An Introduction to the Botanical Type Specimen Register

SMITHSONIAN CONTRIBUTIONS TO BOTANY NUMBER 12 

An Introduction to the Botanical 

Type Specimen Register 

Stanwyn G. Shetler 

with Mary Jane Petrini, Constance Graham Carley, 

M. J. Harvey, Larry E. Morse, Thomas E. Kopfler, 

and Collaborators 

SMITHSONIAN INSTITUTION PRESS 

City of Washington 

1973

ABSTRACT 

Shetler, Stanwyn G., with Mary Jane Petrini, Constance Graham Carley, M. J. 

Harvey, Larry E. Morse, Thomas E. Kopfler, and Collaborators. An Introduc- 

tion to the Botanical Type Specimen Register. Srnithsonian Contributions to 

Botany, number 12, 186 pages, 3 figures, frontispiece, 1973.--In the first part, 

the development of a computer-based system for storing and retrieving infonna- 

tion about botanical type specimens is described from its pilot stage to its present 

operational stage. The concept, purpose, and scope are explained, and the 

operational procedures are outlined. Ways of using and contributing to this 

computerized register of types, both in the short-run and in the longrun, are 

proposed. A statistical summary of the content of the Type Register as of 30 

September 1972 is given. Over 13,000 specimens representing more than 10,000 

taxa have been registered. The second part consists of a Catalog of more than' 

1,000 specimens representing over 600 taxa of the genus Carex (Cyperaceae), 

which are deposited in ten major American herbaria, and the Catalog is cross- 

indexed five different ways: by author, publication date, collector, country, and 

herbarium. An introduction summarizes the preparation and editing of the 

Catalog. This Carex Catalog represents the first published installment of the 

Type Register and as such is intended to serve as an example. 

COLLABORATORS 

Listed here by institution are the persons who have collaborated in the compiling of 

data from their respective herbaria for the Carex Catalog: 

California Academy of Sciences, Golden Gate Park, Sun Francisco, California 94118; 

Stanford University (Dudley Herbarium), Stanford, CaGifornia 94305; University of Cali- 

fornia (Herbarium) , Berkeley, California 94720 

Dr. John H. Thomas, Associate Professor of Biological Sciences and Curator of Dudley 

Herbarium (Stanford), Curator of Botany (Acad-my) , and compiler of data at Berkeley; 

assisted by Dr. Margaret Sharp 

Field Museum of Xatural History, Roosevelt Road at Lakeshore Drive, Chicago, Illinois 

60605 

Dr. Lt'illiam C. Burger, Associate Curator of Vascular Plants: assisted by Mr. Ronald L. 

Liesner 

Harvard University, 22 Divinity Avenue, Cam bridge, Massachusetts 02138 

Dr. Richard 4. Howard, Director, Arnold Arboretum; Dr. Reed C. Rollins, Asa Gray 

Professor of Systematic Botany and Director of the Gray Herbarium 

Missouri Botanical Garden, 2315 Tower Grove Avenue, St. Louis, Missouri 63110 

Dr. IValter H. Lewis, Senior Botanist, Professor of Biology (Washington University); 

assisted by Dr. Raymond F. Altevogt 

h'ew York Botanical Garden, Bronx Park, New York 10458 

Dr. Howard S. Irwin, Director: Dr. Patricia Kern Holmgren, Associate Curator and 

Herbarium Administrator; Dr. Noel H. Holmgren, Associate Curator 

Smithsonian Institution, Washington, D. C. 20560 

Dr. Mason E. Hale, Curator of Cryptogams; Dr. Dan H. Nicolson, Associate Curator of 

Phanerogams 

OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is recorded 

in the Institution's annual report, Smithsonian Year. SI PRESS NUMBER 4783. SERIES 

COVFR DESIGN: Leaf clearing from the katsura tree Cercidiphjlluin japonicum Siebold and 

luccarini. 

Library of Congress Cataloging in Publication Data 

Shetler, Stanwyn G. 

An introduction to the botanical type specimen register. 

(Smithsonian contributions to botany, no. 12.) 

Bibliography: p. 

1, Information storage and retrieval systems-Botany-Type specimens. 2. Carex. I. Title. 11. 

Title: Botanical tjpe specimen register. 111. Series: Smithsonian Institution. Smithsonian 

contributions to botany, no. 12. 

QKlS2747 no. 12 [Z699.5.B6] 58lt.08s[579] 72-11706 

For sale by the Superintendent of Documents, US. Government Printing Office, Washington. D.C. 20402 

Price $2.85 domestic postpaid or $2.50 GPO Bookstore

Foreword 

It always has been the policy of the United States National Herbarium to 

make its collections as easily available as possible. We have welcomed visitors 

and loaned specimens on request since the founding of our herbarium. Now, 

in an effort to make our collections even more accessible, we have undertaken 

a new project to compile a computerized catalog of our type collection of ap- 

proximately 65,000 specimens. Eventually, we hope to broaden this catalog, 

which we are calling the “Botanical Type Specimen Register,” to include the 

type collections of many other institutions so that it will serve as a union listing 

of types. Already we have enlisted the cooperation of other institutions, and 

the computer file presently includes records from more than a score of herbaria. 

The United States National Herbarium, a worldwide collection of plants now 

totaling some 3 million specimens, is administered by the Smithsonian Institu- 

tion’s Department of Botany, a unit of the National Museum of Natural History. 

The Department of Botany has played a pioneering role in the development 

of the Museum’s active program in data processing, The Type Register is the 

Museum’s first operational effort in cooperative, multi-institutional (network) 

data banking and, as such, is of special interest. If this approach to common 

data banking proves successful, it \vill point the way for many cooperative efforts 

in other branches of natural history. 

Although the Type Register is still very much in its infancy, we are zealous 

to demonstrate its potential to the botanical community with a tangible product 

so that we can receive advice and counsel from the community on the basis of 

concrete results while the data bank is still small and susceptible to modifica- 

tion. This publication should prove useful in itself as a catalog of type speci- 

mens of Cayex, particularly to specialists on the family Cyperaceae. The larger 

purpose, however, in issuing a preliminary catalog of limited scope at this time 

is to demonstrate the concept of the Type Register in concrete terms and thereby 

to solicit the collaboration of all plant systematists in molding the Register into 

an effective, scholarly tool for future generations of the profession. 

The computer file presently registers over 13,500 type specimens, representing 

some 10,500 vascular plant taxa. Thus the Cawx Catalog, with its 1,000 speci- 

mens and 600 taxa, is a printout of less than 10 percent of the current, rapidly 

growing file. Less formal and less expensive means of putting out the informa- 

tion will be tried with future installments, and it may become desirable or neces- 

sary at some point to begin publishing in microform. Perhaps the most common 

and economical mode of disseminating the accumulated information will be to 

provide computer printouts to individual users in response to queries for up-to- 

the-minute reports on specific taxonomic groups. Once the data bank is well 

established query service can be provided to any user for a modest fee. 

We welcome your reaction to the concept of the Botanical Type Specimen 

Register on the basis of this sample. Only with the backing of the botanical 

community can we continue to get the necessary financial support to carry on 

the work. 

iii 

EDWARD S. AYENSU, Chairman 

Department of Botany 

30 September 1972

Con tents 

Foreword ............................. ........... .................... 

THE BOTANICAL TYPE SPECIMEN REGISTER 

Stanwyn G. Shetler 

Introduction ......................................................... ..... 

Acknowledgments .................... ........................ 

Concept and Purpose of Type Register . . . . . . , . . . . . . . . . . . . . . . . . . . . . .......... 

Scope of Register . . . . . . . , . . . . . . . . . . , . . . . . . . . ....I..... 

Procedures and Standards ,........ ................................................ 

Source of Data ,,,,.......,........................................ 

Instructions for Contributors . . . , . , . . . . . . . , . . . , , . . 

Record Format and Content ,..... , .,................ ,.......................... 

Level I-Taxon Data ....... . . . .. . ..................................... 

Family , ,......... ,...................................... 

Genus a . . . . . ,.. . ......................It............. 

Species .,......... .............................................. 

Infraspecific Taxon , , . , . . . , ............... ...... .. 

Author ....................................... ...................... 

Title .................................. ................... 

Level 2-Collection Data , . .......... . ................... 

Collector(s) .................................................................. 

Collection Number . , . . . . . . . . , , , . . . , . . . . , . . . . . . . . . . . . , , . . , . . . . , . . . . . . . . . . . . . . . 

Collection Date(s) ,......... , ... ............................................ 

Geographic Data Fields . . . . . . . . . . . . . . . . . . . . . . ................ 

Level 3Specimen Data , . . . , . . . . . . . . . . . , , , , . . . . , , . . . . . . . . . . . . . . , . . . . . . . . . . . , . 

Herbarium Abbreviation , . . . . 

Herbarium Sheet Number , , . , . . . . , . . . , . . . , . , , . . . . . , . . . . . . . . 

Kind of Type , , , , . , . . 

Data Source Code . . , . . . . . . . . . 

Processing System . . . , . . . , . . , , . . . , . . , , . . . . . , , , . 

....................... 

.......................... 

Paper Tape System .......................................................... 

On-Line System , . . . , , , . . , , , , , , , . , , , , , , . , , , , . , . , . . , , , . . , , , . 

Optical Scanning System , , , , , , , , , . . , . , . , , , , . , , . . . . . , , . . . . . . 

Magnetic Tape System , , , , , . , 

Computer Processing System , . , , . 

,,........ 

..................... 

Statistical Summary of Type Register Contents , . . . , , , . . , , , , , . , , . , . , , . . . . . . . . 

Use of Type Register , , , , . . . , . . . , . . . . . . . . . . , . , . . . . . ... . . . , . . . . . . . 

A CATALOG OF THE GENUS Carex (CYPERACEAE) 

Stanwyn G. Shetler (Editor): Mary Jane Petrini, Constance Graham Carley, M. J. Harvey, 

Larry E. Morse (Assistant Editors) ; Thomas E. Kopfler (Programmer) : and Collaborators 

Contributing Institutions , . . . . . . . . . . . . . . . . . . . . ... .................... 

Data-Collection Procedure .......................................................... 

Editorial Process ................................................................... 

Milestone Events ............................... 

Use of Carex Catalog . . . . 

Errata .................................................... ............... 

Statistical Summary of Catalog . . . . . . , . . . . . . . . . . . . . , . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 

Catalog of Specimens ............................................................... 

Author Index .................................................. 

Publication-Date Index . . . . . , . . . . . . . . , . . . . . . . 

Collector Index . , . . . . . , . . . . . . . I . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 

Geographic Index . , . , . . . 

Herbarium Index .... .......................... ............. 

Bibliography . , . . . , , . ................................................ . 

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An Introduction to the Botanical 

Type Specimen Register’ 

Stanwyn G. Shetler, with Mary Jane Petrini, Constance Graham Carley, 

M. J. Harvey, Larry E. Morse, Thomas E. Kopfler, and Collaborators 

The Botanical Type Specimen Register 

The Botanical Type Specimen Register is a 

computer-based system for recording information 

about type specimens of plant species and infraspe- 

cific taxa, which is designed to become a union reg- 

istry of type holdings in the world’s herbaria. It 

introduces a new dimension to the management of 

herbarium collections. Through the use of advanced 

information processing methods, critical data are 

compiled from the herbarium and library and 

merged into a common, machine-searchable file 

from which catalogs can be printed or special 

queries, defined by complex selection criteria, can 

be answered rapidly on demand. As a result, future 

taxonomic investigators will be able to learn the 

whereabouts of type specimens of particular con- 

cern to them and obtain answers to certain basic 

questions without painstaking research or time- 

consuming travel or correspondence. 

Stanwyn G. Shetler, Department of Botany, Smithsonian 

Institution, Washington, D.C. 20.560; same address: May 

Jane Petrini; Constance Graham Carley (nee Graham) 

(present address: 275 Collier Rd., Atlanta, Georgia 30309) ; 

M. J. Harvey (present address: Department of Biology, Dal- 

housie University, Halifax, Nova Scotia) ; Larry E. Morse 

(present address: Biological Laboratories, Harriard University, 

Cambridge, Massachusetts 02138) ; Thomas E. Kopfler. 

~~ 

‘This is No. 70 in the Flora North America Report 

series. 

STANWYN G. SHETLER 

Introduction 

1 

The Type Register was conceived by Mason E. 

Hale, who organized a pilot project in 1968 while 

he was chairman of the Smithsonian’s Department 

of Botany. This was one of several projects that 

Hale initiated as chairman to introduce electronic 

data processing (EDP) to collection management 

in the United States National Herbarium. (For a 

description of his automated system for recording 

specimen exchanges, see Hale and Creighton, 1970; 

the pilot Type Register project is described in 

Shetler et al., 1971.) At the outset, the author and 

Flora North America (FNA) personnel assisted in 

the development of the pilot system, and the proj- 

ect, though separate, has continued until the pres- 

ent to be associated closely with the FNA program 

(Shetler and Meadow, 1971). After launching the 

pilot effort, Hale, while continuing his involvement 

on an advisory level, passed the supervisory re- 

sponsibility to the author, who continues to direct 

the work. 

The project was initiated with special funds. In 

fiscal year 1969 and 1970, limited allocations of 

regular funds of the National Museum of Natural 

History were made through the Department of 

Botany, but regular budgeting did not begin until 

FY-71 (1 July 1970-30 June 1971) when the Mu- 

seum organized a new, Museum-wide program in 

data processing and incorporated the Type Regis-

ter as one of the charter projects. These projects 

are controlled by the respective departments, but 

the overall program is coordinated and managed 

by James F. Mello, Assistant Director. 

The response to the several requests for participa- 

tion during the pilot phase was excellent, showing 

general interest among botanists and convincing 

Hale and others at the Smithsonian of the potential 

value of a type-specimen register and of the bo- 

tanical community’s willingness to collaborate to 

the fullest extent possible in creating a register 

along the lines conceived. From the preliminary 

experiments much was learned about the logistical 

problems and the costs and manpower requirements 

of network data banking. The Smithsonian neces- 

sarily is concentrating present operations on its own 

type collections ot about 65,000 specimens. The 

system is designed, however, to accept data from 

any institution at any time, and as long as the 

inflow remains on a small scale, data from other 

institutions can be added to the central file by the 

paresent staff more or less as they are received. It 

is hoped that as interest in collaboration grows 

support commensurate to the interest will be forth- 

coming both for the central operation and for 

participating institutions. 

The second part of this paper represents the first 

published installment of the Type Register. It is 

a provisional union catalog of the type specimens 

of the genus Carex on deposit in ten major Ameri- 

can herbaria (see “Contributing Institutions”) and 

a concrete example of cooperative data banking. 

In FY-72, the year that ended on 30 June 1972, 

the Type Register project finally was put on a solid 

footing. Midway through this year it was possible 

for the first time in the four-year history of the 

project to staff adequately by employing three full- 

time persons (two assistant editors and a data con- 

version operator) and also to have adequate funds 

to process regularly. Prior to January 1970, the 

pilot project was carried on intermittently as funds 

were available by one to three part-time employees, 

and from January 1970 to December 1971 the proj- 

ect advanced on a more or less continuous, opera- 

tional basis with the assistance of one, two, and 

occasionally three full-time persons, the number 

depending again on available money. During the 

pilot effort, the computer analysis, programming, 

and file processing were done on a part-time basis, 

as needed, by personnel of the Smithsonian Infor- 

ShIITHSOSIAN COSTRIBUTIOSS TO BOTANY 

mation Systems Division, and since the project has 

become operational this work has been performed 

on a similar basis by FNA personnel so as to keep 

the Type Register system compatible with the FNA 

system. 

From the beginning Hale planned for the in- 

clusion of data from an indefinite and constantly 

growing number of other institutions and actively 

sought such collaboration. In one test of the feasi- 

bility of multi-institutional input, he distributed 

a computer-printed set of 52 cards, representing a 

card catalog of the National Herbarium’s complete 

type holdings in the genus Mimulus (Scrophu- 

lariaceae), to each of 50 large herbaria in the 

United States and abroad and solicited their co- 

operation in providing similar data from their own 

type collections, if any, of Mimulus. A second ma- 

jor test involved sending a computer-printed card 

catalog of the National Herbarium’s complete type 

holdings in the family Lamiaceae (Labiatae) to the 

University of California, Los Angeles, where Carl 

Epling’s extensive t) pe collections in this family are 

deposited, and later to the Missouri Botanical Gar- 

den and the New York Botanical Garden. All three 

institutions cooperated in providing data from 

their own collections of types in this family. The 

latter two institutions continued thereafter to col- 

laborate as much as possible on other taxonomic 

groups, and, apart from the Smithsonian itself, 

they have been the institutions with the greatest 

involvement in the Type Register project. 

Certainly, no claim to completeness can be made 

for a catalog that concerns a single genus and only 

one percent (10/1000) of the world’s public insti- 

tutional herbaria (Shetler, 1969). “Usefulness” is 

the pragmatic criterion for compilation and pub- 

lication of the Type Register, however, and useful- 

ness is dependent on critical mass, not absolute 

coverage. Clearly it is unrealistic to think that the 

Type Register could ever achieve absolute com- 

pleteness, registering all type specimens for all pub- 

lished taxa in all of the world’s herbaria, and the 

Register has not been conceived on this false prem- 

ise. The Register is being created on the assump- 

tion that some information is better than no 

information and that a catalog of ten type collec- 

tions is more useful than a catalog of one type 

collection. Although it must be admitted that the 

coverage is very uneven among the ten herbaria 

contributing to the Carex Catalog, for example,

NUMBER 12 

nevertheless this Catalog tells us more than we have 

ever known before about the Cal-ex type collections 

of the participating institutions and provides a 

solid framework to which new information can be 

added as it becomes available from these or any 

other institutions. 

The concept of a type register is not new. Al- 

ready in the mid-l930s, A. S. Hitchcock of the 

Smithsonian Institution, in his capacity as chair- 

man of the Committee on Nomenclature of the 

Botanical Society of America, coordinated the com- 

pilation of information on the location of type 

specimens. Lists of authors of new names, indicat- 

ing the major group(s) of plants they described 

(e.g., phanerogams) and the herbaria where they 

deposited their types, were compiled (Hitchcock 

et al., 1934, 1935). Other members of the Com- 

mittee in 1934 were L. R. Abrams, J. C. Arthur, 

A. TV. Evans, J. &I. Greenman, $1. A. Howe, E. D. 

Nerrill, F. TY. Pennell, and C. L. Shear. “The Com- 

mittee is not attempting to decide what specimens 

are types nor to determine the identity of types,” 

Hitchcock wrote in 1934; “it is attempting only to 

aid botanists in their search for types.” His words 

can only be reiterated in the present context. Other 

recent efforts to catalog types have been made in 

connection with specimen-data retrieval projects in 

the herbarium or museum (e.g., Beschel and Soper, 

1970; Collier, 1971; Crovello, 1972). 

Information processing technology has advanced 

far since the days of the first applications in biol- 

ogy, when the limitations of the computer led to 

some unfortunate consequences, as thoughtfully 

analyzed, for example, in reviews by Wood et al. 

(1963) and Rollins (1966) of some early applica- 

tions in plant taxonomy. We make no pretense of 

having avoided all the pitfalls cataloged by these 

reviewers, but we have tried to make good use of 

their advice. If we have learned anything so far, 

we have learned that no one can design the perfect 

system on the first trial. Every operational system 

is at the same time a pilot system for an even more 

advanced and refined, next-generation system. 

kKsOWLEDChfENTs.-Without the inspiration 

and genius of Rlason E. Hale the Type Register 

would not exist. 1Yhile chairman of the Smith- 

sonian’s Department of Botany (1967-69), he had 

not only the foresight to inaugurate this computer 

application but also the fortitude to persist with 

administrative support for it when others did not 

always share his vision nor his optimism for its 

potential value. The continuing support of Rich- 

ard S. Cowan, former Director, National Museum 

of Natural History, and of Hale’s successor as chair- 

man, Edward S. Ayensu, have been crucial in put- 

ing the project on a stable footing. The Assistant 

Director, James F. Mello, and his assistant, David 

Bridge, deserve much credit for laying the budget- 

ary groundwork that has brought the project to its 

present viable and relatively healthy state, and for 

facilitating its administration. From his vantage 

point as overseer of all EDP projects in the Na- 

tional Museum of Natural History, including the 

Type Register, RIello has provided stimulating and 

\vise counsel, as well as constant encouragement and 

help. 

Many curators have contributed in some way to 

the development of the Type Register thus far, 

especially in the course of the multi-institutional 

data-collecting experiment with MimuEus, con- 

ducted by Hale, and their cooperation is hereby 

gratefully acknowledged. Those curators and their 

assistants who participated directly in the compila- 

tion of data for the Carex Catalog are listed earlier 

as “Collaborators.” Among them, Hale, Irwin, 

Lewis, and Nicolson have taken a dedicated per- 

sonal interest in the success of the Type Register 

from the beginning and have in effect constituted 

a standing advisory editorial board. In their re- 

spective institutions, they have played a role in all 

of the data compilation that has been done for the 

Register thus far, regardless of the taxonomic 

group. As a Smithsonian colleague, Nicolson has 

been a steadfast supporter of the project, showing 

deep interest in the work itself and sharing his time 

and seemingly inexhaustible nomenclatural knowl- 

edge willingly and unselfishly whenever there has 

been need, which often has been daily. John H. 

Thomas raised enthusiasm for the Type Register 

to a new level wrhen, as a result of his collabora- 

tion on the Cal-ex Catalog, he began to ask special- 

ists borrowing from the collections he curates to 

compile data for the Register from type specimens 

loaned to them and to affix an annotation label of 

his own design to each, which reads, “The written 

itiformation on this specimen has been abstracted 

for the TYPE REGISTER PROJECT by - 

on

4 

\Vitli respect to the editorship of the Carex Cata- 

log, Carley, who served the FNA program and the 

Type Register project from December 1969 to April 

1971, and Petrini, who is the senior technical editor 

of the Type Register project, assisted Shetler with 

the clay-today technical editing. Harvey and Morse 

assisted him with the botanical editing during the 

)ear that each spent working at the Smithsonian on 

the FNA program-1969-50 and 197 1-72, respec- 

ti\elj. Harvey provided botanical supervision of 

the data-capture operation during the initial input 

of Cn? ex records from the Missouri Botanical Gar- 

den, New York Botanical Garden, and the U.S. 

National Herbarium, He directed the two-week, 

on-site input effort at the New York Botanical Gar- 

den in June 1970, Tvhich involved other genera 

besides Carex. Cynthia N. Ostroff of the Index 

Sominuin Generzcoi urn project assisted part-time 

at the Smithsonian in the technical editing of the 

Cai ex data from Harvard University. Marilyn An- 

draeson helped with the data compilation at the 

I\lissouri Botanical Garden, while similar assistance 

was rendered by Zella Ellshoff, Robert Helliwig, 

and Gail Johnson at the New York Botanical Gar- 

den. 

At least part of the Carex Catalog was examined 

in near final form by Frederick J. Hermann of the 

U.S. Forest Service Herbarium at Fort Collins, 

Colorado, and Tetsuo Koyama of the New York 

Botanical Garden, specialists on Carex and the 

family Cyperaceae, respectively. Although both 

provided helpful comments, they should not be 

held responsible in any way for the final Catalog, 

because neither was able to devote the enormous 

amount of time that would have been required to 

check the file authoritatively. This responsibility 

rests with the editors and collaborators. 

Several specialists at the New York Botanical 

Garden have provided data to the Type Register 

for groups other than Carex: Caroline Allen (Lau- 

raceae: a few records of selected taxa), Patricia 

Kern Holmgren (Brassicaceae: Draba, Thlaspi), 

Tetsuo Koyama (Cyperaceae: a few records of se- 

lected taxa), John T. Mickel (Schizaeaceae: Ane- 

mia subgenus Coptophyllum and segregate genera), 

and Ghillean T. Prance (Chrysobalanceae, Di- 

chapetalaceae). (See also “Statistical Summary of 

Type Register Contents.”) Holmgren has been the 

one chiefly responsible for coordination at the work- 

ing level of the New York Botanical Garden’s col- 

SMITHSOSIAS CONTRIBUTIONS TO BOTANY 

laboration in the Type Register project. Mildred 

E. klathias, Director, Botanical Gardens-Herba- 

I ium, supervised the compilation of data on types 

of the family Lamiaceae at the University of Cali- 

fornia, Los Angeles, early in the project. Bruce 

RIacBrJde assisted in the compilation of the data 

lor this family and for the genus MimuZus at the 

Missouri Botanical Garden. Other botanists who 

deserve mention for playing a part in the project 

at the Smithsonian are Amy Jean Gilmzrtin, Mon- 

terey Peninsula College, Monterey, California, and 

Milosla\ Kovanda, Czechoslovak Academy of Sci- 

ences, Prague, .izho spent the years 1969-70 and 

1970-5 1, respectively, with the FNA program. 

The FNX Editorial Committee (John H. Bea- 

man, M’alter H. Lewis, John McNeill, John T. 

;\lickel, Peter H. Raven, Stanwyn G. Shetler, Roy 

L. Taylor, John H. Thomas) has taken a deep 

interest in the Type Register project from the out- 

set and provided financial and material support 

through the FNA program, as well as invaluable 

advice and encouragement. A report on progress 

has been given at every meeting of the Committee 

since the Register was organized, and much time 

has been devoted to evaluation of the present and 

future development of the Register. 

The pilot processing system was designed by 

Reginald Creighton, Manager of Information Stor- 

age and Indexing, and programmed by Willard 

Handley, both of the Smithsonian’s Information 

Systems Division. Creighton worked closely with 

Hale to develop the pilot system, which served the 

original purposes well, and the Type Register 

would not exist if it had not been for Creighton’s 

pioneering insight and dedication. When the proj- 

ect became more closely associated with the FNA 

systems effort, Harriet R. Meadow, Systems Devel- 

opment Manager of ENA, designed the present op- 

erational sjstem in its general outlines. Kopfler 

has been responsible for the detailed design and 

FIGURE 

1.-Stages in the preparation of records for the Type 

Rcgister: a, Constance Carley checks nomenclatural data in 

the Gray Herbarium Card Index: b, Mary Jane Petrini 

enters corrections into the computer via a remote typewriter 

terminal connected by telephone (photos a and b by Walter 

G. Peter 111); c, type specimens of the genus Viola (Violaceae) 

are examined and discussed with respect to the problems of 

recording them in the Type Register by (left to right) M. J. 

Harvey, John T. Mickel, and Harriet R. Meadow in the 

herbarium at the New York Botanical Garden (photo by 

New York Botanical Garden).

NUMBER 12 5

programming and for the maintenance and query- 

ing of the machine files. Morse wrote the COBOL 

program for concatenating the data into the para- 

graph form used in the Carex Catalog. Meadow's 

analysis and design resolved some basic unforeseen 

problems that arose after using the pilot system for 

a time, and her efforts resulted in the operational 

system that continues to serve very well. She has 

played a crucial role in the success of the project 

and continues to provide advice and guidance on 

matters of systems development. Several botanists 

who also have experience with computer applica- 

tions have provided valuable advice trom time to 

time: Theodore J. Crovello, University of Notre 

Dame; David J. Rogers, University of Colorado; 

and James H. Soper, National Museum of Natural 

Sciences, Ottawa. 

Technical help in editing and capturing data 

has been given by the following persons, listed in 

the order in which they were hired, some as em- 

ployees of FNA and others as employees of the 

Type Register project: John Bolduc, Nancy How- 

ard, Barbara Bryant, Grace Rickard, Edna Mont- 

ford, Mary Beth Moore, Barbara Halter, Gudrun 

Christenson, and Rita Abessinio. Julia E. Taylor 

and Liliosa Mangosing Evangelista have been cheer- 

fully indispensable in discharging the myriad 

clerical and administrative tasks without which a 

project of this nature could never succeed. 

In addition to receiving regular budgetary sup- 

port on an increasing scale from the Smithsonian 

Institution, the Type Register project has been 

funded in part by the National Science Foundation 

through grants made to the American Institute of 

Biological Sciences (GB-844 1, GN-8 12, GB-26 173) 

and to the Smithsonian Institution (GB-31715 and 

contract C-720) for the FNA program, the Smith- 

sonian Research Foundation (grants Sg062 1054, 

Sg0621054/Cl and Sg0621054/C2), and the Smith- 

sonian Office of Systematics. 

Concept and Purpose of Type Register 

Perhaps 200 million specimens are on deposit in 

the more than one thousand public herbaria in the 

world, and scattered among these vast collections 

may be as many as 4 million type specimens (Shet- 

ler, 1969; Shetler et al., 1971). North American 

herbaria alone probably contain more than a half- 

million types. Likewise the original publications 

SMITHSONIAN COSTRIBUTIONS TO BOTANY 

describing new plant taxa and establishing their 

types are myriad and scattered through the world's 

literature. The taxonomist who wishes to make a 

scholarly study of a group of species, including an 

investigation of their typification, faces the for- 

midable task of locating the relevant original de- 

scriptions and type specimens. Fortunately for him, 

two standard indices, Index Kewensis (Rouleau, 

1970; Meikle, 1971) and the Gray Herbarium 

(Card) Index (Shaw, 1971), and various standard 

library catalogs and union listings are readily avail- 

able to guide him to the pertinent literature. No 

similar indices exist, however, to guide the taxono- 

mist to the pertinent type specimens. To find types 

he first must search the original literature case by 

case for indications or clues and then, through 

travel or correspondence, continue his search in 

herbaria among the specimens themselves. Even 

with the original descriptions in hand the specialist 

often faces great difficulties in trying to determine 

where the types are deposited. The modern litera- 

ture still shows an astonishing lack of standardiza- 

tion in the way types are designated and their 

herbarium deposition indicated. 

'IVhat the taxonomist needs, therefore, is a find- 

efs guide to the type holdings of at least the world's 

major herbaria. This guide should be indexed pri- 

marily by taxon but also cross-indexed several ways, 

and it should include citations of the original pub- 

lications and basic collection data as provided by 

the specimen label and/or published description. 

Such an index ultimately would incorporate and 

enhance the functions of Index Kewensis and the 

Gray Hcibar.ium Index. This type of registry could 

become effective at once for newly published taxa 

if taxonomists would agree to require registration 

of all new taxa and type depositions at the time of 

publication as a condition for effective publication. 

Furthermore, the registry, if computerized, would 

be the logical central repository for specialists' an- 

notations on typification, especially with respect to 

lectotypification and neotypification. Plant taxon- 

omy desperately needs an effective central place and 

straight-forward procedure for registering lecto- 

types, neotypes, and specialists' conclusions about 

other kinds of types. 

The Botanical Type Specimen Register, as con- 

ceived, therefore, is to serve primarily as a finder's 

guide for locating type specimens. The secondary 

function, however, is to serve as a guide to the

SUMBER 12 7 

original descriptions of the registered taxa and to 

provide collection data for the registered specimens. 

The specimen information necessarily is organized 

by taxonomic name. Thus the file is not purely a 

specimen register; the data are organized so as to 

constitute a three-level hierarchy: taxon, collection, 

specimen. A full entry (record) in the Register is, 

therefore, a synthesis of data from the herbarium 

and the library which cannot be completed with- 

out examining the original specimen and the origi- 

nal publication, as well as other specimens and 

publications as necessary. A record can be initi- 

ated with data from either the herbarium or the 

library and supplemented later by data from the 

other. To a degree this happens naturally as other 

institutions contribute to the initial record, but in 

any event years may elapse before all records of a 

given taxonomic group can be brought to relative 

completeness. 

The development of the Type Register ulti- 

mately involves three stages: registration, verifica- 

tion, and validation. 

The immediate objective is to initiate the com- 

puterized working file without getting bogged down 

in time-consuming researches to resolve challenges 

of the validity of the data. Such scholarly research 

is the province of specialists who may need years 

to resolve particularly difficult nomenclatural and 

typification questions, and the preparation of the 

kind of index envisaged here could never be ac- 

complished if all questions had to be answered first 

and the compiled data had to be “perfect.” In the 

first stage, therefore, the primary goal is to record 

or register the facts more or less at face value as 

they are given by the available primary and/or 

secondary sources. Research and editing are kept 

to a minimum except where obvious discrepancies 

can be resolved without extensive investigation. 

The editing is restricted largely to formatting the 

data according to the technical standards of the 

processing system, and to standardizing the use of 

names, titles, and terms in key fields. 

The second stage is to verify all data by firsthand 

examination of the pertinent specimens and origi- 

nal publications. When the data are compiled in 

the first place from the original sources expressly 

for the Type Register, verification is accomplished 

in the process of preparing the data for registra- 

tion, and the two steps merge into a single opera- 

tion. When a secondary source such as an existing 

card file or published index is used, however, there 

is a need to verify the information subsequently by 

checking the specimens and original descriptions. 

Two examples will illustrate. (1) Index Kewensis 

and the Gray Herbal ium Index constitute indis- 

pensable secondary sources of references to original 

publications, but the original publications them- 

selves must be examined in order to verify both 

the existence of the descriptions and the accuracy 

of the citations. (2) Present-day revisions and 

monographs customarily indicate where the key 

type specimens are deposited, but this information 

is not considered verified, for purposes of the Regis- 

ter, until the specimens have been seen in the 

process of compiling the data for the Register, be- 

cause the data must be verified in the context of 

the specific requirements of the Register. This is 

a critical point. 

It should be emphasized here that a taxonomic 

revision or monograph, no matter how carefully 

and authoritatively executed, constitutes a second- 

ary source of information for all taxa treated except 

those being described to science for the first time. 

In tact, as synthetic works, these treatises often 

present only the barest details on type specimens, 

especially tor previously described but even for 

newly described taxa, and the author’s own nomen- 

clatural interpretations frequently are not clearly 

distinguished from the original data. It is not un- 

usual, for example, to discover in the process of 

verifying a record in the Type Register that the 

author of a revision or monograph, in identifying 

what he believed to be the holotype, unwittingly 

designated a lectotype or even a neotype by strict 

application of the international rules of botanical 

ncmenclature. Therefore, except as a reference to 

the original sources, for which it is of course in- 

valuable, the monographic treatise has proved to be 

a disappointing starting point for compiling the 

Type Register; it seldom provides all necessary data 

and often presents summaries of the original facts 

which are telegraphic to the point of being impre- 

cise or even inaccurate. On the other hand, the 

monographic treatise is indispensable in the third 

or validation stage, because it deals in a systematic 

way with the typification of the taxa covered and 

establishes authoritative precedents that must be 

considered in the interpretation of the information 

in the Type Register. Furthermore, short of hav- 

ing all the original references and specimens in

hand, the monograph, which brings together all the 

data for a taxonomic group into one place, is by 

far the best single source of data for the given 

group. 

Verification is not a simple procedure that can 

be accomplished once for all time but an involved, 

virtually never-ending process, which seems to ex- 

pand in direct proportion to the number of speci- 

mens and publications examined. Comparison of 

the original description with data from one or more 

specimens rarely can be made without uncovering 

at least minor discrepancies that must be reconciled. 

As types from additional herbaria are registered it 

often becomes necessary to reexamine the original 

publication and secondary references again and 

again to resolve new discrepancies, and such dis- 

crepancies frequently multiply faster than they can 

be resolved as publication after publication is con- 

sulted. Gazetteers, atlases, biographies and bio- 

graphical dictionaries, personal fieldnotes and 

letters, and even new correspondence with current 

specialists, in addition to the obvious taxonomic 

treatises and reference works, may be employed 

eventually in the course of trying to verify the data 

of an entry in the Register. 

Apart from the facts themselves is the matter of 

interpretation and judgment. The compilation of 

any highly condensed, formatted, and standardized 

file of data such as the Type Register is bound to 

involve much interpretation of fact and judgment 

of what to include and what to exclude. The com- 

puter imposes the additional problem of judging 

how best to format and standardize the data for 

search and retrieval. As new data are provided 

or brought to light, there is a constant need to re- 

evaluate prior interpretations and judgments, and 

this in turn may require reexamination of previ- 

ously consulted literature and specimens. The prob- 

lem is one not only of accuracy and completeness 

rvithin a given record but also of consistency among 

records. How the geographic information is stand- 

ardized in the record for Taxon A, for example, has 

a direct bearing on how the. geographic informa- 

tion is standardized for Taxon B, and decisions 

made for the first case without knowledge of special 

problems to be faced in the second case may have 

to be reevaluated and changed when the two rec- 

ords are considered together. In short, there is no 

n prion' way to set standards for all time. 

S\lITHSOSIAN COETRIBUTIONS TO BOTANY 

The long-term goal of the third and ultimate 

stage in the development of the Type Register is 

to ualzdate the data according to the rules of the 

Zntu natzonal Code of Botanical Nomenclature 

(ZCBIY; Stafleu et al., 1972) and thereby to estab- 

lish the Register as a wholly reliable, authoritative 

index of types of plant species and infraspecific 

taxa. Validation involves typification, specifically 

the designation of kind or status of type, and such 

matters as rank, priority, synonymy and homonymy, 

authorship, and orthography. Up to a point, the 

records can be validated by any botanist or tech- 

nical person skilled in the strict application of the 

proLisions of the ZCBN, because many of the prob- 

lems are purely technical or legal. Indeed, experi- 

ence with the Type Register has proved that a 

trained technical editor frequently makes decisions 

more consistent with the ZCBN than the specialist, 

at least insofar as the objectives of the Register are 

concerned. Such technical validation, while it 

greatly increases the reliability of the data, never- 

theless is without the force of authority that can 

be gained only through the sanction of the taxo- 

nomic authorities themselves. As in all taxonomic 

research, many of the questions that arise regarding 

tjpification have no absolute answer but require 

good judgment by an experienced specialist on the 

basis of all available evidence, and no amount of 

technical expertise could suffice. This type of au- 

thoritative kalidation is needed in the long run if 

the Type Register is ever to take its place as an 

indispensable and thoroughly accepted tool of 

plant systematists, and it is hoped that the special- 

ists will cooperate in validating the information 

in the Register as it becomes available group by 

group in preliminary form. In the short run, how- 

ever, the most that can be achieved is some degree 

of technical validation. The important point to 

stress here is the dynamic state of the file which 

can be updated at any time to accord with current 

knowledge and understanding. 

Like verification, validation is a continuous proc- 

ess that never really ends, because the light of new 

information often requires important reevaluations 

and appropriate changing of the computer file. At 

the same time, a basic threshold can be achieved. 

A record is considered verified at least on an initial 

basis once the original description and all registered 

specimens have been seen in person by someone 

compiling and editing data expressly for the T

ALVBER 12 9 

Register. Likewise a record is considered validated 

at least initialiy once the designation of types has 

been worked out in accordance with the lCBN ex- 

pressly within the framework of Type Register 

specifications and format, These thresholds must be 

attained before the second and third stages of de- 

velopment can be said to have been achieved on a 

minimal basis, Authoritative validation as de- 

scribed above, on the other hand, will require the 

input of many specialists in the years to come and 

is a very long-term proposition. 

The three stages of development may be sum- 

marized as follows: 

Stage 1. Registration.-Creation of the initial 

file, ivhich involves basic standardization of cita- 

tion and geographical fields. 

Stage 2. Verification.-Editing file against pri- 

mary sources in the light of the accumulated data. 

Stage 3. Validation.--Shaping the Register as an 

authoritative tool on typification, fully in accord- 

ance with the ZCBN, by getting input from spe- 

cialists and by incorporating information on lec- 

totypification and neotypification, as well as other 

critical annotations. 

In practice, registration, verification, and valida- 

tion certainly are not sharply delimited phases and 

often merge into each other as a single process. 

Once the original description and specimens are in 

hand one attempts to accomplish as much of the 

entire three-stage process as possible. Verification 

and validation, in particular, tend to overlap; it is 

in fact impossible to accomplish the one without 

to a degree accomplishing the other. From the 

point of view of the daily operation, however, reg- 

istration, verification, and validation represent dis- 

tinct working stages in the creation of the computer 

data base, involving different procedures and per- 

sonnel. Editorially, each stage results in a more 

refined, reliable, and authoritative data base. In 

the first stage, the data can be compiled and regis- 

tered entirely from secondary sources, if necessary, 

although this is not recommended, but neither veri- 

fication nor validation can be accomplished without 

consulting the original sources. Regarding the op- 

erational distinction between verification and vali- 

dation, it should be realized that a technical person 

may be quite competent to verify the accuracy of 

the data but not to validate the type designations 

even with the original sources in hand. Ordinarily, 

technical editors are responsible for verification, 

and onl) when professional botanists or other specialists 

skilled in the application of the ZCBN perform 

this function is it possible to perform the 

k~lidation function at the same time. 

The present computer file of some 13,000 specimen 

entries, constituting the entire Type Register, 

15 a registry ot largely unverified and unvalidated 

records of apparent or piesiimptiue types, and for 

the next several years, at least, effort will continue 

to be concentrated on the rapid compilation and 

input of similar preliminary data from many other 

taxonomic groups and institutions, starting with 

the Smithsonian’s own type collection. The strategy 

is to register the greatest number of taxa and specimens 

in the shortest possible time so as to achieve 

quickly a critical mass of data for producing catalogs 

and answering queries. Clearly the usefulness 

of such a data base will be directly proportional 

to its taxonomic and institutional comprehensiveness. 

Unless efforts to verify and validate the data 

are kept to an essential minimum as new records 

are being processed, there is little chance that a 

comprehensive data base can be created in the 

foreseeable future. The manpower and resources 

simply are not available at present for the massive 

searches in the herbarium and library that would 

be required to bring every new record to the Stage 

2 or Stage 3 level of refinement as it is being entered 

into the file. To a large extent, therefore, 

the Stage 1 Type Register will have to be verified 

and validated through use, through feedback from 

the specialists Lvho discover its shortcomings in the 

course of their research. 

Under no circumstances is the Type Register 

being used or is it intended to be used as a place to 

designate lectotypes and neotypes and thereby to 

set nomenclatural precedents. If the Register is ever 

to be used in this manner, which as indicated earlier 

may prove desirable eventually, the taxonomic 

fraternity will have to make a conscious decision to 

do so. 

When the scope of the task is considered, it is 

not surprising that no one has attempted to compile 

a union catalog of type specimens before now. The 

task can be cut down to size, however, because relatively 

few of the world’s public herbaria are large 

enough to have a significant concentration of type 

specimens. Only about a score of the world’s herbaria, 

for example, contain over two million specimens 

each, and a published index, including

10 

literature citations, to any one of these collections 

would be enormously useful in itself. Each new 

institution to be added to such a base would en- 

hance the catalog greatly and move it one step 

closer to the goal of a worldwide union registry. 

The U. S. National Herbarium is one of the 

score of major herbaria with more than two million 

specimens, and its type collection of 65,000 or more 

specimens certainly constitutes a significant initial 

data base. Furthermore, not only is this type col- 

lection separate from the main herbarium and 

easily accessible, but it also has an associated file 

of cards on which are recorded pertinent data from 

the original publication (see “Source of Data”). 

Without this large, ready-made card file and with- 

out computer technology, which permits the crea- 

tion of a union register on a much more flexible 

and dynamic basis than would otherwise be pos- 

sible, the Type Register doubtlessly would never 

have been conceived or started. The Botanical 

Type Specimen Register is in the first instance, 

therefore, an index (catalog, register) of the U. S. 

National Herbarium’s own type collections. The 

thousands of man-hours that have gone into the 

creation and maintenance of the National Her- 

barium’s type collection and card file have paid off, 

of course, to the many who through the years have 

used the type herbarium on the Smithsonian prem- 

ises. By computerizing this information the Smith- 

sonian’s Department of Botany now makes it 

possible for taxonomists at large to benefit from 

thl:, accumulated data and enormous manpower 

in iestment. 

Scope of Register 

The Register is designed to handle taxa typified 

by specimens, namely, taxa of the rank of species 

or below, and it encompasses all infraspecific taxo- 

nomic levels recognized by the ZCBN. In the fu- 

ture, modifications in design may be desirable if 

not essential to accommodate cases in which the 

type is not a specimen but a description or a figure. 

For the present, however, the object is to register 

specimens, and for this reason data collection usu- 

ally begins with the specimens and proceeds to the 

literature rather than the other way around. There 

are good reasons for arguing on the one hand that 

registration should proceed from the specimen to 

the taxon and on the other hand that the process 

S!vlITHSONIAK CONTRIBUTIONS TO BOTANY 

should be reversed, proceeding from the taxon to 

the specimen. No doubt this publication will stim- 

ulate debate on these alternatives; meanwhile, it 

should be made clear that primarily the first ap- 

proach is being taken. 

\Vitlz one exception, only the original names of 

newly described taxa, i.e., taxa being described to 

science for the first time, are included. The one 

exception is a wholly new name for a previously 

described taxon necessitated because all other possi- 

ble names and combinations would violate the 

international rules. New combinations involving 

previously published epithets are excluded rigor- 

ously insofar as they are Enown to be combina- 

tions; in such cases, only the basionym is entered 

into the Register. In one sense, therefore, the Type 

Register is a basionym file. This approach has been 

taken because it is the only feasible way in the 

foreseeable future to create a stable file with fixed 

points of reference. Eventually, viewed in the long- 

est terms, it will be necessary to link the Type 

Register to a much vaster name list that shows all 

possible synonymy connections among basionyms 

and combinations and thus makes it possible to 

trace the nomenclatural history of a particular spe- 

cies, for example, from modern usage back to origi- 

nal usage. This is far too much to expect of the 

Type Register in itself, however, and for this rea- 

son the design of the Register allows for no syn- 

onymy except for orthographic variants. If a 

taxon originally was published under a generic or 

specific name with a spelling that later was cor- 

rected, then the original spelling is indicated in a 

special field, while the accepted spelling is shown 

in the main taxon field; for example, many species 

have been published in the genus Penstemon under 

the spelling “Pentstemon,” and this spelling is in- 

dicated in the orthographic synonym field, as neces- 

sary. IVithout this approach, the same genus would 

alphabetize in different parts of the file (e.g., 

Aplopnppus vs. Haplopappus). 

Only validly published names are included, but 

the names need not be legitimate, as defined by the 

ZCBN (see also hfcVaugh et al., 1968). 

Taxonomically and geographically, the Register 

is limited only by the availability of data and op- 

erational resources. The present machine file in- 

cludes only vascular plants and primarily flowering 

plants, but it could be expanded at any time to 

include cryptogamic groups if the data and the

NUMBER 12 

resources to input the data, especially personnel, 

were available. The geographic scope already is 

worldwide because the initial source of most rec- 

ords, the U.S. National Herbarium’s type collec- 

tion, is worldwide in scope although particularly 

strong in New World areas. The input is further 

biased geographically at present by the fact that 

the other herbaria which have cooperated thus far 

on the Register also are North American institu- 

tions with principally New IVorld collections (ex- 

cept Arnold Arboretum). 

Inclusion of type photographs has been sug- 

gested several times, but so far this has not been 

done because they present special problems requir- 

ing careful study before the system can be modified 

to accommodate them. Whereas type specimens are 

unique and, even in the case of isotypes and syn- 

types, are distributed to a relatively limited set of 

herbaria among the total, type photographs are not 

unique, and in theory every herbarium can have a 

photograph of any type. The Register soon could 

be overloaded with references to photographs, and 

no purpose would be served. While there is a clear 

need, especially on the part of floristic workers who 

may be able to satisfy their requirements with pho- 

tographs and thus avoid a massive borrowing of 

type specimens, for a central index of negatives on 

deposit at major centers from which type photo- 

graphs could then be purchased, this problem calls 

for separate attention. 

To an extent, the same reasoning applies to type 

fragments because many institutions potentially can 

have fragments of the same specimen. A type frag 

ment has no standing in the ZCBN unless it can 

be interpreted as a formal type of some kind (e.g., 

isotype), and most fragments cannot be dignified 

by such interpretation. In the modern era when 

travel and communication are easy, making the re- 

motest corners of the earth accessible, the informa- 

tional value of the type fragment in one’s own 

herbarium has diminished greatly because the type 

specimen itself can be borrowed or examined by 

personal visit. Thus only in the case of types that 

have been destroyed or of types that for political 

or other reasons are still inaccessible can importance 

be attached to a register of information on the 

whereabouts of type fragments. In other words, the 

taxonomist wants to know, “Where can I find 

a type specimen?” not “Where can I find a type 

fragment?” The latter question will interest him 

only if all efforts to see a type specimen fail or 

prove impractical. For these reasons, type frag- 

ments have been registered sparingly in the present 

file and only when the circumstances seem to war- 

rant doing so. 

Lectotypes and neotypes present a special prob- 

lem that cannot be handled properly with the cur- 

rent system design. The system allows for only a 

single bibliographic citation, namely, the citation 

of the original publication where the taxon was first 

described and the name proposed. In cases of lec- 

totypification or neotypification, however, it is 

necessary to cite also the second, later reference 

where the lectotype or neotype was designated. The 

problem has been largely ignored in this initial 

phase because lectotypification and neotypification 

cannot be documented properly without the direct 

participation of specialists. Identification of lecto- 

types and neotypes is part of the Stage 3 validation 

process described earlier, and by the time this level 

of documentation is possible the system will be 

modified to include a separate file, linked to the 

basic file, for recording lectotypes, neotypes, and 

other pertinent taxonomic or nomenclatural anno- 

tations. Such a file for “remarks” will provide a 

way of recording the names of authorities who have 

validated the data. 

A word is necessary about the relationship of the 

Type Register project to the Index Nominum 

Genericoyum (ZA‘G) project (Cowan, 1970). The 

object of ZArG is authoritative typification of all 

generic plant names. Thus it deals with genera, 

not species, except for type species, and it is not 

concerned with type specimens or collection data 

of any kind. Emphasis is placed on achieving at 

once, before input, the level of validation that the 

Type Register is expected to achieve only in the 

long run. The Register, which is not concerned 

with the typification of genera, and ING are com- 

plementary, therefore, and do not duplicate each 

other in any way (see also p. 16). 

IVith respect to Flora North America, the closest 

links are maintained between it and the Type Reg 

ister project on the one hand and ZhrG on the 

other hand, to ensure that the work of each project 

will complement rather than duplicate the others.

12 SMITHSONIAN CONTRIBUTIONS TO BOTANY 

Procedures and Standards 

SOURCE OF DATA 

The principal source of data at this stage is the 

permanent card file associated with the type col- 

lection of the U.S. National Herbarium (US). The 

card records are converted into machine-readable 

form genus by genus in alphabetical order. During 

the pilot phase cards were pulled from the file by 

family (e.g., Scrophulariaceae), but this approach 

is impractical for the file as a whole, which is ar- 

ranged alphabetically by genus. While that ap- 

proach was being taken, the cards were being 

compared with the specimens in the type collec- 

tion, which are arranged systematically (modified 

Englerian sequence), prior to input. Now that an 

alphabetical rather than systematic approach to the 

file is being taken the specimen-comparison step 

is being postponed until the whole file is in the 

computer and can be sorted systematically by 

family. 

Curators of the U.S. National Herbarium have 

followed the practice of segregating type specimens 

from the general collection since the early part of 

the present century. The practice was first estab- 

lished about 1918 by then-curator Paul C. Standley. 

At the same time an associated card file was started 

to supplement the specimen data with information 

from the literature. The file includes a card for 

every taxon (species, subspecies, variety, form) rep- 

resented in the type collection, and generally the 

responsible curator has had the original publica- 

tion in hand while preparing the card and the 

standard folder for filing the type specimen (s). 

Each card includes the original taxonomic name 

(basionym in cases of later transfer), author, origi- 

nal reference, basic collecting data, and designation 

of kind of type. To re-create this file today from 

the specimens and the literature tvould require at 

least 10 and more likely 20-30 professional man- 

years, and there is no reason to suppose that the 

file could be re-created with any higher professional 

standards or greater degree of accuracy on the av- 

erage than the first time. In short, it is scientifically 

sound as well as eminently practical to create the 

preliminary edition of the Type Register from the 

Smithsonian file as it stands. 

Of the 65,000 specimens in the US type collec- 

tion, about 55,000 are types of phanerogamic species 

and infraspecific taxa, and the other approximately 

10,000 are types of cryptogamic taxa-ferns, mosses, 

and lichens. 

Other institutions can contribute to the Type 

Register in any of a number of ways, as explained 

in the next section. Basically, there are two ways: 

(1) annotation of a printou, listing records already 

registered in the machine file, and (2) submission 

of completed data forms or some equivalent pro- 

cedure for new records not presently registered in 

the machine file. These are complementary pro- 

cedures which must both be used. Institutions with 

ready-made card files like the Smithsonian’s are in 

the best position to contribute in a significant way 

quickly, and their contributions will spare the 

smaller herbaria from repeating costly bibliographic 

research that already has been done somewhere 

else. The cumulative Register provides a basis for 

checking rapidly for isotypes and other “duplicate” 

type material, leaving bibliographic research to be 

performed only for those cases where new taxa are 

to be added to the Register. In other words, to 

conserve effort maximum advantage should be 

taken of the existing file in the process of adding 

new data, especially bibliographic data, and of 

course the larger the machine file becomes the 

greater can be the economy of scholarship on the 

part of newly collaborating herbaria. 

AZonographs and the personal manuscripts or 

files of monographers are obvious sources of au- 

thoritative data for the Register and have been 

used in a few instances, although there are some 

distinct disadvantages in using the monograph as 

the starting point (see p. 7). Future monographers 

should register data routinely for type specimens of 

new taxa prior to, or simultaneously with, publica- 

tion. Likewise, it is hoped that graduate students 

in plant taxonomy will be advised to submit data 

on type specimens examined by them in the course 

of their research. 

For every specimen registered in the file, a code 

is appended at the end of the record which indi- 

cates the source of the data according to a broad 

classification of source categories, summarized later 

under “Data Source Code.” 

INSTRUCTIONS FOR CONTRIBUTORS 

Any herbarium interested in contributing to the 

Register is advised to consult with the staff at the

NUMBER 12 

FIGURE P.-Standard Data Collection Form of Botanical Type Specimen Register. 

13


Smithsonian well in advance of initiating a project 

so that a compatible way of compiling data can be 

devised before the first record is collected. At pres- 

ent the Type Register staff has its hands full with 

the internal Smithsonian file but will do its best 

to cooperate with other potential contributors to 

set up procedures for submitting data. The most 

efficient way of collaborating under present cir- 

cumstances is to submit the records on the standard 

data form (Figure 2) to the Smithsonian for proc- 

essing, but with adequate planning it will be possi- 

ble for another institution to convert its own data 

into machine-readable form in-house before submit- 

ting the data to the Smithsonian and thereby to save 

time and effort in the overall process, at least where 

large herbaria are concerned. Complete records, 

i.e., with all essential data present, submitted by 

other institutions are placed in the queue and proc- 

essed in due course as time permits. Incomplete 

records, particularly those without proper biblio- 

graphic citations, are set aside in an inactive file 

where, for lack of staff to complete them, they may 

remain indefinitely. Potential contributors should 

keep in mind, therefore, that any effort that falls 

short of providing complete records risks becoming 

a wasted effort. 

A collaborating institution should always work 

from the current catalog in the Type Register, if 

one exists, of the genus or other taxonomic group 

concerned. The Carex: Catalog, for example, pro- 

vides a checklist of taxa already registered which 

can be used as a tool by any herbarium wishing to 

search its own collections for type specimens of this 

genus. Working catalogs for other registered groups 

can be provided at cost by the Smithsonian as they 

are needed. The purpose of this procedure is to 

reduce unnecessary effort on the part of both the 

compiler and the editor. In the system, all speci- 

mens pertaining to a given taxon are registered un- 

der a single entry, and thus there is one unit record 

per taxon. It is the editor’s responsibility to pre- 

vent duplications of the same taxa in the first place 

and in the second place to discover and delete the 

occasional duplication that inevitably creeps into a 

file of the size and complexity of the Type Register. 

Individual compilers can do much to assist the 

editor in preventing duplications, however, by 

keeping themselves informed, through working 

catalogs, of the current status of the file, and by 

using procedures that minimize the chances of sub- 

mitting duplicate information for taxa already re- 

corded. 

The procedural details will differ from institu- 

tion to institution, but these general guidelines 

should be followed, unless other special arrange- 

ments have been made beforehand: 

(1) All new records-taxa new to the file-should 

be submitted on the standard data collection form 

(Figure 1) and according to the technical data spec- 

ifications used by the Smithsonian. Be sure that 

the original name and not a later combination is 

being used. 

(2) All additions to existing records-supple- 

mentary publication or collection data, and collec- 

tions or specimens new to the file-should be 

submitted as annotations to the appropriate records 

in a copy of the working catalog; or, if they are 

submitted on standard data forms or in a separate 

typewritten listing, each addition should be prop- 

erly referenced to the record in the file to which 

it belongs. Care should be taken to distinguish 

between specimens that belong to a collection al- 

ready registered in the file and specimens that 

introduce a new collection but to an already regis- 

tered taxon. Both constitute additions but on dif- 

ferent levels. The latter situation arises frequently 

when syntypes are involved, but slight discrepan- 

cies in collection data can easily be overlooked by 

the compiler, leading him to the conclusion that 

the former situation obtains. If the collection data 

recorded in the Register are not identical or at least 

reconcilable with the collection data of the speci- 

men in hand, then the discrepancy should be re- 

solved appropriately or the specimen excluded. In 

annotating the catalog, it is important to be sure 

that the additions are clearly associated with the 

proper collection where two or more collections 

are registered. 

(3) All proposed changes to existing information 

in the file should be submitted with documentation 

as annotations to the appropriate records in a copy 

of the working catalog; or, if they are submitted 

on standard data forms or in a separate typewritten 

listing, each addition should be properly referenced 

to the record in the file to which it pertains. With- 

out documentation, proposed changes raise more 

questions than they answer and complicate the 

work of the editor. 

The most common errors, experience has shown, 

are mistaking (1) a later combination for an ongi-

SUhIBER 12 15 

nal name, (2) the type specimen of a variety or 

form for the type of the species itself, and (3) a 

syntype (or isosyntype) for an isotype. The second 

of these mistakes often turns out to be the ex- 

planation for the situation in which the specimen 

appears to have been collected after the species was 

described, i.e., the collection date is later than the 

publication date. An undetected name transfer 

may lead to puzzling discrepancies or to unnoticed 

duplication. Frequently, for example, a type co:. 

lection passing under an undetected later combina- 

tion becomes the basis for introducing a new taxon 

to the file, while at the same time the collection 

already is properly registered under its basionym. 

A final note should be added about the use of the 

Carex Catalog or any similar working catalog in 

herbaria where type specimens have never been 

identified and segregated into separate folders or a 

separate collection. By means of the collector in- 

dex, it is possible for curators who wish to begin 

segregating types to use such a catalog as a means 

of identifying type specimens within their herbaria 

which belong to collections recorded in the Type 

Register. 

RECORD FORMAT AND CONTENT 

The data content of a unit record in the Regis- 

ter was established largely according to the conven- 

tions long used for the US card file. With the US 

card format as a standard, the data form shown 

in Figure 2 was devised for use by contributors at 

other institutions. This form shows what fields of 

data should be included, distinguishing between 

essential and nonessential fields and indicating cer- 

tain of the basic standards. The fields tagged as 

“essential” constitute the minimum number of data 

elements which the processing system is designed 

either to require or expect. From the botanical- 

content point of view, however, none of the fields 

should be regarded as optional. Every effort should 

be made by the contributor to provide data for all 

fields. Contributors can obtain blank forms at cost 

from the Smithsonian or use facsimiles. 

Before input to the system, all records are edited 

to conform with the field-by-field technical specifica- 

tions that have been established to standardize con- 

tent and format in the system. A copy of these 

specifications can be made available to collaborators 

on request, but most contributors will not want to 

be bothered with all of the technical details of the 

system. For purposes of contributing records, it is 

essential to know only the main conventions and 

standards that govern the content and format of 

the data fields. These basic standards are explained 

below by field, and contributors are strongly urged 

to follow them closely so that the work of the 

editor will be simplified. The rules of standardiza- 

tion have been applied more rigorously and con- 

sistentl) to the Carex Catalog than to any other 

part of the Register, and potential contributors are 

asked to study this Catalog carefully for specific 

examples of how standards have been applied to 

govern content, form, and style. Implicit in this 

Catalog are the answers to many specific questions 

about standardization which cannot be answered 

here. It should be kept in mind, however, that the 

typical edit format is different from the format of 

this published Catalog. In the typical format, the 

information is not strung together (concatenated) 

in paragraph form, but each field is labeled and 

printed by itself with room for annotations. Fur- 

thermore, certain fields of data (e.g., source code) 

have not been printed out in the Carex Catalog. 

The reference works used most frequently in 

the course of compiling and editing data for the 

Type Register are cited in the bibliography. Some 

of these works have been adopted for editorial pur- 

poses as the standard references and authorities for 

verifying and standardizing new data during initial 

input. An “authority” (authority file) is an index, 

thesaurus, or dictionary of terms, names, or titles 

tvhich is used to standardize some category of data, 

eg., B-P-H (Lawrence et al., 1968) for titles of 

botanical periodicals. Published standards have 

been adopted as authorities whenever possible, but 

in some instances it has been necessary to begin 

creating authority files expressly for use in the Type 

Register project. A “standard reference” is an 

authoritative and reliable secondary source that 

provides the editor with a practical means of 

quickly verifying some category or categories of 

incoming data, e.g., a name index such as Index 

Kerciensiy or LVillis’ Dictionary of Flowering Plants 

nnd Fems (7th edition, revised by Airy Shaw, 1966). 

Verification in this editorial sense of double- 

( herking in secondary sources is not to be confused 

Tvith verification in the primary scholarly sense dis- 

cussed earlier in connection with Stage 2 develop- 

ment of the Type Register (p. 7).

16 SXITHSOSIAN CONTRIBUTIONS TO BOTANY 

The Internotional Code of Botanical Nomencla- 

tu)e is, of course, the final authority on all matters 

of typification and nomenclature. (The 1966 edi- 

tion, prepared under the chief editorship of Lan- 

jouw and Stafleu, has been used thus far, but the 

rietver 1972 edition of Stafleu et al. is now availa- 

ble.) Of the essential desk-top references, the one 

most frequently consulted, perhaps, is the seventh 

edition of JVillis’ Dictionary. Also invaluable as 

general reference works are Stearn’s Botanical Latin 

(1966) and Stafleu’s Taxonomic Literature (1967). 

Though hardly desk-top references, Index Kewensis 

and the G?ay He) bai ium Index are indispensable, 

and the Type Register editorial staff is fortunate 

in haking available to it an integrated version of 

the lormer and both the card and book forms of 

the latter. Insofar as possible, the same procedures 

and standards are being applied in both the Type 

Register project ant1 the Flora North America pro- 

gram, so that the data bases will be compatible. 

.luthority files developed especially for the one 

project are being used also for the other as ap- 

propriate. 

The main editorial procedures that have been 

adopted to verify and standardize the ingoing data 

are summarized below field by field. Collaborators 

can do much to increase the reliability and stand- 

ardization of their own data by using these same 

procedures in the process of compilation. The 

editorial burden is eased greatly when the editor 

knows in advance that the essential standards have 

been upheld consistently by the contributor, 

Level 1-Taxon Data 

The following fields of data are recorded only 

once each for every species or infraspecific taxon 

registered in the machine file because these data 

are unique for each taxon. Furthermore, the taxon 

is the unit record, and every taxon is entered into 

the file only once. 

FA%fItu.-The latest edition of Engler’s Syllabus 

der Pflanzenfonizlien (vol. 1 : Melchior and Werder- 

mann, 1954; vol. 2: Melchior, 1964) is the authority 

for the system of families with the exception that 

the accepted family name with a regular ending 

is used in all cases, including the eight cases where 

the Syllabws, following the traditional practice sanc- 

tioned by the ICBA‘ (see list of h’omina Familia- 

rum Coriservandn), uses irregular names. These 

irregular names with their adopted regular equivalents 

are: Compositae/ASTERACEAE, Cruciferae/ 

BRASSICACEAE, Graminae/POACEAE, Guttiferae/CLUSIACEXE, 

Labiatae/LAMIACEAE, Leguminosae/FABACEAE, 

Palmae/ARECACEAE, Umbelliferae/APIACEAE. 

The use of regular family 

names conforms with practice in the FNA program. 

IVillis’ Dictionary is used in conjunction with the 

Syi;obus to determine the family to which a genus 

belongs. 

GEXS AND GENUS SYsoNuhr.-Index Nominum 

Gcne,’icorum, insofar as it is completed, is the final 

authority for generic names, to determine their accepted 

spelling and whether they are validly published. 

IVillis’ Dictionary, which in any case is an 

indispensable authoritative handbook on these 

matters, is consulted for genera not yet covered by 

I*YG. Il’henever the data are being compiled directiy 

from the primary sources, the generic name 

is entered on the data form exactly as it was spelled 

in the original description of the particular species 

or other taxon in queution. If for some reason this 

spelling is a variant of the currently accepted orthography 

of the name, then both spellings are entered 

into the machine file, the accepted spelling 

in the “Genus” field and the orthographic variant 

in the “Genus Synonym” field. In no case is more 

than one spelling permitted in the “Genus” field 

for the species and infraspecific taxa of any particular 

genus of plants. 

SPECIES.-Attempt is made to record the specific 

epithet exactly as it was spelled originally, except 

where a minor change is required by the provisions 

of the ICBhT governing orthography. (A two-word 

epithet, for example, is hyphenated to form a single 

word.) To verify the spelling given on the data 

form, the editor relies upon the Gray Heybarium 

Index and/or Index Kewensis insofar as possible, 

because to check the primary publications in all 

cases is impractical at this stage. These two indices, 

which overlap considerably in coverage, frequently 

provide a check on each other. Unless the taxon in 

question is by definition outside the limits of one 

of the two indices, the second index is checked 

routinely whenever the first does not confirm the 

spelling given on the data form. TVith respect to 

species regarded as being of hybrid origin, the 

standard practice of placing an “X” followed by a 

blank space before the epithet is followed.

SUMBER 12 17 

INFKASPECIFIC TAxos.--An infraspecific taxon of 

any rank recognized by the ICBN can be accommodated 

in the Type Register by entering the infraspecific 

epithet and the appropriate rank 

designator in this field, e.g., VAR GRACILIS or 

SFXI CRASSA. Infraspecific names are entered in 

the Register in the form of trinomial combinations 

Ivith the appropriate rank designator, because this 

form is adequate for nomenclatural purposes; and 

quatlrinomials create problems in the system. The 

Gray Herbarium Index is used to verify infraspecific 

epithets as to rank and spelling, but it covers 

onl) infraspecific taxa of the New World published 

during the past 100 years. For this reason, many 

of the infraspecific names must go unverified at this 

stage when extensive literature research is impractical. 

Following is a list of the infraspecific ranks 

and their standard abbreviations used in the Register: 

subspecies SSP 

variety VAR 

subvariety SVR 

form FOR 

subform SFM 

nothomorph NM. 

.4L1THoK.-This field carries the full last name 

and all initials of each author of the binomial or 

trinomial name under which type specimens are 

being registered. Whether single or multiple au- 

thors, the last name is always placed before the 

initials. When the use of initials only is certain to 

lead to confusion, the full first name also is in- 

cluded, e.g., MACOUN, JOHN vs. MACOUN, 

JAAIES hi., not hIACOUN, J. vs. hIACOUh’, J. Xi. 

The publishing author or the author of the work, 

if different from the author of the name, is always 

included in this field and separated from the author 

of the name by “IN” or “EX” in accordance with 

the rules and recommendations of the ZCBN. No 

authority exists for author names, but an author 

authority file has been started for the Type Register 

on the basis of the Carex Catalog. Further, the 

FNA Author File is well underway, and it is 

planned that the two be compatible and that ulti- 

mately they be merged. Meanwhile, the standard 

references are Barnhart’s Biographical Notes Upon 

Botanists (1965) and Stafleu’s Taxonomic Litera- 

ture (196’i), but all available biographical refer- 

ences are consulted as necessary. (See also under 

“Collector[s].” Many specific problems are en- 

countered in dealing with author names, but the 

details are beyond the scope of this introduction. 

Examples of specific solutions can be found in the 

Carex Catalog. 

TiTLE.-The title of the periodical, monograph, 

or book in which the name of the taxon in question 

ivas first validly published is recorded in this field. 

The title of the article in the periodical or of the 

chapter in the monograph or book is never included 

or given in lieu of the title of the periodical or 

work. In cases where a name was introduced into 

the literature before it was validly published, refer- 

ence is made only to the place of valid publication. 

A name published first without description (nomen 

nzidzim), for example, often is later published valid- 

ly with description. Nomina nuda are not included 

in the Type Register. All titles are abbreviated con- 

sistently. B-P-H is the authority for abbreviating 

titles of periodicals. For titles of monographs and 

books, the Type Register project is developing its 

own authority file, using the principles of abbrevia- 

tion set forth in B-P-H, and, insofar as possible, 

taking advantage of the title abbreviation file de- 

veloped by the ZNG project. An author’s or editor’s 

iiame is not included in this field unless it actually 

is part of the title; otherwise, it is included in the 

previous field as the publishing author or editor. 

Level 2-Collection Data 

Because there may be multiple type collections 

(e.g., syntypes) for any taxon, the collection data 

fields may repeat as a set any number of times. The 

folloiving set of fields is recorded for each collection, 

insofar as the data exist and are available. 

COLI.EcToR(S).-The names of all collectors of the 

type collection being registered are recorded in this 

field exactly as author names are formatted in the 

“Author (s)” field. The name of a collecting expedi- 

tion may be recorded here when individual collec- 

tors cannot be determined. The Type Register 

project is developing its own authority file for col- 

lectors, but meanwhile the following biographical 

indices, in addition to those already mentioned 

under the “Author (s)” field, are being used as 

standard references: “Zndex Herbariorum, Part 11: 

Collectors” (Lanjouw and Stafleu, 1954, 1957; 

Chaudhri et al., 1972-completed for letters A to 

L); “Index to Principal Collections Represented in

the U. S. National Herbarium” (compiled by U. S. 

National Herbarium staff for internal use, 1965). 

COLLECTION NuMBER.-ordinarily this will be the 

collector’s own number, but when there is no trustworthy 

means of determining his number, or if he 

had none, a serial collecting number assigned by an 

institution or expedition may be recorded instead. 

The field is regarded pragmatically as the place for 

a number, any number, that has been associated 

with the collection and which, when combined with 

the name (s) entered in the “ (Collector (s)” field, 

normally will form a unique reference to the collection. 

If a distinction can be made between the 

number of the collector and the number of his 

expedition or institution, then the name of the 

expedition or institution assigning the serial number 

should be placed in the “Collector (s)” field 

unless one or more collectors’ names already have 

been entered there; otherwise, this name should be 

prefixed to the collection number to make clear 

that the number is not the collector’s own. When 

there is absolute evidence that the collection never 

has been numbered in any series, then the abbreviation 

S.N. (sine numei-o), meaning “without 

number,” should be entered in this field. If on the 

other hand the number is merely unknown or there 

is doubt about the existence of a number, then 

dashes (- - -) should be entered in the field. 

COLLECTION DATE(S).-collection date is recorded 

just as accurately as it is known, and if necessary 

two dates or a range of dates are given. All dates, 

whether single or in ranges, take the form: 28 Sep 

1928. On the data form, dashes should be entered 

to indicate that the collection date is unknown, and 

the abbreviation S.D. (sine dato) should be used to 

indicate that the collection is known to be without 

a collection date. In using the latter designation, 

the compiler should be absolutely certain that the 

collection is undatable; otherwise, he should use 

da4ies (- - -). 

GEOGRAPHIC DATA FIELDS.-Four geographic fields 

are used to pinpoint hierarchically the collecting 

locality: (1) country; (2) state, province, department, 

or equivalent; (3) county or equivalent; (4) 

locality. The specific place is recorded in the fourth 

or lowest field more or less in the terms of local 

reference given by the collector himself and should 

include a town, post office, or other place name 

that can be found in an atlas. The locality terms 

in the fourth field are ordered from the largest to 

SMITHSOSIAN CONTRIBUTIOSS TO BOTANY 

the smallest units, and unimportant words are omit- 

ted; latitude and longitude, if given, are placed last 

in this field; and ecological terms, except where re- 

quired to clarify the geographic location, are omit- 

ted. Because locality data from the specimen and 

the original description often are merged into a 

single telegraphic statement, the reference given in 

this fourth field cannot be assumed to be a direct 

quotation. Every effort is made, however, to stay 

close to the words of the original collector and/or 

author of the description, and substantive addi- 

tions or interpolations by the compiler or editor are 

indicated appropriately. It is important to the 

etli tor, therefore, that compilers set apart clearly 

their own comments from the original information. 

Quotation marks are used only when the context 

requires that the exact original words be identified, 

as, for example, when some part of the locality 

statement is so archaic, confusing, or general as to 

appear to contradict the rest of the geographic in- 

formation. 

The three, higher level geographic data fields are 

used to place the locality in its proper geopolitical 

hierarchy. IVhereas the vocabulary used in the 

“Locality” field is standardized for sorting purposes 

but not controlled, the vocabulary used in these 

three fields is controlled as well as standardized for 

purposes of search and retrieval. Insofar as possible, 

current official political units are used in all three 

fields so as to avoid overlapping and inconsistent 

terminology. As the term “geopolitical” implies, 

concessions to age-old geographic designations, as in 

the case of certain islands, are made in a few in- 

stances, and “country” is not always an independent 

political unit in the strictest or most modern sense. 

Such changes are made only within the structure of 

the controlled vocabulary, however, as explained 

below. Island names present a particular problem, 

because often they have long been used in the bio- 

geographic literature but do not fit into a consistent 

geopolitical hierarchy (e.g., Borneo, Madagascar). 

Various stratagems, mostly involving comments in 

the “Locality” field, have been devised to cope with 

the problem of identifying well-known biogeo- 

graphic areas within the file structure of the Type 

Kegister. If the name used in any of the three 

higher level fields is not obviously equivalent to 

the name used originally by the collector, then his 

original designation is included parenthetically

NUMBER 12 19 

with appropriate annotation at the end of the 

“Locality” field. 

A11 available atlases and gazetteers are used as 

standard references, the most valuable being The 

Columbia Lippincott Gazetteer of the World (Selt- 

zer, 1962) and the desk-top Webster’s New Geo- 

graphical Dictionary (1972). The latter, to the 

extent that it covers the geographic units and prob- 

lems encountered, has taken on the force virtually 

of an authority for geographic standardization. The 

authority adopted for the system of classification 

and names of the world’s countries and equivalent 

political units is the National Bureau of Standards’ 

Federal Information Processing Standards Publica- 

tion (FIPS Pub) 10: Countries, Dependencies and 

Areas of Special Sovereignty (1970). With slight 

modifications for the purposes of Type Register, 

this publication is used to control the vocabulary 

used in the first or “Country” field. Any new name 

must be fitted into this system before it can be 

used. 

From the data-processing point of view, the pur- 

pose of the geographic information is to make 

search and retrieval possible at least by country and 

state or province. Thus if the first two fields are 

left blank or if the names are not carefully control- 

led and standardized the retrieval aim is clearly 

thwarted. Collaborators should make every effort 

to provide data in all four geographic fields, but the 

most important ones are the first, second, and 

fourth. 

Level 3-Specimen Data 

Because there may be multiple type specimens 

(e.g., isotypes) in any type collection, the specimen 

data fields may repeat as a set any number of times. 

Ordinarily, an institution will be represented by a 

single type specimen under a given collection, but 

there is no limit to the number of specimens that 

may be registered per collection as long as each 

specimen is uniquely identified. In practice, this 

means that two or more specimens will be cited for 

the same institution onty if they have different 

herbarium sheet numbers or represent different 

kinds of types. The data fields in the specimen 

citation are: Herbarium Abbreviation/Herbarium 

Sheet Number/Kind of Type/ Data Source Code. 

The typical specimen citation takes this form: US 

1727345 HOLOTYPE CF. 

HERBARIUM 

AABBREvIATION.-The standard inter- 

national abbreviations established in the fifth edi- 

tion of “Index Heibariorum, Part I: The Herbaria 

of the SVorld” (Lanjouw and Stafleu, 1964) are 

used to designate the herbaria. 

HERBARIUM SHFET NuniBER.-Many herbaria 

stamp a serial number on every sheet to which an 

herbarium specimen is attached, and that number is 

entered into this field. The field may be left blank 

when the sheet in question lacks a serial number. 

Because a sheet number represents the single most 

effective and reliable means of uniquely identifying 

n specimen, any collaborating herbarium which 

presently does not number its sheets is strongly 

urged to number the sheets of type specimens as the 

data are compiled for the Type Register. From the 

standpoint of the Register, the serial number is a 

completely arbitrary datum and need not belong to 

any general numbering system within the collabo- 

rating institution provided that it is part of a 

unique series. SVhenever there is any choice on the 

matter, a totally numerical series, not a mixed 

alphabetical/numerical (alphanumeric) series, 

should be used, to facilitate proper numerical sort- 

ing b) machine. 

Kmn OF TYPE.-This small field represents the 

purpose of the Type Register and is certain to evoke 

more discussion and controversy than any other 

data field in the unit record. For this reason it is 

vital that every user of the Carex Catalog or any 

other part of the Register understand from the 

outset the limitations of the data recorded in the 

“Kind of Type” field. 

.is emphasized repeatedly, the initial aim of the 

Type Register project is to record the facts just as 

they exist in the presently available sources so as to 

put before the taxonomic user community the 

greatest amount of information in the shortest pos- 

sible time, leaving until later stages the objective of 

methodical, authoritative verification and valida- 

tion. Once comparative data on type specimens are 

available group by taxonomic group on a large 

scale, the specialists themselves, who alone are truly 

qualified to render authoritative decisions on mat- 

ters of nomenclature and typification, can help 

enormously to refine the Type Register data base 

through feedback arising from actual use of the 

file. The consequence of this register-now-validate- 

later approach is seen most often in the imprecise 

if not incorrect terms by which the different kinds

20 

of types are designated. Because the nomenclatural 

rules have changed through the years and the well- 

developed modern terminology is of relatively re- 

cent origin, it is natural that types have not been 

designated according to any consistent standards 

through the years. 

When a type specimen is first registered, its typi- 

fication is designated by whatever term is indicated 

in the data source, which usually is a secondary 

source (card file, specimen file, monograph), unless 

there is firm evidence to indicate otherwise. Thus 

any type designation, whether legal or illegal by 

present nomenclatural rules, may appear in the 

Type Register. 

At the United States National Herbarium, it was 

customary for many years to designate two basic 

categories in the segregated collection of type 

specimens: “type” and “type collection.” Usually, 

“type” has meant what would now be called “holo- 

type,” while “type collection” has embraced syn- 

types, isotypes, and even paratypes according to 

present terminology. “Types” often prove not to be 

holotypes, however, and it would be very wrong to 

draw simple equations between the older and newer 

terms. The collection of type specimens at the New 

York Botanical Garden provides another example 

of the problems with archaic terminology. Here the 

categories “type” and “cotype” were used for many 

years, and now “type” often but certainly not always 

translates to “holotype,” while “cotype” may desig- 

nate any of the kinds of types masquerading under 

“type collection” at the US. 

The authority for designation of kind of type 

is the ICBN. An auxiliary, highly authoritative 

standard reference is “An Annotated Glossary of 

Botanical Nomenclature,” by McVaugh et al. 

(1968). LVhenever the original specimens and liter- 

ature can be examined and the kind of type vali- 

dated in accordance with the rules, proper termi- 

nology is used. By this terminology, the Register 

is designed to include primarily lzolotypes, isotypes, 

syntypes, and isosyntypes. As explained earlier, the 

present file structure is not designed to handle lec- 

totypes and neotypes (or isolectotypes and isoneo- 

types), although these are entered sometimes by 

using the fourth geographic field (“Locality”) as a 

remarks field for the second bibliographic citation. 

Paratypes are excluded unless other, higher order 

types cannot be located and there is reason to be- 

lieve that the paratypes will become important later 

SMITHSONIAN COSTRIBUTIOSS TO BOTANY 

for purposes of lectotypification. Fragments of holo- 

types, isotypes, or syntypes may be included at the 

discretion of the editor (see under “Scope of Reg- 

ister”). 

When a holotype has not been designated, as in 

all the older literature, one usually is faced with a 

“syntype situation,” which often is difficult to re- 

solve precisely on the basis of the ZCBN’s terminol- 

ogy. A syntype, according to the ZCBN (Article 7, 

Note 3), is “any one of two or more specimens cited 

by the author when no holotype was designated, or 

any one of two or more specimens simultaneously 

designated as types,” and an isosyntype is a dupli- 

cate of a syntype (see “Guide for the Determination 

of Tjpes” in the ZCBN). “Duplicate” in this con- 

text is defined as “part of a single gathering made 

by a collector at one time.” In other words, a “dup- 

licate” is one of two or more specimens constituting 

a single “collection,” as this term ordinarily is used 

by plant taxonomists and is being used in the con- 

text of the Type Register. 

The distinction betxveen syntype and isosyntype 

hinges on such relatively subjective criteria as 

“specimen citation” and whether or not the original 

author had the specimens in hand (cf. definitions 

of McVaugh et al., 1968), which are matters for 

specialists to determine. The older literature, where 

the problem of syntypes arises, is well known to be 

less than precise in the manner of citing specimens. 

The editorial staff of the Type Register must re- 

strict its interpretations to the letter of the ZCBA‘, 

and for the most part such fine distinctions as be- 

tween syntype and isosyntype necessarily are defer- 

red for the proper specialists to make at a later 

time. To do otherwise would be to assume the 

specialist’s ro!e and responsibility and to introduce 

false precision at this stage. Accordingly, the term 

syntype is used for both syntypes and isosyntypes 

except in the rare cases where the evidence for the 

isos) nt) pe designation is clear and convincing. 

A final point on the use of the term syntype con- 

cerns the distinction between single collections and 

multiple collections. Throughout the older litera- 

ture there are numerous cases where a single col- 

lection has been designated as the type collection, 

either explicitly or implicitly by virtue of being the 

only collection cited, even though a holotype has 

not been set apart. hiany specialists would single 

out a presumptive holotype in these cases on the 

hsis of the rpecimen (s) which the original author

SUMBER 12 21 

is presumed to have examined firsthand, but the 

Type Register editors cannot and should not make 

authoritative selections in such cases and must regard 

them all as syntypes. The ZCBN does not seem 

to provide term for distinguishing this common 

syntype situation from the other common syntype 

situations in which two or more type collections are 

designated simultaneously. Because it is useful to 

know Lvliether one or more than one type collection 

is cited, in the Type Register project the term “type 

collection” has been given a proper meaning for 

the purpose of distinguishing these two syntype 

situations. Type collection, in this proper sense, 

designates a specimen from a single type collection, 

while “syntype” is reserved for designating a specimen 

from any one of two or more simultaneously 

designated type collections. 

The catchall term type material is used to designate 

any specimen presumed for some reason to be 

a type but for which there is no basis at the time of 

data input to assign a more precise classification. 

Many situations arise in the course of compiling 

and editing data for the Type Register in which 

it would be useful to have a collective term for 

designating a collection as a counterpart to Ihe 

singular term given in the ZCBN for the specimen. 

In fact, the terms “holotype collection” and “syntype 

collection” often are used informally within 

the project as collective counterparts to the singular 

terms holotype/isotype and syntype/isosyntype. 

DATA SOURCE Com.-Source of data is indicated 

according to a classification of source categories, by 

appending the appropriate code at the end of the 

specimen citation. It is impossible to document in 

detail the source of every datum, and any categorization 

of sources is certain to have many imperfections. 

The present classification is only a rough first 

approximation of the kind of documentation needed, 

but presumably it is a strike in the right direction. 

It is drawn up largely from the point of view 

of the central staff and their internal Smithsonian 

operation, and other categories will have to be 

added as other institutions join in the effort. Probably 

codes or numbers will be assigned to individual 

contributors in the future in the manner of Index 

A’ominzrm Genericorurn. It should be noted that 

this is more than a classification of sources; up to a 

point it also is a classification of degree of verification/validation. 

Obviously, a record based on examination 

of the original publication and the original 

specimen is likely to be more reliable than a record 

based solely on secondary sources. Following is the 

classification of data source codes: 

0s 

OP 

TS 

IM G 

ss 

CF 

co 

cs 

CM 

UIL 

Original publication and type Specimen examined by 

person compiling data for Register. 

Original Publication examined by person compiling 

data for Register, but type specimen not seen: 

supplementary information about the specimen, if 

any, derived from secondary source (s) . 

T\pe Specimen examined by person compiling data for 

Register, but original publication not seen: citation 

and other publication data, if any, derived from 

secondary source (s) , including standard indices (In- 

dex Kewensis, Gray Herbarium Index) , monographs 

and revisions, annotations on specimen sheet, card 

files, original dacriptions removed from context of 

publication without exact citation and necessary 

prefatory matter, and the Type Register catalog itself. 

Data derived from most recent MonoGraph of taxo- 

nomic group in question without reference to any 

other source (s) of information. 

Data derived entirely from Secondary Sources. 

Data transcribed directly from a card in the Card 

File of the type collection of the U. S. National 

Herbarium without verification against the original 

publication or type specimen. 

Data from US Card file verified by examination of 

Original publication. 

Data from US Caid file verified by examination of 

t!pe Specimen is) . 

Data from US Card file verified or supplemented by 

consulting latest Monograph of taxonomic group in 

question. 

Source of data UnKnown. 

PROCESSING SYSTEM 

The first step is to convert the data to machine- 

readable form, i.e., to “automate” or “capture” the 

data, so that they can be processed by computer. 

No data conversion (automation, capture) system is 

perfect, and none is capable of handling all appli- 

cations equally well (Shetler, 1972). Several meth- 

ods and media have been tried thus far in the Type 

Register project in an effort to find the data con- 

version system best suited to this data-processing 

application. In general, the aim is to use the system 

that will get the data into the computer with the 

least amount of error and effort. Because the devel- 

opment of data conversion devices and procedures 

continues to evolve rapidly, a flexible approach has 

been taken; the data conversion system is kept as 

independent as possible from the rest of the proces- 

sing system so that a new conversion system can be

22 

adopted at any time with minimal impact on the 

overall Type Register operation. 

The following data conversion systems have been 

used in the chronological order given: 

PAPER TAPE SYsTEM.-Data were mechanically 

encoded on paper tape with a tape-punching typewriter. 

The tapes then were read by the computer 

which converted the holes in the paper tape (i.e., 

mechanical codes) to electronic codes on magnetic 

tape and thus transferred the machine-readable 

data from the medium of capture to the medium of 

computer processing. This system was used for the 

duration of the pilot project, and several thousand 

records, a third of the present file, were captured 

with it. During the pilot phase, while paper tape 

lvas being used, corrections to the machine file 

were made by means of the standard 80-column 

punch card. 

ON-LIKE SYsTE&t.-with a typewriter terminal, 

data were entered via telephone directly on disk 

storage of a remote time-sharing computer under 

the on-line control of a sophisticated text-editing 

program package. This text-editing software permitted 

the terminal operator to direct the computer 

in making any of a whole series of deletions, 

changes, and additions during the input process so 

that maximum editorial accuracy could be achieved 

in the machine-readable data base immediately, before 

it was output onto magnetic tape for subsequent 

processing by the information retrieval system. 

OPTICAL SCANNING SYsTEM.-Data were typed on 

standard forms with an ordinary 10-pitch IBM Selectric 

Typewriter equipped with a head having 

a special optical scanning font. Completed forms 

were scanned by an optical character reader (OCR) 

which encoded the data directly on magnetic tape 

for further computer processing. 

~IAGNETIC TAPE SYsTE;M.-In the system currently 

being used, a typewriter encodes data electronically 

on a magnetic cartridge which is compatible with 

computer tape. The typewriter unit also serves as a 

communication terminal to transmit the data captured 

via telephone directly to the computer, where 

the data are transferred to disk or tape for further 

processing. Processing programs can be controlled 

from this same remote terminal. Of the several 

data conversion systems used, this one seems to offer 

the best compromise of advantages and disadvantages 

in the context of the operation as a whole. 

ShiITHSOSIAN CONTRIBUTIONS TO BOTANY 

COMPUTER PROCESSING SYSTEM.-During the pilot 

phase of the Type Register project, this system consisted 

of a set of specially written COBOL programs, 

designed to run on the Smithsonian’s 

Honeywell 1250 machine, to create and maintain a 

machine file, and to account for collaborating institutions 

to which data-collection cards were sent or 

from which data were received. It was not designed 

as an information retrieval system in the strict 

sense, and in terms of output the system was capable 

chiefly of producing catalogs, either in book or 

card form. The pilot system proved with use to be 

highly specialized with too few capabilities and 

serious weaknesses in file structure such as inability 

to handle syntypes. It served the purpose of a pilot 

system, however, to get a file started and thereby 

define through experience the problems to be 

solved in the operational system. 

After the pilot phase, the specialized COBOL 

programs were abandoned, and the processing system 

was completely redesigned around the commercially 

available, IBM-supported program package 

known as the “Generalized Information System” 

(GIs), which runs on the larger models of 

IBRl’s System/360 and System/370. This generalized 

software performs all the normal functions of 

creating, maintaining, and querying files and of 

generating reports. It is an information retrieval 

system, in the proper sense, with the full capability 

to search, select, and print answers to specific 

queries on demand in addition to the capability for 

producing various types of tabulations, tallies, and 

catalogs. A COBOL preprocessing program (“preprocessor”) 

and a COBOL concatenating program, 

by which, respectively, the data are prepared for 

processing by GIS and the data are joined field by 

field into publishable paragraph output after processing 

by GIS, form a part of the total operational 

system. GIS runs only on IBM equipment and is 

offered by several computer service bureaus in the 

Washington, D.C., area with IBM machines. Type 

Register processing with this system so far has been 

carried out successfully at several different service 

bureaus, and the project basically is independent of 

the computing center. 

One of the most important initial tasks in developing 

the Type Register is to build a file of 

sufficient size to make reliable studies concerning 

such matters as record comprehensiveness, record 

format, field format, need for authority files, and

SUMBER 12 23 

FIGURE %-Two oversimplified schematic representations of the three-level record 

structure in the Botanical Type Specimen Register (from Meadow, 1973a,b). 

report types and formats, This requires a flexible 

information processing system that allows for data 

to be restructured, reports to be reformatted, tallies 

to be made, and edits and mass updates to be made 

without reprogramming. From the output stand- 

point, it must be possible to select records according 

to the content of any data field, to relate records to 

each other on the basis of selected fields, and to sort 

and format selected fields as desired. GIS affords 

all these capabilities. 

The FNA program is using GIS, and it was for 

this reason particularly that the decision was made 

to use GIS in the Type Register project so that 

these botanical data bases would remain fully cont 

patible. The rationale for using a generalized infor- 

mation processing system and a description of the 

use of GIS in the FNA program have been set forth 

in separate papers recently by Harriet R. Meadow 

(1973a, b), who is responsible for the basic design 

of the Type Register processing system. The two 

most important features of GIS, from the point of 

view of biological data retrieval, are its capability 

of handling hierarchical data structures and its 

capability of querying two or more files simultane- 

ously for correlated data. With respect to Type 

Register, for example, the hierarchical feature per- 

mits subordination of two or more specimen collec- 

tions to a single taxon or, in turn, two or more 

specimens to a single collection within a taxon. The 

three-level hierarchy of the Type Register is out- 

lined in Figure 3. 

The second-generation processing system for the 

Type Register by no means represents the ultimate 

system, but it does handle the vast majority of cases 

very well and solves the most bothersome problems 

encountered during the pilot phase with the special- 

ized first-generation system. New problems have 

arisen, however, and with the experience gained by 

using the present system it will be possible to de- 

sign a third-generation system in due course that 

will accommodate all of the special cases that con- 

tinue to be troublesome, e.g., the case of a lectotype 

or neotype that requires a second bibliographic 

citation. 

Statistical Summary of Type Register Contents 

Following is a statistical summary of the records 

on the machine file as of 30 September, 1972:

24 

No. Families 135 

No. Genera 950 

No. Taxa* 10,525 

So. Collections 10,625 

So. Specimens 13,535 

*Species, subspecies, \ arieties, forms, and nothomorphs. 

The figures show that the number of collections 

averages just slightly more than l/taxon, while the 

number of specimens averages about 1.3/taxon. The 

number of specimens will grow rapidly relative to 

the number of taxa as additional herbaria register 

their tjpe holdings of taxa already in the file. 

This is a tally largely of records input from the 

tjpe collection of the U. S. National Herbarium 

(US); i.e., the vast majority of the families and 

genera are represented only by taxa, collections, 

arid specimens in the US type collection. None of 

the 135 families is present solely on the basis of 

tjpes 1 egistered from another herbarium, which is 

to say that at least one taxon in the US collection is 

registered under every family. The inclusion of a 

family does not mean, however, that all US types 

belonging to that family have been recorded. Quite 

the contrar), the project has only begun, and, as in- 

dicated earlier, it now is proceeding alphabetically 

by genus and is still in the letter “C.” This means 

that for the vast majority of families only genera 

starting with “A” or “B” are recorded thus far. 

If a genus is present, however, then all type ma- 

terial in the US collection belonging to that genus 

is registered. In other words, the file is complete to 

the genus level with respect to taxa, collections, and 

specimens in the US type collection. 

Before the alphabetical approach was started, reg- 

istration of US types was essentially completed for 

several families, and in each case the types of one 

or more other herbaria also had been recorded. 

Following is a list of these completed families, show- 

ing for each the number of genera, taxa, and speci- 

mens registered and the abbreviations of the her- 

baria for which the file is relatively complete: 

Chrysobalanaceae (12 genera, 216 taxa, 1,110 specimens), 

many herbaria 

Dichapetalaceae (7 genera, 43 taxa, 153 specimens), man\ 

herbaria 

Lamiaceae (84 genera, 1,234 taxa, 1,851 specimens), LA, MO, 

XY, us 

Ecrophulariaceae (85 genera, 943 taxa, 1,073 specimens) , V5 

Violaceae (17 genera, 232 taxa, 321 specimens), NY, US 

The data for the Chrysobalanaceae and Dichapet- 

alaceae were provided by Ghillean T. Prance of the 

Kew York Botanical Garden from his manuscripts 

SMITHSOXIAX CONTRIBUTIONS TO BOTANY 

of famil) monographs for the Flora Neotropica se- 

ries. All type specimens seen by him in the course 

of his research on these families as of June 1970 are 

recorded, which means that many herbaria are rep- 

resented. With respect to these two families, the 

Type Register is relatively “complete” in the com- 

prehensive, monographic sense in that the file cites 

the significant types, regardless of where they are on 

deposit, as they will be cited in the published 

monograph. (The monographs of these families 

have since appeared-see Prance 1972a, b.) At the 

same time the coverage may not be as thorough for 

any individual herbarium as in the case of the 

other three families-Lamiaceae, Scrophulariaceae, 

Violaceae-for which the data were compiled di- 

rectly from type collections or card files in the her- 

baria indicated. Within the Scrophulariaceae, con- 

tributions to the genus Mimulus have been regis- 

tered by more than a dozen herbaria (CAN, COLO, 

DXO, F, GH, JEPS, RlICH, NY, OSC, PH, uc, 

LA, LTS, 1171s) as a result of Hale’s experi-ent 

(see “Introduction”), and there are miscellaneous 

other contributions to this family recorded from 

110, NY, and a few other herbaria. 

Apart from the families listed and the genus 

Carex, for which the catalog is appended, several 

other groups have been completed in some sense. 

Nearl) half of the US types of Asteraceae (Compos- 

itae) had been recorded when the switch from a 

systematic to an alphabetical approach was made, 

and the file for this family contained as of 30 

September 1972 the following: 255 genera, 2,600 

taxa, and 2,650 specimens. Several important genera 

of the Brassicaceae (Cruciferae) have been complet- 

ed at least for the US, and the tallies for these 

genera as of 30 September 1972 were: 

Arabis (97 taxa, 106 specimens), US 

Draba (119 taxa, 195 specimens), NY, US 

Lepidium (29 taxa, 32 specimens), US 

Lesquerella (38 taxa, 43 specimens), US 

Thlaspi (15 taxa, 59 specimens), F, GH, MO, XY, RM, S, UC, 

US, WTU 

The data for Thlaspi were provided by Patricia 

Kern Holmgren (1971) of the New York Botanical 

Garden from her revision of the genus, during 

which she saw types from the above-indicated ten 

herbaria. She also provided the data from NY for 

the genus Draba. 

Finally, John T. Rfickel of the New York BO- 

tanical Garden provided significant type data on

NUMBER 12 25 

the genus Anemia subgenus Coptophyllum (Schizae- 

aceae) and its three segregate genera Aneimiaebo- 

trys, Coptophyllum, and Trochopteris from his 

monograph of the Anemia (Mickel, 1962), and the 

statistics are: 4 genera, 35 taxa, 81 specimens. 

Use of Type Register 

Some of the main uses of the Type Register will 

be obvious from the discussion in the foregoing 

sections if not from the concept of the Register 

itself, and other uses will become apparent through 

study of the Carex Catalog and its indices. It 

should be emphasized that a catalog of this type 

with the same or other types of indices can be pro- 

duced for any taxonomic group, large or small, 

presently registered, although the data have not 

been edited to the same high degree in any other 

group thus far. The Carex Catalog and each of its 

indices represent outputs to particular queries. 

Many other types of queries are possible, and the 

amount of output depends on the scope of the 

query and the depth and comprehensiveness of the 

data base at the time of querying. For example, the 

request “Print all records of ferns,” would yield a 

relatively small printout at this stage, because only 

a few fern types are registered, but eventually such 

a request could yield an overwhelming printout. In 

querying the file, the user always must exercise dis- 

cretion in framing his requests, and to do this he 

must have a reasonable knowledge of the limits of 

the machine file beforehand or be guided by some- 

one who is familiar with the file. Otherwise he will 

make meaningless or impractical requests. 

By request the file will be queried at cost for 

anyone. Acy kind of query is welcome, and guid- 

ance can be provided in framing queries. It is im- 

portant at this stage to have feedback from poten- 

tial users in the form of requests for file queries so 

that all needs are taken into consideration as the 

Type Register system undergoes further test and 

refinement, particularly with respect to report for- 

mats. Persons wishing to make extensive use of the 

Register should plan to spend time in Washington, 

D.C., working with the project staff at the Smith- 

sonian. The costs and other requirements of such 

an undertaking should be worked out in advance 

by consultation with the staff. 

The Type Register can be queried or sorted by 

taxonomic name, author, book or journal title, 

year of publication, collector, collector’s number, 

date of collection, country, state or province, county, 

herbarium, and kind of type or any combination of 

these. IVith a query or sort on any of these fields 

can be printed other selected fields from the same 

records, as illustrated by the Carex Catalog, which 

is sorted by taxonomic name, and its five indices, 

which are sorted in the lead field by author, collec- 

tor, country, publication date, and herbarium, re- 

spectively. 

With the cooperation of specialists willing to de- 

vote time to editing of groups of interest to them- 

selves, other catalogs can be published. Anyone 

is invited to propose collaborations in publishing 

from the Type Register.

A Catalog of the Genus Carex (Cyperaceae) 

STANWYN G. SHETLER (Editor); MARY JANE PETRINI, CONSTANCE GRAHAM CARLEY, M. J. HARVEY, LARRY E. 

MORSE (Assistant Editors); THOMAS E. KOPFLER (Programmer); AND COLLABORATORS 

Contributing Institutions 

The ten American herbaria that have collaborat- 

ed in the Carex project to provide data on their re- 

sective collections of types in this genus are listed 

here in descending order of number of specimens 

registered. The name of each herbarium is preceded 

by its standard international abbreviation as estab- 

lished in the fifth edition of “Index Herbariorum, 

Part I, The Herbaria of the World” (Lanjouw and 

Stafleu 1964). 

NY Herbarium, New York Botanical Garden, Bronx Park, 

New York 

GH Gray Herbarium, Harvard 

Massachusetts 

University, Cambridge, 

US United States National Herbarium, Smithsonian Institution, 

Washington, D. C. 

CAS Herbarium, California Academy of Sciences, San Francisco, 

California 

F John G. Searle Herbarium, Field Museum of Natural 

History, Chicago, Illinois 

MO Herbarium, Missouri Botanical Garden, St. Louis, 

Missouri 

DS Dudley Herbarium, Stanford University, Stanford, 

UC 

California 

Herbarium, University of California, Berkeley, California 

A Herbarium, Arnold Arboretum, Harvard University, 

Cambridge, Massachusetts 

JEPS Jepson Herbarium, University of California, Berkeley, 

California 

Each institution has contributed to the Carex 

Catalog voluntarily and with little or no outside 

financial support for work on the Type Register. 

The FNA program has made use of the Type Register 

as a pilot project to evaluate various ways of 

collecting, editing, capturing, and processing hierarchically 

structured taxon/collection/specimen data. 

In the course of this work, therefore, small sums of 

“seed” money were allocated from FNA funds to 

support data-gathering for the Carex project at sev- 

26 

eral of the above institutions. The Carex project 

was undertaken in the first place at the suggestion of 

the FNA Editorial Committee. The Committee saw 

in Carex a good model of a large genus with a pre- 

ponderance of North American species which pre- 

sents a full gamut of nomenclatural and taxonomic 

problems to cope with in an information system. 

(The Carex project, insofar as FNA was concerned, 

went beyond the Type Register. While the data 

were being gathered for the Register, other morpho- 

logical data were also being assembled by FNA- 

unpublished Carex data on Sections Montanae and 

Ovules collected by A. J. Gilmartin and M. B. 

Moore-to test the matrix techniques of Morse, 

197 1, for computer-assisted identification and key 

construction.) Once the Carex Catalog was estab- 

lished on the basis of several large herbaria, other 

major herbaria were able to add their data with a 

surprisingly low investment of man-hours. The 

Field Museum, for example, reported just under 35 

man-hours-less than a man-week-to compile its 

data on Carex types, at a cost of under $175 for 

technical labor, but by the time the Museum came 

into the picture it had the benefit of a basic catalog 

already including the contributions of A, GH, MO, 

NY, and US. 

Each institution determined its own extent of 

coverage, and without exception each tried to in- 

clude all types in the general herbarium which al- 

ready had been segregated or otherwise identified 

or which could be identified quickly on the basis of 

the available computer printout. At the same time, 

none of the herbaria makes any claim to absolute 

completeness or thoroughness relative to the num- 

ber of types in the herbarium, including those still 

buried in the general collection. A pragmatic phi- 

losophy prevailed, with each herbarium doing the 

best it could under the constraints of its own local

SUMBER 12 27 

circumstances. It might be assumed in general 

that the coverage is more complete for the herbaria 

like NY and US which had separate type collections 

in the first place and had long-established tradi- 

tions of segregating type specimens from the main 

herbarium as they are recognized, than for the her- 

baria without segregated types, but even this as- 

sumption can be questioned. At CAS, a folder-by- 

folder search of Carex specimens in the herbarium 

with printout in hand turned up as many unrecog- 

nized type specimens as had previously been recog- 

nized and segregated into the type collection. A 

similar search of North American folders at F re- 

sulted likewise in doubling the number of recog- 

nized types. If one pass through the herbarium can 

double the number of types, it certainly cannot be 

concluded that all types have now been found. 

From a purely curatorial point of view, therefore, 

the Carex project was very useful in these instances 

because, as a result, the collection of recognized 

types was increased by 100 percent. 

Clearly it would be unfair to judge the relative 

size or value of the type collection or even just the 

Catex type collection of any of these ten herbaria 

on the basis of the present catalog. Much more 

extensive development of the Type Register is re- 

quired before such judgment will be warranted. 

At the same time, the present ranking is almost 

predictable. An herbarium that is the major de- 

pository for the types of a monographer naturally 

is expected to have a disproportionate number of 

types in the group (s) on which the monographer 

worked. The extensive work of monographer K. K. 

Mackenzie (1 93 1-35, “Cyperaceae-Cariceae,” in 

hrorth American Flora) clearly might be expected, 

therefore, to have put NY in the first position, and 

no one will be surprised to find GH and US close 

behind. Comparison of the NY and US type col- 

lections of Carex provides a good example of the 

influence of a monographer on a type collection. 

Together, NY and US have type specimens for over 

400 taxa. Of these taxa, half are represented by 

type specimens only at NY, while about 40 are 

represented only at US. Thus, NY has five times 

more “unique” Carex taxa in the Type Register 

than US. By contrast comparisons in some other 

groups show NY and US to have about an equal 

number of “unique” representatives in the Regis- 

ter. CAS is expected to be strong in types of west- 

ern taxa. In addition, John Thomas has pointed 

out (personal correspondence) that CAS has a 

complete set ot H. P. Sartwell’s Carices Arnericanae 

SeptentiionaliJ, Part 1 (nos. 1-70, 1848) and Part 

2 (nos. 71-143, 1850), and a “rather good set, but 

not complete,” of S. T. Olney’s Carices Boreali 

Ametzcunae, distributed in 1870 and 1871. Accord- 

ing to Thomas, many CAS specimens bear the an- 

notations of J. W. Stacey, who was connected with 

CAS and published on western sedges. 

Thomas points out these additional interesting 

facts about the other herbaria in the San Francisco 

Bay area. Mackenzie, in the course of doing the 

treatment of Carex for Abrams’ An Zllustrated 

Flom of the Pacific States (Mackenzie, 1923), de- 

termined all the Carex specimens at DS from the 

Pacific States and a number from other regions 

sometime prior to 1920. The Parish Herbarium, 

one of the best early collections of southern Cali- 

fornia plants, is housed at DS. The I. W. Clokey 

Herbarium, including his sedge types, are at UC, 

and of course W. L. Jepson’s material is at JEPS. 

In general, every curator who collaborated in 

the Caiex project felt that his effort had beneficial 

consequences in the herbarium, and there was a 

nearly unanimous opinion that the results were 

well worth the effort from the curatorial stand- 

point alone. 

Data-Collection Procedure 

The Carex Catalog was initiated with data from 

the US and hfO type collections. For the US, rec- 

ords were converted from the existing card file 

(see “Source of Data”), while the MO records were 

captured from data forms filled out at MO in the 

course of a special search of all the Carex type 

folders, which are distributed through the herbar- 

ium in association with the main collections. The 

special search was undertaken specifically as part 

of the Carex project of FNA. 

After a common catalog of the US and MO col- 

lections was compiled, members of the Type Regis- 

ter staff moved their data-conversion operation to 

the New York Botanical Garden for two weeks, 

where, with the assistance of NY personnel, the 

NY Carex data were compiled and input to the 

system in a matter of days. The data were com- 

piled from the specimens in the separate type col-

28 

lectioii, which by and large does not include the 

literature citations, and supplemented with bibli- 

ographic data from reference works, primarily 

Mackenzie’s (193 1-35) monograph of Carex. 

It should be pointed out that the type collec- 

tion at NY was first segregated for security reasons 

during World War 11, and the selection had to be 

done in haste under less than ideal conditions. 

Consequently, many nontype but historically im- 

portant specimens, constituting perhaps as much as 

25 percent of the total type collection, deliberately 

were pulled along with the known or presumptive 

type specimens in the process of going through the 

main herbarium. The person who compiled Cnrex 

data attempted to sort out the nontype material, 

and further culling was done during the editorial 

process. Undoubtedly some nontype material still 

remains in the present Catalog, although it seems 

unlikely that the percentage of such specimens is 

much if any higher for NY at this stage than for 

any of the other nine herbaria. In any event, it 

will be a simple matter to delete nontypes from 

the file as they are discovered and brought to the 

attention of the Type Register editors in the future. 

Once the NY data were merged fully with the 

US and MO data, a union catalog was delivered to 

Harvard University for additions from A and GH. 

.4t Harvard, data on types of taxa already regis- 

tered were annotated in the catalog, and data for 

taxa new to the catalog were compiled on standard 

forms. In general, Harvard provided only specimen 

data, and the bibliographic data were looked up 

and supplied later by the Type Register staff at the 

Smithsonian before annotations and new records 

were captured ant1 merged with the US + MO 

+ h’Y machine file. 

When the annotated catalog was returned from 

Harvard, the X and GH annotations were tran- 

scribed to the main working catalog at the Smith- 

sonian, and then the Harvard catalog, as annotated, 

was sent to t k Field Museum of Natural History. 

Sent with the annotated catalog were photocopies 

of the data forms for the new taxa added by A 

and GH. In this way it was possible to give the 

collaborators at F the benefit of the new data sup- 

plied by A and GH immediately, while the capture 

of these new data was still in progress at the Smith- 

sonian. As a further aid to their work, the col- 

laborators at F were provided with an index by 

SMITHSOXIAN CONTRIBUTIONS TO BOTANY 

tollectoi and collector’s number to the US + h10 

+ NY catalog. 

The collaborators in the San Francisco Bay area 

weie provided with a new catalog incorporating 

the A and GH data along with the US, MO, and 

NY data, and thia catalog also included an index 

by collector and collector’s number. This catalog 

was annotated by them with new data from CAS, 

DS, JEPS, and UC on taxa already registered, and 

data for taxa new to the file were compiled on 

standard forms. The annotated catalog and com- 

pleted forms then were returned to the Smithsonian 

Institution for input to the system. 

’it CAS, DS, F, JEPS, and UC the data were 

compiled in the first instance from the specimens 

in the herbarium, but in most cases the original 

publications also were checked when taxa new to 

the file were involved. Otherwise the bibliographic 

data were obtained from secondary sources. The 

original literature was checked for taxon additions 

without exception at F. 

In summary, a round-robin procedure of sorts 

was used to collect the data. To the extent possible, 

each new collaborating institution was given the 

benefit of the latest cumulative catalog incorporat- 

ing the contributions of previous collaborating 

herbaria. In this way, maximum advantage could 

be taken of previous herbarium and library re- 

search, and duplication of effort was kept to a 

minimum. Once the combined data of CAS, DS, 

TEPS, and UC were returned to the Smithsonian, 

the data-collection phase was closed out and final 

editing began. 

Editorial Process 

All entries were edited in accordance with the 

principles and procedures set forth in the first part, 

and the editors take final responsibility for the 

form and style and all other editorial matters of 

the present Catalog. To integrate new contribu- 

tions into the accumulating data base, differences 

between supposedly identical records from different 

institutions constantly had to be reconciled by turn- 

ing to standard references and the original litera- 

ture. Whenever possible, record content was verified 

by checking the original publication. In the end, 

nearly every original description cited in the Cata- 

log was seen at least once by the editors and in 

many cases several times. In many cases, further- 

more, the designation of kind of type was validated

SUMBER 12 29 

according to the ZCBh’; however, because much of 

the checking of original literature was done by a 

technical editor, untrained in the application of 

the type method and terminology, many of the 

records were verified without being validated, to 

use the distinction defined in an earlier section 

(p. 00). According to this distinction, the develop- 

ment of the Cuiex file of the Type Register can be 

said to have attained the Stage 2 “verification” 

level overall, with some records still at the Stage 1 

“registration” level and with a substantial number 

of others having been “validated” more or less ac- 

cording to Stage 3 standards. Of course the careful 

scrutiny of specialists, with appropriate feedback 

from them, is needed over a period of years before 

the present Cmex file can be said to have truly 

attained Stage 3 development. 

While the bibliographic data could be double- 

checked or supplied (if not provided in the first 

place) by examining the original publication, the 

specimen data could be verified in this way only to 

the extent that the publications gave corroborating 

details. Ultimately, therefore, each contributing in- 

stitution is responsible for the reliability of its own 

specimen data. In the case of NY, however, the 

editors, who assisted in the data collection itself, 

share responsibility for the reliability of the speci- 

men data. In any event, citations always were 

checked against Zndex Kewensis and/or the Gray 

Herbni !urn Zndpx if the original publications could 

not be examined. 

Editing of this type of open-ended file, in which 

considerable subjective judgment is required, is a 

neyer-ending process, and a reasonable degree of 

thoroughness is achieved only after many editorial 

“passes” through the file. The editing proceeded 

in a series of phases and cycles in which all records 

were examined a field at a time, by means of in- 

verted listings or indices, for consistency and ac- 

curacy, and updated printouts were obtained for 

another editorial round. Thus, for example, title 

citations were standardized in one editorial phase, 

while names of authors were standardized in an- 

other phase. The editorial corrections for all fields 

then were merged into a common working hard- 

copy of the file before the process of updating the 

machine file was begun. After one cycle of such 

editing was finished and the file was thought to be 

“clean,” a new catalog and set of indices were 

printed out. Then a whole new cycle of editing 

was begun. This iterative process continued until 

the editing began to yield diminishing returns and 

it became necessary to bring the never-ending proc- 

ess to a reasonable stopping point, which the pres- 

ent Catalog is believed to represent. In such a 

dynamic system, editorial perfection is relative at 

best, and at this stage the editors certainly make 

no claim to perfection in any sense of the word, 

aithough every effort has been made to be thorough 

and consistent. 

Milestone Events 

Computerized data banks are a new development 

in biology, and there is still much to be learned 

about the mechanics of creating and maintaining 

them. The process is all too easily underestimated, 

especially with respect to manpower requirements, 

and overly optimistic timetables are the rule. An 

enormous effort on the part of many individuals 

\vent into the creation of the Cal-ex Catalog over 

a period of more than two years. During this time 

the tedious editorial work seemed to go on end- 

lessly, while technical problems with the systems 

development and processing also came in a steady 

stream. The following chronology of milestones in 

the more than two-year process has a two-fold 

purpose-first, to provide a practical example of 

the laborious steps involved in creating a data 

bank, and, second, to caution against overenthusi- 

asm and oversimplification on the part of others 

contemplating similar efforts. It must be stressed, 

however, that all during the two and one-half 

years other taxonomic groups were being input to 

the Type Register along with Curex, and it never 

was possible to work exclusively on Carex. 

1970 

January Decision made to begin work on genus as part 

of FNA Carex project. Editing of data in US 

card file begun. 

February 

Collection of data begun at MO and first records 

returned to US for input. 

Photocopies of all Carex records in US card file 

sent to MO. 

March h’ew data-capture procedure organized using online, 

text-editing system. 

April ,411 US and MO records input, totaling just over 

200 taxa and about 250 specimens. 

May First printout of US + MO records produced 

and edited, and machine file updated. 

Second printout produced, and copy sent to NY.

30 

June Second printout of US + MO records edited, antl 

machine file updated. 

Members of Type Register staff worked at NY for 

last two weeks during which time all SY 

Cares records (200-k taxa, 300 specimens), 

along with records of other groups, were input 

to the system on-line via telephone to computer 

in Washington, D.C. 

First printout of new taxa added to file by NY 

produced. 

July Report on NY work prepared. 

August Printout of NY additions edited, antl machine 

file updated. 

First printout of US + MO + NY records produced. 

ceptember rS + MO + NY printout edited, antl machine 

file updated. 

cecond printout of joint records produced, totaling 

415 taxa and 544 specimens. 

Decision made to publish a Carex catalog as the 

sample installment of Type Register. 

Novemtier Preliminary report formats defined for catalog 

and indices. 

December Use of on-line system discontinued, and paper 

tape system reorganized and used again. 

1971 

January- Intensive editing and updating of preliminary 

May 

Carex Catalog (US + MO + NY) continued. 

OCR data-capture system inaugurated, and paper 

tape system phased out. 

June 

Sample catalog of 100 Carex records with 4 crossindices 

produced and distributed to a limited 

cross-section of taxonomists. 

Preliminary catalog of Carex displayed at botanical 

meetings in Edmonton, Alberta, and copies 

of sample of 100 records handed out. 

Decision made to hold off publication of Catalog 

July 

at least until GH could be included. 

Final printout of US + hIO + NY catalog 

generated for GH. 

August Copy of latest catalog delivered in person to 

Harvard University for use in compiling data 

from A and GH; specimen data compiled and 

returned to US. 

Copy delivered also to SY for final prepublication 

check: check made and printout returned. 

Copy sent to MO for final prepublication check. 

Copy displayed at annual meeting of American 

Institute of 

Colorado. 

Biological Sciences, Ft. Collins, 

Contributions solicited by letter from F, Sari 

Francisco Bay area herbaria. 

Feptember F indicated willingness 

records. 

to contribute Carey 

October Arrangements made for F to contribute records. 

and necessary materials supplied, including 

index by collector to preliminary catalog. 

Data collection begun at F. 

Editing of A and GH data begun. 

So\ ember 

December 

January 

Februar) 

March- 

April 

May 

June 

August 

SMITHSOSIAN CONTRIBUTIONS TO BOTANY 

Report on progress to meeting of FNA Program 

Council in Miami, Florida. 

Arrangements made for CAS, DS, JEPS, and uc 

to contribute, and necessary materials supplied. 

cop) Of collector index sent to MO. 

Critique Of Sample catalog of 100 Carex received 

from F. J. Hermann. 

Editing of A and GH data completed, and 

records input to system, 

Data collection at F completed, 

Sew cumulative catalog incorporating A, GH, 

hfO, NY, and US produced, with 537 taxa, 542 

collections, and 776 specimens. 

Four indices in preliminary format generated. 

COBOL paragraphing (concatenating) program 

designed, written, and debugged: GIS/COBOL 

interface programmed and tested. 

1972 

All data returned from F to US for input. 

A11 data returned by C.4S, DS, JEPS, and UC. 

Limited input with magnetic cartridge system 

begun. 

Computer program for listing and tallying taxa 

in Type Register tested. 

Data collection phase essentially brought to 

close. 

Sample catalog produced using COBOL con- 

catenation program. 

Annotations from catalog used by F and from 

catalog used by CAS, DS, JEPS, and UC trans- 

ferred to working copy. 

Editing and final data capture continued apace. 

COBOL concatenating program, including GIS 

interface, tested satisfactorily. 

Xew curnulatit e catalog printed, including 600 

taxa, 607 collections, and 854 specimens. 

Nine indices printed for final field-by-field edit- 

ing. 

lt'riting of introductory sections of this paper 

begun. 

New catalog printed, including 609 taxa, 615 

collections, and 1,050 specimens. 

Xew set of nine indices also produced. 

Editing and updating of file continued. 

Final prepublication edition of Catalog printed, 

along with indices, which included 606 taxa, 

612 collections, and 1,059 specimens. 

Final format of Catalog established after various 

tests with COBOL program. 

Several qpes of computer paper tested. 

Camera-ready copy of present Catalog and in- 

dices produced for Smithsonian Institution 

Press, including same data base as final catalog 

in May. 

Manuscript and camera-ready copy of Catalog 

and indices sent to press.

SUMBER 12 31 

Tlie decision to use Carex for the trial publica- 

tion was made in September 1970, exactly two years 

before the manuscript finally went to press. What 

is obvious from this chronology is that for every 

adtli tional collaborating institution the prepara- 

tion ot a final catalog is delayed at least a few 

months. Under the best of circumstances, turn- 

around time between c)cles of data-collecting, edit- 

ing, and processing quickly add up to days, weeks, 

and months. Clearly, the task of creating a data 

base of this type is limitless ultimately, and definite 

bounds must be established if the dynamic process 

is ever to stand still long enough to yield mean- 

ingful products. Desirable as it would have been, 

therefore, to include many more herbaria in this 

Catalog, tlie line had to be drawn finally. If the 

Catalog had been closed off after the NO, KY, and 

US contributions, as originally planned, it would 

have gone to press a year earlier, in mid-1971. The 

addition of seven more herbaria, including several 

key ones, surely justifies the year’s delay, but it is 

doubtful that further delajs could be justified at 

this time for any other herbaria. We believe that 

it is vitally important now to get the concept of 

tlie Type Register arross to the botanical com- 

munity through the medium of the Cnrex Catalog 

without further delay. 

Use of Carex Catalog 

The Carex Catalog consists of the “Catalog of 

Specimens” and cross-indices to the “Catalog of 

Specimens” by five different fields (descriptors): 

(1) “Author Index,” (2) “Publication-Date Index,” 

(3) “Collector Index,” (4) “Geographic Index,” 

and (5) “Herbarium Index.” (Hereafter the term 

“Catalog” is used for the “Catalog of Specimens” 

proper.) The Catalog is arranged alphabetically by 

taxon and includes the full unit-record for each 

taxon, as it is stored in the computer, except for 

the family and genus names, the data source codes, 

and several file-control dates and numbers. The 

family name and genus name, Cyperaceae and 

Carex, respectively, have been omitted because 

they are the same for all taxa and, printed at the 

top of each entry, would constitute unnecesary 

words that would tend only to hide the key words 

for alphabetization, the epithets. The Catalog is 

alphabetized, therefore, by the specific and infra- 

specific epithets. 

The indices, in addition to providing access to 

the Catalog by other criteria than the taxonomic 

name, represent data files in their own right which 

may serve a user’s purpose without his ever taking 

recourse to the main Catalog. They are independ- 

ent data files because they all include the name of 

the taxon and one or more other fields that place 

the key indexing field in the context of related 

data. The data source code has not been included 

record by record because it is usually 0s; further- 

more, the classification of sources has been in use 

for only a short time and was not used throughout 

the development of the Carex Catalog (see “Data 

Source Code”). 

The Catalog provides citations of original au- 

thors ant1 publications, data on type collections, 

and a list of type specimens known to exist in the 

ten herbaria surveyed-all subordinated to the tax- 

onomic name. No taxa are included that are not 

represented by at least one specimen in at least 

one of the ten herbaria. The organization of the 

data in the paragraphed unit-entry is as follows: 

SPECIFIC EPITHET/RANK/INFRASPECIFIC EPITHET/ 

AUTHOR OF NAME, CITATION OF ORIGINAL PUB- 

LICATION. DATE OF PUBLICATION. 

COUNTRY: STATE OR PROVINCE: COUNTY: LOCAL- 

I n (COMMENT IF ANY) (COLLECTOR, COLLEC- 

TOR’S NUMBER. DATE OF COLLECTION) 

HERBARILX ABBREVIATIOS/SHEET NUMBER/ 

KIND OF TYPE 

HERBARIUM ABBREVIATION [etc., for additional 

specimen] 

COUNTRY [etc., for additional collection] 

HERBARIUM ABBREVIATION [etc.] 

The five indices are all cross-referenced to the 

Catalog by means of the specific epithet instead of 

a page or record number because the unit records 

in the Catalog are alphabetical by epithet, allowing 

for fast look-up. Space did not permit printing in- 

fraspecific epithets in four of the indices; therefore, 

the specific epithet is prefixed by an asterisk (*) if 

the record being cross-referenced is not the species 

itself but one of its infraspecific taxa. This device 

should permit the user to get to the desired entry 

almost as quickly as if the infraspecific epithet had 

been printed, as in the “Herbarium Index.” Con- 

tent and format are more or less self-evident in each 

of the indices. It should be emphasized that these 

are only five of an almost infinite number of pos- 

sible indices. Furthermore, they all were generated 

directly from the exact same data base from which

32 

the Catalog itself was produced and have identical 

data wherever content overlaps. 

The purpose of the “Author Index” is to point 

to all taxa in the Catalog described by a particular 

author or combination of authors, and, therefore, 

the Index is alphabetized primarily by author and 

secondarily by specific epithet. Combinations of 

authors are alphabetized as combinations, not as 

individual authors, owing to present system lim- 

itations, A person interested in a particular taxon 

described by a particular author or author combina- 

tion can learn quickly from the Index whether 

there is an entry in the Catalog for this taxon and 

then go to it. Persons wishing to study an author’s 

descriptions chronologically, by year of publication, 

would want to have this Index resequenced with the 

date rather than the epithet being the secondary 

sort-key. 

The primary sort-key of the “Publication-Date 

Index” is the year of publication, from the oldest 

to the youngest, the actual range being from 1803 

to 1971. Within each year the secondary sort-key 

is the specific epithet, which references an entry in 

the Catalog. This Index should be useful to those 

who are interested in nomenclatural priority in the 

genus or in tracing the historical development of 

knowledge about Cnrex. By itself, the Index, which 

includes the name of the author or author combi- 

nation, is a chronological summary of the botanists 

who have described species in the genus as regis- 

tered in the Catalog. 

The “Collector Index” is in effect a list by col- 

lector of collections cited in the Catalog and 

provides a convenient means of checking any her- 

barium for type material of taxa recorded thus far 

in the Carex file of the Type Register. Under the 

name of each collector or combination of collectors 

are listed the numbers and dates of all collections 

in the Catalog, cross-referenced by the specific epi- 

thets. The name is the primary sort-key, and the 

specific epithet is the secondary sort-key. A varia- 

tion of this Index would include a list of all her- 

baria in which the collector’s type specimens are 

deposited. In other words, the kind of guide to 

the location of types that A. S. Hitchcock and his 

colleagues were trying to compile directly in the 

1930s (see “Introduction”) can be produced as a 

by-product of the Type Register data base. Some 

interesting observations can be made by compar- 

SMITHSONIAN CONTRIBUTIOXS TO BOTANY 

ing the “Collector Index” with the “Author In- 

dex.” It becomes clear at once, for example, that 

while some authors collected almost as many new 

taxa as they described, others described many more 

than they collected. Further examination indicates 

that this difference may reflect the difference be- 

tween a floristic taxonomist like M. L. Fernald, who 

tended to describe his own species, and a mono- 

graphic worker like K. K. Mackenzie, who described 

taxa from among specimens of many collectors 

amassed for a study of the genus. 

The “Geographic Index” provides access to the 

collections of the Catalog primarily by country of 

origin and secondarily by state, province, or equiva- 

lent. The specific epithet, which is the cross- 

reference to the Catalog, is the tertiary sort-key. The 

value of this Index will be obvious at once to 

floristic workers who wish to know which taxa in 

their region are typified by specimens collected 

within the region, e.g., within the State of Cali- 

fornia. 

The “Herbarium Index,” alphabetized in the 

first instance by institution, lists alphabetically by 

specific and infraspecific epithets under each of the 

ten institutions the taxa in the Catalog for which 

they hold type specimens. The herbarium sheet 

number, if there is one, and the kind of type are 

listed also. The sheet number is the tertiary sort- 

key. Many of the type designations have not been 

validated, and the problems of validating kind of 

type have been discussed at length earlier in this 

paper (pp. 8 and 19-21). For this reason many 

of the type specimens registered still carry the origi- 

nal designation of the herbarium submitting the 

data. 

The following informal or archaic terms, which 

appear in the Catalag and in the “Herbarium In- 

dex” but which are not sanctioned by the ZCBAI, 

require brief definition of their meaning or ap- 

parent meaning in the Catalog. It should be 

pointed out, however, that the presence of a cor- 

rect term (e.g., holotype, isotype) is no guarantee 

that it has been used correctly. A large number of 

the designations have been validated, and where 

the terminology seems to be used consistently 

within a collection, e.g., where one holotype and 

one or more isotypes are indicated, it can be as- 

sumed that the designations were validated.

SUMBER 12 33 

rotype Presumptive syntype or isosyntype, but may 

be isotype or paratype, if a type at all. 

type 

Presumptive holotype, but may be isotype, 

syntype, isosyntype, or paratype, if a type at 

all. 

type collection Term used in a proper sense for a specimen 

of a single collection cited by original authoi 

without designating a holotype-in this sense, 

a syntype or isosyntype according to ICBN 

(see pp. 19-21); otherwise term flags a presumptive 

syntype, isosyntype, paratype, or 

even isotype, if a type at all. 

type fragment Presumably fragment of the holotype, but 

may be fragment of isotype, syntype, isosyntype, 

paratype, or other kind of type, if of 

type at all. 

type materiul Presumptive type specimen of some kindcatchall 

term. 

The Catalog and Indices were printed by com- 

puter directly from the data base, and not a single 

change has been made. What is presented here is 

exactly what was stored on the machine file as of 

16 June 1972, with the exception of punctuation 

between fields which may have been added in the 

process of concatenation. Because the system pro- 

vided only for printing in upper case letters, possi- 

bilities for variation of typography were limited. 

Boldfacing by overprinting the same words two or 

more times was used for the specific and infraspe- 

cific epithets, and in the process it was necessary 

also to boldface rank designators connecting epi- 

thets. Thus the taxon names stand out from the 

rest of the text and facilitate searching the Cata- 

log. Insofar as practical, the standard conventions 

of punctuation in nomenclatural literature were 

used. 

Errata 

Several errors were detected in the Catalog after 

the camera-ready copy had been produced. By rec- 

ord number in the Catalog, these are: 

No. 2. ABLATA BAILEY, L. H. The citation following 

the author’s name is to the place where the neotype is 

designated, because the original publication did not designate 

a type specimen, and the citation of Bailey’s original publication 

is cited as a parenthetic remark at the end of the 

geographic locality. To be consistent with the rest of the 

Catalog, however, where the citation after the author is 

( I ~ N \ S the citation of the original publication, the editors 

should have reversed the two citations in this record. As a 

result of this editorial error, Bailey in the “Author Index” and 

“Publication-Date Index” appears to have described C. 

ablata in 1935, when Mackenzie designated the neotype, 

rather than in 1888. This error does point up the problem of 

dealing with neotypes, however, and a certain logic can be 

advanced for either way of handling the two citations. 

No. 99. CHIHUAHUAENSIS. Spell CHIHUAHUENSIS, 

omitting second “A.” 

No. 418. PIRCHINCHENSIS. Spell PICHINCHENSIS, 

omitting “R.” 

No. 424. PLUVICA. Spell PLUVIA, omitting “C.” 

So. 446. PURPUREOVAGINATA. Spell PURPUREO- 

VAGINATA, inserting hyphen. 

No. 448. PYCNOTHYSOS. Spell PYCNOTHYRSOS, inserting 

“R.” 

No. 544. TENERA VAR. RICH11 FERNALD, M. L. At 

the end of the geographic locality, MIDDLESEX FALLS 

should read MIDDLESEX FELLS. 

No. 549. TERRAE-NOVAE FERNALD, M. L. The hyphen 

should be removed from the collector’s name GIL- 

BERT-JR., F.A. The practice of joining Jr. to the end of the 

collector’s or author’s last name by a hyphen in this manner 

was required by the specifications of the pilot processing 

s\steni, but this requirement no longer obtains. 

Note: Changes in spelling of specific epithets also 

apply wherever these epithets have appeared in 

the indices. 

Statistical Summary of Catalog 

606 taxa (species, subspecies - SSP, varieties - VAR, 

forms-FOR, nothomorphs-NM.) 

612 collections 

1,059 specimens (sheets)

CATALOG OF SPECIMENS 

10 ABDITA BICKNELL9E.P.9 BULL. TORREY BOT. CLUB 35:492. 19080 

USA: NEW YORK: LONG ISLAND9 RICHMOND HILL (BICKNELLTE.P*T -0-0 

11 MAY 19@4) 

NY TYPE 

-A- 

2. ABLATA BAILEYTL.H.T No AMER. FL. 18:314. 1935. 

CANADA: BRITISH COLUMBIA: VANCOUVER ISLAND, MOUNT MARK; ALTO 

250C FT.; (ORIG. PUB.: BOT. GAZ. 13:82. 1888.1 (MACOUNTJOHNT 

134@1. 26 JUL 1887) 

GH I SONEOTY P E 

NY I SONEOTYPE 

30 ABORIGINUM JONES9M.E.9 BULL. MONTANA STATE UNIV.9 BIOL. SER. 15:69. 

1910 

USA: IDAHO: ADAMS C O O : INDIAN VALLEY: ALTO 2300 FT. 

(JONES,M.E.T --- 12 JUL 1899) 

CAS 242617 ISOTYPE 

DS 149709 I SOTYPE 

NY ISOTYPE 

4. ABRAHSII MACKENZIE,K~K.T BULL. TORREY B0To CLUB 36:482. 1909. 

USA: CALIFORNIA: SAN BERNARDINO C0.t SAN BERNARDINO MOUNTAINST 

BEAR VALLEY (ABRAMStL., 28160 31 JUL 1902) 

DS 55317 ISOTYPE 

F 186491 I SOTYPE 

MY HOLOTYPE 

50 ABRUPTA MACKENZIE,K*K.9 BULL. TORREY 601. CLUB 43:6180 1916. 

USA: CALIFORNIA: BUTTE Coo: STIRLING CITY; ALTO 3000 FT. 

(HELLERTA*AoT 10820. 07 JUN 1913) 


NY HOLOTY PE 

60 ABSCONDITA VARo ROSTELLATA FERNALD9M.L.9 RHODORA, 4423860 1942. 

USA: VIRGIYIA: ISLE OF WIGHT COO: LEE'S MILL (FERNALD9M.L. AND 

LONGIB., 120120 08 JUN 1940) 

GH I SOTYPE 

YO TYPE COLLECTION 

NY 

us 2003161 

TYPE COLLECTION 


70 X ABSCONDITIFORHIS FERNACD9M.L. , RHODORA 44:387. 19420 

USA: VIRGINIA: SUSSEX C O O : NOTTOWAY RIVER9 HUSKE (FERNALDfMoLa 

AND LONGvB.9 12969. 13 JUN 1941) 

GH HOLOTY PE 

GH 

MO 1306480 

I SOTYP E 

ISOTYPE 

34

NUMBER 12 35 

9. 

10 

11. 

12. 

13 

14. 

15- 

16. 

YY I SOTYPE 

us 200 3299 I SOTYP E 

ACCEDENS HOLMTH.T.,T AMER. J. SCI. SER.49 16:457. 1903. 

USA: OREGON: MULTNOMAH COO: SAUVIE ISLAND (COLUMBIA RIVER AT 

MOUTH OF WILLAMETTE RIVER) (HOWELL9T.J. T --- -- MAY 1880) 

GH SY NTY P E 

YO SY NTY P E 

ACROPHILA BLAKETS~T.~ J. ARNOLD ARBOR. 28:114. 1947. 

INDONESIA: WEST NEW GUINEA: 

NEW GUINEA") (BRASSTL.J.~ 

LAKE HABBEMA; (COUNTRY AS "DUTCH 

9515. -- AUG 1938) 

A I SOTYP E 

ACUTA VAR, PALLIOA BOOTT9F.p ILL. GENUS CAREX 4:1669 PL.554. 1867. 

USA: OREGON: "FORT COLVILLE TO ROCKY MOUNTAINS9 WEST KOOTENAY" 

( L Y AL L T DA V I D T --- . -- --- 1861) 

GH TYPE COLLECTION 

ACUTINA BAILEYTLoH*T M E M O TORREY BOT. CLUB 1:52. 1889. 

USA: OREGON: DESCHUTES RIVER (HOWELLTT.J.T 935. 09 MAY 1885) 

F 206585 TYPE MATERIAL 

GH ISOTYPE 

NY TYPE COLLECTION 

us 25164 TYPE COLLECTION 


ACUTINELLA YACKENZIETK.K* T N. AMER. FL. 18:407. 1935. 

USA: OREGON: -- (HENDERSONTL.F.~ 13. -- --- 1883 1 

us 27286 HOLOTYPE 

AOUSTA VAR. GLOMERATA OLNEY 9 s .T EX BAILEY TL oHo 9 BOT. GAZ. 9: 139. 

1884. 

CANADA: NEW BRUNSWICK: KENT COO: SALMON RIVER (FOWLER9J.q __ --- 

---. 

1872) 

GH SYNTYPE 

AENEA FERNALD9M.L. 9 PROC. AMER. ACAD. ARTS 37:480. 1902. 

USA: NEW HAMPSHIRE: GRAFTON COO 2 FRANCONTAT FOREST HILLS HOUSE 

(FAXONTE. AND FAXONTC.E.~ --- 23 JUN 18881 

GH SYNTYPE 

AEQUA CLARKETC.B.T BULL. MISC. INFORM. ADD.SER.8:86. 1908. 

USA: CALIFORNIA: SAN MATE0 CO,: SAY MATE09 CRYSTAL SPRINGS LAKE 

(BAKERvC.F.9 811. 10 MAY 1902) 


NY TYPE COLLECT ION 

AESTIVACIFORMIS MACKENZIETK~K. 9 

1910. 

BULL. TORREY BOT. CLUB 37:238. 

USA: NEH JERSEY: PASSAIC COO: GREENWOOD LAKE (MACKENZIE,K.K. T 

2676. 23 JUN 1907) 

GH I SOTYPE

36 

17. 

18. 

19. 

20 0 

21 0 

22 0 

23. 

24 . 

25. 

YO TYPE MATERIAL 

SMITHSONIAN CONTRIBUTIONS TO BOTANY 

AESTIVALIS CURTIS9M.A. EX GRAY9A.i AMER. J. SCI. ARTS SER.1, 42:280 

1841. 

USA: NORTH CAROLINA: -- ( CURTISTM.A.T --- -- JUL 1841) 


NY TYPE MATERIAL 

AGGLOMERATA MACKENZIE,K.K.r BULL. TORREY BOT. CLUB 33:442. 1906. 

USA: MISSOURI: JACKSON COO: COURTNEY (BUSHTB.F.T 1718. 

25 MAY 1902) 



AGGREGATA MACKENZIETK~KOT BULL. TORREY BOT. CLUB 37:246. 1910. 

USA: MISSOURI: JACKSON COO: COURTNEY (BUSHTBOF~T 1718. 

25 MAY 1902) 

NY TYPE 

AGROSTOIDES YACKENZIE9K.K.9 BULL. TORREY BOT. CLUB 34:607. 1907. 

USA: NEW YEXICO: SOCORRO COO: LUNA; 

--- 28 JUL 1900) 

us 617798 TYPE 


ALTO 6500 FT. (WOOTONTE.O.T 

ALATA TORREYqJ.9 ANN. LYCEUM NATO HIST. NEW YORK 3:396. 1836. 

USA: NORTH CAROLINA: CRAVEN COO: NEW BERN (CROOMTH.B.T -- --- 

---. 

1834) 

NY SYNTYP E 

ALATA VARo FERRUGINEA FERNALO~Y.L.T PROC. AMER. ACAD. ARTS 

37:477T PL.2. 1902. 

USA: OHIO: 0- (SULLIVANTTW.S.T --- . ---I 

GH HOLOTY P E 

AtBIDA BAILEY7L.H.r MEMO TORREY BOT. CLUB 1:9. 1889. 

USA: CALIFORNIA: SANTA ROSA CREEK (BIGELOW9J.M. 9 -0-0 

-- -*- 1853-1854) 


ALBO-NIGRA MACKENZIE9K.K. IN RYDBERG,P.A. 9 FL. ROCKY MOUNT. 

1379 1060. 1917. 

USA: WYOMING: PARK C O O : NEEDLE MOUNTAIN (CARY9M.p 613. 

11 JUL 19101 


ALMA BAILEY,L.H.r MEMO TORREY ROT. CLUB 1:50. 1889. 

USA: CALIFORNIA: -- (PARRYTC~C. AND LEMMON*J.G.r 396. 

-- --- 1876 1 

CAS 

NY 

497554 ISOTYPE 

I SOTY P E 

26, ALOPECOIDEA TUCKERMANTE~T ENUM. CARIC. 18. 1843.

NUMBER 12 57 

21 . 

28- 

29 . 

30. 

31. 

32 

33. 

34. 

35. 

USA: NEW 

F 

f 

F 

F 

F 

F 

GH 

MY 

YORK: YATES COO: PENN YAN (SARTWELL,H*P*, --- . ---I 

32699 I SOTYPE 


56916 ISOTYPE 




I SOTYPE 

I SOTYP E 

ALOPECOIDEA VAR. SPARSI-SPICATA DEWEYtCor AMER. J. SCI. ARTS 

SER.2, 8:350. 

USA: MICHIGAN: 

1849. 

MACOMB COO: WASHINGTON (COOLEY,D.q 74. ---I 

CAS 553879 I SOTYP E 

GH I SOTYPE 

MY I SOTYPE 

AMPHIBOLA STEUDELIE~G.~ SYN. PL. GLUM. 2:234. 1855. 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMOND,T., 437. 

---) 


AHPHIBOLA VAR. TURGIDA FERNALDTM.L*~ RHODORA 44:311. 1942. 

USA: NEW YORK: TOYPKINS C O O : ITHACA (WIEGANDTK.M. AND 

THOMASTC.C.T 1915. 15 JUN 1914) 

GH HOLOTY PE 

AMPLISQUAHA HERMANN,F*J., RHODORA 57:158. 1955. 

USA: GEORGIA: GILMER COO: CHATSWORTH (PYRON,J.H. AND 

MCVAUGH~R.T 2951. 15 MAY 1938) 

us 2231424 TYPE 

ANGUSTIOR MACKENZIETK~K. I N RYDBERG,P.A.q FL. ROCKY MOUNT. 

124, 1060. 1917. 

USA: DISTRICT OF COLUMBIA: -- (STEELE,EoS.r --* . -- --- 1896) 


ANGUSTIOR VARm GRACILENTA CLAUSEN9R.T. AND WAHLTH.A., RHODORA 

41230. 1939. 

USA: PENNSYLVANIA: CENTRE COO: INGLEBY (CLAUSEN,R*T. AND 

WAHLqH.4.9 2532. 06 JUN 19371 

GH TYPE 

ANTHOXANTHERA PRESL,K.B.r REL. HAENK. 3:203. 1828. 

USA: ALASKA: NUTKA SOUNO ("SINUS NUTKA") (HAENKEqT. T --- . ---I 


APERTA BOOTT,F. I N HOOKERTW.J.T FL. BORo-AYER. 2:218. 

1839 ( "1 840") 

USA: WASHINGTON: COLUMBIA RIVER (SCOULER, J. 9 -0- . ---I 

GH SYNTYPE 

APERTA VAR. UMBROSA KUKENTHAL,G.T REPERT. SP. NOV. REGNI VEG.


360 

37. 

38. 

39 . 

40 0 

41 0 

42 

43. 

26: 254. 1929. 

USA: WASHINGTON: KLICKITAT COO: BINGEN (SUKSD0RFtW.N.p 12347. 

---) 

CAS 242962 I SOTYPE 

APERTA VAR. VIRIOANS KUKENTHALt G o t REPERT SP. NOV. REGNI VEG. 

26 : 254. 192 9. 

USA: WASHINGTON: KLICKITAT C O O : BINGEN (SUKSDORF9W.N.9 12348. 

15 SEP-23 OCT 1927) 

CAS 242961 SYNTYPE 

USA: WASHINGTON: KLICKITAT C0.Z BINGEN 4 SUKSD0RFpW.Net 12359. 

23 OCT 1927) 


APODA CL0KEYtI.W.t AMER. J. SCI. SERo5t 3:88t PL.2. 1922. 

USA: IDAHO: CUSTER COO: MACKAY (NELSONgA. AND MACBR1DEtJ.F. 9 

1533. 01 AUG 1911) 

NY I SOTYPE 

uc 90 543 9 HOLOTYP E 

APOOOSTACHYA OHW1,J.t JAP. J. BOT. 7:188. 1934. 

TAIWAN: --: MOUNT NANKO-TAISAN (0HWItJ.t 4182. -- JUL 1933) 


AQUATILIS WAHLENBERGtG.9 KONGL. VETENSK. ACAD. NYA HANDL. 24:165. 

1803 

USA: NEW YORK: SENECA CO.: JUNIUS (SARTWELLtHoPot 56. ---I 


AQUATILIS VAR. SUBSTRICTA KUKENTHALtG. IN ENGLERtH.G.A.9 PFLANZENR. 

49 FAM. 20: 309. 1909. 

USA: NEW YORK: SENECA COO: JUNIUS (SARTWELL,Ho\P.t 56. ---I 

CAS 55401 9 I SOTYPE 

GH I SOTYPE 


ARAPAHOENSIS CLOKEYt 1.W.t RHODORA 21:83. 1919. 

U'SA: COLORADO: BOULOER COO: MOUNT ARAPAHOE; ALTO 11700 FT. 

(CLOKEYtI.W.9 3227. 29 JUL 1918) 

C AS 102030 ISOTYPE 


GH I SOTYPE 

NY I S OTY PE 

uc 905436 HOLOTYPE 

ARCTAEFORMIS YACKENZ1EtK.K. 9 No AMER. FL. 18:97. 1931. 

CANADA: BRITISH COLUMBIA: ELGIN (HENRY9J.K.t 9152. 

C4 JUN 1915) 

NY TYPE 

ARCTICA DEHEYtC., AMER. J. SCI. ARTS SER.1, 27:239. 1835. 

CANADA: SASKATCHEWAN: CARLTON HOUSE (52 51". 9 106 13'W. 1 

(RICHARDSON9J.9 --- . --- b

NUMBER 12 39 

44 . 

45 0 

46 

47. 

480 

49 

50. 

51. 

YY TYPE COLLECT ION 

---. 

ARGYRANTHA TUCKERMANtE. EX DEWEYTC., AMER. J. SCI. ARTS 

SER.29 29:346. 1860. 

USA: 

---) 

MASSACHUSETTS: HAYPSHIRE COO: AMHERST ( TUCKERMANTE. 9 

GH HOLOTYP E 

GH I SOT YP E 

ARISTATA VARo LONGO~LANCEOLATA DEWEYIC.T AMER J SCI ARTS 

SER.2, 18:102. 1854. 

USA: NEBRASKA: BAD LANDS ("MAUVAIS TERRES") (HAYDENTF.V., 

-- --- 1853) 

---. 

GH HOLOTY PE 

ARSENII KUKENTHALTG. T REPERT, SP. NOV. REGNI VEG. 8:3266 1910. 

MEXICO: MICHOACAN: YORELIA (ARSENETG. (FRERE) T 3054. 

16 JUL 1909) 

GH I SOTYPE 

NY 

us 1030011 

I SOTYP E 


ARTITECTA VAR. SUBTILIROSTRIS HERMANNtFoJor RHODORA 40:79. 1938. 

USA: INDIANA: VERMILLION COO: CLINTON (DEAM,C.C.T 54764. 

05 MAY 1934) 

GH HOLOTY PE 

ASSINIBOINENSIS BOOTTTW.T BOT. GAZ. 9291. 1884. 

CANADA: MANITOBA: ASSINIBOINE RAPIDS (YACOUN9JOHNt 52. 

14 JUN 18791 

GH SYNTYP E 

ATHABASCENSIS HERMANNTF.J.v LEAFL. W. BOT. 8:111. 1957. 

CANADA: ALBERTA: JASPER NATIONAL PARK, ATHABASCA RIVER, 

ATHABASCA FALLS ( HERMANNTF. J. T 13498. 28 AUG 1956 1 


ATHROSTACHYA OCNEYTS.T. 

1868. 

IN GRAYvA.9 PROC. AMER. ACAD. ARTS 7:393. 

USA: CALIFORNIA: MARIPOSA COO: YOSEMITE NATIONAL PARK, 

YOSEYITE VALLEY (BOLANDERTH~N~T 6213. 17 JUN 18631 

NY 

us 319165 

SY NTY PE 

SYNTYPE 


YOSEYITE VALLEY (BREUER,W.H., 

MO SY NTYP E 

1650. 17 JUN 1863) 

NY SYNTYPE 

ATRACTOOES HERMANNTF. J.9 J. WASH. ACAD. SCI. 40:283. 1950. 

MEXICO: CHIAPAS: COMITAN (SHARPTA.J.T 45450. 29 APR 1945) 

NY ISOTYPE 

us 2133192 TYPE

40 

520 

53. 

54 0 

55 0 

56 

57, 

58, 

59. 

60- 


ATROFUSCA VAR. DECOLORATA P0RSILDtA.E.t SARGENTIA 4:20. 1943. 

CANADA: NORTHWEST TERRITORIES: MACKENZ IE DISTRICT: GREAT BEAR 

LAKE9 CAPE MCDONNELL (P0RSILDtA.E. AND PORSILDt ROT., 5120. 

02 AUG 1928) 

us 2096188 ISOTYPE 

ATROSQUAUA MACKENZ1EtKoK.t PROC. BIOL. SOC. WASH. 25:51. 1912. 

CANADA: ALBERTA: SMOKY RIVER (H0LLISTERqN.t 14. 05 AUG 1911) 

NY 

us 622651 

I SOTYP E 

HOLOTY PE 

AUREA VAR, ANDROGYNA OLNEY9S.T. IN WATS0NtS.t 

BOT. U.S. GEOL. EXPLOR. 40TH PAR. 371. 1871. 

USA: PENNSYLVANIA: ERIE COO: ERIE9 PRESQUE ISLE (PENINSULA) 

(GAR6ERtAoP.t ---o 09 JUN 1869) 

NY I SOTYPE 

AUROLENSIS STEUDEL, E.G.9 SYN. PL. GLUY. 2:223. 1855. 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMM0NDtT.t 

-s. --- 1832 1 

431. 


AUSTRO-CAROLINXANA BA1LEYtL.H.t BULL. TORREY BOT. CLUB 20:428. 

1893. 

USA: SOUTH CAROLINA: __- . ---I 

PICKENS COO: TABLE MOUNTAIN (6UCKLEYtS.B. t 

YO 1834152 HOLOTYPE 

AUSTROMONfANA PAR1SHtS.B.r BULL. S o CALIF. ACAD. SCI. 4:108t PL.15. 

1905 

USA: CALIFORNIA: SAN BERNARDINO CO.: MILL CREEK FALLS; ALTO 

6000 FT. (PARISHtS.B*t 2485. 03 JUL 1892) 

DS 489410 HOLOTYPE 

AUTUUNALIS MACKENZIE,K.K. 9 No AMER. FL. 18:66. 1931. 

MEXICO: MEXICO: FLOR DE MARIA (PR1NGLEtC.G.t 4275. 

03 OCT 1892) 

F 264169 ISOTYPE 

YO I SOTYPE 


AZTECICA MACKENZ1EtK.K.t No AMER. FL. 18:229. 1935. 

MEXICO: OAXACA! SIERRA DE SAN FELIPE; ALTO 2300 M. 

(PR1VGLEtC.G.t 4839. 19 AUG 1894) 



AZUAYAE STEYERMARK, J.A. 9 PHYTOLOGIA 9:337. 1964. 

ECUADOR: AZUAY: TOREADOR (STEYERMARK9J.A.t 53105. 15 JUN 1943) 


NY I SOTYPE 

us 1933437 ISOTYPE

NUMBER 12 41 

- B- 

61 

62 

63 

64 

65 

66 

67 

680 

BACKANA DEWEYTC., AMER. J. SCI. ARTS SER.19 29:250. 1836. 

CANADA: SASKATCHEWAN: CARLTON HOUSE (52 51'N.r 106 13'W.l 

(RICHARDSONTJIT 417. ---I 


BACKII BOOTTTF. IN HOOKER,W.J., FL. BORorAYER. 2:210. 

1839 ( "1840") 

CANADA: SASKATCHEWAN: CARLTON HOUSE ( 52 51". q 106 13'W. 1 

(RICHAROSONqJ.9 --- . ---I 

GH SYNTYPE 

NY SYNTYPE 

BALTZELLII CH4PMANqA.W. EX DEWEYtC., AMER. J. SCI. ARTS 

SER.2, 3:335. 1847. 

USA: FLORIDA: BEAR CREEK (CHAPMAN P A. We 9 --- . ---I 

NY COTY P E 

BAHBUSETORUH MERR1LLqE.D.r PHILIPP. J. SCI. 13:132. 1918. 

CHINAt KWANGTUNG: LOH-FAU-SHAN (MOUNTAIN) (MERRILLqE.D., 10985. 

12 AUG 1917) 


BANKS11 BOOTTqF.9 TRANS. LINN. SOC. LONDON 20:119. 1846. 

ARGENTINA: T IERSA DEL FUEGO (TERRITORY) : TIERRA DEL FUEGO 

(BANKSqJ. AND SOLANDERTD.C.T --- . -- --- 1769) 

YO 1611724 SYNTYPE 

us 1232938 SYNTYPE 

BARBARA€ DEWEYTC. IN TORREYqJ. IN EYORY,W.H. T 

REP. U.S. MEX. BOUND. SURV.9 BOT. 2(1):231. 1859. 

USA: CALIFORNIA: LOS ANGELES COO : SANTA BARBARA (PARRYqCoC. T 

-_- . -- --- 1850 1 

GH HOLOTY PE 

NY I SOTYPE 

BARRATTII SCHWEINIT2,L.D. AND TORREYqJ. T 

ANN. LYCEUM NATO HIST. NEW YORK 1:361. 1824. 

USA: NEW JERSEY: CAPE MAY COO: CAPE MAY (COLLINSTZ.T --- . ---I 


BARTLETTII O'NEILLTH.T.T PUBL. CARNEGIE INST. WASH. 522:255. 1940. 

BRITISH HONDURAS: CAYO: MOUNT PINE RIDGE (BARTLETTTH.H.T 

11718A. 24 FEB 1931) 


GH I SOTYPE 

NY TYPE 

690 BAYARDI FERNALD1M.L .q RHODORA 44371. 1942.


70. 

71 

72. 

13 . 

14 0 

75 

76 

77. 

78. 

USA: VIRGINIA: SOUTHAMPTON COO: DREWRYVILLE (FERNALD,M.L.; 

LONGTB. AND SMART,R.F.p 5677. 22 JUN 1936) 

GH H01 OTY PE 

BILTHOREANA MACKENZIE,K.K. T BULL. TORREY BOT. CLUB 37:234. 1910. 

USA: NORTH CAROLINA: SATULA MOUNTAIN (---T 2688. 25 MAY 1897) 

GH I SOTYP E 

YY TYPE 

BIPARTfTA VAR. 

1963. 

AUSTROMONTANA HERMANNTF.J.T LEAFL. W. BOT. 10:16. 

USA: COLORADO: BOULDER COO: NEDERLANDV NOWIT RIDGE; ALTO 11500 

FT. (HERMANNIF~J~T 17059. 15 AUG 1961) 


BONANZENSIS BRITTON,N.L.9 BULL. NEW YORK BOT. GARD. 2:160. 1901. 

CANADA: YUKON TERRITORY: BONANZA RIVER ( WILLIAMStR. So T ---. 

18 JUN 1899) 

NY TYPE 

BONPLAND!! VAR. MINOR BOOTTTF. IN GRAYTA.~ 

PROC. ACAD. NATO SCI PHILADELPHIA 1863:77. 1863. 

USA: COLORADO: ROCKY MOUNTAINS; LAT. 39-41 No (HALLqE. AND 

HARBOURTJ.P.T 591. -- --- 1862) 



GH HOLOTYPE 

YO ISOTYPE 

BRACHYPODA HOLMTH.T.T AMER. J. SCI. SER.4, 20:302. 1905. 

USA: OREGON: KLAMATH C0.t CRATER LAKE NATIONAL PARK, CATHEDRAL 

SPRING (COVILLETFOV-T 1455. 11 SEP 1962) 


BRAINERDII MACKENZIE,K.K. T BULL. TORREY BOT. CLUB 40:534. 1913. 

USA: CALIFORNIA: EL DORADO COO: SIERRA NEVADA RANGE, SLIPPERY 

FORD (BRAINERDTE., 1210 19 JUL 1897) 


BREVrCAULIS YACKENZIETK.K.T BULL. TORREY BOT. CLUB 40:547. 1913. 

USA: OREGON: LINCOLN C0.2 

-- MAY 1886) 

YAQUINA BAY (HDWELLTTOJO~ 2994. 

h(Y TYPE COLLECTION 

BREVIS BLAKEfS.T.9 J. ARNOLD ARBOR. 28:lll. 1947. 

PAPUA 4ND NEW GUINEA: PAPUA (TERRITORY): OWEN STANLEY RANGEt 

MOUNT ALBERT EDWARD: (COUNTRY AS "BRITISH NEW GUINEA") 

(BRASS,L.J., 4418. -- MAY-JUL 1933) 

A I SOTYPE 

BREVISQUAMA MACKENZIETK.K.T BULL. TORREY BOT. CLUB 34:152. 19070 

USA: WYOMING: SWEETWATER COO: RED DESERT9 ORENDO BUTTE 

(NELSON9A.q 7124. 11 JUN 1900)

NUMBER 12 43 

19 

80 0 

81 0 

82. 

83 

86 

NY HOLOTYPE 

BREWER1 B00fT~F.q ILL. GENUS CAREX 4:142~ PL.455. 1867. 

USA: CALIFORNIA: SISKIYOU COO: MOUNT SHASTA (BREWER,W.H.* 1422. 

-- --- 1863) 

GH I SOTYPE 

BRONGNIARTI! VARo OENSA BAILEY,L.H., PROC. AMER. ACAD. ARTS 22:137. 

1886 ( "1887") 

USA: CALIFORNIA: MARK WEST CREEK (BIGELOW,J.M., -- --- 

---. 

1 8 53 -1 854 1 

NY SYNTYPE 

BRUNNEA VARo SUBTEIOGYNA KUKENTHALTG. T REPERT. SPo NOV. REGNI VEG. 

8:8. 1910. 

PHILIPPINES: BENGUET: LUZON (ISLAND), MOUNT PULOG 

(MERRILL,E.D.q 6505. -- MAY 1909) 


BUCKLEVI DEWEYqC., AMER. J. SCI. ARTS SER.1, 482143. 1845. 

USA: NORTH CAROLINA: MITCHELL C O O : ROAN MOUNTAIN (BUCKLEY,S.B., 

--- . ---I 


BULBOSTYLIS YACKENZIE9K.K. T BULL. TORREY BOT. CLUB 42:617. 1915. 

USA: TEXAS: TARRANT COO: FORT WCIRTH (RUTHqA., 360. 

12 APR 1913) 

YO 71 01 12 TYPE COLL ECT ION 


BURCHELLIANA BOECKELERqJ.O., LINNAEA 412234. 1877. 

SOUTH AFRICA: --: -- (BURCHELLqW.J.9 1911. ---I 

GH I SOTYPE 

BUSH11 MACKENZ1EqK.K. 9 BULL. TORREY BOT. CLUB 37:241. 1910. 

USA: AQKANSAS: HEYPSTEAD COO: FULTON (BUSHqBeF. T 2514. 

30 APR 1905) 


- C- 

CAESARIENSIS YACKENZ1EtKoK.q No AYER. FL. 18:440. 1935. 

USA: NEW JERSEY: CAMDEN C O O : LAUREL SPRINGS (LONGTB.~ F23212. 

15 JUN 1920) 

GH 1 SOTY PE 

NY TYPE 

CALIFORNICA BA1LEYqL.H.q MEMO TORREY BOT. CLUB 1:9. 1889. 

USA: CALIFORNIA: MENOOCINO COO: YENDOCINO CITY (BOLANDER9H.N. 9 

4741. 01 MAY 1866) 

CAS 383776 TYPE COLLECTION


88 

89. 

90 

91 

92 

93 (. 

94 0 

95 0 

96 

DS 49734 TYPE COLLECTION 



us 29741 TYPE COLLECT1 ON 


CAMPLYOCARPA HOLM,H.T., AMER. J. SCI. SER.4, 20:304. 1905. 

USA: OREGON: KLAMATH COO: CRATER LAKE NATIONAL PARK, CATHEDRAL 

SPRING (COVILLE,F.V.q 1457. 11 SEP 1902) 


CAMPYLOCARPA SSPo AFFlNIS MAGUIRETB. AND HOLMGREN,A.H.( 

LEAFL. W. BOT. 42262. 1946. 

USA: UTAH: JUAR C O O : DEEP CREEK RANGE, INDIAN FARM CREEK 

(MAGUIREvB. AND HOLMGREN9A.H. t 21947. 16 JUL 1943) 


NY HOLOTY PE 

us 1885701 I SOTYPE 

CANESCENS VAR. DISJUNCTA FERNALD,Y.L. 9 PROC. AMER. ACAD. ARTS 

37:4889 PL.5. 1902. 

CANADA: NEW RRUNSWICK: VICTORIA CO. : SERPENTINE RIVER 

(HAYIG~U.~ 84. 24 JUL 1900) 

GH PARATY PE 

CANESCENS VAR. DUBIA BAILEYIL~H. t BOT. GAZ. 9:119. 1884. 

USA: UTAH: BEAR RIVER CANYON; ALTO 10000 FT. ( WATSONvS. T 1231A. 

-- AUG 18691 

NY I SUTYPE 

CANESCENS VARo SPHAEROSTACHYA TUCKERMAN,E., ENUM. CARIC. 19. 1843. 

USA: --: NEW ENGLAND (---, --- . -- --- 1843) 

GH I SOTYPE 


CAREYANA TORREYvJ. EX DEWEY,C.r AYER. J. SCI. ARTS SER.1, 30:60. 

1836. 

USA: NEW YORK: CAYUGA COO: AUBURN (CAREY,J.* --- -- MAY 1832) 

NY HOLOTY P E 

CAROLINIANA BUCKLEY,S.B.v AMER. J. SCI. ARTS SER.1, 45:173. 1843. 

USA: SDUTH CAROLINA: PICKENS COO: TABLE MOUNTAIN (BUCKLEY,S.B., 

_-- . ---I 

GH TY PE C OLL ECT I ON 


CEPHALOPHORA VAR. MAXIMA DEWEYqC., AMER. J. SCI. ARTS SER.19 43:92. 

1842. 

USA: NEW YORK: YATES COO: PENN YAN (SARTWELL,H.Pe, --- . ---I 

GH TYPE 

CHALCIOLEPIS HOLM,H.T.( AMER. J. SCI. SER.4, 16:21,28. 1903. 

USA: COLORADO: MI NERAL COO : PAGOS A PEAK BAKER, C O F 9 226.

NUMBER 12 45 

-- AUG 1899) 

GH SYNTYPE 

YO SYNTYPE 

NY SYNTYPE 


97. CHAPMAN1 SARTWELL9H.P. EX DEWEYqC.9 AMER. J. SCI. ARTS 

SER.29 19:254. 1855. 

USA: FLORIDA: *- (CHAPMAN,A.W., 113. ---I 



98. CHIAPENSIS HERMANNTF.J.T BRITTONIA 19:68. 1967. 

MEXICO: CHIAPAS: CHAYULA (BREEDLOVE,D.E. 9 6714. 30 JUL 1964) 


NY I SOTYPE 


99. CHIHUAHUAENSIS MACKENZIE,K.K., BULL. TURREY BOT. CLUB 35:265. 

1908. 

MEXICO: CHIHUAHUA: PUERTA DE ST. DIEGO; 

(HARTMANTC-V., 620. 12 APR 1891) 

ALTO 6500 FT. 


NY 

us 306281 

HOLOTYPE 

ISOTYPE 

100. CHIKUNGANA BAILEY,L.H., GENTES HERB. 1:13. 1920. 

CHINA: HUPEH AND YONAN: CHIKUNGSHAN ( BAILEYTL.H.T 

13 JUN 1917) 

---. 

NY TYPE 

101. CILIARIS FERNALD9M.L. 

MEXICO: HIDALGO: 

26 AUG 1905) 

9 PROC. AMER. ACAD. ARTS 43:61. 1907. 

LENA STATION (PRINGLETC~G~T 10039. 


CAS 

F 

445943 

202021 

ISOTYPE 

I SOTYPE 

GH HOLOTYPE 

YO I SOTYP E 

hsY 

us 462090 

I SOTYPE 

I SOTYPE 

102- CINNAMOMEA 0LNEYqS.T. 

1868. 

IN GRAYTA~T PROC. AMER. ACAD. ARTS 7:396. 

USA: CALIFORNIA: HUMBOLDT COO: RED MOUNTAIN (BOLANDERTH.N.~ 

6477. -- --- 1866 I 

CAS 553874 TYPE FRAGMENT 

GH 

us 

us 

28457 

319228 




103. CIRCINNATA MEYERTC-A.~ 

MEMO ACAD. IMP SCI ST.-PETERSBOURG DIVERS SAVANS 1 :209~ PL.6.


1040 

1050 

106. 

107- 

168. 

1100 

1110 

1120 

113. 

1831. 

USA: ALASKA: ALEUTIAN ISLANDS9 UNALASKA 4 ISLAND) 

( CH A Y I S S 0 9 L A 9 --- . ---I 

GH ISOTYPE 

CLADOSTACHYA VAR. MAXIMA KUKENTHAL 9 Go IN ENGLER, H .Gm Ao 9 PFL AN ZENR 

49 FAM.20:268. 1909. 

BOLIVIA: --: -- (BANG9M.t 2210. ---I 


CLIVICOLA FERNALD9M.L. AND WEATHERBY,C.A. 9 RHODORA 33:233. 1931 

CANADA: QUEBEC: GASPE COO: MOUNT SAINT PIERRE (FERNALD9M.L. t 

WEATHERBYqCoA. AND STEBBINS9G.L.r 24110 05 JUL 19311 

GH 

us 1839933 

HOLO TYP E 

ISOTYPE 

COLLECTA DEWEY9C.y AMER. J. SCI. ARTS SER.19 11:314. 1826. 

USA: MASSACHUSETTS: HAMPSHT RE COO : WORTHINGTON (DEWEYpC.9 --- ) 

---. 

GH HOLOTYPE 

COLUMBIANA DEWEY9C.p AMER. J. SCI. ARTS SER.1, 30:62. 1836. 

USA: --: COLUMBIA RIVER (SCOULER9J.9 --- . ---I 

NY HOLOTYPE 

COMANS VARo STRICTA CHEESEMANrToF.9 

TRANS. & PROC. NEW ZEALAND INST. 242415. 18920 

NEW ZEALAWD: CANTERBURY (DISTRICT): SOUTH ISLAND9 LAKE TEKAPOi 

JAN 1883) 


ALL 2sor FT. (CHEESEMAN~LF.~ --- 0 -- 

COHMUNIS BAILEY9L.H. 9 MEMO TORREY BOT. CLUB 1 :41 1889. 

USA: NEW YORK: YATES COO: PENN VAN (SARTWELL,H.P.9 108. 

-- --- 1848) 


CONClNNOIDES MACKENZIEqKoK.9 BULL. TORREY BOT. CLUB 33:440. 1906. 

USA: MONTANA: FLATHEAD COO: COLUMBIA FALLS (WILLIAMS9R.S.9 ---. 

07 JUN 1893) 

NY TYPE 

CONFERTIFLORA BOOTT9F. IN GRAYpA.9 MEMO AMER. ACAD. ARTS 

NOS., 6:418. 1859. 

JAPAN: HOKKAIDO (PREFECTURE) : HAKODATE (WRIGHTqC.9 ---. 

0- JUN 1855) 

us 27235 TYPE MATERIAL 

CONJUNCTA BOOTT9F.t ILL. GENUS CAREX 3:1229 PL.392. 

USA: OHIO: FRANKLIN COO: COLUMBUS (SULLIVANT9W.S. 

1862. 

9 --- . ---I 

C AS 383550 SYNTYPE 

GH SY NTY PE 

CONSPECTA YACKENZIE9KoK.9 No AMER. FL. 182294. 1935.

NUMBER 12 47 

1140 

12Qo 

121, 

1220 

MEX ICO: PUEBLA: PUEBLA ( ARSENE9 Go ( FRERE) 9 1359. 01 AUG 1907) 


CONSTANCEANA STACEYTJ.W.T LEAFL. W. BOT. 2:123. 1938. 

USA: W4SHINGTON: YAKIMA C O O : MOUNT ADAMS ("PADDO") 9 WODEN 

VALLEY (SUKSDORFtW.N.9 6864. 16 AUG 1909) 

CAS 242987 HOLOTYPE 

DS 

NY 


I S OTY P E 

CONVOLUTA YACKENZIE9K.K. 9 BULL. TORREY BOT. CLUB 43:428. 1916. 

USA: NEW JERSEY: MORRIS C O O : BUDD'S LAKE (MACKENZIE9K.K.9 20880, 

10 JUN 19C6) 

NY TYPE 

COOLEYI DEWEYtC. IN WOODtA., AMER. J. SCI, ARTS SER.19 48:144. 

1845 

USA: MICHIGAN: MACOMB COO: WASHINGTON (COOLEYtD.9 --- ---I 

GH HOLOTYPE 

COSTATA SCHWEINITZ~L.D.T ANN. LYCEUM NATO HIST. NEW YORK 1:67. 

1824. 

--- NY TYPE COLLECT ION 

USA: PENNSYLVANIA: NORTHAMPTON C O O : EASTON (SCHWEINITZTL.D.T 

---I 

CRANDALLII GANDOGERtM. 9 BULL. SOC. BOT. FRANCE 66:295. 1920. 

USA: COLORADO: SUMMIT COO: GRAYS PEAK (JONEStM.E., 834. 

28 AUG 1878) 


CRAWFORDII FERNALDtM.L.9 PROC. AMER. ACAD. ARTS 37:469* PL.1. 

19C2. 

USA: NEW HAMPSHIRE: COOS C0.z MOUNT WASHINGTON, BETWEEN 

YARSHFIELD AND CRAWFORDS (FAXONvE. AND FAXON9C.E. 9 ---. 

06 JUL 1878) 

GH SYNTYPE 

CRAWFORDII VARo VIGENS FERNALD9M.L. t PROC. AMER. ACAD. ARTS 

37:470~ PL.1. 1902. 

CANADA: QUEBEC: GASPE COO: EAST GASPE (MACOUNTJOHN, 6. 

01 AUG 1882) 

GH SY NTYP E 

CREBRIFLORA WIEGAND9K.M.9 RHODORA 24:197. 1922. 

USA: FLORIDA: GADSDEN COO : APPALACHICOLA RIVER9 CHATTAHOOCHEE 

(CURTISStA.H., 3267. -- SEP 1882) 


GH TYPE 

NY TYPE 

CRINITA VARo BREVICRINIS FERNALDTM.L.T RHODORA 48:54* 1946. 

USA: VIRGINIA: DINWIDDIE COO: ROWANTA (FERNALD9M.L. AND


1250 

126. 

LONGTB.T 8143. 08 JUN 1938) 

GH HOLOTY PE 


CRINITA VARo M!NOR BOOTTTF.T ILL. GENUS CAREX 1:18. 1858. 

USA: NEW YORK: YATES COO: PENN VAN (SARTWELL,H.P*, 78. ---I 

CAS 553883 TYPE COLLECTION 

CRINITA VAR. SIMUCANS FERNALD9Y.L. T PROC. PORTLAND SOC. NATO HIST. 

2:135. 1897, 

USA: MAINE: PISCATAQUIS COO: GREENVILLE (FERNALD,M.L.p 264. 

04 JUL 1894) 

GH SYNTYP E 


CRISTATA SCHWEINITZTL.~. T ANN. LYCEUM NAT. HIST. NEW YORK 1:66. 

1824. 

USA: NEW JERSEY: -- (---T --- . ---I 

GH I SOTY P E 

CRUS-CORVI SHUTTLEWORTHTR-J. EX KUNZETG.~ SUPPL. SCHKUHR'S RIEDGR. 

1281 PL.32. 1844. 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS -- --_ 1832 1 

(DRUMMONDTT., 432. 

GH ISOTYPE 

CRUS-CORVI VAR. VIRGINIANA FERNALDTY.L.T RHODORA 39:3939 PL.476. 

1937. 

USA: VIRGINIA: SOUTHAMPTON COO: DREWRYVILLE (FERNALDvM. Lo 

LONGTB. AND SMART,R.F., 5677. 22-23 JUN 1936) 

GH HOLOTYPE 

YO 1108572 ISOTYPE 

NY I SOTY P E 


CRYPTOLEPIS YACKENZIEtKoK., TORREYA 14: 157. 1914. 

USA: NEW JERSEY: SUSSEX COO: WHITE POND (MACKENZIE,K.K., 4645. 

26 JUN 1910) 

NY TYPE 

CUBENSIS KUKENTHAL, G o 9 REPERT SP. NOY. REGNI VEG. 23:220. 1926. 

CUBA: ORIENTE: PIC0 TURGUINO (EKMANqE*L., 14506. 21 JUL 1922) 

NY I SOTYPE 


CUBENSIS VARo FLACCIDA KUKENTHAL9G.9 REPERT. SP. NOV. REGNI VEG- 

23: 221. 1926. 

HAITI: --: -- (EKMANvE.L., --- 08 AUG 1925) 


CUCHUMATANENSIS STANDLEY,P.C. AND STEYERMARKIJ.A.T CEIBA 4:62. 

1953. 

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUYATANEST TUNIMA;

NUMBER 12 49 

ALTO 3400-3500 Me ( STEYERMARK9J.A. 9 48347. 07 JUL 1942) 

F 1128952 HOLOTYPE 

132- CULHENICOLA STEYERYARKTJ.A., FIELDIANA? BOT. 28:65, FIG.7. 1951. 

VENEZUELA: SUCRE: CERRO fURUMIQUIRE9 EASTERN PEAK; ALTO 2500 Me 

(STEYERYARKIJ~A.~ 62605. 06 YAY 1945) 

F 12661 70 HOLOTY PE 

133. CUHULATA FOR. SOLUTA FERNALDtMoLmr RHODORA 44:285. 1942. 

CANADA: NOVA SCOTIA: QUEENS COO: BROAD RIVER (FERNALD9M.L. AND 

BISSELLtC.H.? 20311. 16 AUG 1920) 

GH HOLOTY PE 

134, CUNEATA OHWIIJ., MEMO COLL- SCI. KYOTO IMP. UNIV.9 SERoB9 BIOL. 

6:256. 1931. 

JAPAN: AOYORI (PREFECTURE) : HONSHU ( I SLAND) 9 AOMORI 

(KINASHI9N.t --- -- JUL 1909) 


135- CURATORIUM STACEYtJoW., LEAFL. We BOT. 2:13. 1937. 

USA: ARIZONA: COCONINO COO: GRAND CANYON NATIONAL PARK? KAIBAB 

TRAIL TO ROARING SPRINGS (EASTWOODvA. AND HOWELL9J.T. 9 1101. 

23 JUN 1933) 



136. CUSICKIh MACKENZIE9K.K. IN PIPER9C.V. AND BEATTIE,R.K.r 

FL. NW. COAST 72.. 1915. 

USA: OREGON: BAKER COO: HEAD OF BURNT RIVER (CUSICKqW.C.9 1331. 

-- JUL 1886) 

NY SY NTY P E 

137. OANAENSIS STACEY?J.W., LEAFL. W. BOT. 2:166. 1939. 

USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA (HOWELL9J.T., 14546. 

11 AUG 1938) 


CAS 2 59875 HOLOTY PE 

GH I SOTYPE 

us 1765700 ISOTY PE 

138- DAVYI YACKENZIE 9K.K. 9 BULL. TORREY BOT. CLUB 43:606. 1916. 

USA: CALIFORNIA: PLACER COO: TRUCKEE RIVER (BURT-DAVY?J.r 3266. 

25-30 JUN 1897) 

GH I SOTYPE 

JEPS 2511 ISOTYPE 

NY 

uc 50814 

I SOT YP E 

HOLOTYPE 

139. X DEAHII HERMANNtFmJ. p RHODORA 40:81. 1938. 

-D-


144. 

1460 

USA: INDIANA: PIKE CO.: OTWELL (HERMANN9F.J.9 6147. 

05 JUL 1934) 


DEBILIFORMIS MACKENZIETK.K.T BULL. TORREY BOT. CLUB 37:244. 

USA: CALIFORNIA: MENDOCINO COO: -- (BOLANDERTH.N-T 6477. 

-- -_- 1866) 

1910. 


OEBILIS VARo INTERCURSA FERNALDTM.L*T RHODORA 44:3@79 PL.713. 

1942. 

USA: VIRGINIA: GREENSVILLE CO.: ORION (FERNALDpM-L. AND 

LONG9B.9 12016. 13 JUN 1940) 

GH HOLOTY PE 


OEBfLIS VAR. PUBERA GRAYfA.9 MAN. BOT. ED059 593. 1867. 

USA: PENNSYLVANIA: CENTRE CO.: BEAR MEADOWS (PORTERfT.C.9 ---* 

--- ) 

GH HOLOTY PE 

DEVIA CHEESEYANTT*F*T TRANS. C PROC. NEW ZEALAND INST. 15:301. 

1883. 

NEW ZEALAND: NELSON (DISTRICT) : SOUTH ISLAND? NELSON 

(CHEESEMAN9T.F.9 83. -- JAN 1882) 

GH ISOTYPE 

DEWEYANA VARo COLLECTANEA FERNALD9M.L. 9 RHODORA 15 :93* 1913. 

CANADA: QUEBEC: BONAVENTURE CO. : CASCAPEDIA RIVER9 GRAND 

CASCAPEOIA (W1LLIAMStE.F.; COLLINS9J.F. AND FERNALD9M.L.r 

--- 12-15 JUL 1905) 

GH HO LO TY P E 

OEHEVANA VARo SPARSIFCORA OLNEYpS-T. EX BAILEY,L.H.r BOT. GAZ. 

13:87* 1888. 

USA: OREGON: MARION CO.: SALEM (HALL9E.9 580. -- --7 1871) 

F 455703 TYPE COLLECT ION 

F 1429766 TYPE COLLECTION 

GH TYPE COLLECT ION 


DIGITALIS VARo ASYNMETRICA FERNALDfM.L.9 RHODORA 43:544. 1941. 

USA: VIRGINIA: SOUTHAMPTON CO. : APPLEWHITE CHURCH 

(FERNALD9M.L. AND LONGpB.9 11791. 08 MAY 1940) 

CA S 336835 I SOTYPE 

GH HOLOTYP E 



DIGITALIS VAR. GLAUCA CHAPMANTA.W.T FL. S. U.S. ED.19 

USA: FLORIDA: MIDDLE FLORIDA (CHAPMANTA*W.T ---a 

541. 1860. 

-- --- 1842 1 

NY 

us 969118 


TYPE COLLECTION

NUMBER 12 51 

149- 

150- 

151. 

1520 

153- 

154. 

1550 

156. 

DIGITALIS VARo MACROPODA FERNALDtM*L.-t RHOOORA 40:400t PL-511- 

1938- 

USA: VIRGINIA: GREENSVILLE COm: -- (FERNAL0tM.L- AND LONGtB-e 

7767. 08 APR 19381 

GH HOLOTYPE 


NY I SOTY PE 


DIVERSISTYLIS ROACHtAa We 9 MADRONO 112277. 1952. 

USA: OREGON: LIYN CO.: CLEAR LAKE JUNCTION (R0ACHtA.W-9 202- 

10 JUN 1949) 


DONNELL-SMITH11 BAILEYvL~H.~ MEY. TORREY BOT. CLUB 1:56. 1889. 

GUATEMALA: ALTA VERAPAZ: PANSAMALA; ALT- 3800 FT. (SMITH, J.0. 

AND TURCKHEIMtHa, 659- -- JUN 1885) 


DOUGLAS11 VAR. DENSISPICATA DEWEYtCat AMERm J. SCI. ARTS 

SER-27 32:41e 1861- 

USA: NEBRASKA: -- (HAYDENPF-V-T 580. ---I 

GH TYPE MATER I AL 

DUDLEY1 MACKENZIE 9K-K. v ERYTHEA 8230. 1922. 

USA: CALIFORNIA: MONTEREY CO.: TASSAJARA HOT SPRINGS 

(ELMERtA-DeE-r 3132- -0 JUN 1901) 

0s 145619 HOLOTYPE 


YO I SOTYP E 

YY I SOTYPE 

X OUMANXI LEPAGEtE., NATURALISTE CANAD. 83:143, FIG-4- 1956. 

CANADA: QUEBEC: VIEUX-COMPTOIR (LEPAGEvE. 9 32078. 30 JUL 1954) 

GH ISOTYPE 


DURANDXI BOECKELER, J.0-t ALLGm BOT- Z. SYST. 2 :189- 1896. 

COSTA RICA: --: CERRO OE BUENA VISTA (PITTIERpH- AND TONDUZTA-, 

3376. 19 JAN 18911 



DURIFOLIA BA1LEYtLeH.t BULL. TORREY BOT. CLUB 20:428. 1893- 

CANADA: SASKATCHEHAN: CARLTON HOUSE 4 52 51"- t 106 13'W. 1 

(RICHARDSON, J 9 -0- -0-1 

NY SY NTY P E 

DUTILLY! 0'NEILLtH.T- AND OUMANtM-t RHODORA 43:413t PL.669- 1941. 

CANADA: MANITOBA: CHURCHILL RIVER, CHURCHILL (DUMAN,Y-, 1506. 

08 AUG 1938) 

GH I SOTYPE


- E- 

151. 

158, 

159. 

160, 

161. 

162 w 

163 w 

164. 

165. 

EASTWOOOIANA STACEYTJ.W.T LEAFL. W. BOT. 2:121. 1938. 

USA: OREGON: GRANT CO.: DIXIE MOUNTAIN (HENDERSONTL.F.T 

25 JUL 1925) 

CAS 1363 86 HOLOTY PE 


5583. 

GH I SOTYP E 

EBENEA RYDBERGTP.A. 9 BULL. TORREY BOT. CLUB 28: 266. 1901. 

USA: COLORADO: EL PAS0 COO: PIKES PEAK (CLEMENTStF.9 _- -_- 

---. 

1906 I 

NY TYPE 

ECHINATA VAR. ORMANTHA FERNALDt M.L. 9 PROC. AMER. ACAD. ARTS 

37:483* PL.4. 1902. 

USA: CALIFORNIA: EL DORADO COO: SIERRA NEVADA RANGE, 

STRAWBERRY CREEK (BRAINERDTE., 160. 18 JUC 18971 

GH HD LO TYP E 

EGGERTII BAILEYTLOH~T BOT. GAZ. 21:6. 1896. 

USA: MISSOURI: BUTLER COO: -- (EGGERT9H.q --- 08 AUG 1893) 


EGGLESTONlI YACKENZIETK~K-T BULL. TORREY BOT. CLUB 422614. 1915. 

USA: COLORADO: GUNNISON C0.2 MOUNT CARBON, KEBLER PASS 

(EGGLESTON,W.W.q 6181. 22 AUG 191C) 

NY I SOTYP E 


EGGLESTONII VARw FESTIVELLIFORMIS HERMANN,F.J.r BRITTONIA 12:78. 

1960. 

MEXICO: NUEVO LEON: GALEANA 4 SCHNEIDER,R.A. 9 954. 25 JUL 1938) 


EGREGIA MACKENZIETK~K~T BULL. TORREY BOT. CLUB 42:414. 1915. 

USA: WASHINGTON: KLICKITAT COO: FALCON VALLEY (SUKSDORFTW.N.T 

5181. 15 JUL 19051 


NY TYPE 

EKHANII KUKENTHALTG., REPERT. SP. NOV. REGNI VEG. 23:221. 1926. 

HAIT I: OUEST: PET IONVILLE ( EKMANT E.L. 7 H1453. 12 AUG 1924) 

GH I SOTYPE 

NY TYPE YATERIAL 

us 1411790 COTYPE 

EKMANIl VAR. HOTTENSIS KUKENTHALvG. AND EKMANTE.L.T ARK. BOT. 

22A( 17) : 9. 1929. 

HAITI: .--: MORNE CALUMETTE; ALTO 1200-1300 M e (EKMAN9E.L. T

NUMBER 12 

H10662. 14 SEP 19281 

GH I SOTYPE 


166- ELBERTANA KELSOvL.9 BIOL. LEAFL. 31~3. 1945. 

USA: COLORADO: LAKE COO: MOUNT ELBERT (KELSOIL~T 4967. 

C1 AUG 1945) 

GH TYPE MATES1 AL 

1670 ELEOCHARLS BAILEY,L.H.t MEM. TORREY 601. CLUB 1:6. 1889. 

CANADA: SASKATCHEWAN: SASKATCHEWAN PLAINS ( YACOUN~JOHNI 1665. 

12 AUG 1872) 

GH I SOTYPE 

168. ELMER1 KUKENTHAL,G., REPERT. SP. NOV. REGNI VEG. 8:326. 1910. 

PHILIPPINES: BENGUET: LUZON (ISLANDIT BAGUIO (ELMERqA.D.E.1 

8444. -- MAR 1907) 



1690 ELROOI JONES9M.E.q BULL. MONTANA STATE UN1V.r BIOL. SER. 15:70. 

1910. 

USA: MONTANA: BEAVERHEAD COO: YONIDA (J0NESvM.E. 7 ---a 

08 JUL 1909) 


NY I SOTYP E 


170. ELYNOIDES HOLM,H.T.T AMER. J. SCI. SER.4, 9:356. 1900. 

USA: COLORADO: MIN'ERAL COO: PAGOSA PEAK; ALT. 12000 FT. 

(BAKER,C.F., 230. -- AUG 1899) 

GH I SOTYPE 



171. ENGELMANNL BAILEY,L.H.9 PROC. AMER. ACAD. ARTS 22:132. 

1886 ("1887") 

USA: COLORADO: EL PAS0 COO: MOUNT FLORA9 "PROBABLY NEAR 

COLORADO SPRINGS" ( ENGELYANNqG. --- . -- --- 1874) 

GH HOLOTYPE 

1720 EPAPILLOSA M4CKENZIE,K.K. IN RYDBERGIP~A~T FL. ROCKY MOUNT. 

138, 1060. 1917. 

USA: UTAH: PIUTE COO: MARYSVALE (JONES,M*E., 53450 

01 JUN 1874) 

YO I S OTY PE 

NY HOLOTY PE 

hsY I SOTYP E 


173. EREMOSTACHYA BLAKEIS-T~T J. ARNOLD ARBOR. 28:99. 1947. 

INDONESIA: WEST NEW GUINEA: LAKE HABBEYA; (COUNTRY AS ItDUTCH 

NEW GUINEA") (BRASS9L.J.r 10255. -- OCT 1938)

54 

174. 

175. 

176. 

117. 

1780 

179. 

180. 

181. 

A I SOTYP E 


ERXLEBENIANA KELSOtL., BIOL. LEAFL. 5121. 19500 

USA: COLORADO: GILPIN COO: ROLLINSVILLE (KELS0tL.v 6362. 

24 JUL 1948) 

GH TYPE MATERIAL 

EURYCARPA H0LYtH.T.t AMER. J. SCI. SER.49 20:303. 1905. 

USA: WASHINGTON: KLICKITAT Coo: FALCON VALLEY (SUKSD0RFyW.N.t 

1284. 26 JUN 1886) 


EURYCARPA VARm ATTENUATA KUKENTHALtGot REPERT. SP. NOV. REGNI VEG. 

26: 254. 1929. 

USA: WASHINGTON: KLICKITAT C O O : FALCON VALLEY (SUKSD0RFtW.N. I 

11551. 21 AUG 1924) 

CAS 246772 TYPE C OLLEC T I ON 

EURYSTACHYA HERMANNTF~J~T LEAFL. We BOT. 8:109. 1957. 

CANADA: ALBERTA: JASPER NATIONAL PARK, MOUNT EDITH CAVELLt 

CAVELL LAKE (HERMANNtF.J.9 13529. 28 AUG 1956) 


GH 

us 2265959 

I SOTYPE 

HOLOTYPE 

EXPLORATORUH NELMES,E.t BULL. MISC. INFORM. 108. 1938. 

MALAYSIA: SABAH (TERRITORY 1: MOUNT KINABALU; ALT. 4000 FT.; 

(COUNTRY AS "BORNEO") (CLEMENStJ. AND CLEMENStMoSot 34297. 

28 JUL 1933) 

GH HOLOTY PE 

NY I SOTYPE 

X EXSALINA LEPAGEtEot NATURALISTE CANAD. 83:133. 1956. 

CANADA: QUEBEC: PIAGOCHIWI RIVER (DUTILLYtA.; LEPAGEtE. AND 

DUMANtMot 32793. 29 AUG 1954) 


-F- 

FARGESII FRANCHETtA., BULL. SOC. PHILOM. PARIS SER.8, 7:34. 1895. 

CHIQA: SZECHWAN: TCHEN-KEOU-TIN (FARGES9R.P. 9 --- . ---I 



FELIPENSIS CLARKEtCoBot BULL. MISC. INFORM. ADD.SER.8:84. 1908. 

MEXICO: OAXACA: SIERRA DE SAN FELIPE; ALTO 10000 FT. 

(PR1NGLEtC.G.t 4838. 25 AUG 1894) 


NY TYPE COLL ECT ION 

182. FENOLERIANA BOECKELERtJ.O.t LINYAEA 39:135. 1875.

NUMBER 12 55 

183, 

184. 

185. 

186, 

187. 

188- 

189, 

190. 

191. 

USA: NEW MEXICO: -- (FENDLERqA.9 878. -0 -0- 1847 

YO 1816497 TYPE COLLECTION 

NY TYPE 

FESTIVA DEWEY,C.T AMER. J. SCI. ARTS SER.1, 29:246. 1836. 

USA: NORTHWEST TERRITORIES: MACKENZIE DISTRICT: GREAT BEAR 

LAKE ( "BEAR LAKE") (RICHARDSON, J 9 --- ---J 

NY SYNTYPE 

FESTIVA VAR. DECUHBENS HOLMTH~T. 9 AMER. J. SCI. SER.4, 16:20,26. 

1903 

USA: COLORADO: MINERAL COO: PAGOSA PEAK (BAKERfC.F.9 232. 

-- AUG 1899) 



FESTIVA VAR. STRICTA BAILEY,L.H.T MEMO TORREY BOT. CLUB 1:51. 

1889 

USA: CALIFORNIA: -- (KELLOGGTA. 

-- --- 1868- 1869 1 

AND HARFORD9W .G.W.T 1073. 


FESTIVELLA MACKENZIE,K.K., BULL. TORREY BOT- 

USA: WYOMING: ALBANY CO.: -- (NELSONTA., 

CLUB 42:609. 1915. 

3275. 02 JUL 1897) 

GH I SOTYPE 

h(Y TYPE 

FETA BAILEYTL.H.T BULL. TORREY BOT. CLUB 20:417. 1893. 

USA: CALIFORNIA: SONOMA C O O : CLOVERDALE SPRING (BOLANDER,H.N., 

50. ---I 

GH HOLOTY PE 

FILIFOLIA VARm EROSTRATA KUKENTHALqG. I N ENGLERTH.G.A.v PFLANZENR. 

41 FAM.20:86. 1909. 

USA: CALIFORNIA: EL DORA00 COO: ECHO LAKE (BRAINERDtE.9 111. 

11 JUL 1897) 

GH I SOTYP E 

FISSA MACKENZ?E,K.K.v No AYER. FL. 18:64. 1931. 

USA: OKLAHOMA: CREEK COO: SAPULPA (BUSHI 6.F. 9 

18 MAY 1895) 

1043. 

YO I SOTY P E 

NY HOLOTY PE 

FISSA VAR. ARISTATA HERMANNvF.J.9 RHODORA 67:198. 1965. 

USA: FLORIDA: SEMINOLE C O O : OVIEDO (RAY9J.D.; WOOD9C.E.; 

SYITH,A.C. AND EAT0NtR.J.t 10750. 26 APR 1961) 

GH HOLOTYPE 

4Y I SOTYP E 


FISSURICOLA MACKENZIETK.K.T MUHLENBERGIA 5:530 1909. 

USA: NEVADA: ELK0 COO: RUBY MOUNTAINST HUMBOLDT RIVER


1920 

193. 

195. 

196, 

1970 

1980 

199. 

200. 

201 0 

(HELLERTA.A.T 9429. 11 AUG 1908) 


FLACCIOULA STEUDEL,E.G*, SYN. PL. GLUM. 2:199. 1855. 

USA: OHIO: MIAMI RIVER VALLEY (FRANK,J.C., 55. -- --- 1835) 

NY TYPE 

FLACCIFOLIA MACKENZ1EtK.K.r ERYTHEA 8:92. 1922. 

USA: CALIFORNIA: SOUTHWEST (PARTI (GRANT,G.B.p ---a 

01 MAY 1902) 


FLACCOSPERHA DEWEYfC.9 AMER. J. SCI. ARTS SER.2r 2:245. 1846. 

USA: FLORIDA: CAMP SABINE (LEAVENWORTH,M.C. T --- . -- --- 1846 1 

GH HOLOTY PE 

NY I SOTYPE 

FLAVA VARo GASPENSIS FERNALDTM.L.T RHODORA 8:200. 1906. 

CANADA: QUEBEC: BONAVENTURE C O O : BONAVENTURE RIVER, BETWEEN 

BALDE AND BAIE DES CHALEURS (COLLINS,J.F.: FERNALDTM.~.. AND 

PEASE,A.S., --- 05-08 AUG 19041 

GH HO L 0 TY P E 

FLAVA VARo RECTIROSTRA GAUDINTJ.F.G.P.I FL. HEtV. 6:97. 1830. 

SWITZERLAND: VALAIS (CANTON) : ZERYATT (---, --- -- AUG 1827) 

GH I S OTY PE 

FOENEA VARo PERPLEXA BAILEYTL.H.T MEMO TORREY BOT. CLUB 1:27. 

1889. 

CANADA: NEW BRUNSWICK: KENT C O O : -- (FOWLERTJ., 

-- --- 

---. 

1871 1 

GH I SOTYPE 

FORMOSA DEWEY9C.v AMER. J. SCI. ARTS SER.1, 8:98. 1824. 

USA: NEW YORK: YATES COO: PENN YAN (SARTWELLTH.P., ---. ---I 



GH I SOTY P E 

FRACTA MACKENZIETK-K. 9 ERYTHEA 8:38. 1922. 

USA: CALIFORNIA: SISKIYOU Coo: MOUNT SHASTA (PRINGLE,C.G.t -0-0 

23 AUG 1881) 

us 81 781 0 HOLOTY PE 

FRANKLIN11 BOOTTTF. IN HOOKERTW. J. 9 FL. BOR.-AMER. 2:2179 PL.218. 

1839 ("1840") 

USA: --: ROCKY MOUNTAINS (DRUMMONDTTOT --- . ---I 

GH I SOTYPE 

VY TYPE 

FULVESCENS YACKENZIETK.K.T BULL. TORREY BOT. CLUB 37:239. 1910. 

ST. PIERRE AND YIQUELON! --: MIQUELON (ISLAND), LANGLADE 

(ARSENE,L.(FRERE), ---a 28 JUL 1902)

NUMBER 12 57 

NY TYPE 

202. FUSCOLUTEA BOECKELERtJmO.9 BOT. JAHRB. SYST. 7:278* 1886. 

MEXICO: SAN LUIS POTOSI: -- (SCHAFFNERTJ~G~T 221. -- --- 1877 1 

YY 

us 397187 

TYPE MATERIAL 


203- FUSCOTINCTA MACKENZIETK~K~T BULL. TORREY BOT. CLUB 36:478. 1909. 

2 

MEXICO: 04XACA: 

1894) 

SIERRA DE SAN FELIPE (PRINGLETC.G.T 

TYPE 

4839. 

251773 TYPE COLLECTION 


19 AUG 

NY 

us 

us 

- G- 

RBERI FERN LDTM.L.T RHODORA 37:253* 19 5. 

USA: PENNSYLVANIA: ERIE CO.: ERIE9 PRESQUE ISLE (PENINSULA) 

( GAR B ER T A. P T --- 09 JUN 18691 

GH I SOTY P E 


2050 GARBERI VAR. BIFARIA FERNALD9M.L. T RHODORA 372253. 1935. 

CANADA: QUEBEC: GASPE C0-z WEST GASPET SAINTE ANNE DES MONTS 

( COLL I NS 9 JmF AND FERNALDq M-L 9 --- 03-17 AUG 1905) 

GH HOLOTYPE 

206. GAYANA VARo HYALINA B A I L E Y T L ~ H ~ PROC. T 

1886 ( "1887") 

AMER. ACAD. ARTS 22:135. 

MEXICO: SONORA: SONORA (THURBERTG-T 652. ---I 

NY TYPE 

207. GEOPHSLA MACKENZIE9KmK.t BULL. TORREY BOT. CLUB 40:546. 1913. 

USA: NEW MEXICO: RIO ARRIBA CO.: TIERRA AMARILLA 

(EGGLESTONTW.W.~ 6584. 18 APR-25 MAY 19111 

us 66@800 TYPE 

208. GEYERS BOOTTTF.~ TRANS. CINN. SOC. LONDON 20:118. 

USA: --: ROCKY MOUNTAINS (GEYERIC-A. T 332. ---I 

1846. 


209. GCAREOSA VARm AMPHIGENA FERNALD9M.L.9 RHODORA 8:47. 1906. 

CANADA: QUEBEC: BONAVENTURE COO: ESCUMINAC BAY1 ESCUMINAC 

( FERNALDT M.L 9 --- 28 JUN 1904) 

GH HOLOTY PE 

NY I SOTYP E 

2100 GLAUCODEA TUCKERMANqE. EX OLNEYTS-T. IN GRAYTA.~ 

PROC. AMER. ACAD. ARTS 72395. 1868. 

USA: MASSACHUSETTS: HAMPSHIRE CO. : MOUNT HOLYOKE (TUCKERMAN9 E.9 

--- -- JUN 18641


GH I SOTYPE 

211. GRACILfOR MACKENZIETKOKO T BULL. TORREY BOTo CLUB 43:6140 1916. 

USA: CALIFORNIA: SONOYA Coo: CLOVERDALE (BOLANDER,HoNo 9 38220 

-- APR 1864) 

CAS 10 30 33 

0s 145620 

GH 

MO 

us 319177 

I SOTYPE 

HO LO TY P E 

I SOTYPE 

I SOTYPE 

ISOTYPE 

2120 GR CILLIHA SCHWEINITZTLO O T ANN. LYCEUM N To HISTo NEW YORK 1:66. 

18240 

USA: PENNSYLVANIA: -- (SCHUEINITZ,L.D.T --- 0 ---I 

GH 

I SOT Y P E 

2130 GRIFFITHII BOOTTIFOT TRANS. LINNo SOCo LONDON 2O:1380 18460 

AFGHANISTAN: --: 0- (GR1FFITHtW.r 78(KEW 607410 ---I 


2140 GRISEA VARo RIGIDA 6AILEYyLoH.y MEMO TORREY B0To CLUB 1:56. 18890 

USA: PENNSYLVANIA: BUCKS COO: SELLERSVILLE (FRETZ,C.D.T ---0 

-- --_ 1884) 

GH HOLOTY PE 

2150 GUATEMALENSIS HERMANN,FoJo, BRITTONIA Z3:14So 19710 

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES, BETWEEN 

TOJIAH AND CHEMALt ALTO 3380 Yo (BEAMANyJoHoy 38800 

31 JUL 1960) 

GH HOLOTYPE 

216. GYHNOCLADA HOLMTHOTOT AMERo J o SCIo SER.4, 14:424. 19020 

USA: OREGON: HURRICANE CREEK: BOGS AT 6000 FTo (cuSIcKyW.co~ 

24870 28 AUG 1900) 


NY ISOTYPE 

GYNOOYNAMA OLNEYySoTo IN GRAYyAoq PROCo AMERo ACADo ARTS 7:394o 

1868. 

USA: CALIFORNIA: MENDOCINO Coo: 

47@@. -- --- 1866) 

MENDOCINO CITY (BOLANDERqHoNoy 


0s 495@0 ISOTYPE 

DS 

NY 


I SOTYP E 

218, HAGIANA KELS0,L.p BIOLo LEAFLo 30:20 19450 

USA: COLORADO: HAGUES PEAKS (KELSOTLO AND KELSO,E~H.T 5250 

C8 AUG 1936) 

- H-

NUMBER 12 

219. 

220. 

221. 

222. 

223. 

224. 

225. 

226. 


HALE1 DEWEYvC., AMER. J. SCI. ARTS SER.29 2:248. 1846. 

USA: LOUISIANA: MISSISSIPPI RIVER (LEAVENWORTH9M.C. AND 

HALEqD.9 683. ---I 

NY SYNTYPE 

HALLIANA BAILEY,CoH.y BOT. GAZ. 9:117. 1884. 

USA: OREGON: -- (HALL9E.q 606. -- --- 1871 1 


HALL11 OLNEY9S.T. IN PORTER9T.C. IN HAYDEN9F.V.9 

ANN. REP. U.S. GEOL. SURV. TERR. 5:496. 1872. 

USA: COLORADO: ROCKY MOUNTAINS: LAT. 39-41 No (HALLvE. AND 

HARB0URqJ.P.r 617. -- --- 1862 1 

F 314892 SYNTYPE 


GH SY NTYP E 

YO SYNTYPE 

NY SYNTYPE 

USA: 

us 

IDAHO: 

29651 SYNTYPE 

PLEASANT VALLEY (PORTERTT.C. 9 --- 26-29 JUN 1871) 

NY SYNTYPE 

HALSEYANA DEWEYTC., AMER. J. SCI. ARTS SER.19 11:313. 1826. 

USA: MASSACHUSETTS: HAMPDEN COO: UESTFIELD (DAVISTE.~ ---. 

---) 

GH 

NY 

HOLOTY PE 

I SOTYPE 

HARFORDII YACKENZIE,Y.K.9 BULL. TORREY BOTo CLUB 43:615. 1916. 

USA: CALIFORNIA: -- (KELLOGG, A. AND HARFORD,W.G.U.9 1073. 

_- --- 1868-1 869) 

NY HOLOTYPE 


HARPER1 FERNALDTM.L.T RHODORA 8:181. 1906. 

USA: GEORGIA: JEFFERSON COO: LOUISVILLE, ROCKY COMFORT CREEK 

(HARPER+R.Y.? 2109. 09 APR 1904) 


GH HOLOTYPE 

NY I SOTYP E 

HASSEI BAILEY9L.H.9 BOT. GAZ. 21:5. 1896. 

USA: CALIFORNIA: SAN BERNARDINO COO: SAN BERNARDINO MOUNTAINS, 

SAN ANTONIO CANYON; ALTO 4500 FT. (HASSE9H.E.f ---. 

-- JUL 1894) 


HATUSIMANA OHWIqJ., JAP. J. BOf. 7:196. 1934. 

TAIWAN: FUKIEN: KAOHSIUNG ("TAKA0")t DAIJURIN (OHWIqJ.9 329. 

-- MAR 1933) 

F 1411493 TYPE YATERIAL 

59


221. 

228. 

229. 

230. 

231 

232. 

233. 

234. 

235. 

HAVOENIANA OLNEY tS.T. IN WATSON, S o 9 

BOT. U.S. GEOL. EXPLOR. 40TH PAR. 366. 1871. 

USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA (BOtANDERtHoN., 5074. 

---) 

GH SYNTYPE 

HAYOENII DEWEYtC.9 AMER. J. SCI. ARTS SER.29 18:103. 1854. 

USA: SOUTH DAKOTA: STANLEY COO: FORT PIERRE (HAYDENtF*V.t 

-4 --- 1853-1 854) 

21. 


HEBETATA BO0TTtF.t ILL. GENUS CAREX 4:172t PL.583. 1867. 

PERU: LIMA: OBRAGILIA? NEAR LIMA (WILKES EXPLOR. EXPED.9 

-- ---. 

--- 1838-1 842 1 

GH HOLOTYP E 

HELLERI MACKENZIEpK.K.9 ERYTHEA 8:80. 1922. 

USA: MEVADA: WASHOE COO: MOUNT ROSE (HELLERt A.A.9 99750 

23 JUL 1910) 


NY TYPE 


HEPBURNII BOOTTtF. IN HOOKER9W.J.t FL. BORo-AMER. 222099 PL.207. 

1839 ( "1 840" 1 

USA: COLORADO: --- ) 

ROCKY MOUNTAINSv SOUTH PARK (DRUMM0ND.T. 9 256. 

GH SYNTY PE 

HETERONEURA BOOTT,W. I N WATS0NtS.t GEOL. SURV. CALIFORNIA9 BOT. 

2:239. 1880. 

USA: CALIFORNIA: LAKE TAHOE TO BEAR VALLEY (KELL0GGtA.t ---. 

03 AUG ---- 1 



HETEROSTACHYA TORREYtJ. EX 0EWEYvC.t AMER. J m SCI. ARTS 

SERo2t 2:248. 1846. 

USA: MICHIGAN: CHIPPEWA COO: DRUMMOND ISLAND (TORREYtJ. 9 &--. 

03 AUG 18391 

NY TYPE 

HINDS11 VARm BREVIGCUHA KUKENTHALtG. IN ENGLER,H.G.Aet PFLANZENR. 

4 9 FAM.20:307. 1909. 

USA: IDAHO: BONNER CO.: HOPE (SANDBERGvJoHot 9330 20 AUG 1892) 

NY ISOTYPE 

HIRSUTA VARm CUSPIDATA DEWEYtC. IN WOODtA., CLASS-BOOK BOT. 758. 

1861. 

USA: XLLINOIS: -- (VASEY9G.t --- . ---I 

YY TYPE COLLECTION

NUMBER 12 61 

2360 

237- 

238- 

239. 

240 0 

241 0 

242 

243 

241. 

HITCHCOCKIANA DEWEYtC.9 AMERm J. SCIm ARTS SER.1, 10:274. 1826. 

USA: MASSACHUSETTS: BERKSHIRE Corn : W I LLI AMSTOWN, 

1823) 

SADDLE 

MOUNTAIN (DAVIS,Emt --- -- --- 

GH HOLOTY PE 

HOLMIANA MACKENZIE+KmKmt BULL. TORREY BOTm CLUB 36:481. 1909. 

USA: MONTANA: JOHN'S LAKE (VREELANDtFmK., 1121. 19 AUG 1901) 

NY TYPE 

HOOD!! VARo NERVOSA BA1LEYtL.H.t MEMm TORREY BOT. CLUB 1 : 1 4 m 1889. 

USA: CALIFORNIA: -- (KELLOGGtA. AND HARFORD+W.G.W.t 1069. 

-- --- 1868-1 869) 


NY SYNTYPE 

HOOD11 VAR. NEUROCARPA PIPERtCmVm, CONTRm Urns. NATL. HERB. 11:167m 

1906. 

USA: CALIFORNIA: -- (KELLOGGtA. AND HARFORDtW.GmW., 1069. 

-- _-- 1868-1869) 


NY SYNTYPE 

HOOKERANA DEWEY9C.t AMER. Jm SCIm ARTS SER.1, 29:248m 1836. 

CANADA: SASKATCHEWAN: CARLTON HOUSE (52 51'N.t 106 13'W.l 

(R1CHARDSONtJ.t --- -0-1 



HORMATHODES FERNALDTY~L.~ RHODORA 8: 165. 1906. 

USA: RHODE ISLAND: PROVIDENCE COm: PROVIDENCE (OLNEYt SmTet ---m 

01 JUL 1867) 

GH SYNTYPE 

HORNSCHUCHIANA VARe LAURENTIANA FERNAL0,M.C. AND WIEGAND,KmMmt 

RHODORA 13: 130. 1911. 

CANADA: NEWFOUNDLAND: PORT AU PORT BAY, TABLE MOUNTAIN 

(FERNALDtMmLm AND WIEGAND,K*Mmt 2897. 16 AUG 19101 

GH HOLOTYP E 

NY I SOTY P E 

HOSTIANA VARo LAURENTIANA FERNALD, M m L m AND W I EGAND9K.M. 9 RHODORA 

26: 122. 1924. 

CANADA: NEWFOUNDLAND: PORT AU PORT BAY, TABLP MOUNTAIN 

(FERNALDtMmL. AND W1EGANDtKmM.t 2897. 16 AUG 1910) 

GH HOLOTYPE 

HOUGHTONIANA TORREYpJm EX DEWEY,Cmt AMER. J. SCIm ARTS 

SER.19 30263. 1836. 

USA: MINNESOTA: CLEARWATER CO.: LAKE ITASCA ("LAKE LA BICHEt 

NEAR SOURCES OF MISSISSIPPI RIVER") (HOUGHTON,D.t ---m 

13 JUL 18321 

NY HOLOTYPE


2450 HUEHUETECA STANDLEYTP~C. AND STEYERMARK9J.A.v 

PUBL. FIELD YUS. NATO HIST., BOT. SER. 23~195, 1947. 

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES 9 CANANA; 

ALTO 2500 M. (STEYERMARKTJOAOT 49055. 18 JUL 1942) 


2460 HYHENODON OHWI TJ. 9 ACTA PHYTOTAX. GEOBOT. 1:298. 1932. 

JAPAN: --• HONSHU ( I SLANDIT OSAWAYURA IN SHIMOTSUKE 

( SEKIMOTOTH. 9 --- 15 JUL 1932) 


2470 ICHANGENSIS CLARKETC.B.T J. LINN. SOC.9 BOT. 36:290. 1903. 

CHINA: HUPEH: -- (HENRYvA.9 7860. -- --- 1885-1888 1 


248. XOAHOA BAILEY9L.H.9 BOT. GAZ. 21:5. 1896. 

USA: IDAHO: BEAVER CANYON (RYDBERGvP.A.9 2339, 07 AUG 1895) 



249. IGNOTA DEWEYTC-T AMER. J. SCI. ARTS SER.29 8:348. 1849, 

USA: LOUISIANA: RAPIDES PARISH: ALEXANDRIA (HALEpD.9 97. 

C AS 553902 TYPE COLLECTION 

---I 

YY TYPE COLLECTION 

2500 ILLINOENSTS DEWEYTC-T AMER. J. SCI. ARTS SER.29 3:245. 1847. 

USA: ILLINOIS: HANCOCK C O O : AUGUSTA (MEAD~S*B.T --- . ---I 


2510 ILLOTA BAILEY9L.H.t MEM. TORREY BOT. CLUB 1:15o 1889. 

USA: COLORADO: ROCKY MOUNTAINS: LAT. 39-41 N o (HALLTE. AND 

HARBOURTJ.P.T 591. -- --- I862 I 



GH HOLOTYPE 

YO I SOTYPE 

2520 INCISOoDENTATA STEUDELTE.G.T SYN. PL. GLUM. 2:189. 1855. 

CHILE: --: -0 ILECHLERTW.~ 1136. -- OCT 1852) 

GH I SOTYPE 

253. INCOHPERTA BICKNELL9E.P. 9 BULL. TORREY BOT. CLUB 35:494. 1908. 

USA: MASSACHUSkTTS: NANTUCKET C O O : NANTUCKET ISLAND 

(BICKNELL,E.P.9 --- 20 JUN 1908) 

NY TYPE 

2540 INCONDITA HERMANNTF. J. T LEAFL. W. BOT. 8:112. 1957. 

CANADA: ALBERTA: RAM RIVER9 NORDEGG (HERMANNTF. J. 9 13347- 

15 AUG 19561 

-I-

NUMBER 12 



255. INCURVIFORMIS MACKENZIE9K.K. I N RYDBERGvP .A. 9 FL. ROCKY MOUNT. 

1209 1060. 1917. 

CANADA: ALBERTA: BANFF NATIONAL PARK9 BANFf; ALTO 

( MACOUNT JOHN 9 --- 0 31 JUL 1891) 

8000 FT. 

GH I SOTYP E 

NY HOLOTYPE 

2560 INFLATA VAR. 

1942. 

ANTICOSTENSIS FERNALDpM.L.9 RHODORA 44:329~ PL.715. 

CANADA: QUEBEC: ANTICOSTI ISLAND, PETITES-RIVIERES 

(MARIE-V ICTORIN, (FRER El AND ROLLAND-GERMAI NT 

(FRERE 1 T 25767. 

20 JUL 1926) 

GH HOLOTYPE 

2570 INOPS BAILEY,L.H., PROC. AMER. ACAD. ARTS 22:126. 1886 ("1887"). 

USA: OREGON: CLACKAMAS COO: MOUNT HOOD (HENDERSONTL.F.T ---. 

-- JUL 1884) 


GH HOLOTY PE 

YY I SOTYPE 

258. INTEGRA MACKENZIE9K.K.9 BULL. TORREY BOT. CLUB 43:608. 1916. 

USA: CALIFORNIA: PLACER COO: SUMMIT; ALTO 7000 FT. 

(HELLERTA.A.T 9841. 16 JUL 1909) 

NY HOLOTY P E 

2590 INTERIWUS YAGUIRETB., BRITTONIA 5: 200. 1944. 

USA: UTAH: CACHE C O O : TONY GROVE LAKE (MAGUIRE96.9 16098. 

05 AUG 19381 


GH I SOTYPE 

NY TYPE 


260. INTERIOR BAILEY9C.H., BULL. TORREY BOT. CLUB 20:426. 1893. 

USA: NEW YORK: YATES COO: PENN YAN (SARTWELL9H.P.9 36. ---I 



NY I SOTYPE 

261. INTERIOR VAR. CHARLESTONENSIS CLOKEY9I.W.9 

BULL. S o CALIF. ACAD. SCI. 38:l. 1939. 

USA: NEVADA: CLARK COO: CHARLESTON PARK (CLOKEY, I.W.9 

19 JUN 19371 

7468. 

CAS 

CAS 

DS 




F 10 76930 I SOTYP E 

GH I SOTYPE 

JEPS 4013 ISOTYPE 

63



MO 1190731 I SOTYPE 


NY ISOTYPE 



2620 INTERIOR VARo JOSSELYNII FERNALDfY.L.9 RHODORA 8:115. 1906. 

USA: MAINE: AROOSTOOK CO,: SAINT JOHN RIVER9 FORT KENT 

( FERNALD9M.L.9 --- 06 JUL 1904) 

GH HOLOTY PE 

NY 

us 

I SOTYPE 

60 5797 I SOTYPE 

263. INTERIOR VAR, KEWEENAWENSIS HERMANN9F.J.9 AMER. MIDL. NATURALIST 

25:19. 1941. 

USA: MICHIGAN: KEWEENAW CO.: 

13 JUL 19361 

EAGLE HARBOR (HERMANN9FoJ.t 7985. 

GH HOLOTY PE 

NY 

us 

I SOTYPE 


264. INTERRUPTA VAR, DISTENTA KUKENTHAL9G.t REPERT. SP. NOV. REGNI VEG. 

26: 254. 1929. 

USA: WASHINGTON: KLICKITAT COO: BINGEN (SUKSDORFqW.N.9 12333. 

22 AUG-05 SEP 1927) 


265. INTUHESCENS FOR0 VENTRIOSA FERNALD9M.L.v RHODORA 44:321r PL.713. 

1942 

USA: VERMONT: ADDISON C O O Z RIPTON (BRAINERD9E.9 ---a 

19 JUL 1898) 

GH HOLOTY PE 

266, fNVOLUCRATELLA MACKENZIEtK.K.9 No AMER. FL. 18:50. 1931. 

MEXICO: SAN LUI S POTOSI : LAS CANOAS ( PRI NGLE TC. Go 9 3126. 

08 JUL 1890) 


GH I SOTYPE 

MO TYPE MATERIAL 

VY 

us 

TYPE COLLECT I ON 


2670 JACINTOENSIS PARISH9S.B.9 BULL. S o CALIF. ACAD. SCI. 4:lOOq PL.16. 

1905. 

USA: CALIFORNIA: RIVERSIDE COO: SAN JACINTO MOUNTAINS9 

TAMARACK VALLEY; ALT, 9000 FT. (HALLTH.M. 9 2483. 

-- JUL-AUG 1901) 

DS 78003 HOLOTYPE 

-J-

NUMBER 12 65 

268. JACOBI-PETER1 HULTENTO.E.G.T ACTA UNIV. LUND. N.S.c 38:30C~ FIG.4. 

1942 

USA: ALASKA: TIN CITY (ANDERSON,J.P., 4871. 19 AUG 1938) 


269. JAMES11 TORREY,J.r ANN. LYCEUM NATO HIST. NEW YORK 3:398. 1836. 

USA: --: ROCKY MOUNTAINS (JAMESTEDWIN, --- . ---I 

NY HOLOTY PE 

270. JAMESONI VARo SUBFULVA KUKENTHALTG.~ REPERT. SP. NOV. REGNI VEG. 

8:7. 1910. 

BOLIVIA: --• -- (BANG1M.p 2376. ---I 


271. JEPSONII HOWELLTJ.T.~ LEAFLO W. BOT. 8:223. 1958. 

USA: CALIFORNIA: TUOLUYNE COO: YOSEMITE NATIONAL PARK, 

TUOLUMNE MEADOWS; ALTO 8800 FT. (JEPSON,W.L.r 4477. 

20 JUL 1911) 

J EPS 20008 ISOTYPE 

NY HOCOTY PE 

272- JONES11 BAILEY,L.H., MEMO TORREY BOT. CLUB 1:16. 1889. 

USA: CALIFORNIA: NEVADA COO: SODA SPRINGS; ALTO 7000 FT. 

(JONESTY.E., ---. 22 JUL 1881) 

NY SYNTYPE 

273. KALOIDES PETRIE,D.p TRANS. & PROC. NEW ZEALAND INST. 13:332. 1881. 

NEW ZEALAND: OTAGO (DISTRICT): SOUTH ISLAND, CARRICK RANGE; 

ALTO 4000 FT. (PETRIETD., --- 0 ---I 

GH I SOTYPE 

274. KATAHDINENSIS FERNACD,M.L.r RHODORA 3:171~ PL.32. 1901. 

USA: MAINE: PISCATAQUIS COO: MOUNT KATAHDIN, DEPOT POND 

( W I L L I 

-K - 

A M S T E F ; C H URC H I L L 7 J R AN D F E R N AL D T M L T 

16 JUL 19C.O) 

GH HOLOTY PE 

NY I SOTYPE 


275- KAUAIENSIS KRAUSSTR., PACIFIC SCI. 4:279. 1950. 

USA: HAWAII: KAUAI COO: KAULUWEHI tROCK,J.F.q 9017. 

-- OCT 1909) 


276. KELLOGGII BOOTTTW. IN WATSONTS., GEOL. SURV. CALIFORNIA, BOT. 

2:240. 1880. 

USA: CALIFORNIA: SIERRA NEVADA RANGE, "LAKE TAHOE TO BEAR 

VALLEY" (KELLOGG, A. 9 --- . ---1 

---

66 

277- 

278 0 

2796 

280. 

281. 

282- 

283. 

204. 

285. 

GH SYNTYPE 


KOKRINENSIS PORSILD,A.E., RHODORA 41:206, PL.551. 1939. 

USA: ALASKA: KOKRINES MOUNTAINS (PORSILD, A.E. AND PORSILDIROT. 9 

711. 23 JUN-05 JUL 1926) 

GH ISOTYPE 

KULfNGANA BAILEYTL~H~T GENTES HERB. 1:13. 1920. 

CHINA: KIANGSI: KULING; ALTO 2500-3500 FT. (BAILEYTL.H.T ---a 

18 JUL 1917) 

NY TYPE 

KURILENSIS OHW1,J.y ACTA PHYTOTAX. GEOBOT. 2:270 19330 

USSR: RUSSIAN SFSR: SAKHALI N OBLAST: KURIL ISLANDS, SHIKOTAN 

(ISLAND), NOTORO; (COUNTRY AS "JAPAN") (UHWI 9 J. 9 813. 

11 AUG 1931) 

F 1406403 TYPE M4TERIAL 

LACINIATA BOOTT cF. T ILL. GENUS CAREX 4:175, PL.594. 1867. 

USA: CALIFORNIA: SACRAMENTO RIVER (RICHTWILLIAMT 

WILKES EXPED. 1241. -- --- 1838-1842) 

NY TYPE 

LACUNARUM HOLYTH.T. T AHER. J. SCI. SER.4, 17:316. 1904. 

-L- 

USA: CALIFORNIA: 

@l JUL 1902) 

SONOMA C O O : SEBASTOPOL (HELLER,A.Ao, 5797. 



YO SYNTYPE 

NY SYNTYPE 

US 430229 SYNTYPE 

LAEVI-CONICA DEWEYTC.~ AMER. J. SCI. ARTS SER.2, 24:470 18570 

USA: NEBRASKA: BIG SIOUX RIVER (HAYDEN9F.V.t --- . ---I 

GH HOLOTYPE 

LAHPROCHLAHYS BLAKETSOT., J. ARNOLD ARBOR. 28:1G4. 1947. 

PAPUA AND NEW GUINEA: PAPUA (TERRITORY): MAFULU; (COUNTRY AS 

"BRIT1 SH NEW GUINEA") (BRASS TL. J. T 5323. -- SEP-NOV 19331 

A I SOTYPE 

CANCIFOLIA CLARKETCOB.~ J. LINN. S0C.r BOT. 36:292. 1903. 

CHINA: HUPEH: -- (HENRY~A~T 5467. ---I 


LANCIFRUCTUS HACKENZIETK.K.T BULL. TORREY BOT. CLUB 43:607. 1916. 

USA: CALIFORNIA: TULARE COO: UPPER KERN RIVER, VOLCANO CREEK 

(HALL9H.M. AND BABCOCKtH.D., 5472. -- JUL 19041 

NY I SOTYPE

NUMBER 12 67 

2860 

287. 

288. 

289- 

290 0 

291 

292 - 

293 

294. 


LANGEANA FERNALDTM~L.~ RHODORA 35:217o 1933. 

CANADA: NEWFOUNDLAND: GARGAMELLE COVE (FERNAL0,M.L.; LONGTB. 

AND FOGG-JR~TJ~M~T 1374. 20 JUL 19291 


GH HOLO TY P E 

LARENSIS STEYERMARKyJoAov FIELDIAVA, BOTo 28:66~ FIG.8. 19510 

VENEZUELA: LARA: BETWEEN BUENOS AIRES AND PARAMO DE LAS ROSAS 

(STEYERMARKTJ.A.T 554700 11 FEB 1944) 



LARICINA MACKENZIETK~KO EX BRIGHTTJoT TRILLIA 9:4,190 19300 

USA: INDIANA: KOSCIUSKO Coo: LEESBURG (DEAMyCoCor 10927. 

05 JUN 19121 

NY TYPE 

LASIOCARPA VARo AMERICANA FERNALDTM~L.~ RHODORA 44:3O40 1942. 

CANADA: NOVA SCOTIA: YARMOUTH COO: ARGYLE (PEASEvAoSo AND 

LONGTB.T 205190 09 JUL 19201 

GH HOLOTYPE 

LATEBRACTEATA WATERFALL,U.To, RHODORA 56: 23. 19540 

USA: OKLAHOMA: YCCURTAIN COO: BROKEN BOW (WATERFALLTU~T~T 

11380. 19 APR 19531 


GH I SOTYPE 

YO 1692174 I SOTYPE 

LAXIFLORA VAR. LEPTONERVIA FERNALDtM.L.9 RHODORA 8:184. 19060 

USA: MAINE: AROOSTOOK CO.: FORT FAIRFIELD (FERNALD,M.L., 146. 

06 JUL 18931 


GH HOLOTY PE 

MO I SOTYPE 

NY ISOTYPE 

LAXIFLORA VARo SERRULATA HERMAVN?F.Jo, RHODORA 40:80. 19380 

USA: INDIANA: CLARK Coo: -- (DEAM~C~C~T 6458. 25 MAY 1910) 


NY I SOTY P E 

LEAVENUORTHII DEWEYTC~T AMERo J o SCI. ARTS SER.2, 2:2460 1846. 

1845 J 


USA: LOUISIANA: -- ( LEAVENWORTHTMOC. T --- -- --4 

LEIOCARPA MEYERTC~A.~ 

MEMO ACADo IMP. SCIo ST.-PETERSBOURG DIVERS SAVANS 1:208t PL.5. 

18310 

USA: ALASKA: SITKA (MERTENS?CoHo, --- . ---I 

GH I SOTYPE

68 

2950 

296- 

297. 

2980 

2990 

3000 

301 

302 


LEIOPHYLLA MACKENZIEqKmKm t Nm AMERm FLm 18:365m 1935- 

CANADA: YUKON TERRITORY: CARCROSS (EASTWOODTAm, 725Am 

16 JUL 1914) 


GH ISOTYPE 


LEMANNIANA VARo SIMPLEX KUKENTHALvGm IN ENGLERqHmGmAmv PFLANZENRm 

49 FAM.20:405* 1909- 

COSTA RICA: --: CERRO DE BUENA VISTA (PITTIERTHm AND TONDUZTA.T 

3381. 19 JAN 18911 


LEMMONI BOOTT,Wm, BOTm GAZm 9:93m 1884- 

USA: CALIFORNIA: SIERRA NEVADA RANGE (LEMMON,JmGmv ---a 

-- --- 1875) 

GH TYPE COLLECT I ON 


LENTICULARIS VARo PAULLIFRUCTUS KUKENTHAL cGm IN ENGLER, HmGm Am 9 

PFLANZENRm 49 FAMm20:308m 1909. 

USA: WASHINGTON: WHI TMAN COm : PALOUSE CREEK ( ELMER9 AmDm Em T 88 1. 

-- JUN 1897) 

NY ISOTYPE 

LEPORINA VARo AMERICANA OLNEYvSmTm EX B A I L E Y ~ L ~ H ~ T 

PROCm AMERm ACAD. ARTS 22:152. 1886 ("1887")m 

USA: OREGON: CLACKAMAS COm: MOUNT HOOD (HALLpEm, 583- 

01 AUG 18711 


F 1425899 TYPE COLLECT ION 

GH TYPE C OLL ECT I ON 



LEPORINELLA MACKENZIEtKmKm, BULL- TORREY BOTm CLUB 43:605m 1916- 

USA: C4LIFORNIA: EL DORADO COm: PYRAMID PEAK (HALL9H.M. AND 

CHANDLERvHmAm T 4716- 01-02 AUG 19031 


GH I SOTYPE 


LEPTOPODA YACKENZIE,K.Km I N RYDBERGTP~A., FL. *ROCKY MOUNT. 

124, 1060. 1917. 

USA: OREGON: CLACKAMAS COm: OSWEGO, ELK ROCK (HELLER9A.A. 9 

10052m 20 YAY 1910) 



NY TYPE 

LIWNOPHILA HERMANNTF- Jo T LEAFLm Wo BOTm 8:28. 19560 

USA: WYOMING: SUBLETTE COm: PINEDALE (HERMANNfFmJm, 12252-

SUMBER 12 69 

21 AUG 19551 


303. LIVIOA VAR. RUFINAEFORMIS FERNALDtMoLo t RHODORA 28: 80 1926. 

CANADA: NEWFOUNDLAND: STRAIT OF BELLE ISLE9 FOUR-MILE COVE 

(FERNAL0tM.L. t WIEGANDvKo Mo AND LONGtBo t 276730 20 JUL 1925) 

GH HOLOTY P E 

3040 LONGICRURIS VARo HENRY1 CLARKEpCoB.9 J o LINN. SOC.9 BOTo 36:2950 

19030 

CHINA: HUPEH: -- (HENRYIA~T 42660 ---I 


3050 LONGICULMIS PETRIEtDot TRANS. 

1882. 

& PROCo NEW ZEALAND INSTo 14:3630 

NEW ZEALAND: OTAGO (DISTRICT): SOUTHLAND SUBOIVI SION: STEWART 

I SLANO PATTERSONS INLET (PETRI ET Do t --- 0 -- JAN 1880) 

GH I SOTYPE 

3060 LONGIROSTRIS VARo MICROCYSTIS BOECKELER,JoOot LINNAEA 41:2410 

18770 

CANADA: MANITOBA: WINNIPEG (BOURGEAU,Eo, ---0 

_- _-- 18 57-1 859 1 


3070 LUNELLIANA MACKENZIETK~K. t BULL. TORREY BOTo CLUB 42:6150 19150 

USA: MISSOURI : JACKSON Coo: OAK GROVE (BUSHtBoFo t 70200 

02 JUN 1913) 

NY TYPE 

3080 LUZULAEFOLIA VARo STROBILANTHA HOLM,HoTor AMERo J o SCIo 

SER.49 20:305. 19050 

USA: CALIFORNIA: NEVADA Coo: OONNER PASS; ALTO 7500 FTo 

(HECLERtAoA.9 7187. 17 AUG 1903) 

c4s 136 ISOTYPE 


3690 LUZULINA OLNEYtSoTo IN GRAYtA.9 PROCo AMER. ACADo ARTS 7:3950 

1868 0 

USA: CALIFORNIA: MENDOCIN0 Coo: YENDOCINO CITY (BOLANDER9HoNop 

4740. -- --- 18661 


0s 76794 ISOTYPE 

GH TYPE C OLL ECT I ON 




3100 MACKENZIANA YEATHERBY 9C.A. 9 CONTRo GRAY HERB. 114:360 19360 

-M-


315* 

318. 

MEXICO: NUEVO LEON: GALEANA (MULLER9C.H. AND MULLERTM.T.T 892. 

28 JUN 1934) 


GH HOLO TYP E 


MACROGLOSSA FRANCHET9A. AND SAVATIERqL.9 ENUM. PL. JAP. 2:148~ 576. 

1879. 

JAPAN: KANAGAWA (PREFECTURE): HONSHU ( ISLAND) 9 YOKOSUKA 

(SAVATIER9L.r 1414. -- --- 1866-1874) 


MACROKOLEA STEUDELTE-G.~ SYN. PL. GLUM. 2:223. 1855. 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMONDtT. 9 420. 

_- --- 1832 1 

NY COTYPE 

WACROSPERUA YACKENZIE9K.K.p BULL. TORREY BOT. CLUB 36:477. 1909. 

YEXICO: OAXACA: SIERRA DE SAN FELIPE; ALTO 6000-7000 FT. 

(PRINGLE~C.G.T 4840. 27 AUG 1894) 

NY TYPE 

MADRENSIS BAILEY~L*H.T ROT. GAZ. 25:270. 1898. 

MEXICO: DURANGO: SIERRA MADRE OCCIDENTAL (ROSE9 J.N.9 2357. 

16 AUG 1897) 

NY ISOTYPE 

lJS 301267 TYPE 

MAGNIFOLIA Y~CKENZIETK~K. I N SMALLTJ~K~T FL. SE. U.S. ED029 1325. 

1913. 

USA: FLORIDA: -- (CHAPMANvA.W.9 --- . ---I 


MANDONIANA BOECKELER9J.O.9 ALLG. BOT. 2. SYST. 2:174. 1896. 

BOLIVIA: --: -- (MANDONTG., 1429. ---I 

NY I SOTYPE 

HARCIOA VARo OEBILIS BAILEYTL.H.T PROC. AMER. ACAD. ARTS 22:136. 

1886 ("1887" 

USA: OREGON: HARNEY C O O : HARNEY VALLEY (HOWELL9T.J.r 937. 

27 MAY 1885) 


NY TYPE 

MARIPOSANA BAILEY9L.H.p BULL. TORREY BOT. CLUB 43:619. 1916. 

USA: CALIFORNIA: TUOLUMNE C O O : YOSEMITE NATIONAL PARKT 

TUOLUMNE MEADOWS (JEPSON? W .L 9 4476. 20 JUL 1911 1 

JEPS 19722 I SOTYPE 

NY HOLOTY PE 

MEAD11 DEWEYqC.9 AMER. J. SCI. ARTS SER.1, 43:9@. 1842. 

USA: ILLINOIS: HANCOCK COO Z AUGUSTA (MEAD9S.B.p --- . ---I 

CAS 553885 ISOTYPE

NUMBER 12 

320- 

321. 

322. 

323. 

324. 

325. 

32b- 

327. 

GH 

YO 

NY 

HOLOTY PE 

I SOTYPE 

ISOTYPE 

MEDITERRANIA YACKENZIETK~K-T BULL. TORREY BOT. CLUB 33:441. 1906. 

USA: DISTRICT OF COLUMBIA: WASHINGTON (STEELE,E.S., ---. 

23 MAY 1898) 

NY TYPE 

MEEKII DEWEYTC., AMER. J. SCI. ARTS SER.29 24:48. 1857. 

USA: NEBRASKA: WHIT€ RIVER (HAYDEN,F.V., --- . ---I 


MELANOPHORA BLAKE,S.T., J- ARNOLD ARBOR- 28: 106. 1947. 

INDONESIA: WEST NEW GUINEA: ORANGE RANGE, MOUNT WILHELMINA; 

(COUNTRY AS "DUTCH NEW GUINEA") ( BRASS, L. J 

MEYER-DREES~E~T 9828. -- SEP 1938) 

A I SOTYPE 

AND 

MELOZITNENSIS PORSILD,A.E., RHODORA 41:209. 1939. 

USA: ALASKA: KOKRINES MOUNTAINS YELOZ ITNA RIVER (PORSI LD, A. E 

AND PORSILDTR.T.T 713. 23 JUN-05 JUL 1926) 

GH I SOTYPE 


MENDOCINENSIS OLNEY 9S.T. EX BOOTTTW. IN WATSONvS.9 

GEOL. SURV. CALIFORNIA, BOT. 2:249. 1880. 

USA: CALIFORNIA: YENDOCINO COO: MENDOCINO CITY (BOLANDERTH.N. T 

4701. -- --- 1866 1 

CAS 553875 TYPE FRAGMENT 


GH HOLOTY PE 

YO I SOTYPE 

NY I SOTYP E 

uc 1098 ISOTYPE 


MERCARENSIS HOCHSTETTER ct OF EX STEUDEL TE .Go T SYN. PL. GLUM. 2 :194. 

1855. 

INDIA: --: NILAGIRI (HOHENACKERTR.F.T 943. -- --- 18511 

A I SOT YP E 

YERRILLII KUKENTHALTG.~ REPERT. SP. NOV. REGNI VEG. 8:7. 1910. 

PHILIPPINES: BENGUET: LUZON (ISLAND), PAUAI (MERRILL,E.D., 

6623. -0 MAY 1909) 



MERRITT-FERNALDII MACKENZIE?K.K., BULL. TORREY BOT. CLUB 49:370. 

1923 

USA: MAIVE: PENOBSCOT COO: ORONO (FERNALD,M.L., ---- 

03 JUL 1897) 

GH HOLO TYP E 

71

72 

328. 

329. 

330 

331, 

332. 

333, 

336, 

335. 

336, 

3370 

3380 


MESOCHOREA MACKENZIE,K.K. y BULL. TORREY BOT. CLUB 37:246. 1910. 

USA: DISTRICT OF COLUMBIA: WASHINGTON (STEELEvEoSo, ---a 

-- -_- 1900 1 

GH ISOTYPE 

MICANS BOOTTTF. I N GRAYvA.9 MEMO AMER. ACAD. ARTS NOS., 6:419. 

1859. 

JAPAN: --: SINODA (WRIGHTTC.T --- . ---I 


MfCRANTHA KUKENTHALTG., BULL. HERB. BOISSIER SER.2, 2:1018. 1902. 

KOREA: --: KAM-OUEN (FAURIE9U.T 919. 28 JUN 19011 


MICROCHAETA HOLM,H.T., AMER. J. SCI. SER.49 17:305. 1904. 

CANADA: YUKON TERRITORY: KLONDIKE INDIAN DIVIDE (MACOUN, JOHN9 

53877. 14 AUG 1902) 

GH I SOTYPE 

HICROGLOCHIN SSPm FUEGINA KUKENTHALqG., BOT. JAHRB. SYST. 27:546. 

1899. 

CHILE: MAGALLANES: TIERRA DEL FUEGO ("FUEGIA"1t ORANGE HARBOR 

I W I LKE S E XPLOR. EXPED. T --- . -- --- 1838-1 842 1 


MfCROPTERA YACKENZI E,K OK T YUHL ENBERGIA 5 : 56 1909. 

USA: NEVADA: 

CAS 

ELK0 COO: DEETH (HELLERTA~A.~ 9067. 


21 JUL 1908) 

NY TYPE 

MICROPTERA VARm CRASSINERVIA HERMANN,FoJ., RHODORA 70:420. 1968. 

USA: COLORADO: OURAY COO: ENGINEER PASS (JOHNSONTW.M.T 594. 

14 AUG 19671 

us 2 543 807 HOLOTY PE 

MILIARIS VARm AUREA BAILEYTL.H.T MEMO TORREY BOT. CLUB 1:37. 

CANADA: NEW BRUNSWICK: KING'S Coo: KENNEBECASIS RIVER 

(FOWLERIJ~T --- . ---I 

1889. 

GH HOLOTY PE 

MIRABILfS DEWEYTC.~ AMER. J. SCI. ARTS SER.1, 30:63. 1836. 

USA: MASSACHUSETTS: FRANKLIN COO: DEERFI EbD (DEWEYTC~T --- ---. 

) 

GH HOLOTY PE 

MIRABILIS VAR. PERLONGA FERNALD~M.L.~ PROC. AMER. ACAD. ARTS 

37:473, PL.2. 1902. 

USA: NEW HAMPSHIRE: HILLSBORO COO: NEW IPSWICH (FERNALDvM.L.9 

---. 05 JUN 1896) 

GH SYNTY PE 

MIRABILIS VARm TINCTA FERNALDTY.L*T PROC. AMER. ACAD. ARTS 37:473.

NUMBER 12 73 

339, 

340 

341 

3420 

343, 

344, 

34s. 

346, 

347. 

1902. 

CANADA: NEU BRUNSWICK: SAINT JOHN RIVER (MACOUNtJOHN, 22. 

04 JUL 1899) 

GH SYNTYPE 

MISANDROIDES FERNALI3,M.L.r RHODORA 17: 158. 1915. 

CANADA: NEWFOUNDLAND: PORT AU PORT BAY, TABLE MOUNTAIN 

(ST.JOHN,H. AND FERNALD9M.L. T 10801. 16-17 JUL 1914) 

GH HOLOTY PE 

NY I SOTY PE 

MISERA BUCKLEY,SoB., AMER. J. SCI. ARTS SER.19 45:173. 1843. 

USA: NORTH CAROLINA: MITCHELL COO: ROAN MOUNTAIN (BUCKLEY cS.B. 9 

--- . ---I 

NY I SOTYPE 

MISERABILIS YACKENZ1ErK.K.r Ne AMER. FL. 18:385. 1935. 

USA: WASHINGTON: CHELAN C O O Z CHIWAUKUM LAKE (EGGLESTON, Wow. 9 

13567. 19-20 AUG 1916) 


MOHRIANA YACKENZIE,K.K.9 No AMER. FL. 18:106. 1931. 

USA: FLORIDA: HARDEE C O O : WAUCHULA (CURTISSrA.H.1 6761. 

15 APR 1901) 

NY 

us 2133195 

HOLOTY PE 

I SOTYPE 

MOLESTA MACKENZIE9K.K. 9 N. AMEQ. FL. 18:151. 1931. 

USA: KANSAS: WYANDOTTE COO: QUINDARO (YACKENZIEqK*K., 

30 MAY 18971 

---. 

NY HOLOTYP E 

HONTANENSIS BA1LEYpL.H.r BOT. GAZ. 17~152. 1892. 

USA: MONTANA: FLATHEAD COO: UPPER MARAIS PASS (CANBY,W.M., 350. 

03 AUG 18831 

NY SYNTYPE 


MONTEREVENSIS YACKENZIE~K.K.T ERYTHEA 8:92. 1922. 

USA: CALIFORNIA: MONTEREY COO: PACIFIC GROVE (SMITH,C.P.t 1055. 

24 JUL 1905) 

GH HOLOTY PE 

MORRISSEYI PORSILD,A.E.~ SARGENTIA 4:21. 1943. 

CANADA: NEWFOUWDLAND: LABRADOR9 CAPE MUGFORD (PORSILD,AoE. 

173. 26 AUG 1937) 


MULTICOSTATA MACKENZIE9K.K. T BULL. TORREY 801. CLUB 43:604. 1916. 

USA: CALIFORNIA: SAN BERNARDINO COO: SAN BERNARDINO MOUNTAINS 

BEAR VALLEY DAY (PARISHTS.B.T 3609. -- JUN 1895) 

0s 4894C9 HOLOTYPE


348. MURICULATA HERMANN9F.J. IN MCVAUGHTR., FIELD t LAB. 17:132. 1949. 

USA: TEXAS: CULBERSON C O O : GUADALUPE MOUNTAINS? MCKITTRICK 

CANYON (M0OREpJ.A. AND STEYERMARK?J.A., 3625. 25 JUL 1931) 


349. NANA BOOTT,F. IN GRAYtAot MEMO AMER. ACAD. ARTS NOS., 6:418. 1859. 

JAPAN: HOKKAIDO (PREFECTURE): HAKODATE (WRIGHTTC. 9 ---. 

-- _-- 1853-1 856 1 

NY 

us 27280 

TYPE MATERIAL 

TYPE MATERIAL 

350. NEBRASKENSIS DEWEYTC.~ AYER. J. SCI. ARTS SER.2, 18:102. 1854. 

USA: NEBR9SKA: -- ( HAY0ENtF.V. T --- . ---I 

NY I SOTY P E 

351. NEBRASKENSIS VARI ERUCAEFORMIS SUKSD0RFjW.N.r WERDENDA 1:5. 1923. 

USA: WASHINGTON: KLICKITAT COO: FALCON VALLEY (SUKSDORF,W.N., 

10249. 22 JUN 1919) 

0s 171453 ISOTYPE 

MO 952735 TYPE COLLECTION 

NY ISOTYPE 


3520 NEBRASKENSIS VARo ULTRIFORMIS BAILEYTL.H.T BOT. GAZ. 21:8. 1896. 

USA: WASHINGTON: ADAMS COO: RITZVILLE (SANDBERG?J.H. AND 

LEIBERGTJ*B.T 194. 09 JUN 1893) 

NY I SOTYPE 

3530 NELSON11 MACKENZ1EtK.K. IN RYDBERG,P.A.v FL. ROCKY MOUNT. 

137t 1060. 1917. 

USA: WYOMING: LA PLATA MINES (NELSONtA. AND NELSONtE., 5264. 

30 AUG 1898) 

GH ISOTYPE 

NY HOLOTY PE 

354. X NEOBIGELOWII LEPAGETE., NATURALISTE CANAD. 91:166. 1964. 

CANADA: QUEBEC: SAGUENAY COO: ROMAINE RIVER (DUTILLYqA. AND 

LEPAGE,E.? 41,305A. 12 AUG 1963) 

GH I SOTYPE 

YY ISOTYPE 

us 243371 9 I SOTYPE 

3550 X NEOFILIPENDULA LEPAGEtE.9 NATURALISTE CANAD. 83:123. 1956. 

CANADA: NEWFOUNDLAND: BONNE BAY? MAIN RIVER, MAIN ARM 

(FERNAL0tM.L.; LONGTB. AND FOGG-JR. 9JoMm~ 1449. 19 AUG 1929) 

GH HOLOTYPE 

356- NEOMEXICANA YACKENZIE*K.K., BULL. TORREY BOT. CLUB 34:153. 1907. 

USA: NEW MEXICO: SANTA RITA DEL COBRA (BIGELOWTJ.M.T 1547. 

- N-

NUMBER 12 75 

357, 

358. 

359, 

360 - 

361 

362 0 

363. 

3640 

365 0 


X NEOPALEACEA LEPAGETE~T NATURALISTE CANAD. 83: 137. 1956. 

CANADA: QUEBEC: PAINT HILLS (DUTILLY, A.; 

DUMANvM.9 32975. 03 SEP 19541 

LEPAGETE. AND 

GH 

us 2176496 

I SOTY P E 

ISOTYPE 

NERVINA BAILEYTL.H. T BOT. GAZ. 10:203. 1885. 

USA: CALIFORNIA: SIERRA NEVADA RANGE9 SUMMIT CAMP (KELLOGGTA. 9 

--- 10 JUL 1870) 

GH HOLOTY PE 


NEUROPHORA MACKENZIE9K.K. IN ABRAYS9L.9 ILL. FL. PACIFIC STATES 

122989 FIG.706. 1923. 

USA: WASHINGTON: CHELAN C O O : CASCADE MOUNTAINS, STEVENS PASS 

(SANDBERG,J.H. AND LEIBERG,J.Bo, 773. 18 AUG 1893) 

GH I SOTYPE 

NY I SOTYP E 

NIGRICANS MEYER9C.A.9 

MEY. ACAD. IMP. SCI. ST.-PETERSBOURG DIVERS SAVANS 122109 PL.7. 

1831. 

USA: ALASKA: ALEUTIAN ISLANDS? UNALASKA ( ISLAND) 

(CHAYISSO9L.A.9 --- . -0-1 

GH I SOTYPE 

NIGRO-HARGINATA SCHWEINITZTL.D.T ANN. 

1:68. 1824. 

USA: NORTH CAROLINA: FORSYTH COO z 

(D0StL.t -a- . ---I 

LYCEUM NATO HIST. NEW YORK 

WINSTON-SALEM ("SALEM") 


NOVAE-ANGLIAE SCHWEINITZtL. Do 7 ANN. LYCEUM NATO HIST. NEW YORK 

1t67. 1824. 

USA: MASSACHUSETTS: BERKSHIRE COO: WILLIAMSTOWN, SADDLE 

MOUNTAIN (DEWEYTC.~ *-- . 

GH I SOTYPE 

-- --- 1822) 

X NUBENS LEPAGETE~T NATURALISTE CANAD. 84:40. 1957. 

CANADA: QUEBEC: JAMES BAY, EASTMAIN (LEPAGETE. T 33131. 

27 JUL 1955) 

GH I SOTYPE 

NUBICOLA MACKENZIE9K.K.9 BULL. TORREY BOT. CLUB 36:480. 

USA: COLORADO: MINERAL COO t PAGOSA PEAK ( BAKER,C.F., 

-- AUG 1899) 

1909. 

232. 

GH I SOTY P E 

NUDATA BOOTTTW. IN WATSON9S.q GEOL. SURV. CALIFORNIA9 BOT. 2z241. 

1880.

76 

USA: CALIFORNIA: MARISS (BOLANDERTHON., 

-- -_- 1860-1 867) 

2299. 



366. NUDATA VAR. FXRHIOR KUKENfHALqG. IN ENGLER9H.G. A. 9 PFLANZENR. 

4 9 FAM*20:337. 1909. 

USA: ARIZONA: WILLOW SPRINGS (PALMERvE.9 546. 0- JUN 1890) 

DS I SOTYPE 

367. NUTANS VAR. JAPONICA FRANCHETTA. AND SAVATIER ,LO 9 ENUM. PL. JAP. 

2:154. 1879. 

JAPAN: KANAGAWA (PREFECTURE) : HONSHU ISLAND) 9 YOKOSUKA 

(SAVATIERvL.9 1404. -- --- 1866- 1874) 


368. NUTTALLII DEWEYTC., AMER. J. SCI. ARTS SEft.1~ 43:92. 

USA: --: ROCKY MOUNTAINS (NUTTALLTT~T 17. ---I 

1842. 

GH HOLOTYPE 

NY I SOTYPE 

369. OAXACANA BAILEY vL.H.9 BOT. GAZ. 25:271. 1898. 

MEXICO: OAXACA: SIERRA DE SAN FELIPE (PRINGLE,C.Gov 4842. 

29 AUG 1894) 

GH I SOTY PE 

YO 

us 

us 

251772 

817656 




370. OBISPOENSIS STACEY1J.W. 9 LEAFL. W. BOT, 1 :240. 1936. 

USA: CALIFORNIA: SAN LUIS OBISPO COO: SAN LUIS OBISPO, STEINER 

CREEK (EASTWOODTA. AND H0WELLvJ.T.r 2271. 07 MAY 1936) 




DS 

F 

270930 

866418 

I SOTYPE 

ISOTYPE 

GH ISOTYPE 

NY 

us 1678188 

I SOTYPE 

ISOTYPE 

371. OBLANCEOLATA KOYAHAtT., WILLDENOWIA 5(3):489. 1969. 

CHINA: KWANGTUNG: CHUNG TUNGv TAI TSANG, YING TAK (TAK9T.W. 

AND CHOW,W*K.( 3202. 20 NOV 1926) 

uc 31 9673 HOLOTYPE 

372. OBOVOXDEA CRONQUISTvA. 9 MADRONO 7 :78. 1943. 

USA: IDAHO: CUSTER COO: STANLEY (CRONQUISTTA. T 2872. 

03 JUL 1941) 

GH I SOTYPE 

-0-

NUMBER 12 77 

373, 

374. 

375, 

376, 

377. 

378. 

379, 

380. 

381. 

YO 1220830 TYPE YATERIAL 

OEDERI VARI ROUSSEAUIANA MARIE-VICTORIN,(FRERE), 

PROC. & TRANS. ROY. SOC. CANADA SER.3, 23(2), SECT.5:262. 

1929 

CANADA: QUEBEC: MONTMAGNY COO: L'ESTUAIRE DU ST. LAURENT, 

BERTHIER-EN-BAS (ROUSSEAU9J. T 24989. 27 JUL 19261 

NY TYPE 

OKLAHOMENSIS YACKENZIE9K.K. T TORREYA 14: 126. 1914. 

USA: OKLAHOMA: CATALE (BUSH,B.Fm, 993. 22 MAY 1895) 


NY TYPE 

OLIGANTHA BOOTT,F., ILL. GENUS CAREX 4:174, PL.589. 1867. 

CHILE: YAGALLANES: TIERRA DEL FUEGO ("FUEG1A")t ORANGE HARBOR 

(WILKES EXPLOR. EXPEDoq --- . -- --- 1838-1 842 1 


OLlGOCARPA VARo LATIFOLIA GRAYvA. EX TORREY,J.t 

ANN. LYCEUM NATO HIST. NEW YORK 32415. 1836. 

USA: NEW YORK: JEFFERSON COO: WATERTOWN (CRAWE,J.B.t --- . ---I 

GH H OL 0 TY P E 

OLYYPICA MACKENZIE,K.K., BULL. TORREY BOT. CLUB 43:610. 1916. 

USA: WASHINGTON: CLALLAY COO : OLYMPIC MOUNTAINS (ELMER, A.D. E. 9 

2700. -- JUN 1900) 

NY SYNTYPE 

ONUSTA MACKENZIE,K.K.r BULL. 

USA: TEXAS: TARRANT C O O : 

TORREY BOT. 

-- (RUTHfA.9 

CLUB 42:618. 1915. 

458. 24 APR 1914) 


NY 

us 504456 

TYPE 

I SOTYPE 

OREGONENSIS OLNEYTS~T. EX BAILEY,L.H.T PROC. AMER. ACAD. ARTS 

22:73. 1886 ("1887") 

USA: OREGON: -- (HALLvE.9 605. -- --- 18711 


NY SYNTYPE 

ORHOSTACHYA WIEGANDTK.M.~ RHODORA 24:196. 1922, 

USA: NEW YAMPSHIRE: GRAFTON COO 2 FRANCONIA, LITTLETON HILL 

(FAXONpE. AND FAXON,C.E*t --- 27 MAY 1896) 

GH HOLOTYP E 

ORONENSIS FERNALD9Y.L 09 PROC. AMER. ACAD. ARTS 

37:471, PL.1, FIG.15,16. 1902. 

USA: MAINE: PENOBSCOT COO: ORONO (FERNALD,M*L.T ---a 

30 JUN 1891) 

GH HOLOTY PE 

NY I SOTYPE


3820 OXYCARPA HOLYtH.T., AMER. J. SCI. SER.49 20:303. 1905. 

USA: WASHINGTON: KLICKITAT C O O : COLUMBIA (SUKSD0RFtW.N.t 816. 

02 JUN 1885) 




3830 OXYLEPIS VAR. PUBESCENS UNDERW0ODtJ.K.t AMER. MIDL. NATURALIST 

33 :635. 1945. 

USA: TENNESSEE: CHEATHAM C O O : PEGRAM (SVENSON,H.K.t 10469. 

12 JUL 1939) 

NY TY PE COLLECT I ON 

384- PACHVCARPA YACKENZIETK.K.V BULL. TORREY BOT. CLUB 43:616. 1916. 

USA: CALIFORNIA: ALPINE C O O : SILVER VALLEY9 BIG TREE ROAD; 

ALTO 8000 FT. (BREWER9W.H.t 1977. 31 JUL 1863) 


385. PACHYSTACHYA CHAM1SSOtL.A. EX STEUDEL9E.G.v SYN. PL. GLUM. 2:197. 

1855. 

USA: ALASKA: ALEUTIAN ISLANDS9 UNALASKA (ISLAND) 

(CHAYI SSOt L *A* T --- . ---I 

GH I SOTYPE 

3860 PACHVSTOMA H0LMtH.T.t AMER. J. SCI. SER.49 20:302. 1905. 

USA: OREGON: KLAMATH COO: CRATER LAKE NATIONAL PARK? ANNA 

CREEK CANYON (C0VILLEtF.V.t 1362. 03 SEP 1902) 


3870 PADDOENSIS SUKSD0RFtW.N.t ALLG. BOT. 2. SYST. 12:43. 1906. 

USA: WASHINGTON: YAKIYA C O O : MOUNT ADAMS ("PADDO") 

(SUKSDORF tW.N. 9 1296. 13 AUG 1897) 



GH I SOTY PE 

NY 

us 529528 

I SOTYPE 

TYPE MATERIAL 

3880 PALAWANENSIS KUKENTHALtG. IN ELMERtA.D.E.9 LEAFL. PHILIPP. BOT. 

4:1169. 1911 

PHILIPPINES: PALAWAN! PUERTO PRINCESA t MOUNT PULGAR 

(ELMERtA.D.E.9 13146. -- MAY 1911) 

GH I SOTYP E 

NY 

us 872800 

TYPE MATERIAL 

TYPE MATERIAL 

3890 PALLESCENS VARo NEOGAEA FERNALDtt4.L.t RHODORA 44:306t PL.712. 

1942 

CANADA: NEWFOUNDLAND: GANDER RIVER VALLEY 9 GLENWOOD 

- P-

NUMBER 12 79 

390. 

391 

392 

393. 

396. 

395. 

396, 

397, 

398, 

(FERNALDtMoL. AND WIEGANDtKoM.? 4918. 12-13 JUL 1911) 


PANSA BAILEYTL~H~T BOT. GAZO 13:82. 1888. 

USA: OREGON: CLATSOP COO: CLATSOP (HENDERSONTL.~ 0) 

31 JUL 1886-20 AUG 1887) 

1482. 

DS 490462 SYNTYPE 

NY SYNTYP E 

PAPULOSA BOOTTpF. 

1859 

I N GRAYTA~T MEMO AMERo ACAD. ARTS NOSOT 6:418. 

-- -4- 

JAPAN: HOKKAIDO (PREFECTURE): HAKODATE (WRIGHT9C.p ---. 

1853- 1856 1 

GH HOLOTYPE 

NY ISOTYPE 


PARCIFLORA BOOTTtF., MEMO AMER. ACAD. ARTS NOS., 6:418. 18590 

JAPAN: HOKKAIDO (PREFECTURE): HAKOOATE (WRIGHTTC.? --- 0 ---I 


PARRYANA DEWEYtC.9 AMER. J. SCI. ARTS SER.11 27:239. 1835. 

CINADA: --: HUDSON BAY (RICHARDSON9J.t --- ---I 


X PATUENSfS LEPAGEtE.9 NATURALISTE CANAD. 89:113~ FIG.1. 1962. 

CANADA: QUEBEC: UNGAVA BAY? LAKE PATU (DUTILLYtA. AND 

LEPAGEtEo? 39329: 19 AUG 1961) 

GH I: SOTYPE 

PAUCICOSTATA MACKENZIEtK.K., ERYTHEA 8:74. 1922. 

USA: CALIFORNIA: MARIPOSA COO: YOSEYITE NATIONAL PARK9 

YOSEMITE VALLEY (BOLANDER?H.N., 6198. -- JUL 1866) 

0s 49738 TYPE COLLECTION 


YO TYPE YATERI AL 


PAUPERCULA VAR, BREVISQUAMA FERNALD9M.L.q RHODORA 20:152. 19180 

CANADA: QUEBEC: ILE AUX COUDRES (MARIE-VICTORIN,(FRERE), 4021. 

-- JUN 19171 

GH HOLOTYPE 

PAUPERCULA VAR. PALCENS FERNALD? M.C. T RHODORA 8:77. 19060 

USA: MAINE: OXFORD COO: BUCKFIELD (ALLEN? J0A.c 21A. 

01 JUL 1878) 

GH HOLOTYPE 

PAYSONIS CLOKEYTIOW.~ AMERo J. SCI. SER.59 3:90~ PL.2. 1922. 

USA: WYOMING: TETON COO: GRAND TETON NATIONAL PARK, JACKSON 

HOLE VALLEY (PAYS0NtE.B. AND PAYSONTL.B., 2224. 06 AUG 1920) 

GH I SOTY PE 

uc 905434 HOLOTYPE

80 

399. 

400 a 

401 a 

402 a 

403 I. 

404. 

605. 

406 0 

407 

ShfITHSONIAN COSTRIBUTIONS TO BOTANY 

PELOCARPA HERMANNTF~J.~ RHODORA 39:492. 1937. 

USA: UTAH: SUYMIT C O O : LAMOTTE PEAK (HERMANNT F.J 0 9 5983. 

15 AUG 1933) 


NY HOLOTYPE 

PERCOSTATA HERMANNTF~J~T J. WASH. ACAD. SCI. 40:282. 1950. 

MEXICO: CHIHUAHUA: MADERA (MULLERTCml4.r 3520. 27 SEP 1939) 

CAS 

us 

369422 

2133207 

ISOTYPE 

TYPE 

PERGLOBOSA MACKENZ I E I KO K. 9 BULL. TORREY BOT. CLUB 34: 606 1907 

USA: COLORADO: SUMMIT C O O : MOUNT BALDY, 

(MACKENZIETK.K.T 167. -- AUG 1901) 

BRECKENRIDGE 


NY HOLOTYP E 

PERILEIA BLAKEIS~T-T J. ARNOLD ARBOR. 28: 102. 1947. 

INDONESIA: WEST NEW GUINEA: LAKE HABBEMA; (COUNTRY AS "DUTCH 

NEW GUINEA") (BRASS,L.J.p 9583. 1- AUG 19381 

A I S OTY P E 

PERLONGA FERNALDTM.L.T PROC. AYER. ACAD. ARTS 43:61. 1907. 

MEXICO: HIDALGO: TRINXDAD IRON WORKS; ALTO 1585 M e 

(PRINGLETC~G., 8863. 02 JUN 1904) 


CAS 1930d5 I SOTYPE 


GH HOLOTY PE 

YO I SOTYPE 

NY ISOTYPE 


PERSTRICTA YACKENZIETK.K.T BULL. TORREY BOT. CLUB 36:479. 19C9. 

MEXICO: NUEVO LEON: MONTERREY (PRINGLE,C.G., 2630. 

05 JUN 1889) 

GH I SOTYPE 

NY TYPE 

PETASATA DEWEYIC., AMER. J. SCI. ARTS SER.1, 29:246. 1836. 

USA: --: ROCKY MOUNTAINS (DRUMMONDTT.T --- . ---I 

NY TYPE 

PETRICOSA DEWEYTC., AMER. J. SCI. ARTS SER.19 29:246. 1836. 

USA: --• ROCKY MOUNTAINS ( DRUMYONDT To T --- ---I 

NY TYPE 

PETRIEI CHEESEMANTT.F.T TRANS. & PROC. NEW ZEALAND INST. 16:413. 

1884. 

NEW ZEALAND: CANTERBURY (DISTRICT): SOUTH ISLAND, BROKEN RIVER 

(CMEESEMANTT.F. T --- -- JAN 1883) 

GH I SOTYPE

NUMBER 12 81 

408. 

409. 

410. 

411. 

412 

413. 

414. 

415. 

416. 

PHAEOCEPHALA PIPER9C.V.9 CONTR. U.S. NATL. HERB. 11:172. 1906. 

USA: OREGON: CLACKAMAS Coo: MOUNT HOOD (HALLtE.9 583. 

01 AUG 1871) 




YO TY PE COLL ECT I ON 


PHAEOLEPfS HOLM,H.T., AMER. J. SCI. SER.49 17:302. 1904. 

USA: OREGON: CROOK COO: BEAR BUTTES (LEIBERGqJoB.9 335. 

26 JUN 1894) 

NY I SOTYPE 

PHALAROIDES VARo PARVULA GROSS9 R. 9 REPERT, SP. NOVo REGNI VEG. 

50 :211. 1941 

ASGENTINA: JUJUY: TILCARA DEPT.? TILCARA (VENTUR1,S.v 6491. 

15 FEB 1927) 

us 1545831 TYPE 

PHILOCRENA KRECZETOWICZTVOI 0 9 

TRUDY SREDNE-AZIATSK. GOSUD. UNIV.9 SER. 8B9 BOT. 17:75. 1934. 

USSR: TADZHIKISTAN: PAYIRS; ALTO 82CC FT. (LIPSKY,V.I.T 2732. 

19 JUL 1899) 


PHYLLOMANICA BOOTTTW. IN WATSON, S.9 GEOL. SURV. CALIFORNIA, BOT. 

2:233. 1880. 

USA: CALIFORNIA: MENDOCINO COO: MENDOCINO CITY (BULANDER9H.N.9 

4746. -- --- 1866 1 

GH HOLOTY P E 

MO I SOTYPE 

NY ISOTYPE 

PHYSOCHLAENA HOLM9H.T.9 AMER. J. SCI. SER.4, 17:317. 1904. 

USA: ALASKA: YUKON VALLEY9 COAL CREEK HILL (FUNSTON9F.9 139. 

30 JUL 1893) 


MO 920815 TYPE COLLECTION 


PICTA BOOTT9F. IN GRAY9A.9 MEMO AMER. ACAD. ARTS N.S.9 

1859. 

JAPAN: HOKKAIDO 4 PREFECTURE): HAKOOATE (WRIGHTTC. 9 


6:418. 

--- . ---I 

PICTA STEUDEL,E.G., SYN. PL. GLUM. 2m4. ia55. 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMOND,T.q 

---) 

---. 


PINETORUM VAR. ELAf!OR KUKENTHALvG. IN ENGLER,H.G.A.T PFLANZENR.

a2 SMITHSONIAN CONTRIBUTIONS TO BOTANY 

417. 

418. 

419. 

420. 

421 . 

422 

423 

424. 

42s. 

4 9 FAM.20: 195. 1909. 

MEXICO: OAXACA: SIERRA DE SAN FELIPEt ALTO 10~000 FT. 

(PRINGLETC~G~T 4685. 08 JUN 1894) 

GH I SOTYP E 

NY ISOTYPE 

PIPER1 MACKENZIETK.K* IN P1PERqC.V. AND BEATTIETRoK.~ FLo NW. COAST 

750 1915. 

CANADA? BRITISH COLUMBIA: VANCOUVER ISLAND, CEDAR HILL 

(MACOUNTJOHNT --- 31 MAY 1887) 

GH I SOTYP E 

PIRCHINCHENSIS VARm SIMPLEX GROSSvR.9 REPERT. SPo NOV. REGNI VEG. 

50:211* 1941. 

COLOMBIA: --: -- (MUTIS,JoCot KILLIP NO.5715. 

_- --_ 1760-1888 I 

us 1563811 TYPE 

PITYOPHILA MACKENZIETK.K.~ BULL. TORREY BOT. CLUB 40:545. 1913. 

USA: NEW MEXICO: RIO ARRIBA C0.Z TIERRA AMARILLA 

(EGGLESTON, We We I 6605 0 18 APR-25 MAY 1911 1 




PLANATA FRANCHETvA. AND SAVATIER*Lov ENUM. PL. JAP. 2:126r 555. 

1879 0 

JAPAN: KANAGAWA (PREFECTURE): HONSHU ( ISLAND) T YOKOSUKA 

(SAVATIERTL., 20590 -- --- 1866-1874) 


PLATVLEPIS MACKENZIEqK.Ko, No AMERo FLo 18:1420 1931. 

USA: WYOMING: BIG HORN CO.: BIG HORN MOUNTAINS, TEN SLEEP LAKES 

(WILLIAMS,T.A., 2951. 19 AUG 1897) 

h(Y HOLOTY PE 

PLATYPHYLLA CAREYTJ., AMERo J o SCI. ARTS SER.29 4:23, 1847. 

USA: NEW YORK: ("NOV. EBOR.") (CAREYTJ~T ---I 


PLECTOCARPA HERMANNTFoJoT LEAFL. Wo BOT. 10:66* 1964. 

USA: MONTANA: GLACIER CO.: GLACIER NATIONAL PARK, LOGAN PASS, 

HIDDEN LAKE; 

21 AUG 1962) 

ALTO 7300 FTo (HERMANNTF.JOT 18120. 

CAS 

us 

41 6360 

2420276 

I SOTYPE 

HOLOTYPE 

PLUVlCA VARI KOOLAUENSIS KRAUSSvRo T PACIFIC SCI 4: 274. 1950. 

USA: HAWAII: HONOLULU CO.: OAHU (ISLAND), KOOLAU MOUNTAINS 

(HOSAKATE~YOT 594. 

us 2074725 

04 JUL 1932) 

TYPE MATERIAL 

--- 0 

PODOCARPA BROWN,R. IN RICHARDSON,Jo IN FRANKLINVJ.9

NUMBER 12 

426. 

427. 

428. 

429. 

430 

431. 

432. 

433 . 

434 

435. 

NARR. JOURNEY POLAR SEA 751. 1819. 

CANADA: --: -- (R1CHARDSONtJ.t --- ---I 


POOOGYNA FRANCHETtA. AND SAVAT1ERtL.t ENUM. PL. JAP. 2:131t 557. 

1879 

JAPAN: TOKYO (PREFECTURE): TOKYO (SAVAT1ERqL.t 1413. 

-- --- 1866- 1874 I 


PORTER1 0LNEYtS.T.t CARIC. BORo-AMER. 12. 1871. 

USA: MAINE: PISCATAQUIS COO: MOUNT KINEOt MOOSEHEAD LAKE 

( P 0 R T E R 9 T C t --- 28 AUG 1871) 

GH HOLOTY P E 

POTOSINA HEMSLEY9W.B.t BIOL. CENTRO-AMER. 3t474. 1885. 

MEXICO: SAN LUIS POTOSI: SAN LUIS POTOSI (SCHAFFNERtJeGot 546. 

-- --- 1877) 

GH ISOTYPE 

PRAECEPTORIUW MACKENZ1EtK.K.t No AMER. FL. 18:95. 1931. 

USA: WASHINGTON: KLICKITAT C O O : SIMCOE MOUNTAINS, GOLDENDALE 

(PECKpM.E.7 13. 13 AUG 1917) 

NY HOLOTY PE 

PRAEGRACILIS B0OTTtW.t BOT. GAZ. 9:87. 1884. 

-- --- 

USA: CALIFORNIA: SAN DIEGO COO: SAN DIEGO (SCOTTt(MISSI, ---a 

1880) 

GH HOLOTYPE 

PRAINII CLARKE,C.B.t 

CHINA: YUNNAN: 

J. LINN. SOC~C 801. 36:305. 

RED RIVER (HENRYtAot 10839. 

1904. 

---I 

MO 

us 458108 

TYPE MATERIAL 

TYPE MATERIAL 

PRAIREA DEWEYtC. IN W0ODtA.t CLASS-BOOK 601. 414. 1845. 

USA: MICHIGAN: -- (---t -9-0 ---I 

GH HOLOTYPE 

PRATENSIS DREJERtSoToN., NATURHIST. TIDSSKR. 3:442. 1841. 

GREENLAND: --: -- (VAHLtJot --- ---I 


PRATENSIS VARe FURVA BA1LEYtL.H. IN MACOUNTJOHN, CAT. CANADIAN PL. 

5:377. 1890. 

CANADA: BRITISH COLUMBIA: VANCOUVER ISLAND, CEDAR HILL 

(MACOUNtJOHNt ---. 3 1 MAY 1887) 

GH I SOTYP E 

PRATICOLA VAR. SUBCORIACEA HERMANNtF.J.t LEAFC. W. BOT. 8:113. 

1957. 

CANADA: ALBERTA: MOUNTAIN PARK, MCCLEOD RIVER (HERMANNtFoJ. 

13453. 26 AUG 1956)


4360 

437. 

438- 

439, 

440 

441 

442. 

443. 



PREISSII ESSENSACH,N.VON IN LEHYANNTJ.T PL. 

AUSTRALIA: --: -- (PREISSTL.T 1825. ---I 

PREISS. 2:94. 1846. 

GH SYNTYPE 

AUSTRALIA: WESTERN AUSTRAL I A: 

-- JUL 18391 

PERTH (PRE ISSTL. T 1861. 

YO 2002968 SYNTYPE 

PRESLII STEUDELTE.G.T SYN. PL. GLUY. 2:242. 1855. 

USA: ALASKA: NUTK,A SOUND ("SINUS NUTKA") (HAENKETT., --- . ---I 


PRINGLEI BAILEYTL.H.T BOT. FAZ. 17:151. 1892. 

MEXICO: SAN LUIS POTOSI: HACIENDA DE ANGUSTURA, 100 MILES EAST 

OF SAN LUIS POTOSI (PRINGLETC.G.~ 3801. 04 AUG 18913 







PRIONPHYLLA HOLM,H.T. T AMER J. SCI SER. 49 14:423. 1902. 

USA: IDAHO: DIVIDE BETWEEN SAINT JOE AND CLEARWATER RIVERS 

(LEIBERGTJ~B~T 125. 10 JUL 1895) 

r\lY TYPE COLLECTION 

PROJECTA MACKENZIE~K~K~T BULL. TORREY BOT. CLUB 35:264. 1908. 

CANADA: NEW BRUNSWICK: KENT COO: KOUCHIBOUGUAC (FOWLERTJ.T 

-- --- 

18721 

GH ISOTYPE 


PROPOSITA YACKENZIE T K O K. I No AMER. FL. 18: 126 1931. 

USA: IDAHO: BLAINE COO: SMOKY MOUNTAINS; ALTO 2700 Ma (MACBRIDE9J.F. AND PAYSON*E.B., 3778. 13 AUG 1916) 


GH I SOTYP E 

NY HOLOTY PE 

X PSEUDO-FULVA FERNALDpM.L., RHODORA 35:231. 1933. 

CANADA: NEWFOUNDLAND: PORT AU PORT BAY9 TABLE MOUNTAIN 

(FERNALD9M.L. AND WIEGAND~K~M~T 4258. 16 AUG 1910) 

GH HOLOTYPE 

PSEUDOJAPONICA CLARKE9C.B. T BULL. MISC. INFORM. ADD.SERo8:81. 

1908. 

USA: CALIFORNIA: NEVADA COO: SIERRA NEVADA RANGE, DONNER LAKE; 

ALTO 2750 M q (HELLER,A.A.T 7187. 17 AUG 1903) 


NY TYPE COLLECTION

SUMBER 12 85 

444 

445 .I 

446 

467 

4480 

449. 

450 

451 

4520 

PTEROLEPTA FRANCHETTA.T NOUV. ARCH. MUS. HIST. NATO SER.39 8?215. 

1896 

CHINA: YUNNAN: -- (DELAVAYTR.P.T 4829. 15 JUL 1889) 

US 1123683 ISOTYPE 

PTYCHOCARPA STEUDEL,E.G., SYN. PL. GLUM. 2:234. 1855. 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMONDqT., 

-- --_ 1832) 

424. 


QURPUREOVAGINATA VARo ITATIAIAE GROSSTR., 

REPERT. SP. NOV. REGNI VEG. 50:212. 1941. 

BRAZIL: RIO DE JANEIRO: ITATIAIA (CHASETA., 8283. 17 JAN 1925) 

!IS 1282178 TYPE 

PURPURIFERA YACKENZIETK.K.T No AMER. FL. 182253. 1935. 

USA: TENNESSEE: CAMPBELL COO: CHASKA (BRIGHTTJ., ---a 

18 MAY 1923) 


PYCNOTHYSOS KUKENTHALTG~T PHILIPPo J. SCI. 6:60. 1911. 

PHILIPPINES: NEGROS OCCIDENTAL: MOUNT CANLAON (VOLCANO) 

(MERRILLTE.D., 543. r.- APR 1910) 


QUADRIFIDA BAILEY,L.H.T PROC. CALIF. ACAD. SCI. SER.2, 3: 104. 

189 1 

USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA, TUOLUMNE RIVER 

(ROLANDER,H.Y. T 5046. -- --- 1866) 


NY SYNTYP E 

QUADRIFIDA VARo LENIS BAILEYTL-HoT PROC. CALIF. ACAD. SCI. 

SER.2, 3:105. 1891. 

USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA, 

(BOLANDERTH~N~T 5046. -- --- 1866) 

TUOLUMNE RIVER 

NY SYNTYPE 

-Q- 

X QUEBECENSIS LEPAGEtE.9 NATURALISTE CANAD. 91:168. 1964. 

CANADA: QUEBEC: SAGUENAY COO: ROMAINE RIVER (DUTILLYtA. AND 

LEPAGE9E.r 41,305. 12 AUG 1963) 

GH I SUTYP E 

NY I SOTYP E 

us 243371 8 I SOTYPE 

QUICHENSIS HERMANNTF~J~T 

J. WASH. ACADo SCI. 40:284. 1950. 

GUATEMALA: QUICHE: NEBAJ (SHARP, A. J. T 45144. 07 FEB 1945) 

us 2133193 TYPE

86 

-R- 


453. RACHILLIS YAGUIREt B.9 BRITTONIA 5: 199. 1944. 

USA: UTAH: GILBERT PEAK (MAGUIREpB. AND MAGUIRE,R.R., 14668. 

16 AUG 1936) 


NY TYPE 


4540 RAMOSII KUKENTHAL~G.T REPERT. SP. NOV. REGNI VEG. 8:8. 1910. 

PHILIPPINES: RIZAL: LUZON ( ISLANDIT MORONG (RAMOSpM.9 

BUR. SCI. 1434. -- AUG 1906) 


4551 RAYNOLOSII DEWEYTC., AMER. J. SCI. ARTS SER.29 32:39. 1861. 

USA: IDAHO: FREMONT C O O : PIERRE'S HOLE AND HENRY'S FORK 

t HAY D EN 9 F V 9 -0- 20 JUN 1860) 

GH SYNTY P E 

456. RETROCURVA DEWEYTC. IN WOODyA., CLASS-BOOK BOT. 423. 1845. 

USA: NEW YORK: JEFFERSON COO: -- (WOODqW.A.9 --- . ---I 

GH HOLOTYPE 

451. RETROCURVA VAR. COPULATA BAILEYTL.H.T HERB. DISTR. N0.161. 1886. 

USA: MICHIGAN: INGHAM COO: LANSING (BAILEYTL.H.T 161. 

01 JUN 1886) 

GH I S OTY P E 

458. RETROFLEXA MUHLENBERGTH. EX WILLDENOWTC.L.T SP. PC. ED041 4:235. 

1805. 

USA: TEXAS: -- (DRUMYONDTT*T --- ---I 

NY TYPE 

459. RHYNCHACHAENIUM CLARKEvC. Be IN MERRILL p E. 0.9 

PUBL. BUR. SCI. GOV. LAB. 35:5. 1906 ("1905"). 

PHILIPPINES: BATAAN: MOUNT MARIVELES; ALTO 1100 Me 

(ELMERTA.D.E.T 6983. -- NOV 1904) 

MY COTYPE 

460. RICHAROSONII BROWNTR. IN RICHARDSONTJ. IN FRANKLINyJ.9 

NARR. JOURNEY POLAR SEA 751. 1819. 

CANADA: --: -- ( R I C HARD SON T J T --- . ---I 


461. RfCHAROSONII FOR. EXSERTA FERNALD,M.L., RHODORA 44:290. 1942. 

USA: ILLINOIS: HANCOCK COO: AUGUSTA (MEADTS~B~T . ---I 

GH HOLOTYP E 

462. ROANENSIS HERYANN~F~J~T 

CASTANEA 12:113. 1948 ("1947"). 

USA: TENNESSEE: CARTER C O O : ROAN MOUNTAIN: ALT.4700 FT.

NUMBER 12 

(BR0WNtD.M.t 255. 02 AUG 1936) 

us 213319C HOLOTYPE 

463. RORAIWENSIS STEYERMARK, J.A. t FIELDIANAt BOT. 28:67t FIG.7. 19510 

VENEZUELA: BOLIVAR: MOUNT SORAIMA; ALTO 2700-2740 M. 

(STEY ERMARK t J A. 7 5887C 28 SEP 1944) 


NY I SOTYPE 

464. ROSAEOIOES HOW€( EoC. IN GORDINIERtHoCo AND H0WEtE.C. 9 

FL. RENSSELAER COO 33. 1894. 

USA: NEW YORK: RENSSELAER COO: LANSINGBURGH (HOWE9E.C.t ---a 

30 MAY 1887) 

NY I SOTY PE 

465. ROSEA VARo ARKANSANA BA1LEYtLoH.t BOT. GAZ. 13:87. 1888. 

USA: ARKANSAS: PULASKI COO: LITTLE ROCK, LA FOURCHE CREEK 

(HASSEtHoEot --- 0 01 MAY 1886) 

GH 

NY 

HOLOTY P E 

I SOTYP E 

466. ROSEA VARo PUSILLA PECKtCsHmt ANNUAL REP. NEW YORK STATE MUS. 

48 132. 1895. 

USA: NEW YORK: LIVINGSTONE COO: PORTAGE (PECKtCoHot ---. 

-- JUN 18941 

NY TYPE 

467- ROSEA VAR. STAMINATA PECK,C.H.t ANNUAL REP. NEW YORK STATE MUS. 

47:164. 1894. 

USA: NEW YORK: OTSEGO Coo: COOPERSTOWN JUNCTION (PECK9C.H.t 5. 

-- JUN 1893) 

NY TYPE 

468. RUBRO-BRUNNEA VAR. ELINEOLATA MERR1LLtE.D.t LINGNAN SCI. J. 13818. 

1934. 

CHINA: KHANGTUNG: LOH-FAU-SHAN (MOUNTAIN) 9 POK-LO (TSU1tT.M.t 

74. -- MAR-APR 1932) 

A I SOTYPE 

GH I SOTYPE 

MO 1260436 ISOTYPE 

NY TYPE 


469. RUGATA FERNALDt MoL t RHODORA 43: 5459 PLo 671. 1941. 

USA: VIRGINIA: SUSSEX COO: HOMEVILLE (FERNALD9M.L. AND LONGtBo t 

117870 

CAS 

F 

07 MAY 1940) 

336836 I SOTYP E 


GH HOLOTYP E 


NY 

us 

ISOTYPE 


87


470. RUGATA OHWI TJO T ACTA PHYTOTAX. GEOBOT. 1:76. 1932. 

JAPAN: --• HONSHU (ISLAND), MOUNT DAIMONJI IN YAMASHIRO 

(OHWI TJO T 29. 04 MAY 1931) 


4710 RUGOSPERMA MACKENZIETK~K., BULL. TORREY BOT. CLUB 42:621, 1915. 

USA: NEW JERSEY: 

-- MAY 1911) 

OCEAN Coo: TUCYERTON (MACKENZIETK~K~T 98710 

NY TYPE 

4720 RUSBYI MACKEYZIETKOK*T SMITHSONIAN MISC. COLLECT. 65(7):20 1915. 

USA: ARIZONA: YAVAPAI COO: -- (RUSBY,H.H., 859. -- --- 1883) 

NY TYPE 

us 30267 TYPE COLLECT1 ON 

4730 RUTH11 MACKENZIETK.K.T 

--_ 0 -- 

N o AMER. FL. 18:112. 1931. 

USA: NORTH CAROL1 NA: BUNCOMBE Corn: CRAGGY MOUNTAIN (RUTH, A. 

JUL 1900) 

9 

YY HOLOTY P E 

474, SAL!NAEFORHIS MACKENZIETK.K.T BULL. TORREY BOT. CLUB 36:477. 19090 

USA: CALIFORNIA: MENDOCINO Coo: MENDOCINO CITY (BOLANDERTHON. T 

4702. -- --- 1866 1 


DS 293480 TYPE COLLECT ION 






6750 SALTAENSIS GROSSTROT REPERT. SPo NOV. REGNI VEGo 50:211. 19410 

ARGENTINA: SALTA: ROSARIO DEPTo: CAMP0 QUIJANO (VENTURITS.T 

8650. -- --- 1929) 

us 1545752 TYPE 

476. SANGUINEA BOOTTTF.T TRANS. LINN. SOC. LONDON 20:137. 1846. 

AFGHANISTAN: --: T- (GRIFFITHTW.T 96(KEW 6094). ---I 


477. SARAWAKETENSIS KUKENTHALTG~T BOT. JAHRBo SYST. 69:2620 19380 

PAPUA AND NEW GUINEA: NORTH-EAST NEW GUINEA (TERRITORY): 

MOROBE DISTRICT: FIN1 STERRE RANGE, MOUNT SARAWAKET 

(CLEMENSTJ. AND CLEMENSTYOS~T 5546. -- MAR 1937) 

A ISOTYPE 

4780 SARTHELLIANA OLNEY9S.T. IN GRAYPA., PROC. AMER. ACAD. ARTS 7:396. 

1868. 

USA: CALIFORNIA: YARIPOSA COO: YOSEYITE NATIONAL PARK, 

-S-

NUMBER 12 89 

4790 

480. 

481 

482 

4830 

4840 

485. 

4860 

481 . 

YOSEMITE VALLEY (BREWER,W.Hoq 1636. -- --- 1863) 

GH HO LOTY P E 

SARTWELLII DEWEYTC.~ AMER. J. SCI. ARTS SER.1, 43:90. 1842. 

USA: NEW YORK: SENECA COO: JUNIUS (SARTWELLIH~P~T 12. ---I 


CAS 553975 TYPE COLLECT ION 


SAVAIIENSIS KUKENTHAL9G.q BERNICE P. BISHOP MUS. BULL. 128:24. 

1935. 

WESTERN SAMOA: --: SAVAII ISLAND (CHRISTOPHERSEN, E.9 800. 

24 SEP 1929) 

NY I SOTY PE 

X SAXENII NMo FERRUGINEA LEPAGE9E.9 NATURALISTE CANAD. 83:142. 

1956. 

CANADA: QUEBEC: JAMES BAY9 FORT GEORGE (DUTILLY9A.G LEPAGE9E. 

AND DUYAN9M.9 32357. 14 AUG 1954) 

GH I SOTYPE 


SAXIMONTANA YACKENZIETKOK-T BULL. TORREY BOT. CLUB 33:439. 1906. 

_-- ) 

USA: COLORADO: LARIMER C O O : FORT COLLINS (BAKER,C.F., ---. 

MO 

NY 

TYPE MATERIAL 

TYPE MATERIAL 

SCABRIOR SARTWELL9H.P. EX DEWEY9C.9 AMER. J* SCI. ARTS 

SER.2, 8:349* 1849. 

USA: NEW YORK: YATES COO: PENN YAN (SARTWELLfH.P.9 72. 

CA S 553877 I SOTYPE 

SCABRIUSCULA MACKENZIEIK~K~T BULL. TOQREY BOT. CLUB 35:268. 1908. 

USA: WASHINGTON: CASCADE MOUNTAINS (CUSICK9W.C. 9 2849. 

30 JUN 1902) 


YO ISOTYPE 

NY 

us 52 8631 

TYPE 

I SOTYPE 

SCAPOSA HOOKERTJ.D.T BOT. MAG. PL.6940. 1887. 

CHINA: KWANGTUVG: LOH-FAU-SHAN (MOUNTAIN) ('FORDTC~T -- --- 

---. 

18831) 


SCHAFFNERI BOOTTrW. IN WATSON,S., PROC. AMER. ACAD. ARTS 18:172. 

1883. 

MEXICO: SAN LUI S POTOSI : SAN LUIS POTOSI (SCHAFFNER. JOG. 9 -- --- 1877 1 

546. 

GH ISOTYPE 

SCHNEIDERI NELMES,E.y BULL. MISC. INFORM. 201. 1939. 

---)


488. 

489. 

490. 

491 

492 

493 

494. 

495. 

496. 

CHINA: YUNNAN: LIKIANG (SCHNEIDERIC~T 2738. 06 AUG 1914) 

GH HOLOTY PE 

SCHUEINITZII OEWEYTC-T AMER. J. SCI. ARTS SER.19 9268. 1825. 

USA: MASSACHUSETTS: BERKSHIRE COO : WILLIAMSTOWN (DEWEYvC. 9 ---. 

---) 

GH HOLOTYP E 

SCIRPIFORHIS YACKENZIE?K.K., BULL. TORREY BOT. CLUB 35:270. 1908. 

CANADA: ALBERTA: BANFF NATIONAL PARK? BANFF (MCCALLA9W.C. 9 

2348. 28 JUL 1899) 

rYY TYPE 

SCIRPOIDEA VAR. GIGAS HOLMTH.T.T AMER. J. SCI. SER.4, 18Z20. 

USA: CALIFORNIA: SISKIYOU COO: MOUNT EDDY (PRINGLE?C.G., 

19 AUG 1881) 

1904. ---. 



SCIRPOIDEA VAR. STENOCHLAENA HOLMTH.T.T AMER. J. SCI. SER.49 18:20. 

1904. 

CANADA: BRITISH COLUMBIA: CHILLIWACK VALLEY (MACOUNT J.M. T 

33728. 12 JUL 1901) 


SCOPARIA VARo CONOENSA FERNALD9M.L.t PROC. AMER. ACAD. ARTS 

37:468? PL.1. 1902. 

USA: MASSACHUSETTS: MIDDLESEX C0.2 MEDFORD (BOOTTTW.T ---. 

26 JUL 1865) 

GH HOLOTY PE 

SCOPARIA VAR. FULVA BOOTTvW. IN WATSONTS., 

GEOL. SURV. CALIFORNIA? BOT. 2:237. 1880. 

USA: CALIFORNIA: ALPINE COO: SILVER VALLEY (6REWERtW.H.t 1969. 

31 JUL 1863) 

GH SYNTYPE 

SCOPARIA VAR. MINOR BOOTT,F., ILL. GENUS CAREX 3:116~ PL.369. 

1862 

USA: NEW HAMPSHIRE: WHITE MOUNTAINS (TUCKERMANTE.~ --- ---I 


SCOPARIA VARo UONILIFORMIS TUCKERMANTE.? ENUM. CARIC. 17. 1843. 

USA: MASSACHUSETTS: MIDDLESEX CO.: CAMBRIDGE (TUCKERMAN,E.r 

--_- ---_ 1 

GH HOLOTY P E 

SCOPARIA FOR. PfRACUlA FERNALD, M.L.9 RHODORA 23:234. 1921. 

CANADA: NOVA SCOTIA: YARMOUTH COO : SAND 8 EACH ( FERNALD? M. L o 

AND LONGVB.T 20296. 14 JUL 1920) 

GH HOLOTY PE 

497. SCOPARIA VARo SUBfURBINAfA FERWALD1M.L. AND WTEGANDTK.M.T RHODORA

NUMBER 12 91 

14:116. 1912. 

CANADA: NEWFOUNDLAND: EXPLOITS RIVER VALLEY, GRAND FALLS AREA, 

RUSHY POND (FERNALD9M.L. AND WIEGANDTK. Me T 4796. 

28 AUG 19111 


GH HOLO TY PE 

NY ISOTYPE 

SCOPARIA VARo TESSELLATA FERNALDTM~L. AND WIEGANDTK*M.~ RHODORA 

12: 135. 1910. 

USA: MAINE: WASHINGTON COO: PEYBROKE (FERNALDTM~L.~ 1464. 

C8 JUL 19091 

GH HOLOTYPE 

NY ISOTYPE 

SCOULERI TORREYTJ., ANN. LYCEUM NAT. HIST. NEW YORK 3:399. 1836. 

USA: BRITISH COLUMBIA: "WESTERN SHORE OF AMERICA, OBSERVATORY 

INLET" (ARM OF PORTLAND INLET) (SC0ULERtJ.q 296. ---I 

NY HOLOTY P E 

SEATONIANA BAILEYTL.H., BOT. GAZ. 25327C. 1898. 

MEXICO: HIDALGO: TULA (PRINGLETC.G.T 7452. 24 JUN 18971 


GH I SOTYPE 

us 30 5734 TYPE C OLLEC T I ON 

SHELDON11 MACKENZ1EvKmK.r BULL. TORREY 601. CLUB 42:618. 1915, 

USA: OREGON: UNION Coo: CLARK'S CREEK (SHELDON,EoP., 8854. 

09 SEP 1897) 

NY I SOTYP E 

NY TYPE 


SICCATA DEWEYTC~T AMER. 

USA: MASSACHUSETTS: 

---) 

J. SCI. ARTS SER.19 1@:278o 1826. 

HAMPDEN COO: WESTFIELD 4 DAVIS,Ee ---o 


SIMULATA MACKENZIE,K.K., BULL. TORREY BOTo CLUB 34:604. 1907. 

USA: WYOMING: ALBANY CO.: CHUG CREEK (NELSON,&., 7316. 

29 JUN 1900) 

NY HOLO TYPE 

SMALLIANA YACKENZIETK~K. T BULL. TORREY BOT. CLUB 36:484. 1909. 

USA: GEORGIA: TATTNALL COO: REIDSVILLE (HARPERTR~M.T 2159. 

26 APR 1904) 


SONOMENSIS STACEYqSeW., LEAFL. We 601. 2:63* 1937. 

USA: CALIFORNIA: SONOMA CO.: SEBASTOPOL, PITKIN MARSH 

(HOWELLTJ.~. AND STACEYTJ.W.~ 13042. 06 JUN 19371 


CAS 246636 I SOTYPE


506- 

507. 

5080 

509. 

5130 

DS 258275 TSOTYPE 



GH I SOTYP E 

NY 

uc 

us 

835699 

1736782 

I SOTY PE 

I SOTYPE 

ISOTYPE 

SOPERI RAUPTH.M.T SARGENTIA 6:129, FIG.12. 1947. 

CANADA: NORTHWEST TERRITORIES: MACKENZIE DISTRICT: BRINTELL 

LAKE (R4UPtH.M. AND SOPERTJ~H., 9534. 18 JUL 1939) 

GH HOLOTY P E 

SPECIFICA BAILEY,L.H.r MEMO TORREY BOT. CLUB 1:21. 1889. 

USA: CALIFORNIA: ALPINE C O O 2 SILVER VALLEY (BREWERtWoHot 1969. 

31 JUL 1863) 



SPECUICOLA HOWELL, J .To t LEAFL. W. BOT. 5: 148. 1949. 

USA: ARIZONA: COCONINO COO: INSCRIPTION HOUSE (HOWELL~J~T~T 

24609. 23 JUN 1948) 


CAS 342553 HOLOTY PE 


GH ISOTYPE 

NY ISOTYPE 

us 2C06386 I SOTYPE 

SPRETA BAILEYTL~H., MEMO TORREY BOT. CLUB 1:6. 1889. 

USA: OREGON: MULTNOMAH C O O : SAUVIE ISLAND (COLUMBIA RIVER AT 

MOUTH OF WILLAMETTE RIVER) (HOWELLTT~J~T --- -- MAY 1880) 



STANDLEYANA STEYERMARK, J. A. 9 CEIBA 3:23. 1952. 

GUATEMALA: JALAPA: AGUACATE (WILL1AMSyL.O.t 13178. 

06 JUL 1947) 


STANTONENSIS JONES,M.E. 9 BULL. MONTANA STATE UN1V.t BIOL. SER. 

15:2Cy PL.3. 1910. 

USA: MONTANA: STANTON LAKE ( WILLIAMS,R.S.', --- 11 AUG 1894) 

NY TYPE 

STELtATA M4CKENZIE,K.K.~ No AMER. FL. 18:226. 1935. 

MEXICO: HIDALGO: IXMIQUILPAN (ROSE, J. No i PAINTER, JoHm AND 

ROS€TJ.S. 9 9019. -- --- 1905) 

NY I SOTYPE 


STELLULATA VAR. CONFERfA CHAPYAN,A.W., FL. S o U.S. ED.1, 534. 

1860.

NUMBER 12 93 

USA: FLORIDA: GADSDEN COO: -- (CHAPMANpA.W.9 --- . ---I 

NY TYPE 

514. STENOPHYCLA VARI DESERTORUH LITVINOVTD.I.T ALLGO BOT. Z. SYST. 

5( BEIH. 1) : 56. 1899. 

USSR: TUSKESTAN: -- (LITVINOV,b.I., 1530 -- --- 1897) 


515. STENOPTERA MACKENZIE9K.K.9 ERYTHEA 8:28. 1922. 

USA: CALIFORNIA: COS ANGELES COO: SAN ANTONIO MOUNTAINS9 ICE 

HOUSE CANYON (JOHNSTONTI.M.T 1505. 31 JUL 1917) 


NY TYPE 

5160 STERILIS VAR. EXCELSIOR BAILEYTL OH. T BULL TORREY BOT. CLUB 20:424. 

1893. 

USA: NEW YORK: SENECA C O O : JUNIUS (SARTWELL9H.P.9 35. ---I 

NY I SOTY P E 

517. STEUDELII KUNTHTC~S. 9 ENUM. PL. 2:480. 1837. 

USA: OHIO: MXAMI RIVER VALLEY (FRANK9J.C. 9 --- . -- --- 1835) 


518. STEVERHARKII STANDLEYTP.C.T PUBL. FIELD MUS. NAT. HIST.? BOT. SER. 

23: 196. 1947. 

GUATEMALA: HUEHUETENANGO: CERRO HUI TZ (STEYERMARK, J. A. 9 48542 

14 JUL 1942) 


519. STIPATA VAR. LAEVIVAGINATA KUKENTHALvG. IN ENGLERvHeG.A.9 

PFLANZENR. 4 9 FAM.20: 172. 1909. 

USA: NORTH CAROL1 NA: BUNCOYBE COO: BILTMORE (NEAR ASHEVILLEI 

(BILTMORE HERBARIUM9 262A. 28 MAY 1897) 


GH TYPE COLLECT I ON 

MO TYPE COLLECTION 


52 8 ST ATA VAR. NAXIHA CHAPMAN9A.W. EX BOOTT9F.9 ILL. GENUS CAREX 

3:1219 PL.391. 1862. 

USA: FLORIDA: APPALACHICOLA (CHAPYAN,A.W., --- . -0-1 


521. STIPATA VARo SUBSECUTA PECKTC.H.T ANNUAL REP. NEW YORK STATE MUS. 

48: 128. 1895. 

USA: NEW YORK: RENSSELAER COO: BERLIN (PECK,C.H.p ---a 

-- JUN 1894) 

WY TYPE COLLECTION 

522. STIPATA VAR. UBERIOR MOHR9C.p CONTR. U.S. 

USA: ALABAMA: MOBILE RIVER (MOHRvC.9 

NATL. HERB. 6:417. 

--- 26 APR 1897) 

1910. 

NY I SOTYPE

94 


5230 STRAHINEA VARo CUMULATA BAILEY, L .He T YEY. TORREY BOT. CLUB 1: 23. 

1889. 

CANADA: NEW BRUNSWICK: KENT COO: -0 (FOWLERTJ., ---a 

-0 JUL 1870) 

GH SY NTY P E 

YO SY NT YPE 

NY SYNTYP E 

STRANINEA VARo ECHINODES FERNALD,M.L., PROC. AMER. ACAD. ARTS 

37:474, PL.20 1902. 

CANADA: ONTARIO: LAYBTON COO: WYOMING (MACOUNvJOHN, 26624. 

24 JUN 1901) 

GH SYNTYPE 

STRAWINEA VARo RENIFORMIS BAILEY,L.H., MEM. TORREY BOT. CLUB 1~73. 

1889 

USA: MISSISSIPPI: OKTIBBEHA COO: STARKVILLE (TRACY,S.M.t 17. 

23 MAY 1888) 

NY SYNTYPE 

STRICTA VARo XEROCARPA BRITTON,N.L., BULL. TORREY BOT. CLUB 22:222. 

1895. 

USA: NEW YORK: STEUBEN COO: PRATTSBURGH (WRIGHT,S.H., .--. 

---) 


STYLOFLEXA BUCKLEYtS.B., AMER. J. SCI. ARTS SER.1, 45:174. 1843. 

USA: NORTH CAROLINA: MACON COO: MOUNTAINS (BUCKLEY,S.B., ---a 

--- ) 



STYLOSA VARe VlRENS BAILEYpLoH., PROC. AMER. ACAD. ARTS 22:79. 

1886 ("1887"). 

USA: OREGON: MULTNOMAH COO: SAUVIE ISLAND (COLUMBIA RIVER AT 

MOUTH OF WILLAMETTE RIVER) (HOWELLIT~J., --- -- MAY 1880) 

GH SYNTYPE 

YO SYNTYPE 

SUB-BRACTEATA MACKENZIE,Y.K., BULL. TORREY BOf. CLUB 43:612. 1916. 

USA: CALIFORNIA: ALAYEDA COO: OAKLAND (BOLANDER,H.N.r ---. 

_- _-- 1860 CA.1 

GH ISOTYPE 

NY HOLOTY PE 


SUBFUSCA ROOTTTW. IN WATSONTS., GEOL. SURV. CALIFORNIA, BOT. 2:234. 

1880. 

USA: CALIFORNIA: LAKE TAHOE TO BEAR VALLEY (KELLOGG,A., ---. 

---) 

GH ISOTYPE 

SUBIWPRESSA CLOKEY,I.W., RHODORA 21:84. 1919.

NUMBER 12 95 

USA: ILLINOIS: MACON COO: -- (CLOKEYTI.W.T 2338. 06 AUG 1915) 


uc 9054 33 HOLOTY P E 

5320 SUBNfGRICANS STACEYTJ.W.T LEAFL. W. BOT, 22167. 1939. 

USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA (HOWELLTJ~T~T 14519. 

11 AUG 1938) 

CAS 

us 

259816 

1765699 

HOLOTYPE 

ISOTYPE 

5330 SUBORBICULATA MACKENZIE9K.K. I N ABRAMS,L.r ILL, FL. PACIFIC STATES 

1:338. 1923. 

USA: WASHINGTON: KL ICKITAT COO : -- (SUKSDORF rW.N. T 1315. 

-- JUL 1883) 

NY TYPE 

534. SUBTRANSVERSA CLARKE9C.B. T PHILIPP. J. SC1 2:108. 1907. 

PHILIPPINES: BENGUET: LUZON I ISLAND), PAUAI (MERRILLqE.D.1 

473C. -9 OCT-NOV 1905) 

YY TYPE YATERIAL 


5350 SUKSDORFII KUKENTHALTG., REPERT. SP. NOV. REGNI VEG. 16:434. 

USA: WASHINGTON: YAKI MA CO. : MOUNT ADAMS ( loPADDO1g 1 

1920. 

(SUKSDORFTW-N., 7383. 27 AUG 1912) 

CAS 

CAS 

152864 

243333 

ISOTYPE 

I SOTYPE 

DS 

0s 



GH I SOTYPE 


NY 

us 1437926 

1 SOTYPE 


5360 SUKSDORFII VARo OVALIS KUKENTHALqG., REPERT. SP. NOV. REGNI VEG. 

16 :434. 1920. 

USA: WASHINGTON: YAKI MA COO : MOUNT AOAMS ( "PADDO") 

(SUKSDORF,W.N., 5259. 21 AUG-20 SEP 1905) 


NY SYNTYPE 

537. SURCULOSA RAYMONDTM.T MEMO JARD. BOT. MONTREAL 52:22. 1959. 

CHINA: KUANGTUNG: SIN-FUNG DISTRICT: SAI-LIN-SHAN VILLAGET 

NGOK SYING SHAN (TAAMTY.W.T 502. 1-16 APR 1938) 

A HOLOTYPE 

538. SYCHNOCEPHALA CAREYTJ~T AYER. J. SCI. ARTS SER.2, 4:24. 1847. 

USA: NEW YORK: JEFFERSON C O O : WATERTOWN (CAREYTJ~T --- . ---I 

GH I SOTYPE

96 

-T- 


5390 TACHIRENSIS STEYERMARKtJoAot FIELDIANA, BOT. 28:68t FIG080 1951. 

VENEZUELA: TACHIRA: COLOMBI AN-VENEZUELAN BOUNDARY t PARAMO DE 

TAMA; ALTO 3045-3475 M. (STEYERMARK9J.A.t 573670 

15 JUL 1944) 



5400 TAHITENSIS BROWNTF.B.H.~ OCCAS. PAP. BERNICE PAUAHI BISHOP MUS. 

9(4):8. 1930. 

FRENCH POLYNESIA: --: SOCIETY ISLANDS t TAHITI t MOUNT OROHENA 

(MACDAYIELStL .Hot 1542. 15 YAY 1927) 

A I SOTY P E 

561. TAHAKXI K0YAMAtT.t BULL. ARTS SCI. DIV. RYUKYU UNIV. 3:75. 1959. 

RYUKYU ISLANDS: OKINAWA (PREFECTURE): OKINAWA (ISLANDIT YONA 

OKINAWA (ISLAND), YONA EXPERIMENTAL FOREST OF RYUKYU 

UNIVERSITY; (COUNTRY AS "JAPAN") 

23 NOV 1958) 

(K0YAMAtT.t ---a 

NY HOLOTY PE 

542, TAHANA STEYERMARKTJ~A~T FIELDIANAt BOT. 28:70. 1951. 

VENEZUELA: TACHIRA: COLOMBIAN*VENEZUALAN BOUNDARY9 PARAMO DE 

TAMA; ALTO 3045-3475 Mo (STEYERYARKtJ0A.t 57401. 

15 JUL 1944) 


543. TENERA DEWEYTC.~ AMER. J. SCI. ARTS SER.19 8:97. 1824. 

USA: MASSACHUSETTS: BERKSHIRE COO: WILLIAMSTOWNt SADDLE 

MOUNTAIN (DEWEYtC. T --- 20 JUN ---- 1 

GH HOLO TY PE 

544. TENERA VARm RICH11 FERNALDtMoLo, PROCo AMER. ACAD. ARTS 

37:475T pL.2. 1902. 

USA: MASSACHUSETTS: YIDDLESEX COO : MIDDLESEX FALLS (R1CHtW.P. t 

--- 05 JUN 1894) 

GH HOL OTY P E 

5450 TENERAEFORWIS MACKENZIE,K.K.T BULL. TORREY BOT. CLUB 4326090 1916. 

USA: CALIFORNIA: BUTTE Coo: JONESVILLE; ALTO 5100 FT. 

(HALLpHoMot 97810 25 JUL 1914) 

NY HOLOTY P E 

546. TENTACULATA VARo ACTIOR BOOTTTF.T ILL. GENUS CAREX 2:94r PL.278. 

1860. 

USA: NEW YORK: YATES COO: PENN YAN (SARTWELLTH.P.~ 1380 ---I 


547- TENUIFLORA VARo SETACEA KUKENTHALtG. IN ENGLERTH~G~A~T PFLANZENR.

NUMBER 12 97 

548. 

549. 

550. 

5511 

552. 

553. 

554. 

555 . 

4 9 FAM.20:224. 1909. 

USA: MICHIGAN: INGHAM COO: LANSING (WHEELERtCoF. 9 

28 JUN 1890) 

GH I SOTYPE 

TERETIUSCULA VAR. AMPLA BA1LEYtL.H.t MEMO TORREY BOT. CLUB 1:53. 

1889 

USA: OREGON: BAKER C O O : HEAD OF BURNT RIVER (CUS1CKtW.C.t 1331. 

-- JUL 1886) 

NY SY NTY P E 

TERRAE-NOVAE FERNALDtMoL.? RHODORA 44:290, PL.711. 1942. 

CANADA: NEWFOUNDLAND: SAINT JOHN BAY? SAINT JOHN ISLAND 

(FERNAL0,M.L.t W1EGANDtK.M.; LONGyB.; GILBERT-JR.,F.A. AND 

HOTCHKISS,N., 27657. 31 JUL 1925) 

GH HOLOTYPE 

TETSUOI 0HWIpJ.q M I X . REP. NATL. SCI. MUS. 5:2t PL.2. 1952. 

RYUKYU ISLANDS: OKINAWA (PREFECTURE): OKINAWA (ISLAND); 

(COUNTRY AS "JAPAN") (AMAN0,T.t 6358. -- MAY 19511 


THURBERI DEWEYpC. IN TORREYtJ. IN EM0RYtW.H.t 

REP. U.S. MEX. BOUND. SURV.7 BOT. 2(11:232. 1859. 

MEXICO: SONORA: MABIBI (THURBER9G.t --- 0- JUN 1850) 

GH HOLOTY P E 

TOJQUIANENSIS STANDLEYtPoC. AND STEYERMARK,J.A.q CEIBA 4:64. 1953. 

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES, BETWEEN 

TOJQUIA AND CAXIN (STEYERMARKtJ*A.t 50150. 06 AUG 1942) 



TOHPKINSI H0WELLtJ.T.t LEAFL. W. BOT. 9:185. 1961. 

USA: CALIFORNIA: FRESNO COO: KINGS RIVER CANYON? COPPER CREEK 

TRAIL (H0WECLtJ.T.t 35333. 06 JUN 1960) 

CA S 428953 HOLOTYPE 


NY I SOTYPE 


TOREAOORA STEYERMARKT J.A.r PHYTOLOGIA 9:338* 1954, 

ECUADOR: AZUAY: TOREADOR: ALTO 3810-3930 Me (STEYERMARK,J.A., 

53095. 15 JUN 1943) 


NY I SOTYPE 

TORTA VAR. STAMINATA PECKpCeH., ANNUAL REP. NEW YORK STATE MUS. 

46: 131. 1893. 

USA: NEW YORK: ONEIDA COO: TABERG (PECKtC.H.9 ---. 

-- JUN 1892) 

NY TYPE COLLECTION

98 

556. 

557. 

558. 

559, 

560 o 

561. 

562. 

563 

5640 

565. 

566- 


TOWNSENDII MACKENZIE,K.K.t No AMER. FL. 18:111. 1931. 

MEXICO: CHIHUAHUA: COLONIA GARCIA; ALTO 2250 M. 

(TOWNSEND,C.H.T. AND BARBER~COM~T 157. 21 JUL 1899) 


NY 

us 568126 

I SOTYPE 

HOLOTYPE 

TRACY1 YACKENZIE,K.K.T ERYTHEA 8:41. 1922. 

USA: CALIFORNIA: HUMBOLDT COO: BALD MOUNTAIN (TRACYTJ~P~T 4547. 

04 JUL 1914) 

NY TYPE 

TRIANGULARIS BOECKELERTJ.~. 7 FLORA 39:226. 1856. 

USA: TEXAS: -- (0RUYMONDtT.T --- . ---I 

NY I SOTY P E 

TRIBULOIDES VARm SANGAMONENSIS CLOKEYtIoW., RHODORA ?1:84. 1919. 

USA: ILLTNOI~: MACON rn.: -- (CLnKFY.1-Wet 2364- 37 AUG 1915) 

uc 5,"3+41 LlnLOTYPE 

X TRICHINA FERYALD, H o L o t 9HOr)OSA 35z2l.9. 19330 

USA: MAINE: ARnOSTOOK CO.: FORT FAIRFIELO (WICLIAMS,E.F.; 

C~~~LINSTJ.F. ANn FERYILD~M*L.T IIC. I9 JUL 1902) 

GH HOLOTYPE 

TRICHOPHYLLA NELYESTE.~ MEMO YUS. NATL. HIST. NATO9 SER. Bt BOT. 

4:lCb. 1955. 

VIET-NAY? NORTH: 

foCTCLOTqDoA+ 

GH 

TONKIN: CHAPlJ: (rOUNTRY AS "TNDOCHINA") 

c324. -- JUI. :"?I 

HT)LOTYDE 

TRISPERYA DEWEYtC.9 AMER. J. SCI. 4qT5 SER.1, 9:ha. 

USA: MASSACHUSETTS: -- (DEWEYTC., -0- . ---I 

1825. 

NY 1 SOTY PE 

TRISPERWA VAR. BILLINGS11 KNIGH1,O.W. T RHOOORA 8 :185. 19C6. 

USA: M4INE: SOMERSET COO: PLEASANT RIDGE9 JEWETT BROOK BOG 

(WARE9R.A.t ROLLINStS. AlVD KVIGHTTO.W., 5066. G5 JUL 19C6) 

GH I SOTYPE 

TSOI MEqRILLT F.D. 4N3 CH?INvY.K. 9 SUYYATSEYIA 2:207. 1935. 

CHIYA: KW4NGTUNI;: YAIN4N (ISLAY@IT DlJhlG KA TO WEY FA SHI 

(CHUIVtY-K. 4Nr) TSOTC.L.T 43683. -- --- 1932-1 933 1 

GH I SnTYPE 

NY ry PF 

us 167'1 2C TVPE "4TESI 4L 

TUHULICOLA MACKENZ1EtK.K. T BULL. TORREY BOT. CLUB 34:154. 1907. 

USA: CALIFORNIA: ALAYEDA COO: LAKE TEYESCAC (BIOLETTI TF-T. T 1. 

25 JUN 18931 

NY HOLO TY P E 

TUNIHANENSIS STANDLEYvP .C. AND STEYERMARKT J. A. 9 CEIBA 4:65. 1953.

NUMBER 12 99 

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUYATANES, TUNIMA 

( STEYERMARK? J.A. T 48334. 07 JUL 1942) 

F 1128966 TYPE 

567. TURGESCENS TORREYTJ.~ ANN. LYCEUM NATO HIST. NEW YORK 3:419. 18360 

USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (INGALLSTTOT --to 

--- ) 

h(Y TYPE 

5680 TURUMIQUIRENSIS STEYERMARKtJoAoq FIELDIANA, BOTo 28:70, FIG.7. 

1951. 

VENEZUELA: SUCRE: CERRO TURUMIQUIRE, RIDGE DIVIOING HEADWATERS 

Of RIC) YANZANARES AND RIO DE AMANA; ALTO 1900-2000 Ma 

(STEYERMARKTJOA~T 62705. 10 YAY 1945) 


GH I SOTYPE 

NY 

us 1933688 

I SOTYPE 

I SOTY PE 

569. ULTRA BA1LEYtLoH.r PROC. AYERo ACAD. ARTS 228830 1886 ("1887"). 

USA: ARIZONA: COCHISE C O O : HUACHUCA ( LEMMON, J.G.9 2. 

21 JUN 1882) 


DS 64032 I SOTY P E 

5710 UNCOMPAHGRE KELSO9L.r BIOL. LEAFLO 38:l. 1947. 

USA: COLORADO: LA PLATA COO: LA PLATA YOUNTAINSt MOUNT 

HESPERUST GOLD KING MINE (KELSOTL., 6058. 03 JUL 1947) 

GH I SOTYPE 

572, UNOERHOODII BRITTDN*N.L.q TORREYA 5: 100 1905. 

JAMAICA: --: SALT HILL MARSH (UNDERWOOD,L.M., 158. 

29 JAN 19031 

YY TYPE 

573. UNILATERALIS MACKENZIETK~K~, 

ERYTHEA 8243. 1922. 

USA: CALIFORNIA: HUYBOLDT Coo: ALTON (TRACY, JoPot 3783. 

21 JUL 1912) 

NY TYPE 

- U-

100 

- V- 


576. VAGANS H0LMrH.T.~ AMER. J. SCI. SER.4, 17:301. 1904. 

USA: OREGON: HARNEY COO: STEEN MOUNTAIN, ANDREWS (LEIBERG,J.B.* 

2558. 10 JUL 1896) 

NY I SOTYP E 

5750 VAGINATA VAR. ALTO-CAULIS DEWEY,C.* AMER. J. SCI. ARTS 

SER 0 2 41: 227. 1866. 

USA: NEW YORK: GENESEE C0.Z BERGEN (PAINE,J.A., --to ---I 

GH HOLOTY PE 

576. VALLICOLA DEWEY,C.r AMER. J. SCI. ARTS SER.2, 32:40. 1861. 

USA: WYOYING: TETON C O O : SNAKE RIVER, JACKSON HOLE VALLEY 

(DEWEYIC.~ 1Q. 18 JUN 18601 

GH HOLOTY PE 

5770 VENOSIVAGINATA STANDLEY,PoC. AND STEYERMARK,J.A.T CEIBA 4267. 

1953. 

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES, CERRO 

HUTTZ, BETWEEN MIMANHUITZ AND YULHUITZ; ALTO 2600 M. 

(STEYERYARK,J.Ao, 48554. 14 JUL 1942) 

F 11 2 9094 HOLOTY PE 

GH I SOTYPE 

578. VERNACULA VAR. HOBSONII YAGUIRE,Boq BRITTONIA 5:199. 1944. 

USA: UTAH: BEAR RiVER RANGE, WHITE PINE LAKE (MAGUIRETB.~ 

H0BSONtD.A. AND MAGUIRE+R.R. 9 14013. 16 JUL 1936) 


GH ISOTYPE 

NY 

us 1872573 

HOLOTY PE 

ISOTYPE 

5?9. VESfCARIA VAR. JEJUNA FERNALD,M.L.t RHODORA 3:530 1901. 

USA: MAINE: AROOSTOOK COO: MADAWASKA LAKE (WILLIAMS,E.F.~ 

-- AUG 1900) 

---. 

GH HO LO TY P E 

5800 VESICARIA VARo LAURENTIANA FERNALDqY.L.9 RHODORA 35:232. 1933. 

CANADA: NEWFOUNDLAND: SAINT JOHN'S (FERNALD7M.L.; LONGvB. AND 

FOGG-JR.,J.M., 1474. 31 JUL 1929) 

GH HOLOTYPE 

us 205@647 I SOTYPE 

5810 VESTITA VAR. KENNEDY1 FERNALDIM~L~T RHODORA 2:170o 1900. 

USA: MASSACHUSETTS: MIDDLESEX COO: WILMINGTON, SILVER LAKE 

(KENNEDYIGSG.~ --- 11 JUN 1899) 

GH HOLOTYPE 

582. VEXANS HERYANNIF-J., RHODORA 57:156. 1955.

NUMBER 12 101 

USA: FLORIDA: HENDRY COO: CLEWISTON (DEAMTC.C.T 61177. 

19 M4R 1941) 

US 2231425 HOLOTYPE 

5830 VICARIA BAILEY9L.H. T HEM. TORREY BOT. CLUB 1:49. 1889. 

USA: OREGON: -- (HALLTE.T --- -- --.II 

r,H HOLOTYPE 

1871 1 

5840 VIOLACEA CLARKE,C.B., BULL. MISC. INFORM. ADD.SER.8:87. 1908. 

USA: CCILORADO: ROCKY MOUNTAINS; 141. 39-41 No (HALLTE. AND 

HARBOURTJ.P., 587. -- --- 1862) 

YO TY PE COLLECT ION 

5850 VIRIOlOR MACKENZ1EvK.K. IN ABRAHS,L.T ILL. FL. PACIFIC STATES 

1.: 331. 1923. 

USA: WASYINGTON: OKANOGAN COO: SHEEP MOUNTAINS ( EGGLEST0NvW.W. 9 

3329. 30 JUL-01 AUG 1916) 

IJ S 886234 TYPE 

5860 VlRIDULA FOR. PVGMAEA LEPAGE9E.p NATURALISTE CANAD. 89:115. 1962. 

CANADA: QUEBEC: UNGAVA BAY, SWAMPY BAY (DUTILLYTA. AND 

LEPAGE9E.r 39274. 16 AUG 1961) 

GH I SOTYP E 

5870 VITIENSIS ST.JOHNTH.~ PACIFIC SCI. 1:116~ FIG.1. 1947. 

FIJI: --: V ITI LEVU (ISLAND) (STIJOHNTH.T 18330. 18 AUG 1937) 


5880 VITREA HOLMTH.T.T AMER. J. SCI. SER.4, 17:302. 1904. 

USA: CALIFORNIA: RIVERSIDE COO: PALM SPRINGS (PARISHTS.B.~ 

4144. 04-13 APR 18961 

F 89120 TYPE HATERIAL 

YO ISOTYPE 


589- VULPINOlOEA VARo PYCNOCEPHALA HERMANN9F.J.v RHOOORA 38:363. 1936. 

USA: MICHIGAN: EMYET COO: BIG STONE BAY (HERHANNvFoJ., 64C8. 

14 AUG 1934) 

GH HOLOTYPE 

5900 UAHUENSIS VARo RUBlGfNOSA KRAUSSTR., PACIFIC SCI. 

4~2579 F16.2934-Do 1950. 

USA: HAWAII: KILAUEA IKI (BEANqR.S.t HOSAKATE~Y. AND 

ST.JOHNTH.T 11228. 21 DEC 1931) 


591. HATSONI OLNEY9S.T. IN WATSONTS., BOT. U.S. GEOL. EXPLOR. 40TH PAR. 

370. 1871. 

USA: NEVADA: ORYSBY COO: CARSON CITY (WATSONTS., 1246. 

-W-

102 

-- MAY 1868) 

GH HOLOTY PE 


5920 WEROERMANNIT GROSSTR. IN WERDERMAYNtE.1 

NOTIZBL. BOT. GART. BERLIN-DAHLEM lot7630 1929. 

CHILE: LLANQUIHUE: ALTO 700 M. (WERDERMANNTEOT 1687. 

-- MAR 1925) 

YY TYPE COLLECTION 

5930 WHIfNEYI OLNEY,S.T. IN GRAY,A., PROC. AMERo ACAD. ARTS 7:394. 

1868. 

USA: CALIFORNIA: TUOLUMNE COO: YOUNT DANA; ALTO 1200C FT. 

(BOLANDERTHON., 5086. -- --- 1866) 

YO SYNTYPE 

NY SYNTYPE 


YOSEYITE VALLEY (BOLANDERIH.N.~ 6198. -- --- 1866 1 



YO SYNTYPE 

NY SY NTYP E 

USA: CALIFORNIA: NEVADA COO: SODA SPRINGS; ACT. 9000 FT. 

(BREWERTU.H.T 1778. 04 JUL 1863) 

YO SYNTYP E 

NY SY NTY PE 

5940 WIEGANDII MACKENZIETK.K*T No AMER. FL. 18:lCS. 1931. 

CANADA: NEUFOUNDLAND: RAY OF ISLANDST HUMBER ARM, CURLING 

(FERNAL0,M.L. AND WIEGANDTK.M.T 2776. 21 JUL 1910) 

GH ISOTYPE 

5950 W1LKESII TORREYTJ. IN WILKESTC. 9 U.S. EXPLOR. EXPED. 17:477r PL.17. 

1854. 

USA: CALIFORNIA: SACRAMENTO RIVER (WILKES EXPLOR. EXPEDOT ---a 

-_ --- 1 8 3 8- 1 842 1 


596. WILLDENOVII VARm PAUCIFLORA 

0LNEYvS.T. EX BAILEY,LoH. IN COULTER9J.M.r 

CONTR. U.S. N4TL. HERB. 2:482. 1894. 

USA: TEXAS: HARRIS COO: HOUSTON (HALLTE.~ --- . -- -0- 

GH TYPE COLLECTION \ 

1872) 

597. WILLOENOWII VARI HEGARRHYNCHA HERYANNTFoJ.~ AMER. MIDL. NATURALIST 

51:277, 1954. 

USA: GEORGIA: JASPER CU.: OCMULGEE (SMITH,S.J. AND DUNCAN,W.H., 

4872. 06 APR 1949) 


598- WILLfAMSIT BRITTONtN*L., BULL. NEW YORK BOT. GARD. 2:159. 1901. 

CANADA: YUKON TERRITORY: DAUSON (WILtIAMS,R.S., ---a 

12 JUN 1899) 

NY TYPE

NUMBER 12 103 

5990 UOODII DEWEYTC., AMER. J. SCI. ARTS SER.2, 2:249. 1846. 

USA: NEW YORK: JEFFERSON COO: PERCH LAKE, PERCH RIVER 

(CRANE, 1.B. AND WOOD,W.A., ---.I. ---I 

GH HOLOTY PE 

600. WRIGHTI! DEWEYTC. I N TORREYTJ. I N EYORY,W.H., 

REP. U.S. MEX. BOUND. SURV.r BOT. 2(1):232. 1859. 

USA: TEXAS: -- (WRIGHT,C., 1561. -- --- 1850 1 


- X- 

601. X XANTHINA FERNALDTM.~., RHODORA 35:230. 1933. 

CANADA: NEWFOUNDLAND: MAIN RIVER (FERNALD,M.L. AND LONGTB.~ 

1455. 27 AUG 1929) 

GH HOLOTY PE 


602. XANTHOCARPA VAR. ANNECTANS BICKNELLTE.P.T BULL. TORREY BOT. CLUB 

23:23. 1896. 

USA: NEW YORK: LONG ISLAND, RICHMOND VALLEY (BRITTONTN.L.T ---. 

06 JUI. 1895) 

NY TYPE 

6630 XERANTICA BAILEYTL~H~T BOT. 642. 17:151. 1892. 

CAN4OA: SASKATCHEWAN: FILE HILLS; 50.5N. 9 1041. ( MACOUNTJOHN, 

--- 04 JUL 1879) 

GH SYNTYPE 

NY SYNTYP E 

604. XEROCARPA WRIGHTTSOH. IN DEWEYTC.? AMER. J. SCI. ARTS 

SER.21 42:334. 1866. 

USA: NEW YORK: STEUBEN COO: PRATTSBURGH (WRIGHT,S.H.r ---. 

---) 


6050 YUKONENSIS BRITTON9N.L.1 BULL. NEW YORK BOT. GARD. 2:159. 1901. 

CANADA: YUKON TERRITORY: BONANZA RIVER (WILLlAMS,R.S., ---a 

18 JUN 1899) 

NY TYPE 

606. ZIZANIAEFOLIA RAYMONDvM. T MEMO JARD. BOT. MONTREAL 53:36. 1959. 

- Y- 

- Z-


CHINA: YUNNANI: -- (TSA1,H.T.p 62809. -- 0-w 1934 1 

4 

HO LO f Y P E

NUMBER 12 105 

AUTHOR 

BAILEY,L.H. 

AUTHOR INDEX 

DATE 

1935 

1889 

1889 

1889 

1893 

1886 

1889 

1884 

1920 

1889 

1889 

1893 

1896 

1889 

1886 

1889 

1893 

1889 

1886 

1889 

1884 

1896 

1889 

1896 

1889 

1886 

1893 

1889 

1920 

1898 

1886 

1916 

1889 

1892 

1896 

1885 

1898 

1888 

1892 

1891 

1891 

1886 

1888 

1898 

1889 

1889 

1893 

1889 

1889 

SPECIES 

ABLATA 

ACUTI NA 

ALBIDA 

ALMA 

AUSTRO-CAROLINIANA 

*BRONGNIARTII 

CALI FORNICA 

*CANEXENS 

CHIKUNGANA 

COMMUNI S 

DONNE LL -SM I THI I 

DUR I FOL I A 

EGGERTI I 

ELEOCHARIS 

ENGELMANN I 

*FEST I VA 

FETA 

*FOENEA 

*GAYANA 

*GR I SEA 

HALL I ANA 

HASSEI 

*HOOD1 I 

IDAHOA 

ILLOTA 

INOPS 

INTERIOR 

JONES I KUL I NGANA 

MADRENS IS 

*MARC1 DA 

MARIPOSANA 

*MI LI ARI S 

MONTANENSI S 

*NEBRASKENSIS 

NERVINA 

OA XAC ANA 

PANSA 

PRINGLE I 

QUADRI F IDA *QUADRIF IDA 

*RETROCURVA 

*ROSEA 

SEATONIANA 

SPECIFICA 

SPRETA 

*STERILI S 

*STRAM1 NEA 

*STRAMINEA

106 

AUTHOR 

BAILEY9L.H. 

BAILEY1L.H. IN MACOUNTJOHN 

BICKNELL,E.P. 

BLAKE, SOT 

BOOTTqF. 

BOOTTqF. IN GRAYTA. 

ROOTT,F. IN H0OKERqW.J. 

BOOTT I W. 


DATE SPECIES 

1886 

1889 

1886 

1889 

1892 

1890 

1908 

1908 

1896 

1947 

1947 

1947 

1947 

1947 

1947 

1877 

1896 

1875 

1886 

1877 

1896 

1856 

1867 

1846 

1867 

1862 

1858 

1846 

1846 

1867 

1867 

1867 

1859 

1846 

1862 

1860 

1863 

1859 

1859 

1859 

1859 

1859 

1839 

1839 

1839 

1839 

1884 

1884 

1884 

*STYLOSA 

* TE R ET I U SCUL A 

ULTRA 

VICARIA 

XERANTI CA 

*PRATENSI S 

ABDITA 

INCOMPERTA 

*XANTHOCARP A 

ACROPHILA 

BREVI S 

ER EMOSTACHY A 

LA MPR OCHL A MY S 

MELANOPHORA 

PERIL E I A 

BURCHELL I ANA 

OURAN~I I 

F ENOL ER IAN A 

FUSCOLUTEA 

*LONGIROSTRI S 

MANDONI ANA 

TR I ANGULAR I S 

*ACUTA 

BANKS1 I 

BREWER1 

CONJUNCTA 

*CR I NITA 

GEYERI 

GRIFF ITHI I 

HE BET AT A 

LAC1 NI ATA 

OL I GANTHA 

PARCIFLORA 

SANGUINE A 

*SCOPARI A 

*TENTACULATA 

*BONPLANDI I 

CONFERTIFLORA 

MICANS 

NANA 

’ PAPULOSA 

PICTA 

APERTA 

BACK I FRANKLI NI I 

HEPBURNI I 

ASS IN I BOIN ENS IS LEMMONI 

PRAEGRACILIS

NUMBER 12 

AUTHOR 

BOOTTTW. I N WATSONTS. 

BROWN, F Bo He 

BROWN,R. IN RICHARDSONTJ. I N FRANKLINVJ. 

CAREY T J 

CHAMISS0,L.A. EX STEUDELTE~G~ 

CHAPMAN T A. W 

CHAPMAN,AoWo EX BOOTTfFo 

CHAPMAN9A.W. EX DEWEYTCO 

CHEE SEMAN 9 T. F 

CLARKE,C.Bo IN MERRILLTEoD. 

CLAUSEN9R.T. AND WAHL9H.A. 

CLOKEYTI.W* 

CRONQUI STT A. 

CURTIS,M.A. EX GRAYqA. 

DEWEY 9 C 

DATE 

1880 

1880 

1880 

1880 

1883 

1880 

1880 

1901 

1895 

1905 

19C 1 

190 1 

1930 

1819 

1819 

1843 

1843 

1843 

1847 

1847 

1855 

1860 

1860 

1862 

1847 

1892 

1883 

1884 

1908 

1908 

1903 

190 3 

1903 

1904 

1908 

1907 

1908 

1906 

1939 

1922 

1919 

1939 

1922 

1919 

1919 

1943 

1841 

1849 

1835 

SPECIES 

HETERONEURA 

KELLOGGI I 

NUDATA 

PHYLLOMANI CA 

SCHAF FNE RI 

*SCOPAR I A 

SUBFUSCA 

BONANZENSI S 

*ST RI CTA 

UNDERWOOD11 

W I LLI AMSI I 

YUKONENSIS 

TAHITENSIS 

POD OC A R PA 

RI CHARDSON I I 

CAROL I N I ANA 

MI SERA 

STYLOFLEXA 

PLATYPHYLLA 

SYCHNOC E PH AL A 

PAC HY S T ACHY A 

*DIGITALIS 

*ST ELL UL ATA 

*ST IPATA 

BALTZELLII 

*COMANS 

DEVIA 

PETRI E I 

AEQUA 

FELIPENSIS 

ICHANGENSI S 

LANC I FOL I A 

*LONGICRURI S 

PRAINI I 

PSEUDOJAPONICA 

SU BT R ANSVE R S A 

VI OL AC EA 

RHYNCHACHAENIUM 

*ANGUSTIOR 

APODA 

ARAPAHOENS I S 

*INTERIOR 

PAYSONI S 

SUBIMPRESSA 

*TRIBULOIDES 

OBOVO I DEA 

AE STIVAL IS 

*ALOPECOIDEA 

ARCTI CA 

107

108 

AUTHOR 

DATE 

DEWEY C o 

1854 

1836 

1845 

1842 

1826 

1836 

1861 

1836 

1846 

1824 

1846 

1826 

1854 

1826 

1836 

1849 

1847 

1857 

1846 

1842 

1857 

1836 

1854 

1842 

1835 

1836 

1836 

1861 

1842 

1825 

1826 

1824 

1825 

1826 

1866 

1861 

1846 

DEWEYvCo IN TORREYvJo IN EMORY,WoHo 1859 

1859 

1859 

DEWEYvCo IN WOODqAo 1845 

1861 

1845 

1845 

DREJER, S .To No 

ESSENBACH,NoVON 

FERNALD9M.L 

IN LEHMANNpJo 

1841 

1846 

1942 

1942 

1902 


SPECIES 

*ARI STATA 

BACKANA 

BUCKL EY I 

*CE PHALOPHORA 

COL L ECT A 

COLUMBIANA 

*DOUGLAS1 I 

FESTIVA 

FLACCOSPERMA 

FORMOSA 

HALE1 

HALSEYANA 

HAYDENI I 

HI TCHCOCK I ANA 

HOOKERANA 

I GNOTA 

ILLINOENSIS 

LAEVI-CON1 CA 

LEAVENWORTHII 

MEAD1 I 

MEEK1 I 

MIRAB IL I S 

NEBRASKENSIS 

NUTTALL I1 

PARRYANA 

PETASATA 

PETRI COSA 

RAYNOLDSII 

SARTWELLII 

SCHWEINITZI I 

SICCATA 

TENERA 

TR I SPERMA 

*UMBELLATA 

*V AG I N AT A 

VALLICOLA 

WOOD1 I 

BARBARAE 

THURBERI 

WRIGHT11 

COOLEYI 

*HI RSUTA 

PRAIREA 

RETROCURVA 

PRATENSI S 

PREISSI I 

*ABSCONDITA 

X ABSCONDITIFORMIS 

AENEA

NUMBER 12 

AUTHOR 

FERN ALO 9 M L 

DATE 

190 2 

1942 

1942 

1902 

1907 

1902 

1902 

1946 

1897 

1937 

1942 

1942 

1913 

1941 

1938 

1902 

1906 

1935 

1935 

190 6 

1906 

1906 

1942 

1906 

1942 

190 1 

1933 

€942 

1906 

1926 

1902 

1902 

1915 

1902 

1942 

191 8 

1906 

1907 

1933 

1942 

1941 

1902 

1921 

1902 

1902 

1942 

1933 

19C 1 

1933 

SPECIES 

*ALATA 

*AMPHIBOLA 

BAYARDI 

*CANESCENS 

CIL IARI S 

CRAWFORDII 

*CRAWFORD11 

*CR IN I T A 

*CRI NITA 

*CRUS-CORVI 

*CUMULATA 

*DEBILI S 

*DEWEYANA 

*DIGITALIS 

*DIGITALIS 

*ECHINATA 

*FLAVA 

GARBERI 

*GARBERI 

*GLAREOSA 

HARPER1 

HORMATHODE S 

*INFLATA 

*I NTER I OR 

* I NTUME SCEN S 

KATAHDINENSIS 

LANGEANA 

*LASIOCARPA 

*LAXI FLORA 

*L I VIDA 

*MIRABILIS 

*MIRABILIS 

MI SANDROIOES 

ORONENS IS *PALLESCENS 

*PAUPERCULA 

*PAUPERCULA 

PER LONG A 

X PSEUDO-FULVA 

*RICHARDSON11 

~UGATA 

*SCOPARI A 

*SCOPARI A 

*STRAMINEA 

*TENERA 

TERRAE-NOVAE 

X TRICHINA 

*VESICARIA 

*V E S I CAR I A 

109

110 

AUTHOR 

FERN ALD 9 M L 

FERNALDT M. L AND W EATHERBY T C. A. 

FERNALDpt4.L. AND WIEGANDTK.M. 

FRANCHET 9 A 

FRANCHETTA. AND SAVATIERTL. 

GANDOGER 9 M 

GAUOIN,J.F.G.P. 

GRAY 9 A. 

GRAYTA. EX TORREYTJ. 

GROSST R. 

GROSSpR. IN WERDERMANNyE. 

HEMSLEY9W.B. 

HERMANN9F.J. 

HERMANNT F. J. IN MCVAUGHTR. 


DATE SPECIES 

1900 

1933 

1931 

1911 

1924 

1912 

1910 

1895 

1896 

1879 

1879 

1879 

1879 

1920 

1830 

1867 

1836 

1941 

1941 

1941 

1941 

1929 

1885 

1955 

1938 

1957 

1950 

1963 

1967 

1938 

1960 

1957 

1965 

1971 

1957 

1941 

1938 

1956 

1968 

1937 

1950 

1964 

1957 

1950 

1948 

1955 

1936 

1954 

1949 

*VE STI TA 

X XANTHINA 

CL I VI COLA 

*HORNSCHUCH I ANA 

*HOST1 ANA 

*SCOPARIA 

*SCOPAR I A 

FARGESI I 

PTERO L E PTA 

MACROGLOSS A 

*NUTANS 

PLANATA 

PODOGYNA 

CRANDALLII 

*FLAVA 

*DEB1 LI S 

*OL IGOCARPA 

*PHALAROIDES 

*PIRCHINCHENSIS 

*PURPUREOVAGINATA 

SALTAENSIS 

WERDERMANNII 

POTOSI NA 

AMPL I SQUAMA 

*ART ITECTA 

ATHABASCENSIS 

ATRACTODES 

*BI PART1 TA 

CHIAPENSIS 

X DEAMII 

*EGGLESTONI I 

EURYSTACHYA 

*FISSA 

GUATEMALENSI S 

INCONDITA 

*INTERIOR 

*LA XIFLORA 

LIMNOPHILA 

*MICROPTERA 

PELOCARPA 

PERCO STATA 

P L E C T OC A R P A 

*PRATI COLA 

QU ICHENS IS 

ROANENSIS 

VE XANS 

*VULPINOIDEA 

*HI LLDENOWI I 

MURICULATA

NUMBER 12 111 

AUTHOR 

HOCHSTETTER,Co Fo EX STEUDEL, EoG. 

HOLM 9 H .T 

HOOKERtJoDo 

HOWE9E.C. IN GOROINIER,HoCo AND HOWE,EoCo 

HOWELL 9 J T o 

HULTENy0.E.G. 

JONES9M.E. 

KELS0,Lo 

KNIGHT,OoW. 

KOYAMAIT. 

KRAUSS, R 

KRECZETOWICZTVOI 

KUKENTHALtGo 

DATE 

1855 

19C3 

1905 

1905 

1903 

1900 

190 5 

190 3 

1902 

1904 

190 5 

1904 

190 5 

190 5 

1904 

1904 

1902 

1904 

1904 

1904 

1904 

1887 

1894 

1958 

1949 

1961 

1942 

1910 

1910 

19 10 

1945 

1950 

1945 

1947 

1906 

1969 

1959 

1950 

1950 

1950 

1934 

1929 

1929 

1910 

1910 

1926 

1926 

1926 

1910 

SPECIES 

MERCARENSI S 

ACCEDENS 

BRAC HYPODA 

CAMPLYOCARPA 

CHALCIOLEPIS 

ELYNU IDES 

EURYC AR PA 

*FESTIVA 

GYMNOCLADA 

LACUNARUM 

*LUZULAEFOLI A 

MI CROC HAET A 

OXYCARPA 

PACHY STOMA 

PHAEOLEPIS 

P HY SOC HL AE N A 

PRIONPHYLLA 

*SC IRPOIDEA 

*SCIRPOIDEA 

VAGANS 

VITREA 

SCAPOSA 

ROSAEOIDES 

JEPSONI I 

SPECUICOLA 

TOMPKINSI 

JACOB1 -PETER I 

ABORIGINUM 

ELROOI 

ST ANTONENS IS ELBERTANA 

ER XL E BEN I ANA 

HAGI ANA 

UNCOMPAHGRE 

*TR I SP ERMA 

OBLANC EOLATA 

TAMAK I KAUA I ENS IS 

*PLUVI CA 

*WAHUENSIS 

PHI LOC RENA 

*AP ERT A 

*APERTA 

ARSENII 

*BRUNNEA 

CUBENSI S 

*CU BENS I S 

EKMANI I 

ELMER1

112 

AUTHOR 

KUKENTHAL ,Go 

KU KE NT H A L 9 G AN D E KM AN 9 E L 

KUKENTHALvGo IN ELMER9 AoDoEo 

KUKENTHALTG. I N ENGLER9HoGoAo 

KUNTH9C.S. 

LEPAGEvEo 

L ITVINOV 9 D o I 

MACKENZIE9K.K. 

DATE 

1929 

1929 

1910 

1910 

1902 

1899 

1911 

1910 

1938 

1935 

1920 

1920 

1929 

191 1 

1909 

1909 

1909 

1909 

1909 

1909 

1909 

1909 

190 9 

1909 

1837 

1956 

1956 

1964 

1956 

1956 

1957 

1962 

1964 

1956 

1962 

1899 

1909 

1916 

1935 

1910 

1906 

1910 

1907 

1931 

1912 

1931 

1935 

1910 

1913 


SPECIES 

* EU RY C AR P A 

*INTERRUPTA 

*JAMESONI 

MERRILL I MICRANTHA 

*M ICROGLOCH I N 

PY C NOTHY SOS 

RAMOSI I 

S ARAW AKETENS I S 

SAVAIIENSIS 

SUKSDORFII 

*SUKSDORFII 

*EKMANI I 

PALAWANENSI S 

*AQUAT I L I S 

*CLADOSTACHYA 

*F I L IFOL I A 

*HINDS1 I 

*LEMANNI ANA 

*LENTICULARIS 

*NUDATA 

*PI NETORUM 

*ST I PATA 

*TENUIFLORA 

STEUDELI I 

X DUMANII 

X EXSALINA 

X NEOBIGELOWI I 

X NEOF IL I PENDULA 

X NEOPALEACEA 

X NUBENS 

X PATUENSIS 

X QUEBECENSI S 

*X SAXENII 

*VI RI DULA 

*ST ENOPHY LL A 

ABRAMSI I 

ABRUPTA 

ACUTI NELLA 

AESTIVALIFORMIS 

AGGLOMERATA 

AGGREGATA 

AGROSTOIDES 

ARCTAEFORMIS 

ATROSQUAMA 

AUTUMNAL IS 

AZTEC ICA 

B I LTMORE ANA 

BRAINERDII

NUMBER 12 

AUTHOR 

MACKENZI Ec K.K. 

DATE 

1913 

1907 

1915 

1910 

1935 

1908 

1906 

1935 

1916 

1914 

1916 

1910 

1922 

1915 

1915 

191 5 

1931 

1909 

1922 

1922 

1910 

1909 

1913 

1916 

1916 

1922 

1909 

1916 

1931 

1916 

1935 

1916 

191 5 

190 9 

1906 

1923 

1910 

19G9 

1935 

1931 

1931 

1922 

1916 

1907 

1909 

1914 

1916 

1915 

1916 

SPECIES 

BREVICAULI S 

BREVI SQUAMA 

BULBOSTYLI S 

BUSHI I 

CAESAR1 ENS I S 

CHI HUAHUAENSI S 

CONCINNOIDES 

CONSPECTA 

CONVOL UTA 

CRY PTOL EP I S 

DAVYI 

DEBIL I FORM I S 

DUDLEY I 

EGGLESTONI I 

EGREG I A 

FEST IVELLA 

FI SSA 

FISSURICOLA 

FLACC I FOL I A 

FRACTA 

FULVE SCENS 

FUS COT I N CT A 

GEOPH I LA 

GR AC I L I OR 

HARFORD I HELLERI 

HOLMIANA 

INTEGRA 

I NVOLUC RAT ELLA 

LANCIFRUCTUS 

L E I OP HY L L A 

LEPORINELLA 

LUNELL I ANA 

MACROSPERM A 

MEDITERRANIA 

MERRITT-FERNALDII 

ME SOC HORE A 

MICROPTERA 

MISERABILIS 

MOHR I ANA 

MOLESTA 

MONTEREYENSIS 

MULT IC OSTATA 

NEOME XICANA 

NUBICOLA 

OKLAHOMENSIS 

OLYMPICA 

ONUSTA 

PACHYCARPA 

113

114 

AUTHOR 

MACK EN21 E 9 K K 

MACKENZIEtKoKo EX BRIGHT,Jo 

MACKENZIE,KoKo IN ABRAMSqLo 

MACKEN2IEtK.K. IN PIPER9C.V. AND BEATTIE, 

MACKENZIE,KoKo IN RYOBERG, PaAo 

MACKENZIE,K.Ko IN SMALL,JoKo 

MAGUIRE,B* 

MAGUIRE 9 B 

AND HOLMGRE N 9 A He 

MAR1 E-VI CTORI N q t FR ERE 1 

MERRILL 9Eo Do 

MERRILL1EoDo AND CHUNtNaKo 

DATE 

1922 

19@7 

19C9 

191 3 

1931 

1931 

1908 

1931 

1935 

1915 

1915 

1931 

1909 

1906 

1908 

1908 

1915 

190 7 

1909 

1935 

1922 

1916 

1916 

1931 

1922 

1907 

1922 

1931 

1930 

1923 

1923 

1923 

191 5 

1915 

1917 

1917 

1917 

1917 

1917 

1917 

1913 

1944 

1944 

1944 

1946 

1929 

1918 

1934 

1935 


SPECIES 

PAUC I COSTATA 

PERGLOBOSA 

PER STRI CTA 

PITYOPHILA 

PLATYLEPIS 

PRAECEPTORIUM 

PROJ ECTA 

PROPOS I TA 

PURPURI FERA 

RUGOSPERMA 

RUSBY I 

RUTH1 I 

SALINAEFORMIS 

SAX1 MONTANA 

SCABRIUSCULA 

SC IRPI FORM I S 

SHELDON1 I 

SIMULATA 

SMALLIANA 

STELLATA 

STENOPTERA 

SUB-BRACTE ATA 

TENERAEFORMIS 

TOWNSENDII 

TRACY I 

TUMULICOLA 

UN ILATERAL I S 

W I EGANDI I 

LARICI NA 

NEUROPHORA 

SUBORBICULATA 

V I RIDIOR 

CUSICK I PIPER1 

AL BO- N I GR A 

ANGUST IOR 

EPAPI LLOSA 

INCURVI FORM1 S 

LEPTOPODA 

NELSON1 I 

MAGN I FOL I A 

INTERIMUS 

RACHI LL I S 

*VERNACULA 

WAMPYLOCARPA 

*OEDER I 

BAMBUS ETOR UM 

*RUBRO-BRUNNEA 

TSOI

NUMBER 12 115 

AUTHOR 

MEYE R T C A 

MOHR T C 

MUHLENBERG 9 H. EX W ILLDENOW T C L. 

NELMES, E. 

0' NE ILL TH. To 

0'NEILLpH.T. AND DUMANTM. 

OHWI T J. 

OLNEY 9 So T 

0LNEYvS.T. EX 6AILEYgL.H. 

0LNEYvS.T. EX BAILEYTL.H. IN COULTERTJ~M. 

0LNEYTS.T. EX BOOTTvW. IN WATSONpS. 

0LNEYvS.T. IN GRAYTA. 

0LNEYgS.T. IN PORTER9T.C. IN HAYDEN9F.V. 

0LNEYyS.T. IN WATSONvS. 

PAR1 SH 9 S B 

PECK 9C.H. 

PETRI E, D 

PI PERqC. V. 

PORS ILDT A. E. 

PRESL TKO B. 

DATE 

1831 

1831 

1831 

1910 

1805 

1938 

1939 

1955 

1940 

194 1 

1934 

1931 

1934 

1932 

1933 

1932 

1952 

1871 

1884 

1888 

1886 

1886 

1894 

1880 

1868 

1868 

1868 

1868 

1868 

1868 

1872 

1871 

1871 

1871 

190 5 

190 5 

1895 

1894 

1895 

1893 

1881 

1882 

1906 

1906 

1943 

1939 

1939 

1943 

1828 

SPECIES 

CIRCINNATA 

LEI OCARPA 

N I GR I C ANS 

*ST I PATA 

RETROFLEXA 

EXPLORATORUM 

SCHNEIDERI 

TR I CHOPHYLLA 

BARTLETTII 

DUTILLY I 

APODOSTACHYA 

CUNE AT A 

HATUSIMANA 

HY MENODON 

KURILENSI S 

RUG AT A 

TETSUOI 

PORTER I 

*ADUSTA 

*DEWEYANA 

*LEPORINA 

OREGONENSI S 

*WILLDENOVI I 

MENDOCINENSIS 

ATHROST ACHY A 

C I NNAMOMEA 

GYNODYNAMA 

LUZUL I NA 

SARTWELLIANA 

WH ITNEY I 

HALL1 I 

*AUREA 

HAYDENIANA 

WATSON I 

AUSTROMONT ANA 

JACINTOENSIS 

*ROSEA 

*ROSEA 

*ST I PATA 

*TORTA 

KALOIDES 

LONGICULMI S 

*HOOD I I 

PHAEOCEPHALA 

*ATROF USCA 

KOKRINENSI S 

MELOZITNENS I S 

MORRISSEYI 

ANTHOXANTHERA

116 

AUTHOR 

RAUP H oM 

RAYMOND, Mo 

ROACH,AoWo 

RYDBERG, Po A. 

SARTWELL9H.P. EX DEWEY,Co 

SCHWEINITZ,LoDo 

SCHWEINITZyLoDo AND TORREY,Jo 

SHUTTLEWORTH,RoJo EX KUNZEqGo 

SToJOHNyH. 

ST AC EY 9 J W 

STANDLEY,P.Co 

STANDLEY,PoCo AND STEYERMARK,JoAo 

STEUDEL,EoGo 

STEYERMARK,JoA. 

SUKSDORF ,WoNo 

TORREY J 

DATE 

1947 

1959 

1959 

1952 

1901 

1855 

1849 

1824 

1824 

1824 

1824 

1824 

1824 

1844 

1947 

1938 

1937 

1939 

1938 

1936 

1937 

1939 

1947 

1953 

1947 

1953 

1953 

1953 

1855 

1855 

1855 

1855 

1855 

1855 

1855 

1855 

1964 

1951 

1951 

1951 

1952 

1951 

1951 

1954 

1951 

1923 

1906 

1836 

1836 

5AlITHSONIAN CONTRIBUTIONS TO BOTANY 

SPEC IE S 

SOPERI 

SURCULO SA 

2 I Z AN I AEFOL I A 

DI VERSI STYL IS 

EBENEA 

CHAPMAN I 

SCABR I OR 

COSTATA 

CR I STAT A 

GRAC I LL I MA 

NIGRO-MARGINATA 

NOVAE-ANGLIAE 

BARRATTI I 

CRUS-CORVI 

VI TI ENS I S 

CONSTANCEANA 

CURATOR I UM 

DANAENS I S 

EA ST WOOD I ANA 

OBISPOENSIS 

SONOMENSI S 

SUBNI GR ICANS 

STEYERMARKI I 

CUCHUMATANENSIS 

HUEHUETECA 

TOJQU I ANENS I S 

TUNIMANENSI S 

VENOSIVAGINATA 

AMPHI BOLA 

AUROLENSIS 

FL ACC I D ULA 

I NC I SO-DENT ATA 

MAC R OKOL E A 

PICTA 

PRESL I PTYCHOCARPA 

A2 UAYAE 

CULMEN ICOLA 

LARENSI S 

RORAIMENSI S 

ST ANDLEY ANA 

TACH1 RENSI S 

TAMANA 

TOREADORA 

TURUMIQUIRENSIS 

*NEBRASKENSIS 

PADDOENSIS 

ALATA 

JAMES I

NUMBER 12 

TORREY 9 J 

AUTHOR DATE 

TORREY, J. EX DEWEY ,C. 

TORREYTJ~ IN WILKESyC. 

TUCKERMAN T E. 

TUCKERMAN,Eo EX DEWEY,C. 

TUCKERMANvE. EX 0LNEYvS.T. IN GRAYTA. 

UNDERWOOD, JoK. 

WAHLENBERG , G o 

WATERFALLTU~T. 

WEATHERBY,C.Ao 

W1EGANDyK.M. 

WRIGHTTS.H. I N DEWEYpC. 

1836 

1836 

1836 

1846 

1836 

1854 

1843 

1843 

1843 

1860 

1868 

1945 

1803 

1954 

1936 

1922 

1922 

1866 

SPECIES 

SCOULERI 

TURGE SCENS 

CAREYANA 

HE TEROSTAC HYA 

HOUGHTONI ANA 

WILKESI I 

ALOPECOIDEA 

*CANESCENS 

*SCOPARIA 

ARGYRANTHA 

GL AUCODEA 

*OXYLEPI S 

AQUATI L I S 

LATE BR ACTE AT A 

MAC KE NZ I AN A 

CR EBR I FLORA 

ORMOST ACHY A 

XEROCARPA 

117

118 

DATE 

1803 

1805 

1819 

1824 

1825 

1826 

1828 

1830 

1831 

1835 

1836 

1837 

1839 

1841 

1842 

TAXUN 

AQUAT IL IS 

R ETPOFLEXA 

PODOCARPA 

R I CHARDSONT I 

BARRATTI I 

COSTATA 

C R I S TATA 

FORMOSA 

GRAC ILL IMA 

NIGPO-MARGINATA 

NOVAE-ANGLIAE 

TENERA 

SCHWEINITZI I 

T R I S PEP MA 

Cot1 ECTA 

HAL SEYANA 

HITCHCOCK IANA 

S ICCATA 

*UMBELLATA 

ANTHOX ANTHER A 

*FLAVA 

CIRCINNATA 

LE I OC AR Pb 

NIGRICANS 

ARCT ICA PARRYANA 

AL AT A 

BACKANA 

CAREYANA 

COLUMB I AN4 

FEST IVA 

HOOKE RANA 

HOUGYTONI ANA 

JAMES11 

M I RABILIS 

*OL I GOCARP A 

P ETA SAT A 

PETR ICOSA 

SCOULER I 

TURGESCENS 

STEUDEL I AP ERTA 

BACK1 I 

FRANKLIN1 I 

HEPBURNI I 

AESTIVALI S 

PR ATENS I S 

*C EPHALOPHORA 

MEAD11 

PUBLICATION-DATE INDEX 

AUTHOR 


WRHLEmERG TG. 

MUHLENBERG TH EX W IL LDENOW T C L 

BROWNTR. IN RICHARDSONqJ. IN FRANKLINfJ. 

BROWNTQ~ IN RICHARDSONpJ. IN FRANKLINTJ. 

SCHWE I NI TZ TL D. 

SCHWEINITZ 9L.D. 

SCHWE I NIT2 TL 0. 

OE WEY TC 

AND TORREY T J 

SC HWE IN I TZ TL 0. 

SCHWE I NIT2 TL 000 

SCHWEINIT2,L.D. 

DEWEY T C 

DEWEY +C. 

OEWEY TC. 

DEWEY T C 

DEWEY TC. 

DEWEYqCo 

DEWEY T Co 

DEWEY 9 c 0 

PR ESL TK *R 

GAUDINTJ.F.G.P. 

MEYER TC. A. 

MEYERTC .A. 

MEYER9C.A. 

DEWEY T Co 

DEWEY 9 C 

TORREY T J. 

DEWEY 9 C 

TORREYtJ. EX DEWEYTC. 

DE WEV TC. 

DEWEY T C 

DEWEY 9 C 

TORREYTJ. EX DEWEYfC. 

TORREY 9 J 

DEWEY TC. 

GRAYTA. EX TORQEYqJo 

DEWEY T C. 

DEUEY TC. 

TORREY T J. 

TORREY + J 0 

KUNTHTCOS. 

BOOTTvF. IN HOOKERTW.J. 

BOOTTIF. IN HOOKERTW.J. 

6013TT~F. IN HOOKERTW.J. 

ROOTTTF. IN HOOKERTW.J. 

CURTISTM.A. EX GRAYTA. 

ORE JER T S. To N o 

DEWEY TC. 

DEWEY TC. 

\

NUMBER 12 119 

DATE 

1842 

1843 

1844 

1845 

1846 

1847 

1849 

18 54 

1855 

1856 

1857 

1858 

TAXON 

NUTTACL I SARTWECL I ALOPECOIDEA 

*CANESCENS 

CAROLINIANA 

MISERA 

* S COP AR I A 

STY1 OFL E XA 

CR US -COR V I 

BUCKLEY I 

COOLEY I 

PRAIREA 

R E T R OCU RV A 

BANK SI I 

FLACCOSPERMA 

GEYERI 

GRIFFITHII 

HALE I 

HETEROSTACHY A 

LE4VENWORTHI I 

PREISSI I 

SANGUI NEA 

WOOD1 I 

BACTZELL I I LL I NOENS I S 

P L ATY P HY L L A 

S YCH NOC E P H AL A 

*ALOPECOIDEA 

I GNOTA 

SCABRIOR 

*ARI STATA 

HAYDENI I 

NEBRASKENSIS 

WILKESI I 

AMPH I BOLA 

AUROLENSI S 

CHAPMAN I 

FL ACC I DUL 4 

INCI SO-DENTATA 

MACROKOCEA 

MERCARENS IS PACHYSTAC HYA 

PIETA 

PRESLI I 

P TY C HOC AR PA 

TRIANGULARIS 

LAEVI-CONICA 

MEEKII 

*CRIN ITA 

AUTHOR 

DEWEY ,Co 

DEWEY T f 

TUCKERMANTE. 

TUCKERMANTE. 

BUCKLEY9 S o R o 

BUCKL EY p S Bo 

TUCKERMAN, Eo 

BUCKLEYTSOB. 

SHUTTLEWORTHTR.J. EX KUNZEtG. 

DEWEY 9 C o 

DEWEYTC. IN WOODPA. 

DEWEYTC. I N WOODTA. 

DEWEYTC. IN WOODTA. 

BOOTT 9 F 0 

DEWEY T C 

BOOTT T F 

B(30Tf 9 F 

DEWEY T C 

TORREYTJ. EX OEWEYTC. 

DEWEYTC. 

ESSENRACHT NmVON IN LEMMA", J 

BOOTT F 

DEWEY rC. 

CHAPMAN, 4oWo EX DEWEY TC 

DEWEY c Co 

CAREY, J o 

CAREY 9 J 

DEWEY 9Co 

DEWEY , t o 

SARTWELLTHOP. EX DEWEYvC. 

DEWEY tCo 

DEWEY ,Co 

DEWEY ,Co 

TORREYTJ. IN WICKEStCo 

STEUDEL,EoGo 

STEUDEL T E. G o 

SARTWELLTHOP. EX DEWEYTC. 

STEUDEL E * Go 

STEUDEL T €0 Go 

STEUDEL T €.Go 

HOCHSTETTERTC~F. EX STEUOELTE.G~ 

CHAYISS0,L.A. EX STEUDELTE~G. 

STEUDEL T E Go 

ST EUDEL 9 E o G 

STEUOELTE.G. 

BO ECK E LER p J 0 

DEWEY ,Co 

DEWEY T C. 

BOOTT T F

120 

DATE 

1859 

1860 

1861 

1862 

1863 

1866 

1867 

1868 

1871 

1872 

1875 

1877 

1879 

1880 

TAXON 

BAR 8 AR A E 

CONFERT I FLORA 

MICANS 

NANA 

P APUL 0s A 

PARC 1 FLORA 

P ICTA 

THURBER I 

WR IGHTI I 

ARGYR ANTHA 

*DIGITALIS 

* S TEL LUL AT A 

*T E NT AC UL AT A 

*DOUGLAS I *HI RSUTA 

RAYNOLDS I I 

VALL ICOLA 

CONJUNCfA 

*SCOPARI A 

*STIPATA 

*BONPLANDI I 

*VAGINATA 

XEROCARPA 

*ACUTA 

BREWER1 

*DEBILIS 

HE BET AT A 

LACINIATA 

OL IGANTHA 

ATHROSTACHYA 

CINNAMOMEA 

GLAUCODEA 

GY NO DY N AM A 

CUZULINA 

SAQTWELLIANA 

WH I TNEY I 

*AUREA 

HAY DEN I AN A 

PORTER1 

WATSON1 

HALL I FENDLER I ANA 

BURCHELL IANA 

*LONG IROSTRI S 

MACR OGL 0s S A 

*NUTANS 

PLANATA 

PO DOGY N A 

HE T E P ONEU R A 

AUTHOR 


DEWEY,C. IN TOQREYyJ. IN EMORY,W.H. 

BOOTTvF. IN GRAYyA. 

BOOTTVF. IN GRAYqA. 

ROOTTtF. IN GRAYtA. 

BOOTTIF. IN GRAY,A. 

ROOTT 9 F 

BOOTT,F. IN GRAYyA. 

DEWEYvC. IN TORREY,J. IN EMORYTW.H. 

DEWEYtC. IN TORREY,J. IN EMORY9W.H. 

TUCKERMAYgEo EX DEWEYqC. 

CHAPMAN, A.W. 

CHAPMAN 9 A. W 

BOOTT T F 

DEWEY 9C. 

DEWEYpC. IN WOODtA. 

DEWEY T C 

DEWEY cC. 

BOOTT T F 

BOOTT 9 F 

CHAPMAN,A.W. EX BOOTTpF. 

BOOTTTF. IN GRAYVA. 

DEWEY TC. 

WRIGHT9S.H. IN DEWEYqC. 

BOOTT 9 F 

BOOTT F 

GRAY, A, 

ROOTT 9 F 

BOOTT,F. 

BOOTT 9 F. 

0LNEYtS.T. IN GRAYtA. 

OLNEY,S.T. IN GRAYvA. 

TUCKERMANtE. EX 0LNEYtS.T. IN GRAYqA. 

OLNEY,S.T. IN GRAYqA. 

OLNEY,S.f. IN GRAYvA. 

OLNEY9S.f. IN GRAYqA. 

0LNEYtS.T. IN GRAYqA. 

OLNEY,S.T. IN WATSONqS. 

OLNEYI SOTO IN WATSON9 S o 

OLNEY 9 5 .T 

OLNEYTS.T. IN WATSON,S. 

0LNEYtS.T. IN PORTER,TbC. IN HAYDEN9F.V. 

BOECKELERtJ.0. 

BOECKELERt J.0. 

BOECKELERtJ.0. 

FRANCHETtA. AND SAVATfERvL. 

FRANCHETt A. AND SAVAfIERpC. 

FRANCHET, A. AND SAVATIER,L. 

FR ANCHET 9 A AND SAVATIER? L. 

BOOTTtW. IN WATSON, So

NUMBER 12 121 

DATE 

1880 

1881 

1882 

1883 

1884 

1885 

1886 

1887 

1888 

1989 

TAXON 

KELLOGGI I 

MENDOCINENSIS 

NUDATA 

P HYLL OM AY I C A 

*SCOPARI A 

SUBFUSCA 

KALOIDES 

LONGICULMIS 

DEVIA 

SCHAFFNER I 

* ADU S T A 

ASSINIBOINENSIS 

*CANE SCENS 

HALL IANA 

LEMMONI 

PETRIEI 

PRAEGRAC I L IS NERVINA 

POT0 S IN A 

*BRONGNIARTI I 

ENGELMbNNI 

FUSCOLUTEA 

*GAYANA 

I NOP S 

*LE POR IN A 

*MARC IDA 

OREGONENSIS 

*R ETROCURV A 

*STYLOSA 

ULTRA 

SCAPOSA 

W E WEYANA 

PANSA 

*ROSE A 

ACUTINA 

ALBIDA 

ALMA 

CALIFORNICA 

COb4MUN I S 

DONNELL-SMITH11 

ELEOCHARIS 

*FEST IVA 

*FOENEA 

*GR IS EA 

*HOOD I 

I LLOTA 

JONES11 

*MILIARIS 

SPECIFICA 

AUTHOR 

BOOTTtW. IN WATSONtS. 

0LNEYtS.T. EX BOOTTtW. IN WATSON, S o 



BOOTT t W I IV WATSON T So 

BOOTTtW. IN WATSONTS. 

PETRI EtD. 

PETRI EtD. 

CHEESEMANt1.F. 

BOOTTtW. IN WATSONqS. 

0LNEYtS.T. EX BA1LEYtL.H. 

BOOTT t W 

BA ILEY 9 L H. 

BAILEYtL OH. 

BOOTT t U. 

C HEE S EMAN 9 To F 

BOOTTtW. 

BAILEY 9L.H. 

HE MSL EY t W B 

BAILEY TL .He 

BAILEY ,L.H. 

BOECKELERt J.0. 

BAILEYtLoH. 

BAILEYtL.H. 

0LNEYtS.T. EX BIILEY,L*H. 

BA I LEY tL OH 

Ol-NEYtS.T. EX BA1LEYtL.H. 

BAILEY tL .He 

BAILEY tL.H* 

BAILEY 9L.H. 

HOOKE R t J 0 

0LNEYtS.T. EX BA1LEYtL.H. 

BAILEY L H. 

RAILEY,L.H. 

BA ILEY ,L .He 

BAILEY tL *H 

BAILEY L .H 

6AILEYtL.H. 

BA ILEY t L e H 

BA1CEYtL.H. 

BAILEY t L. H 

RA1LEYvL.H. 

BAILEY ?Lo H 

BA1LEYtL.H. 

8AILEYqL.H. 

BA1LEYtL.H. 

BA I LEY 9 L H 

BA ILEY t L . H 

BA1LEYtL.H.

122 

DATE 

1889 

1890 

1891 

1892 

1893 

1894 

1895 

1896 

1897 

1898 

1899 

1900 

1901 

1902 

TAXON 

SPSETA 

*STRAMINEA 

*STRAMlYEA 

*TERETlUSCULA 

VICAR1 A 

*PRATENSIS 

QUADRIFIOA 

*QUAOR1 f IDA 

*COMANS 

YONTANENSIS 

PRINGLEI 

X E 9 ANT I C A 

A USTRU-CAROL1 NI ANA 

OURIFOLIA 

FETA 

1 NTE RIOR 

*STER 1L IS 

*TORTA 

ROSAEOI DE S 

*ROSEA 

*WILLDENOVII 

FARGESI I 

*ROSEA 

*STIPATA 

*STR ICTA 

DURANOI I 

EGGERTI I 

HASSEI 

I DAHOA 

MANDONIANA 

*NERRASK ENS IS 

PTE ROLE PT A 

* X ANT HOC AR P A 

*CR IN ITA 

MADRENSI S 

0 AX ACAN A 

SEATON1 ANA 

*MI CR OGLOC H I N 

*STENOPHYLLA 

ELYNOIDE S 

*VESTITA 

BONANZENSIS 

EBENEA 

KATAHDI NENSI S 

*VESTCAR14 

W I LL I AMSI I 

YUKONENSI S 

AENEA 

*ALATA 

AUTHOR 


BA ILEY 9 L oH 

BAILEYTL~H~ 

BA I LEY I Lo H e 

BAILEYTL~H~ 

BAILEY ,L Ho 

BAILEY,LoHo IN MACOUNTJOHN 

BAILEYTLoHo 

BAILEY TLOH 

CHEESCYANy ToFo 

BAILEY TL OH 

RA ILEY TL H 

BA ILEY 9 L oH 

BAILEY TL oHo 

BA ILEY T L Ho 

BAILEY rLoHo 

BAILEY TL Ho 

BA ILEY T L H 

PECKTCOH. 

HOWETEOC. IN GORDINIER,HoCo AND YOWEtEoCo 

PECK, CoHo 

OLNEY,SoTo EX BAILEYyLoHo IN COULTERTJOM. 

FRANCHET, A. 

PECK, C OH 

PECK, C eHo 

BR ITTON T No Lo 

BOECKELERrJoOo 

BAILEY ,L .Ho 

BA ILEY ( L o Ho 

BA ILEY ,L Ho 

BOECKELERt JoOo 

BAILEYTLoHo 

FRANCHET, A. 

BICKNELLTEOP. 

FERNALDT M o t 

BA ILEY TL oHo 

BAILEY TL H 

RA ILEY TL Ho 

KUKENTHALvG. 

L I TV I YOV T D I 

HOLM, H o T 

FE RN4L D T Mo L 

BR I TTONt No Lo 

RYDBERG? Po A. 

FE SNALD T Mo L 

FERNALD,MoLo 

BRITTUN 7 No L o 

RRITTON,N.Lo 

FE RNAL DT M oL 

FERNALD, Mo Lo

YUMBER 12 

DATE 

1902 

1903 

1904 

1005 

1906 

T AXON 

*CANESCENS 

CPAWFORDI I 

*CRAWFORD11 

*€CHI NATA 

GY MNOC L AD A 

MI CRANTHA 

*MIRABILIS 

*MIRABILIS 

ORON ENS I S 

PR ID NPHY L L A 

*SCOPARIA 

*STRAMINEA 

*T EN ERA 

ACCEDENS 

CHALCIOLEPIS 

*FESTIVA 

ICHANGENS I S 

LANCIFOLIA 

*LOWGICRUR IS 

LAC UN A P UM 

MICROCHAETA 

PHAEOLEPIS 

PHYSOCHLAENA 

PRAINII 

*SCIRPOIDEA 

*SC IRPOl OEA 

VAGANS 

VITREA 

AUSTROMONTANA 

BRACHYPOOA 

C A YPLY OC A RP A 

EURY CAR PA 

J AC I NTOENS I S 

*LUZULAEFOLf A 

OXYC ARP A 

PACHYSTOMA 

UNDERWOOD11 

AGGLOMERATA 

CONCINNOIDES 

*FLAVA 

*GLAR EOSA 

HARPER I 

*HOOD1 I 

HO R M AT HOD E S 

*I NTE R I OR 

*LAXI FLORA 

MEDITERRANIA 

PADDOENSI S 

*PAUP ERCUL A 

AUTHOR 

FERNALDTY-L. 

FE QN AL DI M L 

FERNALD, M. L. 

FERNALDt Me L. 

HOLM1 HOT. 

KUKENTHAL ,Go 

FE QN ALD T H L 

FERNAtD, M*L 

FERNALDt Me L. 

HOLM, H T* 

FERNALDt M.L. 

FERNALD9M.L. 

FERNAL0tM.L. 

HOLM9H.T. 

HOLM, HOT. 

HOLM, HOT. 

CLARKE TC R 

CLARKE tC 6 

CLARKE TC B 

HOLM, H .T 

H0LMtH.T. 

HOLM, H.T. 

HOLM 9 HOT. 

CLARKE C*B 

HOLYIH*T* 

HOLM9 H .T* 

HOLM, HOT. 

HOLM,H.T. 

PAR1 SH 9s. 6. 

HOLYT l4.f 

HOLM t H .T 

HOLM 9 H T 

PAR I S H 9 S B 

HOLM, H .To 

HOLM, H T 

HOLM,H.T. 

BRITTON, N. L. 

MACKENZIE,K.K. 

MACKENZ I E TK Y 

FERNAL0tM.L. 

FESNAL0,M.L. 

FERNALD9M.L. 

PI PER TC. V. 

FERNALDTMOL. 

FERNALDt M L. 

FERNALDtM.L. 

MACKENf1EtK.K. 

SUKSD0RFtW.N. 

FERNALDTY-L. 

123

124 

DATE TAXON 

1906 P HA E OC E P H A t A 

RHYNCHAC HAE NI UM 

SAXIMONTANA 

*TR I S PERMA 

1907 AGROSTOIDES 

BREVISQUAMA 

CI LI ARI S 

NEOMEXI C AWA 

P E RGLO BOS A 

P E RL ONG A 

S I YULATA 

SU RTR AN SV ERS A 

TUMUL I COL A 

1908 ABDI TA 

AEQUA 

CHIHUAHUAENSI S 

FELIPENSIS 

I NCOMPERTA 

PROJECTA 

P S EIJ DOJ APON I C A 

SCAB RT USC UL A 

SC IRPIFORMI S 

V I OL AC EA 

1909 ABRAMSI I 

*AQUATILIS 

*CL4DOSTACHYA 

*F I L I FOt I A 

FISSUPICOLA 

FU SCOT I NC T A 

*HINDS1 I 

HOLYIANA 

*L EYANN I AN A 

*LENTICULARIS 

M ACR OSP ERMA 

MI CROPTER A 

NUBI COLA 

*NUDATA 

PERSTRICTA 

*P I NETORUM 

SAL I NAEFORM IS 

SMALL1 ANA 

*STI P A tA 

*T ENU I F L OR A 

1910 ABOR IGINUM 


AGGREGATA 

ARSENII 

B I LT MORE AN A 

*BRUNNEA 

AUTHOR 

PI PERT C o V 

CLARKETC~B. IN MERRILLTEOD. 

MACKENZIE KoKo 

KNIGHTTO~W. 

MACKENZI E T Ko Ko 

MACKENZIErKoKo 

FE RN ALD T M L 

MACKENZI E KO Ko 

MACKENZIETKOK. 

FE RNALD 

M L 


MACKENZI E Ko Ko 

CL ARK E 9 C B 

MACKENZI E T K Ko 

BICKNELLTEOPO 

CLARKE 7 C B 

MACKENZI E 9 Ko Ko 

CLARKE 9 C 6. 

BICKNELL, E *Po 

MACKENZIET Y o K. 

CL AQKE T C 06 

MACKENZIETK.K. 

MACKENZI E T K Ko 

CLARKEvC oBo 

MACKE NZI E K K 

KUKENTHAtrG. IN ENGLERTH*G~A~ 

KUKEN~HALTG. IN ENGLERTHOG-A. 

KUKENTHALTG. IN ENGLERTH~G~A~ 

MACKENZIEt KI Ko 


KUKENTHALTG. IN ENGLERTH~G~A~ 

MACYEN 21 E T K K 

KUKENTHALqG. IN ENGLERTH~G~A~ 

KUKENTHALTG. IN ENGLER,HoGoAo 

MACKENZIE,K*Ko 

MACKENZIE, KoKo 

MACKENZIETK~K. 

KUKENTHALTG. IN ENGLESTH.G*A* 

MACKENZI E TK K 

KUKENTHALTG. IN ENGLER,HoGoAo 

MACKENZIETKOK. 

MACKEYZIETK-K. 

KU KE N TH A L G 

I N ENGL E R T H G A 

KUKENTHALqG. IN ENGLERTHoG-A. 

JONESTMOE. 

MACKENZIETK~K. 

MACKENZIE TK OK 

KUKENTHAL T Go 

MACKENZI E TKOK. 

KUKENTHAL T Go

NUMBER 12 

DATE 

1910 

1911 

1912 

1913 

1914 

1915 

1916 

T AXON 

BUSHI1 

DEB I L I FORM IS ELVER1 

ELRODI 

FULV ESC ENS 

*JAW€ SON1 

MERR ILL I I 

M E S OC HO RE A 

RAMOSI I 

*SCOPARIA 

STANTONENSIS 

*STI PATA 

*HORN SCHUC H I AN A 

PALAW ANEN S I S 

PYCNOTHYSOS 

ATROSQUAM A 

* SCOPAR I A 

BRAINERDII 

BREVICAUL IS 

*DEWEYANA 

GEOPH! LA 

MAGN I FOL I A 

PITYOPHILA 

CRYPfOLEPIS 

OKLAHOMENS IS 

RULBOSTYLIS 

CUSICKII 

EGGLESTONII 

EGREGIA 

FEST IVELL A 

LUNELLIANA 

MISANDROIDES 

ONUSTA 

PIPER1 

RUGOSPERYA 

RUSBYI 

SHELDON I I 

ASRUPTA 

C ONV OL UT A 

DAVYI 

GRAC ?L ?OR 

HARFORD I I NTEGRA 

L ANC I FRUCTUS 

LEPORINELLA 

MAR1 POSANA 

MULT ICOSTATA 

OLYMPIC A 

PACHYCARPA 

AUTHOR 


MACKENZIEtKoK. 

KUKENTHAL t G. 

JONES 9 M e E. 


KUKENTHALtG. 

KUKENTHALtG. 

MACKENZI Et K OK 

KUKENTHAL tG. 

FE RNAtO 9 M. L . AND W I EG AND t K. M 

JONES 9 MoE 

MOHR, C 

FERNALOT M.L* AND WIEGANDT K. Ma 

EL MER t A D E 

KUKENTHAL 9 G I N 

KUKENTHAL (Go 

MACKENZ1EtK.K. 

fEQNALD7M.L. AND WIEGAND9K.M. 

MACKENZI E1K.K. 

MACK ENZ I E 9 K K 

FE RN AL D. M L 

MACKENZI E t K.K. 


MACKEYZ1EtY.K. 

MACKENZIE t K. K. 

MACK EN Z I E t K OK 

MACKENZIE 9 K.K. 

IN SMALL9 J.K. 

MACKEN2IEtK.K. 


MACKENZI E1K.K. 



FERNALOp Ma Lo 

MACK EN2 I E 9 K K 

IN P1PERtC.V. AND BEATTIE9R.K. 


MACKENZIE 9K.K. 




MACK EN Z I E t K K 

IN PIPER7C.V. AND BEATT1EtR.K. 

MACKENZ I E 9 KO K 

MACKENZIE TY.K. 


MACKENZI E t K. K. 

MACKENZIEt K O KO 

MACKENZIE tK O K 

BAILEY,L.H. 



MACKENZI E t K. K 

125

126 

DATE 

1916 

1917 

1918 

1919 

19 20 

1921 

1922 

1923 

1924 

1926 

1929 

TAXON 

SUB-BRACTEATA 

TENERAEFORMI S 

ALBO-NIGRA 

ANGUST I OR 

EPAPILLOSA 

INCURV IFORMI S 

L E PT OPODA 

NELSON1 I 

BAMB US ETORUM 

* P AUPERC IJL A 

ARAPAHOENSI S 

SUBIMPRESSA 

*TR I B ULO I D E S 

CHI K UNGAN A 

CRANDALLJ I 

KULI NGANA 

SUKSDORFI I 

*SUKSDORfI I 

*SCOPARI A 

APOOA 

CREBRI FLORA 

Z)UOL EY I 

FLACCIFOLIA 

FRACTA 

HELL EP I 

MONT E RE Y E N S I S 

ORMO ST AC HY A 

PAUCICOSTATA 

P AY S ON I S 

STENOPTER A 

TRACY1 

UNILATERALIS 

MERRITT-FERNALDII 

*N EBR ASKENS IS 

NEUROPHOP A 

SUBORBICULATA 

VIRIDIOR 

*HOSTIANA 

CUBENSI S 

*CUBENS I 

S 

EKMANI I 

*L IVI DA 

*APERTA 

*APEP TA 

*EKMAN I *EURY CAR P A 

*INTERRUPTA 

*OEDERI 

WEROERMANNI I 

AUTHOR 


MACKENZ I E 9 K OK 

MA CKENZI E T K K 

MACKENZIE9K.K. IN RYDBERG,P.A. 

MACKENZIE,KoK. IN RYDt3ERGtP.A. 

MACKENZIE ,KoKo I N RYDBERGTP A. 

MACKENZIE,K.K. IN RYDBERG,P.A. 

MACKENZI E,K. K. IN RYOBERG, P A. 

MACKENZIE,K.K. I N PYDBERGTP.A. 

MERRILLqEoDo 

FERNALD, Me L 

CLOKEY 1. W 

CLOKEYTIOW. 

CLOKEY 9 I o w. 

BA ILEY T L H 

GANDOGER9M. 

BAILEY 9 L OH 

KUKENTHAL I Go 

KUKENTHALtGo 

FERNALDTM. Lo 

CLOKEY,I.W. 

W I EGAND9K.M. 

YACKENZIE,K.K. 

MACKENZIEIK~K. 




WIEGAN0,K.M. 


CLOKEY 9 I W 

M4CKENZI E T K. K. 

MACKENZI E 9 K. K. 



SUKSDORF9W.N. 

YACKENZIE,K.Ko IN ABRAMSvC. 

MACKENZIETK.KO IN ABRAMSTL~ 

MACKENZIEvK*K. IN ABRAMSTL. 

FERNALD, MoLo AND W IEGAND, K O M. 

KUKENTHAL ,Go 

KUKENTHALTG. 

KUKENTHAL 

FERNALD9M.L. 

KUKENTHAL (Go 

KUKENTHAL ,Go 

KUYENTHALqG. AND EKMAN,E.L. 

KUYENTHAL (Go 

KUKENTHAL 9 G* 

MARIE-VICTORIN, (FPERE) 

GROSSTRO I N WERDERMANNqE.

NUMBER 12 

DATE 

1930 

1931 

1932 

1933 

1934 

1935 

1936 

1937 

1938 

f AXON 

LARI CINA 

TAHITENSIS 

ARCTAEFORMI S 

AUTUMNAL I S 

CL IV ICOLA 

CUNEATA 

FISSA 

INVOLUCRATELLA 

MOHRI ANA 

MOLE STA 

PLATYLEPI S 

PRAECEPTORIUM 

PROPOSITA 

RUTH I TOWNSENDII 

W I EGANDI I 

HYMENODON 

RUGATA 

KURILENSIS 

L ANGE AN A 


X TRICHINA 

*VESICARIA 

X XANTHINA 

APOOOST ACHY A 

HATUSI MAlVA 

PHILOCP ENA 

*RURRO-BRUNNEA 

AB'LATA 

ACUT INELLA 

AZTECICA 

CAESAR I EN SI S 

CONS PECTA 

GARBERI 

*GARBERI 

L E I OPHY L L A 

MI SERABI LI S 

PURPURIFERA 

SAVAIIENSTS 

S T EL L AT A 

TSOI 

MACKENZIANA 

OBISPOENSIS 

WULP INOIDEA 

*CRUS-CORVI 

CURATOR I UM 

P ELOCAR P 4 

SONOMENS1 S 

*ART1 TECTA 

AUTHOR 

MACKENZIE9K.K. EX BRIGHTTJ. 

BROWN 9 F B OH. 

MACKENZI E y K. K 


FERNALD9 M. L ANlr WEATHERBY 9 C.A 

OHWI 9 J. 


MACKENLIE9K.K. 

MACKENZI E 9K. K. 


MA CKE NZ I E 9 K K 

MACKENZI E 9 K.K. 

MACK EN2 I E 9 K K 

MACKENZ I E 9 K OK 

MACKENZ I E 9 K. K 


OHWITJ. 

OHMI T J. 

OHWIT J. 

FERNAL0,M.L. 

FERNALD9M.L. 

FERNALD9M.L. 

FERNALD9M.L. 

F E RNALDT M. L 

OHWIT J. 

OHWI 9 J. 

KR EC 2 ETOWI CZ T V I 

MERRI LL 9 E . Dm 

BAICEY9L.H. 

MACKENZIE 9 K.K 



MACKENZ I E 9 K K 

FERNALD9M.L. 

FERNALD9M.L. 

MACK ENZ I E 9 K .I( 



KUKENTHAL 9 Go 


MERRI LL 9 E D. AND CHUN 9N.K 

W EATHERBY ,C. A* 

STACEY 9 J. We 

HEQMAlVN9 F. J. 

FE RNAL D 9 M L 

STACEY 9J. W. 

HERMANNT F. J. 

ST ACEY T J W 

YERMANNTF. J. 

127

128 

DATE 

1938 

1939 

1940 

1941 

1942 

1943 

1944 

1945 

1946 

1947 

TAXON 

CONSTANCE ANA 

X DEAMII 

*DIGITAL IS 

EASTWOODI ANA 

E XP L OR A TO RUM 

*LAXI FLORA 

SARAWAKETENSI S 

*A NG U ST I OR 

DANA ENS IS 

*INTERIOR 

KOKRINENSIS 

MELOZITNENS IS 

SCHNEIDER I 

SUBN I GR ICANS 

BARTLETTII 

+DIGITAL I S 

DUTILLY I 

*INTERIOR 

*PHACAROI DE S 


* P U RP UR E OV AG I N AT A 

R UGATA 

SALTAENS I S 

+A6 SC OND IT A 

X A B SC OND I T I FORM 'I S 

+PMPH IROLA 

BAYARDI 

*CUMUL AT A 

*DEBILIS 

*I NFCATA 

*INTUYEStENS 

JACOBI-PETER1 

*LASI OCARPA 

*PALL ESCENS 

*RICHARDSON11 

TERR AE-NOVAE 

*ATRO FUS C A 

MORRI SSEYI 

OBDVOI OEA 

I NTEQ I MUS 

RACH ILL I S 

WERNACULA 

FLBERTPNA 

HAG1 ANA 

*OXYL EP IS 

*CA MPYL OC A RPA 

*CRI N I TA 

ACROPHILA 

6REVI S 

AUTHOR 

STACEYTJmW. 

HERMANNqFm Jm 

FERNALD, M m L m 

STACEYVJoWm 

NELMES , E m 

HERMANN, F m 

J m 

KUKENTHALTGm 

CLAUSENTRmTm AND WAHL?HmAm 

STACEYvJmWm 

CLOKEY?I.Wm 

PORSI LDTAm E m 

PORSILD, Ao E m 

NELMES ,E 

STACEY 9 J m W. 

O'NEILL? HoTm 

FEQNALDpMmLm 

O'NEILL9H.T. ANT) DUMANqMm 

HERMANN? Fm J 

GROSS 

GROSS? R m 

GROSS 9 Rm 

FE RNALD 9 M L 

GROSS T R o 

FERNALO, MoLm 

FERNALDTM-Lo 

FERN AL 0 T M L 

FE RNAL 09 M m Lm 

FERNALDT M m Lm 

FERNALD?MmLm 

FERNACDVMm Lm 

FERNALDTMm Lm 

HULTEN, 0. E mG 

FE RN ALD 9 Mo L 

FEQNALD, Y e Lo 

FERNALDV M m L m 

FERNALDTM-Lo 

PORSI LD, A m Em 

PORSI LDTAm E m 

CRONQUIST? A m 

MA GU I R E 7 B 

MAGUI RE 7 R 

MAGUI RE 9 6 

KELSO 9 L 

KELSOTLm 

UNDERWOOD? JmKo 

MAGUIRETBm AND HOLMGREN,AmHm 

FERNALD? M m Lm 

BLAKE? SOT. 

BLAKETSmTm 

SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER 12 

DATE 

1947 

1948 

1949 

1950 

1951 

1952 

1953 

1954 

1955 

1956 

1957 

T AXON 

EREMOSTACHYA 

HUEHUETECA 

LAM PROCHC AMY S 

MELANOPHORA 

PEP ILEIA SOPER I 

STEYERMARKT I 

UNCOY PAHGRE 

VITIENSIS 

RO AN EN S I S 

M UR I CU L AT A 

SP ECUICOLA 

ATP ACTOOE S 

ERXLEBENI ANA 

K AUA I ENS I S 

PERC OST AT A 

*PLUVI CA 

QUICHENSI S 

*WAHU ENS IS CULM EN I COLA 

L ARENS I S 

RORAIMENSIS 

TACHIRENSIS 

TAMANA 


OIVERSI STYLI S 

ST ANOL EYANA 

TETSUOI 

C UCHUMATANENS I S 

TOJQUIANENSIS 

TUN1 MANENSI S 


LATE BRACT EAT A 

TOREADORA 

*WILLDENOW I I 

AMPLISQUAMA 

TRICHOPHYLLA 

VEXANS 

X OUMANII 

X EXSALINA 

L I hNOPHIL A 

X NEOFILIPENDULA 

X NEOPALEACEA 

*X SAXENII 

ATHABASCENSI S 

EURY STACYYA 

I NCONDI T A 

X NUBENS 

*PRATICOLA 

AUTHOR 

BLAKE 9 S o 1. 

STANDLEY 9 P .C AN0 STEYERMAQK 9 J. A. 

BLAKE9S.T. 

BLAKE 9 SOTO 

BLAKE 9 S o f 

RAUP 9 H. M. 

STANDLEY TP .C 

KELSO 9 L 

ST.JOHN9H. 

HERMANNTF. J. 

HERMANN9 F J IN MCVAUGHTR 

HOWELL 9J.T. 

HERMAhSN9 F. J. 

KELSO 9 L. 

KRAUSS TR 

HESMANN9 F. J. 

KRAUSS 9R. 

HERMANNvF. J. 

KRAUSS 9 R 

STEYERMARK9J.A. 


STEYEQMARK 9 J A. 

STEYEP MARK 9 J A. 

SfEYERMARK9J.A. 

STEY ERMARK 9 J A 

ROACH 9 A. W 

STEYERNASK9J.A. 

OHWI 9 J 

STANDLEY9P.C. AN0 STEYERMARK9J.A. 

STANDLEYTP~C~ AN0 STEYERYARK9J.A. 

STANDLEYTP.C. AND STEYERMARK9J.A. 

STANDLEY9P.C. AND STEYERMARK9J.A. 

WATERFALL 9 U. To 

STEYfRMARK9J.A. 

HERMANNfF. J. 

HERMANN9 F J 

NELMESTE. 

HERMANN9F.J. 

L E P AG E 9 E 

LE PAGE 9 E 

HERMANN9 F. J. 

LE PAGE 9 E 

LEPAGE 9 E 

LEPAGEvE. 

HERMANN9F.J. 

HERMANNTF. J. 

HERMA"N9 F. J. 

LE PAGE 9 E 

HERMANNvF. Jo 

129


D4TE 

1958 

1959 

1960 

1961 

1962 

1963 

1964 

1965 

1967 

1968 

1969 

1971 

TAXON 

JEPSONI I 

SURC UL 0 S A 

TAMAKII 

ZIZANIAEFOLIA 

*EGGLESTONII 

TPMPKINSI 

X PATUENSIS 

*VIP IDULA 

*BIPARTITA 

AZUAYAE 


PLECTOC AP PA 

X QUEBECENSIS 

*CISSA 

CHIAPENSIS 

* M I C R OPT E R A 

ORLANCEOLATA 

GUAT EMALENS IS 

AUTHOR 

HO WEL L T J T 

RAYMONDTM~ 

KOYAMA ,To 

RAYMONDTM~ 

HERMANN, Fo J. 

HOWELL T J T 

LEPAGE T E 

LEPAGE TE 

HERMANNTF~Jo 

STEYERMARK, J A. 

LEPAGE TEo 

HERMANN, F J. 

LE PAGE T E 

HERYANNvFo J o 

HEPMANN, F o J o 

HERMANN,FoJo 

KOYAMAyT. 

HERM~NNTF~J~

NUMBER 12 

COLLECTOR INDEX 

NUMBER DATE COLLECTED 

--- 

2688 -- - 

--- 

--- -"- 

ABRAM S , L a 

25 MAY 1897 -- -.-- 1843 

--.L 

-- AUG 1827 

--c 

2816 

ALLEN, Ja Aa 

31 JUL 1902 

21 A 

AMANOtT a 

6358 

ANDERSON, Ja Pa 

01 JUL 1878 

-- MAY 1951 

4871 

ARSENEtG.(FRERE) 

19 AUG$€938 

30 54 16 JUL 1909 

1359 

ARSENEtLa(FRERE1 

01 AUG 1907 

-9- 28 JUL 1902 

BAILEY tLaHa 

--- _-- 13 JUN 1917 

18 JUL 1917 

161 01 JUN 1886 

BAKER 9C. Fa 

811 

226 

230 

232 

232 -- - 

BANG9 Ma 

2210 

2376 

10 MAY 1902 

-- AUG 1899 

-- AUG 1899 -- AUG 1899 

-- AUG 1899 --- 

--- 

--- 

BANKS,Ja AND SOLANDER,DaC* 

1769 

-__ -- --- 

BARTLETT I Ha Ha 

11718A 

BEAMANT JaHa 

24 FEB 1931 

3880 31 JUL 1960 

BEAN9R-S.; HOSAKAvE-Y. AND STaJOHN9Ha 

11228 

B ICKNELL 9 E -P -- t 

21 DEC 1931 

11 YAY 1904 

--9 

B IGELOW 9 J a M a 

--- --- 

20 JUN 1908 

*- --- 18 53- 1854 

-- --- 18 53 -18 54 

1547 -7- 

BI LTMORE HER BAR IUY 

TAXON 

BI LTMOREANA 

*CANE SC E N S 

CR ISTATA 

*FLAVA 

PRAIREA 

ABR4MS I I 

* PAU P ERCUL A 

TETSUOI 

JACOB I -PETER I 

ARSENII 

CONSPECTA 

FULV ESC E NS 

CHIKUNGANA 

KULI NGANA 

*RE TROC URVA 

AE QUA 

CHA LC I OLE P I S 

ELY NO1 DES 

*FESTIVA 

NUB1 COLA 

SAX I MONTANA 

*CL ADOSTACHYA 

*JAM€ SON I 

BANKS1 I 

BAxRTtETTII 

GUATEMALENSIS 

*W A HUENS I S 

ABDI TA 

INCOHPERTA 

ALBIDA 

*BRONGN IARTI I 

NEOMEX I CANA 

131


NUYRER 

B ILTMORE HER BAR I UY 

262A 

BIOLETTI 9F.T. 

1 

BOL ANDE R 9 Ho N 

6213 

4741 

6477 

6477 

50 

3822 

4700 

50 74 

4740 

470 1 

2299 

6198 

4746 

5046 

5046 

4702 

--- 

DATE COLLECTED 

28 YAY 1897 

25 JUN 1893 

1863 

1866 

1866 

1866 

1864 

1866 

1866 

1866 

1860-1867 

1866 

1866 

1866 

1866 

1866 

1860 CA. 

5086 

1866 

6198 

1866 

BOOTT p W --- 

26 JUL 1865 

BOURGEAUpE. --- ..- --- 1857-1859 

BRA INERDT E 

121 

19 JUL 1897 

160 

18 JUL 1897 

111 

11 JUL 1897 

-- - 

19 JUL 1898 

BRASSpLo J o 

9515 

-- AUG 1938 

441 8 

-- YAY-JUL 1933 

10255 -- OCT 1938 

5323 

-9 SEP-NOV 1933 

9583 

-- AUG 1938 

BRASS9L.J. AND MEYER-DSEES,E. 

98 28 -- SEP 1938 

BREEOLOVEpO. E o 

67 14 30 JUL 1964 

BREWER+ W. He 

1650 

1422 

1977 

1636 

1969 

1969 

17 JIJN 1863 -- --- 1863 

31 JlJL 1863 -- --- 1863 

31 JUL 1863 

31 JUL 1863 

TAXON 

*STI PATA 

TUMULICOLA 


CALI FORNICA 

CINNAMOMEA 

DEB1 LIFORMIS 

FETA 

GRACIL IOR 

GYNODYNAMA 

HAYOENI ANA 

L(J ZUL I NA 

YE ND OC I NE NS I S 

NUDATA 

PAUC I COST AT A 

PH YL LOMANI C A 

QUADRIFIDA 

* QU ADR I F I DA 

SALINAEFORMI S 

SU 6-BRACT€ ATA 

WHITNEY I 

WHI TNEYI 

1 

*SCOPARI A 

*LONGIROSTRIS 

BRAINERDI I 

*ECHINATA 

*FILIFOLIA 

*INTUMESCENS 

ACROPHI LA 

BREVIS 

ERE MBST ACHY A 

LAMPROCHLAMYS 

PERILE I A 

YE L ANOPHOR A 

CHIAPENSIS 


BREWER1 

PACHYCARPA 

S ART W E LL I ANA 

* SCOPAR I A 

SPECIFICA

NUMBER 12 133 


BREWER, W Ho 

1778 

BRIGHT, J o 

--- 

BRI TTON ,No L o --- 

BROWY,O* Yo 

25 5 

BUCKLEY 9S.B. 

--- 

--- 

--- 

--- 

-- - 

6URCHEL.L ,W Je 

1911 

BURT-DAVYTJO 

3266 

BUSH9B.F. 

1718 

1718 

2514 

1043 

7020 

99 3 

CAN BY 9 W Y 

350 

CAREY 9 J 

--- 

--- 

--- 

CARY 9 Y. 

61 3 

CHAM ISSO, L Ao 

_-- 

-- - 

_-- 

CHAPYAN1Ao Wo 

--- 

113 

--- 

-- - 

--- 

-- - 

CHASE, A 

8283 

CHEESEMANITOF. --- 

83 --- 

TAXON 

04 JUL 1863 WHITNEY I 

18 YAY 1923 PURPURI FERA 

06 JUL 1895 * XANTHOC AR P A 

02 AUG 1936 ROANENS IS 

25-30 JUN 1897 DAVY I 

25 YAY 1902 

25 YAY 1902 

36 APR 1905 

18 MAY 1895 

02 JUN 1913 

22 MAY 1095 

AUSTRO-CAROL INIANA 

BUCKLEY I 

CAROLINIANA 

MISERA 

STYLOFLEXA 

BURCHELL IANA 

AGGLOYERATA 

AGGREGATA 

BUSH1 I 

FI SSA 

LUNE LL I ANA 

OKLAHOMENSIS 

03 AUG 1883 

-- MAY 1832 

--- 

MONT ANENS IS 

CARE YANA 

PLATYPHYLL A 

--9 

SYCHNOCEPHALA 

11 JUL 1910 ACBO-NIGRA 

CIRCINNATA ~ 

N I GR IC ANS 

PACHYSTACHYA 

BA LT ZE LL I C \A P MAN I 

*DIGITALIS 

MAGNIFOL I A 

*STE L LULATA 

*ST I PATA 

17 JAN 1925 *PURPUREOVAGINATA 

-- JAN 1803 

-- JAN 1882 

-- JAN 1883 

* COMANS 

DEVIA 

PETRI E I

134 


CHRISTOPHERSENT€. 

800 

CHUN9N.K. AND TS0,C.L. 

43680 

24 SEP 1929 

-_ --- 1932-1933 

CLAUSEN9R.T. 

2532 

AND WAHL9H.A. 

06 JUN 1937 

CLEMENSTJ. AND CLEMENSTM.S. 

34297 28 JUL 1933 

5546 -- MAR 1937 

CLEMENTS, Fe --- 

-- -c- 1900 

CLOKEY I I We 

3227 29 JUL 1918 

7468 19 JUN 1937 

2338 06 AUG 1915 

23 64 07 AUG 1915 

COLLINS+JeFe AND FERNAL0pM.L. 

--* 

COLLINS,J.F*; 

--- 

03-17 AUG 1905 

FERNALDTM~L. AND PEASETA~S. 

05-08 AUG 1904 

COLLINS p Z a 

COOL E Y T D 

-- - --- 

74 --- 

_-- 

--- 

COVILLE 1Fe Ve 

1455 

1457 

1362 

CRAWETI~B. AND WOODTWOA. 

11 SEP 1902 

11 SEP 1902 

03 SEP 1902 

CRAWEt J. Be 

-- - --- 

t-- q-- 

CRONQUI STTA. 

2872 

CROOM,HeBe 

c-- 

CURTI S 9 M A. 

--- 

03 JUL 1941 

-- --- 1834 

-- JUL 1841 

CURT1 SS 9 A H 

3267 -- SEP 1882 

676 1 15 APR 1901 

CUSICKTW~C. 

1331 -- JUL 1886 

2487 28 AUG 1900 

2849 3@ JUN 1902 

1331 -- JUL 1886 

DAVIS,€ 

--- --- 


TAXON 

SAVA I ENS1 S 

TSOI 

*ANGUSTIOR 

EXPLORATORUM 

SARAW AK ETENS IS 

EBENEA 

AR AP AHOENS I S 

* INTER IOR 

SUBIMPRESSA 

*TRIBULOIDES 

*GARBERI 

*FLAVA 

BARRATTI I 

* AL OPEC01 DEA 

COOL E Y I 

BR ACHY PODA 

CAYPLYOCARPA 

PACHY STOMA 

WOOD1 I 

*OLIGOCARPA 

OBOVOIDEA 

ALAT A 

, AESTIVALIS 

CREBRIFLORA 

MOHR I ANA 

CUS ICK I I 

GY MN OC L AD A 

SCABRI USCULA 

*TERETIUSCULA 

HAL SE YANA

NUMBER 12 135 

'NUMBER 

DAVIS,E --- 

-- - 

DEAM, C .C 

54764 

10927 

6458 

61 177 

DELAVAY 9 Re P 

ND LEP 

41, 3054 

39329 

41,305 

39 2 74 

GEvE. 

DATE COLLECTED 

-1 --- --- 

1823 

05 MAY 1934 

05 JUN Is12 

25 MAY 1910 

19 MAR 1941 

15 JUL 1889 

08 AUG 1938 

12 AUG 1963 

19 AUG 1961 

12 4UG 1963 

16 AUG 1961 

DUTILLYqA. i LEPAGEvE. AND DUMAN,M. 

32793 

32975 

32357 

29 AUG 1954 

03 SEP 1954 

14 AUG 1954 

EASTWOOD T A 

725A 16 JUL 1914 

EASTWOODTA. AND HOWELL, J O T . 

1101 23 JUN 1933 

2271 07 MAY 1936 

TAXON 

HI TCHCOCKI ANA 

S ICCATA 

*ART ITECTA 

LARICINA 

*LA XI F LORA 

VEXANS 

P TEROLE PTA 

COLLECTA 

YIRABI LI S 

NOVAE-ANGL I AE 

SCHWEINITZI I 

TENERA 

T R I S PERMA 

*UMBELLATA 

VALL ICOLA 


AMPHI BOLA 

AUROLENSI S 

CRUS-CORVI 

FRANKLI NI I 

HEPBURNI I 

MACROKOLEA 

PE T A SAT A 

PETR ICOSA 

PICTA 

PT YCH OC ARP A 

RETROF L E XA 

TRI ANGULAR IS 

DUTILLY I 


X PATUENSIS 

X QUEBECENSIS 

*VI RI DULA 

X EXSALINA 

X NEOPALEACEA 

*X SAXENII 

LE IOPHYLLA 

CURATOR1 UM 

OB I SPOENSI S

136 SMITHSONIAN COiXTRIBUTIONS TO BOTANY 

EGGERT T Ho 


*-- 

EGGLESTONTW~W. 

08 AUG 1893 

6181 22 AUG 1910 

65 84 18 APR-25 MAY 1911 

13567 19-20 PUG 1916 

6605 18 APR-25 YAY 1911 

3329 3@ JUL-C1 AUG 1916 

EKMANT EoLo 

14506 

--- 

21 JUL 1922 

08 AUG 1925 

H1453 12 AUG 1924 

H10662 14 SEP 1928 

ELMER T A .DOE 

3132 

8444 

88 1 

2700 

13146 

-- JUN 1901 

-- MAR 1907 

-- JUN 1897 

-- JUN 190C 

-- YAY 1911 

6983 

ENGELM~NNTG~ --- 

FARGESTROP~ 

-* NOV 1904 

-- --- 1874 

FAURI E T Uo 

--- --- 

919 

FAXONqEo AND FAXON,CoEo 

--- 

--- 

--- 

FENDLER TAO 

878 

28 JUN 1901 

23 JUN 1888 

06 JUL 1878 

27 YAY 1896 

-- --- 1847 

FERNALDqMoLo 

264 -- 

--- 

- 

146 

04 JUL 1894 

28 JUN 1904 

06 JUL 1904 

06 JUL 1893 

-- - 03 JUL 1897 

I-- 

--_ 05 JUN 1896 

30 JUN 1891 

1464 08 JUL 1909 

FERNALD,M.Lo AND BISSELLTCOH. 

203 11 16 AUG 1920 

FERNALD1M.L. AND LONGqBo 

12012 08 JUN 1940 

12969 13 JUN 1941 

8143 08 JUN 1938 

12016 13 JUN 1940 

11791 08 YAY 1940 

T AXON 

EGGERTII 

EGGLESTONI I 

GEOPHI LA 

MI SERABILIS 

PITYOPHILA 

VI RI DIOR 

CUBENSI S 

*CU BENS I S 

EKMANI I 

*EKMANI I 

DUOLfYI 

ELMER I 

*LENT’ICULARIS 

OLYHPI CA 

PALAWANENSIS 

RHYNCHACHAE NIUM 

ENGELMANNI 

F ARGESI I 

M I C RANT H A 

AENEA 

CRAWFORDII 

OR MOST ACHYA 

FENDLE R IANA 

*CRINITA 

*GLAREOSA 

*INTERIOR 

*LAXIFLORA 

MERRITT-FERNALDI I 

*MI RABIL IS 

ORONENSIS 

*SCOPARIA 

*CUMULATA 

*ABSCONDITA 

X A6 SCOND I TI FORM I 

*CRINITA 

*DEB1 LIS 

*DIGITALIS 

S

NUMBER 12 

.NUMBER DATE COLLECTED TAXON 

FERNALDTMOL. AND LONGTB. 

7767 08 APR 1938 *DIGITAL I S 

11787 07 MAY 1940 RUGATA 

20 2 96 14 JUL 1920 *SCOPARIA 

1455 27 AUG 1929 X XANTHINA 

FERNALD 9 YoL AND W I EGAND 9 K. Mo 

2897 

2897 

4918 

16 AUG 1910 

16 AUG 1910 

12-13 JUL 1911 

*HORNSCHUCHI ANA 

*HOST IANA 

*PALL E SCE NS 

42 58 

4796 

2776 

16 AUG 1910 

28 AUG 1911 

21 JUL 1910 

X P SEUDO-FULVA 

*SCOPAR I A 

W I EGANDI I 

FERNALD9M.L.; LONGvB. AND FOGG-JR.,JoYo 

1374 

1449 

1474 

20 JUL 1929 

19 AUG 1929 

31 JUL 1929 

LANGEANA 


*VE S ICARI A 

FERNALDtYoLo: LONG980 AND SMARTTROF. 

5677 22 JUN 1936 

5677 22-23 JUN 1936 

BAYARDI 

*CR US-CO R V I 

FERNALD9M.L.; 

241 1 

WEATHERBY,C.A- AND STEBBINSTGOLO 

05 JUL 1931 CLIVICOLA 

FERNAL0,M.L.; WIEGAND7K.M. AND LONG, Be 

27673 

FERNAL0,Y.L. f 

20 JSJL 1925 

W IEGANDTK~Mo~ LnNGt B. ; GILBERT-JR. 

*LIVIDA 

TF~A. AND HOTCHKI SSqNo 

27657 

FORDTC. -_- 

31 JUL 1925 

-- --- 18833 

T ERR AE-NOV AE 

SCAPOSA 

FOWLER, J o 

_- - 

--- 

--- 

--- 

--- 

FRANKTJOC. 

55 --- 

FRET2 ,C. Do --- 

FUNSTONTF. 

_- --- 1872 

-- --- 1871 

-- --- --- 1872 

-- JUL 1870 

-_ --- 1835 -- -_- 1835 

-- --- 1884 

*ADUSTA 

*FOENEA 

*MILIARIS 

PRO JECTA 

*STRAM1 NEA 

FL ACC I DULA 

STEUDE L I *GRI SEA 

139 

GARBER, AoP 

--- 

GEYERtCoA. 

332 

GRAIVTvG. Bo -- - 

GR I FF I TH t Wo 78(KEW 6074) 

30 JUL 1893 

C9 JUN 1869 

09 JUN 1869 

--- 

01 MAY 1902 

--- 

PYY SOCHL AEN A 

*AUREA 

GARB ER I 

GEYERI 

FLACCIFOLIA 

GR I FF I THI I 

96(KEW 6094) --t SANGUINEA 

137

138 


HAENKE, T --- --- 

HALEID. 

97 

HALLrEo 

5 80 

606 

-- - --- 

--- 

-- --- 

-- --_ 

1871 

1871 

583 01 AUG 1871 

60 5 

-- --c 1871 

583 01 AUG 1871 

-- - -- --- 2871 

--- -- --- 1872 

HALLIE. AND HARBOURTJOP. 

59 1 

-- --- 1862 

61 7 

-- --- 1862 

-* 59 1 

--- 1862 

587 

-- -_- 1862 

HALLTHO Yo 

2483 -- JUL-AUG 1901 

9781 25 JUL 1914 

HALLTHO Y. AND BABCOCK,HoDo 

5472 -- JUL 1904 

HALL9H. Me AND CHANDLERTHOAO 

4716 @I-02 AUG 1903 

HARPER, RoMo 

2109 09 APR 1904 

21 59 26 APR 1964 

H ARTMAN T C .V 

620 12 APR 1891 

HASSE,H.Eo 

--- -- JUL 1894 

-c- 01 YAY 1886 

HAY,G.Uo 

84 

HAYDENvFo Vo -- - 

24 JUL 1900 

_- -_- 1853 

580 

--* 

21 -- --- 1853-1854 

-- - --- 

--- 

--- 

--- 

--- 

-- - 20 JUN 1860 

HELLER, A. A. 

10820 

9429 

9975 

9841 

5797 

07 JUN 1913 

11 AUG 1908 

23 JUL I910 

16 JUL 1909 

01 JUL I902 


TAXON 

ANTHOXANTHERA 

PRESLI I 

IGNOTA 

*DE WE YANA 

HALL I ANA 

*LEPORI NA 

OREGONENSI S 

PH A E OC E PHAL A 

VI CARIA 

*WILLDENOVI I 

WONPLANDII 

HALL I I 

ILLOTA 

VI OLkCEA 

JACI NTOENS I S 

TENERAEFORMI S 

LANCIFRUCTUS 

LEPORINELL A 

HARPERI 

SMALLIANA 


HA SSE I 

+ROSEA 

*CANE SC ENS 

*ARISTATA 

*DOUGLAS1 I 

HAYDENI I 

’ LAEVI-CONICA 

MEEK11 

NE BRASK ENS IS 

RAYNOLDSI I 

ABRUPT A 

FI SSURICOLA 

HELLERI 

INTEGRA 

LACUNARUM

NUMBER 12 


HELLER 9 A. A a 

LO052 

71 87 

9067 

71 87 

HENDERSON,LaF. 

13 

5583 _-- 

140 2 

20 MAY 1910 

17 AUG 1903 

21 JUL 1908 

17 AUG 1903 

-- _-- 1883 

25 JUL 1925 

-- JUL 1884 

31 JUL 1886-20 AUG 1887 

HENRY 9 A. 

7860 _- *-- 1855-1888 

5467 t-- 

4266 

_-- 

10839 --- 

HENRY 9 J eK. 

91 52 

HERMANNt Fa J a 

04 JUN 1915 

13498 

17059 

6147 

13529 

13347 

7985 

12252 

5983 

28 4UG 1956 

15 AUG 1961 

05 JUL 1934 

28 AUG 1956 

15 AUG 1956 

13 JUL 1936 

21 AUG 1955 

15 AUG 1933 

18120 

13453 

6408 

21 AUG 1962 

26 AUG 1956 

14 AUG 1934 

HOHENACK ER 9 R a F. -- --- 

943 

1851 

HOLLI STERvNa 

14 C5 AUG 1911 

HOSAKA, E aY a 

594 04 JUL 1932 

HOUGHTON 9 Oa 

--_ 

HOWE 9 E a C a 

--- 

13 JUL 1832 

30 MAY 1887 

HOWELL 9 Ja f a 

14546 

24609 

14519 

35333 

11 AUG 1938 

23 JUN 1948 

11 AUG 1938 

06 JUN 1960 

HOWELL,J.Te 

13Q42 

HOWELLvTaJa 

AND STACEYIJ~W~ 

06 JUN 1937 

--- 

-- MAY 1880 

93s 09 MAY 1885 

2994 -- MAY 1886 

TA XON 

LE PTOPODA 

*LU ZU LA EFOL I A 

MICROPTERA 

PSEUDO JAPONI C A 

ACUT INELLA 

EA ST WOOD I ANA 

INOPS 

PANSA 

ICHANGENSIS 

LA NC I FOL I A 

*LONGICRURI S 

PRAINI I 

ARCTAEFORMIS 

ATHABASCENSIS 

* 6I PART I TA 

X DEAMII 

EUR Y STACHYA 

INCOND ITA 

*INTER I OR 

LI HNOPHI LA 

PELOCARPA 

P LEC T OC ARP A 

*PR AT I COLA 

*VULP IN01 OE A 

MERCARENSIS 

ATROSQUAMA 

* PLUV IC A 

HOUGHTON I ANA 

ROSAEOI DES 

D'ANAENS IS 

SPECUICOLA 

SUBN IGR ICANS 

TOMPKI NSI 

SONOMENSI S 

AGC EDE N S 

ACUT INA 

B RE V IC AUL I S 

139



HOWELL 9 To J 

93 7 27 YAY 1885 

--- 

-- MAY 1880 

--- -- YAY 1880 

INGALtS ,To 

--- --- 

--- --- 

JAMEStEDWIN 

JEPSONr Wo Lo 

4477 20 JUL 1911 

4476 

JOHNSON t We Mo 

20 JUL 1911 

594 

JOHNSTON? IoYe 

14 AUG 1967 

1505 31 JUL 1917 

JONES? M oE --- 

834 --- 

5345 --- 

KELLOGG? A. --- 

--f 

--- 

12 JUL 1899 

28 AUG 1878 

OR JUL 1909 

01 JUN 1874 

22 JUL 1881 

03 AUG 

--- ---- 

10 JUL 1870 

--- _-- 

1073 

-- --_ 1868-1869 

KELLOGGqA. AND HARFORD, WeGoUo 

1073 

-- --- 1868-1869 

1@69 

-- --_ 1868-1869 

1069 

-- --- 1868- 1869 

YEL SO, L 

4967 C1 AUG 1945 

6362 24 JlJL 1948 

6058 03 JUL 1947 

KELSOtL AND KELSOr E.Ho 

525 08 AUG 193-6 

KENNEDY ,Go Go 

--- 11 JUN 1899 

K I NASHI T No 

--- -- JUL 1909 

KOYAMApT 

--- 23 NOV 1958 

LEAVENWnRTHt MoCo 

--- -- -_- 1846 

--- -- --- 1845 

LEAVENWORTHI MeC AND HALE 7 D 

683 --- 

LECHLERqWo 

1136 -- OCT 1852 

TAXON 

*MARC IDA 

SPRE TA 

*STY LOS A 

TURGESCENS 

JAMES1 I 

JEPSONI I 

MAR1 PQSANA 

*MICROPTERA 

STENOPTERA 

ABOR I G I NUY 

CRANDALL I 

ELRODI 

EPAPILLOSA 

JONES1 I 

HETERONEURA 

KELLOGGI I 

NERVINA 

SU 6 F USC A 

*FESTIVA 

HARFORDI I 

*HOOD I *HOOD1 I 

EL B E RT AN A 

ERXLEBENIANA 

UNCOMPAHGRE 

HAG1 ANA 

*VE ST I TA 

’ CUNEATA 

TA MAK I 

F LACC OSPERMA 

LEAVENWORTHI I 

HALE I 

I NC I SO-DENTATA

NUMBER 12 


LEIBERG9 JOB. 

33 5 

12 5 

2556 

LEMMONT JOG. --- 

2 

LEPAGE t E 

26 JUN 1894 

10 JUL 1895 

19 JUL 1896 

-- --- 1875 

21 JUN 1882 

32078 30 JUC 1954 

33131 

LI PSKY, V. I 

27 JUL 1955 

2732 

LITVINOV,D.I. 

153 

LONG,Be 

19 JUL 1899 

-- --- 1897 

F23212 

LYALL TOAVID --- 

15 JUN 1929 

-- _-- 1861 

YACBRIDF9J.F. AND PAYSON9E.B. 

3778 13 AUG 1916 

YACDANIELS,LeH. 

1542 15 MAY 1027 

MACKEYZIE,K.K. 

2676 

2088 

4645 

*-- 

167 

9871 

23 JUN 1907 

10 JUN 1906 

26 JUN 1910 

30 YAY 1897 

-- AUG 1-901 

-- MAY 1911 

YACOUNIJ~M~ 

33728 12 JUL 1901 

MACOUN, JOHN 

134Ql 

s2 

6 

1665 --- 

53877 

22 --- 

_-- 

26624 --- 

MAGUI RE Be 

26 JUL 1887 

14 JUN 1879 

01 AUG 1882 

12 AUG 1872 

31 JUL 1891 

14 AUG 19n2 

04 JUL 1899 

31 MAY 1887 

31 MAY 188P 

24 JUN 1901 

04 JUL 1879 

16098 05 AUG 1938 

MAGUIREt 8. AN0 HOLMGSEN9A.H. 

21947 16 JUL 1943 

MAGUIRE t 6. AN0 MAGUIRE ,Re Re 

14668 16 AUG 1936 

TAXON 

PHAEOL EP I S 

PRIONPHYLLA 

VAGANS 

LEMMONI 

ULTRA 

X DUMANII 

X NUBENS 

PH I L OC R E N A 

*STENDPHYLLA 

CAESARIENSIS 

*ACUTA 

PROPOS ITA 

TAHITENSIS 

A€ STIVAL IF ORMI S 

CONVOLUTA 

CRYPTOLEPI S 

MOLE STA 

PE RGL 0 60s A 

RUGOSPERMA 

*SC IRPOIOEA 

ABLATA 

ASS I NI BOINENS I S 

*CRAWFORD11 

ELEOCHAR IS 

INCURVIFORMIS 

M I CROC HA E T A 

*MIR4BILIS 

PI PERI 

*PRATENSI S 

*STRAYINEA 

YERANTl CA 

I NT E RI MUS 

*CAMP Y L OC A R P A 

RACHI LL I S 

14 1


NUYBER DATE COLLECTED TAXON 

MAGUIREt 6.; H0BSONtD.A. AND YAGUIREIR~R. 

14013 16 JUL 1936 

MANDON t C 

1429 

--- 

YARIE-VTCTORINT(FRERE~ 

4021 0- JUN 1917 

MAR IE-V I CTORIN 9 (FRER E 1 AND ROLL AND-GERYAIN T ( FRER E 1 

25767 20 JUL 1926 

MCCALLA t W .C 

2348 28 JUL 1899 

MEAD, So Be 

--- -1- 

-_- --c 

--- *-- 

MERRILL t Eo Do 

10985 12 AUG 1917 

6505 -* YAY 1909 

6623 -- MAY 1909 

543 -- APR 1910 

4730 -- OCT-NOV 1905 

MERTENST CoHo 

MOHRTC. --- 26 APR 1897 

YOOREfJo A. 

3625 

AND STEYERMARKTJOA. 

25 JUL 1931 

MULLER9C.H. 

3520 27 SSP 1939 

MULLERTC~HO AND MULLERTY~T. 

892 28 JUN 1934 

MUTI S 9 J C o 

--- --- 

KILLIP NO05715 _- --_ 1760- 1808 

NELSON9 A. 

7124 

32 75 

73 16 

11 JUN 1900 

02 JUL 1897 

29 JUN 1900 

NELSON,A. AND YACBRIDETJ.FO 

1533 01 AUG 1911 

NELSONTA. AND NELSONTEO 

5264 

NUTTALL ,T 

17 

30 AUG 1898 

--- 

OHHI, J o 

4182 -- JUL 1933 

329 -- MAR 1933 

813 11 AUG 1931 

29 04 MAY 1931 

OLNE Y T S 1 

--_ 01 JUL 1867 

WERNACULA 

YANDONIANA 

*PAUPERCULA 

*I NFL AT A 

SC I RP I FORM1 S 

I LLI NOENSI S 

MEAD I *RICHARDSON I 

BAMBUSETORUM 

* BRU NNE A 

YERRILLII 

PYCNOTHYSOS 

SU BT R ANS V ERS A 

LEIOCARPA 

* STI P ATA 

MUR I CU L AT A 

P E RC 0s TAT A 

MACKENZIANA 


BREVISQUAMA 

FESTI VELLA 

S I MULATA 

APODA 

NE LS ON1 I 

NUTTALL I I 

A PODOST ACHY A 

HATUS I MANA 

KURI LENS1 S 

RUGATA 

HORYATHODES

NUMBER 12 

PAINEt J c A. 


--_ __- 

PALMERTE. 

546 -- JUN 1890 

PAR I SH 9 So R 

2485 03 JUL 1892 

3609 -- JUN 1895 

41 44 04-13 ApR 1896 

PARRY qC C o 

--- -- --- 

PARRY9C.C. 

39 6 

18SO 

AND LEMMONt JOG. -- --- 1876 

PAYSONTE~B~ AND PAYSONTL-B. 

2224 06 AUG 1920 

PEASEpAoSo AND LONGvB. 

2C519 

PECK1 C o H o 

--- 

5 

--- 

--- 

09 JUL 1920 

-- JUN 1894 

-- JUN 1893 

-- JUN 1894 

-- JUN 1892 

PECK TM. IF. 

13 13 AUG 1917 

PETELOT CP A. 

5325 

PETRIEt Do _-- 

--- 

-- JUL 1930 

_-- 

-- JAN 1880 

PITTIERqH. 

3376 

AND TONDUZrA. 

19 JAN 1891 

3381 19 JAN 1891 

PORSIL0,A.E. 

173 26 AUG 1937 

PORSILD,A.Eo 

5120 

71 1 

713 

PORTER 9 T C --- 

--- 

-- - 

PREISST Lo 

1825 

1861 

AND P0RSILDqR.T. 

02 AUG 1928 

23 JUN-05 JUL 1926 

23 JUN-05 JUL 1926 

--_ 

26-29 JUN 1871 

28 AUG 1871 

--_ 

-- JUL 1839 

PRINGLE TC. Go 

4275 03 OCT 1892 

4839 19 AUG 1894 

10039 

4838 

26 AUG 1905 

25 4UG 1894 

TAXON 

*VAGINA TA 

*NUDATA 

AUSTROMONTANA 

MU L T I C OSTA TA 

VITREA 

BARBARA E 

ALMA 

PAY SON1 S 

*LASIOCARPA 

*ROSE A 

*ROSEA 

*ST IPATA 

*TORTA 

PRAECEPTOR IUM 

TR ICHOPHYLLA 

KALOI DE S 

LONGICULMIS 

DURANDI I 

*LE MANNI ANA 

MORRI SSEYI 

*ATROFUSCA 

KOKR INENSI S 

MELOZITNENSIS 

*QEBILIS 

HALL I I 

PORTER1 

PREI SSI I 

PREI SSI I 

AUTUMNAL1 S 

AZTECICA 

C I L I ARI S 

FELIPENSIS 

143



--- 

PRINGLE T C G. 

48 39 

23 AUG 1881 

19 AUG 1894 

3126 08 JUL 1890 

40 40 27 AUG 1894 

4842 29 AUG 1894 

8863 02 JUN 1904 

2630 05 JUN 1889 

4685 

3801 

--- 

7452 

08 JUN 1894 

04 AUG 1891 

19 AUG 1881 

24 JUN 1897 

PYRON, J.H. AND MCVAUGHTR~ 

2951 15 MAY 1938 

RAMOSTM. 

BUR. 

RAUPTH. M. 

SCIo 1434 

AND SOPER 9 J.H. 

-- AUG 1906 

9534 18 dUL 1939 

RAY 9 J o D i WOOOTC E : S # IT HI A .C AND EATON , Re J 

10750 

R I CH, W P --- 

R ICH 9 WILL I AM 

WILKES EXPED. 1241 

26 APR 1961 

05 JUN 1894 

-- --- 183 8-1842 

_- - --- 

--- 

RICHARDSON, J o 

41 7 

1- - 

--f 

-- - 

ROACH, A. W 

20 2 

ROCK,J.F. 

9017 

ROSE, J. No 

2357 

R0SEqJoN.i PAINTER,JoH. 

90 19 

RDUSSEAUv J 

24989 

RUSBY 9H.H. 

859 

RUTH, A. 

360 

45 8 

--- 

--9 

10 JUN 1949 

-- OCT 1909 

16 AUG 1897 

AND ROSE, J.S. 

-- 1905 

27 JUL 1926 

-- --- 

1883 

12 AOR 1913 

24 APR 1914 

-- JlJL 1900 

TAXON 

FRACTA 

FUSCOT I NCT A 


MACROSP ERMA 

OAXACANA 

PERLONGA 

PERSTRICTA 

*PI NETORUM 

PR 1 NGLE I 

*SCIRPOIDEA 

SEATONIANA 

AM PL IS QUAM A 

RAMOSI I 

SOPERI 

*FISSA 

*TEN ERA 

LAC1 NI ATA 

ARCTICA 

BACKANA 

BACK I OUR I FOL IA 

FE ST I VA 

HUOKERANA 

PARRYANA 

PODOC ARPA 

RICHARDSON1 I 

DIVERSISTYLIS 

KA UA I E NSI S 

,MADRENS IS 

ST E LL AT A 

*OEDERI 

RUSBYI 

BULBOSTYLIS 

ONUSTA 

RUTH I I

NUMBER 12 

.NUMBER DATE COLLECTED 

RYDBERG9P.A. 

23 39 

SANDBERG9 J*H* 

07 AUG 1895 

933 20 AUG 1892 

SANDBERG9J.H. AND LEIBERGIJ.~~ 

194 09 JUN 1893 

773 

SARTWELL? HOP 0 

18 AUG 1893 

-- - 

7-- 

56 

--- 

56 

--- 

--- 

--- 

108 -- --- 

78 

--- 

--- --- 

--- 

--- 

1848 

36 

12 

72 t-- 

35 

-v- 

138 

--- 

SAVATI ER TL. 

1414 -- --- 1866-1 874 

1404 -- _-- 1866- 1874 

2959 

-- --- 186 6 -1 874 

1413 

-- --- 1966-1 874 

SCHAFFNERpJoG. 

22 1 

-- 1877 

5 46 

_- 

--- 

1877 

546 

-- 

--- 

--- 1877 

SCHNEIDERTCO 

2738 06 AUG 1914 

SCHNEIDER TR. A. 

954 25 JUL 1938 

SCHWEINITZ 9LoDo 

--* --- 

--- q-- 

--- -_ --- 

SCOTT 9 ( MISS 1 

SCOULERT J 

--- 

--- --- 

0-0 

--- 

1880 

296 

SEK IMOTO T H o 

--- 15 JUL 1932 

SHARP9A.J. 

45450 29 APR 1945 

45 144 07 FEB 1945 

SHELDON1E.P. 

88 54 09 SEP 1897 

TAXON 

IDAHOA 

*HINDS1 I 

*NEBRASKENS I S 

NEUROPHORA 

AL OPEC0 IDE A 

AQUATIL IS 

*AQUATICIS 

*CEPHALOPHORA 

C OMMU N I S 

*CR I N I TA 

FORMOSA 

INTER I OR 

SARTWELLI I 

SCABR IOR 

*STERI LI S 

* TE NTAC ULATA 

MACR OGLOS S A 

*NUTANS 

PLANATA 

PODOGYNA 

FUSCOL UTE A 

POTOSI NA 

SCHAF FN ER I 

SCHNE I DER I 

*EGGLESTONI I 

COSTATA 

GRAC I CL I MA 

PRAEGRACIL IS 

A+ ER TA 

COLUMB I ANA 

SC OUL ER I 

HY ME NO DON 

AT R ACT0 D ES 

QU ICHE NS I S 

SHELDON I I 

145

146 


SMITH9C.P. 

1055 

SYITH9J.De 

659 

24 JUL 1905 

AND TURCKHEIM9H. 

-- JUN 1885 

SMITHtSeJ. AND DUNCAN9W.H. 

4872 

STo JOHNvH. 

06 APR 1949 

18330 18 AUG 1937 

STeJOHN9He AND FERNALD9M.L. 

1C801 

STEELE, EeSe --- 

_-- --- 


16-17 JUL 1914 

-- -0- 1896 

23 YAY 1898 

0- f-- 1900 

53105 15 JUN 1943 

48347 07 JUL 1942 

62605 06 MAY 1945 

49055 18 JUL 1942 

55470 11 FER 1944 

5887C 28 SEP 1944 

48 542 14 JWL 1942 

57367 15 JUL 1944 

5740 1 15 JUL 1944 

50 150 06 AUG 1942 

53095 15 JUN 1943 

48334 07 JUL 1942 

62705 10 MAY 1945 

48554 

SUKSDORF9W.N. 

12347 

14 JUL 1942 

--- 

12348 15 SEP-23 OCT 1927 

12359 23 OCT 1927 

6864 16 4UG 1909 

51 81 15 JUL 1905 

1284 26 JUN 1386 

11551 21 AUG 1924 

12333 22 AUG-05 SEP 1927 

10 249 22 JUN 1919 

81 6 02 JUN 1885 

1296 

1315 

13 AUG 1897 

-- JUL 1883 

7383 27 AUG 1912 

5259 

5 ULL I VANT 9 W S 

21 AUG-20 SEP 1905 

--- -c1 

--- --_ 

SVENSON T H 9 K 

10469 12 JUL 1939 


TAXON 

MONT EREY ENS I S 

DONNELL-SM ITHI I 

*WILLDENOW1 I 

VITIENSIS 

MISANDROIDES 

ANGUST I OR 

MEDITERRANIA 

ME SOCHUREA 

AZU AY AE 


CULMEN ICOLA 

HUEHUETECA 

LARENSI S 

RORAIMENSIS 

STEYERMARKI I 

TACH I R ENS1 S 

TAMANA 

TOJQUI ANENS I S 

TO RE ADOR A 

TUNI MANENS IS 


VENOSI VAGI NATA 

*APERTA 

*AP ER TA 

*APERTA 

CUN STANCEANA 

EGREGIA 

EU RY C A RP A 

*EURYCARPA 

* INTERRUPTA 

*NEBRASKENS IS 

OXYC ARPA 

PADDOENSIS 

S UBOR 8 I CU L AT A 

SUKSDORF I I 

*SUKSDORFII 

*ALATA 

CONJUNCTA 

*oXY L EP I S

NUMBER 12 

TAAMpYo Wo 

,NUMBER DATE COLLECTED 

50 2 

TAKpToWo AND CHOWqWoKo 

1-16 APR 1938 

3202 

THURBERt Go 

652 --- 

20 NOV 1926 

--- 

-- JUN 1850 

TORREVT~. 

_-- 03 AUG 1839 

TOWNSfjNDtC.H.T* AND BARBES9C.M. 

157 

1RACYpJ.P. 

21 JUL 1899 

4547 04 JUL 1914 

3783 21 JUL 1912 

TRACY p So Mo 

17 23 MAY 1888 

TSAI ,HOT. 

62809 

_- --- 1934 

TSUI p T oM 74 -- YAR-APR 1932 

TUCKERMANqEo 

--- --- 

--- -- JUN 1864 

-- - --- 

---- _--- 

UN D E R NO 0 0 9 L Y 

158 29 JAN 1903 

VAHL T Jo 

VASEY p G m 

t-- 

--- 

--- 

--- 

8650 

-- --- 

VENTURI 9 So 

6491 15 FEB 1927 

1929 

VREELANDIF~Y~ 

1121 19 AUG 1901 

WAREpRo A. t ROLLINS 7s. AND KNIGHftO.Wo 

5066 

WATERFALL tU.Te 

05 JUL 1906 

11380 

HATSONIS. 

1231A 

1246 

WERDERMANNp E. 

1687 

19 APR 1953 

-- AUG 1869 

-- MAY 1868 

-- MAR 1925 

WHEELER9C.F 

--- 28 JUN 1890 

HI EGANDt Ko Mo AND THOYAS , CoCo 

1915 15 JUN 1914 

TAXON 

SURCULOSA 

OBL ANC EOL ATA 

*GAYANA 

T HUR 6 ER I 

HETEROSTACHYA 

TOWNSENDI I 

TRACY I 

UNILATERALIS 

*STRAMINEA 

ZIZANIAEFOLI A 


AR GVRANTHA 

GL AUCODEA 

*SCOPARIA 

* SCOPAR I A 

UNDERWOOD1 I 

PR AT ENS I S 

*HIRSUTA 

* PHA L ARO I DE S 

SALTAENSIS 

HOLM1 ANA 

*TR IS PERMA 

L~TEBR ACTE AT A 

WANE SC ENS 

WATSON I 

WERDERMANNII 

*TENUI F LORA 

*AMPHIBOLA 

147



WILKES EXPLOR. 

--- 

_I- 

-7- 

--- 

W I LL IAYS T E F 

--- 

W I LLI AMS T Em F. 1 

WILLIAMS 9 € 0 F. i 

110 

WILLIAMS ~L.00 

13178 

WILLIAMSTR.S. 

--- 

--- 

--- 

WILL I AMS TT a A 

2951 

WOODtW. A. 

--- 

--- 

W R I GHT 9 C --- 

-T - 

WOOTON, E o 0. 

EXPED. 

-- --- 1838-1842 

-- --- 1838-1842 

-* --_ 1838- 1842 

-0 --- 1 83 8- 1 842 

-- AUG 1900 

CHURCHILL, JeR. AND FERNALOt MoL 

16 Jut 1900 

COLLINS7J.F. AND FERNALDTM-L. 

12-15 JUL 1905 

19 JUL 1992 

06 JUL 1947 

18 JUN 1899 

07 JUN 1893 

11 AUG 1894 

12 JlJN 1899 

18 JUN 1899 

19 AUG 1897 

28 JUL 1900 

TAXON 

HEBE TATA 

*MICROGLOCH IN 

OL IGANTHA 

WILKESI I 

*VESICARI A 

KATAHDINENSIS 

*DE WE YANA 

X TRICHINA 

ST ANDLEYANA 

BONANZENSI S 

CONCINNOIDES 

STANTONENSIS 

WILL IAMSII YUKONENS I S 

PLATYLEPIS 

RETROC URVA 

AGROSTOIDES 

CONF ERTIFLORA 

MICANS 

NA NA 

PAPULOS A 

PARCIFLORA 

PICTA 

WRIGHT1 I 

*STRICTA 

XEROCARPA

NUMBER 12 149 

COUNTRY 

AFGHANISTAN 

ARGENTINA 

AUSTRAL I A 

BOLIVIA 

BRAZIL 

BRITISH HONDURAS 

CANADA 

GEOGRAPHIC INDEX 

STATE TAXON 

JUJUY 

GR I F F ITH I S A NGU I N E A 

*PHALAROIDES 

SALTA 

TIERRA DEL FUEGO (TER 

SALTAENSIS 

BANKS1 I 

PREISSI I 

WESTERN AUSTRAL I A PREISSI I 

*C L ADO STAC H Y A 

* JAME S ON I 

MANDONIANA 

RIO DE JANEIRO 

*PURPUREOVAGI NATA 

CAY0 

BARTL ETT I PARRY ANA 

PODOCARPA 

ALBERTA 

RICHARDSON1 I 

ATHABASCEN S I S 

ALBERTA 

ATROSQUAMA 

ALBERTA 

EURYSTACHY A 

ALBERTA I NCONDI TA 

ALBERTA 

INCURVIFORMI S 

ALBERTA 

ALBERTA 

*PR AT! COLA 

SC IRPIFORMI S 

BRITISH COLUMBIA 

ABLATA 

BRlTI SH COLUMBIA 

ARCT AE FORM I S 


PIPER1 

BR IT1 SH COLUMBI A 

*PRATENS IS 

BRIT1 SH COLUMBIA 

*SCIRPOIDEA 

MANITOBA 

ASSINIBOINENSIS 

MAN I T 06 A 

DUTILLYI 

MAN I TOBA 

*LONGIROSTRIS 

NEW BRUNSWICK 

NEW BRUNSWICK 

*ADUSTA 

*C ANESCEN S 

NEW BRUNSWICK 

*FOENEA 

NEW BRUNSW I CK 

*MIL IAR I S 

NEW B RUN SWI CK 

*MIRABILIS 

NEW BRUNSWICK 

PROJECTA 

NEW BRUNSWICK 

*STRAMINEA 

NEWFOUNDLAND 

*HORNSCHUCHIANA 

NEW FO UN DL AND 

*$IOSTI ANA 

NEWFOUNDLAND 

LANGEANA 

NEWFOUNDLAND 

*LIVIDA 

NEW FOUNDLAND 

MI SANDROI DE S 

NEWFOUNDLAND 

MORRISSEYI 

NEWFOUNDLAND 

NEW FOUNDLAND 


*PALL E SC EN S 

NE WFOUNDLANO 


NEWFOUNDLAND 

*SCOPARI A 

NEWFOUNDLAND 

NE WFOUNDLAND 

TERRAE-NOVAE 

*VESICARIA

150 

COUNTRY 

CANADA 

CHILE 

CHINA 


STATE TAXON 

NEWFOUNDLAND WIEGANDII 

NEW FOUNDL AND X XANTHINA 

NORTHWEST TERRITORIES *ATROFUSCA 

NORTHWEST TERRITORIES SOPERI 

NOVA SCOTIA *CUMULATA 

NOVA SCOTIA *LASIOCARPA 

NOVA SCOTIA *SCOPARIA 

ONTAR I0 *STRAMINEA 

QUEBEC CLIVICOLA 

QUEBEC *CRAW FORD 1.1 

QUEBEC *DEWEY ANA 

QUEBEC X DUMANII 

QUEBEC X EXSALINA 

QUEBEC *FLAVA 

QUEBEC *GARB ER I 

QUEBEC *GLAREOSA 

QUEBEC *INFLATA 

QUEBEC X NEOBIGELOWII 

QUEBEC X NEOPALEACEA 

QUEBEC X NUBENS 

QUEBEC *OEDERI 

QUEBEC X PATUENSIS 

QUEBEC 

QUEBEC 

*PAUPERCULA 

X QUEBECENSI S 

QUEBEC *X SAXENII 

QUEBEC *VI RI DULA 

SASKATCHEWAN ARCTIC A 

SASKATCHEWAN BACKANA 

S A SK A TC H E WAN 

BACK1 I 

SASKATCHEWAN OUR1 FOL IA 

SASKATCHEWAN EL EOCHAR I 

S A SK A TC H E W AN HOOKERANA 

SASKATCHEWAN XERANTI CA 

YUKON TERRITORY BONANZENSIS 

YUKON TERRITORY LEIOPHYLLA 

YUKON TERRITORY MI CROCHAET A 

YUKON TERRITORY WILLIAMS11 

YUKON TERRITORY YUKONENSIS 

LLANQUIHUE WERDERMANNII 

MAGALLANES vMICROGLOCH1 N 

MAGALL AN E S OLIGANTHA 

HUPEH ICHANGENSI S 

HUPEH LANCIFOLIA 

HUPEH *LONGICRURIS 

HUPEH AND HONAN CHI KUNGANA 

KIANGSI 

K WANGTUNG 

KUL I NGANA 

B AM BU S ETORU M 

KWANGTUNG 

OB LA NC E OL A T A 

S 

INC I SO-DENTATA

NUMBER 12 

COUNTRY 

CHINA 

COLOMBIA 

COSTA RICA 

CUBA 

ECUADOR 

FIJI 

FRENCH POLYNESIA 

GREENLAND 

GUATEMALA 

HAITI 

STATE 

KW ANGTUNG 

KWANGTUN G 

KW AN G T UN G 

KW ANG TUN G 

SZECHWAN 

YUNNAN 

YUNNAN 

YUNNAN 

YUNNAN 

OR I ENTE 

AZUAY 

AZUAY 

ALTA VERAPAZ 

HU EHU ETEN AN GO 

HUEHUETENANGO 

HUEHUETEN ANGO 

HU EHU E l E NANGO 

HUEHUETENANGO 

HUEHUETENANGO 

HUEHUETEN ANGO 

JALAPA 

QUICHE 

OUEST 

INDIA 

INDONESIA WEST NEW GUINEA 

WEST NEW GUINEA 



JAMAICA 

JAPAN 

AOMORI (PREFECTURE) 

HOKKAIDO (PREFECTURE) 


HOKKAIOO (PREFECTURE) 



KANAGAWA (PREFECTURE) 



TAXON 


SCAPOSA 

SURCULOSA 

TSOI 

FARGESI I 

PRAfNII 

PT EROL EPTA 

SCHNEIDERI 

Z I Z AN I AE FOL I A 


DURANDI I 

*LEMANNIANA 

CUBENSIS 

AZUAYAE 

TOREADORA 

VITIENSIS 

TAHITENSIS 

PRATENS I S 

DONNELL-SMITH1 I 


GUATE MAL EN S I S 

HUEHUETECA 

STEYERMARKII 

TOJQU I ANENS I S 



STANDLEY ANA 

QUICHENSIS 

*CUBENS I S 

*EKMANI I 

EKMANI I 

MERCARENSI S 

ACROPHI LA 

ERE MOSTACHYA 

MELANOPHORA 

PERILEI A 

UNOERWOODI I 

HY MENODON 

MICANS 

RUGATA 

CMNEAT A 

CONFERTIFLORA 

NANA 

PAPULOSA 

PARCIFLORA 

PICTA 

MACROGLOSSA 

*NUTANS 

PLANATA 

151

152 

COUNTRY 

JAPAN 

KOREA 

MALAYSIA 

MEXICO 

NEW ZEALAND 

PAPUA AND NEW GUINEA 

PERU 

PHILIPPINES 


STATE TAXON 

TOKYO (PREFECTURE) 

SABAH (TERRITORY) 

CHIAPAS 

CHI APAS 

CHIHUAHUA 

CHIHUAHUA 

CHIHUAHUA 

DURANGO 

HIDALGO 

HIDALGO 

HIDALGO 

HIDALGO 

MEXICO 

MICHOACAN 

NUEVO LEON 

NUEVO LEON 

NUEVO LEON 

OAXACA 

OAXACA 

OA XAC A 

OAXACA 

OAXACA 

OAXACA 

PUEBLA 

SAN LUIS POTOSI 



SAN LUIS POTQSI 


SONORA 

SONORA 

CANTERBURY (DISTRICT) 

CANTERBURY (OISTRICT) 

NELSON (DISTRICT) 

OTAGO (DISTRICT 1 

OTAGO (DISTRICT) 

NORTH -E A ST NEW GUINEA 

PAPUA (TERRITORY] 

PAPUA (TERRITORY) 

LIMA 

BATAAN 

BENGUET 

B ENGU ET 

BENGUET 

BENGUET 

NEGROS OCCIDENTAL 

PAL AWAN 

RIZAL 

PODOGYNA 

MI CR ANTHA 

EXPLORATORUM 

ATR AC TOD E S 

CHIAPENSIS 

CHIHUAHUAENS IS PERCOSTATA 

TOWNSENDII 

MADRENS IS CILIARIS 

PERLONGA 

SEATONI ANA 

STE LLATA 

AUTUMNAL I S 

ARSENI I 

*EGGLESTONII 

MACKENZIANA 

PERSTRICTA 

AZTEC ICA 

FELIPENSIS 

FUSCOTI NCTA 

MACROSPERMA 

OAX AC AN A 

*PINETORUM 

CONSPECTA 

FUSCOLUTEA 

I NVOLUC RAT ELLA 

POTOS INA 

PR INGLEI 

SCHAFFNERI 

*GAYANA 

THURBERI 

*C OMAN S 

PETRIEI 

DEVIA 

KALOI DES 

LONGICULMIS 

SARAHAKETE NSI S 

BREVIS 

LAMPROCHLAMYS 

HE B E TATA 


*BRUNNEA 

ELMER1 

MERRILLII 

SUBTRANSVERSA 

P Y CNO THY SO S 

PALAW ANENS IS 

RAMOSI I

NUMBER 12 153 

COUNTRY STATE 

RYUKYU ISLANDS OKINAWA (PREFECTURE) 

OK1 NA WA ( PREFECTURE 1 

SOUTH AFRICA 

ST. PIERRE AND MIQUELON 

SW ITZERL AND VALAI S (CANTON 1 

TAIWAN 

FUKIEN 

USA 

ALABAMA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ALASKA 

ARIZONA 

AR I ZONA 

ARIZONA 

ARIZONA 

AR I ZONA 

ARKANSAS 

ARKANSAS 


CALIFORNIA 

CALIFORNIA 

CALI FORN IA 

CALIFCRNIA 

CALIFORNIA 

CALIFORNIA 

CALIFORNIA 

CAL I FORN I A 

CALI FORN I A 

CALIFORNIA 

CALIFORNIA 

CALI FCRNIA 


CALI FORN I A 

TAXON 

TAMAK I TETSUOI 

BURCHE LL I ANA 

FULVE SC E NS 

*FLAVA 

APODOSTACHYA 

HATUSIMANA 

*CANESCENS 

COLUMBIANA 

FRANKLIN11 

GEYERI 

JAMES I NUTTALL I PET AS AT A 

PETRI CO SA 

*UMBELLATA 

*ST I P ATA 

ANTHOXANTHERA 

C I RC I NNATA 

JACOBI-PETER1 

KOKRINENSIS 

LEIOCARPA 

MELOZITNENSI S 

NIGRICANS 

P ACHY S T ACHY A 

PHY SOCHLAE NA 

PRESLII 

CURATORIUM 

*NUDATA 

RUSBY I 

SPECUICOLA 

ULTRA 

BUSH1 I 

*ROSEA 

SC OULER I 

ABRAMS I I 

ABRUPTA 

AEQUA 

ALBIOA 

ALMA 

RTHROSTACHYA 


AUSTROMONTANA 

BARBARAE 

BRAINERCI I 

BREWER1 

*BRONGNIART I CALIFORNICA 

C I NNAMOMEA

154 

USA 

COUNTRY 

STATE 

CALI FCRN I A 

CALIFORNIA 


CALIFCRNIA 

CALI F ORNI A 

CAL IF ORN IA 

CALIFORNIA 

CAL I F ORN I A 

CAL IF CRN I A 

CALIFCRNIP 

CALIFORNIA 

CALIFCRNIA 

CALIFCRNIA 



CALIFORNIA 

CALI FCRN IA 

CALIFGRNIA 

CALIFORNIA 

CALI FORN I A 

CALIFCRNIA 

CAL I FCRN I A 

CALIFORNIA 

CALIF CRNI A 

CALI FORN I A 

CALIFORNIA 

CALI FCRNI A 

CALIFCRNIA 

CALIFGRNIA 

CALI FORN I A 

CAL I FORN IA 

CALIFCRNIA 

CALIFORNIA 

CAL IFORN I A 

CALIFCRNIA 


CAL IFORN I A 

CALI FCRN I A 

CALIFORNIA 

CALIFORNIA 

CALIFCRNIA 


CALIFORNIA 

CALI FORN I A 


CALIFORNIA 

CALI F ORN I A 

CAL I F ORN I A 

CALIFCRNIA 

SMJTHSONIAN CONTRIBUTIONS TO BOTANY 

TAXON 

DANAENS I S 

DAVY I 

DEBILIFORMIS 

CUDLEY I 

*ECHINATA 

*FE ST I VA 

FETA 

*FILIFOLlA 

FLACC I FOLIA 

FRACT A 

GRAC I LI OR 

GYNODYNAMA 

HARFORDI I 

HASSEI 

HAYDENIANA 

HETERONEURA 

*HOOD I *HOOD I INTEGRA 

JACINTOENSIS 

JEPSONI I 

JONES I I 

KELLOGGI I 

LAC I N I ATA 

LACUNARUM 

LANC I FRUCTUS 

LEMMONI 

LEPORINELLA 

*LUZULAEF OL I A 

LUZUL I NA 

MAR1 POSANA 

MENDOC INEN S I S 

MONTEREYENSIS 

MULTI COSTATA 

NE RV I N A 

NUDATA 

OBISPOENSIS 

PACHY CARPA 

PAUCICOSTATA 

PHYLLOMANI C A 

PRAEGRACILIS 

P SEUD 0 J A PON I C A 

OUADR I F IDA *OUAORI F IDA SAL I NA EFORM I S 

SARTWELL I ANA 

*SC I RPOI DEA 

*SCOPARI A 

SONOMENSIS

NUMBER 12 

COUNTRY 

STATE 

US A CAL IFORN I A 

CALIFORNIA 

CALIFCRNIA 

CALIFORNIA 

CALIFCRNIA 

CALI FCRNI A 

CAL I FCRN I A 

CALI FCRN I A 

CALIFCRNIA 

CALI FCRN IA 

CALIFCRNIA 

CALIFORNIA 

CALIFORNIA 

CALIFCRNIA 

CALI FORN I A 

COLOR ADO 

COLORADO 

COLORADO 

COLOR bD0 

CO L OR ADO 

COLORADO 

COLOR ADO 

COCORADO 

COLORADO 

COLOR ADO 

COLORADO 

COLOR ADO 

COLOR ADO 

COLOR ADO 

COL OR ADO 

COLORADO 

COLOR ADO 

COLOR ADO 

COLORADO 

COLORADO 

COLORADO 

COLOR ADO 

DISTRICT OF COLUMBIA 



FLORIDA 

FLORI DA 

FLORIDA 

FLORIDA 

FLORIDA 

FLORIDA 

FLORIDA 

FLORIDA 

FLOR I DA 

TAXON 

SPEC1 FICA 

STENOPTERA 

SUB-BRACTEATA 

SUBFUSCA 

SUBNIGR ICANS 

TENERAEFORMIS 

TOMPKI NSI 

TRACY I 

TUMUL I COLA 

UNILATERAL1 S 

VITREA 

WHITNEY I 

WHITNEY I 

WHITNEY I 

WILKESI I 

ARAPAHOENS I S 

*BIPARTITA 

*BONPLANDI I 

CHALC IOLEP I S 

CRANDALLII 

EBENEA 

EGGLESTONII 

ELBERTANA 

ELY NO I DE S 

ENGELMANN I 

ERXLE BENIANA 

*FESTIVA 

HAG1 ANA 

HALL1 I 

HEPBURNII 

ILLOTA 

*MI CROPTERA 

NUBICOLA 

PERGLOBOSA 

SAXIMONTANA 

UNCOMPAHGRE 

V I OLAC EA 

ANGUSTIOR 

MEDITERRAN I A 

M E$ OC HOR E A 

BALTZELLII 

CHAPMAN1 

CREBRIFLORA 

*DIGITALIS 

*FISSA 

FLACCOSPERMA 

MAGNIFOLIA 

MOHRI ANA 

*STELLULATA 

155

156 

USA 

COUNTRY 

STATE 

FLORIDA 

FLORIDA 

GEORGIA 

GEORG I A 

GEORGIA 

GEORGIA 

HAWAII 

HAWAI I 

HAWAI I 

IDAHO 

IDAHO 

IDAHO 

IDAHO 

IDAHO 

IDAHO 

IDAHO 

IDAHO 

IDAHO 

I L LINOI S 

ILLINOIS 

ILL INCI S 

ILLINOIS 

ILLINOIS 

ILLINOIS 

INDIANA 

INDIANA 

INDIANA 

INDIANA 

KANSAS 

LOUI S I ANA 

LOUI S I AN A 

LOUISIANA 

LOUI S I ANA 

LOUISIANA 

LOUISIANA 

LOUI S I ANA 

LOUI SIANA 

LOUISIANA 

LOUISIANA 

MAINE 

MAINE 

MAINE 

MAINE 

MA I NE 

MAINE 

MAINE 

MA I NE 

MAINE 

MAINE 


T AXON 

*STI PATA 

VEXANS 

AMPLI SQUAMA 

HARPER I 

SMALLIANA 

*WI LLDENOWI I 

KAUAIENSIS 

*PLUVICA 

*WAHUENSIS 

ABOR I GI NUM 

APODA 

HALL1 I 

*HINDS11 

IDAHOA 

OBOVOI DEA 

PR I ONPHYLL A 

PROPOSI TA 

RAYNOLDSII 

*HI RSUTA 

ILLINOENSI S 

MEAD1 I 

*RICHARDSON11 

SUB1 MPRESSA 

*TRIBULOIDE S 

*ARTITECTA 

X DEAMII 

LARICINA 

*LAXI FLORA 

MOLES TA 

AMPHI BOLA 

AUROLENSIS 

CRUS-CORVI 

HALE1 

I GNOTA 

LEAVENWORTHII 

MACROKOLEA 

PICTA 

PTYC HOC ARP A 

TURGESCENS 

*CRI NI TA 

*INTERIOR 

KATAHDI NENS I S 

*LA XI FLORA 

MERRITT-FERNALDI I 

ORONENS I S 

*PAUPERCULA 

PORTER1 

*SCOPARIA 

X TRICHINA

NUMBER 12 

COUNTRY 

STATE 

USA MAINE 

MAINE 

MA S S ACHU S ET TS 

MASSACHUSETTS 

MASSACHUSETTS 

MASSACHUSETTS 

MASSACHUSETTS 

MASSACHUSETTS 

MASSACHUSETTS 

MASSACHUSETTS 

MASS ACHUS ETT S 

MASSACHUSETTS 

MASSACHUSETTS 

MASS A CHU S ETT S 

MAS S A CH U S ETT S 

MA S S ACHU SETT S 

MASSACHUSETTS 

MASSACHUSETTS 

MICHIGAN 

MICHIGAN 

M ICHI GAN 

M IC H I GAN 

M ICHI G AN 

MI CHI GAN 

M IC H I GAN 

MICHIGAN 

MI NNE SOTA 

MISSISSIPPI 

MISSOURI 

MI SSOURI 

MISSOURI 

MISSOURI 

MONTANA 

MONTANA 

MONTANA 

MONTANA 

MONTANA 

MONTANA 

N E BRA SKA 

NEBRASKA 

NEBRASKA 

NEBRASKA 

NEBRASKA 

NEVADA 

NEVADA 

NEVADA 

NEVAOA 

NEVADA 

NEW HAMPSHIRE 

TA XON 

*TR I S P ERMA 

*VE SIC ARIA 

AR GY RAN THA 

COLLECTA 

G L AUC ODE A 

HA LSEYANA 

HITCHCOCKI ANA 

INC @MPERTA 

MIRAB IL I S 

NOVAE-ANGLIAE 

SCHWEINITZI I 

*SCOPARI A 

*SCOPARI A 

SI CCATA 

TENERA 

*TENERA 

TR I SPERMA 

*VESTITA 

*ALOPECOIDEA 

COOLEY I 

HET EROST AC HY A 

*INTER I OR 

PRA?REA 

*RETROCURVA 

*TENUIFLORA 

*VULPINUIDEA 

HOUGHTONIANA 

*STRAMINEA 

AGGLOMERATA 

AGGRE GATA 

EGGERTI I 

LUNELLIANA 

CO NC I NNO I D E S 

ELRODI 

HOLM1 ANA 

MONTANENSI S 

PL ECTOCARPA 

STANTONENS IS 

*ARI STATA 

*DqUGLASI I 

LAEVI-CON1 CA 

MEEK1 I 

NEBRASKENSIS 

FISSURICOLA 

HELLERI 

*INTERIOR 

MICROPTERA 

WATSON1 

AENEA 

157

158 

US A 

COUNTRY 

STATE 

NEW HAMPSHIRE 

NEW HAMPSHIRE 

NEW H AMP SHI RE 

NEW HAMPSHIRE 

NEW JERSEY 

NEW JERSEY 

NEW JERSEY 

NEW JERSEY 

NEW JERSEY 

NEW JERSEY 

NEW JERSEY 

NEW MEXICO 

NEW MEXICO 

NEW MEXICO 

NEW MEXICO 

NEW MEXICO 

NEW Y ORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YCRK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW Y ORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NEW YORK 

NORTH CAROL I NA 

NORTH CAROLINA 




TAXON 

CRAWFORDII 

*MIRABILIS 

ORMOST ACHY A 

*SCOPARI A 


BARRATT I CAESARIENSIS 

CO NVOL UT A 

CR I ST ATA 

CRYPTOLEPI S 

RUGOSPERMA 

AGROSTOIDES 

FENDLERIANA 

GEOPH I LA 

NEOMEXICANA 

PITYOPHILA 

ABDITA 

A L OPEC 0 ID E A 

*AMPHIBOLA 

AQUAT IL I S 

*AQUATILI S 

CAREYANA 

*CEPHALOPHORA 

COMMUNI S 

*CRINITA 

FORMOSA 

INTERIOR 

*OLIGOCARPA 

PLATYPHYLLA 

RETROCURVA 

ROSAEOIDES 

*ROSEA 

*ROSEA 

SARTWELLII 

SCABR I OR 

*STERI L I S 

*ST I PATA 

*STRI CTA 

SYCHNOCEPHALA 

*TENTACULAT A 

*TORTA 

*VAGI N AT A 

woo0 I I 

* X ANTHOC AR P A 

XEROCARPA 

AESTIVALIS 

ALATA 

BILTMOREANA 

BUCKLEY I

NUMBER 12 

USA 

COUNTRY 

STATE 






NORTHWEST TERRITORIES 

OHIO 

OHIO 

OHIO 

OHIO 

OKLAHOMA 

OKLAHOMA 

OK LAH CMA 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

OREGON 

PENNSYLVANIA 

PENNSYLVANIA 

PENNSYLVANI A 

PENNSYLVANIA 

PENNSYLVANIA 

PENNSYLVANIA 

PENNSYLVANIA 

RHODE ISLAND 

T AXON 

MI SERA 


RUTH1 I 

*STIPATA 

STYLOFLEXA 

FESTIVA 

*ALATA 

CONJUNCTA 

FLACC IDULA 

ST EUDEL I FI SSA 

LATEBRACTEATA 

OKLAHOMENSIS 

ACCEDENS 

*ACUTA 

ACUT INA 

ACUTI NELLA 

BRACHY PODA 

BREVICAULIS 

CA MPLY OC AR PA 

CUSICKI I 

*DEWEYANA 

DIVERS1 STYLIS 

EAST WOOD I AN A 

GYMNOCLADA 

HALL1 ANA 

INOPS 

*LEPORINA 

LE PTOPODA 

*MARC I DA 

OREGONENSI S 

PACHYSTOMA 

PANS A 

PH AEOCE PHAL A 

PHAEO LE P I S 

SHELDON1 I 

SPRETA 

*STYLOSA 

*TERET IUSCULA 

VAGANS 

VICARIA 

*ANGUSTIOR 

*AUREA 

COSTATA 

*DEB1 LI S 

GARB ER I 

GRACILLIMA 

*GR I SEA 

HORMATHODE S 

159

160 

US A 

COUNTRY 

STATE 

SOUTH CAROLINA 

SOUTH CAROLI NA 

SOUTH DAKOTA 

TENNESSEE 

TENNESSEE 

TENNESSEE 

TEXAS 

TEXAS 

TEXAS 

TEXAS 

TEXAS 

TEXAS 

TEXAS 

UTAH 

UTAH 

UTAH 

UTAH 

UTAH 

UTAH 

UTAH 

VERMONT 

VIRGINIA 

VIRGINIA 

VIRGI NI A 

VIRGINIA 

V I RGI NI A 

VI RGI NI A 

VIRGINIA 

V I RGI NI A 

VIRGINIA 

WASHINGTON 

WASHI NGTON 

WASHINGTON 

WASHINGTON 

WASHINGTON 

WASHI NGTON 

WASH I NGTO N 

WASHI NGTON 

WASHI NGTON 

WASHINGTON 

WASHI NGTON 

WASHI NGTON 

WASHI NGTON 

WASHI NGTON 

WASHINGTON 

WASHI NGTON 

WASHI NGTON 

WASHINGTON 

WASHI NGTON 


TAXON 

AUSTRO-CAROLINIANA 

CAROLI NI ANA 

HAYOEN I *OXYLEPIS 

PURPURI FERA 

ROANENS I S 

BUL BOSTY L I S 

MUR ICULATA 

ONUSTA 

RETROFLEXA 

TRI ANGULAR1 S 

*WILLDENOVI I 

WRIGHT11 

*CAMPY L OC AR PA 

*CANESCENS 

EPAPI LLOSA 

INTERIMUS 

PELOC ARPA 

RACHI LL I S 

*VERNACULA 

rC: INTUMESC EN S 

*ABSCONDITA 

X ABSCOND IT1 FORM1 S 

BAY ARDI 

*CR IN1 TA 

*CRUS-CORVI 

*DE B I L I S 

*DIGITALIS 

*DIGITAL I S 

RUGATA 

APERTA 

*APERTA 

*APERTA 

*APERTA 

C 0 N ST ANC E AN A 

EGREGIA 

EURYCARPA 

* EU RY C AR P A 

*INTERRUPTA 

YLENTICULARIS 

MISERABILIS 

*NEBRA SKENS I S 

*NEBRASKENSIS 

NEUROPHORA 

OLY MPI CA 

OXYCA RPA 

PADDOENS I S 

PRAECEPTORIUM 

SCABRIUSCULA

NUMBER 12 

US A 

USSR 

COUNTRY 

VENEZUELA 

VIET-NAM, NORTH 

WESTEFN SAMOA 

STATE 

WASH I NGTON 

WASHINGTON 

WASHI NGTON 

WASHI NGTON 

WY OM1 NG 

WYOMING 

WYOMING 

WY OM I NG 

WYOMING 

WYOMING 

WYOMING 

WYOMING 

WYOMING 

RUSSIAN SFSR 

TADZHIKI STAN 

TURK ESTAN 

BOLIVAR 

LARA 

SUCRE 

SUCRE 

TACHI RA 

TACHI RA 

TONKI N 

TA XON 

SUBORBICULATA 

SUKSDORFII 

*SUKSDORFII 

V I R I01 OR 

ALBO-NIGRA 

BR EVI SQUAM A 

FE STIVELLA 

LIMNOPHILA 

NELSON1 I 

PAYSONI S 

PLATYLEPIS 

SI MULATA 

VALLI COLA 

KURILENSIS 

PHI LOCRENA 

*STENOPHYLLA 

RORAIMENSIS 

LARENSI S 

CULMENICOLA 


TACHI RENSI S 

TAMANA 

TR I CHOP HY LL A 

SAVAI IENSI S 

161


HERBARIUM INDEX 

SHEET NO. KIND OF TYPE TAXON 

A (ARNOLD ARBORETUM, CAMBRIDGE, MASSACHUSETTS) 

I SOTYPE 

ISOTY PE 

I SOTYPE 

I SOTYPE 

IS OTY P E 

I SOTYPE 

I SOTYPE 

I SOTY PE 

I SOTY PE 

HDLOTYPE 

ISOTYPE 

HOLOTYPE 

CAS (CALIFORNIA ACADEMY OF 

242617 

497554 

553879 

242962 

242960 

242961 

5540 19 

102030 

43088 1 

383776 

334353 

553918 

2320 50 

445943 

553874 

553913 

383554 

242987 

553883 

264973 

204974 

259875 

259874 

336835 

372834 

351155 

130386 

2429 57 

246772 

40 1490 

234898 

102307 

I SOTYPE 

I SOTYPE 

I SOTY PE 

I SOTY PE 

SYNTYPE 

SYNTYPE 

I SDTYPE 

ISOTYPE 

I SOTYPE 


ISQTY PE 

r SOTYPE 

ISOTYPE 

ISOTYPE 

TYPE FRAGMENT 

SYNTYPE 

SYNTYP E 

HOLOTY PE 


SYNTYPE 

SYNTYPE 

HOLOTY PE 

I SOTY PE 

I SOTYPE 

I SOTYPE 

I SOTYPE 

HOLOTYP E 

SYNTYPE 


ISOTYP E 

ISOTYPE 

I SOTYPE 

ACROPHI LA 

BREVIS 

ERE YOST ACHY A 

L AMPROCHL AMY S 

YEL A NOP HOR A 

MERCARENS I S 

PER ILE I A 

R UBR 0-BRUNN E A VAR ELINEOLATA 

SARAWAKETENSIS 

SURCULOSA 

TAHITENSIS 

ZIZANI AEFOLIA 

SCIENCESi SAN FRANCISCO) 

ABORI GI NUM 

ALMA 

ALOPECOIDEA 

APERTA 

APERTA 

APERTA 

AQUA TI L I S 

ARAPAHOENSIS 

BI PARTITA 

CALIFOSNICA 

C AM PYL OC ARP A 

CHA PMANI 

C IL I ARI S 

C IL I ARTS 

CINNAMOMEA 

COMMUNI S 

C ON J U NC T A 

CONSTANCEANA 

CRINlTA 

CUR AT OR I UM 

CURATORIUM 

DANAENSIS 

DANAENS IS 

DIGITALIS 

DIVERS I STYLI S 

OUR AND1 I 

E A ST WOOD I ANA 

EURYCARPA 

E URYC ARPA 

E UQ Y ST ACHY A 

F I S S URI COL A 

FORYOSA 

VAR SPARSI-SPI CATA 

VAR UMBROSA 

VAR VIRIDANS 

VAR VIRlDANS 

VAR SUBSTRICTA 

VAR AUSTROMONTANA 

SSP AFFINIS 

VAR MINOR 

VAR ASYMMETRICA 

VAR ATTENUATA

NUMBER 12 

SHEET NO. KIND OF TYPE TA XON 

CAS (CALIFORNIA ACADEMY OF SCIENCES, SAN FRANCISCO) 

383156 

103C33 

353986 

328017 

103098 

103098 

553902 

404489 

2039 10 

348506 

553999 

272528 

272529 

242959 

477664 

384438 

10 248 1 

264341 

186427 

136 

231121 

384084 

264346 

553885 

553875 

234896 

194659 

235733 

237824 

237908 

351152 

239452 

369422 

155657 

193005 

383889 

416360 

105004 

404488 

102638 

3252 53 

336836 

3838@1 

383407 

553975 

553877 

445940 

246086 

ISOTYPE 

I SOTY PE 

I SOTYPE 

ISOTYPE 

SYNTYPE 

SYNTYPE 

TYPE COLLECT ION 

I SOTYPE 

ISOTYPE 

I SOTY PE 

1 SOTYPE 

I SOTYP E 

I SOTY PE 


I SOTY PE 

ISOTYPE 

I SOTYP E 

SYNTYPE 

I SOTY PE 

ISOTYPE 

i SOTY PE 

I SOTYPE 

I SOTYP E 

I SOTYPE 

TYPE FRAGMENT 

ISOTYPE 

I SOTY PE 

HOLOTY PE 

I S Of Y PE 

I SOTY PE 

I SOTYPE 

ISOTYPE 

I SOTYPE 

I SOTYPE 

ISOTYPE 

I SOTYPE 

ISOTYPE 

TYPE COLL ECT I ON 

I SOTYPE 

ISOTYPE 

I SOTYPE 

I SOTYPE 




f SOTY PE 

I SOTYPE 

HOLOTY P E 

FORMOSA 

GRAC IL I OR 

GYNODYNAMA 

HAGIANA 

HOOD I I 

HOOD I I 

IGNOTA 

I NCONDITA 

lNOPS 

I NT E R I MUS 

J NTERIOR 

VAR NERVOSA 

VAR NEUROCARPA 

INTERIOR VAR CHARLESTONENSI S 

I NT ER I OR VAR CHARLESTONENSIS 

I NTERRUPTA VAR DISTENTA 

JACOBI-PETER1 

L ATE BRACT EAT A 

LEIOPHYLLA 

LEMANNTANA VAR SIMPLEX 

LE PTOPOOA 

LUZ ULAEF OLI A VAR STROBILANTHA 

LUZULAEFOLI A VAR STROBILANTHA 

LUZULI NA 

MACKENZI ANA 

MEAD1 I 

MENDOC I NE NS I S 

MICROPTERA 

MURI CUL AT A 

OBI SPOENSIS 

OBISPOENSIS 

OR I SPOE NS IS 

ONUSTA 

PELOCARPA 

PER COST AT A 

PER LONG A 

P ERL ONGA 

P I TY OPH I L A 

P LEC TOC AR P A 

PRATENSIS 

PR AT I C OL A 

VAR SUBCORIACEA 

PROPOSI TA 

R AC H I LL I S 

RUGATA 

SALINAEFORMIS 

SARTWELLII 

SARTWELLI I 

SCABR IOR 

SEATONIANA 

SONOMENSI S 

163



CAS (CALIFORNIA ACADEMY OF SCIENCES, SAN FRANCISCO) 

246636 

232289 

342553 

342552 

162423 

259816 

152864 

243333 

553943 

428953 

429306 

351161 

348507 

I SOTY PE 

SYN TY PE 

HOLOTYPE 

I SOTY PE 

I SOTYPE 

HOLOTYPE 

I SOTY PE 

I SOTYPE 


HOLOTYPE 

I SOTYPE 

I SOTY PE 

I SOTYPE 

SONOMENSIS 

SPECIFICA 

SPEC UI C OL A 

SPECUICOLA 

SUB1 MPRESSA 

SURNIGRICANS 

SUKSDORFIX 

SUKSDORF I I 

T EN1 AC UL A T A VAR 4LTIOR 

TOMPKINSI 

TOMPKI NSI 

TOWNSEND I I 

VER NACULA VAR HOBSONII 

DS ( DUDLEY HERBAR IUY, STANFORD UNIVERSITY, STANFORD, CALIFORNIA) 

149709 

55317 

64125 

109019 

489410 

49734 

269649 

14561 9 

629609 

144009 

284598 

149706 

145620 

49500 

490408 

278190 

75003 

490443 

13923 

76794 

§4832 

489409 

171453 

270930 

490462 

49738 

55002 

293480 

490735 

258275 

I SOTYPE 

I SOTYPE 

I SOTY PE 

I SOTYPE 

HOLOTY P E 


1 SOTYPE 

HOLOTY P E 

I SOTYPE 

I SOTYPE 

I SOTY PE 

I SOTYPE 

HDLOTY P E 

ISOTYPE 

I SOTYPE 

ISOTYPE 

HOLOTY PE 

I SOTYPE 

I SOT YP E 

I SOTY PE 

I SOTYPE 

HOLOTY PE 

I SOTYPE 

I SOTYPE 

ISOTYPE 

SYNTYPE 


SYNTYPE 


I SOTYPE 

I SOTY PE 

ABORIGI NUM 

ARRAMSI I 

ABRUPT A 

ARAPAHOENS IS 

AUSTROYONTANA 

CALI FORNICA 

C ONSTANC E ANA 

OUDLEY I 

OUDL EY I 

EA STWOOD I ANA 

EGREGEA 

ELRODI 

GRACILIOR 

GYNO DY NA MA 

GYNOOY NAMA 

INTERIOR VAR CHARLESTONENSI S 

J AC 1: NTOENS IS 

LEPORI NELLA 

LEPTOPODA 

LUZllL I NA 

MEN DOC I NE N S I S 

YULTICOSTATA 

NESRASKENSIS VAR ERUCAEFORM IS 

NUOATA VAR FIRMIOR 

OBISPOENSIS 

PANSA 

PAUCICOSTATA 

QUADR IFIDA 

SAL€ NAEFORMIS 

SCABR I USC UL A 

SONOMENSIS

NUMBER 12 


DS (DUDLEY HERBARIUM9 STANFORD UNIVERSITY, STANFORD9 CALIFORNIA) 

374718 

337970 

83850 

171455 

269641 

269625 

63991 

64032 

I SOTYPE 

I SOTY P E 

I SOTY PE 

I SOTYPE 

f SOTY P E 

SYNTYPE 

I SOTYPE 

I S OTY PE 

SONOMENSI S 

SPECUI COL A 

ST E NOPTE R A 

S UK S DORF I J 

SUKSDORF I I 

SUKSDORFII VAR OVALIS 

ULTRA 

ULTRA 

F (FIELD MUSEUM OF NATURAl HISTORY 9 CHICAGO, ILLINOIS 1 

186491 

206585 

32699 

32700 

56916 

349624 

373673 

373679 

1464064 

264169 

12661 84 

999642 

314869 

456934 

1620435 

49642 

2020 21 

26304 

1 I28952 

1266170 

1406416 

751055 

455703 

1429766 

122779 

314892 

456958 

176870 

141 1493 

2831 19 

1128957 

1463659 

314869 

456934 

1076930 

263394 

ISOTYPE 

TYPE MATERIAL 

I SO TYP E 

ISOTYPE 

I SOTYPE 

I SOTYP E 

I SOTY PE 

I SOTYPE 

TYPE MATERIAL 

f SOTYPE 

TYPE MATERIAL 

TYPE MATERIAL 

I SOTYPE 

I SOTYPE 

I S OTY PE 

I SOTYPE 

I SUTYP E 

TYPE MATERIAL 

HOLOTYPE 

HO L D TY P E 

TYPE YATERI AL 

I SOTYPE 



TYPE MATERIAL 

SY NTY PE 

SYNTYPE 

I SOTYPE 

TYPE MATERIAL 

TYPE MATERIAL 

HOLOTYPE 

TYPE MATERI AL 

I SOTY PE 

f SCITY PE 

I SOTY PE 


ABRAMS I I 

ACUTINA 

ALOPECOIDEA 

AL OPEC0 I D E A 

ALOPECOIDEA 

ALOP ECOI DE A 

ALOPECOIDEA 

ALOPECOI DE A 

APODOSTACHYA 

AUTUMNAL I S 

AZUAYAE 

BARTLETTI I 

BONPLANDI I VAR MINOR 

BONPL4NDIl 

CHIAPENSIS 

CHI HUAHUAENS I S 

C ILIARI S 

CREBRI FLORA 

CUCHUMATANENSI S 

CULMENI COLA 

CUNEATA 

X DEAMII 

VAR MINOR 

DEW EYANA 

VAR SPARSIFLORA 

DEHEYANA 


FESTlVA 

HALL 1 I 

HALL I HARPER1 

HATUSIMANA 

HELLERI 

HUE HUE TEC A 

HY YENODON 

ILLOTA 

VAR DECUMBENS 

ILLOTA 

I NT ER I OR 

I NVOLUCRATELLA 

VAR CHARLESTONENSI S 

165



F (FIELD MUSEUM OF NATURAL HISTORY, CHICAGO, ILLINOIS) 

1406403 

129242 

1566419 

1481645 

55470 

267758 

455706 

1425899 

206587 

866418 

455736 

96129 

211365 

223512 

1471 48 9 

309086 

178542 

455706 

1425899 

755322 

1G5551 

1607711 

215918 

1263854 

1489429 

1463953 

30885 

210109 

464432 

907841 

1252385 

1129096 

813737 

1263 85 8 

1263857 

1129085 

1129086 

1266183 

1128966 

12661 50 

1129094 

89120 

309085 

309086 

TYPE MATERIAL 

SYNTYPE 

SYNTYPE 

ISOTYPE 

HOLOTY PE 

I SQTYPE 

TYPE COLL ECT ION 

TYPE COCLECTiON 


I SOTY PE 

S YN TY PE 

TYPE YATER IAL 

TYPE YATERIAL 

TYPE Y4TERIAL 

TYPE Y4TERIAL 

TYPE YATERIAL 

I SOTYPE 



TYPE MATERIAL 




HOLOTYPE 

I SOTYPE 

TYPE MATERIAL 

TYPE M4TERIAL 


SY NTY PE 

I SOTYPE 

HOL OTY P E 

HOLOTYPE 


HO L 0 TY P E 

HOLOTYPE 

TYPE MATERIAL 

TYPE MATERIAL 

TYPE MATERIAL 

TYPE 

HOLOTY PE 

HOLOTY 9 E 

TYPE YATERIAL 

SY NTY P E 

SYNTYPE 

KURILENSI S 

L ACU NARUM 

LAC UNARUM 

L ANGEANA 

L ARENS I S 

LA XI FLORA 

L EPORINA 

LEPORINA 

MARC IDA 

OBISPOENSIS 

OREGONENS IS 

OXYCARPA 

OXY CARPA 

PADOOENSI S 

P ADDOENS I S 

PAUC I COST AT A 

PER LONG A 

PHAEOCEPHALA 

P HA E OC E PH AL A 

P HY SOC HL A E N A 

PR I NGL E I 

PRI NGLE I 

PSEUDOJAPONIC A 

RORA IMENS IS 

RUGATA 

RUGATA 

VAR LEPTONERVI A 

VAR AMERICANA 

VAR AMERICANA 

VAR OEBILIS 

S AL I NAE FORM IS 

SCI RPOIDE A VAR GIGAS 

SCOPAR IA VAR SUBTURBI NATA 

SONOMENSIS 

S TA NDLE YANA 

STEY ERYARK I STI PATA 

V AR L A EV IV AG IN AT A 

TACHIRENSI S 

TAYANA 

TOJQUIANENSIS 

TOJQUIANENSIS 

TORE ADORA 


TURUMIQUIRENS IS 

V E NO S I V AG I N AT A 

VI TREA 

WHITNEY I 

WHITNEY I

NUMBER 12 167 

SHEET. NO. KIND OF TYPE TAXON 

GH (GRAY HERBARIUM, HARVARD UNIVERSITY, CAMBRIDGE, MASSACHUSETTS) 

I SON EOTY P E 

I SOTY PE 

HOLOTYPE 

I SOTY PE 

SY NTY PE 


ISOTYPE 

SYNTYPE 

SYNTYPE 

TYPE CQLL ECT ION 

I SOTYPE 

HOLOTYPE 

I SOTYPE 

I SOTYPE 

HOLOTYP E 

TYPE 

SYNTYPE 

ISOTYPE 

I SOTY PE 

HDLOTYPE 

ISOTYPE 

HOLQTY PE 

I SOTYPE 

HOLOTY PE 

SYNTYPE 



SYNTYPE 

HOLOTYP E 

ISOTYPE 

HOLOTYPE 

ISOTYPE 

HOLOTY P E 

I SOTYPE 

ISOTY PE 

ISOTYPE 

PARATYPE 

I SOTY PE 


TYPE 

SY NTY PE 

HOLOTY PE 


ISOTYPE 

HOLOT Y PE 

HOLOTYPE 

SY NTY PE 

HOLOTYPE 

ABLATA 

A B SC ON0 IT A 

X ABSC ONDI T I FORM IS 

X A BSCONO I T I FOR M I S 

ACCEDENS 

AC UTA 

ACUTINA 

ADUSTA 

AENEA 

AEQlJA 

AESTIVACIFORMIS 

ALATA 

AL OPEC 0 I DE A 

AL OPEC 0 I DE A 

AMPHIBOLA 

ANGUSTIOR 

APERTA 

AQUAT IL IS 

ARAPAHOENS IS 

ARG YR AN THA 

ARGYRANTHA 

ARI STATA 

ARSENII 

ART ITECTA 

ASSINIROINENSIS 

AZTECICA 

BACKANA 

BACK1 I 

BARBARA€ 

BARTLETT I I 

BAYARDI 

B ILTMOREANA 

BONPLANDI I 

SREWERI 

B UR C HEL L I AN A 

CAESARIENS IS 

CANESCENS 

CANE SCEN S 

CAROL IN I AN A 

CEPHALOPMORA 

CHALC IOLEP IS 

C IL I ARIS 

CI NNAMOYEA 

C IRC INNATA 

CLI VICOLA 

COLLECTA 

CONJUNCTA 

C OOCEY I 

VAR ROSTECLATA 

VAR PALLIDA 

VAR GLOMERATA 

VAR FERRUGINEA 

VAR SPARS I-SPI CATA 

VAR TURGIDA 

VAR GRAC ILENTA 

VAR SUB STR IC TA 

V A R LONGO-L ANCEOL ATA 

VAR SUBTILIROSTRIS 

VAR MINOR 

VAR DISJUNCTA 

VAR SPHAEROSTACHYA 

VAR MAXIMA


SHEET Nn. KIND OF TYPE TA XON 

GH (GRAY HERBARIUM, HARVARD UNIVERSITY, CAMBRIDGE, MASSACHUSETTS) 

SYNTYPE 

SYNTYPE 

TYPE 

HOLQTYP E 

SYNTYPE 

I SOTYPE 

ISOTYDE 

YOLOTY PE 

HOLOTY PE 

I SOT Y PE 

I SOTYPE 

HOLOTYPE 

HOLOTY P E 

I SOTYPE 

HOLD T Y P E 


HOLOTYPE 

HOL OTY P E 


I SOTYPE 

ISOTYPE 

I SOTYPE 

HOLOT Y P € 

I SOTY P E 

I SOTY PE 

TYPE Y4TERIAL 

I SOTY PE 

I SOTYPE 

HOLOTYPE 

TYPE YATE RI AL 

I SOTYPE 

HOLOTY PE 


ISDTYPE 

HDLOTY PE 

I SOTYPE 

HOLCITYPE 

HOLOTY PE 

HOLOTYPE 

ISOTYPE 

I SOTY PE 

I SOTYPE 

ISOTY P E 

I SOTYPE 

HOLOTYPE 

HOLOTY PE 

I SOTY PE 

I SOTYPE 

CRA WFORD I I 

CRAWFORDX I 

C REBRI FLORA 

CRINITA 

CRINITA 

C Rf STATA 

CRUS-CORVI 

C RUS -CO RV I 

C UYULA TA 

DANAENS IS 

DAVY I 

DERILIS 

DEB It IS 

DEVIA 

DEUEYANA 

DEWEY ANA 

DIGITALIS 

DIGITAL IS 

DOUGLAS11 

X DUMANII 

DUT ILLY I 

EASTWOODI ANA 

ECHINATA 

EKMANI I 

EKYANI I 

EL9 ERTANA 

ELEOCHARIS 

ELYNOI DES 

ENGE LMANNI 

ERXLEBENI ANA 

EURY STACHYA 

E XP L OR ATORUM 

FEL I PENS IS 

FESTIVELLA 

FETA 

FILIFOLIA 

F I SSA 

FLACCOSPERMA 

FLAVA 

FLAVA 

FOENEA 

FORMOSA 

FRANKLIN11 

GARBERI 

GARBERI 

GLAREOSA 

GLAUCODEA 

GR4CILIOR 

VAR VIGENS 

VAR BREVICRINIS 

VAR SIMULANS 

VAR VIRGINIANA 

FOR SOLUTA 

VAR INTERCURSA 

VAR PUBERA 

VAR COLLECTANEA 



VAR MACROPODA 

VAR DENSISPICATA 

VAR ORMANTHA 

VAR HOTTENSIS 

VAR EROSTRATA 

VAR ARISTATA 

VAR GASPENSIS 

VAR RECTI ROSTRA 

VAR PERPLEXA 

VAR BIFARIA 

VAR AYPHIGENA

NUMBER 12 169 

SHEET. NO. KIND OF TYPE TA XON 

GH (GRAY HERBARIUYv HARVARD UNIVERSITY, CAMBRIDGE, MASSACHUSETTS) 

I SOTY PE 

HOLOTY PE 

HOLOTYPE 


SYNTYPE 

HOLOTYPE 

YO LOTY P E 

SYNTYPE 

HOLOTY P E 

SYNTYPE 


HOLOTY P E 


SYNTYPE 

HOLOTY PE 

HOLOTYPE 

HOLOT Y PE 

ISOTYPE 

I SOTY PE 

HOLO TY P E 

YOLOTY P E 

I SOTY PE 

I SOTYPE 

HOLOTY PE 

HOLOTY PE 

HOLOTYPE 

I S OTY PE 

I SOTYDE 

HOLOTYPE 

SYNTYPE 

I SOTYPE 

HOL Ot Y P E 

HOLOTYPE 

HOLOTVPE 

ISOTYPE 

HOtOTYPE 

HOLOTYPE 

ISOTYPE 

I SOTYPE 



I SOTYPE 

HOLOTYP E 

ISOTYPE 


HOLOTYPE 

HOLOTY PE 

TYPE CDLLECTION 

GRACILLIMA 

GRI SEA 

GUATEMALENSI S 

HALL IANA 

HALL I HALSEYANA 

HARPER I 

HAY DEN I AN A 

HERE TATA 

HEPBURN I H E f E RON EU R A 

HI TCHCOCKI ANA 

HOOK ERANA 

H OR MATH ODES 

HORNSCHUCHI ANA 

HOST IANA 

TLLOTA 

I NC I SO-DE NTATA 

I NCURV I FORM IS 

I NFLATA 

INOPS 

INTERIMUS 

INTERIOR 

INTERIOR 

INTERIOR 

I NTUME SCENS 


KALOIDES 

KATAHDINENSIS 

KELLOGGI I 

KOKRINENSIS 

LAEVI-CONICA 

LANGEANA 

LASI OCARPA 

I- ATEBRACTEATA 

LAX IFLORA 

LAX1 FLORA 

L E IOCARPA 

LEI OPHY LLA 

LEMHONI 

L EPOSINA 

LEPORINELLA 

LIVIDA 

LONG ICULM I S 

LUZULINA 

MACKENZIANA 

MEAOX I 

MEEK1 I 

VAR RIGIDA 

VAR LAURENTI ANA 

VAR LAURENTIANA 

VAR ANT ICOSTENS I S 

VAR CHARLESTONENS IS 

VAR JOSSELYNI I 

VAR KEWEENAWENSIS 

FOR VENTRIOSA 

VAR AMERICANA 

VAR LEPTONERVIA 

VAR SERRULATA 

VAR AMER I CANA 

VAR RUFINAEFORMIS



GH (GRAY HERBARIUM, HARVARD UNIVERSITYI CAMBRIDGE, MASSACHUSETTS) 

I SOTYPE 

H0LC)TYPE 

HOLOTYPE 

I SOTY P E 

I SOTYPF 

HOLOTYPE 

HOLOTY P E 

SY NTY PE 

SYNTYPE 

HOLOTY PE 

HOLOTY P E 

I SOTYP E 

ISOTYPE 

HOLO TY PE 

I SOTYP E 

HOL OTYP E 

I SOTYPE 

I SOTYP E 

ISOTYPE 

I SOTYPE 

ISOTYPE 

H OLOTY PE 

I SOTYPE 

ISOTYPE 

I SOTYPE 

HOLOTY P E 

HOLOTY PE 

HOLOTYP E 

ISOTYPE 

!SOTYPE 

I SOTYPE 

HOLOTY PE 

HOLOTYPE 

I SDTYPE 

HOLOTY PE 

YOLOTY PE 

I SOTYP E 

HOLOTY PE 

I SOTYPE 

I SOTYPE 


HOLOTYPE 

ISOTYPE 

I SOTYPE 

TYPE MATERIAL 

TYPE VAT ER I AL 

HOLOT Y PE 

I SOTYP E 

MELOZITNENSIS 

YENDOCINENSIS 

MERR I TT-FERNALDI I 

ME S OCHOR E A 

M I C ROC H AE 1 A 

MIL1 ARI S 

MIRABILIS 

MIRABIL IS 

MIRABILIS 

M IS ANDRO IDES 

MONTEREYENSIS 

NEL SON1 I 

X NEOBIGELOW I I 


X NEOPALEACEA 

NERV INA 

NEUROPHORA 

NIGRICANS 

NOVAE-ANGL I AE 

X NUBENS 

NU5 ICOLA 

NUTTALL I I 

OAXACANA 

OBISPOENSIS 

OBOVOI DEA 

OL I G UC A R P A 

ORMOST ACHY A 

ORONENSI S 

P AC H Y ST AC HY A 

PADDOENSIS 

PALAWANENSI S 

PALL ESCENS 

PAPULOSA 

X PATUENSIS 

P AUP ERCULA 

P AUP ERCUL A 

PAYSONI S 

P ER L ONGA 

PERSTRI CTA 

PETRIE I 

PHA EOC E PH AL A 

PHY I, LUM AN I C A 

P I NE TOR UM 

P IPERI PLATYPHYLLA 

P 0 DOCAR PA 

PORTER1 

POTOSINA 

VAR AUREA 

VAR PERLONGA 

VAR TINCTA 

VAR LATIFOLIA 

VAR NEOGAEA 

VAR BREVISQUAMA 

VAR PALLENS 

VAR ELATIOR

NUMBER 12 17 1 


GH (GRAY HERBARIUM9 HARVARD UNIVERSITY, CAMBRIDGE9 MASSACHUSETTS) 

HOLOTY PE 

H OLOT Y QE 

ISOTYPE 

SY NTY PE 


ISOTYPE 

T SOTY PE 

HOLO TYP E 

ISOTY P E 

SYNTYPE 

HOLOTYPE 

ISOTYPE 


HOLOTY P E 

HOLOTYP E 

I SOTYPE 

HCILOTYPE 

HOLOT Y P E 

I SOTYPE 

ISOTYPE 

HOLOTY P E 

HOLOTYPE 

HOLOTY P E 

SY NTY PE 

HOLOTVPE 

HOLOTY PE 

HOLOTYPE 

HOLOTYP E 

I SOTYPE 

I SOTYPE 

HOL OTY P E 

ISOTYPE 



SYNTYPE 

SYNTY PE 

SYNTY PE 

I SOTY PE 

I SOTYPE 

ISOTYPE 

I SOTYPE 

YOLOTYPE 

HOLOTY PE 

I SOTYPE 

HOLD TY P E 

HOLOTY PE 

HOLOTYPE 

HOLD TY P E 

PRAEGRACIL IS 

PRA I REA 

PRATENSI S 

PRE ISS I I 

BKI NGLEI 

PROJECTA 

PROPOSITA 


X QUEBECENSIS 

RAYNOLDSI I 

R E T R OC UR VA 

R ETROCURVA 

R IC HARDSON I RICHARDSON I 

ROS EA 

RURRO-BRUNNEA 

RUGATA 

S ARTW ELL I ANA 

X SAXENII 

SCHAFFNERI 

S C H N E I 0 E R I 

SCHWEINITZII 

SCOP AR 1 A 

SCOPARl A 

SCOPARI A 

SCOPAR I A 

SCOP AR I A 

SC OPARI A 

S EATONIANA 

SONOMENSIS 

SOPERI 

SPECUICOLA 

SPRETA 

STI PATA 

STRAMINEA 

STRAMINEA 

STYLOSA 

S UR -BR ACT EAT A 

SUB FUSC A 

SUK SDORF I 

SYCHNOC EPHALA 

TENERA 

TENERA 

T ENU I FL OR A 

TERRAE-NOVAE 

T HU RBE R I 

X TRICHINA 

TRICHOPHYLLA 

VAR FURVA 

VAR COPULATA 

FOR EXSERTA 

VAR ARKANSANA 

VAR ELINEOLATA 

NM. FERRUGINEA 

VAR CONDENSA 

VAR FULVA 

VAR MONILIFORMIS 

FOR PERACUTA 

V A R SU BTUR BIN AT A 

VAR TESSELLA A 

VAR LAEVIVAG NATA 

VAR CUMULATA 

VAR ECHINODES 

VAR VIRENS 

VAR RICH11 

VAR SETACEA

172 SMITHSOKIAN CONTRIBUTIONS TO BOTANY 


GH (GRAY HERBARIUM, HARVARO UNIVERSITY, CAMBRIDGE, MASSACHUSETTS) 

I SOTY PE TRI S PERM A VAR B ILL INGS I I 

1 SOTY PE TSOI 

ISOTY PE TURUYIQUIRENSIS 

HOLOTYPE UMBELLATA VAR VICINA 

I SOTYPE UNCOMPAHGRE 

HOLOTY PE VAG INATA VAR ALTO-CAUL1 S 

HOLOTYPE VALL ICOLA 

ISOTYPE VENOSIVAGINATA 

I SOTY PE V ERN ACUL A VAR HOBSONII 

HOLO TYP E 

HOLOTYP E 

VESICARIA 

VES I CAR I A 

VAR JEJUNA 


HOLOTY PE VESTITA VAR KENNEDY1 

H0LI)TY PE VICARIA 

ISOTYPE VIRIDULA FOR PYGMAEA 

HOLOTY PE VULP INOIDE A VAR PYCNOCEPHALA 

HOLOTYPE 

ISOTYPE 


HOL 0 TY P E 

WA T SON1 

W I EGBND I I 

W I LLDENOVI I 

WOOD I1 V AR PAUC I FLORA 

HOLOTY PE X XANTHINA 

SYNTYPE 

X E RANT I C A 

JEPS (JEPSON HERBAP IUMt UNIVERSITY OF CALIFORNIA, BERKELEY) 

2511 ISOTYPE 

4013 ISOTYPE 

20008 I SOTY PE 


MO (MISSOURI BOTANICAL GARDEN9 ST. LOUIS) 


130648G ISOTY PE 

SYNTYPE 

TYPE MATERIAL 

TYPE VAT ER I AL TYPE COLLECT1 ON 


SYNTYPE 

18341 52 HOLOTYPE 

ISOTYPE 

TYPE COLL ECT I ON 

1611724 SYNTYPE 

ISOTYPE 



DAVY I 

INTERIOR VAR CHARLE STONENSI S 

JEPSONI I 

MAR1 POSANA 

X 

ABSCOND I TA 

ABSCOND IT I FORM IS 

ACCEDENS 


AESTIVALIS 

AGGLOMERATA 

VAR ROSTELLATA 

AQUATILI S 

AT HR OST ACHY A 

AUSTRO-C AROL I N I AN A 

A UTVMN4L I S 

AZT ECI CA 

BANKS1 I 

VAR SUBSTRICTA 

BONPLANDI I 

BUL BOSTY L IS 

CALI FORNICA 

VAR MINOR

NUMBER 12 


YO (YISSOURI BOTANICAL GARDEN, ST. LOUIS 1 

1108572 

1306423 

1129747 

181 6497 

1816496 

114838 1 

1190731 

1201697 

1692 174 

952735 

1220830 

9208 15 

2002968 

1260436 

1306478 

SYNTYPE 

ISOTYPE 

I SOTYPE 

TYPE MATERIAL 

I SOTYPE 

r SOTYPE 

I SOTY P E 



I SOTY PE 


I SOTYPE 

I SOTY P E 

TYPE MATERIAL 

SYNTYPE 

TYPE YATERIAL 

I SOTYPE 


I SOTY P E 

ISOTYPE 

ISOTYPE 

TYPE MATERIAL 

SYNTYPE 

I SOTY P E 

I SOTY PE 


TYPE YATERIAL 

I SOTY PE 

I SOTYPE 




TYPE WATERIAL 

TYPE MATERI AC 

TYPE MATERIAL 

TYPE MATERIAL 

I SOTYPE 


ISOTYPE 


TYPE MATERIAL 

SYNTYPE 


I SOTYPE 

I SOTY PE 

TYPE MATERIAL 

TYPE YATERIAL 

I SOTY PE 

CHALCIOLEPIS 

C IL I AR I S 

C R U S-C ORV I 

DE6 IL IFORM I S 

0 I G ITAL I S 

0 I G I TAL I S 

DUDLEY I 

ELYERI 

ECYNOIDE S 

EPAP ILLOSA 

FENDLERIANA 

F ISSA GRAC IL IOR 

GY MNOCL A DA 

HALL I 

HAY DEN I I 

ILLOTA 

INTERIOR 

INTER I OR 

INTERIOR 

INTERIOR 

I NV OLUC RAT ELL A 

LACUN4RUM 

LAT EBRACTEATA 

LAYIFLORA 

LEPORINA 

LUZUL INA 

MEAD I I 

MENDOCI NENSI S 

N EBRA SK EN S I S 

NUDATA 

0 A XACA N A 

0 BOVO I DE A 

OKLAHOMENSIS 

P A UC I CO ST AT A 

PER GLOBOSA 

PERLONGA 

PHA E OC E PH 4LA P HY L LOMAN IC A 

PHY S OCHL A E N A 

PRAI NI I 

PRE ISSI I 

PRI NGLEI 

RUB RO-B RUNNE A 

RUGATA 

SALI NAEFORMIS 

SAX1 MONTANA 

SCABRIUSCULA 



VAR MACROPODA 

VAR CHARLESTONENSI S 

VAR CHARLESTONENSIS 

VAR CHARLESTONENSIS 

VAR CEPTONERVI A 

VAR AMERICANA 

VAR ERUCAEFORMI S 


173



MO (MISSOURI BOTANICAL GARDEN, ST. LOUIS) 




SY MT Y P E 


SYNTYPE 



I SOTYPE 

SYNTYPE 

SYNTY P E 

SY NT Y PE 

SCAPOSA 

SPR ETA 

STI PATA 

STR AM I NE A 

STYLOFL EXA 

STYLOSA 

SUK SDORF I I 

VIOL ACEA 

VI TREA 

WH I TNEY I 

WHITNEY I 

WHITNEYI 

NY (NEW YORK BOTANICAL GARDEN, NEW YORK CITY) 

TYPE 

IS ONE OTY P E 

I SOTY PE 

HOL OT Y P E 

HOLOTY PE 


ISOTYPE 




TYPE 

SYNTYPE 


ISOTYPE 

I SOTYPE 

I SOTYP E 



I SOTYPE 


I SOTYPE 

TYPE 


I SOTYPE 

SYNTYPE 

SYNTYPE 

I SOTYPE 

I S OTY PE 

I SOTY PE 


ISOTYPE 

SYNTYPE 

VAR LAEVIVAGINATA 

VAR CUMULATA 

VAR VIRENS 

ABDI TA 

ABLATA 

ABORI GI NUM 

ARRAMSI I 

ABRUPTA 

A 6SC OND IT A VAR ROSTELLATA 

X ARSCONDITIFORMIS 

AClJT INA 

AEQUA 

AESTIVALIS 

A GGREG AT A 

ALATA 

ALO I DA 

ALYA 

ALOPECOIDEA 

ALOPECOIDEA 

AMP H 100 L A 

ANGUSTI OR 

APODA 

AQUATIL IS 

ARAPAHOENS IS 

ARCTAEFORYI S 

ARC1 ICA 

ARSENII 

ATHROSTACHYA 


A TQ AC TODE S 

A TP. 0 S Q U A M A 

AUREA 

AUROLENSI S 

AZUAYAE 

BAC K I I 

VAR SPARSI-SPI CATA 

VAR ANDROGYNA

NUMBER 12 



COTY P E 

I SOTY PE 


TYPE 

TYPE 

TYPE 


HOC OTY P E 

SYNTY PE 



TYPE 


HOLOTY PE 

ISOTYPE 


HOLO TYP E 


SYNTYPE 

I SOTYPE 

HOLOT YP E 

TYPE 

I SOTYPE 

HOLOTYPE 

TYPE 

ISOTYPE 

TYPE 



TYPE 

I SOTY PE 

TYPE 

I SOTY PE 


SYNTYPE 

ISOTYPE 



ISOTYPE 

I SOTYPE 

SYNTYPE 

TYPE 


I SOTY P E 

TYPE 

TYPE MATERIAL 

ISDTYPE 

HOLOTY PE 

BALTZELLI I 

BARBARAE 

BARRATTI I 

BARTLETTI1 

B ILTMOREANA 

BOYANZENSI S 

RRE V I CAUL I S 

RREVISQUAMA 

BRONGNI ART1 I 

BUCKL EY I 

BUSHI I 

CALSARIENSI S 

CAL I FORN ICA 

VAR DENSA 

CAMPY LOCAR P A SSP AFFINIS 

CANE SCENS VAR DUBIA 

CANESCENS 

CAREYANA 

CAROL? NI ANA 

CHALC IOLEP IS 

CHI APENSIS 


CHIKUNGANA 

CILI ARIS 

COLUMBI ANA 

CONC INNOIDES 

CONSTANCE ANA 

CONVOLUTA 

COSTATA 

CRANDALLI I 

CR EBR I FLORA 

WAR SPHAEROSTACHYA 

CRUS-CORVI 

CRYPTOLEPIS 

CURENS IS 


CU6 ENS I S 

CUSICKII 

VAR FLACCIDA 

DAVY I 

DEWEYANA 

V AR SPARS I FLORA 

DIGITAL I S 

D IG IT AL I S 

DUDLEY I 

DUR I FOL I A 

EBENEA 

EGGERTI I 

EGGLESTONII 

EGREGTA 

EKMANI I 

ELRODl 

€PAP ILLOSA 

VAR GLAUCA 

VAR MACROPODA 

175



NY (NEW YORK BOTANICAL GARDEfl? NEW YORK CITY! 

I SOTYPE 

I SOTY PE 

TYPE MATERIAL 


TYPE 

SYNTYPE 


TYPE YATERIAL 

TYPE 

HOLOTYPE 

I SOTY PE 

TYPE 

ISOTYPE 

TYPE 

TYPE 


TYPE 

TYPE 


I SOTY PE 


1 S OTY PE 

I SOTYPE 

SYNTYPE 

SYNTYPE 

SYNTYPE 

I SOTYPE 

HO LOT Y P E 

I SOTY PE 


TYPE 

TYPE 

I SOTYPE 


TYPE 

SYNTYPE 

SYMTYPE 

TYPE COLLEC TI ON 

ISOTYPE 

HOLOTY PE 



TYPE 

HOLOTYPE 

I S OTY PE 

HOL 0 TYPE 

TYPE 

ISOTYPE 

EPAPI LLOSA 

EXPLORATORUM 

FARGESII 

FELI PENS1 S 

FENDLER IANA 

f EST IVA 

FESTI VA VAR DECUMBENS 

FESTIVA VAR STRICTA 

FESTIVELLA 

F TSSA 

F ISSA VAR ARISTATA 

FLACCI DULA 

F LACC 0s PE RMA 

f RANKLINI I 

FULVE SC ENS 

FUSCOLUTEA 

FUSC Of I NCT A 

GAYANA 

VAR HYALINA 

GEYERI 

GLAREOS A 

VAR AYPHIGENA 

GRIFFITHII 

GY NOC L AD A 

GY N ODY N AM A 

HALE I 

HALLII 

HALL I HALSEYANA 

H AR FORD I I 

HARPER1 

HASSEI 

HELLERI 

HETEROSTACHYA 

HINDS11 VAR BREVIGLUMA 

HI R SUT A VAR CUSPIDATA 

HOL M I AN A 

HOOD I I VAR NERVOSA 

HOOD I I VAR NEUROCARPA 

H OOKERANA 

HORNSCHUCHI ANA VAR LAURENTIANA 

HOUGHTONI ANA 

IGNOTA 

ILL INOENS IS 

INCOMPERT A 

INCURVIFORMI S 

INOPS 

I NT EGRA 

I NTERI M US 

INTER IOR

NUMBER 12 

SHEEP NO. KIND OF TYPE TAXON 


I SOTYPE 

I SOTYP E 

I SOTY PE 


YO LO TY P E 

HOCOTY PE 

SYNTYPE 

I s OTY PE 

TYPE 

TYPE 

SYNTY P E 

I SOTYPE 

TYPE 

I SOTY Pf 

I SOTYPE 


I SDTY PE 


TYPE 


TYPE 


COTYPE 

TYPE 

ISOTYPE 

I SOTYPE 

TYPE 

HOLOTYPE 

I SOTYPE 

TYPE 

ISOTY PE 


TYPE 

ISOTYPE 

I SOTYPE 

HOLOTYP E 

HOLOTYPE 

SYNTY P E 

TYPE MATERIAL 

I S ClTY P E 

I SOTYPE 

I SOTYPE 

HOLOTYPE 

I SOTYPE 


I SOTY PE 


1 SOTY PE 

I NTERI OR VAR CHARL ESTON ENS IS 

INTERIOR VAR JOSSELYNI I 

I NT E R IOR VAR KEWEENAWENSI S 

I NVOLUCRATELLA 

JAYESII 

JEPSONI I 

JON€ SI I 

K ATAHDINENSIS 

KUL I NGANA 

LAC I NI A TA 

L ACUNARUM 

LANC I FRUCTUS 

LARI CI NA 

LAX I FLORA VAR LEPTONERVI A 

LAX1 FLORA VAR SERRULATA 

LEAVENWORTHI I 

LENT ICUL ARI S VAR PAULL IFRUCTUS 

L E PORT N A VAR AMERICANA 

LE P TOPODA 

L ONGI ROSTRI S VAR MICROCYSTIS 

LUNELLI ANA 

LUZULIYA 

MAC ROKOL E A 

MAC ROSPERMA 

MADRENSIS 

MANDONI ANA 

MARC I DA 

VAR DEBILIS 

MAR I POSANA 

MEADII. 

WED1 TERRANI A 

MENDOC INENSIS 

MERRILLII 

M 1 C ROPTE R A 

MISANOROIDES 

MISERA 

MOHR I ANA 

MOLESTA 

MONTANENSIS 

NANA 

NEBRASKENSIS 

NEBRASKENS IS VAR ERUCAEFORM IS 

NEBRASKENSI S VAR ULTRIFORMIS 

NELSON1 I 


NEOYEXICANA 

NEU ROPHOR A 


NUTTALL I I 

177


SHEET NO, KIND OF TYPE TAX ON 


I SOTYPE 

TYPE 

TYPE 

SYNTYPE 

TYPE 

SYNTYPE 

I SOTYPE 


I SOTYPE 

TYPE MATERIAL 

SYNTYPE 

I S OTY P E 



HOLOTY PE 

HOLOTYPE 

I S OT Y PE 

TYPE 

TYPE 

TYPE 


I SOTYP E 

TYPE MATERIAL 

I SOTYPE 



I SOTYPE 

TYPE MATERIAL 

HOLOTY PE 

HOLOTY PE 

TYPE C3LL ECT ION 



HOLOTY PE 




SYNTYPE 

SYNTYPE 

I SOTY PE 

TYPE 

TYPE 

COTYPE 

I SOTYPE 

ISOTYPE 

T SOTYPE 

TYPE 

TYPE 

OBI SPOENS IS 

OEDERI VAR ROUSSEAUIANA 

OKLAHOMENSIS 

OLY MPICA 

ONUSTA 

OREGONENS IS 

ORONENS IS 

OXYL EP I S VAR PUBESCENS 

PADDOENSIS 

PALA WANENS IS 

PANSA 

PAPULOSA 

PARRYANA 

P AUC ICOSTATA 

PEL OC AR P A 

PERGLOBOSA 

PERLONGA 

PER STR 1 CT A 

PE TA SATA 

PETR ICOSA 

PHA EOC E PH A L A 

PHAEOCEPI S 

PHILOCRENA 

PHYLLOMAN! CA 

PHY SOCHLAENA 

PICTA 

PI NETORUM VAR ELATIOR 

P ITYOPHILA 

PLA TYLE P I S 

PRAECE PTOR I UM 

PR I NGLE I 

PRI ONPHY LLA 

PROJEC TA 

PROPOS ITA 

PSEUDOJ APON I C A 

PTYCHOC ARP A 

P UR P UR I FERA 

QUADRIFIDA 

QUADRI F I DA 

X QUEBECENSIS 

RACHI LL I S 

RETROFLEXA 


RORAIMENSIS 

R OS AEO I OES 

ROSEA 

ROS EA 

ROSEA 

VAR LENIS 

VAR ARKANSANA 

VAR PUSILLA 

VAR STAMINATA

NUMBER 12 

SHEET NO. KIYO OF TYPE TA XON 

NY (NEW YORK BOTANICAL G4RDENv NEY YORK CITY) 

TYPE 

I SDTY PE 

TYPE 

TYPE 

HOLOTY 0 E 


TYPF COLLECTION 


I SOTYPE 

TYPE MATERIAL 

TYPE 

TYPE 



TYPE COLLECTJON 

I SOTY PE 

I: SOTYPE 

HOLOTYPE 

I SOTY PE 

TYPE 


HOLOTY P E 

1 SOTYPE 

ISOTY PE 

TYPE 

I SOTYPE 

fY PE 

TYPE 

I SOTYPE 




ISOTYPE 

SYNTYPE 

SYNTYPE 



HDLOTYPE 

TYPE 

TYPE Y4TERIAL 

ISOTYPE 

SY NTY PE 

HDLOTYPE 

HOLOTY PE 

SYNTYPE 

I SOTYPF 

ISOTYPE 


RUB RO-B RUNNEA 

RUGATA 

RUGOSPERMA 

RUSBYI 

RUTHI I 


SANGUI NEA 

SARTWELLII 

SAVAIIENSIS 

SAXIYONTANA 

SCARRI USCUL A 

SCIRPIFORMIS 

SCIRPOIDEA 

SC I RPO I DE A 

SCfIPARI A 

SCOPAR I A 

SCOPARI A 

SC OULERI 

SHELDONI I 

SHELDONI I 

S ICCATA SIMULAT9 

SONOMENSI S 

SPEC U ICOL A 

STANTONENS IS 

STE L LATA STELLULATA 

STENOPTER A 

STERILI S 

STEUDEL I I 

STI PATA 

STI PATA 

STIPATA 

STQAMINEA 

STRAMINEA 

STRICTA 

STY L OFL EX A 

SUS-RRACTEATA 

SURORBI CULATA 

SUSTPANSVERSA 

SUK SDORF 1 I 

SUKSDORF I 

T4MAKI I 

T ENERAE FORM IS 

TERETIUSCULA 

TOMPKI NSI 

TOR EADORA 

TORTA 


VAR GIGAS 

VAR STENOCHLAENA 

VAR MINOR 

VAR SUBTURBINATA 

VAR TESSELLATA 

VAR CONFERTA 

VAR EXCELSIOR 

VAR LAEV IVAGINATA 

VAR SUBSECUTA 

VAR UBERIOR 

VAS CUMULATA 

VAR RENIFORMIS 

VAR XEROCARPA 

VAR OVALIS 

VAR AMPLA 

VAR STAMINATA 

179




ISOTYPE 

TYPE 

I SOTYPE 

ISOTYPE 

TYPE 

HOLOTYP E 

TY PF 

I SOTY PE 

TYPE 

TYPE 

I SOTY PE 

HOLOTYPE 


SYNTY PE 

SYNTYPE 

SYNfYPE 


TYPE 


TYPE 

SYNTYPE 


TYPE 

uc (UNIVERSITY OF CALIFORNIA, BERKELEY) 

9@5439 

905436 

50814 

9 100 20 

127723 

55234 

1098 

319673 

10 60 

90 5434 

835699 

9c5433 

90 544 1 

HOLDTYPF 

HOLOT Y PE 

HOLOTYPE 

HOLOTYPE 

HOLOTY PE 

HOLRTYPE 

ISOTYPE 

HOLOTY PE 

HOLO TYPE 

HOLOTYPE 

I SOTY PE 

HOLOTYPE 

HOCOTY PE 

TOWNSENDI I 

TRACY1 

TRI ANGULAR1 S 

TR I SPERYA 

TSOI 

TUMULICOLA 

TURGESCENS 

TURUMIQUIRENS IS 

UNDERWOOD1 I 

UN I L AT ER At I S 

VAGANS 

VEQ N AC UL A VAR HOBSONII 

WERDERMANNI I 

WHI TNEY I 

WHITNEYT 

WHITNEY I 

W I LKESI I 

WILLIAYSII 

WRIGHT1 I 

X A N THOC A R P A 

VAR ANNECTANS 

XERANT ICA 

XEROCARPA 

YUYONENSI S 

A PODA 

AR AP AH0 EN S I S 

DAVY I 

INTERIOR VAR CHARLESTONENS IS 

LANC IFRUCTUS 

LEPORINELLA 

MENDQC I NENS I S 

OBL ANCE OL AT A 

PAC H YC A RP A 

PAY SON1 S 

SONOMENS I S 

SUBIMPRESSA 

TRIBULO IDES VAR SANGAMONENSI S 

us (U. S. NATIONAL HERBARIUM, SYITHSONIAN INSTITUTIONV WASHINGTON) 

2003161 TYPE COLLECTION ABSCQNDITA VAR ROSTELLATA 

2003299 I SOTYPE X ABSCONDITIFORMIS 

25 I64 TYP’E COLLECT ION ACUTXNA 

817087 TYPE COLLECTION ACUf INA

NUMBER 12 

SHEET NO0 KIND OF TYPE. TAXON 

us (U. S o NATIONAL HERBARIUM, SYITHSONIAN INSTITUTIONt WASHINGTON) 

27286 

4401 79 

617798 

694342 

858947 

2231424 

865058 

1G30011 

2265958 

319165 

2133192 

2096188 

622651 

817295 

1933437 

2373748 

1232938 

415269 

964504 

711129 

587668 

29741 

319268 

6909 37 

1885701 

368814 

28433 

2460272 

306281 

462090 

28457 

319228 

350c77 

1839933 

20 388 2 2 

2723 5 

1032323 

278555 

278555 

1682487 

13026G2 

17657C.O 

2003164 

2003133 

969128 

176 11 5 1 

817314 

2176489 

HOLOTYPE 


TYPE 

TYPE 


TYPE 

TYPE 


HOLOTY PE 

SYNTYPE 

TYPE 

I SOTY PE 

HOLOTY P E 

YOLOTYPE 

I SOTYPE 

ISOTYPF 

SVNTYPE 



TYPE 

TYPS COLLECTION 




ISOTYPE 

SUN TY P E 

ISCTYPE 

HOLr3TY PE 

I SOTYPE 

(SOTYPE 




I SOTY PE 


TYPE MAT ERI AL 

HOLOTYPE 

I SOTYPE 

SYNTYPE 

I SOTYPE 

TYPE CDLLECT ION 

I SOTY PE 

I SOTYPE 

ISOTYPE 


I SOTYPE 


1 SOTYPE 

ACUTI NELLA 

A GGL OME RA T A 

AGROSTOIDES 

AGRO STOI DE S 

AL 60-M I GR A 

AYPLISQUAMA 

ANTH OXANTHERA 

ARSENII 

ATHABASCENS IS 

A THR OSTACHY A 

ATRACTODES 

ATROFUSCA 

ATROSQUAMA 

AUTUMNAL IS 

A ZUAYA E 

BAY8 USETORUY 

BANKSI I 

6 R A C HY POD A 

BRA I NERD I I 

BRUNNEA 

RULBOSTYL I S 

CAL I FORN ICA 

CAJ_ I FORNICA 

C A Y P L YOC A R P A 

CAM PYL OCARP A 

CHALCI OLEPIS 

C HA P MAN I 

CHI APENS IS 

CHI HUAHUAENS I S 

CILIARIS 

CINNAMOYEA 

CI NNAMOYE A 

CC A DO S T AC HY A 

CLIVICOLA 

COMANS 

CONFERT IFLORA 

CONSPECTA 

CRINITA 

CRI NITA 

C R U S -C 0 RV I 

CUBENSIS 

DANAENS IS 

DEBILIS 

DIGITAL IS 

DIGITAL I S 

DIGITAL1 S 

DONNELL-SYITHI I 

X DUMANII 

VAR DECOLORATA 

VAR SUBTEIOGYNA 

SSP AFFINIS 

VAR MAXIMA 

VAR STRICTA 

VAR BREVICRINIS 

VAR SIMULANS 


VAR INTERCURSA 

VAR ASYMYETRICA 

VAR GLAUCA 

VAR MACROPODA 

181


SHEET NO. KIVD OF TYPE TA XON 

us (U. S. NATIONAL HERBARIUM, SMITHSONIAN INSTITUTION, WASHINGTON) 

579795 

857864 

2466328 

1411790 

1414090 

854950 

1531248 

368818 

270933 

2 2659 59 

2 176495 

1123660 

2 449 5 0 6 

468192 

8 178 10 

397187 

251773 

817237 

6352 5 

660800 

319177 

29651 

28685 

509004 

28206 

802160 

235568 

235569 

2265956 

1872574 

1733722 

605797 

16970 5 7 

30661 

82 5890 

132 5847 

2074700 

430229 

801132 

193201 5 

538796 

29211 

2231577 

800846 

964080 

1746479 

27238 

301267 

TYPE MATERIAL 

TYPE 

HOLOTYPE 

COTYPE 


TYPE YATERIAL 

TYPE YATERIAL 


I SOTYPE 

HOLDTYPE 

I SOTY PE 

I SOTYPE 

ISOTY PE 

TYPE 

HOLOTYPE 



TYPE COLLECTIQN 

TYPE MATERI AL 

TYPE 

ISClTYPE 

SYNTYPE 

I SOTYPE 

ISOTYPE 


TYPE MATERIAL 


TYPE COLLEC T I ON 

HOLOTYP E 

ISOTYPE 

I SOTYPE 

I SOTYPE 

ISOTYPE 


TYPE YATERIAL 

I SOTY PE 

TYPE MATERIAL 

SYNTYPE 

SYNTYPE 

I SOTYPE 

HOLOTY PE 


HOLOTYPE 

SYNTYPE 


I SOTYPE 

TYPE MATERIAL. 

TYPE 

DURANDI I 

EGGLEST'ONII 

EGGLESTONI I 

EKMANI I 

EKYAN I I 

ELMER1 

FLRODl 

ELY NO IDES 

EPAPILLOSA 

EURYSTACHYA 

X EXSALINA 

FARGESI I 

F I SSA 

FLACCIFOLIA 

FRACTA 

FUSCOLUTEA 

FUSCOTI NCT A 

F USC OT I NC T A 

GARBERI 

GEOPHI LA 

GRACILIOR 

HALL I I 

HAR.fORDI I 

HELLERI 

HETERONEURA 

ICHANGENSIS 

I DA HOA 

I DAHOA 

I NC OND I TA 

INTERIMUS 

X N TERI OR VAR CHARLESTONENSIS 

INTERIOR VAR JOSSELYNI I 

INTFRIOR VAR KEWEENAWENSI S 

I NVOLUC RATELLA 

JAME SON I VAR SUBFULVA 

KAT AH01 NENS IS 

KAUAIENSIS 

LACUNARUM 

LANC IFOL I A 

LARENS! S 

LEIOPYYLLA 

LEYMONI 

LI YNOPH I L A 

L ON G ICQ UR I S 

LUZUL INA 

MACKENZ I ANA 

M ACR OGL 0 S S A 

MAORENS IS 

VAR FESTIVELLIFORMIS 

VAR HOTTENSI S 

VAR ARISTATA 

VAR HENRY1

NUMBER 12 183 

SHEET. Nile KIND OF TYPE TAXON 

us (U. S* NATIONAL HERBARIUM, SYITHSONIAN INSTITUTION, WASHINGTON) 

969118 

1789621 

29453 

711171 

27281 

2501314 

30h9 5 

2 543 897 

886422 

2133195 

23257 

2095886 

2728C 

1438017 

2433719 

2 176496 

286861 

31277 

251772 

817656 

16781 88 

30695 

504456 

27292 

415172 

529528 

872860 

31344 

27275 

2133207 

461358 

154583 1 

31374 

15638 11 

660821 

27269 

24202 76 

2074725 

27270 

458 108 

2265957 

865056 

817724 

1123683 

1282178 

1398 8 3C 

243371 8 

2133193 

TYPE YATERIAL 

ISOTYPE 

I SOTYPE 


TYPE MATERIAL 

I SOTYPE 

I SOTYPE 

HOLOTY PE 

HOL13 TY PE 

ISOTYPE 

SYNTYPE 

I SOTYPE 

TYPE M4TERI 4L 


I SOTYPE 

I SOTY PE 

I SOTYPE 

TYPE YATERIAL 



I SOTYP E 

1 SOTYPE 

I SOTYPE 


TYPE 

TYPE YATERIAL 

TYPE Y4TERIAL 

I SOTYPE 

TYPE MATERIAL 

TYPE 

I SOTYPE 

TYPE 

TYPE Y4TERIAL 

TYPE 

TYPE 


HOLOTYPE 

TYPE YATERIAL 

TYPE YAYERI AL 

TYPE YATERIAL 

HOI.OTY PE 

TYPE MATERIAL 


I SOTY PE 

TYPE 

TYPE M4TERIAL 

I S OTY P E 

TYPE 

MAGNIFOLI 4 

MELOZITYENSIS 

MENDOC I NENS IS 

MERR ILL1 I 

MICANS 

MICRANTYA 

MICROGLOCHIN 

M I C ROPT E RA 

MISERABILI S 

MOHR I ANA 

MONTANE NS I S 

MORRI SSEYI 

NANA 

NEBRASKENS IS 

X NEDRIGELOWI? 

X NEOPALEACEA 

NERVINA 

NUTANS 

OAXACANA 

OAX 4CANA 

OB I SPOENSI S 

OL I GANTHA 

ONUSTA 

OXYC4RPA 

PAC HY STOM A 

PADDOENSI S 

PAL A WAN ENS IS 

PAPULOSA 

PARCIFLORA 

PERCOSTATA 

PERLONGA 

P HA L ARO I DE S 

PICTA 

PIRCHINCHENSIS 

PITYOPHILA 

PLAN AT A 

P L E C TOC A RP A 

PLUVICA 

POD OGY NA 

PRAINII 

PRAT ICOLA 

PRESLI I 

PR I NGLE I 

PT EROL EPT A 

PUR P UREOVAG I N ATA 

PYCNOTHY SOS 

X QUEBECENSIS 

QUICHENS I S 

SSP FUEGINA 

VAR CRASSINERVIA 

V AR ERUCAE FORM I S 

VAR JAPONICA 

VAR PARVULA 

VAR SIMPLEX 

VAR KOOLAUENSI S 

VAR SUBCORIACEA 

VAR ITAT IAIAE



us (U. So NATIONAL HERBARIUM, SMITHSONIAN INSTITUTION, WASHINGTON) 

1872576 

626608 

2133190 

1754487 

200 3 132 

30267 

29888 

319226 

154575 2 

2176493 

528631 

305734 

528495 

511177 

1736782 

30329 

20063 86 

452499 

616142 

96909 1 

28683 

1765699 

710428 

143792 6 

1932033 

20923 56 

2604282 

568126 

1675120 

1933688 

1872573 

20 50647 

2 23 14% 5 

886234 

19678 19 

279151 

2074653 

2133191 

2G50636 

TYPE MATERIAL 


HOLOTYPE 

ISOTYPE 

I SOTY PE 


TYPE COLLECT I3N 


TYPE 

I SOTY PE 

I SOTYPE 

TYPE COCC ECT ION 



IS CITY PE 

SYNTYPE 

I SOTYPE 

HOLOTY PE 



ISOTYPE 

I SOTYPE 

TYPE MATERIAL 


I SDTYPE 

TYPE MATERIAL 

ISOTYPE 

YOLOTYPE 

TYPE Y4TER IAL 

I SOTYPE 

I SOTYPE 

I SOTYPE 

HOLOTYPE 

TYPE 

I SOTY PE 

I SOTYPE 



I SOTYPE 

RACHILL I S 

R AMOS I I 

ROANENSI S 

RUBRO-BRUNNEA 

RUG AT A 


P. use yr 

SAL INAEFORY IS 


SACTAENSI S 

X SAXENII NM. FERRUGINEA 

SCABRIUSCULA 

SEATONIANA 

SHELDON I I 

SMALLIANA 

SONOMENSIS 

SPECIFICA 

S PECUI C OL A 

STELLATA 

STE NOPHY L L A VAR DESERTORUM 

STI PATA VAR MAXIMA 

SUB- BR ACTE ATA 

SlJRNIGRI CANS 

SUB TR A N SVE R S A 

S UK S DOR F I I 

TACHIRENSIS 

TETSUOI 

TOYPKI NS I 

TOWNSEND1 I 

TSOI 


VE R N AC UL A 

VES ICARIA 

VEX A M 

V I ? I DI OR 

VITIENSIS 

V I TREA 

WAHUENSIS 

WILLDENOWI I 

X XANTHINA 

VAR HORSONII 


VAR RUBIGINOSA 

V A R M E G A R R H Y NC H A

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Crovello, T. J. 

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An Introduction to the Botanical Type Specimen Register

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