Some caddis flies (Trichoptera) in western Norway, and - Norsk ...

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The Editorial Board and the Publisher much regret the very delayed printing date of this issue. The next one 

(Vol. 26 No. 2) will appear in the first months of 1980.

A Letter from the Editor 

With the present issue a new periodical commences. 

However, in order to indicate that this is not entirely 

a new publication, but a continuation of the previous 

Norwegian Journal of Entomology (Norsk Entomologisk 

Tidsskrift) the volume numbering is retained 

and the old names will appear as sub-headings. 

The change is not the result of a lack of manuscripts 

or interest in the old journal. On the contrary, 

following a period of 58 years as a journal, the periodical 

had obtained a high quality in as well content as 

layout. Particularly the last 12 years under the editorship 

of docent Lauritz S0mme has seen a great expansion 

in numbers of pages printed and in areas covered. 

As an acknowledgement of the great contribution 

made to the journal and to Norwegian entomology 

by Or. S0mme one of his favourite research animals, 

the carabid Pelophila borealis, is retained as a 

frontispiece. 

The expansion of the journal as well as the improvement 

of its layout contributed to the downfall of 

the journal. The expenses increased significantly. 

These were largely covered by contribution from the 

Norwegian Research Council (NAVF), and their recent 

policy has been to support internordic journals. 

With the erection of the new journal Holarctic Eco 

Ole A. Sa:ther 

Editor, Fauna Norvegica ser. B 

logy and the previous Oikos and Entomologica Scandinavica 

much of the need for publishing entomological 

papers in ecology and systematics would be covered. 

However, there still were no place, even on an 

interscandinavian basis, where to publish faunistical 

and zoogeographical papers. In order to satisfy this 

need a new larger faunistical journal, Fauna Norvegica, 

were created and divided into three parts or series 

(entomology series B, ornithology series C, and 

other groups series A.) 

Fauna Norvegica ser. B will emphasize papers 

which are mainly faunistical and zoogeographical in 

scope or content, including checklists, faunal lists, 

type catalogues and regional keys. 

It is obvious, however, that as well ecological information 

as some taxonomical details belong to the 

documentation data of faunistical papers. Otherwise 

these would be merely faunal lists. 

It is a sad and strange coincidence that we in the 

first issue can give an obituary (written by prof. emerit. 

Hans Kauril of the leading zoogeographer of 

Scandinavia for the last decades, Or. Carl H. Lindroth. 

Let ut hope that this new journal will get the 

honour of publishing articles of the same scope and 

vision as many of Dr. Lindroth's papers.

l 

6L6I - S06I QlOJpUn oH IJB3

Carl H. Lindroth d0de den 23. februar i Lund. 

Med hans bortgang har entomologisk og zoogeografisk 

forskning mistet en av sine forgrunnsfigurer. 

Han ble f0dt i Lund i 1905, studerte zoologi 

i Stockholm og Uppsala, og disputerte for 

doktorgraden i Uppsala i 1932. Han tjenestegjorde 

som lektor i biologi, ble dosent ved Stockholms 

h0gskole i 1947 og ble utnevnt til professor 

i entomologi i Lund i 1951. 

Hans vitenskapelige produksjon er meget stor 

og omfatter over 160 arbeider. Noen av de viktigste 

skal nevnes her: Die Insektenfauna Islands 

und ihre Probleme, 1931 (doktoravhandling); 

Die Fennoskandischen Carabidae I-IJI, 1945-49; 

Inheritance of wing dimorphism in Pterostichus 

anthracinus Ill., 1946; The fauna history of 

Newfoundland, 1963; The faunal connections 

between Europe and North America, 1957; The 

ground-beetles of Canada and Alaska, 1961-69; 

Skaftafell, Iceland, a living glacial refugium, 

1965; Surtsey, Iceland; the development of a 

new fauna; Terrestrial invertebrates. 1973. 

Ett av Lindroths vitenskapelige hovedmaJ bJe 

meget tidlig klarlagt. Allerede i sin avhandling 

over Islands fauna anser han overvintringsmulighetene 

for deler av faunaen under den siste istiden 

for a va;re et av de viktigste problemene, 

og i sin store monografi over fennoskandiske carabider 

(del 3, 1949) nevner han som arbeidets 

hovedsp0rsmat: «die postglaziale - und moglicherweise 

die glaziale - Geschichte der fennoskandischen 

Fauna». Ingen bedre enn Lindroth 

selv forsto hvor komplisert et slikt problem kan 

va;re, omfattende en hendelse som ligger tusener 

av ar tilbake i tiden. Derfor anvendte han ogsa 

hele den metod910giske apparatur som star til 

disposisjon for en biolog og zoogeograf for a 

samle og klarlegge bevis. Han hadde en klar 

malsetting og vek ikke unna hverken for anstrengelser 

eller meget tidskrevende oppgaver. 

Det gjaldt f0rst a renske opp i det taksonomiske 

feltet for a va;re sikker pa a kunne arbeide 

med de virkelige enhetene - artene. Dette gjaldt 

mindre den nordeuropeiske, men f0rst og fremst 

den nordamerikanske og kanadiske faunaen. 

Dette store arbeidet munnet ut i en omfattende 

monografi over Canadas og Alaskas jord 

10pere i 6 bind, dertil et antall mere frittstaende 

skrifter. Senere fortsatte han med kartleggingen 

av utbredelsen, artenes spredningsmekanismer 

og aut0kologiske egenskaper. 

Den 0kologiske siden av sp0rsmalet unders0kte 

han igjen pa Island. I Skaftafell ble de 10kalklimatiske 

for hold i na;rheten av breen 

unders0kt, for pa den maten a rekonstruere 

landisens innvirkning pa insektfaunaen i de 

na;rliggende populasjonene (1965). 

Det viktigste argumentet i diskusjonen om refugieproblemet 

viste seg a va;re utbredelsen av 

de vingedimorfe carabideartene. Vingedimorfismen, 

kjent allerede av Paykull og Gyllendal, var 

genetisk bundet. Lindroth Iykkedes i a klarlegge 

dens genetiske karakter (1946). Ved kartleggingen 

av disse artenes utbredelse kom det fram 

at hos visse arter hadde den ikke flygedyktige 

kortvingete formen spredd seg til omrader der 

den mer lettspredde langvingete delen av populasjonen 

ikke hadde nadd fram. Et forhoJd som 

vanskelig lot seg forklare pa annen mate enn at 

disse har overlevet nedisingen, eller en del av 

den perioden pa selve stedet. En verdifull sammenfatning 

om dette problem er publisert i Ambio, 

1972. 

Lindroths forskning og vitenskapelige problemstilling 

ble hurtig utvidet. Problematikken 

om kolonisering av nye omrader og faunainnvandringsproblem 

kom i forgrunnen. Han har 

ofret spesialstudier pa disse problem i flere omganger. 

sa;rlig i forbindelse med forskningarbeidet 

pa Newfoundland og Nord-Amerika. Ballastens 

betydning fra England ble oppdaget som en 

viktig faktor for overf0ringen av europeiske arter 

til Newfoundland og Nord-Amerika. Den 

nye vulkan0ya Surtsey's kolonisering ble en velkommen 

anledning til a kunne f01ge opp innvandringen 

av dyre- og vekstarter fra begynnelsen 

av pa en nydannet 0Y. J dette tilfelle utf0rte 

han et betydningsfullt eksperimenteJt arbeid for 

a teste havstmmmenes og vindenes betydning, 

saltvannets innvirkning pa landorganismenes 

overleving, avstandens rolle til basisomradet, 

samt artssammensetningen i basisomradet 

(1973). 

Hans zoogeografiske unders0kelser som er 

nevnt ovenfor, ble ikke begrenset til Europa, 

men ble ogsa utvidet til Nord-Amerika, Alaska, 

Kodiak og Aleutene. Dette gjaldt fmst og fremst 

den nearktiske faunas forbindelser til palearktis, 

BeringJandets rolle ved utveksling av faunakomponentene 

(1969), levninger fra den siste interglasiaJperiodens 

insektfauna m.m. 

Selv om den faunahistoriske forskningen pa 

denne maten liksom dannet et midtpunkt for 

hans vitenskapelige virksomhet, ble hans vitenskapelige 

palett av problemstillinger og forskningsresultat 

adskillig mer omfattende. Med et 

klart syn for taksonomiske problem har han utredet 

en lang rekke av mangfoldige spmsmal vedmrende 

sin egen spesialgruppe, Carabidae. 

Han appliserte 0kologisk, sa;rskilt eksperimen 

3

tell metodikk i sine zoogeografiske studier og ble 

pa den mMen en av grunnleggerne av 0kologisk 

zoogeografi i Norden. 

Med vaken interesse fulgte han entomologiens 

utvikling fomvrig og sto i mer kontakt med 

kollegene pa omradet. Om dette vitner den lange 

serien av artikler og biografier om nordiske entomologer. 

Han ble reres- og korresponderende 

medlem i en mengde lrerde ·sammenslutninger 

verden over og fikk motta mange internasjonale 

utmerkelser. 

Carl Lindroths vitenskapelige bane ble ikke 

den letteste. Etter doktordisputasen tjenesteg 

4 

jorde han som gymnaslektor pa ulike steder i 18 

ar. For hans forskning innebar dette nattarbeid 

og feriejobbing. 

Som lrerer var han inspirerende og avholdt. 

Hans lrerebok i entomologi er en vakker gave til 

ettertidens studentgenerasjoner. Som populrervitenskapelig 

forfatter og foredragsholder var 

han alltid inspirerende og fengslende, 

Med Carl Lindroths bortgang mistet fagkollegene 

en stimulerende radgiver og venn. For 

hans venner er tapet smertelig. 

Hails Kauri

New records of Coleoptera in Norway 

JOSTEIN ENGDAL AND KARL ERIK ZACHARIASSEN 

INTRODUCTION 

Engdal, J. & K. E. Zachariassen 1979. New records of Coleoptera in Norway. Fauna Norv. 

Ser. B, VoI. 26, 5-7. 

This paper presents alist of 65 records of Coleoptera not previously reported from the respective 

Norwegian districts. One of the species, Masoreus wetterhalli Gyllenhal. is new to 

Norway. The new records are the northernmost in Norway for 12 of the species. 

J. Engdal and K. E. Zachariassen, University of Trondheim, Departement of zoology, Rosenborg, 

N - 7000 Trondheim, Norway. 

In this paper 65 records of beetles are given. 

They all extend the distribution areas of respective 

species, according to Lindroth (J 960) and later 

supplement lists. The records are mainly 

from the provinces of M0re og Romsdal and 

S0r-Tf0ndelag. The district abbreviations are according 

to Strand (J 943). The records were 

made during the years 1972 - 1978, and the material 

is kept in collections of the authors. The 

list is not a result of a systematic collection within 

the areas, but represents more casual finds. 

The main methods used for collecting were 

sweep-netting and picking by hand. 

OBSERVAnONS 

Notiophi/us palustris Duftschmid TEi: Notodden, 

30.8.76. One specimen found in country 

courtyard. I 

Bembidion aeneum German NTy: Ofstadsj0en, 

Vikna, 24.6.76. One specimen under plank at 

seashore. 

Bembidion bruxellense Wesmael NTy: R0rvik, 

Vikna, 25.6.77. One specimen under stone 

on a gravelly grasfield near the seashore. 

Bembidion femoratum Sturm. STy: Slettvik, Agdenes, 

16.6.78. One specimen under stone. 

"Trechus micros Herbst. STy: Rissa, 20.5.78. 

One specimen under stone at seashore. 

Dichirotrichus pubescens v. Paykull. NTy: R0rvik, 

Vikna, 29.6.77. Several specimens under 

stones near the seashore. 

Masoreus wetterhalli Gyllenhal YE: Stavern, 

8.7.78. One specimen under plank on sandy 

shore. 

Lebia chlorocephala HofTmansegg TEi: Notodden, 

30.8.76. One specimen found in meadow 

field. 

Fauna Norv. Ser. B 26, 5-7. Oslo 1979. 

Liodes picea Panzer NTi: Binde, Snasa, I. 9.77. 

One specimen found in meadow field. 

Hister striola C.R.Sahlberg STi: Lade, Trondheim, 

15.5.77. One specimen in pit-fall trap. 

"Cantharis pellucida Fabricius STi: Byneset, 

Trondheim, 20.5.74. One specimen caught 

with sweep-net in meadow. 

Xylobius corticalis v. Paykull VAy: S0gne, 

10.7.78. One specimen under bar on stem of 

oak, Quercus sp. 

Agri/us viridis L. MRi: Verma, 22.7.78. Five 

specimens caught with sweep-net in Salix sp. 

Pityophagus ferrugineus L. STy: Slettvik, Agdenes, 

16.6.78. Several specimens on ground 

under log of pine, Pinus sylvestris. 

" Si/vanus bidentatus Fabricius NTi: Nordli, Lierne, 

24.6.78. One specimen on pile of planks 

at saw mill. 

Dendrophagus crenatus v. Paykull STy: Rissa, 

20.5.78. One specimen under bark on log of 

elm, Ulmus glabra. 

Cerylon histeroides Fabricius STy: Rissa, 

20.5.78. Two specimens under bark on log of 

elm, Ulmus glabra. 

Coccinella hieroglyphica L. STy: Rissa, 20.5.78. 

One specimen caught in flight in sunshine. 

Paramysia oblongoguttata L. STy: Rissa, 

20.5.78. One specimen caught in flight in 

sunshine. 

Chi/ocorus bipustulatus L. TEi: Notodden, 

30.8.76. One specimen caught with sweepnet 

in aspen, Populus tremulae. 

Dorcatoma dresdensis Herbst VAy: BOf0Y, 

S0gne, 9.7.78. One specimen caught in flight 

in the evening. 

Orchesia micans Panzer VAy: BOf0Y, S0gne, 

14.7.78. Several specimens found in tree 

brackets on a trunk. 

Bolitophagus reticulatus L. NTi: Binde, Snasa, 

1.9.77. Several specimens found in tree 

brackets on birch. 

5

Aphodills llel110ralis Erichsson AAy: Laget, Ri One specimen caught with sweep-net in mea 

S0r, 20,7,78. Several specimens in excredow.ments of elk, Alces alces. 

Apionlo/i Kirby MRy: Molde, 4.8.76. One speci 

Tricl/ius fascia/us L. STy: Rissa, 20.6.74. One men caught with sweep-net in meadow. 

specimen caught on thistle, Circilll11 he/erop 

Apiol1 simile Kirby MRy: Molde, 4.8.76. One 

hylllllJl. 

specimen caught with sweep-net in meadow. 

Ce/ollia aura/a L. STy: Rissa, 20.6.74. One spe 

Apion virens Herbst MRy: Molde, 4.8.76. Three 

cimen caught in flight in sunshine. 

specimens caught with sweep-net in mea 

Criocephallls rlls/icus L. HEn: Akrestmmmen, dow. 

24.7.78. One specimen under plank at saw *Po/ydrosus cavinlls L. MRi: Verma, 22.7.78. 

mill. 

One specimen caught with sweep-net in 

Toxo/lIs cursor L. STy: Hambara, Agdenes, birch, Be/ula pllbescens. 

13.6.75. One specimen caught on HieracilllJ1 Po/vdrosis mol/is Strom. STy: Rissa, 20.5.78. 

sp. 

One specimen caught with sweep-net in 

Semano/lIs unda/us L. STi: Jonsvannet, 2.6.78. birch, Be/u/a pllbescells. 

One specimen on pile of planks. 

Polydroslls n!ficomis Bonsdorff MRy: Molde, 

*Cly/us arie/is L. STi: Byneset, Trondheim, 4.8.76. One specimen caught with sweep-net 

1.7.78. One specimen on trunk of deciduous in deciduous forest. 

tree. 

•Barypi/hes pe/lllcidlls Boheman MRy: Molde, 

STy: Hambara, Agdenes, 30.6.77. One specimen 3.8.76. One specimen caught with sweep-net 

caught with sweep-net in meadow. 

in deciduous forest. 

*Leioplls nebuloslls L. STi: Byneset, Trondheim, S/rophosomus melanogramllnfs Forster STi: 

3.7.78. One specimen on log of deciduous Lade, Trondheim, 15.6.72. One specimen in 

tree and one specimen hatched from the same pitfall trap. 

log. 

• Sitona pllncticol/is Stephens MRy: Molde, 

Donacia aqlla/ica L. STy: AfJord, 3.6.78. One 4.8.76. One specimen caught with sweep-net 

specimen on Salix sp. on river bank. 

in meadow. Kortgard, 20.8.73. 

MRi: Verma, 27.7.78. One specimen caught 

with sweep-net in Carex sp. near river. 

• SilO/la slllftcrollS Thunberg MRy: Molde, 

4.8.76. Two specimens caught with sweep 

Donacia vulgaris Zschach. STi: Langtj0rna, net in meadow. 

Femunden, 2.8.77. One specimen on floating • SilO/la sU/llralis Stephens MRy: Molde, 4.8.76. 

water-vegetation. 

Two specimens caught with sweep-net in 

Plateumaris disc%r Panzer STy: Hambara, Ag meadow. 

denes, 15.6.74. One specimen on Carex sp. Dorytomus /aellia/llS Fabricius STy: Rissa, 

near water. 

20.5.78. One specimen caught with sweep 

MRi: Verma, 31. 7.78. One specimen caught net in sallow, Salix caprea. 

with sweep-net in Carex sp. near river. EI/escus bipunc/a/lIs L. STy: Rissa, 20.5.78. One 

Pla/eumaris sericea L. MRi: Verma, 25.7.78. specimen on sallow, Salix caprea. 

Four specimens caught with sweep. net in Micco/rogus picirostris Fabricius MRy: Molde, 

Carex sp. near river. 

4.8.76. One specimen caught with sweep-net 

Plty/odec/a pallidus L. MRy: Molde, 3.8.76. One in meadow. 

specimen caught with sweep-net in sallow, Pissodes gyl/enhali Gyllenhal STy: Rissa, 

Salix caprea. 

20.5.78. One specimen on logs of spruce, Pi 

Galerucel/a sagillariae Gyllenhal STi: Jonsvancea abies. 

net, 7.6.78. Several specimens on water-ve STi: Jonsvannet, 7.6.78. One specimen on pile of 

getation. 

•Lochmaea su/uralis Thomson STy: Hambara, 

Agdenes, 13.6.75. One specimen caught with 

sweep-net in Salix sp. 

Apioll cur/irostre Germar MRy: Molde, 4.8.76. 

planks. 

1 

Hylobius picells DeGeer STy: Slettvik, Agdenes, 

16.6.78. Several specimens on ground under 

log of pine, Pinus sylves/ris. 

I 

•Liosoma deflexllm Panzer STi: Lade, Trond 

Three specimens caught with sweep-net in heim, 22.5.77. Two specimens caught in pit 

meadow. 

fall trap. 

Apion ervi Kirby MRy: Molde, 4.8.76. One spe Zacladus afftllis v. Paykull MRy: Molde, 4.8.76. 

cimen caught with sweep-net in meadow. One specimen caught with sweep-net in mea 

Apionface/um Gyllenhal MRi: Verma, 24.7.78. dow. 

6

Cionus scrophulariae L. STy: Hambara, Agdenes, 

15.6.74. One specimen caught with 

sweep-net in meadow. 

Scoly/us ra/zeburgi E.W.Janson STi: Budal, 

6.7.77. - 5.8.77. 21 specimens caught in 

swarm-traps. 

* Records representing the northernmost find of 

the species. 

DISCUSSION 

Of the above mentioned records, 12 are the northernmost 

for the species in Norway. Most of 

these records represent extensions of the northern 

borders of the known areas of distribution 

of the different species along the coast from the 

province of Hordaland to the province of Sor 

".-- Tf0ndelag. 

However, three of the species are found far 

outside their previously known areas of distribution, 

thus deserving further comments. 

Silvanlls biden/a/lIs is previously recorded 

from the southern parts of Sweden and in Norway 

only from the province of Akershus. The 

specimen from Nordli was found in a saw mill, 

and has probably been transported with timber 

imported from Sweden. 

Leiopus nebulosus is previously known from 

the districts around the Oslo fiord, and west to 

the province of Rogaland. It is possible that the 

species is distributed along the western coast of 

Norway, but that it has been overlooked. If not, 

the long distance between the previously known 

distribution area, and the locality of the present 

find, might indicate that Byneset supports a relict 

populatiol} from the post-glacial warm age. 

, Byneset and the adjacent northern parts of 

Gauldalen valley may support several relict spe- 

I 

cies. Dolmen et at. (] 975) have investigated the 

distribution of Odonata. They have found a distribution 

for three species of damselflies similar 

to that found for L. neblllosus. Further investigations 

may reveal that more species with a predominantly 

southern distribution are established 

in this area. 

Si/ona pllnc/icollis is up to now in Scandinavia 

known to about 600 N. The present collecting 

sites are situated considerably farther north 

(nearly 63 0 N). Since the species in Norway previously 

is known only from the Oslo area, it 

might have reached the western coast via the 

Gudbrandsdalen and Romsdalen valleys, or it 

might have been introduced by human activities 

Masorells wetterllalli is a new species to Norway, 

and its record is published separately (Zachariassen 

1979). 

REFERENCES 

Dolmen, 0 .. Srether. B. & Aagaard, K. 1975. Ferskvannsbiologiske 

undersokelser av tjonner og evjer 

langs elvene i Gauldalen og Orkdalen, Sor-Trondelag. 

K. Ilorske Vide/1.sk. Se/sk. MlIs. Rapp. 

Zoo/. Ser 1975 - 5. 47 pp. 

Lindroth, C. H. (ed.l 1960. Cata/oglls Co/eopterorlll11 

FellllOscalldiae et Dalliae. Entomologiska sallsk.. 

Lund. 

Strand. A. 1943. Inndeling av Norge til bruk ved faunistiske 

oppgaver. Norsk ell/. Tidsskr. 6. 208 

224. 

Zachariassen, K. E. 1979. Masoreus wellerhalli Gyll. 

(Col.. Carabidae) ny art for Norge. Faul1a NorI'. 

ser. B 26, 

Received 20 May 1979. 

7

Noen funn av Coleoptera fra Trondelag og More 

DAGFINN REFSETH 

Refseth, D. 1979. Noen funn av Coleoptera fra Tmndelag og M0re. Fauna Norv. Ser. B, 26. 

8-9. 

Twenty-eight new records of Coleoptera are given from parts of the provinces of M0re og 

Romsdal, S0r-Tmndelag and Nord-Tmndelag, Central Norway. Most of the species also occur 

in neighbouring areas, but for Trechus rivu/aris Gyllenhal and Cholevafagniezi Jeannel 

the new finds represent a considerable extension of the distribution areas in Norway. For T. 

rivu/aris a possible connection with the Swedish population is indicated. 

Dagfinn Refseth, Zoologisk Institutt, Universitetet i Trondheim, Rosenborg, N-7000 Trondheim. 

1 denne artikkelen presenteres 28 funn av biller 

som tidligere ikke er registrert i de respektive 

omradene i M0re og Romsdal, S0r-Tmndelag og 

Nord-Tmndelag (Lindroth 1960, Strand 1970, 

1977, Zachariassen 1977). Funnene er gjort delvis 

ved tilfeldige innsamlinger og delvis ved systematiske 

fangster med fall feller. Blant annet 

har Dag Dolmen stilt til disposisjon materiale 

innsamlet i forbindelse med \BP's inventeringer i 

edell0vskogslokaliteter i M0re, Tmndelag og 

Nordland. 

Materialet oppbevares delvis hos forfatteren, 

delvis ved DKNVS, Museet, Trondheim. 

Omradebetegnelsene er i henhold til Strand 

(1943), med senere korreksjoner av kommunegrenser. 

Carabus nel110ralis Muller. STy: Agdenes, juni 

1975. 

Notiophilus biguttatus Fabricius. MRy: Molde, 

Am, 29 juli 1974. 

Blethisa multipunctata L. STy: Agdenes, Lillevann, 

8 juni 1975. 

Trechus rivularis GyllenhaJ. NTi: Snasa, juJi 

1972 (Leg. D. Dolmen). To eksemplarer ble 

tatt i fall feller pa en fuktig skogslokaJitet med 

svartor (Alnus glutinosa (L.». Arten er sterkt 

bundet til fuktige, gjerne myrlendte, 10Vskogsbiotoper 

og er i Norge tidJigere bare 

funnet pa noen fa lokaliteter i Akershus (Lindroth 

1945, 1960). Arten har et 0stlig utbredelsesm0nster 

i Fennoskandia og forekommer 

i Sverige spredt opp til Norrbotten (Lindroth 

1945). Flygevingene er hos de fleste individene 

reduserte, og selv om makroptere 

individer finnes, tyder den begrensede utbredelsen 

og artens stenotope karakter pa at aktiv 

flyging spiller liten rolle ved spredning 

8 

(Lindroth 1945). Eksemplarene fra Smisa 

mangler flygevinger. og funnet kan derfor 

neppe skyldes nylig innvandring fra 0S1. Men 

den relativt korte avstanden til mermeste 

funnlokalitet i Sverige kan tyde pa at forekomsten 

har sin opprinnelse i den svenske 

popu lasjonen. 

Trechus secalis Paykull. MRi: Rindal, Dalsegg, 

juli 1972 (Leg. D. Dolmen). 

Patrobus assil11i1is Chaudoir. NTi: Stod, 29 juni 

1976. 

Harpalus quadripunctatus Dejean. MRy: 

Molde, 29 juli 1975. 

Harpalus ru./ipes DeGeer. MRy: Molde, Am, 4 

juli 1978. 

Al11ara apricaria Paykull. MRy: Molde, Am, 4 

juli 1978. 

Al11ara aulica Panzer. MRy: Molde, Am, 4 august 

1976. 

Amara bifrol1S Gyllenhal. MRy: Molde, Am, 4 

juJi 1978. 

A l11ara lunicollis Schi0dte. STy: Stadsbygd, 20 

mai 1978. 

Amara nilida Sturm. STy: Rissa, Rams0Y, 20 

mai 1978. 1 

Pterostichus diligens Sturm. MRy: Molde, Am, 

4 juli 1978. 

Agonum piceul11 L. STy: Agdenes, Lillevann, 12 

juni 1975. 

Necrophorus investigator Zettersted1. MRi: Rindal, 

Dalsegg, juli 1972 (Leg. D. Dolmen). 

Necrodes Iilloralis L. STy: Agdenes, Selva, juni 

1974. 

Choleva fagniezi JeanneJ. NTi: Stod, november 

1977, STi: Brekken, 30 September 1978. Arten 

ble tatt i fallfeller i to forskjellige biotoper: 

glissen granskog (Stod) og subalpin bj0rkeskog 

(Brekken). Den ble funnet f0rste gang i 

Fauna Norv. Ser. B 26 .. 8-9. Oslo 1979. 

• 

JI' 

t

" .. 

) I 

Norge i Hordaland i 1968 (Fjellberg 1970) og 

er senere rapportert fra Oppland (Strand 

1977). J Sverige er den spredt utbredt til Halsingland 

(Lindroth 1960). Arten er imidlertid 

neppe sa sjelden som de spredte funnstedene 

tyder pa. J likhet med andre Choleva-arter 

har den gjerne tilhold i smagnagerganger 

(Hansen 1968) og kan derfor vrere vanskelig 

a registrere. 

Catops alpinus GyllenhaL MRi: Rindal, Dalsegg, 

juli 1972 (Leg. D. Dolmen) 

Catops coracinus Kellner. MRi: Rindal, Dalsegg. 


Catops nigrila Erichson. MRi: Rindal, Dalsegg, 


Megatoma undata L NTi: Stod, 8 mars 1977. 

Hippodamia septemmaculata DeGeer. STy: Agdenes, 

8 juni 1975. 

Rhagium mordax DeGeer. STy: Rissa, Asly. 20 

mai 1978. 

Polygraph us poligraphlls L STy: Stadsbygd, 20 

mai 1978. 

Cryptllrglls pllsilllls GyllenhaL STy: Stadsbygd, 

20 mai 1978. 

Dryocoetes alllographlls Ratzeburg. STy: Stadsbygd, 

20 mai 1978. 

REFERENCES 

Fjellberg, A. 1970. Six species of Coleoptera new to 

Norway. Norsk en!. Tidsskr. 17: 105-106. 

Hansen, V. 1968. Biller XXV. Adselbiller, stumpbiller 

mm. Danm. Fauna 77. 

Lindroth, C. H. 1945. Die fennoskandischen Carabidae 

1-11. K. Vet. Vitt. Samh. Hand. (6) 84. Goteborg. 

pp 1-711, 1-279. 

- (ed.) 1960. Catalogus Coleopterorum Fennoscandiae 

et Daniae. Ent Siillsk., Lund. 

Strand, A. 1943. Inndeling av Norge til bruk ved faunistiske 

oppgaver. Norsk ent. Tidsskr. 6: 208-224. 

- 1970. Additions and corrections to the Norwegian 

part of Catalogus Coleopterorum Fennoscandiae 

et Daniae. Norsk ent. Tidsskr. 17: 125-145. 

- 1977. Additions and corrections to the Norwegian 

part of Catalogus Coleopterorum Fennoscandiae 

et Daniae. Second series. Norw. J. Ent. 24: 

159-165. 

Zachariassen, K. E. 1977. Nye funn av Coleoptera i 

Norge. Norw. J. Ent. 24: 147-148. 

Received 1 March 1979. 

9

Notes on the distribution of some Norwegian ant species 

(Hymenoptera, Formicidae). 

PER SVEUM 

Sveum, P. 1979. Notes on the distribution of some Norwegian ant species (Hymenoptera, 

Formicidae) Fauna Norv. ser. B. 26, 10-11. 

New distributional records of eight species occurring in Norway are given. Leptothorax muscorum 

(Nylander), Lasius niger (L.), L. flavus Fabricius, Formica sanguinea Latreille, and 

F. fusca L. occur further north than recorded previously. 

Per Sveum, Saupstadringen 65 B, N-7078 Saupstad, Norway. 

Recent contributions to the ant fauna of Norway 

have been given by Collingwood (1974) 

and Sveum (1978). The species dealt with in the 

present paper have been collected by Dag Dolmen 

(DD), Bodil K. Pedersen (BKP) and the author 

(PS). The nomenclature used follows Collingwood 

(1974). The geographical division follow 

Strand (1943) with names and limits updated. 

Except w hen otherwise stated the species 

are not previously recorded from the provinces 

listed. The material have been deposited in the 

collection of The Royal Norwegian Society of 

Sciences and Letters, The Museum. 

Myrmica rubra (L.) 

MRi: Rindal, Dalsegga 6 July 197\, DD. 

Myrl11ica scabrinodis Nylander 

Nsi: Rana, Hammeren 6 Aug. 1975, PS. The 

only record of the species from Northern 

Norway is from Nsy: Bod0 (Holgersen 1944) 

Sveum (1978) recorded it from: NTy: Leka. 

Although the author has collected several 

years in the province of Nordland, this is the 

only record of the species. The nest was not 

found. The distribution of the known records 

indicate that the species occurs at least as far 

north as Bod0, however it seems to be much 

less common in the north than in the south. 

Myrl11ica lobicornis Nylander 

MRi: EikesdaJ 16 July 1971, DD. 

Leptothorax muscorum (Nylander) 

STi: Trondheim, Lade 7 May \978, PS. Holgersen 

(1944) considered L. l11uscorum as a 

southern species, not exceeding its range 

north of the Dovre Mountains. 

Lasius niger (Ll 

MRi: Rindal, Dalsegga 20 June 1972, DD; 

STi: Orkdal (date unknown), DD, Trondheim, 

inner city 7 May, PS; Frosta, Tautra 29 

10 

May 1978. BKP. The record at Frosta, Tautra 

represent the northernmost record from Norway. 

This is consistent with the assumption 

of Holgersen (1944), although he did not record 

the species north of the Dovre Mountains, 

or give any explanation of his assumption. 

Lasius flavus Fabricius 

NTi: SteinkjeL Sparbu 21 May 1978, BKP. 

The northernmost previous record was 

Trondheim (Holgersen 1944). A single record 

from F0: Polmark (Holgersen 1942) was later 

regarded as doubtful by the same author 

(Holgersen 1944). Accordingly the present record 

should be regarded as the northernmost 

observation from Norway. In Sweden the 

species is found all over the country except in 

the most northern parts (Douwes 1976). 

Formica sanguinea Latreille 

STi: Trondheim. Steinan 14 May 1978. PS. 

This is the first record of the species north of 

the Dovre Mountains. Holgersen (1944) believed, 

without giving any arguments, that the 

species might be found as far north as T[0ndelag. 

It is known that F. sanguinea might found 

its colonies by temporary social parasitism 

and that it is facultative dulotic, The present 

colony was a mixed colony between F. 

sanguinea and F. lemani Bondroit. 1was not 

able to decide what kind of relation excisted 

between the two species. F. lel11ani is distributed 

all over the country. 

Formica fusca L. 

M Ry: Tingvoll, Vulvik 14 July 1971, DD, 

B0fjord, Kallset 22 June 1972, II) act. 1972, 

DD. The species was previously known from 

the southeastern provinces and the western 

coast north to Sogn og Fjordane (SF) (Collingwood 

1974). The present northernmost 

Fauna Norv. Ser. B 26 .. 10-11. Oslo 1979.

• 

.. 

Norwegian record, may indicate a southern 

and coastal distribution. 

ACKNOWLEDG EMENTS 

I wish to thank Mr. Dag Dolmen for permission 

to publish material collected by him when working 

for !BP, and Mrs. Rodil K. Pedersen who 

also contributed specimens. 

REFERENCES 

Collingwood, CA. 1974. A revised lis( of Norwegian 

ants (Hymenoptera, Formicidael. Norsk em. 

Tidsskr. 21. 31-35. 

Douwes, P. 1976. Sveriges myror - illustrerande 

bestamningstabeller over arbetarna. Elltom%gell 

5, 37-54. 

Holgersen, H. 1942. Ants of Northern Norway. 

Trol11so Mus. A'rsh. 63, 1-34. 

Holgersen, H. 1944. Ants of Norway (Hymenoptera, 

Formicidael. Nyu Mag. Naturv. 84, 163-203. 

Strand. A. 1943. Inndeling av Norge til bruk ved faunistiske 

oppgaver. Norsk ell!. Tidsskr. 6, 208-224. 

Sveum, P. 1978. On the biology of ants (Hym., Formicidae) 

in Tremdelag, Norway. Norw. 1. Ell!. 25. 

153-155. 


"

Some caddis flies (Trichoptera) in western Norway, and 

their arrival pattern in light traps. 

TROND ANDERSEN 

Andersen, T. 1979. Some caddis flies (Trichoptera) in western Norway. and their arrival pattern 

in light traps. Faul1a Norwegica, ser. B, 26, 12-17. 

Between 2 and 17 August 1972 3014 specimens of Trichoptera belonging to 42 species 

were collected in two light traps on Oster0y in western Norway. Four species. viz: Oxyelhira 

Irisle/la Klapalek. 1895. Lype phaeopa (Stephens, 1836), Glypholaelius pe/lucidus (Retzius. 

1783), and Phacopleryx brevipellllis (Curtis. 1834). are previously no. recorded from western 

Norway and eight further species are new to outer Hordaland. 

The traps were operated from 21.00 to 05.00 hrs every night. and emptied manually 

every hour. The limnephilids were trapped in greatest number between 23.00 and 03.00 hrs, 

while the non - limnephilids, as a group, were most abundant just after sunset and to some 

degree just before sunrise. However. in Oxyelhiraflavicomis (Pictet. 1834, and Polycel1lropus 

[lavolllaculalus (Pictet, 1834), the two most abundant non - limnephilids in the study. only 

one peak. before midnight. could be discerned. 

Sex differences in the arrival pattern were found in one species. Molal1llodes lil1cla (Zetterstedt, 

1840). The females of this species were trapped in greatest numbers just after sunset, 

while the males were mainly caught after midnight. 

Trond Andersen. Museum of Zoology, N-50 14 Bergen - University. Norway. 

The arrival pattern of Trichoptera at artificial wide and half a meter deep at the trapping site. 

light have been studied, among others, by The bottom is covered with fine sand and some 

Brindle (] 958) in England, Corbet and Tj0nne larger stones. Great areas are grown with Equiland 

(1956) at Lake Victoria in Uganda, and by selum fll/vialile L.. Sparganium anguSI(folium 

Nimmo (1966) near Montreal in Canada. They Michx., Potamogelon natans L. and Nuphar luall 

found that most Trichoptera species showed teum L. 

a bimodal arrival pattern, generalJy with a 

strong peak at dusk and with a less pronounced 

peak at dawn. 

The composition of the Trichoptera fauna in 

western Norway differs quite strongly from 

those studied in the investigations quoted above. 

Therefore the aim of the present investigation 

was twofold, both a study of the caddis flies fauna 

in western Norway and a study of the arrival 

pattern at artificial light of some of the species. 

that is whether their arrival pattern differed 

from the bimodal type mentioned above. 

The light traps used were Robinson traps fit 

ted with mercury vapour bulbs (Philips HPL 

125W). They were placed on the bank, approximately 

100 m apart, and shone freely over the 

river. The traps were switched on at 21.00 hrs 

every evening and emptied manualJy every hour 

until 05.00 hrs in the morning. To minimize the 

bias of the changes in times of sunrise and sun 

set on the arrival patterns obtained, the samp 

ling period was restricted to fourteen consecu 

tive days. At the beginning of the experiment 

(August 2) sunset and sunrise occurred at app 

METHODS AND MATERIAL 

roximately 21.05 hrs and 04.22 hrs, respectively, 

whereas corresponding times at the end of 

The sampling took place between 2 and 17 Au the experiment (August 17) were 20.25 hrs and 

gust I 972. I considered the beginning of August 04.58 hrs. 

the most probable period to get adequate material 

both of limnephilids and non-limnephilids, 

A total of 3014 specimens belonging to 42 

species were collected (Table O. Eighteen Limtwo 

groups which may exhibit different arrival nephilidae species comprising 54,8 % of the indipatterns. 

The material was collected in two light 

traps at the River Fitjo (600 31' 44" N, 50 32' 8" 

viduals were trapped. Seven species were taken 

in hundred specimens or more, viz.: Limnephi 

E) on Oster0Y in Hordaland, western Norway. Ius fla vicornis (Fabricius, 1787) (n = 693), Sle 

The slowly running river is about four meters nophylax permislus McLachlan, 1895 (n = 

12 

Fauna Norv. Ser. B 26: /2-17. Oslo 1979.

.. 

Table L Trichoptera caught in I-hr intervals in two light traps at Fitjo 2 - 17 August 1972. 

Species 21-22 22-23 

d 9 d 9 

Rhyacophila nubila (Zetterstedt, 1840) 9 7 9 2 

Agapetus ochripes Curtis, 1834 I 

Oxyethira distinctella McLachlan, 1880 I I I I 

O.flavicornis (Pictet, 1834) 55 63 80 106 

O.frici O(Japalek, 189 I) 3 2 2 

O. tristella Klapalek, 1895 I 2 

Hydroptila tineoides Dalman, 1819 2 

Wormaldia subnigra McLachlan, 1865 I 

Hydropsyche instabilis (Curtis, 1834) I 

Pleetrocnemia conspersa (Curtis, 1834) 3 I 12 

Polycentropus flavomaculatus (Pictet, 

1834) 13 37 68 37 

P. irroratus (Curtis, 1835) I 

Cyrnusflavidus McLachlan, 1864 

C. trimaculatus (Curtis, 1834) I 

Lype phaeopa (Stephens, 1836) 

Tinodes waeneri(Linnaeus, 1758) 4 11 10 

Agrypnia varia (Fabricius, 1793) 7 I 8 

Apatania stigmatella (Zetterstedt, 1840) 3 

Limnephilus centralis (Curtis, 1834) 

L. coenosus (Curtis, 1834) 

L. extricatus McLachlan, 1865 I 3 I 2 

L. flavicornis (Fabricius, 1787) 16 7 31 28 

L. griseus (Linnaeus, 1758) 

L. incisus Curtis, 1834 

L. lunatus Curtis, 1834 2 6 13 

L. marmoratus Curtis, 1834 2 

L. rhombicus (Linnaeus, 1758) 

L. sparsus Curtis, 1834 

L. vittatus (Fabricius, 1798) 2 3 4 

Glyphotaelius pelluci,dus (Retzius, 1783) 3 2 I 

Phacopteryx brevipennis (Curtis, 1834) 2 I 5 

Rhadicoleptus alpestris (Kolenati, 1848) 

Potamophylax cingulatus (Stephens, 

1837) I 3 

Stenophylax permistus McLachlan, 1895 I 2 6 1 

Micropterna sequax McLachlan, 1875 I 

Goera pilosa (Fabricius, 1775) I 2 

Lepidostoma hirtum (Fabricius, 1775) 2 I 

Athripsodes alboguttatus (Hagen, 1860) 8 41 3 21 

A. cinereus (Curtis, 1834) I 

A. dissimilis (Stephens, 1836) 5 7 I 2 

Oecetis lacustris (Pictet, 1834) 2 

Molannodes tincta (Zetterstedt, 1840) 9 16 7 7 

Hrs 

23-24 00-01 01-02 02-03 

d 9 d 9 d 9 d 9 

6 3 6 I I 

I I 

I 

30 34 17 41 7 23 4 2 

I 

I I 

4 I 3 I 

I 

61 33 35 18 9 5 8 5 

I I 

I 

7 3 6 2 I 3 I 

4 I I 2 I 

2 3 

I 2 

2 6 5 

I 1 2 1 I 

72 66 82 81 66 80 58 60 

I 

I I 

22 26 16 33 13 40 13 22 

I I 2 I 

I 

I 1 

I 2 I I 

19 16 3 15 2 5 I 

2 

I 

3 I 4 13 I 7 

18 15 75 56 74 56 101 55 

2 1 

3 8 3 

I 

4 

I 

1 2 

4 

I 

5 

5 

1 

3 

2 

13 

13 

2 

3 

I 

I 

14 

8 

2 33 

2 

I 

1 

03-04 

d 9 

2 5 

5 3 

I 

2 3 

2 

I 

I 2 

13 28 

3 9 

I 

5 2 

3 2 

I 

6 I 

77 19 

2 

I 

3 2 

51 2 

04-05 

d 9 

I 

I I 

2 

13 

2 3 

I 2 

I 

3 I 

I 

4 

2 

I I

• 

sa 

30 

10 

sa 

30 

10 

A 

21 22 23 24 01 02 03 04 as 

Hrs 

Fig. 3. Catches of dominant non-limnephilidis in l-hr 

intervals in two light traps at Fitjo 2 - 17 August 

1972. Males (white) and females (hatched). A - Oxyethira 

f1avicomis (Pictet, 1834) (n = 196 dd, 275 

9 9)· B - Pulycel1trupus f1avomaculatus (Pictet, 

8 

21 22 23 24 01 02 03 04 as 

Hrs 

1834)(n = 201 dd, 138 Q 9). C - Athripsudes 

albugllttatlls (Hagen, 1860) (n = 18 dd, 82 9 9). 

D - Mulannudes til1cta (Zetterstedt, 1840) (n = 133 

dd, 35 99)· 

15

, the 

median interval of both sexes were 0.00 to The arrival pattern of an insect at light may 

o1.00 hrs. In S. permistus the median interval of 

the females was 01.00 to 02.00 hrs and that of 

the males 02.00 to 03.00 hrs. 

DISCUSSION 

The arrival pattern of Trichoptera on Osteroy 

differs quite strongly from the bimodal pattern 

previously recorded by Brindle (1958), Corbet 

and Tjonneland (I 956), and Nimmo (I 966). On 

Osteroy Limnephilidae species made out nearly 

55 'y, of the material and the family strongly in 

, fluenced the total arrival pattern. In the materials 

of Corbet and Tjonneland (I956), Nimmo 

(I 966) and perhaps also Brindle (1958) limnephilids 

were poorly represented, and the maximum 

between 0.00 and I) 1.00 hrs found on Osteroy, 

therefore lacks in these studies. 

Unlike the limnephilids, many non-limnephilids 

were observed flying during day time in the 

study area. A. alboglltlatlls, as well as other leptocerids, 

swarmed regularly over the river, and 

P../lavomaclIlatlls could be seen in numbers flying 

close to the water surface along the banks or 

between the plants in the river. The dusk peak 

shown by the non-limnephilids in the light trap 

catches probably reflects the end of the activity 

of mainly day-active species. As the daylight is 

falling the efficiency of the traps increases. At 

the same time the activity of the insects is decreasing 

resulting in the dusk peak in the arrival 

pattern of these diurnal species. The reversed 

conditions at dawn should have given a new 

peak. However, low morning temperatures during 

the samplirtg period certainly restricted the 

magnitude of the dawn peak. This peak was 

more strongly expressed in the materials of Cor 

• bet and Tjonneland (1956) and Nimmo (I966). 

Sex difference in the arrival pattern was found 

in M. tincta, the males arriving later than 

the females. Similar patterns were found in 

three species by Corbet and Tjonneland (I956). 

In Trichoptera sex differences in the activity pattern 

is not uncommon. Swarms of caddis flies 

near the water during the day consist predominately 

of males (Hickin 1967l. On Osteroy M. 

til1cla was seen swarming now and then, and 

from these swarms only males were collected. 

differ from its real activity pattern as only the 

nocturnal aspect can be expressed. Muller and 

Ulfstrand (1970) showed that a Trichoptera species, 

Philopotamus mol1tanus (Donovan, 1813), 

appeared to be day-active when collected in suction 

traps, but night-active in light traps. However, 

the arrival patterns of the dominant Limnephilidae 

species suggest that they to some degree 

reflects real activity patterns. 

The use of light traps in faunistical or ecological 

investigations often rises the question to 

Which extent the exact relative abundance of the 

species is reflected in the catches (Southwood 

1975). Even though specimens resting near the 

trap during the night will be attracted to the 

trap, the extent to which a species actually is fly· 

ing during the dark period of the night and 

hence is available for trapping have to be of importance 

for its representativity in the catches. 

REFERENCES 

Andersen, T. 1975. Caddis nies (Trichoptera) from 

Vestfold, south-eastern Norway. Norw. 1. Em. 22, 

155-162. 

Andersen, T. 1978. Influence of temperature on the 

sex ratio of Trichoptera in light-trap catches in 

weslern Norway. Norw. J. EIII. 25, 149-151. 

Brekke, R. 1946. Norwegian caddis flies (Trichopteraj. 

Norsk em. Tidsskr. 7, 155-163. 

Brindle, A. 1958. Night activity ofTrichoptera. Emomologist's 

mall. Mag. 94, 38-42. 

Corbel, P. S. & Tjonneland, A. 1956. The flight activity 

of twelve species of East African Trichoptera. 

Ulliv. Bergell Arb., Nalurv. r. 1955 No 9, 1-49. 

Hickin, N. E. 1967. Caddis larvae. Larvae o{lhe British 

Trichoplera. Hutchinson, London.. 

Muller, K. & Ulfstrand, S. 1970. Die Tagesperiodik 

der Flugaktivitat von Philopotamus montanus 

Don. und Rhyacophila nubila Zetl. (Trichoptera). 

Oikos suppl. 13, 80-86. 

Nimmo, A. P. 1966. The arrival pattern of Trichoptera 

at artificial Iighl near Montreal, Quebec. Quaesl. 

Em. 2, 217-242. 

Solem. J. O. 1970. Trichoptera new to Norway. 

Norsk elll. Tidsskr. 17, 93-95. 

Southwood, T. R. E. 1975. Ecologicalmelhods wilh 

particular re{erellce 10 Ihe sludy ofillsecl populaliollS 

(3 imp.l. Chapman and Hall, London. 


17

Seven species of Lepidoptera new to Norway. 

LEIF AARVIK 

Aarvik, L. 1979. Seven species of Lepidoptera new to Norway. Fauna Narv. Ser. B. 26, 

18-20. 

The following species are reported new to Norway: 

Elatobia fuliginasella (Zeller), Bucculatrix capreella Krogerus, Swammerdamia passerella 

(Zetterstedt), Celypha rasaceana (Schlager), c. rufana (Scopolj), Olethreutes dissalutana 

(Stange), and Cydia illutana (Herrich-Schafferl. Remarks on diagnostic characters, distribution, 

and food plants are given. 

Leif Aarvik, Gammelbruvegen 2, N·2400 Elverum, Norway. 

The material, except when otherwise stated, were 

collected and identified by the author. 

Tineidae 

Elatobiafuliginosella (Zeller) 

4 d d 19, Vestad, Elverum REs (EIS 55), 

13-30 July 1978. One of the males was captured 

by Kai Berggren. All the males were attracted to 

light, whereas the female was netted in the evening. 

E. fuliginosella differs from all known Norwegian 

species of the family by having grey 

hairs on the head. The forewings, too, are grey. 

The genitalia are figured by Rannemann (I 977). 

The distribution is holarctic, but the species is 

rarely encountered. In Sweden it has been recorded 

from the Stockholm area (Sodermanland 

and Uppland) and Gotska Sandon (Svensson 

I 974b). 

In Sweden the larva has been found once, 

under the bark of an old pine log (Svensson 

1974b). 

Lyonetiidae 

Bucculatrix capreella Krogerus 

Id, Vestad, Elverum REs (EIS 55) 27 May 

1978 and I 9, same locality, 23 August 1978. 

The male was captured in a light trap. The female 

was found indoors on a window. 

B. capreella is a polymorphic species, and it 

may resemble other species of the genus. The 

distinct genitalia are figured by Svensson (1970. 

According to Svensson (197 I) it has previously 

been recorded in Finland, Sweden and 

Scotland only. In Sweden it has been found in 

five provinces from Vastergotland to Norrbotten 

(Benander 1953, Svensson 1974a, 1978). 

The biology is not known. 

18 

Yponomeutidae 

Swal1ll11erdamia passerella (Zetterstedt) 

Id, Damtjern, Elverum HEs (EIS 55), 6 June 

1977. 

Some authors regard S. passerella as a form 

of S. pyrella (Villers). S. pyrella is distributed in 

south and central Europe extending as far north 

as Denmark and south Sweden. It feeds on Crataegus, 

Prunus, Cerasus, Malus and Pyrus (Hannemann 

1977). S. passerella is distributed in 

Scotland, Fennoscandia, and the Baltic part of 

the USSR. It feeds exclusively on Betula nana L. 

(Hannemann 1977). The specimen from Elverum 

was captured in a bog with plenty of Betula 

Ilana. 

Hannemann (1977) who figures the genitalia 

of S. pyrella states that there are minor differences 

between the two taxa in the male genitalia. 

In Sweden S. passerella is recorded from 

eight provinces from Varmland to Lule Lappmark 

(Benander 1946, 1953, Svensson I974a, 

1978). 

Tortricidae 

Celypha rosaceana (Schlagerl synonym: e. purpl/rana 

(Raworth) 

Id, Bjelland, Trom0Y AAy (EIS 6), 29 June 

1970, Sigurd Bakke leg. 

e. rosaceana often resembles e. rufana (Scopo 

ID, but the two species can easily be separated by 

examination of their genitalia which are figured 

by Pierce & Metcalfe (1922). The genitalia of e. 

rosaceana show a remarkable similarity to those 

of e. striana (Denis & Schiffermullerl. This species, 

however, is completely different externally. 

e. rosaceana is often found on sea shores (Benander 

\950), but it has also been taken in inland 

habitats. The locality at Trom0Y is situated 

close to the sea. 

Fauna Norv. Ser. B 26: 18-20. Oslo 1979.

Hoverflies (Diptera, Syrphidae) associated with Ramson 

Allium ursinum L. 

TORE R. NIELSEN 

Nielsen, T. R. 1979: Hoverflies (Dipt., Syrphidae) associated with Ramson Allium ursillum 

L. Fauna Norv. Ser, B 26. 21-23. 

Two species of Hoverflies associated with Ramson are reported from the Bergen area. Portevinia 

l11aculata (Fallen) 1817 is new to the Norwegian fauna. while Cheilosia fasciata Schiner 

& Egger 1853 has, according to older literature, been found in Oslo. A few observations 

on the biology of the species are given, and also a key to the adult insects. 

Tore R. Nielsen, Juvelveien 19 D, N-4300 Sandnes, Norway. 

Amongst Hoverflies the genus Cheilosia is 

known to have herbivorous larvae. Quite a few 

This is the more slender of the two species. It 

differs from Portevinia mandata also in having 

species have their larvae in the stems or roots of 

different herbs, a few have leaf-mining larvae, 

hairy eyes (Fig. 1a). Other characters will appear 

from the key, Table I. 

... 

while others eat fresh or rotting tissues of mushrooms. 

Cheilosia fasciata has an early flight period. 

Within the old genus Chei/osia there are two From the material above it seems that in wes 

Norwegian species which, in the larval stage, tern Norway it may be found from the end of 

are associated with Ramson AIlium ursinul11 L.: April till the end of May. The localities at Para 

Ch. fasciata Schiner & Egger 1853 whose larvae dis and in Isdalen have frequently been visited 

are leaf-mining, and Portevinia (previously Chethrough several years, but it has not been posi1osia) 

macu/ata (Fallen) 1817, where the larvae sible to trace the species later than 26 May, and 

are assumed to live in other parts of the host only one year as early as at the end of April. The 

plant. 

length of the flight period seems to be about 

Most of the Chei/osia-species are, as adult in three weeks in years of normal weather condisects, 

shining black. The two species mentioned tions. 

(along with a third Norwegian species, Ch. seThe species has been recorded in open and 

mi!asciata Becker 1894, which mine on Sedum rather humid herb-fields (Ground Elder Aegoporosea 

(L.» differ jn having three pairs of greyish dium podagraria, Lady's Mantle A/chemilla sp. 

abdominal spots. The body-length of both spe and Ramson) in the neighbourhood of rich, decies 

are 8.0-8.5 mm. The material, collected ciduous forests. The adult insects were most of 

1966-71, originates from six localities in the Ber ten seen flying around the Ramson plants, and 

gen area, but the species will most probably be here both of the sexes were often observed sunfound 

in many other localities where Ramson is ning on the leaves. Otherwise the females visited 

found. 

the flowers of Lesser Celandine Ranuncu/us Iicaria 

L., and on sunny days some males might 

be seen hovering in Iow height above the gro 

Cheilosia fasciata Schiner & Egger, 1853 

und vegetation. 

Paradis, Hay: Bergen 19 May 1966 (I d, Towards the end of the flight period the fema 

29 9); 12 May 1967 (I d, 29 9); 28 April les could be seen creeping down on the under 

(I d, 29 9), 30 April (4 dd, 49 9), I May side of the Ramson leaves, where the yellowish 

(9 dd, I 9), 4 May (2 d d) and 22 May 1968 white eggs were deposited on the epidermis. 

(I 9); 2 May (2 d d) and 4 May 1969 (9 dd, Only one egg seemed to be laid on each leaf. Af 

29 9); 5 May (1 d), 10 May (9 dd, I 9) and ter hatching, the larva gnawed through the epi 

23 May 1970 (I d, I 9); 4 May (8 dd, 59 9), dermis and into the green, succulent tissues 

14 May (I 9, A. Fjellberg leg,) and 26 May 1971 above. At first the mines were relatively nar 

(I 9). Isdalen, HOy: Bergen 12 May 1967 (I 9, row. Later they were expanded to large, airfilled 

A. Fjellberg leg.). Lii, Os, HOy: Os 4 May 1969 sacs inside the leaves (Fig. 2). The larvae pupated 

(I d, 99 9, A. Fjellberg leg,). in the soil, just below the surface. 

Fauna Norv. Ser, B 26; 21-23, Oslo 1979. 21

Table I. Determination key for Cheilosia fasciata 

Schiner & Egger and Portevinia maculata (Fallen). 

I. Abdomen black with three pairs of greyish spots 

(spots often less distinct in the females) . . . .. 2 

2. Eyes hairy 3 

- Eyes bare . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 4 

3. Antennae brownish-black. Face (Fig. Ia) just below 

antennae not much hollowed, lower part of 

face with protruding central prominence. Distance 

between lower margin of eye and lower 

margin of face relatively large (about 1/4 of height 

of head). Legs black, except for basal 1/3 

1/2 of tibiae on all legs yellowish. Abdomen (Fig. 

Ib) with long, bristly hairs, in female at least 

along side-margins. Rather slender species Cheilosia 

fasciata * 

4. Third antennal joint orange. Face (Fig. le) just below 

antennae much hollowed, lower part of face 

evenly convex. Distance between lower margin 

of eye and lower margin of face short. Legs black, 

only basal ca. 1/10 of tibiae on all legs yellowish. 

Abdomen (Fig. Id) with short, adpressed hairs. 

Greyish spots in female most readily seen from 

behind. Rather broad species Portevillia maculata 

* Cheilosia semifasciata has, like Ch..fasciata, hairy 

eyes. The male differs, however, from that species 

in having the abdominal spots more diffuse, and 

the longest hairs along the sides of tergite 3 distinctly 

shorter than the height of the abdominal 

spots. (In Ch. fasciata they are longer than the height 

of the spots). 

The female Ch. sem(fasciata is glittering black, 

without abdomimal spots. 

ACKNOWLEDGEMENTS 

I wish to thank Cand. real. Arne Fjellberg for 

having contributed with material of both species 

concerned. 

REFERENCES 

Strand, E. 1903. Norske lokaliteter for Diptera. Forh. 

Christiallia Videllsk. 3, 3-11. 


23

Spiders from Jan Mayen 

Sl2JREN TOFT 

Toft, S. 1979. Spiders from Jan Mayen. Fauna Norv. Ser. B. 26, 24-25. 

A collection of spiders from Jan Mayen, taken mainly by means of pitfall-traps in June 

July 1972, contained live species. of which one, Walckenaera c1avicornis (Emerton), is new 

to the island. Co/linsia holmgreni (Thorell) is by far the dominant species. The fauna comprises 

three holarctic, and two palaearctic species. 

S0ren Toft, Zoological Laboratory, University of Aarhus, DK-8000 Aarhus C, Denmark. 

The spider fauna of the Norwegian arctic island 

Jan Mayen has been dealt with in papers by 

Bristowe (1925, 1948), the latter of which summarizes 

all collections made up to then. Though 

a total of about 800 specimens have been collected, 

only four species are recorded, all belonging 

to the family Linyphiidae: Cof/insia ho(mgreni 

Thorelll, Erigone tiro/ensis L. Koch, Hi/aira frigida 

(ThorelJ) and Meioneta nigripes (Simon). 

Through the courtesy of Professor Niels 

Haarl0v, I have had the opportunity to examine 

a new collection, made by Klaus Vestergaard, in 

June - July 1972. Most of the spiders in this 

collection were taken by means of pitfall-traps, 

of which 30 were used for different periods in 

various localities. Besides this, 281 core samples 

were extracted with a Berlese - extractor, but 

only five of these each contained a single spider. 

As none of the previous collections have used 

pitfall traps, one would expect somewhat different 

species proportions in the present material, 

but this is only partly observable. 

The collection comprised a total of 187 specimens, 

of which 182 were taken in pitfall traps; 

five species could be identified, i.e. the four species 

previously reported from Jan Mayen, and 

one species new to the island: Wa/ckenaera clavicornis 

(Emerton). All five specimens from extraction 

samples were juvenile C. ho/mgreni. Details 

about localities, habitats etc. are given below. 

The material will be kept at the Zoological 

Museum, Copenhagen, Denmark. 

Wa/ckenaera clavicornis (Emerton) 

Material: Id 

Locality: Nordahl Grieg Lia. 

Habitat: In moss (Rhacomitrium) on gravelly 

ground. 

Co/linsia holmgreni (Thorell) 

Material: 76 Q Q 37 d d 43 juv. in pitfall traps, 

5 juv. in core samples. 

24 

Localities: Nordahl Grieg Lia, Tornoe delta, 

Fishburndalen, Kreklinghaugen, Havhestberget, 

Wilezekdalen, Blyttberget, Kvalrossbukta, Sjuhollendarbukta, 

Borgdalen, S0yla, Libergsletta. 

Habitats: The species is distributed all over the 

island, and seems to occur in nearly all available 

habitats. By far the greatest numbers were caught 

in stony areas devoid of yegetation (Wilezek 

dalen). Bristowe also found them under stones. 

However, in a Norwegian high mountain 

area, Hauge et al. (1978) caught the largest numbers 

in eutrophic meadow, but it was also abundant 

in snow-bed. Palmgren (1965, 1976) reports 

it to occur in mountaneous heathland in 

northern Finland. 

Erigone tiro/ensis L. Koch 

Material: 4 Q Q 7 d d I juv. 

Localities: Fishburndalen, Havhestberget, Kvalrossbukta. 

Bristowe (1948) also mentions Sjuhollendarbukta. 

Habitats: In the first mentioned locality the 

trap was situated in a rich vegetation dominated 

by Sibba/dia; the remaining localities were moss 

below bird cliffs. Hauge et al. (1978) report the 

species to be abundant in a wide variety of high 

mountain habitats. In Northern Finland it occurrs 

from the timberline to the highest mountain 

tops (Palmgren 1965, 1976). 

Hilairafrigida (Thorell) 

Material: I Q 2 d d I juv. 

Localities: Nordahl Grieg Lia, Sjuhollendarbukta. 

Habitats: At both localities the traps were situated 

in moss. According to Bristowe (1948) it 

is abundant throughout the island. The species 

has two activity peaks, one in very early spring, 

the other in late autumn (Hauge et al. 1978), 

Fauna Norv. Ser. B 26, 24-25. Oslo 1979.

which may explain the low numbers in the present 

collection. In Northern Finland it is a dominant 

species of mountain heaths (Palmgren 

1965, 1975). 

Meioneta nigripes (Simon) 


I am greatly indebted to Professor Niels Haarl0v 

and Or. Klaus Vestergaard for having the opportunity 

to examine this collection, and for detailed 

information about the sampling sites. 

MateriaL 3 Q Q 6 d d 

Localities: NordahJ Grieg Lia, Wilczekdalen, 

Libergsletta, Fugleberget. Bristowe (1948) further 

mentions Fishburndalen and Bernakrater. 

Habitats: The species has been taken in moss 

and stony areas without vegetation. This is in 

accordance with Hauge et al. (I 978), who caught 

it exclusively in a pioneer ground habitat. 

REFERENCES 

Bristowe, W. S. 1925. The Fauna of the Arctic Island 

of Jan Mayen and its probable Origin. Ann. Mag. 

Nat. Hist. Ser. 9, 15, 480-485. 

Bristowe, W. S. 1948. Spiders from the Arctic Island 

of Jan Mayen. Proc. Zoo/. Soc. Lond. 118, 

223-225 

Bnendegaard,1. 1946. The spiders (Araneina) of East 

Greenland. Medd. Gronland 121 (15), 1-128. 

DISCUSSION 

Brrendegaard, J. 1958. Araneida. The Zoology oficeland 

11/ 54: 113 pp. 

The five species known from Jan Mayen are wi 

Hauge, E., Hiigvar, S. & 0stbye, E. 1978. Pit-fall 

dely distributed in the Northatlantic area. They catches of surface-active arthropods in some high 

all occur in Iceland (Bnendegaard 1958), East mountain habitats at Finse, south Norway. Ill. 

Greenland (Bnendegaard 1946), Scotland The species of Araneida. Norw. J. Ent. 25, 

(Locket & Millidge 1953) and Northern Scandi 207-220. 

navia (Palmgren 1965, 1975, 1976). In the two Holm, A. 1937. Zur Kenntnis der Spinnenfauna 

latter areas they are restricted to high mounta Spitzbergens und der Baren InseJ. Ark. Zoo/. 29 A 

ins. Holm (1967) characterizes C. holmgreni, W. (18),1-13. 

c1avicornis and H. frigida as holarctic, M. nigri 

Holm, A. 1967. Spiders (Araneae) from West Greenland. 

Medd. Gr@nland 184 W, 1-99. 

pes and E. tirolel/sis as palaearctic. 

Locket, G. H. & Millidge, A. F. 1953. British Spiders 

A fauna comprising only five species of spi /1. Ray Society, London. 

ders may seem extremely poor. However, this Palmgren, P. 1965. Die Spinnenfauna der Gegend 

figure is probably not very far from the equili von Kilpisjarvi in Lappland. Acta zool. fenn. 110, 

brium number of species in the area. For compa 1-70. 

rison, 38 species are known from East Green Palmgren, P. 1975. Die Spinnenfauna Finnlands und 

land (Holm 1967), 16 species from Spitsbergen 

(Holm 1937, Jambs-Lyche 1967), both with a 

much larger area, and five species have been recorded 

from Bear Island (Holm 1937). Thus, in 

view of the small area and great isolation of Jan 

Mayen only few more species can be expected. 

The faunal composition has remained stable 

Ostfennoskandiens. VI. Linyphiidae I. Fauna 

Fennica 28. 102 pp. 

Palmgren, P. 1976. Die Spinnenfauna Finnlands und 

Ostfennoskandiens. VII. Linyphiidae 2. Fauna 

Fennica 29. 126 pp. 

Tambs-Lyche, H. 1967. Notes on the distribution of 

some Arctic Spiders. Astarte 28, 1-13. 

through the last 50 years. 


25

Spiders from arctic Fennoscandia 

N. PHILIP ASHMOLE AND BERNARD R. PLANTEROSE 

INTRODUCTION 

Ashmole, N. P. & B. R. Planterose 1979. Spiders from arctic Fennoscandia. Fauna Norv. 

Ser. B, 26, 26-30. 

Spider collections made in the Lofoten Islands in 1975 and in a number of localities in northern 

Norway and Sweden in 1977 are described. Altitudinal information is provided for all 

the montane records and details of the vegetation are given for many of the sites. 

N. Philip Ashmole, Department of Zoology, University of Edinburgh. 

Bernard R. Planterose, Department of Forestry and Natural Resources, University of Edinburgh. 

A few intensive studies of spider communities 

have been made in particular localities in arctic 

Fennoscandia (e.g. Holm 1950, Palmgren 1965, 

Koponen 1976) but there is a paucity of distributional 

data. Recent contributions include a brief 

note by Cooke (\ 967) on collections from several 

parts of Norway; an account by Waaler 

(\ 970) of a collection from S0r-Varanger, North 

Norway; a description by Hauge (\ 97 \) of collections 

made by D.G. Goddard in 1969 in the 

Lyngen Peninsula and near Troms0, North Norway; 

an account by Hauge (\ 977) of spiders 

from birch forests in the Skjomen fjord, northern 

Nordland, Norway; and a list published by 

Holm (1973) of spiders collected in the islands of 

northern Norway by a Swedish expedition in 

1875. 

The collections described here come from two 

sources. In the summer of 1975, in the course of 

a holiday in the Lofoten Islands, NPA spent 

short periods collecting in anum ber of localities. 

In the summer of 1977 BRP led an Edinburgh 

University expedition to northern Fennoscandia 

during which a number of spider collections 

were made, although the primary objective was 

ornithological. 

Different approaches were used in the two 

years, and the data for the two are therefore presented 

in a somewhat different manner. On the 

Lofoten Islands in 1975 collections were made 

in a number of places by a single person, usually 

spending about 30 minutes at each site; altitudes 

were recorded and general notes were made on 

the habitats. These data are presented in the Systematic 

List. 

During the 1977 expedition collections were 

made in well-defined areas that were investigated 

simultaneously by several people. Usually, a 

26 

I sq. metre quadrat was searched intensively; 

within it, vegetation was uprooted, stones were 

carefully removed and any loose surface humic 

matter was sifted. At most sites, less intensive 

searches of the area within about 10 m round 

about the quadrat were also carried out. Estimates 

were made of the percentage' contribution to 

the vegetation cover by the various species within 

each quad rat. 

Since more habitat information is available 

for these sites they are described individually in 

the next section and a list of the spider species 

represented in each is provided. The number 

and sex of the specimens from these is indicated 

in the Systematic List but habitat information is 

not repeated: it can be found by referring to the 

relevant locality and elevation in the Description 

of Sites. 

DESCRIPTION OF SITES 

I. Lofoten Islands, Norway. 

Over 20 sites were sampled, the southernmost 

on Vremy at 67°40'N. and the northernmost on 

Austvag0Y at 68°16'N. Elevations ranged from 

sea level to 510 m. 

11. Ammarnas, Sweden. 

Four sites were sampled in the Vindalalven, just 

north of Ammarniis at 66°05'N I O'E. Elevations 

ranged from 685 m to 880 m. 

The lowest site, in the region known as Hobacken 

at 685 m, was studied for three manhours 

on 30 June 1977. It was in open birch 

forest with a vegetation cover consisting of 40 % 

Juniperus communis, 30% Deschampsia flexuosa, 

10% Vaccinium myrti/lus, 60 % mosses and 

I % lichens. The spider species represented were 

Fauna Norv. Ser. B 26: 26-30. Oslo 1979.

Pa rdosa lugubris (Walckenaer) Oreonetides va· 

ginatus (Thore\l) and Hilaira frigida (ThoreID. 

Sampling at 780 m occupied 4 1/2 manhours 

on 29 June and 2 July 1977. The site was 

just within birch at the tree-line and was dry, 

with 80 % Empetrum nigrum agg., 10% lichens, 

5 % Vaccinium spp., 4 % Arctostaphylos uvaursi 

and I % Carex spp. The first collection, 

made from the moist organic layer of the topsoil, 

contained only a subadult Gnaphosa sp. and 

juvenile Pardosa sp. The second collection was 

made on the surface and was dominated by juvenile 

Iycosids: the only adults were Pardosa 

hyperborea (Thorell) and Alopecosa aculeata 

(C1erck). 

One and a half man-hours were spent sampling 

at 8I 0 m on 29 June 1977 at a site of a very 

different nature: a marshy hollow with a stream 

and willow scrub, just above the birch tree-line. 

Vegetation cover consisted of Salix sp. 20 %, 

Betula nana 10%, Molillia caerulea 80 %, 

Sphagnum sp. 10% and Equisetul11 sp. 0,5 %. 

The spider sample was distinct from that at 780 

m, and more diverse: Pardosa al11entata (C1erck) 

was abundant and Robertus scoticus Jackson, 

Hypoml11a bituberculatum (Wider) and Hi/aira 

pervicax Hull were also represented. 

The highest site was at 880 m, where sampling 

was for I 1/2 man-hours on 5 July 1977. 

This site was just above the dwarf birch/willow 

zone, close to a snow patch. Vegetation cover 

was not complete, with about 20 % bare ground, 

Vaccinium myrtillus 20 %, Betula nana 

5 %, El11petrum nigrum 2 %, lichens J0 % and 

dead grass 40 %. The spiders included a number 

of Pardosa, but only one adult, which was P. 

palustris (L.); bther species found were Meioneta 

nigripes (Simon), Oreonetides vaginatus and 

Lepthyphantes alacris (BlackwaIl). 

Ill. Sareks National Park, Sweden. 

Three samples were taken here, at 67°20'N.. 

17°50'E., the sites ranging in elevation from 815 

m to 1650 m, all on the east facing slope of Sarvatjakka, 

above Pierikjaure. 

The lowest site, at 815 m, was sampled by quadrat 

on 20 July 1977, for 2 1/2 man-hours. 

There was scrubby vegetation with cover comprising 

Empetrum nigrum 60 %, Betula nana 

20 % and Salix sp. (max. height 20 cm) 20 %. 

The spider fauna was fairly diverse, many specimens 

being found under rocks. Pardosa spp. 

were the commonest forms, the only two adults 

being P. eiseni; another species was represented 

only by young individuals. Other species were 

Tiso aestivus (L. Koch), Oedothorax retusus 

(Westring), Trichopterna mengei (Simon), Xysticus 

cristatus (C1erck) and Rhaebothorax moru 

Ius (O.P.-Cambridge). 

The next site, at I 280 m, was also sampled for 

2 1/2 man-hours on the same day. The sampling 

area was close to a melting snow-patch but 

was fairly dry and rocky with lichens; the other 

vegetation consisted mainly of Betula nana with 

some Carex sp. and Deschampsiaflexuosa. Only 

two species of spider could be identified, and 

one of these, Pardosa eiseni, was represented 

only by immature individuals; the other was 

Lepthyphantes bergstroemi Schenkel. 

The highest site, at 1650 m, was sampled for 

40 minutes on the same day. It was by a glacier 

on loose moraine, with scattered Betula nana 

and Salix sp. The six spiders collected were all 

Pardosa, and the three adults were P. eiseni. 

IV. Lyngen Peninsula, Norway. 

Sampling was in the Lyngsdalen, 69°30'N., 

20 o E., around 300 m elevation between the 

snout of the Sydbreen glacier and the top of the 

birch zone. About 2 1/2 man-hours were spent 

collecting on 13 August 1977. 

The substrate was mainly of rocks and boulders 

with patches of smaller gravel; there were 

scattered plants of Salix sp. and Empetrum nigrum. 

It was interesting to find Araneus diadematus 

C1erck abundant among the boulders within 

about 200 m of the glacier. Most of the other 

spiders were Pardosa; the only adults in the 

main collecting site, which was rocky, were P. 

amentata but all those in a braided river area 

with smaller stones a little lower down were P. 

palustris. In the rocky area there was also one 

Erigone dentipalpis. 

Hauge (] 97 J) described a larger collection 

from the Lyngen Peninsula made by Mr. D.G. 

Goddard. Precies localities were not given but it 

is evident that a much wider range of habitats 

was sampled than in 1977. One interesting record 

is of Lycosa tarsalis Thorell ( = Pardosa palustris(L.))«at 

moraine»; our record from a braided 

river close to a glacier confirms the association 

of this species with this kind of habitat. 

V. Ser-Varanger, Norway. 

Collections were made at three sites in an area 

27 km east of Kirkenes, at 69°45'N., 30 0 45'E. 

Elevations ranged from 220 m to 400 m. 

The first site, 20 m from the south bank of 

Holmvatnet and at 220 m, was studied for just 

over 3 man-hours on 31 July 1977. The ground 

27

surface was very dry, without rocks or stones. 

Vegetation consisted of Empetrum nigrum 60 %, 

Vaccinium myrtillus 30 %, Betula nana 10% 

and lichens I I) %. In keeping with the open nature 

of the site and the absence of rocks, the spider 

fauna was dominated by lycosids. Several 

Alopecosa were obtained, but none was adult so 

the species cannot be determined. All the other 

identifiable spiders were Pardosa palustris. Several 

more Pardosa were collected en route to 

the next site, in areas where the vegetation was 

similar but rather more sparse than at the first 

site. These were mainly P. palustris but included 

one P. eiseni. 

The next site was at 400 m, at the top of a hill 

south of Holmvatnet. Sampling occupied just 

over two man-hours on the same day. Theground 

was dry and rocky, with some patches of 

smaller stones and gravel; there were scattered 

plants of Diapensia Japponica, Empetrum nigrum, 

Vaccinium uliginosul11, Betula nana and lichens. 

The spiders fell into two groups. Those 

collected on the surface were almost alllycosids; 

as at the first site, P. palustris was most abundant, 

but here there was also one P. eiseni, subadult 

Alopecosa sp. and subadult Tricca alpigena 

(Doleschall); the only non-Iycosid was Xysticus 

albidus Grese. The other spiders were collected 

under small stones and gravel; apart from one 

Tiso aestivus, they were all Meioneta nigripes. 

The last sample, collected in I I /2 man-hours 

on the same day, was a little to the south of the 

hilltop at 370 m, in a wet gully with a small 

stream. The site was damp and mossy with EI11petrum 

nigrum and patches of Salix sp. and Betula 

nana; part of the area was drier, with some 

Vaccinium spp. The spider fauna was similar to 

that in the previous site, with Alopecosa sp., 

Pardosa palustris and P. eiseni; in addition, however, 

there were two P. amentata, and among 

some big boulders were two Araneus cornutus 

Clerck. 

GNAPHOSIDAE 

SYSTEMATIC LIST 

Zelotes subterraneus (C.L. Koch). F1akstad0ya, Lofoten; 

west slopes of Fisken at L40 m; F; 30 June 75. 

Austvag0Y, Lofoten; on moss near shore close to 

Svolvrer; 25 June 75. 

CLUBIONIDAE 

Clubiona ree/usa a.p.- Cambridge. M, F, 23 and 26 

June 75. Near Svolvrer, Austvag0y, at 68° 15'N.; 

heath near sea Level. 

28 

THOMISIDAE 

Oxyptila trux (Blackwall). FLakstad0ya, Lofoten; moorland 

at Less than 30 m; 3 F; 27-29 June 75. 

Xysticus albidus Grese. S0r-Yaranger, Norway, 400 

m; M; 31 July 77. This specimen was identified 

by Or. Erling Hauge. The species has not previously 

been recorded from Norway. 

Xysticus eristatus (Clerck). Sareks National Park, 

Sweden; 815 m; F; 20 July 77. Flakstad0ya, Lofoten; 

rocky shore near Nusfjord; F; 27 June 75. 

Austvag0y, Lofoten; under rock on slopes of 

Rundfjell at 305 m; M; 26 June 75. 

LYCOSIDAE 

Alopecosa aculeata (C1erck). Ammarnas, Sweden, 

810 m; M; 3 July 77. The identity of this specimen 

was kindly confirmed by Or. Ake Holm. 

Alopecosa pulverulenta (C1erck). Flakstad0ya, Lofoten; 

M and F abundant on moorland at low altitude 

and recorded also on a steep slope at about 

L40 m; 27-30 June 75. 

Pardosa agricola (Thorell). Austvag0Y, Lofoten; on 

wet moss by the shore near Svolvrer; M; 25 June 

75. 

Pardosa al11entata (Clerck). Flakstad0ya, Lofoten; on 

water by lake; M; 30 June 75. Ammarnas, Sweden; 

810 m; M and F abundant; 29 June 77. 

Lyngen Peninsula, Norway; c. 300 m; 3 F; 13 

Aug. 77. S0r-Yaranger, Norway; 370 m; 2 F; 31 

July 77. 

Pardosa eiseni (Thorell). Sareks National Park, Sweden; 

815 m; 2 F; 20 July 77. Same area, but 1650 

m; 2M, F; 20 July 77. S0r-Yaranger, Norway; 

220-400; 3M, 2F; 31 July 77. 

Pardosa hyperborea (ThorellJ. Ammarnas, Sweden at 

780 m; 2 M, F; 2 July 77. 

Pardosa lugubris (Walckenaerl. Ammarnas, Sweden; 

685 m; 3 F; 30 June 77. 

Pardosa palustris (L.). Ammarnas, Sweden; 880 m; 

F; 5 July 77. Lyngen Peninsula, Norway; just below 

300 m; 5 F; 13 Aug. 77. S0r-Yaranger. Norway; 

220-400 m; 9 F; 31 July 77. 

Pardosa sphagnicola (F. OahlJ. Flakstad0ya, Lofoten; 

moorland at less than 30 m; 3 M; 27 June-2 July 

75. Austvag0y, Lofoten; on wet moss near the 

shore and in a marsh, near Svolvrer; M, 2 F; 25 

June 75. 

Pirata piraticus (C1erck). Austvag0y, Lofoten; in a 

ditch near Svolvrer; M; 25 June 75. 

Tricca alpigena (Doleschall). S0r-Yaranger, Norway; 

400 m; subadult F; 3LJuly 77. 

Xerolycosa nemoralis (Westring). Flakstad0ya, Lofoten, 

at 68°03'N.; on east slopes of Fisken at about 

140 m; in area with scree of small stones; 2 M; 30 

June 75. 

AGELENIDAE 

Cryphoeca si/vicola (Koch). Yrer0Y, Lofoten; among 

boulders near beach; F; 3 July 75. Austvag0Y, Lofoten; 

birch/rowan wood at c. 50 m near Svolvrer; 

F; 24 June 75.

THERIDIIDAE 

Robertus arwldineti (O.P. Cambridge). Austvag0y, 

Lofoten; a marsh near Svolvrer at 68° 13'N.; F; 25 

June 75. This may be the northernmost record for 

the species, although Palmgren (I974) indicates 

that it reaches the arctic circle in Finland. 

Robertus lividus (Blackwall). Vref0Y, Lofoten; under 

rock in grass sward close to cliff top at c. 425 m; 

F; 3 July 75. Flakstad0ya, Lofoten; moorland 

close to sea level and hillside at c. 160 m; 6 F; 

27 - 28 June 75. Austvag0Y, Lofoten; a marsh 

near Svolvrer; F; 25 June 75. 

Robertus scoticus Jackson. Ammarniis. Sweden; 810 

m; F; 29 June 77. 

Theridion bel/icosum Simon. Vref0Y, Lofoten; boulders 

near beach; F; 3 July 75. Austvag0y, Lofoten; 

birch/rowan wood at c. 50 m near Svolvrer; 

F; 24 June 75. 

TETRAGNATHIDAE 

Meta merianae (Scopolil. Moskenes0ya, Lofoten; 

among boulders near sea level; F; 28 June 75. 

Flakstad0ya, Lofoten at 68°06'N.; sheltered places 

in moorland below 30 m; M, F; 28-29 June 75. 

Tetragnatha extensa (L.). Flakstad0ya, Lofoten; moorland 

below 30 m; M; 2 July 75. 

ARANEIDAE 

Aranells cornutus Clerck. S0r-Varanger, Norway; 

370 m; 2 F; 31 July 77. 

Araneus diadell1atus Clerck. Lyngen Peninsula. Norway; 

c. 300 m; 5 F; 13 Aug. 77. 

LINYPHIIDAE, Subfamily ERIGONINAE 

Caledonia evansi O.P. Cambridge. Austvag0Y, Lofoten; 

mountainside near Svolvrer at 400 m; F; 24 

June 75. 

Ceratinella brevipes (Westring). Flakstad0ya, Lofoten; 

moorland close to sea level and at 140 m. in 

vegetation; 3F; 28 and 30 June 75. 

Dip/ocepha/us cristatlls (BlackwalI). Flakstad0ya, Lofoten 

at 68°06'N.; sandy beach; 5M, I1 F; 29 June 

75. This appears to be the nothernmost record for 

the species. 

Erigone arctica (White). Flakstad0ya, Lofoten; sandy 

beach near Flakstad; 3 M; 18 F; 29 June 75. 

Erigone atra (Blackwalll. Flakstad0ya, Lofoten; 

sandy beach with E. arclica; M, 2F; 29 June 75. 

Erigone dentipa/pis (Wider). Lyngen Peninsula, Norway; 

c. 300 m; 13 Aug. 77. 

HYPol1ll1la bitubercu/atum (Wider). Ammarniis, Sweden; 

8\ 0 m; F; 29 June 77. 

Oedothorax retusus (Westring). Sareks National 

Park, Sweden; 815 m; F; 20 July 77. Austvag0Y, 

Lofoten; under seaweed on rocky beach near 

Svolvrer; M, 7 F; 25 June 75. 

Pocadicnel1lis pumila (Blackwalll. Flakstad0ya, Lofoten; 

c. 140 m in vegetation on steep slope; F; 30 

June 75. 

Rhaebothorax morulus (O.P. Cambridge) Sareks National 

Park, Sweden; 815 m; F; 20 July 77. 

Savignya frontata (Blackwall). Flakstad0ya, Lofoten; 

under debris on rocky shore; F; 27 June 75. 

SiIol11etopus reussi (Thorell). Flakstad0ya, Lofoten; 

sandy beach; F; 29 June 75. This specimen was 

kindly determined by A. Holm. 

Tiso aestivus (L. Koch). Ammarniis, Sweden; 880 m; 

F; 5 July 77. Sareks National Park, Sweden; 815 

m; 2 F; 20 July 77. Vref0Y, Lofoten; 425 m; under 

rock in grass sward close to cliff top; 3 F; 3 July 

75. S0r-Varanger, Norway; 400 m; 4 F; 31 July 

77. 

Trichopterna mengei (Simon). Sareks National Park, 

Sweden; 815 m; F; 20 July 77. 

Wa/ckenaera antica (Wider). Flakstad0ya. Lofoten; 

moorland close to sea; F; 28 June 75. 

Wa/ckenaera capito (Westring). Austvag0Y, Lofoten; 

510 m on Rundfjell, in area recently exposed by 

melting snow; 2 F; 26 June 75. 

LINYPHIIDAE, Subfamily LINYPHIINAE 

Bo/yphantes /uteo/us (Blackwall). Vref0Y, Lofoten; 

among boulders near beach; F; 3 July 75. 

Centrol11erus arcanus (O.P. Cambridge). Austvag0y, 

Lofoten; 305 m on Rundfjell; F; 26 June 75. 

Centrol11erus incilium (L. Koch). Flakstad0ya, Lofoten; 

at 68°0)' N.; 260 m on East slope of Fisken, 

under rocks in grassy area; F; 27 June 75. This 

may be the northernmost record of this rare species. 

The specimen was determined by P. Merrett. 

Centromerus sy/vaticus (Blackwalll. Flakstad0ya, Lofoten; 

moorland below 30 m; F; 27 June 75. 

Dip/osty/a conc%r (Wider). Austvag0y, Lofoten; 

under debris on rocky shore near Svolvrer; M; 25 

June 75. This may be the most northerly record of 

the species. 

Ha/orates reprobus (O.P. Cambridge). Vrer0Y, Lofoten; 

under debris on rocky shore; M; 3 July 75. 

Flakstad0ya, Lofoten; on rocky shore; M, F; 27 

June 75. 

HiIaira excisa (O.P. Cambridge). Nusfjord, Flakstad0ya, 

Lofoten, at 68°0 I'N.; under debris on rocky 

shore; F; 27 June 75. This may be the most northerly 

record of the species. 

Hi/aira frigida (Thorelll. Ammarnas. Sweden; 685 

m; F; 30 June 77. Vrer0Y, Lofoten; under rock in 

grass sward close to cliff-top at 425 m; 5 F; 3 July 

75. Austvag0Y, Lofoten; 510 m on Rundfjell, 

under rock recently exposed by melting snow; F; 

26 June 75. 

HiIaira pervicax Hull. Ammarnas, Sweden; 810 m; 

F; 29 June 77. 

Lepthyphantes a/aais (Blackwalll. Ammarniis, Sweden; 

880 m; F; 5 July 77. Austvag0Y, Lofoten; 

mountainside near Svolvrer at 270 m and 400 m; 

2 M, F; 24 June 75. Also in the same area, 510 m 

on Rundfjell, under rock recently exposed by 

melting snow; M, 2 F; 26 June 75. 

Lepthyphantes bergstroemi Schenkel. Sareks National 

Park, Sweden; 1280 m; M; 20 July 77. 

29

Lepthyphantes ericaeus (Blackwall). Flakstad0ya, Lofoten, 

at 68°06'N.; sandy beach near Flakstad; F; 

29 June 75. There are very few records of this 

species in Fennoscandia. although it is locally abundant 

in Western Norway (Hauge 1976). 

Lepthyphall/es mengei Kulczynski. Flakstad0ya, Lofoten; 

moorland close to sea level; M; 28 June 75. 

Lepthyphall/es zimmermanni Bertkau. V

Checklist of Norwegian ticks and mites (Acari) 1 

REIDAR MEHL 

INTRODUCTION 

Mehl, R. 1979. Checklist of Norwegian ticks and mites (Acari). Fauna Norvegica Ser. B. 26, 

31-45. 

The purpose of this faunistic checklist is to sum up all the published species of ticks and 

mites from Norway and give references so that the literature will be more accessible. 

The checklist includes 810 species belonging to the following six groups: Mesostigmata 

96, Ixodides 12, Trombidiformes (exc1. HydrachnelIae) 20 I, Hydrachnellae 183, Sarcoptiformes 

(excl. Oribatei) 74, and Oribatei 244. 

Reidar Mehl, Laboratory of Medical Entomology, Department of Environmental Toxicology, 

National Institute of Public Health, Postuttak, N-OsJo I, Norway. 

The knowledge of Norwegian ticks and mites is 

based upon numerous articles, many of which 

are old and often difficult to locate. 

The purpose of this faunistic checklist is to 

sum up all the published species and give references 

so that the literature will be more accessible. 

The result, I hope, will be a useful startingpoint 

for future investigations. A critical revision 

of identifications and nomenclature is beyond 

the scope of this survey. 

This compilation surely is incomplete, and information 

concerning publications and unpublished 

finds of mites from Norway will be 

highly appreciated. 

Mites from Spitzbergen are not included in this 

survey. 

In the old surveys of animals in Norway by 

Stmm (1768) ind Hammer (1775) some species 

of mites and ticks are mentioned, but it is difficult 

to recognize the species and the identifications 

are uncertain. Thor published numerous 

papers on soil mites and water mites from 1897 

until his death in 1937,andin 1943 Natvig published 

a bibliography of Thor's works, but some 

of his latest papers were not included. 

Foreign scientists have contributed knowledge 

to the mite fauna by publishing faunistical 

notes (Oudemans 1927, Willmann 1929, Dalenius 

1960, Forsslund in L0ken 1966), comprehensive 

surveys (Leatherdale 1959, CadwaJladr 

1969, Karppinen 1971) or have included material 

from Norway in their taxonomical works 

(Berlese 1904, 1923, Fain 1969, Fain et al. 

IConstribution No. 149 from Zoological Museum, University 

of Oslo. 

Fauna Norv. Ser. B 26: 31-45. Oslo 1979. 

1974). Leatherdale (1959) gave references (not 

repeated here) to many older faunistical publications 

concerning gall-making insects and mites 

in Norway. 

The Government Entomologists W. M. SCh0yen 

(I891-1912), 1. H. Sch0yen (1913-1941) 

and Fjelddalen (1951, 1953, 1960, 1962) included 

mites of-economic importance in their publications 

on agricultural pests and in their annual 

reports. Fjelddalen (1968) has included some unpublished 

gall-mites from his collections in the 

Checklist of Norwegian names of animals (Norske 

Dyrenavll.BJ 

Parasitic mites from domestic animals and 

humans have been published by Tambs-Lyche 

(I 943a, b), Nrerland (1950), Austad (1968), Tharaldsen 

(1973, 1978) and Mehl (1978). Some parasitic 

species were included in a report of a 

committee for pesticides against ectoparasites 

(Komite 1975). 

During the past ten years several Norwegian 

biologists have been studying mites. Publications 

from three projects are cited: The Norwegian 

International Biological Program (TBP) (Solh0y 

1975, 1976, Solh0Y et al. 1975); Acid Precipitation 

- Effects on Forests and Fish

after the initials means that the species have 

been recorded, but not published. 

In the checklist 1 have used the same scientific 

names as appeared in the literature cited. However, 

some alterations have been made, and some 

synonyms have been added. The list of the Oribatei 

is based on an unpublished list by the late 

Karl-H. Forsslund and corrected in accordance 

with the recent literature. 

When an author has used an other name of 

the species or the genus than the name on the 

list, the synonym is put in parenthesis after the 

reference. 

About 50 000 species of mites are known and 

in Norway we may expect to find about 2 000 

species. This checklist includes 810 species belonging 

to the following six groups: Mesostigmata 

96, Ixodides 12, Trombidiformes (excl. 

Hydrachnellae) 20 I, Hydrachnellae 183, Sarcoptiformes 

(excl. Oribateil 74, and Oribatei 244. 

ABBREVIATIONS: 

AB - Antonio Berlese, AE - Anders Edler, 

AF - Alex Fain, AL - Astrid L0ken, AO 

A. C. Oudemans, CH - Christopher Hammer, 

CS - Christian Stenseth, CW - C. Willmann, 

DAC - D. A. Cadwalladr, DL - Donald Leatherdale, 

DOlO - Dag Olav IOvstedal, EK - E. 

Karppinen, ES - Embrikt Strand, GN - G. 

Nrerland, HS - Hans Stmm, HTL - Hans 

Tambs-Lyche, JF - Jac. Fjelddalen, JT - Jorunn 

Tharaldsen, JIO - Jan IOkland, MM 

Manfred Moritz, OAS - Ole A. Srether, NVH 

- Komite nedsatt av Landbruksdepartementets 

rad for veterinrermedisin, plantevernmidler og 

formidler m.m., PO - Peter Dalenius, RM 

Reidar Mehl, SH - Sigmund Hagvar, SSS - G. 

Schulte, R. Schuster & H. Schubart, ST - Sig 

Thor, THS - T. H. Sch0yen, TS - Torstein 

Solh0y, WMS - W. M. Sch0yen, OH - Oddvar 

Helle 


I am greatly indepted to the following persons 

for valuable information about records of mites 

and literature: Jac. Fjelddalen, Sigmund Hagvar, 

Torstein Solh0y, Christian Stenseth and Jorunn 

Tharaldsen. Valuable technical assistance in preparing 

the manuscript was given by Inger H0iby. 

32 

MESOSTIGMATA 

I. Amblyseius Jugortus Athias-Henriot SH 

1978. 

2. A. obtusus (Koch) SH 1978. 

3. Androlaelaps casali casali (Berlese) AE 

+ RM 1972. 

4. A..fahrenholzi(Berlese) AE + RM 1972. 

5. Cyrtolaelaps minor Willmann AE + RM 

1972. 

6. C. mucronatus (G. & R. Canestrinil AE 

+ RM 1972. 

7. Dermanyssus alaudae (Schrank) RM. 

8. D. americanus Ewing RM. 

9. D. chelidonis Oudemans RM. 

10. D. grochovskae Zemskaya RM. 

11. D. gallinae (De Geed RM 1978. 

12. D. hirundinis (Hermann) RM 1978. 

13. D. quintus Vitzthum RM. 

14. Eulaelaps novus Vitzthum RM. 

15. E. stabularis (c. L. Koch) AB 1904a, 

AE + RM 1972, RM I 972e. 

16. Eviphis ostrinus (c. L. Koch) AE + RM 

1972, SH 1978. 

17. Gamasellus montanus (Willmann) SH 

1978. 

18. Haemogamasus ambulans (Thorell) AE 

+ RM 1972. 

19. H. hirsutus Berlese AE + RM 1972. 

20. H. horridus Michael AE + RM 1972. 

21. H. liponyssoides Ewing RM. 

22. H. nidi Michael AE + RM 1972. 

23. H. nidiformes Bregetova AE + RM 

1972. 

24. H. pantiger Berlese RM 1977. 

25. Halolaelaps marinus (Brady) AO 1927. 

26. Hirstionyssus isabellinus Oudemans AE + 

RM 1972. 

27. H. latiscutatus (de MeiJlon & Lavoipierrel 

AE + RM 1972. 

28. H. sciurinus (Hirst) RM 1971. 

29. H. soricis (Turk) AE + RM 1972. 

30. H. tatricus Mrciak AE + RM 1972. 

31. Hyperlaelaps amphibius Zachvatkin RM 

1970. 

32. H. microti (Ewingl AE + RM 1972. 

33. Hypoaspis aculeifer (Canestrinil SH 1978. 

34. H. krameri (Canestrinil RM. 

35. H. lubrica Voigts & Oudemans RM. 

36. H. marginalis (Willmann) RM. 

37. H. sardoa (Berlese) AE + RM 1972. 

38. Laelaps agilis C. L. Koch AE + RM 

1972. 

39. L. clethrionomydis Lange AE + RM 

1972. 

40. L. hilaris C. L. Koch AE + RM 1972.

6. A. calycophthira Nalepa JF 1968. 

7. A. centaureae(Nalepa) DL 1959. 

8. A. dispar (Nalepa) DL 1959. 

9. A. diversipunctatus (Nalepa) DL 1959. 

10. A. drabae(Nalepa) DL 1959. 

11. A. enanthlls (Nalepa) DL 1959. 

12. A. euaspis (Nalepa) DL 1959. 

13. A..f/liformis (Nalepa) DL 1959. 

14. A. geranii(Canestrini) DL /959. 

/5. A. iteinlls (Nalepa) DL 1959. 

16. A. kochi(Nalepa& Thomas) DL 1959. 

17. A.leionotus Nalepa DL 1959 (A. 

lissonotlls (Nalepa)). 

18. A. nervisequa Canestrini JF 1968. 

19. A. platanoidea Nalepa JF 1968. 

20. A. pseudoplatani Corti JF 1968. 

21. A. rhodiolae(Canestrinil DL 1959. 

22a. A. rudis longisetosus (Nalepa) DL I959. 

22b. A. rudis typicus (Nalepa) FL 1888 DL 

1959. 

23. Aceria saxifragae (Rostrup) DL 1959. 

24. A. scaber(Nalepa) JF 1962. 

25. A. stenaspis Nalepa JF 1968. 

26a. A. tetanothrix (Nalepa) DL 1959. 

26b. A. tetanothrix laevis (Nalepa) DL 1959. 

27. A. tiliae(Pagenstecherl DL 1959. 

28. A. vinosa Roivainen DL 1959. 

29. Alicorhagia plumipilus Thor ST 1931 b 

(Epistomalycus). 

30. Amorphacarlls elongatus (Poppe) RM 

1972. 

31. A. parvisetoslls Lukoschus & 

Driessen RM 1972. 

32. Anystis baccarum (L.) ST Ilf·12b AO 

1927. 

33. Bdella longicornis (L.) ST 1904. 

34. B. strandi Berlese AB 1923 ( + var. 

vistosa). 

35. Bimichaelia setigera (Berlese) ST 1931 b. 

36. B. subnuda (Berlese) ST 1931 b. 

37. Biscirus intermedius Thor ST 1928b. 

38. Bonzia halacaroides Oudemans AO 

1927, ST + CW 1941. 

39. Brachytydeus breviclllus (c. L. Koch) ST 

1931a, 1933. 

40. Bryobia praetiosa C. L. Koch JF 1952. 

41. Calepitrimerlls alchemillae (Lira) DL 

1959. 

42. Callyntrotus polygoni (Liro) DL 1959. 

43. Cecidophyes atrichus (Na[epa) DL 1959. 

44. C. betulae(Nalepa) DL 1959. 

45. C.ga!ii(Karpelles) DL 1959. 

46. C. nlldlls (Nalepa) DL 1959. 

47. C. ribis(Westwood) DL 1959,JF 1962 

(Cecidophyopsis). 

48. C. tetratrichus (Nalepa) DL 1959. 

34 

49. Cecidophyopsis ribis (Westwood) JF 

1962. 

50. entrotrombidium schneideri Kramer AO 

1927. 

51. Cheletomorpha venustissima (c. L. 

Koch) ST I912c. 

52. Cheyletiella blakei(Smiley) RM 1978. 

53. C. parasitivorax (Megnin) RM 1978. 

54. C. yasguri(Smiley) RM 1978. 

55. Cheyletus eruditlls (Schrank) ST 1912c. 

56. Cocceupodes clavifrons (Canestrini) ST 

1934a. 

57. C. curviclava Thor ST 1934b. 

58. C. mollicelllls(C. L. Koch) ST + CW 

1941. 

59. Coccotydells globifer Thor ST I931 a, 

1933. 

60. C. tenuiclavigerThor ST 1931a, 1933. 

61. Cunaxa setirostris (Hermann) ST 1912c. 

62. C. taurus (Kramerl ST 1912c. 

63. Cyta latirostris (Hermann) ST 1904. 

64. Demodex bovis Stiles OH 1972, NVH 

1975. / 

65. Demodex canis Leydig OH 1972, NVH 

1975. 

66. D. cati Megnin OH 1972. 

67. D. caprae Railiet GN 1950, NVH 1975. 

68. D. equi Railliet NVH 1975. 

69. DJolliculorum (Simon) OH. 

70. Enemothrombium caligigerlll11 

(Berlese) AB 1910. 

71. Eotetranychlls telarius L. JF 1968. 

72. Epitrimerus trilobus Nalepa JF 1968. 

73. Eriophyes exilis (Nalepa) DL 1959. 

74. E. gibbosus Nalepa JF 1962 DL 1959. 

75. E.goniothoraxNa1epa DL 1959. 

76. E. gracilis (Nalepa) JF 1962. 

77. E. inangulis(Nalepa) DL 1959. 

78a. E. laevis alni-incanae Nalepa DL 1959. 

78b. E. laevis typicus Nalepa DL 1959. 

79. E. leiosoma (Nalepa) DL 1959. 

80. E. loewi Nalepa JF 1968. 

81. E. macrochelus pseudoplatani Corti DL 

1959. 

82. E. maerorrhynchus cephaloneus 

Nalepa DL 1959. 

83. E. mali Nalepa JF 1962. 

84. E. malimarginemtorquens (Liro) DL 

1959. 

85. E. malinus Nalepa DL 1959 JF 1962. 

86. E. paderineus Nalepa DL 1959. 

87.E.padiNalepa DL1959JF1962. 

88. E. pyri(Pagenstecherl DL 1959 JF 1962. 

89. E. silvicola (Canestrinil JF 1962. 

90a. E. similis Na1epa DL 1959 JF 1962. 

90b. E. similis prunispinosae Nalepa DL 1959.

91. E. sorbeus (Nalepa) DL J959. 

92. E. sorbi

175. Smaris expalpis (Hermann) ST 1900b. 

176. Speleorchestesformicorum Tnigardh ST 

1913b. 

I 77. Spinbdella reducta Thor ST 19 30a. 

178. Tanaupodus passimpilosus Berlese AO 

1927. 

179. Tarsolarkus articulosus Thor ST 1912d. 

180. Tarsonemus pallidus Banks 1F 1952. 

181. Tetranychus cinnabarinus Boisd 1F 1968. 

182. T. urticaec. L. Koch1F 1952. (T. 

althaeae) CS 1976. 

183. Triophtydeus pinicolus (Oudemans) ST 

1932b, 1933. 

184. Trombicula zachvatkini Schluger RM. 

185. Trombidiumfilipes C. L. Koch ST 1900a. 

186. T.fuliginosum Hermann ST 1900a. 

187. T. holosericeum (L.) ST 1900a. 

188. Tydeus aberrans Oudemans ST 1932b, 

1933, AO 1932. 

189. T. albofasciatus C. L. Koch ST 1933. 

190. T. croceus(L.) ST 1912a, 1933(T. 

foliorum (Schrank)). 

191. T. interruptus Thor ST 1932b, 1933. 

192. T. subalpinus Thor ST 1932b, 1933. 

193. T. thori Oudemans ST 1932b, AO 1931. 

194. T. totensis Thor ST 1932b, 1933. 

195·. Vasates anthobius (Nalepa) DL 1959. 

196. V. comatus (Nalepa) 1F 1962. 

197. V.fockeui (Na1epa & Trouessartl DL 

19591F 1962. 

198. V. masseei (Nalepa) 1F 1962. 

199. V. myrsinitis Roivanen DL 1959. 

200. V. schlechtendaJi (Na1epal 1F 1962. 

20 I. Villersia vietsi Oudemans AOI927. 

HYDRACHNELLAE 

36 

I. Acercus torris (Muller) ST, OAS 1965. 

2. A. triangularis Piersig ST. 

3. Arrhenurus albator (Muller) 10 1964. 

4. A. bicuspidator Berlese ST,10 1964. 

5. A. caudatus De Geer ST. 

6. A. claviger Koenike ST. 

7. A. conicus Piersig ST. 

8. A. coronatorThor ST. 

9. A cuspidifer Piersig ST. 

10. A. emarginator Muller ST. 

11. A. errator Thor ST. 

12. A.finmarchicus Thor ST 1923. 

13. A.forpicatus Neuman 'ST. 

14. A. fragilis Thor ST. 

15. A. globator Muller ST. 

16. A. integrator(Muller) ST. 

17. A. kjermanni Neuman ST. 

18. A. maculator (Muller) ST. 

19. A. mediorotundatus Thor ST. 

20. A. membranatorThor ST. 

21. A. paluster Thor ST. 

22. A. primitivus Thor ST 1923. 

23. A. pustulator (Muller) ST. 

24. A. regulator Thor ST. 

25. A. stecki Koenike ST. 

26. A: stjoerdalensis Thor ST. 

27. A. truncatellus (Muller) ST. 

28. Atax crassipes (Muller) ST. 

29. A. kochiThor ST. 

30. Atractides spinirostris Thor ST. 

31 . Aturus scaber Kramer ST. 

32. Bandakia concreta Thor ST 1913a. 

33. Brachypoda versicolor(Muller) ST,10 

1964, OAS 1965. 

34. Curvipes alatus Thor ST. 

35. C. annulatus Thor ST. 

36. C. carneus C. L. Koch ST. 

37. C. clavicornis(Muller) ST. 

38. C. coccineus C. L. Koch ST,10 1964 

(Piona). 

39. C. coccinoides Thor ST. 

40. C. controversiosus Piersig ST. 

41. C. disparilis Koenike ST. 

42. C. laminatus Thor ST. 

43. C. luteolus Koch ST. 

44. C. neumani Koenike ST. 

45. C. pauciporus Thor ST. 

46. C. raropalpis Thor ST. 

47. C. rotundoides Thor ST. 

48. C. rotundus Kramer ST. 

49. C. stjoerdalensis Thor ST. 

50. C. uncatus Koenike ST. 

51. Diplodontus despiciens (Muller) ST, AO 

1927 10 1964 (Hydrodromal. 

52. Drammenia clongata Thor ST 1913b. 

53. D. crassipalpis Thor ST 1913b. 

54. Euthyas longirastris Piersig ST. 

55. Eylais angustipons Thor ST. 

56. E. cornuta Thor ST. 

57. E. discreta Koenike ST. 

58. E. duplex Thor ST. 

59. E. extedens (Muller) ST,10 1964, OAS 

1965. 

60. E.foraminipons Thor ST. 

61. E. infundibulifera Koenike ST. OAS 

1967. 

62. E. latipons Thon ST. 

63. E. muelleri Koenike ST,10 1964 (E. 

setosa muellerO. 

64. E. mutila Koenike ST. 

65. E. neglecta Thor ST. 

66. E setosa Koenike ST,10 1964. 

67. E. spinipons Thor ST. 

68. E. tenuipons Thor ST. 

69. E. undulosa thienemanni(Koenike) 10 

1964.

70. Feltria composita Thor ST. 

7 I. Forelia liliacea (Mliller) ST (A cercus), 

OAS 1965. 

72. Frontipoda musculus (Muller) ST. 

73. Gnaphiscus setosus Koenike ST. 

74. HJartdalia runcinata Thor ST 190 I a. 

75. Huitfeldtia rectipes Thor ST,OAS 1967. 

76. Hydrachna binominata Thor ST. 

77. If. biscutata Thor ST. 

78. If. cruenta (Muller) 10 1964. 

79. If. geographica (Muller) ST. 

80. If. globosa (De Geer) ST. 

81 . If. koenikei Thor ST. 

82. If. uniscutata Thor ST. 

83. Hydrochoreutes acutus Thor ST. 

84. If. incertus Piersig ST. 

85. If. ungulatus Koch ST,OAS 1965. 

86. Hydrophantes octoporus Koenike ST. 

87. Hydrophantes clypeatus Thor ST. 

88. If. dispar(von Schaub) ST. 

89. If. draco Thor ST. 

90. If. ruber(De Geer) .ST, OAS 1965. 

91. Hygrobates albinus Thor ST 1927b. 

92. If. calliger Piersig ST I 927b. 

93. If.jluviatilis (Stmm) HS 1768, ST 1928a. 

94. If. longipalpis (Hermann) ST 1927b, J0 

1964, OAS 1965. 

95. If.longiporus Thor ST 1927b. 

96. If. nigromaculatus (Lebert) ST I 927b. 

97. If. processifer Thor ST 1905d, ST 1923. 

98. If. reticulatus Kramer ST. 

99. If. squamifer Thor ST 1927b. 

100. If. trigonicus Koenike ST 1927b. 

101. If. walteriThor ST 1923. 

102. Kongsbergia materna Thor ST. 

103. Lebertia angulata Thor ST 1907d. 

104. L. brevipora Thor ST 1900e, 1906b. 

105. L. contracta Thor ST 1906b. 

106. L. expansa Thor ST 1923. 

107. L.fimbriata Thor ST 1900e, 1905a. 

108. L. glabra Thor ST 1900e, 1907a. 

109. L. insignis Neuman ST 1900e, 1906a. 

110. L. obscura Thor ST 1906a. 

I 11. L. oudemansi Koenike ST. 

112. L. porosa Thor ST 1906a. 

113. L. prolongata Thor ST 1923. 

114. L. stigmatifera Thor ST 1907d. 

115. L. theodorae Thor ST 1907d. 

116. Lebertia vietsiThor ST 1923. 

117. L. vigintimaculata Thor ST. 

118. Limnesia connata Koenike ST. 

119. L. histrionica Hermann ST,10 1964 (L. 

fulgida C. L. Koch) 

120. L. koenikei Piersig ST. 

121. L. lorea Thor ST. 

122. L. maculata (Muller) ST, J0 1964, OAS 

1965. 

123. L. undulata (Muller) ST. 

124. Limnochares aquatica (L.) ST (L. 

holosericea Latreille) 10 1964. OAS 1965. 

125. LJania bipapillata Thor ST. 

126. Megapus nodipalpis Thor ST. 

127. M. spinipes (c. L. Koch) ST. 

128. M. tener Thor ST. 

129. Mesobatesforcipates Thor ST 190 I a. 

130. Midea elliptica Muller ST. 

131. M. orbiculata (Muller) J0 1964. 

132. Mideopsis orbicularis (Muller) ST. 

133. Neumania deltoides (Piersig) ST (N. 

mirabilis (Neuman», OAS 1965. 

134. N. setosa Thor, 190 I ST. 

135. N. spinipes (Muller) ST. 

136. N. vernalis (Mi.iller) ST, OAS 1965. 

I 37. Oxus koenikei Thor ST. 

138. O. ovalis (Muller) ST. 

139. O. plantaris Thor ST. 

140. Panisus michaeli Koenike ST. 

141. P. thoracatus (Piersig) ST. 

142. Piona bipora Thor ST 1923. 

143. P. bullata Thor ST. 

144. P. conglobata (c. L. Koch) ST (Curvipes), 

10 1964, OAS 1965. 

145. P. latipes Muller ST. 

146. P. longipalpis (Krendowsky) ST (Curvipes 

brunzella Thor), 10 1964, OAS 1965. 

147. P. nodata(Muller) ST(Curvipesfuscatus 

Herman). OAS 1965. 

148. P. ornata (c. L. Koch) ST,10 1964 

(A cercus). 

149. P. scaura Koenike ST. 

150. P. variabilis (c. L. Koch) ST (Curvipes 

rufus C. L. Koch), OAS 1965. 

151. Pionacercus leuckarti Piersig ST. 

152. P. norvegicus Thor ST. 

153. P. scutatus Thor ST. 

154. P. sinuosus Thor ST. 

155. P. uncinatus Koenike ST. 

156. Pionides ensifer Koenike ST. 

157. Pionopsis lutescens Hermann ST,OAS 

1965. 

158. Pseudothyas trabecula Thor ST. 

159. Pseudoxus integer Thor ST. 

160. Rivobates norvegicus Thor ST. 

161. Sperchon brevirostris Koenike ST. 

162. S. elegans Thor ST. 

163. S. glandulosus Koenike ST. 

164. S. lineatus Thor ST. 

165. S. multiplicatus Thor ST. 

166. S. norvegicus Thor ST 1923. 

167. S. papillosus Thor ST 1901a. 

168. S. setigerThor ST. 

169. S. squamosus Kramer ST. 

170. S. tenuabilis Koenike ST. 

37

171. S. thori Koenike ST. 

30. Hypodectes propus (Nitzsch) RM. 

172. Sperchonopsis verrucosa (Protz) ST. 31. Knemidocoptes laevis gallinae (Railliet) 

173. Teutonia cometes (Koch) J0 1964. 

THS 1929. 

174. T. primaria Koenike, ST. 

32. K. mutans (Robin & Lanquetin) OH 1972. 

175. T. subalpina Thor ST. 

33. Kuzinia laevis (Dujardin) RM. 

176. Thyas barbigera Viets J0 1964. 

34. Labidophorus sciurinus (C L. Koch) RM 

177. T. delltata Thor ST. 

1971. 

178. T. pachystoma Koenike J0 1964. 

35. Leporacarus brevicaudatus (Ullrich) RM. 

179. T. pustulosa Thor ST. 

36. L. gibbus (Pagenstecher) RM. 

180. T. stoW Koenike, ST. 

37. Listrophorus americanus Radford RM. 

181. Thyas valvata Thor ST. 

182. Unionicola intermedia (Koenike) J0 

38. L. dozieri Radford RM. 

39. L. leuckarli Pagenstecher RM. 

1964. 

40. L. mustelae Megnin RM. 

183. Zschokkae oblonga Koenike ST. 

41. L. pagenstecheri Hailer RM J971. 

42. L. synaptomys Fain, Whitaker, McDaniel & 

Lukoschus AF et a1. 1974. 

SARCOPTIFORMES (excI. OribateO 

43. Melesodectes auricularis Fain & Lukoschus 

RM 1972. 

I. Acarus siro L. WMS 1894-1913, THS 44. Myocoptes j. japonensis Radford RM. 

1914-1941, RM 1978. 

45. M. musculinus (C L. Koch) RM. 

2. Alloptes bisetatus (Hailer) RM. 

46. Monojoubertia hemiphylla (Robin) RM. 

3. Apodemopus apodemi (Fain) RM. 

47. M. mierophylla (Robin) RM. 

4. Brephosceles formosus Cerny RM. 

48. Neottialges evansi Fain RM. / 

5. Bucholzia fusca Nitzsch RM. 

49. N. vitzthumi Fain, RM. 

6. Buchovskiata charadrii (Canestrini) RM. 50. Notoedres cali (Hering) NVH. 

7. B. squatarolae (CanestrinD RM. 

51. N. muris Megnin RM. 

8. Caparinia tripilis (MichaeI) RM I972b. 52. Orycteroxenus soricis (Oudemans) RM. 

9. Carpoglyphus lactis (L.) THS 1917 (c. pas 

53. Otodectes cynolis (Hering) RM 1972. 

sularum), RM 1978. 

54. Picidectes RM. 

10. Chirodiscoides caviae Hirst RM. 

55. Pomeranzevia numenii (Canestrini) RM. 

11. Chorioptes bovis (Hering) OH 1972, NVH 56. Proctophyllodes picae C L. Koch, RM. 

1975. 

57. Psorergates RM. . 

12. Criniscansor apodemi Fain, Munting & Lu 58. Psoroptes cuniculi (Delafond) RM. 

koschus RM. 

59. P. equi (Hering) NVH 1975. 

13. Dermacarus caucasicus Fain RM. 

60. P. ovis (Hering) GN 1950. 

14. D. hypudei septentrionalis Fain AF 1969. 61. Rhizoglyphus echinopus (Fumouze & Ro 

15. D. norvegicus Fain, Philips & Wilson AF et bin) THS 1926. 

al. 1979. 

62. Sarcoptes scabiei (L.) ES 1876, RM 1978. 

16. Dermatophagoides chelidonis (Hull) RM. 63. Sokoloviana rehbergi (Canestrini & Berlese) 

17. D. farinae Hughes RM 1973. 

RM. 

18. D. pteronyssinus (Trouessart) RM 1973. 64. Sulanyssus caput-medusae (Trouessart) 

19. D. mieroceras Griffiths & Cunnington RM. RM. 

20. Euroglyphus maynei (Cooreman) RM. 65. Trichoecius apodemi Fain, Munting & Lu 

21. E. longior (Trouessart) RM. 

koschus RM. 

22. Gliricoptes betulina Kok, Lukoschus & Fain 66. T. tenax (MichaeI) RM. 

RM. 

67. Tyrophagus longior (Gervais) WMS 1907, 

23. Glycyphagus destructor (Schrank) THS 1911, THS 1916-26, RM. 

1915, 1916, 1922 (G. spinipes) RM 1978. 68. T. putrescenliae (Schrank) RM. 

24. G. domesticus (De Geer) WMS 1904-12, 69. Thecarthra setigera maritimae W. Dubinin 

THS 1915-30, RM 1978. 

RM. 

25. Gohieria fusca (Oudemans) RM. 

70. Xenocastor fedjushini Zachvatkin RM. 

26. Harpyrrhyncus rubeculinus Cerny & Six1 71. Xenoryctes krameri (MichaeI) RM. 

RM. 

72. X. punctatus Fain RM. 

27. Histiophorus capitis Dubinina RM. 

73. Zachvatkinia stercorarii W. Dubinin RM. 

28. H. fedjushini Dubinina RM. 

29. H. latus Dubinina RM. 

74. Zachvatkiniana lutrae Volgin RM. 

38

ORIBATEI 

28. C. foveolata Hammer TS 1976b. 

29. C. horrida (Hermann) ES 1910 and ST 

1. Achipteria coleoptrata (L.) ES 1910 (Oriba 

1937 (Nothrus), TS 1975. 

tes ovalis), CW 1929, ST 1937, AL 1966, 

30. C. invenusta (MichaeI) AL 1966, TS + D00 

OAC 1969, EX 1971. 

1979. 

2. A. italicus Oudemans ( =A. willmanni van 

31. C. lapponica (Tragardh) ST 1937, OAC 

der Hammen) ST 1937, AL 1966, OAC 

1969, EK 1971, TS 1975,1976. 

1969, TS + 000 1979 (Para chipteria). 

32. C. spinifer (c. L. Koch) ST 1937, DAC 

3. A. magna (Sellnick) EK 1971. 

1969, EK 1971, TS+000 1979. 

4. A. punctata (Nicolet) ST 1937, OAC 1969, 

EK 1971, TS 1976a (Parachipteria). 

33. Carabodes affinis Ber1ese ST 1937. 

5. A. sellniki van der Hammen ST 1937 (A. ni 

34. C. areolatus Berlese ST 1937, EK 1971. 

tens (Nicolet». 

35. C. coriaceus C. L. Koch ST 1937. 

6. Aerogalumna longiplumus (Berlese) AL 36. C. femoralis (Nico1et) ST 1937, EK 1971, 

1966. 

SH 1978. 

7. Adoristes ovatus (c. L. Koch) CW 1929, ST 37. C. forsslundi Sell nick EK 1971, SH 1978. 

1937, OAC 1969. 

38. C. labyrinthicus (MichaeI) CW 1929, ST 

8. A. poppedOudemans) EK 1971 (in ST 1937 1937, EK 1971, TS 1975. 

as part of A. ovatus sensu EK), TS + 000 39. C. marginatus (MichaeI) ST 1937, DAC 

1979. 

1969, EK 1971, TS et al. 1975, TS 1976, 

9. Ameronothrus bilineatus (Michael) SSS SH 1978, TS + D00 1979. 

1975. 

40. C. minusculus Berlese ST 1937, EK 1971, 

10. A. Iineatus (ThoreU) ES 1910, SSS 1975. TS et al. 1975, TS 1976a, TS + D00 1979. 

11. A. maculatus (Michael) SSS 1975, 41. C. supalpinus Thor ST 1937. 

TS + 000 1979. 

42. C. subarcticus Tragardh EK 1971, SH 

12. A. marinus (Banks) SSS 1975. 

1978. 

13. A. nigrofel11oratus (L. Koch) SSS 1975. 43. Cepheus cepheiformis (Nicolet) ST 1937. 

14. Astegistes pilosus (c. L. Koch) ES 1910 (Li 

44. C. dentatus (MichaeI) ST 1937, AL 1966. 

acarus bicornis). 

45. C. latus C. L. Koch ST 1937. 

15. Autogneta parva Forss1und EK 1971. 46. Ceratoppia bipilis (Hermann) ES 1910 (No 

16. A. traegaardhi Forsslund. A. longilamellata taspis) ST 1937, EK 1971, TS 1976, 

(MichaeI) in ST 1937 is probably A. trae 

TS + D00 1979. 

gaardhi sensu EK 1971. 

47. C. sphaerica C. L. Koch DAC 1969. 

17. Belba compta (Kulczynskj) ST 1937 (Da 

48. Ceratozetes cisalpinus Berlese ST 1937, EK 

maeus), OAC 1969, EK 1971. 

1971. 

18. B. corynopus Hermann ST 1937 (Notaspis). 49.C. gracilis (MichaeI) ST 1937, EK 1971, 

19. B. crispata'l.Kulczynskj) ST 1937. 

TS + D00 1979. 

20. B. moll/ana (Kulczynskj) ST 1937 (Damae 

50. C. medioeris Berlese ST 1937. 

us). 

51. C. parvulus Sellnick ST 1937 (Allozetes). 

21. Brachychochthonius immaculatus Fors 52. C. thienemanndWillmann) OAC 1969, EK 

slund OAC 1969 (B. semiornatus G. O. 1971. 

Evans), MM 1976b. 

53. Chamobates cuspidatiformes (Tragardh) 

22. B. zelawaiensis Sellnick OAC 1969, AL 1966, TS 1976a, TS + D00 1979. 

TS + 000 1979. 

54. C. cuspidatus (Michael) ST 1973, TS 

23. Caleremaeus monilipes (MichaeI) ST 1937, 1976a, TS + D00 1979. 

OAC 1969, EK 1971. 

55. Chamobates lapodarius (Lucas) AB 1923, 

24. Calyptozetes sarekensis (Tragardh) ST ST 1937. 

1937, PO 1960, AL 1966, DAC 1969, EK 56. C. schuetzii (Oudemans) ST 1937, AL 

1971, TS 1975. 

1966, and 1969 (C. borealis (Tragardh», 

25. Camisia biurus (c. L. Koch) ST 1937 (C. EK 1971, TS 1975. 

segnis), AL 1966, DAC 1969 (c. segllis), 57. Cultroribula juncata (MichaeI) ST 1937. 

EK 1971, TS 1975, TS 1976a, TS+000 58. Cymbaeremaeus cymba (Nicolet) ES 1910, 

1979. 

ST 1937. 

26. C. biverrucata (c. L. Koch) ST 1937 (Not 

59. Damaeus c1avipes (Hermann) ES 1910, ST 

hrus) 

1937 (Notaspis), AL 1966, TS 1976a (Para 

27. C. borealis (Thorell) EK 1971. 

damaeus). 

39

180. P. borealis (Tragardh) ST 1937, OAC 1969, 213. S. striculus (C L. Koch) CW 1929, ST 

EK 1971. 

1937 (Hoploderma), AL 1966, OAC 1969 

181. P. clavatus Parry TS + 000 1979. 

(Hoploderma), EK 1971, TS 1976, 

182. P. globosus (C L. Koch) EK 1971, TS + 000 1979. 

TS + 000 1979. 

214. Suctobelba acutidens Forsslund TS + 000 

183. P. italicus (Oudemans) ES 1910 (Hoplo 

1979. 

derma dasypus), EK 1971. 

215. S. intermedia Willmann EK 1971. 

184. P. laevigatus (C L. Koch) ST 1937 (Hoplo 

216. S. longirostris Forsslund OAC 1969, EK 

dermal. 

1971. 

185. P. Iigneus Willmann EK 1971. 

217. S. perforata (Strenzke) OAC 1969, 

186. P. pavidus (Berlese) ST 1937, EK 1971. TS + 000 1979. 

187. P. piger(Scopoli) ST 1937, OAC 1969, EK 218. S. sarekensis Forsslund EK 1971, 

1971, TS 1975. 

TS + 000 1979. 

188. P. rectisetosus Parry TS + 000 1979. 218. S. similis (Forsslund) TS 1976, TS + 000 

189. P. subglobosus Berlese AB 1923. 

1979. 

190. P. tardus Forss1und EK 1971. 

220. S. subcornigera Forsslund EK 1971, TS 

191. Platynothrus capillatus (Berlese) EK 1971, 1976, TS + 000 1979. 

TS 1976 (== Ovonothrus septentrionalis 221. S. trigona Michael ST 1937. 

(Sellnick)). 

222. Synchthonius crenulatus (Jacot) OAC 1969 

192. P. peltifer (C L. Koch) CW 1929, ST 1937, (S. boschmai van der Hammen sensu Mo 

PO 1960, AL 1966, OAC 1969, EK 1971, ritz 1976 b). 

TS 1975, TS 1976, TS + 000 1979. 

193. P. punctatus (C L. Koch) ST 1937. AL 

223. Tectocepheus cuspidentatus 

TS+0001979. I 

Kniille 

1966, EK 1971, TS 1975. 

224. T. sarekensis Tragardh PO 1960, EK 1971. 

194. P. thori (Berlese) AB 1904a (Angelia), ST 225. T. velatus (Michael) CW 1929, ST 1937, 

1937 (Heminothrus), AL 1966, OAC 1969, AL 1966, OAC 1966, EK 1971, TS 1975, 

EK 1971. 

TS 1976, SH 1978, TS + 000 1979. 

195. Porobelba spinosa (Sellnick) ST 1937 (Da 

226. Tegoribates latirostris (C L. Koch) PO 

maeus), TS 1975. 

1960, EK 1971. 

196. Pseudotritia monodactyla Willmann ST 227. Thyrisoma lanceolatum (Michael) PO 1960 

1937. 

(Oribella), TS + 000 1979 (Banksinoma). 

197. Punctoribates punctum (L. Koch), ST 1937. 228. Trhypochthonius cladonicola (Willmann) 

198. Quadroppia quadricarinata (Michael) ST ST 1937. 

1937 (Oppia), OAC 1969 (Oppia) EK 1971, 229. T. tectorum (Berlese) EK 1971. 

TS 1976, TS + 000 1979. 

230. Trichoribates incisellus (Kramerl EK 1971. 

199. Rhysotritia ardua (C L. KQch) ES 1910 231. T. monticola (Tragardh) AL 1966. 

(Ph thiracarus), ST 1937, EK 1971. 232. T. numerosus (Sellnick) EK 1971. 

200. R. duplicata Grandjean TS + 000 1979. 233. T. oblongus (L. Koch) ST 1937 (Murcia). 

201. Scapheremaeus palustris (Sell nick) ST 234. T. setiger (Tragardh) TS 1975. 

1937. 

235. T. tectopedacutus (Thor) ST 1937 (Murcia). 

202. S. frigidus (Ber]ese) AB 1908. 

236. T. trimaculatus (C L. Koch) ST 1937 (Mur 

203.S. laevigatus (C L. Koch) ST 1937, AL cia), AL 1966, EK 1971, TS+000 1979. 

1966, EK 1971. 

237. Trimalaconothrus glaber (Michael) ST 

204. S. latipes (C L. Koch) ST 1937, EK 1971. 1937. 

205. S. pallidulus (C L. Koch) PO 1960, OAC 238. T. indusiatus Berlese, ST 1937. 

1969, EK 1971. 

239. T. novus Sellnick CW 1929, ST 1937, EK 

206. Scutovertex bidactylus Coggi OAC 1969. 1971. 

207. S. minutus (C L. Koch) ST 1937, EK 1971. 240. Tritegeus bifidatus (Nicolet) ST 1937, 

208. Sphaerozetes orbicularis (C L. Koch) ST TS + 000 1979 (T. bisulcatus Grandjean). 

1937. 

241. Tropacarus carinatus (C L. Koch) ST 

209. S. piriformis (Nicolet) ST 1937, TS 1976, 1937, EK 1971. 

TS + 000 1979. 

242. Xenillus tegeocranus (Hermann) ES 1910 

210. Steganacarus applicatus (Sellnick) ST 1937 (Cepheus), ST 1937, EK 1971, TS + 000 

(Hoploderma), EK 1971. 

1979. 

211. S. magnus (Nico]et) ST 1937 (Hoploderma). 243. Zetorchestes micronychus (Berlese) ST 

212. S. spinosus (Sellnick) ST 1937 (Hoploder 

1937. 

mal. 

42

( Short communicatiOn) 

NEW RECORDS OF ISOPERLA DlFFORMIS 

(KLAPALEK, 1909)(PLEC., PERLODIDAE) 

FROM WESTERN NORWAY. 

TROND ANDERSEN 

Three records of Isoperla difJormis from outer Hordaland 

are given. 

Trond Andersen, Museum of Zoology, N-5014 

Bergen - University, Norway. 

In his study on the distribution and ecology of 

the Norwegian Plecoptera, Lillehammer (I 974) 

stated that lsoperla difformis (KlapaIek, 1909) 

«has only once been taken in western Norway». 

According to his map of the distribution the locality 

is in the Stavanger area. However, Brekke 

(I 94 I), in his review of the Norwegian stoneflies, 

recorded /. difformis from Hordaland (Fana). 

A female of /. difformis, identified by 

Brekke, is deposited in the collection at the Museum 

of Zoology in Bergen. The label reads: 

Skjold, Fana, 9 May 1937, N.Knaben leg. 

On 22 May 1973 I collected a micropterous 

male of /. difformis at Fitjo (UTM: 

32VLN098 I 56), HOy: Oster0Y (60 m a.s.U. The 

specimen was collected by sweeping the vegetation 

along a small, calm river. Later the same 

day I collected a second specimen of/. difJormis, 

a female, at Kleppe (UTM: 32VLNI11161), 

HOy: Oster0Y (250 m a.s.U. This specimen also 

was taken by sweeping the vegetation along 

another, more rapidly running stream. 

/. difformis is distributed throughout Central 

and Northern Europe (Illies 1967). It is common 

and widely distributed in the southern and central 

parts of Sweden and occurs, but rarely, also 

in the northern mountainous parts (Brinck 

1949). In Norway the species is common in the 

inner Oslofjord area, and is also found sporadically 

in other parts of southern and northern 

Norway (Lillehammer 1974) From my findings 

it can be concluded that /. difJormis is a species 

also inhabiting the outer parts of western Norway. 

However, as a rather large lsoperla material 

from outer as well as inner Hordaland has 

been identified without encountering more than 

three specimens, it seems obvious that I. difJormis 

is very rarely found within this area. 

46 

REFERENCES 

Brekke, R. 1941. The Norwegian stonellies. Plecoptera. 

Norsk ent. Tidsskr. 6, 1-24. 

Brinck, P. 1949. Studies on Sweedish stoneflies (Plecoptera). 

Opusc. ent., suppl. 11, 250 pp. 

lilies, J. 1967. Plecoptera. In: lilies, J. 

(ed.),Limnofauna Europaea. G. Fischer Verlag, 

Stuttgart, pp. 220-229. 

Lillehammer, A. 1974. Norwegian stoneflies. It. Distribution 

and relationship to the environment. 

Norskent. Tidsskr. 21.195-250. 


MASOREUS WETTERHALLl GYLLENHAL 

(COL., CARABIDAE) NY ART I NORGE. 

KARL ERIK ZACHARIASSEN 

The carabid beetle Masoreus wetterhalli Gyllenhal, is 

reported found in Norway for the first time. One 

female specimen was found on saI)dy ground near 

the sea in Stavern, Vestfold county, on 8 July 1978. 

Karl Erik Zachariassen, Zoologisk institutt, Universitetet 

i Trondheim, NLHT; Rosenborg, 7000 Trondheim. 

Lopebillen Masoreus wetterhalli Gyllenhal, lever 

pa torre, sandige lokaliteter (Landin, 1970), 

gjerne pa spredt bevokst flyvesand mer havet 

(Hansen, 1968). Arten er vidt utbredt i Danmark 

og i det sydlige Sverige, bt.a. i Bohusliin (Lindroth, 

1960), men er hittil ikke rapportert fun net 

i Norge. 

Ett eksemplar av M. wetterhal/i ble 8.7.1978 

funnet av forfatteren pa Risoya ved Stavern. Dyret 

ble fun net under en planke, som la pa spredt 

bevokst sand, noen fa meter fra stranden. Dyret 

var en hunn med lengde 4,8 mm. Det oppbevares 

i forfatterens samling. 

Lokaliteten der dyret ble funnet er dessverre i 

ferd med a bli odelagt, idet bukten funnstedet 

ligger ved er under gjenfylling. Det er imidlertid 

l1ere Iignende biotoper i omradet, der arten ogsa 

kan vrere a finne. 

REFERANSER 

Hansen, V. 1968. Biller XXIV. Sandspringere og Lobebiller. 

Danm. Fauna 76, 451 pp. 

Landin, B. -0. 1970. Fiiltfauna/lnsekter 2. Natur 

och Kultur. KunskapsfOrlaget, Stockholm. 

Lindroth, C.H. (ed.) 1960. Catalogus Coleopterorum 

Fennoscandiae et Daniae. Entomologiska Siillsk., 

Lund. 


Fauna Norv. Ser. B 26. Oslo 1979.

BEMBIDJON NIGRICORNE GYLLENHAL vis 10S og nesten t0rr. Vegetasjonen der dyra ble 

(COL., CARABIDAE) NY FOR NORGE. funnet var sparsom med flekkvis sauesvingel 

(Festuca ovina) og meget lav mose. Billene ble 

JOHAN ANDERSEN 

tatt under lauv eller barkflak (overskyet veer 

OG KARL ERIK ZACHARIASSEN 

med vind). Undervegetasjonen rundt de sparsomt 

bevokste feltene bestod av tildels tette 

Bembidion nigricorne Gyllenhal, (CoI.Carabidae) is klynger av f0ss1yng (Calluna vulgaris), mens 

reported new to Norway from S0rumsand, Akershus 

skogen utgjordes av gran og furu. B.nigircorne 

province. Four specimens were found in a habitat typical 

for the species. 

ble funnet sammen med Amara tibialis Paykall. 

Den beskrivelsen av habitatet for B. nigri 

Johan Andersen, Institutt for biologi og geologi, Unicorne som er gitt her, er i bra samsvar med den 

versitetet i Troms0, N-9000 Troms0, 

Lindroth (I 945) gir for arten ellers i Fennoskan 

Karl Erik Zachariassen, Zoologisk Institutt, Universitetet 

i Trondheim, NLHT, Rosenborg, N-7000 

dia. 

Trondheim. 

B. nigricorne ansees for a veere en 0stlig art i 

Fennoskandia og Danmark, men den er vidt ut 

I et sandtak pa S0rumsand, Akershus fylke, fant bredt fra s0rlige del av Sverige og Finland nord 

den ene av forfatterne (K.E.Z.) I. august 1973 et til Kalix (Lindroth pers. com.). Funnet ved S0 

eksemplar av Bembidion nigricorne Gyllenhal. rumsand er det vestligste i Fennoskandia, men 

Dette er det f0rste funn av arten i Norge. Stedet dette forandret ikke noe vesentlig pa det 0stlige 

ble atter bes0kt 12/5 -79. Det ble da ialt samlet utbredelsesm0nster arten viser i den nordlige de 

fire sma, m0rkt metalliske eksemplarer av Bemlen 

av sitt utbredelsesomrade. 

bidion sp. Et av disse eksemplarene var B. lampros 

Herbst, de 0vrige tre var B. nigricorne. LITTERATUR 

Tre av eksemplarene av B. nigricorne (de funnet 

12/5 - 79) ble tatt i utkanten av sandtaket 

eller i soleksponert felt i en skogsglenne mer 

dette. I begge tilfelle bestod jordbunnen av mo 

Lindroth, C.H. 1945. Die fennoscandischen Carabidae. 

I. GfJleborgs K. Velensk. - o. Vitlerh. samh. 

Handl. Ser. B4, 709 pp. 

renesand, delvis med noe grus og stein pa overflaten. 

Jorda varierte en del i fuktighet og var del 

Received 10. July 1979. 

Fauna Norv. Ser. B 26. Oslo 1979. 

47

( Bokanmeldelser ) 

Frolich, G., R. Beyer und E.-M. Kleiner 1979. Phytopathologie 

und Pflanzenschutz. Wiirterbiicher der 

Biologie. VEB Gustav Fisher Verlag Jena (295 sider, 

62 fig., Bestellnr. 532869 I, pris 19DM). 

Gustav Fisher forlag er i gang med a gi ut ein serie 

sma, hendige ordb0ker i biologi, - «Det biologiske 

fagomracte i leksikalsk framstilling». 

Det undrar kanskje somme at ei bok i serien med 

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Her tar vi ikkje standpunkt til kva 

som er rett sprakbruk, men merkjer oss at boka omfattar 

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Entomologien kjem da inn som ei viktig grein av fagomradet. 

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48 

opplegget ser det ut til at hovudvekta er lagt pa generelt 

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Holozyklie - Gegens.: Anholozyklie, Antagonismus 

- Gegens.: Synergismus, Optimum - Gegens.: Pessimum. 

I det heile er dette ei bok som er innhaldsrik etter 

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La oss som d0me slutte av med «Terminalia: Endsegmente 

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Analsegmente. Oppslag pa Analsegment gir tilvising 

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til a tvile pa om det var det stafetten starta med. 

Gudmund TaksdaI

Some caddis flies (Trichoptera) in western Norway, and - Norsk ...

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