233.0000
Family
Chrysobalanaceae R.Br.; J. H. Tuckey, Narr. Exped. Zaire: 433 (1818), nom. cons.
20 genera, ca. 540 spp., mainly tropical America. Phylogeny see Sothers et al. 2016.
Species listed in database: 547
= Chrysobalaneae DC.; Prodr. 2: 525 (1825)
= Chrysobalaninae Focke; Engl. & Prantl, Nat. Pflanzenfam. III, 3: 12 (1888)
= Chrysobalanineae J.Presl; Nowoceská Bibl. [Wsobecný Rostl.] 7: 467, 469 (1846)
= Chrysobalanoideae Arn.; Botany: 107 (1832)
= Couepieae Prance & F.White; Phil. Trans. Roy. Soc. Lond., Ser. B, 320: 78 (1988)
= Grangeriinae Bonne; Res. Pédic. Fl. Rosacées: 356 (1928)
= Hirtellaceae Horan.; Char. Ess. Fam.: 152 (1847)
= Hirtelleae Bonne; Res. Pédic. Fl. Rosacées: 356 (1928)
= Hirtellinae Focke; Engl. & Prantl, Nat. Pflanzenfam. III, 3: 12 (1888)
= Licaniaceae Martinov; Tekhno-Bot. Slovar: 336 (1820)
= Parinarieae Prance & F.White; Phil. Trans. Roy. Soc. Lond., Ser. B, 320: 78 (1988)
233.0028
Genus
Couepia Aubl.; Hist. Pl. Guian. Franc. 1: 519, t. 221 (1775)
Distribution: Tropical America from Mexico to S-Brazil
Species listed in database: 63
= Dulacia Neck.; Elem. 2: 414 (1790)
= Acia Schreb.; Gen. Pl.: 458 (1791), nom. superfl.
= Pleragina Arruda; H. Koster, Trav. Brazil: 499 (1816), nomen
species :
Couepia belemii
Prance; Fl. Neotrop. 9: 228 (1972)
Distribution: NE-Brazil (Bahia); SE-Brazil (Espírito Santo)
84 BRA
species :
Couepia bernardii
Prance; Brittonia 33(3): 347 (1981)
Distribution: Peru; N-Brazil (Amazonas)
83 PER 84 BRA
species :
Couepia bondarii
Prance; Fl. Neotrop. 9: 240 (1972)
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Couepia bracteosa
Benth.; J. Bot. [Hooker] 2: 215 (1840)
Distribution: Colombia (Vaupés); Guyana; Surinam; French Guiana; Peru; Bolivia (Pando); N-Brazil (Amapa, Pará, Amazonas, Rondônia)
82 CLM FRG GUY SUR 83 BOL PER 84 BRA
Synonyms: hidden
= Couepia bracteosa var. grandifolia Benoist; Bull. Mus. Natl. Hist. Nat. 29: 597 (1923)
= Couepia bracteosa var. minor Ducke; Arch. Jard. Bot. Rio de Janeiro 5: 117 (1930)
= Moquilea bracteosa (Benth.) Walp.; Repert. Bot. Syst. 2: 7 (1843)
= Moquilea rufa Barb.Rodr.; Hort. Flumin. 1893: 165 (1895)
species :
Couepia brevistaminea
Barb.Silva & Antar; Phytotaxa 471(1): 39 (2020)
Distribution: Brazil (Goiás, Tocantins)
84 BRA
species :
Couepia carautae
Prance; Fl. Neotrop. 9S: 70 (1989)
Distribution: SE-Brazil (Espírito Santo)
84 BRA
species :
Couepia caryophylloides
Benoist; Bull. Mus. Hist. Nat. Paris 28: 253 (1922)
subspecies :
subsp. caryophylloides
Distribution: Surinam; French Guiana; N-Brazil (Amapa, Pará, Amazonas)
82 FRG SUR 84 BRA
subspecies :
subsp. glabra
Prance; Fl. Neotrop. 9S: 72 (1989)
Distribution: N-Brazil (Pará)
84 BRA
species :
Couepia cataractae
Ducke; Arq. Serv. Florest. 1(1): 31 (1939)
Distribution: N-Brazil (Pará, Rondônia)
84 BRA
species :
Couepia chrysocalyx
(Poepp.) Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 42 (1867)
Distribution: Colombia (Amazonas, Caquetá, Cauca, Guainía, Putumayo, Vaupés); Venezuela (Amazonas); Ecuador; Peru; N-Brazil (Pará, Amazonas, Acre)
82 CLM VEN 83 ECU PER 84 BRA
Synonyms: hidden
= Moquilea chrysocalyx Poepp.; E. F. Poepp. & S. L. Endl., Nov. Gen. Sp. Pl. 3: 75 (1845)
species :
Couepia cidiana
Prance; Acta Amazonica 13(1): 21 (1983)
Distribution: N-Brazil (Pará)
84 BRA
species :
Couepia coarctata
Prance; Fl. Neotrop. 9S: 77 (1989)
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Couepia comosa
Benth.; J. Bot. [Hooker] 2: 215 (1840)
Distribution: Venezuela (Bolívar); Guyana
82 GUY VEN
Synonyms: hidden
= Moquilea comosa (Benth.) Walp.; Repert. Bot. Syst. 2: 6 (1843)
species :
Couepia eriantha
Spruce ex Hook.f.; Mart., Fl. Bras. 14(2): 45 (1867)
Distribution: N-Brazil (Pará, Amazonas)
84 BRA
species :
Couepia excelsa
Ducke; Arch. Jard. Bot. Rio de Janeiro 5: 116 (1930)
Distribution: French Guiana; N-Brazil (Roraima, Amapa, Pará, Amazonas)
82 FRG 84 BRA
species :
Couepia exflexa
Fanshawe & Maguire; Bull. Torrey Bot. Club 75: 376 (1948)
Distribution: Guyana; French Guiana
82 FRG GUY
species :
Couepia froesii
Prance; Fl. Neotrop. 9: 258 (1972)
Distribution: N-Brazil (Pará, Amazonas)
84 BRA
species :
Couepia glabra
Prance; Acta Amazon. 2: 10 (1972)
Distribution: N-Brazil (Amazonas)
84 BRA
species :
Couepia grandiflora
(Mart. & Zucc.) Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 46 (1867)
Distribution: Bolivia (Santa Cruz); N-Brazil (Tocantins, Rondônia); NE-Brazil (Maranhao, Piauí, Bahia); WC-Brazil (Mato Grosso, Goiás, Brasilia Distrito Federal, Mato Grosso do Sul); SE-Brazil (Minas Gerais, São Paulo); S-Brazil (Paraná); Paraguay (Amambay, Canindeyu, Concepcion)
83 BOL 84 BRA 85 PAR
Synonyms: hidden
= Couepia formosana Taub.; Bot. Jahrb. Syst. 21: 429 (1896)
= Couepia suberosa Pilg.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 538 (1923)
= Moquilea grandiflora Mart. & Zucc.; Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 1: 388 (1832)
species :
Couepia guianensis
Aubl.; Pl. Gui. 1: 519, 207 (1775)
subspecies :
subsp. guianensis
Distribution: Colombia (Amazonas, Antioquia, Caquetá, Chocó, Guainía, Meta, Santander, Vaupés); Venezuela (Amazonas, Bolívar, Delta Amacuro?, Falcon?, Tachira); Guyana; Surinam; French Guiana; Ecuador; Peru; N-Brazil (Amapa, Pará, Amazonas, Rondônia); NE-Brazil (Maranhao)
82 CLM FRG GUY SUR VEN 83 ECU PER 84 BRA
Synonyms: hidden
= Acia amara Willd.; Sp. Pl. 3: 717 (1800), nom. superfl.
= Acia parillo Oken; Allg. Naturgesch. 3(3): 2043 (1841)
= Acioa amara Steud.; Nomencl. Bot. 1: 9 (1821)
= Couepia leptostachya Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 44 (1867)
= Couepia surinamensis Kleinh.; Recueil Trav. Bot. Néerl. 22: 390 (1925)
= Couepia versicolor Benoist; Bull. Mus. Natl. Hist. Nat. 29: 596 (1923)
= Moquilea couepia Steud.; Nomencl. Bot., ed. 2, 2: 159 (1841)
subspecies :
subsp. divaricata
(Hub.) Prance; Brittonia 33(3): 351 (1981)
Distribution: Venezuela (Amazonas, Delta Amacuro); Guyana; Surinam; N-Brazil (Amapa, Pará, Amazonas, Rondônia); NE-Brazil (Maranhao)
82 GUY SUR VEN 84 BRA
Synonyms: hidden
= Couepia divaricata Huber; Bol. Mus. Paraense Hist. Nat. Ethnogr. 6: 75 (1910)
= Couepia divaricata var. strictiuscula Huber; Bol. Mus. Paraense Hist. Nat. Ethnogr. 6: 76 (1910)
= Parinari krukovii Gleason; Bull. Torrey Bot. Club 60: 353 (1933)
subspecies :
subsp. glandulosa
(Miq.) Prance; Brittonia 33(3): 350 (1981)
Distribution: Colombia; Venezuela (Amazonas, Bolívar, Delta Amacuro); Guyana; Surinam; French Guiana; N-Brazil (Roraima, Amapa, Pará, Amazonas); NE-Brazil (Maranhao)
82 CLM FRG GUY SUR VEN 84 BRA
Synonyms: hidden
= Couepia glandulosa Miq.; Stirp. Surinam. Select.: 28 (1851)
= Couepia myrtifolia Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 44 (1867)
= Couepia thyrsiflora Hook.f.; Mart., Fl. Bras. 14(2): 45 (1867)
= Moquilea glandulosa (Miq.) Walp.; Ann. Bot. Syst. 2: 463 (1852)
species :
Couepia habrantha
Standl.; Publ. Field Mus. Nat. Hist., Chicago, Bot. Ser. 17: 249 (1937)
Distribution: Colombia (Amazonas); Venezuela (Amazonas); Guyana; French Guiana; N-Brazil (Pará, Amazonas)
82 CLM FRG GUY VEN 84 BRA
species :
Couepia hallwachsiae
D.Santam. & Lagom.; Phytotaxa 233(1): 70 (2015)
Distribution: Costa Rica
80 COS
species :
Couepia hondurasensis
Prance; Kew Bull. 54(1): 110 (1999)
Distribution: Honduras
80 HON
Remarks: known only from the type collection
species :
Couepia impressa
Prance; Fl. Neotrop. 9: 255 (1972)
subspecies :
subsp. impressa
Distribution: NE-Brazil (Pernambuco, Alagoas)
84 BRA
subspecies :
subsp. cabraliae
Prance; Rev. Brasil. Bot. 2(1): 33 (1979)
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Couepia insignis
Fritsch; Ann. naturh. Hofmus. Wien. 5: (1890) xi.
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Couepia janzenii
D.Santam. & Lagom.; Phytotaxa 233(1): 73 (2015)
Distribution: Costa Rica; Panama
80 COS PAN
species :
Couepia joaquinae
Prance; Kew Bull. 46(1): 107 (1991)
Distribution: French Guiana; N-Brazil (Amapa)
82 FRG 84 BRA
species :
Couepia krukovii
Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 17: 250 (1937)
Distribution: Colombia (Amazonas); Bolivia (Beni); N-Brazil (Amazonas, Rondônia)
82 CLM 83 BOL 84 BRA
species :
Couepia latifolia
Standl.; Publ. Field Mus. Nat. Hist., Chicago, Bot. Ser. 17: 251 (1937)
Distribution: Colombia (Vaupés); Peru; Bolivia (Beni); N-Brazil (Amazonas, Acre)
82 CLM 83 BOL PER 84 BRA
species :
Couepia leitaofilhoi
Prance; Kew Bull. 54(1): 108 (1999)
Distribution: SE-Brazil (São Paulo)
84 BRA
species :
Couepia longipetiolata
Prance; Rev. Brasil. Bot. 2(1): 31 (1979)
Distribution: NE-Brazil (Bahia)
84 BRA
species :
Couepia macrophylla
Spruce ex Hook.f.; Mart., Fl. Bras. 14(2): 41 (1867)
Distribution: Panama; Colombia (Amazonas); Ecuador; Peru; N-Brazil (Amazonas, Acre)
80 PAN 82 CLM 83 ECU PER 84 BRA
Synonyms: hidden
= Couepia speciosa Pilg.; Bot. Jahrb. Syst. 37: 539 (1906)
species :
Couepia magnoliifolia
Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 43 (1867)
Distribution: French Guiana; N-Brazil (Amazonas)
82 FRG 84 BRA
species :
Couepia maguirei
Prance; Fl. Neotrop. 9: 214 (1972)
Distribution: Colombia (Guainía); Venezuela (Amazonas, Bolívar); N-Brazil (Amazonas)
82 CLM VEN 84 BRA
species :
Couepia marleneae
Prance; Acta Amazonica. 4(1): 17 (1974)
Distribution: N-Brazil (Amazonas)
84 BRA
species :
Couepia martinii
Prance; Fl. Neotrop. 9: 238 (1972)
Distribution: French Guiana
82 FRG
species :
Couepia meridionalis
Prance; Fl. Neotrop. 9: 256 (1972)
Distribution: SE-Brazil (São Paulo)
84 BRA
species :
Couepia monteclarensis
Prance; Fl. Neotrop. 9S: 66 (1989)
Distribution: NE-Brazil (Bahia); SE-Brazil (Minas Gerais, São Paulo, Rio de Janeiro)
84 BRA
species :
Couepia morii
Prance; Kew Bull. 47(2): 254 (1992)
Distribution: N-Brazil (Amazonas)
84 BRA
species :
Couepia multiflora
Benth.; J. Bot. [Hooker] 2: 215 (1840)
Distribution: Guyana; N-Brazil (Roraima)
82 GUY 84 BRA
Synonyms: hidden
= Moquilea multiflora (Benth.) Walp.; Repert. Bot. Syst. 2: 7 (1843)
species :
Couepia nutans
Prance; Brittonia 31(2): 248 (1979)
Distribution: Colombia (Valle)
82 CLM
species :
Couepia obovata
Ducke; Archiv. Inst. Biol. Veg., Rio de Joneiro 2: 35 (1935)
Distribution: Colombia (Vaupés); Venezuela (Amazonas); Guyana; Surinam; French Guiana; Ecuador; Peru; Bolivia (Pando); N-Brazil (Pará, Amazonas)
82 CLM FRG GUY SUR VEN 83 BOL ECU PER 84 BRA
species :
Couepia osaensis
Aguilar & D.Santam.; Phytoneuron 2014-64: 2 (2014)
Distribution: Costa Rica
80 COS
species :
Couepia ovalifolia
(Schott) Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 48 (1867)
Distribution: NE-Brazil (Pernambuco, Bahia); SE-Brazil (Minas Gerais, Espírito Santo, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Chrysobalanus ovalifolius Schott; K. P. J. Spreng., Syst. Veg. 4(2): 406 (1827)
= Couepia hypoleuca Miq.; Linnaea 19: 438 (1846)
= Moquilea hypoleuca (Miq.) B.D.Jacks.; Index Kew. 2: 262 (1894)
species :
Couepia oxossii
Amorim & Asprino; J. Torr. Bot. Soc. 145(3): 272 (2018)
Distribution: Brazil (Bahia)
84 BRA
species :
Couepia paraensis
(Mart. & Zucc.) Benth.; J. Bot. [Hooker] 2: 216 (1840)
subspecies :
subsp. paraensis
Distribution: Venezuela (Amazonas, Anzoategui, Apure, Bolívar, Tachira); Peru; N-Brazil (Pará, Amazonas, Tocantins, Rondônia); WC-Brazil (Mato Grosso, Goiás)
82 VEN 83 PER 84 BRA
Synonyms: hidden
= Moquilea paraensis Mart. & Zucc.; Abh. Akad. Muench. 1: 390 (1832)
= Couepia vismioides Klotzsch ex Hook.f.; Mart., Fl. Bras. 14(2): 48 (1867)
subspecies :
subsp. glaucescens
(Spruce ex Hook.f.) Prance; Fl. Neotrop. 9: 212 (1972)
Distribution: Colombia (Arauca, Guainía, Vichada); Venezuela (Amazonas, Apure, Bolívar, Falcon, Guarico, Monagas, Tachira); Guyana; Peru; Bolivia (Beni, Pando); N-Brazil (Roraima, Pará, Amazonas, Tocantins, Acre, Rondônia); WC-Brazil (Mato Grosso)
82 CLM GUY VEN 83 BOL PER 84 BRA
Synonyms: hidden
= Couepia glaucescens Spruce ex Hook.f.; Mart., Fl. Bras. 14: II. 49
= Couepia duckei Huber; Bol. Mus. Paraense Hist. Nat. Ethnogr. 5: 371 (1909)
subspecies :
subsp. cerradoana
Prance; Fl. Neotrop. 9: 213 (1972)
Distribution: N-Brazil (Pará, Tocantins); NE-Brazil (Maranhao); WC-Brazil (Mato Grosso)
84 BRA
species :
Couepia parvifolia
Prance; Fl. Neotrop. 9: 258 (1972)
Distribution: SE-Brazil (Rio de Janeiro)
84 BRA
species :
Couepia pernambucensis
Prance; Fl. Neotrop. 9: 256 (1972)
Distribution: NE-Brazil (Pernambuco, Bahia, Alagoas)
84 BRA
species :
Couepia polyandra
(Kunth) Rose; U. S. Dept. Agric. Contrib. Natl. Herb. 5: 196 (1899)
Distribution: Mexico (Campeche, Chiapas, Colima, Guanajuato, Guerrero, Jalisco, Michoacan, Nayarit, Oaxaca, Puebla, Queretaro, Quintana Roo, Sinaloa, Tabasco, Veracruz, Yucatan); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Cuba [c]
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 81 cub
Synonyms: hidden
= Couepia dodecandra (Moc. & Sessé ex DC.) Hemsl.; Hooker´s Icon. Pl. 27: t. 2620 (1901)
= Couepia floccosa Fritsch; Ann. K. K. Naturhist. Hofmus. 5: 12 (1890)
= Couepia kunthiana (Mart. & Zucc.) Benth.; J. Bot. [Hooker] 2: 216 (1840), nom. superfl.
= Grymania polyandra (Kunth) C.Presl; Epimel. Bot.: 193 (1851)
= Hirtella dodecandra Moc. & Sessé ex DC.; Prodr. 2: 529 (1825)
= Hirtella polyandra Kunth; Humb., Bonpl. & Kunth, Nov. Gen. Sp. 6: 246 (1824)
= Moquilea kunthiana Mart. & Zucc.; Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 1: 390 (1832), nom. superfl.
species :
Couepia rankiniae
Prance; Kew Bull. 47(4): 640 (1992), as ´rankinae´
Distribution: French Guiana; N-Brazil (Amazonas)
82 FRG 84 BRA
species :
Couepia reflexa
Ducke; Arch. Jard. Bot. Rio de Janeiro 5: 118 (1930)
Distribution: N-Brazil (Pará)
84 BRA
species :
Couepia robusta
Huber; Bol. Mus. Paraense Hist. Nat. Ethnogr. 6: 76 (1910)
Distribution: N-Brazil (Amapa, Pará, Amazonas); NE-Brazil (Maranhao)
84 BRA
species :
Couepia rufa
Ducke; Arch. Mus. Nac. Rio de Janeiro 22: 66 (1919)
Distribution: NE-Brazil (Pernambuco, Bahia, Alagoas); SE-Brazil (Minas Gerais)
84 BRA
Synonyms: hidden
= Pleragina rufa Arruda; Trav. Brazil: 499 (1816), nom. inval.
= Couepia glazioviana Pilg.; Notizbl. Bot. Gart. Berlin-Dahlem 8: 537 (1923)
species :
Couepia sandwithii
Prance; Fl. Neotrop. 9: 216 (1972)
Distribution: Venezuela (Bolívar); Guyana
82 GUY VEN
species :
Couepia schottii
Fritsch; Ann. K. K. Naturhist. Hofmus. 5: 13 (1890)
Distribution: NE-Brazil (Bahia); SE-Brazil (Espírito Santo, Rio de Janeiro)
84 BRA
Synonyms: hidden
= Chrysobalanus macrophyllus Schott; K. P. J. Spreng., Syst. Veg. 4(2): 407 (1827)
species :
Couepia scottmorii
Prance; Fl. Neotrop. 9S: 68 (1989)
Distribution: Panama; Colombia
80 PAN 82 CLM
species :
Couepia spicata
Ducke; Archiv. Inst. Biol. Veg., Rio de Janeiro 2: 36 (1935)
Distribution: N-Brazil (Amazonas)
84 BRA
species :
Couepia stipularis
Ducke; Archiv. Inst. Biol. Veg., Rio de Janeiro 4: 3 (1938)
Distribution: N-Brazil (Amazonas)
84 BRA
species :
Couepia subcordata
Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 46 (1867)
Distribution: Colombia (Amazonas); Ecuador; Peru; N-Brazil (Pará, Amazonas)
82 CLM 83 ECU PER 84 BRA
Synonyms: hidden
= Couepia amazonica Fritsch; Ann. K. K. Naturhist. Hofmus. 5: 12 (1890)
species :
Couepia trapezioana
Cuatrec.; Brittonia 8: 197 (1956)
Distribution: Colombia (Amazonas); Peru; N-Brazil (Amazonas)
82 CLM 83 PER 84 BRA
species :
Couepia uiti
(Mart. & Zucc.) Benth. ex Hook.f.; Mart., Fl. Bras. 14(2): 47 (1867)
Distribution: Bolivia (Santa Cruz); N-Brazil (Rondônia); NE-Brazil (Piauí, Pernambuco, Bahia, Alagoas, Sergipe); WC-Brazil (Mato Grosso, Goiás, Mato Grosso do Sul); SE-Brazil (Minas Gerais, São Paulo); Paraguay (Concepcion)
83 BOL 84 BRA 85 PAR
Synonyms: hidden
= Moquilea uiti Mart. & Zucc.; Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 1: 390 (1832)
= Couepia martiana Hook.f.; Mart., Fl. Bras. 14(2): 47 (1867)
= Couepia paraguariensis Hassl.; Repert. Spec. Nov. Regni Veg. 7: 375 (1909)
= Couepia dahlgrenii Standl.; Publ. Field Mus. Nat. Hist., Bot. Ser. 17: 249 (1937)
species :
Couepia ulei
Pilg.; Verh. Bot. Ver. Brandenb. 47: 149 (1905)
Distribution: Colombia (Amazonas); Venezuela (Apure); Ecuador; Peru; N-Brazil (Amazonas, Acre, Rondônia)
82 CLM VEN 83 ECU PER 84 BRA
species :
Couepia venosa
Prance; Fl. Neotrop. 9: 251 (1972)
Distribution: SE-Brazil (Minas Gerais, São Paulo, Rio de Janeiro)
84 BRA
species :
Couepia williamsii
J.F.Macbr.; Candollea 5: 366 (1934)
Distribution: Colombia (Amazonas); Peru; N-Brazil (Amazonas, Acre)
82 CLM 83 PER 84 BRA
