Family 270. RUBIACEAE Acranthera Arnott MINDANAO ...

Aidia foveata Ridsd., Blumea 41 (1996) 166. --Type from Sabah.Borneo (Kalimantan, Sarawak, Sabah) and Philippines. MINDANAO, RS Williams 2161.Aidia pulcherrima (Merr.) Ridsd., Blumea 41 (1996) 173; --Gymnopachis pulcherrima (Merr.)Tirveng. in Tirveng. & Sastre, Bull. Mus. Nat. Hist. Nat., B, Adansonia, 4e sér., 8 (1986) 285;--Randia pulcherrima Merr., PJS 3 c (1908) Bot. 164; EPFP 3 (1923) 528. –Lectotype(Tirvengandum & Sastre, 1986): Clemens 765 (not seen); syntypes: FB 7863 (US*),Elmer 9127(A, L, NY*, US*);LUZON: Laguna, Quezon (Lucban; Mt Banahaw), Camarines, CATANDUANES,MINDANAO: Lanao del Sur. Medium altitude primary forests.Aidia racemosa (Cav.) Tirveng., Nord. J. Bot. (1983) 455; --Tirveng. & Sastre, Bull. Mus. Nat.Hist. Nat., B, Adansonia, 4e sér., 8 (1986) 262; --Ridsd., Blumea 41 (1996) 149; --Petungaracemosa (Cav.) K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 4 4 (1891) 80; --Merr., PJS 2 c(1907) Bot. 426; --Elmer, LPB 3 (1911) 1007; ibid. 4 (1912) 1345; --Merr., EPFP 3 (1923) 533;--Randia racemosa Roxb., Hort. Beng. (1814) 15, nom. nud.; Fl. Ind. ed. 2 (1824) 144; --Fern.-Villar,Novis. App. (1880) 108; --Merr., EPFP 3 (1923) 528; --Stylocoryna racemosa Cav. Icon. 4(1798)45, t. 368. –Type: Nee sn, 1793 (MA, cited but not seen by Ridsdale, 1996).Petunga roxburghii DC, Prodr. 4 (1830) 399; --Hook.f., Fl. Brit. Ind. 3 (1880) 120.Nicobar Isls, Peninsular Thailand, Vietnam, Hainan, Taiwan, Peninsular Malaysia, Java,Borneo, Lesser Sunda Isls, Philippines, Sulawesi, Christmas Is, New Guinea, SolomonIsls, Australia (Northern Territory; Queensland; Western Australia), Nauru, Samoa,Tonga, Wallis-Futuna Isls, Caroline Isls, Marianas, Marshall Isls.N LUZON (Cagayan) to PALAWAN and MINDANAO, in most islands and provinces.Chiefly in secondary forests at low altitudes.Amaracarpus BlumeAmaracarpus pubescens Blume, Bijdr. 16 (1826) 954.ssp. microphyllus (Miq.) AP Davis, Blumea 49 (2004) 35.Amaracarpus apoensis Elmer, LPB 8 (1915) 2804; --Merr. EPFP 3 (1923) 568. –Type: Elmer 10737 (PNH,holo; iso: A, L, NY*, P). Mindanao: Davao del Sur prov., Mt Apo, Todaya, May 1909.Philippines, Sulawesi, Moluccas (Halmahera; Seram).MINDANAO: Davao del Sur (Mt Apo). In primary forests, altitude about 1050 m.Andria Puttock, ined.Andria loheri (Merr.) Puttock, ined.; --Randia loheri Merr., PJS 27 (1925) 57; EPFP 4 (1925) 254.--Type: BS 12067 Loher, cited by Merrill as ‘12967’ in the protologue (A*, iso). Luzon: Rizal prov.,Montalban, Paningtingan. Oct 1909.LUZON: Rizal. Merrill (1925) listed two collections, from rather adjacent localities on thefoothills of the Sierra Madre.Antherostele BremekampRevision:Bremekamp, CEB. 1940. Antherostele genus novum Rubiacearum Urophyllo affine. Journal ofthe Arnold Arboretum 21: 25–31.

Antherostele banahaensis (Elmer) Bremek., J. Arnold Arb. 21 (1940) 27, 28; --Urophyllumbanahaense Elmer, LPB 1 (1906) 70; --Merr., EPFP 3 (1923) EPFP 522. –Type: Elmer 7746 ‘7946’in the protologue (L*, holo). Luzon: Quezon prov., Lucban, May 1906.LUZON: Isabela (Palanan), Rizal (Mt Lumutan), Quezon (Mt Binuang; Mt Banahaw deLucban), Laguna (Mt Makiling), Sorsogon (Mt Bulusan). In damp primary forests at lowand medium altitudes, ascending to 1000m.Antherostele callophylla Bremek., J. Arnold Arb. 21 (1940) 29. –Type: BS 33326 Ramos (A*,holo). Luzon: Ilocos Norte prov., Mt Palimlim. Aug 1918.LUZON: Ilocos Norte (Mt Palimlim). Habitat not noted.Antherostele grandistipula (Merr.) Bremek., J. Arnold Arb. 21 (1940) 30; --Urophyllumgrandistipulum Merr. PJS 8 c (1913) Bot. 61; --Merr., EPFP 3 (1923) EPFP 522. –Type: BS 15372Ramos (US*, holo). Leyte: Leyte prov., mountains near Dagami, in forests, 13 Aug 1912.LUZON: Camarines, MINDORO, CATANDUANES, LEYTE: Leyte (Mtns near Dagami),Southern Leyte (Cabalian), SAMAR: Catubig River. In damp primary forests at low ormedium altitudes.Antherostele luzoniensis (Merr.) Bremek., J. Arnold Arb. 21 (1940) 30; --Urophyllumluzoniense Merr., PJS 12 c (1917) Bot. 161; --Merr., EPFP 3 (1923) EPFP 523. –Type: BS 26522Ramos & Edaño (US*, holo; iso in A*). Luzon: Aurora prov., Mt Dingalan, in forest, altitude at least300m, 8 Sept 1916.LUZON: Isabela (Mt Moises), Aurora (Mt Dingalan). Primary forests, probably above300m.Antirhea Commerson ex JussieuRevisions:Jansen, ME. 1984. A synopsis of Guettardella Benth. and the Old World species of Antirhea(Rubiaceae: Guettardeae). Blumea 29: 565–588.Chaw, Shu-Miaw & SP Darwin. 1992. A systematic study of the Paleotropical genus Antirhea(Rubiaceae: Guettardeae). Tulane Studies in Zoology & Botany 28: 25–118.Antirhea attenuata (Elmer) Chaw in Tulane Stud. Zool. Bot. 28 (1992) 75; --Timonius attenuatusElmer, LPB 1 (1906) 34. –Type: FB 1867 Ahern’s Collector (NY*, US*, isolecto). Luzon: Rizal prov.,Bosoboso, Sept 1904Antirhea hexasperma (Roxb.) Merr., EPFP 3 (1923) 540; --Guettardella hexasperma (Roxb.) ME Jansen,Blumea 29 (1984) 575. –Type:LUZON: Ilocos Norte, Cagayan, Rizal, Batangas, Camarines. Lowland forests.Antirhea benguetensis (Elmer) Valeton, Beih Bot. Centralbl. 34, 2, (1916) 44, in obs.; --Merr.,EPFP 3 (1923) 540; --Timonius benguetensis Elmer, LPB 1 (1906) 35. –Type: Elmer 6396 (NY*,US*, iso). Luzon: Benguet prov., Twin Peaks, on bluffs along Bued Rivers, May 1904.Guettardella microphylla (Bartl. ex DC) ME Jansen, Blumea 29 (1984) 577; --Antirhea microphylla sensuMerr., EPFP 3 (1923) 540, pro parte.LUZON: Benguet, Nueva Ecija, Rizal. Low altitude thickets or forests, c. 300m.Antirhea bombysia Chaw in Tulane Stud. Zool. Bot. 28 (1992) 78. –Type: PNH 12400 Sulit (L*, holo).Palawan: Victoria Mtns, SE slopes, stunted ridge forest, altitude 1000m, 18 May 1950.PALAWAN: Victoria Mtns. Ridge forest, presumably on ultramafics.Antirhea caudata (ME Jansen) Chaw in Tulane Stud. Zool. Bot. 28 (1992) 80; --Guettardellacaudata ME Jansen, Blumea, 29 (1984) 572. –Type: PNH 12315 Sulit (L*, holo; iso L*). Palawan:Victoria Mtns, SE base, dense forest along Karaniogan River, altitude 100m, 10 May 1950.

Antirhea philippinensis Elmer, LPB 3 (1911) 1009; ibid. 4 (1912) 1329, non Rolfe. As to (cite collections).Guettardella microphylla (Bartl. ex DC) ME Jansen, Blumea, 29 (1984) 577.PALAWAN.Antirhea edanoi Chaw in Tulane Stud. Zool. Bot. 28 (1992) 83. –Type: BS 44015 Ramos & Edaño(NY*, iso). Tawi-tawi, Jul-Aug 1924Antirhea livida Merr. EPFP 3 (1923) 540, pro parte,SULUAntirhea foveolata Chaw in Tulane Stud. Zool. Bot. 28 (1992) 85. –Type:Antirhea microphylla sensu Merr. (1923) non Bartl. ex DC,LUZONAntirhea hexasperma (Roxb.) Merr., EPFP 3 (1923) 540. –Type:Guettardella hexasperma (Roxb.) ME Jansen, Blumea 29 (1984) 575.Non-Philippine!Antirhea livida Elmer, LPB 4 (1912) 1327. –Type: Elmer 12768 (BISH*, L*, NA*, NY*, U*, US*, iso).Palawan: Palawan prov., Puerto Princesa, Mt Pulgar (=Thumb Peak), ‘good moist soil with gravellysubsoil in humid woods or forests along the river at 250ft’, Apr 1911.Guettardella livida (Elmer) ME Jansen, Blumea 29 (1984) 576.PALAWAN:Antirhea microphylla (Bartl. ex DC) Merr., EPFP 3 (1923) 540; --Guettardella microphylla (Bartl.ex DC) ME Jansen, Blumea 29 (1984) 577. –Type:LUZON:Antirhea paxillata Chaw in Tulane Stud. Zool. Bot. 28 (1992) 100.LUZON: Mt BabatginAntirhea philippinensis (Benth.) Rolfe, J. Linn. Soc. 21 (1884) 312; --Guettardella philippinensisBenth. in Hook., Kew J. 4 (1852) 197. –Type:BOHOLAntirhea ramosii Chaw in Tulane Stud. Zool. Bot. 28 (1992) 103. –Type: BS 46597 Ramos & Edaño(NY*, iso). Luzon: Cagayan prov., Peñablanca, Apr 1926.Antirhea microphylla sensu Merr. (1923), pro parte, non Bartl. ex DC.LUZONAntirhea tayabensis Chaw in Tulane Stud. Zool. Bot. 28 (1992) 109. –Type: BS 28869 Ramos &Edaño (NY*, holo). Luzon: Quezon prov., Mt Binuang, May 1917.Antirhea livida Merr., EPFP 3 (1923) 540, pro parte, non Elmer (1912), as to BS 28514, 28869 & 28982Ramos & Edaño.LUZONAntirhea ternata Chaw in Tulane Stud. Zool. Bot. 28 (1992) 113.SIARGAO.Anthocephalus A Richard =NeolamarckiaLiterature:Argostemma Wallich

Bakhuizen van den Brink, RC Jr. 1953. Notes on Malaysian Rubi aceae. Blumea 7: 329-338.Bremer, B. 1989. The genus Argostemma (Rubiaceae-Argostemmateae) in Borneo. Annals of theMissouri Botanical Garden 76: 7-49.Argostemma arachnosum Merr., PJS 20 (1922) 465; EPFP 3 (1923) 504. –Type: BS 36830Ramos & EdañoMINDANAO: Zamboanga. Along streams at low altitudes.Argostemma bryophilum K Schum. in K Schum. & Lauterb. Nachtr. Fl. Deutsch. Sudsee (1905)393. –Type from New Guinea.Argostemma maquilingense Elmer, LPB 9 (1934) 3242. –Type: Elmer 17637 (PNH, holo, lost; iso BISH*, L*,MO*, NY*, US*). Luzon: Laguna prov., Mt Makiling, Jun-Jul 1917.Argostemma rupestrinum Elmer, LPB 9 (1934) 3244. Type: Elmer 16175 (PNH, holo, lost; iso MO*, NY*,US*). Luzon: Sorsogon prov., Mt Bulusan, May 1916.Borneo (Sarawak), Philippines, New Guinea.LUZON: Laguna (Mt Makiling), Sorsogon (Mt Bulusan). Ecol.Argostemma distichum Valeton, Nova Guinea 8 (1911) 447.subsp. quadripetalum (Elmer) Bakh.f., Blumea 7 (1953) 333; --Argostemma quadripetalumElmer, LPB 1 (1908) 361; (1906) 63, sphalm. ‘quadripetiolata’ --Merr., EPFP 3 (1923) 504. –Type:Elmer 7944.LUZON, LEYTE, MINDANAO. Along shaded streams and in dense forests at mediumaltitudes, ascending to 1200m.Argostemma neesianum Walp., Nov. Act. Acad. Nat. Cur. 19 (1843) Suppl. 1: 349; --Merr., EPFP3 (1923) 503. –Type: FB 3368 Ahern’s Collector.LUZON: Pangasinan, Bataan, Rizal, Laguna. On damp shaded ledges and banks alongsmall streams at low altitudes.Argostemma solaniflorum Elmer, LPB 1 (1906) 2; --Merr. PJS 2 (1907) Bot. 305, 3 (1908) Bot.436. –Type: Elmer 6211 (US*, iso). Luzon: Benguet prov., Sablan, damp shady streambanks, Apr1904.Ryukyus, Taiwan, Lanyu, and the Philippines.BATAN ISLS; N LUZON and MINDANAO and the SULU Archipelago. Chiefly in themossy forest at and above an altitudes of 900m, but also along mountain streams atlower altitudes. According to Merrill (1923), not certainly distinct from the Malayan A.montanum Blume.Argostemma stenophyllum Merr., PJS 10 c (1915) Bot. 100; EPFP 3 (1923) 504. –Type: BS20566 Ramos (US*, iso). Luzon: Laguna prov., San Antonio, on rocks along the river, 10 Mar 1913.LUZON: Laguna, CATANDUANES. On damp banks along shaded streams at low ormedium altitudes.Argostemma timorense Benn., Pl. Jav. Rar. i. (1838) 95.Argostemma pedicellatum Elmer, LPB 9 (1934) 3243. –Type:Philippines, Lesser Sunda Isls (Timor).LUZON, LEYTE, MINDANAO. Along shaded streams and in dense forests at mediumaltitudes, ascending to 1200m.Argostemma wallichii Walp., Nov. Act. Acad. Nat. Cur. 19 (1843) Suppl. 1: 350; --Merr., EPFP 3(1923) 504. –Type:LUZON: Benguet, Bulacan, Rizal, Laguna, Batangas. On damp banks and cliffs alongsmall shaded streams at low altitudes.

Unplaced:Argostemma umbellatum Elmer, LPB 9 (1934) 3245, nom. illeg., non Geddes, 1927. –Type:Atractocarpus Schechlter & K KrauseLiterature:Ridsdale, CE. 1979. The taxonomic position of Sulitia (Rubiaceae). Blumea 25: 301–303Atractocarpus obscurinervius (Merr.) Puttock, Austral. Syst. Bot. 12 (1999) 304; --Sulitiaobscurinervia (Merr.) Ridsd., Blumea 25 (1979) 301; --Gardenia obscurinervia Merr., PJS11 c (1916) Bot. 32; EPFP 3 (1923) 531. –Type: BS 23492 Ramos (A*, L*, NY*, US*, iso).Luzon: Sorsogon prov., Mt Pocdal, 1 Aug. 1915.Sulitia longiflora Merr., PJS 29 (1926) 494. –Type: FB 29445 Sulit (not seen). Mindanao: Sultan Kudarat(formerly part of a bigger Cotabato) prov., near Lebak, Apr & May 1903. Reportedly epiphytic.LUZON: Laguna, Sorsogon, CATANDUANES, MINDANAO: Sultan Kudarat. In primaryforest at low altitudes.Badusa A GrayLiterature-Ridsdale, CE 1982. A revision of the genus Badusa (Rubiaceae, Condamineeae, Porlandiinae).Blumea 28:145–150.Soejarto, D. et. al. 1996. The true provenance and identity of Badusa philippica (Rubiaceae).Taxon 45:487–492.Badusa palawanensis Ridsd., Blumea 28 (1982) 146. Type: PNH 80766 HG Gutierrez & RA Espiritu(L*). Palawan: Palawan prov., Quezon, Lipuun Point, rocky summit of limestone hill, altitude 500ft [=m], 24Mar 1963.PALAWAN: Quezon, Lipuun Point. Rocky summit of limestone hill.Badusa corymbifera (G Forst.) A Gray, Proc. Amer. Acad. Arts 4 (1860) 308; --Soejarto et al.,Taxon 45: 487-492.Badusa philippica (Cav.) Vidal, Rev. Pl. Vasc. Filip. (1888) 150. --Type: Nees sn, prope Manila in Sta. Cruzde Laguna (not traced in MA).Literature:Darwin, S. 1979. Allertonia 2(1): 22.Bikkia ReinwardtBikkia philippinensis Valeton, PJS 23 (1923) 6; --Merr., EPFP 3 (1923) 492.CEBU, SULU ARCHIPELAGO (SIBUTU), MINDANAO: Surigao. In thickets along theseashore.Boholia MerrillBoholia nematostylis Merr., PJS 29 (1926) 493. –Type: BS 43107 (A*, US*, iso). Bohol: Bilar, dampforest, altitude c. 600m, Aug 1923.Sumatra, Lesser Sunda Island and C Philippines.BOHOL (Bilar; Kalingohan). Presumably on limestone.Borreria GFW Meyer =Spermacoce

Caelospermum BlumeRevision:Johansson, JT. 1988. Revision of the genus Caelospermum (Rubiaceae, Rubioideae,Morindeae). Blumea 33: 265–297Caelospermum volubile (Merr.) JT Johansson, Blumea 33 (1988) 281, Fig. 11; --Morindavolubilis (Blanco) Merr., PJS 1 (1906) Suppl. 137; EPFP 3 (1923) 574. –Type: FB 2290 Meyer (K,lecto, selected by Johansson, 1988; isolecto: F, NY). Luzon: Bataan prov., Mt Mariveles, LamaoRiver, Dec 1904.LUZON: Zambales, Bataan, Laguna, CATANDUANES, MINDORO. Lowland secondaryforests to above 100m. Fl. Jul, Aug, Dec.Canthium LamarckLiterature:Bridson, DM. 1992. The genus Canthium (Rubiaceae-Vanguerieae) in tropical Africa. Kew Bull.47.Canthium amplifolium (Elmer) Co, comb. nov.; --Plectronia amplifolia Elmer, LPB 9 (1934) 3261.–Type: Elmer 16739 (BISH*, L*, NA*, NY*, US*, iso). Luzon: Sorsogon prov., Irosin, Mt Bulusan, Jul 1916.LUZON: SorsogonCanthium brunneum (Merr.) Merr., PJS 35 (1928) 7; --Plectronia brunnea Merr., PJS 17 (1920,publ. 1921) 441; EPFP 3 (1923) 535. –Type: BS 28791 Ramos & Edaño (US*, iso). Luzon: Quezonprov., Mt Binuang, May 1917.LUZON: Quezon, Camarines. Mossy forest, altitude c. 1000m.Canthium confertum Korth., Ned. Kruidk. Arch. 2 (1851) 235; --KM Wong in FS Ng (ed.) Tree Fl.Malaya 4 (1989) 342.?Canthium villarii Vidal, Phan. Cuming. Philipp. (1885) 120, 182. –Type:Peninsular Malaysia, Borneo and Philippines.Canthium culionense (Elmer) Merr., PJS 35 (1928) 7; --Plectronia culionensis (Elmer) Merr.,iso). Culion: Halsey, in dry forests, Dec 1902.CULION. Lowland dry secondary forests and thickets.EPFP (1923) 535; --LaCanthium dicoccum (Gaertn.) Merr., PJS 35 (1928) 8; --Plectronia dicocca (Gaertn.) Merr.,EPFP (1923) 536; --Psydrax dicoccos Gaertn., Fruct. 1 (1788) 125, t. 26.Canthium didymum Gaertn.India to Malaya.LUZON: Benguet, Pangasinan, Zambales, Bataan, Rizal, Batangas, MINDORO,PALAWAN, TICAO, GUIMARAS, NEGROS, MINDANAO, BUCAS GRANDE. Lowlandprimary and secondary forests. (Merrill, 1923).Malesian materials hitherto distributed and interpreted as C. dicoccum (=Psydraxdicocca Gaertn.) were possibly misidentified according to Bridson, Kew Bull. 40 (1985),as this taxon is restricted to Sri Lanka and S India. Canthium didymum Gaertn. is asynonym of Psydrax dicocca Gaertn., the type species of Psydrax. In Malaya, severalentities previously annotated in herbaria as C. didymum have now been placed underPsydrax by KM Wong (Tree Fl. Malaya 4 (1989) 400). Similar case for some Philippineentities currently under Canthium?

Canthium edanoi Co, nom nov.; --Canthium longistipulum Merr. & Quisumb., non Merr., Mitt. Inst.Allg. Bot. Hamburg 7 (1937) 291. --Type: PNH 151 Edaño (L*). Palawan: Brookes Point,Mantalingahan Range, Mt Balabag, mossy forest, altitude 1573m, 13 May 1947.PALAWAN: Brookes Point, Mt Mantalingahan.Canthium ellipticum (Merr.) Merr., PJS 35 (1928) 8; --Plectronia elliptica Merr., PJS 12 c (1917)166; EPFP 3 (1923) 536. –Type: BS 29594 Ramos & EdañoLUZON: Nueva Ecija, Rizal. Medium altitude forests.Canthium elmeri Merr., PJS 35 (1928) 8; --Plectronia gracilipes Elmer, LPB 3 (1911) 1008, nonKurz; --Merr., EPFP 3 (1923) 536. –Type: Elmer 12550SIBUYAN. Forests along streams, altitude c. 225m.Canthium fenicis (Merr.) Merr., PJS 35 (1928) 8; --Plectronia fenicis Merr., PJS 8 c (1913) Bot. 166;EPFP 3 (1923) 536. –Type: BS 12693 Fenix (not seen). Luzon: Benguet prov., Sablan, growing onslopes, 23 Nov 1910.LUZON: Benguet, Tayabas, CEBU (Danao). Medium altitude secondary forests.Canthium glandulosum (Blanco) Merr., PJS 35 (1928) 8; --Plectronia glandulosa (Blanco) Merr.,Sp. Blancoanae (1918) 365; EPFP 3 (1923) 536; --Ixora glandulosa Blanco, Fl. Filip. (1837) 61.–Type: None extant; ‘illustrative specimen’: Merrill Sp. Blancoanae 897 (US*).Polyozus bipinnatus Blanco, Fl. Filip. ed. 2 (1845) 43; ibid. ed. 3, 1 (1877) 79. –Type: None extant;‘illustrative specimen’: Merrill Sp. Blancoanae 897 (US*).Plectronia viridis Merr., PJS 1 (1906) Suppl. 131; --Elmer, LPB 3 (1911) 1008. –Type: Merrill 3945 (US*,LUZON: Cagayan, Bataan, Rizal, Quezon, MINDORO, SIBUYAN, NEGROS,MINDANAO. Low and medium altitude forests.iso). Luzon: Bataan proCanthium gynochthodes Baill., Adansonia 12 (1878) 199; --Plectronia gynochthodes (Baill.)Merr., EPFP 3 (1923) 536. –Type:Lanyu, Lutao, and the Philippines.N LUZON to PALAWAN and MINDANAO, in most islands and provinces. Low andmedium secondary forests generally, ascending to 1500m.Canthium horridum Blume, Bijdr. (1826) 966; --KM Wong in FS Ng (ed.). Tree Fl. Malaya 4 (1989)341; --Plectronia horrida (Blume) Benth. & Hook.f. ex Kurz in J. As. Soc. Beng. 46, 2 (1877) 153;--Merr., EPFP 3 (1923) 537.Canthium pauciflorum Blanco, Fl. Filip. (1837) 165. –Type: Not extant. ‘Illustrative specimen’: Merrill Sp.Blancoanae 143 (US*)..Burma, Indochina, Peninsular Malaysia, Java, Borneo, Philippines.LUZON: Pangasinan, Zambales, Bulacan, Rizal, Camarines, Sorsogon. Lowland drythickets.Canthium leytense (Merr.) Merr., PJS 35 (1928) 8; --Plectronia leytensis Merr., PJS 8 c (1913) 166;EPFP 3 (1923) 537. –Type: BS 15383 Ramos (US*, iso). Leyte: Leyte prov., mountains back ofDagami, in the mossy forest, Aug 1912.LUZON: Quezon (E coast), LEYTE. Primary forests, probably at c. 1000m.Canthium megacarpum (Merr.) Merr., PJS 35 (1928) 8; --Plectronia megacarpa Merr., PJS 4 c(1909) Bot. 326; EPFP 3 (1923) 537. –Type: FB 8232 Curran & Merritt (US*, iso). Luzon: Zambalesprov., near Sta Cruz, altitude 100m, Dec 1907.LUZON: Benguet, Cagayan, Zambales, Quezon. Lowland forests.Canthium merrillianum Mabb., Taxon 26 (1977) 532; --Canthium cordatum (Merr.) Merr., PJS 35(1928) 7, nom illeg., non Dillwyn, 1839; --Plectronia cordata Merr., PJS 12 c (1917) Bot. 166; EPFP3 (1923) 535. –Type: BS 26549 Ramos & Edaño (A*, US*, iso). Luzon: Aurora prov., Mt Dingalan,

Aug-Sept 1916.LUZON: Aurora. Forested slopes, altitude c. 2000m.Canthium monstrosum (A Rich.) Merr., PJS 35 (1928) 8; --Plectronia monstrosa A Rich., Mém.Rub. (1830) 109; --Merr., PJS 8 c (1913) Bot. 48; Sp. Blancoanae (1918) 365; EPFP 3 (1923) 538.–Type: Perrottet sn (P, 2 sheets, cited by Merrill, 1913).Canthium bipinnatum (Blanco) Merr., Govt. Lab. Publ. (Philip.) 27 (1905) 53; --Ronabea bipinnata Blanco,Fl. Filip. (1837) 162. –Type: Not extant. “Illustrative specimen”: Merrill Sp. Blancoanae 248, leg. MRamos (US*). Luzon: Rizal prov., Antipolo, 19 Jan 1913.Canthium mite Bartl. ex DC., Prodr. 4 (1830) 474; --Plectronia mitis (Bartl. ex DC.) Elmer, LPB 1 (1906) 28.Canthium arboreum (Blanco) Vidal, Phan. Cuming. Philip. (1885) 119, 181; Rev. Pl. Vasc. Filip. (1886) 155;--Ronabea arborea Blanco, Fl. Filip. ed. 2 (1845) 114; idem. ed. 3, 1 (1877) 204. –Syntype: Cuming776 (L*, NY*). Luzon: Quezon prov.Philippines, N Sulawesi.LUZON: Ilocos Norte, Ilocos Sur, Abra, La Union, Apayao, Mountain Province, Benguet,Cagayan, Isabela, Nueva Vizcaya, Pangasinan, Zambales, Bataan, Pampanga,Bulacan, Rizal, Laguna, Cavite, Quezon, LEYTE, NEGROS. Low and medium altitudesecondary forests, often common.Canthium oblongifolium Quisumb. & Merr., PJS 37 (1928) 200. –Type: BS 46109 Edaño (NY*,US*, iso). Panay: Capiz prov., Haman-ay, forested slopes at low altitudes, 13 Oct 1925.PANAY: Capiz. Lowland forest.Canthium obovatifolium (Merr.) Merr., PJS 35 (1928) 8; --Plectronia obovatifolia Merr., PJS 12 c(1917) Bot. 167; EPFP 3 (1923) 538. –Type: BS 26526 Ramos & Edaño (A*, US*, iso). Luzon:Aurora prov., Mt Dingalan (Mt Dalingdingan in the protologue), forested slopes at medium altitudes,9 Sept 1916.LUZON: Aurora. Medium altitude forests.Canthium oligophlebium (Merr.) Merr., PJS 35 (1928) 8; --Plectronia oligophlebia Merr., PJS 17(1920) 442; EPFP 3 (1923) 538. –Type: BS 29342 Ramos and Edaño (A*, US*, iso). Luzon: Rizalprov., Mt Susong Dalaga, mossy forest at the summit, apparently at or above 900m altitude, 7 Aug1917.LUZON: Rizal. Known only from the type collection.Canthium paucinervium (Merr.) Merr., PJS 35 (1928) 8; --Plectronia paucinervia Merr., PJS 8 c(1913) Bot. 51; EPFP 3 (1923) 538. –Type: Merrill 8263 (not seen). Mindanao: Zamboanga, SaxRiver mtns back of San Ramon, in damp forest, altitude c. 900m, 29 Nov 1911.LUZON: Camarines, MINDANAO: Zamboanga, BUCAS GRANDE. Mossy forest, altitudec. 900m at the type locality. Possibly at lower altitude elsewhere.Canthium pedunculare Cav., Icon. 5 (1799) 21, t. 436. –Type:Plectronia peduncularis (Cav.) Vidal, Cat. Pl. Prov. Manila (1880) 33; --Elmer, LPB 1 (1906) 28; idem. 4(1912) 1346; --Merr., PJS 1 (1906) Suppl. 131; idem. 8 c (1913) Bot. 50; Fl. Manila (1912) 454; Sp.Blancoanae (1918) 364; EPFP 3 (1923) 538.Canthium monoflorum Blanco, Fl. Filip (1837) 166. –Type: Not extant. ‘Illustrative specimen’: Merrill Sp.Blancoanae 380 (US*). Palawan: Taytay, dry thickets and open places, May 1913Plectronia lycioides (A Rich.) Elmer, LPB 1 (1906) 28; --Canthium lycioides A Rich., Mém. Rub. (1830) 108.–Type:LUZON: Pangasinan, Bataan, Bulacan, Rizal, PALAWAN. Lowland dry thickets.Canthium ramosii (Merr.) Merr., PJS 35 (1928) 9; --Plectronia ramosii Merr., PJS 17 (1920, publ.1921) 443; EPFP 3 (1923) 539. –Type: BS 28973 Ramos & Edaño (NY*, US*, iso). Luzon: Quezonprov., Umiray, forest along the Umiray River at low altitudes, 1 Jun 1917.LUZON: Quezon (Umiray). Lowland forests.

Canthium sarcocarpum (Merr.) Merr., PJS 35 (1928) 9; --Plectronia sarcocarpa Merr., PJS 10 c(1915) Bot. 114; EPFP 3 (1923) 539. –Type: BS 10968 Ramos (not seen). Luzon: Laguna prov.,San Antonio, in thickets, Aug 1910.LUZON: Laguna, Quezon, Sorsogon. Lowland forests.Canthium subcapitatum (Merr.) Merr., PJS 35 (1928) 9; --Plectronia subcapitata Merr., PJS 17(1920) 443; EPFP 3 (1923) 539. –Type: BS 28117 Fenix (L*, NY*, US*, iso). Luzon: Apayao prov.,Ngagan, on forested slopes, 10 May 1917.LUZON: Apayao. Medium altitude forests.Canthium subsessilifolium (Merr.) Merr., PJS 35 (1928) 9; --Plectronia subsessilifolia Merr., PJS12 c (1917) Bot. 168; EPFP 3 (1923) 539. –Type: BS 27205 Ramos (A*, US*, iso). Luzon: IlocosNorte prov., in thickets at low altitude, 2 Mar 1917.LUZON: Ilocos Norte. Lowland thickets.Canthium trichophorum Quisumb. & Merr., PJS 37 (1928) 201. –Type: BS 46759 Ramos &Edaño (A*, NY*, iso). Luzon: Isabela prov., San Mariano, Ambulawan, along forested streams,altitude c. 400m, 16 Mar 1926.LUZON: Isabela (San Mariano, Ambulawan; Mt Moises). Along forested streams,altitude 400-900m.Canthium wenzelii (Merr.) Merr., PJS 35 (1928) 9. --Plectronia wenzelii Merr., PJS 9 c (1914) Bot.387; EPFP 3 (1923) 539. –Type: Wenzel 399 (A*, US*, iso). Leyte: Leyte prov., 30 Aug 1913.LEYTE. Lowland forest, altitude c. 60m.Ceriscoides (Hooker filius) TirvengadumLiterature:Wong, KM. (1984): The genera of Peninsular Malaysian Rubiaceae formerly confused withRandia. Malayan Nat. J. 38: 1–57.Ceriscoides curranii (Merr.) Tirveng., Nordic J. Bot. 3 (1983) 456; --Gardenia curranii Merr., Sp.Blancoanae (1918) 363. –Type:LUZON: Pangasinan, Bataan, Rizal, Batangas. In thickets and forests at low altitudes.Chassalia Commerson ex JussieauLiterature:Deb, DB & B. Krishna. 1982. Taxonomic studies of the genus Chassalia Comm. ex Poir. in Indiaand adjoining regions. Bulletin of the Botanical Survey of India 24 xxxxx.Chassalia curviflora (Wall.) Thwaites, Enum. Pl. Zeylan. (1859) 150; --Elmer, LPB 1 (1906) 3;idem. 3 (1911) 1033; --Merr., EPFP 3 (1923) 565; --Psychotria curviflora Wall. in Roxb. Fl. Ind. 2(1824) 167. –Type from Sri Lanka.Chasalia lurida (Blume) Miq., Fl. Ind. Bat. 2 (1857) 282; --Merr., PJS 10 c (1915) Bot. 140; --Psychotrialurida Blume, Bijdr. (1826) 959. –Type from Java.var. ophioxyloides (Wall.) DB Deb & B Krishna, Bull. Bot. Surv. India 24 (1982, publ. 1983) 222.India, Sri Lanka, E Himalayas, Bangladesh, Nicobar Isls, Burma, Thailand, Borneo,Philippines.MINDORO, PALAWAN, BALABAC, TICAO, BANTON, SIBUYAN, PANAY, LEYTE,MINDANAO: Cotabato, BALUT. Lowland primary forests.Chassalia obscurinervia Elmer, LPB 3 (1911) 1033. –Type: Elmer 11200 (BISH*, GH*, NY*, US*,

iso). Mindanao: Davao del Sur prov., Mt Apo, Todaya, ‘dry forests of fertile soil along the SibulanRiver and the Baracatan Creek from 1250-2500ft’, Jul 1909.MINDANAO: Davao del Sur (Mt Apo).Coelospermum Blume =CaelospermumCoprosma JR & G ForsterRevision:Heads, 1996. Candollea 51:Coprosma granadensis (Mutis ex L.f.) Heads, Candollea 51 (1996) 388; --Nertera granadensis(Mutis ex L.f.) Druce, Bot. Exch. Club Brit. Isles Rep. 4 (1917) 637; --Yang TYA in Huang, TC (ed).Fl. Taiwan, ed. 2, 4 (1998) 306, pl. 138, photo 186.Nertera depressa Banks & Solander ex Gaertn., Fruct. 1 (1788) 124; --Merr., EPFP 3 (1923) 571.Coprosma dentata (Elmer) Heads, Candollea 51 (1996) 389; --Nertera dentata Elmer, LPB 1 (1906) 15. –Type:var. granadensis.Temperate S hemisphere to tropical Asia, Madagascar, Australia, New Zealand, Pacificislands, Andes to Mexico.LUZON: Abra, Mountain Province, Benguet, Zambales, Laguna, Quezon, MINDORO,NEGROS, MINDANAO: Davao. In mossy forests on the tops of the higher mountains,sometimes in shaded ravines as low as 1500m.Coptosapelta KorthalsLiterature-Valeton, Th. 1922. Die Gattung Coptosapelta Korth. Rec. Trav. Bot. Neerl. 19: 281–292. pl. X,XI.Coptosepelta olaciformis (Merr.) Elmer, LPB 5 (1913) 1856; --Merr., EPFP 3 (1923) 507; --Randia olaciformis Merr., PJS 3 c (1908) Bot. 163. –Type:LUZON, LEYTE, MINDANAO. In primary forests at low and medium altitudes.Cowiea WernhamCowiea philippinensis Merr., PJS 20 (1922) 459; EPFP 3 (1923) 533. –Type: BS 36819MINDANAO: Zamboanga. Damp forests along streams at low altitudes.Damnacanthus GaertnerLiterature-Robbrecht, E. et al. 1991. The genera Mitchella and Damnacanthus. Evidence for their alliance,comments on the campylotrophy in the Rubiaceae and the circumscription of theMorindeae. Blumea 35: 307–345.Damnacanthus indicus Gaertn., Fruct. 3 (1805) 18, t. 182;India (Assam), Bangladesh, Burma, S China, Taiwan, C & S Japan, Korea (Cheju Do),Philippines.LUZON: Quezon (Mt Banahaw). On forested slopes, 1400-2000m.Dentella JR & G Forster

Literature:Airy Shaw. 1934. Kew Bull ?: 290Dentella repens (L.) Forst., Char. Gen. (1776) 26, t. 13; –Merr., Fl. Manila (1912) 447, EPFP 3 (1923)492; --Oldenlandia repens L., Mant. 1 (1767) 40..Mauritius, Sri Lanka, India, E Himalayas, Bangladesh, S China, Taiwan, Burma,Thailand, Peninsular Malaysia, Java, Philippines, Sulawesi, Moluccas, NorthernTerritory, Queensland, New Caledonia.Throughout the Philippines in open waste places in and about towns at low and mediumaltitudes. Introduced in Mexico.Diplospora de CandolleLiterature:Ali, SI & E Robbrecht. 1991. Remarks on the tropical Asian and Australian taxa included inDiplospora or Tricalysia (Rubiaceae-Ixoroideae-Gardenieae). Blumea 35: 279–305.Diplospora fasciculiflora (Elmer) Elmer, LPB 3 (1911) 113; --Merr., EPFP 3 (1923) 534; --Tricalysia fasciculiflora (Elmer) Merr., PJS 10 c (1915) Bot. 13; EPFP 3 (1923) 534, pro parte.; --Randia fasciculiflora Elmer, LPB 1 (1906) 4,–Type: Elmer 6900 (NY*, iso). Luzon: Bataan prov.,Mt Mariveles, Nov 1904.Tricalysia fasciculiflora (Elmer) Merr. var. oblongifolia Merr., EPFP 3 (1923) 534. –Type: FB 1117 Ramos(US*, iso). Luzon: Rizal prov., Bosoboso, Jul 1906LUZON: Mountain Province, Benguet, Cagayan, Nueva Ecija, Bataan, Bulacan,Rizal, Batangas, Quezon, Camarines, CULION, PALAWAN, MASBATE, MINDANAO:Cotabato. Low and medium altitude primary forests, ascending to 1600m.Diplospora puberula (Merr.) Ali & Robbrecht, Blumea 35 (1991) 299; --Tricalysia puberula Merr.,PJS 10 c (1915) Bot. 113; EPFP 3 (1923) 535. –Type: BS 20431 Ramos (US*, iso). Luzon:Laguna prov., San Antonio, Feb 1913.LUZON: Isabela, Laguna, CATANDUANES. Lowland primary forests, ascending to300m.Diplospora tinagaoensis (Elmer) Ali & Robbrecht, Blumea 35 (1991) 299; --Tricalysiatinagaoensis Elmer, LPB 1 (1906) 34; --Merr., EPFP 3 (1923) 535. –Type: Ahern 417, leg. Quadras(US*, iso). Tinago Island, Mar-May 1901.Phaleria axillaris Elmer, LPB 8 (1915) 2840. –Type:NEGROS, LEYTE, SAMAR, MINDANAO: Zamboanga, Lanao del Sur, Agusan delNorte, DINAGAT, TINAGO. Low and medium altitude primary forests, ascending to900m.Tricalysia negrosensis Elmer, LPB 2 (1908) 518; --Merr., EPFP 3 (1923) 535. –Type: Elmer 9685NEGROS. Primary forests, altitude c. 1050m. Presumably to be placed in Diplosporasensu str. (fide Ali & Robbrecht, Blumea 35 (1991) 303).Generic position to be determined:Diplospora sessilis Elmer, LPB 4 (1912) 1329. –Type: Elmer 13139 (L*, NY*, US*, iso).Tricalysia sessilis (Elmer) Merr., EPFP 3 (1923) 535.PALAWAN. On gravel bar along Iwahig River, altitude c. 150m.Literature:Discospermum Dalzell

Ali, SI & E Robbrecht. 1991. Blumea 35: 279.Discospermum whitfordii (Elmer) Ali & Robbrecht, Blumea 35 (1991) 301; 302; --Randiawhitfordii (Elmer) Merr., PJS 1 (1906) Suppl. 130; EPFP 3 (1923) 529; --Gardenia whitfordii Elmer,LPB 1 (1906) 4. --Type: Elmer 6643 (NY*, iso). Luzon: Bataan prov., Mt Mariveles, common in thewoods at 600m, Nov 1904.Philippines.Dolicholobium A GrayRevision:Jansen, ME & CE Ridsdale. 1983. A revision of the genus Dolicholobium (Rubiaceae). Blumea29: 251-311.Dolicholobium philippinense Trelease, LPB 3 (1911) 984; --Jansen & Ridsd., Blumea 29(1983) 278. –Type:Dolicholobium hirsutum Elmer, LPB 9 (1934) 3247.LUZON, SIBUYAN, SAMAR. In rainforest understorey, but often in more open situationslike streambanks or exposed ridges; on well-drained to swampy soils, sea level - 1500(-1900) m.Fagerlindia TirvengadumRevision:Ridsdale, C.E. (1985). The genus Fagerlindia (Rubiaceae) in the Philippines. Blumea. 31: 239–244.Fagerlindia emanuelssoniana Ridsd., Blumea 31 (1985) 243, t. 2. --Type: SMHI 1238 JDransfield (L*, holo). Palawan: Km 36, Puerto Princesa to Roxas, (erroneously cited in protologueas Mt Victoria). Extremely degraded forest on ultramafic rocks, alt. 10m, 12 May 1984.PALAWAN. Apparently restricted to ultramafic rocks.Fagerlindia microcarpa (Bartl. ex DC) Ridsd., Blumea 31 (1985) 243; --Randia microcarpa(Bartl. ex DC) Merr., EPFP 3 (1923) 527, comb. Illeg., non Sessé & Mociño (1888). –Type: Haenkesn (G-DC)Randia cumingiana Vidal, Phan. Cuming. Philip. (1885) 179; Rev. Pl. Vasc. Filip. (1886) 153; --Merr., Philip.Bur. Forestry Bull. 1 (1903) 53; PJS 1 (1906) Suppl. 130; Fl. Manila (1912) 449; --Elmer, LPB 1(1906) 32. –Type: Cuming 1366 (L, lecto). Syntypes: Vidal 388 (L), Herb. Propr. 39LUZON: widespread, in most provinces, GOLO, MINDANAO: Zamboanga. Lowlandthickets and secondary forests, common. (Merr., 1923)Randia ticaensis Merr., PJS 5 c (1910) Bot. 245; --Merr., EPFP 3 (1923) 528. –Type: BS 12547Rosenbluth (US*, iso). Ticao: Linadlaran Point, on steep hillside, altitude c. 30m, 24 Dec 1908.TICAO. Lowland forests. Allied to R. cumingiana Vidal (=Fagerlindia microcarpa).Galium LinnaeusLiterature:Royen, P. van. 1983. Galium in The Alpine Flora of New Guinea 4: 2763–2776.Galium philippinense Merr., PJS 1 (1906) Suppl. 238; ibid. 5 c (1910) Bot. 391; --Elmer, LPB 3(1911) 1046. –Type: Elmer 6557.Galium ciliare Elmer, LPB 1 (1906) 4, non Hook.f.LUZON: Mountain Province, Benguet. Open grassy slopes, 1600-2400m.

Galium subtrifidum Reinw. ex Blume, Bijdr. 16 (1826) 944.Galium gaudichaudii DC, Prodr. 4 (1830) 607; Benth. Fl. Austral. 3 (1866) 446; --Elmer, LPB 1 (1906) 4; --Merr., PJS 5 c (1910) Bot. 391; EPFP 3 (1923) 575.N Sumatra, Java, N Philippines, Lesser Sunda Isls (Buru), New Guinea.LUZON: Mountain Province, Benguet. On rocks, ledges, and open banks, altitude 1300-2000m.Gardenia LinnaeusLiterature:Wong, KM. 1982. Notes on Gardenia and Acranthera (Rubiaceae) from Peninsular Malaysia.Gardens Bulletin Singapore 35: 21–32.Gardenia longiflora Vidal, Rev. Pl. Vasc. Filip. (1886) 153; --Merr., EPFP 3 (1923) 530, nom. illeg.,non Ruiz & Pavon 1799, nec (Salisb.) WT Aiton, 1810.LUZON: Cagayan, Isabela, Rizal, Laguna, Tayabas, Camarines, Sorsogon,BUSUANGA, PALAWAN, SAMAR, SIBUYAN, MINDANAO: Zamboanga. In primaryforest at low altitudes.Gardenia pseudopsidium (Blanco) Fern.-Villar, Novis. App. (1880) 109; --Merr., EPFP 3(1923) 531; --Sulipa pseudopsidium Blanco, Fl. Filip. (1837) 497.Gardenia barnesii Merr., Publ. Govt. Lab. (Philipp.) 17 (1904) 47. –Type: FB 163 Barnes (NY*, US*, iso).Luzon: Bataan prov., Lamao River, Jan 1904.Gardenia segmenta Elmer, LPB 4 (1912) 1331; --Merr., EPFP 3 (1923) 531. –Type: Elmer 13153 (A*, L*,NY*, US*, iso). Palawan: Mt Pulgar. May 1911.Gardenia megalocarpa Merr., PJS 20 (1922) 463; EPFP 3 (1923) 530. --Type: BS 38919 Ramos (A*, US*,iso). Mindoro: Mindoro Occidental prov., Paluan. In forest at low altitudes, Apr 1921.LUZON: Cagayan, Benguet, Nueva Vizcaya, Pangasinan, Tarlac, Rizal, Bataan,Batangas, Laguna, Quezon, Camarines, MINDORO, BUSUANGA, PALAWAN,MASBATE, NEGROS, SAMAR, MINDANAO: Cotabato, Davao. In primary forest at lowaltitudes, common.Gardenia ramosii Merr., PJS 29 (1926) 489. –Type: BS 43323 Ramos (A*, iso). Bohol:BOHOLNon-naturalized cultivated species:Gardenia jasminioides (L.) Ellis, Philos. Trans. 51 (1761) 935.Gardenia Auga (L ) Merr., Interp. Herb. Amb. (1917) 485; EPFP (1923) 531.Gardenia florida L., Sp. Pl. ed. 2 (1762) 305;--Merr. Fl. Manila (1912) 449.Cultivated for ornamental purposes, but nowhere spontaneous. Only the doublefloweredform occurs in the Philippines.Gardenia philastrei Pierre ex Pit. in Lecomte et al. (ed.) Fl. Indo-Chine 3 (1923) 253.Geophila D DonGeophila repens (L.) IM Johnst., Sargentia 8 (1949) 281.Geocardia repens (L.) Bakh.f. in CA Backer & RC Bakhuizen van der Brink, Bekn. Fl. Java 15(173) (1956)144; --Carinta repens (L.) LB Sm. & Downs, Sellowia 7 (1956) 88; --Psychotria repens (L.) L.,Amoen. Acad. 5 (1760) 377; --Rondeletia repens L., Syst. Nat. ed. 10, 2 (1759) 928.Geophila herbacea (Jacq.) K.Schum. in HGA Engler & KAE Prantl, Nat. Pflanzenfam. 4(4) (1891) 119; --Kuntze, Rev. Gen. Pl. (1891) 300; --Merr. EPFP 3 (1923) 565.

Geocardia herbacea (Jacq.) Standl., Contr. U. S. Natl. Herb. 17 (1914) 445.Carinta herbacea (Jacq.) W.Wight, Contr. U. S. Natl. Herb. 9 (1905) 216.Mapouria herbacea (Jacq.) Muell.-Arg. in CFP von Martius & auct. suc. (eds.), Fl. Bras. 6(5) (1881) 427.Cephaelis herbacea (Jacq.) Kurz, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 46(2) (1877) 140.Psychotrophum herbaceum (Jacq.) Crantz, Inst. Rei Herb. 2 (1766) 259.Psychotria herbacea Jacq., Enum. Syst. Pl. (1760) 16.Pantropic.BATAN, LUZON: Bataan, Laguna, Quezon, Sorsogon, BALABAC, BILIRAN, LEYTE,MINDANAO, BASILAN, SULU. Chiefly in forest at low altitudes.Greenea Wight & ArnottGreenea hirsuta Elmer =Timonius hirsutus (Elmer) Merr.Greeniopsis MerrillGreeniopsis discolor Merr., PJS 12 c (1917) Bot. 163; --EPFP 3 (1923) 504. --Type: BS 26448Ramos & Edaño.LUZON: Nueva Ecija. In primary forest at or above 300m altitudes.Greeniopsis euphlebia Merr., PJS 17 (1920, publ. 1921) 319; --EPFP 3 (1923) 504. --Type: BS35134 Ramos & Pascasio.BUCAS GRANDE. Lowland open forests.Greeniopsis megalantha Merr., PJS 12 c (1917) Bot. 164; --EPFP 3 (1923) 505. --Type: BS 34587Ramos & Pascasio.MINDANAO: Surigao del Norte. Low and medium altitudes forests.Greeniopsis multiflora (Elmer) Merr., PJS 4 c (1909) Bot. 325; --Elmer, LPB 3 (1911) 981; --EPFP3 (1923) 505; --Mussaendopsis multiflora Elmer, LPB 1 (1906) 14. –Type: FB 490 JF Quadras (notseen). Dinagat, Mar-May 1905.Greeniopsis sibuyanensis Elmer, LPB 3 (1911) 982. –Type:Greeniopsis philippinensis Merr., PJS 4 c (1909) Bot. 324; --EPFP 3 (1923) 505. –Type: Merrill 5216 (L* 2sheets, NY*, US*, iso). Samar: Eastern Samar prov., Borongan, along border of a nipa swamp, Oct1906.Greeniopsis pubescens Merr., PJS 5 c (1910) Bot. 230; --EPFP 3 (1923) 505. –Type:LUZON: Cagayan, Isabela, Camarines, Sorsogon, LEYTE, SAMAR, MINDANAO:Surigao, DINAGAT. Low altitude forests. Often on ultramafics.Guettarda LinnaeusGuettarda speciosa L., Sp. Pl. (1753) 991; --Merr., Fl. Manila (1912) 450,Tropical Asia and Australia to Polynesia.Throughout the Philippines along seashore.Guettardella Champion & Bentham =AntirheaGynopachis Blume =Aidia

Gynochthodes BlumeGynocthodes lenticellata (C Robinson) Merr., EPFP 3 (1923) 571. --Type: FB 13689 Curran.NEGROS. Lowland forests.Gynocthodes mindanaensis Merr., PJS 17 (1920, publ. 1921) 425; EPFP 3 (1923) 571. –Type:BS 34677 Ramos & Pascasio.MINDANAO: Surigao. Forests, altitude 400-700m.Gynocthodes nigra (Merr.) Merr., EPFP 3 (1923) 571; --Tetralopha nigra Merr., PJS 8 c (1913) Bot.58. –Type: BS 15877 Fenix (L*, US*, iso). Mindanao: Agusan del Norte prov., near Butuan, 30 Aug1912.PANAY, MINDANAO: Agusan del Norte. Lowland forests along streams.Gynocthodes philippinensis (Elmer) Merr., EPFP 3 (1923) 571; --Tetralopha philippinensis Elmer, LPB 3 (1911) 10Tetralopha polillensis C Robinson, PJS 6 c (1911) Bot. 226. –Type:LUZON: Apayao, Rizal, Quezon, Camarines, POLILLO, SIBUYAN, SIARGAO, BUCASGRANDE. Lowland forests.Hedyotis LinnaeusHedyotis atropurpurea Merr., PJS 17 (1920, publ. 1921) 426; EPFP 3 (1923) 495. –Type: BS 28578Ramos & Edaño.LUZON: Quezon (Mt Binuang). Forests, altitude c. 1000m.Hedyotis bambusetorum Merr., PJS 20 (1922) 469; EPFP 3 (1923) 495. –Type: Merrill 9214.PALAWAN. Low altitude bamboo thickets along trails.Hedyotis brachyantha Merr., PJS 17 (1920, publ. 1921) 427; EPFP 3 (1923) 496. –Type: BS 28769Ramos & Edaño.LUZON: Quezon (Mt Binuang). Forested slopes, altitude c. 200m.Hedyotis cagayanensis Merr., PJS 5 c (1910) Bot. 231; EPFP 3 (1923) 496. –Type: BS 7502Ramos.LUZON: Apayao, Cagayan. Low altitude forests.Hedyotis camarinensis Merr., PJS 20 (1922) 468; EPFP 3 (1923) 496. –Type: BS BS 33604 Ramos& EdañoLUZON: Camarines Norte (Paracale). Lowland clearings near thickets.Hedyotis capitellata Wall. ex G Don, Gen. Hist. 3 (1834) 527.Oldenlandia capitellata (Wall. ex G Don) Kuntze, Revis. Gen. Pl. 1 (1891) 292.var. pubescens Kurz, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 46(2) (1877) 135.Hedyotis kingiana Elmer, LPB 4 (1912) 1334; --Merr., EPFP 3 (1923) 498. –Type: Merrill 796.Hedyotis prainiana Elmer, LPB 1 (1906) 17, non King.Burma, Philippines.PALAWAN. Low altitudes, in thickets and among coarse grasses.Hedyotis cardiophylla Quisumb. & Merr., PJS 37 (1928) 202. –Type: BS 45585 Ramos & Edaño

LUZON: Quirino (Mt Alzapan).Hedyotis catanduanensis Merr., PJS 17 (1920, publ. 1921) 428; EPFP 3 (1923) 496. –Type: BS30443 Ramos.CATANDUANES. Low altitude primary forests.Hedyotis caudata Merr., PJS 8c (1913) Bot. 33; EPFP 3 (1923) 496. –Type: Loher 6419 (not seen).Without locality data.LUZON: Probably Rizal. Possibly from medium altitude forest.Hedyotis diffusissima Merr., PJS 17 (1920, publ. 1921) 429, EPFP 3 (1923) 497. –Type: BS 28387FenixLUZON: Ilocos Norte, Apayao. Primary forests, altitude c. 1000m.Hedyotis edanoi Quisumb. & Merr., PJS 37 (1928) 203. –Type: BS 47327 Ramos & Edaño.LUZON: Isabela (Mt Moises).Hedyotis eucapitata Merr., PJS 2 c (1907) Bot. 303; EPFP 3 (1923) 497. –Type: Merrill 5726.MINDORO: Mindoro Oriental (Mt Halcon). Mossy forests, altitude 1500-1800m.Hedyotis humilis Merr., PJS 8 c (1913) Bot. 33; EPFP 3 (1923) 497. –Type: BS 14691 Ramos (notseen). Camiguin: Camiguin prov., Mt Mahinog, in mossy forests, Apr 1912.CAMIGUIN, MINDANAO: Agusan del Norte (Mt Urdaneta =Hilong-hilong). Mossy forestsabove 1000m.Hedyotis laxiflora Merr., PJS 17 (1920, publ. 1921) 430., EPFP 3 (1923) 498, non Walp. 1852. –Type: BS 30313 Ramos.CATANDUANES. Forested ravines, altitude c. 400m.Hedyotis longipedunculata Merr., PJS 17 (1920, publ. 1921) 430; EPFP 3 (1923) 498. –Type: BS30551 Ramos.CATANDUANES. Low altitude forests along streams.Hedyotis luzoniensis Merr., PJS 12 c (1917) Bot. 160; EPFP 3 (1923) 498. –Type: BS 26580Ramos & Edaño (US*, iso). Luzon:LUZON: Mountain Province, Aurora (Mt Dingalan). Forests, altitude at least 300m.Hedyotis macgregorii Merr., PJS 5 c (1910) Bot. 160; EPFP 3 (1923) 498. –Type: BS 8490McGregor (US*, iso). Luzon: Benguet prov., Pauai, Jun 1909.Luzon: Benguet. Mossy forest, altitude c. 2100m.Hedyotis mindorensis Quisumb., PJS 41 (1930) 361, t. 25. --Type: BS 46363 Ramos (PNH, holo,possibly lost; iso in A*). Mindoro: Mindoro Oriental prov., Puerto Galera, forested slopes, altitude700m, Nov-Dec 1925.Mindoro: Mindoro Oriental (Puerto Galera). Known only from the type collection.Hedyotis montana Merr., PJS 2 c (1907) Bot. 304; EPFP 3 (1923) 498. –Type: Merrill 5782.MINDORO: Mindoro Oriental (Mt Halcon). Open heaths, altitude c. 2400m.Hedyotis perhispida Elmer, LPB 4 (1912) 1333; --Merr., EPFP 3 (1923) 498; --Lasianthus hispidusElmer, LPB 1 (1906) 10, non H. hispida Retz. –Type: Merrill 767 (NY*, US*, iso). Palawan:Palawanprov., Ewiig (=Iwahig) River, Feb 1903.

Exallage perhispida (Elmer) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Tweede Sect. 48(2)(1952) 142.PALAWAN (Iwahig River; Thumb Peak). Lowland streamside thickets or gravelly creeks,up to c. 75m.Hedyotis radicans (DC) Miq., Fl. Ind. Bat. 2 (1857) 181; --Merr., EPFP 3 (1923) 499. –Type: Haenkesn (not seen).BATANES, LUZON: Cagayan, Camarines, Sorsogon, PANAY, NEGROS, MINDANAO:Agusan del Norte, Surigao del Norte. Low altitude forests.Hedyotis scaberrima Merr., PJS 17 (1920, publ. 1921) 432; EPFP 3 (1923) 500. –Type: BS 30940Ramos & Edaño (A*, US*, iso). Panay: Capiz prov., Jamindan, thickets along streams, 27 May1918.PANAY: Capiz (Jamindan). Low altitude thickets along streams.Hedyotis sibuyanensis Elmer, LPB 3(1911) 977; --Merr., EPFP 3 (1923) 500. –Type: Elmer 12311(PNH, holo, lost; iso GH*, L*, MO*, NY*, US*, iso). Sibuyan: Mt Giting-giting, Magallanes(=Magdiwang), clayey soil of a forested ridge, altitude 1500ft, Apr 1910.SIBUYAN. Ridge forest, altitude c. 450m.Hedyotis simplex Merr., PJS 17 (1920, publ. 1921) 433; EPFP 3 (1923) 500. –Type: BS 30389Ramos (US*, iso). Catanduanes: in forests, Nov 1917.CATANDUANES, BILIRAN (Mt Suiro), MINDANAO: Zamboanga. Primary forests,altitude 300-600m.Hedyotis subevenosa Merr., PJS 17 (1920, publ. 1921) 432; EPFP 3 (1923) 500. –Type: Weber1563 (A*, US*, iso). Luzon: Cagayan prov., Abulug River, Jan 1912.LUZON: Cagayan. Low and medium altitude forests, said to be common at altitude 180-250m.Hedyotis subvelutina Elmer, LPB 3(1911) 978; --Merr., EPFP 3 (1923) 500. –Type: Elmer 12441SIBUYAN. Streamside thickets, altitude c. 225m.Hedyotis trisecta Elmer, LPB 9 (1934) 3249. --Type: Elmer 14885 (PNH, holo, lost; MO*, NY*, US*,iso). Luzon: Sorsogon prov., Mt Bulusan (Irosin).Unplaced:Hedyotis bartlingii Merr., PJS 5 c (1910) Bot. 388; EPFP 3 (1923) 495. –Type: Haenke snLUZON: Mountain Province, Benguet, Bulacan, Quezon, MINDORO, PANAY. Openplaces at low and medium altitudes, ascending to 1300m.Hedyotis microphylla Merr., PJS 1 (1906) suppl. 239; EPFP 3 (1923) 498, non Kunth (1820). –Merrill 4693LUZON: Ifugao, Mountain Province, Benguet. Mossy forests, altitude 2000-2600m.Hedyotis oligantha Merr., PJS 17 (1920, publ. 1921) 431; EPFP 3 (1923) 498. –Type: BS 29425Ramos & Edaño.LUZON: Ilocos Norte, Bataan, Pampanga, Rizal. Shaded banks in forests, altitude 300-900m.Hedyotis phanerophlebia Merr., PJS 8 c (1913) Bot. 34; EPFP 3 (1923) 499. --Type: FB 4552Mearns & Hutchinson (US*, iso). Mindanao: Misamis Occidental prov., Mt Malindang, in forests,

altitude c. 1800m, May 1906.MINDANAO: Misamis Occidental (Mt Malindang). Forest, altitude c. 1700m.Hedyotis pilosissima Merr., PJS 5 c (1910) Bot. 232; EPFP 3 (1923) 499. –Type: Merrill 6704 (notseen). Panay: Capiz prov., Dumarao. Damp shaded ravines along streams, altitude c. 100m, 25 Mar 1910.LUZON: Bulacan, Laguna, Quezon, Camarines Sur (Maagnas), Albay, CATANDUANES,PANAY, SAMAR, MINDANAO: Agusan, Zurriago del Norte. Lowland forests and dampravines.Hedyotis pinatubensis Elmer, LPB 10 (1939) 3778. --Type: Elmer 22194 (MO*, NY*, iso). Luzon:Pampanga prov., Mt Pinatubo (Camp Stotsenburg). May 1927.Hedyotis whiteheadii Merr., PJS 2 c (1907) Bot. 303; EPFP 3 (1923) 501. –Type: Merrill 5733 (US*,iso). Mindoro: Mindoro Oriental prov., Mt HalconMINDORO: Mindoro Oriental (Mt Halcon). Dense mossy forests on exposed ridges,altitude c. 2400m.Hydnophytum JackLiterature:Huxley, CR & MH Jebb. 1991. The tuberous epiphytes of the Rubiaceae 1: A new subtribe- TheHydnophytinae. Blumea 36: 1–20.Huxley, CR. 1993. The tuberous epiphytes of the Rubiaceae. 6: a taxonomic history of theHydnophytinae. Blumea 37.Hydnophytum angustifolium Merr., PJS 3 c (1908) Bot. 162; EPFP 3 (1923) 568.MINDANAO: Zamboanga, Lanao del Sur. Epiphytic in forests, altitude 200-600m.Hydnophytum brachycladum Merr., PJS 10 c (1915) Bot.142; EPFP 3 (1923) 568. Type: BS 924Mangubat.MINDORO. Epiphytic in lowland forests.Hydnophytum formicarum Jack, Trans. Linn. Soc. 14 (1823) 124; --Merr., EPFP 3 (1923) 568.Hydnopytum orbiculatum Elmer, LPB 5 (1913) 1860; --Merr., EPFP 3 (1923) 569. –Type: Elmer 14046Indochina, Sumatra, Peninsular Malaysia, Borneo, Philippines.LUZON: Quezon, POLILLO, MINDANAO: Agusan del Norte. Mangrove to ridge forestsc. 600m.Hydnophytum intermedium Elmer, LPB 3 (1911) 1039; --Merr., EPFP 3 (1923) 569. --Type: Elmer10782.MINDANAO: Davao del Sur (Mt Apo). Epiphytic in damp forests, altitude c. 1100m.Hydnophytum leytense Merr., PJS 8 c (1913) Bot. 390; EPFP 3 (1923) 569. –Type: Wenzel 45LUZON: Laguna, Sorsogon, LEYTE. Epiphytic in primary forests,altitude 200 to 400m.Hydnophytum membranaceum Merr., PJS 10 c (1915) Bot.143; EPFP 3 (1923) 569. –Type: FB3720 MerrittMINDORO, DINAGAT. On trees at low altitudes, the type from the mangrove swamps.Hydnophytum mindanaense Elmer, LPB 3 (1911) 1039; --Merr., EPFP 3 (1923) 569. –Type:Elmer 10874

MINDANAO: Davao, Cotabato. Epiphytic in lowland forests, ascending to 450m.Hydnophytum mindorense Merr., PJS 10 c (1915) Bot.143; EPFP 3 (1923) 569. –Type: Merrill6182LUZON: Quezon, MINDORO. Epiphytic in primary lowland forests.Hydnophytum nitidum Merr., PJS 2 c (1907) Bot. 307; EPFP 3 (1923) 569; Elmer, LPB 3 (1911)1039. –Type: Merrill 6181LUZON: Bataan, MINDORO. Epiphytic in primary forests, altitude 900-1400m.Hydnophytum philippinense Becc., Malesia 2 (1885) 149, t. 33, Fig.14-19; --Rolfe, J. Bot. 23(1885) 213; --Vidal, Rev. Pl. Vasc. Filip. (1886) 158; --Elmer, LPB 1 (1906) 7; --Merr., EPFP 3(1923) 569. –Type: RS Williams 2178.LUZON: Camarines, MINDANAO: Zamboanga, Davao, Agusan del Norte, Surigao delNorte. Epiphytic in lowland forests, sometimes in mangroves.Hymenodictyon WallichHymenodictyon orixense (Roxb.) Mabb., Taxon 31 (1982) 66.Hymenodictyon excelsum (Roxb.) Wall. in Roxb. Fl. Ind. 2 (1824) 149; -Merr., EPFP 3 (1923) 506.Sri Lanka, W Himalayas, India, E Himalayas, Nepal, Bangladesh, Burma, Thailand,Laos, Cambodia, Vietnam, Peninsular Malaysia, Java, Philippines.LUZON: Abra, Ilocos Sur, Pangasinan, Nueva Ecija, Tarlac, Pampanga, Bulacan, Rizal.PALAWAN, GUIMARAS, MINDANAO: Davao; BASILAN. Deciduous lowland forest,frequently on limestone, often about cliffs near the sea, rarely apparently in lowlandevergreen forest.Hypobathrum BlumeHypobathrum coriaceum Merr., PJS 29 (1926) 490. –Type: BS 42615 (UC*, holo; iso: A*, US*).Bohol: Bohol prov., Bilar. On forested slopes, altitude c. 600m, Sept 1923.BOHOL. Medium altitude forest, altitude c. 600m.Hypobathrum longifolium (DC) KM Wong in F Ng (ed.), Tree Fl. Malaya 4 (1989) 354; --Higginsia longifolia Blume ex DC., Prodr. 4 (1830) 399; --Petunga longifolia DC, Prodr. 4 (1830)399; --Merr., EPFP 3 (1923) 533.Peninsular Thailand, Sumatra, Peninsular Malaysia, Java, Borneo, S Philippines,Sulawesi.MINDANAO: Agusan del Norte. Lowland secondary forests.Hypobathrum multibracteatum Elmer, LPB 9 (1934) 3250. –Syntypes, : Elmer 15925 (NY*) May1916; Elmer 15674 (NY*). Luzon: Sorsogon prov., Irosin, Mt BulusanLUZON:Hypobathrum purpureum (Elmer) Merr., EPFP 3 (1923) 534; --Tricalysia purpurea Elmer, LPB 1(1906) 70. –Type:LUZON, ALABAT, SAMAR, LEYTE. In primary forests at low and medium altitudes.Hypobathrum purpuricarpum (Elmer) Ridsd., ined.Randia purpuricarpa Elmer, LPB 9 (1934) 3267.

Involucrella (Hooker filius) ined.Involucrella coronaria (Kurz) ined.Hedyotis coronaria (Kurz) Craib, Fl. Siam. 2 (1932) 38; --Scleromitrion coronarium Kurz, J. Asiat. Soc.Bengal, Pt. 2, Nat. Hist. 46(2) (1877) 136.Oldenlandia connata (Hook.f.) K Schum. in HGA Engler & KAE Prantl, Nat. Pflanzenfam. 4(4) (1891) 25;--Hedyotis connata Wall., Cat. (1828) No. 856, nom. nud., --Hook.f., Fl. Brit. Ind. 3 (1880) 62,sphalm. As ‘coronata’; --Merr., EPFP 3 (1923) 496.Assam, Burma, China (Yunnan), Peninsular Malaysia, Philippines.LUZON: Pangasinan, Nueva Ecija, Bataan, Rizal, Laguna, MINDANAO: Davao. Lowlandforests along streams.Ixora LinnaeusLiterature:Bremekamp, CEB. 1937. The Malaysian species of the genus Ixora (Rubiaceae). Bulletin duJardin Botanique du Buitenzorg, ser. 3, 14: 197–367.Bremekamp, CEB. 1940. The Malaysian species of the genus Ixora (Rubiaceae). Additions andemendations. Bulletin du Jardin Botanique du Buitenzorg, ser. 3, 14: 276–299.Ixora amplexifolia K Schum. & Lauterb., Fl. Deutsch. Sudsee, (1901) 572; --Bremek., Bull. Jard.Bot. Buitenz. Ser. 3, 14 (1937) 347. –Type from New Guinea.Ixora auriculata Elmer, LPB 3 (1911) 1018; --Merr., EPFP 3 (1923) 546. --Type: Elmer 12483 (L*, NY*, US*,iso). Sibuyan: Magallanes (=Magdiwang), Mt Giting-giting. Densely wooded flats of woodlandstreams, 750-1000 ft., May 1910.Ixora inaequifolia C Robinson, PJS 6 c (1911) Bot. 222; --Merr., EPFP 3 (1923) 548.Philippines, New Guinea.LUZON: Cagayan, Isabela, Nueva Ecija, Rizal, Camarines, POLILLO, SIBUYAN. Inprimary forests at low and medium altitudes.Ixora angustilimba Merr., PJS 17 (1920, publ. 1921) 320; EPFP 3 (1923) 546. --Type: BS 34588Ramos & Pascasio (US*, iso). Mindanao: Surigao del Norte prov.BUCAS GRANDE, DINAGAT, MINDANAO: Surigao del Norte. On dry forested slopesand ridges at low altitudes.Ixora bartlingii Elmer, LPB 3 (1911) 1020; --Merr., EPFP 3 (1923) 546; --Bremek., Bull. Jard.Bot. Buitenz. Ser. 3, 14 (1937) 299. –Type: Elmer 12323 (PNH, holo, lost; A*, BISH*, MO*, NY*,iso). Sibuyan: Magallanes (=Magdiwang), Mt Giting-giting, wet alluvial soil of dense forest near thePauala River at 1000ft, Apr 1910.Philippines, N Sulawesi.LUZON: Isabela, Zambales, Bataan, Batangas, Cavite, Laguna, Rizal, Quezon,Camarines, MINDORO; SIBUYAN; MASBATE; BILIRAN; MINDANAO: Cotabato. Inforests at low and medium altitudes.Ixora bibracteata Elmer, LPB 1 (1906) 8; --Merr., EPFP 3 (1923) 546. –Type: HN Whitford 789 (notseen). Luzon: Quezon prov., Tinnan River, Sept 1904.LUZON: Ilocos Norte, Quezon, Nueva Ecija, Rizal. In primary forests, ascending to400m.Ixora capitulifera Merr., PJS 5 c (1910) Bot. 233; EPFP 3 (1923) 546. --Type: BS 686 Foxworthy(NY*, US*, iso). Palawan: Mt Victoria, in forests along streams, altitude c. 1050m, 23 Mar 1906.PALAWAN. Along streams in forests, c.1000m.

Ixora chartacea Elmer, LPB 5 (1913) 1861; --Merr., EPFP 3 (1923) 547. --Type: Elmer 13806 (BISH*,L*, MO*, NY*). Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta (=Mt Hilong-hilong),in dry, humus-covered soil of forested ridges at 3250ft, in the depression of Duros and KawilananPeak, Sept 1912var. chartaceaMINDANAO: Agusan del Norte. In forests, c.1000m.var. membranacea Elmer, LPB 5 (1913) 1862; --Merr., EPFP 3 (1923) 547. –Type: Elmer 13767LUZON: Sorsogon, MINDANAO: Agusan del Norte, SIARGAO. Medium altitude primaryforests, ascending to 900m.Ixora confertiflora Merr., PJS 17 (1920, publ. 1921) 321; EPFP 3 (1923) 547. --Type: BS 35206Ramos & Pascasio (NY*, US*). Dinagat, May 1919.DINAGAT. Low altitude primary forests.Ixora crassifolia Merr., PJS 5 c (1910) Bot. 233; EPFP 3 (1923) 547. –Type: FB 9439 Whitford &Hutchinson (US*, iso). Mindanao: Zamboanga del Sur prov., Port Banga, in dipterocarp forest,altitude 30-50m, Feb 1908.BASILAN, MINDANAO: Zamboanga, Lanao. Low altitude primary forests.Ixora cumingiana Vidal, Phan. Cuming. Philipp.(1885) 183; --Merr., EPFP 3 (1923) 547. –Type:Cuming 1233 (L*.MO*, syntypes). Luzon.N LUZON to PALAWAN and MINDANAO, in most all islands and provinces. Low altitudeprimary forests.Ixora ebracteolata Merr., PJS 5 c (1910) Bot. 234; EPFP 3 (1923) 548. –Type: FB 5845 Curran(NY*, US*). Luzon: Zambales prov., Mahumaling, on dry cogonales (Imperata grassland) slopes,Jan 1907LUZON: Cagayan, Isabela, Pangasinan, Zambales, Quezon, Sorsogon, SIBUYAN. Lowaltitude forests.Ixora filipes Valeton, Icon. Bog. (1913) t. 359; --Bremek., Bull. Jard. Bot. Buitenz. Ser. 3, 14 (1937) 307.Ixora gracilipes Merr., PJS 10 c (1915) Bot. 119., EPFP 3 (1923) 548, nom. illeg. –Type: BS 20723 Escritor(not seen). Luzon: Quezon prov., Mt Cadig, near Guinayangan, Mar 1913.Philippines, Talaud, Sulawesi.LUZON: Quezon (Mt Cadig), Camarines, SAMAR. Lowland forests.Ixora fulgens Roxb., Hort. Beng. (1814) 10, nom. nud.; Fl. Ind. Ed. Carey I (1820) 378; --Merr., EPFP3 (1923) 548.Peninsular Malaysia and archipelago.LEYTE; MINDANAO: Zamboanga, Misamis, Lanao del Sur, JOLO. Low altitude primaryforests.Ixora gigantifolia Elmer, LPB 5 (1913) 1864; --Merr., EPFP 3 (1923) 548. –Type: Elmer 14206hilong), Oct 1912.Ixora platyphylla Merr., PJS 10 c (1915) Bot. 120. --Type: FB 18250 Miranda (not seen). Mindanao: SouthCotabato prov., Glan, Jun 1912.MINDANAO: South Cotabato, Agusan del Norte, SIARGAO. Low altitude forests toc.1000m.(BISH*, L*, NY*, US*, iIxora ilocana Merr., PJS 12 c (1917) Bot. 168; EPFP 3 (1923) 548. --Type: BS 27325 Ramos (US*,iso). Luzon:LUZON: Ilocos Norte. In forests along streams at low altitudes.

Ixora intermedia Elmer, LPB 4 (1912) 1336; --Merr., EPFP 3 (1923) 549. Type: Elmer 12957 (BISH*,L*, MO*, NY*, US*, iso). Palawan: Palawan prov., Puerto Princesa, Mt Pulgar [=Thumb Peak],‘stony soil of dense woods along a dry stream bed at 50ft’, Apr 1911.PALAWAN. In forests at low altitudes.Ixora irosinensis Elmer, LPB 9 (1934) 3251.LUZON:Ixora javanica (Blume) DC, Prodr. 4 (1830) 487; --Bremek., Bull. Jard. Bot. Buitenz. Ser. 3, 14 (1937)253; --Corner, Wayside Trees 1 (1940) 206; --KM Wong in FSP Ng (ed). Tree Fl. Malaya 4 (1989)360.Ixora magnifica Elmer, LPB 5 (1913) 1865; --Merr., EPFP 3 (1923) 550. –Type: Elmer 13424 (BISH*,L*, MO*, NY*, US*, iso). Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta (=Hilonghilong),very moist fertile soil of dense wood or forests from 500-1500ft altitude, Aug 1912.var. javanicaIndia, Tenasserim, Indochina, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines.PALAWAN, MINDANAO: Agusan del Norte, Davao. Lowland primary forests, ascendingto 500m.Ixora leucocarpa Elmer, LPB 4 (1912) 1337; --Merr., EPFP 3 (1923) 549. –Type: Elmer 12894(BISH*, L*, NY*, US*, iso). Palawan: Puerto Princesa, ‘in fertile adobe or otherwise fertile soil of lowlightly flats of woodland, or in humid forest at a somewhat higher elevation 250 to 750ft altitude’,Mar 1911.PALAWAN. Primary forests ascending to 300m altitude.Ixora leytensis Elmer, LPB 1 (1906) 69; --Merr., EPFP 3 (1923) 549. --Type: Elmer 7044 (not seen).LEYTE, SAMAR. Lowland forests.Ixora longissima Merr. PJS 5 c (1910) Bot. 235; EPFP 3 (1923) 549. –Type: FB 16975 Rosenbluth(not seen). Leyte: without definite locality, Mar 1909.LEYTE, BILIRAN, SAMAR. In primary forests ascending to 900m.Ixora longistipula Merr., PJS 5 c (1910) Bot. 236; EPFP 3 (1923) 549. –Type: FB 13625 Curran &Foxworthy (US*, iso). Negros: Negros Occidental prov., Mt Marapara, Sept 1909MINDORO: Mindoro Oriental (Mt Halcon), PANAY, NEGROS: Negros Occidental. Lowand medium altitude primary forestsIxora luzoniensis Merr., PJS 17 (1920, publ. 1921) 434; EPFP 3 (1923) 549. –Type: BS 29956 Fenix(US*, iso). Luzon: Pangasinan prov., San Isidro Labrador, in forests, altitude 350-400m, 4 Nov 1917.LUZON: Ilocos Norte, Pangasinan, Cagayan (Pamplona), Zambales. Lowland forests.Ixora macgregorii C Robinson, PJS 6 c (1911) Bot. 223; --Merr., EPFP 3 (1923) 549. --Type: BS10219 Mcgregor (L*, NY*, US*, iso). Polillo. Oct-Nov 1909.POLILLO. Lowland forests.Ixora macrophylla Bartl., in DC, Prodr. 4 (1830) 487; --Merr., EPFP 3 (1923) 549. –Type:Ixora arborea Blanco, Fl. Filip. (1837) 61; ed. 2 (1845) 45; ed. 3, 1 (1877) 78, non Roxb. –Type: non extant;Illustrative specimen: Merrill Sp. Blancoanae 734 (US*). Luzon: Batangas prov., Mt Batulao, inforest, 25 Jul 1914.Philippines, Sulawesi.BATAN and BABUYAN ISLS to MINDANAO and BASILAN, in most or all islands andprovinces. Lowland forests.

Ixora mearnsii Merr., PJS 5 c (1910) Bot. 235; EPFP 3 (1923) 549. –Type: BS 2999 Mearns (notseen). Luzon: Aurora prov., Casiguran, 1 Jun 1907.LUZON: Aurora (Casiguran; Baler). Lowland forests.Ixora mindanaensis Merr., PJS 5 c (1910) Bot. 237; EPFP 3 (1923) 550. –Type: FB 9010 (Whitford &Hutchinson (not seen). Mindanao: Zamboanga del Sur prov., Port Banga, 29 Nov 1907.MINDANAO: Zamboanga del Sur, Misamis Occidental (Malabug River, trail to MtMalindang), Lanao del Sur (Lake Lanao, Camp Keithley). Low and medium altitudeforests.Ixora myriantha Merr., PJS 20 (1922) 461; EPFP 3 (1923) 550. –Type: Merrill Philip. Pl. 1700 (L*,MO*, NY*, US*, iso). Samar: no further locality data. Apr 1914.SAMAR. In damp forests at low altitudes. Possibly not distinct from I. cumingiana Vidal,differing mainly in the crowded shorter flowers.Ixora oblongifolia Elmer, LPB 5 (1913) 1867; --Merr., EPFP 3 (1923) 550. –Type: Elmer 13775hilong), humus-covered soil of steep densely wooded slopes at 3000ft along Dalahion River, Sept1912LUZON: Camarines, MINDANAO: Agusan del Norte, Surigao del Norte. In forests up toan altitude of 900m.(BISH*, L*, MO*, NY*,Ixora palawanensis Merr., PJS 5 c (1910) Bot. 238; EPFP 3 (1923) 550. –Type: BS 793 Foxworthy(US*, iso). Palawan: Forested ravines c. 1.5 miles NW of Iwahig, 22 Apr 1906.CULION, PALAWAN. Lowland forests.Ixora philippinensis Merr., PJS 5 c (1910) Bot. 238, Fl. Manila (1912) 452; EPFP 3 (1923) 550; --Bremek., Bull. Jard. Bot. Buitenz. Ser. 3, 14 (1937) 343. –Type: FB 2299 Meyer (NY*, US*, iso).Luzon: Bataan prov., Mt Mariveles, Lamao River, Dec 1904.Ixora philippinensis Merr. var. brevituba Merr., PJS 5 c (1910) Bot. 240; EPFP 3 (1923) 551. --Type: FB15527 Merritt & Darling (not seen). Luzon: Ilocos Norte prov., Pasuquin, on limestone formation,hillsides, altitude c. 100m, 3 Nov 1908.Ixora filmeri Elmer, LPB 4 (1912) 1338. –Type: Elmer 12719 (BISH*, NA*, NY*, US*, iso). Palawan: BrookesPoint, Addison Peak, Mar 1911.Ixora littoralis Merr., PJS 5 c (1910) Bot. 240; EPFP 3 (1923) 549. --Type: BS 1274 McGregor (not seen).Bohol: Tagbilaran, on beach cliff, Jul 1906Lutao (off Taiwan), Philippines, Banguey Is (off Sabah), N Sulawesi, Moluccas (SulaIsls).BATAN and BABUYAN Islands and N LUZON to PALAWAN, BALABAC, BOHOL,MINDANAO and BASILAN in most or all islands and provinces. Common in thicketsand secondary forests at low altitudes, including coastal bluffs and inland limestoneformation.Ixora pilosa Merr., PJS 8 c (1913) Bot. 39; EPFP 3 (1923) 551. –Type: BS 13371 Ramos (not seen).Luzon: Quezon prov., Tagcawayan, Mar 1911.LUZON: Quezon. Lowland forests.Ixora propinqua Merr., PJS 8 c (1913) Bot. 39; EPFP 3 (1923) 551. --Type: Merrill 8254 (not seen).Mindanao: Zamboanga, Sax River Mtns back of San Ramon, in damp forest, altitude c. 800m, Nov1911.MINDANAO: Zamboanga. In primary forests, c. 800m.Ixora salicifolia (Blume) DC, Prodr. 4 (1830) 487; --Merr., EPFP 3 (1923) 551; --Bremek., Bull. Jard.Bot. Buitenz. Ser. 3, 14 (1937) 240.

Java, Borneo, Philippines.LUZON, PALAWAN, MASBATE, PANAY, BILIRAN, LEYTE, BUCAS GRANDE,SIARGAO, DINAGAT, MINDANAO. In primary forests at low altitudes.Ixora samarensis Merr., PJS 10 c (1915) Bot. 121; EPFP 3 (1923) 551. –Type: BS 15399 Ramos(US*, iso). Samar: Western Samar prov., Catbalogan, Jul 1912.SAMAR. Lowland primary forests.Ixora sparsiflora Elmer, LPB 1 (1906) 8; --Merr., EPFP 3 (1923) 551. --Type: Elmer 6641 (NY*, iso).Luzon: Bataan prov., Mt Mariveles, Nov 1904.LUZON: Bataan.In primary forests at low or medium altitudes.Ixora tenuipedunculata Merr., PJS 17 (1920, publ. 1921) 321; EPFP 3 (1923) 551. --Type: BS34475 Ramos & Pascasio (Iso in A*, US*). Mindanao: Surigao prov., no definite locality cited, damp forest atlow altitudes, 30 Apr 1919.MINDANAO: Surigao. In primary forests at low altitudes.Cultivated:Ixora chinensis Lam., Encycl. 3 (1879) 344; --Merr., EPFP 3 (1923) 547; Fl. Manila (1912) 452,India to S China, widely cultivated in the tropics for ornamental purposes.Ixora coccinea L.. Sp. Pl. (1753) 110; --Merr., EPFP 3 (1923) 547; Fl. Manila (1912) 452.Native to India. Cultivated throughout the tropics for ornamental purposes.Ixora finlaysoniana Wall., Cat. (1832) No. 6166 nom. nud.; --G Don Gen. Syst. 3 (1832) 512; --Merr.Fl. Manila (1912) 451; EPFP 3 (1923) 547.Commonly cultivated in Manila and other large towns, but apparently never producingfruits and nowhere spontaneous.Ixora rosea Wall. in Roxb. Fl. Ind. 1 (1820) 398;Cultivated in the larger towns for its pink flowers. Synonymized under Ixora stricta Roxb.(= I. chinensis Lam.) but Merrill (1923) consider it to be distinct.Knoxia LinnaeusLiterature:Bhattacharjee, R & DB Deb. 1985. A revision of Knoxia (Rubiaceae). J. Econ. Tax. Bot. 6: 73–95.Puff, C & E Robbrecht. 1989. A survey of the Knoxieae (Rubiaceae-Antirheoideae). Bot. Jahrb.Syst. 110: 511–588.Knoxia sumatrensis (Retz.) DC, Prod. 4 (1830) 569.var. sumatrensisKnoxia corymbosa Willd., Sp. Pl. 1 (1797) 582; --Merr., EPFP 3 (1923) 535.Tropical Asia to Australia.LUZON, CULION, PALAWAN, MINDANAO. In open places, on banks, etc., sometimesat sea level, more commonly at medium altitudes, ascending to 1500m.Lasianthus JackLiterature:Igersheim & E. Robbrecht. 1993. Opera Bot. Belg. 6: 77.

Lasianthus acuminatissimus Merr., PJS 20 (1922) 467; --Type: BS 38058 Ramos & EdañoS China and the Philippines: LUZON. In mossy forests, c. 1800m.Lasianthus attenuatus Jack, Trans. Linn. Soc. 14 (1823) 126.Lasianthus bordenii Elmer, LPB 1 (1906) 11; --Merr. EPFP 3 (1923) 565. –Type: Elmer 6997 (NY*, iso). Luzon: Bataan prov., MLasianthus copelandii Elmer, LPB 1 (1906) 10; --Merr. EPFP 3 (1923) 566. –Type: EB Copeland sn (notseen). Negros: Gimogon River, in forests, Jan 1905.Lasianthus wallichii Wight, Calcutta J. Nat. Hist. 6 (1846) 503; --HJ Li & HY Liu, Fl. Taiwan ed.2, 4 (1998)295, pl. 133, photo 180.Trop. & Subtrop. Asia: Assam, E Himalayas, Nepal, Tibet, C & S China, Hainan, Taiwan,Ryukyu Isls, Burma, Thailand, Laos, Cambodia, Vietnam, Sumatra, Peninsular Malaysia,Java, Lesser Sunda Isls, Borneo, Philippines, Sulawesi, Moluccas, New Guinea.LUZON, MINDORO, POLILLO, CATANDUANES, BILIRAN, LEYTE, NEGROS, PANAY,MINDANAO. In primary forests at low altitudes to 900mLasianthus biflorus (Blume) M Gangop. & Chakrab., J. Econ. Taxon. Bot. 16 (1992) 338; --Litosanthes biflora Blume, Cat. Gew. Buitenz. (1823) 21; --Merr. EPFP 3 (1923) 568.China (S Yunnan), Hainan, C & S Taiwan, Thailand, Sumatra, Peninsular Malaysia,Java, Lesser Sunda Isls, Borneo, Philippines, Sulawesi.LUZON, MINDORO, CATANDUANES, LEYTE, NEGROS, MINDANAO. In damp primaryforests, chiefly at medium altitudes, ascending to 1400m.Lasianthus chinensis (Champ.) Benth., Fl. Hongk. (1861) 160; --Nonatelia chinensis (Champ.)Kuntze, Revis. Gen. Pl. 1 (1891) 291; --Mephitidia chinensis Champ., Hooker's J. Bot. Kew Gard.Misc. 4 (1852) 196.S & C China, Hainan, Taiwan, Thailand, Laos, Cambodia, Vietnam, PeninsularMalaysia, Philippines.PALAWAN.Lasianthus clementis Merr., PJS 3 c (1908) Bot.164; --Merr. EPFP 3 (1923) 565.Lasianthus bulusanensis Elmer, LPB 9 (1934) 3252.Philippines, Sulawesi.LUZON: Sorsogon, LEYTE, MINDANAO. In primary forests, ascending to 700m.Lasianthus cyaneus (Elmer) Merr., EPFP 3 (1923) 566.LUZON, MINDANAO. In primary forests, 800-1300m.Lasianthus cyanocarpus Jack, Trans. Linn. Soc. London 14 (1825) 125; --Chao, Fl. Taiwan 4(1978) 289; --HJ Li & HY Liu, Fl. Taiwan ed. 2, 4 (1998) 280, pl. 125; --Merr. EPFP 3 (1923) 565.Nonatelia inaequalis (Blume) Kuntze, Revis. Gen. Pl. 1 (1891) 291; --Mephitidia inaequalis (Blume) DC,Prodr. 4 (1830) 453; --Lasianthus inaequalis Blume, Bijdr. (1826) 996; --Type: Merrill 863 (notseen). Palawan.India, Andaman Isls, Nicobar Isls, Burma, Thailand, Vietnam, S China, Taiwan, RyukyuIsls, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, Moluccas, New Guinea.LUZON: Cagayan, Rizal, Laguna, Quezon, Camarines, CATANDUANES, BUSUANGA,PALAWAN, BILIRAN, NEGROS, LEYTE, PANAY, MINDANAO: Bukidnon, Agusan. Indamp primary forests at low altitudes.Lasianthus fordii Hance, J. Bot. 23 (1885) 324; --Chao, Fl. Taiwan 4 (1978) 292; --HJ Li & HY Liu, Fl.Taiwan ed. 2, 4 (1998) 284, pl. 126, photo 178.Lasianthus submembranifolius Elmer, LPB 5 (1913) 1871; --Merr. EPFP 3 (1923) 567.Lasianthus zambalensis Elmer, LPB 9 (1934) 3209.Mephitidia tashiroi (Matsum.) Nakai, Trees Shrubs Japan (1922) 396; --Lasianthus tashiroi Matsum., Bot.Mag. Tokyo 15 (1901) 37; --Merr. EPFP 3 (1923) 567.

S Japan, Ryukyu Isls, S & C China, Hainan, Taiwan, Thailand, Cambodia, Vietnam,Java, Philippines, Sulawesi, New Guinea.LUZON (Zambales), NEGROS, MINDANAO. Chiefly in the mossy forests, altitude 1000-2000m.Lasianthus furcatus (Miq.) Bremek., Candollea 16 (1957) 94, in adnot.; --Canthium furcatum Miq.,Fl. Ned. Ind. 2 (1857) 252.Geniostoma acuminatissima Merr., PJS 20 (1922) 432; EPFP 3 (1923) 309.Java, Philippines?, New Guinea, Bismarck Archipelago.Lasianthus hirsutus (Roxb.) Merr., J. Arnold Arb. 33 (1952) 229; --Triosteum hirsutum Roxb., Fl.Ind. 1 (1820) 538.Lasianthus everettii Merr., PJS 3 c (1908) Bot. 265.Trop. & Subtrop. Asia: India, Bangladesh, Nicobar Isls, Burma, Thailand, Vietnam, SChina, Hainan, Taiwan, Ryukyu Isls, Sumatra, Peninsular Malaysia, Java, Borneo,Philippines, Moluccas, New GuineaLasianthus hispidulus (Drake) Pit. in H Lecomte, Fl. Indo-Chine 3 (1924) 393; --Mephitidiahispidula Drake, J. Bot. (Morot) 9 (1895) 239.Lasianthus bunzaensis Simizu, Trans. Nat. Hist. Soc. Formosa 34 (1944) 301; --Chao, Fl. Taiwan 4(1978) 286; --HJ Li & HY Liu, Fl. Taiwan ed. 2, 4 (1998) 277, pl. 122.S & China, Thailand, Vietnam, Hainan, Taiwan, Ryukyu Isls, Sumatra, Borneo,Philippines.Lasianthus lucidus Blume, Bijdr. (1826) 997.Nonatelia lucida (Blume) Kuntze, Revis. Gen. Pl. 1 (1891) 291; --Mephitidia lucida (Blume) DC., Prodr. 4(1830) 453.var. lucidusIndia (Assam), Burma, S China (Yunnan), Hainan, Vietnam, Thailand, PeniunsularMalaysia, Java, Lesser Sunda Isls, Philippines, Sulawesi, Moluccas, New Guinea.Lasianthus microphyllus Elmer, LPB 5 (1913) 1870; --Merr. EPFP 3 (1923) 566; --Chao, Fl.Taiwan 4 (1978) 298; --HJ Li & HY Liu, Fl. Taiwan ed.2, 4 (1998) 290, pl. 130.Taiwan. Malesia: Java, Philippines.LUZON, MINDORO, PANAY, LEYTE, NEGROS, MINDANAO. Chiefly in the mossyforest, 1000-2200m.Perhaps better treated as a variety of L. fordiiLasianthus rigidus Miq., Fl. Ind. Bat. 2 (1857) 321Lasianthus humilis Elmer, LPB 2 (1908) 519; --Merr. EPFP 3 (1923) 565. Type: Elmer 9678, Negros.China (S Yunnan), Assam, Sumatra, Java, Philippines.NEGROS. On forested ridges, c.1000m.Lasianthus sikkimensis Hook.f., Fl. Brit. Ind. 3 (1880 180.Lasianthus mindananensis Merr., PJS 20 (1922) 466; --Merr. EPFP 3 (1923) 567. --Type: BS 38795 Ramos& Edaño.Assam, E Himalayas, Bangladesh, C & S China, Hainan, Taiwan, Thailand, Vietnam,Philippines.MINDANAO, In damp forests, c. 1000m.Lasianthus stipularis Blume, Bijdr. (1826) 997; --Merr. EPFP 3 (1923) 567; --Mephitidia stipularis(Blume) DC, Prodr. 4 (1830) 453.

var. stipularisSumatra, Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi.MINDORO, LEYTE, MINDANAO. In damp primary forests at low altitudes.Lasianthus trichophlebus Hemsl. ex FB Forbes & Hemsl., J. Linn. Soc., Bot. 23 (1888) 388.var. trichophlebus.S China (Guangdong), Hainan, Thailand, Vietnam, Taiwan, Sumatra, PeninsularMalaysia, Java, Philippines, Sulawesi.var. latifolius (Miq.) H Zhu, Syst. Geogr. Pl. 72 (2002) 93; --Lasianthus sylvestris var. latifolius Miq.,Ann. Mus. Bot. Lugduno-Batavi 4 (1869) 248.Lasianthus tomentosus var. hiiranensis (Hayata) H Zhu, Acta Bot. Yunnan. 20 (1998) 156; --Lasianthushiiranensis Hayata, Icon. Pl. Formos. 9 (1920) 62; --Chao, Fl. Taiwan 4 (1978) 295; --HJ Li & HYLiu, Fl. Taiwan ed.2, 4 (1998) 287, pl. 128, photo 179.Hainan, SE Taiwan, Thailand, Vietnam, Sumatra, Peninsular Malaysia, Java,Philippines.Lasianthus verticillatus (Lour.) Merr., Trans. Am. Phil. Soc. n. s. 24, 2 (1935) 372.Lasianthus morus Elmer, LPB 1 (1906) 73; --Merr. EPFP 3 (1923) 567. –Type: Elmer 7050 (NY*, iso). Leyte:Leyte prov., Palo, wooded ravines along the Bangon River, Jan 1906.Lasianthus obliquinervis Merr., PJS 1 (1906) Suppl. 136; --Chao, Fl. Taiwan 4 (1978) 301; --HJ Li & HY Liu,Fl. Taiwan ed.2, 4 (1998) 293, pl. 132.India, Andaman Isls, Nicobar Isls, S & C China, Hainan, Taiwan, Ryukyu Isls, Burma,Thailand, Laos, Cambodia, Vietnam, Sumatra, Peninsular Malaysia, Java, Lesser SundaIsls, Borneo, Philippines, Sulawesi.BABUYAN ISLS, LUZON, MINDORO, LEYTE, SAMAR, MINDANAO. In primary forests,mostly at low and medium altitudes, ascending to 1400m.Notes- On-going revision is being done by Zhu Hua, where according to him about 21species occur in the Philippines. But it could be more.Litosanthes Blume =LasianthusLucinaea de Candolle =SchraderaLudekia RidsdaleRevision:Ridsdale, CE. 1978. A revision of the tribe Naucleeae s. s. (Rubiaceae). Blumea 24: 307–366.(note: Starts with p. 335).Ludekia bernardoi (Merr.) Ridsd., Blumea 24 (1978) 335; --Neonauclea bernardoi (Merr.) Merr., J.Wash. Acad. Sci. 5 (1915) 539; --EPFP 3 (1923) 511; Nauclea bernardoi Merr., PJS 10 c (1915)102. –Type: FB 20446 Bernardo (not seen). Luzon: Cagayan prov., May 1913.LUZON: Cagayan, Laguna (Mt Makiling), Camarines, MASBATE, LEYTE, SAMAR,MINDANAO: Zamboanga, Davao. Lowland primary forests. (Merrill 1923).Revision:Metadina Bakhuizen filius

Ridsdale, CE. 1978. A revision of the tribe Naucleeae s. s. (Rubiaceae). Blumea 24: 307–366.Metadina trichotoma (Zoll. & Mor.) Bakh.f., Taxon 19 (1970) 472; --Ridsd., Blumea 24 (1978)350-351.Adina zschokkei Elmer, LPB 3 (1911) 987; --Merr., EPFP 3 (1923) 507. –Type: Elmer 12176India (Assam), Bangladesh, Burma, Thailand, Sumatra, Peninsular Malaysia, Java,Borneo, Philippines, Sulawesi, Moluccas, New Guinea.PALAWAN, SIBUYAN. Lowland thickets.Mitracarpus Zuccarini ex Schultes & Schultes filiusMitracarpus hirtus (L.) DC, Prodr. 4 (1830) 572; --Spermacoce hirta L., Sp. Pl. ed. 2 (1762) 148.Native of tropical America, introduced elsewere.Mitragyna KorthalsRevision:Ridsdale, CE. 1978. A revision of Mitragyna & Uncaria. Blumea 24: 43–100.Mitragyna diversifolia (Wall. ex G Don) Havil., J. Linn. Soc. Bot. 33 (1897) 71, pro parte; --Ridsdale, Blumea 24 (1978) 65; --Stephegyne diversifolia (Wall. ex G Don) Brandis, Forest Fl. NWIndia (1874) 263; --Nauclea diversifolia Wall. ex G Don, Gen. Hist. 3 (1834) 467.Mitragyna rotundifolia (Roxb.) Kuntze, Rev. Gen. Pl. (1891) 289; --Merr. Fl. Manila (1912) 446; EPFP 3(1923) 508; --Nauclea rotundifolia Roxb., Fl. Ind. 2 (1824) 124.Nauclea adina Blanco, Fl. Filip., ed. 2 (1845) 102.Stephegyne tubulosa Fern.-Villar in Blanco, Fl. Filip., ed. 3, 4(13A) (1880) 104.Stephegyne parvifolia Vidal, Sinopsis Atlas (1883) t. 56, Fig. A.Bangladesh, Burma, Thailand, Cambodia, Laos, Vietnam, S China, Peninsular Malaysia,Java, Philippines.LUZON: Abra, Benguet, Nueva Vizcaya, Pangasinan, Zambales, Nueva Ecija,Pampanga, Bataan, Bulacan, Rizal, Batangas, MINDORO, MASBATE, GUIMARAS.Low and medium altitude dry thickets.Mitragyna speciosa (Korth.) Havil., J. Linn. Soc. Bot. 33 (1897) 69; --Elmer, LPB 1 (1906) 12;--Merr., EPFP 3 (1923) 508; --Ridsdale, Blumea 24 (1978) 65; --Stephegyne speciosa Korth.,Verh.Nat. Geschaft Bot. (1839-43) 160.Nauclea luzoniensis Blanco, Fl. Filip., ed. 2: 102 (1845).Peninsular Thailand, Sumatra, Peninsular Malaysia, Borneo, Philippines, New Guinea.LUZON: Cagayan, MINDORO, MINDANAO: Davao, Agusan. Lowland forests.Morinda LinnaeusLiterature:Wong, KM. 1984. A synopsis of Morinda (Rubiaceae) in the Peninsular Malaysia. Malay. Nat. J.38: 89–98.Igersheim, A. & E. Robbrecht. 1993. The character states and relationships of thePrismatomerideae (Rubiaceae-Rubioideae). Opera Bot. Belg. 6: 61–79.Johansson, JT.1994. The genus Morinda (Morindeae, Rubioideae, Rubiaceae) in New Caledonia:taxonomy and phylogeny. Opera Bot. 122.Morinda bartlingii Elmer, LPB 4 (1912) 1340; --Merr., EPFP 3 (1923) 572. –Type: Elmer 13037.PALAWAN. Lowland thickets. According to Merrill (1923), very close to M. celebica Miq.

Morinda celebica Miq., Fl. Ind. Bat. 2 (1857) 246; --Merr., EPFP 3 (1923) 572.Borneo, Philippines, Sulawesi.BUSUANGA, PALAWAN, SULU ARCHIPELAGO (UBIAN)Morinda citrifolia L., Sp. Pl. (1753) 176; --Merr., Fl. Manila (1912) 453; EPFP 3 (1923) 573var. citrifoliaTrop. & E. Asia to SW. Pacific. 36 CHH 38 KZN NNS OGA TAI 40 ASS IND SRL 41 CBD?MYA NCB THA VIE 42 BOR CKI JAW LSI MLY MOL PHI SUL SUM XMS 43 BIS NWG SOL50 NTA QLD WAU (60) nwc PHX SAM TOK ton VAN (61) coo mrq sci tua tub (62) crl mrnmrs (63) haw (79) mxt (80) blz cos hon nic pan (81) jam lee trt win (83) clm.India to Indochina, Malesia, Australia and Polynesia. Throughout the Philippines.Lowland thickets, chiefly along the seashore.var. bracteata (Roxb.) Kurz, J. Asiat. Soc. Bengal 46(2) (1877) 151; --Hook.f., Fl. Brit. Ind. 3 (1880)156, nom. illeg.; --Morinda bracteata Roxb., Fl. Ind. 2 (1824) 198, nom. illeg.; --Merr., Fl. Manila(1912) 453; EPFP 3 (1923) 572.Malesia (widespread). Throughout the Philippines in most islands and provinces.Lowland thickets and secondary forests.Morinda coriacea Merr., PJS 17 (1920, publ. 1921) 435; EPFP 3 (1923) 573. –Type: BS 34439Ramos & PascasioMINDANAO: Surigao del Norte. Lowland thickets and forests.Morinda elliptifolia Quisumb. & Merr., PJS 37 (1928) 204. –Type:MINDANAO:Morinda microcephala Bartl. ex DC., Prodr. 4 (1830) 449.Morinda microphylla D.Dietr., Syn. Pl. 1 (1839) 784.Philippines. 42 PHI.Morinda nitida Merr., PJS 17 (1920, publ. 1921) 435; EPFP 3 (1923) 573. –Type: BS 35074 Ramos &PascasioBUCAS GRANDE. Lowland dry forest.Morinda parvifolia Bartl. in DC, Prodr. 4 (1830) 499; --Merr., EPFP 3 (1923) 573. –Type: Haenkesn.Morinda cumingiana Vidal, Phan. Cuming. Philip. (1885) 184.Lucinaea cumingiana Vidal, Phan. Cuming. Philip. (1885) 216.SE China, Hainan, Taiwan, Vietnam, and the Philippines.BABUYAN ISLS, LUZON: Ilocos Norte, Cagayan. Lowland thickets.Morinda philippinensis Elmer, LPB 3 (1911) 1044; --Merr., EPFP 3 (1923) 573. –Type: Elmer12460N Borneo, Philippines.MINDORO, PALAWAN, SIBUYAN. Lowland thickets and forests.Morinda platyphylla Merr., PJS 11 c (1920) Bot. 33; EPFP 3 (1923) 573. –Type: BS 23414 RamosLUZON: Sorsogon. Forest, altitude c. 1000m.Morinda umbellata L., Sp. Pl. (1753) 176; --Merr., EPFP 3 (1923) 574.subsp. umbellata.

Sri Lanka, India, S China, Hainan, Taiwan, Ryukyus, Thailand, Vietnam, Sumatra,Peninsular Malaysia, Java, Borneo, Philippines, Bismarck Archipelago, Solomon Isls,Australia (Queensland), Marianas, Samoa.LUZON: Cagayan, Zambales, Sorsogon, MINDORO, CULION. Lowland thickets andforests.Doubtful taxa:Morinda coreia Buch.-Ham., Trans. Linn. Soc. London 13: 537 (1822).Morinda tinctoria Roxb., Hort. Beng. (1814) 15, nom. nud.; Fl. Ind. 2 (1824) 197; --Merr., EPFP 3 (1923) 572.Sri Lanka, India, Thailand, Vietnam, Java.LUZON: Benguet, NEGROS. Thickets and forests up to 500m altitude.According to Merrill (1923) identification of Philippine materials not certain as thematerial is imperfect.Mussaenda Burman ex LinnaeusRevisions:Jayaweera, DMA. 1963. The Rubiaceous genus Mussaenda: The morphology of the Asiaticspecies. Journal of the Arnold Arboretum 44: 111–126.Jayaweera, DMA. 1963. The Rubiaceous genus Mussaenda: The species of the PhilippineIslands. Journal of the Arnold Arboretum 45: 101–139.Mussaenda acuminatissima Merr., PJS 17 c (1920) 436; EPFP 3 (1923) 517; --Jayaweera, J. Arn.Arb. 45 (1964) 104, Fig. 4 o-q. --Type: BS 33133 (NY*, lecto, designated by Jayaweera, 1964).Luzon: Ilocos Norte prov., Mt Nangapatan, dry slopes, altitude 700m, Aug 1918.LUZON: Ilocos Norte (Mt Nangapatan). Known only from the type collection.Mussaenda albiflora Merr., PJS 5 c (1910) Bot. 241; EPFP 3 (1923) 518; --Jayaweera, J. ArnoldArb. 45 (1964) 105, Fig. 1 h-i, 3 a-e. --Type: FB 17358 Curran (NY*, lecto, designated byJayaweera, 1964; isolecto: US*). Negros: Negros Occidental prov., Faraon, Sept 1909.NEGROS: Negros Occidental (Faraon; Iglamgam); PANAY: Aklan (Libacao), Capiz(Jamindan), Antique. Humid gullies to c. 300m. Fl. Apr-Jun, Sept; fr. Sept.Mussaenda anisophylla Vidal, Phan. Cuming. Philipp. (1885) 178; --Merr., EPFP 3 (1923) 518; --Jayaweera, J. Arnold Arb. 45 (1964) 108, Fig. 3 j-s. --Lectoype: Cuming 918 (BM, designated byJayaweera, 1964; isolecto: G). Luzon: Albay prov.LUZON: Kalinga (Balbalan), Pangasinan (Mt San Isidro Labrador), Bataan (MtMariveles, Lamao River), Rizal (Antipolo), Laguna (Mt Makiling; San Antonio), Batangas,Quezon (Lucban), Albay, ALABAT, MINDORO: Mindoro Oriental (Mt Halcon; Mt Ilong),NEGROS: Gimogaan River, DINAGAT. Near water courses in deeply shaded ravines inmossy forests, medium altitudes up to 800m.Mussaenda attenuifolia Elmer, LPB 5 (1913) 1874; --Merr., EPFP 3 (1923) 518 (sphalm.‘attenuatifolia’ ); --Jayaweera, J. Arnold Arb. 45 (1964) 109, Fig. 1 f. --Type: Elmer 13304 (A, lecto;isolecto: GH, L*, NA*, NY*, US*). Mindanao: Agusan del Norte prov., Mt Urdaneta (=Hilonghilong).Fairly well-scattered in wet earth and upon rock ledges of the wooded banks of theCatangan Creek, Jul 1912MINDANAO: Agusan del Norte (Mt Urdaneta, =Hilong-hilong). Along wooded streamembankments, altitude 150-450m. Known only from the type collection.Mussaenda benguetensis Elmer, LPB 1 (1906) 13; EPFP 3 (1923) 518; --Jayaweera, J. ArnoldArb. 45 (1964) 111, Fig. 1l, 3 x, y, z, aa. --Type: Elmer 5935 (PNH, holo; iso in NY*, US*). Luzon:Benguet prov., Baguio, Mar 1904.

LUZON: Abra, Mountain Province, Ifugao, Benguet. In thickets on open slopes, inravines, etc., altitude 1200-2200m.Mussaenda chlorantha Merr., PJS 8 c (1913) Bot. 47; EPFP 3 (1923) 518; --Jayaweera, J. ArnoldArb. 45 (1964) 112, Fig. 1b-c; 2 d-e. --Type: Merrill 770 (US*, lecto, cited by Jayaweera, 1964).Luzon: Benguet prov., Mt Tonglon (=Mt Sto Tomas), in forests, altitude c. 1800m, May 1911.LUZON: Benguet (Mt Sto Tomas; Twin Peaks), Pampanga (Mt Pinatubo, CampStotsenburg). Mossy forests, 1800-2000m. Fl. May.Mussaenda grandifolia Elmer, LPB 1 (1906) 12; --Merr., EPFP 3 (1923) 518; --Jayaweera, J. ArnoldArb. 45 (1964) 115, Fig. 1q; 4 a-c. –Type: Merrill 817 (US*, lecto, designated by Jayaweera, 1960-61). Palawan: Palawan prov., Point Separation, 20 Feb 1903.PALAWAN. Lowland forests. Fl. & fr: Feb and Mar. Known from two collections.Mussaenda lanata C Robinson, PJS 6 c (1911) Bot. 47; --Merr., EPFP 3 (1923) 518; --Jayaweera,J. Arnold Arb. 45 (1964) 116, Fig. 1a; 2 a-c. –Type: Merrill 6681 (US*, lecto). Luzon: Benguet prov.,Antamok, May 1909.LUZON: Ilocos Sur (Concepcion, Mt Tirad; Cervantes Trail), Abra, Mountain Province,Benguet (Sabangan; Sablan; Baguio; Itogon; Antamok), Zambales. Streamside thickets,altitude 900-1600m. Fl. Mar, Apr, May, Aug, Oct, Dec; fr. Oct, Nov, Dec.Mussaenda macrophylla Wall. var. brevipilosa Jayaweera, J. Arnold Arb. 45 (1964) 117.–Type: BS 3770 Fenix (NY, holo; iso US). Batan, Jun 1907.Mussaenda macrophylla Merr., PJS 3 c (1908) Bot. 437; EPFP 3 (1923) 518, non Wall. in Roxb. (1824).BATAN, CALAYAN, CAMIGUIN DE BABUYANES, BOHOL, CAMIGUIN DEMINDANAO, SIARGAO. Lowland thickets and forests.Mussaenda magallanensis Elmer, LPB 3 (1911) 996; --Merr., EPFP 3 (1923) 519; --Jayaweera,J. Arnold Arb. 45 (1964) 119, Fig. 1d; 2 f-h. –Type: Elmer 12451 (A, lecto; isolecto: BISH*, L*, GH,MO*, NY*, US*). Sibuyan: Magallanes (=Magdiwang), ‘light woods along the coast or in the firsthills’, May 1910.LUZON: Isabela (San Mariano; Palanan), Laguna (Mt Banahaw), MINDORO: MindoroOccidental (Sablayan, Ligaya), Mindoro Oriental (Puerto Galera; Calapan; Baco River;Bongabon and Pinamalayan; SIBUYAN (foothills of Mt Giting-giting), PANAY: Capiz,LEYTE: Leyte (Dagami). Low altitudes from sea-level to 300m, along rivers, mountainslopes, brushlands and thickets. Fl. Jan-May, Aug, Oct-Dec; fr. Jan, Aug-Nov.Mussaenda milleri Elmer, LPB 10 (1939) 3780, not validly published, no Latin diagnosis; --Jayaweera, J. Arnold Arb. 45 (1964) 121. –Type: Elmer 22694 (no materials extant, possibly lost).Luzon: Nueva Ecija prov., Camp Labi, ‘upon the summit mountain ridge at c. 3500ft [=1067m], Mar1937. --Neotype (designated by Jayaweera, 1964): BS 26468 Ramos & Edaño (NY*). Luzon:Nueva Ecija prov., Mt Umingan (=Mingan), Aug-Sept 1916.LUZON: Nueva Ecija (Mingan Mtns), altitude c. 1000m. Ridge summit vegetation,altitude c. 1000m. Fl. Mar; fr. Aug-Sept. Apparently known only from this general locality.Mussaenda multibracteata Merr., PJS 11 c (1916) Bot. 34; EPFP 3 (1923) 519; --Jayaweera, J.Arnold Arb. 45 (1964) 123, Fig. 4 f-n. –Type: BS 23585 Ramos (NY*, lecto, designated byJayaweera, 1964; isolecto: US*). Luzon: Sorsogon prov., Mt Pocdal, on damp slopes in openforests, 10 Aug 1915.LUZON: Cagayan (Mt Dos Cuernos), Isabela (Mt Moises; Palanan), Aurora (Casiguran),Quezon (Mt Tulaog; Mt Binuang), Camarines Norte (Minasag River), Camarines Sur(Her-it River; Mt Isarog), Albay (Mayon Volcano), Sorsogon, CATANDUANES (MtTagmasuso; Bato trail to Viga). Along streams, damp slopes, forests, from sea-level to2100m. Fl. May-Aug; fr. Jul-Sept, Nov, Dec.

Mussaenda nervosa Elmer, LPB 3 (1911) 994; --Merr., EPFP 3 (1923) 519; --Jayaweera, J. ArnoldArb. 45 (1964) 124, Fig. 1e; 2 i, j, k, ka. –Type: Elmer 10510 (A, lecto, designated by Jayaweera,1964; isolecto: BISH*, GH, L*, MO*, NY*, US*). Mindanao: Davao, Todaya, Mt Apo, ‘open exposedhill at 4000ft [=1219m]’, May 1909LUZON: Nueva Ecija, MINDANAO: Bukidnon, Davao del Sur (Mt Apo). Degraded forestsup to c. 1200m.Mussaenda palawanensis Merr., PJS 10 c (1915) Bot. 103; EPFP 3 (1923) 519; --Jayaweera, J.Arnold Arb. 45 (1964) 125, Fig. 1k. –Type: BS 15531 Fenix (US*, lecto, designated by Jayaweera,1964). Palawan: Alphonso XIII (=Quezon), Jul 1912.PALAWAN (widespread). Lowland thickets and old clearings, up to c. 150m.Mussaenda philippica A Rich., Mém. Rubiac. (1830) 165; Mem. Soc. Hist. Nat. Paris 5 (1834) 245;--Merr., EPFP 3 (1923) 519; --Jayaweera, J. Arnold Arb. 45 (1964) 128, Fig. 1j; 3 f-i. –Type:Perrottet sn.Mussaenda philippica A Rich. forma aurorae (Sulit) Jayaweera, J. Arnold Arb. 45 (1964) 131; --Mussaendaphilippica A Rich. var. aurorae Sulit, Philip. J. Forestry 2 (1939) 39, pl. 3, Fig. 1. –Type: FB 24876Mabesa (US*, isolecto). Luzon: Laguna prov., Mt Balong bulo, Jun-Dec 1915.Mussaenda grandiflora Rolfe, J. Linn. Soc., Bot. 21 (1884) 311, nom. illeg.Philippines, Solomon Isls, Carolines (Palau; Yap), Fiji Isls.LUZON: Cagayan to PALAWAN and MINDANAO. Common in thickets and secondaryforests at low and medium altitudes.Mussaenda philippinensis Merr., PJS 3 c (1908) Bot. 264; EPFP 3 (1923) 520; --Jayaweera, J.Arn. Arb. 45 (1964) 132, Fig. 1n-p; 3 t-w. --Type: Merrill 4139 (NY*, lecto, designated byJayaweera, 1964; isolecto in L*, US*). Semirara Is., Jul 1905.LUZON: Apayao (Mt Duraragan), ALABAT, MINDORO: Mindoro Oriental (Mt Yagaw, Eslope; Oyon Creek), SEMIRARA, PANAY. Partial shade along creeks and ridges ofsecond-growth forests, altitude 10-300m. Flowering in Apr-Jul, fruiting Jun, Nov andDec.Mussaenda pinatubensis Elmer, LPB 10 (1934) 3210; --Jayaweera, J. Arnold Arb. 45 (1964) 134.–Type: Elmer 21978 (PNH, holo; iso: A, GH, L*, NY*, US*). Luzon: Pampanga prov., CampStotsenburg (lower slopes of Mt Pinatubo), on hot and dry river embankments, altitude c. 1065m,May 1927.LUZON: Pampanga (Mt Pinatubo). Known only from the type collection. Fl. May.Mussaenda scandens Elmer, LPB 3 (1911) 992; --Merr., EPFP 3 (1923) 520; --Jayaweera, J. ArnoldArb. 45 (1964) 135, Fig. 1g; 2 l, m, n, o, p. –Type: Elmer 11291 (GH, lecto, designated byJayaweera, 1964; isolecto: A, L*, MO*, NA*, NY*, US*). Mindanao: Davao del Sur prov., Mt Apo,Todaya, ‘densely wooded ravine of the Baracatan creek at 2750ft [=838m]’, Aug 1909.Mussaenda villosa Merr., PJS 5 c (1910) Bot. 243; --Elmer, LPB 3 (1911) 992, non Wall.MINDANAO: Zamboanga del Sur (Malangas), Davao del Sur (Mt Apo), Surigao delNorte. Medium altitude forests along streams, altitude 600-800m. Fl. May, Jul, Aug;fr. Aug, Oct, Nov.Mussaenda setosa Merr., PJS 10 c (1915) Bot. 104; EPFP 3 (1923) 520; --Jayaweera, J. Arnold Arb.45 (1964) 137, Fig. 1m; 4 d-e. –Type: Merrill 9496 (US, lecto, chosen by Jayaweera, 1964; isolectoin L*). Palawan: Malampaya Bay, Mt Capoas, Apr 1913.PALAWAN: Mt Capoas. Forested ridge, c. 700m. Fl. Apr. Only known from the typecollection.Mussaenda vidalii Elmer, LPB 3 (1911) 993; --Merr., EPFP 3 (1923) 520; --Jayaweera, J. Arnold Arb.45 (1964) 138, Fig. 2 q-x. –Type: Elmer 11309 (GH, lecto, designated by Jayaweera, 1964;

isolecto: A, L*, MO*, NY*, US*). Mindanao: Davao del Sur prov., Mt Apo, Todaya, ‘humid woods at3750ft [=1143m], Aug 1909.LEYTE: Leyte (Ormoc, Antilao River; Lake Danao), SAMAR: Province unknown (CatubigRiver; Mt Cansayao); Western Samar (Loquilocon; Bagacay), MINDANAO: Bukidnon(Pigtaoranan; Tangculan & vicinity, Davao del Sur (Mt Apo). Low to mediumaltitudes, 150-600m, along edges of humid forests as well as on second-growthforests of open rolling country. Fl. Mar-Aug; fr. Mar, Jun & Jul.Mycetia ReinwardtRevisions:Deb, D. 1986. Taxonomic revision of Mycetia Reinw. (Rubiaceae) in Indian subcontinent. Bulletinof the Botanical Survey of India. 28: 114–132.Gideon, OG. et al. 1988. Notes on Mycetia (Rubiaceae). Floribunda 1.Mycetia cauliflora Reinw., Syll. Ratisb. 2 (1828) 9; --Steen., Mountain Fl. Java (1972) pl. 47-10; --KMWong in FS Ng (ed.) Tree Fl. Malaya 4 (1989) 379.Mycetia lateriflora (Blume) Reinw. ex Korth., Nederl. Kruidk. Arch 2 (1850) 118; --Merr., EPFP 3 (1923) 521;--Bertiera latiflora Blume, Bijdr. (1826) 988.Adenosacme apoensis Elmer, LPB 3 (1911) 1001. –Type: Elmer 10504 (BISH*, L*, MO*, NY*, US*, iso).Mindanao: Davao del Sur prov., Mt Apo, Todaya, humus-covered fertile soil in moist woods at4000ft, May 1909.S Thailand, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi, LesserSunda Isls (Flores).MINDORO, MINDANAO (Lanao, Bukidnon, Davao del Sur). In damp rainforestundergowth, altitude 600-1200m.Mycetia javanica (Blume) Korth., Nederl. Kruidk. Arch. 2 (1850) ex Valeton in Icon. Bogor. 3(1908) 183; --Merr., PJS 10 c (1915) Bot. 105; EPFP 3 (1923) 520; --Bertiera javanica Blume, Bijdr.(1826) 987. –Type from Java.Adenosacme longifolia Rolfe, J. Bot. 23 (1885) 213; --Vidal, Rev. Pl. Vasc. Filip. (1886) 152, non ?Wall., nonRondeletia longifolia Wall.Adenosacme mindanaensis Elmer, LPB 3 (1911) 1002. –Type: Elmer 10831 (BISH*, L*, MO*, NY*, US*,iso). Mindanao: Davao del Sur prov., Mt Apo, Todaya, on very wet, densely shaded ravines or onseepageledges along the Sibulan River at 3250ft, Jun 1909.Nicobar Isls, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, New Guinea.LUZON: Laguna (Los Baños; Mt Banahaw), Albay, Sorsogon, MINDORO: MindoroOriental, PANAY, LEYTE, SAMAR, MINDANAO: Zamboanga, Lanao del Sur (CampKeithley), Bukidnon, Davao, Agusan del Norte, Surigao. In very damp forests alongstreams at low and medium altitudes.Myrmecodia JackRevision:Huxley, CR & MH Jebb. 1991. The tuberous epiphytes of the Rubiaceae 1 : A new subtribe- TheHydnophytinae. Blumea 36: 1–20.Huxley, CR & MH Jebb. 1993. xxxxxxxxx Blumea 37 (1993) 271-334.Myrmecodia tuberosa Jack, Trans. Linn. Soc. London 14 (1823) 123.Mymecodia echinata Gaudich., Bot. Freycinet. Voy. (1826) 472, t. 96; --Merr., EPFP 3 (1923) 570.Vietnam, Sumatra, Peninsular Malaysia, Java, Lesser Sunda Isls, Borneo, Philippines,Sulawesi, Moluccas, New Guinea, Bismarck Archipelago, Australia (Queensland),Solomon Isls.LUZON, Merrill 3394, Whitford 1144, Copeland. Epiphytic in mangrove swamps,

apparently also occurring in primary forest at medium altitudes, ascending to 1000m.‘apoensis’; --Huxley & Jebb, Blumea 37 (1993) 280.Myrmecodia apoensis Elmer, LPB 3 (1911) 1040; --Merr., EPFP 3 (1923) 569.Myrmecodia urdanetensis Elmer, LPB 5 (1913) 1876. Elmer 14083.Myrmecodia tuberosa Jack ‘apoensis’LUZON, MINDANAO: Davao del Sur (Mt Apo), Agusan del Norte (Mt Urdaneta, =MtHilong-hilong). Epiphytic on exposed ridges and mossy forests, c. 1200-1600m.‘sibuyanensis’; --Huxley & Jebb, Blumea 37 (1993) 280.Myrmecodia sibuyanensis Elmer, LPB 3 (1911) 1041.Myrmecodia sorsogonensis Elmer, LPB 9 (1934) 3253.LUZON, SIBUYAN, PANAY, MINDANAO. Epiphytic in primary forests, 225-1000m.Myrmeconauclea MerrillRevision:Ridsdale, CE 1978. A revision of the tribe Naucleeae s. s. (Rubiaceae). Blumea 24: 307–366.Myrmeconauclea strigosa (Korth.) Merr., PJS 17 (1920, publ. 1921) 375; EPFP 3 (1923) 517.Neonauclea strigosa (Korth.) Merr., J. Wash. Acad. Sci. 5 (1915) 542.Bancalus strigosus (Korth.) Kuntze, Revis. Gen. Pl. 1 (1891) 276.Nauclea strigosa Korth., Verh. Nat. Gesch. Ned. Bezitt., Bot. (1842) 157.Sarcocephalus fluviatilis Elmer, LPB 4 (1912) 1357.Borneo, Philippines.LUZON, PALAWAN.MyrmephytumRevisions:Huxley, CR & MH Jebb. 1991. The tuberous epiphytes of the Rubiaceae 1 : A new subtribe- TheHydnophytinae. Blumea 36: 1–20.Huxley, CR & MH Jebb. 1991. The tuberous epiphytes of the Rubiaceae 3 : A revision ofMyrmephytum to include Myrmedoma. Blumea 36: 43–52.Myrmephytum beccarii Elmer, LPB 3 (1911) 1037; --Merr. EPFP 3 (1923) 568; --Huxley & Jebb,Blumea 36 (1991) 47, Fig. 2-b. --Type: Elmer 11467-a (Sibuyan: Mt Giting-giting, along PaualaRiver, thicket on dry, gravelly stream bed, altitude 750 ft, May 1910.SIBUYAN, BILIRAN. In thickets, alt. 225 m.Myrmephytum selebicum (Becc.) Becc., Malesia 2 (1884) 93, t. 8 & 9.; --Huxley and Jebb,Blumea 36 (1991) 46, Fig. 2-a.Philippines, Sulawesi.MINDANAO: Agusan del Norte.Myrmephytum ‘sp. 3’ Huxley and Jebb, Blumea 36 (1991) 51.Based on BS 925 Mangubat (K). Mindoro: Mindoro Mindoro Oriental prov., Calapan.Nauclea LinnaeusRevision:Ridsdale, CE. 1978. A revision of the tribe Naucleeae s. s. (Rubiaceae). Blumea 24: 307–366.

Nauclea officinalis (Pierre ex Pit.) Merr. & Chun, Sunyatsenia 5 (1940) 188; --Sarcocephalusofficinalis Pierre ex Pit. in H Lecomte, Fl. Indo-Chine 3 (1922) 26.China (Guangdong), Hainan, Cambodia, Laos, Thailand, Sumatra, Peninsular Malaysia,Java, Borneo, ?Philppines.Nauclea orientalis (L.) L., Sp. Pl. ed. 2 (1762) 243; --Merr., J. Wash. Acad. Sci. 5 (1915) 535;Sarcocephalus orientalis (L.) Merr., PJS 3 c (1909) Bot. 436; --Bancalus orientalis (L.) Kuntze,Revis. Gen. Pl. 1 (1891) 276; --Cephalanthus orientalis L., Sp. Pl. (1753) 95.Sarcocephalus undulatus (Roxb.) Miq., Fl. Ned. Ind. 2 (1856) 133; --Nauclea undulata Roxb., Hort. Beng.(1814) 14, nom. nud., Fl. Ind. 2 (1824) 117; --Merr., EPFP 3 (1923) 517.Nauclea elmeri Merr., J. Wash. Acad. Sci. 5 (1915) 535; EPFP 3 (1923) 515.Sarcocephalus ovatus Elmer, LPB 1 (1906) 33. –Type: FB 3150 Ahern’s collector (NY*, US*, iso). Luzon:Rizal prov., Bosoboso, May-Jun 1905Nauclea lutea Blanco, Fl. Filip.: 141 (1837).Sri Lanka, Burma, Thailand, Cambodia, Sumatra, Java, Lesser Sunda Isls, Borneo,Philippines, Sulawesi, Moluccas, New Guinea, Bismarck Archipelago, Solomon Isls,Australia (Northern Territory, Queensland).BATAN to MINDANAO, in most islands and provinces. Chiefly in secondary forests atlow and medium altitudes.EPFP 3 (1923) 515; --ANauclea robinsonii Merr., J. Wash. Acad. Sci. 5 (1915) 537; EPFP 3 (1923) 517; --Sarcocephaluspubescens C Robinson, PJS 6 c (1911) Bot. 225, non Valeton. –Type: BS 6917 Robinson (NY*,US*, iso). Polillo:POLILLO, LEYTE, SAMAR. Low altitude secondary forests.Nauclea subdita (Korth.) Steud, Nom. Bot. ed. 2, 2 (1841) 186; --Ridsd., Blumea 24 (1978) 329.Sarcocephalus subditus (Korth.) Miq., Fl. Ned. Ind. 2 (1856) 134; --Platanocarpum subditum Korth., Observ.Naucl. Indic. (1839) 19.Nauclea junghuhnii (Miq.) Merr., J. Wash. Acad. Sci. 5 (1915) 536; EPFP 3 (1923) 516; --Sarcocephalusjunghuhnii Miq., Fl. Ned. Ind. 2 (1856) 134.Nauclea multicephala (Elmer) Merr., J. Wash. Acad. Sci. 5 (1915) 536; EPFP 3 (1923) 516; --Sarcocephalusmulticephalus Elmer, Leafl. Philipp. Bot. 5 (1913) 1896.Nauclea glaberrima Blanco, Fl. Filip., ed. 2 (1845) 100.India (Assam), Sumatra, Peninsular Malaysia, Java, Lesser Sunda Isls, Borneo,Philippines, Sulawesi, Moluccas.LUZON: Isabela, Laguna, Quezon, Camarines, Albay, Sorsogon, MASBATE, NEGROS,LEYTE, SAMAR, MINDANAO: Zamboanga, Misamis, Lanao, Bukidnon, Davao, Surigao.In primary and secondary forests, ascending to 900 m.Neolamarckia BosserNeolamarckia cadamba (Roxb.) Bosser, Bull. Mus. Natl. Hist. Nat., B, Adansonia 6 (1984) 247.Anthocephalus cadamba (Roxb.) Miq., Fl. Ind. Bat. ii. 135;Samama cadamba (Roxb.) Kuntze, Revis. Gen. Pl. 1 (1891) 296; --Sarcocephalus cadamba (Roxb.) Kurz,Prelim. Rep. Forest Pegu, App. A (1875) lxxviii; --Nauclea cadamba Roxb., Fl. Ind. 2 (1824) 121.Sri Lanka, India, E Himalayas, Bangladesh, Nicobar Isls, Thailand, Laos, Cambodia,Vietnam, Sumatra, Java, Lesser Sunda Isls, Borneo, Philippines (introduced), Sulawesi,New Guinea.Neonauclea MerrillRevisions:Ridsdale, CE. 1978. A revision of the tribe Naucleeae s. s. (Rubiaceae). Blumea 24: 307–366.

Ridsdale, CE. 1978. A revision of Neonauclea (Rubiaceae). Blumea 34: 177–275Neonauclea bartlingii (DC) Merr., J. Wash. Acad. Sci. 5 (1915) 539; --EPFP 3 (1923) 511; --Ridsd.,Blumea 34 (1989) 218, Fig. 15c (map); --Sarcocephalus bartlingii (DC) Miq., Fl. Ind. Bat. 2 (1857)133; --Fern.-Villar, Novis. App. (1880) 104; --Nauclea bartlingii DC, Prodr. 4 (1830) 344. –Type:Haenke sn (L, photo). Luzon: near Sorsogon.Neonauclea ategii (Elmer) Merr., J. Wash. Acad. Sci. 5 (1915) 539; --EPFP 3 (1923) 511; --Nauclea ategiiElmer, LPB 5 (1913) 1877. –Type: Elmer 13910 (BISH*, BO, L*, NY*2 sheets, US*, iso). Mindanao:Agusan del Norte, Mt Urdaneta (=Hilong-hilong), ‘stony soil among boulders along the woodedbanks of Dalahion River at 1000ft’, Sept 1912.Neonauclea auriculata Quisumb. & Merr., PJS 37 (1928) 205. --Type: BS 49281 Ramos & Edaño (BO, L*,NY*, US*, iso). Mindanao: Davao Oriental prov., Mati, Mar-Apr 1927.var. bartlingiiLUZON: Nueva Vizcaya, Bataan, Pampanga, Bulacan, Rizal, Laguna, Cavite, Batangas,Quezon, Camarines, Albay, MINDORO, MINDANAO: Davao Oriental, Agusan delNorte. Lowland to medium altitude secondary forests, along streams.var. cumingiana (Vidal) Ridsd., Blumea 34 (1989) 219, pl. 9, Fig. 15b (map); --Nauclea cumingianaVidal, Phan. Cuming. Philip. (1885) 176. --Lecto: Cuming 833 (L*, isolecto L*, NY*). Luzon: Lagunaprov., ‘mtns of St Cristoval and Majaijai’ (=Mt San Cristobal and Banahaw); --Syntypes: Cuming1522; Vidal 365, 367 (L).Neonauclea vidalii (Elmer) Merr., J. Wash. Acad. Sci. 5 (1915) 542; --Nauclea vidalii Elmer, LPB 1 (1906)16. –Type: Elmer 6138 (BO, L*, NY*, iso). Luzon: Benguet prov., Sablan, Apr 1904,Neonauclea cordatula (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 539; --EPFP 3 (1923) 512; --Naucleacordatula Merr., PJS 8 c (1913) Bot. 40. --Type: FB 22219 Cenabre (L*, iso). Cebu: Cebu prov.,Pagulason, on ridges, altitude c. 1000m, Dec 1910.LUZON: Ilocos Norte, Ilocos Sur, Pangasinan, Abra, Mountain Province, Benguet,Nueva Vizcaya, Pampanga, Rizal, CEBU. Medium altitudes, chiefly in disturbedforests, ascending to 2100m.Neonauclea calycina (Bartl. ex DC) Merr., J. Wash. Acad. Sci. 5 (1915) 539; EPFP 3 (1923) 512;--Ridsdale, Blumea 34 (1989) 200, Fig. 7 (map). --Nauclea calycina Bartling ex DC, Prodr. 4 (1830)346. –Type: Haenke sn (L, photo).Nauclea blancoi Vidal, Phan. Cuming. Philip. (1885) 175. –Type Cuming 890 (not seen).Neonauclea monocephala (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 541; EPFP 3 (1923) 514; --Naucleamonocephala Merr., PJS 8 c (1913) Bot. 44. –Type: Merrill 2980, leg. S Gracia (NY*, US*, iso; L*,photo). Luzon: Zambales prov., Botolan, Jun 1903.Burma, Peninsular Thailand, Cambodia, Laos, Vietnam, Sumatra, Peninsular Malaysia,Java, Lesser Sunda Isls, N Borneo, Philippines, Sulawesi.LUZON: Cagayan to Sorsogon, most provinces, MINDORO, PALAWAN,POLILLO,TICAO, MASBATE, LEYTE, PANAY, NEGROS, BASILAN, MINDANAO:Zamboanga, Agusan del Norte. In primary and secondary forests at low and mediumaltitudes.Neonauclea circumscissa Ridsd., Blumea 34 (1989) 217, Fig. 15a (map); 16, pl. 8. --Type: PNH18407 DR Mendoza (L*, holo). Luzon: Albay prov., Mt Mayon, May-Jun 1953.LUZON: Batangas, Quezon, Camarines, Albay, Sorsogon.Neonauclea formicaria (Elmer) Merr., EPFP 3 (1923) 512; --Ridsdale, Blumea 34 (1989) 214, Fig.15a (map); --Nauclea formicaria Elmer, LPB 3 (1911) 989. --Type: Elmer 11034 (BISH*, L*, NY*,US*, WRSL, iso). Mindanao: Davao del Sur prov., Mt Apo, Todaya, ‘in shrubberies of gulches in thecogonregion at 1000ft’, Jun 1909.PANAY: Antique, Capiz, Iloilo, GUIMARAS, NEGROS, CEBU, BOHOL, BILIRAN,SAMAR, MINDANAO: Zamboanga del Sur, Misamis Occidental, Davao, Bukidnon,Agusan del Norte, Surigao del Sur. Low and medium altitude shrubberies, thickets

and open forests associated with cogonal grasslands.Neonauclea glabra (Roxb.) Bakh.f. & Ridsd., Blumea 34 (1989) 240, pl. 13, Fig. 27 (map); --Nauclea glabra Roxb., [Hort. Beng. (1814) 86, nom. nud.] Fl. Ind. ed. 1, 2 (1824) 121. –Type from --Neonauclea nitida (Havil.) Merr., J. Wash. Acad. Sci. 5 (1915) 541; --Nauclea nitida Havil., J. Linn Soc. Bot.33 (1897) 53; --EPFP 3 (1923) 514. --Type: Callery 55 (P). Photo of type in L*.Neonauclea ovata (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 541; --EPFP 3 (1923) 514; --Nauclea ovataMerr., PJS 8 c (1913) Bot. 42. --Type: BS 14597 Ramos (L*, photo; US*, iso). Camiguin: Camiguinprov., in forests along streams near the old volcano, 27 Apr 1912.Neonauclea venosa (Merr.) Merr. , J. Wash. Acad. Sci. 5 (1915) 542; --EPFP 3 (1923) 515; --Naucleavenosa Merr., PJS 8 c (1913) Bot. 45. --Type: FB 9183 Whitford & Hutchinson (not seen).Mindanao: Zamboanga del Sur prov., Port Banga, Jan 1908.Neonauclea kobbei Elmer, LPB 8 (1919) 3100. –Type: Calicosa sn, Aug 1914Philippines, Moluccas, New Guinea, Australia (Queensland).LUZON: Laguna, Quezon, MASBATE, NEGROS, CAMIGUIN, BASILAN,MINDANAO: Zamboanga del Sur. Lowland forests, ascending to 600m.Neonauclea jagorii (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 540; EPFP 3 (1923) 513; --Ridsd.,Blumea 34 (1989) 221, pl. 15c , Fig. 17; --Nauclea jagorii Merr., PJS 4 c (1909) Bot. 326. –Type:Jagor 987 (B, lost). Samar; --neotype: FB 31136 Fontanoza (NY).LUZON: Camarines Sur, SAMAR. Lowlands, along riverbanks.Neonauclea kentii (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 540; EPFP 3 (1923) 513; --Ridsd.,Blumea 34 (1989) 228, Fig. 21a; --Nauclea kentii Merr., PJS 8 c (1913) Bot. 43. –Type: BS 15440Reillo (L*, 2 sheets, iso). Basilan: Basilan prov., in forests, Aug 1912.Neonauclea mindanaensis (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 541; EPFP 3 (1923) 514; --Naucleamindanaensis Merr., PJS 8 c (1913) Bot. 43. –Type: Copeland 1630 (not seen). Mindanao:Zamboanga, San Ramon, Feb 1905.Neonauclea media auct. non (Havil.) Merr.; --Seeber, Weidelt & Banaag, Dendr. Charact. Imp. For. Trees EMindanao (1979) 360, Fig. 218.BASILAN, MINDANAO: Zamboanga, Lanao del Sur, E Mindanao. Lowland forests,chiefly along streams.Neonauclea lanceolata (Blume) Merr., J. Wash. Acad. Sci. 5 (1915) 504; --Ridsd., Blumea 34(1989) 225.subsp. gracilis (Vidal) Ridsd., Blumea 34 (1989) 225, Fig. 19c (map); --Neonauclea gracilis (Vidal)Merr., J. Wash. Acad. Sci. 5 (1915) 540; EPFP 3 (1923) 513; --Nauclea gracilis Vidal, Phan.Cuming. Philip. (1885) 176; --Type: Cuming 835 (K, L*, 2 sheets).Neonauclea oligophlebia Merr., PJS 12 c (1917) Bot. 159; EPFP 3 (1923) 514. –Type: Wenzel 1608 (A*,BM, iso). Leyte: Leyte prov., Sept 1915.Neonauclea philippinensis (Vidal) Merr., J. Wash. Acad. Sci. 5 (1915) 542; --Nauclea philippinensis (Vidal)Havil., J. Linn. Soc., Bot. 33 (1897) 52; --Adina philippinensis Vidal, Rev. Pl. Vasc. Filip. (1886)148.Peninsular Malaysia, Borneo, Philippines, Sulawesi, Moluccas, New Guinea.LUZON: Nueva Vizcaya, Bataan, Rizal, Laguna, Camarines, Albay,Sorsogon, MINDORO, PALAWAN, LEYTE, SAMAR, MINDANAO: Bukidnon. Chieflyin primary forests at low altitudes.Neonauclea media (Havil.) Merr., J. Wash. Acad. Sci. 5 (1915) 541; EPFP 3 (1923) 513; --Ridsd.,Blumea 34 (1989) 230, Fig. 21c (Map); --Nauclea media Havil., J. Linn. Soc. Bot. 33 (1897) 56.–Type: Callery sn (P).Nauclea latifolia Blanco, Fl. Filip. (1837) 144. –Type: not extant. “Illustrative specimen”: Merrill Sp.Blancoanae 542 (US*). Luzon: Batangas prov., Mt Batulao, forest slopes, altitude 49m, 12 Aug1914.LUZON: Ilocos Sur, Cagayan, Bataan, Rizal, Laguna, Quezon, Batangas, MINDORO,PANAY, GUIMARAS, MINDANAO. Lowland forests.

Neonauclea pseudocalycina Ridsd., Blumea 34 (1989) 204, pl. 2, Fig 10 (map). --Type fromSabah.Borneo (Sabah; Kalimantan), SW Philippines.TAWI-TAWI.Neonauclea puberula (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 542; EPFP 3 (1923) 514; --Ridsd., Blumea 34 (1989) 229, Fig. 21b; --Nauclea puberula Merr., PJS 8 c (1913) 41. –Type:Elmer 7319 (BO, iso). Luzon: Sorsogon prov., Sorsogon, Nov 1905.Neonauclea forsteri Merr. EPFP 3 (1923) 512, non (Seem. ex Havil.) Merr. 1915.LUZON: Sorsogon, MINDORO: Mindoro Oriental (near Calapan), PANAY,GUIMARAS, ?MINDANAO. Low altitude forests. The record from Mindanao istentatively based on Ahern 3624 with no further locality.Neonauclea reticulata (Havil.) Merr., J. Wash. Acad. Sci. 5 (1915) 542; EPFP 3 (1923) 514; --Ridsd., Blumea 34 (1989) 215, pl. 8, Fig. 15a (map); --Nauclea reticulata Havil., J. Linn. Soc. Bot.33 (1897) 62. –Type: Vidal 2955 (K, holo).Taiwan (S tip), Lanyu and the Philippines.BATAN and BABUYAN ISLS, LUZON: Apayao, Abra, Benguet, MountainProvince, Ilocos Sur, Pangasinan, Cagayan, Nueva Vizcaya, Zambales, Nueva Ecija,Bulacan, Rizal. In forested ravines and on slopes at low and medium altitudes.Neonauclea truncata (Hayata) Yamamoto, J. Soc. Trop. Agr. Taiwan 7 (1935) 149; --Ridsdale,Blumea 34 (1989) 213, pl. 8. –Type from Taiwan.Taiwan and N Philippines.BATAN.Neonauclea wenzelii (Merr.) Merr., J. Wash. Acad. Sci. 5 (1915) 542; EPFP 3 (1923) 515; --Ridsd.,Blumea 34 (1989) 228, Fig. 20, pl. 21a; --Nauclea wenzelii Merr., PJS 9 (1914, publ. 1915) 386. –Type: Wenzel 348 (A*, BM, US*, iso). Leyte: Leyte prov., 10 Jul 1913.LEYTE, SAMAR. Lowland forests.Nertera Banks & Solander ex Gaertner =CoprosmaOldenlandia LinnaeusOldenlandia apoensis Elmer, LPB 1 (1906) 17. –Type: Copeland 1131 (NY*, US*, iso). Mindanao:Davao, Mt Apo, altitude c. 2000m, Apr 1904.Hedyotis apoensis (Elmer) Elmer, LPB 3 (1911) 976; --Merr., EPFP 3 (1923) 495.LEYTE, MINDANAO: Misamis Occidental, Bukidnon, Davao del Sur (Mt Apo). Forests,altitude 1200-2000m.Oldenlandia auricularia (L.) K Schum. in HGA Engler & KAE Prantl, Nat. Pflanzenfam. 4(4) (1891)25; --Exallage auricularia (L.) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., TweedeSect. 48(2) (1952) 142; --Metabolus auricularius (L.) Blume ex Bremek., Meded. Bot. Mus. Herb.Rijks Univ. Utrecht 56 (1939) 439; --Hedyotis auricularia L., Sp. Pl. (1753) 101; --Merr., EPFP 3(1923) 495subsp. venosa (Blume) ined.,Hedyotis auricularia subsp. venosa (Blume) Deb, Fl. Tripura 2 (1983) 52; --Hedyotis venosa (Blume) Korth.,Ned. Kruidk. Arch. 2(2) (1851) 160; --Metabolos venosus Blume, Bijdr. (1826) 991.Trop. & Subtrop. Asia: Sri Lanka, India, E Himalayas, Bangladesh, C & S China, Hainan,Burma, Thailand, Vietnam, Sumatra, Peninsular Malaysia, Java, Lesser Sunda Isls,Borneo, Philippines.

LUZON: Ilocos Norte, Benguet, Bulacan, Rizal, Camarines, Albay, MINDANAO:Zamboanga, Lanao, Surigao. Low and medium altitude thickets and open sites.Oldenlandia banksii Elmer, LPB 1 (8 Apr 1906) 19. –Type: CS Banks sn (not seen). Negros: NegrosOccidental prov., Mt Kanlaon, Mar 1902.Hedyotis banksii (Elmer) Elmer, LPB 3 (1911) 976; --Merr., EPFP 3 (1923) 495.Hedyotis yoderi (Elmer) Elmer, LPB 3 (1911) 976; --Merr., EPFP 3 (1923) 501; -- Oldenlandia yoderi Elmer,LPB 1 (8 Apr 1906) 17. –Type: Yoder sn (not seen). Panay: Antique prov., Mt Madiaas, Apr 1905.Hedyotis elmeri Merr., PJS 1 (15 Apr 1906) suppl. 127; EPFP 3 (1923) 497. –Syntypes: Elmer 6834, 6979,9198LUZON: Zambales, Bataan, Rizal, Laguna, Quezon, Camarines, Albay, Sorsogon,PANAY, NEGROS, LEYTE. Mossy forests, altitude 1000-1800m.Oldenlandia benguetensis Elmer, LPB 1 (1906) 18. –Type: Elmer 6243 (NY*, US*, iso). Luzon:Benguet prov., Mt Sto Tomas, May 1904.Hedyotis benguetensis (Elmer) Elmer, LPB 3 (1911) 976; --Merr., EPFP 3 (1923) 496.Philippines, Sulawesi.LUZON: Ilocos Norte, Abra, Mountain Province, Benguet. Mossy forest, altitude1400-2400m.Oldenlandia biflora L., Sp. Pl. (1753) 119; --Merr., EPFP 3 (1923) 492.Thecagonum biflorum (L.) Babu, Bull. Bot. Surv. India 11 (1969, publ. 1971) 214; --Gonotheca biflora (L.)Masam., Sci. Rep. Kanazawa Univ., Biol. 4 (1955) 78; --Hedyotis biflora (L.) Lam., Tabl. Encycl. 1(1792) 272.Tropical and subtropical Asia to W Pacific: Nicobar Isls, Burma, Vietnam, PeninsularMalaysia, Java, Philippines, Sulawesi, New Guinea, Solomon Isls, Australia(Queensland), Fiji, Gilbert Isls, New Caledonia, Samoa, Tonga, Vanuatu, Caroline Isls,Marianas, Marshall Isls.Throughout the Philippines. Low and medium altitude weed in waste places,cultivated sites, settlements, etc. Common.Oldenlandia ciliata Elmer, LPB 1 (1906) 20; --Merr., EPFP 3 (1923) 493. –Type: Elmer 6615 (notseen). Luzon: Benguet prov., Baguio, in loose soil amongst limestone boulders.LUZON: Mountain Province, Benguet. Altitude 1200-1600m, on limestone boulders andledges, recent slides, etc.Oldenlandia corymbosa L., Sp. Pl. (1753) 119; --Merr., Fl. Manila (1912) 447; EPFP 3 (1923) 493; --Hedyotis biflora var. corymbosa (L.) Kurz, J. Asiat. Soc. Bengal 46(2) (1877) 133; --Gerontogeacorymbosa (L.) Cham. & Schltdl., Linnaea 4 (1829) 154; --Hedyotis corymbosa (L.) Lam., Tab.Encyc. 1 (1791) 272Africa, tropical and subtropical Asia, introduced elsewhere.Throughout the Philippines. Often common in and near settlements, gardens, wasteplaces, etc. Note: According to the Global Rubiaceae Checklist, identity of Philippineplants so-named uncertain.Oldenlandia cristata (Willd. ex Roem. & Schult.) ined.Spermacoce cristata Willd. ex Roem. & Schult., Syst. Veg. 3 (1818) 530.Exallage costata (Roxb.) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Tweede Sect. 48(2)(1952) 142; --Oldenlandia costata (Roxb.) K.Schum., Bot. Tidsskr. 24 (1902) 330; --Hedyotiscostata (Roxb.) Kurz, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 45(2) (1876) 135, nom. illeg.; --Spermacoce costata Roxb., Fl. Ind. 1 (1820) 376.Exallage philippensis (Willd. ex Spreng.) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk.,Tweede Sect. 48(2) (1952) 142; --Hedyotis philippensis (Willd. ex Spreng.) Merr. ex C Robinson,PJS 6 c (1911) Bot. 222; --Merr., PJS 8 c (1913) Bot. 37; EPFP 3 (1923) 499; --Spermacocephilippensis Willd. ex Spreng., Syst. Veg. (ed. 16) 1 (1824) 401. –Type: ‘Malaspina’ (B-Willd.*).

Luzon: without definite locality.Hedyotis philippensis var. meyeniana (Walp.) Merr., PJS 8 c (1913) Bot. 38; --EPFP 3 (1923) 499; -- Spermacoce meyeniansn (B, cited by Merrill, 1913). Luzon: Laguna prov.Hedyotis asperrima (Merr.) Merr., EPFP 3 (1923) 495; --Hedyotis philippensis var. asperrima Merr., PJS 8 c(1913) Bot. 38. –Type: Merrill 7252 (L*, iso). Palawan: Malampaya Bay.Hedyotis prostrata (Blume) Korth., Nederl. Kruidk. Arch. 2 (1851) 160; --Merr., PJS 8 c (1913) Bot. 36, pl. 1;--Metabolos prostratus Blume, Bijdr. (1826) 991. –Type from Java.Exallage rigida (Blume) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Tweede Sect. 48(2)(1952) 142; --Oldenlandia rigida (Blume) Kuntze, Revis. Gen. Pl. 1 (1891) 292, nom. illeg.; --Scleromitrion rigidum (Blume) Kurz, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 46(2) (1877) 136; --Hedyotis rigida (Blume) Walp., Ann. Bot. Syst. 2 (1852) 772; --Hedyotis rigida (Blume) Miq., Fl.Ned. Ind. 2 (1857) 181, nom. illeg.; --Merr., EPFP 3 (1923) 500; Backer & Bakhuizen f., Fl. Java 2(1963) 287; --Metabolus rigidus Blume, Bijdr. (1826) 992. –Type from Java.Hedyotis gitingensis Elmer, LPB 3 (1911) 979; --Merr., EPFP 3 (1923) 497, syn. nov.; –Type: Elmer 12434.Hedyotis leucocarpa Elmer, LPB 2 (1908) 524; --Merr., EPFP 3 (1923) 500. –Type: Elmer 9434 (L*, MO*,NY*, US*, iso). Negros: Negros Oriental prov., Cuernos Mtns, in damp soil of shaded woods,altitude 4000ft, Mar 1908.Oldenlandia vestita (R.Br. ex G.Don) Drake, J. Bot. (Morot) 9 (1895) 211; --Hedyotis vestita R Br. in Wall.Cat. n. 847, nom. nud.; ex G Don, Gen. Syst. (1834) 526.Hedyotis costata (Roxb.) Kurz, J. As. Soc. Beng. 45² (1876) 135; --Merr., EPFP 3 (1923) 497.Hedyotis pulgarensis Elmer, LPB 4 (1912) 1333; --Merr., EPFP 3 (1923) 499, syn. nov. –Type: Elmer 13212(L*, MO*, NY*, US*, iso). Palawan: Puerto Princesa, Mt Pulgar (=Thumb Peak), May 1911.E Himalayas, Assam, Bangladesh, Burma, Nicobar Isls, Thailand, Sumatra, PeninsularMalaysia, Java, Borneo, Philippines, Moluccas, New Guinea, Australia (Queensland).Throughout the Philippines in most islands and provinces: LUZON: Pangasinan,Benguet, Isabela, Nueva Vizcaya, Zambales, Bataan (Mt Mariveles), Pampanga(Mt Arayat), Bulacan, Rizal, Laguna, Camarines, Sorsogon, POLILLO, MINDORO,BUSUANGA, CULION, PALAWAN, SIBUYAN, PANAY, NEGROS: Negros Occidental(Mt Kanlaon), Negros Oriental (Cuernos Mtns), CAMIGUIN, BASILAN, MINDANAO:Lanao, Bukidnon, Surigao. Chiefly in low and medium altitude forests, thickets andgrasslands, to 1000m altitude, often common.Oldenlandia diffusa (Willd.) Roxb., Hort. Beng. (1814) 11; --Merr., Fl. Manila (1912) 447; EPFP 3(1923) 493; --Hedyotis diffusa Willd., Sp. Pl. 1 (1797) 566Sri Lanka, India, E Himalayas, Bangladesh, China, Taiwan, Korea, Japan, Nicobar Isls,Burma, Thailand, Sumatra(?), Peninsular Malaysia, Borneo, Philippines.Throughout the Philippines. Habitat and occurrence of the preceding.Oldenlandia filifolia Elmer, LPB 1 (1906) 64; --Merr., PJS 1 (1906) Suppl. 126. –Type: Merrill 3295.Hedyotis filifolia (Elmer) Merr., EPFP 3 (1923) 497.LUZON: Bataan. On coastal bluffs.Oldenlandia herbacea (L.) Roxb., Fl. Ind. 1 (1820) 445; --Merr., Fl. Manila (1912) 448; EPFP 3(1923) 493; --Hedyotis herbacea L., Sp. Pl. (1753) 102.Tropical Africa and Asia. LUZON: Cagayan, Nueva Vizcaya, Zambales, Bulacan, Rizal,GUIMARAS.Oldenlandia horneriana Miq., Fl. Ind. Bat. 2 (1857) 190; --Merr., EPFP 3 (1923) 494.Hedyotis horneriana (Miq.) Fukuoka, South East Asian Stud. 8 (1870) 335, non Miq. (1857).Sumatra, ?Philippines.LUZON: Pangasinan, based on Cuming 974. According to Merrill (1923), knownonly from the original description.Oldenlandia laevigata (Miq.) Kuntze, Revis. Gen. Pl. 1 (1891) 292; --Hedyotis laevigata Miq., Fl.Ned. Ind. 2 (1857) 178.

Philippines.Where?Oldenlandia ovatifolia (Cav.) DC, Prodr. 4 (1830) 427; --Merr., PJS 8 c (1913) Bot. 46; EPFP 3(1923) 494; --Hedyotis ovatifolia Cav., Icon. 6 (1801) 52, t. 573, Fig. 1.Oldenlandia nudicaulis Roth, Nov. Pl. Sp. (1821) 95; --Fern.-Villar, Novis. App. (1880) 107; --Elmer, LPB 1(1906) 20; --Merr., PJS 1 (1906) Suppl. 127.Sri Lanka, India, W & E Himalayas, Nepal, Peninsular Malaysia, Java, Philippines.LUZON: Bataan, Bulacan, Laguna, MINDANAO: Davao. Lowland grasslands andclearings.Oldenlandia pterita (Blume) Miq., Fl. Ind. Bat. 2 (1857) 193; --Merr., EPFP 3 (1923) 494.Thecagonum pteritum (Blume) Babu, Bull. Bot. Surv. India 11 (1969 publ. 1971) 214; --Hedyotis pteritaBlume, Bijdr. (1826) 972.India, Burma, Thailand, Vietnam, Java, Lesser Sunda Isls, Philippines, Sulawesi.LUZON: Cagayan, PALAWAN, NEGROS, MINDANAO: Zamboanga, Davao. Lowlandclearings, plantations, etc., seen in limestone areas.Oldenlandia radicans (Bartl. ex DC.) Kuntze, Revis. Gen. Pl. 1 (1891) 292.Exallage radicans (Bartl. ex DC.) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Tweede Sect.48(2) (1952) 142; --Metabolos radicans Bartl. ex DC., Prodr. 4 (1830) 435.Hedyotis ramosii Merr., PJS 8 c (1913) Bot. 35; EPFP 3 (1923) 500. –Type: BS 15337 Ramos (L*, US*, iso).Leyte: Leyte prov., mtns back of Dagami, in forests, Aug 1912.Hedyotis irosinensis Elmer, LPB 9 (1934) 3248. --Type: Elmer 17394 (NY*, US*, iso). Luzon: Sorsogon prov., Mt Bulusan, SepBorneo(?), Philippines, New Guinea, Australia (Queensland). LUZON: Pangasinan,Bulacan, Rizal, Quezon, Sorsogon (Mt Bulusan), CATANDUANES, LEYTE, SAMAR,MINDANAO: Surigao del Norte. Low altitude forests.Oldenlandia strigulosa Bartl. ex DC., Prodr. 4 (1830) 427.Hedyotis strigulosa (Bartl. ex DC.) Fosberg, Smithsonian Contr. Bot. 45 (1980) 28.Tropical & subtropical Asia to W Pacific: India, S China, Taiwan, Korea, Japan, RyukyuIsls, Philippines, New Guinea, Solomon Isls, Caroline Isls, Marianas.Philippine distribution:Ophiorrhiza LinnaeusLiterature:Darwin, SP. 1976. The Pacific species of Ophiorrhiza L. (Rubiaceae). Lyonia 1.Ophiorrhiza argostemmoides Elmer, LPB 9 (1934) 3254. –Type: Elmer 16345 (PNH, holo, lost;MO*, NY*, US*, iso). Luzon: Sorsogon prov., Irosin, Mt Bulusan, Jun 1916.LUZON.Ophiorrhiza biflora Elmer, LPB 1 (1906) 21; --Merr., EPFP 3 (1923) 501. –Type: Copeland sn (notseen). Mindanao: Davao, Mt Apo, on mossy logs, altitude c. 2000m, Oct 1904.NEGROS, MINDANAO: Davao del Sur (Mt Apo). Mossy forests, altitude c. 1800m.Ophiorrhiza caespitulosa Elmer, LPB 2 (1908) 523; --Merr., EPFP 3 (1923) 502. –Type: Elmer10332 (not seen).Ophiorrhiza caespitulosa var. fulva Elmer, LPB 5 (1913) 1880. –Type:PANAY, NEGROS, MINDANAO (Agusan del Norte). Damp primary forests, altitude c.

1050m.Ophiorrhiza camiguinensis Elmer, Leafl. Philipp. Bot. 5: 1881 (1913).Philippines (Mindanao). 42 PHI.Ophiorrhiza ciliata Elmer, LPB 9 (1934) 3256. –Type: Elmer 17544 (NY*, US*, iso). Luzon: Lagunaprov., Mt Makiling, Jun-Jul 1917.LUZON.Ophiorrhiza curtiflora Elmer, LPB 5 (1913) 1882; --Merr., EPFP 3 (1923) 502. –Type: Elmer14208 (BISH*, MO*, NY*, US*, iso). Mindanao: Agusan del Norte prov., Mt Urdaneta (=Hilong-hilong),Cabadbaran, in wet, deeply humus-covered soil of an alluvial nature in a dense forested depression at3500ft N of Duros.MINDANAO: Agusan del Norte (Mt Hilong-hilong). Primary forest, altitude c. 1050m.Ophiorrhiza davaensis Quisumb., PJS 41 (1930) 363, t. 26. --Type: BS 49307 Ramos & Edaño(PNH, holo, possibly lost; iso in A*). Mindanao: Davao Oriental prov., Mati. Mar-Apr 1927.MINDANAO: Davao Oriental (Mati). Along stream banks and open places at low alt.Known only from the type collection.Ophiorrhiza dolichophylla Merr., PJS 20 (1922) 470; --EPFP 3 (1923) 502. –Type: BS 36565Ramos & Edaño (US*, iso). Luzon: Zamboanga District, Mt Tubuan, Oct 1919.MINDANAO: Zamboanga. Boulders along small streams.Ophiorrhiza inaequifolia Elmer, LPB 9 (1934) 3257. –Type:Philippines (Luzon). 42 PHI.Ophiorrhiza involucrata Elmer, LPB 1 (1908) 351; --Merr., EPFP 3 (1923) 501. –Type: Elmer 7603(not seen).LUZON: Apayao, Quezon, POLILLO. Low and medium altitde forests along streams,ascending to 750m.Ophiorrhiza lancilimba Merr., PJS 29 (1926) 491.BOHOL.Ophiorrhiza linearifolia Merr., PJS 10 c (1915) Bot. 99; --EPFP 3 (1923) 502. –Type: BS 16109Reillo (US*, iso). Basilan: Basilan prov., Comalarang, in forests, 29 Aug 1912.BASILAN. Lowland forests, apparently along streams.Ophiorrhiza liukiuensis Hayata, Icon. Pl. Formosan. 2 (1912) 89.Ophiorrhiza japonica var. kuroiwai (Makino) Ohwi, Acta Phytotax. Geobot. 7 (1938) 195; --Ophiorrhizakuroiwae Makino, Bot. Mag. Tokyo 20 (1906) 5, nom. nud.; --HJ Li & HY Liu, Fl. Taiwan ed.2, 4(1998) 311, pl. 141, photo 187.Ophiorrhiza mungos Merr., EPFP 3 (1923) 502, non L., Sp. Pl. (1753) 150Ryukyus, Taiwan (Lanyu and Lutao), PhilippinesBATAN. The plant is characteristically purplish when dry.Ophiorrhiza macgregorii Merr., PJS 17 (1920, publ. 1921) 437; --EPFP 3 (1923) 502. –Type: BS23024 McGregor (US*, iso). Luzon: Laguna prov., Jun-Aug 1915LUZON: Laguna, Quezon. Primary forests, altitude 200-300m.Ophiorrhiza maquilingense Elmer, LPB 9 (1934) 3258. –Type: Elmer 17536 (PNH, holo, lost;

MO*, NY*, US*, iso). Luzon: Laguna prov., Mt MakilingLUZONOphiorrhiza mungos L., Sp. Pl. (1753) 150; --Merr., EPFP 3 (1923) 502, excl. Batan plants.Sri Lanka, India, Nicobar Isls, Burma, Sumatra, Peninsular Malaysia, Java, ?Philippines.N LUZON to PALAWAN and MINDANAO. According to Merrill (1923), not certainlydistinct from the Linnean species. Plant is said to be characteristically purplish whendry.Ophiorriza oblongifolia DC, Prodr. 4 (1830) 415; --Merr., EPFP 3 (1923) 502. –Type:Ophiorrhiza triandra Blanco, Fl. Filip., ed. 2 (1845) 65.LUZON: Isabela to Sorsogon, MINDORO, PALAWAN, POLILLO, BILIRAN, PANAY,NEGROS, MINDANAO. In forests, chiefly along streams at low and medium altitudes.Ophiorrhiza oblongilimba Merr., PJS 12 c (1917) Bot. 164; --EPFP 3 (1923) 503. –Type: BS 27139RamosLUZON: Ilocos Norte. Lowland forests along streams.Ophiorrhiza ovata Merr., PJS 17 (1920, publ. 1921) 438; --EPFP 3 (1923) 503. –Type: BS 23163McGregorLUZON: Laguna, Quezon. Low and medium altitude forests, along rocky streamembankments.Ophiorrhiza pubescens Elmer, LPB 1 (1906) 68; --Merr., EPFP 3 (1923) 503. –Type: Elmer 14971LUZON: Laguna, Sorsogon, SAMAR, MINDANAO: Agusan del Norte. Open grasslandsalong streams at low and medium altitudes.Ophiorrhiza pubiflora Merr., PJS 17 (1920, publ. 1921) 438; --EPFP 3 (1923) 503. –Type: BS 29253Ramos & EdañoLUZON: Rizal. Forests, apparently at c. 1000m.Ophiorrhiza pumila Champ. ex Benth. in Hook. J. Bot. Kew Gard. Misc. 4 (1852) 169; --Chao, Fl.Taiwan 4 (1978) 319 --HJ Li & HY Liu, Fl. Taiwan ed.2, 4 (1998) 311, pl. 142.Japan, S China, Taiwan, Ryukyu Isls, Japan (Yakushima), Vietnam, ?Philippines.Ophiorrhiza pulgarensis Elmer, LPB 4 (1912) 1342; --Merr., EPFP 3 (1923) 503. –Type: Elmer12949PALAWAN. Forests, up to 450m.Ophiorrhiza sorsogonensis Elmer, LPB 9 (1934) 3260.LUZON.Ophiorrhiza stenophylla Merr., PJS 17 (1920, publ. 1921) 436; --EPFP 3 (1923) 494, erroneouslyas ‘Oldenlandia stenophylla’; non Ophiorrhiza stenophylla Valeton (1912). –Type: BS 22591 Ramos &Deroy.LUZON: Cavite. On rocks along streams, altitude c. 600m.Ophiorrhiza tenuis Merr., PJS 17 (1920, publ. 1921) 440; --EPFP 3 (1923) 503, nom. illeg., non Ridl.(1912). –Type: BS 29785 Ramos & EdañoLUZON: Rizal. Forests, apparently at c. 1000m.

Ophiorrhiza undulata Merr., PJS 8 c (1913) Bot. 46; --EPFP 3 (1923) 503. –Type: Weber 1445(US*, iso). Mindanao: Davao, Mt Binutun, on banks of small streams in bamboo forests, 26 Oct1911.MINDANAO: Zamboanga, Davao. Along small streams in bamboo forests, altitude c.550m.Ophiorrhiza venosa Merr., PJS 2 c (1907) Bot. 304; --EPFP 3 (1923) 503. –Type: Merrill 5628MINDORO, SIBUYAN, PANAY, SIARGAO. Lowland primary forests.Ophiorrhiza zambalensis Elmer, LPB 9 (1934) 3212. –Type:LUZON:Unplaced taxa:Ophiorrhiza bicolor Elmer, LPB 9 (1934) 3211, 3212, nom. illeg., non Craig (1932). –Type: Elmer21958 (PNH, holo, lost; iso MO*, NY*, US*). Luzon: Pampanga prov., Mt Pinatubo, Camp Stotsenburg,May 1927.Dubious taxa:Ophiorrhiza acuminata DC, Prodr. 4 (1830) 416; --Merr., EPFP 3 (1923) 501.“The type ia a specimen collected at Sorsogon, LUZON, by Haenke. I know the speciesonly by description. Endemic.” –Merrill, l.c.Oxyceros LoureiroOxyceros bispinosa (Griff.) Tirveng., Nordic J. Bot. 3 (1983) 466; --Randia bispinosa (Griff.) Craib,Fl. Siam. 2 (1932) 99; --Stylocoryna bispinosa Griff., Not. Pl. Asiat. 4 (1854) 260.Randia uncaria Elmer, LPB 1 (1906) 30; --Merr., EPFP 3 (1923) 529. –Type: Elmer 7001 (NY*, iso). Luzon:Bataan prov., Mt Mariveles, Nov 1904.Randia williamsii Elmer, LPB 3 (1911) 1004; --Merr., EPFP 3 (1923) 529. –Type: Elmer 12474 (not seen).Burma, Thailand, Vietnam, Borneo, Philippines.LUZON: Zambales, Bataan, Laguna, Camarines, Sorsogon, MINDORO,PALAWAN, SIBUYAN, PANAY, LEYTE, SAMAR, BASILAN, MINDANAO: Zamboanga.Low and medium primary forests.Paederia LinnaeusLiterature:Deb, DB. & K Roy. 1983. Taxonomic revision of Paederia L. (Rubiaceae) in India and adjoiningregions. J. Econ. Taxon. Bot. 4.Puff, C., E Robbrecht & L Andersson. 1991. Relationships and evolution in Paederia L.(Rubiaceae-Paederieae). A comparison classical approach, numerical evaluation andcladistic analysis. Opera Bot. Belg. 3: 337–358.Puff, C. 1991. Revision of the genus Paederia L. (Rubiaceae-Paederieae) in Africa andMadagascar. Opera Bot. Belg. 3: 293–322.Paederia foetida L., Mant. Pl. (1767) 52; --Blanco, Fl. Filip. (1837) 159; ibid. ed. 2 (1845) 112; ibid. ed.3, 1 (1877) 201, t. 54; --Fern.-Villar, Novis. App. (1880) 112; --Vidal, Sinopsis Atlas (1883) 29, t. 57,Fig. K; --Elmer, LPB 1 (1906) 28; --Merr., Interp. Herb. Amboin. (1917) 489; EPFP 3 (1923) 570.Paederia scandens (Lour.) Merr., Contrib. Arn. Arb. 8 (1934) 163.Paederia tomentosa Blume, Bijdr. (1826) 968; --Merr., Fl. Manila (1912) 455.Sri Lanka, India, E Himalayas, Nepal, Bangladesh, Burma, Tibet, China, Thailand,Laos, Cambodia, Vietnam, Hainan, Taiwan, Ryukyu Isls, Korea, N & C Japan, Sumatra,

Peninsular Malaysia, Java, Lesser Sunda Isls, Borneo, Philippines, Sulawesi, Moluccas,Christmas Is. Introduced in Mauritius, Reunion, Bismarck Archipelago, Hawaii andFlorida.Throughout the Philippines. Low and medium altitude thickets.Paederia verticillata Blume, Bijdr. (1826) 968; --Merr., EPFP 3 (1923) 570.Paederia verticillata Blume var. tomentella Elmer, LPB 5 (1913) 1885; --Merr., EPFP 3 (1923) 570.Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi.Pavetta LinnaeusRevisions:Bremekamp, CEB. 1935. A monograph of the genus Pavetta L. Repert. Sp. Nov. Reg. Veg. 37:1–208.Bremekamp, CEB. 1939. A monograph of the genus Pavetta L.: additions and emendations.Repert. Sp. Nov. Reg. Veg. 47: 12–28; part 2, 81–98.Pavetta basilanensis Bremek., Fedde, Repert. 37 (1934) 87. –Type:BASILANPavetta batanensis Bremek., Fedde, Repert. 47 (1939) 24. –Type: BS 3748 Fenix (GH*, iso).Batanes. May-Jun 1907.BATANPavetta brachyantha Merr., PJS 8 c (1913) Bot. 47; EPFP 3 (1923) 544. –Type: RS Williams 1077(US*, iso). Luzon: Benguet prov., Baguio, Jun 1904.LUZON: Mountain Province, Benguet. In ravines, altitude 1300-2000m.Pavetta cumingii Bremek., Fedde, Repert. 37 (1935) 109. –Type: Cuming 1322 (L*, NY*, iso). Luzon:Albay prov.LUZON:Pavetta dolichostyla Merr., PJS 1 (1906) Suppl. 239; EPFP 3 (1923) 544. –Type: FB 3714 MerrittMINDORO. Lowland forests.Pavetta elmeri Merr., PJS 17 (1920, publ. 1921) 475; EPFP 3 (1923) 544. –Type: Elmer 6394LUZON: Benguet. Thickets or forests, altitude c. 1400m.Pavetta indica L., Sp. Pl. (1753) 110; --Merr., EPFP 3 (1923) 544.var. glabrescens (Kurz) Deb & Rout, J. Bombay Nat. Hist. Soc. 89 (1992, publ. 1993) 351.Pavetta indica var. glabra Bremek., Repert. Spec. Nov. Regni Veg. 37 (1934) 119, nom. inval.Pavetta barnesii Elmer, LPB 1 (1906) 2. –Type: Elmer 6788 (NY*, iso). Luzon: Bataan prov., Mt Mariveles,Nov 1904.India, E Himalayas, Bangladesh, Nepal, S China, Hainan, Peninsular Malaysia, Borneo,Philippines.Throughout the Philippines in most islands and provinces. Lowland primaryforests, often common.Pavetta leytensis Bremek., Fedde, Repert. 47 (1939) 26. –Type: Wenzel 1731 (GH, holo?). 10 Apr1906.LEYTE:Pavetta luzonica Bremek., Fedde, Repert. 37 (1934) 109. –Type: BS 94 Adduru (A*, holo?). Luzon:

Cagayan prov., Peñablanca, Littoc, 28-31 May 1917.LUZON:Pavetta membranacea Blanco, Fl. Filip. (1837) 59; --Merr., EPFP 3 (1923) 545. –Type: BS 7634RamosPavetta sambucina Blanco, Fl. Filip., ed. 2 (1845) 41, nom. illeg.LUZON: Ilocos Norte, Bataan, Rizal. Low or medium altitude thickets or forests.Pavetta mindanaensis Bremek., Fedde, Repert. 37 (1934) 88. –Type: Elmer 11233 (NY*, iso).Mindanao: Davao del Sur prov., Mt Apo, Todaya, dry fertile soil of wooded flat lands at 1500ft, Jul 1909Pavetta parvifolia Vidal, Phan. Cuming. Philipp. (1885) 183; --Merr., EPFP 3 (1923) 545. –Type:Cuming 1394LUZON: Nueva Ecija, Zambales, MINDORO. Lowland thickets.Pavetta phanerophlebia Merr., PJS 17 (1920, publ. 1921) 478; EPFP 3 (1923) 545. –Type: BS 326BermejosPALAWAN. Low and medium altitude forests.Pavetta subferruginea Merr., PJS 17 (1920, publ. 1921) 477; EPFP 3 (1923) 545. –Type: Merrill 831PALAWAN. Lowland forests.Pavetta williamsii Merr., PJS 17 (1920, publ. 1921) 475; EPFP 3 (1923) 545. –Type: RS Williams2631MINDANAO: Davao, DINAGAT. Forests, altitude up to 900m.Pertusadina RidsdaleRevision:Ridsdale, C.E. 1978. A revision of the tribe Naucleeae s. s. (Rubiaceae). Blumea 24: 307–366.Pertusadina multifolia (Havil.) Ridsd., Blumea 24 (1978) 356; --Metadina multifolia (Havil.) Ridsd.,Gard. Bull. Sing. 25 (1970) 250; --Adina multifolia Havil., J. Linn. Soc. Bot. 33 (1897) 45; --Merr.,EPFP 3 (1923) 507.Philippines, Moluccas, New Guinea.LUZON, MINDORO, SIBUYAN, TICAO, SAMAR, MINDANAO. In forests at low andmedium altitudes.Petunga de Candolle =Philippine spp. now in Hypobathrum and Aidia, q.v.Plectronia Linnaeus =CanthiumPleiocarpidia SchumannRevisions:Bremekamp, CEB. 1940. A monograph of the genus Pleiocarpidia K. Sch. (Rubiaceae), RecueilTrav. Bot. Neerl. 37: 198–236.Bremekamp, CEB. 1940. A preliminary account of the Philippine species of Urophyllum Wall.,Pleiocarpidia K Schum., & Praravinia Korth. (Rubiaceae). Journal of the ArnoldArboretum 21: 40–41.

Pleiocarpidia lanaensis Merr., PJS 20 (1922) 462; EPFP 3 (1923) 524; --Bremek., J. Arnold Arb.21 (1940) 40. –Type: Clemens 882 (not seen). Mindanao: Lanao del Sur prov., Jan 1907.MINDANAO: Lanao del Sur, Bukidnon (Mt Candoon). In forests along streams at lowand medium altitudes, ascending to 800m.Praravinia KorthalsRevision:Bremekamp, CEB. 1940. A preliminary account of the Philippine species of Urophyllum Wall.,Pleiocarpidia K Schum., & Praravinia Korth. (Rubiaceae). Journal of the ArnoldArboretum 21: 41–47.Praravinia acuminata (Merr.) Bremek., J. Arnold Arb. 21 (1940) 43; --Urophyllum acuminatumMerr., PJS 1 (1906) Suppl. 129; EPFP 3 (1923) 521. –Type: Whitford 236 (NY*, US*,syntypes).Luzon: Bataan prov., Mt Mariveles, Lamao River, May 1904. The US sheet annotated by R.Buchner as holo.Praravinia streptopodium Elmer, LPB 1 (1906) 40, non Wall. ex Hook. (1880).LUZON: Mountain Province (Mt Masapilid), Isabela (Mt Moises), Nueva Ecija (MtUmingan), Rizal (Montalban; Mt Lumutan; Mt Irig (=Irid); Mt Canumay). In primary forest,500-1300m.Praravinia affinis (Merr.) Bremek., J. Arnold Arb. 21 (1940) 46; --Urophyllum affine Merr., PJS 17(1920) 481; EPFP 3 (1923) 521. –Type: BS 28716 Ramos & Edaño (A*, iso). Luzon: Quezonprov., Mt Binuang, on forested slopes, apparently at low altitudes, May 1917. Iso sheet in Asterile; a syntype cited in the original protologue is BS 28482 Ramos & Edaño (A*, staminate),alsocollected on Mt Binuang in May 1917.LUZON: Quezon (Mt Binuang; Mt Tulaog). According to Bremekamp, BS 10211McGregor (L) from POLILLO might belong here, but is much less hairy. In damp primaryforest, at low altitudes.Praravinia everettii Merr., PJS 10 c (1915) Bot. 107; --Bremek., J. Arnold Arb. 21 (1940) 44;--Williamsia everettii (Merr.) Merr., EPFP 3 (1923) 524. –Type: FB 7294 Everett (not seen),Negros:Negros Occidental prov., Mt Silay, on ridges in forests, altitude 700m, 9 May 1907.NEGROS: Negros Occidental (Mt Silay), Negros Oriental (Cuernos Mtns). Primaryforests, altitude c. 700m.Very near P. lucbanensis (Elmer) Bremek.Praravinia glabra (Merr.) Bremek., J. Arnold Arb. 21 (1940) 44; --Williamsia glabra Merr., PJS 10 c(1915) Bot. 108; EPFP 3 (1923) 524. –Type: BS 9484 Robinson (L). Luzon: Quezon prov.,Siniloantrail, in forests, altitude c. 200m, Aug 1909.LUZON: Nueva Ecija (Mt Umingan), Quezon (Siniloan trail), Laguna (San Antonio;unspecified locality for BS 23047 McGregor). Low and medium altitude primaryforests.Bremekamp also cites here a collection from ALABAT, BS 48249 Ramos & Edaño, withoblong leaves, probably representing an undescribed species.Praravinia loheri (Merr.) Bremek., J. Arnold Arb. 21 (1940) 45; --Williamsia loheri Merr., PJS 27(1925) 58; EPFP 4 (1925) 254. --Type: Loher 12309 (not seen). Luzon: Rizal prov., Montalban,Oct1909.

LUZON: Kalinga (Mt Masingit, Lubuagan), Quirino (Mt Alzapan), Quezon (Mt Binuang),Rizal (Montalban).According to Bremekamp, closely related to P. sablanensis (Elmer) Bremek.Praravinia longistipula (Merr.) Bremek., J. Arnold Arb. 21 (1940) 46; --Williamsia longistipulaMerr., PJS 17 (1920, publ. 1921) 485; EPFP 3 (1923) 524. –Type: Merrill 7287 (K, US*, iso).Mindanao: Agusan del Norte prov., Agusan River at Waloe, in damp forest, altitude c. 50m, 2Oct1910.MINDANAO: Agusan del Norte. Low altitude primary forest. Apparently known only fromthe type collection.Praravinia lucbanensis (Elmer) Bremek., J. Arnold Arb. 21 (1940) 43; --Urophyllum lucbanenseElmer, LPB 1 (1906) 71; --Merr., EPFP 3 (1923) 521, pro parte. –Type: Elmer 7945 (A, L*,NY*,US*, iso). Luzon: Quezon prov., Lucban, overhanging stream banks of wooded ravines,altitude 850m, May 1906.LUZON: Nueva Ecija (Mt Umingan), Rizal (Montalban; Mt Angilog), Laguna (SanAntonio), Quezon (Lucban), POLILLO, CATANDUANES. Low and medium altitudeprimary forests, usually along water courses.Praravinia microphylla (Merr.) Bremek., J. Arnold Arb. 21 (1940) 43; --Urophyllum microphyllumMerr., PJS 12 c (1917) Bot. 161; EPFP 3 (1923) 523. –Type: BS 26389 Ramos (US*, holo).Luzon: Nueva Ecija prov., Mt Umingan, in forest near the mountain summit, altitude 1000m, 20Aug. 1916.LUZON: Nueva Ecija (Mt Umingan). In primary forest, c.1000m. Known with certaintyonly from this locality. The altitude given for the type is way below that of the mountain’ssummit which is c. 1900m.A collection from Quirino prov., Mt Alzapan (BS 45581 Ramos & Edaño, A, staminate)is doubtfully referred here by Bremekamp, who also notes that it could perhaps be astunted P. acuminata (Merr.) Bremek.Praravinia mimica (Merr.) Bremek., J. Arnold Arb. 21 (1940) 44; --Williamsia mimica Merr., PJS26 (1925) 495; EPFP 4 (1925) 251. --Type: BS 38790 Ramos & Edaño (US*, holo; iso: A*). Mindanao:Bukidnon prov., Mt Candoon, damp forested slopes, altitude c. 1000m, Jun-Jul 1920.MINDANAO: Bukidnon (Mt Candoon). Forested slopes, altitude c. 1000m.Praravinia mindanaensis (Elmer) Bremek., J. Arnold Arb. 21 (1940) 44; --Williamsiamindanaensis Elmer, LPB 5 (1913) 1904; --Merr., EPFP 3 (1923) 524. –Type: Elmer 13561(US*,holo; iso: L*, MO*, NY*). Mindanao: Agusan del Norte prov., Mt Urdaneta (=Hilonghilong),Cabadbaran, Aug. 1912.?Williamsia caudata Merr., PJS 9 c (1914) Bot. 389. –Type: Wenzel 686 (A*, US*, iso). Leyte: Leyte prov.,Buenavista near Jaro, 18 May 1914.PANAY, BILIRAN, LEYTE: Leyte (Buenavista; Abuyog), Southern Leyte (Cabalian),SAMAR: Northern Samar (Catubig River), MINDANAO: Zamboanga, Lanao delSur (Lake Lanao, Camp Keithley), Davao (Mt Apo, Todaya), Davao Oriental (MtGalintan), Agusan del Norte (Mt Urdaneta, =Mt Hilong-hilong, Cabadbaran), Surigaodel Norte. Low and medium altitude primary forests, ascending to 900m.According to Bremekamp, specimens from outside Mindanao may necessitate thereinstatement of Merrill’s epithet caudata.Praravinia multinervia (Merr.) Bremek., J. Arnold Arb. 21 (1940) 46; --Williamsia multinerviaMerr., PJS 10 c (1915) Bot. 107; EPFP 3 (1923) 524. –Type: Merrill 8085 (L*, iso). Mindanao:Zamboanga, Sax River Mtns back of San Ramon, in damp forests, altitude 600-900m, 28 Nov

1911.BASILAN, MINDANAO:Zamboanga (Sax River Mtns), Zamboanga del Sur (Malangas).Primary forests, altitude 400-900m.Praravinia negrosensis (Merr.) Bremek., J. Arnold Arb. 21 (1940) 46; --Urophyllum negrosenseMerr., PJS 5 c (1910) Bot. 247; EPFP 3 (1923) 524. –Type: FB 5550 Everett (not seen).Negros:Negros Occidental prov., Himugaan River, in ravines, altitude c. 50m, 25 Oct 1906.NEGROS: Negros Occidental (Himugaan River). In primary forests at low altitudes. Inaddition, Bremekamp cites Herre 1140 from Negros Oriental (Cuernos Mtns, altitude1200-1800m), which comes very near to P. multinervia (Merr.) Bremek.Praravinia panayensis (Merr.) Bremek., J. Arnold Arb. 21 (1940) 46; --Williamsia panayensisMerr., PJS 17 (1920, publ. 1921) 484; EPFP 3 (1923) 524, non Urophyllum panayense Merr.,ibid.482. –Type: BS 31043 Ramos & Edaño (US*, holo; iso: A*, K). Panay: Capiz prov., Jamindan.Indamp forests along small streams, Apr-May 1918.PANAY: Capiz (Jamindan). Low altitude forests along streams. Three collections citedby Bremekamp, all from Jamindan .Praravinia pubescens (Quisumb. & Merr.) Bremek., J. Arnold Arb. 21 (1940) 47; --Williamsiapubescens Quisumb. & Merr., PJS 37 (1928) 208. –Type: BS 47275 Ramos & Edaño (A*, 2sheets,US*, iso). Luzon: Isabela prov., Mt Moises, altitude 1200m, Mar 1926. Note: Buchnerannotates theUS sheet as holo, contrary to statement in the original publication, i.e. in PNH).LUZON: Isabela (Mt Moises). Altitude 1200m. Aside from the type, two other collectionsfrom this locality: Clemens 16852 (not seen), and 17000 (K, NY*).Praravinia quadribracteolata (Merr.) Bremek., J. Arnold Arb. 21 (1940) 43; --Urophyllumquadribrateolatum Merr. PJS 17 (1920, publ. 1921) 483; EPFP 3 (1923) 523. --Type: BS28161Fenix (A*, holo). Luzon: Apayao prov., Ngagan, in damp forest, 8 May 1917.LUZON: Apayao (Ngagan), Quezon (Infanta-Siniloan Trail). In damp forest at low ormedium altitudes.Praravinia sablanensis (Elmer) Bremek., J. Arnold Arb. 21 (1940) 45; --Williamsia sablanensis(Elmer) Merr., PJS 3 c (1908) Bot. 165; --Elmer, LPB 3 (1911) 1000, excluding Mindanaospecimens cited; --Merr., EPFP 3 (1923) 524, as to specimens cited from Luzon only; --Urophyllumsablanense Elmer, LPB 1 (1906) 39. –Type: Elmer 6131 (US*, holo, fide Buchner in annot.;iso: NY*). Luzon: Benguet prov., Sablan, Apr 1904.LUZON: Abra (Mt Posuey, =Posooy), Benguet (Sablan), La Union (Tonglon, =Mt Sto.Tomas), Cagayan, Isabela (Palanan), Nueva Vizcaya, Rizal (Mt Angilog), Laguna,Quezon. Low and medium altitude forests.Praravinia stenophylla (Merr.) Bremek., J. Arnold Arb. 21 (1940) 44; --Williamsia stenophyllaMerr., PJS 27 (1925) 59; EPFP 4 (1925) 254. --Type: Loher sn, (not seen). Luzon: Nueva Vizcayaprov., Caraballo Mtns, Mar 1915.LUZON: Nueva Vizcaya (Caraballo Mtns). Habitat not known.Difference with P. glabra (Merr.) Bremek. is apparently slight. P. stenophylla hasnarrower and not so completely glabrous leaves.Praravinia triflora (Quisumb. & Merr.) Bremek., J. Arnold Arb. 21 (1940) 47; --Williamsia triflora

Quisumb. & Merr., PJS 37 (1928) 210. –Type: BS 45458 Ramos & Edaño (NY*, US*, iso).Luzon: Aurora prov., Casiguran, forested streams at low altitudes, 5 May 1925. Note: Buchnerannotates the US sheet as holo, contrary to statement in the original publication, i.e. in PNH).LUZON: Aurora (Casiguran). Low altitude forest. Apparently known only from the typecollection.Praravinia viridescens (Elmer) Bremek., J. Arnold Arb. 21 (1940) 43; --Williamsia viridescensElmer, LPB 9 (1934) 3215. –Type: Elmer 22078 (L*, K, MO*, NY*, US*, iso). Luzon:Pampanga prov., Mt Pinatubo, Camp Stotsenburg, altitude 1000m, pistillate, May 1927.LUZON: Pampanga (Mt Pinatubo), Bataan (Mt Mariveles).Prismatomeris ThwaitesRevisions:Johansson, JT. 1987. The revision of the genus Prismatomeris Thw. (Rubiaceae, Morindeae).Opera Botanica 94.Igersheim, A & E Robbrecht. 1993. The character states and relationships of thePrismatomerideae (Rubiaceae-Rubioideae). Opera Bot. Belg. 6: 61–79.Prismatomeris brachypus Ridl., Kew Bull. (1940) 602.E. Philippines.Prismatomeris obtusifolia Merr., PJS 26 (1925) 492; EPFP 4 (1925) 251. --Type: BS 41193 (A).Busuanga.BUSUANGA, PALAWAN. In thickets near the sea.Prismatomeris tetrandra (Roxb.) K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 4 (1891) 138; --Merr., EPFP 3 (1923) 572.Prismatomeris tetranda (Roxb.) K Schum. var. philippinensis Ridl., Bull. Misc. Inform. Kew 1939 (1940) 603.subsp. tetrandaIndia (Assam), E Himalayas, Bangladesh, S China, Hainan, Burma, Thailand, Vietnam,Philippines.LUZON (Sierra Madre Range from Cagayan to Quezon, Camarines), SAMAR,LEYTE, MINDANAO: Surigao. In primary forests at low and medium altitudes.Psilanthus Hooker filiusPsilanthus mabesae (Elmer) J-F Leroy, Bull. Mus. Natl. Hist. Nat., B, Adansonia 3 (1981, publ.1982) 256; --Canthium mabesae (Elmer) Merr., PJS 35 (1928) 8; --Coffea mabesae (Elmer) J-FLeroy, J. Agric. Trop. Bot. Appl. 9 (1962) 419; --Plectronia mabesae Elmer, LPB 8 (1919) 3099;--Merr., EPFP 3 (1923) 537. –Type: Elmer 17427 (NY*, US*, iso). Luzon: Rizal prov., Montalban,Mar 1917.Canthium lycioides A.Rich. ex DC., Prodr. 4 (1830) 475; --Plectronia lycioides (A.Rich. ex DC.) Elmer, LPB 1(1906) 28.Philippines, Lesser Sunda Isls, New Guinea, Queensland (Torres Strait Isls). LUZON:Ilocos Norte, Bataan, Rizal, Laguna, Batangas, PANAY. Lowland secondary forests andthickets.Psychotria LinnaeusRevision:Sohmer, SH & AP Davis. 2007. The Genus Psychotria (Rubiaceae) in the Philippine Archipelago.Sida, Botanical Miscellany No. 27: i-viii + 1-247.

Psychotria acuminatissima Elmer, LPB 1 (1908) 353; --Merr., EPFP 3 (1923) 552; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 228, Fig. 124. –Type: Elmer 8752 (US*, lecto; isolecto: A, K, MO,NY*, W). Luzon: Benguet prov., Baguio, limestone shrubbery below Antop barrio, altitude c.1000m,Mar 1907.Grumilea acuminatissima (Elmer) Merr., PJS 10 c (1915) Bot. 132.LUZON: Benguet (Baguio; Road to Asin, Mountain Trail =Halsema Road), Aurora(Mt Alzapan), Nueva Vizcaya (Caraballo Mtns), Rizal (Montalban). Shrubby forests,including those on limestone outcrops, altitude c. 1000m. Flowering: unknown; Fruiting:Jan, Mar, Dec.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1925.Psychotria alabatensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 142, Fig. 74. --Type: BS48006 Ramos & Edaño (K, holo; iso: A, BM, K, NY*, UC). Alabat: without further localitydata, Sept.-Oct. 1926.LUZON: Quezon (Umiray), ALABAT (Villa Norte to Barangay Bacung, Sitio Canumay),CATANDUANES (Mt Nagpakdit). Low elevation (including sea-level) moist forest.Flowering: Mar; Fruiting: May, Jun, Sept to Oct.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in Alabat in 1996.Psychotria alzapaniensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 123, Fig. 64. --Type:BS 45688 Ramos & Edaño (BM, holo; iso in K, NY, UC). Luzon: Aurora prov., MtAlzapan, on mossy forest slopes at c. 1600-1700m (‘5000 ft’), 20 May 1925.LUZON: Aurora (Mt Alzapan). Forest, including ‘mossy forest’, c. 1600-1700m.Flowering: Apr to Jun.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)], orpossibly Extinct [EX]. Only collected once in 1925.Psychotria amaracarpoides (Merr.) Merr., EPFP 3 (1923) 552; --Sohmer & AP Davis, Sida Bot.Misc. 27 (2007) 71, Fig. 34; --Grumilea amaracarpoides Merr., PJS 10 c (1915) Bot. 134. –Type:FB 8120 Curran & Merritt (US, lecto; isolecto: NY). Luzon: Zambales prov., Mt Tapulao (=HighPeak), Nov-Dec 1907.LUZON: Abra (Mt Posooy), Mountain Province (Mt Masapilid), Nueva Vizcaya (CaraballoMtns), Zambales (Mt Tapulao), Bataan (Mt Mariveles; Lamao River), Rizal (MtTocduanbanoy), MINDORO: Mindoro Occidental (Paluan), Unspecified locality for FB8638 Merritt. Primary forests, 1000-1600m. Flowering unknown; Fruiting: Jan-Mar., Jun,Oct.-Dec.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1926.Psychotria amplissima Merr., PJS 17 (1920, publ. 1921) 444; EPFP 3 (1923) 552; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 125, Fig. 65. –Type: BS 15198 Ramos (US*, lecto). Leyte: Leyteprov., Dagami, along streams in forests, 9 Aug. 1912.LEYTE (Dagami; unspecified locality for Wenzel 1628 & 1751), SAMAR: NorthernSamar (Catubig River), Western Samar (Mahagna, Tarabucan; Mt Sohoton, BabasianNa Daku River), MINDANAO: Agusan del Norte (Diuata Mtns), Surigao (Sukailung).Moist forests, sometimes near streams, low elevations (eg. 150m). Flowering: Aug;Fruiting: Feb ruary to Apr.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1970.

Psychotria arborescens Elmer, LPB 3 (1911) 1030; --Merr., EPFP 3 (1923) 552; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 94, Fig. 48; --Grumilea arborescens (Elmer) Merr., PJS 10 c(1915) Bot. 130. –Type: Elmer 11263 (US, lecto; isolecto: A*, BISH, BM, GH*, K, L*, MO,NY*, W). Mindanao: Davao del Sur prov., Mt Apo, Todaya, altitude 4000 ft, Aug 1909.BILIRAN (Mt Suiro), LEYTE: Southern Leyte (Cabalian), prov. unknown (Mt Abucayan),MINDANAO: Davao (Mt Apo, Todaya). Primary forests, altitude c. 1000-1200m. Possiblylower in Leyte. Flowering: unknown; Fruiting: Aug.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1954.Psychotria balabacensis Merr., PJS 10 c (1915) Bot. 127; EPFP 3 (1923) 553; --Quisumb., PJS 41(1930) 365, t. 27; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 115, Fig. 60. --Type: BS 21605Escritor (PNH, holo, lost; lecto in UC, isolecto: BM, P, US*). Balabac. Jan 1913.CAMIGUIN DE BABUYANES (Balatubat), MINDORO: Mindoro Oriental (Pinamalayan),PALAWAN (Bonabona), URSULA, BALABAC (Mt Caunayan), LUMBUCAN. Forests atlow altitude. Moist forest, low altitudes. Flowering: May, Sept; Fruiting: Sept.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1957.Psychotria banahaensis Elmer, LPB 1 (1906) 26; --Merr., EPFP 3 (1923) 553; --Sohmer & AP Davis, Sida Bot. Misc.prov., Mt Banahaw, Oct 1904.Grumilea banahaensis (Elmer) Merr., PJS 10 c (1915) Bot. 132.var. banahaensisLUZON: Cagayan (N of Bagio Cove; Isabela (San Mariano, Bgy. San Jose, SitioKapuntian; Palanan, Dicotcotan (=San Isidro), Diago; Mt Yalobo region), Aurora(Casiguran; San Luis, Bgy. Diteki), Nueva Ecija (Mt Umingan =Mingan), Rizal (Mt Angilo;Balacbac), Quezon (Mt Binuang; Lucban; Atimonan, Quezon National Park; Unisan;Guinayangan, Hinabaan; Tagkawayan, Bgy. Bagong Silang II; Mt Cadig), CamarinesSur (Sarapan; Botol River), Albay (Mt Masaraga), POLILLO, ALABAT, MINDORO:Mindoro Oriental (Mt Halcon), PALAWAN (Quezon, Tabon), MANGSEE, SIBUYAN (MtPunong, near trail to Barrio Jaosan; Magdiwang, Hawasan, Ating River; Mt Giting-giting,Camp 1 above Magdiwang on ridge leading to Mayos Peak), MASBATE (Milagros, Bgy.Matipuron, Sitio Kalunakan, 7-R Ranch), NEGROS: Negros Oriental (Calo), SAMAR:Western Samar (Mt Sohoton; Paranas, Bgy. Tinane, Sitio Campo Uno), MINDANAO:Surigao del Norte (Mt Kabatuan). In forests, including lowland dipterocarp forest, primaryor secondary (logged-over) forest, sometimes by streamsides, altitude (35-) 140-650m.Flowering: Feb, Mar; Fruiting: Feb to May, Aug to Nov.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1993.var. montalbanensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 188, Fig. 100. --Type:Jacobs 7846 (L*, holo; iso: PNH). Luzon: Aurora prov., Sierra Madre Mtns NNE of Dingalan,altitude 1000-1100m, 18 Mar 1968.LUZON: Aurora (Sierra Madre Mtns NNE of Dingalan), Rizal (Montalban). Forest,altitude 1000-1100m. Flowering: Unknown; Fruiting: Mar.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1968.Psychotria barkleyi Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 26, Fig. 8; --Grumilealongipedunculata (Elmer) Merr., PJS 10 c (1915) Bot. 133; EPFP 3 (1923) 562; --Psychotrialongipedunculata Elmer, LPB 3 (1911) 1027; non (Gardener) Muell.-Arg. (1881). –Type: Elmer12532-a (US*, lecto; isolecto: A*, BISH, BM, GH*, K, L*, MO, NY*, W). Sibuyan: Mt

Giting-giting, along Patos River in very humid, alluvial forest at 250m, May 1910.LUZON: Camarines Norte (Bicol National Park, NE), SIBUYAN (Mt Giting-giting)Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1994.Psychotria bataanensis Elmer, LPB 1 (1906) 23; --Merr., PJS 1 Suppl. 1 (1906) 134; EPFP 3(1923) 553; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 89, Fig. 45. –Type: Elmer 6980 (K,lecto; isolecto: NY*). Luzon: Bataan prov., Mt Mariveles, Nov 1904.Grumilea bataanensis (Elmer) Merr., PJS 10 c (1915) Bot. 132.Grumilea alvarezii (Merr.) Merr., PJS 10 c (1915) Bot. 132; --Psychotria alvarezii Merr., PJS 8 c (1913) Bot.56. –Type: FB 22189 Alvarez (not seen, presumably lost or destroyed). Luzon: Nueva Ecija prov.,Dec 1910.Grumilea rizalensis (Merr.) Merr., PJS 10 c (1915) Bot. 130; --Psychotria rizalensis Merr., PJS 8 c (1913)Bot. 57; EPFP 3 (1923) 562. –Type: Loher 6345 (K, lecto). Luzon: Rizal prov., Montalban, Jul 1905.LUZON: Nueva Ecija ((Mt Umingan; unspecified locality for FB 22189 Alvarez), Bataan(Mt Mariveles; Lamao River; Mt Kuyapo), Rizal (Mt Lumutan; Montalban). Montaneforest on ridges, 700-1600m. Flowering: Jan, Oct; Fruiting: Jan, May, Jul to Dec.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1947.Psychotria bayombongensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 203, Fig. 109.–Type: Vidal 3068 (K, holo; iso: A). Luzon: Nueva Vizcaya prov., Bayombong, Mar 1886.LUZON: Kalinga (Balbalan), Cagayan (Claveria), Nueva Vizcaya (Bayombong). Primaryforest, altitude unknown. Flowering: May; Fruiting: unknown .Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [EN B1ab(iii)] orpossibly Extinct [EX]. Last collected in 1948.Psychotria boholensis Merr. ex Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 131, Fig. 69.–Type: BS 43201 Ramos (PNH, holo; iso: BM, UC, W). Bohol: Bohol prov., Kalingohan(?), rocky forest, 29 Jan 1923.BOHOL: Bohol (Kalingohan; Bilar). Relatively dry or well-drained forest, includingsecondary forest; recorded on limestone, altitude 600-700m. Flowering: unknown;Fruiting: Jan, Mar, Aug, Oct.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1993.Psychotria bontocensis Merr., PJS 9 c (1914, publ. 1915) Bot. 456; EPFP 3 (1923) 553; --Sohmer& AP Davis, Sida Bot. Misc. 27 (2007) 166. –Type: Vanoverbergh 2610 (not seen). Luzon:Mountain Province, Bauco, along streams, altitude c. 1500m, 9 Mar 1913.LUZON: Kalinga (Mt Masingit, Lubuagan), Mountain Province (Bauco; Bontoc; MtMasapilid), Ifugao (Banawe, Batad), Benguet (Pauai), Rizal (Balacbac), ROMBLON(based on Elmer 12165, I suspect this was actually from Mt Giting-giting in Sibuyan),MINDANAO: Agusan (without further locality data, FB 24524 Miras & Oliveros). Montaneforests, in ravines and by streams, altitude 1300-1800m. Note: Pauai in Benguet is atleast 2200m. Flowering: Mar to Jun, Fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1967.Psychotria cadigensis Merr., PJS 12 c (1917) Bot. 175; EPFP 3 (1923) 553; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 23, fig. 5. –Type: BS 25484 Yates (US*, lecto, here designated).Luzon: Quezon prov., Mt Cadig, in forests, altitude 350-500m, Dec. 1916.LUZON: Quezon (Mt Cadig). Forest, altitude 350-500m. Flowering: Dec; Fruiting: Dec.The type number cited by Merrill in the original protologue was not seen by Sohmer& Davis, hence their designation of the syntype, BS 25533 Yates, as lecto (BM, lecto;

isolecto at K,P). A duplicate of the type number has been located at US.Conservation status (Sohmer & AP Davis, 2007): Extinct (EX). Known only from twocollections from Mt Cadig collected in 1916.Psychotria capizensis Merr., PJS 17 (1920, publ. 1921) 445; EPFP 3 (1923) 553; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 170, Fig. 90. –Type: BS 30657 Ramos & Edaño (A*, lecto;isolecto: K, US*). Panay: Capiz prov., Mt Madia-as, in forests, altitude not indicated, 13 May1918.PANAY: Capiz-Antique (Mt Madiaas). A recent collection, Busemeyer et al. 350 (PNH,CMNH, PUH, MU), is from a primary forest near the peak, altitude 1430m.probably at medium altitudes, otherwise unknown. Flowering: unknown; Fruiting: Apr-May.Conservation status: Critically Endangered [CR B1ab(iii)] or possibly Extinct [EX]. Knownfrom two collections, one of which was last collected in 1992.Psychotria cardiophylla Merr., PJS 17 (1920, publ. 1921) 446; EPFP 3 (1923) 553; --Sohmer & APDavis, SidaBot. Misc. 27 (2007) 164, Fig. 86. –Type: BS 29166 Ramos & Edaño (A*, lecto; isolecto: K, P).Luzon: Quezon prov., Infanta-Siniloan Trail, along small streams in forests, 12 Jun 1917.LUZON: Quezon (road between Infanta-Siniloan). Forests along streams, low altitudes.Flowering: unknown; Fruiting: Jun.Conservation status (Sohmer & AP Davis, 2007): Extinct [EX]. Known only from the typespecimen collected in 1917.Psychotria casiguraensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 208, Fig. 112. –Type:BS 45291 Ramos & Edaño (BM, holo; iso: K, NY*, P, US). Luzon: Aurora prov., Casiguran, alongstream in forest, 4 May 1925.LUZON: Aurora (Casiguran), Quezon (Infanta-Siniloan Trail). Low elevation moist forest,including streamside forests. Flowering: unknown; Fruiting: May, Jun.Conservation status (Sohmer & AP Davis, 2007): Extinct [EX]. Last collected in 1925.Psychotria castroi Merr. & Quisumb. ex Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 98,Fig. 50. --Type: PNH 5855 Castro & Añonuevo (L*, holo; iso: K). Samar: Eastern Samar prov.,Borongan, Pinangasan. Primary forest, altitude 150m, 23 Jun 1948.SAMAR: Eastern Samar (Borongan, Pinangasan; Mt Apoi). Low altitude primary forest,150m. Flowering: unknown; Fruiting: Jun.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1969.Psychotria catanduaniensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 184, Fig. 98.--Type: BS 75589 Ramos & Edaño (NY*, holo). Catanduanes: Masanganbahi, forested streams,altitude 520 ft, 21 Aug 1928.LUZON: Camarines Sur (Lagonoy), CATANDUANES (Masanganbahi), PANAY: Capiz(Hamanay?), Aklan (Libacao). Forests, including streamside forest, sea level to c. 160m.Flowering: unknown; Fruiting: Apr, May, Aug.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1928.Psychotria cephalophora Merr., PJS 3 c (1908) Bot. 438; EPFP 3 (1923) 554; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 60, Fig. 27; –Type: BS 4048 Fénix (US*, lecto, designated bySohmer, 2007). Camiguin de Babuyanes: in forests, 27 Jun 1907.Grumilea cephalophora (Merr.) Merr., PJS 10 c (1915) Bot. 133.CAMIGUIN DE BABUYANES, CATANDUANES, PANAY: Aklan (Libacao). Forest at low

and medium altitude. Flowering unknown; fruiting: Jun.Specimens identified as this species from Taiwan (eg. Chang 8721, K) are similar to thePhilippine species but are not conspecific.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)].Psychotria christarusselliae Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 39, Fig. 15, --Type: BS 43230 Ramos (P, holo; iso: A, BM, K). Bohol: without precise locality.BOHOL. Ecology not known. Flowering: unknown; Fruiting: Aug-Oct.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly extinct [EX]. Known only from the type specimen collected in 1948.Psychotria conglomeratiflora Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 120, Fig. 62. --Type: PNH 6306 Suliof Kansulaban Creek, in dipterocarp forest, altitude c. 200m, 26 Apr1948.SAMAR: Western Samar (Bagacay, bank of Kansulaban Creek), Eastern Samar(Borongan, Tagaslian). Primary dipterocarp forest, altitude 150-200m. Flowering:unknown; Fruiting: Apr to Jun.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly extinct [EX]. Last collected in 1948.Psychotria cordatula Merr., PJS 17 (1920, publ. 1921) 447; EPFP 3 (1923) 554; --Sohmer & APDavis, SidaBot. Misc. 27 (2007) 166, Fig. 87. –Type: BS 37854 Ramos & Edaño (A*, lecto; isolecto: K, US*).Luzon: Mountain Province, Mt Masapilid, mossy forest, altitude c. 1250m, 15 Mar 1920.LUZON: Mountain Province (Mt Masapilid), Ifugao (Banawe, Bayninan; Mt Hagada, Bgy.Pula). Mid-montane forest, inclusing secondary forest, altitude 1250-1600m. Flowering:unknown; Fruiting: Mar, Jul, Aug.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1992.Psychotria crassifolia Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 205; --Sohmer & AP Davis, SidaBot. Misc. 27 (2007) 155, Fig. 82. –Type from Borneo.Psychotria malayana auctt., non Jackvar. crassifoliaBorneo, Philippines. PALAWAN [Puerto Princesa (road bet. Puerto Princesa and Roxas,Km post 36; Mt Beaufort; Mt Pulgar; Bacungan; Malinao Mtn; Irawan,Tatanarum &Venture Area; Iwahig, Lapu-lapu; Iwahig River), Aborlan (Mt Iraan; Victoria Mtns),Quezon (Tabon), Mt Gantung; Mt Mantalingahan], MINDANAO: Bukidnon (vicinity ofTanculan), Davao Oriental (Mt Galintan). Forests, sometimes in secondary forest; oftenrecorded on rocks and on steep slopes; altitude (10-)250-650m. Flowering: Feb to Jun;Fruiting: Jan, Mar, May to Aug.Conservation status (Sohmer & AP Davis, 2007): Least Concern [LC]. Last collected in1996.Psychotria culionensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 157, Fig. 83. --Type: BS41319 Ramos (L*, holo; iso: K, PNH). Culion, no further data, Oct 1922.CULION. Ecology unknown. Flowering: Oct; Fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)], orpossibly Extinct [EX]. Only known from the type specimen collected in 1922.Psychotria crispipila Merr., PJS 1 (1906) suppl. 240; EPFP 3 (1923) 554; --Sohmer & AP Davis, Sida Bot. Misc. 27 (200Mountain Province, Mt Data, in mossy forest, altitude 2200m, Nov 1905.Grumilea crispipila (Merr.) Merr., PJS 10 c (1915) 133.Psychotria fusca (Merr.) Merr., EPFP 3 (1923) 556; --Grumilea fusca Merr., PJS 12 c (1917) 171. –Type: BS

26232 Ramos & Edaño (not seen). Luzon: Nueva Ecija prov., Mt Umingan (=Mingan), 14 Aug 1916.LUZON: Abra (Mt Posuey), Mountain Province (Mt Data; Mt Pukis), Ifugao (Mt Polis),Benguet (Mt Tabayoc (=Tabeyo); Mt Pulag; Mt Libung; Bontoc Trail (=Halsema Road),between km post 88 to 59; Pauai; Baguio; Mt Tonglon (=Mt Sto.Tomas); Libo-libo (Loher1514); without further locality data (BS 5581 Ramos), Aurora (Sierra Madre WSW ofBaler), Nueva Ecija (Mt Umingan =Mingan), Rizal (Bosoboso), Camarines Sur (MtPotianay), PANAY: Antique (Mt Madia-as, Culasi, Hangod Tubig). Montane forests,including drier types, chiefly between 1695-2500m, much lower elevation in Rizal prov.Flowering: Mar, Jul, Nov; Fruiting: Jan to Mar, Jun, Sept, Oct.Conservation status (Sohmer & AP Davis, 2007): Near Threatened (NT). Last collectedin 1992.Psychotria cuernosensis Elmer, LPB 2 (1908) 521; --Merr., EPFP 3 (1923) 554; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 45, fig. 18. –Type: Elmer 10076 (L, lecto; isolecto: A, BISH, BM,GH, K, MO, NY, P, US, W). Negros: Negros Oriental prov., Cuernos Mtns, mossy forests at c.1500m, May 1908.Psychotria mearnsii Merr., PJS 10 c (1915) 126. –Type: FB 4770 Mearns & Hutchinson (US*, lecto). Mindanao: Misamis OcMINDORO: Mindoro Oriental (north coast, Subaan River headwaters, inland from SanTeodoro; Mt Halcon; Mt Ilong), PANAY: Antique (Mt Madiaas; Mt Baloy; unspecifiedlocalities), Capiz (Mt Bulilao), NEGROS: Negros Occidental (Mt Canlaon; Mt Malbug),Negros Oriental (Matangwa River; Cuernos Mtns), LEYTE (unspecified locality),MINDANAO: Misamis Occidental (Mt Malindang), Bukidnon (Mt Candoon; Mt Kitanglad),Davao del Sur (Mt Apo), South Cotabato (Mt Apo, Kidapawan side), Agusan (MtMalaampayan), Surigao del Norte (Kabatuan). Primary and secondary forest, includinghigh altitude ‘moss forest’, altitude 500-2000m.Conservation status (Sohmer & AP Davis, 2007): Least concern (LC). Last collected in1992.Psychotria cuneata Elmer, LPB 9 (1934) 3261; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 63,Fig. 29, --Type: Elmer 17170 (UC, lecto; isolecto in A, BISH, BM, GH, K, L*, NA*, NY*, MO, P,1916.LUZON: Sorsogon (Mt Bulusan), TABLAS (San Andres, Mt Pan-ayong). Forests,including disturbed, ravines and ridges, recorded on limestone at 450m altitude inTablas.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1996.PNH, US). Luzon: SorsPsychotria diffusa Merr., PJS 1 (1906) suppl. 134; EPFP 3 (1923) 554; --Sohmer & AP Davis, SidaBot. Misc. 27 (2007) 52, Fig. 22. –Type: Elmer 6811 (NY*, lecto; Isolecto in K). Luzon: Bataanprov., Mt Mariveles, Nov. 1904.Grumilea microphylla (Elmer) Merr., PJS 10 c (1915) 130; --Psychotria microphylla Elmer, LPB 2 (1908)522; --Merr., EPFP 3 (1923) 559. –Type: Elmer 9505 (L*, lecto; isolecto: A, BISH, BM, K, MO, NY*,US*, W). Negros: Negros Oriental prov., Cuernos Mtns. Moss-covered forests at 4500ft, Mar 1908.var. diffusaWidely distributed throughout the Philippines. LUZON: Mountain Province (Mt Data; MtPolis), Ifugao (Banawe), Benguet (Lepanto; Mt Lusung; Sablan; Baguio; Mt Sto Tomas),Cagayan (Baguio Cove), Isabela (Mt Moises; Sierra Madre at Sangcad near Palanan),Quirino (Mt Alzapan), Bataan (Mt Mariveles), Rizal (Mt Irid; Mt Lumutan; Montalban;Rolay; Oriud; Angilog), Laguna (Paete), Quezon (Infanta; Real, National BotanicGarden; Mt Banahaw; Lucban; Tagkawayan), Camarines Sur (Mt Potianay; Mt Isarog),Sorsogon (Castilla; Mt Bulusan), POLILLO, CATANDUANES, MINDORO: MindoroOccidental (Mt Calavite), Mindoro Oriental (Mt Halcon), PALAWAN, PANAY: Antique-Capiz (Mt Madia-as), NEGROS: Negros Occidental (Mt Kanlaon National Park), Negros

Oriental (Cuernos Mtns), BILIRAN, LEYTE: Leyte (Mt Mamban), SAMAR: WesternSamar (Mt Sohoton), MINDANAO: Davao Oriental (Mt Galintan). Humid forest, includingsecondary forest, altitude 500-2500m.Conservation status (Sohmer & AP Davis, 2007): Near Threatened (NT). Last collectedin 1996.var. agusanensis (Elmer) Merr., PJS 10 c (1915) Bot. 123; EPFP 3 (1923) 555; --Sohmer & APGH, K, MO, NY, P, UC, US, W). Mindanao: Agusan del Norte prov., Mt Urdaneta (= MtHilong-hilong), Cawilanan Peak, wind-swepy forested ridge at c. 1300m, Sept 1912.Psychotria ovalis Elmer, LPB 3 (1911) 1023; --Merr., EPFP 3 (1923) 560. –Type: Elmer 10567 (L, lecto;isolecto: A, BISH, BM, GH, K, MO, NY, US, W). Mindanao: Davao del Sur prov., Mt Apo, Todaya,dense ridgeforests S of Mt Calelan, altitude c. 1400m, May 1909.LUZON: Camarines Norte (Sipocot National Park), Albay (Mt Malinao, Tiwi side),MINDANAO: Bukidnon (Mt Kitanglad; Tangculan), Agusan del Norte (San Mateo, sitioTungao), Davao Oriental (Mt Kampalili). Forests, altitude 350-1800m.Conservation status (Sohmer & AP Davis, 2007): Near Threatened (NT). Last collectedin 1995.Davis, Sida Bot. Misc.var. cervina Elmer, LPB 2 (1908) 592; --Merr., EPFP 3 (1923) 555; --Sohmer & AP Davis, Sida Bot.Misc. 27 (2007) 55, Fig. 24. –Type: Elmer 9800 (A, lecto; isolecto: BM, K, L, MO, NY, US, W).Negros: Negros Oriental prov., Cuernos Mtns, in forest of wind-swept ridge at c. 1300m, Apr1908.NEGROS: Negros Oriental (Cuernos Mtns). Only known from the type collection.Conservation status (Sohmer & AP Davis, 2007): Extinct (Ex). Only collected once in1908.Psychotria dingalanensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 171, Fig. 91. --Type:Jacobs 7654 (K, holo; iso L*). Luzon: Aurora prov., Sierra Madre Mtns NNE of Dingalan, primarydipterocarp forest, altitude 300-400m, 12 Mar 1968.LUZON: Aurora (Sierra Madre Mtns of Dingalan), primary dipterocarp forest, altitude300-400m. Flowering: Mar; Fruiting: Mar.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)],or possibly Extinct [EX].Psychotria elliptifolia Elmer, LPB 1 (1906) 66; --Merr., EPFP 3 (1923) 555; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 47, Fig. 19. –Type: Elmer 7943 (A, lecto; isolecto: K, L, NY, US, W).Luzon: Quezon prov., Mt Banahaw, wet mossy region at 1600m, Apr. 1906.Distribution?Conservation status (Sohmer & AP Davis, 2007): Vulnerable (VU B1ab(iii)). Lastcollected in 1994.Psychotria elliptilimba Merr., PJS 17 (1920, publ. 1921) 448; EPFP 3 (1923) 555; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 92, Fig. 46. –Type: BS 30534 Ramos (US, lecto; isolecto: A*, K, P).Catanduanes: Sto. Domingo River on low elevation forested slopes, 3 Dec. 1917.CATANDUANES. Only known from the type collection.Conservation status (Sohmer & AP Davis, 2007): Extinct (Ex). Only collected once in1917.Psychotria elmeriana Hochr., Candollea 5 (1934) 270; --Sohmer & AP Davis, Sida Bot. Misc. 27non Krause (1909). –Type: Elmer 14199 (A, lecto; isolecto: BISH, BM, GH, K, L, MO, NY, P, UC,US, W). Mindanao: Agusan del Norte prov., Mt Urdaneta (= Mt Hilong-hilong). Wet, stony forestedslopes at c. 1800m, Oct. 1912.MINDORO: Mindoro Oriental (Laylay, 10 km from San Teodoro), MINDANAO: Agusan(2007) 41, Fig. 17; --Ps

del Norte (Mt Urdaneta, =Mt Hilong-hilong). Forest, c. 1800m.Conservation status (Sohmer & AP Davis, 2007): Vulnerable (VU B1ab(iii)). Lastcollected in 1985.Psychotria fenicis Merr., PJS 17 (1920, publ. 1921) 449; EPFP 3 (1923) 555; --Sohmer & AP Davis,Sida Bot.Misc. 27 (2007) 176, Fig. 94. –Type: BS 29914 Fenix (US*, lecto; Isolecto: L*). Luzon: Pangasinanprov., Labrador, Mt San Isidro Nov. 1917.LUZON: Pangasinan (Labrador, Mt San Isidro), Zambales (Mt Tapulao =High Peak;Masinloc, Coto Mines; San Marcelino, Mt Aglao; Mt Pinatubo; Subig =Subic), Bataan (MtNatib: slopes at Tawawa; Bagac side of Mt Natib, Bgy. Banawang, Sitio Ulingan; withoutfurther locality data for FB 23223 Alhambra & Canlas). Dry forest or well drained ridgeforest, including secondary forest; recorded on rocky soils, altitude 200-400m. Flowering:May, Nov; Fruiting: Apr, Nov-JanConservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1992.Psychotria foxworthyi Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 145, Fig. 76. --Type: BS657 (NY*, holo; iso GH, K). Palawan: without further data, Mar-Apr 1906.PALAWAN. Ecology unknown. Flowering: Mar, Apr; Fruiting: Mar, Apr.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [EX]. Known only from type specimen collected in 1906.Psychotria frakei Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 147, Fig. 78. --Type: PNH 38038Frake (A, holo; iso L*). Mindanao: Zamboanga del Norte prov., locality not legible, 2 Jan 1958.MINDANAO: Zamboanga del Norte (Tulaln; Dikus). Primary or secondary forests,altitude 300-600m. Flowering Jan; fruiting: Nov-JanConservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1958. Three collections cited by Sohmer & Davis.Psychotria galintanensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 188, Fig. 101. --Type:BS 48899 Ramos & Edaño (UC, holo; iso NY*). Mindanao: Davao Oriental prov., Mt Galintan, onforest slopes at 1000m, 13 Jun 1927.MINDANAO: Davao Oriental (Mt Galintan). Forest, low to medium elevation, eg. c.1000m. Flowering: unknown; Fruiting: Jun.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [EX]. Only known from the type specimen collected in 1927.Psychotria gitingensis Elmer, LPB 3 (1911) 1024; --Merr., EPFP 3 (1923) 556; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 157, Figs. 84, 85. --Type: Elmer 12431 (BM, lecto; isolecto: A,BISH, GH, K, L*, MO, NY*, P, US*, W). Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), onforested ridge, altitude 2750 ft, May 1910.Grumilea similis (Elmer) Merr., PJS 10 c (1915) Bot. 134; --Psychotria similis Elmer, LPB 3 (1911) 1025; --Merr., EPFP 3 (1923) 563; –Type: Elmer 12439 (GH*, lecto; isolecto: BISH*, L*, NY*, US*).Sibuyan: Magallanes (=Magdiwang), Mt Giting-giting, ‘in red soil compressed in between rocks of awind-swept forested ridge at 3000ft, May 1910.Grumilea versicolor (Elmer) Merr., PJS 10 c (1915) Bot. 130; --Psychotria versicolor Elmer, LPB 4 (1912)1352; --Merr., EPFP 3 (1923) 564. –Type: Elmer 12763 (A*, lecto; isolecto: BISH, BM, GH, K, L*,MO, NY*, P, US*, W). Palawan: Palawan prov., Puerto Princesa, Mt Pulgar (=Thumb Peak), banksof Iwahig River, Mar 1911.Psychotria lucida Merr., PJS 17 (1920, publ. 1921) 452. –Type: BS 29716 Ramos & Edaño (US*, lecto;isolecto: A*, K, NY, P). Luzon: Rizal prov., Mt Lumutan, Jul 1917.Psychotria malayana auctt., non JackPhilippines and possibly Borneo. CALAYAN, LUZON: Ilocos Norte (Bangui to Claveria),Mountain Province (as former Lepanto District for Vidal 1488), Cagayan (Baguio

Cove), Pangasinan (Bolinao, Luciente I), Nueva Ecija (Caraballo Mtns), Aurora (Baler,Cemento, on road c. 8 km from Baler), Rizal (Mt Angilog; Mt Irid; Mt Batay; Mt Lumutan;Montalban), Batangas (no further locality data for Cuming 1526), Quezon (QuezonNational Park, Pinagbanderaan Trail), Camarines Norte (Paracale), Albay (no furtherlocality data for Vidal 3054), POLILLO, MINDORO: Mindoro Occidental (Mt Iglit-Baco National Park, Calintaan, Barangay Poypoy, Sitio Sapaton), Mindoro Oriental(Puerto Galera; Mt Halcon; Pinamalayan; Bongabon and Pinamalayan; Mt Yagaw,E slopes), PALAWAN [Puerto Princesa (Irawan River valley; base of Mt Beaufort,Tatanarom), Aborlan (Victoria Mtns; Trident Mining area), Brookes Point, Pulot), MtGantung; Mt Mantalingahan], SEMIRARA, TABLAS (San Andres, Bgy. Marigondon Sur,Sitio Barkwang); SIBUYAN (Mt Giting-giting; Mt Punong; Jaosan-Punong River trail;Magdiwang, SE of Binalabag), MASBATE (Masbate, Milagros, Bgy. Matipuron, SitioKalunakan, 7-R Ranch), PANAY: Antique (Mt Madia-as; San Remigio; unprecise localityfor BS 32511 McGregor), Magosolon Aklan (Libacao; Mt Timbaban; Mt Macosolon=Magosolon), Capiz (Mt Bulilao), Iloilo (Mt Salibongbong; Miag-ao), GUIMARAS(Buenavista, Bgy. Tubod), DANJUGAN (off Cauayan, Negros Occidental prov.), CEBU(Alcoy, Manguera Forest; Mantalongon; Carmen, Corte, Malac), LEYTE: Leyte (Palo; MtAbucayan; unprecise locality for Wenzel 1297, 1361, 1376 & 1416), SAMAR: provinceuknown (Quinay, for Vidal 1481c), Western Samar (Basey, Bgy. Hinamok, Pinabacdao,near Lakes, vicinity of km 56 and Bgy. Cambayan; Paranas, Bgy Lokilokon, SitioKapinyahan; San Juanico), Eastern Samar (Balangiga, Bgy. I, Sitio Cantinoc), TAWI-TAWI (Birad-dali; Languyan), JOLO (Mt Dajo), BASILAN, MINDANAO: Zamboanga(without further locality data, Merrill 11621), Misamis (without further locality data, FB17986 Miranda), Agusan del Norte (without further data, FB 20734 Rafael & Ponce),Surigao del Sur (Adlay), Surigao (unprecise localities for BS 34616 & 36416 Ramos &Pascasio), DINAGAT (Loreto; Cambinliw, Santiago), PONEAS ISLET (near SIARGAO),BUCAS GRANDE (N end, Bgy. Salug, Sitio Lawigan). Moist forest, including beach andmangrove forest, and secondary forest, sometimes on cultivated lands, on well-drainedsoils and limestone, altitude 0-1000m. Flowering: Jan to Sept; fruiting all year round.Conservation status (Sohmer & AP Davis, 2007): Least Concern (LC). Last collected in1996.Psychotria gracilipes Merr., PJS 8 c (1913) 52; EPFP 3 (1923) 556; --Sohmer & AP Davis, Sida Bot.Misc. 27 (2007) 226, Fig. 123;–Type: Weber 1572 (A*, lecto; isolecto: GH*, K, P, US*). Luzon: Cagayan prov., Abulug River, rocky,forested hillside on limestone, altitude unknown, 5 Feb 1912.Grumilea gracilipes (Merr.) Merr., PJS 10 c (1915) Bot. 132.Psychotria isarogensis (Merr.) Merr., EPFP 3 (1923) 556; --Grumilea isarogensis Merr., PJS 10 c (1915) Bot. 136. –Type: BS 15prov., Mt Isarog, mossy forest, altitude unknown, 29 Nov 1913.LUZON: Cagayan (unspecified locality for FB 14776 Darling), Quirino (Mt Alzapan),Camarines Sur (Mt Isarog), Sorsogon (unspecified locality for BS 23700 Ramos),BILIRAN (Mt Suiro), CAMIGUIN, MINDANAO: Davao Oriental (Mt Mayo; Mt Kampalili),Agusan del Norte (Mt Urdaneta (=Mt Hilong-hilong), Cabadbaran). Moist to wet forest,altitude c. 700-1800m. Flowering: Mar, Apr; Fruiting: Jan, Apr to Aug, Oct.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1988.Psychotria hilonghilongensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 220, Fig. 119;--Type: PNH 10760 Mendoza & Convocar (A, holo; iso L*). Mindanao: Agusan del Norte prov., MtHilong-hilong, Cabadbaran, mossy forest ridge, altitude 1050m, 29 Apr 1949.MINDANAO: Agusan del Norte (Mt Hilong-hilong), in mossy forest, altitude 1050m.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].

Only known from the type specimen collected in 1949.Psychotria ilocana (Merr.) Merr., EPFP 3 (1923) 556; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 220, Fig. 120; --(K, lecto; isolecto: A*, NY*, US*). Luzon: Ilocos Norte prov., Bangui, low altitude forest, Feb 1917.LUZON: Ilocos Norte (Bangui; Burgos), Zambales (without further locality data, for BS4704 Ramos). Dry forest, low elevation. Flowering: unknown; Fruiting: Feb, Mar.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [EX]. Last collected in 1918.Psychotria iridensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 134, Fig. 70. --Type: BS48443 Ramos & Edaño (A, holo; iso: BM, K, NY*, UC). Luzon: Rizal prov., Mt Irid, on damp forestslopes, 13 Nov. 1926.LUZON: Ifugao (Mt Bobor, Barangay Batad), Rizal (Mt Angilo; Mt Irid; unspecifiedlocality for Loher 6402). Forests, including disturbed and possibly secondary forest,1000-1700m. Flowering: Apr; Fruiting: Feb, Apr, Nov.Conservation status (Sohmer & AP Davis, 2007): Vulnerable (VU B1ab(iii)). Lastcollected in 1994.Psychotria iwahigensis Elmer, LPB 4 (1912) 1351; --Merr., EPFP 3 (1923) 556; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 31, Fig. 11; --Type: Elmer 13052 (NY, lecto; isolecto: BM, L).Palawan: Mt Pulgar, in forests at c. 100m, Apr 1911.PALAWAN (Cleopatra Ranges, Barangay Tanabag, Sitio Calabayog; Mt Pulgar; Rizal,Barangay Ransang, Balen-balen). Low altitude humid forests c. 100m. Floweringunknown; fruiting Apr, Jun.Conservation status (Sohmer & AP Davis, 2007): Vulnerable (VU B1ab(iii)). Lastcollected in 1996.Psychotria ixoroides Bartl. ex DC, Prodr. 4 (1830) 522; --Merr., EPFP (1923) 556; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 41, Fig. 17. --Type: Haenke, sn. (PR, holo). Luzon: without preciselocality.LUZON: Ilocos Norte (Burgos), Isabela (Palanan Bay), Pangasinan (Mt San IsidroLabrador), Nueva Ecija (Mt Umingan), Bataan (Mt Mariveles; unspecified locality),Bulacan (Angat; Norzagaray), Rizal (Mt Susong Dalaga; Mt Irid; Tanay, San Isidro;unspecified locality), Quezon (Atimonan, Quezon National Park; Malikboy; Pagbilao;Tagkawayan; Mt Cadig), Camarines Sur (vicinity of Bicol National Park; Mt Asog), Albay(unspecified locality), Sorsogon (Mt Bulusan and Bulusan Lake; Casiguran; Castilla),POLILLO, ALABAT, CATANDUANES (Simamla), MINDORO: Mindoro Occidental(Paluan), Mindoro Oriental (Inland from San Teodoro, at and above Subaan Reserve;Puerto Galera), PALAWAN (Taytay, Lake Manguao; Puerto Princesa; Mt Victoria,Aborlan, Mt Iraan; Quezon, Bundog near Sumindap River; San Antonio Bay, BrookesPoint, lower slopes of Mt Mantalingahan; Rizal, Ransang, Madubis; BALABAC (PasigBay; Panaan), PANAY: Antique (Valderrama, lower slopes of Mt Baloi), Aklan (Libacao),CEBU: Cebu (Alcoy, Manguera Forest), BILIRAN (Mt Suiro; Naval, Libtong), LEYTE:Leyte (Palo; Dagami, Mt Top), SAMAR: Eastern Samar (Balangiga), Western Samar(Paranas, Campo Uno), MINDANAO: Davao Oriental (Mt Galintan), Surigao (unspecifiedlocality; Sukailang); Surigao del Sur (Hinatuan), DINAGAT (Mabini, Tubajon; unspecifiedlocality), SIARGAO and adjacent PONIAS Islet. Disturbed vegatation generally likedlogged-over and secondary forest and coconut plantation, reported on limestone andlateritic soils, from sea-level to 700m. Flowering Jan-Jun; fruiting: Feb, Aug., Sept.Conservation status (Sohmer & AP Davis, 2007): Least Concern (LC). Last collected in1997.

Psychotria lagunensis (Merr.) Merr., EPFP 3 (1923) 556; --Sohmer & AP Davis, Sida Bot. Misc.27 (2007) 174, Fig. 92; --Grumilea lagunensis Merr., PJS 12 c (1917) Bot. 169. --Type: BS 23826Ramos (K, lecto; Isolecto: US*). Luzon: Laguna prov., San Antonio, in damp forest, 19 Oct 1915.LUZON: Quirino (Mt Alzapan), Rizal (Montalban; Mt Susong Dalaga; Mt Lumutan; MtIrid), Laguna (Majayjay; San Antonio), Quezon (Mt Binuang; Tagkawayan), CamarinesNorte (Paracale), Camarines Sur (Sarapan), POLILLO (Anibawan), ALABAT,CATANDUANES (no further locality data, 3 collections). Moist forest and forestedslopes, altitude 70-1532m. Flowering: Mar; Fruiting: Jun to Dec.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [BU B1ab(iii)]. Lastcollected in 1928.Psychotria lanaensis (Merr.) Merr., EPFP 3 (1923) 557; --Sohmer & AP Davis, Sida Bot. Misc. 27(2007) 52; --Grumilea lanaensis Merr., PJS 10 c (1915) Bot. 136, --Type: Clemens 924 (not seen).Mindanao: Lanao del Sur prov., Camp Keithley. Swamps at c. 750m, Jun 1907.LUZON: Ifugao (Banawe: Bayninan), MINDORO: Mindoro Occidental (Mt Calavite),SIBUYAN: Mt Giting-giting, DINAGAT: Krominco, Mt Redondo, open pit mine. Primaryand disturbed forests, including swamp forest, altitude 650-700m.Conservation status (Sohmer & AP Davis, 2007): Near Threatened. Last collected in1991.Psychotria lancilimba Merr., PJS 17 (1920, publ. 1921) 451; EPFP 3 (1923) 557; --Sohmer & APDavis, SidaBot. Misc. 27 (2007) 103. --Type: BS 29713 Ramos & Edaño (US, lecto). Luzon: Rizal prov., MtLumutan, Jul 1917.LUZON: Benguet (Baguio), Nueva Vizcaya (Caraballo Mtns), Zambales (without furtherlocality data, for BS 5018 Ramos), Rizal (Mt Irid; Mt Batay; Mt Lumutan; Mt Mapatag).Lower montane forest, altitude 600-1000m. Flowering: Jul; fruiting: Jul.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1926.Psychotria leptothyrsa Miq., Ann. Mus. Bot. Lugtd.-Bat. 4 (1868) 208; --Merr., EPFP 3 (1923)557.Coelospermum ahernianum Elmer, LPB 1 (1906) 3. –Type: Ahern 196 (US*, iso). Luzon: Camarines Surprov., Pasacao (Dalupaon), Jan 1902.DistributionPsychotria lianoides Elmer, LPB 3 (1911) 1029; --Merr., EPFP 3 (1923) 557; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 47, Fig. 20. --Type: Elmer 11645 (P, lecto; isolecto: A, BISH*, BM, GH, K,L*, MO, NA*, NY* 2 sheets, US, W). Mindanao: Davao del Sur prov., Mt Apo, Todaya. Humid forestalongBaruring River on trail to Talon at c. 1300m, Sept 1909.MINDANAO: Davao del Sur (Mt Apo). Humid forest, altitude 1200-1300m.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Only known from the type specimen collected in 1909.Psychotria linearis Bartl. ex DC, Prodr. 4 (1830) 522; --Merr., EPFP (1923) 557; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 58, Fig. 25. --Type: Haenke sn (PR, holo). Luzon: “in ins. Manilaprope Sorzogon”.Psychotria depauperata Merr., PJS 12 c (1917) Bot. 174; EPFP 3 (1923) 554. –Type: Merrill 10487 (US,lecto; isolecto: A, BM, K, L, NY, P, PNH). Alabat: back of Sangirin on dry ridges in virgin forest areaat c. 100m, 23 Dec. 1916.Amaracarpus longifolius Elmer, LPB 1 (1906) 1. –Type: Merrill 4000 (PNH?, holo; iso US*). Luzon: Quezonprov., Atimonan, Mar 1905.LUZON: Nueva Ecija, Rizal (Mt Susong Dalaga; Mt Irid; Tanay, San Isidro), Quezon(Mt Pular; Mauban; Atimoman), Camarines (no further locality data), ALABAT, SAMAR:

Western Samar (Mt Sohoton; Basey). Relatively dry ridge forest, altitude 100-500m.Some of the cited localities are in predominantly limestone areas. Fl. Aug, Nov; fr. Jan,Mar, Aug-Oct, Dec.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1970.Psychotria loheri Elmer, LPB 1 (1908) 356; --Merr., EPFP 3 (1923) 557; --Sohmer & AP Davis, SidaBot. Misc. 27 (2007) 234, Fig. 128; –Type: Elmer 8584 (K, lecto; isolecto: A, L*, NY*, US*, W).Luzon: Benguet prov., Mt Sto Tomas, ‘dense oak woods at 7500 ft’, Mar 1907.Grumilea loheri (Elmer) Merr., PJS 10 c (1915) Bot. 132.Psychotria vanoverberghii Merr., PJS 9 c (1914, publ. 1915) Bot. 457; EPFP 3 (1923) 563. –Type:Vanoverbergh 2818 (US*, lecto; isolecto: K). Mountain Province, Bauco, in forests from 1300-1600m, Apr 1913.LUZON: Ilocos Norte (Mt Darna), Abra (unspecified locality for BS 7062 Ramos),Mountain Province (Bauco; Mt Data; unspecified locality for Vanoverbergh 2861),Benguet (Baguio; Mt Sto Tomas), Zambales (Sta Cruz, Acoje Mine concession area.Sta Cruz), LEYTE: Leyte (Buenavista near Jaro). Mostly in the higher montane sites,including mossy forests, altitude 1300-2200m. Presumably at considerably much lowerelevations in Zambales province and Leyte.Psychotria longipedicellata Elmer, LPB 1 (1906) 22; --Merr., EPFP 3 (1923) 557; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 63, Fig. 28. –Type: HN Whitford 907 (K, lecto; isolecto: NY*, P,US*). Luzon: Quezon prov., Lagumanoc, Sept 1904. –Syntype: FB 1727 WW Clark (NY*, US*).Burias, Jun 1904.Grumilea longipedicellata (Elmer) Merr., PJS 10 c (1915) Bot. 133.LUZON: Aurora (Mt Dingalan), Quezon (Lagumanoc), BURIAS (no further locality data).Low altitude forests. Flowering: Jun; Fruiting: Sept, Nov.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [(CR B1ab(iii)] orExtinct [Ex]. Last collected in 1910.Psychotria longissima Quisumb. & Merr., PJS 37 (1928) 206; --Sohmer & AP Davis, Sida Bot.Misc. 27 (2007) 179, Fig. 95. –Type: BS 45681 Ramos & Edaño (UC, lecto; isolecto: A, K, NY*,US*). Luzon: Aurora prov., Mt Alzapan, forested slopes, altitude c. 1300m, 7 Jun 1925.LUZON: Aurora (Mt Alzapan). Forests, altitude c. 1300m. Flowering: May, Jun; Fruiting:unknown.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [(CR B1ab(iii)] orExtinct [Ex]. Known only from the type specimen collected in 1925.Psychotria luzoniensis (Cham. & Schltdl.) Fern.-Villar, Novis. App. (1880) 157; --Merr., EPFP3 (1923) 558; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 201; --Coffea luzoniensis Cham. &Schltdl., Linnaea 4 (1829) 32. –Type: Chamisso sn, (LE, holo; iso: K). ‘Luçonia’.Grumilea luzoniensis (Cham. & Schltdl.) Merr., PJS 10 c (1915) Bot. 131, as ‘luçoniensis’.Psychotria tacpo (Blanco) Rolfe, J. Linn. Soc. (Bot.) 21 (1884) 312; --Paederia tacpo Blanco, Fl. Filip. (1837)160. –Type: Unknown.Widely distributed throughout the Philippines. LUZON: Ilocos Norte (Pagudpud, BgyDampig, Sitio Nalanus; Bangui; Mt Palimlim; Mt Pico de Loro; Nueva Era to MtSemenublan; Mt Burnay; Mt Bubungkayo en-route from Sto. Niño to Miner’s camp;unspecified locality for FB 13972 Merritt & Darling), Abra (Sallapadan, PoblacionCangal; Masissiat), Ilocos Sur, Kalinga (Lubuagan, Mabuntat), Mountain Province(Bontoc), Benguet (Sablang =Sablan; Trinidad to Kapangan; Baguio; between Baguio &Sablan; Tonglon =Mt Sto Tomas; La Union (Sta. Rita; Bauang), Cagayan (Mt Bawa; MtCaboloan; Peñablanca), Isabela (Ilagan to Echague), Nueva Vizcaya (Carasacas;vicinity of Dupax; unspecified locality), Pangasinan, Zambales (unspecified locality forBS 34666 Edaño; HERMANA MAYOR IS; Mt Pinatubo), Bataan (Limay; Mt Mariveles),

Pampanga (Arayat; Mt Pinatubo, Camp Stotsenburg), Rizal (Manila; Quezon City:Balara, Novaliches, & San Francisco del Monte; Pasig; Antipolo; Bosoboso; San Mateo(Comision Flora Forestal 1455), Montalban; Mt Mapatag; Mt Angilog; Mt Irid; MtPinagbayanan & San Andales =San Andres; Mt Susong Dalaga; Teresa, Kay Ungulan),Laguna (Los Baños; Mt Makiling; Calauan; Sta. Maria; Pagsanjan), Quezon (Infanta;Pagbilao; Guinayangan, Salakan), Cavite (Mt Palay-palay National Park; PalicpicanRiver; Pico de Loro), Batangas (Mt Maculot; Mataas na Kahoy; inland from MalabrigoPoint), Camarines Sur (Pili), Albay (Daraga), Sorsogon (Castilla; Mt Bulusan, Irosin),POLILLO, ALABAT (Bgy. Bacung, Sitio Canumay), MINDORO: unspecified prov. &locality for FB 19932 Amarillas; Mindoro Occidental (Abra de Ilog, S (?) Alatin; Iriron),Mindoro Oriental (Subaan River, inland from San Teodoro; Puerto Galera, Mt Minolo &Mahabang Parang Cove; Mt Halcon; Pinamalayan), PALAWAN (Quezon, Lipuun Peak;unspecified locality for BS 561 Foxworthy), MALAPACKUN, MARINDUQUE (Torrijos,Bgy. Dampulan, Sitio Maulawin, Hawilian), MASBATE (Bgy. Tugbo; also unspecifiedlocalities for Merrill 3395, FB 12563 Rosenbluth), SEMIRARA (Caluya, Pungtod),PANAY: Antique (unspecified locality for BS 32459 McGregor), Iloilo (Miag-ao;unspecified locality for BS 18068 Robinson), GUIMARAS (Buenavista), NEGROS:Negros Occidental (Luzuriaga), Negros Oriental (San Jose, Calo; Cuernos Mtns).Forest, including primary, disturbed, and secondary forest; recorded from limestone andwell-drained substrates, altitude 0- c. 2000m but mostly below 700m.Conservation status (Sohmer & AP Davis, 2007): Least Concern [LC]. Last collected in1996.Psychotria macgregorii Merr., PJS 5 c (1910) Bot. 389; EPFP 3 (1923) 558; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 223, Fig. 121;–Type: Merrill 6495 (published as 8495) (US*, lecto;isolecto: NY*). Luzon: Benguet prov., Mt Pulog (=Pulag), mossy forest above 2400m, May 1909.Grumilea macgregorii (Merr.) Merr., PJS 10 c (1915) Bot. 133.LUZON: Ifugao (Mt Polis; Sumigar, Mt Pimmage), Mountain Province (Mt Data; MtAmuyao, Barlig), Benguet (Mt Tabayoc =Tabeyo; Mt Pulag; Bokod, Lusod; Mt Pauai;Mt Tonglon =Sto.Tomas), Benguet-Nueva Vizcaya prov. borders (for FB 15841 Curran& Merritt), Nueva Ecija (Mt Umingan =Mingan); Rizal (Montalban, for Loher 1515),PALAWAN (Mt Manalsal; Mt Mantalingahan), DINAGAT (Mt Redondo, KROMINCO).Montane forest, including mossy forest, with one record from grassland; on ultramaficson Dinagat, altitude (?700-)1500-2800m. Flowering: Jan, Mar to Jun; Fruiting: Jan toJun, Sept.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1996.Psychotria madulidii Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 60, Fig. 26. –Type: BS78408 Edaño (A, holo). Luzon: Cagayan prov., Mt Cagua, Oct-Nov 1929.LUZON: Cagayan (Mt Cagua), PALAWAN (Mt Mantalingahan). Ecology unknown.Flowering unknown; Fruiting: Oct, Nov.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1929.Psychotria magnifolia Merr., PJS 17 (1920, publ. 1921) 453; EPFP 3 (1923) 558; --Sohmer & APDavis, SidaBot. Misc. 27 (2007) 120, Fig. 63. –Type: BS 28828 Ramos & Edaño (US, lecto; isolecto: A*, K, P).Luzon: Quezon prov., Mt Binuang, along small streams in damp forests, 11 May 1917.LUZON: Quezon (Mt Binuang), Rizal (precise locality not known, based on Loher 15011(UC). Moist forest, reported along streams, elevation unknown. Flowering: unknown;Fruiting: May.

Conservation status (Sohmer & AP Davis, 2007): Extinct [EX]. Last collected in 1917.Psychotria manillensis Bartl. ex DC. Prodr. 4 (1830) 522; --Merr., EPFP 3 (1923) 559; --Sohmer &AP Davis, Sida Bot. Misc. 27 (2007) 138, Figs. 73a, 73b. –Type: Haenke sn (PR, holo). Luzon:Sorsogon prov., without precise locality.Psychotria propinqua (Merr.) Merr., EPFP 3 (1923) 561; --Grumilea propinqua Merr., PJS 12 c (1917) Bot.170. –Type: BS 26570 Ramos & Edaño (K, lecto; isolecto: NY*). Luzon: Aurora prov., Mt Dingalan,27 Jan 1916.Psychotria ridsdalei Sohmer & AP Davis, ined., --Type: SMHI 1648 Ridsdale (L, holo; iso L)Ryukyus,Taiwan (Lutao and Lanyu), and the Philippines.BATAN (Mt Iraya; Mt Matarem; along road from Itbud to Mahatao, DALUPIRI, CALAYAN(Balatubat, Salpad), CAMIGUIN DE BABUYANES (Mt Malabsing; Camiguin Volcano),LUZON: Ilocos Norte (Bangui; Pagudpud, Dampig, Nalamus; Mt Pico de Loro; MtBubungkayo, from Sto Niño to Miner’s Camp, Mt Darna; Mangoratao; Dumalneg, Cabel,Simuba Mtns, Apayao (Mt Magnas), Ilocos Sur (Burgos), Cagayan (Sta. Praxedes,San Juan; Claveria, Kilkiling, Mt Taggat), Nueva Ecija (Mt Umingan, =Mingan), Bataan(Mt Mariveles, Lamao River), Bulacan (Angat Warershed area, Norzagaray, BarangaySan Lorenzo); Rizal (Mt Irid; Montalban; Mt Mapatad, San Andres; Mt Tokduanbanoy),Laguna (Mt Makiling), Cavite (Ternate, Mt Palay-palay/Mataas na Gulod National Park,Ternate, Tala), Quezon (Mt Banahaw, Dolores, Kinabuhayan), POLILLO (Karlagan),ALABAT, CATANDUANES (Kasalugan), MINDORO: Mindoro Oriental (Puerto Galera;Pinamalayan; Mansalay, Mt Naapug; Mt Yagaw, SE slopes), MASBATE (Milagros,Barangay Matipuron, Sitio Kalunakan, 7-R Ranch), PALAWAN (Puerto Princesa,Cabayugan; Mt Cleopatra Ranges; Quezon; Mt Balagbag), BALABAC, PANAY: Iloilo(Miag-ao), MINDANAO: South Cotabato. Primary forests (including remnant dipterocarpforest), disturbed forest, regrowth and regenerating forest, at the margins of forest (eg.near meadows and fields), and secondary forest; recorded on limestone, altitude 50-900m. Flowering: Mar to Sept; Fruiting Jan, Aug, Nov.Conservation status (Sohmer & AP Davis, 2007): Least Concern [LC]. Last collected in1996.Psychotria mariguidonensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 208, Fig. 113; --Psychotria longipetiolata Merr., PJS 17 (1920, publ. 1921) 451, non Thwaites (1849). –Type: BS30418 Ramos (K, lecto). Catanduanes: Sto. Domingo River and Mt Mariguidon, on forestedslopes, Dec 1917.LUZON: Ifugao (Banawe, Mt Taggutu), Isabela (Palanan, Dibungko), CATANDUANES(Mt Mariguidon). Lowland forest near sea level at Palanan, Isabela, possibly 1000-1200m in Ifugao, reported at c. 180m by Sohmer & Davis. Flowering unknown; Fruiting:Apr, Aug-Dec.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1991.Psychotria membranifolia Bartl. ex DC, Prodr. 4 (1830) 522; --Merr., PJS 10 c (1915) Bot. 124;EPFP 3 (1923) 559; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 115, Fig. 61; --Chasaliamembranifolia (Bartl. ex DC) Elmer, LPB 3 (1911) 1032. –Type: Haenke sn (PR, holo). Luzon:without further locality data.Psychotria membranifolia var. elmeri Merr., PJS 10 c (1915) Bot. 124; EPFP 3 (1923) 559. –Type: Merrill4083 (K, lecto; isolecto: NY). Luzon: Quezon prov., Atimonan.Psychotria chasalioides Merr., PJS 10 c (1915) Bot. 125; EPFP 3 (1923) 554. –Type: Elmer 12314 (MO,lecto; isolecto: A, BISH, L*, NY*, US*). Sibuyan: Mt Giting-giting, Apr 1910.New Guinea and probably many other places in Malesia. Found on nearly all theislands and island groups of the Philippines, although extirpated in many lowland areas.PALAUI, LUZON: Ilocos Norte (Mt Palimlim; Bangui to Claveria; unspecified localityfor BS 7640 Ramos), Apayao (Mt Magnas), Cagayan (Gonzaga; Pinagteponan River),

Isabela (Malunu; Palanan), Aurora (Casiguran), Nueva Ecija (Mt Umingan=Mingan),Pampanga (Mt Pinatubo, Camp Stotsenburg), Rizal (Mt Irid; Montalban; Mt Mapatag,San Andres), Laguna (Los Baños; Mt Makiling; Majayjay; Mt Banahaw), Quezon(Umiray; Mt Tulaog; Mt Binuang; Lucban; Mt San Cristobal, Dolores, Sta. Lucia;Kinabuhayan; Atimonan, Quezon National Park; Camarines Norte (Paracale),Camarines Sur (Buhi; Agosahis), Albay (Mayon Volcano), Sorsogon (Mt Bulusan),MINDORO: Mindoro Occidental (Mt Sablayan), Mindoro Oriental (San Teodoro,Sibuang; Mt Halcon; Mt Ilong), SIBUYAN (Mt Giting-giting and vicinity), MASBATE(Masbate, Tugbo), CATANDUANES (Viga-Gigmoto area), MARINDUQUE (Dampulan,Matalim), PALAWAN: Palawan (Aborlan, Malasgao and Panalingajan River; Quezon,Sumindap River, Barangay Bundog; Mt Gantung), PANAY: Aklan (Libacao), Capiz(Jamindan; Mt Kinablan), CEBU (Argao, Poblacion Alcoy), BOHOL (Bilar, Logarita;Valencia; Dimyao), BILIRAN, LEYTE: Leyte (Ormoc, Tongonan; Ormoc, Mt Janagdan;Ormoc, Mt Mamban; SAMAR: Northern Samar (Catubig River), Western Samar (MtMalingon, Paranas; Loquilocon; Mt Concord; Sohoton, Rawis, Eastern Samar (Mt Apoi;Oras; Borongan, Tagaslian; Balangiga),MINDANAO: Zambonga del Norte (Siuya), Zamboanga del Sur (Malangas), Zamboanga(San Ramon), Agusan del Norte (Mt Urdaneta (=Mt Hilong-hilong), Cabadbaran;Butuan), Surigao del Norte (Placer; Mt Kabatuan), BALUT, DINAGAT (Cambinliw,Santiago, Loreto), PONEAS ISLET off SIARGAO (Sunkoi Hill), SIARGAO.Mostly in primary forests (including mixed dipterocarp forest, and swamp forest) but alsofound in secondary forest (e.g. emerging from logged-over forest), quite often found nearrivers; coral limestone, clay, and ultramafic rocks, altitude 50-1300 (-1500)m. Flowering:Feb to May; Fruiting: Mar to Aug; Oct to Dec.Conservation status (Sohmer & AP Davis, 2007): Least Concern [LC]. Last collected in1996.Psychotria mendozii Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 214, Fig. 116. --Type: PNH10789 Mendoza & Convocar (A, holo; iso: K, L*). Mindanao: Agusan del Norte prov., Mt Hilonghilong,mossy forest slopes, altitude 1333m, 21 Apr 1949.BILIRAN (Mt Suiro, N slopes), MINDANAO: Agusan del Norte (Mt Hilong-hilong).Forests, including mossy forest, and forested slopes; mid to higher elevations (eg. c.1333m). Flowering: Apr and May; Fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected collected in 1954.Psychotria merrittii Merr., PJS 26 (1925) 493; EPFP 4 (1925) 251; --Sohmer & AP Davis, Sida Bot.Misc. 27 (2007) 168, Fig. 88. –Type: BS 40599 Ramos & Edaño (P, lecto; isolecto: A*, K, US*).Mindoro: Mindoro Oriental prov., Mt Halcon, on forested slopes at c. 1400m, 16 Mar 1922.MINDORO: Mindoro Oriental (Mt Halcon: N face, Lantuyan; Subaan River, inland fromSan Teodoro). Usually at relatively high altitude (1350-1450m) mossy and stunted ridgeforests, also at low elevtion (eg. 150-200m). Flowering: Mar to May; Fruiting: Apr.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected collected in 1991.Psychotria merrittioides Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 142, Fig. 75. --Type:PNH 19774 Edaño (PNH, holo; iso: K, L*, PNH). Luzon: Apayao prov., Mt Magnas, summit ofmountain at ‘2900m’ (altitude very excessive, most probably in feet).LUZON: Apayao (Mt Magnas). In forest. Flowering unknown; Fruiting: Jun.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Only collected once in 1953.

Babatngin), Isabela (Mt Moises; Palanan; 20km S of Palanan), Quezon (Mt Binuang),SAMAR: Western Samar (Mt Concord). Mossy forests and on forested slopes, altitude730-1100m, in Benguet up to 2400m. Flowering: Mar; Fruiting: May, Jun.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1981.Psychotria obscurinervia Merr., PJS 17 (1920, publ. 1921) 455; --Sohmer & AP Davis, Sida BotMisc. 27 (2007) 34, Fig. 13. –Type: BS 33077 Ramos (A, lecto; isolecto: BM, K, P, US). Luzon:Ilocos Norte prov., between Bangui and Claveria in low altitude forest, 30 Aug 1918.Psychotria vulcanica Elmer, LPB 9 (1934) 3264. –Type: Elmer 14431 (A, lecto; isolecto: BISH, GH, K, L,MO, NY, P, UC, US). Luzon: Sorsogon prov., Mt Bulusan. Forest, altitude c. 200m, Oct 1915.DistributionConservation status (Sohmer & AP Davis, 2007):Psychotria oconnori Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) ; --Soejarto et al. (1995)Prelim. Checklist Fl. Pl. Palawan.PALAWAN: Mt Mantalingahan. Based on BS 77556 Edaño.Conservation status (Sohmer & AP Davis, 2007):Psychotria olsenii Sohmer & AP Davis, Sida, Bot. Misc. 27 (2007) 20, Fig. 3, --Type: S Olsen758 (L*, holo). Tawi-tawi: Tarawakan, rainforest, altitude 50m, 14 Nov 1961, fr.TAWI-TAWI (Tarawakan). Lowland rainforest, altitude 50m. Flowering: unknown;Fruiting: Nov.Conservation status (Sohmer & Davis): Critically Endangered [CR B1ab(iii)], or possiblyextinct (EX). Only known from the type specimen collected in 1961.Psychotria palawanensis Elmer, LPB 4 (1912) 1348; --Merr., EPFP 3 (1923) 560; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 136, Fig. 71;–Type: Elmer 12737 (A, lecto; isolecto: BISH, BM,GH, K, L*, MO, NY*, US*, W). Palawan: Mt Pulgar, in a slightly wooded flat at 76m, Mar 1911.Grumilea palawanensis (Elmer) Merr., PJS 10 c (1915) Bot. 134.PALAWAN (Mt Pulgar, =Thumb Peak; Tagimbong). Low and medium altitude forest,including those on well-drained soil. Flowering: Mar; fruiting: Mar.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1961.Psychotria palimlimensis Sohmer & AP Davis, Sida, Bot. Misc. 27 (2007) 223, Fig. 122, --Type:BS 33364 Ramos (A, holo; iso: K, P). Luzon: Ilocos Norte prov., Mt Palimlim, Aug 1918.LUZON: Ilocos Norte (Mt Palimlim; Mt Nagapatan; Mt Pico de Loro; Mt Darna). Forests,altitude 1000m or less. Flowering: unknown; Fruiting: Feb, Mar, Aug.Conservation status (Sohmer & Davis): Critically Endangered [CR B1ab(iii)]. Lastcollected in 1953.Psychotria pallidifolia Merr., PJS 17 (1920, publ. 1921) 456; EPFP 3 (1923) 560; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 191, Fig. 102. –Type: BS 30510 Ramos (US*, lecto; isolecto: K).Catanduanes: Mt Mariguidon and vicinity of Cololbong, on forested slopes, Nov-Dec 1917.LUZON: Cagayan (Kilingkiling River), Isabela (Palanan, Diguyo), Zambales (Subic),Rizal (Mt Lumutan), Camarines Norte (Paracale), Camarines Sur (Kamugong River;Mt Potianay; Mt Bagacay); CATANDUANES (Mt Abucay; Mt Mariguidon; unspecifiedlocalities for BS 30320, 30567 & 30446 Ramos, BS 75440 Ramos & Edaño). Forest,including dipterocarp forest, altitude 50-400m. Flowering: Apr, Sept-Dec; Fruiting: Sept-Nov.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1992

Psychotria paloensis Elmer, LPB 1 (1906) 65, as ‘paloense’; --Merr., EPFP 3 (1923) 560; --Sohmer& AP Davis, Sida Bot. Misc. 27 (2007) 194, Fig. 104. –Type: Elmer 7056 (A, lecto; isolecto: K,NY*). Leyte: Leyte prov., Palo, gently sloping wooden ravine, Jan 1906.Grumilea paloensis (Elmer) Merr., PJS 10 c (1915) Bot. 131.var. paloensisLUZON: Sorsogon (Mt Bulusan, Irosin), PALAWAN (Mt Mantalingahan; Tigaplan River),NEGROS: Negros Occidental (Mt Canlaon), LEYTE: Leyte (Buenavista near Jaro; Palo),CAMIGUIN (Mambajao), MINDANAO: Misamis (without further locality data for FB30446 Ponce & Abejero), Lanao del Sur (near Lumbatan & Sawir), Davao (Mt Apo,Todaya), Agusan del Norte (Cabadbaran, between Duros & Cawilanan; unspecifiedlocality for Weber 1007). Forests, altitude 200-1212m. Flowering: Apr, Aug to Nov;Fruiting: Jan, Apr, May, Sept, Oct.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1992.var. bukidnonensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 196, Fig. 105. --Type: PNH9977 Sulit (A, holo; iso PNH). Mindanao: Bukidnon prov., Mt Katanglad (=Kitanglad), S slope ofsouth peak in dense forest above Bgy Kibangay, altitude 1900m, 7 Mar 1949.MINDANAO: Bukidnon (Mahilucot River; Mt Candoon; Mt Kitanglad), Davao (Katigan,probably Katingan, Davao del Sur prov. for Kanehira 2553 & 2564, NY), Agusan delNorte (San Mateo, Butvun, Berdee Line). Montane forests, altitude c. 1900m.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1961.var. ovatifolia Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 197, Fig. 107. --Type: BS 39564Ramos (US*, holo; iso A, K, L, P). Mindoro: Mindoro Occidental prov., Paluan, Apr 1921.MINDORO: Mindoro Occidental (Paluan), Mindoro Oriental (San Teodoro, Subaan Riverand Sibuang River). Forest, often on steep slopes, altitude 300-630m.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1986.var. subelliptifolia Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 199, Fig. 108. --Type: PNH11913 Edaño (A, holo; iso: L*, PNH). Leyte: Leyte prov., Ormoc, Lake Danao, along forested ridge,altitude 666m, 21 Mar 1950.LEYTE: Leyte (Ormoc, Lake Danao), forested slopes, altitude 666m. Flowering:unknown; Fruiting: Mar.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Known only from type specimen collected in 1950.var. wenzeliana Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 201, Fig. 106. --Type: Wenzel 3261(MO, holo; iso: A, GH, K, NY, UC). Mindanao: Surigao del Norte prov., possibly Placer municipality,“Turugaun Turbid Placer?”, in forest, altitude 150m, 19 Apr 1928.MINDANAO: Surigao del Norte (Placer; unspecified localities for Wenzel 3120). Lowaltitude forests (eg. c. 150m). Flowering: Apr, Aug; Fruiting: Apr, Aug.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1928.Psychotria panayensis Merr., PJS 17 (1920, publ. 1921) 456; EPFP 3 (1923) 560; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 147, Fig. 77. –Type: BS 31198 Ramos & Edaño (K, lecto; isolecto:US). Panay: Capiz prov., Jamindan, moist forests along small streams, Apr-May 1918.LUZON: Camarines Sur (Botol River), MINDORO: Mindoro Oriental (Mt Ilong), PANAY:Capiz (Jamindan). Moist forest; recorded along small streams; elevation unknown.

Flowering: Feb to May; Fruiting: unknown. Known only with certainty from Panay.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1948.Psychotria papillata (Merr.) Merr., EPFP 3 (1923) 560; --Sohmer & AP Davis, Sida Bot. Misc. 27(2007) 68, Fig. 31; --Grumilea papillata Merr., PJS 10 c (1915) 138. –Type: BS 1701 Ramos (MO,lecto; isolecto: A, BM, GH, L*, NY*, P, SING, US*). Samar: prov. Unknown, Cauayan Valley, inforests, along small streams, Apr 1914.CATANDUANES (Mt Abucay; also unspecified locality for BS 30517 Ramos), SAMAR:prov. not known (Cauayan River); Western Samar (Loquilocon. Forests, below 1000m.Flowering: unknown; Fruiting: Apr-Jun, Nov, Dec.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1948.Psychotria paracalensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 20, fig. 4, --Type: BS33697 Ramos & Edaño (A, holo; iso in BM, P). Luzon: Camarines Norte prov., Paracale, Nov-Dec1918.LUZON: Camarines Norte (Paracale). Habitat not known. Flowering: unknown; Fruiting:Nov, Dec.Conservation status (Sohmer & Davis): Extinct (EX). Known only from the type specimencollected in 1918.Psychotria patulinervia Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 206, Fig. 111, --Type:PNH 1420 Edaño (L*, holo). Mindanao: Davao, Mt Apo, Miran River, in forest along stream at1916m, 9 Nov. 1946.MINDANAO: Bukidnon (Tangculan), Davao del Sur (Mt Apo). Forest, and streamsideforest, in high montane forest on Mt Apo (1916-2123m). Flowering: Aug to Nov; Fruiting:May to Jul, Oct.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1946.Psychotria pauciflora Bartl. ex DC, Prodr. 4 (1830) 522; --Merr., PJS 10 c (1915) Bot. 129; EPFP 3(1923) 560; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 232, Fig. 127. –Type: Haenke sn, (PR, holo). Luzon: withLUZON: Kalinga (Mt Masingit, Lubuagan), Benguet (unspecified locality for Loher 1511),Cagayan (Bauan, Mt Tabuan), Bulacan (Mt Biak na Bato, San Miguel; Angat WatershedArea, Norzagaray), Rizal (Tanay, Bgy. Sampaloc, Sitio Bayucan), Quezon (Mtns W ofPhilippine National Botanic Garden), MINDORO: Mindoro Oriental (Mt Halcon).Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1996.Psychotria paucinervia Merr., PJS 12 c (1917) Bot. 176; EPFP 3 (1923) 561; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 34, Fig. 12. –Type: BS 27055 Ramos (US, Lecto; isolecto: K). Luzon:Abra prov., Mt Posooy, near summit of mountain in forests, 4 Feb 1917.Conservation status (Sohmer & AP Davis, 2007):Psychotria phanerophlebia Merr., PJS 5 c (1910) Bot. 243; EPFP 3 (1923) 561; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 131, Fig. 68. –Type: BS 7404 (US, lecto*; isolecto: K). Luzon:Cagayan prov., near Pamplona, 17 Mar 1909.Grumilea phanerophlebia (Merr.) Merr., PJS 10 c (1915) 131.LUZON: Apayao (BS 28060 Fenix, cited by Merrill, 1923), Cagayan (near Pamplona;unspecified locality for BS 14506, cited by Merrill, 1923). Low altitude forests.Conservation status (Sohmer & AP Davis, 2007): Extinct [EX]. Now apparently knownonly from the type collected in 1909. Two collections other than the type not accounted

for by Sohmer & Davis.Psychotria pilosella Elmer, LPB 1 (1908) 353; --Merr., EPFP 3 (1923) 561; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 106, Fig. 53. –Type: Elmer 9213 (US*, lecto; isolecto: A, K, L*, MO, NY*,W). Luzon: Quezon prov., Mt Banahaw, in forested ravine at 850m, May 1907.Grumilea pilosella (Elmer) Merr., PJS 10 c (1915) 131.Psychotria heteromera Merr., PJS 17 (1920, publ. 1921) 449; EPFP 3 (1923) 556. –Type: BS 28773 Ramos& Edaño (K, lecto; isolecto: BM, P, US). Luzon: Quezon prov., Mt Binuang, in forest, c. 1000m,May1917.ssp. pilosella var. pilosellaLUZON: Cagayan (Claveria, Kilkiling, Mt Taggat), Quirino (Mt Alzapan), Rizal (Lagunade Bay), Laguna (Majayjay), Quezon (Mt Binuang; Mt Pular; Infanta; Real, PhilippineNational Botanic Garden; Infanta-Siniloan trail), Camarines Norte (Paracale; MinasagRiver), Camarines Sur (Mt Potianay; Kamugong River; Mt Bagacay), CATANDUANES(Viga-Gigmoto area), SAMAR: Eastern Samar (Borongan, Pinangasan; Balangiga,Barangay 1, Sitio Cantinoc). Moist forest, including dipterocarp forest, altitude 150-1000m. Flowering: Jan, Feb, May, Jun, Dec; Fruiting: May-Dec.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1996.ssp. pilosella var. samarensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 108, Fig. 56.--Type: PNH 14346 Sulit (A, holo; iso: K, L*). Samar: Northern Samar prov., Mt Cansayao,Catarman. Primary dipterocarp forest, altitude 200m, 3 Apr 1951.SAMAR: Northern Samar (Mt Cansayao), Western Samar (Mt Mahagna, Turabucan,Oquendo; Sohoton, Rawis). Moist forest, altitude 50-250m. Flowering: Apr; fruiting:unknown.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1971.ssp. erythrotricha (Elmer) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 109, Fig. 57; --Psychotria erythrotricha Elmer, LPB 5 (1913) 1891. –Type: Elmer 13486 (K, lecto; isolecto: A,BISH*, BM, GH, L*, MO, NY* 2 sheets, P, UC, US*, W). Mindanao: Agusan del Norte prov., MtUrdaneta (= Mt Hilong-hilong), altitude c. 1200m, Aug 1912.Grumilea erythrotrica (Elmer) Merr., PJS 10 c (1915) Bot. 133.var. erythrotrichaLUZON: Isabela (Mt Moises; Palanan), Aurora (without further locality data, based onFB 30972 Oro), Camarines Norte (Bicol National Park), PANAY: Antique (Sibalom,Imparayan, Mt Poras), LEYTE (no further locality data, based on Wenzel 530), SAMAR:Province unknown (Mt Comoral), Western Samar (Loquilocon; Mt Sohoton, Babasianna Daku River; Mt Concord), MINDANAO: Agusan del Norte (Mt Hilong-hilong,Cabadbaran), Surigao del Norte (without further locality data, based on BS 34702Ramos & Pascasio), DINAGAT (BS 83791 Ramos & Convocar), BUCAS GRANDE(BS 35049 Ramos & Pascasio). Moist forest, including disturbed forest, 300-1200m.Flowering: Apr-May, Aug; fruiting: Jun-Aug, Nov.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1969.var. camarinensis (Merr.) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 111, Fig. 58; --Psychotria camarinensis (Merr.) Merr., EPFP 3 (1923) 553; --Grumilea camarinensis Merr., PJS 10c (1915) Bot. 135. –Type: BS 1552 Ramos (BM, lecto; isolecto: A, L*, MO, NY*, P). Luzon:Camarines Sur prov., Mt Iriga, 29 Nov 1913.LUZON: Laguna (Mt Banahaw), Quezon (Mt Binuang), Camarines Sur (Mt Isarog; MtIriga; Mt Potianay), BILIRAN (Mt Suiro; Mt Mamban), LEYTE: Leyte (Mt Dagami; central

divide, without further locality data, based on FB 16906 Rosenbluth),SAMAR: Western Samar (Mt Calbiga), Eastern Samar (Mt Apoi). Forest on slope,altitude 900-1500m. Flowering: Apr to May; Fruiting: Feb, Aug.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1969.Psychotria pinnatinervia Elmer, LPB 1 (1906) 26; --Merr., EPFP 3 (1923) 561; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 179, Fig. 96. –Type: Whitford 932 (US*, lecto; isolecto: K, NY*, P).Luzon: Quezon prov., Mt Banahaw, Oct 1904.Grumilea pinnatinervia (Elmer) Merr., PJS 10 c (1915) Bot. 131.LUZON: Ifugao (Banawe, Lugu), Cagayan (Mt Cagua; Mt Dos Cuernos; Mt Balatongan),Isabela (Mt Moises), Aurora (Mt Alzapan), Rizal (Montalban; Mt Paningtingan; MtLumutan; Balacbac; Antipolo; Bosoboso; Tanay, San Andres; Mt Irid; Mt Angilog),Laguna (Mt Makiling), Quezon (Mt Banahaw; Dolores; Lucban), Camarines Sur (MtIsarog National Park, Ocampo), POLILLO (Barrio Apad), MINDORO: Mindoro Occidental(Paluan), LEYTE: Leyte (Mt Abucayan). Drier parts of montane forests, altitude 600-1600m. Flowering: Feb to May; Fruiting: Jan, Feb, May to Aug, Oct to Dec.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1994.Psychotria piperi Merr., PJS 17 (1920, publ. 1921) 457; EPFP 3 (1923) 561; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 26, Fig. 7. –Type: BS 513 Piper (not seen by Sohmer & Davis).Mindanao: Surigao del Norte prov., Hinatuan, 16 May 1911.SAMAR: Western Samar (Mt Calbiga; Loquilocon, Wright; Paranas, Campo Uno),MINDANAO: Agusan del Sur (Mambalili, ERDS Experimental Forest), DINAGAT(Mabini, Tabajon; unspecified locality). Dipterocarp forests, altitude 100-400m. FloweringJan; Fruiting Apr, May.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1996.Psychotria plumeriaefolia Elmer, LPB 3 (1911) 1026; --Merr., EPFP 3 (1923) 561; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 211; --Grumilea plumeriaefolia (Elmer) Merr., PJS 10 c (1915) Bot.31. –Type: Elmer 12550 (US, lecto; isolecto: A, BISH, BM, K, L, MO, US, W). Sibuyan: Mt Gitinggiting,Magallanes (=Magdiwang), along wooded banks at c. 250m, Apr 1910.MINDORO: Mindoro Oriental (Banus River), SIBUYAN (Mt Giting-giting). Lowlandforests, including forests along streams, altitude 100-250m.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1964.Psychotria pubilimba Quisumb., PJS 41 (1930) 366, t. 28; --Sohmer & AP Davis, Sida Bot. Misc.27 (2007) 181, Fig. 97. –Type: BS 47221 Ramos & Edaño (A*, lecto; isolecto in A*, BM, NY*).Luzon: Isabela prov., San Mariano, along streams in damp forests at c. 700m, 1 Mar 1926.LUZON: Cagayan (Ilang River), Isabela (San Mariano; Mt Moises; Palanan; Digallorin=Digollorin, Divinisa campsite). Streamside forest (along watercourses), or on forestedslopes, altitude (50-)610-1128m. Flowering: Apr, May; Fruiting: Mar to May.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1992.Psychotria pulgarensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 103, Fig. 53; --Psychotriarepens Elmer, LPB 4 (1912) 1349; --Merr., EPFP 3 (1923) 562, non (L.) L.,1760. –Type: Elmer 12991 (L, lecto; isolectin moist, dense forest, altitude c. 80m, Apr 1911.PALAWAN (Mt Pulgar), BALABAC (Mt Caunhayan; unspecified locality for BS 49793Ramos & Edaño). Moist forest, c. 80m. Flowering: Apr; Fruiting: unknown.

Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1927.Psychotria pygmaea Merr., PJS 17 (1920, publ. 1921) 458; EPFP 3 (1923) 561; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 94, Fig. 47. –Type: BS 29271 Ramos & Edaño (US*, lecto).Luzon: Rizal prov., Mt Susong Dalaga. In forests, 14 Aug 1917.LUZON: Rizal (Mt Susong Dalaga). Forest, altitude not known. Flowering: unknown;fruiting: Aug.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [EX]. Only known from the type specimen collected in 1917.Psychotria pyramidata Elmer, LPB 4 (1912) 1350; --Merr., EPFP 3 (1923) 562; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 150, Fig. 80; --Grumilea pyramidata (Elmer) Merr., PJS 10 c (1915) Bot. 134. –Type(=Thumb Peak), shaded forest near watercourses from c. 200-800m, May 1911.BUSUANGA, PALAWAN (Mt Capoas; Mt Pulgar; Irawan River Valley; Quezon, Bundog,Sumindap River; Ransang, Aga; Mt Mantalingahan). Undisturbed or disturbed moistforests, including dipterocarp forest and dipterocarp-swamp forest, altitude 150-1200m.Flowering: Jan, Apr, May; Fruiting: Apr-Jun, Aug.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1996.Psychotria pyramidata x gitingensisPALAWAN: Mt Beaufort. SMHI 265 (L, PNH).Conservation status (Sohmer & AP Davis, 2007):Psychotria quisumbingiana Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 136, Fig. 72. –Type: BS 43188 Ramos (UC, holo; iso: BM, K). Bohol: Bohol prov., Sevilla, along stream in theforest, 15 Sept 1923.BOHOL: Bohol (Sevilla; Valencia). Humid forest, recorded near stream, altitude 300-700m. Flowering: unknown; Fruiting: Aug-Oct.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)], orpossibly Extinct [EX]. Last collected in 1923.Psychotria radicans (Merr.) Merr., EPFP 3 (1923) 562; --Sohmer & AP Davis, Sida Bot. Misc. 27(2007) 29, Fig. 9; --Grumilea radicans Merr., PJS 10 c (1915) Bot. 139. –Type: Wenzel 1193 (BM,lecto; isolecto MO). Leyte: Buenavista, near Jaro, in forests, c. 500m, 10 Oct 1914.LEYTE (Buenavista, near Jaro; unspecified locality), SAMAR: Western Samar (Basey).Forests, altitude 500-1000m. Flowering unknown; fruiting Oct.Conservation status (Sohmer & AP Davis, 2007): Vulnerable (VU B1ab(iii)). Lastcollected in 1929.Psychotria ramosii Merr., PJS 5 c (1910) Bot. 244; EPFP 3 (1923) 562; --Sohmer & AP Davis, SidaBot. Misc. 27 (2007) 170, Fig. 89. –Type: BS 7499 Ramos (US*, lecto; isolecto K). Luzon: Cagayanprov., near Pamplona, 17 Mar 1909.LUZON: Cagayan. Low altitude forest. Flowering: Mar; fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Extinct [EX]. Known only from the typespecimen collected in 1909.Psychotria ramosissima Elmer, LPB 1 (1908) 355; --Merr., EPFP 3 (1923) 562; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 50, Fig. 21. –Type: Elmer 9178 (US*, lecto; isolecto A, K, L*, MO,NY*, W). Luzon: Quezon prov., Lucban. Dense shrubby forest of gulches at c. 200m, May 1907.LUZON: Sorsogon (Lake Bulusan), BOHOL: Bohol (unspecified locality), LEYTE:Leyte (unspecified locality), SAMAR: Northern Samar (Catubig River); Western Samar

(Wright). Forests, altitude 100-600m. Flowering: Feb-May, Jul, Aug., Fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1947.Psychotria reynosoi Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 191, Figs. 2e, 103, --Type:BS 44808 Ramos & Edaño (UC, holo; iso: A, K, L*, NY*, P, US*). Luzon: Zambales prov., MtLUZON: Cagayan (Mt Balatongan; Mt Cagua; unspecified locality), Isabela (Mt Moises;San Mariano: San Jose, Kapuntian and Agal; Palanan Forest Dynamics Plot); Quirino(Mt Alzapan), Aurora (Aurora National Park, San Luis, Bgy. Diteki, Sitio Dikong);Quezon (Mt Binuang), Zambales (Mt Marayep), Camarines Sur (Bagacay), NEGROS:Unspecified locality, but possibly Negros Occidental by Everett (FB 7258). Forestedslopes, including good condition secondary forests, 400-1300m. Flowering: Mar, May;fruiting: Jun, Jul, Dec, JanConservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1995.Psychotria rigidaefolia (Elmer) Merr., PJS 10 c (1915) Bot. 128; EPFP 3 (1923) 562; --Sohmer &AP Davis, Sida Bot. Misc. 27 (2007) 29; --Randia rigidaefolia Elmer, LPB 1 (1908) 352. –Type:LUZON: Quezon, Camarines Sur (Mt Potianay), CATANDUANES (Mt Abucay;unspecified locality), BILIRAN (Mt Suiro), LEYTE (Dagami), SAMAR: Western Samar(Mt Calbiga). Humid low altitude forests. Flowering unknown; fruiting Aug.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1954.Marayep. On forested sElmer 9118 (A*, iso). LPsychotria rubiginosa Elmer, LPB 1 (1906) 25; --Merr., PJS 1 (1906) Suppl. 135; EPFP 3 (1923) 562; --Sohmer & AP Daisolecto: NY*). Luzon: Bataan prov., Mt Mariveles, Nov 1904.Grumilea rubiginosa (Elmer) Merr., PJS 10 c (1915) Bot. 131.var. rubiginosaLUZON: Ilocos Norte (Mt Palimlim), Zambales (Sta. Cruz, Acoje Mine concession area),Bataan (Mt Mariveles; Lamao River), Rizal (Mt Pinagbayanan; Mt Irid; Montalban; MtCanumay; Mt Tokduanbaboy; Mt Lumutan; unspecified locality for Loher 14301 & 6389;also for Vidal 3070, viz. ‘district of Morong’), CEBU: Cebu (Talamban, Tebunan). Forest;ridges and ravines, altitude 600-900m. Flowering: Jan, Mar to May; Fruiting: Mar, May,Aug.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1993.var. jacobsii Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 129, Fig. 67, --Type: Jacobs 7847 (L*,holo; iso A, CANB, K). Luzon: Aurora prov., mtns NNE of Dingalan, low mixed mossy primary forestat 1000-1100m, 18 Mar 1968.LUZON: Ilocos Norte (Mt Tigbi, Nueva Era), Aurora (Sierra Madre Mtns NNE ofDingalan), NEGROS: Negros Occidental (Mt Dumalablab, Talisay municipality). Humidmixed forest, altitude 1000-1100m. Flowering: Feb, Mar; fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1992.Psychotria sarcocarpa Merr., PJS 8 c (1913) Bot. 53; EPFP 3 (1923) 563; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 31, Fig. 10. –Type: Merrill 8077 (US*, lecto; isolecto A, BM, K, L*, P).Mindanao: Zamboanga del Sur prov., Sax River Mtns, in wet, dense forest along small streams at650-1000m, 27 Nov 1911.BASILAN (Udiongan, Ayala), MINDANAO: Zamboanga del Sur (Sax River Mtns).Forests, including disturbed dipterocarp forest; recorded from wet streamside forests,

altitude 600-1000m. Flowering unknown; fruiting Jun, Nov, Dec.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU B1ab(iii)]. Lastcollected in 1996.Psychotria scaberula Merr., PJS 17 (1920, publ. 1921) 459; EPFP 3 (1923) 563; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 89, Fig. 44. –Type: BS 35159 Ramos & Pascasio (US, lecto;isolecto K). Dinagat, precise locality not known. Forest at low altitudes, 11 May 1919.DINAGAT (without further locality data). Two collections cited by Sohmer & Davis(2007). Low altitude forests. Flowering: unknown; Fruiting: May.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct (EX). Last collected in 1931.Psychotria sibuyanensis Elmer, LPB 3 (1911) 1028; --Merr., EPFP 3 (1923) 563; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 23, Fig. 6. –Type: Elmer 12355 (BM, lecto; isolecto BISH, GH, K,L*, MO, NY*, P, US*, W). Sibuyan: Mt Giting-giting, forested ridge at c. 700m, Apr 1910.LUZON: Cagayan (Claveria; Sta. Praxedes), SIBUYAN (Mt Giting-giting), PANAY:Antique (Mt Balingaso, 10km NE of Libertad), Capiz (Jamindan), MINDANAO: Surigaodel Norte (Tubud Placer: Twagaau & Mawpayang), DINAGAT (Cambinliw, Santiago,Loreto; Anahaw, Kaantojan). Primary forests, including forest with dipterocarps anddipterocarp forests, also in open and exposed places such as ridges, 150-700m.Flowering Jan-May; fruiting Apr to Sept.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1995.Psychotria sohotonensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 228, Fig. 125. --Type:PNH 117798 Madulid et al. (L*, holo). Samar: Western Samar prov., Mt Sohoton, Babasian NaGatti, 12km, Apr 1970.SAMAR: Western Samar (Mt Sohoton). Moist forest, altitude not known but probably atlow to mid elevations. Flowering: Apr; Fruiting: unknown.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)], orpossibly Extinct [EX]. Last collected in 1970.Psychotria sorsogonensis Elmer, LPB 9 (1934) 3264; --Sohmer & AP Davis, Sida Bot. Misc. 27(2007) 153, Fig. 81. –Type: Elmer 16772 (A, lecto; isolecto BISH, BM, GH, K, L*, MO, NY*, P, UC,US*). Luzon: Sorsogon prov., Mt Bulusan, Irosin. Deeply shaded forest at 606m, Jul 1916.LUZON: Sorsogon (Mt Bulusan). Humid forest, altitude c. 600m. Flowering: unknown;Fruiting: Jul.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)], orpossibly Extinct [EX]. Only known from the type specimen collected in 1916.Psychotria subalpina Elmer, LPB 1 (1906) 64; --Merr., EPFP 3 (1923) 563; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 231, Fig. 126;–Type: Elmer 7942 (K, lecto; isolecto A, L*, NY*, US*).Luzon: Quezon prov., Mt Banahaw, Lucban, mossy forest , altitude 1500m, May 1906.Grumilea subalpina (Elmer) Merr., PJS 10 c (1915) Bot. 131.LUZON: Ilocos Norte (Mt Palimlim; Bangui to Claveria), Ifugao and Mountain Province(Mt Polis), Aurora (Mt Alzapan), Nueva Vizcaya (Caraballo Mtns), Laguna (MtBanahaw), Quezon (Mt Banahaw; Banahaw de Lucban; Kinatakutan), Camarines Sur(Mt Isarog, W slope, Naga, Bgy. Panicuason), MINDORO: Mindoro Oriental (Mt Halcon),SIBUYAN (Mayos Peak), PANAY: Aklan (Mt Baloi, Libacao, Bgy. Dalagasaan, SitioTibulong), Capiz (Mt Madiaas, E slopes), NEGROS: Negros Occidental (Mt Canlaon),CAMIGUIN, MINDANAO: Misamis Occidental (Mt Malindang, Bgy. Lake Duminagat toLiboron), Bukidnon (Mt Lipa; Mt Kitanglad, Impasug-ong, Intavas), Davao (Mt McKinley).Humid forests, primary and disturbed, including ‘mossy’ and ‘elfin’ forest, altitude 1200-

2100m. Flowering: Mar to Jul; Fruiting: Jan to Oct.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1995.Psychotria subcucullata Merr., PJS 29 (1926) 492; --Sohmer & AP Davis, Sida Bot. Misc. 27(2007) 71, Fig. 33. –Type: BS 43144 Ramos (UC, lecto; iso A*, K, P, US*). Bohol: Kalingohan,rocky slopes in forest, altitude 300-700m, 20 Aug. 1923.BOHOL (Kalingohan; Bilar). Forest, on rocky slopes, 300-700m. Flowering unknown;Fruiting: Aug-Oct.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)], orpossibly Extinct [EX]. Last collected in 1923.Psychotria subsessiliflora Elmer, LPB 1 (1906) 24; --Merr., EPFP 3 (1923) 563; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 77, Fig. 37;–Type: HN Whitford 722 (P, lecto; isolecto US*).Luzon: Quezon prov., Atimonan, Aug-Sept 1904.Grumilea subsessiliflora (Elmer) Merr., PJS 10 c (1915) Bot. 131. Psychotria edanoi Merr., EPFP 3 (1923)555 as ‘edanoii’; --Grumilea brachybotrys Merr., PJS 12 c (1917) Bot. 172, non Psychotriabrachybotrya Ridl. (1909). –Type: BS 26256 Ramos & Edaño (PNH, holo; iso K). Luzon: NuevaEcija prov., Mt Umingan (=Mingan), Aug 1916.var. subsessilifloraLUZON: Ilocos Norte (unspecified locality for BS 32960 Ramos), Apayao (unspecifiedlocality for BS 28058 Fenix), Cagayan (Tamdagan River; Baguio Cove; Mt Caboloan; MtBabatngin; Pagikpik), Isabela (Mt Moises; Palanan), Bataan, Laguna (San Antonio),Quezon (Real, Philippine National Botanic Garden, Lalawinan River; Lucban; Atimonan,Quezon National Park, Pinaglabanan Trail; Mt Cadig), Camarines Norte (Paracale;Bicol National Park; Basud, Tuaca, Mt Nilisan), Camarines Sur (Mt Potianay), Albay(Mayon Volcano), Sorsogon (Mt Bulusan: Irosin and Casiguran), POLILLO (Anibawan),ALABAT, CATANDUANES (San Miguel, Barangay Sulung, Mt Saan), MINDORO:Mindoro Oriental (Mt Halcon; Naujan, Busay; Laylay, 10km from Swedish MatchHilleshog Tree Farm), NEGROS: Negros Occidental (Victoria Mtns), BILIRAN (MtSuiro), SAMAR: Western Samar (Mt Sarawag), CAMIGUIN (Guinsiliban, BarangayBotay, Sitio Sinablayan), MINDANAO: Zamboanga del Norte (Tulalu). Forests, includingsubmossy secondary forest, and plantations, altitude 35-900m. Forests, including submossysecondary forest, and plantations, altitude 35-900m. Flowering Feb to May, Sept;Fruiting all year round.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1996.var. carinata (Quisumb. & Merr.) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 79, Fig. 38;--Psychotria carinata Quisumb. & Merr., PJS 37 (1928) 205. --Type: BS 45278 Ramos & Edaño(US, lecto; isolecto A, BM, NY*, US*). Luzon: Aurora prov., Casiguran, in damp forest, 28 May1925.LUZON: Cagayan (Gonzaga; Mt Cagua; Mt Balatongan), Isabela (San Mariano), Aurora(Casiguran; Mt Dingalan), Camarines Norte (Mt Kadig), Camarines Sur (Sarapan),CATANDUANES (Dakulangpatag; unspecified locality for BS 75439 Ramos & Edaño).Low altitude forests. Flowering: unknown; Fruiting: May to Sept.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1961.Psychotria sulitii Merr. & Quisumb. ex Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 100,--Type: PNH 6408 Sulit (A, holo; iso: K, L). Samar: Western Samar prov., Mt Calbiga, Wright.Midmontane forest, altitude c. 300m, 23 May 1948.SAMAR: Western Samar (Mt Calbiga). Forest, c. 300m. Flowering: May; Fruiting: May.

Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Only collected once in 1948.Psychotria surigaoensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 100, Fig. 51. --Type:BS 34581 Ramos & Pascasio (BM, holo; iso K, PNH). Mindanao: Surigao del Norte prov., Apr1919.MINDANAO: Surigao del Norte (unspecified locality of type), DINAGAT (Loreto,Santiago, Canbilio). Forest, including dipterocarp forest, often near rivers, altitude c.50m. Flowering: Apr; Fruiting: Aug to Sept.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1991.Psychotria tayabensis Elmer, LPB 1 (1908) 354; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007)79, Fig. 39 –Type: Elmer 7581 (US*, lecto; isolecto A, K, L*, MO, NY*, W). Luzon: Quezon prov.,Lucban, May 1907.Grumilea tayabensis (Elmer) Merr., PJS 10 c (1915) Bot. 133.var. tayabensisLUZON: Ilocos Norte (Mt Quebrada), Apayao (Mt Magnas), Ifugao (Sumigar, Mt Nabito),Cagayan (Claveria; Baguio Cove), Rizal (Montalban; Mt Lumutan; Tanay, San Isidro),Laguna (Mt Banahaw; Majayjay; unspecified locality for BS 6070 Robinson), Quezon(Real, Lalawinan River; Lucban; Tagkawayan), Camarines Sur (Botol River), Sorsogon(Lake Agangay; Mt Bulusan: Irosin and Talatala Ridge, vicinity of Aguingai Valley),MINDORO (unspecified locality for FB 6883 Merritt), CATANDUANES (unspecifiedlocality for BS 75193 Ramos & Edaño), PANAY: Aklan (Libacao), NEGROS: NegrosOccidental (Mt Canlaon, Wasay trail), Negros Oriental (Tanjay, Lake Balinsasayao),BOHOL (Demyao=Dimyao; Kalingohan), LEYTE (unspecified locality for Wenzel 1620).Drier forests, sea-level to 1300m. Flowering: Feb, May, Sept; Fruiting: Mar to Aug, Oct,Nov.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1996.var. castanea (Merr.) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 82, Fig. 40; --Psychotriacastanea Merr., PJS 17 (1920, publ. 1921) 446; EPFP 3 (1923) 554. –Type: BS 33212 Ramos (A,lecto; isolecto: BM, K, P, US). Luzon: Ilocos Norte prov., Mt Nagapatan, Aug 1918.LUZON: Ilocos Norte (Mt Nagapatan; Bangui to Claveria; Mt Semenublan, E of NuevaEra; Mt Quebrada). Drier forests, on slopes, altitude c. 700m.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN]. Last collected in1975.var. euphlebia (Merr.) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 84, Fig. 43; --Grumileaeuphlebia (Merr.) Merr., PJS 10 c (1915) Bot. 133. --Psychotria euphlebia Merr., PJS 8 c (1913)Bot. 55; EPFP 3 (1923) 555;–Type: BS 14582 Ramos (not seen). Luzon: Cagayan prov., AbulugRiver, Jan 1912.Psychotria tricarpa Merr., PJS 17 (1920, publ. 1921) 460; EPFP 3 (1923) 563. –Type: BS 31224 Ramos &Edaño (A*,lecto; isolecto K, US*). Panay: Capiz prov., Jamindan, Apr-May 1918.LUZON: Apayao (Mt Magnas), Cagayan (Claveria, Taggat Watershed; Abulug River;Mt Caboloan; unspecified locality for FB 26976 Velasco), Isabela (Mt Moises; Palanan,Sangcad), Quezon (Mt Cadig), Camarines Sur (Sarapan; Mt Bagacay), ALABAT(unspecified locality for BS 48228 Ramos & Edaño), PANAY: Capiz (Jamindan). Forests,including mixed forests with dipterocarps, altitude 400-1000m. Flowering: Dec; Fruiting:Apr to Jul.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collected

in 1995.var. fasciculiflora (Merr.) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 84, Fig. 41; --Psychotriafasciculiflora (Merr.) Merr., EPFP 3 (1923) 555; --Grumilea fasciculiflora Merr., PJS 10 c (1915)135; --Type: BS 17519 Ramos (BM, lecto; Isolecto: K, US*). Samar: Yabong, on forested slopes,15 Mar 1914.SAMAR: Northern Samar (Catubig River), Western Samar (Mt Sarawag), prov. unknown(Mt Pulog). Forests, altitude 100-200m. Flowering: unknown; Fruiting: Mar.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN]. Last collected in1951.var. pinagteponanensis (Merr.) Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 88, Fig. 42;--Type: BS 78343 Edaño (A, holo). Luzon: Cagayan prov., Pinagteponan River, Oct 1929.LUZON: Cagayan (Pinagteponan River; Sta. Praxedes, Barangay San Juan), CamarinesSur (Agosahis). Forests, including mixed forest with dipterocarps, altitude c. 500m.Flowering: unknown; Fruiting: Oct.Conservation status (Sohmer & AP Davis, 2007): Vulnerable [VU]. Last collected in1995.Psychotria urdanetensis Elmer, LPB 5 (1913) 1893; --Merr., EPFP 3 (1923) 563; --Sohmer & APDavis, Sida Bot. Misc. 27 (2007) 74, Fig. 35. –Type: Elmer 13703 (UC, lecto; isolecto A, BISH,BM,GH, K, L, MO, NY, P, US, W). Mindanao: Agusan del Norte prov., Mt Urdaneta (= Mt Hilonghilong),Cabadbaran, humid forest near Lake Danao, altitude c. 1600m, Sept. 1912;Grumilea urdanetensis (Elmer) Merr., PJS 10 c (1915) 130.MINDANAO: Agusan del Norte (Mt Urdaneta, = Mt Hilong-hilong). Humid forest,including mossy forest, altititude 1500-1600m. Flowering: unknown; Fruiting: Sept.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [Ex]. Known only from the type collected in 1912.Psychotria velutina Elmer, LPB 5 (1913) 1894; --Merr., EPFP 3 (1923) 564; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 176, Fig. 93. –Type: Elmer 13615 (UC, lecto; isolecto A, BISH, BM, GH,K, L*, MO*, NY*, P, US*, W). Mindanao: Agusan del Norte prov., Mt Urdaneta (= Mt Hilong-hilong),Cabadbaran, in moist fertile soil of dense forest at 3500 ft, Aug 1912.Grumilea velutina (Elmer) Merr., PJS 10 c (1915) Bot. 33.LUZON: Quezon (Malbog), MINDANAO: Agusan del Norte (Mt Urdaneta, = Mt Hilonghilong).Humid forest, c. 1060m. Flowering: unknown; Fruiting: Aug.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Last collected in 1923.Psychotria weberi Merr., PJS 8 c (1913) Bot. 54; EPFP 3 (1923) 564; --Sohmer & AP Davis, SidaBot. Misc. 27 (2007) 217, Fig. 118. –Type: Weber 1573 (US*, lecto; isolecto A*, GH*, K, P, UC).Luzon: Cagayan prov., Abulug River, rocky hillsides, altitude c. 30m, Jan 1912.Grumilea weberi (Merr.) Merr., PJS 10 c (1915) Bot. 133.LUZON: Cagayan (Abulug River; Peñablanca, near Lagum), Isabela (Mt Cresta; MtMoises), Aurora (Casiguran, Mamator River), Quezon (Real, vicinity of PhilippineNational Botanic Garden), Camarines Norte (Mt Labo; Basud, Tuaca, Mt Nilisan),Camarines Sur (Kinaclan River; Ponawa River). Humid forest, including dipterocarpforest and secondary forest; along stream or on slopes, altitude 30-610m. Flowering: Aprto Jun; Fruiting: May, Jun, Sept.Conservation status (Sohmer & AP Davis, 2007): Endangered [EN B1ab(iii)]. Lastcollected in 1991.Psychotria wenzelii (Merr.) Merr., EPFP 3 (1923) 564; --Sohmer & AP Davis, Sida Bot. Misc. 27

(2007) 94, Fig. 49; --Grumilea wenzelii Merr., PJS 10 c (1915) Bot. 140. –Type: Wenzel 1142(MO*, lecto; isolecto A*, BM). Leyte: Leyte prov., Buenavista near Jaro, in forests, altitude c. 500m,1 Oct 1914.LEYTE. Forest, 500-700m. Flowering: unknown; Fruiting: Oct.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [Ex]. Known only from the type collected in 1914.Psychotria yagawensis Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 66, Fig. 30. --Type: PNH18687 Conklin (A, holo; iso L*, PNH). Mindoro: Mindoro Oriental prov., Mt Yagaw, SE slope,altitude 330m, 17 Jul 1953.MINDORO: Mindoro Oriental (Mt Yagaw). Forests, altitude c. 330m. Flowering:unknown; Fruiting: Jul.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)] orpossibly Extinct [Ex]. Known only from the type collected in 1953.Psychotria yatesii (Merr.) Merr., EPFP 3 (1923) 564; --Sohmer & AP Davis, Sida Bot. Misc. 27(2007) 217, Fig. 117; --Grumilea yatesii Merr., PJS 12 c (1917) 172. –Type: BS 25518 Yates (K,lecto; isolecto: A*, P, US*). Luzon: Quezon prov., Mt Cadig, low altitude forests, Dec 1916.LUZON: Isabela (Palanan, Digallorin =Digollorin, Divinisa camp site), Quezon (MtBinuang; Mt Cadig), Laguna (San Antonio), Camarines Sur (imprecise locality for FB23787 Alvarez; Sarapan; Mt Bagacay), Sorsogon (Mt Bulusan), POLILLO (Sta. Maria),SAMAR: Western Samar (Bagacay), Eastern Samar (Oras). Forest, once recorded onsteep sea cliff; also on ultramafics, altitude 50-200m. Flowering: Mar to May; Fruiting:Apr, Dec.Conservation status (Sohmer & AP Davis, 2007): Near Threatened [NT]. Last collectedin 1992.Psychotria yenii Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 68, Fig. 32. --Type: Yen 173(BISH, holo). Mindanao: South Cotabato prov., Tasaday Forest, dipterocarp forest at 1220m,Aug or Dec 1972.MINDANAO: South Cotabato (Tasaday Forest). Known only from the type collection.Conservation status (Sohmer & AP Davis, 2007): Critically Endangered [CR B1ab(iii)].Only collected once in 1972.Potential New Species:Psychotria sp. A , Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 236. --Vouchers: PPI 13205Gaerlan, Romero & Sagcal (BRIT). Palawan: Rizal municip., Ransang, Kumaraka; --PPI 38209 Pipoly &Romero et al. (BRIT), Quezon municip., Bgy. Bundog.Psychotria sp. B , Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 236. --Vouchers: SMHI 2160Podzorski (K, L). Palawan: Brookes point municip., Pulot, c. 50km N of town; --SMHI 1000 Ridsdale (K, L),Brookes Point municip., Pulot, Massin River, 12km N of Brookes Point.Psychotria sp. C, Sohmer & AP Davis, Sida Bot. Misc. 27 (2007) 237. --Vouchers: PPI 18318Garcia, Fuentes & Romero (BRIT, K). Luzon: Cagayan prov., Claveria municip., Bgy. Kilkiling, TaggatWatershed Area,UNKNOWN SPECIES—SPECIES INCOGNITAE:Psychotria cagayanensis Merr., PJS 8 c (1913) Bot. 51; EPFP 3 (1923) 553; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 237; –Type: FB 15491 Bernardo (not seen). Luzon: Cagayan prov.,Casambalangan, in forest, altitude c. 10m, Apr 1910Grumilea cagayanensis (Merr.) Merr., PJS 10 c (1915) Bot. 133.LUZON: Cagayan. Lowland primary forests. According to Merrill’s original description,

this taxon resembles P. pinnatinervia and P. luzoniensis, and that the pyrenes are notridged or grooved. Probably belonging to Sohmer and Davis’ LuzoniensisPsychotria irosinensis Elmer, LPB 9 (1934) 3262; --Sohmer & AP Davis, Sida Bot. Misc. 27 (2007)238. –Type: Elmer 17356 (not seen). Luzon: Sorsogon prov., Irosin, Mt Bulusan, Sept 1916.According to Elmer’s description, this taxon has unbranched inflorescences but isotherwise similar to P. manillensis and P. pallidiflora. Species group unknown.Psychotria samarensis Merr., PJS 12 c (1917) Bot. 174; EPFP 3 (1923) 562; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 238. –Type: FB 23576 Lasquety (not seen). Samar: Eastern Samarprov., Salcedo, on ridges, altitude c. 200m, 11 Mar 1915.SAMAR. Now known only from description.Psychotria voluta Elmer, LPB 4 (1912) 1347; --Merr., EPFP 3 (1923) 564; --Sohmer & AP Davis,Sida Bot. Misc. 27 (2007) 238. –Type: Elmer 13253 (not seen). Palawan: Palawan prov., PuertoPrincesa, Mt Pulgar (=Thumb Peak), Apr 1911.PALAWAN: Now only known from description. A member of the Ixoroides SpeciesGroup, but identity and relationship with other climbing species of this group not certain.Psydrax GaertnerRandia Linnaeus= Philippine species previously referred here arenow assigned to the following genera, viz.Aidia, Andria, Fagerlindia, Oxyceros, and TarennoideaDon’t know where the ff. were transferred--Randia lanceolata Merr., PJS 8 c (1913) Bot. 57; EPFP 3 (1923) 527. –Type: FB 11626 Fischer (notseen). Luzon: Cagayan prov., Abulug River, near Tauit, on sandstone cliff along the river,apparently in situations subject to overflow during times of high water, altitude c. 30m, Feb 1912.LUZON: Apayao, Cagayan. On rocky river banks subjected to periodic flashfloods. Lowaltitudes. Similar to R. stenophylla Merr., but nearly glabrous, with shining leaves, andwith very different fruits.Randia mindorensis Elmer, LPB 1 (1906) 28; --Merr., EPFP 3 (1923) 527. –Type: Merrill 3326 (NY*,US*, iso). Mindoro: Mindoro Oriental prov., Puerto Galera, Oct 1903.MINDORO: Mindoro Oriental (Puerto Galera). Lowland forests.Randia rostrata Merr., PJS 17 (1920, publ. 1921) 460; --Merr., EPFP 3 (1923) 528. –Type: BS 35102Ramos & Pascasio (A*, US*, iso). Bucas Grande, dry forests, 11 Jun 1919.BUCAS GRANDE. Lowland forests.Randia samalensis Elmer, LPB 1 (1906) 29; --Merr., EPFP 3 (1923) 528. –Type: Copeland 847 (notseen). Samal, Apr 1904.SAMAL (near Davao). Lowland forests.Randia stenophylla Merr., PJS 5 c (1910) Bot. 245, nom. illeg., non R. stenophylla Krause, 1909; --Merr., EPFP 3 (1923) 528. –Type: FB 7170 Curran (not seen). Luzon: Bulacan prov., Norzagaray,on rocky river banks, 16 Jun 1907.LUZON: Bulacan. On rocky river banks subjected to periodic flashfloods. Low altitudes.

Rothmannia ThunbergRothmannia graciliflora (Merr.) Ridsdale, ined.Randia graciliflora Merr., PJS 10 c (1915) Bot. 109; EPFP 3 (1923) 527. –Type: FB 17988 Miranda (L*, NY*,US*, iso). Mindanao: Misamis Occidental prov., Mt Malindang, in forests along streams, altitude 120m, 26Feb 1913.MINDANAO: Misamis Occidental (Mt Malindang). Lowland forests along streams,altitude c. 120m.Rothmannia lagunensis (Merr.) Ridsd., ined.Gardenia lagunensis Merr., PJS 10 c (1915) Bot. 110; EPFP 3 (1923) 530. –Type: Baker 3176 (not seen).Luzon: Laguna prov., hills back of Paete, 10 Apr 1914.LUZON: Cagayan, Isabela, Bulacan, Rizal, Laguna, Quezon, BUSUANGA. Lowlandprimary forests.Rothmannia leytensis Ridsd., ined. Based on: Wenzel 641 (A*). Leyte: Leyte prov., 8 Mar 1907.LEYTE.Rothmannia merrillii (Elmer) Ridsd., ined.; --Gardenia merrillii Elmer, LPB 1 (1906) 5; --Merr.,EPFP 3 (1923) 530. –Type: Merrill 669, flowering type (not seen). Culion: Halsey, Feb 1903; FB1226, fruiting type (not seen). Mindoro: Lugin, Jan 1903.Gardenia negrosensis Merr., PJS 10 c (1915) Bot. 111; EPFP 3 (1923) 530. –Type: FB 7255 Everett (notseen). Negros: Negros Occidental prov., vicinity of Cadiz and Gimagaan River.LUZON, GOLO, MINDORO, CULION, BUSUANGA, PALAWAN, ROMBLON, PANAY,NEGROS, LEYTE, MINDANAO. In thickets, secondary and primary forest at lowaltitudes.Rubia LinnaeusLiterature-Puff, C. 1984. Journal of South African Botany 50: 347.Rubia cordifolia L., Syst. ed. 12 (1767) 229; --Merr., EPFP 3 (1923) 575.Greece, subtropical and tropical Africa (except Sahara and sub-Saharan regions),temperate/subtropical Asia from Afghanistan, China, Mongolia to the Russian Far East,tropical Asia from Sri Lanka, India, Himalayan region, SE Asia and Malesia.LUZON: Mountain Province, Ifugao, Benguet, Rizal, Laguna, Quezon,MINDANAO: Lanao, Davao. In thickets and forests, chiefly above 1000m, extending to2400m, and often common in the mossy forest.Philippine plants so-named not certainly corresponding to the Linnaean species.Rubia philippinensis Elmer, LPB 9 (1934) 3214. –Type:LUZON.Saprosma BlumeSaprosma philippinensis Elmer, LPB 3 (1911) 1035; --Merr. EPFP 3 (1923) 568. –Type: Elmer9222 (PNH, holo, lost; iso MO*, NY*, US*). Luzon: Quezon prov., Lucban, “well-drained soil of lightwoods at 800m”, May 1907.LUZON: Quezon Sorsogon. In forests, c. 750m.

Saprosma ternatum (Wall.) Hook.f. in G Bentham & JD Hooker, Gen. Pl. 2 (1873) 131; --Serissaternata (Wall.) Kurz, Rep. Veg. Andaman Is. (1870) 40; --Paederia ternata Wall. in W Roxburgh, Fl.Ind. 2 (1824) 520.India (Assam), Bhutan, Bangladesh, W and S-C China, Hainan, Burma, Thailand,Laos, ?Philippines.Schradera VahlRevisions:Puff, C, L Andersson, U Rohrhofer & A. Igersheim. 1993. The tribe Schradereae (Rubiaceae)reexamined. Bot. Jahrb. Syst. 114: 449–479.Puff, C, J Greimler & R Buchner. 1998. Revision of Schradera (Rubiaceae-Schradereae) inMalesia. Blumea 43: 287–335.Schradera elmeri Puff, Buchner & Greimler, Blumea 43 (1998) 315; --Lucinaea involucrata Elmer,LPB 3 (1911) 997; --Merr., EPFP 3 (1923) 520. –Type:MINDANAO: Davao (Mt Apo:Todaya, Lake Linao and Gumate District, E slopes of MtApo), Davao Oriental (Mt Kampalili), Bukidnon (Mt Kitanglad; Mahilucot River). Exposedridges, mossy forests, 750-1900m. Flowers in Mar, Jun, Jul and Oct; fruits in Mar, Junand Jul.Schradera monocephala (Merr.) Puff, Buchner & Greimler, Blumea 43 (1998) 299; --Lucinaeamonocephala Merr., PJS 5 c (1910) Bot. 241; EPFP 3 (1923) 520. --Type: BS 10034 Ramos(?PNH, holo, presumed lost by Puff et al., loc. cit.; an isotype however, is extant in US*[barcode 00137914], and is here designated as lectotype. BS 1111 Ramos (BM, K), was earlierdesignated by Puff et al as neotype. The above collections from Luzon: Laguna prov., Paete, nearDahican River.Lucinaea epiphytica Elmer, LPB 5 (1913) 1873; --Merr., EPFP 3 (1923) 520. --Type: Elmer 14147 (?PNH,holo, probably lost). Mindanao: Agusan del Norte prov., Mt Urdaneta (=Mt Hilong-hilong),Cabadbaran.LUZON: Laguna (Paete), Sorsogon (Mt Bulusan), MINDORO: Mindoro Oriental (MtYagaw), PALAWAN (Mt Beaufort); CATANDUANES, BILIRAN (Mt Suiro), LEYTE,NEGROS, MINDANAO: Lanao, Agusan del Norte (Mt Urdaneta (= Mt Hilong-hilong),Cabadbaran). In primary montane forests with Agathis, 620 to 850m. Flowering: Mar toSept, Fruiting: Nov.Scleromitrion (Wight & Arnott) MeissnerScleromitrion angustifolium (Cham. & Schltdl.) Benth., Hooker's J. Bot. Kew Gard. Misc. 4(1852) 172; --Oldenlandia angustifolia (Cham. & Schltdl.) Kuntze, Revis. Gen. Pl. 1 (1891) 292;--Sclerococcus angustifolius (Cham. & Schltdl.) Bartl. ex DC., Prodr. 4 (1830) 436; --Hedyotisangustifolia Cham. & Schltdl., Linnaea 4 (1829) 153.E Himalayas, Burma, Nicobar Isls, Sumatra, Peninsular Malaysia, Java, Philippines,Moluccas.Scleromitrion pauciflorum (Bartl. ex DC.) Miq., Fl. Ned. Ind. 2 (1857) 187; --Hedyotis paucifloraBartl. ex DC, Prodr. 4 (1830) 419.Philippines. 42 PHI. Provisionally accepted.Scleromitrion strictum (L.f.) ined.Tardavel stricta (L.f.) Hiern, Cat. Afr. Pl. 1 (1898) 503; --Borreria stricta (L.f.) K.Schum. in HGA Engler &

KAE Prantl, Nat. Pflanzenfam. 4(4) (1891) 143; --Bigelovia stricta (L.f.) Blume, Bijdr. (1826) 945; --Spermacoce stricta L.f., Suppl. Pl. (1782) 120.Oldenlandia tenelliflora (Blume) Kuntze, Revis. Gen. Pl. 1 (1891) 292; --Scleromitrion tenelliflorum (Blume)Korth., Ned. Kruidk. Arch. 2 (1851) 155; --Hedyotis tenelliflora Blume, Bijdr. (1826) 971; --Merr.,EPFP 3 (1923) 500.S-C China, Hainan, Japan, Ryukyu isls, Taiwan, Nicobar Isls, Thailand, PeninsularMalaysia, Java, Borneo, Philippines, New Guinea, ?Queensland.LUZON: Ilocos Norte, Benguet, Sorsogon, MINDORO, PANAY, LEYTE,MINDANAO. Low and médium altitudes, ascending to 1500m, along streams,clearings, etc.Scleromitrion verticillatum (L.) ined.Hedyotis verticillata (L.) Lam., Tabl. Encycl. (1791) 271; --Merr., EPFP 3 (1923) 501; --Oldenlandiaverticillata L., Mant. (1767) 40.Scleromitrion hispidum (Retz.) Korth., Ned. Kruidk. Arch. 2 (1851) 155; --Hedyotis hispida Roth, Nov. Pl. Sp.(1821) 95; --Oldenlandia hispida (Retz.) Lam., Encycl. 4 (1798) 536; --Hedyotis hispida Retz.,Observ. Bot. 4 (1786) 23.Hedyotis verticillata var. scaberrima Hatus., Trans. Nat. Hist. Soc. Taiwan 26 (1936) 224.Trop. & Subtrop. Asia to NW. Pacific. Assam, Bangladesh, E Himalayas, India,Nepal, Burma, Thailand, Vietnam, S China, Hainan, Taiwan, Bonin Isls, Sumatra,Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi, Moluccas, Caroline Isls.N LUZON to MINDANAO, in most islands and provinces. Lowlands inthickets, old clearings and other open sites.Scyphiphora CF GaertnerScyphiphora hydrophyllacea Gaertn.f., Fruct. 3 (1805) 91, t. 196; --Merr., EPFP 3 (1923) 533.India, Sri Lanka, Andaman Isls, Peninsular Malaysia, Philippines, N Australia, NewCaledonia.N LUZON to PALAWAN and MINDANAO, in mangrove swamps and along mosttidal streams subject to the influence of salt or brackish water.Spermacoce LinnaeusRevision:Cabral, EL & NM Bacigalupo. 1996. Revision of Borreria section Pseudodiodia (RubiaceaeSpermacoceae). Opera Botanica Belgica 7: 309–327.Spermacoce articularis L.f., Suppl. (1872) 118;Borreria articularis (L.f.) FN Williams, Bull. Herb. Boiss. sér. 5, 2 (1905) 956.India, Burma, Indochina, S China, Taiwán, Ryukyu Isls, Malesia.Tropical & subtropical Asia: Sri Lanka, India, E Himalayas, Nepal, S China, Hainan,Taiwan, Ryukyu Isls, Burma, Vietnam, Peninsular Malaysia, Borneo, Philippines,Sulawesi. Introduced in Tanzania, Mauritius, Madagascar and Australia (NorthernTerritory, Queensland).Spermacoce buruensis Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 261.Exallage buruensis (Miq.) Bremek., Verh. Kon. Ned. Akad. Wetensch., Afd. Natuurk., Tweede Sect. 48(2)(1952) 142; --Hedyotis buruensis (Miq.) Valeton ex Merr., EPFP 3 (1923) 496.Hedyotis magallanensis Elmer, LPB 3 (1911) 976; --Merr., EPFP 3 (1923) 498. –Type: Elmer 12205.Hedyotis parva Merr., PJS 3 c (1908) Bot. 162; --Elmer, LPB 3 (1911) 975. –Type:Philippines, Sulawesi, Moluccas (Ambon, Buru).

SIBUYAN, PANAY. Low altitude forests, ascending to 400m.Spermacoce gracilis Ruiz & Pav., Fl. Peruv. 1 (1798) 61.Borreria gracilis LO Williams, Phytologia 26 (1973) 487 (1973), homonym.Spermacoce exilis (LO Williams) CD Adams ex WC Burger & CM Taylor, Fieldiana, Bot., n.s., 33 (1993)316; --Borreria exilis LO Williams, Phytologia 28 (1974) 227.Pantropic.Spermacoce hispida L. Sp. Pl. (1753) 102; DC. Prodr. 4 (1830) 555. –Merr., Fl. Manila (1912) 455. –Type:Borreria articularis var. hispida (L.) Sivar. & Manilal, Int. Quart. J. Pl. Sci. Res. 2 (1975) 89, nom. inval.; --Borreria hispida (L.) K Schum. in HGA Engler & KAE Prantl, Nat. Pflanzenfam. 4(4) (1891) 144,nom. illeg.; --Merr., EPFP 3 (1923) 574.Spermacoce mutilata Blanco, Fl. Filip., ed. 2: 43 (1845).W Himalayas, India, Bangladesh, E Himalayas, Nepal, Thailand, Vietnam, S China,Hainan, Taiwan, Ryukyus, Taiwan, Peninsular Malaysia, Java, Philippines, SolomonIsls.BATAN to BASILAN. Open dry places at low and medium altitudes.Spermacoce manillensis Kuntze, Revis. Gen. Pl. 3(2) (1898) 123.Spermacoce discolor (DC.) Fern.-Villar in FM Blanco, Fl. Filip., ed. 3, 4(13A) (1880) 113, nom. illeg.; --Borreria discolor DC., Prodr. 4 (1830) 545; --Bigelovia discolor Bartl. ex DC., Prodr. 4 (1830) 545.Philippines. 42 PHI.Spermacoce ocymoides Burm.f., Fl. Ind. (1768) 34, t. 13, Fig.. 1; --Merr., Fl. Manila (1912) 455.Borreria ocymoides (Burm.f.) DC, Prodr. 4 (1830) 544; --Merr., EPFP 3 (1923) 574.Spermacoce laevis Spreng., Syst. Veg. 1 (1824) 401, nom. illeg., non Lam. (1791).Tropical Asia to SW Pacific: Sri Lanka, India, Burma, Peninsular Malaysia, Java,Philippines, New Guinea, Solomon Isls, Fiji, Samoa, Vanuatu. Introduced in theneotropics and tropical Africa.LUZON: Bataan, Bulacan, Rizal, BASILAN. Lowland thickets.Spermacoce pusilla Wall. in W Roxburgh, Fl. Ind. 1 (1820) 379.Borreria pusilla (Wall.) DC., Prodr. 4 (1830) 543; --Bigelovia pusilla (Wall.) Spreng., Syst. Veg. 1 (1824) 405.Trop. Asia: Sri Lanka, India, Nepal, W & E Himalayas, Burma, ?Sumatra, PeninsularMalaysia, ?Borneo, Philippines. Introduced in tropical Africa.Spermacoce ramanii Sivar. & RV Nair, Taxon 35 (1986) 367.Borreria ramanii (Sivar. & RV Nair) MR Almeida, Fl. Maharashtra 3A (2001) 6.Spermacoce lasiocarpa R Br. ex Wall., Numer. List (1829) 832, nom. nud.Trop. Africa, Madagascar, Sri Lanka, India, Nepal, Himalayas, S China, Taiwan, Japan,Burma, Thailand, Vietnam, Peninsular Malaysia, Philippines.Spermacoce remota Lam., Tabl. Encycl. 1 (1792) 273.Borreria remota (Lam.) Bacigalupo & EL Cabral, Darwiniana 37 (1999) 334.Indigenous to tropical and subtropical America, introduced in the Philippines.Spermacoce scaberrima Blume, Bijdr. (1826) 946.Borreria scaberrima (Blume) Koord., Exkurs.-Fl. Java 3 (1912) 282.Spermacoce muriculata Blanco, Fl. Filip., ed. 2 (1845) 44.Sumatra, Peninsular Malaysia, Java, Philippines.

Streblosa KorthalsRevision:Bremekamp, CEB. 1947. A monograph of the genus Streblosa Korth. [Rubiaceae]. Journal of theArnold Arboretum 28: 145-185.Streblosa axilliflora Merr., PJS 10 c (1915) Bot. 141. –Type: BS 22018 Ramos (US*, iso). Luzon:Camarines Sur prov., Mt Isarog, Nov 1913.Streblosa glabra Valeton ex Winkler in Engl., Bot. Jahrb. 44 (1910) 567; --Elmer, LPB 4 (1912) 1356; --Merr., EPFP 3 (1923) 564.Streblosa axilliflora Merr. var. angustifolia Bremek., J. Arnold Arb. 28 (1947) 184.Streblosa axilliflora Merr. var. latifolia Bremek., J. Arnold Arb. 28 (1947) 184.Streblosa axilliflora Merr. var. laxiflora Bremek., J. Arnold Arb. 28 (1947) 184.LUZON, CATANDUANES, SAMAR, PALAWAN. In primary forest at low altitudes. (S.glabra, fide Merr. 1923).Streblosa palawanensis Bremek., J. Arnold Arb. 28 (1947) 182.Streblosa palawanensis var. elmeri Bremek., J. Arnold Arb. 28 (1947) 183.Streblosa palawanensis var. merrillii Bremek., J. Arnold Arb. 28 (1947) 183.PALAWAN.Sulitia Merrill =AtractocarpusLiterature-Bridson. 1979. Kew Bull. 34: 377.Tarenna GaertnerTarenna acuminata Merr., PJS 17 (1920, publ. 1921) 462; EPFP 3 (1923) 525. –Type: BS 13904RamosLUZON: Apayao. Lowland forests.Tarenna arborea (Elmer) Elmer, LPB 4 (1912) 1359; --Merr., EPFP 3 (1923) 525; --Randia arboreaElmer, LPB 3 (1911) 1005. –Type: Elmer 10978LUZON: Laguna, Camarines, MINDANAO: Davao. Forests, up to 500m altitude.Tarenna bakeri (Merr.) Bremek., Fedde, Repert. 37 (1934) 187; --Pavetta bakeri Merr., PJS 10 c(1915) Bot. 117; EPFP 3 (1923) 544. –Type: Baker 3214 (not seen). Luzon: Laguna prov., hillsback of Paete, 10 Apr 1914.LUZON: Laguna, Quezon. Low or medium altitude primary forests.Tarenna bartlingii Bremek., Repert. Spec. Nov. Regni Veg. 47 (1939) 97.Tarenna pubescens (Bartl. ex DC.) Bremek., Repert. Spec. Nov. Regni Veg. 37: 203 (1934), nom. illeg., nonCraib (1932); --Pavetta pubescens (Bartl. ex DC.) Merr., PJS 17 (1920 publ. 1921) 479; EPFP 3(1923) 545; --Webera pubescens Fern.-Villar, Novis. App. (1880) 108; --Stylocoryna pubescensBartl. ex DC., Prodr. 4 (1830) 377; --Miq., Fl. Ind. Bat. 2 (1857) 206; --Elmer, LPB 1 (1906) 33. –Type: Cuming 1236LUZON: Ilocos Norte. Lowland thickets or secondary forests.Tarenna catanduanensis Merr., PJS 17 (1920, publ. 1921) 463; EPFP 3 (1923) 525. –Type: BS30387LUZON: Camarines, CATANDUANES. Lowland forests.Tarenna cumingiana (Vidal) Elmer, LPB 5 (1913) 1898; --Merr., EPFP 3 (1923) 525; --Webera

cumingiana Vidal, Phan. Cuming. Philip. (1885) 178; Rev. Pl. Vasc. Filip. (1886) 152; --Elmer, LPB 1(1906) 40. –Type: Cuming 685Borneo, Philippines.LUZON: Quezon, Camarines, Albay, Sorsogon, MINDORO, BILIRAN, LEYTE,MINDANAO: Agusan del Norte, Surigao del Norte. Lowland primary forests.Tarenna elongata Merr., PJS 17 (1920, Publ. 1921) 464; EPFP 3 (1923) 525. –Type: BS 35218Ramos & Pascasio.DINAGAT, BUCAS GRANDE. Lowland forests.Tarenna fragrans (Blume) Koord. & Valeton, Bijdr. Boom. Java 8 (1902) 77; --Merr., EPFP 3(1923) 525; --KM Wong in FS Ng (ed.) Tree Fl. Malaya 4 (1989) 411; --Webera fragrans Hook.f., Fl.Brit. Ind. 3 (1880) 103; --Fern.-Villar, Novis. App. (1880) 108; --Stylocoryna fragrans Blume,Bijdr. (1826) 982.Sumatra, Peninsular Malaysia, Java, Borneo, Philippines. PALAWAN. Lowland forests.According to Merrill (1923), identification of Palawan material is not certainly correct.Tarenna littoralis Merr., PJS 17 (1920, publ. 1921) 465; EPFP 3 (1923) 526. –Type: Merrill 9418LUZON: Pangasinan, Zambales, APO (off the W coast of Mindoro), CORON,PALAWAN, DUMARAN, BURIAS, SULU ARCHIPELAGO (UBIAN). Lowland forests,often on or near the seashore.Tarenna loheri (Merr.) Bremek., Repert. Spec. Nov. Regni Veg. 37 (1934) 197; --Pavetta loheriMerr., PJS 27 (1925) 55; EPFP 4 (1925) 254. --Type: Loher 13151 (not seen). Luzon: Rizal prov.,Montalban, Paningtingan, Mar 1913.LUZON: Rizal.Tarenna luzoniensis (Vidal) Bremek., Fedde, Repert. 37 (1934) 198; --Pavetta luzoniensis (Vidal)Merr., PJS 17 (1920, publ. 1921) 479; EPFP 3 (1923) 544; --Webera luzoniensis Vidal, Phan.Cuming. Philip. (1885) 179; Rev. Pl. Vasc. Filip. (1886) 152; --Elmer, LPB 1 (1906) 40; --Merr., PJS1 (1906) Suppl. 133. –Type: Cuming 1323.Tarenna eucrantha (Elmer) Merr., PJS 17 (1920, publ. 1921) 471; EPFP 3 (1923) 525; --Pavetta eucranthaElmer, LPB 3 (1911) 1015; ibid. 5 (1913) 1886. –Type: Elmer 12492LUZON: Ilocos Norte, La Union, Pangasinan, Apayao, Cagayan, Nueva Vizcaya,Bataan, Rizal, Laguna, Batangas, PALAWAN, SIBUYAN, LEYTE, SAMAR,MINDANAO: Zamboanga, Lanao del Sur, Agusan del Norte, DINAGAT, SIARGAO.Low and médium altitude forests.Tarenna meyeri Elmer, LPB 5 (1913) 1898; --Bremek., Fedde, Repert. 37 (1934) 199; --Pavettameyeri (Elmer) Elmer, LPB 3 (1911) 1015; --Merr., PJS 17 (1920, publ. 1921) 479; EPFP 3 (1923)545; --Webera meyeri (Elmer) Merr., PJS 1 (1906) Suppl. 133; --Ixora meyeri Elmer, LPB 1 (1906)68. –Type: FB 2764 Meyer (not seen). Luzon: Bataan prov., Lamao Forest Reserve, Feb 1905LUZON: Bataan (Mt Mariveles, Lamao Forest Reserve). Primary forest, altitude c. 650m.Note: The type material cited for the basionym Ixora meyeri Elmer is Elmer 7958 fromQuezon province (Mt Banahaw), collected in Apr 1906. Why this disparity?Tarenna multinervia (Merr.) Bremek., Fedde, Repert. 37 (1934) 200; --Pavetta multinervia Merr.,PJS 17 (1920, publ. 1921) 475; EPFP 3 (1923) 545. –Type: FB 9013 Whitford & HutchinsonMINDANAO: Zamboanga. Lowland primary forests.Tarenna nitida Merr., PJS 17 (1920, publ. 1921) 466; EPFP 3 (1923) 526. –Type: FB 23388 ContrerasNEGROS. Lowland forests.Tarenna obtusifolia Merr., PJS 17 (1920, publ. 1921) 467; EPFP 3 (1923) 526. –Type: BS 1257

McGregorBOHOL, BUCAS GRANDE. Thickets and forests near the seashore.Tarenna palawanensis (Elmer) Merr., PJS 17 (1920, publ. 1921) 471; EPFP 3 (1923) 526; --Pavetta palawanensis Elmer, LPB 4 (1912) 1344; --Merr., PJS 10 c (1915) Bot. 117. –Type: Elmer12940 (L*, NY*, US*, iso). Palawan: Palawan prov., Puerto Princesa, Mt Pulgar (=Thumb Peak), inhumus-covered compact soil of a densely wooded flat at 250ft, Apr 1911.PALAWAN. Lowland primary forests, to 300m altitude.Tarenna pangasinensis Merr., PJS 20 (1922) 458; EPFP 3 (1923) 526. –Type: FB 28419 ZamucoLUZON: Pangasinan. Near streams at low altitude.Tarenna scaberula (Merr.) Merr., PJS 17 (1920, publ. 1921) 471; EPFP 3 (1923) 526; --Pavettascaberula Merr., PJS 10 c (1915) Bot. 118. –Type: BS 13114 Ramos & Foxworthy (not seen).Luzon: Quezon prov., Dayap, in forests, altitude c. 30m, 9 Mar 1911.LUZON: Cagayan, Laguna, Quezon, Camarines. Low and medium altitude forests.Tarenna stenantha Merr., PJS 17 (1920, publ. 1921) 468; EPFP 3 (1923) 526. –Type: BS 29003Ramos & EdañoLUZON: Ilocos Norte, Bataan, Batangas, Quezon, MINDORO. Low and medium altitudeforests.TarennaRandia pubifolia Elmer, LPB 5 (1913) 1895; --Merr., EPFP 3 (1923) 528. –Type: Elmer 14060MINDANAO: Agusan del Norte. Humid forests, altitude c. 1000m.“May be the fruiting specimen of Tricalysia negrosensis Elmer”, fide Elmer, LPB 5 (1913)1895.Tarennoidea Tirvengadum & SastreLiterature:Tirvengadum, DD & C Sastre. 1979. La signification taxonomique des modes des ramification deRandia et genres affines. Mauritius Inst. Bull. 8: 77–98.Tarennoidea wallichii (Hook.f.) Tirveng. & Sastre, Mauritius Inst. Bull. 8 (1979) 90; --Tarennawallichii (Hook.f.) Ridl., Fl. Mal. Pen. 2 (1923) 106; --KM Wong in FS Ng (ed.) Tree Fl. Malaya 4(1989) 413, 415; --Randia wallichii Hook.f., Fl. Brit. Ind. 3 (1860) 113.Stylocoryna incerta (Koord. & Valeton) Elmer, LPB 1 (1906) 33; --Tarenna incerta Koord. & Valeton, Bijdr.Boom. Java 8 (1902) 269; --Merr., EPFP 3 (1923) 526.Tarenna ebracteata (Elmer) Elmer, LPB 5 (1913) 1898; --Randia ebracteata Elmer, LPB 4 (1912) 1354.Tarenna kotoensis (Hayata) Masamune, Trans. Nat. Hist. Soc. Formos. 20 (1932) 462; --Hatusima, Mem.Fac. Agr. Kagoshima Univ. 5 (1966) 52; --Chomelia kotoensis Hayata, Icon. Pl. Formosan. 9 (1920) 58.Tropical and subtropical Asia: India, Sri Lanka, Bangladesh, Laos, Vietnam, Taiwan,Ryukyu Isls, Japan, Thailand, Peninsular Malaysia, Java, Borneo, Philippines.BATANES, LUZON (widespread), MINDORO, CORON, PALAWAN, JOLO,MINDANAO: Bukidnon. Chiefly in low and medium altitude secondary forests.Tetralopha Hooker filius =GynochthodesTimonius de CandolleTimonius appendiculatus Merr., PJS 4 c (1909) Bot. 327; EPFP 3 (1923) 541. –Syntypes: BS 2164

Ramos (NY*, US*). Luzon: Rizal prov., Bosoboso, Jan 1904; --BS 2651Ramos (NY*, US*). Luzon:Rizal prov., May 1907.LUZON: Rizal, Batangas, Camarines, Sorsogon, MINDORO (Mt Agong). Mediumaltitude primary forests.Timonius arboreus Elmer, LPB 1 (1906) 72; --Merr., EPFP 3 (1923) 541. –Type: Elmer 7960 (notseen). Luzon: Quezon prov., Mt Banahaw, forests, altitude 500m, Apr 1906.Timonius gamillii Elmer, LPB 4 (1912) 1359; ibid. 3 (1911) 1013 (staminate fl.). –Type: Elmer 12469 (A*,GH*, L*, NY*, US*). Sibuyan: Mt Giting-giting, Magallanes (=Magdiwang), “in moist gravelly soilofwooded banks of the Pauala River at 1000ft”, May 1910.Taiwan (Lanyu and Lutao), Philippines.LUZON: Isabela, Nueva Ecija, Zambales, Bataan, Rizal, Laguna, Camarines,Sorsogon, MINDORO, PALAWAN, SIBUYAN, TICAO, PANAY, GUIMARAS,NEGROS, LEYTE, MINDANAO: Agusan del Norte, Surigao. Low and mediumaltitude primary forests.Timonius auriculatus Merr., PJS 17 (1920, publ. 1921) 480; EPFP 3 (1923) 541. –Type: BS 35190RamosDINAGAT. Lowland forests.Timonius caudatifolius Elmer, LPB 5 (1913) 1898; --Merr., EPFP 3 (1923) 541. –Type: Elmer14190 (BISH*, L*, MO*, NA*, NY*, US*, iso). Mindanao: Agusan del Norte prov., Cabadbaran, MtUrdaneta (=Hilong-hilong), ‘lax epiphytic shrub upon moss covered ascending tree trunks 15ftabove the very humus wet stony soil in humid forests at 5250ft or some distance above the lagoon’,Oct 1912.S Philippines, N Sulawesi.MINDANAO: Agusan del Norte (Mt Hilong-hilong). Exposed ridges in themossy forest, altitude c. 1550m.Timonius confertiflorus Merr., PJS 10 c (1915) Bot. 114; EPFP 3 (1923) 541. –Type: Merrill Phil.Pl. 1698, leg. M. Ramos (L*, NY*, US*, iso). Samar: Yabong, in dry thickets, 16 Mar 1914.SAMAR, MINDANAO: Surigao. Lowland dry thickets.Timonius epiphyticus Elmer, LPB 3 (1911) 1011; --Merr., EPFP 3 (1923) 541. –Type: Elmer 10706BASILAN, MINDANAO: Lanao del Sur, Bukidnon, Davao. Primary forests, altitude 800-1050m.Timonius ferrugineus Merr., PJS 10 c (1915) Bot. 115; EPFP 3 (1923) 541. –Type: Merrill 9210 (L*,NY*, US*, iso). Palawan: Palawan prov., Taytay, in forests immediately back of mangroves, 30 Apr1913.PALAWAN. Primary forests behind mangroves.Timonius finlaysonianus (Wall. ex G Don) Hook.f., Fl. Brit. India 3 (1880) 127; --Guettardafinlaysoniana Wall. ex G.Don, Gen. Hist. 3 (1834) 552.Sumatra, Peninsular Malaysia, Java, Borneo, Philippines.Timonius gracilipes Merr., PJS 8 c (1913) Bot. 60; EPFP 3 (1923) 541. –Type: BS 11110 Ramos(US*, iso). Cebu: Cebu prov., Buacao, dry hill thickets, Mar 1912.CEBU. Low and medium altitude thickets.Timonius hirsutus (Elmer) Merr., PJS 17 (1920, publ. 1921) 480; EPFP 3 (1923) 542; --Greeneahirsuta Elmer, LPB 5 (1913) 1857. –Type: Elmer 13476Timonius trichophorus Merr., PJS 9 c (1914, publ. 1915) Bot. 388. –Type:

LEYTE, SAMAR, MINDANAO: Agusan del Norte, BUCAS GRANDE. Primary forests upto 900m altitude.Timonius lanceolatus Merr., PJS 17 (1920, publ. 1921) 318; EPFP 3 (1923) 542. –Type: BS 35211Ramos & PascasioMINDANAO: Surigao del Norte, DINAGAT. Lowland forests.Timonius longiflorus (Merr.) Merr., EPFP 3 (1923) 542; --Greenea longiflora Merr., PJS 4 c (1909)Bot. 323. –Type: FB 9430 Whitford & Hutchinson (US*, iso). Mindanao: Zamboanga, Tibucuy, inforest, altitude c. 20m, Feb 1908.MINDANAO: Zamboanga. Lowland primary forests.Timonius longistipulus Merr., PJS 8 c (1913) Bot. 59; EPFP 3 (1923) 542. –Type: BS 15371Ramos (L*, US*, iso). Leyte: Leyte prov., Mtns back of Dagami, on trees in the mossy forests, 13Aug 1912.LEYTE. Mossy forests, altitude at or above 1000m.Timonius obovatus Elmer, LPB 1 (1906) 36; --Merr., EPFP 3 (1923) 542. –Type: Merrill 1048, leg. SGracia (not seen). Luzon: Aurora prov., Aug 1902LUZON: Aurora, SIBUYAN. Low and medium altitude forests.Timonius oligophlebius Merr., PJS 11 c (1916) Bot. 34; EPFP 3 (1923) 542. –Type: BS 23367Ramos (BISH, 3 sheets*, L*, NY*, US*, iso). Luzon: Sorsogon prov., Mt Kililibong, in forests near thesummit, apparently at altitude c. 1000m, 17 Aug 1915.LUZON: Sorsogon. Forests, altitude c. 1000m.Timonius pachyphyllus Merr., PJS 26 (1925) 494; EPFP 4 (1925) 251. --Type: BS 40794 Ramos(not seen). Luzon: Rizal prov., Mt Angilog (=?Angelo or Angilo), 20 Apr 1922.LUZON: Rizal. Forested slopes, c. 1200m.Timonius palawanensis Elmer, LPB 4 (1912) 1360; --Merr., EPFP 3 (1923) 542. –Type: Elmer12705 (A*, BISH*, L*, NY*, US*, iso). Palawan: Palawan prov., Brookes Point, ‘on earthen banks ofwell-shaded woods at 100ft altitude along a rivulet’, Feb 1911.PALAWAN. Lowland primary forests.Timonius panayensis Merr., PJS 17 (1920, publ. 1921) 317; EPFP 3 (1923) 542. –Type: BS 35445Martelino & Edaño (L*, US*, iso). Panay: Aklan prov., Libacao, May-Jun 1919.PANAY: Aklan (Libacao), Iloilo (Mt Bulilao), Mt Salibongbong. Forests along streams,ascending to 600m.Timonius philippinensis Merr., Philipp. Bur. Forestry Bull. 1 (1903) 54; --EPFP 3 (1923) 542. –Syntypes: Ahern 463 (NY*, US*), 456 (not seen). Dinagat, 1902.MINDORO, SAMAR, MINDANAO: Surigao del Norte, DINAGAT, BUCAS GRANDE.Lowland forests.Timonius pulgarensis Elmer, LPB 4 (1912) 1360; --Merr., EPFP 3 (1923) 542, erroneouslyas ‘palawanensis’. –Type: Elmer 13201 (L*, iso). Palawan: Palawan prov., Puerto Princesa, Mt Pulgar(=Thumb Peak), May 1911PALAWAN. Dense thickets near the summit of Mt Pulgar, altitude c. 1300m.Timonius quadrasii Elmer, LPB 1 (1906) 36; --Merr., EPFP 3 (1923) 542. –Type: Ahern 545, leg. JFQuadras (US*, iso). Mindanao: Zamboanga del Sur prov., Tetuan, Mar-Jun 1901.MINDANAO: Zamboanga del Sur (Tetuan). Probably from lowland forests.

Timonius quinqueflorus Merr., PJS 10 c (1915) Bot. 116; EPFP 3 (1923) 543. –Type: BS 20885Escritor (L*, US*, iso). Luzon: Quezon prov., Guinayangan, 8 Mar 1913.LUZON: Quezon (Guinayangan). Low or medium altitude forests.Timonius rotundus Merr., PJS 17 (1920, publ. 1921) 317; EPFP 3 (1923) 543. –Type: BS 35176Ramos & PascasioDINAGAT. Lowland forests.Timonius samarensis Merr., PJS 12 c (1917) Bot. 165; EPFP 3 (1923) 543; --Timoniusmacrophyllus Merr., PJS 5 c (1910) Bot. 246; --Elmer, LPB 3 (1911) 1010, non Valeton. –Type: FB12856 (US*, iso). Samar: Western Samar prov., near Catbalogan, hillsides along streams, altitude c.200m, 7 Feb 1909.SAMAR, BUCAS GRANDE. Lowland forests.Timonius ternifolius (Bartl.) Fern.-Villar, Novis. App. (1880) 109; --Elmer, LPB 1 (1906) 36;--Merr., EPFP 3 (1923) 543; --Petesia ternifolia Bartl. ex DC, Prodr. 4 (1830) 395. –Type: Haenke sn (notseen).BUBUYAN ISLS (CAMIGUIN), LUZON: Ilocos Norte, BUSUANGA, CULION,PALAWAN, PANAY. Lowland forests and thicketsTimonius urdanetensis Elmer, LPB 5 (1913) 1899; --Merr., EPFP 3 (1923) 543. –Type: Elmer14197MINDANAO: Agusan del Norte (Mt Hilong-hilong). Exposed ridges in the mossy forest,altitude c. 1650mTimonius valetonii Elmer, LPB 3 (1911) 1012; --Merr., EPFP 3 (1923) 543. –Type: Elmer 12342SIBUYAN, MINDANAO: SurigaoPossible ined. names:Timonius stenophyllus Merr. & Quisumb. Based on: PNH 6417 Sulit & Conese (L*). Samar:Western Samar prov., Wright, Mt Calbiga, near Km 62, forest on flat ridge, altitude c. 300m, 25 May1948.SAMAR: Western Samar (Wright, Mt Calbiga).Timonius sulitii Merr. & Quisumb. Based on PNH 6429 Sulit & Conese (L*). Samar: Eastern Samarprov., Taft, Mt Calbiga, dipterocarp forest along road, Km 64, altitude c. 250m, 18 May 1948.SAMAR: Eastern Samar (Taft, Mt Calbiga).Tricalysia A. Richard ex de Candolle=Philippine spp. now in Diplospora, DiscospermumTricalysia sorsogonensis Elmer, LPB 9 (1934) 3269, not validly published, no Latin descr. --Voucher: Elmer 17164 (iso in NY*).In Hypobathreae. Generic position yet to be determined. Although in habit very similarto Tricalysia and Diplospora, fruit characters (pendulous seeds with superior embryoradicle, folded exotesta with elongated cells thickened along radial walls) leave no doubtabout the placement in Hypobathreae. –Ali & Robbrecht, op. cit.Uncaria SchreberRevision:Ridsdale, CE. 1978. A revision of Mitragyna & Uncaria. Blumea 24: 43–100.

Uncaria acida (Hunter) Roxb., Hort. Beng. (1814) 86; Fl. Ind. ed. 1, 2 (1824) 129; --Merr., EPFP 3(1923) 508; --Ridsdale, Blumea 24 (1978) 80. –Type from Penang, Malaysia.var. acidaBurma (Mergui), ?Thailand, S Vietnam, Sumatra, Peninsular Malaysia, Java, LesserSunda Isls, Borneo, Philippines, Sulawesi.Uncaria attenuata Korth., Verh. Nat. Gesch. Ned. Bot. (1840) 170; --Ridsdale, Blumea 24 (1978) 79.Uncaria canescens Elmer, LPB 1 (1906) 37; --Merr., EPFP 3 (1923) 509, non Korth. (1840).Uncaria attenuata ssp. bulusanensis Ridsd. ex Phillip. & Heming., Phytochem. 14 (1975) 1855, nom. nud; --Uncaria bulusanensis Elmer, LPB 9 (1934) 3270. –Type: Elmer 14917 (L*, MO*, NY*, U*, US*, iso).Luzon: Sorsogon prov., Irosin, Mt Bulusan, Nov. 1915.Andaman Isls, Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi.Uncaria callophylla Blume ex Korth., Verh. Nat. Gesch. Ned. Bot. (1840) 170; --Ridsdale, Blumea24 (1978) 85.Uncaria luzoniensis Merr., PJS 27 (1925) 57; EPFP 4 (1925) 254. --Type: Loher 14457 (UC; holotypefragment: L* ). Luzon: Rizal prov., Mabiluang, Oct 1913.Sumatra, Peninsular Malaysia, Borneo, Philippines, Moluccas, New Guinea.Uncaria cordata (Lour.) Merr., Interp. Herb. Amb. (1917) 479; EPFP 3 (1923) 509; --Ridsdale,Blumea 24 (1978) 75; --Restiaria cordata Lour., Fl. Cochinch. (1790) 639.var. cordataforma cordataS Burma, Peninsular Thailand, Cambodia, Laos, S Vietnam, Sumatra, PeninsularMalaysia, Java, Lesser Sunda Isls, Borneo, Philippines, Sulawesi, Moluccas, NewGuinea.forma sundaica Ridsd., Blumea 24 (1978) 76.Sumatra, Peninsular Malaysia, Java, Borneo, Philippines, Sulawesi.var. ferruginea (Blume) Ridsd., Blumea 24 (1978) 76forma insignis (Bartl.) Ridsd., Blumea 24 (1978) 77.Uncaria insignis DC, Prodr. 4 (1830) 348; --Merr., EPFP 3 (1923) 509.Nauclea luzonensis D Dietr., Syn. Pl. 1 (1839) 791.Sumatra, Borneo, Philippines, Sulawesi, ?Moluccas.Uncaria lanosa Wall. in W Roxburgh, Fl. Ind. 2 (1824) 131.var. lanosaforma philippinensis (Elmer) Ridsd., Blumea 24 (1978) 89; --Uncaria philippinensis Elmer, LPB 1(1906) 38; --Merr., EPFP 3 (1923) 510. –Syntypes: RC McGregor 322 (NY*, US*). Mindoro:Mindoro Oriental prov., Baco River, Apr-May 1905; --HN Whitford 762 (US*). Luzon: Quezon prov.,Infanta, Sept 1904.S Taiwan, Philippines, N Sulawesi.forma setiloba (Benth.) Ridsd., Blumea 24 (1978) 89, Fig. 9.6; --Uncaria setiloba Benth. in Hooker'sJ. Bot. Kew Gard. Misc. 2 (1843) 223. –Type from Ambon.Uncaria glabrata Fern.-Villar in FM Blanco, Fl. Filip., ed. 3, 4 (13A) (1880): 105.Uncaria florida Vidal, Phan. Cuming. Philip. (1885) 176. . –Type: Cuming 1504 (L*, lecto; isolecto: K, L*, NY,2 sheets*). Philippines. No further locality data.Philippines, Moluccas.

var. ferrea (Blume) Ridsd., Blumea 24 (1978) 87, Fig. 10:6; --Uncaria ferrea (Blume) DC., Prodr. 4(1830) 348; --Nauclea ferrea Blume, Bijdr. (1826) 1014.Uncaria lanosa forma ferrea (Blume) Ridsd., Blumea 24 (1978) 87.Nicobar Isls, Andaman Isls, Burma, Thailand, Sumatra, Peninsular Malaysia, Java,Lesser Sunda Isls, Borneo, Philippines, New Guinea.PALAWANUncaria longiflora (Poir.) Merr., Interp. Herb. Amb. (1917) 480; EPFP 3 (1923) 509; --Ridsdale,Blumea 24 (1978) 82.Uncaria pachyphylla Merr., PJS 10 c (1915) Bot. 102. –Type: BS 16652 Ramos (US*, iso). Luzon: Lagunaprov., San Antonio, Sept 1912.var. longifloraPeninsular Thailand, Sumatra, Peninsular Malaysia, Borneo, Sulawesi, Moluccas, NewGuinea, ?Solomon Isls. No material seen from Java or Lesser Sunda Isls.var. pteropoda (Miq.) Ridsd., Blumea 24 (1978) 82; --Uncaria pteropoda Miq., Fl. Ind. Bat. 2 (1857)143. –Type from Sumatra.Uncaria laevifolia Elmer, LPB 5 (1913) 1902; --Merr., EPFP 3 (1923) 509. –Type: Elmer 14178 (BO, L*, NY*,2 sheets). Mindanao: Agusan del Norte prov., Cabadbaran, Mt Urdaneta [=Mt Hilong-hilong], Oct1912.Sumatra, Peninsular Malaysia, Borneo, ?Philippines.Uncaria nervosa Elmer, LPB 3 (1911) 985; --Merr., EPFP 3 (1923) 510; --Ridsd., Blumea 24 (1978)79. --Type: Elmer 11794 (K, L*, MO*, NY*, US*, iso). Mindanao: Davao del Sur prov., Mt Apo,Todaya, ‘in densehumid forested flats at 4000ft [=1219 m], south of the Baruring River’, Sept 1909.Java, Borneo, Philippines, Moluccas, New Guinea, Solomon Isls.Uncaria perrottetii (A Rich.) Merr., PJS 8 c (1913) Bot. 60; EPFP 3 (1923) 510; --Ridsd., Blumea24 (1978) 85, Fig. 9:8; --Sabicea perrottetii A Rich., Mém. Soc. Hist. Nat. Paris 5 (1834) 228. –Type: Perrottet sn (P, holo; iso: L*). Philippines: “Manille”.Uncaria hookeri Vidal, Phan. Cuming. Philip. (1885) 177; Rev. Pl. Vasc. Filip. (1886) 149; --Havil., J. Linn.Soc. Bot. 33 (1897) 86, t. 4, Fig. 13-18; --Elmer, LPB 1 (1906) 37. –Syntypes: Cuming 619; 1128(K). Luzon: without definite locality; --Vidal 1469 (L*). Luzon: Bulacan prov., Angat.Philippines. LUZON: Ilocos Norte (Mt Piao), Ilocos Sur, La Union, Pangasinan,Mountain Province, Cagayan, Bataan (Dinalupihan), Pampanga (Mt Abu), Bulacan(Norzagaray), Rizal (Montalban; Bosoboso), Laguna (Los Baños; Calauan),MINDANAO: Lanao, Davao. Lowland and medium altitude forests.Uncaria roxburghiana Korth., Verh. Nat. Gesch. Ned. Bot. (1840) 172; --Ridsdale, Blumea 24(1978) 89.Uncaria brevicarpa Elmer, LPB 9 (1934) 3271. --Type: Elmer 16842 (BO, K, L, MO*, NY*, U*, US*, iso).Luzon: Sorsogon prov., Irosin, Mt Bulusan, Aug 1916.Sumatra, Peninsular Malaysia, Borneo, Philippines.Uncaria velutina Havil., J. Linn. Soc. Bot. 33 (1897) 84; --Merr., EPFP 3 (1923) 509; --Ridsdale,Blumea 24 (1978) 80. –Type: Cuming 1503 (K, L*, syntype). Mindoro.Uncaria canescens var. velutina Ridsd. ex Phillip. & Heming., Phytochem 14 (1975) 1855, nom. nud.Uncaria clavisepala Elmer, LPB 1 (1908) 350; --Nauclea clavisepala Merr., PJS 3 c (1908) Bot. 266. –Type: Elmer 8262 (BO, K, NY*, iso). Luzon: Laguna prov., Los Baños, Apr 1906.Philippines.Urophyllum Jack ex Wallich

Revisions:Bremekamp, CEB. 1940. On Urophyllum Wall. (Rubiaceae) and its nearest allies. Recueil Trav.Bot. Neerl. 37: 171–197.Bremekamp, CEB. 1940. A preliminary account of the Philippine species of Urophyllum Wall.,Pleiocarpidia K Schum., & Praravinia Korth. (Rubiaceae). Journal of the Arnold Arboretum21: 32–40.Urophyllum acuminatissimum Merr., PJS 10 c (1915) Bot. 106; EPFP 3 (1923) 521; --Bremek., J.Arnold Arb. 21 (1940) 39. –Type: BS 20423 Ramos (not seen). Luzon: Laguna prov., San Antonio,in forest, Feb 1913.LUZON: Laguna (San Antonio), Quezon, Camarines Norte (Paracale), Sorsogon. Inprimary forests, at low altitudes. An easily recognizable species, of which the flowers arestill unkown.Urophyllum bataanense Elmer, LPB 1 (1906) 40; --Merr., EPFP 3 (1923) 522; --Bremek., J. ArnoldArb. 21 (1940) 40. –Type: Elmer 6810 (NY*, iso). Luzon: Bataan prov., Mt Mariveles, Nov 1904.LUZON, MINDORO, SIBUYAN, LEYTE, PANAY, NEGROS. In primary forests chiefly atmedium altitudes, ascending to 1200m in the mossy forests.Notes: Bremekamp (1940) is of the opinion that specimens collected in parts of Luzonother than Bataan, as well as from Mindoro, Bohol, and Jolo belong in the neighborhoodof U. mindorense Merr.Urophyllum caudatum Merr., PJS 17 (1920, publ. 1921) 481; EPFP 3 (1923) 522; --Bremek., J.Arnold Arb. 21 (1940) 38; --Type: BS 33348 Ramos (A*, US*, iso). Luzon: Ilocos Norte prov., Mt Palimlim,on forested slopes, altitude 900m, 20 Aug. 1918.LUZON: Ilocos Norte (Mt Palimlim), PANAY: 32302 McGregor. In primary forests,extending into the mossy forests, 900-1100m.Urophyllum elliptifolium Merr., PJS 5 c (1910) Bot. 247; EPFP 3 (1923) 522; --Elmer, LPB 3 (1911)998; --Bremek., J. Arnold Arb. 21 (1940) . --Type: FB 3871 Curran. Palawan: Mt Pulgar, onforestedslopes, altitude 700-1300m, Feb 1906.PALAWAN: Mt Pulgar. In forests, altitude 700-1300m. Merrill compares this with theBornean U. subanurum Stapf.Urophyllum leytense Merr., PJS 8 c (1913) Bot. 62; EPFP 3 (1923) 522; --Bremek., J. Arnold Arb.21 (1940) 40. --Type: BS 15289 Ramos (not seen). Leyte: Leyte prov., Mountains back of Dagami,in forests, 13 Aug 1912.NEGROS: Negros Oriental (Cuernos Mtns), LEYTE: Leyte (Mtns back of Dagami),Southern Leyte (Cabalian). In forests, probably at or above 900m.Urophyllum memecyloides (C Presl) Rolfe, J. Bot. 23 (1885) 119; --Bremek., J. Arnold Arb. 21(1940) 38; --Cymelonema memecyloides C Presl, Epim. (1851) 210. –Type: Cuming 1678. Samar.Urophyllum arboreum C Robinson, PJS 6 c (1911) Bot. 227; --Merr., EPFP 3 (1923) 521, non (Reinw. exBlume) Korth., Nederl. Kruidk. Arch. 2 (1851) 194.LUZON: Rizal (Mt Irig =Irid), Laguna (Infanta-Siniloan trail; Paete; San Antonio; Cavinti),Quezon (Mt Binuang; Lucban), Camarines Sur (Maagnas), POLILLO, CATANDUANES,MINDORO: Mindoro Oriental (Mt Halcon), SAMAR: Without further locality data (Cuming1678); Northern Samar (Catubig River), LEYTE (Mt Abucayan; without precise locality(Wenzel 85)), PANAY: Capiz side of Mt Madiaas, NEGROS: Negros Oriental (CuernosMtns), CAMIGUIN, MINDANAO: Zamboanga del Sur (Malangas), Bukidnon (MtDumalucpican), Agusan del Norte (Mt Urdaneta (=Hilong-hilong), Cabadbaran), Surigaodel Norte. Forests at low and medium altitudes.

Urophyllum mindorense Merr., PJS 20 (1922) 464; EPFP 3 (1923) 523; --Bremek., J. Arnold Arb.21 (1940) 39. –Type: BS 39379 (A*, iso). Mindoro: Mindoro Occidental prov., Mt Calavite, forestedslopes, altitude 600-700m, Apr 1921.MINDORO: Mindoro Occidental (Mt Calavite, also BS 39398 Ramos in addition to thetype collection). Known only from the type locality.Urophyllum panayense Merr., PJS 17 (1920, publ. 1921) 482; EPFP 3 (1923) 523; --Bremek., J.Arnold Arb. 21(1940) 39. –Type: BS 31421 Ramos & Edaño (A*, US*, iso). Panay: Capiz prov., Jamindan, inforests along small streams, 22 May 1918.PANAY: Capiz (Jamindan, BS 31318 Ramos & Edaño), Aklan (Mt Macosolen,=Magosolon BS 30819 Ramos & Edaño). In primary forests along streams at low ormedium altitudes.Urophyllum platyphyllum Elmer, LPB 3 (1911) 999; --Merr., EPFP 3 (1923) 523; --Bremek., J.Arnold Arb. 21 (1940) 39; --Type: Elmer 12363 (A*, GH*, L, MO*, NY*, US*, iso). Sibuyan: MtGiting-giting, Magallanes (=Magdiwang), moist gravelly soil of humid forested gulleys at 1750 ft onthe España side of the trail leading over the ridge, Apr 1910.SIBUYAN (Mt Giting-giting), CATANDUANES (BS 30548 Ramos). In forest at lowand medium altitudes. Apparently related to U. macrophyllum (Blume) Korth., but stillimperfectly known.Urophyllum reticulatum Elmer, LPB 3 (1911) 999; --Merr., EPFP 3 (1923) 523--Bremek., J. ArnoldArb. 21 (1940) 39; --Type: Elmer 12506 (A*, GH*, L, MO*, NY*, US*). Sibuyan: Mt Giting-giting,Magallanes (=Magdiwang), stony moist soil on a forested ridge, altitude 4000 ft, May 1910.SIBUYAN: Mt Giting-giting. On ridges in the mossy forest, c. 1200m. Known only fromthe type collection.Urophyllum subglabrum Merr., PJS 12 c (1917) Bot. 162; EPFP 3 (1923) 523; --Bremek., J. ArnoldArb. 21 (1940) 38. –Type: BS 26507 Ramos & Edaño (US*, holo; iso: A, NY*). Luzon: Nueva Ecijaprov., Mt Umingan (=Mingan), dry slopes, altitude c. 100m, 6 Sept. 1916.LUZON: Nueva Ecija (Mt Umingan, BS 29196), Rizal (Mt Irig =Irid, BS 41960 Ramos).In primary forests at low medium altitudes.Urophyllum urdanetense Elmer, LPB 5 (1913) 1900; --Merr., EPFP 3 (1923) 524; --Bremek., J.Arnold Arb. 21 (1940) 38. –Type: Elmer 13788 (US*, holo, fide Buchner; iso A, L*, NY*, 2 sheets,U*. Mindanao: Agusan del Norte prov., Mt Urdaneta (=Hilong-hilong), Cabadbaran, Oct. 1912.MINDANAO: Agusan del Norte (Mt Urdaneta, =Mt Hilong-hilong). Mossy forest, altitude1500-1650m.Known from two collections in the type locality. Very near U. memecyloides (C Presl)Rolfe but imperfectly described. Nearly related, but apparently distinct from the twopreceding species is a plant colleted from JOLO: Mt Dajo, BS 4391 (L), staminate.Ined.names & possible novelties:Urophyllum sp. aff. reticulatum, Bremek., J. Arnold Arb. 21 (1940) 39.PANAY: Capiz (Jamindan). Based on BS 31165 Ramos & Edaño (A, c.fr.) and BS 31000(L, fl. buds). According to Bremekamp, ‘doubtless a good species…’ but inavailability offlowers prevents him from formally naming it.Urophyllum sp. aff. leytense, cited in Bremek., J. Arnold. Arb. 21 (1940) 40. Based on BS 40963Ramos (A), female, Mindoro: Mindoro Oriental prov., Pinamalayan; and BS 40595 (A), young buds, Mindoro:Mindoro Oriental prov., Mt Halcon.

Urophyllum halconense Merr., in sched., cited in Bremek., J. Arnold. Arb. 21 (1940) 40. Based on BS40707 Ramos & Edaño (A), fr.; also BS 40603 (A), sterile. Mindoro: Mindoro Oriental prov., Mt Halcon.Urophyllum sp., cited in Bremek., J. Arnold. Arb. 21 (1940) 40. Based on BS 40885 & 41022 Ramos (A,L)., male. Mindoro: Mindoro Oriental prov., Pinamalayan..A very curious plant with solitary subsessile flowers, provided with a 6-merous calyx andcorolla and surrounded by a fairly large involucel; the buds are too young for a gooddescription.Vidalasia TirvengadumVidalasia morindifolia (Elmer) Tirveng., Biogeographica (The Hague) 74 (1998) 166; --Gardeniamorindaefolia Elmer, LPB 1 (1906) 67; --Merr., EPFP 3 (1923) 530. –Type: BS 32706 Ramos (notseen).Gardenia pubifolia Merr., PJS 10 c (1915) Bot. 112; --Merr., EPFP 3 (1923) 531. –Type: BS 19343 Wester(US*, iso). Mindanao: Davao Oriental prov., Mati, 19 Aug 1912.LUZON: Ilocos Norte, La Union, Bataan, Quezon, Laguna; PALAWAN, TICAO,MINDANAO: Davao Oriental. In thickets and low altitudes forest, often along seashoreon cliffs.Villaria RolfeVillaria acutifolia (Elmer) Merr., PJS 5 c (1910) Bot. 248; EPFP 3 (1923) 532; --Gardenia acutifoliaElmer, LPB 1 (1906) 6. --Syntypes: Copeland 437 (NY*, US*, type specimen for flower). Mindanao:Davao, Mar 1904; --FB 457 JF Quadras (not seen, type specimen for fruit). Dinagat, Feb 1901.DINAGAT, MINDANAO: Davao. Forest at low altitudes.Villaria fasciculiflora Quisumb. & Merr., PJS 37 (1928) 207. --Type: BS 46775 Ramos & Edaño(NY*, US*, iso). Luzon: Isabela prov., San Mariano.LUZON: Isabela (San Mariano and Palanan).Villaria glomerata (Bartl. ex DC) Mulyan. & Ridsd., Reinwardtia 12 (2004) 195; --Hypobathrumglomeratum (Bartl. ex DC) K Schum. in Engl. & Prantl, Nat. Pflanzenfam. 4 4 (1891) 156; --Merr. Fl.Manila (1912) 450; EPFP 3 (1923) 533;LUZON, MINDANAO, In thickets, hedges, and secondary forests at low altitudes.Villaria odorata (Blanco) Merr., Sp. Blancoanae (1918) 363; EPFP 3 (1923) 532.Gardenia rolfei (Vidal) Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966) 50; --Villaria rolfei Vidal, Phan.Cuming. Philip. (1885) 180; --Merr., EPFP 3 (1923) 532. --Type: Cuming 1271 (iso L*, 2 sheets).Gardenia elliptica Elmer, LPB 1 (1906) 6. –Type: HN Whitford 705 (US*, iso). Luzon: Quezon prov., Atimonan, Aug-Sept 19Lanyu and the Philippines. BATAN, BABUYAN ISLS, LUZON: Cagayan, Isabela, Aurora,Quezon, Laguna, Camarines, Albay, Sorsogon, SAMAR, MINDANAO: Surigao. Inthickets along the seashore, occasionally also inland thickets.Villaria philippinensis Rolfe, J. Linn. Soc. Bot. 21 (1884) 311, t. 10, Fig. 1-8; --Merr., EPFP 3 (1923)532.LUZON: Cagayan, Aurora, Quezon, Camarines, CATANDUANES, LEYTE, SIARGAO,DINAGAT. Secondary forests at low altitudes.Revisions:Wendlandia Bartling ex de Candolle

Cowan, JM. 1932. The genus Wendlandia. Notes from the Royal Botanic Garden Edinburgh 16:233–314.Cowan, JM. 1936. The Malaysian species of Wendlandia (Rubiaceae). Bulletin du JardinBotanique du Buitenzorg 3, 14: 8-46.Wendlandia brachyantha Merr., PJS 1 (1906) Suppl. 128; EPFP 3 (1923) 505. –Type: Whitford 449(US*, iso). Luzon: Bataan prov., Mt Mariveles. On ridges on the mossy forest, altitude c. 1050m,Jul 1904.LUZON: Bataan. Known only from the type collection.Wendlandia glabrata DC, Prodr. 4 (1830) 411; -- Merr., EPFP 3 (1923) 505.India to Taiwan, Peninsular Malaysia, Philippines. BABUYAN ISLS, LUZON: IlocosNorte, Abra Ilocos Sur, Mountain Province, Benguet, Nueva Vizcaya, Zambales,Pampanga, Rizal, Camarines, Sorsogon, MINDORO. Medium altitude forests,ascending to 2100m.Wendlandia luzoniensis DC, Prodr. 4 (1830) 412; Merr., EPFP 3 (1923) 506.var. luzoniensisBATANES, LUZON: La Union, Pangasinan, Zambales, Bataan, Rizal, Laguna,Camarines, CULION, PALAWAN, CAMIGUIN, MINDANAO: Davao. Low andmedium altitude forests. According to Merrill, very close to W. paniculata DC.var. membranifolia (Elmer) Cowan, xxxxxWendlandia membranifolia Elmer, LPB 1 (1908) 357; --Merr., EPFP 3 (1923) 506.LUZON: Abra, Pangasinan, Mountain Province, Benguet, Cagayan, Nueva Vizcaya,Nueva Ecija, Bulacan, Rizal, MINDANAO: Bukidnon. Low and medium altitudeforests. Possibly a mere pubescent form of the preceding.var. syringoides (Cowan)Wendlandia syringoides (Cowan) Cowan, Notes Bot. Gard. Edinburgh 18 (1934) 187.var. williamsii (Merr.) CowanWendlandia williamsii Merr., PJS 10 c (1915) Bot. 101; EPFP 3 (1923) 506. –Type: Williams 2084 (NY*, 2sheets; US*, iso). Mindanao: Zamboanga, Sax River, altitude c. 130m, 1 Feb 1905.MINDANAO: Zamboanga. Low altitude forests.Wendlandia nervosa Merr., PJS 3 c (1908) Bot. 263; EPFP 3 (1923) 506. –Type: BS 5007 Ramos(NY*, US*, IT). Luzon: Zambales prov., Nov. –Dec. 1907.LUZON: Zambales. Forests, probably above 1000m.Wendlandia philippinensis Cowan, Notes Roy. Bot. Gard. Edinburgh 16 (1932) 274. –Type: Merrill4409 (NY*, iso). Luzon: Benguet prov., Daklan to Kabayan, Oct.-Nov. 1905.Wendlandia sibuyanensis Cowan, Notes Roy. Bot. Gard. Edinburgh 16 (1932) 299. –Type: Elmer12345 (A*, NY*). Sibuyan: Mt Giting-giting, Magallanes (=present day Magdiwang), moist gravellysoil of wooded banks along Pauala River, alt. 1000ft, Apr 1910.Wendlandia uvariifolia Hance, J. Bot. 8 (1870) 73; --Merr., EPFP 3 (1923) 506.S China, Taiwan, Indochina, Philippines. LUZON: Ilocos Sur, Cagayan. Low altitudethickets and second growth forests.Wendlandia warburgii Merr., PJS 26 (1925) 495; EPFP 4 (1925) 251. --Type: Warburg 12016.Luzon: Isabela prov. Apr 1888.

LUZON: Isabela.Williamsia Merrill =PraraviniaXanthophytum BlumeRevision:Axelius, B. 1990. The genus Xanthophytum (Rubiaceae). Taxonomy, phylogeny andbiogeography. Blumea 34: 425–497Xanthophytum ferrugineum (DC) Merr., in Mitt. Inst. Allg. Bot. Hamb. 7 (1937) 270; --Axelius,Blumea 34 (1990) 462, t. 23; --Metabolus ferrugineus DC, Prodr. 4 (1830) 436. –Type: Haenke sn(Herb Prague, cited by Merrill, 1909). Luzon: without definite locality.Xanthophytum fruticulosum Reinw. ex Blume Bijdr. (1826) 989; --Merr., EPFP 3 (1923) 492.Java, Borneo, Philippines. LUZON: Ilocos Norte, Laguna, Quezon, Camarines,Sorsogon, POLILLO, PANAY, MINDANAO: Agusan del Norte, Surigao del Norte. Lowand medium altitude damp primary forests. (Distribution based on Merr., 1923).Cultivated:Carphalea JusseauCarphalea kirondon Baill., Bull. Mens. Soc. Linn. Paris 1 (1878) 188.Cinchona LinnaeusCinchona calisaya Wedd., Ann. Sci. Nat., Bot., III, 10 (1848) 6.Cinchona officinalis L., Sp. Pl. (1753) 172.Cinchona pubescens Vahl, Skr. Naturhist.-Selsk. 1 (1790) 19.Coffea LinnaeusCoffea arabica L., Sp. Pl. 1 (1753) 172.Coffea excelsa Cheval, Rev. Cult. Colon. xii. 258; et in Compt. Rend. Acad. Sci. Par. cxl. 517.Coffea liberica Hiern, Trans. Linn. Soc. London, Bot. 1(4) 1876 [1880, publ. Dec 1876] 171, t. 24.Coffea robusta L Linden, Cat. Pl. Econ. (1900) 64.Hamelia JacquinHamelia patens Jacq., Enum. Syst. Pl. (1760) 16.Pentas BenthamPentas lanceolata (Forssk.) Deflers, Voy. Yemen (1889) 142.Posoqueria AubletPosoquieria longiflora Aubl., Hist. Pl. Guiane 1 (1775) 134.Pseudomussaenda flava Verdc., Kew Bull. 6 (1951, publ. 1952) 378.

Mussaenda flava (Verdc.) Bakh.f. in Backer, Beknopte Fl. Java 15 (Fam. 173) (1956) 72.Native of tropical Africa. Sometimes planted here as an ornamental.

Family 270. RUBIACEAE Acranthera Arnott MINDANAO ...

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