Contents

Foreword ....................................................................................................... ix
Introduction
The Genus Begonia in Borneo ................................................................. 1
Borneo ..................................................................................................... 2
History ..................................................................................................... 5
Begonia Diversity .................................................................................. 11
How to Recognise a Begonia? ............................................................... 12
Pollination and Dispersal ....................................................................... 14
Grouping Begonias ................................................................................ 16
Where to Find Begonias? ....................................................................... 18
Conservation ......................................................................................... 23
What Next? ............................................................................................ 24

A Selection of Species
Begonia adliniana ................. 28 Begonia congesta .................. 82
Begonia alabensis ................. 30 Begonia conipila ................... 84
Begonia amphioxus ............... 32 Begonia conniegeriae ............ 86
Begonia andersonii ............... 34 Begonia corrugata ................ 88
Begonia apiensis ................... 36 Begonia crockerensis ............ 90
Begonia awongii ................... 38 Begonia cyanescens .............. 92
Begonia bakunensis .............. 40 Begonia danumensis ............. 94
Begonia baramensis .............. 42 Begonia dimorpha ................ 96
Begonia basintaliana ........... 44 Begonia divergens ................. 98
Begonia belagaensis ............. 46 Begonia doloisii .................. 100
Begonia benaratensis ............ 48 Begonia erythrogyna ........... 102
Begonia bengohensis ............ 50 Begonia eutricha ................. 104
Begonia bernadusii ............... 52 Begonia flammea ................. 106
Begonia beryllae ................... 54 Begonia flavovirens ............. 108
A Guide to Begonias of Borneo

Begonia bintang .................... 56 Begonia fuscisetosa ............. 110

Begonia bosuangiana ........... 58 Begonia gibbsiae ................. 112
Begonia bruneiana ................ 60 Begonia gomantongensis .... 114
Begonia burbidgei ................ 62 Begonia gueritziana ............ 116
Begonia burttii ...................... 66 Begonia gusilii .................... 118
Begonia calcarea .................. 68 Begonia hexaptera .............. 120
Begonia chaiana ................... 70 Begonia hullettii .................. 122
Begonia chakensis ................ 72 Begonia incompta ............... 124
Begonia chlorocarpa ............ 74 Begonia inobongensis ......... 126
Begonia chlorosticta ............. 76 Begonia inostegia ............... 128
Begonia chongii .................... 80 Begonia inostegioides .......... 129

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 Begonia jamiliana ............... 130 Begonia penrissenensis ....... 206
Begonia joffrei .................... 132 Begonia peridoticola ........... 208
Begonia johariana .............. 134 Begonia piring .................... 210
Begonia jugamensis ............ 136 Begonia plieranensis ........... 212
Begonia julaihiana .............. 138 Begonia postarii .................. 214
Begonia juliasangii ............. 140 Begonia prasinimarginata .. 216
Begonia kachak ................... 142 Begonia propinqua .............. 218
Begonia kasutensis .............. 144 Begonia punchak ................. 220
Begonia keithii .................... 146 Begonia rambutan ............... 222
Begonia kiamfeei ................. 148 Begonia ramlanii ................ 224
Begonia kibambangensis .... 150 Begonia rhodochaeta .......... 226
Begonia kinabaluensis ........ 152 Begonia rhodoneura ........... 228
Begonia kipandiensis .......... 154 Begonia rhodotricha ........... 230
Begonia labiensis ................ 156 Begonia rhombipetala ......... 232
Begonia lailana ................... 158 Begonia roseopunctata ....... 234
Begonia lambii .................... 162 Begonia rotundibracteata ... 238
Begonia lambirensis ............ 164 Begonia rubida ................... 240
Begonia lamriana ............... 166 Begonia ruthiae ................... 242
Begonia lazat ...................... 168 Begonia sabahensis ............. 244
Begonia linauensis .............. 170 Begonia sarangica .............. 246
Begonia lucychongiana ....... 172 Begonia serapatensis .......... 248
Begonia madaiensis ............ 174 Begonia setiamensis ............ 250
Begonia magentifolia .......... 176 Begonia sibutensis .............. 252
Begonia malachosticta ........ 178 Begonia simunii .................. 254
Begonia mariaensis ............. 180 Begonia sirukitii .................. 256
Begonia matangensis .......... 182 Begonia speluncae .............. 258
Begonia melanosticta .......... 184 Begonia stenogyna .............. 260
Begonia melinauensis ......... 186 Begonia subisensis .............. 262
Begonia murumensis ........... 188 Begonia taniana .................. 264
Begonia nagaensis .............. 190 Begonia tenuissima ............. 266
Begonia niahensis ............... 192 Begonia tomaniensis ........... 268 A Guide to Begonias of Borneo
Begonia nothobaramensis ... 194 Begonia umbratica .............. 270
Begonia oblongifolia ........... 196 Begonia vaccinioides .......... 272
Begonia paoana .................. 198 Begonia vanderentii ............ 274
Begonia papyraptera .......... 200 Begonia vespipropinqua ..... 276
Begonia payung .................. 202 Begonia vulgare .................. 278
Begonia pendula ................. 204 Begonia xiphophylla ........... 280
A Checklist of Begonias in Borneo ............................................................. 282
Further Reading .......................................................................................... 285
Acknowledgements .................................................................................... 289
Photo Credits .............................................................................................. 290
Index .......................................................................................................... 291
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