First Contact - WWF

FIRST CONTACT 

in the Greater Mekong 

new species discoveries

1 

Acknowledgements 

The author would like to thank the following for their new discoveries, 

expert advice and generous contribution to this publication. 

Prof Dr Norhayati Ahmad: Universiti Kebangsaan Malaysia / Dr Dirk Ahrens: Natural 

History Museum / Dr Per Alström: Swedish University of Agricultural Sciences / Dr 

Arthur Anker: Smithsonian Tropical Research Institute / Dr Raoul Bain: America Museum 

of Natural History / Dr Aaron Bauer: Villanova University / Julianne Becker: WWF Greater 

Mekong Programme / Daicus Belabut: Universiti Malaya / Dr Diana Bell: University of 

East Anglia / Prof Wolfgang Böhme: Museum Alexander Koenig / Stuart Chapman: WWF 

Greater Mekong Programme / Dr Hongwei Chen: South China Agricultural University 

/ Nick Cox: WWF Greater Mekong Programme / Dr Gabor Csorba: Hungarian Natural 

History Museum / Trinh Viet Cuong: Fauna & Flora International / Dr Patrick David: 

Muséum National d’Histoire Naturelle / Pete Davidson: BirdLife International / Dr Peter 

Davie: Queensland Museum / Dr Jenny Daltry: Fauna and Flora International / Pakawin 

Dankittipakul: Chiang Mai University / Dr Mary Dawson: Carnegie Museum of Natural 

History / Dr Louis Deharveng: Museum National d’Histoire Naturelle / Dr Aljos Farjon: 

Royal Botanic Gardens Kew / Dr Carl Ferraris: Smithsonian Institution / Dr Charles M 

Francis: Canadian Wildlife Service / Nicole Frisina: WWF Greater Mekong Programme 

/ Gina Fullerlove: Royal Botanic Gardens Kew / Dr Mike Gee: Plymouth Marine Laboratory 

/ Dr Penelope Greenslade: Australian National University / Dr L Lee Grismer: La Sierra 

University / Cristian J Grismado: Museo Argentino de Ciencias Naturales / Dr Patrick 

Grootaert: Royal Belgian Institute of Natural Sciences / Marc-Alexander Gross: WWF 

Greater Mekong Programme / Dr Antonio Guillén-Servent: Instituto de Ecología / Prof 

Markku Häkkinen: Helsinki University Botanical Garden / Hong Hoangminh: WWF Greater 

Mekong Programme / Dr Mark Hughes: The Herbarium Singapore Botanic Gardens / 

Veronica Hunter: WWF-US / Dr Peter Jäger: Arachnology Research Institute and Natural 

History Museum Senckenberg / Dr Jiang Jianping: Chinese Academy of Sciences / Dr 

Darren Yeo Chong Jinn / National University of Singapore / Dr Rudy Jocqué: Royal 

Museum for Central Africa / Dr Maurice Kottelat: National University of Singapore / Dr 

Ulrich Kuch: Senckenberg Museum of Natural History / Dr Alan E Leviton: California 

Academy of Sciences / Dr Ai-Ping Liang: Institute of Zoology Chinese Academy of Sciences 

/ Dr Barney Long: WWF-US / Dr Wilson R Lourenço: Muséum National d’Histoire Naturelle 

/ Dr Darrin Lunde: American Museum of Natural History / Josef Margraf: Nature Products 

/ Dr Ivan N Marin: A N Severtzov Institute of Ecology and Evolution / Prof Matsui Masafumi: 

Kyoto University / Dr David J Middleton: Royal Botanic Garden Edinburgh / Christopher 

Milensky: Smithsonian Institution / Dr Hiroyuki Motomura: The Kagoshima University 

Museum / Dr Mark Newman: Royal Botanic Garden Edinburgh / Dr Heok Hee Ng: 

University of Michigan / Chan Kin Onn: Universiti Kebangsaan Malaysia / Prof Somsak 

Panha: Chulalongkorn University / Dr Olivier S G Pauwels: Smithsonian Institution / Le 

Khac Quyet: Fauna and Flora International / Dr Martín J Ramírez: Museo Argentino de 

Ciencias Naturales / Dr John H Rappole: Smithsonian National Zoological Park / Dr 

Dave Redfield: Florida State University / René Ries / Prof Robert W Murphy: University 

of Toronto / Prof Robert W Sites: University of Missouri / Dr Tyson Roberts: Smithsonian 

Tropical Research Institute / Dr Andrew J Ross: The Natural History Museum / Dr 

Andrew Short: University of Kansas / Dr Pipat Soisook: Prince of Songkla University / Dr 

Somran Suddee: Royal Botanic Gardens Kew / Montri Sumontha: Ranong Fisheries 

Department / Dr Heok Hui Tan: National University of Singapore / Philip Thomas: Royal 

Botanic Garden Edinburgh / Nguyen Quang Truong: Vietnamese Academy of Science 

and Technology / Kampol Udomrittiruj / Dr Chavalit Vidthayanon: WWF Greater Mekong 

Programme / Dr Harold Voris: Field Museum / Dr Gernot Vogel: Society for Southeast 

Asian Herpetology / Dr Van Wallach: Harvard University / Nicholas Wilkenson / Beck 

Woodrow: FSC / Dr Wolfgang Wüster: Bangor University / Dr Herbert Zettel: Natural 

History Museum Vienna / And special thanks to Dr Amy Lathrop: Royal Ontario Museum 

and Dr Thomas Ziegler: Vietnam Nature Conservation Project Cologne Zoo. 

The material and geographical designations in this report do not imply the 

expression of any opinion whatsoever on the part of WWF concerning the legal 

status of any country, territory, or area, or concerning the delimitation of its 

frontiers or boundaries. 

View across the Mekong River 

from Thailand / Lao PDR border 

© Gerald S Cubbit / WWF-Canon.

Executive Summary 

Some of the richest and most valuable habitats on Earth can be found in 

the Greater Mekong, a region comprising Cambodia, Lao People's 

Democratic Republic, Myanmar, Thailand, Vietnam and Yunnan Province 

of the People's Republic of China. 

A staggering 16 WWF Global 200 ecoregions, critical landscapes of 

international biological importance, can be found here. These precious 

landscapes are home to an estimated 20,000 species of plant, 1,200 bird 

species, 800 species of reptiles and amphibians, and 430 mammal 

species, including Asian elephants, tigers and one of only two populations 

of the critically endangered Javan rhino in the world. In addition to rare 

Irrawaddy dolphins, the Mekong River basin is estimated to house at least 

1,300 species of fish, including the Mekong giant catfish, one of the 

largest freshwater fish in the world. By length, the Mekong is the richest 

waterway for biodiversity on the planet, fostering more species per unit 

area than the Amazon. Many of the species are endemic to the region. 

Such is the extent of the Greater Mekong's biodiversity that new species 

continue to be described by science even today. Between 1997 and 2007 

“ 

at least 1,068 new species have been discovered in the Greater 

Mekong, two new species a week on average every year for the 

past 10 years (see Appendix). 

Opposite page, from left to right: Platostoma 

cambodgense, new plant species © Royal Botanic Gardens 

Kew; Leptobrachium smithi, new frog species © Chan Kin 

Onn; Pontonides ankeri, new shrimp species © Ivan Marin; 

Gumprecht’s green pitviper (Trimeresurus gumprechti), new 

species © Piyawan Niyomwan; Pseudopoda confusa, new 

spider species © Peter Jäger. 

This report celebrates the unique and fascinating species that can be found 

in this extraordinary region that lies between China to the east and India to 

the west. It also highlights many vital habitats that face growing pressures 

as a consequence of unsustainable development. The Greater Mekong is 

ranked as one of the top five most threatened biodiversity hotspots in the 

world by Conservation International. Accelerating economic development, 

population growth and increased consumption patterns of the wider Asia 

Pacific region are exposing the Greater Mekong to a range of destructive 

activities and practices. These include forest conversion for agricultural 

plantations, unsustainable logging and the illegal timber trade, wildlife trade, 

over fishing, dam and road construction, and mining. The region is also 

expected to be among the most vulnerable to, and hardest hit by global 

climate change, which will amplify the impacts of these threats. 

Achieving a balance between pursuing development and conserving natural 

resources presents the greatest challenge facing the region today. 

Economic development and environmental protection must be mutually 

supportive to provide for human security needs, reduce poverty and ensure 

the survival of the Greater Mekong's astonishing array of species and 

natural habitats. 

There is no time to lose. WWF has realised the need for a new layer of 

strategic actions to augment our longstanding implementation of field-level 

projects in the Greater Mekong. With governments and industry, WWF will 

work to conserve and sustainably manage 600,000 km 2 of transboundary 

forest and freshwater habitats in this unique and diverse land. 

2

3 

Greater Mekong 

Greater Biodiversity 

Geography 

The Greater Mekong comprises Cambodia, Lao People's Democratic Republic, Myanmar, 

Thailand, Vietnam and Yunnan Province of the People's Republic of China. Flowing 

through these countries some 4,500 kilometres is the mighty Mekong River, a regional 

life force that emerges from the vast and towering mountains surrounding the Tibetan 

plateau and ends in the tranquil waterways of the Mekong Delta in southern Vietnam, 

where it disperses into the South China Sea. The river is the critical link that unites 320 

million people 1 and supports an extraordinary level of species diversity and endemism, 

together creating one of the most culturally vibrant and biologically important regions in 

the world. 

Some of the richest and most valuable habitats on Earth can be found extending out 

across the Greater Mekong's impressive and diverse geographic landscape, from 

isolated massifs, plateaus and limestone karsts, to sweeping expanses of lowlands, 

fertile floodplains and deltas. Forests range from evergreen and semi-evergreen, mixed 

deciduous to deciduous dipterocarp, down to panoramic grasslands, swamp forests 

and mangroves. Freshwater habitats include fast-flowing rocky mountain streams and 

expansive wetlands, such as Tonlé Sap in Cambodia, Southeast Asia's largest 

freshwater lake. 

The Greater Mekong's high variation in geography and climatic zones supports an 

immense amount of biodiversity. The region features 16 WWF Global 200 ecoregions 2 , 

critical landscapes of international biological importance. This represents the greatest 

concentration of ecoregions on mainland Asia. 

The Greater Mekong is home to 

the world’s remaining 

Indochinese tigers © Elizabeth 

Kemf / WWF-Canon.

Biodiversity 4 

The Greater Mekong harbours 430 mammal species 3 , including the Asian 

elephant, Irrawaddy dolphin and Javan rhino, and is one of the last 

strongholds for the critically endangered Indochinese tiger with a 

population of 700-1,225 4 . Vietnam has 25 species and subspecies of 

primates including the critically endangered grey-shanked douc, 

endangered black-shanked douc and endangered red-shanked douc. 

Cambodia supports the largest remaining blocks of tropical dry forest in 

the region and with it comes a diverse range of mega-fauna. 

As many as 70 mammal species are endemic to the Greater Mekong 5 . 

Endemic species are those found only within a specific area, and in many 

of the remote habitats of the Greater Mekong, pockets of unique biodiversity 

have evolved and survived in total isolation, reliant on the continued 

existence of the habitats in their immediate vicinity. 

The region is home to 24 'Centres of Plant Diversity' as defined by the 

World Conservation Union (IUCN) 6 . The total vascular plant diversity of the 

Greater Mekong may be as high as 20,000 species, with conservative 

estimates suggesting that approximately 50 per cent of flowering and 

seed-bearing plants are endemic to the region 7 . So extraordinarily high is 

the plant diversity that experts believe that the complex merging of floras 

in the highlands of the area has no parallel in any other part of the world 8 . 

More than 1,200 bird species have been recorded in the Greater Mekong, 

of which approximately 10 per cent are endemic 9 . The region contains all 

or part of seven 'Endemic Bird Areas' defined by BirdLife International 10 , 

with Vietnam alone hosting an estimated 850 species 11 . 

Reptiles number nearly 520 species, 200 species of which are endemic 12 . 

The region also supports the most diverse non-marine turtle fauna in the 

world. Approximately 280 amphibian species are found here, with more 

than 150 being endemic 13 . Lao PDR supports over 165 species of 

amphibians and reptiles, including species such as the Rock and Burmese 

pythons, and King cobras. The country of Myanmar is no stranger to deadly 

snakes, with 46 dangerously venomous species of snake known to be 

resident 14 . 

The Mekong River basin is estimated to house over 1,300 species of fish, 

including the endemic Mekong giant catfish, one of the largest freshwater 

fish in the world, the giant Mekong barb and several species of giant 

stingray. By length, the Mekong is the richest waterway for biodiversity on 

the planet, fostering more species per unit area than the Amazon. Many of 

the species are endemic to the region. 

‘Camera trap’ photo of an 

Indochinese tiger moving 

through the Lower Mekong 

Dry Forests © WWF Greater 

Mekong Programme.

5 

Mekong Magic 

A decade of remarkable discoveries 

Despite explorations dating back to the mid-nineteenth century, knowledge of the biodiversity of 

the Greater Mekong remains relatively poor. The turbulent history of colonialism followed by war 

and conflict resulted in little research on flora and fauna. Beginning in the early 1990s, political 

stability and the development of a network of protected areas allowed new scientific information to 

emerge, at times resulting in the revision of previous species descriptions 15 . 

Discoveries of new mammal species during the 1990s focused the attention of the global 

conservation community once again on the extraordinary diversity of the Greater Mekong. After 50 

years in which only one new large mammal species had emerged worldwide, three new hoofed 

mammals were identified in the same region of Vietnam within four years 16 . Amongst these was the 

discovery of the saola in 1992, a bovine dwelling in the evergreen forests of the Annamite 

Mountains of Lao PDR and Vietnam which received widespread international interest and triggered 

a series of scientifically explosive expeditions into the jungles of the region. 

Unlike other places on Earth where much of the fauna and flora is known, even today new species 

continue to be discovered in the Greater Mekong. Between 1997 and 2007 at least 1,068 have been 

officially described by science as being newly discovered species. This includes 519 plants, 279 

fish, 88 frogs, 88 spiders, 46 lizards, 22 snakes, 15 mammals, four birds, four turtles, two 

salamanders and a toad (see Appendix). In addition to this number are perhaps thousands of 

new invertebrate species. 

The extent of the Greater Mekong's unique habitats has also hidden many species from scientific 

discovery. In the Greater Annamites ecoregion for example, extraordinary conditions allowed 

rainforest to persist during the last ice age, giving the forest and its species thousands of additional 

years of refuge to evolve in isolation. The result is one of the highest rates of endemism in a 

continental setting globally.

Cyrtodactylus phongnhakebangensis, 

named after the Phong Nha Ke Bang National 

Park, Vietnam, where this new species was 

discovered © Thomas Ziegler. 

Despite the wealth of recent discoveries, the biological 

diversity of the region has not yet been completely 

documented. Scientists believe we have only just 

scratched the surface. 

Lower Mekong Dry Forests, 

Cambodia, a WWF priority 

ecoregion © Nick Cox / 

WWF Greater Mekong 

Programme. 

6

7 

Amphibians 

The countries of the Greater Mekong have done much to further Southeast 

Asia's reputation as a "lost world" for amphibians. An astonishing 91 new 

species of amphibian have been described within the Greater Mekong region 

since 1997. The rate of frog discovery has been truly phenomenal; in Vietnam 

alone the known frog diversity has doubled since 1999 17 as a result of intensive 

scientific study on the fauna of the country. 

In 1998, the number of species in the Asian toadfrog genus Leptobrachium 

increased from four to six in the Mekong region with the discovery of two new 

species in Vietnam 18 . Leptobrachium xanthospilum is named after the Greek 

xanthos, meaning yellow, and the Greek spilos, meaning spot, referring to the 

conspicuous yellow spots this species bears. Leptobrachium banae, found by 

local Ba Na hunters in Gia Lia Province, has red-orange spots on the back, 

with bright red and black bands on the limbs. 

Two further species of frog were collected from the Song Gam and Tam Dao 

mountain ranges of northern Vietnam 19 . The highly endemic Leptolalax sungi 

was collected some 925 metres up and only found in the vicinity of a single 

stream passing through the east side of the village of Tam Dao, Vinh Phu 

Province. The species has iridescent 

gold-green eyes. Also with piercing gold 

eyes, Leptolalax nahangensis was 

found at the entrance of a cave in Na 

Hang Nature Reserve, Tuyen Quang 

Province. 

. 

Left: Rhacophorus 

cyanopunctatus 

© Chan Kin Onn. 

Theloderma licin © Daicus Belabut. 

Leptolalax sungi © Robert W Murphy. Leptolalax nahangensis © Amy Lathrop. 

Leptobrachium banae © Amy Lathrop. 

Leptobrachium xanthospilum © Amy Lathrop.

There has been an abundance of discoveries too in the Rana and 

Rhacophorus genera, known as pond and flying frogs respectively. In 2003, 

six new species of cascade frog were identified in Vietnam 20 . Three of 

these new species occur in montane forests in northern Vietnam (Rana 

bacboensis, Rana hmongorum and Rana daorum) and two are known 

only from the Tay Nguyen Plateau of Vietnam's Central Highlands (Rana 

banaorum and Rana morafkai). 

The unique Rana morafkai frogs are unusual in that they often turn brown 

at night, but during the day their entire body becomes green. In 

Cambodia, a new species of rhacophorid frog, Chiromantis samkosensis, 

was identified in 2007 from Phnom Samkos in the northwestern section of 

Chiromantis samkosensis © L Lee Grismer. 

the Cardamom Mountains and is distinguished from other species of Asian 

Chiromantis by having green blood and turquoise bones amongst other 

unique characteristics 21 . A rare rhacophorid in the region, Rhacophorus 

cyanopunctatus or the blue-spotted tree frog, can be found perching on 

leaves or branches beside slow moving streams in Phanom, Thailand 22 . 

Other newly identified residents of Thailand include the smooth-skinned 

wart frog, Theloderma licin 23 , and 'Smith's litter frog', Leptobrachium smithi 24 . 

The frog fauna of Lao PDR is poorly known relative to that of neighbouring 

China, Thailand, and Vietnam, but scientists have still been able to identify 

at least 46 species 25 . 

Rana daorum © Amy Lathrop. 

Leptobrachium smithi © Chan Kin Onn. 

Rana megatympanum © Amy Lathrop. 

Rana banaorum © Robert W Murphy. 

8

9 

Invertebrates 

Scientists say there may have been several thousands of new discoveries 

over the past 10 years in the Greater Mekong, most of which are 

invertebrates, representing the largest group of new species finds 26 . To count 

them all might well be an impossible task. This group ranges from the 

microscopic, to the parasitic, from the aquatic to the multi-legged, from 

timid to truly terrifying - it certainly is an exciting field of research. 

Most remarkable perhaps is the impressive list of at least 88 new species of 

spider, the majority of which have been found as a result of expeditions into 

Lao PDR, Thailand and Yunnan Province in China. At the forefront of this 

effort has been scientist Dr Peter Jäger, who described Heteropoda maxima 

from caves in Khammouan Province, Lao PDR, in 2001 27 . This find was 

particularly significant as with a colossal legspan of up to 30 centimetres, 

the species is the largest huntsman spider in the world 28 . 

The medium-sized spider species Pseudopoda confusa, described in 2006 

from northern Lao PDR, was collected by hand in the forests of Luang Nam 

Tha and Muang Sing districts. 

Top right: Storenomorpha 

anne © Peter Jäger. Right: 

Pseudopoda confusa 

© Peter Jäger. Left: 

Heteropoda maxima © 

Petra & Wilfried / 

Creative Commons.

“…individuals were quite aggressive 

and have bitten the two authors in their fingers, 

“Scientists say there may have been 

which was moderately painful.” several thousands of new discoveries 

Description of Heteropoda dagmarae (Jäger and Vedel, over 2005) the past 10 From years northern in the and Greater central Lao PDR, the large and aggressive Heteropoda dagmarae was 

Mekong, most of first which encountered. are inverte- Officially recorded as a new species in 2005, the nocturnal spider was 

brates” found in forests where it skillfully ambushes its prey from shrubs, trees, or bamboo around 

2-4 metres above the ground29 . 

These finds make an important contribution to the records of known spider species, 

particularly from Lao PDR, a country scientists say has been neglected in the past; in 

2006 the World Spider Catalog listed just five known species of spider from Lao PDR, 

compared with 2,428 from China 30 . 

Elsewhere, a shocking pink, spiny new species of "dragon millipede", Desmoxytes 

purpurosea, was described in 2007 from Lansak district, Uthaithani Province, Thailand. 

Several millipedes were found sitting and moving on limestone rocks and on the leaves of 

Arenga pinnata palms. Scientists suggest the stark bright colour is to alert would-be 

predators of the toxic animal, and they would do well to heed this warning - the millipede 

has glands that produce cyanide as a defensive mechanism. The species joins 

twenty-three other dragon millipedes of the genus Desmoxytes known from a large area in 

Southeast Asia, from southeastern China, south through Myanmar, Thailand and 

Vietnam 31 . A further four of these deadly dragon millipedes were described from Vietnam 

in 2005. 

Above, left: Desmoxytes 

purpurosea, new species 

of highly toxic shocking 

pink millipede © Somsak 

Panha. Left: Heteropoda 

dagmarae © Peter Jäger. 

10

11 

Plants 

There has been an unrivalled 519 plant discoveries in the Greater Mekong over the past ten 

years 32 , an incredible botanical bounty. 

The Chinese province of Yunnan, well known as the 'Kingdom of Plants' with as many as 

16,000 different species, has recently revealed some vibrant species of wild banana, 

including the purple Musa yunnanensis. This plant grows abundantly in the Mekong River 

watersheds on slopes from 500-1,800 metres, with its seeds being dispersed by monkeys 

and bats 33 . 

Among the new finds in the region have also been new trees. In the forests of northern 

Vietnam, Xanthocyparis vietnamensis, or the Golden Vietnamese cypress, was identified in 

2002 34 . Located on karst limestone mountains in Ha Giang Province, near the Chinese 

border, the conifer is closely related to the North American Nootka cypress. This species 

was first listed as Critically Endangered on the IUCN Red List in 2002, but today an 

estimated 560 individuals are known. This has given new hope for the species, but it 

continues to be extremely rare, only existing in an area of 10 km 2 . Although deforestation is 

occurring in the area due to logging, many trees are difficult to access as they grow on 

ridges and summits, aiding the survival of this species. 

Other plant species enjoy a broader distribution, for example the ghostly Anisochilus harmandii 

identified in 2004 can be found in Cambodia, Lao PDR and Thailand 35 . 

Despite harbouring as many as 11,000 species of flowering plants, some scientists 

consider Lao PDR to be one of the most botanically unexplored countries in Asia. There 

have been further significant finds recently however, for example in 2007 the beautiful plant 

Aeschynanthus mendumiae was recorded 850 metres up on the southeastern slopes of 

Phou Yang, in Nakai Nam Theun, Khammouan Province 36 . Also in Nakai Nam Theun, the 

delicate blue flowering Gentiana khammouanensis was discovered the same year 37 . Even 

with such a large number of hidden botanical secrets uncovered, scientists say that they 

still know very little about the plant species of the Greater Mekong. 

Anisochilus harmandii 

© Royal Botanic 

Gardens Kew. 

Plectranthus albicalyx 

© Royal Botanic 

Gardens Kew. 

Xanthocyparis 

vietnamensis conifer 

© Aljos Farjon.

Aeschynanthus mendumiae 

© Royal Botanic Garden Edinburgh. 

New species of wild banana, 

Musa yunnanensis 

© Nature Products. 

B i r d s 

A small jungle bird reminiscent of a wren, Jabouilleia naungmungensis, or the Naung Mung Scimitar-Babbler, 

was described from a remote part of Myanmar in 2005 38 . At the base of the Himalayas, in a temperate 

rainforest, a team of scientists first discovered the species, which has a long curved bill and relatively large 

feet. The only other member of this genus, the short-tailed scimitar-babbler (Jabouilleia danjoui), is native 

to parts of Lao PDR and Vietnam, although what is probably a third species, as yet unnamed, was recently 

found in north Vietnam 39 . 

Further new bird species discovered include the black-crowned barwing, Actinodura sodangorum 40 , and the 

golden-winged laughingthrush, Garrulax ngoclinhensis 41 . Both species were discovered in the Central 

Annamites, Vietnam, becoming the first new bird species to be identified in mainland Southeast Asia for 

over 30 years. The chestnut-eared laughingthrush, Garrulax konkakinhensis, was discovered in Vietnam in 

2001 42 . 

These new species join an already impressive number of birds found in the Greater Mekong. 

Left: Gentiana 

khammouanensis 

© Royal Botanic Garden 

Edinburgh. Right: Jabouilleia 

naungmungensis, the Naung 

Mung Scimitar-Babbler © 

Christopher Milensky. 

12

13 

Reptiles 

There have been 22 new snake additions to the animal kingdom over the past decade 

from the Greater Mekong, including species of pitvipers found throughout the Mekong 

countries of Cambodia, Lao PDR, Myanmar, Thailand, Vietnam and Yunnan Province in 

China. 

The species Trimeresurus vogeli was found in the rafters of a restaurant in the 

Headquarters of the Khao Yai National Park in Thailand 43 . The province of Nakhon Si 

Thammarat, Thailand, was the location for the discovery of the Siamese Peninsula pitviper, 

Trimeresurus fucatus, in 2004 44 . The Latin name fucatus is translated as "with 

make-up", in reference to the red and white hues on the cheeks of actors in the ancient 

Latin theatre, and similarly the white and red postocular streak is present in some 

males of this snake species. The same year, the Latin name nebularis was given to 

another new pitviper. Meaning "from the clouds", the name alludes to the cloudy montane 

rainforests, or cloud forests, inhabited by this species 45 . 

Another snake from Thailand, Enhydris chanardi, discovered in 2005 46 , is a late addition 

to the 22 other Oriental-Australasian species of aquatic snake. Most known localities 

for this species are in the freshwater environments of metropolitan Bangkok. Unlike 

vipers and elapids, the snake has venom-injecting fangs located at the back of its mouth. 

The country of Cambodia made a small but significant contribution to new snake 

discoveries in 2002, with the pocket-sized wolf snake Lycodon cardamomensis from the 

Cardamom mountain range, the most remote and least known part of Cambodia 47 . 

New members of the colubridae family of snakes have been described from Vietnam. 

The white-lipped keelback (Amphiesma leucomystax) surfaced in several locations in 

2007 48 including within Vietnam's Green Corridor, an area renowned for its high biodiversity. 

The species tends to live by streams where it catches frogs and other small animals. It 

has a beautiful yellow-white stripe that sweeps along its head and red dots cover its 

body. The discovery added to the country's already abundant number of ten species of 

Amphiesma. 

Siamese Peninsula 

pitviper,Trimeresurus 

fucatus © Gernot Vogel / 

Zootaxa 727: 1-63.

In 2005, a new species of krait was discovered in Lao Cai and Yen Bai 

Provinces, northern Vietnam 49 . The new snake, Bungarus slowinskii, is a 

member of the Elapidae, a highly venomous family of snakes that includes 

the black mamba, cobras, fierce snake and sea snakes. The snakes have 

long and slender bodies with smooth scales, but vary in colouration. Bungarus 

slowinskii has black and white rings covering the length of its body and tail. 

Another elapid, the Burmese spitting cobra (Naja mandalayensis) emerged 

in 2000 50 . The species is aggressive, with a tendency to spit venom when 

threatened. Although endemic to the arid region in central Myanmar, the 

species is closely related to the Thai spitting cobra. 

Red River krait 

(Bungarus slowinskii) 

© Q T Nguyen. 

Vogel’s green pitviper 

(Trimeresurus vogeli) 

© Montri Sumontha. 

Gumprecht’s green pitviper 

(Trimeresurus gumprechti) 

© René Ries. 

Andrea’s keelback 

(Amphiesma andreae) 

© Thomas Ziegler. 

Above: Burmese spitting cobra, Naja mandalayensis © California Academy of Sciences. 

White-lipped keelback 

(Amphiesma leucomystax) 


Kopstein’s bronzeback snake 

(Dendrelaphis kopsteini) 

© Chan Kin Onn. 

Cameron Highlands pitviper 

(Trimeresurus nebularis) 

© Gernot Vogel / Zootaxa 

727: 1-63. 

Cardamom wolf snake 

(Lycodon cardamomensis) 

© Jenny Daltry, FFI. 

14

15 

Gekko scientiadventura © Thomas Ziegler. 

Cnemaspis aurantiacopes © L Lee Grismer. 

Cnemaspis nuicamensis 

© L Lee Grismer. 

More than twice as many lizards have been described from the region than snakes over 

the past decade. At least 46 lizards join the ranks of the Greater Mekong's known reptile 

species. 

Recently in 2007, four new gekko species were recorded in forests in southern Vietnam 51 . 

Scientists suggest that the presence of these species in small, isolated mountains in the 

Mekong Delta flood plain and on one of the many small islands in Rach Gia Bay indicates 

that additional new species are likely to occur in these and similar underexplored regions 

in southern Vietnam 52 . 

Four new turtle species have also been recorded from the Greater Mekong in the last 10 

years. 

Cnemaspis tucdupensis 

© L Lee Grismer. 

Lygosoma boehmei 


Cnemaspis caudanivea 

© L Lee Grismer.. 

Far left, top: Gekko 

Left: Cyrtodactylus 

phongnhakebangensis 


tiadventura © Thomas 

Ziegler.

Fish 

Over 1,300 fish species occur in the Mekong River basin 

and each new scientific survey of the river and tributaries 

identifies new, often endemic, fish species with additional 

surveys of limestone caves, rapids, peat swamps and 

waterfalls, all adding to the known fish diversity of this 

mighty river. 

Between 1997 and 2007, an impressive 279 new species 

of fish were identified in the Greater Mekong. Amongst 

these was Polynemus bidentatus, or the Toothy Blackhand 

Paradise Fish, described in 2006 from the Cho Gao Canal 

in the Mekong River Delta, Vietnam 53 . 

In the Tapi basin, Thailand, Ellopostoma mystax was 

described in 2002 54 . Named after the Latin mystax, 

meaning moustache, there is a unique dark margin over 

the snout of the species. 

In addition, an abundance of catfish, cyprinids, fighting 

fish, gobies, loaches, stingrays and numerous other new 

species have been identified. 

Polynemus bidentatus 

© Vidthayanon Chavalit / 

WWF. 

Ellopostoma mystax 

© Kampol Udomrittiruj. 

Gagata gasawyuh 

© Vidthayanon 

Chavalit / WWF. 

Mekong River near Luang 

Prabang, Lao PDR © Peter 

Denton / WWF-Canon. 

16

17 

Mammals 

Southeast Asia is home to more than 500 species of mammal 55 , 430 of 

which can be found in the Greater Mekong 56 . Globally, new mammal 

discoveries are extremely rare, but the Greater Mekong is proving to be an 

anomaly, being a particular hotspot for new mammal finds. 

In 1998, scientists from WWF, Vietnam's Ministry of Agriculture and Rural 

Development and Da Nang University, described the Dark Annamite Muntjac, 

Muntiacus truongsonensis, from the Truong Son mountain range 57 . Although, 

no living specimen of the deer has been observed, scientists at the Institute 

of Zoology, University of Copenhagen, were able to analyse genetic tissue 

samples to confirm the muntjac as a new species. Possibly the smallest 

muntjac, weighing about 15 kilograms, the species is half the size of the 

common muntjac. Local villagers call the deer samsoi cacoong, meaning 

"the deer that lives in the deep, thick forest." In Myanmar, yet another 

species of muntjac, Muntiacus putaoensis, has been described by 

scientists in recent years 58 . 

Unlike some global regions that have yielded disappointing results when it 

comes to small mammal discoveries, the Greater Mekong has excelled. 

No less than 13 new species of small mammalia have been found here over 

the past decade, including bats, rats and shrews. In Vietnam and 

neighbouring Lao PDR, the Annamite striped rabbit was identified in 2000 59 . 

The furry black and brown striped species resemble the endangered Sumatran 

striped rabbit, the only other known striped rabbit. The find was extremely 

significant as it extended the known range of the genus Nesolagus more 

than 1,500 kilometres north from the island of Sumatra into mainland 

Southeast Asia. Although similar, genetic data indicates that the Sumatran 

and mainland Annamite populations have been isolated for millions of years. 

Above: Annamite 

striped rabbit, 

Nesolagus timminsi 

© Trinh Viet Cuong, 

Fauna and Flora 

International. 

Left: New rat 

species,Tonkinomys 

daovantieni 

© Darrin Lunde.

In 2007, the woolly bat Kerivoula titania was described from Seima Biodiversity Conservation 

Area, in Cambodia 60 . The species is believed to have a wide distribution across mainland 

Southeast Asia, as additional specimens have been found in 12 further locations in Lao PDR, 

Myanmar, Thailand and Vietnam. It is known from a variety of different forested habitats. Named 

after Titania, the Queen of the Fairies from William Shakespeare's "A Midsummer Night's Dream", 

the name was chosen to reflect the nymph-like nature of this forest bat. The woolly bat, Kerivoula 

kachinensis, was identified in 2004 from Myanmar 61 , but has since been revealed to inhabit a 

range of countries including Cambodia, Lao PDR, Thailand 62 and Vietnam 63 . Scientists fear that 

the new roundleaf bat, Hipposideros khaokhouayensis, may be at risk, as even though the 

species was found in a National Biodiversity Conservation Area, this habitat is being affected by 

infrastructure development 64 . 

Out of the several million species of animal that inhabit the planet, only a few thousand are 

mammals. With 15 new forest-dependent species identified in just 10 years, the Greater Mekong 

region certainly is an exciting and unexplored region for mammologists. 

Far left: 

Kerivoula 

kachinensis 

woolly bat © 

Pipat Soisook. 

Left: Kerivoula 

titania woolly 

bat © Gabor 

Csorba. 

Laotian rock rat (Laonastes aenigmamus) 

© David Redfield. 

A distant relative comes to dinner 

Described as a species new to science in 

2005, the Laotian rock rat (Laonastes 

aenigmamus) 65 or Kha-nyou, was first 

encountered by scientists on sale at an 

outdoor food market in Lao PDR. It was so 

unlike anything else, that they believed the 

species to be the first discovery of a new 

mammal family since 1974. Further 

investigation revealed however that Kha-nyou 

was in fact a striking example of the ''Lazarus 

effect'', whereby a species family that was 

formerly thought to be long extinct is 

rediscovered in the present day. It transpired 

that this particular species was remarkably 

the sole survivor of an ancient group of 

rodents understood to have disappeared 11 

million years ago 66 . The first report of a living 

Kha-nyou, except by hunters, was by Dr David 

Redfield who travelled to Lao PDR 

specifically to find and photograph a living 

specimen 67 . 

18

19 

With Great Biodiversity 

comes Great Responsibility 

What does the future hold for the many diverse and extraordinary species in 

the Greater Mekong? Will scientists continue to unearth fascinating new 

discoveries or will hundreds of species, many rare, silently disappear as 

more natural resources are consumed at unsustainable levels? 

Over the past 40 years the wider Asia Pacific region has trebled its 

consumption of natural resources 68 . This has resulted in 13 Asia Pacific 

countries having large ecological footprints, consuming more resources than 

are domestically available 69 . By now the impacts of this unsustainable 

development are evident in the Greater Mekong, with significant natural 

resources being exported to China. 

With the onset of peace in the 1990s, the area is now undergoing greater 

change than ever before. The Mekong countries are gradually shifting from 

subsistence farming to more diversified economies, and to more open, 

market-based systems, making the Greater Mekong a new frontier of Asian 

economic growth. According to the Asian Development Bank the Greater 

Mekong has the potential to be one of the world's fastest growing areas 70 , 

but despite the region's increasing economic transformation it remains poor. 

Population growth and economic development are putting considerable 

pressure on forest and marine habitats, species survival and the availability 

of freshwater. Widespread poverty is also driving the illegal trade in timber, 

wildlife and marine fish. The situation is becoming urgent. The Greater 

Mekong forms a large proportion of the Indo-Burma hotspot, spanning more 

than 2 million km² of tropical Asia. The Indo-Burma hotspot ranks as one of 

the top five most threatened biodiversity hotspots in the world, with only 

five per cent of its natural habitat remaining 71 . 

The Mekong River is the direct source of livelihoods and food for 60 million 

people living in its catchment. Cambodians alone capture two million tonnes 

of fish a year from the Mekong. The New Scientist recently reported that 

due to declining fish catches an astounding seven million water snakes 

were being harvested annually in Cambodia's Tonlé Sap, the largest 

freshwater lake in Southeast Asia 72 . Yet even though overfishing is a big 

threat to the wildlife of the Greater Mekong, there are even greater, more 

serious threats. Currently, there are 150 large hydropower dams at 

different stages of planning in the Greater Mekong 73 . Beyond direct 

biodiversity and habitat loss, dams can have a significant impact on fisheries 

and hydrology, as well as river and coastal erosion. 

Unique and irreplaceable rainforest habitats are also being eroded. Since 

the 1990s, net forest loss in Southeast Asia has totalled 2.7 million 

hectares annually 74 . The main cause of forest loss in the Greater Mekong 

is the establishment of plantations for the production of cacao, cashew 

nuts, coconut, coffee, palm oil, rubber, sugarcane, and tea 75 . Experts 

estimate that in recent years such conversion for cash crop plantations in 

Cambodia, Lao PDR, Myanmar, and to a lesser extent, the highlands of 

Vietnam and southern Yunnan Province, China, has had a much stronger 

impact on the remaining forest cover than logging and timber 

exploitation 76 . 

Opposite page: Unsustainable forestry 

practices continue in Southeast Asia 

© Alain Compost / WWF-Canon.

“Laos and Cambodia appear to be facing waves of 

forest conversion to cash crop plantations, 

often uncontrolled and linked to incentives 

for foreign investment and to foreign demand.” 

- European Commission (2007) 

20

21 

Forest clearance for plantations, as well as illegal, unregulated and poorly 

regulated logging, has opened up previously remote areas, exacerbating 

the illegal trade in timber and wildlife. 

A devastating 70 per cent of endemic mammals in the Greater Mekong are 

globally threatened 77 . These include the saola and five species of primate 

endemic to Vietnam that are listed among 25 of the world's most 

endangered primates. In addition, species such as the tiger, Asian elephant, 

banteng and gaur, are all severely threatened by overexploitation. 

Achieving a balance between pursuing development and conserving natural 

resources presents the greatest challenge facing the region today. Our 

future depends on finding ways to take better care of the ecosystems that 

support life on Earth, and on forging collaborations that can deliver big 

results for conservation and development. What is fundamentally clear is 

that poverty and the environment are inextricably linked. Economic 

development and environmental protection therefore must be mutually 

supportive to provide for human security needs, reduce poverty and ensure 

the survival of the Greater Mekong's astonishing array of species and 

natural habitats. 

Above: The skins of Indochinese tigers and other 

rare cats are openly displayed for sale in Cholon 

District, Ho Chi Minh City, Vietnam, October 2002 © 

Adam Oswell / WWF-Canon. Parts of at least 107 

tigers were recently observed in Myanmar’s wildlife 

markets during 12 surveys undertaken by TRAFFIC, 

the wildlife trade monitoring network 78 . 

Opposite page: Red River krait (Bungarus 

slowinskii), new snake species from Vietnam © Q T 

Nguyen.

The situation in Vietnam is critical. 

With 10 species recently vanishing 

altogether, a further 900 species 

are threatened with extinction. 

Source: IUCN (2008) 

22

23 Conclusions 

As the Greater Mekong becomes one of the fastest growing regions in the 

world, the challenge facing its governments is clear: they must sustain 

economic growth while simultaneously ensuring that natural ecosystems 

remain viable. 

The six nations along the Mekong River are caught up in an immense and 

irreversible current of change, being driven to a large degree by China's 

massive economic development. Transformation born of the Greater Mekong 

Subregion’s plan to promote free flows of goods and people across national 

borders is yielding economic benefits and regional integration on an 

unprecedented scale. Yet this same plan also threatens much of the 

region's unique biological and cultural diversity. 

The Greater Mekong Subregion grouping of countries is committed to 

increasing cooperation for accelerated economic development, as 

facilitated by the Asian Development Bank. Economic activity and 

associated investments in infrastructure development is concentrated along 

three "economic corridors" that criss-cross the region and has the 

potential both to lift the region's rural populations out of poverty but also to 

exacerbate existing threats, ultimately depleting the natural resource base 

upon which long-term development of the region depends. WWF believes 

that the Greater Mekong Subregion holds the key to both economic 

development and ensuring the integrity of conservation landscapes remains 

intact. 

Once-remote areas are poised to become development frontiers. 

Cross-border transport links and more stable political and investment 

climates are transforming rural backwaters in parts of resource-rich 

Cambodia and Lao PDR into investment hotbeds. Neighbouring countries 

are clambering over one another to secure concessions while start-up costs 

remain low. 

There is no time to lose. It is clear that the Greater Mekong Subregion’s 

road to regional economic development is neither sustainable nor 

equitable. At its current rate, it will result in irreversible grand-scale losses 

of biological and cultural diversity. A report by Oxfam 79 published in 2007 

concluded "the ability of natural resources to continue to support poor 

peoples' livelihoods in the Mekong is at a crisis point. Forests and rivers 

are in a state of rapid ecological decline caused by human 

over-exploitation". 

Many of WWF's established priority conservation landscapes rest squarely 

at the epicentres of the region's development plans, and so we have 

realised the need for a new layer of strategic actions to augment our 

longstanding implementation of field-level projects. WWF will work with 

governments and industry to develop an agreement that formalises 

the conservation of 600,000 km 2 of transboundary forest and 

freshwater habitats in the Greater Mekong and promotes sustainable 

landscape management as well as sets regional standards for sustainable 

infrastructure and climate change adaptation measures. 

W WF believes that real progress can be made in tackling huge 

poverty-impacting issues in the Greater Mekong, like global warming, 

deforestation, the illegal wildlife and timber trade, poor infrastructure 

development, and thereby ensure the availability of livelihoods, subsistence 

and freshwater to millions of people throughout the region. Companies that 

share WWF's commitment to sustainability and adopt challenging targets 

for change are already experiencing how partnerships with WWF can result 

in a win-win situation for the environment and profit margins. 

International problems always have a local dimension, and through efforts 

made locally by everyone across the globe, a significant difference can be 

made in this unique and diverse land.

Chiromantis samkosensis, 

new frog species © L Lee Grismer. 

24

25 

Recommendations 

Governments of the Greater Mekong should: 

Commit to a transboundary agreement spanning the countries of the Greater Mekong - Cambodia, Lao 

PDR, Myanmar, People's Republic of China, Thailand and Vietnam - that conserves and sustainably 

manages 600,000 km 2 of Biodiversity Conservation Landscapes. 

Governments in the Greater Mekong and in consumer countries should: 

Wildlife trade: Strengthen policy and legal frameworks and the capacity of national enforcement 

agencies needs to be increased to effectively reduce illegal wildlife trade. Long-term education and 

awareness campaigns are required to change consumer behaviour to help ensure that the 

consumption of wildlife is both legal and sustainable and without adverse impacts to wild populations 

of plants and animals. 

Climate change: Ensure that conservation, resource management, and development actions are 

climate smart, and directed to understanding how climate change will affect the region and assessing 

which ecosystems, resources, and people are most vulnerable. This information could then be used 

to creatively devise and mainstream proactive adaptation strategies into conservation, management, 

and development planning. The success of these efforts will depend on collaborative processes that 

build local adaptive capacity. 

Infrastructure: Develop guidelines and sustainability assessment tools that can be integrated into 

existing planning procedures and processes and move the input of sustainability considerations to 

earlier stages of the development cycle. Those initiatives will assist in energy master planning, transport 

master planning, water resources planning, economic development planning, ensuring that the 

biodiversity and natural resources aspects are adequately taken into account. It will also contribute 

to demonstrating that understanding natural processes will allow to plan better energy 

infrastructures, thus improving the living conditions of the people of the Greater Mekong and 

benefiting the economy.

“The Indo-Burma hotspot ranks 

as one of the top five 

most threatened hotspots 

in the world, with only 

five per cent of 

its natural habitat remaining.” 

- Conservation International (2007) 

Governments of timber product importing countries should: 

Introduce legislation that makes it illegal to import wood products from illegal sources. Ensure this 

legislation covers all wood products, including secondary-processed products and paper. Place the 

onus on companies to demonstrate compliance with the requirements of the law. Provide for a 

standardised legal verification system that companies must apply, including effective traceability and 

chain of custody systems, such as those promoted by the Forest Stewardship Council (FSC). Ensure 

regional political initiatives e.g. East Asia FLEG (Forest Law Enforcement and Governance) and EU 

FLEGT, adopt similar appropriate measures. 

Forest products industry should: 

Respect High Conservation Value Areas (HCVAs) and ensure illegal and unsustainable products are 

excluded from entering the global supply chain. Operations, exports and imports of products, should be 

certified by the FSC, Roundtable on Sustainable Palm Oil (RSPO) or Rainforest Alliance, to ensure wood, 

paper, palm oil and other plantation commodities are from a legal and sustainable source and not 

inadvertently contributing to habitat or species destruction in the Greater Mekong. Forest industries can 

seek further WWF guidance and establish greater market-links by joining the RSPO or Global Forest & 

Trade Network (GFTN), now present in 30 countries around the globe, including in China, Lao PDR, 

Vietnam and most major timber producing and consuming nations. 

Banks and lending institutions should: 

Move beyond the Equator Principles and establish and implement environmental reporting, 

assessment, management, and risk evaluation systems, if they are to be a reliable, effective and 

profitable catalyst for sustainable development in the Greater Mekong. 

Left: Forests of the Greater Annamites, 

Vietnam, a priority WWF ecoregion © WWF 

Greater Mekong Programme. 

26

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78 Chris R. Shepherd and Vincent Nijman (2008): The wild cat trade in Myanmar. TRAFFIC 

Southeast Asia, Petaling Jaya, Selangor, Malaysia. 

79 Cornford, J. & Matthews, N. (2007) Hidden Costs: The underside of economic transformation 

in the Greater Mekong Subregion. Australia: Oxfam Australia.

Appendix 

Note: New species will invariably arise as a result of the scientific 

community reviewing and reclassifying previously known species (e.g. 

subspecies). A selection of the species listed in the Appendix will have been 

newly described using this method. 

AMPHIBIANS 

FROGS 

Species Author Year Country 

Amolops bellulus Li u, Yang, Ferraris & Matsui 2000 Myanmar / Yunnan (China) 

Amolops caelumnoctis Rao & Wilki nson 2007 Yunnan (China) 

Amolops cremnobatus 

Amolops minutus 

Inger & Kottelat 

Orlov & Cuc 

1998 

2007 

Lao PDR / Vi etnam 

Vietnam 

Amolops panhai 

Amolops spinapectoral is 

Matsui & Nabhitabhata 

Inger, Orlov & Darevsky 

2006 

1999 

Thailand 

Vietnam 

Amolops splendissimus Orlov & Cuc 2007 Vietnam 

Amolops tuberodepressus Li u & Yang 2000 Yunnan (China) 

Ansonia inthanon Matsui, Nabhi tabhata & Panha 1998 Thailand 

Ansonia kraensis Matsui, Khonsue & Nabhitabhata 2005 Thailand 

Brachytarsophrys platyparietus Rao & Yang 1997 Yunnan (China) 

Chirixalus ananjevae Matsui, Or lov 2004 Vietnam 

Chirixalus punctatus Wilkinson, Win, Thin, Lwin, Shein & 2003 Myanmar 

Tun 

Chiromantis samkosensis Grismer, Neang, Chav & Holden 2007 Cambodia 

Fejervarya triora Stuart, Chuaynkern, Chan-ard, Inger 2006 Thailand 

Huia absita Stuart & Chan-ard 2005 Lao PDR 

Huia melasma Stuart & Chan-ard 2005 Thailand 

Leptobrachium banae Lathrop, Murphy, Orlov & Ho 1998 Vietnam / Lao PDR 

Leptobrachium buchardi Ohler, Teynié & David 2004 Lao PDR 

Leptobrachium huashen Fei L, Ye C, Ji ang J, Xie F, Huang Y 2005 Yunnan (China) 

Leptobrachium mouhoti Stuart, Sok & Neang 2006 Cambodia / Vietnam 

Leptobrachium smithi Matsui, Nabhi tabhata & Panha 1999 Thailand 

Leptobrachium xanthospilum 

Leptolalax fuligi nosus 

Lathrop, Murphy, Orlov & Ho 

Matsui 

1998 

2006 

Vietnam 

Thailand 

Leptolalax melanoleucus 

Leptolalax nahangensis 

Matsui 

Lathrop, Murphy, Orlov & Ho 

2006 

1998 

Thailand 

Vietnam 

Leptolalax pluvialis Ohler, Marquis, Swan & Grosjean 2000 Vietnam 

Leptolalax solus Matsui 2006 Thailand 

Leptolalax sungi Lathrop, Murphy, Orlov & Ho 1998 Vietnam 

Leptolalax tuberosus Inger, Orlov & Darevsky 1998 Vietnam 

Limnonectes bannaensis Ye, Fei, Xie & Jiang 2007 Yunnan (China) 

Megophrys auralensis Ohler, Swan & Daltry 2002 Cambodia 

Megophrys lekaguli Stuart, Chuaynkern, Chan-ard, Inger 2006 Thailand 

Microhyla marmorata 

Microhyla nanapollexa 

Bain & Nguyen 

Bain & Nguyen 

2004 

2004 

Vietnam 

Vietnam 

Microhyla pulverata Bain & Nguyen 2004 Vietnam 

Odorrana aureola Stuart, Chuaynkern, Chan-ard, Inger 2006 Thailand 

Odorrana jingdongensis Fei,Ye & Li 2001 Yunnan (China) 

Odorrana junlianensis Huang, Fei & Ye 2001 Yunnan (China) / Vietnam / Lao 

PDR 

Ophryophryne gerti Ohler 2003 Vietnam / Lao PDR 

Ophryophryne hansi Ohler 2003 Vietnam 

Ophryophryne synoria 

Oreolalax nanjiangensis 

Stuart, Sok & Neang 

Fei &Ye 

2006 

1999 

Cambodia 

Yunnan (China) 

Philautus abditus Inger, Orlov & Darevsky 1999 Vietnam 

Philautus cardamonus Ohler, Swan & Daltry 2002 Cambodia 

Philautus petilus Stuart & Heatwole 2004 Lao PDR 

Philautus supercornutus Orlov, Ho & Nguyen 2004 Vietnam 

Philautus truongsonensis Orlov & Ho 2005 Vietnam 

Rana archotaphus Inger & Chanard 1997 Thailand 

Rana attigua Inger, Orlov & Darevsky 1999 Vietnam / Lao PDR 

Rana bacboensis Bain, Lathrop, Murphy & Ho 2003 Vietnam 

Rana banaorum Bain, Lathrop, Murphy & Ho 2003 Vietnam 

Rana banjarana Leong & Lim 2003 Thailand 

Rana bannanica 

Rana bolavensis 

Rao & Yang 

Stuart & Bain 

1997 

2005 


Lao PDR 

Rana compotrix Bain, Stuart, Orlov 2006 Lao PDR 

Rana cucae Bain, Stuart & Orlov 2006 Vietnam

Rana daorum Bain, Lathrop, Murphy & Ho 2003 Vietnam 

Rana faber Ohler, Swan & Daltry 2002 Cambodia 

Rana gigatympana Orlov, Ananjeva, & Ho 2006 Vietnam 

Rana heatwolei Stuart & Bain 2005 Lao PDR 

Rana hmongorum Bain, Lathrop, Murphy & Ho 2003 Vietnam 

Rana indeprensa Bain & Stuart 2005 Thai land 

Rana iriodes Bain & Nguyen 2004 Vietnam 

Rana khalam Stuart, Orlov, & Chan-ard 2005 Vietnam / Lao PDR 

Rana lini Chou 1999 Yunnan (China) 

Rana megatympanum Bain, Lathrop, Murphy & Ho 2003 Vietnam 

Rana morafkai Bain, Lathrop, Murphy & Ho 2003 Vietnam 

Rana mortenseni Ohler, Swan & Daltry 2002 Cambodia / Thailand 

Rana orba Stuart & Bain 2005 Vietnam 

Rana trankieni Orlov, Le & Ho 2003 Vietnam 

Rana vitrea Bain, Stuart, Orlov 2006 Lao PDR 

Rhacophorus baliogaster Inger, Orlov & Darevsky 1999 Vietnam / Lao PDR 

Rhacophorus cyanopunctatus Manthey & Steioff 1998 Thai land 

Rhacophorus duboisi Orler, Marquis, Swan & Grosjean 2000 Vietnam / Yunnan (China) 

Rhacophorus exechopygus Inger, Orlov & Darevsky 1999 Vietnam / Lao PDR 

Rhacophorus hoangl ienensis Orlov, Lathrop, Murphy & Ho 2001 Vietnam 

Rhacophorus htunwini 

Wilkinson, Thin Thin, Kyi Soe Lwin, & 

Awan Khwi Shein 

2005 Myanmar 

Rhacophorus jaruj ini Matsui & Panha 2006 Thai land 

Rhacophorus kio Ohler & Delor me 2006 Yunnan (China), Lao PDR, 

Thai land 

Rhacophorus orlovi Ziegler & Köhler 2001 Lao PDR / Thailand / Vietnam 

Theloderma licin McLeod & Norhayati 2007 Thai land 

Theloderma ryabovi Orlov, Dutta, Ghate, Kent 2006 Vietnam 

Vibrissaphora echinata Dubois & Ohler 1998 Vietnam 

Vibrissaphora ngoclinhensis Orlov 2005 Vietnam 

Vibrissaphora promustache Rao, Wilki nson & Zhang 2006 Yunnan (China) 

Xenophrys daweimontis Rao & Yang 1997 Yunnan (China) 

Xenophrys lekaguli Stuart et al 2006 Thai land 

Subtotal 88 

TOADS 

Bufo crocus Wogan et al 2003 Myanmar 

Subtotal 1 

SALAMANDERS 

Paramesotriton laoensis Stuart & Papenfuss 2002 Lao PDR 

Tylototriton vi etnamensis Böhme, Schöttler, Nguyen & Köhler 2005 Vietnam 

Subtotal 2 

TOTAL NUMBER OF AMPHIBIANS 91 

BIRDS 


Actinodura sodangorum Eames et al 1999 Vietnam 

Garrulax konkakinhensis Eames & Eames 2001 Vietnam 

Garrulax ngoclinhensis Eames et al 1999 Vietnam 

Jabouilleia naungmungensis Rappole et al 2005 Myanmar 

TOTAL NUMBER OF BIRDS 4 

FISH 


Acrochordichthys gyrinus Vidthayanon & Ng 2003 Thai land 

Acropoma argentistigma Okamoto & Ida 2002 Thai land 

Acrossocheilus xamensis Kottelat 2000 Lao PDR 

Akysis clavulus Ng & Freyhof 2003 Vietnam 

Akysis ephippi fer Ng & Kottelat 1998 Cambodia / Lao PDR 

Akysis ful iginatus Ng & Rainboth 2005 Cambodia 

Akysis hardmani Ng & Sabaj 2005 Thailand 

Akysis longi fili s Ng 2006 Myanmar 

Akysis pictus Ng & Kottelat 2004 Myanmar 

Akysis prashadi Ng & Kottelat 2004 Myanmar 

Akysis recavus Ng & Kottelat 1998 Lao PDR / Thai land 

Akysis similis Ng & Kottelat 1998 Vietnam 

Akysis subti lis Ng & Kottelat 1998 Lao PDR / Thai land 

Akysis varius Ng & Kottelat 1998 Cambodia/ Lao PDR / Thailand 

Akysis vespa Ng & Kottelat 2004 Myanmar 

Amblyceps carinatum Ng 2005 Myanmar 

Amblyceps mucronatum Ng & Kottelat 2000 Thailand / Lao PDR 

Amblyceps serratum Ng & Kottelat 2000 Cambodia / Lao PDR 

Amblyceps foratum Ng & Kottelat 2000 Thailand / Cambodia 

Amblyceps platycephalus Ng & Kottelat 2000 Thailand 

Amblyceps variegatum Ng & Kottelat 2000 Thailand 

Amblyrhynchichthys micracanthus Ng & Kottelat 2004 Cambodia / Lao PDR / Thailand / 

Vietnam 

Ayarnangra estuarius Roberts 2001 Myanmar 

Bagrichthys majusculus Ng 2002 Thailand 

Bagrichthys obscurus 

Ng 1999 Cambodia / Lao PDR / Thailand / 

Vietnam 

Balantiocheilos ambusticauda Ng & Kottelat 2007 Cambodia / Thailand / Vietnam 

Balitora nantingensis Chen, Cui & Yang 2005 Yunnan 

Bangana elegans Kottelat 1998 Lao PDR 

Batasio elongates Ng 2004 Myanmar 

Batasio ferumi natus Ng & Kottelat 2007 Myanmar 

Batasio tigrinus Ng & Kottelat 2002 Thailand 

Belodontichthys truncatus Kottelat & Ng 1999 Cambodia / Lao PDR / Thailand / 

Vietnam 

Betta pallida Schindler & Schmidt 2004 Thailand 

Betta pi Tan 1998 Thailand 

Botia kubotai Kottelat 2004 Myanmar 

Botia udomritthiruji Ng 2007 Myanmar 

Brachygobius mekongensis Larson & Vidthayanon 2000 Cambodia / Lao PDR / Thailand 

Caelatoglanis zonatus Ng & Kottelat 2005 Myanmar 

Celestichthys margaritatus Roberts 2007 Myanmar 

Cephalopholis polyspila Randall & Satapoomin 2000 Thailand 

Ceratoglanis pachynema Ng 1999 Thailand 

Channa ornatipinni s Britz 2007 Myanmar 

Channa panaw Musikasinthorn 1998 Myanmar 

Channa pulchra Britz 2007 Myanmar 

Chaudhuria fusipinnis Kottelat & Bri tz 2000 Lao PDR 

Cl upisoma nujiangense Chen, Ferraris & Yang 2005 Yunnan 

Cl upisoma roosae Ferraris 2004 Myanmar 

Coius pulcher Kottelat 1998 Lao PDR 

Crossocheilus atrilimes Kottelat 2000 Lao PDR 

Danio kyathit Fang 1998 Myanmar 

Danio maetaengensis Fang 1997 Thailand 

Danio roseus Fang & Kottelat 2000 Lao PDR / Thai land 

Danio fangfangae Kottelat 2000 Lao PDR 

Danio gibber Kottelat 2000 Lao PDR 

Danio salmonata Kottelat 2000 Lao PDR 

Danionella mirifica Britz 2003 Myanmar 

Devario acrostomus Fang & Kottelat 1999 Lao PDR 

Devario apopyris Fang & Kottelat 1999 Lao PDR 

Devario leptos Fang & Kottelat 1999 Lao PDR 

Di enbienia namnuaensis Nguyen & Nguyen 2002 Vietnam 

Doryichthys contiguus Kottelat 2000 Lao PDR 

Ellopostoma mystax Tan & Lim 2002 Thailand 

Erromyzon compactus Kottelat 2004 Vietnam 

Eutropiichthys salweenensis Ferraris & Vari 2007 Myanmar / Thailand 

Feia ranta Winterbottom 2003 Vietnam

Gagata gasawyuh Roberts & Ferraris 1998 Myanmar 

Gagata melanopterus Roberts & Ferraris 1998 Myanmar 

Garra bispinosa Zhou, Pan & Kottelat 2005 Yunnan 

Garra flavatra Kullander & Fang Fang 2004 Myanmar 

Garra micropulvinus Zhou, Pan, & Kottelat 2005 Yunnan 

Garra nigricollis Kullander & Fang Fang 2004 Myanmar 

Garra poecilura Kullander & Fang Fang 2004 Myanmar 

Garra propulvinus Kullander & Fang Fang 2004 Myanmar 

Garra rakhinica Kullander & Fang Fang 2004 Myanmar 

Garra robustus Zhang & Chen 2002 Yunnan 

Garra spilota Kullander & Fang Fang 2004 Myanmar 

Garra vittatula Kullander & Fang Fang 2004 Myanmar 

Garra cyrano Kottelat 2000 Lao PDR 

Garra theunensis Kottelat 1998 Lao PDR 

Gerres chrysops Iwatsuki, Kimura & Yoshino 1999 Thai land 

Gerres infasciatus Iwatsuki & Kimura 1998 Thai land 

Glyptothorax panda Ferraris & Britz 2005 Myanmar 

Helicophagus leptorhynchus Ng & Kottelat 2000 Thai land / Vietnam 

Hemiarius verrucosus Ng 2003 Lao PDR / Thailand 

Hemibagrus imbrifer Ng & Ferraris 2000 Myanmar / Thailand 

Hemibagrus spilopterus 

Hemibagrus variegatus 

Ng & Rainboth 

Ng & Ferraris 

1999 

2000 

Cambo dia 

Myanmar / Thailand 

Hemimyzon ecdyonuroides Freyhof & Herder 2002 Vietnam 

Hemimyzon nanensi s Doi & Kottelat 1998 Thai land 

Hemimyzon confluens Kottelat 2000 Lao PDR 

Hemimyzon khonensis Kottelat 2000 Lao PDR 

Hemimyzon papilio Kottelat 1998 Lao PDR 

Himantura kittipongi Vidthayanon & Roberts 2006 Thai land 

Homaloptera confuzona Kottelat 2000 Lao PDR 

Homaloptera parclitella Tan & Ng 2005 Thai land 

Indostomus crocodylus Britz & Kottelat 1999 Thai land 

Indostomus spinosus Britz & Kottelat 1999 Cambo dia / Lao PDR / Thailand / 

Vietnam 

Kryptopterus dissi tus Ng 2001 Cambo dia / Lao PDR / Thailand 

Kryptopterus gemi nus 

Ng 2003 Cambo dia / Lao PDR / Thailand / 

Vietnam 

Kryptopterus paraschilbeides Ng 2003 Cambo dia / Vietnam 

Laocypris hispida Kottelat 2000 Lao PDR 

Macropodus erythropterus Freyhof & Herder 2002 Vietnam 

Makararaja chindwinensis Roberts 2007 Myanmar 

Microrasbora kubotai Kottelat & Witte 1999 Thai land 

Microrasbora nana Kottelat & Witte 1999 Myanmar 

Monotrete turgidus Kottelat 2000 Lao PDR 

Myripristis astakhovi Kotlyar 1997 Vietnam 

Mystacol eucus ectypus Kottelat 2000 Lao PDR 

Mystus armiger Ng 2004 Thai land 

Mystus castaneus Ng 2002 Thai land 

Mystus falcari us Chakrabarty & Ng 2005 Myanmar 

Nemacheilus arenicolus Kottelat 1998 Lao PDR 

Nemacheilus banar Freyhof & Serov 2001 Vietnam 

Nemacheilus cleopatra Freyhof & Serov 2001 Vietnam 

Neolissochilus baoshanensis Chen & Yang 1999 Yunnan 

Neolissochilus heterostomus Chen & Yang 1999 Yunnan 

Neolissochilus subterraneus Vidthayanon, and Kottelat 2003 Thai land 

Odontamblyopus rebecca Murdy & Shibukawa 2003 Vietnam 

Odontobutis aspro Kottelat 1998 Lao PDR 

Onychostoma fusiforme Kottelat 1998 Lao PDR 

Onychostoma meridionale Kottelat 1998 Lao PDR 

Oreoglanis hypsiurus Ng & M. Kottelat 1999 Lao PDR 

Oreoglanis immaculatus Kong, Chen & Yang 2007 Yunnan 

Oreoglanis infulatus Ng & Freyhof 2001 Vietnam 

Oreoglanis jingdongensis Kong, Chen & Yang 2007 Yunnan 

Oreoglanis lepturus Ng & Rainboth 2001 Lao PDR 

Oreoglanis macronemus Ng 2004 Lao PDR 

Oreoglanis setiger Ng & Rainbo th 2001 Lao PDR /Yunnan (China) 

Oryzias haugiangensis Roberts 1998 Vietna m 

Oryzias pectoralis Roberts 1998 Lao PDR 

Oryzias uwai Roberts 1998 Mya nmar 

Osteochilus striatus Kottelat 1998 Lao PDR 

Pangio lumbriciformis Britz & Maclaine 2007 Mya nmar 

Pangio signicauda Britz & Maclaine 2007 Mya nmar 

Papuligobius uniporus Chen & Kottelat 2003 Lao PDR 

Paralepidocep halus guishanen sis Li 2004 Yunnan 

Parambassis pulcinell a Kottelat 2003 Mya nmar 

Paraprotomyzon ni ulanji angensis Lu, Lu & Mao 2005 Yunnan 

Parasikukia maculata Doi 2000 Tha iland 

Parasphaerichthys lineatus Britz & Kottelat 2002 Mya nmar 

Pareuchilog lanis abbreviatus Li, Zhou, Thomson, Zhang & Ya ng 2007 Yunnan 

Pareuchilog lanis prolixd orsal is Li, Zhou, Thomson, Zhang & Ya ng 2007 Yunnan 

Pillaia kachi nica Kul lander, Britz & Fang 2000 Mya nmar 

Polydactylus siame nsis Motomura, Iwatsuki & Yo shi no 2001 Tha iland 

Polynemus bidentatus Motomura & Tsukawaki 2006 Vietna m 

Poropuntius genyognathus Roberts 1998 Mya nmar 

Poropuntius hathe Roberts 1998 Tha iland 

Poropuntius heterolepid otus Roberts 1998 Tha iland 

Poropuntius melanogrammus Roberts 1998 Tha iland 

Poropuntius scapanognathus Roberts 1998 Tha iland 

Poropuntius angustus Kottelat 2000 Lao PDR 

Poropuntius conste rnans Kottelat 2000 Lao PDR 

Poropuntius lo boch eiloides Kottelat 2000 Lao PDR 

Poropuntius solitus Kottelat 2000 Lao PDR 

Pseudecheneis maurus Ng & Ta n 2007 Vietna m 

Pseudecheneis sympelvicus Roberts 1998 Lao PDR 

Pseudobagarius filifer Ng & Rainbo th 2005 Cambodia 

Pseudobagarius inermis Ng & Kottelat 2000 Lao PDR 

Pseudobagarius nitidus Ng & Rainbo th 2005 Lao PDR 

Pseudobagrus n ub ilosus Ng & Freyhof 2007 Vietna m 

Pseudolaguvia tenebricosa Britz & Ferra ris 2003 Mya nmar 

Pseudomystus sobrinus Ng & Freyhof 2005 Vietna m 

Pseudomystus bomboides Kottelat 2000 Lao PDR 

Psilorhynchus robustus Conway & Kottelat 2007 Mya nmar 

Pterocryptis buccata Ng & Kottelat 1998 Tha iland 

Pterocryptis inusitata Ng 1999 Lao PDR 

Pterocryptis crenula Ng & Freyhof 2001 Vietna m 

Pterocryptis verecu nda Ng & Freyhof 2001 Vietna m 

Puntius didi Kul l&er & Fang 2005 Mya nmar 

Puntius tiantian Kul l&er & Fang 2005 Mya nmar 

Puntius rhombeus Kottelat 2000 Lao PDR 

Rasbora amplistriga Kottelat 2000 Lao PDR 

Rasbora septentrionalis Kottelat 2000 Lao PDR 

Rhinogobius b oa Chen & Kottelat 2005 Vietna m 

Rhinogobius maculicervix Chen & Kottelat 2000 Lao PDR 

Rhinogobius milleri Chen & Kottelat 2003 Lao PDR 

Rhinogobius n ammaensis Chen & Kottelat 2003 Lao PDR 

Rhinogobius sulcatus Chen & Kottelat 2005 Vietna m 

Rhinogobius variolatus Chen & Kottelat 2005 Vietna m 

Rhinogobius vermiculatus Chen & Kottelat 2003 Lao PDR 

Rhinogobius virg igena Chen & Kottelat 2005 Vietna m 

Rhinogobius a lbimaculatus Chen & Kottelat & Miller 1999 Lao PDR 

Rhinogobius lineatus Chen & Kottelat & Miller 1999 Lao PDR 

Rhinogobius taenig ena Chen & Kottelat & Miller 1999 Lao PDR 

Rhodeus laoensis Kottelat, Doi & Musikasinthorn 1998 Lao PDR 

Scaphognathops theunensis Kottelat 1998 Lao PDR 

Schistura bannaensis Chen, Yan g & Qi 2005 Yunnan 

Schistura cryptofasciata Chen, Kon g & Yang 2005 Yunnan 

Schistura disparizona Zhou & Ko tte lat 2005 Yunnan

Schistura pridii Vidthayanon 2003 Thai land 

Schistura spekuli Kottelat 2004 Vietnam 

Schistura amplizona Kottelat 2000 Lao PDR 

Schistura antennata Freyhof & Serov 2001 Vietnam 

Schistura aramis Kottelat 2000 Lao PDR 

Schistura athos Kottelat 2000 Lao PDR 

Schistura atra Kottelat 1998 Lao PDR 

Schistura bachmaensis Freyhof & Serov 2001 Vietnam 

Schistura bai rdi Kottelat 2000 Lao PDR 

Schistura bol avenensis Kottelat 2000 Lao PDR 

Schistura carbonaria Freyhof & Serov 2001 Vietnam 

Schistura cataracta Kottelat 1998 Lao PDR 

Schistura clatrata Kottelat 2000 Lao PDR 

Schistura coruscans Kottelat 2000 Lao PDR 

Schistura crabro Kottelat 2000 Lao PDR 

Schistura dal atensis Freyhof & Serov 2001 Vietnam 

Schistura deansmarti Vidthayanon & Kottelat 2003 Thai land 

Schistura defectiva Kottelat 2000 Lao PDR 

Schistura dorsizona Kottelat 1998 Lao PDR 

Schistura ephelis Kottelat 2000 Lao PDR 

Schistura finis Kottelat 2000 Lao PDR 

Schistura fusinotata Kottelat 2000 Lao PDR 

Schistura globiceps Kottelat 2000 Lao PDR 

Schistura huongensis Freyhof & Serov 2001 Vietnam 

Schistura imitator Kottelat 2000 Lao PDR 

Schistura implicata Kottelat 2000 Lao PDR 

Schistura irregularis Kottelat 2000 Lao PDR 

Schistura isostigma Kottelat 1998 Lao PDR 

Schistura kaysonei Vidthayanon & Jaruthanin 2002 Lao PDR 

Schistura khamtanhi Kottelat 2000 Lao PDR 

Schistura kloetzliae Kottelat 2000 Lao PDR 

Schistura kongphengi Kottelat 1998 Lao PDR 

Schistura kontumensis Freyhof & Serov 2001 Vietnam 

Schistura latidens Kottelat 2000 Lao PDR 

Schistura leukensis Kottelat 2000 Lao PDR 

Schistura macrocephalus Kottelat 2000 Lao PDR 

Schistura melarancia Kottelat 2000 Lao PDR 

Schistura namboensis Freyhof & Serov 2001 Vietnam 

Schistura nomi Kottelat 2000 Lao PDR 

Schistura novemradiata Kottelat 2000 Lao PDR 

Schistura nudidorsum Kottelat 1998 Lao PDR 

Schistura obeini Kottelat 1998 Lao PDR 

Schistura personata Kottelat 2000 Lao PDR 

Schistura pertica Kottelat 2000 Lao PDR 

Schistura pervagata Kottelat 2000 Lao PDR 

Schistura porthos Kottelat 2000 Lao PDR 

Schistura procera Kottelat 2000 Lao PDR 

Schistura psittacul a Freyhof & Serov 2001 Vietnam 

Schistura punctifasciata Kottelat 1998 Lao PDR 

Schistura quaesita Kottelat 2000 Lao PDR 

Schistura quasimodo Kottelat 2000 Lao PDR 

Schistura rikiki Kottelat 2000 Lao PDR 

Schistura russa Kottelat 2000 Lao PDR 

Schistura sertata Kottelat 2000 Lao PDR 

Schistura sigi llata Kottelat 2000 Lao PDR 

Schistura sokolovi Freyhof & Serov 2001 Vietnam 

Schistura sombooni Kottelat 1998 Lao PDR 

Schistura speisi Vidthayanon & Kottelat 2003 Thai land 

Schistura suber Kottelat 2000 Lao PDR 

Schistura susannae Freyhof & Serov 2001 Vietnam 

Schistura tenura Kottelat 2000 Lao PDR 

Schistura thanho Freyhof & Serov 2001 Vietnam 

Schistura tizardi Kottelat 2000 Lao PDR 

Schistura tubulinaris Kottelat 1998 Lao PDR 

Schistura xhatensis Kottelat 2000 Lao PDR 

Schistura yersini Freyhof & Serov 2001 Vietnam 

Sectori a megastoma Kottelat 2000 Lao PDR 

Serpenticobitis cingulata Roberts 1997 Lao PDR / Thai land 

Serpenticobitis zonata Kottelat 1998 Lao PDR 

Sewelli a albisuera Freyhof 2003 Vietnam 

Sewelli a breviventralis Freyhof & Serov 2000 Vietnam 

Sewelli a patella Freyhof & Serov 2000 Vietnam 

Sineleotris namxamensis Chen & Kottelat 2004 Lao PDR / Vietnam 

Sinocyclocheilus guishanensis 

Sinocyclocheilus 

wumengshanensis 

Li 

Li, Mao & Lu, 

2003 

2003 

Yunnan 

Yunnan 

Sundasalanx mekongensis Britz & Kottelat 1999 Cambodia / Lao PDR / Thailand / 

Vietnam 

Tanichthys micagemmae Freyhof & Herder 2001 Vietnam 

Tetraodon abei 

Tonlesapia tsukawakii 

Tor ater 

Roberts 

Motomura & Mukai 

Roberts 

1998 

2006 

1999 

Lao PDR 

Cambodia 

Lao PDR 

Tor yingjiangensis Chen & Yang 2004 Yunnan 

Triplophysa laticeps Zhou & Cui, 1997 Yunnan 

Triplophysa nujiangensa Chen, Cui & Yang 2004 Yunnan 

Triplophysa xiangshuingensis Li, 2004 Yunnan 

Troglocyclocheilus 

khammouanensis 

Kottelat & Bréhi er 1999 Lao PDR 

Vanmanenia crassicauda 

Vanmanenia serrilineata 

Kottelat 

Kottelat 

2000 

2000 

Lao PDR 

Lao PDR 

Wallago micropogon Ng 2004 Vietnam / Lao PDR / Thailand 

Yunnanilus bajiangensis Li 2004 Yunnan 

Yunnanilus longibarbatus Gan, Chen & Yang 2007 Yunnan 

TOTAL NUMBER OF FISH 279 

MAMMALS 


Chodsi goa caovansunga 

Crocidura kegoensis 

Lunde, Musser & Son 

Lunde, Musser & Ziegler 

2003 

2004 

Vietnam 

Vietnam 

Hi pposideros khaokhouayensis Guillén-Servent & Francis 2006 Lao PDR 

Hi pposideros rotalis Francis, Kock & Habersetzer 1999 Lao PDR 

Hi pposideros scutinares Robinson et al 2003 Lao PDR / Vietnam 

Kerivoula kachinensis Bates, Struebig, Rossiter, Kingston, 

Tigga, Sai, Khin 

2004 Cambodia / Lao PDR / Myanmar 

/ Thailand 

Kerivoula titania Bates, Struebig, Hayes, Furey, Mya 2007 Cambodia/ Lao PDR / Myanmar 

Mya, et al 

/ Thailand / Vietnam 

Laonastes aenigmamus Jenkins et al 2005 Lao PDR 

Muntiacus putaoensis Rabinowitz, Than Myint, & Saw Tun 

Khaing 

1999 Myanmar 

Muntiacus truongsonensis Giao, Tuoc, Dung, Wikramanayake, 1998 Vietnam 

Amato, Arctander & MacKinnon 

Murina harrisoni 

Myotis annamiticus 

Csorba, Bates 

Kruskop & Tsytsulina 

2005 

2001 

Cambodia 

Vietnam 

Nesolagus timminsi Averianov, Abramov & Tikhinov 2000 Vietnam / Lao PDR 

Saxatilomys paulinae Musser, Smith, Robi nson & Lunde 2005 Lao PDR 

Tonkinomys daovantieni Musser, Lunde & Son 2006 Vietnam 

TOTAL NUMBER OF MAMMALS 15 

PLANTS 


Acer yangbi ense Chan & Yang 2003 Yunnan (China) 

Acronema brevipedicellatum Pan & Watson 2004 Yunnan (China) 

Actinodaphne menghaiensis Li 2005 Yunnan (China)

Aeschynanthus mendumi ae Middleton 2007 Lao PDR 

Aeschynanthus minutifolius Middleton 2007 Thai land 

Agrostis sinorupestri s Liu ex Phillips & Lu 2006 Yunnan (China) 

Ailanthus vietnamensis Sam & Noot 2007 Vietnam 

Ainsliaea fulvi oides Chuang 2004 Yunnan (China) 

Ainsliaea gongshanensis Chuang 2004 Yunnan (China) 

Ainsliaea lijiangensis Chuang 2004 Yunnan (China) 

Aleuritopteris gongshanensis Zhang 2004 Yunnan (China) 

Alocasia hypnosa Yin, Wang & Xu 2005 Yunnan (China) 

Alphonsea sonlaensi s Bân 2000 Vietnam 

Alseodaphne huanglianshanensis Li & Shui 2004 Yunnan (China) 

Alysicarpus bracteus Gao 2006 Yunnan (China) 

Amitostigma wenshanense Chen, Shui & Lang 2003 Yunnan (China) 

Amorphophal lus xiei Li & Dao 2006 Yunnan (China) 

Ancylostemon hekouensis Shui & Chen 2006 Yunnan (China) 

Anisochilus harmandii Doan ex Suddee & Paton 2004 Cambodia/Lao PDR/ Thailand 

Anoectochilus annamensi s Averyanov 2005 Vietnam 

Anoectochilus papillosus Averyanov 2007 Vietnam 

Aquilaria rugosa Le-Cong & Kessler 2005 Vietnam 

Argostemma fasciculata Sridith & Larsen 2004 Cambodia 

Arisaema averyanovii Nguyen & Boyce 2005 Vietnam 

Arisaema lidaense Murata & Wu 2003 Yunnan (China) 

Arisaema menghaiense Yin, Li & Xu 2004 Yunnan (China) 

Arisaema menglaense Ji, Li & Xu 2004 Yunnan (China) 

Arisaema muratae Gusman & Yin 2007 Yunnan (China) 

Arisaema rostratum Nguyen & Boyce 2005 Vietnam 

Aristolochia hansenii Phuphathanaphong 2006 Thai land 

Aristolochia kongkandae Phuphathanaphong 2006 Thai land 

Aristolochia perangustifolia Phuphathanaphong 2006 Thai land 

Aristolochia poomae Phuphathanaphong 2006 Thai land 

Aristolochia yalaensis Phuphathanaphong 2006 Thai land 

Artabotrys hienianus Bân 2000 Vietnam 

Artabotrys phuongianus Bân 2000 Vietnam 

Artabotrys taynguyenensis Bân 2000 Vietnam 

Artabotrys tetramerus Bân 2000 Vietnam 

Artabotrys vietnamensis Bân 2000 Vietnam 

Arthromeris cyrtomioides Lu & Xu 2007 Yunnan (China) 

Artocarpus Thailandicus Berg 2005 Thai land 

Arundina caespitosa Averyanov 2007 Vietnam 

Arundinella suniana Philli ps & Chen 2005 Yunnan (China) 

Aspidistra alata Tillich 2007 Vietnam 

Aspidistra arnautovii Tillich 2005 Vietnam 

Aspidistra atroviolacea Tillich 2005 Vietnam 

Aspidistra bicolor Tillich 2005 Vietnam 

Aspidistra bogneri Tillich 2005 Vietnam 

Aspidistra campanulata Tillich 2007 Vietnam 

Aspidistra carnosa Tillich 2005 Vietnam 

Aspidistra connata Tillich 2005 Vietnam 

Aspidistra cryptantha Tillich 2007 Vietnam 

Aspidistra foli osa Tillich 2005 Vietnam 

Aspidistra geastrum Tillich 2005 Vietnam 

Aspidistra grandifl ora Tillich 2007 Vietnam 

Aspidistra lateralis Tillich 2005 Vietnam 

Aspidistra locii Arnautov & Bogner 2004 Vietnam 

Aspidistra lutea Tillich 2005 Vietnam 

Aspidistra marasmioides Tillich 2005 Vietnam 

Aspidistra opaca Tillich 2005 Vietnam 

Aspidistra petiol ata Tillich 2005 Vietnam 

Aspidistra recondi ta Tillich 2007 Vietnam 

Aspidistra renatae Bräuchler 2005 Vietnam 

Aspidistra stricta Tillich 2005 Vietnam 

Aspidistra superba Tillich 2005 Vietnam 

Aspidistra tubiflora Tillich 2006 Vietnam 

Aspidistra umbrosa Tilli ch 2007 Vietnam 

Asplenium kukkonenii Viane & Reichstein 2003 Yunnan (China) 

Asplenium semivarians Viane & Reichstein 2003 Yunnan (China) 

Astragalus chengkangensis Podlech & Xu 2007 Yunnan (China) 

Astragalus gongshanensis Podlech & Xu 2007 Yunnan (China) 

Astragalus supralaevis Podlech & Xu 2007 Yunnan (China) 

Begoni a bataiensis Kiew 2005 Vietnam 

Begoni a cladotricha Hughes 2007 Lao PDR 

Begoni a crocea Peng 2006 Yunnan (China) 

Begoni a crystallina Shui & Chen 2005 Yunnan (China) 

Begoni a cucphuongensis Nguyen & Tebbitt 2005 Vietnam 

Begoni a glutinosa Kiew 2007 Vietnam 

Begoni a guani ana Ma & Li 2006 Yunnan (China) 

Begoni a hahiepiana Nguyen & Tebbitt 2006 Vietnam 

Begoni a hayamiana Tanaka 2007 Myanmar 

Begoni a huangii Shui & Chen 2005 Yunnan (China) 

Begoni a kingdon-wardii Tebbitt 2007 Yunnan (China) 

Begoni a kui Peng 2007 Vietnam 

Begoni a longistyla Shui & Chen 2005 Yunnan (China) 

Begoni a phamiana Kiew 2007 Cambodia / Vietnam 

Begoni a platycarpa Shui & Chen 2005 Yunnan (China) 

Begoni a poi lanei Kiew 2007 Vietnam 

Begoni a rhynchocarpa Shui & Chen 2005 Yunnan (China) 

Begoni a tetralobata Shui 2007 Yunnan (China) 

Berberis micropetala Ying 1999 Yunnan (China) 

Bistorta albiflora Miyamoto & Ohba 2002 Yunnan (China) 

Boesenbergia regalis Kharuk & Tohdam 2003 Thailand 

Brassaiopsis ficifolioides Wen & Lowry 2006 Vietnam 

Brassaiopsis gigantea Wen & Lowry 2006 Vietnam 

Brassaiopsis pseudoficifolia Lowry & C.B.Shang 2006 Yunnan (China) 

Bromheadia annamensis Averyanov & Averyanova 2006 Vietnam 

Bromus pseudoramosus Keng ex Li u 2002 Yunnan (China) 

Buddleja microstachya Liu & Peng 2006 Yunnan (China) 

Bulbophyllum dulongjiangense Jin 2006 Yunnan (China) 

Bulbophyllum gunnarii Averyanov 2005 Vietnam 

Bulbophyllum guttulatoides Averyanov 2005 Vietnam 

Bulbophyllum lockii Averyanov & Averyanova 2006 Vietnam 

Bulbophyllum paraemarginatum Averyanov 2007 Vietnam 

Bulbophyllum sinhoense Averyanov 2007 Vietnam 

Caesal pinia yunnanensis Li, Zhang & Chen 2006 Yunnan (China) 

Calamus banlingensis Yang, Yang & Lu 2007 Yunnan (China) 

Calamus evansii Hend 2007 Lao PDR 

Calamus hukaungensis Hend 2007 Myanmar 

Calamus mi nor Hend 2007 Lao PDR 

Calamus spicatus Hend 2007 Myanmar 

Calanthe dulongensis Li, Li & Dao 2003 Yunnan (China) 

Calanthe duyana Averyanov 2006 Vietnam 

Calocedrus rupestris Averyanov, Nguyên & Lôc 2004 Vietnam 

Camell ia hamyenensis Tran & Le 2005 Vietnam 

Camell ia rubriflora Tran & Hakoda 1998 Vietnam 

Campylandra siamensis Yamashita & Tamura 2001 Thailand 

Carex bijiangensis Liang & Zhang 2006 Yunnan (China) 

Carex globulosa Phulphong & Si mpson 2007 Thailand 

Carex obliquitruncata Tang & Liang 2006 Yunnan (China) 

Castanopsis pseudohystrix Phengklai 2004 Thailand 

Castanopsis thaiensis Phengklai 2004 Thailand 

Caulokaempferia amplexicaulis Suksathan 2005 Thailand 

Caulokaempferia appendiculata Larsen & Triboun 2003 Thailand 

Caulokaempferia bracteata Larsen & Larsen 2003 Thailand 

Caulokaempferia jirawongsei Picheans & Mokkamul 2004 Thailand 

Caulokaempferia khaomaenensis Picheans & Mokkamul 2004 Thailand 

Caulokaempferia laoti ca Picheans & Mokkamul 2006 Lao PDR 

Caulokaempferia limiana Mokkamul & Picheans 2004 Thailand

Caulokaempferia pedemontana Tri boun & K.Larsen 2005 Thai land 

Caulokaempferia phuluangensis Picheans & Mokkamul 2004 Thai land 

Caulokaempferia violacea Larsen & Triboun 2003 Thai land 

Cephalostachyum scandens Hsueh & Hui 1997 Yunnan (China) 

Cheirostylis calcarata Jin & Chen 2007 Yunnan (China) 

Cheirostylis foliosa Averyanov 2007 Vietnam 

Cheirostylis latipetala Averyanov & Averyanova 2006 Vietnam 

Cheirostylis malipoensis Jin & Chen 2007 Yunnan (China) 

Cheirostylis serpens Averyanov 2005 Vietnam 

Chiloschista rodriguezii Cavestro & Ormerod 2005 Thai land 

Cinnamomum bhamoensi s Gangopadhyay 2006 Myanmar 

Cinnamomum blandfordii Gangopadhyay 2006 Myanmar 

Cladrasti s chingii Duley & Vincent 2003 Yunnan (China) 

Cleisostoma melanorachi s Averyanov & Averyanova 2006 Vietnam 

Cleisostoma subulifolium Averyanov & Averyanova 2006 Vietnam 

Clematis hagiangensis Do 2006 Vietnam 

Clematis pingbianensis Wang 2003 Yunnan (China) 

Clematis vietnamensis Wang & Do 2006 Vietnam 

Coelogyne tsii Jin & Li 2006 Yunnan (China) 

Coelogyne weixiensis Jin 2005 Yunnan (China) 

Colocasia bicolor Long & Cao 2003 Yunnan (China) 

Colocasia menglaensis Yin, Li & Xu 2004 Yunnan (China) 

Colocasia yunnanensis Long & Cai 2006 Yunnan (China) 

Conioselinum reflexum Pimenov & Kljuykov 2003 Yunnan (China) 

Cordisepalum phalanthopetalum Staples 2006 Myanmar / Thailand 

Cornus eydeana Xiang & Shui 2003 Yunnan (China) 

Corydalis heterothylax Wu ex Su & Li dén 2007 Yunnan (China) 

Corydalis ischnosiphon Lidén & Su 2007 Yunnan (China) 

Corydalis mediterranea Su & Li dén 2007 Yunnan (China) 

Crepidium klimkoanum Margońska 2005 Thai land 

Cryptocarya biswasii Gangopadhyay 2006 Yunnan (China) 

Cucumis debilis De Wilde & Duyfjes 2007 Vietnam 

Curcuma anti naia Chaveer & Tanee 2007 Thai land 

Curcuma larsenii Maknoi & Jenji tt 2006 Thai land/Lao PDR/ Vietnam 

Cymbi dium aestivum Liu & Chen 2004 Yunnan (China) 

Cymbi dium changningense Liu & Chen 2005 Yunnan (China) 

Cymbi dium concinnum Liu & Chen 2006 Yunnan (China) 

Cymbi dium gaoligongense Liu & Zhang 2003 Yunnan (China) 

Cymbi dium lushuiense Liu, Chen & Shi 2005 Yunnan (China) 

Cymbi dium micranthum Liu & Chen 2004 Yunnan (China) 

Cymbi dium multiradi catum Liu & Chen 2004 Yunnan (China) 

Cymbi dium quinquel obum Liu & Chen 2006 Yunnan (China) 

Cymbopogon minor Sun & Zhang ex Philli ps & Chen 2005 Yunnan (China) 

Cyperus simaoensis Qi an 2001 Yunnan (China) 

Cypripedium malipoense Chen & Liu 2004 Yunnan (China) 

Dalzellia angustissima Kato 2006 Thai land 

Dalzellia kailarsenii Kato 2006 Thai land 

Dalzellia ranongensis Kato 2006 Thai land 

Dalzellia ubonensis Kato 2006 Thai land 

Dasymaschalon tueanum Bân 2000 Vietnam 

Dendrobium bifurcatum Yukawa 2003 Vietnam 

Dendrobium chapaense Averyanov 2006 Vietnam 

Dendrobium farinatum Schildhauer & Schraut 2004 Vietnam 

Dendrobium menglaensis Jin & Li 2006 Yunnan (China) 

Dendrobium schrauti i Schildhauer 2006 Vietnam 

Dendrobium trankimi anum Yukawa 2004 Vietnam 

Dendrobium trantuanii Perner & Dang 2003 Vietnam 

Dendrobium tuananhii Averyanov 2004 Vietnam 

Dendrobium vietnamense Averyanov 2005 Vietnam 

Dendrocalamus khoonmengii Sungkaew, Teerawat. & Hodkinson 2007 Thai land 

Diospyros phuketensis Phengklai 2005 Thai land 

Diospyros ranongensis Phengklai 2005 Thai land 

Diphasiastrum wilceae Ivanenko 2003 Yunnan (China) 

Di schidia cornuta Livsh 2005 Lao PDR 

Di schidia dohtii Tran & Livsh 2005 Lao PDR/Vietnam 

Di stichochlamys rubrostriata Kress & Rehse 2003 Vietnam 

Dracaena impressivenia Yan & Guo 2006 Yunnan (China) 

Eclipta angustata Umemoto & Koyama 2007 Thailand / Vietnam / Yunnan 

(China) 

Elatostema attenuatoi des Wang 2006 Yunnan (China) 

Elatostema binerve Wang 2006 Yunnan (China) 

Elatostema jianshanicum Wang 2003 Yunnan (China) 

Elatostema maguanense Wang 2006 Yunnan (China) 

Elatostema melanophyllum Wang 2006 Yunnan (China) 

Elatostema septemflorum Wang 2006 Yunnan (China) 

Elatostema shuii Wang 2003 Yunnan (China) 

Elatostema trichotomum Wang 2003 Yunnan (China) 

Elatostema wangii Lin & Duan 2003 Yunnan (China) 

Enicosanthum daclacense Bân 2000 Vietnam 

Epigeneium forrestii Ormerod 2007 Yunnan (China) 

Epigeneium gaol igongense Yu & Zhang 2005 Yunnan (China) 

Epipactis alatus Averyanov & Efimov 2006 Vietnam 

Eria chlorantha Averyanov & Averyanova 2006 Vietnam 

Eurya j intungensis Li 2000 Yunnan (China) 

Eurya l uchunensis Wang & Wang 2005 Yunnan (China) 

Eurya subcordata Li 2000 Yunnan (China) 

Eurya taronensis Li 2000 Yunnan (China) 

Fagopyrum capillatum Ohnishi 1998 Yunnan (China) 

Festuca scabriflora Liu 2002 Yunnan (China) 

Ficus Thailandica Berg & Gardner 2007 Thailand 

Fissistigma taynguyenense Bân 2000 Vietnam 

Galium shanense Bhattacharjee 2006 Myanmar 

Gastrochilus alatus Jin & Chen 2007 Yunnan (China) 

Gastrochilus mal ipoensis Jin & Chen 2007 Yunnan (China) 

Gastrodia major Averyanov 2006 Vietnam 

Gastrodia punctata Averyanov 2006 Vietnam 

Gastrodia theana Averyanov 2005 Vietnam 

Gastrodia tonkinensis Averyanov & Averyanova 2006 Vietnam 

Gentiana khammouanensi s Hul 2007 Lao PDR 

Glyceri a ovatiflora Keng ex Tzvelev 2006 Yunnan (China) 

Glyptopetalum reticulinerve Wu ex Fan & Xu 2007 Yunnan (China) 

Gomphogyne cirromitrata De Wilde & Duyfjes 2007 Thailand 

Gomphogyne heterosperma De Wilde & Duyfjes 2007 Thailand 

Gomphostemma grandiflorum Doan ex Suddee & Paton 2007 Vietnam 

Goniothalamus albi florus Bân 2000 Vietnam 

Goniothalamus gracilipes Bân 2000 Vietnam 

Goniothalamus lii Hou & Shui 2003 Yunnan (China) 

Goniothalamus macrocalyx Bân 2000 Vietnam 

Goniothalamus ninhianus Bân 2000 Vietnam 

Goniothalamus takhtajanii Bân 2000 Vietnam 

Goniothalamus vietnamensis Bân 2000 Vietnam 

Goodyera myanmarica Ormerod & Kumar 2006 Yunnan (China) 

Goodyera rhombodoides Averyanov 2007 Vietnam 

Gynostemma pentaphyl lum De Wilde & Duyfjes 2007 Thailand 

Habenaria harderi Averyanov & Averyanova 2006 Vietnam 

Habenaria tuanae Rice 2007 Yunnan (China) 

Hamularia puluongensis Averyanov & Averyanova 2006 Vietnam 

Hanseniella smitinandii Kato 2004 Thailand 

Hedychium khaomaenense Picheans & Mokkamul 2005 Thailand 

Hedychium thaianum Mokkamul & Picheans 2005 Thailand 

Hemipi lia discolor Averyanov & Averyanova 2006 Vietnam 

Hemsleya kunmingensis Li & Li 2007 Yunnan (China) 

Hetaeria youngsayei Ormerod 2004 Thailand 

Heterostemma lobulatum Li & Konta 2002 Yunnan (China) 

Holcoglossum auriculatum Liu, Chen & Jin 2005 Yunnan (China) 

Holcoglossum weixiense Jin & Chen 2003 Yunnan (China)

Hoya bal aensis Kidyoo & Thaithong 2007 Thailand 

Hoya weebella Kloppenburg 2005 Thailand 

Huperzia muscicola Chu 2006 Yunnan (China) 

Hydrobryum chiangmaiense Kato 2004 Thailand 

Hydrobryum kaengsophense Kato 2004 Thailand 

Hydrobryum khaoyaiense Kato 2004 Thailand 

Hydrobryum l oeicum Kato 2004 Thailand 

Hydrobryum somranii Kato 2004 Thailand 

Hydrobryum tardhuangense Kato 2004 Thailand 

Hypericum fosteri Robson 2005 Yunnan (China) 

Hypericum wardianum Robson 2005 Myanmar / Yunnan (China) 

Iguanura polymorpha Lim 1998 Thailand 

Impatiens armeniaca Huang 2003 Yunnan (China) 

Impatiens austroyunnanensis Huang 2003 Yunnan (China) 

Impatiens daguanensis Huang 2003 Yunnan (China) 

Impatiens deqinensis Huang 2003 Yunnan (China) 

Impatiens lancisepala Huang 2003 Yunnan (China) 

Impatiens laojunshanensis Huang 2003 Yunnan (China) 

Impatiens latipetala Huang 2003 Yunnan (China) 

Impatiens longirostris Huang 2003 Yunnan (China) 

Impatiens malipoensi s Huang 2003 Yunnan (China) 

Impatiens multiramea Huang 2003 Yunnan (China) 

Impatiens phahompokensis Shimi zu & Suksathan 2004 Thailand 

Impatiens phengkl aii Shimi zu & Suksathan 2004 Thailand 

Impatiens pianmaensis Huang 2003 Yunnan (China) 

Impatiens salwinensi s Huang 2003 Yunnan (China) 

Impatiens suij iangensis Huang 2003 Yunnan (China) 

Impatiens sunii Huang 2003 Yunnan (China) 

Impatiens tribounii Shimi zu & Suksathan 2004 Thailand 

Impatiens wenshanensis Huang 2003 Yunnan (China) 

Impatiens xishuangbannaensis Huang 2003 Yunnan (China) 

Impatiens yongshanensis Huang 2003 Yunnan (China) 

Impatiens yui Huang 2003 Yunnan (China) 

Impatiens zixishanensis Huang 2003 Yunnan (China) 

Isodon colaniae Hara ex Suddee & Paton 2004 Lao PDR 

Ixora phuluangensis Chamch 2005 Thailand 

Jasminum kaulbackii Green 2003 Myanmar 

Juncus fugongensis Bao 2003 Yunnan (China) 

Juncus jinpingensis Bao 2003 Yunnan (China) 

Juncus lancangensis Qian 2001 Yunnan (China) 

Juncus megalophyllus Bao 2003 Yunnan (China) 

Juncus yui Bao 2003 Yunnan (China) 

Kamettia chandeei Middleton 2005 Thailand 

Kandelia obovata Sheue, Liu & Yong 2003 Vietnam 

Kedarnatha vaginata Pimenov & Kljuykov 2004 Myanmar 

Khaosokia caricoides Simpson, Chayam & Parn 2005 Thailand 

Khmeriosicyos harmandii De Wilde & Duyfjes 2004 Cambodia 

Kopsia vidalii Middleton 2005 Vietnam 

Lecanorchis vietnami ca Averyanov 2005 Vietnam 

Lespedeza lichiyuniae Nemoto, Ohashi & Itoh 2007 Yunnan (China) 

Ligularia pi anmaensis Chen ex Ming 2004 Yunnan (China) 

Lindenbergia luchunensis Tao & Shui 2006 Yunnan (China) 

Liparis filiformis Averyanov 2005 Vietnam 

Liparis petraea Averyanov & Averyanova 2006 Vietnam 

Liparis ri vularis Averyanov 2007 Vietnam 

Liparis rockii Ormerod 2007 Yunnan (China) 

Liparis superposita Ormerod 2007 Yunnan (China) 

Liparis tripartita Averyanov & Averyanova 2006 Vietnam 

Liparis yuana Ormerod 2007 Yunnan (China) 

Listera fugongensis Jin 2007 Yunnan (China) 

Lithocarpus loratifolius Phengklai 2004 Thailand 

Litsea cangyuanensis Li & Li 2006 Yunnan (China) 

Litsea phuwuaensis Ngernsaengsaruay 2004 Thailand 

Lycianthes baviensis Hop 2004 Vietnam 

Lysimachia gesnerioides Shui & Zhang 2006 Vietnam / Yunnan (China) 

Magnolia amabilis Sima & Wang 2006 Yunnan (Chi na) 

Magnolia carpunii Romanov & Bobrov 2003 Yunnan (Chi na) 

Magnolia citrata Nooteboom & Chalermglin 2007 Thailand 

Magnolia glaucophyl la Sima & Yu 2007 Yunnan (Chi na) 

Mall otus gl omerulatus Welzen 2004 Thailand 

Mall otus mirus Sierra 2007 Thailand 

Manglietia hongheensis Shui & Chen 2003 Yunnan (Chi na) 

Manglietia rufisyncarpa Law, Zhou & Wang 2007 Yunnan (Chi na) 

Melanosciadium genufl exum Pimenov & Kljuykov 2006 Yunnan (Chi na) 

Melodorum kontumense Bân 2000 Vietnam 

Metathelypteris deltoideofrons Ching ex Chu & Lu 2006 Yunnan (Chi na) 

Michelia multi tepal a Zhou & Jian 2007 Yunnan (Chi na) 

Microsorum siamensis Boonkerd 2006 Thailand 

Miliusa bannaensis Hou 2004 Yunnan (Chi na) 

Mimulicalyx paludigenus Tsoong ex Li & Cai 2005 Yunnan (Chi na) 

Miscanthus lutarioriparius Liu ex Chen & Renvoi ze 2006 Yunnan (Chi na) 

Mitrella touranensis Bân 2000 Vietnam 

Mitrephora sirikitiae Weeras, Chalermgli n & Saunders 2006 Thailand 

Mosla bracteata Doan ex Suddee & Paton 2007 Vietnam 

Musa exotica Valmayor 2001 Vietnam 

Musa lutea Valmayor, Danh & Häkkinen 2004 Vietnam 

Musa paracoccinea Liu & D.Z.Li 2002 Yunnan (Chi na) 

Musa tonkinensis Valmayor, Danh & Häkkinen 2005 Vietnam 

Musa viridi s Valmayor, Danh & Häkkinen 2004 Vietnam 

Musa yunnanensis Häkkinen & Wang 2007 Yunnan (Chi na) 

Musella splendida Valmayor & Danh 2004 Vietnam 

Neohouzeaua fimbriata Dransfield, Pattanavibool & 

Sungkaew 

2003 Myanmar 

Nyssa yunnanensis Yin ex Qin & Phengklai 2007 Yunnan (Chi na) 

Oberonia huensis Averyanov 2005 Vietnam 

Oberonia multidentata Averyanov 2007 Vietnam 

Oberonia trichophora Averyanov 2007 Vietnam 

Ophiorrhiza larseniorum Schanzer 2005 Thailand 

Ophiorrhiza pseudofasciculata Schanzer 2005 Thailand 

Orchidantha foetida Jenjittikul & Larsen 2003 Thailand 

Orchipedum echinatum Averyanov & Averyanova 2006 Vietnam 

Orthosiphon lanatus Doan ex Suddee & Paton 2005 Vietnam 

Orthosiphon pseudoaristatus Suddee 2005 Thailand 

Orthosiphon rotundifolius Doan ex Suddee & Paton 2005 Vietnam 

Orthosiphon truncatus Doan ex Suddee & Paton 2005 Vietnam 

Panisea vinhii Averyanov & Averyanova 2006 Vietnam 

Paphiopedilum × cribbii Averyanov 2006 Vietnam 

Paphiopedilum × yingjiangense Liu & Chen 2007 Yunnan (Chi na) 

Paphiopedilum hangianum Perner & Gruss 1999 Vietnam 

Paphiopedilum smaragdinum Liu & Chen 2003 Yunnan (Chi na) 

Paphiopedilum thaianum Iamwiriyakul 2006 Thailand 

Paracladopus chiangmaiensis Kato 2006 Thailand 

Paris caobangensi s Ji, Li & Zhou 2006 Vietnam 

Passiflora xishuangbannaensis Krosnick 2005 Yunnan (Chi na) 

Pedicularis yaoshanensis Wang 2006 Yunnan (Chi na) 

Peliosanthes divaricatanthera Tanaka 2004 Yunnan (Chi na) 

Peliosanthes pachystachya Chen & Shui 2003 Yunnan (Chi na) 

Pennil abium yunnanense 

Persea bhaskarii 

Chen & Luo 

Gangopadhyay 

2004 

2006 

Yunnan (Chi na) 

Myanmar 

Persea russellii Gangopadhyay 2006 Myanmar 

Persea sharmae Gangopadhyay 2006 Myanmar 

Phaeanthus vietnamensis Bân 2000 Vietnam 

Phalaenopsis malipoensis Liu & Chen 2005 Yunnan (Chi na) 

Phoebe lummaoensis Gangopadhyay 2006 Myanmar 

Phoebe prazeri Gangopadhyay 2006 Myanmar 

Phreatia al bofarinosa Ormerod 2005 Thailand

Phreatia palmifrons Ormerod 2005 Thai land 

Phyllanthus pseudoparvifolius Mitra & Sanjappa 2003 Myanmar 

Physospermopsis siamensis Esser & Watson 2006 Thai land 

Pinus eremitana Businsky 2004 Vietnam 

Piper betloides Chaveerach & Tanomtong 2007 Thai land 

Piper dominantinervium Chaveerach & Mokkamul 2006 Thai land 

Piper phuwuaense Chaveerach & Tanee 2006 Thai land 

Piper pilobracteatum Chaveerach & Sudmoon 2006 Thai land 

Platanthera epiphytica Averyanov & Efimov 2006 Vietnam 

Platostoma becquerelii Suddee & Paton 2005 Cambodia 

Platostoma cambodgense Suddee & Paton 2005 Cambodia / Thailand/ Vietnam 

Platostoma grandiflorum Suddee & Paton 2005 Cambodia/ Thailand 

Platostoma kerrii Suddee & Paton 2005 Cambodia/ Thailand 

Platostoma mekongense Suddee 2005 Thai land 

Platostoma rubrum Suddee & Paton 2005 Lao PDR/Thailand 

Platostoma taylori i Suddee & Paton 2005 Cambodia 

Plectranthus albicalyx Suddee 2004 Thai land 

Plectranthus gigantifolius Suddee 2004 Thai land 

Plectranthus tomentifol ius Suddee 2004 Thai land 

Poa eragrosti oides Liu 2002 Yunnan (China) 

Poa sunbisini i Soreng & Zhu 2006 Yunnan (China) 

Poa zhongdianensis Liu 2002 Yunnan (China) 

Podocarpium lancangense Qi an 2003 Yunnan (China) 

Pogostemon globulosus Phuong ex Suddee & Paton 2007 Vietnam 

Pogostemon litigiosus Doan ex Suddee & Paton 2007 Vietnam 

Pogostemon nelsonii Doan ex Suddee & Paton 2007 Vietnam 

Polyalthi a barenensis Bân 2000 Vietnam 

Polyalthi a bracteosa Bân 2000 Vietnam 

Polyalthi a kanchanaburiana Khumchompoo & Thongpukdee 2005 Thai land 

Polyalthi a praeflorens Bân 2000 Vietnam 

Polypleurum erectum Kato 2006 Thai land 

Polypleurum longi caule Kato 2006 Thai land 

Polypleurum longi foli um Kato 2006 Thai land 

Polypleurum longi stylosum Kato 2006 Thai land 

Polypleurum phuwuaense Kato 2006 Thai land 

Polypleurum rubroradicans Kato 2006 Thai land 

Polypleurum wongprasertii Kato 2006 Thai land 

Potentilla assimilis Soják 2006 Yunnan (China) 

Potentilla millefoliolata Soják 2006 Yunnan (China) 

Pouzolzia Thailandica Fri is & Wilmot-Dear 2006 Thai land 

Primul a calyptrata Gong & Fang 2003 Yunnan (China) 

Pterospermum mengii Wilki e 2007 Cambodia / Vietnam 

Pueraria xyzhui Ohashi & Iokawa 2006 Yunnan (China) 

Ranunculus ailaoshanicus Wang 2007 Yunnan (China) 

Renanthera sinica Liu & Chen 2003 Yunnan (China) 

Rhododendron longil obum Gao & Li 2003 Yunnan (China) 

Rhododendron truncatovarium Gao & Li 2005 Yunnan (China) 

Rhododendron yaoshanense Gao & Zhang 2007 Yunnan (China) 

Roscoea cangshanensis Luo, Gao & Lin 2007 Yunnan (China) 

Rothmanni a daweishanensis Shui & Chen 2003 Yunnan (China) 

Saccolabiopsis viridiflora Averyanov 2005 Vietnam 

Saxifraga xiaozhongdianensis Pan 2007 Yunnan (China) 

Sciaphila arcuata Averyanov 2007 Vietnam 

Sciaphila stell ata Averyanov 2007 Vietnam 

Scutellaria attenui foli a Suddee & Paton 2006 Cambodia 

Selaginella hengduanshanicola Chu 2006 Yunnan (China) 

Selaginella rubella Chu 2006 Yunnan (China) 

Selaginella xichouensis Chu 2006 Yunnan (China) 

Shangrilaia nana Al-Shehbaz, Yue & Sun 2004 Yunnan (China) 

Shuteria lancangensis Qi an 2003 Yunnan (China) 

Sinobaiji ania smitinandii De Wilde & Duyfjes 2006 Thai land 

Sirindhornia mirabilis Pedersen & Suksathan 2003 Thai land 

Sirindhornia pulchella Pedersen & Indhamusika 2003 Thai land 

Solms-laubachia zhongdianensis Yue, Al-Shehbaz & Sun 2005 Yunnan (China) 

Sorbus amoena McAllister 2005 Yunnan (China) 

Sorbus apiculata McAllister 2005 Yunnan (China) 

Sorbus bulleyana McAllister 2005 Yunnan (China) 

Sorbus carmesina McAllister 2005 Yunnan (China) 

Sorbus coxii McAllister 2005 Yunnan (China) 

Sorbus ellipsoidalis McAllister 2005 Yunnan (China) 

Sorbus fansipanensis McAll 2005 Vietnam 

Sorbus glabriuscula McAllister 2005 Yunnan (China) 

Sorbus pseudohupehensis McAllister 2005 Yunnan (China) 

Sorbus pseudovi lmorinii McAllister 2005 Yunnan (China) 

Sorbus rubescens McAllister 2005 Yunnan (China) 

Spatholirion puluongense Averyanov 2006 Vietnam 

Stahlianthus pedicellatus Chaveer & Mokkamul 2007 Thailand 

Strobilanthes abbreviata Deng & Wood 2006 Cambodia/ Myanmar / Thailand / 


Strobilanthes euantha Wood 2003 Myanmar / Yunnan (China) 

Strobilanthes lihengiae Deng & Wood 2006 Yunnan (China) 

Strobilanthes vallicola Deng & Wood 2006 Yunnan (China) 

Sunipia nigricans Averyanov 2007 Vietnam 

Syzygium bubengense Chen 2006 Yunnan (China) 

Taxus florinii Spj ut 2007 Yunnan (China) 

Tectaria luchunensis Wu 2002 Yunnan (China) 

Terniopsis brevis Kato 2006 Thailand 

Terniopsis ramosa Kato 2006 Thailand 

Terniopsis ubonensis Kato 2006 Thailand 

Teucrium petelotii Doan ex Suddee & Paton 2007 Vietnam 

Thawatchaia trilobata Kato, Koi & Kita 2004 Thailand 

Thepparatia Thailandica Phuphathanaphong 2006 Thailand 

Thladiantha angustisepala De Wilde & Duyfjes 2006 Lao PDR / Thai land 

Thrixspermum hiepii Averyanov & Averyanova 2006 Vietnam 

Thrixspermum stelidioides Averyanov & Averyanova 2006 Vietnam 

Thrixspermum tsii Chen & Shui 2005 Yunnan (China) 

Tirpitzia bilocularis Suksathan & Larsen 2006 Thailand 

Tolypanthus pustul atus Barlow 2005 Thailand 

Trachycarpus geminisectus Spanner, Gibbons, Nguyen, Anh 2003 Vietnam 

Trevesia vietnamensis Wen & Lôc 2007 Vietnam 

Trias nummularia Averyanov & Averyanova 2006 Vietnam 

Trichosanthes dolichosperma Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes erosa Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes inthanonensis Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes kostermansii Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes pallida Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes phonsenae Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes pubera Duyfjes & Pruesapan 2004 Thailand 

Trichosanthes si amensis Duyfjes & Pruesapan 2004 Thailand 

Typhonium baoshanense Dao & Li 2007 Yunnan (China) 

Utricularia jackii Parnell 2005 Thailand 

Ventilago zhengdei Fan 2004 Yunnan (China) 

Vernonia chiangdaoensis Koyama 2005 Thailand 

Vernonia doichangensis Koyama 2004 Thailand 

Vernonia kradungensis Koyama 2005 Thailand 

Vernonia namnaoensis Koyama 2004 Thailand 

Vernonia ngaoensis Koyama 2004 Thailand 

Vernonia pseudobirmanica Koyama 2003 Thailand 

Vernonia pseudosutepensis Koyama 2005 Thailand 

Viola binchuanensi s Huang 2003 Yunnan (China) 

Viola dimorphophylla Chen & Yang 2005 Yunnan (China) 

Viola jizushanensis Huang 2003 Yunnan (China) 

Viola subdelavayi Huang 2003 Yunnan (China) 

Viscum macrofalcatum Han & Zhang 2006 Yunnan (China) 

Wallichia lidiae Hend 2007 Myanmar 

Wallichia marianniae Hodel 1997 Thailand

Wikstroemia fuminensis Qi & Wang 2004 Yunnan (China) 

Wrightia siamensis Middleton 2007 Thailand 

Xyloselinum leonidii Pimenov & Kljuykov 2006 Vietnam 

Xyloselinum vietnamense Pimenov & Kljuykov 2006 Vietnam 

Ypsilandra jinpingensis Chen, Shui & Yu 2003 Yunnan (China) 

Yushania shangrilaensis Demoly 2006 Yunnan (China) 

Yushania velutina Demoly 2006 Yunnan (China) 

Zehneria brevirostris De Wilde & Duyfjes 2004 Thailand 

Zehneria hermaphrodita De Wilde & Duyfjes 2004 Thailand 

Zehneria sphaerosperma De Wilde & Duyfjes 2004 Thailand 

Zehneria tenuispica De Wilde & Duyfjes 2004 Thailand 

Zeuxine bidupensis Averyanov 2006 Vietnam 

Zingiber kelantanense Lim 2003 Thailand 

Zingiber phumiangense Chaveer & Mokkamul 2007 Thailand 

Zingiber raja Lim & Kharuk 2003 Thailand 

TOTAL NUMBER OF PLANTS 519 

REPTILES 

LIZARDS 


Acanthosaura nataliae Orlov, Nguyen & Nguyen 2006 Vietnam 

Bronchocela orlovi Haller mann 2004 Vietnam 

Bronchocela vietnamensis Haller mann & Orlov 2005 Vietnam 

Calotes htunwini Zug & Vindum 2006 Myanmar 

Calotes irawadi Zug, Brown, Schulte, & Vindum 2006 Myanmar 

Cnemaspis aurantiacopes Gri smer & Tri 2007 Vietnam 

Cnemaspis caudanivea Gri smer & Tri 2007 Vietnam 

Cnemaspis chanthaburiensis Bauer & Das 1998 Cambodia 

Cnemaspis nuicamensis Gri smer & Tri 2007 Vietnam 

Cnemaspis tucdupensis Gri smer & Tri 2007 Vietnam 

Cyrtodactylus aequalis Bauer 2003 Myanmar 

Cyrtodactylus annandalei Bauer 2003 Myanmar 

Cyrtodactylus ayeyarwadyensis Bauer 2003 Myanmar 

Cyrtodactylus badenensis N. Ngoc et al., 2006 Vietnam 

Cyrtodactylus brevidactylus Bauer 2002 Myanmar 

Cyrtodactylus chanhomeae Bauer, Sumontha, Pauwels 2003 Thailand 

Cyrtodactylus chrysopylos Bauer 2003 Myanmar 

Cyrtodactylus cryptus Heidrich, Rösler, Thanh, Böhme & 

Ziegler 

2007 Vietnam 

Cyrtodactylus gansi Bauer 2003 Myanmar 

Cyrtodactylus nigriocularis N. Ngoc et al., 2006 Vietnam 

Cyrtodactylus paradoxus Darevsky & Szczerbak 1997 Vietnam 

Cyrtodactylus 

Ziegler, Rösler, Herrmann, 

2002 Vietnam 

phongnhakebangensis 

& Vu, 

Cyrtodactylus russell i Bauer 2003 Myanmar 

Cyrtodactylus slowinskii Bauer 2002 Myanmar 

Cyrtodactylus sumonthai Bauer, Pauwels & Chanhome 2002 Thailand 

Cyrtodactylus thirakhupti Pauwels, Bauer, Sumontha, 

Chanhome 

2004 Thailand 

Cyrtodactylus tigroides Bauer, Sumontha, Pauwels 2003 Thailand 

Cyrtodactylus wakeorum Bauer 2003 Myanmar 

Dibamus deharvengi Ineich 1999 Vietnam 

Dibamus kondaoensis Honda, Ota, Hikida & Darevsky 2001 Vietnam 

Dixonius hangseesom Bauer, Sumontha, Grossman, 

Pauwels & Vogel 

2004 Thailand 

Dixonius vi etnamensis Das 2004 Vietnam 

Gekko scientiadventura Rösler, Zi egler, Vu, Herrmann, & 

Böhme 

2005 Vietnam 

Goniurosaurus araneus Gri smer, Viets, & Boyle 1999 Vietnam 

Hemidactylus aquilonius McMahan & Zug 2007 Myanmar 

Hemidactylus thayene Zug & McMahan 2007 Myanmar 

Leptoseps tetradactylus Darevsky & Orlov 2005 Vietnam 

Lygosoma boehmei Ziegler, Schmi tz, Heidrich, Vu & 

Nguyen 

2007 Vietnam 

Paralipinia rara Darevsky & Orlov 1997 Vietnam 

Ptyctolaemus collicristatus Schulte & Vi ndum 2003 Myanmar 

Sphenomorphus cryptotis Darevsky, Orlov & Ho 2004 Vietnam 

Sphenomorphus devorator Darevsky, Orlov & Ho 2004 Vietnam 

Takydromus hani Chou, Nguyen & Pauwels 2001 Vietnam 

Tropidophorus matsuii Hikida, Orlov, Nabhitabhata & Ota 2002 Vietnam 

Tropidophorus murphyi Hikida, Orlov, Nabhitabhata & Ota 2002 Vietnam 

Tropidophorus noggei Ziegler, Vu & Bui 2005 Vietnam 

Subtotal 46 

SNAKES 

Amphiesma andreae Ziegler & Le 2006 Vietnam 

Amphiesma leucomystax David, Bai n, Nguyen, Orlov, Vogel, 

Vu & Ziegler 

2007 Vietnam 

Boiga bourreti Tillack, Ziegler & Le 2004 Vietnam 

Bungarus slowinskii Kuch, Kizirian, Nguyen, Lawson, 

Donnelly, & Mebs 

2005 Vietnam 

Calamaria thanhi Ziegler & Le 2005 Vietnam 

Dendrelaphis kopsteini Vogel & Van Rooijen 2007 Thailand 

Enhydris chanardi Murphy & Voris 2005 Thailand 

Enhydris vorisi John C. Mur phy 2007 Myanmar 

Lycodon cardamomensis Daltry & Wüster 2002 Cambodia 

Lycodon zawi Slowinski, Pawar, Win, Thin, Gyi, Oo 

& Tun 

2001 Myanmar 

Naja mandalayensis Slowi nski & Wüster 2000 Myanmar 

Oligodon jintakunei Pauwels, Wallach, David & 

Chanhome 

2002 Thailand 

Opisthotropis daovantieni Orlov, Darevsky, Murphy 1998 Vietnam 

Opisthotropis maculosus Stuart & Chuaynkern 2007 Thailand 

Sinonatrix yunnanensis Rao & Yang 1998 Myanmar 

Triceratolepidophis sieversorum Ziegler, Herrmann, David, 

Orlov & Pauwels 

2000 Vietnam / Lao PDR 

Trimeresurus fucatus Vogel, David & Pauwels 2004 Thailand / Myanmar 

Trimeresurus gumprechti David, Vogel, Pauwels & Vidal 2002 China, Lao PDR, Myanmar, 

Thailand, Vietnam 

Trimeresurus nebularis Vogel, David & Pauwels 2004 Thailand 

Trimeresurus truongsonensis Orlov, Ryabov, Bui & Ho 2004 Vietnam 

Trimeresurus vogeli 

David, Vidal & Pauwels 2001 Cambodia, China, Lao PDR, 

Thailand, Vietnam 

Typhlops roxaneae Wallach 2001 Thailand 

Subtotal 22 

TURTLES 

Chitra vandijki McCord & Pritchard 2002 Myanmar 

Cyclemys atripons Iverson & McCord 1997 Thailand / Cambodia / Vietnam 

Cyclemys pulchristriata Fritz, Gaulke & Lehr 1997 Vietnam 

Mauremys pritchardi McCord 1997 Myanmar 

Subtotal 4 

TOTAL NUMBER OF REPTILES 72 

SPIDERS 


A. inthanonensis Dankittipakul, Wang 2003 Thailand 

A. siamensis Dankittipakul, Wang 2003 Thailand 

Allagelena monticola Chami-Kranon, Likhitrakarn & 

Dankittipakul 

2007 Thailand 

Asiacoelotes sparus Dankittipakul, Chami-Kranon & 

Wang 

2005 Thailand 

C. lannaensis Schwendi nger 2005 Thailand 

C. siamensis Schwendi nger 2005 Thailand

Clubiona aculeata Zhang, Zhu & Song 2007 Yunnan 

Clubiona lamina Zhang, Zhu & Song 2007 Yunnan 

Clubiona tengchong Zhang, Zhu & Song 2007 Yunnan 

Coelotes suthepicus Dankitti pakul, Chami-Kranon & 

Wang 

2005 Thailand 

Coelotes Thailandensis Dankitti pakul, Wang 2003 Thailand 

Coronilla lanna Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Cydrela deciduas Dankitti pakul & Jocqué 2006 Thailand 

Cydrela pristine Dankitti pakul & Jocqué 2006 Thailand 

D. anthonyi Dankitti pakul, Wang 2003 Thailand 

D. paralateralis Dankitti pakul, Wang 2004 Thailand 

D. pseudolateralis Dankitti pakul, Wang 2004 Thailand 

D. subulatus Dankitti pakul, Wang 2003 Thailand 

Deinopis liukuensis Yin, Gri swol d & Yan 2002 Yunnan (China) 

Draconarius 

globul atus 

Chami-Kranon, Sonthichai & Wang 2006 Thailand 

Draconarius abbreviatus Dankitti pakul, Wang 2003 Thailand 

Draconarius australis Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius elatus Dankitti pakul, Wang 2004 Thailand 

Draconarius lateralis Dankitti pakul, Wang 2004 Thailand 

Draconarius monticol a Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius montis Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius papai Chami-Kranon, Sonthichai & Wang 2006 Thailand 

Draconarius phuhi n Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius promontories Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius schwendingeri Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius silva Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius silvicola Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Draconarius tentus Dankitti pakul, Sonthichai & Wang 2006 Thailand 

Evarcha bicuspidata Peng, Li 2003 Vietnam 

Heptathela abca Ono 1999 Vietnam 

Heptathela cucphuongensis Ono 1999 Vietnam 

Heptathela tomokunii Ono 1997 Vietnam 

Heteropoda dagmarae Jäger & Vedel 2005 Lao PDR 

Heteropoda maxima Jäger 2001 Lao PDR 

Liphistius isan Schwendinger 1998 Thailand 

Liphistius phileion Schwendinger 1998 Thailand 

Liphistius phuketensis Schwendinger 1998 Thailand 

Liphistius sayam Schwendinger 1998 Thailand 

Liphistius lahu Schwendinger 1998 Thailand 

Lysitel es conicus Tang, Yin, Peng, Ubick & Gris wold 2007 Yunnan (China) 

Lysitel es davidi Tang, Yin, Peng, Ubick & Gris wold 2007 Yunnan (China) 

Lysitel es dentatus Tang, Yin, Peng, Ubick & Gris wold 2007 Yunnan (China) 

M. flammea Ono 2004 Vietnam 

M. karubei Ono 2003 Vietnam 

M. septemmaculata Ono 2004 Vietnam 

M. thinhi Ono 2003 Vietnam 

M. vietnamensis Ono 2003 Vietnam 

Macrothele yani Xu, Yin & Griswold 2002 Yunnan (China) 

Mallinella nomurai Ono 2003 Vietnam 

Mallinella paradi sea Ono 2004 Vietnam 

Neriene yani Chen & Yin 1999 Yunnan (China) 

Pimoa lihengae Griswol d, Long & Hormiga 1999 Yunnan (China) 

Psechrus khammouan Jäger 2007 Lao PDR 

Psechrus luangprabang Jäger 2007 Lao PDR 

Pseudicius wenshanensis He & Hu 1999 Yunnan (China) 

Pseudopoda ameli a Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda cangschana Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda confusa Jäger, Pathoumthong & Vedel 2006 Lao PDR 

Pseudopoda contenti o Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda contrari a Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda daliensis Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda digitata Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda fissa Jäger & Vedel 2005 Vietnam 

Pseudopoda gemina Jäger, Pathoumthong & Vedel 2006 Lao PDR 

Pseudopoda gongschana Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda houaphan Jäger 2007 Lao PDR 

Pseudopoda interposita Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda namkhan Jäger, Pathoumthong & Vedel 2006 Lao PDR 

Pseudopoda obtuse Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda rivicola Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda roganda Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda saetosa Jäger & Vedel 2007 Yunnan (China) 

Pseudopoda shuqiangi Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda sinapophysis Jäger & Vedel 2007 Yunnan (Chi na) 

Pseudopoda yinae Jäger & Vedel 2007 Yunnan (Chi na) 

Sanmenia gongshan Yang, Zhu & Song 2006 Yunnan (Chi na) 

Sesieutes thakek Jäger 2007 Lao PDR 

Sinothomisus liae Tang, Yin, Griswold & Peng 2006 Yunnan (Chi na) 

Smodicinodes schwendingeri Benjamin 2002 Thailand 

Storenomorpha anne Jäger 2007 Lao PDR 

Storenomorpha paguma Grismado & Ramirez 2004 Vietnam 

Utivarachna rama Chami-Kranon, Li khitrakarn & 

Wongsawad. 

2007 Thailand 

Weintrauboa yunnan Yang, Zhu & Song 2006 Yunnan (Chi na) 

TOTAL NUMBER OF SPIDERS 88 

GRAN D TOTAL 1,068 

Cover - Main photo: Gumprecht’s green pitviper 

(Trimeresurus gumprechti) © René Ries. 

From top to bottom: Stone forest near Kunming, 

Yunnan Province, China © Janet Jent / WWF-Canon; 

Aeschynanthus mendumiae, new plant species © 

Royal Botanic Garden Edinburgh; Theloderma licin, 

new frog species © Daicus Belabut; Desmoxytes 

purpurosea, new species of highly toxic shocking 

pink millipede © Somsak Panha; Aerial view of the 

Mekong Delta, southern Vietnam © Elizabeth Kemf / 

WWF-Canon; Annamite striped rabbit, Nesolagus 

timminsi © Trinh Viet Cuong, Fauna and Flora 

International.

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network active in 100 countries. 

WWF’s mission is to stop the degradation of the planet’s natural environment 

and to build a future in which humans live in harmony with nature, by: 

- conserving the world’s biological diversity 

- ensuring that the use of renewable natural resources is sustainable 

- promoting the reduction of pollution and wasteful consumption 

WWF Greater Mekong 

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Hanoi 

Vietnam 

Tel: +(84 4) 3719 3111 

Fax: +(84 4) 3719 3102 

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Produced by the green room: research & text by Christian Thompson, design & layout by Torva Thompson 

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