeBecHeBarcruue. Jlucrrs B qr{cJre 16 22, rlelrbHbte, or ?JururrruqecKr4x 

Ao luueftHo-JraHrlerHblx, rlo Kpalo HeflcHo rturbqaro-3y6varrte, pexe 

uetrluorcpafrHbre, lrpu ocHoBaHLrr.r KJU{HoBnAHbre, K BepxyruKe rrpnocrpeHHbre 

IllJrH TyrroBaTble, B lla3yxax c yKopoqeHHbtMrr cTepr4JtbHbtMr4 Be- 

ToqKaMK; HaxHtae Lr cpeAHue go 45 urr,r Al. u 8 MM ul{p., cyrrportrBHbre, 

3[[I4tlTr,rqecKrre, [poAoJrroBaro-3JrJrr.{[Tr,rqecxne u.rrn ysro-o6parHoxiiqe- 

B?rAHbre, Ha rreperuKax,qo 15 MMAJT.; BepxHlre,- 6olee MeJtKr.re r{ y3Kue, 

o qepeAH bre, cr41xqr4e. Kucru BepxyueqHble, oArlHoqHbre, KoJloco Br4AHbre, 

rycrbre, TyfloBarbre. I-[oeroHoxru roHKue, I 1.5 (2) MM AJr., [pr4uBerHuru 

;ruHefiHo-JraHuerHbte utu larrlr.eit:r,arc, n 2-2.5 pa3a AJrr{HHee uBeroHoxex. 

tlaure.{xa 2-2.5 (3) uu AJr., Ha tlrv.nu t/, parAeneua Ha ocrpbre 

aftqesraAHo-JraHrrerHbre, KoporKo xeJre3rrcro ouyueHHbre Aonu. BeHqrlr 

TeMHo-cr4Hr{ir, 6.5 7.5 MM An.; rpy6ra cJrerKa u3orHyrar, orolo 3 mpr 

AJr.; Jre[ecrKu 3.5-4.5 MM AJr., y:rorfi qenn4Ho-JraHrlerHbre, [ptrocrpeH- 

Hrte. Tn.r[uxu c rduno-ct4HvrMvt rrbrJrbHKKaMr,I, paBHbr BeHqr,rKy. Kopo- 

6o.mu (Hespenue) xeJre3ucro ouyurdHHrre; crol6nx 5.5-6 urra an. (Cru. 

pr,rcyHor). 

T u u ; Mouronua, Xo6.u,occrufi afiuaK, Mosrollcruft A;rraft, nepx. 

p. Earu-rol, ceB. MaKpocKJroH xp. Apuranrrru-Hypy 6na: nep. Eaau-Xyrsn,46"22' 

c. u., 9l'10' s.4., h = 1900-2100 M. KaMeHHcrbIe" MeclaMrr 

cHJrbHo 3axycrapeHHbre cKJroHbr c Juniperus sabina, cKaJlbr. 14 VII 2003, 

!. A. feprraan (SSBG, r{3ornrr - LE). 

PoAcrno. fz6pu4oreuHufi nu4, [poMe)Kyror{unff rrrexgy V. porphyriana 

Pavl. u V. laeta Kar. et Kir. Or [epBoro Bl4pla orrl4lraercr 6olee 

KoporKI,IM otlylueHueM, ToHKrrMI4 cre6mun, 6onee ysrnruu a MHoforruc- 

JIeHHbIMI{, n nepxuefi .{acrI,t cre6Jtf, orrepeAHblMr4 Jrvc'tbflMt4, HalruqueM B 

210

i \ 

(-R..- \, 

t\ 

t i 

I ii 

', ,-r4.il 

,v ,rtl 

-J 

I ilr .u" 

. / 

Puc. Veronica smirnovii Kossaczev et D. German 

,l '- o6urufi Br{a pacreHr4r (no rrany), 1a - neHqnx 

2tr

fia3yxax JurcrbeB yKopoqeHHblx crepnnbHhtx Beroqex, 6olee g.lruuHuruu 

TIBeTOHOXKaMT4 ; OT BTOpOf O -- XeJIe3r4CTbrM OrryureHr{eM, cy[poTI,rBH br- 

Mr,r B Hr4xHeft qacru cre6ns JrlrcrbrMtr 14 BoJrocr4crbrun rcopo6o.rxauu, 

BUA HassaH B qecrb nccJreAoBarenr Qlopn Alrar C. B. Crrar.rpHora. 

Anropu npu3HareJrbHu A. lI. lllnaaxony r.r P. B. fuor.ueny 3a opra- 

Hu3aur.rlo lKcrreauul{oHHbrx r4ccJreaogaHLtft . 

Jlureparypa 

Koca.r€n lI. A. K npo6irenae rra6pu4oreHHoro nn4oo6pasonaHr,rr B poAe 

Veronica L. // 3sofforlrroHHar 6nororuq: Marepualrr II Mex4ynapogHofi 

rouQepeuqnr.r (Torr,lcr,2+26 orrq6pa 

2001 r.) / floa peA. B. H. CrerHns. Torr,rcr, 2002. C.365. 

K o c a q d n 11. A. O6sop ceKrlr.rn Pseudolysimachium W. D. J. Koch po4a 

Veronica L. (Suophulariaceae) no $lope Anraficrofi ropnoft crpaxu // 

Turczaninowia. 2003. Bun. 6, J\b l. C. ll-33. 

Anraft crnfi rocyAapcrBeuHufi yHurepcr,rrer 

656000, Eapuayl, np. JleHr.tra 6l 

E-mail: pkosachev@yandex.ru; german@bio.asu.ru

H. H. Llnerdn N. Tzvelev 

O PONE SPHAERAIITHUS L. I{ CLICTEME 

cJroxHouBETHhrx (ASTERACEAE) 

DE GENERE SPHAERANTHUS L. ET ASTERACEARUM 

SYSTEMATE 

B 1990 roAy B Aerrbre Bonru A. H. EaprraraHurvr, B. B. CuHqruHoit(I4ucrr.rryr 

3Koirornu Bonxcrcoro 6acceftHa) u B. H. flruuueuro (Acrpaxancxufi 

rocyAapcrBeuuufi yHunepcrarer) 6ru HafiAeu oprrnrHrurbHefi trufi flpeAcrasureilr 

Asteraceee, orrl4cansnft Havru uoA Ha3BaHueM Sphaeranthus volgensis 

Tzvel. (EapuuH14 !p.,1991 : 1770). Ero ulorHue qfiuesrAnne o6urue couserufl.7-l2 

MM Al. BHeruHe rroxoxr{ Ha roJroBKoBrrAHHe cor-lBerlrfl HeKoropbrx 

Br.rAoB Cephalaria Schrad. ex Roem. et Schult. (Dipsacaceae), uo s AeftcrBlrreJrbHocrli 

cocroflT u3 MHofoqI{cJIeHHbIx oqenb MeJrKr4x couBerufr-rcop- 

3r4HoK, cocroflrrrr{x ras 6 4llrraopQnux (2 o6oelorurx lr 4 necruqHux) qserroe 

c o6eprKofi r.r3 4 HeoAnHaroBbrx Jrr,lcroqKos. folosroBlrAHbre o6u1ue 

couBerr4r, B cBoro oqepeAb, co6pann B pbrxJryn o6nucrneunyro MereJrKy. 

K coxaneuulo, HaM Tof.qa ocraracb Hel{3Becrgoft ornocurelbHo Ho- 

Bas MoHorpaSur po4a Sphaeranthus L: (Ross-Craig,1954), r xoropoft 

uMelorcfl He rolbKo oqeHb [oApo6uue orvcanvrfl, Bcex Br,rAoB poAa, Ho 

r{ orlurrHo BbrrroJrHeHHbre pucyHKlr rrx c AeraJrrMr,r crpoeHur cousernfr. 

Hauu yxe 6una orne.{eua sua.ri{reJrbHar 6nu:ocrr BoJrxcKol'o BrlAa K 

nraqy S. strobiliferus Boiss. et Noe (1856, in Boiss. Diagn. Pl. Or. Nov., 

ser.2,3 : 6), uenecruoMy nrrtub nr 2 uecrouaxoxAeHuft r I4paxe. Cpanlrvruat 

Harrr BrrA c pr,rcyHKoM S. strobiliferus r aroft noHorpaQnu (Ross- 

Craig, 1954: tab. 3522 A), Mbr He cMorlr.r o6Hapyxnu raxux-ln6o pasttu,rui4 

Melrqy 3Tr4Mn Br4AaMu, I,r rroroMy S. volgensis, rro-Br{AnMoMy, cle- 

Ayer c{I,Iraru 6onee rro3AHHM urHoHrrMoM S. strobiliJbras. B 4enlre Bonfrr 

3ToT BHA Tporrr4qecKoro poAa BprA Jrr,r flBrfleTcr TpeTHqHbrM peJruK- 

ToM H, cKopee, 6lur saHeceH cloAa locJre rrocrreAHero oireAeHeHr,rr BoAo- 

[JraBarcLqItMI4 IITLIqaMU, HeKOTOpbre r,r3 KOTOpbIX 3IIMyIOT S 6aCCeftUe 

Turpa r,r ErQpara lrJur upoJreraror qepe3 Hero. 

Po!, Sphaeranthus L. rprrHaAJrexur r rpu6e Inuleae Cass. ceneficrsa 

cJroxHouBerHbrx, r{3 roropofi rrHorAa BbrAeJrflercr r oco6yru TporrnqecKylo 

u cy6rpounvecryrc rpr.r6y Plucheeae (Cass. ex Dumort.) Anderb. 

BMecre c prAoM Apyrr4x poAoB, H3 Koropbrx x Sphaeranthus sauiotee 

6rnsox pog Pterocaulon Ell., orrrqaflcb rJraBHbrM o6pasou rrpr{cyrc'l'BueM 

xoporuo pa3BlrToro [aIIIIyca I43 rrleTr.rHoK Ha ceMsHKax. o.reHl strcoKafl 

o6ulax cfierllraJrr{3agut Sphaeranthus He Bbr3brBaer couuesufi. 

Orono 50 elr.qos 3Toro poAa paclpocrpaHeHbr B TpoluqecKr,rx n orqacru 

cy6rponr4qecKr,rx o6nacrsx 6onrurefr vacrra ASpr,rru, IOxHofi Asuu 

n Ceaepuofi Ancrpanr4r,r 14 ToJrbKo BoJrxcKoe MecroHaxoxAeHue 

S. strobiliJbral HaxoAtrrcfl B rrpeAeJrax yMepeHHoreurroft soHu. 

2r3

MoHorpa$ po4a A. Pocc-Kpaftr (Ross-Craig, 1954:5 6) c.rnraer, rro 

B rrpeAeJrax poga Sphaeranthus HeBo3MoxHo flprlHqrb xarue-nu6o nuyrpr{poAoBbre 

rrogpa3AeJreHr{r, orpaHuq[Brrrrrcb JmlJJb BbtAeJIeHr4eM 4 rpyun 

Br{AoB, Ha3blBaeMbrx ercl (aJlbrHcauu> 6eg HasnaHlrft. Ha naru B3r.Jrflrl. 

HarHque HeMHofr4x ([epexoAHbrx> MexAy 3Tr.{Mr.t rpy[IraMtr Br4AoB He 

MOXeT CJryXr4Tb OCHOBaHneM Anr oTKa3a or BHyTpHpoAoBbrx rroApa3Aeteuuit, 

lrpeAJralaBuuxcfl Apyfr4Mu aBTopaMH. He clyuafino 6olee pauuufi 

uouorpa$ po4a B. Po6uHc (Robyns, 1924) pasaeurr rrot poA Ha 

2 uogpoga u 4 cexguu, a oAuH rr3 3Tux rroApo4oB Aa)Ke BbtAe[rJtcr B oco- 

6uft po4 Oligolepis Wight (1846,Icon. Pl. Ind. Or. 3 : tab. ll49;id. 1847, 

Calcutta Journ. Nat. Hist. 7 : l6l). Mn rarxe cqr{raeM, qro poA Aocra- 

ToqHo rrerKo AeJu4Tcs. sa 2 rroApoAa, u3 Koropbtx oAnH ,- rloApoA Sphaeranthus, 

r4Meer rrpeltMyulec'r'BeHHo rxapoBt{AHbre roJroBKH, BHeutHe oqeHb 

rroxoxfle Ha yMeHbueHHbre roJroBxu Echinops L., a gpyrofi , Pseudosphaeranthus 

Robyns (= Oligolepis Wight), BKrroqarcutrfi nonxcrnfi nug, 

6onlrue noxox Ha roJroBKr{ HeKoropbrx poAoB ceueficrra Dipsacaceae. 

Hnxe uu [pnBoAlrM cnucoK ocHoBHbrx Br4AoB poAa (ucxlroqar pqA 

Br{AoB, upuHqrbtx vroHorpaQoru poAa, Ha Haur B3rJuA, He Bcer-Aa Aocra- 

Tor{Ho o6ocnosaHHo, 3a pa3HoBr4AHocrr,r Apyrl4x Br,rAoB, r.r rpex arppu- 

KaHcKr{x BHAoB, onucaHHbtx rrocJre BbrxoAa uoHor.paQun) n rropa4re 

rrpeAJraraeMofi Hapru cucreMbl, B ocHoBHoM coorBerc.l.Byrouleti cucrene 

B. Po6uuca (Robyns, 1924) c yqeroM ua3rauufi, ycraHoBJreHHux 6olee 

paHHuMrr aBTopaMrtr. floApo.q Sphaeranthu,s rro coBoKyrrHoc.fr{ rrptr3HaKoB 

(orcyrcrrue o6ueft o6eprxn y ocHoBaHr{r loJloBoK, 6o.rrluroe KoJrr4qecrBo 

rrJrocKl4x H MaJIo orJrr4rranulnxcr Apyf or Apyra nucrovroa o6ep- 

TKH Kop3trHox, o6uvHo 6onturoe qucro uBerKoB B Kop3r4HKax) nroxHo 

cql,ITarb 6olee upuulrrrlBHbrM, H Mbr rroMeuraeM ero B Har{aJre cucreMbr. 

PoA Sphaeranthus L.,1753, Sp. Pl;927; id., 1154, Gen. Pl., ed. 5: 399. 

Typus: S.indicusL. 

flo4po4 L Sphaeranthus. 

O6urrae couBer?rr -. uapoBuAHbre, pexe lJrnuncouAaJrr,Hble nJrr,r rruluHApr4qecK[e 

roJroBKLr, y ocHoBaHrlt 6er yrelnueHHblx Kporcx1l{x Jlr4c- 

ToqKoB, pexe c He6ollruuutr JurcroqKaMu. Jlncroqrn o6eprru Kop3u- 

HoK B qucJre (3) 6- l8 (24), n"rrocrue u rtorrrt,t oAHHaKoBbre. flecrnqHnx 

uBerKoB B Kop3HHKax 6,18 (24). 

Typus: generistypus. 

Cexuus l. Sphaeranthus. 

fonosxn rJrapoBr4AHbre, peAKo 3JlJlr{rrcolrAaJrbHbre, y ocnonauur 6e: 

yBenuqeHHblx Kporoulr4x lucrorlKoB. O6uee uBeroJroxe or uapoBlrAHof 

o Ao 3JrJlr{rrCOrlAZUrbHOf o. 

Typus: generistypus. 

l, S. aJricanus L. -- 2. S. indicus L. - 3. S. gazaensis Bremek. 

4. S. senegalensis DC. 5. S. confertiJblius Robyns. - 6. ^S. bullatus 

214

Mattf. - 7. S. angolensis O. Hoffm. - 8. S. polycephalus Oliv' et 

Hiern. - 9. S. flexuosus O. Hoffm. -- 10. S. humilis O' Hoffm. 

ll. S. chandteri Ross-CraiC. - 12.5. greenwayi Ross-Craig' -- 13. S. napierae 

Ross-Craig. 

Eorrruar qacrb BuaoB ceKIrI,rIl cocraBJlter BIIoJIHe ecrecrBeHHyrcl 

rpyuny. OAHaxo S. conferti/b/lzs nueer 6onee yrroe o6uee qBeroJloxe 

u 6olee MHoroql4cJreHHble IrBerKI,t B Kop3uHKax (uecutuuux 6onee 20, 

o6oerrorr,rx 6 li 6onrure), S. greenwayi uueer o6eprxu Kop3lrnoK I{3 

3 nncroqroso 6 uecrltuHltx uBerKoB u I o6oeuolnft useror. Etrle 6olee 

yKloHqercf, or ocHoBHoro TI4rIa s. napierae, uuerculuft AoBoJlbHo clrJrb- 

Ho cniuocHyroe o6qee IrBeroloxe n 3-4 tmcro'rra o6eprrn Kop3l{HoK, 

Bo3MoxHo, 3acJryxl{Bat BbrAereHI,Ifl n ocobyto ceKul4to. 

Cexqus 2. Polycephalas (Forssk.) DC., 1836, Prodr.5 :370. - 

Polycephalos Forssk., 1775, Fl. Aegypt.-Arab. : 154. 

Typu s: S. suqveolens(Forssk.) DC. 

Kax npegu4yqat ceKqut, Ho Kpoloulue Jll4crorrKu roJloBoK 6olee 

pa3Br,rrH r,r BbrcrylrarcT y ux ocHoBaHI,Ifl, a 6orosrte rucrorrKu o6epror 

KOp3trHOK rracTo HeMHOTO BAOnb CJIOXeHHbIe. 

14. S. suaveolens s.l. (nrnnvaer HecxoJlbxo BIdAoB, IIpIlHfl'rbIx MoHorpaQovr 

poAa 3a pasHouuguocru). 

3ra cexuur 3aMerHo yKJloHflerct B cropoHy ceKulrll Pauciflori cJIeAy- 

Krurero uoApoAa, rc roropoft u orHoclrr ee BI4AbI Po6r,rHc (Robyns, 1924 

:179), xorfl rro ApyI'I4M tIpI,I3HaKaM oHa tBHo 6oree 6nusKa x tnttoeoil 

ceKuuu. 

Cerqns 3. CylindrocephaliRobyns, 1924, Bull. Misc. Inform. (Kew) 

1924 : 179 (

oJroBoK. Jlucro.rrtl o6epror Kop3r,rHoK B q[cJre 2-4 (7), r{3 Koropbtx 

6oxosHe o6rrquo BAoJIb cJroxeHHHe, KLIJreBarbte. a HapyxHbte o6rI.tno 

cr,rJrbHo paculrpeHHbre u [ptrKpbrBarou-u4e co6on Kop3r4HKr,r. flecruqulrx 

rrBerKoB B Kop3lrHKe o6H.rHo 4-5, pe4ro l-3 nnvr 5-9. 

Cexunr 4. Pauciflori O. Hoffm., 1895, Bot. Jahrb. 20: 227. 

Oligolepis Wight, 1846, l. c.:tab. 1149, s. str. (typus: S. amaranthoides). 

Sphaeranthus sect. Oocephali Robyns, 1942, L s. : 179 (

eceptaculo communi paulo convexo. Involucri phylla in numero 2. -- 

T y p u s : S. oppositi/b/rzs Ross-Craig. 

Jlucrrr cy[porrlBHbre. foflosxt{ [oqrl,I rIoJIyuapoBtIAHbIe, AoBoJlbHo 

MiLIIOIlBerKOnrre, 6lu: ocHoBaHnfl c KpyrIHbIMt I Kporou{,IMLI n[cTor{KaM[, 

c HeMHOIO BbrnyKJrbrM o6ulurr,r IIBeroJIoxeM. Jlucroqxoe 

nor 2. - T n n : S. oppositifolias Ross-Craig. 

36. S. oppositifulius. 

o6eprxn Kop3l,l- 

Mn c.r[rae[r, qro eAr4HcreeHuufi Br,IA poAa c cyupornBHblMtr JrI,IcrbflMr4 

3acnyxr,rBaer BbrgeJreHur b oco6yto ceKrlrirc, a, Bo3Mo)I(Ho, H B oco- 

6ufi uoApoA, Kporrae rofo, oH r{Meer BocxoArull,Ie cre6lu c MHoroqucreHHbrMrr 

[a3yrrrHbrMr4 no6erauu, Bo3MoxHo, flB lrfl.flcb MHof o JIeTHL{Ko M. 

flo crpoenuro o6uero uBeronoxa roJIoBoK on 6lulor 

Ayulefi ceKul,ttt. 

K BIAaM rIpeAbI- 

Cexqus 7. Cristati Tzvel. sect. nova. -- Capitula conico-ovata, receptaculo 

communi cylindrico. Involucri phylla in numero 2, lata, dorso 

cristadentatapraedita. -T y p u s: S. cristatus Ross-Craig. 

f olosrcu Ko HrlqecKll-tfi qeurquue, c uLIJuIHApI4trecKI,IM o6ulr.I u uBero- 

JroxeM. Jlt,tcro.rros o6eprrn 2, uruporux, Ha c[rlHKe c ay6varrrrr,r rpe6- 

HeM. -T u u: S. cristatus Ross-Craie. 

37. S. cristatus. 

EAuucrnenulrft sr.rA ceKrrr,ru eule Ma-rro Lr3yqeH, Ho rro $opue o6qefo 

uBeroJtoxa, Iloxoxero Ha uBeroJro)Ke ceKuHr.r cylindrocephali, w ttucroqKoB 

o6eprru Kop3rrHoK oH BpflA Jrlr Moxer 6url rroueureH B ceKqI4K) 

Pauciflori, c Br,rAaMH roropofi r,rMeer MHoro o6ulero. 

flpegcraueHrrr o crrcreMe cJroxHorlBerHbrx (Asteraceae) 3a nocJre- 

AHr4e AecflTr{nerr{fl cyulecrBeHHo r,r3MeHlrJrr{cb, HraBHbrM o6pasou Ha oc- 

Ho BaH?rr{ MoJreKyJrrpHo -reHerlrqecKr,rx r4ccJreAo vauuir, o r{eM MoxHo cy- 

Ar4Tb rro BbrrreA[urM B nocJreAHee BpeMq Ha pyccKoM r3brKe pa6oravr 

P. B. Kauemna (2000) u 9. Axe$$pu (2002). flpan4a, 3 ocrosHux noApa3AeneHnfl 

ceraeficrsa - rroAceMeficrsa Asteroideae (Cass.) Lindl., 

Carduoideae Cass. ex Sweet n Cichorioideae (Juss.) Chev. ra 6onruraHcrso 

yxe AaBHo ycraHoBJreHHux rpu6 coxpaHunncb, xorfl irr4clo rrogreNreficrs 

yBeJrr,r{eHo Ao 5 y,{xeQ$pn (2002), [pl4Hr,rMarouero eule noAceMeficrsa 

Barnadesioideae (D. Don) Bremer et Jansen n Mutisioideae (Cass.) Lindl., 

I,I Aaxe 4o l0 (Panero, Funk, 2002). I,I: 5 uostrx noAcenaeftcrB rrocJre- 

AHlrx aBTopos 3 rsrqrcrcq MoHorI-InHbrMr,r, a BbrAeneHHue us Mutisioideae 

no4ceueftcrsa GochnatioideaePanero et Funk u Pertyoideae Panero 

et Funk coAepxar uo 5-6 rpo[uqecKr,rx u cy6rpour{qecKrrx poAoB. 

flo uonerylrpHo-reHeruqecKuM AaHHbrM (Panero, Funk, 2002 : 911), 

rpyilrraMu, paHee Bcero orAeJIItBIrMMUcfl or o6tr1elo crBoJra cJroxHoqBer- 

Hbrx, rBJrrrorcfl rroAceMeficrra Barnqdesioideae, Goghnatioideae u 

217

Mutisioideae, Koropbre, KaK ora3aJlocb, rloKa3blBator HeKoropoe poAcrBo 

K ceprefrcrear4 Calyceraceae u Goodeniaceue. ?lo o6croare.urcrso 

AaJro ocHoBaHr4fl MHor-HM aBTopaM, B ToM trHcJIe 

.{xe$$pu (2002), tto- 

Mecrurb ?Trr rroaceMeftcrsa s HaqaJre lnuefiHofi cncreMbr ciroxHouBe'r- 

Hbrx, rrocJre Koropbrx r{Ayr rroAceMeficrna Carduoideae w Cichorioideoe, 

a 3aHuMaB[Iee HaqaJIr'HOe uOJTOXeHUe B I]pe]KHLIx CHcTeMaX noi{Ceuefrcrso 

Asterotdeae oKa3aJrocb B KoHue cl,tcreMbr. 

Xors raroe pacrroJroxeHue uoAcenaeiiclB B Hacrofluree BpeMf, rloJryquro 

rIIHpoKoe paclpocTpaHeH[e B rr,rreparype, Mbr Bce xe cqHraeM ero 

HeAocraroqHo o6ocHosaHHbrM 14 upeAflorrnraeM npr,rH H Malr rr r.r Hefi Hyto 

cncreMy, 6.rtn:ryrc K cr{creMe H3BecrHoro cfiequaJrr4cra rro cJroxHouBer- 

HbrM A. Kpouxuacra (Cronquist, 1955), craBuBrrero Ha rrepBoe Mecro 

uau6o:ree rtpuMr4rnBHyrcl uo coBoKyllHocrrr rrpr{3HaKoB rpu6y Heliantheae 

Cass. ue no4cerraefrcrsa Asteroideae. ?ta crrcreMa 6rrLra rrpnun'ra 

uauu (I{nenes. 1994 : 5) n crogre ). 

Karoerl xe ocHoBaHHr, \rrfl rroMe[IeHlrt Heliantheae B Harrale cucre- 

Mbr cnoxHorlseruHx? 3ro npex4e Bcero rrpaBr4JrbHble (KoJloKoJrbt{arbreD 

BeHr{l{Kr{ rpy6uarux rJBerKoB, AoBoJrbHo rxrrpoKoe pac[pocrpaHeHne B 

[peAeirax rpn6u [phuBerHr4Kon Ha o6uleM uBeroJroxe H rra[rryca !r3 

qeuJyeBuAHbrx crpyrffyp (a ue us BoJrocKoBr.rgHbrx ureruHox), a laxxe 

xeJrroro rudrMeHTa B rlBerKax. Orcyrcrnue 3Tr{x nprr3Haxon y Barnadesioideae 

u Mutisioideae 3acraBfirro .{xe$$pu r,r HeKoropbrx Apyl-ux aB- 

TopoB cqrrrarb r4x He rrpuMr4Tr,rBHhrMr,r, a BTopr4rrHbrrvrn:

peAKoB c surolropSHbrM BeHqHKon, blu:rcnx x Calyceraceae uilu Loheliaceae. 

Xenrar oKpacxa rrBerKoB rr B Apyrlrx ceMeficrsax [orpbrroce- 

MeHHbrx pacreuufi BbrrJuArrr 6ouee upuvu4TrrBHbrM rrpr,r3HaKoM no cpaB- 

HeHr4to c gpyrofi orpacrofi - pe3yJrbraroM rrpucyrcrBr4r B qBerKax rtr4r- 

MeHToB aHTorruaHoBoft rpyulu LrJrr,I orcyrcrBlreM BctKrIx nIlrMeHToB B 

crry.rae 6enofi oKpacKr,r. Orcyrcrnue alr6uHocos y xeirroqBerKoBbrx 

BI,IAoB TaKXe MoxHo cql4Tarb IroATBepxAeHueM orHocurenrHofi lrpnMr{- 

TI,IBHOCTI4 }KCJITbIX IHTMCHTOB, 

flpu.rnua onru6oqnoft TpaKToBKn sBoJrloqnoHHblx prAoB orAeJlbHbrx 

[pn3HaKoB (cerraoQra.uesa) y clroxnoqBerHbrx rrpeAcraBJrf,ercr HaM Aocra- 

ToqHo scHofi. Pacnonoxerure coBpeMeHHbrx raKcoHoB y ocHoBaHur KJra- 

.qucrr{qecKoro (ApeBa), To ecrb san6onee paHHee orAeJreHr4e nx or o6ulefo 

clBona 3BoJrrcqnoHupylou1ux t[ulyuon, HuKaK He o3Haqaer, qro 

Bce r4x rlpn3HaKrr [plrMHTr{BHbr. Hanporun, atu 6onee ApeBHHe u 6olee 

o6oco6leuHbre raKcoHbr He Moryr He 6rrrr oqeHb BbrcoKo c[eutraJru3r4poBaHHbrMr{ 

u rroroMy paHo rrpeKparuBuruMl{ 4anrHefiuryrc 3BoJrrouurc. 

Tarue raKcoHbr Moryr r,rMerb orAeJrbHbre !r Aax(e oqeHb upr.rMrdrrrBHbre 

rIpH3HaKn (xax upoauleHue rerepo6arnun), Ho oHr.r AoJr)KHbl 6nrr gurcoKo 

c[euuairr,r3npoBaHHblMn B Apyrnx orHorueHr{rx. flolrouy Barnctdesioideae 

u Mutisioideae B xohe csoeft esorrcur{r4 BrroJrHe MofJr?l He3aB}.Icr{Mo 

or Apyrr4x fpy[rr cJroxHouBerHbrx [orepqrb rrptruoerHr{Ku o6ulero 

IlBeroJloxa, KaK r,I upno6pecru rrarrnyc r43 rtleruHoK u HerrpaB[JlbHbre 

BeHquKH qBerKoB. Orueruu, cro Barnadesioideae, KaK 14 rpyurra poAoB 

noAceueftcrsa Mutisioideae, BbrqerfleMar B rrocJreAHee BpeMq s oco6oe 

roAcelreficrso Gochnatioideae (Benth, et Hook. f.) Panero et Funk (2002: 

913), r,rvrenr roJrbre pbrJrbrla -- [pr,r3Har rBHo BTopuqusrfi no cpaBHeHuro 

c pbrJrbrraMrr Apyrrdx rpyln cJroxHouBerHux. Cpe4r.r Gochnatioideae 

o6Hqnrl n 6o.rree BbrcoKo c[euuaJrr,r3r4poBaHHbre tro cpaBHeHuro c rpaBa- 

MIa xrr3HeHHue $opuur AepeBbeB H KycrapHnKon. Ha Ky6e n oKpec'rHocrqx 

fasaHrr HaM rrpuxoAr,rJrocb Br,rAerb Gochnatia ilic'iJblia Less. 3ro 

cr{JIbHo pa3BerBJreHHoe AepeBue 2-3 M sHc. c KoJrIoq[Mu Koxr4crbrMn 

JurcrbqMr,r, oqeHb rroxoxL4Mr Ha Jlncrbr llex aqudblium L., u KpylrHbrMr{ 

oAnHoqHbrMr4 Kop3HHKaMu c 6enosarsrMr4 r.IBelKaMr4 r{ rrarr[ycoM r{3 ule- 

TnHoK. Bce ero ttpH3HaKH cBl4Aerelbc'rByrcT o sHcoroft cfleuttaJtr,r3aur4r,r. 

flo-suAlrlloray, r4 Bo MHornx Apyrr4x cJryqarx, Llcrlonb3yf, MoJreKyJrrp- 

HO-IeHeTnqecKr{e KJIaALlCTr4qeCKHe CXeMbI, CJreAyeT He Sa6nSarl, qTo co- 

BpeMeHHbIe raKcoHbr, paHbllre rcerlo o6oco6ueuruecr or o6t:1elo crBoJra 

r,r rroroMy nau6onee ApeBHHe rro Bo3pacry BcerAa rBJl{K)Tct Bbrcorocrreur{aJrrr3r{poBaHHbrMrr 

(:rolrcuuoHHo rrpoABr.rHytrura) 14 Moryr noo6rqe 

He rrMerb rlpuMltrtrBHbrx lpl43HaKos. Her ocnoraHnfi lroMeularr, I4x B 

HaqaJIe c[cTeM, ecII4, KoHeqHO, TaKCOHbI B Cl4CTeMaX paCI]OJraI'a1.b uO 

creueHr4 y6unauua [puMI4TuBHbrx oco6egHocrefi, a He lo ux Bo3pac'fy. 

Tax ua o6ulena KJraAucrr4qecKoM (ApeBeD rroKpbrroceMeHHbrx rrocJteAHero 

BpeMeHu 6rus ocuosaHr4s aroro (ApeBil) pacrroJraranrcr rarne ceueffcr- 

2t9

Ba KaK Nymphaeaceae, Ceratophyllaceae u Amborellaceae, Koropbre 

oqeHb BbrcoKo cnequaJrr,r3r.rpoBaHbr, ll o6ulze [peAKI{ rroKpbITOCeMeHHblx, 

ecnv o*t4 cyulecrBoBatu, 6rl'nw coBceM He floxo)Kli Ha Hr,rx. flplrsuarcn 

locJreAHux Moryr 6nrr pacceaHHblMrr uo caMbrM pa3ruqHbrM KJraAaM 

(ApeBa), a B [paB[JIbHOCTr,r [pLrHHMaeMoro MHOTUMTI aBTOpaMrl (r rorr,r 

rurcJre, 

lxeQSpra, 2002) npr.rHuurra [apcr4Mouuu (upuHqun axoHouun) 

Mbl oqeHb coMHeBaeMct. 

klcxoln H3 cxa3aHHoro, Mbr cKJroHHbr rrpr4Hr{Marb 4la poccuftcrux 

cJroxHouBersnx luneftHyro ctrcreMy, o6parHyrc ctrcreMe {xer[$pu 

(2002:10 ll) u 6rur:ryru cucreMe A. Kpournracra (Cronquist, 1955), 

Koropaq BbrrJuAtrr cJreAynqr.rM o6pasorra. 

floAceueftcrso l. Asteroideae (Cass.) Lindl 

Tpu6a l. Heliantheae Cass. (incl. Ambrosioideae Raf., Coreopsideae 

Lindl., Helenieae Lindl., Tageteae Cass., Eupatorieae Cass.). 

Tpu6a 2. Inuleae Cass. (incl. Plucheeae (Cass. ex Dumort.) Anderb.). 

Tpu6a 3. Senecioneae Cass. 

Tpu6a 4. Calenduleae Cass. 

Tpn6a 5. Anthemideae Cass. 

Tpn6a 6. Astereae Cass. 

Tpu6a 7. Gnaphalieae (Cass.) Lecoq et Jullet (incl. Filagineae (O. Hoffm.) 

Poljak.). 

flo4ceueiicrso 2. Cichorioideae (Juss.) Chev. 

Tpu6a 8. Cichorieae Lam. et DC. 

Tpu6a 9. Gundelieae DC. ex Lecoq et Jullet. 

Tpra6a 10. Arctotideae Cass. 

floAceueftcrso 3. Carduoideae Cass. ex Sweet. 

Tpra6a ll. Cynareae Lam. ex DC. 

flo4ceueftcrso 4. Mutisioideae (Cass.) Lindl. 

Tpu6a 12. Mutisieae Cass. 

flogcenaeiicrno Barnadesioideae, Ha Har[ B3rJnA, cJreAyer fioMexlarb 

B KoHrIe cucreMbr nocne Mutisioideae vrnu Gochnatioideae, ecJrr,r rrocJre- 

AHee uoAceMeftcreo BbrAeJrrrb r,r3 Myrr,r3ueBHx. Ero BbrcoKar cfleurraJru- 

3arp4r, y Hac He Bbr3btBaer couHeHllft, o qeM cBr4AereJrbcrByror rrper{My_ 

ulecrBeHHo KycrapHr4xoBa.f, xtr3HeHHar Sopua, gayry6rre rrBerKrr, rracro 

.quN,ropQHbre ceMrHKr,r c IIaIIIIycoM r43 rqerrrHoK, Koropbre nrolyr 6r,lrr 

uepr4crbrMr,r. Koporxae rr yroJrrqeHHbre pbrJrbua 6er xarux-nn60 sofioc- 

KOB r,IJII4 COCOqKOB TaKXe CneAyeT cqr,rTaTb BTopr,rqHbrM upnsuarolr 6o_ 

ree sHcoroft c[eqqiltu3auuu. KoneqHo, Hr4 oAHo uoqcerueficrno or ApyfUX 

HE BbIBONI,ITCfl. 

P. B. KarvrenuH (2000) npuHrlMaer B KaqecrBe eilIe oAHoro noaceueftcrsa 

Ambrosioideae Raf. rpynny 4eficrnnromHo oqeHb opnnrHaJrbHbrx 

poAoB, r43 Koropbrx s Poccul,r Bcrpeqarorc , Amhrosia L. u Xanthium L., 

220

Ho MbI, cneAyf, Axe00pu (2002) I,I ApyrI{M aBropaM, upeArotrltraeM 

BKrroqarb ero n rplr6y Heliantheae, c xoropoil srn poAbr recHo cBr3a- 

Hbr qepe3 pogu lva L. u Cyclachaena Fresen. Moxuo upe4cranurr ce6e, 

rrro Kop3r,rHKr4 Tr,rfifiqHbrx Heliantheae B xoAe 3Bonror{uu cr.rJrbHo yMeHb- 

IULIJII4Cb, HO yBenHqUnUCb B KOIUqeCTBe, a JIOXHOt3bIqKOBble IIBeTKI4 n 

I]aI]IIyc peAyur.rpoBarurcb, B pByJrbTaTe qero Bo3HuKJrr,r poAa poAcTBa 

Ivo. 3areu Kop3r,rHKr{ craJrr,r pa3AeJrbHorroJrbrMu, fipuqeM Kop3r4HKr4 c 

THquHorrHbrMrr rrBerKaMr.r coxpaHrrJur crpoeHrre Kop3r4HoK lva, a xopzuu- 

Kn c fiecrnqHblMrr qBerKaMIa cI4JIbHo BI{AoI43MeH[Jlr4cb, Irpr4ctloco6uerurrcb 

K BbrcoKoc[euu:Lrrr43rrpoBanHofi grcsogooxoplrl{ y X ant h ium. floxoxee 

Ha[paBJreHrre 3BoJrlourrr4 nMeJro Mecro r,r B Apyrr.rx rpu6ax Asteroideae 

(n rpu6e Inuleae oHo rrprBeJro r $oprrar.rpoBaHr4lo poga Sphaeranthus, 

z rpv6e Anthemideae - K poAy Artemisia L., upegraun Koroporo, 

Bo3MoxHo, 6urtu KaKr{e-To Br.rAbr poAa Chrysanthemum L.), oAHaKo 

y Heliantheae oHo 3arxJro oco6eHuo AaJreKo. I,Iurepecuo, uro y Heliantheae 

wst BcrpeqaeMcf, rr c AoBoJrbHo rurrporrdM pacrrpocrpaHeHueM eule 

OAHOTO BTOpr4rrHOrO [prEHaKa -- Cy[pOTHBHbTX Jrr,rCTbeB, Ha Ha[r B3fJrrA 

(!,renen, 1993 : 54), flBJrrroqerocr pe3yJrbraroM oAHof,pycHoft uonuueprrsarrulr 

$urorraepon. B 4pyrux rplr6ax cyflpornBnbre Jrucrbr Bcrpeqa- 

Iorcr oqeHb peAKo, a n rpu6e Inuleae oAHr.rM r,r3 raKrlx ucxnroqeHlrii rs- 

Jrrercr yuo urHyrurfi orl'urc S p hae r an t hus o pp o s i t ifu lius . 

Cronr yfloMrHyrb, rrro ocHoBHar Jrr{Teparypa rro cr4creMarrlKe cJrox- 

HOTIBeTHbTX BrrJrOTb AO HaCTOTUIeTO BpeMeHr{ [pnBeAeHa B rltrrr{poBaH- 

Hofi naura crarbe 9. !xe$$pu (2002). 

Pa6ora BbrrroJrHeHa upu Qr.lnanconoft noAAepxKe rro rpaH'ry (BeAyulue 

HayqHbte uKoJIbl). 

Jlureparypa 

Eapnlrn A. H., Il r4rrlrfleHKo B. H., Cr,rHrKnHa B. B., Inen 

e n H. H. Sphaeranthus (Asteraceae) - nor'ui,r poA Anr $nopu CCCP ll 

6or. xypn. 1991. T. 76, J\b 12. C.1768-1771. 

Ixe Q @p n 9. CucrepraruKa cJloxHolrBernsrx(Compositae)s na.{are XXI 

seua ll 6or. xypn. 2002.T.87, Ns IL C. l-15. 

K a lr e I lr n P. B. CnoxnouBerHbre (rparxraft o6sop clrcrenarr). CfI6.; Eapuayn, 

2000. 60 c. 

I n e n e e H. H. 3sorrcqr.r' Qr.rrorraepa y Bbrcmux patennir ll Brotr. MOI4n. 

Ora. 6r.ron. 1993. T. 98, sun. 2. C. 53,60. 

I{nenen H. H. flpe4racrroBue ll @rcpa enponeficxoft.{acrx CCCP. Cn6., 

1994. T.7. C. 5-6. 

Candolle A. P. de. Compositae // Prodromus systematis naturalis regni 

vegetabilis. Parisiis, 1836. T. 5. P. 4-706. 

C ro nqu is t A. Phylogeny and taxonomy of the Compositae //Amer. Midl. 

Naturalist. 1955. Vol. 53. P. 475-511. 

Hof f mann O.Compositaeafticanae.ll//Bot.Jahrb. 1895. Bd20. S.219- 

221

P a n e r o J. L., Funk V. A. Toward a phylogenetic subfamilies classihcation 

for the Compositae (Asteraceae) // Proceed. Biol. Soc. Washington. 2002. 

Vol. ll5, N 4. P. 909-922. 

R o b y n s W. Revision of the genus Sphaeranthus // Bull. Misc. lnform. Kew 

(Kew Bull.), 1924. N 5.P.177-199. 

Ro s s -C ra ig S.A revision ofthe genus Sphaeranthus // Hooker's lcones 

Plantarum, ser. 5. 1954. Vol. 6, pt l. P. l-90. 

Eoranu.recrcufi uucruryr uu. B. JL Kouapoaa PAH 

197 37 6 C.-flerep6ypr, yrr. ftporb. flonora, 2 

E-mail: tzvelev@DG4972.spb. edu

A. C.3epnon 

A. Zernov 

HOBbIE TAKCOHbI LI HOBbIE KOMEI4HAIIHLT B POAAX 

PACTEHI,Ift OJIOPbI KABKA3A 

TAXA NOVA ET COMBINATIONES NOVAE IN GENERIBUS 

PLANTARUM FLORAE CAUCASI 

Bo nperraa peBu3HLI Slopu Cerepo-3anaAHol'o Kasra:a Bo3HI4KJIa 

Heo6xoAlrvrocrb n3MeHeHI4g cl{creMarl,tKl,t I.r raKcoHoMtrll ptrAa poAoB I,I 

BT4AOB H3 pa3Hbrx Ceneficrn, [peAcraBileHHhlx Ha rroil reppuropuu. 

Typha L. sect. Rohrbachia (Kronf. ex Riedl) A. Zernov comb. et stat. 

nov. -'- Typha L. sect. Bracteolatae Graebn subsect. Rohrbachia Kronf. 

ex Riedl. Rohrhachia (Kronf. ex Riedl) Mavrodiev, 2001, Eor. xypH. 

86,9: 120. 

E. B. Manpoaraen (2001) crpaBeAJIHBo orMeqaer, uro Typha mininn 

Funk ra 6nt,tsrlre nn4rt uoptfofior[qecrl,I o6oco6neHll or Apyfux BllAoB 

ceKur4rr Bracteolatae Graebn. B csr:n c 3TI4M BbIAeJIeHI{e nx s oco6lte 

TaKcoHbI B[oJrHe o[paBAaHHo, Ho MHe rlpeAcraBJlgercq 6oJlee [paBuJrb- 

HbrM paccMarpr,rBarb rpyffIy poAcrBa T. minima B paHre ceKuI,IH. 

Ruscus L. sect. Platyruscus (A. Khokhr. et V. Tichomirov) A. Zernov 

comb. et stat. nov. --- Platyruscus A. Khokhr. et V. Tichomirov, 1993, 

Eruirn. Mocr. o6ul. r,rcn. trptrp., orA, 6uo.rr. 98, 4 :92. -- Ruscus L. ser. 

Simplices P. F, Yeo, 1968, Notes Roy. Bot. Gard. Edinb. 28 : 250. 

OrnoureHug B poAe Ruscus s. l. K s ceMeftcrse Ruscaceae B uerloM 

uo4po6Ho paro6paHu A. fI. XoxpflKoBbIM Ia B. H. Tuxomuponr,lu 

(1993). Ha ocsone ananr43a rlplt3HaKoB pa3Hblx BUAoB oHK rIpI4luJIH K 

BbrBoAy o Heo6xoAuMocrtr BbtAeJIeHI,lr oco6oro poAa - Platvruscus. 

B raqegrse ocHoBHbIx Al,IarHocl'I4trecKux [pI,I3HaKoB orMeqeHbr upocrble 

rro6eru (uuerorca BBI,IAy HeMerarnrop$uruponaHHue) u rpyuultfi Kpolotl1ufi 

nucr couBerufi. B ro xe BpeMt raxofi saxHlrft npH3HaK, KaK Hailutrue 

SyurquouzutbHo oAHorIoJIbIx IIBeTKoB co cso6oAHltMI,I Jlucl'o qKa- 

Mn oKoJrorlBerHnKa, oKa3brBaercfl o6lrlnu .qnq o6oHx poAoB. flo 4py|r,rna 

rrpH3HaKaM TaKxe ua6rrcAaerct nepeKpbtBanL4e Ruscus s. str. n Platyruscus. 

Bce eto, na uofi B3rJuA, cBuAereJlbcrByer B rroJrb3y eArrHc'tBa 

poga Ruscus, r{ [o3BoJr-f,er rlpl{3Harb B ero cocraBe ABe ceKqllu Ruscus 

w Platyruscus. 

Cephalanthera Rich. sect. Cucullatae A. Zernov sect. nova. - Corollae 

labium calcare brevi conico. 

T y p u s: C. cucullara Boiss. et Heldr. ex Reichenb. fil. 

Sectio monotypica. 

fy6a c KoporKr,rM KoHr,rqecKuM ufioprreM. 

T u u : C. cucullata Boiss. et Heldr. ex Reichenb. fiI. 

Mosorunsaf, ceKuur. 

223

Tpa4uquouno pop, Cephalanthera upuHuMaercs 6es nltAeneHl{t Karux-nn6o 

BHyrpr,rpoAoBbrx raxcoHos. OAHaxo C. cucullata orruqaercq 

or ocraJrbHbrx Br,rAoB poAa HaJu4trueM Iunopua B ocHoBaHI,Iu ry6rt ororouBeTHnKa 

r4 cr{3oBaTblMl4, peAKO paccTaBJIeHHblMIr, MeJIKIiMU, AO 4 CM 

qil. u 2 cM [Hp., Jr]rcrbf,Ml,I. 3ru npusHaKu rIo3BoJItIor BbIAeJInrb 

C. cucullata n oco6yrc ceKLII4Io. HHorAa Ans BocroqHoro Cpegu3eMHo- 

Mopbr, Kasxa:a n Malofi A:uu npunoAurcs HecKoltro 6lu:lxrtx BIi- 

AoB--- C. epipactoides Fisch. et C. A. Mey., C. Jloribunda Woronow, 

C. kurdica Bornm. ex Kraenzlin. ApryrvreHraqlrfl cBeAeHrrs ux HagsaHl'lfr 

B cr{HoHHMbr x C. cucullata I\aHa vruoii pauee (3epHon, 2000). 

Polygonum L. subgen. Bistorta (Scop.) A. Zernov comb. et stat. 

nov. - Bistorta Scop., 1754, Meth. Pl.:24. -- Polygonunz L. sect. 

Bistorta (Scop.) DC., 1805, in Lam. et DC., Fl. Fr., ed.3,3:364. 

Polygonum L. subgen. Bistorta (Scop.) A. Zernov sect. Vivipara 

(Tzvel.) A. Zernov comb. nova. Bistorta Scop. sect. Vivipara Tzvel., 

1987, Hosocrlr cucr. Bbrcru. pacr. 24 :75, 

Bo:uoxuo, uMeloularct reHAeHqI{t pa3AeneHl{t poga Polygonum s.l. 

Ha HecKoJrbKo MeJIKIIX poAoB Bo MHofoM o6ocHoeaHua. Ho ec.rtu oAgu 

r,r3 Hrrx BrronHe o6oco6reHH, KaK, Hartpt,tuep, Reynoutria,ro Apyrue nMeror 

rrepeKpbrBaroul[ecr [pr.r3HaKu: Bist or ta, Ac onogon, P ersicari a. floc- 

JleAHHe TaKCOHbr, Ha HaIx B3fJItA, lIpaBHJIbHee [pI{HHMaTb B paHfe I]OApoAoB 

poga Polygonum. 

Arabis hirsuta L. var. sagittata (DC,) A. Zernov comb. nov. A. sctgittata 

(Bertol.) DC,, 1821, Regn. Veg. Syst. Nat. :221. -- A. sagittata 

(Bertol,) DC. var. subglabrata DC,, 1821, l. c.:222. ? A. abietina 

Bornm., 1936, Feddes Repert. (Beih.), 89 :26. 

MHe npegcraBJrflercr 6olee npanuJIbHbIM rlpllHrrMarb A. hirsuta s 

rrrnpoKoM o6leue, He BbrAeJr.rlfl I{3 Hero MeJIK[e $oprr,ru (A. sagittata 

(Bertol.) DC; A. planisiliqua (Pers.) Reichenb.; A. gerardii (Bess.) Bess. 

et W. D. J. Koch), orucaHHbre rar oco6lte nuArt. I4x o6oco6neuue 

6lrno ocHosaHo rrperrMyulecrBeHHo Ha Sopue JlrlcrbeB (Candolle, l82l). 

Kax uor

subglabrata s A. hirsuta B KaqecrBe pa3HoBHAHocrI,I [paBl{JlbHblM Ha3Ba- 

Hr,reM uocreAHefi, cotrracHo crarbe I1.6 (fpeftrep 14 Ap., 2001), AoJt]I(Ho 6btrr, 

A. hirsuta var. sagittata. 

Crambe L. sect. Tatariae (Khalilov) A. Zernov comb. et stat. nov. '.-' 

Crambe L. sect. Crambe subsect. Tatariae Khalilov, 1991, Eor. xypn.76, 

1l : 1613. 

V suAoa rrofr cercqun aepxHttfi qJtsHI,IK crpyttoqKa c verltprur pe6paMu 

u cerqaro-xlrJlrqarofi IloBepxHocruo. CIoAa orHoctrcs: C. sleve' 

niana Rupr., C. tataria Sebeyk, C. grandiflora DC. u C. litwinowii 

K.Cross. Ceruux Tatariae 6lutxa K ceKuur4 Crambe, Ho y BttIloB llocrequeft 

nepxul,rfi qneHur crpyqoqKa 6es pe6ep, c tuaAxofi unu 6yropuaroft 

IloBepxHocrbKl. 

Saxifraga moschata Wulf. var. pontica (Albov) A. Zernov comb. et 

stat. nov. - Saxifraga pontica Albov, 1895, Tp. Tratfr. 6or. caAa l, flpurox. 

I (Mar. @n. Kolx.) :97. 

B orlrnque or f. JI. KyApauronoil (1995), csoAqulefi S. pontica n cu- 

HoHrrMbr x S. moschata, fl. 9uvrrato Bo3MoxHbIM pacreHl4t c KoJtoHoBtrA- 

HbrMr{ rycro o6[ucrseHHuruu uo6eraMu BblAeJltrb B KaqecrBe oco6ofi 

pa3HoBr4AHOCru. 

Saxifraga subverticillata Boiss. subsp. colchica (Albov) A. Zernov 

comb. et stat. nov. --- S. colchica Albov, 1895, Tp. Tu$r.6or. caAa, I, 

npun. I (Mar.

crr,r Koropofi upnlncrurarra 3-5 MM AJI., a uo6ern ouyueubl peAKLIMI{ 

6elrrur,r BoJrocKaMI,I. 

Pacreuuq c KpyrrHbrMr,r [pI4Jlt{crH[KaM?I Ir l'oJIbIMu ttoberanau Bcrpeqarorcr 

rrper4MyxlecrBeHHo n cy6a.rtnnnftcKoM I,t zurr'unicrou Ilotcax, a pac- 

TeHr4g c MelrKHMr,r rrprIJII{crHI{KaMr4 v oly[reHHblMu uo6eraun B lrecHoM 

florce. Ho na6nruAaercr [epeKpbrBaH[e AaHHbIx IIpr{3HaKoB, trro IIo3BoJIteT 

paCCMaTpI'IBaTb 3TH paCTeHI'{q B Kaqeg[Be BbICOI'HLIX IIOABI{AOB. 

Vicia truncatula Fisch. ex Bieb. subsp. balansae (Boiss.) A. Zernov 

comb. nova. V. balansae Boiss.. 1872. Fl. Or.2: 569. V. ctbbreviata 

Fisch. ex Spreng. subsp. halansae (Boiss.) Stankev., 1988, C6opH. Hayq. 

rp. rpuKJL 6or. reH. u cen. I 17 : 89. 

Vicia balansae He Moxer paccMarpnBarbcs KaK IIoABI.{A V. abliret'iuta 

BuAa, orHocrulerocfl n 4pyrofi ceKIrKu. TarcosotuuqecKue orHouevus, 

V. abbreviata u V. truncatula paccvorpeHtt vtuofr pauee (3epuoe, 

2000). 

Vicia tenuifolia Roth subsp. subalpina (Grossh.) A. Zernov comb. et 

stat. nov. V. variabilis Freyn et Sint. ex Freyn var. subalpina Grossh., 

1930,

ostrata [oqerrHble qeuyK reuuo-6ypue, JII4croqKn JIaHIIerHble, couBerl4t 

o6nqHo KucreBu.4Hbre, y ocHoBaHI{s He pa3BerBJreHHne. OAuaro nprn 

BKrroqeHr4r,r F. oxycarpa Willd. var. rostrata (Guss.) K.Koch s F. excelsior 

s raqecrBe pa3HoBI,IAHocru rlpaBuJlbHbtM Ha3BaHI4eu rtocleAuefr, 

cofJracHo crarbe ll.6

Jlnreparypa 

A a u 6 o n H. M. Marepnann 4.nr $nopu KolxuAu // Tp. TuQn. 6or. ca4a. 

1895. T. I. flpnnoxenne |. C. l-287. 

f pocc re fi u A. A. @nopa Kasrasa. M.; Jl., 1952.Vls[,.2.T.5.453 c. 

3 e p H o n A. C. Pacrenlra Cenepo-3anaAHoro 3aranxasrs. M., 2000. 130 c. 

l4wxa H H rr K a s H. H. Celleftcrso Oleaceae no $nope Kasxaga. l. Poarr 

Jasminum u Fraxinus // 6or. xypn. 1998. T. 83, Ne 9. C. 106-115. 

Kylp r ruo B a f. JI. Koscnexraugor ceueftctv Saxifragaceaen Parnassiac:eae 

Qnopsr Kasrasa // Eor. xypn. 1995. T. 80, Ne 8. C. 104-108. 

KysneuoB H.V. Oleaceael/Kyrnduos H. ll., Eyur H. A., @orvrran A. B. 

Flora caucasica critica. Marepnalu 4la rfnopn Kanrasa. IOpree, 1903. 

9acrr 4, sun. l. C.231-269. 

ManpoAres E. B. Rohrbachia - nosui po4cerueficrnaThyphaceae ll 

Eor. xypH. 2001. T. 86, Ne 9. C.120-124. 

fl ecx o ra 14. M. TaxcoHoMr,rr u KapnoJrortrrJrbHrHoK (Linaria Mill.) Bocro.{Hoft 

Enponn n Kaexasa. Auc. ... rasA. 6uol. nayx. M., 2003.219 c. 

XoxprKoB A. n., TlrxoMr,rpoB B. H. O HoBoM poge Platyruscus 

A. Khokhr. et V. Tichom. (Ruscaceae) // Eron. MOI4[. Ora. 6non. 1993. 

T. 98, sun. 4. C.9l-93. 

Candolle A. P. de. Regni vegetabilis systemanaturale. Parisiis, 1821. 

Yol.2. 745 p. 

F r e y n J. F. Plantae novae orientales. Il // Osterr. Bot. Zeitschr; 1892. Bd 42. 

s. 80-84. 

Grierson A. C. J. InulaL. /P. H. Davis. (ed.). Flora of Turkey and the 

East Aegean lslands. Edinburgh, 1975. Vol. 5. P. 54-73. 

Mocrcoscruft neAaroruqecrufi rocyAapcrreuuuft yHzBepcrzrer 

129278, r. MocKna, yn. Ku6anrvr4qa, A. 6, ropn. 5, 

ra$e4pa 6oraHnru 

E-mail: falco@pues.ru

I. A. JlassKon 

G. Lazkov 

o HoBbrx LI PEAKI4X BI4AAX AIIS @JrOPbI KbIP[bI3CTAHA 

DE SPECIEBUS PRO FLORA KYRGYZSTANIA NOVIS ET RARIS 

flpn onpegeJreHlln Marepl{tuloB, co6paHuutx B 3Kc[eAuIrHu Ha reppl4ropr{u 

KmpruscraHa s 2003 r., a raKxe upocMorpe paHee co6paHHoro 

MarepuaJra, xpaHqulefocr e fep6apr,rn Euonoro-tloqBeHHoro l4Hcl'lrryra 

HAH KuprHscrofi Pecny6lnru (FRU), 6ulu o6uapyx(eHbl He IIpHBo- 

AuBuldecfl paHee AJIf, pecny6rulrcn nu4u. ,{na ApyrKX, yxe l43BecrHblx I'I3 

KrrpmcraHa BLrAoB, yKa3aHbI HoBbte MecroHaxox LeHvLr., 3HaqurerbHo 

pacurl,tpsrcque t,tx apeair. Bce qnruponaHHble o6pa:qu [epeAaHbl n fep- 

6apwtrt Eorasuqecxoro rlHcrnryra I,IM. B. JlKoltaposa PAH (LE), ay6- 

JrrrKarbr HaxoAqrcs n fep6apuu Fraonoro-noqBeHHoro nHcruryra HAH 

KP (FRU), AHHoruponaHHrrfr crIHcoK HaxoAoK [pnBoAllrcfl HI{xe. 

|. Delphinium knorringianunz B. Fedtsch. 

HosHft BIIA ArIr 3auaAsoro TsHs-IIIaHq. Elur ussecreu n Kltprlt:rcraHe 

roirbKo c ceBepHoro MaKpocKJIona Alaftcroro xpe6ra (flaxorraona, 

1972). Hauu HafiAeH creAylottruft o6paseu:

4. Trollius komarovii Pachom. 

Honufi BUA Arur flcreucxoro xpe6ra:

AepHeHHat, rIoJlblHHo-Mf,TJrItKOBat pa3HoTpaB., IIBeT UBeTKa po3ol 

cupeueeuft, 11 V 1963, V6yxeena, fop6yuona>' 

PacreHur c TyprecraucKofo tr Alaficroro xpe6ron HecKoJrbKo O'IJUI- . 

qaroTcr oT Trrrrr4qHoro A. aulieatensis 3-5, a ue 2 3 uaparuu [I,tcTOtIKoB 

u 6Onee CLIJTbHbtM OApeBeCHeHlIeM H, BO3MOXHO, OTHOC'TC' r 6.1USxOfr 

K HeMy pace. .{nr BbucHeHI4s 3Toro Bollpoca rpe6yrcrcs AzurluefiUlue 

UCCJIEII.OBAHHfl. 

8. A. compositus Pavl. 

Honufi Br4r, Nrfl$lopu Krtprnscrana: . 

Pauee sHA 6HJr H3BecreH ToJIbKo I43 TI{rIoBoro MecroHaxoxAerrfl.Ha 

Ta"raccropr xpe6re n Kupruscr

13. Phlomoides lanatiJblia Machmedov 

6u'n ugeecreH ToJrbKo c roxnoft oKoHeqHocru @eprancroro xpe6ra 

(Maxuegon, 1986). Hauu safiAeH n cpegnefi efo qacrr,r: (yuleJrbe peKH 

Ypyu6aur, BAoJrb Aoporr.r Ha rrepeBaJr, l0 VI 2003, f. A. Jlarrroe. 

H. B. Kenxe6aesa)). 

14. Echinops tschimganicus B. Fedtsch. 

flepaaa HaxoAKa Br4ra Ha Cerepuorr,r TrHl-[IIaue. paHee sraA 6trr 

rr3BecreH ToJrbKo na 3anaAHo|r TrHr-lrlase u xpafiHefi socroqHofi roqrofi 

ero pac[pocrpaHeH[r rBJrrJrocb yuleJrbe peru Kapa-Ap.ra, Ha rox- 

HoM MaKpocKJroHe Knprurcxoro xpe6ra (HuxnrraHa, 1965). Hauu safr_ 

AeH ropa3Ao BocroqHee:

K o sarr e B c K as C. C. Poa Hepatica Mill. - fle.reHo.Ittnqa // Onpe.uelnreJrb 

pacrenr.Ifi Cpeanefi Asuu. Taurenr, 1972. T. 3. C. 193. 

Jlagrros f. A. Hosrrft nn4 u $nopucrl,rqecKlte HaxoAKtt s cetreficrse 

Asteraceae nr Knprnrun // Eor. xypn. 1999. T. 84. Ns 5. C.125-126. 

MaxueAoB A M. HosHftsLIApoAa Phlomoides Moench (Lamiaceae)ut 

CpeAneft Asnn ll Hosocrn crcreMarnKlt Bbtctutlx pacrenufi. JI., 1986. T. 23. 

c. l l5-l16. 

Hnrnrr.rHa E. B. Pon Astragalus L. -Acrparan//

Vl. O. Eysynona l. Buzunova 

TI4fTOBbIE OEpA3rIbr TAKCOHOB CEMEIZCTBA ROSACEAE, 

ofItICAHHbIX I43 KI4TA.fl, XPAHtt{UECfl B rEp6Apnn 

EOTAHHIIECKOIO ITHCTHTYTA LIM. B. JI. KOMAPOBA 

(LE). r. rroAcEMErlcrso sprRAEorDEAE AGARDH 

SPECIMINA TYPICA TAXORUM FAMILIAE ROSACEAE 

E CHINA DESCRIPTORUM IN HERBARIO INSTITUTI 

BOTANICI NOMTNE V. L. KOMAROVTT (LE) CONSERVATA. 

I. SUBFAMILIA SPIRAEOIDEAE AGARDH 

B Hacrosulefi crarre uy6luxyrcrcr cBeAeHur o xpaHfluruxca n fep- 

6apww. EoraHnqecroro nHcrr{ryra nu.B. JI. Korraapona (LE) rr{rroBbtx 

o6pasqax 20 raxcoHos us 3 po4oo uogcemefrcrna Spiraeoideae cevertcrsa 

Rosaceae, ouucaHHbrx c reppuropr,ru Kurar. 

OcHony KoJrJreKrIrl4 cocraBJuror c6opur Bn.urcoua (E. H.Wilson) r,r 

feHplr (A. Henry) ns I{eurpalbHofo Knrar. fep6apuue o6pa:{tr Bo- 

TaHuqecKoro HHcrrrryra rBJrqtorcfl, s 6orlurrlucrBe cnyqaeB, Ay6leralul 

c6opon, xpaHquruxcr r Apuolr4 ap6operyue, fapnapacrufi yuunepcurer, 

CIIIA (A) ra JlouaoHe (K). Mn ornocnu lrx K KareropLrv

Paratypus: (LE). 

flo nporonory:

flo nporonory: . 

10. S. dasyantha Bunge, 1833, Enum. Pl. China Bor. : 23. 

Syntypi: Cea. Kuraft,

cKr{M crroco6oM 3ruxerKofi 

r,r HaArrucblo ua Heft, cAenannofi pyxofi Maxculaosnua . 

16. S. myrtillodes Rehder, 1913, in Sarg., Pl, Wilson. 1,3 :440, 

Syntypi: Kuraft, CuvyaHb, ((Western Szechuan, Vl 1908, [fl.], N 989, E. H. Wilson>; 

(ibid., lX 1908, [fr,], N 989, id.>; Kr.rraft, Xy6rft, - (LE). 


17. S. prostrataMaxim.,lST9, Tp. flerep6. 6or. caAa 6, I : 184, nom 

illeg., non Schur, 1866. (= Spiraeapjassetzkii Buzunova,2004, Hosocrn 

crcr. Bbrcur. pacr. 36 :284). 

237

Lectotypus (Buzunova, hoc loco): Knraft, [IIeHlcn, 

(LE). 

Syntypi: Knrafi, fanucy, ; ; ; Kurafi, llleurur, ; Kurafi, Xy6rft, - (LE). 


18. S. pubescensTurcz., 1832, Bull. Soc. Nat. Moscou, 5 : 190. 

Syntypi (4 nucra): Klrrafi, Xr6rft, (ln montosis lapidosis China borealis, 1831, Ui.l, 

Kuznetzov> (LE). 

flo nporonoly: . 

Lectotypus:KW(?). 

19. S. sublobata Hand.-Mazz., 1933, Symb. Syn. 7 : 451. 

I s o t y p u s : Kuraft, Cn.ryaHt, (LE). 

flo nporoaory:

H. H. LluxauuunaR 

N.Imchanitzkaia 

TIIILIOHKAIII{fl HA3BAHI4IZ TAKCOHOB CEMEIZCTBA 

RUBIACEAE EPA3LIJII{I4, COEPAHHbIX O. CEJIJIO 

14 OTIIICAHHbIX A. IITAMHCCO U A. IITJIEXTEHAAJIEM. 

I. TIOACEMEIZCTBO RUBIOIDEAE: COCCOCYPSELUM 

P. BROWNE, FARAMEA AUBL. I{ RUDGEA SALISB. 

TYPIFICATIO NOMINUM TAXORUM E FAMILIA RUBIACEAE 

BRASILIAE A F. SELLOWIO COLLECTORUM ET 

A. CHAMISSO ET D. SCHLECHTENDALIO DESCRIPTORUM. 

I. SUBFAMILIA RUBIOIDEAE: COCCOCYPSELUM 

P. BROWNE, FARAMEA AUBL. ET RUDGEA SALISB. 

Hauu upogoJrxeHo Haqaroe paHee (I,lrraxaHl4qKat, 1996 a r) usyve- 

HHe xpaHrulefrcfl B fep6apuw Borauu'recrofo I,IHc't[Tyra uM. B. JI. Ko- 

MapoBa PAH (LE) IIeHHefiuefi fep6apHofi KoTJIeKIII4I{ A. lll3v6."o 

(). OHa nrtrcqaer B ce6f, rep6apHlte o6pasult 6oLtlruoro 

rrHcra raKcoHoB ceMeficrBa Rubiaceae (l0l rnA I{3 38 poAoB), n rorra 

rtl{cJre rr.rrrbr 59 slaAon, orll4caHHblx fllauncco, a raKxe Lllaur'rcco coBMecrHo 

c,{. IlftrexreHAaJreM B x(ypHaJle IIo c6opaM HeMeqKoI'o 

nyreurecreeHHL4Kav KoJUteKTopa

B .qaHHoft crarbe [puBeAeHbI cBeAeHHt o rltrloBblx uarepualax l8 

BrrAoB ceMeficrBa Rubisceae (uo4ceueftcrso Ruhioideae) uz poAoB 

Coccocypselum P. Browne (rpu6a Coccocypseleae Bremek')' Faramea 

Aubl. (rpn6a Coussareeae Hook. f ') u Rudgea Salisb. (rpra6a P'sychotrieae 

A. Rich. ex Dumort.), Kolopble [peAcraBJleHlt 30 xpanrquMucs 

s LE runosutrtn o6pasqarvru. 16 I43 3Tl4x BltAoB 6ltltu onncanul lllauucco 

ra IIInexreHAaJreM (Chamisso, Schlechtendal, 1829a, 1829b)' a raKxe 

IIIsNausss (Chamisso, 1834) uo rep6apurtu 3K3eMIIJIrpaM, co6paHHbIM 

Cerrro e 18l4-1829 rr. n Epa:ulrr4v, a 2 sv1a (Schwenkf'eldia aurea 

spreng. = coccocypselum aureum. (Spreng.) Cham. u Mussaenda coriocea 

Spreng. = Rudgea coriacea (Spreng.) Cham.; cM. Aaree) ory6Jfl{KoBaHbr ' 

Illnpenrelervr (Sprengel, 1820, l82l)' 

Alr xaxAoro raKcoHa rlpl,tBoAflTcg JlarsHcKoe Ha3BaHI'Ie c HoMeHK- 

JraTypHoft rII4TaTOfr, Ha3BaHUe [puHtToe B HacTottqee BpeMrI' KaTeropl4fl 

T:aIA U KOIUTIeCTBO fep6apHblx JII4CTOB, IIOJIHLIfi 'IeKCT Opl4fl{HanbHoil 

rep6apHofr 3TnKerKu n LE (aonolHeHut, Heo6xoAHMbIe AJII IIoHI{MaHufl 

TeKCTa, 3aKJrIOqeHbI B KBaAparHbIe cKo6KI4), UHTara I{3 llporoJlofa' a 

TaKXe rpuMeqaHl4e, Kacarculeecfl Aar v MecT c6opa o6pa3l-loB, HoMeHK- 

Jrarypbl r4Jrr.{ TaKcoHo MAV, A TaKXe reorpa(bl4 qecKo f o pacl IpocTpaHeHl4fl 

BT4AOB H T. A. POAII I,l BI4AbI B npeAeJlax poAoB pacflonoxeHu r alQa- 

BHTHOM rropflAKe. A.rts raxAot'o poAa yKa3aHa tpn6a B coorBercrBl4n'c 

KJraccr4onKatll4eft ceMefr cr Ba Rub iac e a e, ilpe[JroxeHHoft Po66pexron 

(Robbrecht, 1988). 

Fam. Rubiaceae 

Subfam. Rubioideae 

L Coccocypselum P. Browne, nom. cons. 

l. Coccocypselum aureum (Spreng.) Cham' et Schltdl., 1829, Linnaea 

4,2:139; Steyerm., 1967, Mem. New York Bot. Gard. 17, I :306 (var. 

aureum); DC., 1830, Prodr. 4:397; K. Schum., 1889, in Mart., Fl. Bras. 

6,6:312. -- Schwenklbldia aurea Spreng., 1820, Neue Entd' I : 280; id', 

1825, Syst, Veg., ed. 16,1 :764. 

Ouucan u3 Epa3urura. Neotypus (Imchanitzkaja, hoc loco): 

(LE!)' 

flo uporolory: . 

flplrue.raHue. IIlllpsHls.lrl (Sprengel, 1820, l. c.) I uporo.uol-e oIIucaH- 

Horo rrM us Epasurtuu no cOopau Certno su1a SchwenkJbldia aurea He 

yKa3ar runa. Illnvscco I4 LLIrexreHAaJIb (Chamisso, Schlechtendal, l. c.) 

orHecJrr,r ouncaHnmfi llltrpeureleu BuA K poAy Coccocypselum. OHu sn\e- 

Jru rnnoBofi Marepr{aJl S. aurea, cyAt uo [oMerKe (

Spr. N. Entd. 1. p. 280!)) npn o6napoAoBaHrltt xou6nuaunu C. aureum, 

a raKxe yKa3aHua Mfl B:aAa: ); rlocregnr,rft, 

xpaunBurnfict n B n fep6apnu lUupeHteu, 6tul yrpaqeH Bo Bper"rx 

Bropoft utrporofi BofiHbr. Illyrvrau (Schumann, l. c.) cnycra 60 tler so 

selum (C. pedunculare u C. pulchellum; cM [anee) e Caurr-flerep6ypr 

K. A. Tpunaycy; c [ocJreAHr,rM oH BeJr aKrr{BHyro fleperrr4cry u o6ueuunatct 

gy6nuKaraMr.r pacreunfi (I,IuxaHnqra41996a). HasnaHue poAa Ha 

3TI4KCTKE HCOTUTIA, B OI.JII4.IHC OT IIPOTOJIOTA, IPI,IBCACHO IIIANAKCCO 6CS 

BOrrpocLITeIbHofo 3HaKa, rrocKoJrbKy, 6narogapx HzulHqr.rrc rIBeTKa, Irpn_ 

HaArrexHocrl o6pasqa K poAy Coccocypselum iln, Hefo crana oqeBuAuofi. 

OrcyrcrByer raKxe [oMerKa (unra , ), corrpo- 

BOXAaIOUIaq Ha3BaHLre HoBOro TaKCoHa, OrruCaHHorO lllnNausss n IIIIex- 

TeHAaneM, KaK B opr,rrr,rualrsofi uy6luxauun, TaK 14 ua :ruxerre. ne_ 

KaHAoJrb (Candolle, l. c.) ue Br,rAen rr4rroBoro o6parqa C.? erythrocephalum. 

flocroltry Becb rrepBoHaqallnrrff Marepr{an, Ha Ko.ropoM 

ocHoBaHo outacaH]re BHAa, B B 6lrn yrpaqeH Bo BpeMfl Bropofi unponoit 

BoftHbr, xpauntluitcx r LE rep6apHuft ox:eururp C. erythrocephalunt 

o6osua.{eH HaMr.{ KaK Heorr'fi 3Toro Br{Aa. Illyrraan (Schumann, l. C. : 3lg) 

no rrpuBeJr 1trx C. erythrocephalum o6pau1br, co- 

6paHHue Ceruo a (), a 

TaK)r(e (rrpeArroJroxr,rreJrbHo)) n

Bartl. ex DC. (1830, l. c.: 396) (= C.nummulariaeiblium auct, mult.' non 

C. nummularifulium Cham. et Schltdl. 1829). C. nummulari/bliumCham. et 

Schltdl., B rroHIrMaHI4u Crefieprvrapra (Steyermark, l. c. : 305), - caMocrosrerrrHrrft 

6pasnltcxufi BI,IA; oH orr[qaerct or C. hirsutum v or C. guianense 

(Aubl.) K. Schum. (1889, in Mart., Fl. Bras. 6, 6 : 315; Beuecyua, 

fafrana, Cypuuau, fruaHa, o-n Tpuuugag), c roropuuu 6oraHI'IKI4 llpex- 

Ae efo cMerur4Biull4, tlOprrlofi JIucToBbIx rIJIacrnHoK n 6onee KoporKllMl4 

qepeurKaMr,r. lllyrraau (Schumann, L c. :276,315) ao > 

orHec (C. nummularidblium>> Cham. et Schltdl. B crIHoHI{Mbl AByx BlrAoB - 

p. max. p. (incl. typo) r Lipostoma capitatum (Graham) D' Don (= C. campanulillorum 

(Hook) Cham. et Schltdl. fide DC., 1830, l. c.:673), 

p. min. p.) - rc C. guianense.Por,LipostomaD.Don He Irpu3HaH 6o'tasu- 

KaMu r4 orHeceH uMIr B cIrHoHuuw CoccocypselumP. Browne. 

C6opu Cenlo, [pnBeAeHHbIe lllyrraanou AJII yKa3aHHbIx Bblure Blr- 

AoB, rrpor4cxoAflT u3 llrraroB Puo-Ae-Xasefrpo, 3cunpury-Caury u Muuac-Xepafic 

u 6autu cAeJlaubr rIM Bo BpeMt nepBoro (1814 1815 rr.), 

Broporo (1816 r.) I4 rperbero (1818-1819 rr.) nyreurecrsptfi rto Epasrl- 

:ruu (cu. Herter, 1945). 

4. C. pedunculare Cham. et Schltdl., 1829, Linnaea 4,2 : I42; DC., 

1830, Prodr, 4:397; K. Schum., 1889, in Mart., Fl. Bras. 6,6:312. 

Onucan u3 Epa3rnulr. Lect o ty p u s (Imchanitzkaja, hocloco): 

Isolecto typus:

Oparueur -- G - DC, n. v. (microfiche4:497.6, card N 710!) 

( (LE!). 

Oparvreur -- G DC, n. v. (microfiche 4: 497. 7, card N 710!) 

(

Berolinensi (sub nom. F. monteviden.rr"s DC.)> (LE!); uocreAHuil He uueer 

orHorrreHr{.N K TrrrroBoMy Marepr4iury T. montevidenss, frocKoJrbKy 3Tor 

Br{A, corJracHo [poloJrory, orrr4caH no rep6apHnM sK3eMrrJupaM c rrJro- 

"qaMra ((Specimina nostra fructifera ...))). K roMy xe, cyAfl rro HoMepy Ha 

3rr4KerKe, yra:anunfi arrure o6paleq 6un co6paH Cel:ro s 1829 r'. no 

BpeMr [rroro rryrerxecrBur rro EparnJlr{r,l B [rrare Cau-flayly (cu. 

Herter, 1945 : 134) u, oqeB[AHo, rrpr4cJraH s B nocne o6Hapo4onauua 

T. montevidense. 

3. T. stipulaceum Cham. et Schltdl., 1829, Linnaea 4, | : 31. 1= porameq 

stipulacea (Cham. et Schltdl.) DC., 1830, Prodr. 4 : 497; Miill. 

Arg., 1875, Flora (Regensb.) 58, 30:472; id., 1881, in Mart., Fl. Bras. 

6,5 : 138). 

OrrucaH n: Epasulrln. Neotypus (lmchanitzkaja, hoc loco): 

(LE!) (Typus - B, Hb. 

Chamisso, yrpa.reu). 

flo nporonory:

6uuauuro F. monteviden.rf.r (Cham. et Schltdl.) DC', otty6;tuKoBaHHylo K 

roMy BpeMeHn Art srolo BrAa !,erau4oLteu (Candolle, 1830' l. c.) B 

. 9ro xe Kacaerct AByx Apyrl{x BI4AoB Tetramerium 

(7. latifulium w T. stipulaceum; cM. Bbl111e), ollucaHHblx lllaNancco I'I 

I I InexreHAa-rreM oAHoBpeMeHHo c T. rn ont e v idens e (Chamisso, Schlechtendal, 

1829a, L c.), ro oHI{, KaK I,I KoM6I4Haquu F. latiJblia (Cham' et 

Schltdl.) DC. u F. stipulacea (Cham. et Schltdl.) DC. (Candolle, l. c.), 

lllauncco He 6rrnn yrloMtHyrbl. YrasaHHHe Bbl[re BHAbI Fqramea oluecenu 

,Lf,erauAoJreM (Candolle, l.c.: 496) r > poga 

Faramea. 

4. Faramea marginata Cham., 1834, Linnaea 9,2 : 221 (

noftHrr, HaMr,r B xa.recrBe Heornna o6o:Ha.reH o6paseu C. gardenioides 

noA N 270, xpauntluilcs, s LE u uolnocrbro coorBercrayrculnfr lrporo- 

Jlofy; rrocnegunfr , no-nnAr.rMoMy, rBJrtercf, Ay6[r.rKaroM cl4HTt4rra froro 

BLr4.a, xpaHnBtrrerocr n B n yrpaveHHoro Bo BpeMr noftHr,r. Mrcuep 

(Mtiller, 1881, l. c. : 207) He Br4Aer runa C. gardenioides. Bo r oH cocnancr Jrr4rrrb Ha rrpororror fror.o Br,rAa (). cyAr uo HoMepy Ha 3rlrKerre Heor.urra, 

sTor Br4A uor 6rrrl co6paH Cel.uo s l8l5-1817 ru. Bo BpeMr ero B.roporo 

rryrerrrecrBr{r uo Epa:unuu (crra. Herter, 1945). 

2. C. jasminoides Cham., 1834, Linnaea 9,2:222, non Welw. ex 

Hiern, 1898. 1= Rudgea jasminoides (Cham.) Miill. Arg., 1876, Flora 

(Regensb.) 59,29: 452;id,.,1881, in Mart., Fl. Bras.6,5: l9l; L. B. Sm. 

et Downs, 1956, SellowiaT :62, fig. 20, f i). 

Orrncau uz Bpaznnwu. L e c t o t y p u s (Imchanitzkaja,hoc loco): 

novum c TeM xe caMbIM Tr,r[oM, KaK I{ y He3aKOHHO|O HA3BAHVfl (CTaTbt 

58. I ) (LE!). 

flo lpotonot'y: 

flpuvreuaune. Mrclulep (Mtiller, 1881, l. c') He an4ert 

'twa C. nutjor. 

Bo oH cocraJrcf, Ha flporonol']roro srAa ((Habitat 

: Sello>). 

5. C- minor Cham., 1834, Linnaea 9,2: 227 . (= Rudgea minor (Cham.) 

Standl., 1936, Publ. Field Colimb. Mus. ll, 5 : 264; Miill. Arg., l88l' in 

Mart., Fl. Bras. 6, 5 : 188 et 189, in

6url ucnonb3oBaH (

y3KOJIr{CTHOfi rfOprr4Ofi C KOpOTKIIMII TbI!{IZHKaMU, BK[IOrreHHblMn n rpy6- 

Ky BeHqr4Ka. OAHaro IITsMugss coo6xlul B tlporoJrofe raKxe 06 I43y'{eH- 

HoM r.rM eAnHcrBeHHoM sK3eMIIJItpe y3KoJlucrHofi xoporxocro.lr6uxoaoii 

Sopnu C. parquioides c BKrIoqeHHbIM B rpybry BeHqtrKa croJr6uKoM 

(

.ren rep6apublft ?K3eMrrtp R. parvfolla Sello N B 1779 c l2l5; rrocJre- 

4unfi, no-nnAr,rMoMy, Ay6[nKar cr.rHTrrrra 3Toro Br4Aa, yrpavenHoro r B, 

rroJrHocrbrc) coorBercrByer rrporoJrory. Mtonlep ((Mtiller, 1881, l. c.) ne 

Br.rAeJr rvrna C. parviJblia. OH npueel lrnurb rrporoJror 3roro BHAa 

(: Sello, ...>). Cym uo HoMepy Ha 

3rrKerKe Heorr4na, oH 6brn co6pau Ceuo s 1819-1820 rr'. Bo BpeMr rpe- 

Tbelo rryreurecrBr4f, no Eparulun B rrrrare CaH-llayly (crra. Herter, 

le4s). 

8. C. viburnoides Cham., 1834, Linnaea 9,2 :228. (= Rudgea 

viburnoides (Cham.) Benth., 1850, Linnaea23,4:458; Miill. Arg., 1876, 

Flora (Regensb.) 59, 29 :454; id., 1881, in Mart., Fl. Bras, 6,5 :204, tab. 

27). 

Onucau us Eparululr. Neotypus (Imchanitzkaja, hoc loco): 

(LE!) (Typus -- B, Hb. 

Chamisso, yrpa.reH). 

flo nporolory:

Ouncasue BuAa ocHoBaHo Ha rep6apuux 3K3eMIIJIspax c uBerKaMI4 14 

rrJroAaMH. flo uporonory: . Turosoil 

Marepr4uu yKasaHHblx Bblue BHAOB co.ffea 6ul yrpa'reu, 3a LISKJIIOqeHUeM 

eArrHcrBeHHoI'o o6pa3ua C. nodosa, coI'JIacHo yKa3aHI4IO Eesraua 

(Bentham, L c. :457), xpaHtlqeroct r K (); llocJleAHnft, Ay6rrKar cuHrl{rla 

C. nodosa, yrpaqeHHoro B B, o6o3Ha'IeH HaMu KaK Heorurl (N e o t y - 

p u s (Imchanitzkaja, hoc loco): (K, herb. Hook.))' 

Bu;rll CoJJbs luzoniensis Cham. et Schltdl. (>) u C. ma' 

riniana Cham. et Schltdl. (Chamisso, Schlechtendal, 1829a: 32; 35), ottr'tcaHHbre 

lllauucco ll IlllexresAareM rlo c6opaM lllauucco n I4' O. ?rurrroJrbua 

Ha o-Bax Jlycon ((Du:rnunuHcKue o-Ba) ra Oaxy (faraftcxue o-na) 

B KpyrocBerHoM Iryrerrecrnura (1815-1818 rr.) Ha 6pl4re no4 

KoMaHAoBaHueu O. E. Koqe6y (I4rraxaHnIlxaq, 1996n : 20), npuuarlexar 

K pogy Psychotria (P. luzoniensls (Cham. et Schltdl.) Fern.-Vill. v P. ma' 

riniana (Cham. et Schltdl.) Fosberg). 

Hlrxe upune4eH 6pa:untcKuft BI.IA Rudgea, ueorunurfuquponaHHltil 

HaMr{ repoapHbrMl{ 3K3eMrIJItpaM[ I,I3 KOJIJIeKUI4II4 I-LIaMucco, xpaurqefrca 

n LE. 

Rudgea coriacea (Spreng.) K. Schum., 1891, in Engl. u. Prantl, Nat. 

Pflanzenfam. 4, 4 '. 116; L. B. Sm, et Downs, 1956, Sellowia 7: 61, frg. 

20, d. -- Mussaenda coriacea Spreng., 1821, Neue Entd. 2 : 145. -- 

Rondeletia coriacea (Spreng.) Spreng., 1825, Syst. Veg., ed. 16, | : 707 

(

flpuuevaHue. BuA M ussaenda coriacea orrucau I I InpeHrenerr,a (Sprengel, 

1821, l. c.) no sK3eMturrpy c rrBerKaMr4, co6pauHorr,ry n Eparr,r.nnu, 

6e: yr

Candolle A. P.de. Rubiaceae ll Ptodromus systematis naturalis regni 

vegetabilis. Parisiis, I 830. Pars 4. P. 341-622, 672-67 3. 

C h a m i s s o A. Spicilegium plantarum e familiis jam prius recensitis praesertim 

Brasiliensium. Rubiaceae ll Linnaea. 1834. Bd 9, H. 2. S. 214-261. 

Chamisso A., Schlechtendal D. F'. L. De plantis in expeditione 

speculatoria Romanzofftana obsevatis. Rubiaceae. Machaoniu. - Tribus lV. 

Psychotriaceue ll Linnaea.1829a. Bd 4, H. l. S. l-36. 

Chamisso A., Schlechtendal D. F. L. De plantis in expeditione 

speculatoria Romanzoffiana obsevatis. Rubiaceae ll Linnaea. 1829s. Bd 4' 

H.2. S. t29-202. 

H e r t e r W. Auf den Spuren der Naturforscher Sellow und Saint-Hilatre ll 

Bot. Jahrb. 1945. Bd 74, H. L S. I 19-149. 

H i e p k o P. The collections of the Botanical Museum Berlin-Dahlem (B) and 

their history // Englera. 1987. N 7. P.219-252. 

L o r e n c e D. H. A nomenclator of Mexican and Central American Rubiuceue 

// Monogr. Syst. Bot. Missouri Bot. Gard. 1999. Vol. 73. P.l-177. 

M til le r J., A rgo v ien s i s Rubiaceae // Martius C. I'-. Ph. Flora Brasiliensis. 

Monachii, 1881. Vol. 6, pt 5. P. l-470. 

Robbrecht E. Tropical woody Rubiaceae. Characteristic features and 

progressions. Contribution to a new subfamilial classification // Opera Bot. 

Belg. 1988. Vol. 1. P. l-271. 

Schlechtendal D. F-. L. De plantis Mexicanis a G. Schiede M. Dt'e' 

collectis ll Linnaea. 1835. Bd 9, H. 5. S. 589-610. 

Sch lech tend al D. F. L., Chami ss o A. Plantarum Mexicauarum a 

cel..viris Schiede et Deppe collectarum recensio brevis // Linnaea. 1830' Bd 

5, H. l.5.72-203. 

Schlech tendal D. F. L., Chamisso A..Plantarum Mexicanaruma 

cel. viris Schiede et Deppe collectarum recensio brevis. Addenda ll Linnaea. 

r831. Bd 6, H. 3. S. 385-430. 

S ch u m ann K. Rubiaceae // Martius C. F. Ph. Flora Brasiliensis. Monachii, 

1889. Vol. 6, pt 6. P.125442. 

Smith L. B., Downs R. J. Resumo preliminar das Rubiriceas de Santa 

Catarina // Sellowia. 1956. Anos 7-8. N 7. P. l3-86. 

Sprengel K. Neue Entdeckungen im Eanzen Umfang der Pflanzenkunde. 

Leipzig,1820. Bd 1. 452 S.; 1821. Bd 2. 363 S. 

Steyermark J. A. Rubiaceae // B. Maguire (ed.). The botany of the 

Guayana Highland - Part VII // Mem. New York Bot. Gard. 1967 . Y ol. 17 , 

N t. P.230-436. 

U r b a n L Friedrich Sellow (1789-1831) / Vitae itineraque collectorum botanicorum 

notae collaboratorum biographicae, florae Brasiliensis ratio edendi 

chronologica... // Martius C. F-. Ph. Flora Brasiliensis. Monachii, 1906. 

Vol. l, pt l. P. 105-lll. 

U r b a n l. Biographische Skizzen. L Friedrich Sellow (1789-18-1 l) tl Bot. 

Jahrb. 1893. Bd 17, H. l-2. S. 177-198. 

EoraHuqecruft uHcruryr nu. B. JI. Korraapona 

PAH 

197 37 6, C.-flerep6ypr, yr. IIpo$. flonoea, 2

M. C. Honoci!.nona, 

IO. P. Pocxon 

M. Novosselova, 

Yu. Roskov 

Tr4noBbrE o6pA3rlbr TAKCOHOB CEMETICTBA FABACEAE 

KI4TAfl, XPAHqIUUECfl B fEpBApI4I,I EOTAHI4TTECKOTO 

r4HCTtrTyTA trM. B. JI. KOMAPOBA (LE). rr 

SPECIMINA TYPICA TAXORUM E FAMILIA FABACEAE 

CHINAE IN HERBARIO INSTITUTI BOTANICI NOMINE 

v. L. KOMAROVII (LE) CONSERVATA. rr 

Hacrorulaa crarbfl coAepxr4T pe3yJrbrarbt AallHefturelo t4ccre/loBa- 

HHr rlutoBbrx MarepuaJros ceveftcraa Fahaceae (po4rr Campylotropis, 

C aragana, C olut ea, Gue ldens taedtia, H edysarum, Lespe deza, T'he rm op,s is, 

Vicia) uz Kuraq, xpaHrult4xcfl n fep6apr.rn EoraHn.recxoro uHcruTyra 

nu. B. JL Kouapoaa PAH (LE). .{anuue r43JroxeHH B c;reAyrouleM lroprAKe: 

HoMeHKJrarypHar rlr,rrara rulnQraqrapyeMor.o Ha3BaHr4r, rrptrHrroe 

B HaCTOgUIee BpeMr Ha3BaHr4e, yKa3aHr,re KaTefopt4r4 Tr4[a, KoluqecTBa 

JrrrcroB (eclu ux 6olrure oqHoro) H KpylrHolo pernoHa c6opa rr4rroBoro 

o6pa3rla, TeKcr orr4KerKu rr,rrroBoro o6pa:qa (n xanu.rxax), qurara 

locus classicus t43 rrporoJtofa, rrpr4Mer{aH[r. 

B page cryqaeB o6pa:qu, lruKerKrr Koropbrx noJrHocrbro coBnaAa- 

IOT C eAr4HCTBeHHbrM rrpOUr4TnpOBaHHbrM B rrpoToJtofe lK3eMrrJrrpoM. 

orHeceHbr K Karefoptrr4 crrHTr4rra, a He u3orlrfla. 3To o6lflcr.lqercq ret4, 

qro Mbr He r{MeeM cneAeuufr o KoJurqecrBe rep6apHux Jrr{croB (.rry6Lre- 

ToB), Bo3MoxHo xpauruluxcr B ApyrKX rep6apnax, a rar)r(e He 3HaeM, 

6ulru nr.r rarue-lu6o H3 STtlx 3K3eMtrJrflpos o6o:Ha.reHbr aBTopaMn Br.t- 

TOB KAK I'OJ]OTI,ITIbI. 

Bce uepe.rucneHHbre B crarbe rep6apuue lK3eM[Jlfl pbr orcxaHr4poBaHbr 

B coorBercrBlr[ c MeroAr4Kofi co3AaHr.rr Bl4pryirnbHor-o rep6apua (Pocxon, 

1997). I{ut[ponre nso6paxeHr{r rr,trroB Ha KoMr]aKr-Alrcre AocryrrHbr rro 

crequrlrrbHoMy 3arrpocy, aApecoBaHHorray n fep6apufi EI,IH PAH. 

I,Icc:reAosaHr4e BbrrroJrH eHo rrpn ltoAAepxxe Poccniicroro @ ou4a $yH - 

AaMeHrarrbHbtx plccrreAosaHuit (npoeHru N! 02-04-48479 n J\b 00-07- 

90271); aBTopaM TaKxe rrpr4qrHo Bbrpa3lrrb cBoro r4cKpeHHrcro 6narogapsocu 

(DouAy co4efrcrnuf, oreqecrBeHuofi Hayre. 

l. Campylotropis argentea Schindl., 1912, Feddes Repert. ll :426. 

Lectotypus (Roskov, hoc loco): IOro-3anaAntrfi Kuraft, 

(LE). 

flo lporo,rory: . 

2. C. gracilis Ricker, 1946, Journ. Wash. Acad. Sci. 36 : 38 (= C. mscrocarpa 

(Bunge) Rehd. frde P. Y. Fu, 1995, in Fl. Reip. pop. Sin. 4l ll4). 

256

Isotypus: I{eHrpalbHhlft Kurafi,

flo nporoaory: > (LE). 

Syntypi: IOro-3an. Knrafi, r. 

13. C. kozlolfitil(sst., 1909, Tp. flerep6. 6or. caAa 29,2:283, tab. 

ll, fig. A. 

Holotypus: IOro-3au. Kurafi,

B rperquHax cKaJr, pocr go 2 apurun, 5 IX 1900, [veg. et fr.], N 526, 

B. (D. JlaAurzu> (LE). 

flo uporonory:

Syntypus: Ceu.-3an. Kuraft, (LE). 

flo nporonory: < Specimina vidi: l. Potanin, l8 Vl 1884, Chinae borealis prov. 

schansi, in rupibus angust. meridionalium ad portas yen-men-kuan montium Mao-touschan. 

2. [id.] l3 vl 1884, schansi, in parte superiore faucis angust. septentrionalium 

montis Utaischan in rupibus apertis siccis>. 

flpnue.raHne. Ha erurerKe cuHTr.rrra Nb 2 nueercr AorronHr4TeJrbHat 

HaArrr{cb, cAeJraHHaq, uo sceft BepoflrHocrr.r, pyroft B. JI. Korrlapona 

(13 VI U-tai-scharu>. 

17. C. rosea Turcz. ex Kom., 1909, Tp. flerep6. 6or. caAa 29,2:205, 

tab. 5, fig.C. 

Syntypi (2 nucra)'. Ces. Ku.rai, (LE). 

Syntypi: Kraraft, Bnyrp. Mourolur, > (LE); Ces. Knrafi, ((montes 

Pohua-shan prope Pekin, 1874, [fr.], N 14, Dr. Bretschneider> (LE); Cen.-3an. K'rrafi, 

(LE); Cen.-3an. Kr.rrafi, (LE); Ces. 

Karaft, (LE). 

flo nporonory:

flo nporonory: ,. 

19. Gueldenstaedtiamaritima Maxim., 1879, Bull. Soc. Nat. Moscou 

54. 1 :7. 

Lectotypus (Roskov, hoc loco): Bocr. Krlraft,

Syntypus: Bocr. Kurafi, (LE). 

flo nporono.y:

M. C. Honoc€.nona, 

A. K. Csrrnn 

M. Novosselova, 

A. Svtin 

TIrrroBbIE oEPA3Ubr TAKCOHOB POAOB ASTRAGALUS L. 

IT OXYTROPIS DC. (FABACEAE) KI4TAfl LI MOHTOJII{4 

xPAHfITI[ECfl B TEPEAPI{n EOTAHUqECKOIO 

IIHCTIITYTA IIM. B. JI. KOMAPOBA (LE) 

SPECIMINA TYPICA TAXORUM E GENERIBUS ASTRAGALUS 

L. ET OXYTROPIS DC. (FABACEAE) CHINAE ET MONGOLIAE 

IN HERBARIO INSTITUTI BOTANICI NOMINE 

v. L. KOMAROVI (LE) CONSERVATA 

B Hacrosqeft crarre orpaxeHbl pe3yJlbrarbl I'IccJIeAoBaHIlt rHrIoBbIx 

Mal'epuaJloB ceMeftcTBa Fabaceae I,I3 Kurat n Monrolul't, xpaHflIquxct B 

fep6apun Eoranu.recroro l4Hcrnryra IrM. B. JL Korraapona PAH (LE)' 

.I[aHHue l43noxeHbr B cJIeAyIoqeM lopflAKe: HoMeHKJIarypHafl ul4Tara ruuraQraqupyeuoro 

Ha3BaHI4t, [plrntroe B Hacroflulee BpeMt Ha3BaHI,le, yKa- 

3aHr4e KaTeropl[r T!I[a, KornqecTBa JII,iCTOB (eCnra nx 6ollure oAHoI'o) n 

KpyrrHoro peruoHa c6opa ruuonoro o6pa3lla, TeKcr 3TI,IKeTKI{ TLIIIOBOIO 

o6parua (r raertuxax), r\v:Tara locus classicus I43 llporoJlofa, [plaMerraHl4t. 

HccreAogaHne BbrrroJrHeHo upu rloAAepxKe Poccnficroro


flpuuevaune. Ha grurerKe [epBoro cvrHTurra aMeercr KapaHAarrrHas 

HaArIHcb' cAeJIaHHat, HacKoJrbKo Mbr MoxeM ey!.r4Tb, pyroft A. EyHre: 

6. A. hancockii Bunge ex Maxim., 1880, Bull. Acad. Sci. P6tersb. 26: 471. 

Lec to t yp u s (Sytin, hocloco): Cer' Ktlrafi,

Lectotyp us et isole cto typi (Wenninger, l99l): K,P. 

flo rporonory: . 

flpuuevauue. Paruuqa B Aare c6opa Ha 9'rlrKerKe cuHTr,rrra t,r B rrpo- 

ToJroI'e, Bepof,THo, o6rrcngercr oueqatrofi upu tty6lruraquu. 

12. Oxytropis arenaria Jurtz., 1964, Hosocrl4 cr.rcr. Bbrcu. pac;t. 1964: 

210. 1= O. oxyphylla (Pall.) DC. frde fpy6on, 1998, n Pacr. I{eurp. A:run 

86 : 6l; = O. hailarensrs Kitag. fide Z. W. Zhang, 1998, in Fl. Reip. Pop. 

Sin. 42, 2:125). 

H olo typ u s: Ces.-Bocr. Kurafi, (LE 

cum isotypo). 

fIo rpoloLroly:

16. O. mandshuricaBunge,l8T4, M6m. Acad. Sci. P6tersb. ser.7,22, 

1 :35. (= O. coerulea (Pall.) DC. firde Z.W.Zhang,1998, in Fl. Reip. 

Pop. Sin. 42,2 : 62). 

L e c t o t y p u s (flarnona, 1989): pocclrficrcufi'lanrnlrft Boctor, 

(coast of Manchuria,lat.44-45 N, 1859, [alab. et fl.], C.Wilford> (LE). 

Syntypi: Cen. Krraft, (LE); (LE); (LE). 

flo nporolory:

H o lotypus: MoHronl,rl,

Syntypi: Ces.-3an. Kurafi,

JI. B. Op.nona L. Orlova 

Tr4noBbtE o6pA3rlbr HEKOTOPbTX TAKCOHOB POAA 

ABIES MILL. (PINACEAE), XpAHguIt4ECfl B fEpSApI4I,I 

EOTAHI'IqECKOfO I4HCTITyTA I,IM. B. JI. KOMAPOBA (LE) 

SPECIMINA TYPICA TAXORUM NONNULLORUM E GENERE 

ABIES MILL. (PINACEAE) tN HERBARIO INSI'ITUTT 

BOTANTCT NOMINE V. L. KOMAROVTI (t,E) CONSERVATA 

B uacroauqefr clarle rry6Lruxyurcr AaHHbte o xpauqullrxct e fep6apur.r 

Eoraur.rqecKol'o r4Hc'lr{fyla nru. B. JL Konrapona (LE)-rntroBbtx Marepr,raJrax 

24 raxconos pot\a Abies Mill. n paHre BrrAa n pa3HoBu4Hoc- 

Tr-r, orrr.rcaHHbrx t{3 pa3JtrrrrHblx peluouon Poccnu (Kauxa:, ALnai.r, Clr- 

6upv u la,rrrunli Bocror), I-{eHrpaLt,uoi.r ra lOxuoii Enpoulr, IOr-o-Bocro.rnofr 

r.r MaLrofr Asnn, ASpr.rxu (Arxlrp) u Ces. Auepuxn. B npuue- 

AeH HOM Ht{Xe a H HOTr{pOBaH HOM CllHCKe HA3BAHl,jfl 1'aKCOHOB paCtIOJlOXe- 

Hu n alrQaor.rrHoM troprAre. llu KaxAol'o Ha3BaHHr upl{BeAeHbr Ho[,reH- 

KJra-fypHafl urilara, coBpeMeHHoe Ha3BaHHe pacleHur{ (n xpylLrlrx cxo6xax), 

xalelopur'rll\ta, lroJlHbrr"I lexc-r' r'ep6apuoil trurerKt{, cBegeHrln o 

Apyl'r4x fep6apulx, B Ko'ropbtx xpaHu'l'cfl 

'r'urrosoit Malepr4aJr (c yr

Sint. ex Boiss.) Coode et Cullen, 1965, Notes Roy' Bot. Gard. Edinb. 26 : 

167; Farjon, 1998, World Checkl. Conif. : 115, l2l). 

Onncan rc TypIrHIr. Lectotypus (Farjon' 1993: lll. ): 

6. A. delavayi Franch., 1899, in Journ. Bot. (Morot) l3 : 255; Farjon, 

1998, World Checkl. Conif. : ll0. 

OuncaH us [Oro-3ar. Kurar. Typus:

Onucan c o-Ba Caxanuu. Lecto typus (Farjon,1993:124, 

): ; >; 3) ; 

4) ; 5)

Ouucan us Ces. Auepurn. 

Syntypus:

Onucan I,I3 Arrxfipa. 

Syntypi (2): - (LE!). 

flo nporonory: ' 

20. A. sachalinensis F. Schmidt, 1868, M6m. Acad. Sci. P6tersb. (Sci. 

Phys. Math.), s6r. 7, 12,2 : 175. - A. veitchii Lindl. var. sachalinensis 

F. Schmidt, 1868, l. c. i 175,tab.4, fig., nom. altern. 

Onucan c o-Ba CaxanraH, 

Lectotypus (S. Lipschitz, 1960, in sched., (specimen authenticum>>, 

Orlova, hoc loco):


23. A. sibirica Ledeb. f. alpina Poljak., 1931, XypH. Pyccr. 6or. o6rq. 

16, 5-6 : 476 (= A. sibirica Ledeb., 1833, Fl. Alt. 4 :202). 

Ouucan cA.lrar. Le c t o t y p u s (Kamelin, in sched., Orlova, hoc 

loco):

T. B. Eropona 

XPOHOJIOTI4IIECKI{I)I NEPETIEHb AAT, 

COAEPXAIIILIXCfl B PAEOTE (MEXAYHAPOAHbIII 

KO.IIEKC EOTAHI,IqECKOIT HOMEHKIIATYPbI 

(CEHT-JIyUCCKI,Ifi KOAEKCD 

T. Egorova 

INDEX CHRONOLOGICUS DIERUM QUI OPUS 

(INTERNATIONAL CODE OF BOTANICAL NOMENCLATURE 

(SAINT LOUIS CODE> COMPREHENDIT 

Hacrorulax crarbfl, LtMeloulafl c[paBoqHblft xaparrep, BKJIIoqaer 

xpoHorofllqecKufi [epeqeHb Bcex AaT, coAepxaulHxct B HoMeHKlaryp- 

Hbrx cTarbrx rIocJIeAHefo u3AaHnt KHI{[?I

KOAeKCa v3baTA AATA LI COOTBeTcTByIoqee IpaBLIJIO, Kacalouleect Ha3Ba- 

Hr.rft pa3HoBu4nocrefi KyJr brr4Br4 pyeMbrx pacreHr4ft . 

flpegcrauxer r{3BecrHbre rpyAHocrrr noJlb3oBaHr4e rroMerqeHHoii B 

Toruficrou KoAeKce (Greuter et al.,1994; fpoftrep H Ap., 1996) xpoHororfiqecKoft 

ra6lr,rrleft (crp.23-24), B Koropofi [pr4 Aarax AaHbr ccbrJr- 

KI4 Ha oAHy ulu 6olee crarefi. ,{na rumleHur Heo6xoAr.rMofo rrpaBl.r- 

Jra llpuxoAHTcr B ueJroM pqAe cJtyqaeB [pocMaTpr4BaTb MHoIHe cTaTbn, 

rrro cBr3aHo c orrpeAeJreHHoir 3arparofi npeueuu. Kponae rofo, KaK obrJro 

orMeqeHo BbIUte, B nocJreAueM, ceur-JlyuccKoM KoAeKce, r,rMerorct Ho- 

BbIe AaTbr l1 cooTBeTcTByrouue I1IM IrpaBHJra. 

flpr,rao4uunfi sgect leperleHb Aar Hoctrr Jttltub BcrroMofareJ]bHbtfr 

xapaKTep I1I He MoxeT 3aMeHuTb TeKcT CaMOfO KOAeKCa, B KOTOpOM rpa- 

BLlJra, BKJlK'qarcUIne AaTbr, qacTo r4JlJuocTptrpoBaHbr rrpllMepaMu. 

I uar 1753 r. - Aara ny6luraqnu l-ro ulgaHur pa6orn K. JInHuee 

(C. Linnaeus) , paccMarpuBaercr KaK Aara, c 

KoTopoft HaqHHaercr AeficrBr,r'renbHoe o6Hapo4oBaHue Ha3BaHnfi TaKco- 

HOB, rrpr4HaAJrexauprx r Spermatophyta [cerraeHHbre pacTeHflfl : f oJroceMeH- 

HHe I/r roKpbrroceMeHHbtel n Pteridophyta. (Crarsr l3.l (a)). 

.{ra ronaa l-ro HgAaHna

'reNa 1753 r., Moxer 6url ocyqecrBJleHo c ltcrIoJIb3oBaHHeM cJIoBa (ex). 

Ho eclu paHr Lr raKcoHoMlaqecKoe rloJroxeHlre Ha3BaHLIt, flotBl'IB[rel'ocr 

.qo r4cxoAHoft Aarbr, 6lrnn ugnaeseHbl aBTopoM, Koropblfi BrlepBble 

A,eficrBnreilbHo o6HapoAoBaJI 3To Ha3BaHI,Ie, To B cKo6Kax I4 c rIocJIeAy- 

KrrqttM cJroBoM (ex)) Moxer 6blft Ao6as:IeHa oaMl,ulllt aBTopa Ao l{oxoA- 

Hoft Aarbr. (Crau 46.5, npnuep 22). 

.[o I nnnapr 1890 r. ncrIoJIb3oBaHI,Ie B KaKoft-fiu6o ueiloft uy6nuraqr.ru 

(cu. crarbrc 35.5) ronxo oAHoro BHyrpl4BllAoBoro paHra paccMarpnBaeTcr 

KaK TIpHHflTI{e paHra pa3HoBI,tAHocTl,I, ecJII,l 3TO He npOTlIBOpeqr,rr 

yKa3aHr4rM CaMOIO aBTOpa B TOft Xe caMofi uy6rltKallur.I. (Crarrr 

35.4). 

.(o I mrnapr 1908 r. ouy6JII,IKoBaHHat LlJurlocrpalrufl c aHaJILI3oM 

v$rvr, Arr'. 6eccocyqucrbrx pacreHr,rft, eAuncrseHHuft pucyHoK, Ha KoropoM 

rroKa3aHbr AeTaJIIt, c[oco6crBylorque I4AeHTuQI,lraquu, fBJltlorcf 

rrpl{eMJIeMHMH UpnMeHI4TeJIbHO K CTaTbe 42, AUeCTO OUI'/;CAHHfl rtnr \UarHo3a. 

(Crarrr 42.3). lB yroMsHyrofi crarbe 42, manntru o6pa3oM B 

flaparpaoe 42.1, t'ouopurcr 06 ycJroButx AeficrBltreJrbHot'o o6HapoAo- 

BAHIIa Ha3BaHHr pOAa H BI4Aa IIOCpeACTBOM eAHHOI'O pOAO-BI,TAOBOI'O 

orlrcaHr,rr --'

Hvrro HJrv Ar,rafHo3y, r,rnJrlocrpaqnefi. nnn pHcyHI(oM, rroKa3brBaroultrMr4 

cyqecTBeHHbre npr{3HaKr4, Jtu6o ccbulKofi Ha paHee ra a$Qerruruo o6sapoAoBaHHbre 

HJurrccrpauuro rrJur plrcyHoK. (Craur 38.I). 

C I rnnapn 1935 r. Ha3BaHr{e HoBoro rarcoHa pacrennft (rpoue ro- 

AOpOCJrefi r,I BCex ucKorraeMbrx raKcoHoB lo6 emux zpynnax pocmeuuil ctw. 

damu-- I nueapn 1958 z. u I nneapn 1996 2., coorBercrBeHHol cquraercf, 

AeficrBtrreJrbHo o6HapoAoBaHHbrM, ecJrr4 oHo cofipoBoxAaefrr Jrarr4HcKr4M 

OrrucaHr,reM r,rJrtr Ar.rarHo3oM, JI1460 CCbUTKOft Ha paHee n r$$eruEn_ 

Ho o6uapogoBaHHoe Jrarr,rHcKoe o[ucaHr4e r,rJur Ar{arHo3. (Crarlr 36.1). 

{o I ruoapn 1953 r. o6uapogoBaHr{e [Ha:aaHur HoBoro raxcoua] 

[ocpeAcrBoM Hecrupaeuofi anrorpaQnu rBJuercr e$serrvrnnHu. HecrupaeMar 

anrorpasur, [poH3BeAeHHar rro3AHee 3TOft Aarbr, He flBJrrercfl 

a$SerrrarHo o6uapogoraHuoft. (Crarrr 30.1). 

C I rHnapr 1953 r. o6Hapo4onaHr,re Ha3BaHr,rq HoBoro raKcoHa B Top- 

IOBbTX KaTalofax uJlr{ HeHayyHux uusopMarIHOHHbtX VrcAAHLLflx, a a 

I nueapn 1973 z. r,r B o6MeHHbrx crrncKax ceMrH He rBJrrercr s0serrran- 

Huu o6HapoAoBaHueM. (Crarrs 30.3). 

C I rurnapr 1953 r. pacrpocrpaHeHr,re fleqarHoro MarepraaJra, cor]po- 

BOXAarOIIIefo 3KcHKaTbr, He rBJrserc_f, :ssexrunuuM ooHapoAoBaHueM 

[ncne4crnne sroro He rBJ'ercr e00ex'anuo o6uapoAonaHHbrM r.r Ha3Ba- 

HI{e HoBoro raKcoHa' orII4caHI'Ie Koroporo coAep)Io4Tct B gToM lreqarHoM 

uarepuale]. Ecnu ne.rarHrrfi uarepnaJr pac[pocTp ajilfleTcr., KpoMe ror.o, 

orAenbHo or SKcr.rKar [r. e. n [eqarHbrx BbluycKax, KaKr.rMu qBurorcfl, 

Harrpr,rMep, BbrrrycKtr n3p.aHva (ctlucoK pacreunft rep6apraa snopm 

CCCPDI,'ro 3ro rBJrrercr a$serrunuuu o6Hapo4oBaHr,reM. (Crarir 30.4 

n upr,ruevauue l). 

C I rnnapr 1953 r. Arfl AeftcrBr4rerbHofo o6uapogoraHls,HasBaHVa 

TaKCOHa rrocpeAcrBoM ccbrJrKrr Ha paHee u e$$errunHo o6Hapo40BaHHOe 

onucaHue r4ltu Arlatno3 3Ta ccbIJIKa AoJIXHa 6r,rrr no.unoft n npauofi, 

KaK 3ro orrpeAeJreHo B crarbe 33.3 (Crarrfl,32.3). lTercr crarln 33.3 

rrpHBeAeH B cJre.(yrcuem a6sar1e]. 

c I nnnapr 1953 r. HoBar KoM6uHarr4fl r4Jrr{ 3aMeHrroulee Ha3BaHne 

(nomen novum), oc'oBaHHbre Ha paHee r.r AefrcrBr,rreJrbHo o6HapoAoBaH- 

HOM HA3BAHNId, HE f,BJIfl K)TC' ACfi CTBI,ITCNbUO O6UAPO4OBAHHbIMU, ECJIII 

6aluouuu, (r. e. cuHour,rM, Hecyulufi Ha3BaHr{e nnu euurer) r,lJrr,r 3aMeHreublfi 

cuuour,rru (ror4a [peAraraercr HoBoe Ha3BaHHe) He yKa3brBaerct 

flcHo 14 He AaeTCr [OnHar v [rpflMafl. ccbrnKa Ha efo aBTopa u rraecro [r. e. 

ucro'ruur] Aeficrrurelruoro o6napoAoBaHqr c yKa3aHHeM crpaHr.rubr 

utu ra6tut\br, a raKxe qaru. (Crarlr 33.3). [flpnueurx aro upanulo, 

Heo6xoAHMO yqurblBarb B onpeAeJreHHbrx cryqarx rroJro)KeHr,rs crareft 

33.2,33.4 u 33,61. 

c I rnnapr 1953 r. Hr{ oAHo r,r3 AByx r,rJrr,r 6oJrbruero rrlrcJra HasBaHr4ft, 

OAHOBPEMCHHO IPCANAIACMbIX AN' OAHOIO I,I TOIO XC TAKCOHA O,IIHITM 14 

280

TeM xe aBTopoM (TaK Ha3brBaeMbre aJrbrepHarr,rBHbre Hasnauua) He rB- 

Juercr AeftcrBurelbuo o6Hapo4oBaHnbrM. 3ro npanulo He rrpnMeHflerct 

B Tex cfiyqarx, KorAa oAHa Ir ra xe xona6uHallur oAHoBpeMeHHo r4crroJrb3yeTcs 

B pa3Hbrx paHfax - T. e. AJI' BHyTpr{Br4AOBbrx TaKCOHOB B 

[peAeJrax Bla4awwt AJrfl rro.qpa3AeJreHrlft poAa B lrpeAerax po4a (cu. coseru 

22 A. 1,22 A. 2 v.26 A. l-3). (C'rarw 34.2). 

C I rnnapr 1953 r. HoBoe Ha3BaHVe uilLr HoBafl rou6usaqur, o6HapoAoBaHHbre 

6eg scnoro yxa3aHlrr paHra raKcoHa, He flBJurorcr AeficrBrreirbHo 

o6napogoraHHbrMrr. (Crarra 35. I ). 

,{o I nnnapr 1953 r. HoBoe Ha3BaHwe vilv HoBaq roM6uHaqns, o6HapoAoBaHHbre 

6e: scuoro yKasaHuff paHra, fBJurorcr AeficrBr,rrenbso o6- 

HapoAoBaHHbrMH rrpta ycJroB?rr4, qro BbrrroJrHeHbt Bce Apyrr.re rpe6oBaHfit 

Alr AeficrnnTeJlbHoro oonapogona[vrfl; o1vt, oAHaKo, He uMelor crrJrbr B 

BO[pOCax rlpl,Iopr.rTeTa, KpoMe cryqaeB oMoHHMtrr{. Taroe [6espanronoe] 

HoBoe Ha3BaHue Moxer cryxl,ITb 6asuoHuMo[{ AJrr rrocJreAyrculux KoM- 

6unal\uit B orrpeAeJreHHbrx paHrax r,tJII{ 3aMeHreMbIM clrHoHtrMoM AJrfl 

nomina nova raKxe B orrpeAeJreHHbtx paHrax. (Crarla 35.3). 

C I rnnapr 1958 r. Ha3BaHr,re HoBoro raKcoHa coBpeMeHHbrx BoAopocneft 

crruTaercs AeftcrBurelbHo o6uapo4oBaHHbrM, ecJln oHo cofipo- 

BoxAaerct JIarI,tHcKI{M orII4caHI{eM r4Ju4 AnafHogou, rtu6o ccunxofi na 

panee n eQQerrn*ro o6napogoraHHoe naruHcKoe oilucaHr{e }rJrr{ Alrafuos. 

(Crarrs.36.2). 

C I rnnapt 1958 r. Ha3BaHr4e HoBoro raKcoHa HencKonaeMbrx BoAopocnefi 

Br.rAoBoro nlu 6onee Hu3Koro paHra cquraercq AeftcrsnrentHo 

o6Hapo4oraHHbrM, ecJrr,r oHo co[poBoxAaerct, B AorroJIHeHt{e K JrarLrHcKoMy 

orII{caHHIo HJIU AlrarHo3y, I4JlJllocrpa\ueit r{JIr{ pl4cyHKoM, rroKa- 

3bIBarcIrIr{MH orJlllrtlrrerbHue uopSoloruqecKue rrpr,{3HaKr,r, ;ru6o ccruxoft 

Ha paHee u e$Qerrrarno o6uapogonaHHbre nJrJlrocrparruro uJrn pucyHoK. 

(Crarra 39.1). 

C I rnnapr 1958 r. o6napogonaHr{e Ha3BaHr{tr HoBolo raKcoHa B paHre 

po.qa vrrr4 HvDKe poAa rBJrrercr Ae[crsr4reJrbHbrM ToJlbKo B ToM cJryqae, 

KorAa yKrrbrBaercf, Tlrrr Ha3BaHr.rr (cu. crarruT*10, Ho cM. crarbK) 

H. 9, npuueqaHr,re l, o nasnaru4rx HeKoropux ru6pr,r4oo). [Tuuon He 

r,rMeroT Ha3BaHlrfl HoTopoAoB r,r HoTOTaKCOHOB B paHfe [oApa3AeJreHHt 

poAal. (Crarua 37.1). [Cna. raKxe crarbro 37.2u rpr{Mep L n damot",- 

I nneapn 1990 e. u I nueapn 2001 z.l. 

C I rnnapr 1973 r. o6uapo4oraHue Ha3BaHr4f, HoBoro raKcoHa B 06- 

MeHHbrx crrucKax ceMflH He rBJrrercr e$Qerrunuuu o6Hapo4oBaHr4eM. 

(Crarra 30.3). 

C I rnnapr 1973 r. Ha3BaHrle HoBolo raKcoHa, Arr.f, Koroporo He 

6rrnn suuolHeHhr oAHoBpeMeHHo pa3JtnrrHbre ycnoBlar AeficrBr4TerbHoro 

o6Hapo4o BaHHfl., He rBJrrercr 4efi cr:nurenruo o6napo4oBaHHbrM, ecJrr,r 

He AaHa [oJrHar v npfllaafl ccbrJrKa (crarrr 33.3) na ny6luxaquu, r.Ae 3TH 

281

pe6onaHur 6tt;rn sttttorHeHbl paHee. .{arofi Taxofo Ha3BaH[r flBJttercfl, 

Aara, B Koropyrc 6nro sunoJrHeHo rrocreAHee ycJroBI,Ie 4eftcraure.rrluoro 

o6HapoAoBaHr4r. (Craru 45.1). 

C I mroapr 1990 r. nexrorurru$r{Kaur4r r{Jr[ Heorunu$raraqrax Ha3Ba- 

HVtf. BldLA UJur BHyTp?rBr4AOBOfO TaKCOHa rrOcpeAcTBoM ?K3eMlrJrflpa HJrr,r 

Heouy6luxosaHHofr r,rJrJrrocrpaur4u He rBJurercr 4eftcrnureJrlHoft, ecJrH He 

yKa3au fep6apufi r.rlr.{ yqpexAeH[e, B KoropoM xpaHr.rrcf, coorBerc.rByrouluii 

rnn. (Cra'rtr^ 9.20). [B orHorueHnri JrerrorlrrruQurcaquu rocpeAcrBoM 

Heouy6lnxosaHHofr uJrJtrocrpar-([]r cM. crarbrc 9.10 ra c-rararc 9.2, 

Irpr{MeqaHr.,re 2; ca. maKJrce npasuno, scmynuswee s culy c I aueapn 

2001 z.: o ueo6xoAnMocrtl r,tcrroJrb3oBaHur repMnHoB (([erTorr,ur) 14 

>]. 

C I rnnapr 1990 r. yra3aHr4e rr4ua uprr o6uapo4onaHur Ha3BaHr{t 

HoBot'o raKcoHa B paHre poAa uJrr4 s 6olee Hrl3KoM paHl.e AoJtxHo 

BKJrroqarb oAHo r,r3 cnoB - - (typus) ulu

Te)), Koropar, crana ucrloJlb3oBarbcs B pyccKofi :rureparype 3aAoJIro Ao 

[prdBeAeHHoil sr[xe Aaru. Ctvt. TaKxe crarlto 7,10], 

C I rnnapn 2fi)l r. nerroruuu$HKaul4t uJln Heo'ruun$nraum Ha3Ba- 

HVtfl. BUilA r4Jrn BHyTpI4BI4AOBOTO TaKCOHa He CTInTaeTCS BbIIIOJIHeHHOIi, 

ecru IIpn rlrulrsuraqnH He 6bul I4crIoJIb3oBaH TepML{H , ero coKpaIUeHHar Sopua IrJlI4 3KBlrBaJIeHT B coBpeMeHHoM 

r3brKe (Ho cvr. crarblo 9.8). (Crarrt 9.2l). [B yrrorrlauyrofi crarte 9.8 u 

B rrpnMepe 3 roropurca o roM, tlTo I{cIIoJIb3oBaHLIe repMKHa, He rlpeA- 

Ha3HaqeHHOrO AJrf o6ogHaqenut o[peAeneHHoil KaTefopliI,I THIla 

(naupnuep, ynorpebleuue repMI4Ha (oIeKTorr.IlD .4rtfl o6o3HaqeHut rKIIa, 

xoropuil B AeftcrBI{TeJIbHOgI'I{ rBJrtercfl Heorl[loM, IIJII{ TepMI,IHa (l'oJlo- 

Tr{rrD BMecTo TepMr{Ha (neKroTn[))). paccMaTpHBaeTcf, KaK oullt6Ka, IIOA- 

rexaqas ucupaaleHurc]. 

Jlnreparypa 

Arexcees 8.6., f y6anos 14. A., Tlrxo M Hpos B. H. EotasnqecKar 

HoMeHKrrarypa. M., 1989. 167 c. 

f p6ftrep 8., M arHun .[,x. r.r4p. Mex4ynapoAuufi KoAeKc OoraHnqecxoft 

nouenxlarypn (Cenr-Jlyr.rccrufi roAexc), npuHrrufi LllecrnaAuarstu 

Mex4ynapogsnu 6orasuqecKr4M KoHrpeccoM, CeHr-JIync, Muccypra, 

r,uorrb-aBrycr 1999 r. flep. c aurn. CII6., 2001. 210 c. 

f poftrep B.,6app r,r @. P. uAp. MexAyHapognmft KoAeKc6oraunqecroti 

HoMeHKJrarypu (Toxnftcxuft roAexc), npunrrufi flrrnaAuarrtu MexqyuapoAHbrM 

6oransqecxnM KonrpeccoM, l{oxoraua, aBrycr-ce}tra6pr 1993 r. 

flep. c anu. Cft6., 1996. 19l c. 

Greuter W., Barry F. R. etal. International codeof botanical nomenclature 

(Tokyo code), adopted by the Fifteenth lnternational Botanical 

Congress, Yokohama, August-September I 993. Kbnigstein, Germany, I 994. 

xv + 389 p. (Regn. Veg. Vol. l3l). 

Greuter W., McN eill J. et al. lnternational code of botanical nomenclature 

(Saint Louis code), adopted by the Sixteenth International Botanical 

Congress, St Louis, Missouri, July-August 1999. Kiinigstein, Germany, 

2000. xviii + 474 p. (Regn. Veg. Vol. 138). 

Voss E. G., Bu rdet H. M. et al. International code of botanical nomenclature, 

adopted by the Thirteenth Intemational Botanical Congress, Sydney, 

August 1981. Utrecht, Antwerpen, 1983. xv + 47?p. (Regn. Veg. Vol. lll). 

Eoraxuqecrcnfi uncrnryr uivr. B. JI. Kouapona PAH 

197376, C.-flerep6ypr, yn. flpoQ. flonosa,2 

E-mail: egorova@DG4972. spb.edu

lt. O. Eyrynona 

HOMEHKJIATYPHbIE 3AMETKII 

NOTULAE NOMENCLATURALES 

sPrRAllA PJASSETZKII BUZUNOVA (ROSACEAE) - HOBOE 

HA3BAHI4E,IIJI' SPTRAEA PROSTRATA MAXIM 

sprRAEA PJASSETZKI BUZUNOVA (ROSACEAE) - NOMEN 

NOVUM PRO SPECIE SPIRAEA PROSTRATA MAXIM. 

Spiraea piassetzkii Buzunova nom, nov. - Spiraea prostrata Maxim., 

1879, Tp. flerep6. 6or. caAa 6, 1 : 184, nom. illeg., non Schur, 1866; 

T. T. Yii et L. T. Lu,1974, in Fl. Reip. Pop, Sin. 36 : 53. 

Eorannqecrltft nHcrnryr uu. B. JI. Korraapona PAH 

197376 C,-flerep6ypr, yn. flpoQ. flouora, 2 

E-mail: buzunova@DG4972.spb.edu 

B. B. Es.nr 

V. Byalt 

TETRAGONIA PHILIPPI BYALT (TETRAGONIACT]AE) _ HOBOE 

HA3BAHI4E AJlf TETRAGONIA ROBUSTA PHIL. 

TETRAGONTA PHILIPPI BYALT (TETRAGONIACEAE) - NOMEN 

NOVUM PRO SPECII] TETRAGONIA ROBUSTA PHIL. 

B 1894 r'. @ururnnu (Philippi, 1893-94) onucar HosHft BI{A r43 poAa 

Tetragonia -- T. robusta Phil., pacrpocrpaHeHHblft n Kappucal Eaxo s 

rrpoBHHrIHr,r Araxanaa (9r,trln). OAHaro 3To Ha3BaHue orasanocl 6olee 

no3AHrrM oMoHr,tMoM T. robusta Fenzl (Fenzl, 1862), orllcaHHoro H3 

HauaxyaraHAa B IOIKH. Atfpuxe. HarsaHue T. robusta Phil. ssil.flercq 

He3aKoHHbrM, r{ rro3ToMy MbI rIpeAJIaraeM [epeuMeHoBarb nuq, u T. philippi 

Byalt nom. nov. 

Tetragonia philippi Byalt nom. nov. -- Tetragonia robusta Phil., 1894, 

Anal. Univ. Chil. 85 : 169, nom. illeg., non Fenzl, 1862; C, M. Pizarro, 

1960, Espec. Pl. Descr. Phil, : 54. 

T y p u s: Se encuentra en el Carrizal Bajo de la provincia de Atacama, 

R. A. Philippi (holotypus -- SGO: 48403). 

Hasnasne AaHo B rraMnrb P. (DunnurIu (R. A. Philippi), KpyllHeftIuero 

3HaroKa $.nopm Qnlu n XIX eere. 

284 

l. Buzunova

Jlnreparypa 

F e n z I E. Genus Tetragonia L. // Harvey w. A., Sonder O. w. Flora capensis. 

Cape Town, 1862' Vol. 2. ix + 621 p' 

ehilipii R. A. plantasnuevas chilenas//Anal. Univ. Chile. 1893-1894. 

Vol. 85. P. 167-195. 

Eoranu.recKl,Ifi I,IHcrI'ITyr I{M. B. JI' Koruraposa PAH 

lg73'76 C.-fterep6ypr, yn' flpo$. flonora,2 

E-mail: byalt@Dc4972. spb.edu 

A. K. Crrrnn 

A. Sytin 

O AATE TTYEJTI4KAqUH PAEOTbI A. A. EYHIE 

DEDIEA.BUNGEIoPERtS(BETTRAGZURKENNTNISSDERI.-LOR 

RUSSLANDS UND DER STEPPEN CENTRAL-ASIENS> 

,{aroft ny6lurauuu pa6oru A. A. Eynre

AJIOABIITHbIIZ YKA3ATEJIb HA3BAHI{IT TAKCOHOB 

INDEX ALPHABETICUS TAXORUM NOMINUM 

Xnpuum urpu$rom orMeqeHbr Ha3BaHr,rt B[epBbIe onucblBaeMbIx 

TaKCOHOB U HOBbre Ha3BAHnfl. AJlfl LI3BeCTHbIX paHee TaKCOHOB, a TaKXe 

crpaHrrrlbt, r4e ny6nnrynrcfl 3TI1I Ha3BaHIlIr. Kpyxo.moM (o) orMeqe- 

Hbr crpaHr,rqbr, Ha Koropbrx AaHbI Kaprbl apeaJroB, Kpecr[KoM (+) 

crpaHrrubr, Ha Koropbrx lloMeuleHbr uro6paxeuur 

TAKCOHOB, 

coorBercrBynuux 

Acantholimon 19 

Acantholimon kokandense 38, 39 

-- linczevskianum 37+, 38 

- minshelkense 38, 39 

Aconogon 224 

Actinoscirpus 4l 

Allium 96, 99, 101 

-- subgen. Rhizirideum 96 

* sect. Allium 101 

- sect. Brevispatha 102 

- sect. Cepa 102 

- sect. Codonoprasum 102 

- sect. Oreiprason 96 

- aegyptiacum 101 

--- ascalonicum l0l, 102 

-- callimischon subsp. haemostictum 102 

- cepa [01 

var. schallot 102 

-'curtum subsp. aegyptiacum l0l 

-- -- subsp. palaestinum 102 

-- - var. negevense 102 

- durandoi 101 

- flavum var. pilosum 102 

- flexuosum 102 

--globosum 96, 97 

- haemostictum 102 

- junceum subsp. tridentatum l0l 

- - var. tridentatum 101 

.'- kurrat l0l, I02 

- marschallianum 96,97 

- palaestinum 102 

286 

parvillorum l0l 

-' phanerantherum subsp. deciduum l0l 

'.- - var. deciduum 101 

-- pilosum 102 

- porrum l0l 

var. kurrat 102 

- proponticum var. parviflorum l0l 

- psebaicum 96 

- sanctum 102 

var. negevense 102 

savranicum 96 

- saxatile 96, 97 

- sphaerocephalon subsp. durandoi l0l 

- var. durandoi l0l 

- staticiforme subsp. llexuosum 102 

- f. flexuosum 102 

- stearnianum subsp. vanense 102 

--- vanense 102 

- villosiusculum 102 

Amborellaceae 220 

Ambrosia 200 

Androcoma 5l 

Antirrhinum linaria 185 

- albifrons 200 

-- arvense 204 

- canadense 201 

dalmaticum 196 

genistifolium 194 

- chalepense 206 

- incarnatum 202 

--- macrourum 191

- micranthum 204 

-- odorum 190 

- pelisserianum 205 

- pyramidatum 189 

reflexum 199 

-- repens 202 

-- simplex 204 

Arabis abietina 224 

- gerardii 224 

hirsuta 244, 225 

- - var. sagittata 224,225 

- planisiliqua 224 

- s'dFittatl224 

- - var. subglabrata 224,225 

Archaeopteridopsida 7 

Archaeopteris 7 

Arenaria glutinosa 107, 109 

Artemisia 221 

- incana 151 

- sect. Laguropsis 149 

- sect. Megalocystis 150, l5l 

- sect. Poterion 150 

-- sect. Xiphidium 123, 124,138, l4l,147 

- ser. Angustati 150 

.- ser. Arbusculi 126,127 

ser. Densiflori 150 

ser. Macrotropidi 127, 128 

- ser. Mugodsharici 140 

-- ser. Subauriculati 150 

-- ser. Viminei 136 

- ser. Virgati 140 

- aktiubensis 139, 140 

- albicaulis 146 

-- anthylloides l5l, 157 

-- arbuscula 126 

Asteraceae 217 

-- subfam. Ambrosioideae 220 

-- subfam. Asteroideae 217, 218, 220, 221 

- subfam. Barnadesioideae 217 . 218,220 

- subfam. Carduoideae 217.218.220 

-- subfam. Cichorioideae 217, 218, 220 

-- subfzrm. Gochnatioideae 217, 218, 220 

- subfam. Mutisioideae 217. 2I8. 220 

- subfam. Pertyoideae 217 

-- trib. Ambrosioideae 220 

- trib. Anthemideae 218, 220,221 

- trib. Arctotideae 220 

- trib. Astereae 220 

- trib. Calenduleae220 

- trib. Cichorieae 218. 220 

--- trib. Coreopsideae 220 

- trib. Cynareae 220 

.*' trib. Eupatorieae 220 

-- trib. Filagineae 220 

-- trib. Gnaphalieae 220 

-- trib. Gundelieae 220 

- trib. Helenieae 220 

-- trib. Heliantheae 218, 220,221 

- trib. lnuleae 213,220,221 

- trib. Mutisieae 220 

- trib. Plucheeae 213. 220 

- trib. Senecioneae220 

- arbuscula auct. 126 

- aulieatensis 230 

--- austrouralensis 125, 128, I29+,132, 

134, t39', 147 

- brachylobus 126, 138, 139', 140, 147 

-- calycinus 149 

-.-- cinereus 146 

- compositus 231 

-- cornutus 126,136,137', 138, I4l, l4'l 

- daghestanicus 150-152 

- dictiophysus 154 

- ebenoides 151 

-.- fruticosus 138 

- fruticosus auct. 136 

- halicacabus 157 

- helmii 132 

- igoschinae 124, 126,127, 147 

-- kaghysmani 152, 154, 155+ 

-- karakuschensis 152, 156+, 157 

-- karelinianus 125, 128, 129+, 130-132, 

r34.139" 147 

- - var. karelinianus 132 

-- - var. violascens 132 

- karelinianus auct.127, I32 

- khoshjailensis 150 

- kustanaicus 143,144 

-- macroceras 132 

- macrolobus var. p. nigropilosus 128, 

130 

- - var. nigropilosus 128, 130, 13l 

- trib. Tageteae 220 

- macrolobus auct. 128 

Astragalus 123, 124, 147, 149-l5l 

- subgen. Astragalus 149 

- subgen. Cercidothrix 124,149 

* sect. Anthylloidei 149-151 

-- sect. Grammocalyx 150 

- sect. Halicacabus 149-l5l 

macropus 126, 133+, l4l-144, 145', 

t46. t47 

- macropus auct. 142 

-mesites 150, 152, 153+, 154 

- 140 

- mugosaricus 124, 140, 147 

287

-* oropolitanus 126,137,138, 145", 147 caldwellii 90 

-- ortholobus 128. 132. 134 

- compactus 89 

- pallescens 123, 126, 142, 145', 146, 147 -- desoulavii 88 

- physaloides 149 

--.fluviatilis 87 

-- pseudanthylloides 150 

- - subsp. yagaraST 

- pseudomacropus 125, 133+, 142, 144, - glaucus 84, 86, 87 

t45", t47 

- grandispicus 90 

- pseudotataricus 125, 114-136, I47 - koshewnikowii 90 

-- pycnolobus l3l 

- laticarpus 90 

- raddei 151 

- medianus 90 

- stenocarpus 231 

-- maritimus 82, 8,1-89 

- stenoceras I27, 128 

- - subsp. paludosus 89 

- storozhevae I25, 137", 142, 145,147 - -- var. desoulavii 88 

-- subulatus 135 

-- maritimus auct. 87 

- - var. y. tataricus 134 

- nobilis 85,87 

var. y. uralensis l4l 

- novae-angliae 90 

- subuliformis 135 

- paludosus 84, 89 

- --- var. (. tartaricus 134 

-- planiculmis 81, 90 

- subuliformis auct. 134, 135 

- popovii 89 

- suffruticosus 138 

--robustus 89 

surugensis 150 

- schmidii 89 

- szovitsii l5l 

-- strobilinus 90 

- tataricus 134 

- yagaru 87 

- temirensis 125, 127 , 128, 137', 147 Botrychiaceae 8 

- tenuifolius 132 

Botrychium 7-10,20 

- ucrainicus 125, 134-136, I47 

- subgen. Botrychium 8-10, 13 

- uninodus 231 

- subgen. Osmundopteris 8, 9, 12 

- uralensis 138 

-'subgen. Sceptridium 8, 9, l8 

- varius 126,133+,138, 139', 140--145, - sect. Botrychium 13 

-- vimineus 136 

- sect. Elongata l8 

-- virescens 128. 130. l3l 

--- sect. Hiemobotrychium 8, l8 

- virgatus 140 

- sect. Lanceolata 9, 13 

- var. B. uralensis 141 

- sect. Lanuginosae 9 

- wagneri 149,157 

- sect. Multifida l8 

- willisii 150 

- sect. Osmundopteris 12 

- wolgensis 132 

- alaskense 9. 11. l3-15 

- zederbaueri 149 

- -- var. pavlovii 14 

-- angustisegmentum 13 

Bellis 218 

-- anthemoides 11. 12. 19 

Bistorta 224 

-- boreale 9, 12 16 

- sect. Yivipara 224 

- - subsp. obtusilobum 14 

Blysmus 40, 4l 

- - var. intermedium 15 

Bolboschoenus 40, 41, 80-84, 86-88 --- boreale auct. 13, l4 

-- sect. Bolboschoenus 83-86,88 

- crassifolium 14 

-- sect. Browningia 86 

- crassinervium 9, 12, 14, l5 

-- sect. Lentischoenus 83, 84, 88 

- - var. obtusilobum 14, l5 

- sect. Mediani 90 

- crenulatum 16 

- subsect. AIIines 89 

- dissectum l8 

--- subsect. Bolboschoenus 88 

- x intermedium 14, l5 

-- subsect. Fluviatiles 87 

- lanceolatum 9, 11, 13, 14, 16 

- subsect. Glauci 87 

- lanceolatum auct. 13 

- subsect. Popoviorum 89 

- lanuginosum 8, 9 

- affinis 85, 89,90 

- lunaria 9-ll, 14-17 

288

- - var. incisum 16 

var. lunaria 16, 17 

- -- var. przevalskii 17 

*- -- var. rhombeum 16 

- lunarioides 8. 18 

--- matricariac 19, 20 

- matricariifolium 9-11, 13, l7 

- mindagense 16 

- multilidum 9-ll, 17 -20 

-.- subsp. robustum 18 

-.obliquum 18 

- onondagense 16 

-- pinnatum 14, 15 

- ramosum 17 

- robustum 10. 18, 19 

-- rutaccum 19 

- rutaefolium var. robustum 18 

silaifolium 19 

- simplex 9, 10, 17 

- strictum ll, 13 

- ternatum 10, 18, 19 

- tunux 16 

- virginianum 9, 12 

- - subsp. curopaeum 12 

- - subsp. virginianum 12 

-- - var. europaeum l2 

- virginianum auct. 12 

- yaaxudakeit 16 

Botrypus 8, l2 

-- strictus 13 

Bryodesma 23, 26 

- shakotanensis 26 

- sibericum 26 

Calyceraceae 219 

Campanulaceae 218 

Carduus edpressus subsp. novorossicts 2tl 

- laciniatus 227 

- -- subsp. novorossicus 227 

- novorossicus 227 

Caryophyllaceae 106, 109 

Cephalanthera 224 

- sect. Cucullatee 223 

-- cucullata 223.224 

- epipactoides 224 

- floribunda 224 

-- kurdica 224 

Cephalaria 213 

Cerastium 108 

Ceratophyllaceae 220 

Chrysanthemum 221 

Coronilla 225 

- balansae 225 

l9 3arar Ne 915 

- cappadocica var. balansae 225 

- orientalis 225 

- - subsp. belansre 225 

- -- var. balansae 225 

Crambe sect. Crambe 225 

- sect. Trtrriae 225 

- subsect. Tataiae 225 

- grandiflora 225 

- litwinowii 225 

steveniana 225 

- tataria225 

Cyclachaena 221 

Cyperaceae 40, 48, 51, 55, 80, 82 

- trib. Fuirencae 82 

- trib. Scirpeae 82 

Delphinium knorringianum 229 

Dichodon 108 

Dipsacaceae 214 

Echinops 214 

- tschimganicus 232 

Eremurus 103-105 

- azerbaidshanicus 104 

-- spectablis 104 

tauricus 104 

-- zangezuricus 103-105 

Eriophorum 40,67-69 

- subgen. Japonicus 65 

- sect. Japonica 65 

- cyperinus 6l 

- maximowiczii 66 

-japonicum 66-67,72 

Euphorbia 159 

- subgen. Esula 159, 160 

- sect. Murtekias 159 

- sect. Myrsiniteae 159, 160 

- sect. 160 

- sect. Paralias 159, 160 

- sect. Tithymalus 159 

--- subsect. Myrsiniteae 159, 160, 163 

- ser. Biglandulosae 162 

- ser. Denticulatae 167 

- ser. Myrsiniteae 163, 168 

-- $ Myrsiniteac 159 

- anacampseros 161, 163, 166, 168 

- - var. tmolea 166 

- armena 16l, 163, 164, 166 

- biglandulosa 162 

- broteri 163 

- cilicica 167 

- corsica 16l. 167. 168 

-- craspedia 16l, 167 

289

curtifolia 165 

- denticulata 16l,167 

- fontqueriana 16l,166, 167 

- marschalliana 16l,164,166, 168 

- - subsp. armena [63 

- - subsp. marschalliana 164 

- - subsp. hirsutum 106 

-- - subsp. umbellatum 106 

- var. glandulosum 109 

var. glutinosum 106, 107, 109 

var. tenerrinum 106 

- -- var. umbellatum 106, 107, 109 

subsp. woronowii 164 

-'monostyla 160, 162-164, 166 

Hulthemia persica l5l 

--myrsinites 160, 16l, 163-166, 168 llex aquifolium 219 

-- - subsp. litardierei 166 

- -- subsp. rechingeri 167 

- oxyphylla 160, 163 

-- pectinata 167 

lnula grandiflora 227 

- orientalis 227 

- - subsp. grandiflora 227 

lsontopsida 22 

phlomos 162 lsolepis 40, 4l 

- pontica 165 

- pungens 162 

lva22I 

- rechingeri 16l, 167, 168 

- rigida 160, 162,163, 168 

- rotundata 167 

- spinidens 161, 163, 165 

- veneris 16I,162 

- woronowii 164 

Japanobotrychium 8 

arisanense 8 

Juniperus sabina 210 

Kickxia aegyptiaca 195 

Lamiaceae trib. Marrubieae 178 

Ferula 170 Lapsana 218 

petiolaris 172 Lepidodendron 7 

- turcomanica 174 

Festuca schischkinii 28, 30 

- subgen. Leucopoa 28 

- tzveleviane 28,29+ 

Fraxinus excelsior 226,227 

- - var. oxycarpa 226 

- oxycarpa226 

- - var. rostrata 226,227 

- rostrata 226 

Gladiolus 96, 98 

caucasicus 98, 99 

- communis 98. 99 

- murgusicus 98 

Gochnatia ilicifolia 219 

Goodeniaceae 218 

Hedysarum cumuschtanicum 231 

pskemense 231 

Helminthostachys 8 

Hepatica falconeri22g 

Holosteum 106, 108 

-- glutinosum 106-109 

liniflorum 106-109 

- marginatum 108-ll0 

- polygamum 106, 108, 109 

- umbellatum 106-109 

- - subsp. glutinosum 106, 109 

Leutea 170, 172,174, I77 

-- cupularis 172 

- petiolaris 170, l7l+, 172,174,177 

sclerophylla 174, l'17 

-- turcomanica I74,175+, 177 

Linaria 182-184 

- sect. Arvenses 184,203, 205 

sect. Chaenorhinum 182 

...' sect. Corrugat:as 200,202 

-- sect. Cymballaria 182 

-- sect. Cretaceae 183, 197, 198 

--'sect. Diffusae 197,199 

- sect. Discoideae 185 

-- sect. Elatinoides 182 

- sect. Grandes 184 

--- sect. Kickxia 182 

- sect. Leptoplectron 201 

- sect. Linaria 182-184, 195 

- sect. Linariastrum 182, 184, 203 

- sect. Macrocentrum 205,206 

- sect. Minutiflorae 199 

- sect. Oblongae 193 

- sect. Pelisserianae203,204 

-- sect. Repentes200,202 

-- sect. Speciosae 193, 196, 197, 198, 200 

- sect. Supinae 193,203 

sect. Versicolores 200, 20t,205 

'- subsect. Cretaceae 197 

290

- subsect. Laeves 183, 185, 190' 193 

- subsect. Linaria 183, 185, 189 

- subsect. Macrocentrum 205 

- subsect. Pyramidatae 189 

- subsect. Tuberculatae 185 

- ser. Cretaceae 197 

- grex Discoideae 185 

- grex Oblongae 193 

- g Arvenses 203 

-- $ Diffusae 199 

- $ Discoideae 183, 184 

- $ Grandes 184 

- $ Leontorrhinum 183 

- $ Licorrhinum 183 

- $ Oblongae 183 

- $ Speciosae 193 

- $ Versicolores 200,201, 205 

- acutiloba 183, 186 

- adzharica 194 

- albifrons 200 

- altaica 190 

-- angustissima 187 

- araratica 188 

- armeniaca 205.206 

- arvensis 203,204 

- - subsp. simplex 204 

- - var. micrantha 204 

- - var. simplex 204 

- baxanensis l9l,192 

- bessarabica 186, 187 

- biebersteinii 186, 187 

- - subsp. maeotica 187 

- - subsp. ruthenica 186 

--- - var. ruthenica 186 

- bipartita 202 

- bipartita auct.202 

- canadensis 183, 201, 206 

- caucasigena 194 

- chalepensis 206 

- confertiflora 109 

- corifolia 200-202 

- corrugata 201,202 

-.cretacea 183, 197-199 

- - var! creticola 198 

- creticola 197, 198 

- dalmatica 194. 196.227 

- - var. grandiflora 195 

- - var. parviflora 196,227 

-debilis l9l,l92 

-- decipiens 199 

-- depauperata 195 

-- dulcis 190 

- elymaitica 189 

- euxina 194 

.- flava 199 

-- floridiana 201 

- genistifolia 183, 193-197 

- - subsp. dalmatica 194 

- -- subsp. euxina 194 

- - var. p*vif7on227 

- grandiflora 183, 195, 196 

- grandiflora auct. 196 

- griffithii 190 

-- grossheimii 187, t88 

- iberica 194 

- imeretica 194 

-- incarnata 202,206 

- incompleta l9l, 192 

- iralica 187 

- italica auct. 187 

- kantschavelii 194 

- kopetdaghensis 189 

- kurdica 183, 188, 189 

- - subsp. araratica 188 

- - subsp. aucheri 189 

- - subsp. eriocalyx 189 

- - subsp. pycnophylla 189 

- lenkoranica 189 

- lineolata 189 

--P elymaiticalS9 

- loeselii l9l 

- loeselii auct. 190 

- macrophylla 183, 197, 198 

- macroura 183, l9l, 192 

- x vulgaris 192 

- maeotica 186 

- megrica 188 

- menisperma 198 

- meyeri 192, 193 

- micrantha 204 

- minutiflora 200 

- monspessulana 202 

- odora 183. 190. 191 

-- - subsp. dulcis 190 

- ordubadica 188 

- pelisseriana 203,205 

- petraea 194 

pontica 194 

-- pyramidalis 189, 190 

- - subsp. kopetdaghensis 189 

- - subsp. lenkoranica 189 

- reflexa 199 

- repens 200,202 

--0 monspessulana203 

-- rupestris 193 

- ruthenica 186 

291

- sabulosa 182. 197 

- scenoreina 194 

- schelkownikowii 187 

- schirvanica 192 

-.- simplex 204 

-- sofiana 194 

- sl.Iiata202 

- syspirensis 194, I95 

- tesquicola 186 

- texana 201 

- uralensis 190 

- versicolor 200 

- viscosa 200 

- volgensis 183, 190, l9l 

- vulgaris 183-186 

- - var. acutiloba 186 

-- - var. latifolia 186 

-- - var. vulgaris 185, 186 

- - f. latifolia 185 

- vulgaris auct. 187 

- zalagezura 195, 196 

Lobeliaceae 219 

Lycopodioides 22-24 

- subgen. Lycopodioides 23, 24 

- subgcn. Tetragonostachys 23, 24, 26 

-- sect. Diplostachyum 24 

-- sect. Lycopodioides 24 

- sect. Rosuharc24.25 

- sect. Stachygynrudrum 24, 25 

--aitcNmnii 25 

- borealis 25 

- denticulata 24 

- flabellata 25 

- helvetica 24 

- rossii 25 

- rupestris 26 

- sanguinolenta 25 

- shakotanensis 26 

- siberica 26 

- tamariscina 25 

Lycopodiophyta 7, 22 

Lycopodiopsida 22 

Lycopodium boreale 25 

-- helveticum 24 

- sanguinolentum 25 

- selaginoides 24 

Mankyua 8 

Maximoviczia 65, 68 

- japonica 66 

Maximowiczia 65 

Maximowicziella 65. 68 

-japonica 66, 68 

292 

Nemocharis 44, 69 

radicans 70 

- sylvatica 73 

Neuroloma angrenicum 230 

Nuttallanthus 183, 201 

- canadensis 183, 201 

Nymphaeaceae 220 

Oleaceae 226 

Olgaea vvedenskyi 232 

Oligolepis 214-216 

Ophioglossaceae 7, 8, 20 

Ophioglossales 7 

Ophioglossopsida 7 

Ophioglossum 8 

Ornithogalum 103 

- subgen. Beryllis 103, 104 

-- arcuatum 103, 104 

- magnum 103, 104 

Osmunda lanceolata l3 

- lunaria 16 

- - var. matricariifolia 17 

- matricariae 19 

- niultifida 19 

- ramosa 17 

ternata 18 

- virginiana 12 

Osmundopteris 8, 9, 12 

Persicaia 224 

Peucedanum 170 

cupufare 172,173+ 

-- involucratum 170, 174 

.- sclerophyllum 170, 174, 176+,177 

- turcomanicum 170, l'12, 174 

Phlomidoschema 178 

Phlomoides adylovii 32, 33+ 

'- alaica 34 

- kurpsaica 30, 3l+ 

'- lanatifolir 232 

-- urodonta 32 

Platyruscus 223 

Plumbaginaceae 38, 39 

Poaceae 28, 30 

Polygonum 224 

-- subgen. Bistorte 224 

-- sect. Bistorta 224 

- sect. Vivipara 224 

Polypodiophyta 7, 20 

Potentilla malacotricha 230 

- stanjukoviczii 230 

Primula eugcniae 36, 38 

-- macrocalyx 38

- pskemensis 35+, 36 

Pseudosphaeranthus 214 

Pterocaulon 213 

Ranunculus longilobus 229 

Ruscus 223 

- sect. Platyruscus 223 

- sect. Ruscus 223 

- ser. Simplices 223 

Reigera 84, 88 

Reynoutria 224 

Rohrbachia 223 

Rosa 112, l13, ll5, l16 

- sect. Caninac 113, l16, l19, 120 

- sect. Cassiorhodon I 16 

- sect. Cinnamomeae l12, ll3, l16, t20 

- subsect. Rubiginosae I 19 

- subsect. Vestitae ll6 

-* cycl. Balcaricae ll6 

- cycl. Oxyodontae 112, l15 

- cycl. Pendulinae lL2, ll4 

- acicularis 113 

- adenophylla l15, l16, 121 

- x altidaghestanica 118 

- balcarica 116, l18, l2l 

- x baxanensis l18 

- beggerana l13 

- cinnamomea I14, t20 

- --var. glabrifolia 120 

- donetzica I 14 

- forrestiana l13 

- gadzhievii l19 

- giraldii l13 

- glabrifolia I 14, I I 5, Il7, 120 

- var. glabrifolia ll5 

- var. pubescens l15, 120 

- glauca 113 

- gorenkensis l16 

- haemathodes ll8 

- majalis 114-l 17, 120, l2l 

-- majalis auct. 120 

- obtegens 116, l18, l2l 

- oxyacantha l13 

- oxyodon ll2-115, ll7-l19 

- - var. pleiantha I l8 

- x oxyodontoides ll8 

- pendulina l12, 113 

-- phoenicia l13 

- pratorum 115, 120 

-prokhanovii ll2,ll3, l17, l19 

- roopae ll9, l20 

- sempervirens I 13 

- sosnovskyana l12, l13, tl7, 119 

- spinosissima I l3 . 

webbiana 113 

Ruscaceae 223 

Saxifraga colchica 225 

- moschata 225 

- - var. pontica 225 

- pontica 225 

- subverticillata 225 

-- --- subsp. colchica 225 

Scabiosa bipinnata 227 

- - var. calcarea 227 

- imeretica 227 

- ochroleuca 227 

- - var. calcarea 227 

- sosnowskyi 227 

Sceptridium 8, 9,18 

- multifidum [9 

- obliquum 18 

- robustum 18 

-- ternatum 18 

Schoenoplectus 40, 41, 80,82, 85 

- subgen. Bolboschoenus 85 

- sect. Bolboschoenus 85 

- fluviatilis 87 

- lacusrtis 41. 82 

- maritimus 89 

- mucronatus 4l 

novae-angliae 90 

- robustus 89 

- tabernaemontani 82 

- triqueter 82 

Scirpus 40-44, 50 52, 57-59, 67 -69,73, 

75,80-82,86,88 

'- subgen. Bolboschoenus 85 

- subgen. Euscirpus 43,44,49 

- subgen. Japonici 65 

-- subgen. Phyllothryon 85 

- subgen. Scirpus 43, 69 

-- subgen. Taphrogeton 43, 69 

'- sect. Actinoscirpus 43, 5l 

- sect. Androcoma 5l 

-- sect. Androcoma auct. 50 

- sect. Baeothryon44 

- sect. Beetlea 42,50,52,63 

- sect. Bolboschoenus 43, 81, 85 

- sect. Dichostylis 43 

- sect. Euscirpus 43, 69 

-- sect. Filipes 62 

- sect. Heterogenes 42, 50 

- sect. Japonici 42, 50,65,68 

- sect. Lineatus 50. 5l 

293

- sect. Maritimi 81, 85, 86, 88 

- sect. Paniculatocorymbosi 42, 4'1, 49, 

50. 65. 69 

- sect. 47 

- sect. Phyllantheli 8l 

- sect. Pseudoeriophorum 65, 68 

- sect

- - var. wichurae 52, 54 

- - f. borealis 52 

--- f. concolor 52, 55 

- -- f. cylindricus 52, 55 

- wichurae auct. 58 

Scrophulariaceae trib. Antirrhineae I 82' 

195 

Seidlia 44, 69 

- atrovirens 74 

radicans 70 

sylvatica 73 

Selaginella 22-24 

- subgen. Ericetorum 23 

- subgen. HeterostachYs 23 

- subgen. Selaginella 22,23 

- subgen. Stachygynandrum 23, 24 

-- subgen. TetragonostachYs 23, 26 

--- sect. Rosulatae 25 

- subser. Rosulatae 25 

- aitchisonii 25 

- borealis 25 

- deflexa22 

-- helvetica 24 

- involvens auct.25 

- mongholica var. rossii 25 

- rossii 25 

- rupestris 23 

- - var. shakotanensis 26 

- - f. siberica 26 

- rupestris auct.26 

- - sanguinolenta 25 

- selaginoides 22, 24 

- siberica 26 

spinosa 22, 24 

- tamariscina 25 

Selaginellaceae 22, 24 

Selaginellales 22 

Selaginellopsida 22 

Sideritis 178,179 

- sect. Hesiodia 178 

- sect. Marrubiastrum 178 

- sect. Sideritopsis 179 

- balansae 178 180 

macrostachyos 178 

- woronowii 179 

Sphaeranthus 213, 214, 216, 218, 221 

- subgen. Pseudosphaeranthus 215 

-- subgen. Sphaeranthus 214 

-. sect. Cristati 217 

sect. Cuspidella 216 

-- sect. Cylindrocephali 215,216 

- sect. 215 

--sect. Oocephali 216 

- sect. (Oocephalae> 216 

- sect. Oppositifolii 216 

- sect. Pauciflori 216, 217 

-- sect. Platycephalae 216 

- sect. Polycephalos 215 

- sect. Sphaeranthus 214 

- $ Multiflori 216 

- $ Pauciflori 216 

- africanus 214,215 

- amaranthoides 215, 216 

- angolensis 215 

- angustifolius 216 

-- bullatus 214 

- chandleri 215 

-- confertifolius 214, 215 

- cristatus 217 

- cyathuloides 216 

- epigaeus 216 

- frrscheri 216 

- flexuosus 215 

.- foliosus 216 

- gazaensis 214 

- greenwayi 215 

- humilis 215 

- incisus 216 

- indicus 214 

kirkii 216 

- mimetes 216 

- napierae 215 

-- neglectus 216 

- oppositifolius 217, 221 

- peduncularis 216 

-- peguensis 216 

- polycephalus 215 

- randii 216 

salinarum 216 

- senegalensis 214 

- spathulatus 216 

sphenocleoides 215 

-- steetzii 216 

strobiliferus 2I3,216 

-'- stuhlmannii 216 

-- suaveolens 215 

talbotii 216 

-- ukambensis 215 

- variabilis 215 

volgensis 213 

zavattarii 216 

Spiraea pjassetzkii 284 

- prostrata 284 

Stachygynandrum 24,25 

- sanguinolentum 25 

tamariscinum 25 

29s

Stachys 178,179 

- sect. Campanistrum 180 

- sect. Pseudosideritopsis lEO 

-.sect. Sideritopsis 178, 179, 180 

- arabica var. minor l8l 

- iberica 178 

- obscura 180 

- pseudosideritis 179 

- sparsipilosa 178 

- woronowii 178, 179, 180 

- zoharyana 180, l8l 

Taphrogeton 44, 69 

Tetragonia 284 

- philippi 284 

- robusta 284 

Tithymalus 159 

- subgen. Murketias 160 

- anacampseros 

166 

- armenus 163 

- broteri 163 

- corsicus 167 

- craspedius 

167 

- fontquerianus 166 

-- marschallianus 164 

-- marschallianus auct. 163 

-- monostylus 163 

- myrsinites 165 

- rechingeri 167 

- rigidus 162 

- spinidens 

165 

- veneris 162 

Trichanthemis glabriflora 232 

- karataviensis 232 

Trichophorum 40, 41, 44, 50, S7 

- alpinum 44, 58 

- cyperinum 58 

- lineatum 50, 58 

Trollius komarovii 230 

Typha sect. Bracteolatae 223 

- sect. Rohrbachit223 

- subsect. Rohrbachia 223 

- minima 223 

Veronica 209 

- sect. Pseudolysimachium 209 

-- laeta 210 

- porphyriana 210 

-- smirnovii 209.211+ 

- spicata subsp. kamelinii 209 

Vicia abbreviata 226 

-- -- subsp. balansae 226 

- balansae 226 

- grossheimii 226 

-- tenuifolia 226 

-- - subsp. subalpina 226 

-- truncatula 226 

-- - subsp. balansae 226 

- variabilis 226 

-- - var. subalpina 226 

Xanthium 220

AJIOABI,ITHbIft YKA3ATE.IIb 

HOBbIX HA3BAHI{fi TIXCOHON 

INDEX ALPHABETICUS TAXORUM 

NOMINUM NOVORUM 

Acantholimon linczevskianum Lazkov sp. nova 38 

Allium aegyptiacum (Tiickholm et Drar) Seregin comb. et stat. nov. l0l 

- cepa L. var. schallot Seregin nom. nov. 102 

durandoi (Batt. et Trabut) Seregin comb. et stat. nov. l0l 

- haemostictum (Stearn) Seregin comb. et stat. nov. 102 

- junceum Smith var. tridentatum (Kollmann, N. Ozhatay et Koyuncu) Seregin comb. 

et stat. nov. 101 

- parviflorum (Kollmann) Seregin comb. et stat. nov. l0l 

- phanerantherum Boiss. et Hausskn. var. deciduum (Kollmann et Koyuncu) Seregin 

comb. et stat. nov. l0l 

- porrum L. var. kurrat (Schweinf. ex K. Krause) Seregin comb. et stat. nov. 102 

- psebaicum Mikheev sp. nova 96 

sanctum Scregin nom. nov. 102 

- - var. negevense (Kollmann) Seregin comb. nova 102 

- staticiforme Smith subsp. flexuosum (d'Urv.) Seregin comb. et stat. nov. 102 

- vanense (Kollmann et Koyuncu) Seregin comb. et stat. nov. 102 

- villosiusculum Seregin nom. nov. 102 

Arabis hirsuta L. var. sagittata (DC.) A. Zernov comb. nova 204 

Astragalus karelinianus M. Pop. var. violaceus Knjasev et Kulikov var. nova 132 

- pseudomacropus Knjasev et Kulikov sp. nova 142 

Bolboschoenus (Aschers.) Palla sect. Browningia Tatanov sect. nova 86 

- sect. Mediani Tatanov sect. nova 90 

- subsect. Affrnes Tatanov subsect. nova 89 

- subsect. Fluviatiles Tatanov subsect. nova 87 

- subsect. Glauci Tatanov sect. nova 87 

-- subsect. Popoviorum Tatanov subsect. nova 89 

Botrychium alaskense var. pavlovii Tzvel. var. nova 14 

- lunaria var. przevalskii Tzvel. var. nova 17 

Carduus adpressus C. A. Mey. subsp. novorossicus (Portenier) A. Zernov comb. nova 

227 

Cephalanthera Rich. sect. Cucullatae A. Zernov sect. nova 223 

Coronilla orientalis Mill. subsp. balansae (Boiss.) A. Zernov comb. et stat. nov. 225 

Crambe L. sect. Tatariae (Khalilov) A. Zernov comb. et stat. nov. 225 

Festuca tzveleviana Lazkov sp. nova 28 

Fraxinus excelsior L. var. oxycarpa (Willd.) A. Zernov comb. et stat. nov. 226 

297

Gladiolus murgusicus Mikheev sp. nova 98 

lnula orientalis Lam. subsp. grandiflora (Willd.) A. Zernov comb. et stat. nov.227 

Leutea sclerophylla (Boiss. ct Hausskn.) V. Vinogr. comb. nova 174 

- turcomanica (Schischk.) V. Vinogr. comb. nova 174 

Linaria Mill. sect. Cretaceae Pesskova sect. nova 197 

. sect. Linaria subsect. Laeves Kuprijan. ex Pesskova subsect. nova 190 

--. genistifolia (L.) Mill. var. parviflora (Albov) A. Zernov comb. nova 227 

Lycopodioides Boehmer ex Ludwig subgen. Tetragonostachys (Jermy) Tzvel. cornb. 

nova 26 

- sect. Rosulatae (A. Br.) Tzvel. comb. nova 25 

-- sect. Stachygynandrum (P. Beauv. ex Mirbel) Tzvel. comb. nova25 

- aitchisonii (Hieron.) Tzvel. comb. nova 25 

- rossii (Baker) Tzvel. comb. nova25 

-- shakotanensis (Franch. ex Takeda) Tzvel. comb. nova26 

-- siberica (Milde) Tzvel. comb. nova26 

-- tamariscina (P. Beauv.) Tzvel. comb. nova 25 

Phlomoides adylovii Lazkov sp. nova 32 

kurpsaica Lazkov sp. nova 30 

Polygonum L. subgen. Bistorta (Scop.) A. Zernov comb. et stat. nov. 224 

-- -- sect. Vivipara (Tzvel.) A. Zernov comb. nova 224 

Primula pskemensis Lazkov sp. nova 36 

Rosa glabrifolia c. A. Mey. ex Rupr. var. pubescens Buzunova et Kamelin var. nova 120 

Ruscus L. sect. Platyruscus (A. Khokhr. et v. Tichomirov) A. Zernov comb. et star. nov. 

223 

Saxifraga moschata Wulf. var. pontica (Albov) A. Zernov comb. et stat. nov. 225 

-- subverticillata Boiss. subsp. colchica (Atbov) A. Zernov comb. et stat. nov. 225 

Scabiosa bipinnata K. Koch var. calcarea (Albov) A. Zernov comb. nova227 

Scirpus L. sect. Beetlea Egorova sect. nova 50 

sect. Ternatani Egorova sect. nova 62 

Sideritis L. sect. Sideritopsis (Bhattacharjee) Krestovsk. comb. nova 179 

Sphaeranthus L. sect. Cristati Tzvel. sect. nova 2l'1 

-- sect. Oppositifolii Tzvel. sect. nova 216 

Spiraea pjassetzkii Buzunova nom. nov. 284 

Stachys L. sect. Pseudosideritopsis Krestovsk. sect. nova 180 

Tetragonia philippi Byalt nom. nov. 284 

Typha L. sect. Rohrbachia (Kronf. ex Riedl) A. Zernov comb. et stat. nov. 223 

Veronica smirnovii Kossaczev et D. German sp. nova 209 

vicia tenuifolia Roth subsp. subalpina (Grossh.) A. Zernov comb. et stat. nov. 226 

- truncatula Fisch. ex Bieb. subsp. balansae (Boiss.) A. Zernov comb. nova226

Новости систематики высших растений Том 36 2004 ОТ ...

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