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The Genus Coccothrinax in Cultivation - International Palm Society

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

<strong>The</strong> <strong>Genus</strong><br />

<strong>Coccothr<strong>in</strong>ax</strong><br />

<strong>in</strong> <strong>Cultivation</strong><br />

CHRIS MIGLIACCIO<br />

Dept. of Natural Sciences,<br />

Health & Wellness,<br />

Miami-Dade College,<br />

300 NE 2nd Avenue,<br />

Miami, Florida, 33132<br />

USA<br />

chris.migliaccio@mdc.edu<br />

AND<br />

ARMANDO REYES<br />

3641 SW 21st Street,<br />

Miami, Florida, 33145<br />

USA<br />

kronos5@msn.com<br />

1. A portion<br />

of the<br />

extensive<br />

collection of<br />

<strong>Coccothr<strong>in</strong>ax</strong><br />

at Fairchild<br />

Tropical<br />

Botanic<br />

Garden.<br />

For many years, only a handful of <strong>Coccothr<strong>in</strong>ax</strong> species have ever been cultivated<br />

outside of the Caribbean bas<strong>in</strong>. In the last five years, over two dozen additional<br />

species have become available to collectors. Although little is known about the<br />

needs of these species <strong>in</strong> cultivation, some of these palms are very beautiful and<br />

deserve wider use.<br />

PALMS 49(2): 57–71<br />

57


PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

<strong>Palm</strong>s of the genus <strong>Coccothr<strong>in</strong>ax</strong> are species<br />

from coastal scrub and open forest from the<br />

Caribbean islands and border<strong>in</strong>g lands (Uhl &<br />

Dransfield 1987). While most forest species are<br />

found <strong>in</strong> the mid-canopy, a few are emergent<br />

palms. All are solitary, with the exception of<br />

C. pseudorigida and C. fagildei, which cluster.<br />

Currently, there are 50 accepted species <strong>in</strong> the<br />

genus. Considerable taxonomic confusion<br />

exists because many pockets of endemism<br />

occur <strong>in</strong> eastern Cuba, perhaps due to the rich<br />

assemblages of soil types and microclimates,<br />

and most species seem to possess limited<br />

distribution ranges, mak<strong>in</strong>g f<strong>in</strong>d<strong>in</strong>g the palms,<br />

much less collect<strong>in</strong>g ripe seeds, difficult.<br />

When Henderson, Galeano and Bernal<br />

published the Field Guide to the <strong>Palm</strong>s of the<br />

Americas <strong>in</strong> 1995, they reduced the number of<br />

species to 14 and <strong>in</strong>dicated that the genus was<br />

<strong>in</strong> dire need of revision. However, Moya and<br />

Leiva’s checklist (2000) enumerated 38 species<br />

for Cuba alone. When the second largest<br />

center of distribution, Hispaniola, is<br />

considered, the number of accepted species<br />

2. <strong>Coccothr<strong>in</strong>ax</strong><br />

cr<strong>in</strong>ita, a<br />

species known<br />

for its long<br />

fibers cover<strong>in</strong>g<br />

the trunk.<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

may be even greater. It is easy to see that this<br />

genus does <strong>in</strong>deed need a great deal of work<br />

to sort out the species. Complicat<strong>in</strong>g matters<br />

further is the fact that anecdotal evidence<br />

strongly suggests that a significant amount of<br />

hybridization has occurred among the species<br />

cultivated <strong>in</strong> South Florida, a situation mak<strong>in</strong>g<br />

taxonomic work based upon these <strong>in</strong>dividuals<br />

challeng<strong>in</strong>g, to say the least. A complete<br />

taxonomic review of <strong>Coccothr<strong>in</strong>ax</strong> would clarify<br />

the taxonomy of exist<strong>in</strong>g species and would be<br />

particularly welcome <strong>in</strong> the light of the current<br />

collect<strong>in</strong>g activity <strong>in</strong> Cuba and the Dom<strong>in</strong>ican<br />

Republic that may very well result <strong>in</strong> the<br />

discovery of new species.<br />

This article focuses on the cultivation of some<br />

of the attractive, but little known, taxa that<br />

have recently come <strong>in</strong>to South Florida. Because<br />

of the confusion regard<strong>in</strong>g names, the<br />

taxonomy of the genus cannot be ignored<br />

completely. This paper thus relies more closely<br />

on the work of Cuban botanists most familiar<br />

with the species <strong>in</strong> the wild than on other<br />

published accounts.<br />

For many years, only a handful of <strong>Coccothr<strong>in</strong>ax</strong><br />

species have ever been cultivated. In this paper,<br />

we def<strong>in</strong>e “cultivation” to mean species found<br />

<strong>in</strong> the landscape trade or <strong>in</strong> private collections<br />

(botanic gardens and <strong>in</strong>dividual collectors).<br />

3. <strong>Coccothr<strong>in</strong>ax</strong><br />

miraguama is<br />

appreciated for its<br />

attractive, persistent<br />

leaf base fibers.<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

4. <strong>The</strong> trunk of<br />

<strong>Coccothr<strong>in</strong>ax</strong><br />

spissa is<br />

swollen.<br />

Interest<strong>in</strong>gly, a few species are widely grown,<br />

while the many more species are practically<br />

unknown outside their countries of orig<strong>in</strong>.<br />

Some of these species deserve wider use<br />

because of their beauty.<br />

Presently, only about eight taxa are known to<br />

be grown for the commercial landscape<br />

market; a handful are be<strong>in</strong>g tried <strong>in</strong> the Canary<br />

Islands, but another 30 have gradually begun<br />

to be grown <strong>in</strong> South Florida. Proximity to<br />

wild seed sources, however irregular and<br />

limited <strong>in</strong> quantity, and similar climate and<br />

soil type, make South Florida well suited for<br />

cultivat<strong>in</strong>g and study<strong>in</strong>g the genus. <strong>The</strong>ir slow<br />

growth rate makes them challeng<strong>in</strong>g<br />

candidates for long term study, and for the<br />

same reason, few of the newly available species<br />

are likely to be embraced by the landscape<br />

trade. Several species that have moderately fast<br />

growth rates hold some promise. Warm and<br />

dry regions of Australia and Hawaii could also<br />

be well suited for grow<strong>in</strong>g this genus, but low<br />

soil pH <strong>in</strong> both locations limits <strong>Coccothr<strong>in</strong>ax</strong><br />

cultivation. Few species grow well <strong>in</strong> either<br />

location. Likewise, southern California does<br />

not seem to be a good habitat for most species<br />

because of its long and cool w<strong>in</strong>ters.<br />

For many years, <strong>Coccothr<strong>in</strong>ax</strong> argentata, C.<br />

barbadensis, C. cr<strong>in</strong>ita (Fig. 2), C. miraguama<br />

(Fig. 3) and C. spissa (Fig. 3) were the only<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

5. A suspected hybrid between<br />

<strong>Coccothr<strong>in</strong>ax</strong> and Zombia.<br />

species widely found <strong>in</strong> cultivation, although<br />

a handful of names (C. argentea, C. martii, C.<br />

alta) were <strong>in</strong>discrim<strong>in</strong>ately applied to plants<br />

available at local palm sales. Occasionally<br />

plants identified as C. boschiana, C. clarensis,<br />

C. cupularis, C. gracilis, C. fragrans, C. <strong>in</strong>aguensis,<br />

C. jamaicensis, C. readii and C. yuruguana have<br />

been sold at Fairchild Tropical Botanic Garden<br />

or South Florida <strong>Palm</strong> <strong>Society</strong> sales. However,<br />

very few other species have appeared <strong>in</strong><br />

cultivation until the past three years when<br />

collectors from around the Caribbean bas<strong>in</strong><br />

began palm prospect<strong>in</strong>g <strong>in</strong> Cuba and the<br />

Dom<strong>in</strong>ican Republic. As a result, seeds and<br />

seedl<strong>in</strong>gs of numerous taxa have begun to<br />

appear at local palm sales and offered on the<br />

Internet by commercial seed and plant<br />

vendors. However, because the taxonomy is<br />

<strong>in</strong> disarray and there are no mature specimens<br />

of these new “immigrants” <strong>in</strong> South Florida,<br />

it is important to acknowledge that<br />

identification of these newly offered species is<br />

tentative at best. In a few cases, quantities of<br />

“fresh seeds” have been offered from locations<br />

where the palms simply do not exist and/or<br />

many months after the plants f<strong>in</strong>ished their<br />

annual fruit<strong>in</strong>g.<br />

Collections of seeds from palms <strong>in</strong> the type<br />

localities and extensive photo collections have<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

Table 1. Species of <strong>Coccothr<strong>in</strong>ax</strong> <strong>in</strong> cultivation.<br />

<strong>Coccothr<strong>in</strong>ax</strong> species Native to Characteristics<br />

argentata P<strong>in</strong>e rocklands of South Florida <strong>in</strong>clud<strong>in</strong>g Florida Lax leaves dark green above and silver below; high<br />

Keys; coastal hammocks of southern Florida Keys; drought and salt tolerance; slow grow<strong>in</strong>g.<br />

Bahamas; Yucatan.<br />

barbadensis Coastal scrub from Lesser Antilles south to Large green leaves, may be slightly silver below;<br />

Tr<strong>in</strong>idad and Margarita Island, Venezuela. fastest grow<strong>in</strong>g of genus (seed to 6 m [20 ft.] <strong>in</strong> ten<br />

years).<br />

borhidiana Scrub forest on karst near coast, Matanzas, Cuba. Exceptionally stiff leaves with short petioles; leaves<br />

reta<strong>in</strong>ed for years – superficially resembl<strong>in</strong>g a<br />

m<strong>in</strong>iature Copernicia macroglossa; erect <strong>in</strong>florescences;<br />

South Florida specimens are f<strong>in</strong>ally produc<strong>in</strong>g large<br />

purple fruit after about 18 years of cultivation from<br />

seed; salt and drought tolerant.<br />

cr<strong>in</strong>ita Low serpent<strong>in</strong>e hill soils <strong>in</strong> central and western Large green leaves; long arch<strong>in</strong>g <strong>in</strong>florescences above<br />

Cuba; critically endangered. the crown with large burgundy fruit; persistent leaf<br />

sheaths form thick layer of fibers on trunk; critically<br />

endangered <strong>in</strong> the wild (


PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

6. Community pot<br />

of seedl<strong>in</strong>gs of<br />

Coccoth<strong>in</strong>rax<br />

argentata mak<strong>in</strong>g<br />

vigorous growth.<br />

been made to help identify the plants us<strong>in</strong>g the<br />

older <strong>Coccothr<strong>in</strong>ax</strong> literature. Buyers should<br />

realize that all identifications should be<br />

considered tentative until more extensive field<br />

work is done and the genus is formally revised.<br />

In the meantime, growers should keep accurate<br />

records of seed sources and their own<br />

cultivation experiences to contribute to a<br />

grow<strong>in</strong>g body of <strong>in</strong>formation on this genus of<br />

attractive palms.<br />

In the last few years, small quantities of seeds<br />

and nursery-grown seedl<strong>in</strong>gs labeled as C.<br />

alexandri, C. argentea, C. camagueyana, C.<br />

fragrans, C. ekmanii, C. garciana, C. gundlachii,<br />

C. hiorami, C. macroglossa, C. moaensis, C.<br />

montana, C. munizii, C. borhidana, C. nipensis,<br />

C. pauciramosa, C. pseudorigida, C. salvatoris<br />

and C. saxicola, have become available to<br />

growers. Because these species are “new to the<br />

trade” little is known about their cultural<br />

requirements. This paper, therefore, presents<br />

what is known at this time and identifies what<br />

may be some horticulturally valuable species<br />

worthy of greater use <strong>in</strong> the subtropical and<br />

tropical landscapes.<br />

Species <strong>in</strong> <strong>Cultivation</strong><br />

Because Florida is close to the center of<br />

distribution of the genus and has the wet/dry<br />

season and alkal<strong>in</strong>e soil preferred by most<br />

members of the genus, more <strong>Coccothr<strong>in</strong>ax</strong> (ca.<br />

40 taxa) are cultivated here than anywhere<br />

else. <strong>The</strong> majority of species are recent arrivals,<br />

and few fruit<strong>in</strong>g specimens exist. Anecdotal<br />

evidence suggests that what have been<br />

recognized as <strong>Coccothr<strong>in</strong>ax</strong> species (particularly<br />

those <strong>in</strong> the C. argentata – C. readii – C.<br />

<strong>in</strong>aguensis – C. litoralis and C. barbadensis – C.<br />

alta – C. readii – C. proctorii – C. jamaicensis<br />

groups outside of Cuba) can form fertile<br />

hybrids. This evidence suggests that each<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

group may be one species that displays a wide<br />

range of characteristics across its range rather<br />

than a guild of multiple species. Additionally,<br />

an <strong>in</strong>tergeneric and sterile <strong>Coccothr<strong>in</strong>ax</strong> ×<br />

Zombia hybrid has flourished at Fairchild<br />

Tropical Botanic Garden <strong>in</strong> Miami for decades<br />

(Fig 5). Obviously the application of DNA<br />

analysis to all <strong>Coccothr<strong>in</strong>ax</strong> could help clarify<br />

the picture.<br />

Table 1 presents the species that have been<br />

grown for the longest time <strong>in</strong> South Florida,<br />

as well as <strong>in</strong> Hawaii, Australia, the Canary<br />

Islands and elsewhere. <strong>Palm</strong>s known as C. alta<br />

and C. argentea are also <strong>in</strong> cultivation, but all<br />

the mature plants the authors have seen have<br />

horticultural orig<strong>in</strong>s and resemble either C.<br />

argentata or C. barbadensis, or share a mixture<br />

of characteristics that make identification<br />

challeng<strong>in</strong>g.<br />

Cultivat<strong>in</strong>g <strong>Coccothr<strong>in</strong>ax</strong><br />

<strong>Coccothr<strong>in</strong>ax</strong> are best grown from freshly<br />

collected ripe seeds. After remov<strong>in</strong>g the pulp,<br />

soak the seeds 1–3 days and discard any<br />

“floaters.” Dried seeds may be soaked for over<br />

a week to rehydrate them. Sow the seeds<br />

shallowly <strong>in</strong> a 10–15 cm deep community pot<br />

of well dra<strong>in</strong>ed medium kept <strong>in</strong> a warm, sunny<br />

space. Fresh seeds germ<strong>in</strong>ate best, but seeds<br />

can take up to a year to germ<strong>in</strong>ate. Be patient<br />

and do not throw the seeds out until they rot.<br />

If the soil mix breaks down, remove the<br />

ungerm<strong>in</strong>ated seeds, replace the mix and<br />

resow.<br />

7. <strong>The</strong> root<br />

mass of<br />

<strong>Coccothr<strong>in</strong>ax</strong><br />

hiorami is<br />

large and<br />

benefits from<br />

a large,<br />

roomy pot.<br />

64


PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

8. <strong>Coccothr<strong>in</strong>ax</strong><br />

boschiana is<br />

slow-grow<strong>in</strong>g<br />

but attractive,<br />

even as a<br />

young plant.<br />

Community pots may be divided when<br />

seedl<strong>in</strong>gs are at the two-leaf stage. Seedl<strong>in</strong>gs<br />

should be placed <strong>in</strong> pots deep enough to<br />

accommodate the long roots (Fig. 6)<br />

<strong>Coccothr<strong>in</strong>ax</strong> germ<strong>in</strong>ates remotely, so be sure to<br />

plant the seedl<strong>in</strong>g so the roots are below the<br />

soil and the bud of the seedl<strong>in</strong>g is above<br />

ground. Once aga<strong>in</strong>, a well-dra<strong>in</strong>ed medium is<br />

best and may be amended with crushed<br />

limestone rock or dolomite for dra<strong>in</strong>age and<br />

to ma<strong>in</strong>ta<strong>in</strong> an alkal<strong>in</strong>e root zone.<br />

<strong>Coccothr<strong>in</strong>ax</strong> benefits from over-pott<strong>in</strong>g – that<br />

is, plant<strong>in</strong>g <strong>in</strong> larger pots than most palms,<br />

because of their proportionately greater root<br />

mass (Fig. 7). Growth rates can be <strong>in</strong>creased by<br />

mov<strong>in</strong>g seedl<strong>in</strong>gs and juvenile plants <strong>in</strong>to pots<br />

one size greater than the usual sequence. Keep<br />

potted <strong>Coccothr<strong>in</strong>ax</strong> off the ground on benches<br />

to ensure good air circulation around the<br />

plants. Most species (except C. cr<strong>in</strong>ita, C.<br />

montana and C. yunquensis) will tolerate full<br />

sun even as seedl<strong>in</strong>gs. Growers <strong>in</strong> subtropical<br />

65


PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

9. <strong>Coccothr<strong>in</strong>ax</strong> borhidiana is a handsome palm for gardens.<br />

regions will also need to provide cold<br />

protection for these small potted palms,<br />

especially from dry w<strong>in</strong>ds and cold ra<strong>in</strong> that<br />

encourage fatal fungal <strong>in</strong>fections. Regular<br />

applications of a balanced time release palm<br />

fertilizer and micronutrient sprays will also<br />

promote growth and strengthen plants aga<strong>in</strong>st<br />

conditions of low humidity and cool w<strong>in</strong>ter<br />

temperature. Patience is also a requirement for<br />

grow<strong>in</strong>g palms <strong>in</strong> this genus as most are slowgrow<strong>in</strong>g,<br />

particularly for the first 3–5 years.<br />

Because most species are found <strong>in</strong> sunny,<br />

seasonally dry climates and <strong>in</strong> well dra<strong>in</strong>ed<br />

alkal<strong>in</strong>e soils, similar landscape conditions will<br />

promote the best growth when the palms are<br />

ready to be planted. Dig a large hole to<br />

accommodate future root growth and then<br />

water and mulch the newly planted palms to<br />

mimic their habitat. Plant<strong>in</strong>g at the beg<strong>in</strong>n<strong>in</strong>g<br />

of the ra<strong>in</strong>y season will help young plants get<br />

established. Keep mulch away from the trunk<br />

to avoid fungal <strong>in</strong>fection and fertilize lightly<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

10. <strong>Coccothr<strong>in</strong>ax</strong> camagueyana has silvery leaves with bluish cast.<br />

for the first six months with a palm fertilizer<br />

with N-P-K ratio of 2-1-3 and micronutrients.<br />

Liquid fertilizers may be used for plants <strong>in</strong> the<br />

ground or <strong>in</strong> pots but are unlikely to have<br />

much effect on slow-grow<strong>in</strong>g palms such as<br />

these.<br />

Many of the very attractive Cuban species are<br />

found <strong>in</strong> serpent<strong>in</strong>e soils rich <strong>in</strong> nickel and/or<br />

other heavy metals such as iron, chromium,<br />

cobalt and manganese. <strong>The</strong>se soils also feature<br />

high magnesium/calcium ratios with<br />

correspond<strong>in</strong>gly low levels of calcium,<br />

potassium and phosphorus. In fact, calcium<br />

levels <strong>in</strong> serpent<strong>in</strong>e soils are virtually zero.<br />

Serpent<strong>in</strong>e soils can be either acidic or alkal<strong>in</strong>e<br />

with older serpent<strong>in</strong>e areas tend<strong>in</strong>g to be more<br />

acidic and newer ones slightly more alkal<strong>in</strong>e.<br />

For the serpent<strong>in</strong>e species (C. sp. “azul”, C.<br />

camagueyana, C. clarensis, C. garciana, C.<br />

moaensis, C. pseudorigida and C. yuruguana)<br />

long-term conta<strong>in</strong>er culture may be the only<br />

viable option unless local soils are serpent<strong>in</strong>e<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

as well. Alternatively, us<strong>in</strong>g an iron and<br />

magnesium amendment once a month seems<br />

beneficial.<br />

Species worthy of wider cultivation<br />

<strong>The</strong> follow<strong>in</strong>g species are worthy of wider<br />

cultivation because of the characteristics listed<br />

and their prospective adaptability to<br />

subtropical or tropical conditions on well<br />

dra<strong>in</strong>ed alkal<strong>in</strong>e soils.<br />

C. alexandri – 13–20 m, robust, Cuban, coastal<br />

palm with large crown of stiff leaves with silver<br />

undersides.<br />

C. boschiana (Fig. 8) – slender, Dom<strong>in</strong>ican<br />

Republic, coastal species with leaves golden<br />

green above and silver below; heavy, coarse<br />

fiber; slow-grow<strong>in</strong>g.<br />

C. brevicr<strong>in</strong>is – Cuban palm with short trunk,<br />

short, coarse fibers and dense crown<br />

<strong>in</strong>termediate <strong>in</strong> “hair<strong>in</strong>ess” between C.<br />

borhidiana (Fig. 9) and C. cr<strong>in</strong>ita (Fig. 2); from<br />

a different part of Cuba with different soil (C.<br />

cr<strong>in</strong>ita comes from serpent<strong>in</strong>e area, C.<br />

brevicr<strong>in</strong>is from alkal<strong>in</strong>e calcareous soils.<br />

C. camagueyana (Fig. 10) – drought tolerant,<br />

serpent<strong>in</strong>e species good for pot culture,<br />

moderate growth rate, stiff leaves with silverblue<br />

cast on upper surface; similar to plants<br />

known as C. “azul” (Fig. 11).<br />

C. fragrans – Dom<strong>in</strong>ican Republic and Cuban<br />

species with stout trunk; close aff<strong>in</strong>ity to C.<br />

spissa.<br />

C. garciana – serpent<strong>in</strong>e species from Cuba with<br />

sp<strong>in</strong>y red leaf fibers to 2 cm wide, rigid round<br />

leaves on short petioles; exceptionally slow<br />

growth rate and soil requirements may limit<br />

this to long-term pot culture.<br />

C. gundlachii – robust, Cuban species with<br />

large, stiff leaves, erect <strong>in</strong>florescence and heavy,<br />

coarse fiber.<br />

C. litoralis – coastal Cuban species found <strong>in</strong><br />

sandy, wet, littoral scrub; high salt tolerance<br />

and moderate growth rate.<br />

C. macroglossa (Fig. 12) – robust, Cuban palm<br />

with large black fruit, stiff leaves that are silver<br />

underneath, coarse fibers.<br />

C. montana – an understory palm from the<br />

moist mounta<strong>in</strong>s of the central Dom<strong>in</strong>ican<br />

Republic; dark green leaves deeply divided;<br />

fast-grow<strong>in</strong>g; white fruit.<br />

11. <strong>The</strong> species known to growers as <strong>Coccothr<strong>in</strong>ax</strong> “azul” is one of the bluest species.<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

12. <strong>Coccothr<strong>in</strong>ax</strong> macroglossa has stiff leaves that are silvery underneath.<br />

C. munizii (Fig. 13) – a th<strong>in</strong>-stemmed, Cuban<br />

species with small, wedge-shaped leaves; t<strong>in</strong>y<br />

fruit; a good plant<strong>in</strong>g companion with cactus.<br />

C. pauciramosa – medium-sized, coastal, Cuban<br />

species with erect <strong>in</strong>florescence and heavy,<br />

coarse fiber; easy to cultivate.<br />

C. salvatoris – Cuban brackish forest species<br />

with a compact crown, wide leaf segments,<br />

heavy fiber and starburst-like leaves.<br />

Many questions rema<strong>in</strong><br />

A handful of other species (C. clarensis, C.<br />

cupularis (Fig. 14), C. ekmanii, C.<br />

guantanamensis, C. hiorami, C. <strong>in</strong>aguensis, C.<br />

jamaicensis, C. pseudorigida, C. readii, C. saxicola,<br />

C. yunquensis, and C. yuraguana) are so poorly<br />

represented <strong>in</strong> cultivation that identification<br />

of some of the plants may be questionable,<br />

particularly if the plants were grown from<br />

seeds collected <strong>in</strong> botanic gardens.<br />

Furthermore, the follow<strong>in</strong>g taxa rema<strong>in</strong> poorly<br />

known altogether or unknown <strong>in</strong> cultivation<br />

outside their country of orig<strong>in</strong> (Cuba, unless<br />

otherwise noted): C. acunana, C. baracoensis, C.<br />

bermudezii, C. concolor (Haiti, ext<strong>in</strong>ct?), C.<br />

elegans, C. fagildei (cluster<strong>in</strong>g with up to 100<br />

heads!), C. leonis, C. microphylla, C. moaensis,<br />

C. muricata, C. nipensis, C. orientalis, C. pumila,<br />

C. rigida, C. savannarum, C. tr<strong>in</strong>itensis, and C.<br />

victor<strong>in</strong>i) that their horticultural potential<br />

cannot be evaluated at this time.<br />

A further comment<br />

As we worked on this article, we realized that<br />

cultivat<strong>in</strong>g <strong>Coccothr<strong>in</strong>ax</strong> serves as a metaphor<br />

of life itself and offers many mean<strong>in</strong>gful<br />

lessons – patience is an important virtue,<br />

actions often take a long time to bear fruit and<br />

no matter how much we would like to see<br />

th<strong>in</strong>gs happen, Nature moves accord<strong>in</strong>g to its<br />

own timetable. <strong>Coccothr<strong>in</strong>ax</strong> seedl<strong>in</strong>gs need the<br />

right nutrients, just the right amount of water<br />

and sunlight and a healthy environment. <strong>The</strong>y<br />

can tolerate a fair amount of pests, <strong>in</strong>clud<strong>in</strong>g<br />

overprotective “parents.” Whenever our<br />

conversation turns to which species just<br />

produced its first divided leaf after years of<br />

growth, we are rem<strong>in</strong>ded of these lessons.<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

13. <strong>Coccothr<strong>in</strong>ax</strong> munizii is a<br />

slender palm with stiff<br />

leaves from the driest part<br />

of southeastern Cuba.<br />

Summary<br />

As more species of <strong>Coccothr<strong>in</strong>ax</strong> are cultivated,<br />

we hope that commercial growers and<br />

collectors will share their experiences <strong>in</strong> palm<br />

forums and <strong>in</strong> the literature so that these<br />

attractive palms will f<strong>in</strong>d greater use <strong>in</strong><br />

landscapes. We hope to see more <strong>in</strong> situ and<br />

ex situ conservation efforts, particularly for the<br />

Cuban species with limited distributions, <strong>in</strong><br />

order to buffer the effects of human activity,<br />

particularly coastal tourism, <strong>in</strong> Cuba and the<br />

Dom<strong>in</strong>ican Republic. Additionally, consistent<br />

practice of susta<strong>in</strong>able collection techniques<br />

and appropriate horticultural methods can also<br />

ensure the survival of <strong>Coccothr<strong>in</strong>ax</strong> <strong>in</strong> the wild<br />

and <strong>in</strong> cultivation. <strong>The</strong>se practices <strong>in</strong>clude<br />

collect<strong>in</strong>g a few seeds from many <strong>in</strong>dividuals<br />

rather than all the seeds of a few specimens,<br />

harvest<strong>in</strong>g only ripe seeds and process<strong>in</strong>g them<br />

promptly, match<strong>in</strong>g the cultural conditions to<br />

the plant’s own habitat as closely as possible,<br />

as well as carefully monitor<strong>in</strong>g these palms<br />

from sow<strong>in</strong>g to plant<strong>in</strong>g <strong>in</strong> the landscape. It<br />

is particularly important to obta<strong>in</strong> as much<br />

knowledge as possible of the ecology of these<br />

palms <strong>in</strong> the wild, as this <strong>in</strong>formation will help<br />

us determ<strong>in</strong>e each species’ suitability for<br />

cultivation and techniques of conservation.<br />

Acknowledgments<br />

<strong>The</strong> authors thank Dr. Scott Zona, <strong>Palm</strong><br />

Biologist at Fairchild Tropical Botanic Garden<br />

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PALMS Migliaccio & Reyes: <strong>Coccothr<strong>in</strong>ax</strong> Vol. 49(2) 2005<br />

14. <strong>Coccothr<strong>in</strong>ax</strong><br />

cupularis is still<br />

uncommon outside<br />

botanical gardens. This<br />

specimen grows at<br />

Fairchild Tropical<br />

Botanic Garden, Miami,<br />

Florida.<br />

and Paul Craft, President of IPS for their<br />

ongo<strong>in</strong>g encouragement of this project,<br />

assistance with translations of orig<strong>in</strong>al<br />

descriptions and their editorial suggestions.<br />

Many collectors and horticulturalists k<strong>in</strong>dly<br />

shared their experiences; notable among them<br />

were Mike Harris, Loxahatchee, Florida, USA;<br />

Dale Holton, Lake Worth, Florida, USA; Marcel<br />

Moreno, Miami, Florida, USA; Ken Neugent<br />

and Mike Davenport, Fairchild Tropical<br />

Botanic Garden, Miami, Florida, USA; Carlo<br />

Morici, Tenerife, Canary Islands; and the late<br />

Howard Waddell, Miami, Florida USA. All<br />

photos were taken by Chris Migliaccio unless<br />

otherwise noted.<br />

LITERATURE CITED<br />

MOYA L., C.E. AND A. T. LEIVA S. 2000. Checklist<br />

of the palms of Cuba, with notes on their<br />

ecology, distribution and conservation.<br />

<strong>Palm</strong>s 44: 69–84.<br />

71

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