A Nomenclator

of Mexican

and

Central

American

Rubiaceae

David H. Lorence

tn^\ ^llk- Missouri

(3 ^ Botanical
Garden
Press
 MONOGRAPHS IN SYSTEMATIC BOTANY
FROM THE MISSOURI BOTANICAL GARDEN

AUTHOR'S ADDRESS

David H. Lorence
National Tropical Botanical Garden
P.O. Box 340
Lawai, Hawaii 96765

VOLUME EDITOR
Marshall R. Crosby

MANAGING EDITOR
Amy Scheuler McPherson

Copyright © 1999 by Missouri Botanical Garden Pres;

P.O. Box 299
St. Louis, Missouri 63166-0299, U.S.A.
All rights reserved.
Printed in U.S.A.
 Contents

Introduction

Format

Acknowledgments

References Cited
INTRODUCTION

iomenclatural compilation for ongoing revisionary and floristic studies

1 Rubiaceae, including treatments for the Flora Mesoamericana, Flora
of Chiapas, Flora de Oaxaca, and Flora de Veracruz projects. Its primary purpose is to provide a
comprehensive nomenclarural index of the published names of Rubiaceae from all of Mexico and the
countries comprising Central America, and to indicate their current taxonomic and nomenclarural status.
Names of taxa described from other areas, principally South America, the West Indies, and the southwest
United States, have also been included if authentic records of their occurrence in Mexico or Central
America exist. Names that have been erroneously applied to taxa in this region are also included. Taxa
that are known from cultivation, have become naturalized, or have escaped from cultivation in the region
are also included, and their status is noted. This work includes ca. 2341 entries of specific and infraspecific
names published during the period from 1753 through 1997, plus several more published to mid-1998.
The following herbaria were inventoried for Rubiaceae types from the region: ENCB, F, MEXU, MO,
P, US, and XAL. In addition, type material of selected taxa was borrowed from a number of other herbaria
including: CAS, GH, K, TEX-LL, UC, and WIS. Black and white photos have been taken of most types
and sets of duplicate prints deposited at MEXU, PTBG, and US. The original literature sources in which
the names were published have been checked in nearly all cases.
All type collections have been seen by the author, unless indicated by "n.v." (non visi). Article 9.1 of
the International Code ofBotanical Nomenclature [Tokyo Code] (Greuter et al, 1994) states "A holotype
of a name of a species or infraspecific taxon is the one specimen or illustration used by the author, or
designated by the author as the nomenclarural type." Names published prior to the Code, however, are
frequently associated with typification problems. It was often the practice of various authors (e.g., Ruiz
Lopez and Pavon, or Sesse and Mocino) to base their descriptions and sometimes also their illustrations
primarily or entirely on plants studied in the field, or even on a mixture of specimens of the same taxon.
In these cases a type was not designated and apparently does not exist, which I have noted in the text.
Alternatively, authors would sometimes cite a locality (usually a country or island) and collector's name,
suggesting that they were referring to a single specimen, although this is difficult to prove.
Recommendation 9A.4 of the Code states that if a single specimen is cited in the protologue, without
mention of a specific institution housing it, a specimen at the author's home institution can be regarded
as the holotype until it can be proven that more than one specimen exists. In these cases I have cited
locality and collector information from the protologue and simply noted "type," followed by the probable
herbarium in which the type material may be sought, usually the author's home institution. Monographic
or revisionary studies may be required to determine the status of any original collections and whether or
not designation of a lectotype or neotype is necessary.
The Sesse and Mocino herbarium is housed at Madrid (MA) with duplicates in several other herbaria.
Sesse and Mocino had a large set of colored plates drawn from living specimens in Mexico and Puerto
Rico, the majority of which are now at the Hunt Institute for Botanical Documentation (HU). They were
copied while on loan to A. P. de Candolle, and sets of these outlines or "caiques" were distributed to at
least 12 herbaria, including G-DC. For further information, see Taxonomic Literature (TL-2) by Stafleu
and Cowan (5: 525-528. 1985). I have not seen type material for most names published by Sesse and
Mocino, and the published diagnoses are usually too brief to permit accurate identification of the species.
Consequently the status of many Sesse and Mocino names must remain uncertain pending study of the
type material and original illustrations, which is beyond the scope of this work. Types of the taxa
described by A. P. de Candolle based on Haenke collections are deposited at PR rather than at G-DC (see
Stafleu & Cowan, 1976: 438).
All cited locality data have been taken directly from the labels of the type specimens. In cases where
the type—or a photo or microfiche of it—was not seen, locality information was taken directly from the
protologue. Lectotypes are designated herein for the first time for non-lectotypified names in cases where
the syntypes have been seen.
Microfiches, mostly of type specimens in the de Candolle herbarium (G-DC), are cited in one of two
ways. The microfiche number is equivalent to the volume, page number, and species number based on
their enumeration in volume 4 of de Candolle's Prodromus containing the Rubiaceae treatment (1830).
For example, the type ofCoutarea mexicana is Martius s.n. (Holotype G-DC), and the microfiche citation
is 4: 350.3. Alternatively, the IDC (Inter Documentation Company AF) series is used. Here the G-DC
herbarium is item number 800 and each fiche has an index number. For example, the type of Contarea
mexicana is located on fiche number 680, in the second row, the seventh from the left, so the resulting
IDC number is 800.680.II.7.
Herbarium acronyms are abbreviated following Index Herbariorum (Holmgren et al., 1990).
Abbreviations for authors of botanical names follow Brummitt and Powell (1992). Serial publications have
been abbreviated following Botanico-Periodicum-Huntianum (Lawrence et al., 1968) and Botanico-
Periodicum-Huntiamim/Supplementum (Bridson & Smith, 1991). Book titles and other non-periodical
abbreviations follow Taxonomic Literature (TL-2) by Stafleu and Cowan (1976-1988). Other
abbreviations frequently used herein are: ft. (feet); m (meters); Distr. (Distnto or District); Mun.
(Municipio or Municipality); names of months are always abbreviated by the first three letters in English
(i.e., Jan for January).
Taxonomic concepts utilized in this work generally follow those presented in the most recently
published monographs, revisions, and floristic treatments available. The nomenclature for the tribe
Spermacoceae (including the genera Crusea. Diodia. Enwdia, Galiaiuhe, Miimcarpus, Richardia,
Spermacoce, Staelia, and Tobagoa) follows the annotations and advice of C. Dennis Adams (BM), who
is preparing the treatment for Flora Mesoamericana. Much of this information is summarized in the
extremely useful checklist of neotropical Rubiaceae published by Andersson (1992). In other cases,
however, my own taxonomic judgments are presented. Any comments, corrections, and additions from
readers will be greatly appreciated.

FORMAT

At the beginning of each generic listing the name of each genus and its type species are always written
in normal Roman type. Typifications for these generic names mostly follow Index Nominwn Generic orum
iPLintarum) (Farr et al., 1979).
For each entry, an infrageneric name written in boldface type is accepted as valid, whereas an italicized
name is considered to be a synonym of the name in boldface type preceded by an em dash (—) at the end
of each entry. Each name is cross-referenced with its basionym, which is also italicized. Blocking names
have been indicated in cases where generic transfers have necessitated new names. Other generic
combinations (homotypic synonyms) that appear in this catalog are indicated by the generic name(s) listed
in italics just preceding the accepted name in boldface at the end of each citation. The following examples
illustrate these points.

(1)The accepted name (boldface type) is followed by the author(s) and publication citation, the basionym
(italics) its publication citation, and type information; this is followed by generic names (also in italics)
under which other combinations (=homotypic synonyms) are listed in this catalog:

Elaeagia myriantha (Standi.) C. M. Taylor & Hammel, Novon 3:490.1993. Sickingia myriantha Standi.,
Publ. Field Columbian Mus., Bot. Ser. 7: 27. 1930. Type: COLOMBIA: Magdalena, Las Nubes road,
region of Santa Marta, 1200 m, 3 Dec 1898, H. H. Smith 1810 (Holotype F 138660; isotypes MO, NY,
n.v., US, n.v., W, n.v., Y, n.v.). Holtonia, Simira.
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIA

(2) A homotypic synonym (italics) is followed by its publication citation, the basionym (italics) and its
publication citation and type information, and another genus under which a homotypic synonym exists
(italics). Finally, this is cross-referenced to the accepted name (boldface):

Holtonia myriantha (Standi.) Standi., Trop. Woods 30: 37. 1932. Sickingia myriantha Standi., Publ. Field
Columbian Mus., Bot. Ser. 7: 27. 1930. Type: COLOMBIA: Magdalena, Las Nubes road, region of
Santa Marta, 1200 m, 3 Dec 1898, H. H. Smith 1810 (Holotype F 138660; isotypes MO, NY, n.v., US,
n.v., W, n.v., Y, n.v.). Simira — Elaeagia myriantha (Standi.) C. M. Taylor & Hammel

ACKNOWLEDGMENTS

I sincerely thank the following individuals and institutions whose assistance has been instrumental in
completing this work: John Dwyer for inspiring this nomenclator and for making available to me notes
on the types he examined at BM and K; C. Dennis Adams (BM) for welcome advice and assistance with
Spermacoceae; Tim McDowell for information regarding Exostema and Syringantha; Robert M. Rhode
for information regarding Chiococca; Marshall R. Crosby, Gerrit Davidse, and Amy McPherson for
helpful discussions, comments, and suggestions; Lennart Andersson and Charlotte Taylor for their detailed
reviews, which substantially improved the manuscript; Janet Leopold and Teri Teasdale for help with
often tedious manuscript preparation; Rick Hanna for locating many obscure references; and Barbara
Mack for updating and formatting the manuscript for publication. I also thank the curators of the herbaria
cited within for loaning specimens and for providing work space and access to their collections. During
my tenure as Investigador at the Herbario Nacional de Mexico (MEXU), the Instituto de Biologia of the
Universidad Nacional Autonoma de Mexico provided financial support, enabling me to study and
photograph types of Mexican and Central American Rubiaceae at the F, MO, P, and US herbaria.
Additional financial support enabling me to study and photograph types at P was provided by a grant from
CONACYT (Mexico). The Missouri Botanical Garden provided facilities and support during my many
visits there. Finally, I am grateful to William L. Theobald and the late William McK. Klein, Jr., former
directors of the National Tropical Botanical Garden, for supporting various phases of work on this project.
My sincere gratitude is extended to all who have helped me.

REFERENCES CITED

Anderson, W. R. 1972. A monograph of the genus Crusea (Rubiaceae). Mem. New York Bot. Gard. 22:
1-128.
Andersson, L. 1992. A Provisional Checklist of Neotropical Rubiaceae. Scripta Botanica Belgica 1:
1-199.
Andersson, L. & C. M. Taylor. 1994. Rubiaceae-Cinchoneae-Coptosapelteae. In G. Harling & L.
Andersson (editors), Flora of Ecuador 50. No. 162 (1-4). Berlings, Arldv, Sweden.
Blackwell, W. FL, Jr. 1968. Revision of Bouvardia (Rubiaceae). Ann. Missouri Bot. Gard. 55: 1-30.
Boom, B. M. 1984. A revision of Isertia (Isertieae: Rubiaceae). Brittonia 36: 425^54.
Bridson, D. & B. Verdcourt. 1988. In R. M. Polhill (editor), Flora of Tropical East Africa, Rubiaceae, part
2: 415-747. Rotterdam, Brookfield: A. A. Balkema.
Bridson, G D. R. & E. R. Smith. 1991. Botanico-Periodicum-Huntianum/Supplementum. Pittsburgh: Hunt

Brummitt, R. K. & C. E. Powell. 1992. Authors of Plant Names. Kew: Royal Botanic Gardens.
Burger, W. C. & C. M. Taylor. 1993. Family # 202 Rubiaceae. In W. C. Burger (editor), Flora
Costaricensis. Fieldiana, Bot., n.s. 33: 1-333.
Darwin, S. P. 1976. The genus Lindenia (Rubiaceae). J. Arnold Arbor. 57: 426-449.
Dempster, L. T. 1973a. The polygamous species of the genus Galium (Rubiaceae) section Lophogalium,
of Mexico and Southwestern United States. Univ. Calif. Publ. Bot. 64: 1-36.
Dempster, L. T. 1973b. Note on the type of Rubia acuminata Mart. & Gal. Brittonia 25: 313.
Dempster, L. T. 1976. Galium mexicanum of Central American and western North America. Madrono 23:
378-386.
Dempster, L. T. 1978. The genus Galium (Rubiaceae) in Mexico and Central America. Univ. Calif. Publ.
Bot. 73: 1-33.
Dwyer, J. D. 1980. Rubiaceae. In R. E. Woodson, Jr., R. W. Schery & collaborators, Flora of Panama-Part
IX. Ann. Missouri Bot. Gard. 67: 1-522.
Elias, T. S. 1976. A monograph of the genus Hamelia (Rubiaceae). Mem. New York Bot. Gard. 26:
81-144.
Farr, E. R., J. A. Leussink & F. A. Stafleu. 1979. Index Nominum Genericorum (Plantarum). 3 vols. The
Hague: Bohn, Scheltema & Holkema.
Fosberg, F. R. 1974. Studies in American Rubiaceae 2. Ayuque, Balmea stormae, an endangered Mexican
species. Sida 5: 268-270.
Fosberg, F. R. & M.-H. Sachet. 1989a. Three cultivated ixoras (Rubiaceae). Baileya 23: 74-85.
Fosberg, F. R. & M.-H. Sachet. 1989b. Lectotypification of Ixora coccinea L. (Rubiaceae). Taxon 38:

Greuter, W., F. R. Barrie, H. M. Burdet, W. G. Chaloner, V. Demoulin, D. L. Hawksworth, P. M.

Jorgensen, D. H. Nicolson, P. C. Silva, P. Trehane & J. McNeill. 1994. International Code of Botanical
Nomenclature (Tokyo Code). Regnum Veg. 131.
Hamilton, C. W. 1989. A revision of Mesoamerican Psychotria subgenus Psychotria (Rubiaceae). Ann.
Missouri Bot. Gard. 76: 67-111, 386-429, 886-916.
Holmgren, P. K., N. H. Holmgren & L. C. Barnett. 1990. Index Herbariorum. 8th edition. Bronx: New
York Botanical Garden.
Howard, R. 1989. Rubiaceae. In Flora of the Lesser Antilles 6: 389-471. Jamaica Plain: Arnold
Arboretum, Harvard University.
Kirkbride, J. H., Jr. 1969. A revision of the Panamanian species ofRondeletia (Rubiaceae). Ann. Missouri
Bot. Gard. 55:372-391.
Kirkbride, J. H., Jr. 1997. Manipulus rubiacearum-VI. Brittonia 49: 354-379.
Lawrence, G. H. M., A. F. Giinther Buchheim, G. S. Daniels & H. Dolezal. 1968. Botanico-Periodicum-
Huntianum. Pittsburgh: Hunt Botanical Library.
Lewis, W. H. 1961. Merger of the North American Houstonia and Oldenlandia under Hedyotis. Rhodora
63:216-223.
Lewis, W. H. 1968. Notes on Hedyotis (Rubiaceae) in North America Ann Missouri Bot. Gard. 55:
31-33.
Lewis, W. H. & R. L. Oliver. 1974. A revision of Richardia (Rubiaceae). Brittonia 26: 271-301.
Little, E. L. 1959. Proposal to conserve the generic names 8364, Laugeria Vahl emend. Hook. f. against
Laugieria Jacq. and Laugeria L. (all of Rubiaceae). Taxon 8: 27-28.
Lorence, D. H. 1993. In C. E. Jarvis, F. R. Barrie, D. M. Allan & J. L. Reveal, A List of Linnaean Generic
Names and their Types. Regnum Veg. 127. [Typifications of Morinda royoc p. 68, and Randia mitis
P80.]
Lorence, D. H. & J. D. Dwyer, 1987. New taxa and a new name in Mexican and Central American Randia
(Rubiaceae, Gardenieae). Bol. Soc. Bot. Mexico 47: 37-^8.
Lorence, D. H. & J. D. Dwyer. 1988. A revision of Deppea (Rubiaceae). Allertonia 4: 389-436.
McDowell, T. 1996a. Syringantha coulteri (Hooker f.) T. McDowell, A new combination, and remarks
on the relationships of the monotypic Mexican genus Syringantha Standley (Rubiaceae). Novon 6:
273-279.
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

McDowell, T. 1996b. Exostema (Rubiaceae): Taxonomic history, nomenclature, position and subgeneric
classification. Opera Bot. Belg. 7: 277-295.
Mena V., P. 1986. A revision of the genus Arcytophyllum (Rubiaceae: Hedyotideae). Mem. New York
Bot. Gard. 60: 1-26.
Natali, A. 1993. In C. E. Jarvis, F. R. Barrie, D. M. Allan & J. L. Reveal, A List of Linnaean Generic
Names and their Types. Regnum Vegetabile 127. [Typification of Sherardia an>ensis p. 88.]
Nicolson, D. 1977. Notes for the flora of Dominica. Spermacoce confusa and Schradera exotica. J. Arnold
Arbor. 58: 445^47. [Typification of Spermacoce confusa.}
Puff, C. 1991. The genus Paederia L.: A multidisciplinary study. Opera Bot. Belg. 3: 1 376.
Rendle, A. B. 1934. Linnaean species of Spermacoce. J. Bot. 72: 329-333.
Rendle, A. B. 1936. Spermacoce remota Lam. J. Bot. 74: 10-12.
Reveal, J. 1993. In C. E. Jarvis, F. R. Barrie, D. M. Allan & J. L. Reveal, A List of Linnaean Generic
Names and their Types. Regnum Vegetabile 127. [Typification of Cephalanthus occidentalism p. 31.]
Reveal, J., C. R. Broome, M. L. Brown & G. F. Frick. 1987. On the identities of Maryland plants
mentioned in the first two editions of Linnaeus' Species plantarum. Huntia 7: 209 245.
[Lectotypification of Mitchella repens, p. 215.]
Rogers, G. K. 1987. The genera of Cinchonoideae (Rubiaceae) in the Southeastern United States. J.
Arnold Arbor. 68: 137-183.
Schumann, K. 1891. Rubiaceae. In A. Engler & K. Prantl, Die natiirlichen Pflanzenfamilien IV (4): 1-156.
Leipzig: Wm. Engelmann. [Lectotypification of Crusea, p. 141.]
Smith, A. C. 1974. Studies of Pacific Island plants. XXVII. The genus Gardenia (Rubiaceae) in the Fijian
region. Amer. J. Bot. 61: 109-128.
Stafleu, F. & R. Cowan. 1976-1988. Taxonomic Literature. Vols. 1-7. Utrecht: Bonn, Scheltema &
Holkema.
Standley, P. C. 1926. Rubiaceae. In Trees and Shrubs of Mexico. Contr. U.S. Natl. Herb. 23: 1349-1394.
Standley, P. C. 1934. Rubiaceae. In North American Flora 32(3-4): 159-300.
Standley, P. C. 1938. Rubiaceae. In Flora of Costa Rica. Publ. Field Mus. Nat. Hist., Bot. Ser. 18:
1264-1380.
Standley, P. C. & L. O. Williams. 1975. Rubiaceae. In Flora of Guatemala. Fieldiana: Bot. 24(11): 1-274.
Steyermark, J. A. 1972. Rubiaceae, Part IX. In B. Maguire and collaborators, The Botany of the Guyana
Highland. Mem. New York Bot. Gard. 23: 227-832.
Steyermark, J. A. 1974. Rubiaceae. In Flora de Venezuela 9(1-3): 1-2070.
Taylor, C. M. 1989. Revision ofHillia sect. Ravnia (Rubiaceae: Cinchonioideae). Selbyana 11: 26-34.
Taylor, C. M. 1994. Revision of Hillia (Rubiaceae). Ann. Missouri Bot. Gard. 81: 571-609.
Taylor, C. M. & D. H. Lorence. 1985. Lectotypification of Palicourea galeottiana M. Martens
(Rubiaceae) and a new name for this common species. Taxon 34: 667-669.
Taylor, C. M. & D. H. Lorence. 1993. On the status of Randia armata (Sw.) DC. (Rubiaceae: Gardenieae).
Taxon 42: 865-867.
Terrell, E. E. 1980. Two new species and a new combination in Houstonia (Rubiaceae) from Mexico.
Brittonia 32: 490-494.
Terrell, E. E. 1996. Revision of Houstonia (Rubiaceae-Hedyotideae). Syst. Bot. Monogr. 48: 1-118.
Terrell, E. E. & W. H. Lewis. 1990. Oldenlandiopsis (Rubiaceae), a new genus from the Caribbean Basin,
based on Oldenlandia callitrichoides Grisebach. Brittonia 42: 185-190.
Verdcourt, B. 1975. Studies in the Rubiaceae-Rubioideae for the 'Flora of Tropical East Africa': I. Kew
Bull. 30: 247-326.
Verdcourt, B. 1976. In R. M. Polhill (editor), Flora of Tropical East Africa. Rubiaceae, Part 1: 1^14.
London: Crown Agents for Overseas Governments. [Lectotypifications for Oldenlandia corymbosa L.
on p. 308 and O. lancifolia on p. 292.]
Verdcourt, B. 1983. Notes on Mascarene Rubiaceae. Kew Bull. 37: 521-547. [Spermacoce typifications.]
M > \1 F \ ( LATURALINDEX

Aeginetia Cav., Anal. Ci. Nat. 3: 129. 1801. Alibertia premontana C. M. Taylor, Novon 6:
Type not designated. 215. 1996. Type: COSTA RICA: Alajuela,
Aeginetia longiflora Cav., Anal. Ci. Nat. 3: 130. Reserva Biologica Monteverde, Valle del rio
1801. Type: MEXICO: Between Queretaro Penas Blancas, fila del Toro, serdero Pipilacha
and Guanajuato, Oct, Nee (Holotype y Campo Tres, 18°N, 84°W, 900-1100 m, 21
presumably MA, n.v.).—Bouvardia Jan 1991, W. Haber & E. Cruz 10630
longiflora (Cav.) Kunth (Holotype MO 4234216; Isotype CR, n.v.).
-ieginetici multiflora Cav., Anal. Ci. Nat. 3: 131, Alibertia utleyorum (Dwyer) C. M. Taylor,
t. 28, fig. 2. 1801; Icon. 6: 52, t. 572, fig. 2. Novon 6: 215. 1996. Duroia utleyorum Dwyer
1801. Type: MEXICO: Queretaro, Nee s.n. in W. C. Burger & C. M. Taylor, Fieldiana
(Holotype MA, n.v. (photo MO)). Bot., n.s. 33: 127. 1993. Type: COSTA RICA:
—Bouvardia multiflora (Cav.) Schult. & Puntarenas, adjacent to airfield, Rincon de
Schult. f. Osa, 6-7 Feb 1974, R. Liesner 1774
(Holotype CR, n.v.; Isotype MO 3030387).
Aegiphila Jacq. [Verbenaceae]
K wphila hnjjmumuoides Standi. & Steyerm., s Standi, Ann. Missouri Bot. Gard.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 227. 27: 344. 1940. Type: Allenanthus
1947. Type: GUATEMALA: Huehuetenango, erythrocarpa Standi.
Steyermark 48843 (Holotype F, n.v.). Allenanthus erythrocarpus Standi., Ann.
—Hoffmannia nicotianifolia (M. Martens & Missouri Bot. Gard. 27: 344. 1940. Type:
Galeotti) L. O. Williams PANAMA: Code, Vicinity of El Valle de
Anton, ca. 600 m, 17 Sep 1939, P. H. Allen
Alibertia A. Rich., Mem. Soc. Hist. Nat. Paris 5: 1999 (Holotype MO 1195153 (photos MEXU,
234, t. 21, fig. 1. 1830. Type: Alibertia edulis PTBG); Isotypes F 1015649, GH, n.v., US
(Rich.) A. Rich, ex DC. (photos MEXU, PTBG)).
Alibertia edulis (Rich.) A. Rich, in DC, Prodr. Allenanthus hondurensis Standi., Ceiba 1: 45.
4: 443. 1830. Genipa edulis Rich., Act. Soc. 1950. Type: HONDURAS: Morazan, Faldas
Hist. Nat. Paris 1: 107. 1792. Type: FRENCH del Cerro Majicaran, aguas arriba del Rio de
GUIANA: Guyane, Richard s.n. (Holotype P, la Onlla, 950 m, 27 Nov 1948, A. Molina R.
n.v.; Isotype F 971706 (photo PTBG)). 1750 (Holotype F 152284 (photo PTBG);
Cordiera, Gardenia. Isotypes EAP, n.v., US (photos MEXU,
Alibertia garapatica K. Schum. in Mart., Fl. PTBG)).
Bras. 6(6): 384. 1889. Type not cited. Allenanthus hondurensis var. parvifolia L. O.
Alibertia longistiptdata L. Riley, Kew Bull. Williams, Phytologia 25: 461. 1973. Type:
1927: 122. 1927. Type: PANAMA: Panama, MEXICO: Chiapas, Mun. Venustiano
Old Panama, 16 Jun 1924, L. A. M. Riley 145 Carranza, Rancho Carmen along the road
(Holotype K, n.v.; Isotype US (photos from Acala to Venustiano Carranza, 2600 ft.,
MEXU, PTBG)).—Alibertia edulis (Rich.) 23 Aug 1966, R. M. Lauglin 1647 (Holotype F
A. Rich ex DC. 1662766 (photos MEXU, PTBG)).
Alibertia panamensis L. Riley, Kew Bull. 1927:
123. 1927. Type: PANAMA: Between Alseis Schott in Spreng., Syst. Veg. 4(2): 404.
Panama and Sabana, 14 Jun 1924, L. A. M. 1827. Type: Alseis floribunda Schott
Riley 111 (Holotype K, n.v.; Isotypes US, 2 Alseis blackiana Hemsl., Diag. PI. Mex. 30.
sheets (photos MEXU, PTBG)).—Alibertia 1879. Type: PANAMA: In dense woods,
edulis (Rich.) A. Rich, ex DC. Manma [?] Station, P. R. R., Jun 1862, S.
 L NOMENCLATOR OF MEXICAN / ) CENTRAL AMERICAN RUBIACEAE

Hayes 665 (Holotype K, n.v.; Isotype P Agricola and Alto Piedra (above Santa Fe)
(photos MEXU, PTBG)). and Rio Dos Bocas ca. 5-8 km from Escuela,
Alseis dariensis Dwyer, Ann. Missouri Bot. 730-770 m, 26 Jul 1974, T. B. Croat 25937
Gard. 67: 27. 1980. Type: PANAMA: Darien, (Holotype MO 2212775 (photo PTBG);
Rio Tuquesa, middle Tuquesa Mining Co. Isotypes F (photo PTBG), GH, n.v., MO
camp called Charco Peje, 250 m, S. Mori 2212776 (photo PTBG)).
7001 (Holotype MO, not located 6/96).
—Wittmackanthus stanleyanus (Schomb.) Amphidaysa Standi., Field Mus. Nat. Hist., Bot.
Ser. 11: 180. 1936. Type: Amphidaysa
Alseis hondurensis Standi., Trop. Woods 16: ambigua (Standi.) Standi.
48. 1928. Type: HONDURAS: La Ceiba, Dec Amphidaysa ambigua (Standi.) Standi., Publ.
1927, H. Johansen 9 (Holotype F 572615 Field Columbian Mus., Bot. Ser. 11: 181.
(photo PTBG)). 1936. Sabicea ambigua Standi., Publ. Field.
Alseis schippii Lundell, Lloydia 4: 56. 1941. Mus. Nat. Hist., Bot. Ser. 7: 49. 1930. Type:
Type: BRITISH HONDURAS (BELIZE): COLOMBIA: Without precise locality, 1847,
Toledo Distr., Machaca, 50 ft., 4 Nov 1932, Triana s.n. (Holotype K, n.v. (photo F)).
W. A. Schipp 1230 (Holotype TEX-LL Amphidaysa spathulata Dwyer, Ann. Missouri
(photos MEXU, PTBG); Isotype F (photo Bot. Gard. 67: 33. 1980. Type: PANAMA:
PTBG)).—Alseis hondurensis Standi. Darien, Cerro Campamento, south of Cerro
Alseis yucatanensis Standi., Publ. Field Pire, cloud forest, 20-27 Mar 1968, J. A.
Columbian Mus., Bot. Ser. 8: 50. 1930. Type: Duke 15615 (Holotype MO 1964213 (photos
MEXICO: Yucatan, without precise locality, MEXU, PTBG)).
1917-1921, G. F. Gaumer 24247 (Holotype F
552253 (photo PTBG); Isotype US (photos Anisomeris C. Presl, Symb. Bot. 2: 5, t. 54.
MEXU, PTBG)). 1834. Type: Anisomeris spinosa C. Presl.
[Herein treated as a synonym of Chomelia
Amaioua Aubl., Hist. PI. Guiane 13: 1775. Jacq.]
Type: Amaioua guianensis Aubl. Anisomeris brachypoda (Donn. Sm.) Standi.,
Amaioua corymbosa Kunth, Nov. Gen. Sp. 3: Publ. Field Columbian Mus., Bot. Ser. 4: 293.
419, pi. 294. 1820. Type: VENEZUELA: In 1929. Chomelia brachypoda Donn. Sm., Bot.
locis frondosis Novae-Andalusia prope Gaz. (Crawfordsville) 47: 255. 1909. Type:
Cumanacoa, Humboldt & Bonpland s.n. GUATEMALA: Alta Verapaz, Sasis, banks of
(Holotype presumably P-Bonpland, n.v.; Rio Ogeqaj, 900 m, May 1908, H von
Isotype B-W (photo MO)). Tuerckheim 2253 (Holotype US; Isotypes F
Amaioua hirsuta Poepp. & Endl., Nov. Gen. Sp. (photos MEXU, PTBG), US).—Chomelia
PI. 3: 25, pi. 230 1841. Type: Chili, Perou, et brachypoda Donn. Sm.
Fleuve de La Plata, 1836, Poeppig 2823 Anisomeris englesingii (Standi.) Standi., Publ.
(Holotype W, n.v., fragment F; Isotype F Field Columbian Mus., Bot. Ser. 8: 360. 1931.
(photo PTBG)).—Duroia hirsuta (Poepp. & Chomelia englesingii Standi., Trop. Woods
Endl.) K. Schum. 16: 45. 1928. Type: NICARAGUA: Region of
Amaioua magnicarpa Dwyer, Ann. Missouri Braggman's Bluff (Kukalaya River), 200-600
Bot. Gard. 67: 29. 1980. Type: PANAMA: ft., 2 Dec 1927, F. C Englesing 49 (Holotype
Colon, Santa Rita Ridge, 23 Mar 1972, J. D. F 572606 (photo PTBG)).—Chomelia
Dwyer 9552 (Holotype MO 2162079 (photo recordii Jacq.
PTBG); Isotype MO 4199142 (photo PTBG)). Anisomeris microloba (Donn. Sm.) Standi.,
Amaioua pedicellata Dwyer, Ann. Missouri Publ. Field Columbian Mus., Bot. Ser. 4: 293.
Bot. Gard. 67: 30. 1980. Type: PANAMA: 1929. Chomelia microloba Donn. Sm., Bot.
Veraguas, along road between Escuela Gaz. (Crawfordsville) 31: 114. 1901. Type:
 COSTA RICA: Forets de Santo Domingo de
Golfo Dulce, Feb 1896, A. Tonduz 9874
(Holotype US; Isotypes F, MO (photos Standley & J. Valerio 49196 (Holotype US
MEXU, PTBG), P (photo MEXU), US, 3 1305902).—Chione sylvicola (Standi.) W. C.
sheets).—Chomelia microloba Donn. Sm.
Anisomeris pringlei (S. Watson) Standi., Publ.
Field Columbian Mus., Bot. Ser. 4: 293. 1929. Anotis DC, Prodr. 4: 431. 1830. Type not
Chomelia pringlei S. Watson, Proc. Amer. designated.
Acad. Sci. 26: 137. 1891. Type: MEXICO: Anotis cervantesii (Kunth) DC, Prodr. 4: 432.
San Luis Potosi, Tamasopo Canyon, 6 Aug 1830. Hedyotis cervantesii Kunth in Humb.,
1890, C. G. Pringle 3209 (Holotype US Bonpl. & Kunth, Nov. Gen. Sp. 3: 390. 1820.
48535; Isotypes F (photos MEXU, PTBG), Type: MEXICO: Without precise locality, A.
MO (photos MEXU, PTBG), P (photos Bonplands.n. (Holotype P-Bonpland, n.v.;
MEXU, PTBG)).—Chomelia pringlei S. Isotype B-W (photo MO)).—Hedyotis
Watson cervantesii Kunth
Anisomerisprotracta (Bartl. ex DC.) Standi., Anotis longiflora Benth., PI. Hartw. 23. 1839.
Publ. Field Columbian Mus., Bot. Ser. 4: 293. Type: MEXICO: Without precise locality,
1920. Guettarda protracta Bartl. ex DC, 1837, T. K. Hartweg 206 (Holotype K, n.v.;
Prodr. 4: 457. 1830. Type: MEXICO: Without Isotype GH, n.v.). Bouvardia.—Bouvardia
precise locality, Haenkes.n. (Holotype multiflora (Cav.) Schult. & Schult. f.
presumably PR, n.v.).—Chomelia protracta
(Bartl. ex DC.) Standi. Antirhea Comm. ex Juss., Gen. PL 204. 1789.
Anisomeris purpusii Brandegee, Univ. Calif. Type: Antirhea borbonica J. F. Gmelin
Publ. Bot. 6: 70. 1914. Type: MEXICO: Antirhea aromatica Cast-Campos & Lorence,
Chiapas, nearTonala, Sep 1913, C. A. Purpus Ann. Missouri Bot. Gard. 72: 268 Jig. I.
6940 (Holotype UC 173028, n.v.; Isotypes F 1985. Type: MEXICO: Veracruz, Mun.
(photo PTBG), US (photo PTBG)). Chomelia. Jalcomulco, ca. 10 km al S de Apazapan,
—Chomelia spinosa Jacq. Barranca de Monterrey, al S de Cuetzalan,
Anisomeris recordii (Standi.) Standi., N. Amer. entre Cuetzalan y Apazapan, 350 m, 17 Sep
Fl. 32: 227. 1934. Chomelia recordii Standi.. 1983, G. Castillo-Campos 2957 (Holotype
Trop. Woods 7: 9. 1926. Type: XAL; Isotypes MEXU, MO, XAL, plus about
GUATEMALA: Izabal, high forest between 50 more distributed to a wide range of
Los Andes and Entre Rios, lower Rio herbaria).
Motagua Valley, Record 8862 (Holotype Y, Antirhea lucida (Sw.) Benth. & Hook, f., Gen.
n.v.; Isotype NY, n.v.).—Chomelia recordii PI. 2: 100. 1873. Laugeria lucida Sw., Prodr.
Standi. Veg. Ind. Occ. 48. 1788. Type: JAMAICA:
Anisomeris rudis Standi., Field Mus. Nat. Hist., Without precise locality, Swartz s.n.
Bot. Ser. 17: 394. 1938. Type: HONDURAS: (Holotype presumably S, n.v.).
Comayagua, summit of ridge above El —Stenostomum lucidum (Sw.) C. F. Gaertn.
Achote, above the plains of Siguatepeque, Antirhea panamensis Standi., N. Amer. Fl. 32:
1800 m, 1 Aug 1936, T. G. Yunker, R. F. ' 264. 1934. Type: PANAMA: San Bias
Dawson & H. R. Youse 6253 (Holotype F District, 21-28 Apr 1933, G. P. Cooper 238
857678 (photo PTBG); Isotype MO (Holotype F 686345 (photo PTBG); Isotype F
1115471).—Chomelia rudis (Standi.) 709955). Chomelia.—Chomelia microloba
Donn. Sm.
Anisomeris sylvicola (Standi.) Standi., N. Amer. Antirhea trichantha (Griseb.) Hemsl., Biol.
Fl. 32: 225. 1934. Chomelia ? sylvicola Centr.-Amer., Bot. 2: 42. 1881. Pittoniotis
Standi., J. Wash. Acad. Sci. 18: 182. 1928. trichantha Griseb., Bonplandia 6: 8. 1858.
A NOMENCLATOR OF MEXICAN AND CENTRAL

Type: PANAMA: Without precise locality, m, 14 Jan 1905, W. R. Maxon & R. Hay 3275
Duchaissing s.n. (Holotype presumably (Holotype US 473244 (photos MEXU,
GOET or P, n.v.).—Pittoniotis trichantha PTBG); Isotype US (photos MEXU, PTBG)).
Griseb. —Rondeletia aetheocalymna Donn. Sm.
Arachnothryx albida (Lundell) Borhidi, Acta
Appunia Hook. f. in Benth. & Hook, f., Gen. PI. Bot. Hung. 35: 309. 1989. Rondeletia albida
2: 120. 1873. Type: Appunia tenuiflora Lundell, Wrightia 5: 323. 1976. Type:
(Benth.) Hook. f. MEXICO: Chiapas, Carelas, near Motozintla,
Appunia guatemalensis Donn. Sm., Bot. Gaz. 2176 m, Apr 1945, E. Matuda 5509 (Holotype
(Crawfordsville) 48: 294. 1909. Type: TEX-LL (photos MEXU, PTBG); Isotypes F,
GUATEMALA: Izabal, silva prope pagam MEXU, TEX-LL).—Rondeletia laniflora
martimam Livingston, Jun 1906, H. von Benth.
Tuerckheim IIf-230 (Holotype US 943501 Arachnothryx aspera (Standi.) Borhidi, Acta
(photos MEXU, PTBG)). Morinda. Bot. Acad. Sci. Hung. 28: 68. 1982.
Appunia parviflora Lundell, Contr. Univ. Rondeletia aspera Standi., N. Amer. Fl. 32:
Michigan Herb. 4: 28. 1940. Type: BRITISH 54. 1918. Type: COSTA RICA: San Jose,
HONDURAS (BELIZE): Stann Creek Distr.. Pont du Rodeo, Pacacol (Villa Colon), Jan
Silk Grass Creek Reserve, 5 Sep 1939, P. H. 1891, H. Pittier 3243 (Holotype US 941408
Gentle 2986 (Holotype MICH, n.v.; Isotypes (photo PTBG)).—Rondeletia aspera Standi.
A, n.v., F (photo PTBG), US (photos MEXU, Arachnothryx belizensis (Standi.) Borhidi, Acta
PTBG)).—Psychotria erecta (Alibi.) Standi. Bot. Acad. Sci. Hung. 28: 70. 1982.
& Steyerm. Rondeletia belizensis Standi., Carnegie Inst.
Appunia seibertii Standi., Ann. Missouri Bot. Wash. Publ. 461: 91. 1935. Type: BRITISH
Gard. 24: 208. 1937. Type: PANAMA: HONDURAS (BELIZE): Jasinto Hills, 300
r Rio ft., 28 Aug 1933, W. A. Schipp 1201
Trinidad, 26-50 m, 7-10 Aug 1935, R. J. (Holotype F 683600 (photos MEXU, PTBG);
Seibert 624 (Holotype MO 1110146 (photos Isotypes GH, n.v., K (photo PTBG), MO).
MEXU, PTBG); Isotypes A, n.v., F). —Rondeletia belizensis Standi.
Arachnothryx belizensis (Standi.) Borhidi var.
Arachnothryx Planch., Fl. Serres 5: tab. 442. longiloba (Lundell) Borhidi, Acta Bot. Hung.
1849. Type: Arachnothryx leucophylla 35: 310. 1989. Rondeletia belizensis Standi,
(Kunth) Planch. Type designated by Borhidi, var. longiloba Lundell, Wrightia 5: 323. 1976.
1982. [Herein treated as a synonym of Type: GUATEMALA: Alta Verapaz,
Rondeletia L.] Rubelsanto, 2.3 km ESE of Balastrera, 20 Jul
Arachnothryx acuminata (Oerst. ex Standi.) 1985, C. L. Lundell & E. Contreras 19545
Borhidi, Acta Bot. Hung. 35: 310. 1989 [as (Holotype TEX-LL (photos MEXU, PTBG);
acumunata]. Sommera acuminata Oerst. ex Isotypes F, MO (photo PTBG)).—Rondeletia
Standi., J. Wash. Acad. Sci. 17: 340. 1927. belizensis Standi.
Type: MEXICO: Oaxaca, Amatlan, F. Arachnothryx bertieroides (Standi.) Borhidi,
Liebmann 11712 (Holotype C (photo MEXU); Acta Bot. Acad. Sci. Hung. 28: 70. 1982.
Isotype US (photos MEXU, PTBG)). Rondeletia bertieroides Standi., Publ. Field
Rondeletia.—Rondeletia scabra Hemsl. Columbian Mus., Bot. Ser. 4: 267. 1929.
Arachnothryx aetheocalymna (Donn. Sm.) Type: PANAMA: Bocas del Toro, Buena
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 70. Vista Camp, Chinqui Trail, 900 m, 1928, G
1982. Rondeletia aetheocalymna Donn. Sm., P. Cooper 598 (Yale 12231) (Holotype F
Bot. Gaz. (Crawfordsville) 42: 298. 1906. 579614 (photo PTBG); Isotype NY, n.v.).
Type: GUATEMALA: Alta Verapaz, trail i bertieroides Standi.
between Sepacuite and Secanquim, 550-1000
Arachnothryx bourgaei (Standi.) Borhidi, Acta Arachnothryx calycosa (Donn. Sm.) Borhidi,
Bot. Acad. Sci. Hung. 28: 69. 1982. Acta Bot. Acad. Sci. Hung. 28: 70. 1982.
Rondeletia bourgaei Standi., N. Amer. Fl. 32: Rondeletia calycosa Donn. Sm., Bot. Gaz.
56. 1918. Type: MEXICO: Veracruz, region (Crawfordsville) 56: 59. 1913. Type: COSTA
de Orizaba, Monte San Cristobal, 8 Jul 1866, RICA: Forets de la Palma, 1460 m, 25 Sep
E. Bourgeau 2618 (Holotype US 49964 1898, A. Tonduz 11633 (Holotype US 941406
(photos MEXU, PTBG); Isotypes P, 4 sheets (photos F, MEXU, PTBG)).—Rondeletia
(photos MEXU, PTBG)).—Rondeletia calycosa Donn. Sm.
laniflora Benth. Arachnothryx capitellata (Hemsl.) Borhidi, Acta
| Standi.) Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
Bot. Hung. 33: 301. 1987. Rondeletia brenesii Rondeletia capitellata Hemsl., Diag. PI. Mex.
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. 28. 1879. Type: MEXICO: Without precise
18: 1370. 1938. Type: COSTA RICA: locality, Jurgensen 355 (Lectotype K, here
Camino de San Ramon a La Calera, 20 Mar designated (photo PTBG); Isolectotypes G-
1929, A. M. Brenes 6757 (Holotype F 854972 DEL (n.v., fragment F), K (photo PTBG)).
(photo PTBG)).—Rondeletia brenesii Standi. Syntypes: MEXICO: Oaxaca, Joya, F.
Arachnothryx brevicorolla (J. H. Kirkbr.) Liebmann 202 (K (photo PTBG); without
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69. precise locality, Jurgensen 3 76 (K (photo
1982 [as brt alk ifolia PTBG)).—Rondeletia capitellata Hemsl.
Dwyer & Hayden subsp. brevicorolla J. H. Arachnothryx chinajensis (Standi. & Steyerm.)
Kirkbr., Ann. Missouri Bot. Gard. 55: 383. Borhidi, Acta Bot. Hung. 33: 301. 1987.
1963. Type: PANAMA: Bocas del Toro, Rondeletia chinajensis Standi. & Steyerm.,
cloud forest above Quebrada Huron on Cerro Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.
Bonyik, 166^00 m, 13 Apr 1968, J. H. 1947. Type: GUATEMALA: Alta Verapaz,
Kirkbride, Jr. &J.A. Duke 597 (Holotype Cerro Chinaja, between Finca Yalpemech and
MO 1946832 (photo PTBG); Isotypes NY, Chinaja, above source of Rio San Diego,
n.v., US (photos MEXU, PTBG)). 150-700 m, 1-2 Apr 1942, J. A. Steyermark
—Rondeletia bertieroides Standi. 45637 (Holotype F 1134210 (photos MEXU,
Arachnothryx buddleioides (Benth.) Planch., Fl. PTBG)).—Rondeletia chinajensis Standi. &
Serres 5: tab. 442. 1849. Rondeletia Steyerm.
buddleioides Benth., PI. Hartweg. 69. 1840. Arachnothryx cooperi (Standi.) Borhidi, Acta
Type: MEXICO: Oaxaca, in planitie Llano Bot. Acad. Sci. Hung. 28: 69. 1982.
Verde, T. K. Hartweg 502 (Holotype K (photo Rondeletia cooperi Standi., Publ. Field Mus.
PTBG); Isotypes NY, n.v. (fragment US Nat. Hist., Bot. Ser. 4: 267. 1929. Type:
(photos MEXU, PTBG), P (photos MEXU, PANAMA: Bocas del Toro, Buena Vista
PTBG)).—Rondeletia buddleioides Benth. Camp, on Chiriqui Trail, 1250 ft., Jan-Mar
Arachnothryx buddleioides (Benth.) Planch, var. 1928, G. P. Cooper 600 (Yale 12233)
aspera (J. H. Kirkbr.) Borhidi, Acta Bot. (Holotype F 579596 (photo PTBG); Isotype
Hung. 33: 301. 1987. Rondeletia buddleioides NY, n.v.).—Rondeletia cooperi Standi.
Benth. var. aspera J. H. Kirkbr., Ann. Arachnothryx cordovana (Standi. & Steyerm.)
Missouri Bot. Gard. 55: 379. 1969. Type: Borhidi, Acta Bot. Hung. 33: 301. 1987.
PANAMA: Bocas del Toro, between Criollo Rondeletia cordovana Standi. & Steyerm.,
(just above Buena Vista) and Quebrada Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 26.
Higueron on Chiriqui Trail, 18 Apr 1968, 1943. Type: GUATEMALA: Zacapa, in
J. H. Kirkbride, Jr. & J. A. Duke 780 ravine bordering Quebrada Alejandria,
(Holotype MO 1942973 (photos MEXU, summit of Sierra de las Minas, vicinity of
PTBG)).—Rondeletia buddleioides Benth. Finca Alejandria, 2500 m, 13 Oct 1939, J. A.
Steyermark 29906 (Holotype F 1042952
A NOMENCLATOR OF MEXICAN AND CENTRAL

TY elongata (Bartl. ex DC.) Planch.,

Fl. Serres 5: tab. 442. 1849. Rondeletia
elongata Bartl. ex DC, Prodr. 4: 409. 1830.
I | Borhidi, Type: MEXICO: Guerrero, Haenke s.n.
Acta Bot. Acad. Sci. Hung. 28: 70. 1982. (Holotype presumably PR, n.v.; Isotype W,
Rondeletia costaricensis Standi., N. Amer. Fl. n.v. (photo MO)).—Rondeletia leucophylla
32: 61. 1918. Type: COSTA RICA: Buena Kunth
Vista, San Carlos, 900 m, Apr 1903. H. Pittier Inn hnothryx falciformis (Lundell) Borhidi,
16696 (Holotype US 764141 (photos MEXU, Acta Bot. Hung. 35: 310. 1989. Rondeletia
PTBG)).—Rondeletia costaricensis Standi. falciformis Lundell, Wnghtia 5: 324. 1976.
Arachnothyx darcyi (Dwyer) Borhidi, Acta Type: MEXICO: Hidalgo, km 332 of Mexico
Bot. Acad. Sci. Hung. 28: 70. 1982. City-Laredo highway, above Chapulhuacan,
Rondeletia darcyi Dwyer, Ann. Missouri Bot. 3800 ft., 13 Jul 1943, C L Lundell 12234
Card. 67: 467. {980. Type: PANAMA: Colon, (Holotype TEX-LL (photos MEXU, PTBG);
road bank just W of Portobelo, 12 Apr 1970, Isotype F (photo PTBG)).—Rondeletia
W. G. D 'Any 4079 (Holotype MO 2238438
(photo PTBG); Isotype MO (photo PTBG)). Arachnothryx galeottii (Standi.) Borhidi, Acta
—Rondeletia secunda Standi. Bot. Acad. Sci. Hung. 28: 70. 1982.
Arachnothryx dariensis (Standi.) Borhidi, Acta Rondeletia galeottii Standi., N. Amer. Fl. 32:
Bot. Acad. Sci. Hung. 28: 68. 1982. 59. 1918. Type: MEXICO: Without precise
Rondeletia dariensis Standi., N. Amer. Fl. 32: locality, H. Galeotti s.n. (Holotype US
53. 1918. Type: PANAMA: Darien, Boca de 454830 (photos MEXU, PTBG)).
Paurando, on Sambii River, southern Darien, —Rondeletia galeottii Standi.
20 m, Feb 1912,//. Pittier 5684 (Holotype US ;
•....: ;

867136 (photos MEXU, PTBG); Isotype F Acta Bot. Hung. 33: 302. 1987 [as
(photo PTBG)).—Rondeletia dariensis gonzalaeoides]. Rondeletia gonzaleoides
Standi. Standi., Publ. Field Columbian Mus., Bot. Ser.
Arachnothryx deamii (Donn. Sm.) Borhidi, Acta 8: 160. 1930. Type: MEXICO: Chiapas,
Bot. Acad. Sci. Hung. 28: 68. 1982. Pueblo Nuevo, 2200 m, 1840, J. Linden 149
Bouvardia deamii Donn. Sm., Bot. Gaz. (Holotype LE, n.v. (fragment F, photos
(Crawfordsville) 49: 455. 1910. Type: MEXU, PTBG); probable Isotype P (photos
GUATEMALA: At base of Rillo de las MEXU, PTBG)).—Rondeletia £
Nabajas, about V* mile front side of Fiscal,
3700 ft., 5 Jun 1909, C C. Deam 6190 Arachnothryx gracilis (Hemsl.) Borhidi, Acta
(Holotype US 579579 (photos MEXU, Bot. Acad. Sci. Hung. 28: 69. 1982.
PTBG); Isotypes A, n.v., F (photo PTBG), Rondeletia gracilis Hemsl., Diag. PI. Nov.
GH, n.v., MO (photos MEXU, PTBG), US Mexic. 53. 1880. Type: GUATEMALA: Alta
(photos MEXU, PTBG)).—Rondeletia Verapaz, 3600 ft., H. von Tuerckheim 404
deamii (Donn. Sm.) Standi. (Holotype K (photo PTBG)).—Rondeletia
Arachnothryx edwardsii (Standi.) Borhidi, Acta gracilis Hemsl.
Bot. Hung. 33: 301. 1987. Rondeletia oica (Standi.) Borhidi,
edwardsii Standi., Trop. Woods 37: 31. 1934. Acta Bot. Hung. 33: 302. 1987. Rondeletia
Type: HONDURAS: Comayagua, Pito Solo, gracilispica Standi., Publ. Field Mus. Nat.
Lake Yojoa, 2000 ft, 29 Aug 1932,/ B. Hist., Bot. Ser. 8: 342. 1931. Type: MEXICO:
Edwards P476 (Holotype F 662645 (photos Oaxaca, Sierra San Pedro Nolasco, Talea, etc.,
MEXU, PTBG); Isotype A, n.v.). 1843-33, C. Jurgensen 935 (Holotype K
i Standi. (photo PTBG; fragment F, photos MEXU,
PTBG))—Rondeletia gracilispica Standi.
 'Ha (Dwyer & Hay den) Isotypes B (presumably destroyed, photo
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 70. MO), P (photos MEXU, PTBG)).
1982. Rondeletia hameliifolia Dwyer & —Rondeletia laniflora Benth.
Hayden, Ann. Missouri Bot. Gard. 54: 144. Arachnothryx leptodictya (B. L. Rob.) Borhidi,
1967. Type: PANAMA: Veraguas, Santiago, Acta Bot. Acad. Sci. Hung. 28: 68. 1982.
Aug 1961, J. D. Dwyer 1350 (Holotype MO Rondeletia leptodictya B. L. Rob., Proc.
1824657 (photo PTBG); Isotype NY, n.v.). Amer. Acad. Sci. 45: 402. 1910. Type:
—Rondeletia hameliifolia Dwyer & Hayden MEXICO: [Boundary between Michoacan
Arachnothyx heteranthem (Brandegee) and Guerrero] Rives du Rio Petatlan, 500 m,
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69. 24 Nov 1898, E. Langlasse 666 (Holotype
1982. Rondeletia heteranthera Brandegee, GH, n.v.; Isotypes F (photos MEXU, PTBG),
Univ. Calif. Publ. Bot. 4: 387. 1913. Type: G, n.v., P (2 sheets, photos MEXU, PTBG),
MEXICO: Veracruz, Banos del Carrizal, Aug US (photos MEXU, PTBG)).—Rondeletia
1912, C. A. Purpus 6148 (Holotype UC leucophylla Kunth
155217, n.v.; Isotypes F, MO).—Rondeletia IT leucophylla (Kunth) Planch., Fl.
heteranthera Brandegee Serres 5: tab. 442. 1849. Rondeletia
Irachnothryx izabalensis (Standi. & Steyerm.) leucophylla Kunth in Humb., Bonpl. & Kunth,
Borhidi, Acta Bot. Hung. 33: 302. 1987. Nov. Gen. Sp. 3: 395. t. 290. 1820. Type:
Rondeletia izabalensis Standi. & Steyerm., MEXICO: Inter Alto del Peregrino et Flumen
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 286. Papagallo, Humboldt & Bonpland s.n.
1940. Type: GUATEMALA: Izabal, Escoba, (Holotype P-Bonpland, n.v.; Isotypes B, n.v.,
across the bay (west) from Puerto Barrios, P (photos MEXU, PTBG)).—Rondeletia
near sea level, 3 May 1939, P. C. Standlex leucophylla Kunth
72891 (Holotype F 991611 (photos MEXU, Arachnothyx licbmannii (Standi.) Borhidi,
PTBG)).—Rondeletia izabalensis Standi. & Acta Bot. Hung. 33: 302. 1987. Rondeletia
liebmannii Standi., J. Wash. Acad. Sci. 17:
Arachnothryxjurgensenii (Hemsl.) Borhidi, 336. 1927. Type: MEXICO: Oaxaca, without
Acta Bot. Acad. Sci. Hung. 28: 70. 1982. precise locality, 1841-43, F. Liebmann 11S34
Rondeletia jurgensenii Hemsl., Diag. PI. Nov (Holotype US 1315231 (photos MEXU,
Mexic. 29. 1879. Type: MEXICO: Oaxaca, PTBG); probable Isotype P (photos MEXU,
Sierra de San Pedro Nolasco, Jurgensen 248 PTBG)).—Rondeletia capitellata Hemsl.
(Holotype K (photo PTBG)).—Rondeletia Arachnothryx linguiformis (Hemsl.) Borhidi,
jurgensenii Hemsl. Acta Bot. Acad. Sci. Hung. 28: 70. 1982.
Arachnothryx kirkbridei (Dwyer) Borhidi, Acta Rondeletia linguiformis Hemsl, Diag. PI.
Bot. Acad. Sci. Hung. 28: 70. 1982. Nov. Mexic. 29. 1879. Type: GUATEMALA:
Rondeletia kirkbridei Dwyer, Ann. Missouri Without precise locality, Skinner s.n.
Bot. Gard. 67: 469. 1980. Type: PANAMA: (Holotype K (photo PTBG)).—Rondeletia
Veraguas, 16 km NW of Santa Fe on road to linguiformis Hemsl.
Calovebora (Panama Hwy. 35), 300-500 m Arachnothryx longipetiolata (Lundell) Borhidi,
16 May 1975, S. Mori & J. Kallunki 6112 Acta Bot. Hung. 35: 310. 1989. Rondeletia
(Holotype MO 2300094 (photo PTBG)). longipetolata Lundell, Wrightia 5: 325. 1976.
—Rondeletia bertieroides Standi. Type: MEXICO: Chiapas, Mt. Ovando, near
Arachnothryx laniflora (Benth.) Planch., Fl. Escuintla, 14 Nov 1945, E. Matuda 6146
Serres 5: tab. 442. 1849. Rondeletia laniflora (Holotype TEX-LL; Isotypes MEXU, TEX-
Benth., PI. Hartw. 85. 1841. Type: LL).—Rondeletia buddleioides Benth.
GUATEMALA: In montibus las Nubes, T. Arachnothryx lundelliana (Standi.) Borhidi,
Hartweg 584 (Holotype K (photo PTBG), Acta Bot. Hung. 33: 302. 1987. Rondeletia
fragment US (photos MEXU, PTBG); lundelliana Standi., Contr. Univ. Mich. Herb.
A NOMENCLATOR OF MEXICAN AND CEN

4: 31. 1940. Type: BRITISH HONDURAS Rondeletia myriantha Standi. & Steyerm. var.
(BELIZE): El Cayo Distr., Vaca, 21 Apr armentalis L. O. Williams, Phytologia 26:
1938, P. H. Gentle 2504 (Holotype F 941520 127. 1973. Type: GUATEMALA: Quiche,
(photos MEXU, PTBG); Isotypes A, n.v., Nebaj, 5600 ft., 22 Nov 1934, A. F Skutch
TEX-LL ex MICH (photos MEXU, PTBG), 1776 (Holotype F 815811 (photos MEXU,
US (photos MEXU, PTBG)).—Rondeletia PTBG)).~Rondeletia myriantha Standi. &
stenosiphon Hemsl. Steyerm. var. armentalis L. O. Williams
Arachnothryx macrocalyx (Standi. & Steyerm.) Arachnothryx nebulosa (Standi.) Borhidi, Acta
Borhidi, Acta Bot. Hung. 33: 302. 1987. Bot. Hung. 33: 302. 1987. Rondeletia
Rondeletia macrocalyx Standi. & Steyerm., nebulosa Standi., Trop. Woods 37: 32. 1934.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254. Type: HONDURAS: Tegucigalpa, La Aurora
1947. Type: GUATEMALA: Izabal, Cerro Trail, Rosario, San Juancito, 5400 ft., 2 Apr
San Gil, 300-900 m, 25 Dec 1941, J. A. 1932, J. B. Edwards P10 (Holotype F 662646
Steyermark 41864 (Holotype F 1134224 (photos MEXU, PTBG); Isotype A, n.v.).
(photos MEXU, PTBG); Isotype F (photos —Rondeletia nebulosa Standi.
MEXU, PTBG)).—Rondeletia macrocalyx Arachnothryx nicaraguensis (Oerst.) Borhidi,
Standi. & Steyerm. Acta Bot. Hung. 35: 310. 1989. Rondeletia
Arachnothryx mexiae (Standi.) Borhidi, Acta nicaraguensis Oerst., Vidensk. Meddel.
Bot. Hung. 33: 302. 1987. Rondeletia mexiae Dansk Naturhist. Foren. Kjobenhavn 1852:
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. 43. 1853. Type: NICARAGUA: Matagalpa,
22: 55. 1940. Type: MEXICO: Oaxaca, Distr. Segovia, in monte Pantasmo, Jan 1848,
Choapan, Santa Maria, Montana Santa Maria, Oersted s.n. (Holotype C (photos F, MO, US),
1500 m, 1 Apr 1938, Y. Mexia 9261 fragment US).—Rondeletia nicaraguensis
(Holotype F 982856 (photos MEXU, PTBG); Oerst.
Isotypes GH, n.v., MO (photos MEXU, Arachnothryx nitida (Hemsl.) Borhidi, Acta Bot.
PTBG), US (photos MEXU, PTBG)). Acad. Sci. Hung. 28: 69. 1982. Rondeletia
—Rondeletia scabra Hemsl. nitida Hemsl., Diag. PI. Nov. Mexic. 29.
Arachnothryx minor (Lundell) Borhidi, Acta 1879. Type: MEXICO: Chiapas, Fl. rosatres,
Bot. Hung. 35: 310. 1989. Rondeletia minor forets des pins, Chiapas, in fevr., Linden 1660
Lundell, Wrightia 5: 326. 1976. Type: (Lectotype K, here designated (photo PTBG);
MEXICO: Veracruz, about 10 miles north of Isolectotypes G-DEL, n.v., fragment F, P
Teziutlan, 6 Jan 1964, D. S. Correll & H. B. (photos MEXU, PTBG)). Syntype: MEXICO:
Correll 28736 (Holotype TEX-LL (photos Chiapas, Ghiesbreght 728 (K (photo PTBG),
MEXU, PTBG); Isotype MEXU). GH, n.v.). Linden 1660 was designated as
—Rondeletia gracilispica Standi. lectotype, as it is the better specimen with
Arachnothryx myriantha (Standi. & Steyerm.) more duplicates.—Rondeletia nitida Hemsl.
Borhidi, Acta Bot. Hung. 33: 302. 1987. Arachnothryx oaxacana (Standi.) Borhidi, Acta
Rondeletia myriantha Standi. & Styerm., Bot. Hung. 33: 302. 1987. Rondeletia
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 288. oaxacana Standi., Field Mus. Nat. Hist., Bot.
1940. Type: GUATEMALA: Suchitepequez, Ser. 11: 255. 1936. Nomen superfl., based on
Finca Moca, 3200 ft., 31 Oct 1934, A. F. same type as Rondeletia mexicana (Turcz.)
Skutch 1569 (Holotype F 813463 (photos Standi. Type: MEXICO: Oaxaca, without
MEXU, PTBG); Isotypes A, n.v., US (photos precise locality, 3000 ft., H. Galeotti 2664
MEXU, PTBG)).—Rondeletia myriantha (Holotype P (photos MEXU, PTBG),
Standi. & Steyerm. fragment F (photos MEXU, PTBG)).
Arachnothryx myriantha (Standi. & Steyerm.) —Rondeletia mexicana (Turcz.) Standi.
Borhidi var. armentalis (L.O. Williams) Arachnothryx ovandensis (Lundell) Borhidi,
Borhidi, Acta Bot. Hung. 35: 310. 1989. Acta Bot. Hung. 35: 310. 1989. Rondeletia
 ovandensis Lundell, Wrightia 5: 326. 1976. PTBG); Isotypes F, MEXU, TEX-LL).
Type: MEXICO: Chiapas, Mt. Ovando, near —Rondeletia pyramidalis Lundell
Escuintla, 24 Oct 1941, E. Matuda 6067 Arachnothryx rekoi (Standi.) Borhidi, Acta Bot.
(Holotype TEX-LL (photos MEXU, PTBG); Acad. Sci. Hung. 28: 69. 1982. Rondeletia
Isotypes F (photo PTBG), MEXU, TEX- rekoi Standi, Proc. Biol. Soc. Wash. 8: 126.
LL).—Rondeletia ovandensis Lundell 1918. Type: MEXICO: Oaxaca, Las Pilas
Arachnothryxpansamalana (Standi.) Borhidi, (Cerro Espino), 400 m, 10 Oct 1917, B. P.
Acta Bot. Acad. Sci. Hung. 28: 69. 1982. Reko 3490 (Holotype US 867145 (photos
Rondeletia pansamalana Standi., N. Amer. Fl. MEXU, PTBG)).—Rondeletia rekoi Standi.
32: 58. 1918. Type: GUATEMALA: Alta Arachnothryx rubens (L. O. Williams) Borhidi,
Verapaz, Pansamala, 3800 ft., Apr 1886, H. Acta Bot. Hung. 35: 311. 1989. Rondeletia
von Tuerckheim 897 (Holotype US 49985 rubens L. O. Williams, Phytologia 26: 128.
(photos MEXU, PTBG); Isotype GH, n.v.). 1973. Type: GUATEMALA: Quiche, Nebaj,
—Rondeletia pansamalana Standi. 6000 ft, 19 Nov 1934, A. F. Skutch 1725
Arachnothryxphyllocephala (Standi. & L. O. (Holotype F 815804 (photos MEXU, PTBG)).
Williams) Borhidi, Acta Bot. Hung. 33: 302. —Rondeletia rubens L. O. Williams
1987. Rondeletia phyllocephala Standi. & Arachnothryx rufescens (B. L. Rob.) Borhidi,
L. O. Williams, Ceiba 3: 63. 1952. Type: Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
HONDURAS: Cortes, faldas hiimedas de Rondeletia rufescens B. L. Rob, Proc. Amer.
Montana Sta. Ana, matorrales hiimedos de Acad. Arts 45: 402. 1910. Type:
Rio Sta. Ana, 210 m, 6 Dec 1950, A. Molina GUATEMALA: Alta Verapaz, Coban, 4800
R. 3608 (Holotype US 2216049 (photos ft. Mar 1881, H. von Tuerckheim 582
MEXU, PTBG); Isotype F (photos MEXU, (Holotype GH, n.v.; Isotypes P (photos
PTBG))—Rondeletia thiemei Donn. Sm. MEXU, PTBG), US, 2 sheets (photos MEXU,
•\ x platysepala (Standi.) Borhidi, PTBG)).—Rondeletia gracilis Hemsl.
Acta Bot. Acad.'Sci. Hung. 28: 70. 1982. Arachnothryx salicifolia (Dwyer & Hay den)
Rondeletia platysepala Standi., Ann. Missouri Borhidi, Acta Bot. Acad. Sci. Hung. 28: 70.
Bot. Gard. 27: 343. 1940. Type: PANAMA: 1982. Rondeletia salicifolia Dwyer &
Code, El Valle, North Rim, 600-1000 m, 14 Hayden, Phytologia 15: 58. 1967. Type:
May 1939, P. H. Allen 1791 (Holotype F PANAMA: Panama, Cerro Jefe in "Clusia
989484 (photo PTBG); Isotypes F (photos forest" at 2700-3000 ft, 27 Jan 1966, E. L.
MEXU, PTBG), G (n.v.), GH, n.v, MO Tyson, J. D. Dwyer & K. Blum 3319
(photos MEXU, PTBG), NY, n.v.). (Holotype MO 1824503 (photo PTBG);
—Rondeletia bertieroides Standi. Isotypes F, US (photos MEXU, MO)).
Arm hnotkryxpotycephala (Standi.) Borhidi, —Rondeletia bertieroides Standi.
Acta Bot. Hung. 35: 310. 1989. Rondeletia Arachnothryx secunda (Standi.) Borhidi, Acta
polycephala Standi, J. Wash. Acad. Sci. 17: Bot. Acad. Sci. Hung. 28: 70. 1982.
337. 1927. Type: MEXICO: Oaxaca, without Rondeletia secunda Standi, Contr. U.S. Natl.
precise locality, F. Liebtnann 11826 Herb. 18: 141. 1916. Type: PANAMA: San
(Holotype C, fragment US (photos MEXU, Bias, forests around Puerto Obaldia, San Bias
PTBG)).—Rondeletia mexicana (Turcz.) coast, 0-50 m, Aug 1911, H. Pittier 4279
Standi. (Holotype US 679392 (photos MEXU,
Arachnothryxpyramidalis (Lundell) Borhidi, PTBG); Isotype NY, n.v.).—Rondeletia
Acta Bot. Hung. 35: 310. 1989. Rondeletia secunda Standi.
pyramidalis Lundell, Wrightia 5: 327. 1976. Arachnothryx secundiflora (B. L. Rob.) Borhidi,
Type: MEXICO: Chiapas, San Luis, near Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
Siltepec, 1300 m, 24 Jan 1945, E. Matuda Rondeletia secundiflora B. L. Rob, Proc.
5292 (Holotype TEX-LL (photos MEXU, Amer. Acad. Arts 45: 403. 1910. Type:
A NOMENCLATOR OF MEXICAN AND CENTRAL /

GUATEMALA: Along road from Patin to MEXU, PTBG)).—Rondeletia standleyana

Escuintla, 1860, S. Hayes s.n. (Holotype GH, A. R. Molina
n.v.).—Rondeletia secundifiora B. L. Rob. Arachnothryx stenostachya (Standi.) Borhidi,
Arachnothryx septicidalis (B. L. Rob.) Borhidi, Acta Bot. Hung. 33: 303. 1987. Rondeletia
Acta Bot. Acad. Sci. Hung. 28: 69. 1982. stenostachya Standi., Publ. Field Mus. Nat.
Rondeletia septicidalis B. L. Rob., Proc. Hist., Bot. Ser. 18: 1372. 1938. Type: COSTA
Amer. Acad. Arts 45: 403. 1910. Type: RICA: La Hondura de San Jose, 1300 m, 15
MEXICO: Chiapas, Chicharras, 3000-6000 Aug 1933, M. Valerio 769 (Holotype F
ft., 6 Feb 1896, E. W. Nelson 3755 (Holotype 681816 (photos MEXU, PTBG)).
US 256722 (photos MEXU, PTBG); Isotype
GH, n.v.).—Rondeletia septicidalis B. L. Arachnothryx tacanensis (Lundell) Borhidi,
Rob. Acta Bot. Hung. 33: 303. 1987. Rondeletia
Arachnothryx silvicola (L. O. Williams) tacanensis Lundell, Bull. Torrey Bot. Club 66:
Borhidi, Acta Bot. Hung. 35: 311. 1989. 603. 1939. Type: MEXICO: Chiapas, Volcan
Rondeletia silvicola L. O. Williams, Tacana West, 2800 m, 30 Mar 1939, E
Phytologia 26: 128. 1973. Type: Matuda 2928 (Holotype TEX-LL (photos
HONDURAS: Atlantida, Lancetilla Valley, MEXU, PTBG); Isotypes A, n.v., F (photos
near Tela, 20-600 m, 6 Dec 1927-20 Mar MEXU, PTBG), GH, n.v., MEXU, US
1928, P. C. Standley 56885 (Holotype F (photos MEXU, PTBG)).—Rondeletia
581955 (photos MEXU, PTBG)). tacanensis Lundell
—Rondeletia silvicola L. O. Williams Arachnothryx thiemei (Donn. Sm.) Borhidi,
Arachnothryx skutchii (Standi. & Steyerm.) Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
Borhidi, Acta Bot. Hung. 33: 302. 1987. Rondeletia thiemei Donn. Sm., Bot. Gaz.
Rondeletia skutchii Standi. & Steyerm., Publ. (Crawfordsville) 42: 299. 1906. Type:
Field Mus. Nat. Hist., Bot. Ser. 22: 289. 1940. HONDURAS: Santa Barbara, Rio Chameleon,
Type: GUATEMALA: Suchitepequez, Finca 400 m, 16 Dec 1888, C. Thieme 5276
Moca, 4200 ft., 10 Jan 1935, A. F. Skutch (Holotype US 941403 (photos MEXU,
2110 (Holotype F 934088 (photos MEXU, PTBG)).—Rondeletia thiemei Donn. Sm.
PTBG); Isotypes A, n.v., US (photos MEXU, Arachnothryx torresii (Standi.) Borhidi, Acta
PTBG)).—Rondeletia skutchii Standi. & Bot. Hung. 33: 303. 1987. Rondeletia torresii
Steyerm. Standi., J. Wash. Acad. Sci. 18: 163. 1928.
Arachnothryx stachyoidea (Dorm. Sm.) Borhidi, Type: COSTA RICA: Alajuela, Viento
Acta Bot. Acad. Sci. Hung. 28: 69. 1982. Fresco, 1600-1900 m, 13 Feb 1926, P. C.
Rondeletia stachyoidea Donn. Sm., Bot. Gaz. Standley & R. Torres R. 47839 (Holotype US
(Crawfordsville) 42: 298. 1906. Type: 1305295 (photos MEXU, PTBG)).
GUATEMALA: Alta Verapaz, Semococh, —Rondeletia torresii Standi.
600-900 m, 25 Apr 1905, R. S. Hay s.n. trot hnothryx tuxtlensis (Lorence & Cast.-
(Holotype US 473726 (photos MEXU, Campos) Borhidi, Acta Bot. Hung. 35:311.
PTBG)).—Rondeletia stachyoidea Donn. 1989. Rondeletia tuxtlensis Lorence & Cast.-
Sm. Campos, Biotica 13: 148,7?g. /. 1988. Type:
Arachnothryx standleyana (A. R. Molina) MEXICO: Veracruz, Mun. Mecayapan,
Borhidi, Acta Bot. Hung. 33: 302. 1987. Volcan San Martin Pajapan al sur del Ejido La
Rondeletia standleyana A. R. Molina, Ceiba Valentina, 700 m, 8 May 1985, J. I. Calzada
1: 262. 1951. Type: HONDURAS: Morazan, 11183 (Holotype XAL; Isotypes F, MEXU,
in pine forest above Zambrano, 1200 m, 20 MO (photo PTBG)).—Rondeletia tuxtlensis
Jul 1949, L. O. Williams & A. Molina R. Lorence & Cast.-Campos
14417 (Holotype US 2216018 ex EAP Arachnothryx uxpanapensis (Lorence & Cast.-
(photos MEXU, PTBG); Isotype F (photos Campos) Borhidi, Acta Bot. Hung. 35: 311.
 1989. Rondeletia uxpanapensis Lorence & Almaraz 4244 (Holotype MEXU (photo
Cast.-Campos, Biotica 13: 150, fig. 2. 1988. PTBG); Isotypes CHAPA, PTBG).
Type: MEXICO: Veracruz, Mun. —Rondeletia wendtii Lorence & Cast.-
Hidalgotitlan, zona de Uxpanapa, Rio Campos
Solosuchil a 5-6 km al ESE de Hermanos Arachnothryxyucatanensis (Lundell) Borhidi,
Cedillo, 100-200 m, 24 Mar 1982, D. Lorence Acta Bot. Hung. 35: 311. 1989. Rondeletia
et al. 3909 (Holotype MEXU (photo PTBG); yucatanensis Lundell, Wrightia 5: 329. 1976.
Isotypes BM, CHAPA, ENCB, MO, NY, Type: MEXICO: Yucatan, Chichen Itza, 28
PTBG, UC, XAL).—Rondeletia Apr 1950, J. T. Baldwin, Jr. J4230 (Holotype
uxpanapensis Lorence & Cast.-Campos TEX-LL).—Rondeletia leucophylla Kunth
Arachnothryx villosa (Hemsl.) Borhidi, Acta Arachnothryx zolleriana (Standi. & Steyerm.)
Bot. Acad. Sci. Hung. 28: 69. 1982. Borhidi, Acta Bot. Hung. 33: 303. 1987.
Rondeletia villosa Hemsl., Diag. PI. Nov. Rondeletia zolleriana Standi. & Steyerm.,
Mexic. 27. 1879. Type: MEXICO: Oaxaca, Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 292.
San Pedro Nolasco, etc., Jurgensen 806 1940. Type: GUATEMALA: San Marcos,
(Lectotype K, here designated (photo PTBG); between Finca El Porvenir and Loma Corona,
Isolectotypes G-DEL, n.v. (fragment F), K 9 mi NW of El Porvenir, southwest-facing
(photo PTBG), P (photos MEXU, PTBG)). slopes of Volcan Tajumulco, 1300-2000 m,
Syntypes: MEXICO: Oaxaca, Trapiche de 14 Mar 1940, J A. Steyermark 37738
Concepcion, F. Liebmann 70 (K (photo (Holotype F 1039668 (photos MEXU,
PTBG); G-DEL, n.v., fragment F); Tabasco, PTBG)).—Rondeletia brandegeeana
Linden 145 (K (photo PTBG); probably G-
DEL, n.v., fragment F, P (photos MEXU,
PTBG)). Jurgensen 806 was designated Arcytophyllum Willd. ex Schult. & Schult. f. in
lectotype, as it is the more complete specimen Roem. & Schult., Syst. Veg. 3: 5. 1827. Type:
with three duplicates.—Rondeletia villosa Arcytophyllum blaerioides Willd. ex Schult.
Hemsl. & Schult. f.
Irat hnotkryx vulcanicola (Standi. & Steyerm.) Arcytophyllum lavarum K. Schum. ex Standi.,
Borhidi, Acta Bot. Hung. 33: 303. 1987. Contr. U.S. Natl. Herb. 18: 127. 1916. Type:
Rondeletia vulcanicola Standi. & Steyerm., PANAMA: Chinqui, Summit of Chinqui
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 291 Volcano, 3374 m, 12 Mar 1911, W. R. Maxon
1940. Type: GUATEMALA: Quezaltenango, 5349 (Holotype US (photos MEXU, PTBG);
Finca Pireneos, lower south-facing slopes of Isotype B, presumably destroyed (photo
Volcan Santa Maria de Jesus and Calahuache, MO)). Mallostoma lavarum Dorm. Sm.,
1300-1500 m, 31 Dec 1939,7. A. Steyermark Enum. PI. Guatem. 5: 36. 1899, nom. nud.
33220 (Holotype F 1041786 (photos MEXU, (based on Arcytophyllum lavarum K. Schum.,
PTBG)).—Rondeletia secundiflora B. L.
Rob.
Arcytophyllum muticum (Wedd.) Standi., J.
Arachnothryx wendtii (Lorence & Cast.- Wash. Acad. Sci. 18: 163. 1928. Hedyotis
Campos) Borhidi, Acta Bot. Hung. 35: 311. mutica Wedd., Chloris And. 2: 43, pi 50.
1989. Rondeletia wendtii Lorence & Cast.- 1857. Type: COLOMBIA: "Cundinamarca,
Campos, Biotica 13: 154, fig. 3. 1988. Type: Prov. Bogota," Goudot s.n. (Lectotype P, n.v.;
MEXICO: Oaxaca, Distr. Juchitan, Mun. de Isolectotype F, n.v.; designated by Mena V.,
Sta. Maria Chimalapa, Arroyo Chocolin, 1986).
Poblacion de Nicolas Bravo, cerca del Rancho Arcytophyllum recurvatum Suess., Bot. Jahrb.
de Augustin Montero, 3-4 km al sur de la Syst. 72: 286. 1942. Type: COSTA RICA:
Poblacion de Rio Alegre (Veracruz), 150 m Cartago, "Chirripo Grande, 3450 m," K.
21 Oct 1983, T. Wendt, A. Montero & I. Suessenguth 1313 (Holotype B, presumably
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

destroyed).—Arcytophyllum muticum MEXICO: Michoacan, Palo Verde, Pedregales

(Wedd.) Standi. de Jicalan et San Pedro, near Uruapan, 20 (or
Arcytophyllum shannonii (Donn. Sm.) Standi., 21) Sep 1941, M. Martinez 3400 (Lectotype
Contr. U.S. Natl. Herb. 18: 128. 1916. MEXU; Isolectotypes A, n.v., F, MEXU, US,
Mallostoma shannonii Donn. Sm., Bot. Gaz. 3 sheets (photos MEXU, PTBG)); designated
(Crawfordsville) 18: 203. 1893. Type: byFosberg, 1974.
GUATEMALA: Chimaltenango, Chicoy,
9000 ft., Mar 1892, W. C. Shannon s.n.
(Holotype US 48603 (photos MEXU, PTBG);
Isotype US 943475).—Hedyotis serpyllacea
Schltdl. Basanacantha altiscandens Ducke, Archiv, bast
Biol. Veg. Rio Janiero 2: 172. 1935. Type:
Asemnantha Hook. f. in Benth. & Hook., Gen. BRAZIL: Amazonas, B. Maguire et al. 29469
PL 2: 106. 1873. Type: Asemnantha (Holotype RB 24415, n.v.; Isotype NY, n.v.).
pubescens Hook. f. —Randia altiscandens (Ducke) C. M. Taylor
Asemnantha pubescens Hook. f. in Benth. & Basanacantha echinocarpa (Sesse & Moc. ex
Hook, f., Gen. PI. 2: 106. 1873. Type: DC.) Bullock, Hook. Ic. PI. 3: t. 3298. 1935.
MEXICO: Yucatan, Jul, Linden 1264 bis Randia echinocarpa Sesse & Moc ex DC,
(Holotype K, n.v.; Isotypes F (fragment, photo Prodr. 4: 385. 1830. Type: MEXICO: Fl.
PTBG), P). Mexic. Ic. 469 (presumably MA, n.v., caique
G-DC, n.v., copies MEXU, US). Genipa.
Augusta Pohl, PI. Bras. Icon. Descr. 2: 1. 1828 —Randia echinocarpa Sesse & Moc. ex DC.
(sero)-Feb 1829, nom. cons., not Augusta Basanacantha grandifolia Donn. Sm., Bot. Gaz.
Leandro, 1821. Type: Augusta lanceolata (Crawfordsville) 55: 436. 1913. Type:
Pohl, typ. cons. COSTA RICA: Forets de Santo Domingo de
Augusta rivalis (Benth.) J. H. Kirkbr., Brittonia Golfo Dulce, Feb 1896, A. Tonduz 9878
49: 358. 1997. Lindenia rivalis Benth., PI. (Lectotype US 941393, here designated
Hartweg. 84. 1841. Type: GUATEMALA: In (photos MEXU, PTBG); Isolectotypes F
rivis, la Vera Paz, T. Hartweg 581 (Holotype (photos MEXU, PTBG), US 577282).
K, n.v. (photo MO)). Siphonia. Syntype: COSTA RICA: Bords du Rio
Augusta rivalis (Benth.) J. H. Kirkbr. f. glabra Corozal pres Santo Domingo, Golfo Dulce,
(S. Darwin) J. H. Kirkbr., Brittonia 49: 358. Apr 1896, A. Tonduz 9982 (US; F (photos
1997. Lindenia rivalis Benth. f. glabra S. P. PTBG, MEXU)).—Randia grandifolia
Darwin, J. Arnold Arbor. 57: 439. 1976. (Donn. Sm.) Standi.
Type: MEXICO: Chiapas, Chicomucelo [as Basanacantha lasiantha Standi., Contr. U.S.
"Chicomuselo"], 800 m, 14 Jul 1941, £. Natl. Herb. 18: 134. 1916. Type: PANAMA:
Matuda 4449 (Holotype US 2087017 (photos Panama, Alajuela, Chagres Valley, 30-100 m,
MEXU, PTBG); Isotypes A, n.v., F, MO 12-15 May 1911,//. Pittier 3473 (Holotype
(photos MEXU, PTBG), NY, n.v.).—Augusta US 678517 (photos MEXU, PTBG)).
rivalis (Benth.) J. H. Kirkbr. —Randia lasiantha (Standi.) Standi.
Basanacantha monantha (Benth.) Hook. f. ex
Balmea Martinez, Anales Inst. Biol. Univ. Nac. Hemsl., Biol.Centr. Amer. Bot. 2: 39. 1881.
Mexico 13: 37. 1942; Bull. Torrey Bot. Club Randia monantha Benth., PI. Hartweg. 84:
69: 438. 1942. Type: Balmea stormae 1841. Type: GUATEMALA: Rio de los
Martinez Esclavos, T Hartweg 582 (Holotype K
Balmea stormae Martinez, Anales Inst. Biol. (photos F, MEXU); Isotypes B, presumably
Univ. Nac. Mexico 13: 31 Jigs. 1-4. 1942; destroyed (photo MO), BM, P (label states
Bull. Torrey Bot. Club 69: 438. 1942. Type:
 "Mexique," photos MEXU, PTBG)). Bathysa C. Presl, Bot. Bemerk. 84. 1844. Type:
—Randia monantha Benth. Bathysa stipulata C. Presl
Basanacantha pittieri Standi., Contr. U.S. Natl. Bathysa panamensis Dwyer, Ann. Missouri Bot.
Herb. 18: 134. 1916. Type: COSTA RICA: Gard. 55: 34. 1968. Type: PANAMA: Darien,
Gent Junction, pres Matina, Fev 1900, H. Rio Ucurganti, 7 Jul 1967, N. Bristan 1187
Pittier 16036 (Holotype US 578473 (photos (Holotype MO 1968219 (photo PTBG);
K, MEXU, PTBG); Isotypes F, P (fragment F, Isotype GH, n.v.).—Status uncertain.
photos MEXU, PTBG)).—Randia pittieri Bathysa veraguensis Dwyer, Arm. Missouri
(Standi.) Standi. Bot. Gard. 67: 40, fig. 9. 1980. Type:
Basanacantha pringlei S. Watson, Proc. Amer. PANAMA: Veraguas, Cerro Tute, 1200 m 28
Acad. Arts 18: 98. 1883. Type: MEXICO: Sep 1972, E. A.Lao & A. Gentry 531
Coahuila, Jimulco, mountains, 9 Apr 1885, (Holotype MO 2164349 (photo PTBG)).
C. G Pringle 142 (Holotype GH, n.v.;
Isotypes A, n.v., BM, F (photos MEXU, Belicea Lundell, Contr. Univ. Michigan Herb. 8:
PTBG), MEXU, P (photos MEXU, PTBG), 86. 1942. Type: Belicea hoffmannioides
US (photos MEXU, PTBG)).—Randia Lundell. [Herein treated as a synonym of
pringlei (S. Watson) A. Gray Morinda L.]
Basanacantha reticulata S. Watson, Proc. Belicea hoffmannioides Lundell, Contrib. Univ.
Amer. Acad. Arts 21: 379. 1886. Type: Michigan Herb. 8: 87. 1942. Type: BRITISH
MEXICO: Coahuila, S. Lorenzo de Laguna, HONDURAS (BELIZE): Toledo Distr.,
Feb-Oct 1880, E. Palmer 393 (Holotype US Swasey Branch, Monkey River, near falls, 7
47705 (photos MEXU, PTBG)).—Randia Mar 1942, P. H. Gentle 3945 (Holotype TEX-
pringlei (S. Watson) A. Gray LL, n.v.; Isotypes A, n.v., MO (photo
Basanacantha spinosa (Jacq.) K. Schum. var. PTBG)).—Morinda asperula Standi.
guatemalensis K. Schum. ex Loes., Verhandl.
Bot. Ver. Brandenburg 65: 110. 1923. Type: Berghesia C. G. D. Nees, Linnaea 20: 701.
GUATEMALA: "La Antigua, above 1847. Type: Berghesia coccinea C. G. D.
Alotengo," E. Seler & C. Seler 2570 Nees. [Genus of uncertain status. According
(Holotype B, presumably destroyed). to Nees's protologue description, Berghesia
—Randia armata (Sw.) DC. belongs to "tribu Cinchoneae," the capsule is
Basanacantha subcordata Standi, Contr. U.S. 2-locular, and each locule is 1-seeded.]
Natl. Herb. 18: 133. 1916. Type: COSTA Berghesia coccinea C. G. D. Nees, Linnaea 20:
RICA: Buissons et bois des chemins a Nicoya, 702. 1847. Type: MEXICO, without precise
May 1900, A. Tonduz 13934 (Holotype US locality, De Berghes s.n. (Holotype possibly
861251 (photos MEXU, PTBG); Isotypes B, LE, M, or GZU, n.v.).—Status uncertain.
destroyed (fragment F, photos MEXU,
PTBG), GH, n.v., LE, n.v. (fragment F, photo Bertiera AubL, Hist. PL Guiane 1: 180. 1775, as
PTBG), P (photos MEXU, PTBG; fragment F, "Berthiera." Type: Bertiera guianensis Aubl.
photos MEXU, PTBG)). Randia.—Randia Bertiera aequaliramosa Steyerm., Mem. New
monantha Benth. York Bot. Gard. 17: 319. 1967. Type:
Basanacantha tetracantha (Cav.) Hook. f. in PANAMA: Bocas del Toro, Region of
Jacks., Ind. Kew. 277. 1893. Mussaenda Almirante, Buena Vista Camp on Chiriqui
tetracantha Cav., Icon. PL 4: 20, pi. 435 Trail at 1250 ft., Jan-Mar 1928, G Proctor-
1799. Type: MEXICO: [Probably Guerrero], Cooper 586 (Holotype NY, n.v.; Isotypes F,
cerca Acapulco, Nee s.n. (Holotype MA, n.v. MO (photos MEXU, PTBG), US).—Bertiera
(photos F, MEXU, MO)).—Randia guianensis Aubl.
(Cav.) DC.
 i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Bertiera angustifolia Benth., Bot. Voy. Sulphur um mexicanum Standi.. Publ. Field
103. 1844. Type: COSTA RICA: Cocos Mus. Nat. Hist., Bot. Ser. 22: 108. 1940.
Island, Barclay s.n. (Holotype BM, n.v.). Type: MEXICO: Chiapas, Rvina [Ruina],
Bertiera guianensis Aubl., Hist. PL Guiane 1: Palenque, 10-12 Jul 1939, E. Matuda 3670
180, pi. 69. 1775. Type: FRENCH GUIANA: (Holotype F 1011462 (photo PTBG); Isotypes
"prope Aroura," Aublet s.n. (Holotype A, n.v., TEX-LL (photos MEXU, PTBG), US
presumably BM, n.v.). (photos MEXU, PTBG)).—Blepharidium
Bertiera guianensis var. leiophylla Steyerm., guatemalense Standi.
Mem. New York Bot. Gard. 17: 319. 1967.
Type: PANAMA: Chiriqui, vicinity of San Borojoa Cuatrecasas, Ser. Bot. Aplicada, Serv.
Felix, 0-120 m, 17 Dec 1911, H. Pittier 5178 Devulg., Seer. Agric. Ganad. (Call, Colombia)
(Holotype NY, n.v.; Isotype US (photos 2: 474. 1949. (Preprint of Rev. Acad.
MEXU, PTBG)).—Bertiera guianensis Aubl. Colombia Ci. Exact. 7: 474. 1950.) Type:
Bertiera oligosperma Wernham, J. Bot. 50: 162. Borojoa patinoi Cuatrecasas.
1912. Type: NICARAGUA: Seemann 129 Borojoa atlantica Dwyer, Ann. Missouri Bot.
(Holotype BM, n.v.).—Status uncertain. Gard. 67: 46. 1980. Type: PANAMA: Colon,
Standi., N. Amer. Fl. 32: 2-3 miles up from the Rio Guanche, 10-20 m,
180. 1934. Type: PANAMA: Darien, Cana 19 Jan 1973, H. Kennedy & R. Foster 2197
and vicinity, 2000-5600 ft., 17 Apr-8 Jun (Holotype MO 2163702 (photo PTBG);
1908, R. S. Williams 832 (Holotype US Isotype MO (photo PTBG)).
678312 (photos MEXU, PTBG); Isotype NY, Borojoa panamensis Dwyer, Phytologia 17:
n.v.).—Raritebe palicoureoides Wernham 446. 1968. Type: PANAMA: Code, Cerro
subsp. dwyeranum J. H. Kirkbr. Pilon, El Valle, 4 Jan 1968, J. A. Duke & B. R.
Bertiera tenuis Lundell, Wrightia 4: 49. 1968. Lallathin 15014 (Holotype MO 1968459
Type: GUATEMALA: Alta Verapaz, Sebol, (photo PTBG); Isotype MO).
in high forest about 800 m west of village, Borojoa patinoi Cuatrec, Ser. Bot. Aplicada,
bordering arroyo, 15 Apr 1964, E. Contreras Serv. Devulg., Seer. Agric. Ganad. (Cali,
4318 (Holotype TEX-LL (photos MEXU, Colombia) 2: 474. 1949. (Preprint of Rev.
PTBG); Isotype F).—Bertiera guianensis Acad. Colombia Ci. Exact. 7: 474. 1950.)
Aubl. Type: COLOMBIA: Valle, Rio Calima
Bertiera viburnoides (Standi.) J. H. Kirkbr., (region del Choco); margen derecha, entre
Brittonia 31:311. 1979. Raritebe viburnoides Quebrada de la Brea y La Esperanza, 25 m, 19
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. May 1946, J. Cuatrecasas 21137 (Holotype F
7: 56. 1930. Type: COLOMBIA: "Caucas, La 1251558 (photo PTBG); Isotypes F (3 sheets),
Gallera, Mica Valley," Killip 7934 (Holotype US (photo PTBG)).
US, n.v., fragment F, n.v.; Isotypes G, n.v., Y,
n.v.). Borreria G. Mey., Prim. Fl. Esseq. 79, t. 1. 1818.
Type: Borreria suaveolens G. Mey., excl. ref.
Browne Jam. 141, no. 4, nom. et typ. cons.
[Herein treated as a synonym of Spermacoce
L. In cases where generic transfers have not
Blepharidium guatemalense Standi., J. Wash. yet been published, the correct genus is given
Acad. Sci. 8: 59. 1918. Type: GUATEMALA: in boldface type.]
Alta Verapaz, forest along Saclac River, Borreria arenicola Brandegee, Univ. Calif.
below Secanquim, 300 m, 7 May 1905, H. Publ. Bot. 10: 417. 1924. Type: MEXICO:
Pittier 266 (Holotype US 472925 (photos Chiapas, in sandy arroyo near Jalisco, Aug
MEXU, PTBG); Isotype US (photos MEXU, 1923, C. A. Purpus 8968 (Holotype UC
PTBG)).
 220266, n. Kunth in Humb., Bonpl. & Kunth, Nov. Gen.
(DC.) Alain Sp. 3: 344. 1820. Type: MEXICO: Jalisco,
Borreria aspera M. Martens & Galeotti, Bull. Volcan de Jorullo, 3000 ft., Humboldt &
Acad. Roy. Bruxelles 11: 130. 1844. Type: Bonpland s.n. (Holotype P-Bonpland, n.v.
MEXICO: Oaxaca, Juquila, 4000 ft., H. (photo MO)).—Diodia apiculata (Willd. ex
Galeotti 2625 (Holotype BR, n.v.). Roem. & Schult.) K. Schum., vel sp. aff.
—Spermacoce suaveolens (G. May.) Kuntze Borreria exilis L. O. Williams, Phytologia 28:
Borreria asperifolia (M. Martens & Galeotti) 227. 1974. Type: COSTA RICA: Cocos
B. L. Rob., Proc. Amer. Acad. Arts 45: 409. Island, Wafer Bay, 28 June 1932, J. T. Howell
1910. Spermacoce asperifolia M. Martens & 10178 (Holotype F 1031197 (photo PTBG)).
Galeotti, Bull. Acad. Roy. Bruxelles 11: 132. Nom. nov. for Borreria gracilis L. O.
1844. Type: MEXICO: Oaxaca, pres de Williams, Phytologia 26: 487. 1973.
1'Ocean Pacifique, H. Galeotti 2626 —Spermacoce exilis (L. O. Williams) C D.
(Holotype BR, n.v.; Isotype P, n.v. (photo
MO)).—Staelia scabra (C. Presl) Standi. Borreria fruticosa Standi., J. Arnold Arbor. 11:
Borreria assurgens (Ruiz & Pav.) Griseb., Abh. 42. 1930. Type: HONDURAS: Comayagua,
Konigl. Ges. Wissensch. Gottingen 19: 159. Vicinity of Siguatepec, 1080-1400 m., 14-27
1874. Spermacoce assurgens Ruiz & Pav., Feb 1928, P. C. Standley 56240 (Holotype F
Syst. Veg. Fl. Peru 1: 60, t. 92b. 1798. Type: 581434 (photo PTBG); Isotype US (photos
PERU: Habitat m ruderalis Limae Chancay et MEXU, PTBG)).—Spermacoce
Huanuci, Ruiz Lopez & Pavon s.n. (syntype Borreria fruticosa Standi, var. pubescent Standi.
MA, n.v. (photo MO)); Plukenet Aim. 33, tab. & L. O. Williams, Ceiba 1: 165. 1950. Type:
136/4 (syntype, n.v.).—Spermacoce HONDURAS: El Paraiso, 2 miles W of
assurgens Ruiz & Pav. Guinope, 1400 m, 16 Aug 1946, L. O.
Borreria auriculata Brandegee, Univ. Calif. Williams & A. Molina R. 10315 (Holotype US
Publ. Bot. 10: 417. 1924. Type: MEXICO: 1206971 (photos MEXU, PTBG); Isotype F
Chiapas, meadows near Jalisco, Aug 1923, (photo PTBG)).—Spermacoce
C A. Purpus 9218 (Holotype UC, n.v.; Borreria gracilis L. O. Williams, Phytologia 26:
Isotype F (photo PTBG)).—Crusea 487. 1973. Not Borreria gracilis Scheele,
psyllioides (Kunth) W. R. Anderson 1843, nor Miq. ex Hook., 1855. Type:
Borreria bartlingiana DC, Prodr. 4: 544. 1830. COSTA RICA: Cocos Island, Wafer Bay, 28
Type: MEXICO: Without precise locality, Jun 1932, J. T. Howell 10178 (Holotype F
Haenke s.n. (Holotype presumably PR, n.v.). 1031197 (photo PTBG)).—Spermacoce exilis
—Spermacoce latifolia Aubl. (L. O. Williams) C D. Adams
Borreria chiapensis Brandegee, Univ. Calif. Borreria graminifolia M. Martens & Galeotti,
Publ. Bot. 10: 417. 1924. Type: MEXICO: Bull. Acad. Roy. Bruxelles 11: 129. 1844.
Chiapas, Hacienda Monserrate, Sep 1923, Type: MEXICO: El Sabino, pres de Zimapan,
C. A. Purpus 9216 (Holotype UC 223384, 6500 pieds, H. Galeotti 2599 (Holotype BR,
n.v.; Isotypes F (photo PTBG), NY, n.v., US n.v.).—Spermacoce verticillata L.
(photos MEXU, PTBG)).—Crusea setosa Borreria haenkeana DC, Prodr. 4: 547. 1830.
(M. Martens & Galeotti) Standi. & Steyerm. Type: MEXICO: Without precise locality or
Borreria densiflora DC, Prodr. 4: 542. 1830. collector, presumably Haenke (G-DEL, n.v.
Type: JAMAICA: Without precise locality, (photo MO)).—Borreria suaveolens G. Mey.
Swartzs.n. (Holotype G-DC, n.v. (microfiche Borreria heterophylla Brandegee, Univ. Calif.
4. 542.3)).—Spermacoce densiflora (DC) Publ. Bot. 10: 418. 1924. Type: MEXICO:
Chiapas, in sandy arroyo near Jalisco, Aug
1923, C. A. Purpus 8969 (Holotype UC
220267, n.v.).—Spermacoce prostrata Aubl.
 L NOMENCLATOR OF MEXICAN AND CENTRAL

B, •! , laevigata M. Martens & Galeotti, Bull. Isotype US (photos MEXU, PTBG)).

Acad. Roy. Bruxelles 11: 130. 1844. Type: —Spermacoce nesiotica (B. L. Rob.) G. A.
MEXICO: Michoacan, Vallee de Morelia
(Valladolid de Michoacan), or Mexico, la Borreria oaxacana M. Martens & Galeotti, Bull.
plaine de Jordana, pres Toluca, 8000 pieds, H. Acad. Roy. Bruxelles 11: 130. 1844. Type:
Galeotti 2586 (Holotype BR, n.v.; Isotype P, MEXICO: Oaxaca, pres de la ville d'Oaxaca,
n.v. (photo MO)).- 5000 pieds, H. Galeotti 2629 (Holotype BR,
n.v).—Spermacoce verticillata L.
Borreria laevis (Lam.) Griseb., Abh. Kon. Borreria ocymoides (Burm.f.) DC, Prodr. 4:
Gewiss. Gottingen 7: 231.1857. Spermacoce 544. 1830. Type: Tropical Asia, collector
laevis Lam., Tabl. Encycl. et Meth. Bot. 1: unknown, (Holotype presumably G, n.v.).
273., n. 1435. 1791. Type: SANTO Spermacoce ocymoides Burm.f, Fl. Ind. 34.
DOMINGO: Without precise locality, Martin 1768. This name has been erroneously applied
s.n. (Holotype P-LA, n.v.).—Spermacoce
tenuior L. Note: Although Borreria laevis with - 1 flowers,
sensu Griseb. equals Spermacoce assurgens
Ruiz & Pav., the type of the basionym,
Spermacoce laevis Lam., equals the type of Borreria ovalifolia M. Martens & Galeotti, Bull.
Spermacoce tenuior L. Acad. Roy. Bruxelles 11: 129. 1844. Type:
Borreria latifolia (Aubl.) K. Schum. in Mart., MEXICO: Veracruz, Mirador, 5000 pieds, H.
Fl. Bras. 6(6): 61., t. 80. 1888. Spermacoce Galeotti 2606 (Holotype BR, n.v.).
latifolia Aubl., Hist. PI. Guiana 55. 1775. —Spermacoce ovalifolia (M. Martens &
Type: FRENCH GUIANA: Cayenne, Aublet Galeotti) Hemsl.
s.n. (Holotype P, n.v.; Isotype MA, n.v. Borreria patula M. Martens & Galeotti, Bull.
(photo MO)).—Spermacoce latifolia Aubl. Acad. Roy. Bruxelles 11: 128. 1844. Type:
Borreria longiseta M. Martens & Galeotti, Bull. MEXICO: Veracruz, Mirador, 3000 pieds, H.
Acad. Roy. Bruxelles 11: 128. 1844. Type: Galeotti 2608 (Holotype BR, n.v.; Isotype K,
MEXICO: Veracruz, Zacuapan et Mirador n.v.).—Spermacoce prostrata Aubl.
[label on isotype at P states "cordilleres pres Borreriapodocephala DC, Prodr. 4: 542. 1840.
de Vera Cruz, 1000 pieds"], H. Galeotti 2611 Type: MEXICO or CUBA: Without precise
(Holotype BR, n.v.; Isotypes K, n.v., P)). locality or collector, based on living material
—Status uncertain. According to C. D. (IDC 800.719.11.1 (photo MO)).
Adams (pers. comra. 1996) the description —Spermacoce verticillata L.
does not agree with the type, which actually Borreria procera Schltdl., Linnaea 9: 593. 1834.
represents a specimen of Spermacoce confusa Type: MEXICO: Veracruz, Hacienda de la
Rendle. Laguna, Schiede & Deppe 250 (Holotype
Borreria malacophylla Standi. & L. O. HAL, n.v.).—Spermacoce suaveolens (G.
Williams, Ceiba 1: 165. 1950, nom. illeg., not Mey.) Kuntze
Borreris malacophylla K. Schum., 1903. Borreria pumilio Standi, Ann. Missouri Bot.
Type: HONDURAS: El Paraiso, near Galera, Gard. 27: 342. 1940. Type: PANAMA: Code,
moist thicket, 900 m, Nov-Dec 1943, P. C. Nata, ca. 50 m, 12 Sep 1938, P. H. Allen 822
Standley 15584 (Holotype US 2215979 (Holotype MO 1808165 (photos MEXU,
(photos MEXU, NY, PTBG); Isotype F). PTBG); Isotypes F (photo PTBG), GH, n.v.,
—Spermacoce assurgens Ruiz & Pav. US (photo PTBG)).—Spermacoce
Borreria nesiotica B. L. Rob., Proc. Amer. Borreria radicosa Brandegee, Univ. Calif. Publ.
Acad. Arts 45: 409. 1910. Type: MEXICO: Bot. 10: 416. 1924. Type: MEXICO: Chiapas,
Socorro Island (Revillagigedo Group), 1897, meadows near Jalisco, Aug 1923, C. A.
A. W. Anthony s.n. (Holotype GH, n.v.; Purpus 9223 (Holotype UC 223392, n.v.;
 Isotypes GH, n.v., US (photos MEXU, Borreria tonalensis Brandegee, Univ. Calif.
PTBG)).—Spermacoce densiflora (DC.) Publ. Bot. 6: 191. 1915. Type: MEXICO:
Chiapas, Sierra de Tonala, C. A. Purpus 7369
Borreria rhadinophylla B. L. Rob., Proc. Amer. (Holotype VC, n.v.; Isotype GH, n.v.).
Acad. Arts 45: 409. 1910. Type: BRITISH —Spermacoce
HONDURAS (BELIZE): On dry sandy pine Borreria vegeta Standi. & Steyerm., Publ. Field
ridges, 23 Oct 1905, M. E. Peck 180 Mus. Nat. Hist, Bot. Ser. 23: 21. 1943. Type:
(Holotype GH, n.v.).—Mitracarpus GUATEMALA: Dept. Jutiapa, between
rhadinophyllus (B. L. Rob.) L. O. Williams Jutiapa and La Burrera, NE of Jutiapa,
Borreria scabiosoides Cham. & Schltdl., 800-850 m, 1 Nov 1940, P. C. Standley
Linnaea 3:318. 1828. Type: BRAZIL: Rio de 76007 (Holotype F 1091300 (photos MEXU,
Janeiro, Sellow s.n. (presumably B, PTBG)).—Spermacoce
destroyed?).—Spermacoce scabiosoides Borreria verticillata (L.) G. Mey., Prim. Fl.
(Cham. & Schltdl.) Kuntze Esseq. 83. 1818. Spermacoce verticillata L.,
Borreria setosa M. Martens & Galeotti, Bull. Sp. PI. 102. 1753. Type: JAMAICA: Hort.
Acad. Roy. Sci. Bruxelles 11:131. 1844. Cliff. 33: (BM, n.v.).—Spermacoce
Type: MEXICO: Oaxaca, dans les forets de verticillata L.
palmiers et sur les dunes de la cote d'Oaxaca, Borreria verticillata (L.) G. Mey. var.
baignee par l'Ocean Pacifique, pres du thymiformis B. L. Rob., Proc. Amer. Acad.
Rancho du Rio Grande, Sep 1839, H. Galeotti Arts 45: 410. 1910. Type: MEXICO: Oaxaca,
2627 (Holotype BR, n.v.; Isotype P, n.v. 18 miles SW of city of Oaxaca, 7000-8000
(photo MO)).—Crusea setosa (M. Martens & ft., 10-20 Sep 1894, E. W. Nelson 1410
Galeotti) Standi. & Steyerm. (Holotype GH, n.v.; Isotype US 574465
Borreria suaveolens G. Mey., Prim. Fl. Esseq. (photos MEXU, PTBG)).—Spermacoce
SI.,pi. 1. 1818. Type: SURINAM: In arenosis verticillata L.
continentis circa Arouabischkreck, G. Meyer
s.n. (Holotype GOET, n.v.).—Spermacoce Bouvardia Salisb., Parad. Lond.pl. 88. 1807.
suaveolens (G. Mey.) Kuntze Type: Bouvardia tnphylla Salisb.
Borreria subulata DC. Prodr. 4: 543. 1830. Bouvardia alexanderae A. Carter, Madrono 13:
Type: MEXICO: Without precise locality, 3 142, figs. 1, 2. 1955. Type: MEXICO: Baja
Sep 1827, Berlandier 826 (Lectotype G-DEL, California: Distr. del Sur, Arroyo del Salto, E
n.v. (photos MO, US), B (specimen annotated of La Paz, 30 Mar 1949, A. Carter 2577
by de Candolle but not cited in protologue); (Holotype UC 985926, n.v.; Isotypes F,
Isolectotype BM, n.v.; designated by MEXU, US (photos MEXU, PTBG)).
Anderson, 1972). Crusea.—Crusea Carterella.—Hedyotis alexanderae (A.
diversifolia (Kunth) W. R. Anderson Carter) W. H. Lewis
Borreria tampicana DC, Prodr. 4: 544. 1830. Bouvardia angustifolia Kunth in Humb., Bonpl.
Type: MEXICO: Tamaulipas, Tampico, 1827, & Kunth, Nov. Gen. Sp. 3: 384. 1819. Type:
Berlandier 37 bis (Holotype G-DC, n.v. MEXICO: Moran (?), A. Bonplands.n.
(photo MO); Isotype BM, n.v.). (Holotype P-Bonpland, n.v. (photo MO);
—Spermacoce prostrata Aubl. Isotype F (photo PTBG)).—Bouvardia
Borreria tenella (Kunth) Cham. & Schltdl., ternifolia (Cav.) Schltdl.
Linnaea 3: 317. 1828. Spermacoce tenella Bouvardia bicolor Kunze, Linnaea 20: 24. 1847.
Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Type: Based on plants grown from Mexican
& Sp. 3: 345. 1820. Type: Locality unknown, seed (the description is taken as the type;
Humboldt s.n. (Holotype B-W, n.v. (photo designated by Blackwell, 1968).—B
MO)).—Spermacoce suaveolens (G. Mey.) multiflora (Cav.) Schult. & Schult.
 i NOMENCLATOR OF MEX

Bouvardia bouvardioides (Seem.) Standi., N. MO (photo PTBG), US (photo PTBG)).

Amer. Fl. 32: 102. 1921. Hedyotis Arachnothryx.—Rondeletia deamii (Donn.
bouvardioides Seem., Bot. Voy. Herald 296. Sm.) Standi.
1856. Type: MEXICO: NW Mexico Bouvardia dictyoneura Standi., N. Amer. FL
[Durango], Sierra Madre, Seemann s.n. 32: 109. 1921. Type: MEXICO: Chiapas,
(Holotype BM, n.v.: Isotypes C, n.v., Chicharras, 3000-6000 ft., 6 Feb 1896, E. W.
fragment F (photo PTBG), GH, n.v.). Nelson 3757 (Holotype US 256554 (photos
Bouvardia capitata Bullock, Hook. Ic. PI. t. MEXU, PTBG); Isotype GH, n.v).
3296. 1935. Type: MEXICO: Mexico, Distr. Bouvardia discolor Hook. & Arn., Bot. Beechey
Temascaltepec, Palmar, barranca, 7 Jul 1934, Voy. 428. 1840. Type: MEXICO: Oaxaca,
G. B. Hinton 6319 (Holotype K, n.v.; Isotypes inter Tehuantepec et Voca [Boca] del Monte
BM, n.v., F (photo PTBG)). in dit. Oaxaca maxime australis, Sept., G.
Bouvardia cataphyllaris Bullock, Hook. Ic. PL t. Andrieux 334 (Holotype K, n.v.; Isotype P
3297. 1935. Type: MEXICO: Mexico, Distr. (photos MEXU, PTBG)). Not Rondeletia
Temascaltepec, Temascaltepec, 1780 m, 28 discolor Kunth in Humb., Bonpl. & Kunth,
Jul 1932, G. B. Hinton J131 (Holotype K, 1820—Rondeletia leucophylla Kunth
n.v.; Isotypes BM, n.v., F (photo PTBG), G, I I.oes., Verh. Bot. Ver.
n.v., GH, n.v., MICH, n.v., MEXU, MO Brandenburg 65: 106. 1923. Type:
(photos MEXU, PTBG), US (photos MEXU, GUATEMALA: Huehuetenango, Distr.
PTBG)).—Bouvardia chrysantha Mart. Nenton, near Uaxackanal and toward Quen
Bouvardia cavanillesii DC, Prodr. 4: 366. 1830. Santo, 1100 m, 9 Jul 1896, E. Seler & C. Seler
Nom. superfl., based on Acginctia muhitlora 2883 (Holotype B, presumably destroyed
Cav.—Bouvardia multiflora (Cav.) Schult. (photo MO); Lectotype US 1205579 (photos
& Schult. f. MEXU, PTBG); Isolectotypes F (fragment),
Bouvardia chrysantha Mart., Del. Hort. GH, n.v.; designated by Blackwell, 1968).
Monac. Sem. 1848. 4. 1848. Type: MEXICO: —Bouvardia glabra Pol.
[Hidalgo], near Santiaguillo, Kanvinsky s.n. Bouvardia endlichii Loes., Repert. Sp. Nov. 18:
(Holotype presumably BR, n.v.). 357. 1922. Type: MEXICO: Durango, Endlich
Bouvardia conzattii Greenm., Proc. Amer. 176a (Holotype B, presumably destroyed
Acad. Arts 39: 92. 1903. Type: MEXICO: (photos F, MO, US)).—Bouvardia ternifolia
Oaxaca, San Felipe, 1750 m, Jul-Aug 1900, (Cav.) Schltdl.
C. Conzatti & V. Gonzales 1067 [published as Bouvardia erecta (DC.) Standi., N. Amer. Fl.
"1076"] (Holotype GH, n.v.). 32: 110. 1921. Catesbaea erecta DC, Prodr.
Bouvardia cordifolia DC, Prodr. 4: 366. 1830. 4: 401. 1830. Type: MEXICO: Without
Type: MEXICO: Without precise locality, precise locality, Sesse & Mocino 560
Sesse & Mocino s.n. (Holotype MA, n.v.; Fl. (Holotype MA, n.v.; probable Isotype F
Mexic. Ic. G-DC, n.v. (photo MO)). (photo PTBG); Fl. Mexic. Ic, MA (photo
Bouvardia corymbosa Oerst., Vidensk. Meddel. MO)).
Naturhist. Foren Kjobenhavn 1852: 46. 1853. Bouvardia flava Decne., Fl. Serres 1: 215, pi.
Type: NICARAGUA: Without precise 38. 1845. Type: Based on a greenhouse-
locality, Oersted 11039 (Holotype C, n.v.). grown plant (protologue illustration is the
—Bouvardia leiantha Benth. type; designated by Blackwell, 1968).
Bouvardia deamii Donn. Sm., Bot. Gaz. —Bouvardia laevis M. Martens & Galeotti
(Crawfordsville) 49: 445. 1910. Bouvardia flos-joannis K. Schum., Bull. Herb.
GUATEMALA: Fiscal, Rillo de las Nabajas, Boissier3: 621. 1895. Type: MEXICO: Prope
3700 ft., 5 Jun 1909, C. C. Deam 6190 Puebla, Jul, E. Seler & C. Seler 846 (Holotype
(Holotype US 579579 (photo PTBG); B, presumably destroyed; Isotype GH, n.v.).
Isotypes A, n.v., F (photo PTBG), GH, n.v., —Bouvardia erecta (DC) Standi.
 K. Schum. var. latifolia Bouvardia hintonii Bullock, Kew Bull. 1937:
Loes., Repert. Sp. Nov. 18: 358. 1922. Type: 307. 1937 [published as "hinton!"]. Type:
MEXICO: Puebla, N of Tehuacan, 1725 m, MEXICO: Mexico, Distr. Temascaltepec,
Endlich 1966 (Holotype B, presumably Nanchititla, barranca, 27 Jul 1935, G. B.
destroyed).—Bouvardia erecta (DC.) Standi. Hinton et al. 8107 (Holotype K, n.v.; Isotypes
Bouvardia glaberrima Engelmann in Wisl., A, n.v., BM, n.v., F, G, n.v., GH, n.v.,
Mem. Tour. N. Mex. 106. 1848. Type: MEXU, MO (photos MEXU, MO), US
MEXICO: Cosihuiriachi, Wislizenus 161 (photos MEXU, PTBG)).—Bouvardia
(Holotype MO (photos MEXU, PTBG); loeseneriana Standi.
Isotype GH, n.v.).—Bouvardia ternifolia Bouvardia hintoniorum B. L. Turner, Brittonia
(Cav.) Schltdl. 38: III,fig. 1. 1986. Type: MEXICO:
Bouvardia glabra Pol., Linnaea41: 565. 1877. Guerrero, ca. 42.5 miles SW of Mexico
Type: COSTA RICA: San Jose, Aug 1875, Highway 95 along Milpillas-Atoyac road via
Polakowsky 337 (Holotype B, destroyed Puerto del Gallo, 11.5 miles SW of Filo de
(photos F, G, MO, US); Lectotype F (photo Caballo and 6.3 miles SW of Carrazal de
PTBG), fragment of holotype; designated by Bravo, ca. 8500 ft., 16 Oct 1975, J. L. Reveal
Blackwell, 1968). etal. 4232 (Holotype TEX, n.v.; Isotypes
Bouvardia glabra Pol. var. gracilis Pol., MEXU, MO (photo PTBG)).
Lmnaea 41: 566. 1877. Type: COSTA RICA: Bouvardia hirtella Kunth in Humb., Bonpl. &
San Jose, cultivated, Jun 1875, Polakowsky Kunth, Nov. Gen. Sp. 3: 384. 1819. Type:
172 (Holotype B, presumably destroyed MEXICO: Humboldt & Bonpland s.n.
(photos F, MO)).—Bouvardia glabra Pol. (Holotype P-Bonpland, n.v. (photos F, MO,
Bouvardia glabra Pol. var. obtusa Loes., Verh. US)).—Bouvardia ternifolia (Cav.) Schltdl.
Bot. Ver. Brandenburg 65: 106. 1923. Type: Bouvardia houtteana Schltdl. ex Planch., Fl.
GUATEMALA: Quezaltenango, near Serres 10: 149, /. 55. 1855. Type: The
Almolonga, E. Seler & C. Seler 2920 protologue illustration; designated by
(Holotype B, presumably destroyed). Blackwell, 1968.—Bouvardia ternifolia
—Bouvardia glabra Pol. (Cav.) Schltdl.
Bouvardia gracilipes B. L. Rob., Proc. Amer. Bouvardia hypoleuca Benth., PI. Hartw. 288.
Acad. Arts 45: 404. 1910. Type: MEXICO: 1848. Type: MEXICO: Ad Aguas Cahentes,
Nayarit, Tepic, 5 Jan-6 Feb 1892, E. Palmer 1837, T. Hartweg J605 (Holotype K, n.v.).
1971 (Holotype GH, n.v; Isotypes F, US —Bouvardia ternifolia (Cav.) Schltdl.
(photos MEXU, PTBG)). Bouvardia induta (B. L. Rob.) Standi., N. Amer.
Bouvardia gracilis A. Gray, Proc. Amer. Acad. Fl. 32: 109. 1921. Bouvardia longiflora var.
Arts 22: 306. 1887. Type: MEXICO: induta B. L. Rob., Proc. Amer. Acad. 45: 404.
Chihuahua, shade of cliffs, Mapula Mtns., 23 1910. Type: MEXICO: Chiapas, Jun-Jul
Aug 1887, C G. Pringle 1255 (Holotype GH, 1864-70, Ghiesbreght s.n. (Holotype GH,
n.v.; Isotypes F (photo PTBG), MEXU). middle collection, n.v.).—Bouvardia glabra
—Bouvardia multiflora (Cav.) Schult. & Pol.
Schult. f. Bouvardia jacquinii Kunth in Humb., Bonpl. &
Bouvardia heterophylla Standi., N. Amer. Fl. Kunth, Nov. Gen. Sp. PI. 3: 385. 1820. Norn,
32: 107. 1921. Type: GUATEMALA: Santa superfl., based on Ixora americana Jacq.
Rosa, Santa Rosa, 1000 m, May 1892, Heyde —Bouvardia ternifolia (Cav.) Schltdl.
& Lux 3137 (Holotype US 398167 (photos Bouvardia jacquinii Kunth var. exogyna DC,
MEXU, PTBG); Isotype GH, n.v.). Prodr. 4: 365. 1830. Type: Based on a plant
—Bouvardia multiflora (Cav.) Schult. & grown in Salm-Dyck's garden (Holotype G-
Schult. f. DC, n.v.).—Bouvardia ternifolia (Cav.)
Schltdl.
 t NOMENCLATOR OF MEXICAN AND CENTRAL

i Kunth var. ovata DC, Bouvardia longiflora (Cav.) Kunth in Humb.,

Prodr. 4: 365. 1830. Norn, superfl., based on Bonpl. & Kunth, Nov. Gen. Sp. 3: 386. 1820.
Bouvardia triphylla Salsib.—Bouvardia Aeginetia longiflora Cav., Anal. Ci. Nat. 3:
ternifolia (Cav.) Schltdl. 130, t. 28, f. 1. 1801. Type: MEXICO:
Bouvardia karwinskyi Standi., Publ. Field Between Queretaro and Guanajuato, collector
Mus. Nat. Hist., Bot. Ser. 8: 155. 1930. Type: unknown (Holotype MA, n.v.). Anotis.
MEXICO: Tamaulipas, between Victoria and Bouvardia longiflora var. induta B. L. Rob.,
Rio Blanco, Karwinsky 312 (Holotype LE, Proc. Amer. Acad. Arts 45: 404. 1910. Type:
n.v., fragment F). MEXICO: Chiapas, without precise locality,
Bouvardia laevis M. Martens & Galeotti, Bull. Jun-Jul 1864-70, Ghiesbreght s.n. (Holotype
Acad. Roy. Sci. Bruxelles 11: 236. 1844. GH, n.v.).—Bouvardia glabra Pol.
Type: MEXICO: Veracruz, au bord des ravins Bouvardia longiflora var. latifolia M. Martens
de la colonie de Zacuapan, 3000 pieds, Apr & Galeotti, Bull. Acad. Bruxelles 11: 236.
1840, H. Galeotti 2600 (Holotype BR, n.v. 1844. Type: MEXICO: Oaxaca, montagnes de
(fragment F); Isotype P (photos F, MO, US)). Calpulalpan, 7000-8000 pieds, H. Galeotti
Bouvardia langlassei Standi, N. Amer. Fl. 32: 2659 (Holotype BR, n.v.).—Bouvardia
110. 1921. Type: MEXICO: Guerrero, Yestla longiflora (Cav.) Kunth
[Yextla], 1500 m, 8 Jun 1899, E. Langlasse Bouvardia macdougallii Lorence, Novon 4:
1049 (Holotype US 386252 (photos MEXU, 1 \9,fig. 1. 1994. Type: MEXICO: Oaxaca,
PTBG); Isotypes B, n.v., F (fragment), G, Cerro Atravesada-Cerro Azul, "Arroyo
n.v., GH, n.v., P, n.v.). Rana," Dec 1956, T. MacDougall s.n.
Bouvardia latifolia Standi., N. Amer. Fl. 32: (Holotype MEXU (photo PTBG)).
111. 1921. Type: MEXICO: Guerrero, Mont I maeilenta S. F. Blake, Contr. Gray
de la Piedra, pres Aguila, 450 m, 16 Jul 1898, Herb. 53: 65. 1918. Type: MEXICO: Oaxaca,
E. Langlasse 246 (Holotype US 385804 Miahuatlan, Cerro El Zopilote, 2100 m, 18
(photos MEXU, PTBG; fragment F); Isotypes May 1917, C. Conzatti & B. Reko 3288
B, presumably destroyed (fragment F), GH, (Holotype GH, n.v.).—Bouvardia
n.v., P, n.v.).—Bouvardia multiflora (Cav.) chrysantha Mart.
Schult. & Schult. f. Bouvardia macrantha Standi., N. Amer. Fl. 32:
Bouvardia leiantha Benth., PI. Hartw. 85. 107. 1921. Type: MEXICO: Puebla,
1841. Type: GUATEMALA: In arvis prope Tlacuiloltepec, 7000-8000 ft., Jul 1909, C. A.
TejaretChimaltenango, Jul-Aug 1841, T. Purpus 3981 (Holotype US 841341 (photos
Hartweg 583 (Holotype K, n.v.; Isotypes B, MEXU, PTBG); Isotypes BM, n.v., F, GH,
presumably destroyed (photo MO), GH, n.v., n.v., MO (photos MEXU, PTBG), UC, n.v.).
P, n.v., W, n.v.). —Bouvardia multiflora (Cav.) Schult. &
Bouvardia linearis Kunth in Humb., Bonpl. & Schult. f.
Kunth, Nov. Gen. Sp. PI. 3: 383. 1819. Type: Bouvardia matudae Lundell, Lloydia 2: 105.
MEXICO: Without precise locality, Humboldt 1939. Type: MEXICO: Chiapas, Mt. Ovando,
& Bonpland s.n. (Holotype P-Bonpland, n.v. Feb 1939, E Matuda 2667 (Holotype MICH,
(photos F, MO, US)).-Bouvardia ternifolia n.v.; Isotypes A, n.v., F, MO (photos MEXU,
(Cav.) Schltdl. PTBG), US (photos MEXU, PTBG)).
Bouvardia loeseneriana Standi., Field Mus. —Bouvardia dictyoneura Standi.
Nat. Hist., Bot. Ser. 11: 186. 1936. Type: Bouvardia microphylla Schltdl., Linnaea 26:
MEXICO: Guerrero, Texquitzin, near 112. 1853. Type: MEXICO: Without precise
Chilapa, 28 Oct 1829, L. Schultze 89 locality, Schiede s.n. (Holotype B, presumably
(Holotype B, presumably destroyed (photo F), destroyed).—Bouvardia ternifolia (Cav.)
fragment F (photo PTBG)). Schltdl.
 26: 85. 1853. Type: MEXICO: Without
. Bouvardia pachecoana Standi. & Steyerm.,
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 22.
precise locality or collector, herb. Lipsky 1943. Type: GUATEMALA: San Marcos,
(presumably LE, n.v.).—Bouvardia laevis M. Barranco Eminencia, road between San
Martens & Galeotti Marcos and San Rafael Pie de la Cuesta,
Bouvardia multiflora (Cav.) Schult. & Schult. 2500-2700 m, 6 Feb 1941, P. C. Standley
f, Mant. Syst. Veg. 3:118. 1827. Aeginetia 86226 (Holotype F 1093896 (photo PTBG);
multiflora Cav., Anal. Ci. Nat. 3: 131, /. 28, f. Isotypes A, n.v., US (photos MEXU, PTBG)).
2. 1801; Icon. 6: 52, t. 572, f. 2. 1801. Type: —Bouvardia dictyoneura Standi.
MEXICO: Queretaro, Nee s.n. (Holotype MA, Bouvardia pallida Standi., N. Amer. Fl. 32: 102.
n.v. (photo MO)). 1921. Type: EL SALVADOR: Volcan de San
Bouvardia myrtifolia Schltdl., Linnaea 26: 121. Salvador, 1600-1800 m, 7 Apr 1922, P. C.
1853. Type: MEXICO: Hacienda de Standley 22977 (Holotype US 1138686
Cocoyotle, Schiedes.n. (Holotype B, (photos MEXU, PTBG); Isotype GH, n.v.).
presumably destroyed).—Bouvardia —Bouvardia bouvardioides (Seem.) Standi.
chrysantha Mart. Bouvardia purpusii Brandegee, Univ. Calif.
Bouvardia nubigena Standi. & Steyerm., Publ. Publ. Bot. 10: 415. 1924. Type: MEXICO:
Field Mus. Nat. Hist., Bot. Ser. 22: 382. 1940. Chiapas, Hacienda Monserrate, Sep 1923,
Type: GUATEMALA: Jutiapa, Volcan de C A. Purpus 9251 (Holotype UC 223423,
Suchitan, NW of Asuncion Mita, 2050 m, 18 n.v.; Isotypes F, GH, n.v., MO (photos
Nov 1939, J. A. Steyermark 31901 (Holotype MEXU, PTBG), US (photos MEXU, PTBG)).
F 1043221 (photos F, PTBG)).—Bouvardia —Bouvardia induta (B. L. Rob.) Standi.
laevis M. Martens & Galeotti Bouvardia quaternifolia DC, Prodr. 4: 365.
Bouvardia oaxacana Standi., J. Wash. Acad. 1830. Type: MEXICO: Circa urbem Mexici,
Arts 13: 7. 1923. Type: MEXICO: Oaxaca, Alaman s.n. (Holotype G-DC, n.v.).
Distr. Juquila, de Santa Cruz al Aguacate, 500 —Bouvardia ternifolia (Cav.) Schltdl.
m, 24 Dec 1921, C. Conzatti 4513 (Holotype Bouvardia quinqueflora Dehnh. in Schltdl.,
US 1110842 (photos MEXU, PTBG); Isotype Linnaea 26: 92. 1853. Type: MEXICO:
Without precise locality or collector (location
Bouvardia obov , Bonpl. & unknown).—Bouvardia chrysantha Mart.
Kunth, Nov. Gen. Sp. PI. 3: 385. 1820. Type: Bouvardia quinquenervata Standi., N. Amer.
MEXICO: Distr. Federal, entre Chapultepec et FL 32: 108. 1921. Type: MEXICO: Chiapas,
Tezcoco, Jun 1803-04, A. Bonpland s.n. San Bartolome, 22 Mar 1904, E. A. Goldman
(Holotype presumably P-Bonpland, n.v.). 769 (Holotype US 470574 (photos MEXU,
'louvardia orizabensis Standi., Publ. Field Mus. PTBG)).
Nat. Hist., Bot. Ser. 8: 334. 1931. Type: Bouvardia rekoi Standi., N. Amer. Fl. 32: 108.
MEXICO: Veracruz, Orizaba, 1854, M. 1921. Type: MEXICO: Oaxaca, Cafetal
Botteri 604 (Holotype P, n.v. (fragment F); Montecristo (Cerro Espino), 1000 m, 7 Dec
Isotype G, n.v.).—Bouvardia ternifolia 1917, B. P. Reko 3650 (Holotype US 867149
(Cav.) Schltdl. (photos MEXU, PTBG)).
lime i A. Gray, PL Wright. 5: 67. Bouvardia rosea Schltdl., Linnaea 26: 116.
1853. Type: MEXICO: Sonora, mountain 1853. Type: MEXICO: Guanajuato, 22 miles
valley from San Pedro to Santa Cruz, Sep west of Xichu, road from Xichu to San Luis
1851, C. Wright 1117 (Holotype GH, n.v.; de la Paz, 2300 m, 14 Jun 1957, R. McVaugh
Isotypes MO (photos MEXU, PTBG), US 14819 (Neotype MICH, n.v.; designated by
(photos MEXU, PTBG)).—Bouvardia Blackwell, 1968).
ternifolia (Cav.) Schltdl. Bouvardia rosei Standi., N. Amer. Fl. 32: 109.
1921. Type: MEXICO: Durango, Sierra
 tNOMENCLATOROF D CENTRAL AMERICAN RUBIACEAE

Madre, 16 Aug 1897, J. N. Rose 3516 Bouvardia steyermarkii Standi., Publ. Field
(Holotype US 302493 (photos MEXU, Mus. Nat. Hist., Bot. Ser. 22: 383. 1940.
PTBG); Isotype F). Type: GUATEMALA: Zacapa, Sierra de las
Bouvardia rzedowskii Terrell & S. D. Koch, Minas, between Rio Hondo and Finca
Novon4: 179, fig. 1. 1994. Type: MEXICO: Alejandna, 1700-2000 m, 11 Oct 1939, J. A.
Oaxaca, ladera caliza con vegetacion de Steyermark 29671 (Holotype F 1043386
encinar, 2300 m, 6 km al SW de (photos F, PTBG)).—Bouvardia dolichantha
Tamazulapan, sobre la carretera a Chilapa, 12
Aug 1977, J. Rzedowski 35142 (Holotype Bouvardia strigosa Benth., PI. Hartweg. 75.
ENCB; Isotypes ARIZ, n.v., ASU, n.v., 1841. Type: GUATEMALA: Quezaltenango,
MICH, n.v., XAL). [Volcan] Zunil, Nov., T. Hartweg 530
Bouvardia salvadorensis Steyerm., Ceiba 4: (Lectotype K (photo PTBG), here designated;
302. 1955. Type: EL SALVADOR: Isolectotypes G-DEL, n.v. (fragment F), P
Sonsonante, Rio Acachapa near headquarters (photos MEXU, MO, PTBG), W, n.v. (photos
of Hacienda Las Tablas, Balsam Range, 570 F, MO)). Syntype: GUATEMALA: Volcan de
m, May 1941, J. M. Tucker 1368 (Holotype F Fuego, 6000-6400 ft., Salvin s.n. (K (photo
1492726 (photo PTBG); Isotypes MICH, n.v., PTBG),fragment F). Rogiera.—Rondeletia
UC, n.v., US (photos MEXU, PTBG)). strigosa (Benth.) Hemsl.
—Bouvardia multiflora (Cav.) Schult. & Bouvardia subcordata Standi, N. Amer. Fl.
Schult. f. 32: 105. 1921. Type: MEXICO: Sinaloa, near
Bouvardia scabra Hook. & Arn., Bot. Beechey Colomas, 13 Jul 1897,7. N. Rose 1628
Voy. 427. 1840. Type: MEXICO: Nayarit, (Holotype US 300476 (photos MEXU,
between San Bias and Tepic, Sinclair s.n. PTBG); Isotype F).
(Holotype K, n.v.). Bouvardia tenuijlora Schltdl., Linnaea 26: 97.
Bouvardia scabrida M. Martens & Galeotti, 1853. Type: Based on living plants in
Bull. Acad. Roy. Sci. Bruxelles 11: 237. 1844. cultivation (B, presumably destroyed).
Type: MEXICO: Oaxaca, ravin de Yavezia, —Bouvardia ternifolia (Cav.) Schltdl.
cordillere orientale, 6000-7000 pieds, Dec Bouvardia tenuifolia Standi., N. Amer. Fl. 32:
1840, H. Galeotti 2624 (Holotype BR, n.v.; 104. 1921. Type: MEXICO: Jalisco, grassy
Isotype P, n.v. (photo MO)).—Bouvardia hillsides near Guadalajara, Oct 1889, C. G.
ternifolia (Cav.) Schltdl. Pringle 2292 (Holotype US 49988 (photos
Bouvardia schiedeana Schltdl., Linnaea 26: MEXU, PTBG); Isotypes A, n.v., BM, n.v.,
123. 1853. Type: MEXICO: Barranca de BR, n.v., F, G, n.v., LE, n.v., M, n.v., MICH,
Tioselo, Aug, Schiedes.n. (Holotype B, n.v., MO, MSC, n.v., P, n.v., US, n.v., W,
presumably destroyed).—Bouvardia
multiflora (Cav.) Schult. & Schult. f. Bouvardia ternifolia (Cav.) Schltdl., Linnaea
Bouvardia splendens J. Graham, Bot. Mag. pi. 26: 98. 1853. Ixora ternifolia Cav., Ic. 4: 3, /.
3781. 1840. Type: Based on greenhouse- 305. 1797. Lectotype: Protologue illustration
grown plants (Holotype K, n.v.).—Bouvardia of Ixora ternifolia Cav., Ic. 4: 3, /. 305. 1797
ternifolia (Cav.) Schltdl. (based on a plant grown at MA from Mexican
Bouvardia standleyana W. H. Blackw., Ann. seed; designated by Blackwell, 1968).
Missouri Bot. Gard. 55: 13. 1968. Type: Bouvardia ternifolia (Cav.) Schltdl. var.
MEXICO: Michoacan, Coalcoman, S angustifolia (Kunth) B. L. Rob., Proc. Amer.
Naranjillo, 1360 m, 26 Nov 1938, G. B. Acad. Sci. 45: 405. 1910. Bouvardia
Hinton et al. 12677 (Holotype US 2020693 angustifolia Kunth in Humb., Bonpl. &
(photos MEXU, PTBG); Isotypes K, n.v., Kunth, Nov. Gen. Sp. 3: 384. 1819. Type:
MICH, n.v., UC, n.v.). MEXICO: Moran (?), A. Bonpland s.n.
(Holotype P-Bonpland, n.v. (photo MO);
 Isotype F (photo PTBG)).—Bouvardia (photo PTBG), GH, n.v.).—Bouvardia
ternifolia (Cav.) Schltdl. xylosteoides Hook. & Arn.
Bouvardia tolucana Hook. & Am., Bot. Bouvardia viminalis Schltdl., Linnaea 26: 120.
Beechey Voy. 427. 1840. Type: MEXICO: 1854. Type: MEXICO: Oaxaca, Monte Alban,
Mexico, Toluca, Andheux 332 (Holotype K, 6000 ft., 4 Sep 1894, C G. Pringle 4888
n.v.; Isotype W, n.v.).—Bouvardia ternifolia (Neotype MO 174074 (photos MEXU,
(Cav.) Schltdl. PTBG); Isoneotypes BM, n.v., BR, n.v., F,
Bouvardia triflora Kunth in Humb., Bonpl. & GH, n.v., M, n.v., MEXU, 2 sheets, MSC,
Kunth, Nov. Gen. Sp. 3: 386. 1820. Type: n.v., P, n.v., UC, n.v., US (photos MEXU,
MEXICO: In temperatis, Humboldt & PTBG), W, n.v.; designated by Blackwell,
Bonplands.n. (Holotype P-Bonpland, n.v. 1968).
(fragment P, n.v.)). Houstonia.—Bouvardia Bouvardia violacea Rzed., Ciencia (Mexico) 19:
multiflora (Cav.) Schult. & Schult. f. 82. 1959. Type: MEXICO: San Luis Potosi, J.
Bouvardia triflora Kunth var. hirsuta M. Rzedowski 7680 (Holotype MEXU; Isotype
Martens & Galeotti, Bull. Acad. Bruxelles 11: ENCB).—Bouvardia rosea Schltdl.
236. 1844. Type: MEXICO: Oaxaca, Bouvardia xestosperma (B. L. Rob. &
cordillere orientale, dans les montagnes du Greenm.) Terrell & S. D. Koch, Novon 4:
Rincon, 5000 pieds, H. Galeotti 2658 bis 181. 1994. Oldenlandia xestosperma B. L.
(Holotype BR, n.v.).—Bouvardia multiflora Rob. & Greenm., Proc. Amer. Acad. Arts 32:
(Cav.) Schult. & Schult. f. 41. 1896. Type: MEXICO: Oaxaca, open
Bouvardia triphylla Salisb., Parad. Lond.,p/. 88. glades, Sierra de San Felipe, 8000 ft., Jun
1807. Norn, superfl., based on Ixora ternifolia 1894, C. G. Pringle 4692 (Lectotype GH,
Cav.—Bouvardia ternifolia (Cav.) Schltdl. n.v.; Isolectotypes BM, n.v., CM, n.v., F, K,
Bouvardia venosissima Lundell, Bull. Torrey n.v., MEXU, 2 sheets, MICH, n.v., MO
Bot. Club 66: 602. 1939. Type: MEXICO: (photos MEXU, PTBG), MSC, n.v., PH, n.v.,
Chiapas, Mt. Tacana, 1400 m, 17-23 Mar US, 2 sheets, VT, n.v.; designated by Terrell,
1939, E. Matuda 2748 (Holotype MICH, n.v.; 1980).
Isotype MEXU).—Bouvardia dictyoneura Bouvardia xylosteoides Hook. & Arn., Bot.
Standi. Beechey Voy. 428. 1840. Type: MEXICO:
Bouvardia versicolor KerGawl., Bot. Reg. 3: pi. Oaxaca, on slightly elevated mountains about
245. 1817. Type: Based on a greenhouse- Mitlam [Mitla], Andrieux 335 (Holotype K,
grown plant (the protologue illustration is n.v.; Isotype G-DEL, n.v., fragment F).
taken as the type; designated by Blackwell,
1968).—Bouvardia multiflora (Cav.) Schult. Buena Cav., Anal. Hist. Nat. (Madrid) 2: 278, /.
& Schult. f. 23. 1800. Type: Buena panamensis Cav. Not
Bouvardia versicolor Ker Gawl. var. graciliflora Buena Pohl, 1826. [Herein treated as a
A. Gray in S. Watson, Proc. Amer. Acad. Arts synonym of Gonzalagunia Ruiz & Pavon.]
22: 416. 1887. Type: MEXICO: Jalisco, Rio Buena panamensis Cav., Anal. Hist. Nat.
Blanco, E. Palmer 154 (Syntype GH, n.v.); (Madrid) 2: 279. 1800. Type: PANAMA:
Tequila, E. Palmer 369 (Syntype GH, n.v.); Without precise locality, Nee s.n. (Holotype
Chapala, E. Palmer 708 (Syntype GH, n.v.). MA, n.v.; Isotype F (photo PTBG)).
—Bouvardia multiflora (Cav.) Schult. & Gonzalea.—Gonzalagunia panamensis
Schult. f. (Cav.) K. Schum.
Bouvardia villosa Standi., N. Amer. Fl. 32: 107. Buena skinneri Oerst., Vidensk. Meddel. Dansk
1921. Type: MEXICO: Oaxaca, Alturas de Naturhist. Foren. Kjobenhavn 1852: 48. 1853.
Matatlan, 1800 m, C. Conzatti I486 TYPE: NICARAGUA: Masaya, toppen af
(Holotype US 764094 (photos MEXU, Vulkanen Masaya, 2000-3000 ft., Oersted
PTBG)); Isotypes G-DEL, n.v., fragment F 11134 (Syntypes C, n.v. (photos A, F), US);
 \. NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Oersted 11695 (Syntype C, n.v.).

—Cosmibuena grandiflora (Ruiz & Pav.)
Rusby
H. Galeotti, type not designated.
Calderonia Standi., J. Wash. Acad. Sci. 13: 290. —Hoffmannia discolor (Lem.) Hemsl.
1923. Type: Calderonia salvadorensis Standi. Campylobotrys discolor Hook., Bot. Mag. t.
[Herein treated as a synonym of Simira Aubl.] 4530. 1850. Not Campylobotrys discolor
Calderonia klugei Standi., J. Wash. Acad. Sci. Lem. 1847. Type: Based on cultivated
15: 6. 1925. Type: PANAMA: Panama, near
Chepo, 1924, H. C Kluge 19 (Holotype US Bahia, Brazil, but most likely from Mexico,
1206424 (photos MEXU, PTBG)). Sickingia. type not designated. Later published by
—Simira klugei (Standi.) Steyerm. Hooker under the new name Higginsia regalis
Calderonia salvadorensis Standi., J. Wash. Hook, f, Bot. Mag. t. 5280. 1861.
Acad. Sci. 13: 290. 1923. Type: EL —Hoffmannia regalis (Hook, f.) Hemsl.
SALVADOR: San Salvador, Jul 1922, 5. Campylobotrys ghiesbreghtii Lem., 111. Hortic.
Calderon 761 (Holotype US 1151718 (photos 8: pi. 279. 1861. Type: Based on living
MEXU, PTBG)). Sickingia.—Simira material collected in Mexico by Ghiesbreght,
salvadorensis (Standi.) Steyerm. type not designated.—Hoffmannia
ghiesbreghtii (Lem.) Hemsl.
Callicocca Schreb., Gen. 126. 1789. Type not Campylobotrys pyrophylla Linden, Cat. 16: 2.
designated. [Herein treated as a synonym of 1861, nom. nud—Hoffmannia refulgens
Psychotria L.] (Hook, f.) Hemsl.
Callicocca ipecacuanha Brot., Trans. Linn. Soc.
London 6: 137. 1802. Type: BRAZIL: Carterella Terrell, Brittonia 39: 249. 1987.
Without precise locality, Gomez s.n. Type: Carterella alexanderae (A. Carter)
(presumably LISU, n.v.). Cephaelis. Terrell. [Herein treated as a synonym of
—Psychotria ipecacuanha (Brot.) Stokes Hedyotis L.]
Carterella alexanderae (A. Carter) Terrell,
Calycophyllum DC, Prodr. 4: 367. 1830. Brittonia 39: 250. 1987. Bouvardia
alexanderae A. Carter, Madrono 13: 142, fig.
(Vahl) DC. {Macrocnemum candidissimum 1, 2. 1955. Type: MEXICO: Baja California
Vahl). Sur: Arroyo del Salto, E of La Paz, 24° 12*N,
Calycophyllum candidissumum (Vahl) DC, 110°7.5'E, 30 Mar 1949, A. Carter 2577
Prodr. 4: 367. 1830. Macrocnemum (Holotype UC 985926, n.v.; Isotype US
candidissimum Vahl, Symb. 2: 38. pi. 30. (photo PTBG)).—Hedyotis alexanderae (A.
1791. Type: TRINIDAD: Vicinia St. Martha Carter) W. H. Lewis
von Rohr s. n. (Holotype C, i
Casasia A. Rich in Sagra, Cuba II: 9, t 49.
Campylobotrys Lem., Fl. Serres 3: Misc. 37 (ad 1850. Type: Casasia calophylla A. Rich.
260). 1847. Type: Campylobotrys discolor Casasia chiapensis Miranda, Ceiba 4: 142.
Lem. [Herein treated as a synonym of 1954. Type: MEXICO: Chiapas, Rancho el
Hoffmannia Sw.] Ocote, 30 km al NW de Ocozocuautla, 650 m,
Campylobotrys argyroneura Lindl., Belg. 21 May 1950, F. Miranda 7743 (Lectotype
Hortic. 8: 166. 1858. Type: Based on MEXU; designated by Lorence & Dwyer,
cultivated material collected by Ghiesbreght 1987). Syntype: MEXICO: Chiapas, F.
in Chiapas, Mexico, type not designated, Miranda 6624 (F). —Randia matudae
a discolor (Lem.) Hemsl. Lorence & Dwyer
Cascarilla (Endl.) Wedd., Ann. Sci. Nat., Bot. (Holotype US 2215926 (photos MEXU,
ser. 3, 10: 10. 1848. Not Adanson 1763. Type PTBG); Isotype US (photos MEXU, PTBG)).
not designated. (Based on Cinchona sect. —Psychotria borucana (A. R. Molina) C. M.
Cascarilla Endl.) Taylor & W. C. Burger
Cascarilla bullata Wedd., Ann. Sci. Nat., Bot. Cephaelis bristanii Dwyer & Hayden, Ann.
ser. 3, 10: 11. 1848. Type: PERU: Puuo, Prov. Missouri Bot. Gard. 55: 34. 1968. Type:
de Carabaya, Jan-Feb 1847, Weddel 4358 PANAMA: Darien, Cerro Pirre, Bristan 575
(Holotype P-Ju, n.v.; Isotypes G, n.v., K, n.v.) (Holotype MO 1961544)).—Psychotria
Dwyer (1980) erroneously recorded this
species from Panama, although it is actually icunponutans Dwyer & Hayden, Ann.
endemic to the southern Andes (L. Andersson, Missouri Bot. Gard. 55: 35. 1968. Type:
pers. coram. 1996) —Ladenbergia bullata PANAMA: Colon, Cerro Jefe, Dwyer &
(Wedd.) Standi. Gauger 7346 (Holotype MO).—Psychotria
camponutans (Dwyer & Hayden) Hammel
Cassupa Humb. & Bonpl., PI. Aequin. 1: 43, r Cephaelis chiapensis (Standi.) Standi., Publ.
12. 1806. Type: Cassupa verrucosa Humb. & Field Columbian Mus., Bot. Ser. 4: 295. 1929.
Bonpl. [Herein treated as a synonym of Isertia Evea chiapensis Standi., Contr. U.S. Natl.
Schreb.] Herb. 23: 1392. 1926. Not Psychotria
Cassupa panamensis Standi., Contr. U.S. Natl. chiapensis Standi., 1926. Type: MEXICO:
Herb. 18: 135. 1916. Type: PANAMA: Colon, Chiapas, Cerro del Boqueron, Sep 1913, C A.
along the Rio Fato, 10-100 m, Jul-Aug 1911, Purpus 6928 (Holotype US 567214 (photos
H. Pittier 3889 (Holotype US 678965 (photo MEXU, PTBG); Isotypes F, GH, n.v., MO
PTBG); Isotype US).—Isertia laevis (Triana) (photos MEXU, PTBG)).—Psychotria

Cephaelis chiriquiensis Standi., Ann. Missouri

iL., Sp. PI. 1: 109. 1753. Type: Bot. Gard. 28: 469. 1941. Type: PANAMA:
Catesbaea spinosa L. Chiriqui, vicinity of Bajo Chorro, 1900 m,
Catesbaea erecta DC, Prodr. 4: 401. 1830. 20-22 Jul 1940, R. E. Woodson, Jr. & R. W.
Type: MEXICO: Without precise locality, Schery 610 (Holotype MO 1209028 (photos
Sesse & Mocino s.n. (presumably MA, n.v., MEXU, PTBG); Isotypes F, GH, n.v.).
fragment F); Fl. Mexic. Ic. (G-DC, n.v.)). —Psychotria chiriquiensis (Standi.) C. M.
—Bouvardia erecta (DC.) Standi. Taylor
Cephaelis chlorochlamys Standi., Publ. Field
Cephaelis Sw., Prodr. Veg. Ind. Occ. 3: 45. Mus. Nat. Hist., Bot. Ser. 18: 1278. 1938.
1788 (nom. cons). Type: Cephaelis muscosa Type: COSTA RICA: San Jose, La Hondura
(Jacq.) Sw., type cons. [Herein treated as a de San Jose, 15 Aug 1933, J. Valerio 704
synonym of Psychotria L.] (Holotype F 681815 (photo PTBG)).
Cephaelis axillaris Sw., Prodr. Veg. Ind. Occ. 3: —Psychotria hazenii Standi.
45: 1788. Type: St. Christopher [St. Kitts], Cephaelis correae Dwyer & Hayden, Ann.
Swartzs.n. (Holotype presumably S, n.v.). Missouri Bot. Gard. 55: 36. 1968. Type:
Not Psychotria axillaris Willd. 1798. Evea, PANAMA: Code, mountains N of El Valle de
Tapogomea.—Psychotria aubletiana Anton, 2500-3000 ft., 28 May 1967, W. H.
Steyerm. Lewis, B. MacBryde, R. L. Oliver & J. E.
Cephaelis borucana A. R. Molina, Ceiba 4: 31. Ridgway 1759 (Holotype MO 1887361
1951. Type: COSTA RICA: Puntarenas, forest (photos MEXU, PTBG); Isotypes GH, n.v.,
near Esquinas Experimental Station, area US, n.v., UC, n.v.).—Psychotria correae
between Rio Esquinas and Rio Palmar Sur de (Dwyer & Hayden) C. M. Taylor
Osa, 200 ft., 22 May 1950, P. H. Allen 5540
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Cephaelis costaricensis Schltdl., Linnaea 28: Cephaelis gaugeri Dwyer, Ann. Missouri Bot.
546. 1856. Type: COSTA RICA: Type not Gard. 67: 71. 1980. Type: PANAMA:
designated.—Psychotria elata (Sw.) Hammel Panama, Cerro Jefe, near junction of roads to
Cephaelis croatii Dwyer, Ann. Missouri Bot. Cerro Jefe and Altos de Pacora, 18 Dec 1974,
Gard. 67: 64. 1980. Type: PANAMA: Darien, S. Mori & J. Kallunki 3779 (Holotype MO
upper gold mining camp of Tyler Kittredge, 2240077 (photos MEXU, PTBG)).
headwaters of Rio Tuquesa, ca. 2 air km from —Psychotria camponutans (Dwyer &
continental divide, 26 Aug 1974, T. B. Croat Hayden) Hammel
27204 (Holotype MO 2201416 (photos Cephaelis gentryi Dwyer, Selbyana 2: 57. 1977.
MEXU, PTBG); Isotypes F, GH, n.v.). Type: ECUADOR: Los Rios, Rio Palenque
—Psychotria gentryi (Dwyer) C. M. Taylor Science Center, km 56 of Quevedo-Santo
Cephaelis dichroa (Standi.) Standi., Publ. Field Domingo Road, C. Dodson 5169 (Holotype
Columbian Mus., Bot. Ser. 4: 296. 1929. Evea MO; Isotypes RPSC, n.v., SEL, n.v., US,
dichroa Standi., Contr. U.S. Nat. Herb. 18: n.v.).—Psychotria gentryi (Dwyer) C. M.
124. 1916. Type: PANAMA: Chiriqui, humid Taylor
forest of Cuesta de Las Palmas, southern Cephaelis glomerulata Dorm. Sm., Bot. Gaz.
slopes of Cerro de la Horqueta, 1700-2100 m, (Crawfordsville) 16: 12. 1891. Type:
17-19 Mar 1911, H. Pittier 3218 (Holotype GUATEMALA: Dept. Livingston, Rio Dulce,
US 677615 (photos MEXU, PTBG); Isotype Mar 1880, J. D. Smith 1637 (Holotype US
F)).—Psychotria dichroa (Standi.) C. M. 943529 (photos MEXU, PTBG); Isotype US
Taylor (photos MEXU, PTBG)).—Psychotria
Cephaelis dimorphandrioides Dwyer, Ann. glomerulata (Donn. Sm.) Steyerm.
Missouri Bot. Gard. 67: 66. 1980. Type: Cephaelis guapilense (Standi.) Standi., Publ.
PANAMA: Veraguas, Valley of Rio Dos Field Columbian Mus., Bot. Ser. 4: 295. 1929.
Bocas on road between Alto Piedra (above Evea guapilensis Standi., J. Wash. Acad. Arts
Santa Fe) and Calovebora, 350-400 m, 29 Sci. 15: 104. 1925. Type: COSTA RICA:
Aug 1974, T. B. Croat 27470 (Holotype MO Limon, vicinity of Guapiles, 300-500 m, P. C.
2201358 (photos MEXU, PTBG)). Standley 37025 (Holotype US 1153029
—Palicourea dimorphandrioides (Dwyer) (photos MEXU, PTBG)).—Psychotria
C. M. Taylor guapilensis (Standi.) C. W. Ham.
Cephaelis discolor Pol., Linnaea 41: 572. 1877. Cephaelis hirsute M. Martens & Galeorti, Bull.
Type: COSTA RICA: Angustura, 1875, H. Acad. Roy. Bruxelles 11: 135. 1844. Type:
Polakowsky 384 (Holotype B, presumably MEXICO: Oaxaca, la Chinantla, 4000 pieds,
destroyed (photos F, MO, US), fragment F H. Galeotti 7185 (Holotype BR, n.v.).
(photo PTBG)). Not Psychotria discolor —Psychotria poeppigiana Mull. Arg.
Huber, 1901, nor P. discolor (Griseb.) Rolfe, Cephaelis insueta Dwyer, Ann. Missouri Bot.
1893—Psychotria guapilensis (Standi.) Gard. 67: 73. 1980. Type: PANAMA:
Hammel Veraguas, 11 km from Escuela Agricola Alto
Cephaelis dressleri Dwyer, Ann. Missouri Bot. de Piedra, forest along Rio Dos Bocas,
Gard. 67: 68, fig. 15. 1980. Type: PANAMA: tropical wet forest, Atlantic slope, 15 Nov
San Bias, forest south of Puerto Obaldia, 18 1974, S. Mori & J. Kallunki 3108 (Holotype
Aug 1971, T. B. Croat 16801 (Holotype MO MO 2220981 (photo PTBG)).—Psychotria
2138462 (photo MO)).—Psychotria dressleri insueta (Dwyer) C. W. Ham.
(Dwyer) C. W. Ham. Cephaelis ipecacuanha (Brot.) A. Rich., Bull.
Cephaelis elata Sw., Prodr. Veg. Ind. Occ. 45. Fac. Med. 4: 92. 1818. Callicocca
1788. Type: JAMAICA, Swartz s.n. ipecacuanha Brot., Trans. Linn. Soc. London
(presumably S, n.v.).—Psychotria elata 6: 137. 1802. Type: BRAZIL: Without precise
(Sw.) Hammel
 locality, Gomez s.n. (presumably LISU, n.v.).
—Psychotria ipecacuanha (Brot.) Stokes
Cephaelis kennedyae Dwyer, Ann. Missouri A. Gentry & S. Mori 13895 (Holotype MO
Bot. Gard. 67: 75. 1980 [as kennedyi}. Type: 2276226 (photos MEXU, PTBG)).
PANAMA: Panama, 6 km from Panamerican —Psychotria dressleri (Dwyer) C. W. Ham.
Highway on road from El Llano to Carti- Cephaelis phoenicia (K. Schum.) Dorm. Sm.,
Tupile, H. Kennedy 1779 (Holotype MO Enum. PI. Guat. 5: 39. 1899. Uragoga
2161974 (photo PTBG); Isotype GH, n.v.). Phoenicia K. Schum. in Engl. & Prantl, Nat.
—Psychotria muscosa (Jacq.) Steyerm. Pflanzenfam. 4, 4: 120. 1891. Type: COSTA
( 'cphcichs lansupulata Standi., J. Wash. Acad. RICA: Suerre, Llanuras de Santa Clara, 300
Sci. 18: 281. 1928. Type: COSTA RICA: m, Feb 1896, J. D. Smith 6584 (Syntype US,
Cartago, vicinity of Orosi, 30 Mar 1924, P. C. n.v.); same locality, J. D. Smith 6586
Standley 39695 (Holotype US 1153168 (Syntype US, n.v.).—Psychotria elata (Sw.)
(photos MEXU, PTBG)). Not Psychotria Hammel
latistipula Benth. in Hook., 1849. Cephaelis pittieri K. Krause, Bot. Jahrb. Syst.
—Psychotria cartagoensis Nepokroeff 54, Beibl. 119: 45. 1916. Type: COSTA
Cephaelis nana (Standi.) Standi., J. Wash. Acad. RICA: Puntarenas, Cafias Gordas, 1100 m,
Sci. 17: 171. 1927. Evea nana Standi., J. Feb 1897, H. Pittier 11162 (Holotype B,
Wash. Acad. Sci. 15: 105. 1925. Type: presumably destroyed (photos F, MO;
PANAMA: Canal Zone, hills north of Fnjoles, fragment F); Isotype US (photos MEXU,
19 Dec 1923, P. C Standley 27550 (Holotype PTBG)). not Psychotria pittieri Standi. 1916.
US 1153871 (photos MEXU, PTBG); Isotype —Psychotria aurantibracteata C. M. Taylor
F). Not Psychotria nana K. Krause 1908. Cephaelis psychotriifolia Seem., Bot. Voy.
—Psychotria guapilensis (Standi.) Hammel Herald 138. 1852 [aspsychotriaefolia]. Type:
Cephaelis nicaraguensis Standi., Trop. Woods PANAMA: Panama, damp forest near Cruces;
16: 46. 1928. Type: NICARAGUA: Region of and Darien, Chirambira, Seemann 482
Braggmann's Bluff, on bank of Kukalaya (Holotype BM, n.v. (photo MO); Isotype K
River near Camp 14, 200 ft., 8 Dec 1927, [as Seemann 482bis]).—Psychotria
F. C. Englesing 58 (Holotype F 572595 psychotriifolia (Seem.) Standi.
(photo PTBG)).—Psychotria guapilensis Cephaelis punicea Vahl, Eclog. Amer. 1: 19.
(Standi.) Hammel 1796. Type: JAMAICA: Without precise
Cephaelis ostreophora Wernham, J. Bot. 55: locality, von Rohr s.n. (n.v.).—Psychotria
284. 1917. Type: COLOMBIA: Bogota, elata (Sw.) Hammel
Servita y Villavicencio, Triana s.n. (Holotype Cephaelis rigidifolia Dwyer & Hayden, Ann.
BM, n.v.; Isotype F).—Psychotria Missouri Bot. Gard. 55: 37. 1968. Type:
ostreophora (Wernham) C. M. Taylor PANAMA: Panama, Cerro Jefe, ca. 2700 ft.,
Cephaelis panamensis Dwyer, Ann. Missouri 20 Aug 1967, J. D. Dwyer 8075A (Holotype
Bot. Gard. 67: 77. 1980. Type: PANAMA: MO 2165097 (photos MEXU, PTBG)).
Colon, Rio Guanche, ca. 2.5 km upriver from Palicourea.—Status uncertain.
bridge on road to Portobelo, along small creek Cephaelis sessilifolia Standi., Publ. Field
to the S of the river, 10-100 m, 14 Dec 1974, Columbian Mus., Bot. Ser. 4: 295. 1929.
S. Mori & J. Kallunki 3714 (Holotype MO Type: MEXICO: Jalisco, Quimixto, trail from
2353037 (photos MEXU, PTBG); Isotypes San Pedro el Tuito, 60 m, 2 Dec 1926, Y.
MO 2353038 (photos MEXU, PTBG), PMA Mexia 1240 (Holotype F 579912 (photo
22701, n.v).—Rudgea panamensis (Dwyer) PTBG); Isotypes A, n.v., MO (photos MEXU,
C. M. Tayor PTBG), US (photos MEXU, PTBG)). Not
Cephaelis panchocoensis Dwyer, Ann. Missouri Psychotria sessilifolia M. Martens & Galeotti,
Bot. Gard. 67: 78. 1980. Type: PANAMA: 1844—Psychotria molinarum Lorence
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Cephaelis tetragona Donn. Sm., Bot. Gaz. •'••us occidentalis I var. salicifolius
(Crawfordsville) 61: 376. 1916. Type: (Humb. & Bonpl.) A. Gray, Syn. PI. 1: 29.
COSTA RICA: Forets de Tuis, Oct 1897, 1878. Cephalanthus salicifolius Humb. &
Tonduz 11354 (Syntype US 943500 (photos Bonpl., PL Aequin. 2: 63. 1809. Type:
MEXU, PTBG)), same locality, Tonduz 12997 MEXICO: [Probably Guerrero], near
(Syntype US). Not Psychotria tetragona Acapulco, Humboldt & Bonpland s.n.
Seem., 1867.—Psychotria chiapensis Standi. (Holotype presumably P-Bonpland, n.v.).
Cephaelis tomentosa (Aubl.) Vahl, Eclog.
Amer. 1: 19. 1796. Tapogomea tomentosa Cephalanthus peroblongus Wernham, J. Bot.
Aubl., Hist. PI. Guiane 1: 160, pi 61. 1775. 55: 176. 1917. Type: MEXICO: Nayarit,
Type: FRENCH GUIANA: In sylvarum locis Tepic, 31 May-3 Jun 1837, G. W. Barclay
apertis, Aublet s.n. (Holotype presumably P, 1193 (Holotype BM, n.v.; Isotype US (photos
n.v.; Isotype B, presumably destroyed (photo MEXU, PTBG)).—Cephalanthus salicifolius
MO)). Not Psychotria tomentosa Mull. Arg. Humb. & Bonpl.
ex Mart., 1881.—Psychotria poeppigiana Cephalanthus salicifolius Humb. & Bonpl., PL
Mull. Arg. Aequin. 2: 63. 1809. Type: MEXICO:
Cephaelis tonduzii K. Krause, Bot. Jahrb. Syst. [Probably Guerrero] near Acapulco,
54, Beibl. 119: 45. 1916. COSTA RICA: Bei Humboldt & Bonpland s.n. (Holotype
Tuis, 650 m, Nov 1897, A. Tonduz 11461 presumably P-Bonpland, n.v.).
(Holotype B, presumably destroyed (photos F,
PTBG; fragment F (photo PTBG)). Not Chimarrhis Jacq., Sel. Stirp. Amer. 61. 1763.
Psychotria tonduzii Standi., 1925. Type: Chimarrhis cymosa Jacq.
—Psychotria guapilensis (Standi.) Hammel Chimarrhis decurrens Steyerm., Ceiba 3:18.
Cephaelis vultusmimi Dwyer, Ann. Missouri 1952. Type: PANAMA: Code, El Valle, 600
Bot. Gard. 67: 81, fig. 17. 1980. Type: m, 28 Jun 1946, P. H. Allen 3566 (Holotype F
PANAMA: Colon, Santa Rita, E. Ridge, 23 1394240 (photo PTBG); Isotypes F, 2 sheets,
Mar 1972, A. Gentry & J. D. Dwyer 4811 MO).—Allenanthus erythrocarpa Standi.
(Holotype MO 2164626 (photo PTBG); Chimarrhis latifolia Standi., Publ. Field
Isotype MO (photo PTBG)).—Psychotria Columbian Mus., Bot. Ser. 4: 265. 1929.
poeppigiana Mull. Arg. Type: PANAMA: Chiriqui, Progreso,
Cephaelis williamsii Standi., Publ. Field Jul-Aug 1927, G. P. Cooper & G. M. Slater
Columbian Mus., Bot. Ser. 8: 185. 1930. 260 (Holotype F 573160 (photo PTBG);
Type: PERU: Loreto, Manfina, on upper Rio Isotype US (photos MEXU, PTBG)).
Nany, 29 Jun 1929, L. Williams 1114 Chimarrhis parviflora Standi., Trop. Woods
(Holotype F 604625). Not Psychotria 11: 26. 1927. Type: PANAMA: Bocas del
williamsii Standi., 1930.—Psychotria Toro, Changuinola Valley, 1927, G. P.
zevallosii C. M. Taylor Cooper & G. M. Slater 120 (Holotype US
1315890 (photos MEXU, PTBG); Isotypes A,
Cephalanthus L., Sp. PI. 1: 95. 1753. Type: n.v., F, NY, n.v.).
Cephalanthus occidentalis L.
Cephalanathus berlandieri Wernham, J. Bot. Chiococca P. Browne, Civ. Nat. Hist. Jamaica
55: 175. 1917. Type: MEXICO: Berlandier 164. 1756. Type: Chiococca alba (L.) Hitchc.
1620 (Holotype BM, n.v.).—Cephalanthus Chiococca alba (L.) Hitchc, Annual Rep.
Missouri Bot. Gard. 4: 94. 1893. Lonicera
Cephalanthus occidentalis L., Sp. PL 1: 95 alba L., Sp. PL 175. 1753. Type: JAMAICA
1753. Type: America septentrionali, Herb. or BARBADOS: According to Howard
Linn. 118.1 (Lectotype LINN, (1989), a specimen in the Sloan herbarium
by Reveal, 1993). (BM) may be an appropriate lectotype.
Chiococca axillares Sesse & Moc., La Chiococca jefensis Dwyer, Ann. Missouri Bot.
Naturaleza ser. 2, 1: 36. 1888. Type: Gard. 67: 88. 1980. Type: PANAMA:
MEXICO: Habitat in Tepetlpae prope Sancti Panama, La Eneida, region of Cerro Jefe, R.
Angeli montibus, Sesse & Mocino s.n. Dressier 3296 (Holotype MO).—Elaeagia
(Holotype presumably MA, n.v.; Fl. Mex. Ic. nitidifolia Dwyer
50 (G-DC, n.v.)).—Status uncertain. Chiococca macrocarpa M. Martens & Galeotti,
Chiococca belizensis Lundell, Amer. Midi. Bull. Acad. Roy. Bruxelles 11: 230. 1844.
Naturalist 29: 492. 1943. Type: BRITISH Type: MEXICO: Veracruz, bois de Mirador,
HONDURAS (BELIZE): Toledo Distr., Cow 3000 pieds, H. Galeotti 7064 (Holotype
Pen, near Monkey River, 2 Sep 1942, P. H. presumably BR, n.v.).—Chiococca alba (L.)
Gentle 4115 (Holotype MICH, n.v.; Isotypes Hitchc.
A, n.v., F (photo F), MO (photos MEXU, Chiococca mexicana Lundell, Wrightia 5: 71.
PTBG), NY, n.v., TEX-LL, PTBG), US 1974. Type: MEXICO: Puebla, Mun. de
(photos MEXU, PTBG)). Zacapoaxtla, Tespilco, 1800 m, 23 Apr 1970,
Chiococca caputensis Lorence & C. M. Taylor, F. Ventura A. 955 (Holotype TEX-LL (photos
Novon 5: 210, fig. IB, C. 1995. Type: MEXU, PTBG); Isotype ENCB).—Chiococca
PANAMA: Panama, Cerro Jefe, NE of pachyphylla Wernham
Panama City, 9°15'N, 79°30'W, 12 Jul 1986, Chiococca nocturna Sesse & Moc., Fl. Mexic,
G. McPherson 9744 (Holotype MO 3624428; ed. 2, 59. 1894. Type: MEXICO: Habitat in
Isotype PMA, n.v.). Tuxtlae montibus, floret Julio, Sesse &
Chiococca coriacea M. Martens & Galeotti, Mocino (Holotype presumably MA, n.v.). Not
Bull. Acad. Roy. Bruxelles 11: 231. 1844. Chiococca nocturna Jacq. 1761.—Status
Type: MEXICO: Veracruz, environs de Vera uncertain, probably Cestrum nocturnum L.
Cruz, J. Linden 423 (Holotype presumably (Solanaceae), as protologue states leaves
BR, n.v.; Isotype MICH, n.v.). alternate, flowers nocturnal, smelling of
Chiococca durifolia Dwyer, Ann. Missouri Bot. jasmine.
Gard. 67: 87. 1980. Type: PANAMA: Code, Chiococca oaxacana Standi., Publ. Field Mus.
woods adjacent to chicken farm, La Mesa, Nat. Hist., Bot. Ser. 22: 383. 1940. Type:
above El Valle, 3 Jan 1974, J. D. Dwyer MEXICO: Oaxaca, Cerro de San Antonio,
11865 (Holotype MO 2188155 (photos 1650 m, 26 Jun 1906, C. Conzatti 1418
MEXU, PTBG)).—Chiococca belizensis (Holotype F 579006 (photos F, PTBG);
Lundell Isotypes GH (fragment), US).
Chiococca filipes Lundell, Contr. Univ. Chiococca pachyphylla Wernham, J. Bot. 51:
Michigan Herb. 7: 53. 1942. Type: MEXICO: 323. 1913. Type: MEXICO: Veracruz, entre
Chiapas, Nuevo Amatenango, 1300 m, 17 Jul Acatlan et Chiconquiaco, Schiede & Deppe
1941, E. Matuda 4783 (Holotype MICH, n.v.; (Holotype K, n.v.).
Isotypes A, n.v., MO (photos MEXU, PTBG), Chiococca paniculata Sesse & M09., Fl. Mexic,
NY, n.v., TEX-LL (photos MEXU, PTBG), ed. 2, 58. 1894. Type: MEXICO: Habitat in
US (photos MEXU, PTBG)).—Chiococca calidis Cordovae montibus, floret Augusto,
alba (L.) Hitchc. Sesse & Mocino (Holotype presumably MA,
Chiococca henricksonii M. C. Johnst., n.v.). Not Chiococca paniculata Hoffmanns,
Madrono 28: 30. 1981. Type: MEXICO: ex Roem. & Schult., 1819.—Status
Coahuila, 1-2 km N. of Puerto Colorado, near
crest at SW end of Sierra de la Fragua, near Chiococca petenensis Lundell, Wrightia 5: 7.
26°45TSf, 102°30*W, 2 Sep 1941, /. M. 1972. Type: GUATEMALA: Peten, Dolores,
Johnston 8738 (Holotype TEX 301584, n.v.; in high forest on Santo Toribio trail, about
Isotypes F (photo PTBG), GH (photo PTBG), 200 m west, 7 Jun 1961, E. Contreras 2439
MICH, n.v.). (Holotype TEX-LL (photo F)).
 i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Chiococca petrina Wiggins, Contr. Dudley 265 (Holotype DS 20437, n.v.; Isotype UC
Herb. 3: 76,pi. 20, f. 5-13. 1940. Type: (n.v.)).—Status uncertain, probably
MEXICO: Sonora, gorge in andesitic rock 23 Chiococca pueblensis Lundell
miles S. of Divisadero, 26 Sep 1934, /. L. Chiococca pueblensis Lundell, Wnghtia 5: 7.
Wiggins 7493 (Holotype DS 263330, n.v.; 1972. Nom. nov. for Chiococca pubescens
Isotypes F, TEX-LL, n.v., US (photos MEXU, Standi., not Chiococca pubescens Humb. &
PTBG)). Bonpl. ex Roem. & Schultes, 1819.
Chiococca phaenostemon Schltdl., Linnaea 9: Chiococca racemosa L., Syst. Nat. ed. 10, 2:
594. 1834. Type: MEXICO: Veracruz, "In 917. 1759. Nom. illegit., citing Lonicera alba
sylvis Jalapae, Majo florens, ad Hac. d. 1. L—Chiococca alba (L) Hitchc.
Laguna Jul. fructifer," collector not cited Chiococca racemosa Sesse & Moc., Fl. Mexic,
(Holotype presumably HAL, n.v.). Note: no ed. 2, 58. 1893, nom. illegit. Not Chiococca
actual specimens or collector(s) were cited in racemosa L., 1759, nor C. racemosa Schltdl.
Schlechtendal's protologue (1834), merely "In & Cham., 1830. Type: MEXICO: Sesse &
sylvis Jalapae Majo florens, ad Hac. d. 1. Moqiho (presumably MA, n.v.).—Status
Laguna Jul. fructifer." There is a Schiede 255 uncertain, probably Chiococca alba (L.)
specimen at Paris (with a duplicate at MO) in Hitchc.
Schlechtendal's hand stating only "Mexique." Chiococca racemosa L. var. yucatana Loes.,
Schlechtendal's types are mostly at HAL, so Repert. Sp. Nov. 18: 361. 1922. Type:
the holotype [or possibly syntypes] is MEXICO: Yucatan, im Distrikt Ticul, auf der
presumably deposited there. Schiede is known Hacienda Yaxche, E. Seler & C Seler 5591
to have collected at Hacienda de la Laguna, (Holotype B, presumably destroyed).
Veracruz, and therefore the Schiede 255 —Chiococca alba (L.) Hitchc.
specimens at P & MO are very likely original Chiococca rubriflora Lundell, Wrightia 5: 7.
material, probably isotypes. However, 1972. Type: GUATEMALA: Peten, west of
mention of "Jalapa" suggests that km 165 of Poptun road, 10 Sep 1969, E
Schlechtendal may have been referring to Contreras 9110 (Holotype TEX-LL (photos
another collection (syntype) as well. This is a MEXU, PTBG); Isotype F (photos F, PTBG)).
vining species with a short, internally villous Chiococca semipilosa Standi. & Steyerm., Publ.
corolla rube and exserted stamens with Field Mus. Nat. Hist., Bot. Ser. 22: 279. 1940.
filaments villous in the basal half. Collections Type: GUATEMALA: Chiquimula, Cerro
from southern Mexico and Central America Caracol, 1.5 miles north of Quezaltepeque,
erroneously identified as Chiococca 1200-1400 m, 7 Nov 1939, J. A. Steyermark
phaenostemon have the habit of a small tree, a 31406 (Holotype F 1039351 (photo PTBG)).
glabrous corolla, and actually represent C Chiococca sessilifolia Miranda, Anales Inst.
pachyphylla Wernham. Biol. Mexico 21(2): 306,/ 4. 1950 [1951].
Chiococca pubescens Standi., Contr. U.S. Natl. Type: MEXICO: Chiapas, en borde de bosque
Herb. 20: 209. 1919. Type: MEXICO: Puebla, bajo hacia Las Vistas, al N de Berriozabal, 24
vicinity of San Luis Tultitlanapa, Jul 1908, Sep 1950, F. Miranda 6672 (Holotype
C. A. Purpus 3334 (Holotype US 840975 MEXU).
(photos MEXU, PTBG); Isotypes F, P). Not
Chiococca pubescens Humb. & Bonpl. ex Bull. Acad. Bruxelles 11: 231. 1844. Type:
Roem. & Schultes, 1819.—Chiococca MEXICO: Veracruz, environs de Xalapa,
4000 pieds, / Linden 48 (Holotype BR, n.v).
Chiococca pubescens Standi, var. peninsularis —Status uncertain, probably Chiococca
Wiggins, Contr. Dudley Herb. 4: 24. 1950. phaenostemon Schltdl.
Type: MEXICO: Baja California, Sierra San Chiococca steyermarkii Standi., Publ. Field
Francisquito, 18 Oct 1890, T. S. Brandegee Mus. Nat. Hist., Bot. Ser. 22: 280. 1940.
 Type: GUATEMALA: Dept. San Marcos, Rio Chione chiapasensis Standi., Publ. Field Mus.
Vega, near San Rafael and Guatemala-Mexico Nat. Hist, Bot. Ser. 22: 110. 1940. Type:
boundary, Volcan de Tacana, 2500-3000 m, MEXICO: Chiapas, Palenque, Rvina [Ruina],
20 Feb 1940, J. A. Steyermark 36237 river side in virgin forest, Jul 1939, E. Matuda
(Holotype F 1043222 (photos F, PTBG); 3675 (Holotype F 1011456 (photo PTBG);
Isotype F 1043229). Isotypes A, n.v., US (photos MEXU, PTBG),
Chiococca vestita Lundell, Wrightia 5: 8. 1972. MEXU).
Type: GUATEMALA: Dept. Peten, Poptun, 2 Chione costaricensis Standi., Publ. Field Mus.
km east, in hammock in pineland, 13 Jul Nat. Hist., Bot. Ser. 22: 111. 1940. Type:
1959, C L. Lundell 16425 (Holotype TEX- COSTA RICA: Alajuela, Canton de San
LL, n.v. (photo F)).—Chiococca semipilosa Carlos, Villa Quesada, Caribbean rain forest,
Standi. & Steyerm. 825 m, 14 Mar 1939, Austin Smith F1778
Chiococca vestita Lundell var. glaberrima (Holotype F 980765 (photo PTBG); Isotype
Lundell, Wrightia 5: 8. 1972. Type: US (photos MEXU, PTBG)).—Chione
GUATEMALA: Dept. Peten, Poptun, 2 km sylvicoia (Standi.) W. C. Burger
east, in hammock in pineland, 13 Jul 1959, Chione dariensis Dwyer, Ann. Missouri Bot.
C L. Lundell 16425A (Holotype TEX-LL, Gard. 67: 94. 1980. Type: PANAMA: Darien,
n.v. (photo F)).—Chiococca semipilosa Rio Balsa, between Manene and Tusijuanda,
Standi. & Steyerm. 26 Jul 1967, J. Duke 13571 (Holotype MO
1968196 (photos MEXU, PTBG); Isotypes
Chione DC, Prodr. 4: 461. 1830. Type: Chione MO, 2 sheets, US (photo PTBG)).
glabra DC. Chione guatemalensis Standi. & Steyerm.,
Chione allenii L. O. Williams, Phytologia 25: Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 174.
462. 1973. Type: COSTA RICA: Golfo Dulce 1940. Type: GUATEMALA: Dept. Izabal,
area, vicinity of Esquinas Experiment Station, Rio Dulce, between Livingston and 6 miles up
sea level, 8 Jul 1949, P. H. Allen 5321 river, on north side, 1-25 m, 14 Apr 1940,
(Holotype F 1679433; Isotypes MO (photo J. A. Steyermark 39375 (Holotype F 1035017
PTBG), TUL (photo PTBG)).—Chione (photo PTBG)).
sylvicoia (Standi.) W. C. Burger Chione mexicana Standi., J. Wash. Acad. Sci.
Chione buxifolia Dwyer & Hayden, Phytologia 17: 340. 1927. Type: MEXICO: Veracruz,
15: 55. 1967. Type: PANAMA: Panama, Zacuapan, open forests, Jun 1926, C. A.
Cerro Jefe, E. Tyson et al. 3291 (Holotype Purpus 10757 (Holotype US 1266079 (photos
MO 2210380 (photo PTBG); Isotypes F, NY, MEXU, PTBG); Isotypes A, n.v., F, 2 sheets,
n.v.). MO, 2 sheets (photos MEXU, PTBG)).
Chione campanensis Dwyer, Ann. Missouri Chionepanamensis Steyerm., Ceiba 3: 19.
Bot. Gard. 67: 92. 1980. Type: PANAMA: 1952. Type: PANAMA: Chiriqui, Boquete
Cerro Campana, trail to Campana Ridge, 2300 region, Cerro Horqueta, 6500 ft., 1 Jul 1940,
ft., 18 Sep 1968, A R. Wilbur, R. Foster, C. von Hagen & W. von Hagen 2137
H. Kennedy & R. Dressier 1020 (Holotype (Holotype F 1397595 (photo PTBG); Isotype
MO 2224284 (photo PTBG)). MO, n.v.).—Chione sylvicoia (Standi.) W. C.
Chione chambersii Dwyer & Hayden, Ann.
Missouri Bot. Gard. 54: 138. 1967. Type: Chione sylvicoia (Standi.) W. C. Burger,
PANAMA: Comarca de Bani, area near Selbyana 12: 138. 1991. Chomelia ? sylvicoia
Puerto Armuelles, between Canazo and Standi., J. Wash. Acad. Sci. 18: 182. 1928.
Cocos, 17 Jun 1957, W. I. Stern & K L. Type: COSTA RICA: Heredia, Yerba Buena,
Chambers 144 (Holotype MO 1739838; northeast of San Isidro, 2000 m, 22-28 Feb
Isotype GH (photo PTBG)).—Chomelia 1926, P. C. Standley & J. Valerio 49196
i Donn. Sm. (Holotype US 1305902 (photo PTBG)).
 i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Chomelia Jacq., Enum. PI. Carib. 12. 1760. C, n.v. (photo US), fragment US (photo
Type: Chomelia spinosa Jacq. PTBG)).—Chomelia spinosa Jacq.
Chomelia atlantica Dwyer, Ann. Missouri Bot. Chomelia grandicarpa Dwyer, Ann. Missouri
Gard. 67: 96. 1980. Type: PANAMA: Colon, Bot. Gard. 67: 97. 1980. Type: PANAMA:
swamp forest and bordering terra firma, ca. Colon, 2-3 miles up Rio Guanche, lowland
0.5 km from the Rio Buenaventura, near rain forest, 10-20 m, 19 Jan 1973,
Portobelo, 30 Jan 1973, H. Kennedy & S. Gra H. Kennedy & R. Foster 2174 (Holotype MO.
2242 (Holotype MO 2161971 (photo PTBG); 2465399 (photo PTBG); Isotype MO).
Isotype MO)).—Chomelia barbellata Standi. Chomelia leucophylla Dwyer, Ann. Missouri
Chomelia barbata Standi, J. Wash. Acad. Sci. Bot. Gard. 67: 98. 1980. Type: PANAMA:
13: 7. 1923. Type: MEXICO: Oaxaca, Darien, slopes of Cerro Chucula, drainage of
Juquila, vicinity of Chacahua, 5 m, 17 Dec Rio Pavarando, 11 Feb 1972, A. Gentry 4262
1921, C. Conzatti 4475 (Holotype US (Holotype MO; Isotype MO). Not Rondeletia
1110841 (photo US)). leucophylla Kunth, 1820.—Rondeletia
Chomelia barbellata Standi., Field Mus. Nat. dwyeri Lorence
Hist., Bot. Ser. 13(7): 120. 1936. Type: Chomelia microloba Donn. Sm., Bot. Gaz.
PERU: San Martin, Klug 3920 (Holotype F, (Crawfordsville) 31: 114. 1901. Type:
n.v.). COSTA RICA: Santo Domingo de Golfo
Chomelia brachyloba Standi., Contr. U.S. Natl. Duke, Feb 1896, A. Tonduz 7048 (Herb. Nac.
Herb. 18: 136. 1916. Type: PANAMA: Costa Rica 9874) (Holotype US 943460
Panama, Sabana de San Juan Corso, near (photo PTBG); Isotypes A, n.v., MO (photos
Chepo, 60-80 m, Oct 1911, H. Pittier 4673 MEXU, PTBG), P (photos MEXU, PTBG),
(Holotype US 679833 (photo PTBG)). US (3 sheets; photos PTBG)). Anisomeris.
—Chomelia spinosa Jacq. Chomelia panamensis (Standi.) Dwyer, Ann.
Chomelia brachypoda Donn. Sm., Bot. Gaz. Missouri Bot. Gard. 67: 100. 1980. Antirhea
(Crawfordsville) 47: 255. 1909. Type: panamensis Standi., N. Amer. Fl. 32: 264.
GUATEMALA: Alta Verapaz, Sasis, banks of 1934. Type: PANAMA: San Bias District,
Rio Ogewaj, 900 m, May 1908, H. von 21-28 Apr 1933, G. P. Cooper 238 (Holotype
Tuerckheim II2253 (Holotype US 941389 F 686345 (photo PTBG); Isotype F 709955).
(photos PTBG, US); Isotypes F (photos —Chomelia microloba Donn. Sm.
MEXU, PTBG), US (photo PTBG), W). Chomelia peninsularis Dwyer, Ann. Missouri
Anisomeris. Bot. Gard. 67: 101. 1980. Type: PANAMA:
Chomelia coclensis Dwyer, Ann. Missouri Bot. Chiriqui, Burica Peninsula, along Quebrada
Gard. 67: 97. 1980. Type: PANAMA: Code, Punta de Piedra, 2 mi SE of Puerto Armuelles,
La Mesa above El Valle, 800 m, T. B. Croat 0-100 m, 1 Mar 1973, T. B. Croat 22440
25396 (Holotype MO 2210285; Isotype MO (Holotype MO 2785727 (photo PTBG);
2219529).—Malanea colombiana Standi. Isotype F).—Chomelia microloba Donn. Sm.
Chomelia englesingii Standi., Trop. Woods 16: Chomelia pringlei S. Watson, Proc. Amer.
45. 1928. Type: NICARAGUA: Region of Acad. Arts 26: 137. 1891. Type: MEXICO:
Braggman's Bluff, Kukalaya River bank west San Luis Potosi, Tamasopo Canyon, 6 Aug
of Camp 14, 2 Dec 1927, F. C Englesing 49 1890, C. G. Pringle 3209 (Holotype GH, n.v.;
(Yale 1233) (Holotype F 572606 (photo Isotypes A, n.v., F (photos MEXU, PTBG),
PTBG)). Anisomeris.—Chomelia recordii MO (photos MEXU, PTBG), P (photos
Standi. MEXU, PTBG), US). Anisomeris.
Chomelia filipes Benth. in Oerst., Vidensk. Chomelia protracta (Bartl. ex DC.) Standi.,
Meddel Dansk Naturhist. Foren. Kjobenhavn Contr. U.S. Natl. Herb. 23: 1384. 1926.
1852: 41. 1853. Type: NICARAGUA: Guettarda protracta Bartl. ex DC. Prodr. 4:
Granada, Oersted s.n. (Holotype presumably 457. 1830. Type: MEXICO: Without precise
 locality, Haenke s.n. (Holotype presumably Chomelia venulosa W. C. Burger & C. M.
PR, n.v.). Anisomeris, Pittoniotis. Taylor, Fieldiana, Bot, n.s. 33: 101. 1993.
Chomelia psilocarpa Dwyer & Hayden, Ann. Type: COSTA RICA: Alajuela, Reserva
Missouri Bot. Gard. 54: 139. 1967. Type: Biologica Monteverde, E. Bello 414
PANAMA: Canal Zone, Barro Colorado (Holotype CR, n.v.; Isotypes F, n.v., MO).
Island, SE shoreline from lab, Aug 1965,
M. V. Hayden 143 (Holotype MO 1831286 Choristes Benfh., PI. Hartw. 63. 1840. Type:
(photo PTBG); Isotypes MO, 3 sheets). Choristes obtusiflora Benth. [Herein treated as
Chomelia purpusii (Brandegee) Rusby, Bull. a synonym of Deppea Cham. & Schltdl.]
Torrey Bot. Club 52: 138. 1925. Anisomeris Choristes cornifolia (Benth.) Benth., PI. Hartw.
purpusii Brandegee, Univ. Calif. Publ. Bot. 6: 64. 1840. Rondeletia cornifolia Benth., PI.
70. 1914. Type: MEXICO: Chiapas, Tonala, Hartw. 39. 1840. Type: MEXICO:
Sep 1913, C. A. Purpus 6940 (Holotype UC, Michoacan, prope Morelia, T. Hartweg 299
n.v.; Isotypes F (photo PTBG), US (photo (Holotype K (photo MEXU); Isotypes BM, P,
PTBG))./ " - US (fragment), W, 2 sheets).—Deppea
cornifolia (Benth.) Benth.
i Standi., Trop. Woods 7: 9. Choristes obtusiflora Benth., PI. Hartw. 63.
1926. Type: GUATEMALA: Izabal, Los 1840. Type: MEXICO: Oaxaca, in plamtie
Andes to Entre Rios, lower Rio Motagua Llano Verde, T. Hartweg 473 (Holotype K;
Valley, 1 Mar 1926, S. J. Record 8862 Isotypes P, F, W, 2 sheets).—Deppea
(Holotype US 1209919 (photo PTBG); obtusiflora (Benth.) Benth.
Isotype NY, n.v.). Anisomeris.
Chomelia recordii Standi, var. paucipubescens Cigarrilla Aiello, J. Arnold Arbor. 60: 109.
Dwyer, Ann. Missouri Bot. Gard. 67: 103. 1979. Type: Cigarilla mexicana (Zucc. &
1980. Type: PANAMA: Canal Zone, near Mart, ex DC.) Aiello. [Herein treated at a
archeological site, edge of Madden Lake, A. synonym of Nernstia Urb.]
Gentry & E. Tyson 5038 (Holotype MO (not Cigarrilla mexicana (Zucc. & Mart, ex DC.)
located 6/96)).—Chomelia recordii Standi. Aiello, J. Arnold Arbor. 60: 110. 1979.
Chomelia rudis (Standi.) Lorence, Novon 7: Coutarea mexicana Zucc. & Mart, ex DC,
48. 1997. Anisomeris rudis Standi., Field Mus Prodr. 4: 350. 1830. Type: MEXICO: Without
Nat. Hist, Bot. Ser. 17: 394. 1938. Type: precise locality, Martius s.n. in 1829
HONDURAS: Comayagua, summit of the (Holotype G-DC, n.v. (microfiche IDC
ridge above El Achote, above the plains of 800.680: II.7)).—Nernstia mexicana (Zucc.
Siguatepeque, 1800 m, 1 Aug 1936, G. T. & Mart, ex DC.) Urb.
Yuncker, R. F. Dawson & H. R. Youse 6253
(Holotype F 857678 (photo PTBG); Isotype Cinchona L., Sp. PI. 1: 172. 1753. Type:
\I< ) 1 Cinchona officinalis L.
Chomelia spinosa Jacq., Enum. PI. Carib 12 Cinchona caribaea Jacq., Enum. PI. Carib. 16.
1760. Type: COLOMBIA: Cartagena, in 1760. Type: CUBA: Thickets near Havana,
sylvaticis ad pendem montis de La Po'pa, Jacquin s.n. (location unknown).—Exostema
Jacquin s.n. (location unknown). Ixora. ' caribaeum (Jacq.) Roem. & Schultes
Chomelia sylvicola Standi., J. Wash Acad Sci Cinchona grandiflora Ruiz & Pav., F. Peruv. 2:
18: 182. 1928. Type: COSTA RICA: Heredia, 54, pi. 89. 1799. Type: PERU: Huanuco,
Yerba Buena, northeast of San Isidro 2000 m Andes de Peru, ad Pozuzo et San Antonii de
22-28 Feb 1926, P. C Standley & J Valeria ' Playa grande, Ruiz Lopez & Pavon s.n.
49196 (Holotype US 1305902 (photo (Holotype presumably MA, n.v.; Isotype B,
PTBG)). Anisomeris.—Chione sylvicola destroyed (photo P)).—Cosmibuena
(Standi.) W. C. Burger grandiflora (Ruiz & Pav.) Rusby
Cinchona macrocarpa Vahl, Schr. Naturh. Ges. Huitamulco et Cuapa, Schiede & Deppe 1273
^Copenhagen 1(1): 20. 1793. Type: (Holotype presumably HAL, n.v.).—Status
COLOMBIA: In regno Santa Fe, Ortega s.n. uncertain, probably Coccocypselum
(Lectotype BM, n.v.; designated by hirsutum Bartl. ex DC.
Andersson in Andersson & Taylor, 1994: 16). Coccocypselum cordifolium Nees & Mart.,
—Ladenbergia macrocarpa (Vahl) Klotzsch Nova Acta Phys.-Med. Acad. Caes. Leop.-
Cinchona officinalis L., Sp. PI. 1: 172. 1753. Carol. Nat. Cur. 12: 14. 1824. Type: BRAZIL:
Type: ECUADOR: Habitat in Loxa Pemviae, Ad viam Felisbertiam, Wied-Neuwied
Quinquina La Condamine, Hist. Acad. Roy. (location unknown).
Sci. Paris 1738: 226, t. 5-6. 1738. (For Coccocypselum glabrum Bartl. ex DC, Prodr. 4:
typification see Andersson & Taylor, 1994: 397. 1830. Type: PANAMA: Without precise
locality, Haenke s.n. (Holotype presumably
Cinchona pubescens Vahl, Skr. Naturhist. PR, n.v., fragment G-DC, n.v. (microfiche 4:
Selsk. Kjoebenhavn 1: 119. 1790. Type: 397.14)). Tontanea.—Coccocypselum
Probably ECUADOR: precise locality hirsutum Bartl. ex DC. var. glabrum (Bartl.
unknown, J. deJussieu s.n. (Holotype C, n.v. ex DC.) L. O. Williams
(photos F, MO); Isotype P-JU 9900, n.v.). Coccocypselum guianense (Aubl.) K. Schum.
Cinchona rosea Ruiz & Pav., Fl. Peruv. 2: 54. in Mart., Fl. Bras. 6(6): 315. 1889. Tontonea
1799. Type: PERU: Huanuco, Pozuzo, Ruiz guianensis Aubl., Hist. PI. Guiane 1: 108, tab.
Lopez & Pavon s.n. (Lectotype MA, n.v.; 42. 1775. Type: FRENCH GUIANA: Sylvis
Isolectotypes BM, n.v., G, n.v., G-DC, n.v., Cayenne, Aublet s.n. (Holotype presumably P
K, n.v., MA, n.v., P, n.v.; designated by or BM, n.v.).
Andersson in Andersson & Taylor, 1994: 12). Coccocypselum herbaceum P. Browne, Civ.
—Macrocnemum roseum (Ruiz & Pav.) Nat. Hist. Jamaica 144, pi. 6 J. 2. 1756. Type:
Wedd. JAMAICA: Cooler mountains of Liguancee
Cinchona succirubra Pav. ex Klotzsch, Abh. and Mount-Diable, P. Browne (LINN, n.v.).
Akad.Berl. 1857: 60. 1858. Type: Tontanea.
ECUADOR: Bolivar, San Antonio on trail to Coccocypselum hirsutum Bartl. ex DC, Prodr.
Guaranda, [Tafalla in] Pavon 464 (Holotype 4: 396. 1830. Type: Precise locality unknown,
B, presumably destroyed (photo F); Isotypes probably Trinidad, Haenke s.n. (Holotype
G, n.v., K, n.v., MA, n.v.).—Cinchona presumably PR, n.v., fragment G-DC, n.v.
pubescens Vahl (microfiche 4: 396.5)). Tontanea.
Coccocypselum hirsutum var. glabrum (Bartl.
Coccocypselum P. Browne, Civ. Nat. Hist. ex DC.) L. O. Williams, Phytologia 25: 462.
Jamaica 144. 1756. [as Coccocipsilum], nom. Coccocypselum glabrum Bartl. ex DC, Prodr.
et orthogr. cons. Type: Coccocypselum repens 4: 397. 1830. Type: PANAMA: Without
Sw., Prodr. Veg. Ind. Occ. 31. 1788, not precise locality, Haenke s.n. (Holotype
Kunth, 1819. presumably PR, n.v., fragment G-DC, n.v.
Coccocypselum canescens Willd. ex Cham. & (microfiche 4: 397.14)).
Schltdl., Linnaea4: 139. 1829. Nom. Meg., Coccocypselum hispidulum (Standi.) Standi.,
not Coccocypselum canescens Willd. ex Publ. Field Columbian Mus., Bot. Ser. 4: 281.
Schult. & Schult., 1827. Type: VENEZUELA: 1929. Tontanea hispidula Standi., N. Amer.
Buenavista prope Caracas, Humboldt s.n. Fl. 32: 147. 1921. Type: PANAMA: Chagres,
(Holotype B-W 2874, n.v). Tontanea. Isthmus of Panama, Jan-Feb 1850, A. Fendler
—Coccocypselum lanceolatum (Ruiz & 179 (Holotype US 42058 (photo MO); Isotype
Pav.) Pers. US).
Coccocypselum ciliatum Cham. & Schltdl.,
Linnaea 6: 414. 1831. Type: MEXICO: Inter
Coccocypselum lanceolatum (Ruiz & Pav.) Trans. Linn. Soc, Ser. 2: 171, tab. 24. 1876.
Pers., Syn. PL 1: 132. 1805. Condalia Type: SIERRA LEONE: Daniell A608
lanceolata Ruiz & Pav., PL Fl. Peruv. 1: 54. (Holotype BM, n.v. (photo MO)).
1798. Type: PERU: In nemoribus Cuchero et Occasionally cultivated in Mexico and Central
Chinchao, Ruiz & Pavon s.n. (Holotype
presumably MA, n.v.). ( offea mexicana DC, Prodr. 4: 501. 1830.
Coccocypselum macropodum (Ruiz & Pav.) Type: MEXICO: Without precise locality,
Britton, Bull. Torrey Bot. Club 18: 107. 1891. Haenke s.n. (Holotype presumably PR, n.v.,
Psychotria macropoda Ruiz & Pav., Fl. fragment G-DC, n.v. (microfiche 4: 501.25)).
Peruv. 2: 63, pi 211, fig. b. 1799. Type: —Rudgea cornifolia (Humb. & Bonpl.)
PERU: Huanuco, nemoribus secus Pillao ad Standi.
Iscutunam, Ruiz & Pavon s.n. (Holotype Coffea obovata Cham. & SchltdL, Linnaea 6:
presumably MA, n.v.).—Geophila 412. 1832. Type: MEXICO: Veracruz, m
macropoda (Ruiz & Pav.) DC. umbrosis prope Misantla, Schiede & Deppe
Coccocypselum pleuropodum (Donn. Sm.) 1268 (Holotype presumably HAL, n.v.).
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. —Status uncertain, probably Hoffmannia
4: 281. 1929. Geophila pleuropoda Donn. sp.
Sm., Bot. Gaz. (Crawfordsville) 52: 50. 1911. Coffea rosea Moc. & Sesse ex DC, Prodr. 4:
Type: GUATEMALA: Alta Verapaz, 499. 1830. Type: MEXICO: Without precise
Secanquim, trail to Sepacuite, 4000 ft., 5 Feb locality, Sesse & Mocino s.n. (presumably G-
1905, G. P. Golls.n. (Holotype US 860647 DC, n.v.; Fl. Mex. Ic. (G-DEL, n.v. (photo
(photos MEXU, PTBG)).—Coccocypselum MO), caique 479 (US)).—Status uncertain,
cordifolium Nees & Mart. probably I
Coccocypselum rothschuhii Loes., Bot. Jahrb.
Syst. 60: 370. 1926. Type: NICARAGUA: Condalia Ruiz & Pav., Prodr. II,pi. 2. 1794.
Matagalpa, Canada Yasica, 950 m, Rothschuh Nomen rejic, not Condalia Cav. (1799)
179 (Holotype B, presumably destroyed). (Rhamnaceae). Type: Condalia repens Ruiz &
—Coccocypselum cordifolium Nees & Mart.
Coccocypselum P. Browne.]
Coffea L., 172. 1753. Type: Coffea Ruiz & Pav., Fl. Peruv.
Chil. 1: 54. 1798. Type: PERU: In nemoribus
Coffea arabica L., Sp. PL 1: 172. 1753. Type: Cuchero et Chinchao, Ruiz Lopez & Pavon
Habitat in Arabia felici, Herb. Clifford 9, s.n. (Holotype presumably MA, n.v.).
Coffea 1 (Lectotype BM, n.v.; designated by —Coccocypselum lanceolatum (Ruiz &
Bridson & Verdcourt, 1988: 713). Widely Pav.) Pers.
cultivated in Mexico and Central America.
Coffea canephora Pierre ex Froehner, Notizbl. Condaminea DC, Prodr. 4: 402. 1830.
Bot. Gard. Berlin 1: 237. 1897. Type: Lectotype: Condaminea corymbosa (Ruiz &
GABON: Gabun, Herb. L. Pierre n. 247 Pav.) DC.
(Holotype presumably P, n.v.). Occasionally Condaminea corymbosa (Ruiz & Pav.) DC,
cultivated in Mexico and Central America. Prodr. 4: 402. 1830. Macrocnemum
Coffea lanceolata Cham. & SchltdL, Linnaea 6: corymbosum Ruiz & Pav., Fl. Peruv. Prodr. 2:
412. 1831. Type: MEXICO: Veracruz, prope 4S, pi. 189. 1799. Type: PERU: In Andium
Jalapam, Schiede & Deppe 1269 (Holotype
presumably HAL, n.v.).—Status uncertain,
possibly Chiococca sp.
Coffea liberica W. Bull ex Hieron., Retail List
New, Beautiful and Rare Plants 97: 4. 1874;
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Condaminea petiolata Dwyer, Ann. Missouri Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.
Bot. Gard. 67: 117. 1980. Type: PANAMA 22: 51. 1940. Type: MEXICO: Chiapas,
Panama, Rio Maje, near new Bayano Lake, Volcan Tacana, 2000^039 m, Aug 1938, E.
<100 m, 5 May 1976, T. B. Croat 34706 Matuda 2327 (Holotype F 980255 (photo F);
(Holotype MO 2398904 (photo PTBG): Isotypes A, n.v., NY, n.v., US (photos
Isotype MO). Rondeletia [H MEXU, PTBG)).
Cosmibuena ovalis Standi., Contr. U.S. Natl.
Cordiera A. Rich., Mem. Soc. Hist. Nat., Paris Herb. 18: 137. 1916. Type: PANAMA: Code,
5: 222, t. 20. 1830. (Herein considered as a vicinity of Ola, 100-350 m, 7-9 Dec 1911,//.
synonym of AI'ibertia A. Rich.) Pittier5074 (Holotype US 715202 (photos F,
Cordiera edulis (Rich). Kuntze, Rev. Gen. PI. 1: MEXU, PTBG); Isotype NY, n.v.).
279. 1891. Genipa edulis Rich., Act. Soc. —Cosmibuena grandiflora (Ruiz & Pav.)
Hist. Nat. Paris 1: 107. 1792. Type: FRENCH Rusby
GUIANA: Guyane, without precise locality, Cosmibuena paludicola Standi., Contr. U.S.
Richard s.n. (Holotype P, n.v.; Isotype F Natl. Herb. 18: 137. 1916. Type: PANAMA:
971706 (photo PTBG)).—Alibertia edulis Colon, vicinity of Viento Frio between Playa
(Rich.) A. Rich. Damas and Viento Frio, along beach and near
sea level, in mangrove formation, 7-8 Aug
Cosmibuena Ruiz & Pav., Fl. Peruv. 3: 2. 1802, 1911, H. Pittier 4107 (Holotype US 679204
nom. cons. Type: Cosmibuena obtusifolia (photos F, MEXU, PTBG); Isotypes GH, n.v.,
Ruiz & Pav., nom. illeg., type. cons. [= NY, n.v.)—Cosmibuena macrocarpa
Cosmibuena grandiflora (Ruiz & Pav.) (Benth.) Klotzsch
Rusby]. Not Cosmibuena Ruiz & Pav., 1794, Cosmibuena skinneri (Oerst.) Hemsl., Biol.
Centr.-Amer., Bot. 2: 12. 1881. Buena
iz & Pav.) Rusby, skinneri Oerst., Vidensk. Meddel. Dansk
Bull. New York Bot. Gard. 4: 368. 1907. Naturhist. Foren Kjobenhavn 1852: 48. 1853.
Cinchona grandiflora Ruiz & Pav., Fl. Peruv. Type: NICARAGUA: In monte ignivomo,
2: 54,/?/. 89. 1799. Type: PERU: Huanuco, Masaya, Jan 1848, Oersted 11134 (Holotype
Andes de Peru, ad Pozuzo et San Antonii de C, n.v. (photos A, F, MO); Isotype US (photos
Playa grande, Ruiz Lopez & Pavon s.n. MEXU, PTBG))—Cosmibuena grandiflora
(Holotype presumably MA, n.v.; Isotype B, (Ruiz & Pav.) Rusby
destroyed (photo F)). Cosmibuena valerii (Standi.) C. M. Taylor,
Cosmibuena holdridgei Monach., Phytologia 3: Ann. Missouri Bot. Gard. 79: 897. 1992.
64. 1949. Type: GUATEMALA: Hillia valerii Standi., J. Wash. Acad. Sci. 18:
Huehuetenango, Barillas to Tokija, Jul 1948, 164. 1928. Type: COSTA RICA: San Jose,
L. R. Holdridge s.n. (Monachino 519) vicinity of La Palma, on the road to La
(Holotype NY 3187, n.v. (photo NY); Isotype Hondura, 1500-1700 m, 17-18 Jul 1923,
NY, n.v. (photo NY)).—Cosmibuena W. R. Maxon & A. D. Harvey 8092 (Holotype
matudae (Standi.) L. O. Williams US 1181776 (photos F, MEXU, PTBG)).
Cosmibuena macrocarpa (Benth.) Klotzsch ex
Walp., Repert. Bot. Syst. 6: 69. 1846. Buena Cosmocalyx Standi., Publ. Field Mus. Nat. Hist.,
macrocarpa Benth., Bot. Voy. Sulphur 104, /. Bot. Ser. 8: 56. 1930. Type: Cosmocalyx
38, f. 1-14. 1845. Not Buena macrocarpa spectabilis Standi.
Wedd. 1869. Type: COLOMBIA: Narino, Cosmocalvx spectabilis Standi., Publ. Field
Island of Gorgona, Hinds 353 (Holotype K, Mus. Nat. Hist., Bot. Ser. 8: 56. 1930. Type:
n.v. (photo NY)). MEXICO: Yucatan, without precise locality,
Cosmibuena matudae (Standi.) L. O. Williams, 1917-1921, G. F. Gaumer 24270 (Holotype F
Fieldiana, Bot. 31: 45. 1965. Hillia matudae
 vicinity of Yaviza, 100 ft., 9-11 Jun 1947,
P. H. Allen 4576 (Holotype F 1397594 (photo
PTBG); Isotypes MO, NY, n.v.).—Coussarea
Coussarea Aubl., Hist. PI. Guiane 1: 98. 1775. paniculata (Vahl) Standi.
Type: Coussarea violacea Aubl. Coussarea durifolia Dwyer, Ann. Missouri
Coussarea austin-smithii Standi., Publ. Field Bot. Gard. 67: 124. 1980. Type: PANAMA:
Mus. Nat. Hist, Bot. Ser. 18: 1286. 1938. Panama, along El Llano-Carti Tupile road, 12
Type: COSTA RICA: Region of Zarcero, miles above Panamerican Highway, 200-500
4700 ft., 25 Aug 1937, A. Smith 4226 m, 26-27 Mar 1973, R. Liesner 1180
(Holotype F 905233 (photos MEXU, PTBG)). (Holotype MO 2219526 (photo PTBG)).
Coussarea caroliana Standi., Publ. Field Mus. Coussarea enneantha Standi, J. Wash. Acad.
Nat. Hist., Bot. Ser. 22: 178. 1940. Type: Sci. 18: 282. 1928. Type: PANAMA: Darien,
COSTA RICA: Alajuela, Canton San Carlos, Cana and vicinity, 2000-6500 ft., 17 Apr-8
Suerre, 1000 m, 2 Mar 1939, A. Smith HI645 Jun 1908, R. S. Williams 841 (Holotype US
(Holotype F 981032 (photo PTBG); Isotype 678317 (photos MEXU, PTBG)).
US (photos MEXU, PTBG)). Coussarea euryphylla Standi., Publ. Field Mus.
Coussarea cerroazulensis Dwyer & Hayden, Nat. Hist., Bot. Ser. 4: 294. 1929. Type:
Ann. Missouri Bot. Gard. 53: 374. 1966. PANAMA: Bocas del Toro, Buena Vista
Type: PANAMA: Panama, Cerro Azul, 600 camp on Chiriqui trail, region of Almirante,
m, 6 Aug 1961, J. D. Dwyer 1368 (Holotype Jan-Mar 1928, G. P. Cooper 228 (Holotype F
MO 1810161 (photos MEXU, PTBG)). 579282). Dukea, Raritebe.—Raritebe
Coussarea chiriquiensis (Dwyer) C. M. Taylor palicoureoides Wernham subsp.
in W. C. Burger & C. M. Taylor, Fieldiana, dwyerianum J. H. Kirkbr.
Bot., n.s. 33: 113. 1993. Rudgea chiriquiensis Coussarea garciae Standi., Publ. Field Mus.
Dwyer, Ann. Missouri Bot. Gard. 67: 476. Nat. Hist., Bot. Ser. 22: 180. 1940. Type:
1980. Type: PANAMA: Chiriqui, Cerro COLOMBIA: Choco, Bahia Solano, along
Colorado, along road above San Felix, 29 km Quebrada Jellita, 50-100 m, 22 Feb 1939,
above bridge over Rio San Felix (7.9 km E. P. Killip & H. Garcia 33524 (Holotype F
above turnoff to Escopeta), 1500 m, 14 Jul 999258 (photo PTBG); Isotype US (photo
1976, T. B. Croat 37071 (Holotype MO PTBG)).
2798805 (photo PTBG); Isotype MO). Coussarea hondensis (Standi.) C. M. Taylor &
—Psychotria nebulosa (Dwyer) C. M. Taylor W. C. Burger, Selbyana 12: 138. 1991.
Coussarea colonis Standi., Publ. Field Mus. Nat. Psychotria hondensis Standi., J. Wash. Acad.
Hist., Bot. Ser. 22: 179. 1940. Type: Sci. 18: 183. 1928. Type: COSTA RICA: Bois
PANAMA: Bocas del Toro, Buena Vista, de Rio Hondo, versant atlantique, 100 m, Aug
region of Almirante, Jan-Mar 1928, G. P. 1901,//. Pittier 16161 (Holotype US 764151
Cooper 228 (Holotype F 579282 (photo (photos MEXU, PTBG)).
PTBG)). Norn, superfl. for Coussarea Coussarea imitans L. O. Williams, Phytologia
euryphylla Standi., 1929 —Raritebe 26: 488, fig. 1973. Type: GUATEMALA:
palicoureoides Wernham subsp. Izabal, along trail beginning from mile 33.23
dwyerianum J. H. Kirkbr. between Dartmouth and Moralaes toward
Coussarea curvigemma Dwyer, Phytologia 38- Lake Izabal, Montana de Mico, 30-150 m, 7
215. 1978. Type: PANAMA: Barro Colorado Apr 1940, J. A. Steyermark 39013 (Holotype
Island, Wheeler Trail 850, 1 Jun 1971, T. B. F 1042963 (photos MEXU, PTBG)).
Croat 14863 (Holotype MO 2050013 (photo Coussarea impetiolaris Donn. Sm., Bot. Gaz.
PTBG)). (Crawfordsville) 37: 418. 1904. COSTA
Coussarea dariensis Steyerm., Ceiba 3: 20. RICA: La Colombiana, Llanuras de Santa
1952. Type: PANAMA: Darien, Rio Chico, Clara, 200 m, Jun 1899, H. Pittier 7582
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

(Holotype US 943498 (photos MEXU, Isotype MO 4043108 (photo PTBG)).

PTBG); Isotype US 943497 (photos MEXU, —Psychotria nebulosa (Dwyer) C. M. Taylor
PTBG)). Coussarea neei Dwyer, Ann. Missouri Bot.
Coussarea jefensis Dwyer, Ann. Missouri Bot. Gard. 67: 132. 1980. Type: PANAMA:
Card. 67: 128. 1980. Type: PANAMA: Veraguas, 6-7 km W of Santa Fe on new road
Panama, Cerro Jefe, ca. 1000 m, 25 Aug past agriculture school, 2900 ft., 17 Feb 1974,
1975, S. Mori & L. Mi 7927 (Holotype MO M. Nee 9820 (Holotype MO 2241791 (photo
2285634 (photo PTBG)). PTBG); Isotype MO 2241792 (photo PTBG)).
Coussarea latifolia Standi., J. Wash. Acad. Sci. Coussarea nigrescens C. M. Taylor & Hammel,
18: 281. 1928. Type: COSTA RICA: Forets Selbyana 12: 135, fig. 1. 1991. Type: COSTA
de Tsaki, Talamanca, 200 m, Apr 1895, A. RICA: Heredia, Finca La Selva, the OTS field
Tonduz 9574 (Holotype US 938840 (photos station near Puerto Viejo de Sarapiqui, near
MEXU, PTBG)). the junction of the Rios Puerto Viejo and
Coussarea loftonii (Dwyer & Hayden) Dwyer, Sarapiqui, 100 m, 5 Nov 1980, B. Hammel
Ann. Missouri Bot. Card. 67: 129. 1980. 10383 (Holotype DUKE, n.v.).
Faramea loftonii Dwyer & Hayden, Coussarea paniculata Standi., J. Wash. Acad.
Phytologia 15: 56. 1967. Type: PANAMA: Sci. 18: 282. 1928, based on Froelichia
Panama, Cerro Jefe in Clusia forest at paniculata Vahl, Eclog. Amer. 1: 3. 1796. Not
2700-3000 ft., 27 Jan 1966, E. L. Tyson, J. Froelichia paniculata Moench., 1794. Type:
Dwyer & K. Blum 3323 (Holotype MO TRINIDAD: Ryan s.n. (C, n.v.).
1824505 (photo PTBG); Isotype MO (photo Coussarea psychotrioides C. M. Taylor &
PTBG)). Hammel, Selbyana 12: 135, fig. 2. 1991.
Coussarea mediocris Standi. & Steyerm., Publ. Type: COSTA RICA: Heredia, Finca La
Field Mus. Nat. Hist., Bot. Ser. 23: 248. 1947. Selva, the OTS field station near Puerto Viejo
Type: GUATEMALA: Huehuetenango, Sierra de Sarapiqui, near the junction of the Rios
de los Cuchumatanes, vicinity of Maxbal, Puerto Viejo and Sarapiqui, 100 m, 12 Jul
about 17 miles north of Barillas, 1500 m, 1979, J. H. Beach 1467 (Holotype DUKE,
15-26 Jul 1942, J. A. Steyermark 48732 n.v.; Isotypes CR, n.v., F, MO).
(Holotype F 1134264 (photos MEXU, Coussarea roseocremea Dwyer, Ann. Missouri
PTBG); Isotype US (photos MEXU, PTBG)). Bot. Gard. 67: 133. 1980. Type: PANAMA:
Coussarea mexicana Standi., Publ. Field Mus. Panama, El Llano-Carti Highway, 6-10 km
Nat. Hist, Bot. Ser. 17: 274. 1937. Type: north of El Llano, 13 Apr 1973, R. L. Dressier
MEXICO: Oaxaca, Ubero, 30-60 m, Jun 4334 (Holotype MO 2126258 (photo PTBG)).
1937, L. Williams 9442 (Holotype F 897721 —Psvchotria roseocrema (Dwyer) C. M.
(photos MEXU, PTBG); Isotypes P (photos Taylor
MEXU, PTBG), US (photos MEXU, PTBG)). Coussarea talamancana Standi., Publ. Field
Coussarea morii Dwyer, Ann. Missouri Bot. Mus. Nat. Hist., Bot. Ser. 18: 1288. 1938.
Gard. 67: 130. 1980. Type: PANAMA: Type: COSTA RICA: Talamanca Valley, Jun
Darien, Rio Tuquesa, at middle Tuquesa 1927, G. P. Cooper T4 (Holotype F 572698
Mining Company Camp called Charco Peje, (photo PTBG)).
tropical wet forest, 250 m, 8 Jul 1975, S. Mori Coussarea veraguensis Dwyer, Ann. Missouri
7058 (Holotype MO 2821790 (photo PTBG)). Bot. Gard. 67: 134. 1980. Type: PANAMA:
Coussarea nebulosa Dwyer, Ann. Missouri Bot. Veraguas, NW of Santa Fe, 1 km from
Gard. 67: 131. 1980. Type: PANAMA: Escuela Agricola Alto de Piedra, on the slopes
Chiriqui, uncut cloud forest at Monte Rey of Cerro Tute, S. Mori & J. Kallunki 4800
near Boquete, 20 Jul 1971, T. B. Croat 15868 (Holotype MO 2276605; Isotype MO
(Holotype MO 2162995 (photo PTBG); 2276606).—Coussarea caroliana Standi.
Coussarea villosula Dwyer, Ann. Missouri B< hexandra Jacq., Enum. PI. Carib. 16. 1760.
Gard. 53: 105. 1966. Type: PANAMA: Type: COLOMBIA: Cartagena in fruticosis,
Darien, Cerro Pirre, from Rio Pirre S to El Jacquin s.n. (Holotype presumably C, n.v.).
Real, 750-1030 ft., J. A. Duke 5338 Coutarea latiflora Sesse 8c Moc. ex DC, Prodr.
(Holotype MO). 4: 350. 1830. Type: MEXICO: Based on an
unpublished illustration [caique] (Fl. Mexic.
.1923. Ic. G-DC (photo MO)).—Hintonia latiflora
(Sesse & Moc. ex DC) Bullock
Coutarea lumaeana Baill., Adansonia 12: 301.
Coutaportla ghiesbreghtiana (Baill.) Urb., 1879. Type: GUATEMALA:
Symb. Antill. 9: 147. 1923. Portlandia
ghiesbreghtiana Baill., Adansonia 12: 300. (Holotype P (photos MEXU, MO, PTBG)).
1879. Type: MEXICO: Oaxaca, pres de Portlandia.—Hintonia lumaeana (Baill.)
Hacienda de Huijastla, au bord des ravins, Bullock
1842-43, Ghiesbreght 27 (Holotype P (photos Coutarea mexicana Zucc. & Mart, ex DC,
MEXU, PTBG); Isotypes P, 3 sheets (photos Prodr. 4: 350. 1830. Type: MEXICO: Without
MEXU), US (photos MEXU, PTBG)). precise locality, Martius s.n. in 1829
Coutaportla guatemalensis (Standi.) Lorence, (Holotype G-DC, n.v. (microfiche IDC 800.
Syst. Bot. 11: 210. 1986. Portlandia 680: II.7)). Cigarilla, Portlandia.—Nernstia
guatemalensis Standi., J. Wash. Acad. 18: mexicana (Zucc. & Mart, ex DC.) Urb.
162. 1928. Type: GUATEMALA: Alta Coutarea octomera Hemsl, Biol. Centr.-Amer.,
Verapaz, Quebradas Secas, 2500 ft., 1 Jun Bot. 4: 101. 1887. Type: MEXICO: Yucatan,
1920, H. Johnson 282 (Holotype US 1081354 Cozumel Island, Apr 1895, G. F. Gaumer 148
(photos MEXU, PTBG), fragment F). (Holotype K, n.v.).—Hintonia octomera
Coutaportla pailensis Villarreal, Sida 12: 223, (Hemsl.) Bullock
fig. I. 1987. Type: MEXICO: Coahuila, Mun. Coutareapterosperma (S. Watson) Standi., N.
Ramos Arizpe, Sierra de la Paila, Canon de Amer. Fl. 32: 127. 1921. Portlandia
Loma Prieta, al NW de Hipolito, 1800 m, J. A. pterosperma S. Watson, Proc. Amer. Acad.
I Warreal et al. 3305 (Holotype MEXU, n.v.; Arts 24: 52. 1889. Type: MEXICO: Sonora,
Isotypes ANSM (n.v.), ENCB, n.v., TEX, Guaymas, 1887, E. Palmer 298 (Holotype US
47978 (photos MEXU, PTBG); Isotype US
47979 (photos MEXU, PTBG)).—Hintonia
Coutarea Aubl., Hist. PI. Guiane 1: 314. 1775. latiflora (Sesse & Mocino ex DC.) Bullock
Type: Coutarea speciosa Aubl. Coutarea speciosa Aubl., Hist. Pi. Guiane 1:
Coutarea acamptoclada B. L. Rob. & Millsp., 314, t. 122. 1775. Type: FRENCH GUIANA:
Engl. Bot. Jahrb. 36: Beibl. 80: 28. 1905. Aublet s.n. (presumably P, n.v.). Portlandia.
Type: MEXICO: Yucatan, im Walde bei —Coutarea hexandra (Jacq.) K. Schum.
Uman, 7 Apr 1903, C. Seler & E. Seler 4044
(Holotype F (photo PTBG); Isotype B, Crucianella L., Syst. PI. 108. 1753. Lectotype:
presumably destroyed).—Hintonia octomera Crucianella latifolia L.
(Hemsl.) Bullock Crucianella americana Mill., Gard. Diet., ed. 8,
Coutarea flavescens Mo?. & Sesse ex DC, no. 5. 1768. Type: MEXICO: Veracruz,
Prodr. 4: 350. 1830. Type: MEXICO: Without Miller s.n. (Holotype BM, n.v.).—Status
precise locality, Sesse & Mocino s.n. (Fl. uncertain, probably Spermacoce sp.
Mexic. Ic. G-DC, n.v, (photo MO)). Crucianella hispida Mill., Gard. Diet., ed. 8, no.
—Coutarea hexandra (Jacq.) K. Schum. 4. 1768. Type: MEXICO: Veracruz, Miller
Coutarea hexandra (Jacq.) K. Schum. in Mart. s.n. (Holotype BM, n.v. (photo US)).
Fl. Bras. 6(6): 196. 1889. Portlandia —Crusea hispida (Mill.) B. L. Rob.
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Crusea Schltdl. & Cham., Linnaea 5: 165. Jan Crusea calocephala DC, Prodr. 4: 567. 1830.
1830. Not Crusea A. Rich., Dec 1830. Type: MEXICO: Without precise locality,
Lectotype: Crusea hispida (Mill.) B. L. Rob. "Pavon " [Sesse, Mocino et al], (Presumably
(Lectotype designated by Schumann, 1891: Herb. Dunat, location unknown (n.v.),
141). fragment P, n.v.; Isotype P, n.v.; see
Crusea allococca Ton. & A. Gray, Proc. Amer. Anderson, 1972: 97).
Acad. Arts 19: 78. 1883, nom. superfl. illeg., Crusea chiapensis Brandegee, Univ. Calif. Publ.
based on Diodia tricocca A. Gray (See Bot. 6: 374. 1917. Type: MEXICO: Chiapas,
Anderson, 1972: 116 for discussion.). C. A. Purpus 7526 (Holotype UC, n.v.).
—Richardia tricocca (Torr. & A. Gray) —Crusea longiflora (Willd. ex Roem. &
Standi, subsp. tetracocca (M. Martens & Schultes) W. R. Anderson
Galeotti) W. H. Lewis & R. L. Oliv. Crusea coccinea DC, Prodr. 4: 567. 1830.
Crusea andersoniorum Lorence, Novon 7: 57, Type: MEXICO: Nova Hispana, without
fig. 6. 1997. Type: MEXICO: Oaxaca, Distr. precise locality, "Pavon 588" [=Sesse, Mocino
de Miahuatlan, crest of ridge and steep SW et al. 588] (Lectotype MA (n.v.), fragment G-
facing slope 90-100 km SE of Sola de Vega DC, n.v.; Isolectotype F 850991 (photo
along road to Puerto Escondido, 1830 m, 25 PTBG); designated by Anderson, 1972: 39).
Oct 1984, D. E. Breedlove 62063 (Holotype Crusea coccinea DC. var. breviloba Loes.,
CAS 727669 (photo PTBG)). Verh. Bot. Vereins Prov. Brandenburg 65:
Crusea aspera Bartl., Ind. Sem. Hort. Gotting 115. 1923. Type: GUATEMALA:
1838. Type: Probably MEXICO: Herb. Huehuetenagno, Jacaltenango, San Martin, 9
Fischer, a garden specimen grown during Sep 1896, E. Seler & C Seler 3113 (Holotype
Bartling's tenure (Lectotype GOET, n.v.; B, presumably destroyed; Isotype GH, n.v.)
designated by Anderson, 1972: 111). Crusea coccinea DC. var. chiriquensis W. R.
—Crusea hispida (Mill.) B. L. Rob. var. Anderson, Mem. New York Bot. Gard. 22: 45.
hispida 1972. Type: PANAMA: Chiriqui, Casita Alta
Crusea brachyphylla Schltdl. & Cham., Linnaea to Cerro Copete, 2300-3300 m, 10 Jul 1040,
5: 165. 1830. Type: MEXICO: Veracruz: Woodson & Schery 358 (Holotype F 1089337
Serro [Cerro] Colorado, Aug 1828, Schiede & (photo PTBG); Isotypes GH, n.v., MO).
Deppe 524b (Holotype presumably HAL, n.v.; Crusea coccinea DC. var. pubescens Greenm.,
Isotype LE, n.v., fragment F) —Crusea Proc. Amer. Acad. Arts 32: 306. 1897. Type:
longiflora (Willd. ex Roem. & Schultes) MEXICO: Oaxaca: NW slope of Mt.
W. R. Anderson Zempoaltepec, 8000-10000 ft., 10 Jul 1894,
Crusea brachyphylla Schltdl. & Cham. f. E. W. Nelson 699 (Lectotype GH, n.v.;
microcephala Loes., Verh. Bot. Vereins Prov. Isolectotype US (photo PTBG); designated by
Brandenburg 65: 115. 1923. Type: MEXICO: Anderson, 1972: 39).—Crusea coccinea DC.
Michoacan, Patzcuaro, Monte Calvario, 1 Nov
1895, E. Seler & C Seler 1211 (Holotype B, Crusea coronata B. L. Rob. & Greenm., Proc.
presumably destroyed; Isotype GH, n.v.). Amer. Acad. Arts 29: 387. 1894. Type:
—Crusea longiflora (Willd. ex Roem. & MEXICO: Jalisco, Tequila, 3 Oct 1893, C G.
Schultes) W. R. Anderson Pringle 5424 (Holotype GH, n.v.; Isotype US
Crusea calicola Greenm., Proc. Amer. Acad. (photos MEXU, PTBG)).
Arts 32: 305. 1897. Type: MEXICO: Oaxaca, Crusea cruciata S. Watson, Proc. Amer. Acad.
dry calcareous hills, Las Sedas, 6000 ft., 8 Sep Arts 25: 152. 1890. Type: MEXICO: Jalisco,
1894, C. G. Pringle 4869 (Holotype GH, n.v.; barranca near Guadalajara, 19 Oct 1889, C. G.
Isotypes BM, n.v., BR, n.v., F, MICH, n.v., Pringle 2969 (Holotype GH, n.v.; Isotype F)).
MO (photos MEXU, PTBG), NY, n.v., UC —Crusea setosa (M. Martens 8c Galeotti)
(photo PTBG), US (photos MEXU, PTBG)). Standi. & Steyerm.
Crusea cruciata S. Watson var. villosior Crusea hispida (Mill.) B. L. Rob. var.
Greenm., Proc. Amer. Acad. Arts 32: 306. grandiflora (Paul G. Wilson) W. R.
1897. Type: MEXICO: Jalisco, Guadalajara, Anderson, Mem. New York Bot. Gard. 22:
Jul-Oct 1886, E. Palmer 901 (Lectotype GH, 115. 1972. Crusea grandiflora Paul G.
n.v.; Isolectotype US; designated by Wilson, Kew. Bull. 13: 161. 1958. Type:
Anderson, 1972: 83).—Crusea setosa (M. MEXICO: Mexico, G. B. Hinton 4848
Martens & Galeotti) Standi. & Steyerm. (Holotype K, n.v.).
Crusea diversifolia (Kunth) W. R. Anderson, Crusea hispidula Bartl., Ind. Sem. Hort.
Mem. New York Bot. Gard. 22: 58. 1972. Gotting. 1839: 3. 1839. Type: Herb. Fischer, a
Spermacoce diversifolia Kunth in Humb., garden specimen grown during Bartling's
Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 341. tenure (Holotype GOET (n.v.)).—Crusea
1818 [1819]. Type: MEXICO: Guanajuato, calocephala DC.
Santa Rosa de la Sierra, Humboldt & Crusea longibracteata Benth. in Oerst,
Bonplands.n. (Holotype P-Bonpland, n.v. Vidensk. Meddel. Dansk. Naturhist. Foren.
(photos F, MO)). Kjoebenhavn 1852: 28. 1853. Type:
Crusea elata Brandegee, Univ. Calif. Publ. Bot. NICARAGUA: Granada, Jun 1846, Oersted
6: 68. 1914. Type: MEXICO: Chiapas, Sierra s.n. (Holotype, K, n.v.; Isotype US)).
de Tonala, Sep 1913, C. A. Purpus 6640 —Crusea setosa (M. Martens & Galeotti)
(Holotype UC 172353 (n.v.); Isotypes BM, Standi. & Steyerm.
n.v., F, GH, n.v., NY, n.v., US (photos Crusea longiflora (Willd. ex Roem. &
MEXU, PTBG)). Listed in synonymy under Schultes) W. R. Anderson, Mem. New York
Crusea calocephala by Anderson (1972), but Bot. Gard. 22: 89. 1972. Spermacoce
it actually represents a valid species (W. R. longiflora Willd. ex Roem. & Schultes, Syst.
Anderson, pers. comm. 1984). Veg. 3: 531. 1818. Type: MEXICO:
Crusea gentryi Standi., Publ. Field Mus. Nat. Michoacan, near Ario [de Rosales], and
Hist, Bot. Ser. 22: 112. 1940. Type: Mexico City, Humboldt & Bonpland s.n.
MEXICO: Sonora: Alamos Distr., Quirocoba, (Holotype B-W, n.v.).
12 Nov 1923, H. S. Gentry 766M (Holotype F Crusea lucida Benth., Bot. Voy. Sulphur 109.
968108 (photo PTBG); Isotype MICH, n.v.). 1845. Type: MEXICO: Without precise
—Crusea lucida Benth. locality, southern Mexico, Sinclair s.n.
Crusea grandiflora Paul G. Wilson, Kew Bull. (Holotype K, n.v. (photo MICH)).
13: 161. 1958. Type: MEXICO: Mexico, Crusea megalocarpa (A. Gray) S. Watson,
Distr. Temascaltepec, Tenayac, 1540 m, 28 Proc. Amer. Acad. Arts 26: 137. 1891.
Sep 1933, G. B. Hinton 4848 (Lectotype K, Spermacoce megalocarpa A. Gray, Proc.
n.v.; designated by Anderson, 1972). Amer. Acad. Arts 21: 381. 1886. Type:
—Crusea hispida (Mill.) B. L. Rob. var. MEXICO: Chihuahua, southwestern
Chihuahua, wet places in the higher
Crusea guatemalensis Gand., Bull. Soc. Bot. mountains above Batopilas, Aug-Nov 1885,
France 65: 34. 1918. Type: GUATEMALA: E. Palmer 185 (Holotype GH, n.v.; Isotypes
Alta Verapaz, H. von Tuerckheim s.n. NY, n.v., US (photos MEXU, PTBG)).
(Holotype presumably P, n.v.).—Crusea Crusea palmeri A. Gray, Proc. Amer. Acad.
calocephala DC. Arts 21: 381. 1886. Type: MEXICO:
Crusea hispida (Mill.) B. L. Rob., Proc. Amer. Chihuahua, southwestern Chihuahua, 25 miles
Acad. Arts 45: 409. 1910. var. hispida. S of Batopilas, Aug-Sep 1885, E. Palmer 70
Crucianella hispida Mill, Gard. Diet., ed. 8, (Holotype GH, n.v.; Isotypes BM, n.v.,
no. 4. 1768. Type: MEXICO: Veracruz, MEXU, NY, n.v., US (photos MEXU,
grown at Chelsea from seed sent by Houstoun, PTBG)).—Crusea psyllioides (Kunth) W. R.
Miller s.n. (Holotype BM, n.v. (photo US)). Anderson
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

Crusea parviflora Hook. & Arn., Bot. Beechey n.v.).—Crusea diversifolia (Kunth) W. R.
Voy. 430, pi. 99c. 1840 (text), 1841 (plate). Anderson
Type: MEXICO: Guerrero, Acapulco, Sinclair Crusea subulata (DC.) A. Gray var. leiocarpa
s.n. (Holotype K, n.v.). Greenm., Proc. Amer. Acad. Arts 40: 33.
Crusea psyllioides (Kunth) W. R. Anderson, 1904. Type: MEXICO: San Luis Potosi,
Mem. New York Bot. Gard. 22: 75. 1972. Alvarez, 28 Sep-3 Oct 1902, E. Palmer 147
Spermacoce psyllioides Kunth in Humb., (Holotype GH, n.v.); Isotypes F, MO (photos
Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 346, pi. MEXU, PTBG), NY, n.v., US (photos
278. 1818 [1819]. Type: MEXICO: MEXU, PTBG)).—Crusea diversifolia
Michoacan, near Ario [de Rosales], Humboldt (Kunth) W. R. Anderson
& Bonplands.n. (Holotype P-Bonpland, n.v. Crusea tricocca (Torr. & A. Gray) A. Heller,
(photo MICH)). Contr. Herb. Franklin & Marshall College 1:
Crusea rosea Loes., Repert. Spec. Nov. Regni 97. 1895. Diodia tricocca Torr. & A. Gray, Fl.
Veg. 18: 362. 1922. Type: MEXICO: N. Amer. 2: 30. 1841. Type: U.S.A.: Texas,
Morelos: barranca bei Cuernava Drummond 117 (Holotype F; Isotype G, n.v.).
[Cuernavaca], 14 Oct 1904, E. Seler & C. —Richardia tricocca (Torr. & A. Gray)
Seler 4181 (Lectotype GH, n.v.; designated by Loes. subsp. tricocca
Anderson, 1972: 111).—Crusea hispida Crusea villosa S. Watson, Proc. Amer. Acad.
(Mill.) B. L. Rob. var. hispida Arts 25: 152. 1890. Type: MEXICO: Jalisco,
Crusea rubra (Jacq.) Schltdl. & Cham., Linnaea rocky hillsides near Guadalajara, 9 Oct 1889,
5: 165. 1830. Spermacoce rubra Jacq., PL C. G. Pringle 2448 (Holotype GH, n.v.;
Rar. Hort. Schoenbr. 3: 3,pi. 256. 1798. Isotypes BM, n.v., MEXU, MO (photos
Type: Unknown (see Anderson, 1972: 110). MEXU, PTBG), NY, n.v., US (photos
—Crusea hispida (Mill.) B. L. Rob. var. MEXU, PTBG)).—Crusea psyllioides
(Kunth) W. R. Anderson
Crusea setosa (M. Martens & Galeotti) Standi. Crusea violacea Brongn. ex Neumann, Rev.
& Steyerm., Publ. Field Mus. Nat. Hist., Bot. Hort. II. 4: 368. 1846; 5: 61. 1846. Type:
Ser. 23: 22. 1943. Borreria setosa M. Martens Unknown (see Anderson, 1972: 97).—Crusea
& Galeotti, Bull. Acad. Roy. Sci. Bruxelles calocephala DC.
11:131. 1844. Type: MEXICO: Oaxaca, dans Crusea wrightii A. Gray, PL Wright. 2: 68.
les forets de palmiers et sur les dunes de la 1853. Type: MEXICO: Sonora, prairies near
cote d'Oaxaca, baignee par l'Ocean Pacifique, Santa Cruz, Sep 1851, C. Wright 1121
pres du Rancho du Rio Grande, Sep 1839, H. (Holotype GH, n.v.; Isotypes GH, n.v., US
Galeotti 2627 (Holotype BR, n.v.; Isotype P, (photos MEXU, PTBG)).
Crusea wrightii A. Gray var. angustifolia
Crusea subalata Hook. & Arn., Bot. Beechey W. R. Anderson, Mem. New York Bot. Gard.
Voy. 431. 1840. Type: MEXICO: Nayarit, 22: 108. 1972. Type: MEXICO: Guerrero, 6.3
between San Bias and Tepic, Sinclair s.n. miles east of Chilpancingo on road to Tixtla,
(Holotype K, n.v.).—Crusea parviflora at km 12, 1600 m, W. R. Anderson & C W.
Hook. & Arn. Laskowski 4334 (Holotype MICH, n.v.;
Crusea subulata (DC.) A. Gray, Proc. Amer. Isotype MO (photos MEXU, PTBG)).
Acad. Arts 19: 78. 1883. Borreria subulata
DC, Prodr. 4: 543. 1830. Type: MEXICO: Dasycephala DC, Prodr. 4: 565. 1830. (Diodiae
Without precise locality, Berlandier 826 sect. 2). Type: Not designated.
(Lectotype [annotated by de Candolle but not Dasycephala spicata (Miq.) Benth. in Benth. &
cited in protologue, according to Anderson, Hook, f, Gen. PL 2: 144. 1873. Diodia
1972: 58] G, n.v. (photo F); Isolectotype BM, spicata Miq., Stirp. Surin. Sel. 179, pi 52.
1851. Type: SURINAM: Sylvarum prope
 Bergendaal, Oct, Focke s.n. (presumably U, Deppea Cham. & Schltdl., Linnaea 5: 167.
n.v.; this is Miquel's lectotype according to 1830. Type: Deppea erythrorhiza Cham. &
Howard, 1989).—Diodia spicata Miq. Schltdl.
Deppea amaranthina Standi. & Steyerm., Publ.
Declieuxia Kunth in Humb., Bonpl. & Kunth, Field Mus. Nat. Hist., Bot. Ser. 22; 281. 1940.
Nov. Gen. Sp. 352,/?/. 25/. 1818. Type: Type: GUATEMALA: Chiquimula, Cerro
Declieuxia fruticosa (Willd. ex Roem. & Brujo, in vicinity of Rio Negro, below
Schult.) Kuntze. Montana Montenegro, near village of Brujo,
Declieuxia fruticosa (Willd. ex Roem. & 1500-2000 m, 1 Nov 1939, J. A. Steyermark
Schult.) Kuntze, Rev. Gen. 1: 279. 1891. 30926 (Holotype F 1035809 (photos F,
Houstonia fruticosa Willd. ex Roem. & MEXU, PTBG)).
Schult., Syst. Veg. 3: 527. 1818. Type: Deppea anisophylla L. O. Williams, Phytologia
VENEZUELA: Monagas, Caripe, Humboldt 26: 490. 1973. Type: GUATEMALA:
s.n. (presumably B-W, n.v.). Suchitepequez, Finca Moca, 1000 m, 7 Jan
Declieuxia fruticosa (Willd. ex Roem. & 1935, A. F. Skutch 2072 (Holotype F 815818
Schult.) Kuntze var. mexicana (DC.) Standi., (photos MEXU, PTBG); Isotype US).
Field Mus. Nat. Hist, Bot. Ser. 12: 378. 1936. Deppea cornifolia (Benth.) Benth., PI. Hartw.
Declieuxia mexicana DC, Prodr. 4: 479. 349. 1857. Rondeletia cornifolia Benth., PI.
1830. Type: MEXICO: Without precise Hartw. 39. 1840. Type: MEXICO:
locality, Haenke s.n. (Holotype presumably Michoacan, prope Morelia, T. Hartweg 299
PR, n.v.).—Declieuxia fruticosa (Willd. ex (Holotype K (photo MEXU); Isotypes BM, G-
Roem. & Schult.) Kuntze DEL, n.v., fragment F, P (photos MEXU,
Det lieuxia galeottii M. Martens in M. Martens PTBG, US), US, fragment (photos MEXU,
& Galeotti, Bull. Acad. Roy. Soc. Bruxelles PTBG), W, 2 sheets (photos PTBG)).
11: 231. 1844. Type: MEXICO: Oaxaca, Choristes.
Llano Verde. 3000-7000 ft., Apr 1840, H. Deppea costaricensis Pol., Linnaea 41: 566.
Galeotti 2603 (Holotype BR (photo PTBG); 1877. Type: COSTA RICA: Desengano,
Isotypes BR, n.v., F (photos PTBG), US). prope Alajuela, 27 Jun 1875, H. Polakowski
Hedyotis galeottii (M. Martens) Terrell & 148 (Holotype B, presumably destroyed
(photos F, MEXU, US); Isotypes BM, W
Declieuxia mexicana DC, Prodr. 4: 479. 1830. (photo PTBG), fragment F).—Deppea
Type: MEXICO: Without precise locality, grandiflora Schltdl.
He k (Holotype presumably PR, n.v.). Deppea ehrenbergii Standi., Publ. Field
—Declieuxia fruticosa (Willd. ex Roem & Columbian Mus., Bot. Ser. 8: 161. 1930.
Schult.) Kuntze Type: MEXICO: Without precise locality, C.
Ehrenberg 1318 (Holotype B, presumably
Dentella J. R. & G. Forst., Char. Gen. PI. 25, t. destroyed (fragment F (photos MEXU,
13. 1776. Type: Dentella repens (L.) J. R & PTBG)).
G. Forst. Deppea erythrorhiza Cham. & Schltdl.,
Dentella repens (L.) J. R. & G. Forst., Char. Linnaea 5: 168. 1830 (Jan 1830). Type:
Gen. PL 26. 1776. Oldenlandia repens L., MEXICO: Veracruz, in umbrosis prope
Mant. 40. 1767. Type: INDIA: (Lectotype Hacienda de la Laguna, Aug 1829, C. J. W.
LINN 155/2, n.v.; designated by Verdcourt, Schiede & F. Deppe 388 (Holotype HAL
1983). Naturalized in Mexico at least in (photo PTBG); Isotype HAL (photo PTBG)).
Colima and Guerrero; native to Australasia. Deppea excelsa (Kunth) Standi., N. Amer. Fl.
32: 89. 1921. Psychotria (?) excelsa Kunth in
Humb., Bonpl. & Kunth, Nov. Gen. Sp. 3:
355, t. 282. 1819. Type: MEXICO: Veracruz,
A NOMENCLATOR OF MEXICAN AND CENTRAL

Xalapa, A. Bonpland 1445 (Holotype al este de Tenango de Doria hacia al Cirio,

presumably P-Bonpland, n.v.; Isotypes B 1700-1800 m, 19 Feb 1981, R. Hernandez M.
(photo MO), P (photos MEXU, PTBG)). & D. Rodriguez 5466 (Holotype MEXU;
—Hoffmannia excelsa (Kunth) K. Schum. Isotypes MO (photo PTBG), XAL).
Deppea fiava (Brandegee) L. O. Williams, Deppea hintonii Bullock, Kew Bull. 5: 309.
Fieldiana, Bot. 24(11): 58. 1975. 1937 [as D. hintoni]. Type: MEXICO:
Plocaniophyllon flavum Brandegee, Univ. Mexico, Distr. of Temascaltepec, Tejupilco,
Calif. Publ. Bot. 6: 69. 1914. Type: MEXICO: barranca, 1340 m, 22 Sep 1933, G B. Hinton
Chiapas, Finca Mexiquito, Jun 1913, C. A. 4824 (Holotype K (photo PTBG); Isotypes
Purpus 7019 (Holotype UC 173077, n.v.; BM, F (photo PTBG), MO (photo PTBG),
Isotypes F (photos MEXU, PTBG), US US).
(photos MEXU, PTBG)).—Plocaniophyllon Deppea inaequalis Standi. & Steyerm., Publ.
flavum Brandegee Field Mus. Nat. Hist., Bot. Ser. 22: 384. 1940.
Deppea floribunda Hemsl, Diagn. PI. Nov. Type: GUATEMALA: San Marcos, above
Mexic. 31. 1879. Type: MEXICO: Veracruz, Finca el Porvenir along Rio Cabus to within 2
Orizaba, Botteri 937 (Lectotype P (photos miles of Cueva de las Palomas, south-facing
MEXU, PTBG, US); Isolectotypes BM, F, slopes of Volcan Tajumulco, 1300-1500 m,
fragment, P (photos MEXU, PTBG), W 16 Mar 1940, J. A. Steyermark 37974
(photo PTBG); designated by Lorence & (Holotype F 1043181 (photos F, PTBG)).
Dwyer, 1988).—Deppea grandiflora Schltdl. Deppea longipes Standi., Conrr. U.S. Natl.
Deppea grandiflora Schltdl., Linnaea 19: 748. Herb. 18: 138. 1916. Type: PANAMA.
1847. Type: MEXICO: Without precise Chiriqui, east slopes of Chiriqui Volcano,
locality, Temperirites (presumably Veracruz), Camp Aguacatl, 2100-2200 m, 10-13 Mar
Leibold235, 253, 263 (Syntypes HAL (photos 1911,//. Pittier 3070 (Holotype US 677458
PTBG); a lectotype cannot be selected with (photos MEXU, PTBG, US)).—Deppea
certainty; for discussion see Lorence & grandiflora Schltdl.
Dwyer, 1988). Deppea lundellii Dwyer, Phytologia 54: 277.
Deppea guerrerensis Dwyer & Lorence, 1983. Type: GUATEMALA: Baja Verapaz,
Allertonia4: 428, fig. 4a-c. 1988. Type: Nino Perdido, bordering Rio San Jose, C. /.
MEXICO: Guerrero, vicinity of Omiltemi, 20 Lundell & E. Contreras 21025 (Holotype
miles west of Chilpancingo on logging road, TEX-LL; Isotypes MEXU, MO (photo
2500-2700 m, 31 Jul 1957, R. M. Straw & PTBG)). In the protologue Dwyer states "(LL;
D. P. Gregory 1081 (Holotype MEXU; holotype, MO)." This is probably a
Isotype ENCB). typographical error, as the MO specimen was
Deppea hamelioides Standi., Publ. Field annoted by Dwyer as isotype, and the
Columbian Mus., Bot. Ser. 4: 269. 1929. specimen at TEX-LL is presumably the
Type: MEXICO: Jalisco, San Sebastian, holotype.—Pinarophyllon bullatum Standi.
Segundo Arroyo, Sierra Madre Mountains, Deppea macrocarpa Standi., Publ. Field
1500 m, 22 Jan 1927, Y. Mexia 1546a Columbian Mus., Bot. Ser. 4: 268. 1929.
(Holotype F 579899 (photo PTBG); Isotypes Type: MEXICO: Nayarit, Cerro de San Juan,
A, n.v., BM, US (photos F, MEXU, PTBG)). west of Tepic, 1000 m, 19 Sep 1926, Y. Mexia
Deppea hedyotidea DC, Prodr. 4: 618. 1830 727 (Holotype F 579908 (photos MEXU,
(Sep 1830), nom. superfl., based on the type PTBG); Isotypes GH, n.v., MO, US, 2 sheets
of D. erythrorhiza Cham. & Schltdl. (photos PTBG)).—Rondeletia jurgensenii
—Deppea erythrorhiza Cham. & Schltdl. Hemsl.
Deppea hernandezii Lorence, Allertonia 4: Deppea martinez-calderonii Lorence,
^5, fig. 4d-g. 1988. Type: MEXICO: Allertonia 4: 399, figs, le, 2. 1988. Type:
Hidalgo, Mun. de Tenango de Doria, a 6 km MEXICO: Oaxaca, Distr. de Ixtlan, Sierra de
 Juarez, Ruta 175 a 2.5 km al norte de Puerto n.v., MO, P (photos F, MEXU, PTBG, US),
Eligio, 780 m, 4 Ago 1985, D. Lorence et al. US (photos MEXU, PTBG)).
4714 (Holotype MEXU; Isotypes ENCB, Deppea purpurascens Lorence, Allertonia 4:
MEXU, MO, UC, W, XAL). 402, fig. 3. 1988. Type: MEXICO: Veracruz,
Deppea miahuatlanica Lorence, Novon 7: 48, Mun. de Soteapan, Cumbres de Bastonal, ca.
fig 2. 1997. Type: MEXICO: Oaxaca, Distr.' 10-11 km al sureste de Tebanca (lado sureste
de Miahuatlan, Mun. de San Jeronimo de Lago de Catemaco) por camino, 1200 m,
Coatlan, 17.9 km al SW de San Jeronimo 26 Abr 1982, D. Lorence, R. Cedillo T & G.
Coatlan, carretera Miahuatlan-Piedra Larga, Ibarra M. 4141 (Holotype MEXU; Isotypes
1800 m, 16° 12'N, 96°57'W, 13 Dec 1987, R. CHAPA, MEXU, MO).
Torres C. & A. Campos 10847 (Holotype Deppea purpusii Standi., N. Amer. Fl. 32: 88.
PTBG 15197; Isotype MEXU). 1921. Type: MEXICO: Veracruz, Sierra
Deppea microphylla Greenm., Proc. Amer. Madre between Misantla and Naolinco, Aug
Acad. Arts 41: 249. 1905. Type: MEXICO: 1912, C. A. Purpus 6260 (Holotype US
Hidalgo, river bluffs below the Trinidad Iron 464734 (photos MEXU, PTBG); Isotypes
Works, 5700 ft., 26 Apr 1904, C. G. Pringle BM, F (photo PTBG), GH, n.v., MO (photo
8907 (Holotype GH (n.v.); Isotypes F (photos PTBG)).
MEXU, PTBG), MEXU, MO, P (photos Deppea schultzei Lorence, Allertonia 4: 422.
MEXU, PTBG), US, 2 sheets (photos MEXU 1988. Type: MEXICO: Guerrero,
PTBG), W (photo PTBG)). Malinaltepec, Mar 1930, L. Schultze 363
Deppea oaxacana Lorence, Allertonia 4: 419 (Holotype B, destroyed, fragment F (photos
fig. 5. 1988. Type: MEXICO: Oaxaca, Distr. MEXU, PTBG)). New name for Edithea
de Mixe, a 9 km de la enrrada a Villa Alta, floribunda Standi., Trop. Woods 34: 1. 1933.
este entronque a 10 km al suroeste de Not Deppea floribunda Hemsl., 1879.
Totontepec, 2230 m, 13 Die 1985, R. Torres Deppea scoti (J. H. Kirkbr.) Lorence, Allertonia
C. & C. Martinez 7956 (Holotype MEXU- 4: 398. 1988. Omiltemia scoti J. H. Kirkbr.,
Isotypes BM, ENCB, F, MEXU, MO, UC W Syst. Bot. 9: 411. 1984. Type: MEXICO:
XAL). ' ' Puebla, above Teotklan del Camino on road to
Deppea obtusiflora (Benth.) Benth., PI. Huautla de Jimenez, 2000-3250 m, 3 Aug
Hartweg. 349. 1857. Choristes obtusiflora 1961, C. E. Smith, Jr., F. A. Peterson & N.
Benth., PI. Hartw. 63. 1840. Type: MEXICO: Tejeda 413 7 (Holotype US; Isotype GH
Oaxaca, in planitie Llano Verde, T Hartweg (photo PTBG)).
473 (Holotype K (photo PTBG); Isotypes F Deppea splendens Breedlove & Lorence,
(fragment from P), P (photos F, MEXU Phytologia 63: 43, fig. 1. 1987. Type:
PTBG, US), W, 2 sheets (photos PTBG)). MEXICO: Chiapas, Mun. of Motozintla de
Deppea panamensis Dwyer, Ann. Missouri Bot Mendoza, southwest (side) of Cerro Mozotal,
Card. 67: 145. 1980. Type: PANAMA: 11 km northwest of junction of road to
Panama, 5-10 km northeast of Altos de Motozintla along road to El Porvenir and
Pacora, ca. 750 m, 7 Mar 1975, S. Mori & J Siltepec, 2100 m, 18 Sep 1976, D. E.
Kallunki 4965 (Holotype MO 2257147- Breedlove 40258 (Holotype CAS; Isotypes
Isotypes MO, 2 sheets).—Elaeagia MEXU, MO (photo PTBG)).
myriantha (Standi.) C. M. Taylor & Hammel Deppea stenosepala Standi., Publ. Field
Deppea pubescens Hemsl., Diagn PI Nov Columbian Mus., Bot. Ser. 8: 161. 1930.
Mexic. 31. 1879; Biol. Centr.-Amer., Bot 2- Type: MEXICO: Oaxaca, Tlaxiaco a
27. 1881. Type: MEXICO: Mexico, Santa Chicahuastla, 1844,//. Galeotti 2685
Gertrudis, F. Liebmann 65 (=Plantae (Holotype LE, n.v., fragment F; Isotype P,
Mexicanae Liebmann 11382) (Holotype K n.v., fragment F)).—Rondeletia jurgensenii
(photo PTBG), fragment F); Isotypes GH,
 *. NOMENCLATOR OF MEXICAN AND CENTRAL

i Benth., PI. Hartw. 349. F 838717 (photo PTBG)). 1938.—Didymaea

1857. Type: MEXICO: Chiapas, San Bartolo, alsinoides (Schltdl. & Cham.) Standi.
Feb 1840, Linden 1644 (Holotype K; Isotype Didymaea floribunda Rzed., Bol. Soc. Bot.
LE, n.v., fragment F). Mexico 44:12 Jig. 1. 1983. Type: MEXICO:
Deppea umbellata Hemsl., Diagn. PI. Nov. Distr. Federal, Del. Villa A. Obregon, Limbo,
Mexic. 31. 1879. Type: MEXICO: Veracruz, 2700 m, 4 Aug 1979, A. Ventura A. 3479
region de Orizaba a Izhuatlancillo, 15 Mai (Holotype ENCB).
1866, E. Bourgeau 2428 (Lectotype P (photos Didymaea hispidula L. O. Williams, Fieldiana:
MEXU, PTBG); Isolectotypes F, K (photo Bot. 24(11): 61. 1975. Type: GUATEMALA:
PTBG), P, 3 sheets (photos MEXU, PTBG), San Marcos, Sierra Madre Mountains about 6
US, 2 sheets; designated by Lorence & km (airline) north of San Marcos, 2700 m, 13
Dwyer, 1988). Dec 1963, L. O. Williams, A. Molina R. & T.
Williams 25896 (Holotype F 1655451 (photo
Didymaea Hook. f. in Benth. & Hook, f, Gen. PTBG))
PI. 2(1): 150. 1873. Type: Didymaea Didymaea linearis Standi., Publ. Field
Columbian Mus., Bot. Ser. 4: 297. 1929.
Didymaea alsinoides (Schltdl. & Cham.) Type: MEXICO: Jalisco, Real Alto, Pena
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. Gorda, Sierra Madre Occidental, 2500 m, 21
18: 1291. 1938. Nertera alsinoides Schltdl. & Feb 1927, Y. Mexia 1726 (Holotype F 579896
Cham.,Linnaea6:413. 1831. Type: (photo PTBG); Isotypes A, n.v., MO (photos
MEXICO: Cuesta Grande de Jalacingo, MEXU, PTBG), US (photos MO, PTBG;
Schiede & Deppe 1271 (Holotype presumably fragment F)).
HAL, n.v.). Didvmaea mexicana Hook. f. in Benth. &
Didymaea alsinoides (Schltdl. & Cham.) Standi, Hook, f, Gen. PI. 2: 150. 1873. Syntypes:
var. australis Standi., Publ. Field Mus. Nat. MEXICO: Veracruz, Pico de Orizaba, 10000
Hist., Bot. Ser. 18: 1292. 1938. Type: COSTA ft., Bates s.n. (K, n.v.); Orizaba, Linden 1403
RICA: Charpinteira [La Carpintiera], 1700 m, (K, n.v.), Galeotti 4405 (K, n.v.), Bourgeau
Nov 1908, A. Brade & A. C. Brade 2143 3050 (K, n.v.), F. Mueller 1519 (K, n.v.), F.
(Holotype F 838717 (photo PTBG)). Liebmann Rub. Mexic. 255 (K, n.v.).
—Didymaea alsinoides (Schltdl. & Cham.) Didymaea microflosculosa L. O. Williams,
Standi. Fieldiana: Bot. 24(11): 61. 1975. Type:
Didymaea alsinoides (Schltdl. & Cham.) Standi, GUATEMALA: San Marcos, near Aldea
var. mollis Standi, Publ. Field Mus. Nat. Fraternidad, between San Rafael Pie de la
Hist., Bot. Ser. 18: 1292. 1938. Type: COSTA Cuesta and Palo Gordo, west-facing slopes of
RICA: San Jose, Cerro de Las Vueltas, the Sierra Madre Mountains, 1800-2400 m,
2700-3000 m, 29 Dec 1925-1 Jan 1926, P. C. 10-18 Dec 1963, L. O. Williams, A. Molina R.
Standley & J. Valeria 43949 (Holotype US & T. Williams 25944 (Holotype F 1655450
1253422 (photos MEXU, PTBG); Isotype F (photo PTBG)).
(photo PTBG)).—Didymaea alsinoides Didymaea microphylla L. O. Williams,
(Schltdl. & Cham.) Standi. Fieldiana: Bot. 24(11): 62. 1975. Type:
Didymaea australis (Standi.) L. O. Williams, GUATEMALA: Quezaltenango, narrow ridge
Fieldiana: Bot. 24(11): 61. 1975. Didymaea of Volcan Santo Tomas, 2500-3700 m, 22 Jan
alsinoides (Schltdl. & Cham.) Standi, var. 1940, J. A. Steyermark 34790 (Holotype F
australis Standi., Publ. Field Mus. Nat. Hist., 1038453 (photo PTBG)).
Bot. Ser. 18: 1292. Type: COSTA RICA:
Charpinteira [La Carpintiera], 1700 m, Nov Didymochlamys Hook, f, Icon. PI. pi 1122.
1908, A. Brade & A. C. Brade 2143 (Holotype 1872. Type: Didymochlamys whitei Hook. f.
Didymochlamys connellii N. E. Br., Trans.
Linn. Soc. Lond. (Bot.), ser. 2, 6: 34, pi. 5,fig-
1-9. 1901. Type: VENEZUELA: Bolivar,
Monte Roraima, 1050 m, McConnell & Diodia maritima Thonn. ex Schumach., Besk.
Quelch 714 (location unknown). Guin. PI. 75. 1827. Syntypes: AFRICA: in
sabulosis subimmersis ad littora oceani in
Diodia L., Sp. PI. 1: 104. 1753; Gen. PI. ed. 5: Guinea et ad Dakar prom, viridis Africa,
45. 1754. Type: Diodia virginiana L. Perrottet s.n. and Leprieur s.n. (location
Diodia apiculata (Willd. ex Roem. & Schult.) unknown).—Diodia serrulata (P. Beauv.) G.
K. Schum. in Engler, Bot. Jahrb. Syst. 10:
313. 1889. Spermacoce apiculata Willd. ex Diodia ocymifolia (Willd. ex Roem. & Schult.)
Roem. & Schult., Syst. Veg. 3: 531. 1818. Bremek., Rec. Trav. Bot. Neerl. 31: 305.
Type: BRAZIL: Hoffmanns egg s.n. 1934. Spermacoce ocymifolia Willd. ex
(Holotype:B-W2626,n.v.) Roem. & Schult., Syst. Veg. 3: 530. 1819 [as
Diodia aspera Brandegee, Univ. Calif. Publ. ocimifolia]. Type: WEST ESDIES: Without
Bot. 6: 68. 1914. Type: MEXICO: Chiapas, precise locality, 1768, Rudolphi s.n
Sierra de Tonala, C A. Purpus 6830 (presumably B-W, n.v.).—Spermacoce
(Holotype UC 172537, n.v.). Mixed ocymifolia Willd. ex Roem. & Schult.
collection, with elements of both D. apiculata Diodia prostrata Sw., Prodr. Veg. Ind. Occ.
and D. teres. 218. 1788. Type: JAMAICA: Swartz s.n.
Diodia brasiliensis Spreng. var. angulata (Presumably herb. Heretier, S, n.v., fragment
(Benth.) Standi., Carnegie Inst. Wash. Publ. G-DC, n.v. (microfiche 4: 562.9)).—Diodia
461: 90. 1935. Triodon angulation Benth., PI. teres Walter
Hartweg. 70. 1840. Type: MEXICO: Oaxaca, Diodia prostrata Sw. var. longiseta DC, Prodr.
Teotolcingo, in montibus Chinantla, T. 4: 562. 1830. Type: MEXICO: Without
Hamveg 507 (Holotype K, n.v.).—Galianthe precise locality, Haenke s.n. (Holotype
brasiliensis (Spreng.) E. L. Cabrera & presumably PR, n.v., fragment G-DC, n.v.
Bacigalupo ssp. angulata (Benth.) E. L. (microfiche 4: 562.9)).—Diodia teres Walter
Cabrera & Bacigalupo. Diodia pulchella Brandegee, Univ. Calif. Publ.
Diodia crassifolia Benth., Bot. Voy. Sulphur Bot. 10: 418. 1924. Type: MEXICO: Chiapas,
108. 1845. Type: MEXICO: Nayarit, San Bias Hacienda Monserrate, Sep 1923, C. A. Purpus
[near Tepic], Barclay s.n. (Holotype BM, 9221 (Holotype UC 223390, n.v.; Isotypes
MO (photo PTBG), US (photos MEXU,
Diodia denudata Standi., J. Wash. Acad. Sci. PTBG)).—Diodia apiculata (Willd. ex Roem.
15: 105. 1925. Type: PANAMA: Panama, Rio & Schult.) K. Schum.
Tapia, 7 Dec 1923-11 Jan 1924, P. C. Diodia rigida (Willd. ex Roem. & Schult.)
Standley 28123 (Holotype US 1154022 Cham. & Schltdl., Linnaea 3: 341. 1828.
(photos MEXU, PTBG)).—Diodia spicata Spermacoce rigida Willd. ex Roem. &
Miq. Schult., Syst. 3: 531. 1818. Not Diodia rigida
Diodia erecta Sesse & M09., Fl. Mexic, ed 1 Salisb. 1796. Type: VENEZUELA: Silla de
26. 1893 (La Naturaleza ser. II, 2. App. 2 Caracas, Humboldt & Bonpland (Holotype B-
1893). Type: MEXICO: Habitat in Oppidis S. W 2628, n.v. (microfiche IDC 7440. 174.
Bemardi de Xochiatlam et Hueitlalpae. Floret 111,4)).—Diodia apiculata (Willd. ex Roem.
Augusto ad Octobrim, Sesse & Mocino s.n. & Schult.) K. Schum.
(presumably MA, 1 Sw., Prodr. Veg. Ind. Occ.
Diodia indecora DC, Prodr. 4: 565. 1830. 30. 1788. Type: JAMAICA: No specimen
Type: MEXICO: Cordiere de Guehila, Oct cited. A Swartz specimen would be a probable
1827, M. Berlandier 1186 (Holotype G-DC, neotype according to Howard (1989).
A NOMENCLATOR OF MEXICAN AND CENTRAL

Diodia serrulata (P. Beauv.) G. Taylor in Exell,

Cat. PL S. Tome 220. 1944. Spermacoce
serrulata P. Beauv., Fl. Oware Benin 1: 39, t.
23. 1805. Type: AFRICA: Nigeria, Palisot de Diphragmus C. Presl, Bot. Bermerk. 80. 1844.
Beauvois s.n. (presumably P, n.v.). Type: Diphragmus scaber C. Presl. [Herein
Diodia spicata Miq., Stirp. Surin. Sel. 179, pi. treated as synonym of Staelia Cham. &
52. 1851. Type: SURINAM: Sylvarum prope SchlechtendaL]
Bergendaal, Oct, Focke [1140] (presumably Diphragmus scaber C. Presl, Bot. Bermerk. 81.
U, n.v.; this is Miquel's lectotype according to 1844. Type: MEXICO: Habitat in Mexico,
Howard, 1989). Dasycephala. precise locality and collector not cited.
Diodia teres Walter, Fl. Carol. 87. 1788. Type: —Staelia scabra (C. Presl) Standi.
U.S.A.: In sabulosis sylvarum a Virginia ad
Carolinam, Walter s.n. (location unknown.) Dressleriopsis Dwyer, Ann. Missouri Bot. Gard.
Diodia teres Walter var. angustata A. Gray, 67: 153. 1980. Type: Dressleriopsis
Syn. Fl., ed. 2. 1(2): 35. 1886. Type: panamensis Dwyer [Herein treated as
MEXICO: Sonora, between, San Pedro and synonym of Lasianthus Jack.]
Santa Cruz, Sep 1851, C. A. Wright 1122 Dressleriopsis panamensis Dwyer, Ann.
(Holotype GH, n.v.).—Diodia teres Walter Missouri Bot. Gard. 67: 154. 1980. Type:
Diodia teres Walter subsp. angustata (A. Gray) PANAMA: Panama, near La Eneida, beyond
Steyerm., Acta Bot. Venez. 6: 184. 1971. Cerro Jefe, 25 Jul 1968, R. Dressier 3560
Diodia teres Walter var. angustata A. Gray, (Holotype MO 2267568 (photo PTBG);
Syn. Fl. ed. 2 1(2): 35. 1886. Type: MEXICO: Isotypes MEXU, US (photos MEXU, PTBG)).
Sonora, between San Pedro and Santa Cruz, —Lasianthus panamensis (Dwyer) Robbr.
Sep 1851, C. A. Wright 1/22 (Holotype GH,
n.v.).—Diodia teres Walter Duggena Vahl in H. West, Bidr. Beskr. Ste.
Diodia teres Walter subsp. prostrata (Sw.) Croix 269. 1793. Nomen nudum. Type: Not
Steyerm., Acta Bot. Venez. 6: 185. 1971. cited. [Herein treated as a synonym of
Diodia prostrata Sw., Prodr. Nov. Gen. Sp. Gonzalagunia Ruiz & Pav.j Note: Standley
PI. 30. 1788. Type: JAMAICA: Loefling (Contr. U.S. Natl. Herb. 18: 124-125. 1916)
(location unknown).—Diodia teres Walter transferred a number of names from Gonzalea
Diodia tetracocca (M. Martens & Galeotti) and Gonzalagunia to Duggena, citing
Hemsl., Diag. PI. Nov. 1:32. 1879. Duggena Vahl (in West 1793) as the oldest
Spermacoce ? tetracocca M. Martens & available generic name. However, Howard
Galeotti, Bull. Acad. Roy. Soc. Bruxelles 11: (1989: 417) notes that when H. West proposed
132. 1844. Type: MEXICO: Michoacan, the name, he "gave no generic description for
Santa Maria, cerca Morelia, H. Galeotti 2587 Duggena, referring only to Barleria hirsuta
(Holotype BR, n.v.; Isotype K, n.v.). Jacq. [Acanthaceae] while supplying the new
—Richardia tricocca (Torr. & A. Gray) name Duggena richardii. " Consequently
Standi, subsp. tetracocca (M. Martens & Duggena Vahl in H. West is an invalid name.
Galeotti) W. H. Lewis & R. L. Oliv. The reason Duggena was attributed to Vahl is
Diodia tricocca Torr. & A. Gray, FL N. Amer. that West (loc. cit. p. 263) stated the new
2: 30. 1841. Type: U.S.A.: Texas, Drummond names given in italics are Vahl's.
117 (Holotype GH, n.v.; Isotype G, n.v.). Duggena bracteosa (Donn. Sm.) Standi., Contr.
—Richardia tricocca (Torr. & A. Gray) U.S. Natl. Herb. 18: 125. 1916. Gonzalea
Standi. bracteosa Donn. Sm., Bot. Gaz.
Diodia villosa Sesse & Moc. ex DC, Prodr. 4: (Crawfordsville) 33: 252. 1902. Type:
562. 1830. Type: MEXICO: Sesse & Mocino COSTA RICA: Suerre, Llanuras de Santa
sn. (presumably Herb. Dunat & Herb. Clara, 300 m, Apr 1896, J. D. Smith 6583
 (Holotype US 943468 (photos MEXU,
PTBG); Isotypes F, n.v., NY, n.v., US (photos
MEXU, PTBG)).—Gonzalagunia bracteosa
(Donn. Sm.) B. L. Rob. Duhamelia Pers., Syn. 1: 203. 1805, nom.
Duggena hayesii (Wernham) Standi., Contr. superfl. Based on Hamelia patens Jacq. 1760.
U.S. Natl. Herb. 18: 125. 1916, comb, inval. [Herein treated as a synonym of Hamelia
Gonzalea hayesii Wernham, J. Bot. 51:219. Jacq.]
1913. Type: PANAMA: Panama, in woods Dunhameliapatens (Jacq.) Pers., Syn. 1: 203.
near Panama, Hayes s.n. (Holotype BM, n.v.). 1805. Type: SANTO DOMINGO: Jacquin
—Gonzalagunia panamensis (Cav.) K. s.n. (location unknown).—Hamelia patens
Jacq.
Duggena nivea (Bartl. ex DC.) Standi., Contr.
U.S. Natl. Herb. 18: 125. 1916, comb, inval. Dukea Dwyer, Ann. Missouri Bot. Gard. 53:
Gonzalea nivea Bartl. ex DC, Prodr. 4: 436. 360-367. 1966. Type: Dukea panamensis
1830. Type: MEXICO: Without precise Dwyer. [Herein treated as a synonym of
locality, Haenke s.n. (Holotype presumably Raritebe Werhnam.]
PR, n.v.). The specimen Lesta & Gomes 42 Dukea blumii Dwyer, Ann. Missouri Bot. Gard.
(W, n.v. (photo MO)) is not the type. 53: 365. 1966. Type: PANAMA: Colon,
—Rondeletia buddleoides Benth. around Dos Bocas, Rio Fato Valley, H. Pittier
Duggena ovatifolia (Dorm. Sm.) Standi., Contr. 4223 (Holotype: MO 1227470 (photo PTBG);
U.S. Natl. Herb. 18: 126. 1916, comb, inval. Isotype US). Raritebe.—Raritebe
Gonzalea ovatifolia Donn. Sm., Bot. Gaz. palicoureoides Wernham subsp.
(Crawfordsville) 27: 336. 1899. Type: dwyerianum J. H. Kirkbr.
COSTA RICA: Cartago, Atirro, J. D. Smith rienensis Dwyer, Ann. Missouri Bot.
6599 (Lectotype US 943467 (photos MEXU, Gard. 53: 365. 1966. Type: PANAMA:
PTBG); Isolectotypes F, n.v., GH, US). Darien, Chepigana District, Cana-C UJM I tail
—Gonzalagunia ovatifolia (Donn. Sm.) (Camp 2), M. E. Teny & R. A. Terry 1487
B. L. Rob. (Holotype MO 1196543 (photo PTBG);
Duggena panamensis (Cav.) Standi., Contr. U.S. Isotype F)). Raritebe.—Raritebe
Natl. Herb. 18: 126. 1916, comb, inval. Buena palicoureoides Wernham subsp.
panamensis Cav., Anal. Hist. Nat. 2: 279. dwyerianum J. H. Kirkbr.
1800. Type: PANAMA: Lancon, Nee s.n. Dukea euryphylla (Standi.) Dwyer, Ann.
(Holotype MA, n.v.; Isotype F (photo Missouri Bot. Gard. 53: 366. 1966. Coussarea
PTBG)).—Gonzalagunia panamensis | ( a. » euryphylla Standi., Publ. Field Columbian
K. Schum. Mus., Bot. Ser. 4: 294. 1929. Type:
Duggena rudis Standi., Contr. U.S. Natl. Herb. PANAMA: Bocas del Toro, Almirante (or
18: 125. 1916. Type: PANAMA: Panama, Buena Vista Camp on Chiriqui Trail), G. P.
shady places along Rio Chagres at El Vigia, Cooper 228 (Holotype F, n.v.). Raritebe.
12 Jan 1911, H. Pittier 2378 (Holotype US —Raritebe palicoureoides Wernham subsp.
676602 (photo PTBG)).—Gonzalagunia dwyerianum J. H. Kirkbr.
rudis (Standi.) Standi. Dukea panamensis Dwyer, Ann. Missouri Bot.
Duggena thyrsoidea (Donn. Sm.) Standi., Contr. Gard. 53: 364. 1966. Type: PANAMA:
U.S. Natl. Herb. 18: 126. 1916, comb, inval. Darien, Cana and vicinity, 2000-6500 ft., 17
Gonzalea thyrsoidea Donn. Sm., Bot. Gaz. Apr-8 Jun 1908, E. L. Tyson, J. Dwyer & K.
(Crawfordsville) 13: 188. 1888. Type: Blum 3326 (Holotype MO 2084467 (photo
GUATEMALA: Alta Verapaz, Berge bei PTBG)). Raritebe.—Raritebe palicoureoides
Pansamala, 3800 ft., May 1887, H. von Wernham subsp. dwyerianum J. H. Kirkbr.
Tuerckheim 1249 (Holotype US 941409
A NOMENCLATOR OF MEXICAN AND CENTRAL

Dukea trifoliata Dwyer & Hayden, Ann. (Holotype CR, n.v.; Isotypes MO 3030387
Missouri Bot. Gard. 54: 139. 1967. Type: (photo PTBG), TEX-LL).—Alibertia
PANAMA: Code, El Valle, potato farm utleyorum (Dwyer) C. M. Taylor
above village of El Valle, 12 Jul 1962, J. D.
Dwyer 1933 (Holotype MO 3831231 (photos Edithea Standi., Trop. Woods 34: 1. 1933. Type:
MO, PTBG)). Raritebe.—Raritebe Edithea floribunda Standi. [Herein treated as a
palicoureoides Wernham subsp. synonym of Deppea Cham. & Schltdl.]
dwyerianum J. H. Kirkbr. Edithea floribunda Standi, Trop. Woods 34: 1.
Dukea victoriae Dwyer, Ann. Missouri Bot. 1933. Not Deppea floribunda Hemsl., 1879.
Gard. 53: 364. 1966. Type: PANAMA: Type: MEXICO: Guerrero, Malinantepec,
Darien, Cerro Pirre, cloud forest, Oct 1962, Mar 1930, L. Schultze 363 (Holotype B,
J. A. Duke 6099 (Holotype MO 1840647 presumably destroyed, fragment F (photos
(photo PTBG); Isotype MO 1832075 (photo MEXU, PTBG)).—Deppea schultzei Lorence
PTBG)). Raritebe.—Raritebe palicoureoides
Wernham subsp. dwyerianum J. H. Kirkbr. Eizia Standi., Publ. Field Mus. Nat. Hist., Bot.
Ser. 22: 50. 1940. Type: Eizia mexicana
Duroia L. f, Suppl. PI. 209. 1781. Nomen. cons. Standi.
Type: Duroia eriopila L. f. Eizia mexicana Standi., Publ. Field Mus. Nat.
Duroia costaricensis Standi., Conrr. U.S. Natl. Hist., Bot. Ser. 22: 50. 1940. Type: MEXICO:
Herb. 20: 208. 1919. Type: COSTA RICA: Chiapas, Mt. Tacana, 1000-2000 m, Aug
Marais de Sierpe, Mar 1892, H. Pittier 6803 1938, E. Matuda 2403 [sphalm. as 2402 in
(Holotype US 938658 (photos MEXU, protologue] (Holotype MICH, n.v. (fragment
PTBG)). The name Duroia hirsuta (Poepp. & and photo F); Isotypes A, n.v., TEX-LL
Endl.) K. Schum., an Amazonian species, was (photos MEXU, PTBG), US, 2 sheets (photos
misapplied to this species by Dwyer (1980: MEXU, PTBG)).
155).
Duroia genipifolia Standi. & Steyerm., Publ. Elaeagia Wedd., Monogr. Cinch. 94. 1849.
Field Mus. Nat. Hist., Bot. Ser. 22: 186. 1940. Type: Elaeagia utilis (Goud.) Wedd.
Type: GUATEMALA: Izabal, Rio Dulce, Elaeagia asperula Standi, ex Steyerm., Acta
between Livingston and 6 miles up river, on Biol. Venez. 4: 37. 1964. Type: COLOMBIA:
north side (right side going up river), sea Valle, Rio Calima (region del Choco), La
level, 14 Apr 1940, J. A. Steyermark 39382 Trojita, 5-50 m, 19 Feb-10 Mar 1944, J.
(Holotype F 1035218 (photo PTBG)). Cuatrecasas 16647 (Holotype VEN, n.v.;
—Randia genipifolia (Standi. & Steyerm.) Isotype F (photo PTBG)).
Elaeagia auriculata Hemsl., Diag. PI. Nov.
Duroia panamensis Dwyer in Dwyer & Hayden, Mexic. 32. 1879. Type: COSTA RICA:
Ann. Missouri Bot. Gard. 55: 38. 1968. Type: Without precise locality, alt. 4500 ped.,
PANAMA: Bocas del Toro, Duwebdulup Endres 95 (Holotype K, n.v.)
Peak, N of Rio Teribe across from W. Huron Elaeagia karstenii Standi., Publ. Field
(behind chiefs house), 300-900 ft., 13 Apr Columbian Mus., Bot. Ser. 8: 50. 1930. Type:
1968, J. H. Kirkbride &J.A. Duke 571 VENEZUELA: Near Colonia Tovar,
(Holotype MO 1968346); Isotype MO). 1854-55, A. Fendler 594 (Holotype GH, n.v.,
—Hippotis albiflora H. Karst. fragment F; Isotypes MO, NY, n.v.).
Duroia utleyorum Dwyer in W. Burger & C. M. Elaeagia myriantha (Standi.) C. M. Taylor &
Taylor, Fieldiana, Bot., n.s. 33: 127. 1993. Hammel, No von 3: 490. 1993. Sickingia
Type: COSTA RICA: Puntarenas, Rincon de myriantha Standi., Publ. Field Columbian
Osa, stream and slopes adjacent to Airfield, Mus., Bot. Ser. 7: 27. 1930. Type:
20-300 m, 6-7 Feb 1974, R. L. Liesner 1774 COLOMBIA: Magdalena, Las Nubes road,
 region of Santa Marta, 1200 m, 3 Dec 1898, Ernodea Sw., Prodr. 29. 1788. Type: Ernodea
H. H. Smith 1810 (Holotype F 138660; littoralis Sw.
Isotypes MO, NY, n.v., US, n.v., W, n.v., Y, Ernodea littoralis Sw., Prodr. Veg. Ind. Occ.
n.v.). Holtonia, Simira. 29, pi. 223, tab. 4. 1788. Type: JAMAICA:
Elaeagia nitidifolia Dwyer, Ann. Missouri Bot. Not designated; Swartz referred to the Sloan
Gard. 67: 157. 1980. Type: PANAMA: and Browne descriptions. According to
Panama, region of Cerro Jefe, ca. 1000 m, 3 Howard (1989) a lectotype needs to be
Oct 1969, M. D. Correa A., R. L. Dressier, N. designated.
Escobar & W. H. Lewis 1608 (Holotype MO
2424954 (photo PTBG); Isotype MO 2424954 Evea Aubl., Hist. PI. Guian. 1: 100, t. 39. 1775.
(photo PTBG)). nom. rej. vs. Cephaelis Sw. 1788, nom. cons.
Elaeagia uxpanapensis Lorence, Bol. Soc. Bot. Type: Evea guianensis Aubl. [Herein treated
Mexico 45: 66, fig. 1. 1983 (1984). Type: as a synonym of Psychotria L.]
MEXICO: Oaxaca, Mun. de Matias Romero Evea axillaris (Sw.) Standi., Contr. U.S. Natl.
(Zona de Uxpanapa), a 2 km SE del Herb. 18: 123. 1916. Cephaelis axillaris Sw.,
aserradero La Floresta sobre el camino a Rio Prodr. 3: 45. 1788. Not Psychotria axillaris
Verde, a 15 km S de Esmeralda, 250 m, 25 Willd., 1798. Type: WEST INDIES: St.
May 1981, T. Wendt, S. Koch, A. Villalobos & Christopher, Swartz s.n. (Holotype
J. Garcia 3299 (Holotype MEXU; Isotypes presumably S, n.v.). Tapogomea.
CHAPA (33 others were also distributed). —Psychotria aubletiana Steyerm.
Evea chiapensis Standi., Contr. U.S. Natl. Herb.
Ereicoctis Kuntze, Rev. Gen. PI. 1: 281. 1891. 23: 1392. 1926. Not Psychotria chiapensis
Type: Not designated. Standi, 1926. Type: MEXICO: Chiapas,
Ereicoctis acerosa (A. Gray) Kuntze, Rev. Gen Cerro del Boqueron, C. A. Purpus 6928
PI. 1: 281. 1891. Hedyotis acerosa A Gray (Holotype US 567214 (photos MEXU,
PI. Wright. 1: 81. 1852. U.S.A.: Texas, high PTBG); Isotypes F, MO (photos MEXU,
prairies, from Live Oak Creek to the Las PTBG)). Cephaelis, Psychotr ~ ' '
Moras, Jun 1849, C. Wright 237 (Holotype
GH, n.v.; Isotype US).—Hedyotis acerosa A Evea dichroa Standi., Contr. U.S. Natl. Herb.
Gray 18: 124. 1916. Type: PANAMA: Chinqui,
Ereicoctis cervantesii (Kunth) Kuntze, Rev. Cuesta de las Palmas, S slope of Cerro de la
Gen. PI. 281. 1891. Hedyotis cervantesii Horqueta, 1700-2100 m, 17-19 Mar 1911, H.
Kunth in Humb., Bonpl. & Kunth, Nov. Gen Pittier 3218 (Holotype US 677615 (photos
Sp. 3: 390. 1820. Type: MEXICO: A. MEXU, PBTG); Isotype F). Cephaelis.
Bonpland s.n. (presumably P-Bonpland, n.v.; —Psychotria dichroa (Standi.) C. M. Taylor
Isotype B, presumably destroyed (photo Evea elata (Sw.) Standi., Contr. U.S. Natl. Herb.
MO)).—Hedyotis cervantesii Kunth 18: 123. 1916. Cephaelis elata Sw., Prodr.
Veg. Ind. Occ. 45. 1788. Type: JAMAICA:
Enthahs P. Browne, Civ. Nat. Hist. Jamaica Swartz s.n. (presumably S, n.v.).—Psychotria
165, t. 17, fig. 3. 1756. Lectotype: Enthalis elata (Sw.) Hammel
fruticosa L. Evea guapilensis Standi., J. Wash. Acad. Sci.
Erithalis fruticosa L., Syst. Nat., ed 10 2- 930 15: 104. 1925. Type: COSTA RICA: Limon,
1759. Type: JAMAICA: Not designated vicinity of Guapiles, 300-500 m, 12-13 Mar
based on Browne, Civ. Nat. Hist. Jamaica t. 1934, P. C. Standley 37025 (Holotype US
E7,fig. 3. According to Howard (1989), a 1153029 (photos MEXU, PTBG)). Cephaelis.
Browne figure or specimen would be a' —Psychotria guapilensis (Standi.) Hammel
probable lectotype. Evea ipecacuanha (Brot.) Standi., Contr. U.S.
Natl. Herb. 18: 123. 1916. Callicocca
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

ipecacuanha Brot., Trans. Linn. Soc. London designated by McDowell (1996b: 289)).
6: 137. 1802. Type: BRAZIL: Without precise Howard (1989) stated that no type specimens
locality, Gomez s.n. (presumably LISU, n.v.). are known and a neotype needs to be
Cephaelis.—Psychotria ipecacuanha (Brot.) designated. Later, T. McDowell, in his review
Stokes of the genus, selected a lectotype from among
Evea nana Standi., J. Wash. Acad. Sci. 15: 105. original material.
1925. Cephaelis nana (Standi.) Standi. Type: Exostema coulteri Hook. f. ex Hemsl., Diag. PI.
PANAMA: Canal Zone, hills N of Frijoles, 19 Nov. 32. 1879, [as Exostemma]. Type:
Dec 1923, P. C. Standley 27550 (Holotype US MEXICO: Hidalgo, Zimapan, Coulter 209
1153871 (photos MEXU, PTBG); Isotype F). (Holotype K, n.v.; Isotype MO).
Psychotria guapilensis (Standi.) —Syringantha coulteri (Hook. f. ex Hemsl.)
Hammel T. McDowell
Evea tomentosa (Aubl.) Standi, Contr. U.S. bandl., N. Amer. Fl. 32:
Natl. Herb. 18: 123. 1916. Tapogomea 126. 1921. Type: MEXICO: Oaxaca, Villa
tomentosa Aubl., Hist. PI. Guiane 160. 1775. Alta, 3000 ft., Aug, H. Galeotti 2664
Type: FRENCH GUIANA: Aublets.n. (Holotype US 572997; Isotypes P, 2 sheets
(Holotype presumably P or BM, n.v.; Isotype (photos MEXU, PTBG)).—Exostema
B, presumably destroyed (photo MO)). mexicanum A. Gray
Cephaelis.—Psychotria poeppigiana Muell. Exostema longicuspe Oerst., Vidensk. Meddel.
Arg. Dansk Naturhist. Foren. Kjobenhavn 1852:
48. 1853. Type: COSTA RICA: In insula San
Exandra Standi., J. Wash. Acad. Sci. 13: 292. Lucas, Aug 1849, Oersted 11403 (Holotype
1923. Type: Exandra rhodoclada Standi. C, n.v. (photo US)).—Exostema caribaeum
[Herein treated as a synonym of Simira Aubl.] (Jacq.) Roem. & Schult.
Exandra rhodoclada Standi., J. Wash. Acad. Exostema mexicanum A. Gray, Proc. Amer.
Sci. 13: 292. 1923. Type: MEXICO: Oaxaca, Acad. Arts 5: 180. 1861. Type: MEXICO:
between La Venta and Niltepec, 200 ft., 14 Jul [Presumably Veracruz] Prov. Huasteca,
1895, E. W. Nelson 2796 (Holotype US Wartenberg, near Tantoyuca, 1858, L. C
229232 (photos MEXU, PTBG); Isotype F Ervendberg 125 (Holotype GH, n.v.; Isotype
(photos MEXU, PTBG)).—Simira P (photos MEXU, PTBG)).
rhodoclada (Standi.) Steyerm. Exostema occidentale Benth., Bot. Voy. Sulphur
104. 1844. Type: COLOMBIA: Narino, Isla
Exostema (Pers.) Bonpl. in Humb. & Bonpl., PL Gorgona, Barclay 354 or Hinds 354
Aequin. 1: 131. 1807 ("1808"). Lectotype: (Holotype K, n.v.).—Rustia occidentalis
Exostema caribaeum (Jacq.) Roem. & Schult. (Benth.) Hemsl.
(designated by Rogers, 1987: 165).
Exostema canescens Bartl. ex DC, Prodr. 4: Faramea Aubl., Hist. PI. Guian. 1: 192, t. 40.
359. 1830. Type: MEXICO: Without precise 1775. Type: Faramea corymbosa Aubl.
locality, Haenke s.n. (Holotype presumably Faramea altipetens Dwyer, Ann. Missouri Bot.
PR, n.v., fragment G-DC, n.v. (microfiche 4: Gard. 67: 162. 1980. Type: PANAMA:
359.3)).—Status uncertain ("scarcely of this Darien, Cerro Tacarcuna, south face of
genus" according to Standley, 1926). westernmost peak, lower montane wet forest,
Exostema caribaeum (Jacq.) Roem. & Schult., 1500 m, 30 Jan 1975, A. Gentry & S. Mori
Syst. Veg. 5: 19. 1819. Cinchona caribaea 14013 (Holotype MO 2272402 (photo
Jacq., Enum. PI. Carib. 16. 1760. Type: PTBG)).
DOMINICA: Without precise locality, De Faramea belizensis Standi., Carnegie Inst.
Ponthieu s.n. (lectotype BM, n.v.; illustrated Wash. Publ. 461: 90. 1935. Type: BRITISH
in Lambert, Desc. Cinchona 24, t. 4. 1797; HONDUDRAS (BELIZE): Camp 36, British
 Honduras-Guatemala survey, 2800 ft., 1 Jul Faramea cobana Donn. Sm., Bot. Gaz.
1934, W. A. Schipp S-721 (Holotype F 733535 (Crawfordsville) 57: 422. 1914. Type:
(photo PTBG); Isotypes A, n.v., F, G, n.v., GUATEMALA: Alta Verapaz, Coban, 1550
MO (photos MEXU, PTBG)).—Faramea m, Jul 1912, H. von Tuerckheim 2474
occidentalis (L.) A. Rich. (Holotype US 943628 (photos MEXU,
Faramea bocataurensis Dwyer, Ann. Missouri PTBG)).
Bot. Gard. 67: 163. 1980. Type: PANAMA: Faramea correae C. M. Taylor, Novon 6: 289,
Bocas del Toro, eastern slope of La Zorra to fig 1C, D. 1996. Type: PANAMA: Panama,
Divide, mossy cloud forest, 19 Apr 1968, Parque Nacional Altos de Campana, sendero
J. H. Kirkbride & J. A. Duke 806 (Holotype de Interpretacion, 1 km al E del campamento
MO 1968230 (photos MEXU, PTBG); Isotype de los guardaparques de INRENARE, 8°40'N,
MO 2165813 (photos MEXU, PTBG)). 79°55'W, 800-900 m, 9 Sep 1993, M. D.
Faramea brachysiphon Standi., Publ. Field Correa A., E. Montenegro & E. Hidalgo 9983
Columbian Mus., Bot. Ser. 8: 62. 1930. Type: (Holotype PMA 38229, n.v.; Isotypes MO
BRITISH HONDURAS (BELIZE): 4641338,4658116).
Middlesex, 200 ft., 17 Sep 1928, W. A. Schipp Faramea eurycarpa Donn. Sm., Bot. Gaz.
345 (Holotype F 603047; Isotypes A, n.v., (Crawfordsville) 44: 113. 1907. Type:
GH, n.v., MO (photos MEXU, PTBG), US COSTA RICA: Forets de Las Vueltas,
(photos MEXU, PTBG)). Tucumque, Dec 1898, A. Tonduz 12880
Faramea bullata Standi., Publ. Field Columbian (Holotype US 943627 (photos MEXU,
Mus., Bot. Ser. 4: 294. 1929. Type: PTBG); Isotypes B, presumably destroyed
PANAMA: Bocas del Toro, region of (fragment F), GH, 2 sheets (photos PTBG), P,
Almirante, Kankintoe 10 miles above 2 sheets (photos MEXU, PTBG)).
Holstein, Jan-Mar 1928, G. P. Cooper 507 Faramea hondurae Standi., J. Wash. Acad. Sci.
(Holotype F 579385 (photo PTBG); Isotype 18: 169. 1928. Type: COSTA RICA: San
NY, n.v.).—Faramea trinervia K. Schum. Jose, La Hondura, 1400 m, 16 Mar 1924,
Faramea capulifolia Dwyer in W. C. Burger & P. C. Standley 37890 (Holotype US 1153105
C. M. Taylor, Fieldiana, Bot, n.s. 33: 132. (photos MEXU, PTBG)).
1993. Type: PANAMA: Darien, Cerro Pirre, Faramea jefensis Dwyer & Hayden, Phytologia
ridgetop at 1200 m, J. P. Folsom 4262 15: 56. 1968. Type: PANAMA: Panama,
(Holotype MO 2620580). Note: the following eastern slope of Cerro Jefe in Clusia forest,
collections were also annotated as "type" by 2700 ft., 8 Feb 1966, E. L. Tyson et al. 3396
J. D. Dwyer but are not types: COSTA RICA: (Holotype MO 1936328 (photo PTBG);
Puntarenas, Monteverde Cloud forest Reserve, Isotypes F, GH, n.v., NY, n.v., US (photo
1500-1600 m, 8 Nov 1986, W. A. Haber ex E. PTBG)).
Bello C. 6235 (MO 36482259, photo PTBG). Faramea killipii Standi., Publ. Field Columbian
PANAMA: Darien, Cerro Pirre, 10-20 Jul Mus., Bot. Ser. 7: 73. 1930. Type:
1977, J. P. Folsom 4435 (MO 2620712 (photo COLOMBIA: Norte de Santander, Killip &
PTBG)). Smith 20569 (Holotype US, n.v.).
Faramea caputanguis Dwyer & Hayden, Ann. Faramea liebmannii Standi., J. Wash. Acad.
Missouri Bot. Gard. 55: 29. 1968. Type: Sci. 17: 343. 1927. Type: MEXICO: Oaxaca,
PANAMA: Panama, Cerro Jefe, 2900 ft., 29 Tepitapa, F. Liebmann 11404 (Liebm.
Jul 1967, J. D. Dwyer & G. Gauger 7375 Rubiaceae Mexic. 105) (Holotype presumably
(Holotype MO 1961869; Isotypes GH, n.v., C, n.v.; Isotype US).
K, n.v., MO (photos MEXU, PTBG), NY, ' Faramea Hesneri Dwyer, Ann. Missouri Bot.
n.v., UC, n.v., US).—Faramea papirifolia Gard. 67: 167. 1980. Type: PANAMA:
(Standi, ex Steyerm.) C. M. Taylor Panama, road to Carti (San Bias), 19 km north
of El Llano, ca. 500 m, 13 Mar 1973, P. Busey
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

913 (Holotype MO 2106781 (photos MEXU, 30-50 m, 18, 22 May 1946, J. Cuatrecasas
PTBG)). 21078 (Holotype US 2825318, n.v.; Isotypes
Faramea loftonii Dwyer & Hayden, Phytologia COL, n.v., VEN, n.v.).
15: 56. 1967. Type: PANAMA: Panama, Faramea papulata Dwyer & Hayden,
Cerro Jefe, in C/usia forest, 2700-3000 ft., 27 Phytologia 15: 57. 1967. Type: PANAMA:
Jan 1966, E. L. Tyson, J. D. Dwyer & K. Blum Panama, Cerro Jefe, in Clusia forest,
3323 (Holotype MO 1824505 (photo PTBG); 2700-3000 ft., 27 Jan 1966, E. L. Tyson, J.
Isotype MO (photo PTBG)).—Coussarea Dwyer & K. Blum 3284 (Holotype MO
loftonii (Dwyer & Hayden) Dwyer 1831279 (photos MEXU, PTBG)).
Faramea luteovirens Standi., Contr. U.S. Natl. Faramea parvibracteata Steyerm., Mem. New
Herb. 18: 138. 1916. Type: PANAMA: Colon, York Bot. Gard. 17: 376. 1967. Type:
Loma de la Gloria, near Fato (Nombre de VENEZUELA: Terr. Fed. Amazonas, Cafio
Dios), 10-104 m, Aug 1911, H. Pittier 4098 Tamatama, Rio Orinoco, 150 m, 12 Sep 1954,
(Holotype US 679194 (photos MEXU, S. Silverio Level 145 (Holotype NY, n.v.).
PTBG)). Faramea pauciflora Dwyer, Ann. Missouri
Faramea multiflora A. Rich., Mem. Soc. Hist. Bot. Gard. 67: 172. 1980. Type: PANAMA:
Nat. Paris 5: 176. 1834. Type: FRENCH Veraguas, Valley of Rio Dos Bocas along
GUIANA: Richard s.n. (Holotype presumably road between Escuela Agricola Alta Piedra
P,n.v.). and Calovebora, 15.6 km northwest of Santa
Faramea myrticifolia Dwyer in W. C. Burger Fe, 450-550 m, 31 Aug 1974, T B. Croat
& C. M. Taylor, Fieldiana: Bot., n.s. 33: 134. 27575 (Holotype MO 2201361 (photo
1993. Type: COSTA RICA: Limon, Las PTBG); Isotype MO (photo PTBG)).
Brisas de Pacuarito, Siquirres, 300 m, 18 Apr Faramea permagnifolia Dwyer ex C. M.
1985, L. D. Gomez, C. Herrera, E. Herrera & Taylor, Novon 6: 298, fig. IB. 1996. Type:
W. Murillo 23401 (Holotype CR, n.v.; Isotype COSTA RICA: Puntarenas, Osa Peninsula, in
MO 3648258 (photo PTBG)). forest ca. 9 km W of Rincon on road to
Faramea occidentalis (L.) A. Rich., Mem. Soc. Rancho Quemado, 8°39'N, 83°32'W, 200 m,
Hist. Nat. Paris 5: 176. 1834. Ixora 26 May 1986, B. Hammel, M. Grayum & G.
occidentalis L., Syst., ed. 10, 2: 893. 1759. de Nevers 15212 (Holotype CR, n.v.; Isotypes
Type: JAMAICA: Based on P. Browne, Hist. MO 4658114, MO 4658115).
Jam. t.6,f.2 (LINN, n.v.), and Plumier, Icon. Faramea quercetorum Standi., J. Wash. Acad.
tab. 156, f. 2 (ed. Burmannus). Steyermark Sci. 18: 168. 1928. Type: COSTA RICA: San
(1974: 947) suggested the type was from Jose, oak forest near Quebradillas, north of
Jamaica, thus selecting the Browne reference Santa Maria de Dota, 1800 m, 24 Dec 1925,
over that of Plumier within the references P. C. Standley 42999 (Holotype US 1253066
given by Linnaeus. (photos MEXU, PTBG)).—Faramea ovalis
Faramea ovalis Standi., Contr. U.S. Natl. Herb. Standi.
18: 138. 1916. Type: PANAMA: Chiriqui, Faramea sanblasensis C. M. Taylor, Novon 6:
forests along Rio Ladrillo and vicinity, above 300, fig. I A. 1996. Type: PANAMA:
El Boquete, 1200-1300 m, W. R. Maxon 5397 Comarca de San Bias, Cerro Brewster,
(Holotype US 675764 (photos MEXU, headwaters of Rio Cangandi, 9°18'N,
PTBG)). 79° 16'W, 630 m, 24 Apr 1985, G. de Nevers,
Faramea papirifolia (Standi, ex Steyerm.) H. Herrera, B. Hammel & S. Charnley 5491
C. M. Taylor, Novon 6: 210. 1996. Cephaelis (Holotype PMA 38230, n.v.; Isotype MO
papirifolia Standi, ex Steyerm., Acta Biol. 4658117).
Venez. 4: 21. 1964. Type: COLOMBIA: Faramea scalaris Standi., Contr. U.S. Natl.
Valle, Rio Calima (region del Choco), margen Herb. 13: 139. 1916. Type: PANAMA:
derecha, lomas frente a quebrada de la Brea, Chiriqui, humid forest between Altos de las
 Palmas and top of Cerro de la Horqueta, Faramea talamancarum Standi., Publ. Field
2100-2268 m, 18 Mar 1911, H. Pittier 3265 Columbian Mus., Bot. Ser. 4: 332. 1929.
(Holotype US 677656 (photos MEXU, Type: PANAMA: Talamanca Valley, 3 Feb
PTBG)). 1921, M. A. Carleton 135 (Holotype F 578962
Faramea schultesii Standi., Bot. Mus. Leafl. (photo PTBG); Isotype US (photos MEXU,
Harvard Univ. 9: 194. 1941. Type: MEXICO: PTBG)).—Faramea multiflora A. Rich.
Oaxaca, Distr. Choapam [Choapan], Cerro de Faramea terryae Standi., Publ. Field Mus. Nat.
Lalana, near San Juan Lalana, in forest, long. Hist., Bot. Ser. 22: 189. 1940. Type:
95°45', lat. 17°25\ 1300 m, 11 May 1939, PANAMA: Darien, Chepigana Distr.,
R. E. Schultes & B. P. Reko 855 (Holotype F Cana-Cuasi Trail (Camp 2), 3000 ft., 10 Mar
1016864 (photo PTBG)). 1940, M. E. Terry & R. A. Terry 1479
Faramea sessifolia P. H. Allen, Rain Forests (Holotype F 1034536 (photo PTBG); Isotype
Golfo Dulce 409. 1956. Type: COSTA RICA: MO (photo PTBG)).
Esquinas Forest, area between the Rio Faramea trinervia K. Schum. & Donn. Sm.,
Esquinas and Palmar, 200 ft., 22 May 1950, Bot. Gaz. (Crawfordsville) 31: 115. 1901.
P. H. Allen 5539 (Holotype US 2215891 Type: COSTA RICA: Talamanca, forets de la
(photos MEXU, PTBG); Isotypes F, 2 sheets). Boca de Zhorquin, Mar 1894, A. Tonduz 8571
Faramea standleyana L. O. Williams, (Holotype US 943623 (photos MEXU,
Phytologia 26: 490. 1973. Type: PTBG)).
GUATEMALA: Izabal, Cerro San Gil, rocky Faramea trinervia K. Schum. & Donn. Sm. var.
slopes along Rio Frio, 75 m, 17 Dec 1941, suerrensis Donn. Sm., Bot. Gaz.
/ A. Steyermark 39962 (Holotype F 1134267 (Crawfordsville) 31: 115. 1901. Type:
(photo PTBG)).—Faramea occidentalis (L.) COSTA RICA: Suerre, Llanuras de Santa
A. Rich. Clara, 300 m, Feb 1896, J. D. Smith 6589
Faramea stenophylla Standi., J. Wash. Acad. (Holotype US 943626 (photos MEXU,
Sci. 18: 169. 1928. Type: HONDURAS: PTBG); Isotype US 943624 (photos MEXU,
Cuyamel, on hills, 29 Mar 1924, M. A. PTBG)).—Faramea suerrensis (Donn. Sm.)
Carlcton 592 (Holotype US 1208399 (photos Donn. Sm.
MEXU, PTBG)). Faramea woodsonii Standi., Ann. Missouri
Faramea stenura Standi., Publ. Field Bot. Gard. 28: 470. 1921. Type: PANAMA:
Columbian Mus., Bot. Ser. 4: 331. 1929. Code, North rim of El Valle de Anton, near
Type: HONDURAS: Dept. Atlantida, in wet Cerro Turega, 650-700 m, 30 Jun 1940, R. E.
Woodson, Jr. & R. W. Schery 164 (Holotype F
Lancetilla Valley, 600 m, 13 Mar 1928, P. C. 1085608 (photo PTBG); Isotypes F, GH, n.v.,
Standley 56743 (Holotype F 581582 (photo MO (photo PTBG)).
PTBG); Isotype US 1409383 (photos MEXU, Faramea zetekii Standi., Contr. Arnold Arbor. 5:
PTBG)). 147. 1933. Type: PANAMA: Canal Zone,
Faramea suerrensis (Dorm. Sm.) Donn. Sm., Barro Colorado Island, shore of cove W of
Bot. Gaz. (Crawfordsville) 44: 112. 1907. Fuertes House, 21 Feb 1932, R. H.
Faramea trinenia K. Schum. & Donn. Sm. Woodworth & P. A. Vestal 641 (Holotype A,
var. suerrensis Donn. Sm., Bot. Gaz. n.v.; Isotypes F, MO (photo PTBG)).
(Crawfordsville) 31: 115. 1901. Type: —Faramea occidentalis (L.) A. Rich.
COSTA RICA: Suerre, Llanuras de Santa
Clara, 300 m, Feb 1896, J. D. Smith 6589 Ferdinandusa Pohl, PI. Bras. Icon. Descr. 2: 8.
(Holotype US 943626 (photos MEXU, 1828-1829. (Substitute name for Ferdinandea
PTBG); Isotype US 943624 (photos MEXU Pohl, 1827, not Ferdinanda Lagasca, 1816).
PTBG)). Type: Ferdinandusa speciosa Pohl
 L NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Ferdinandusa panamensis Standi. & L. O. southern California), Douglas (Holotype PH,

Williams, Ceiba 3: 34. 1952. Type: n.v.).
PANAMA: Bocas del Toro, vicinity of Galium aschenbornii S. Schauer, Linnaea 20:
Chinqui Lagoon, Fish Creek, 15 Apr 1941, H. 701. 1847. Type: MEXICO: Hidalgo,
von Wedel 2232 (Holotype F 1120423 (photo Zimapan, Aschenborn 92 (Holotype
PTBG); Isotypes GH, n.v., MO, US (photos presumably B, n.v., fragment JE, n.v.).
MEXU, PTBG)). Galium aparine L., Sp. PI. 108. 1753. Type:
EUROPE: (LINN, n.v.). Naturalized in
Froelichia Vahl, Eclog. Amer. 1: 3. 1796. Type: northern Mexico.
Froelichia paniculata Willd. [Herein treated as Galium carmenicola Dempster, Madrono 23:
a synonym of Coussarea Aubl.] 313. 1974. Type: MEXICO: Coahuila, Mina
Froelichia paniculata Vahl, Eclog. Amer. 1: 3. El Popo, ca. 2 km south of Canon El Diablo
1796. Type: TRINIDAD: Rynan s.n. (C, n.v.). on east slope of Sierra del Carmen, M.
—Coussarea paniculata (Vahl) Standi. Johnston et al 11921 (Holotype UC, n.v.;
Isotypes MEXU, TEX, n.v.).
Galianthe Griseb., Abh. Konigl. Ges. Galium carterae Dempster, Brittonia 22: 184.
Wissensch. Gottingen 24: 157. 1879. Type: 1970. Type: MEXICO: Baja California del
Galianthe fastigiata Griseb. Sur, Sierra de la Giganta, head of south
Galianthe brasiliensis (Spreng.) E. L. Cabral & branch of Canada de Teojo in vicinity of
Bacigalupo subsp. angulata (Benth.) E. L. Portezuelo de Peloteado (southwest of Notri),
Cabral & Bacigalupo, Ann. Missouri Bot. A. Carter & Medellin-Leal 4656 (Holotype
Gard. 84: 863. 1997. Triodon angulatum UC, n.v.).
Benth., PI. Hartweg. 70. 1840. Type: Galium coronadoense Dempster, Univ. Calif.
MEXICO: Oaxaca, Teotolcingo in montibus Publ. Bot. 64: 31. 1973. Type: MEXICO:
Chinantla, T. Hartweg 507 (Holotype K, n.v.). Coronado Islands, South Island, R. Moran
Diodia, Triodon. 8317 (Holotype UC M243903, n.v.; Isotypes
DS.i . SD. :
Galium L., Sp. PI. 1: 105. 1753. Type: Galium Galium correllii Dempster, Sida 3: 249. 1968.
Type: U.S.A.: Texas, Val Verde Co., Eagle
Taxon41:563. 1992). Nest (Langtry) Canyon, E edge of Langtry, 9
Galium aculeolatum Dempster, Univ. Calif. May 1967, D. S. Correll 34155 (Holotype
Publ. Bot. 64: 27. 1973. Type: MEXICO: Baja UC, n.v.; Isotypes ENCB, TEX-LL, n.v., US
California del Sur, north ridge of Cerro (photo PTBG)).
Azufre, R. Moran 18755 (Holotype UC Galium dempsterae B. L. Turner, Madrono 30:
200580, n.v.). 31. 1983. Type: MEXICO: Nuevo Leon, NW
Galium andrewsii A. Gray, Proc. Amer. Acad. slope of Cerro Pena Nevada, ca. 7 km NE of
Arts 6: 537. 1865. Syntypes: U.S.A.: San Antonio Pena Nevada (ca. 30 km NW of
California, without precise locality, Andrews Doctor Arroyo), Jul 1977, C. Wells & G.
s.n. (GH, n.v.), Shelton s.n. (GH, n.v.). Nesom 514 (Holotype TEX-LL, n.v.).
Galium angulosum A. Gray, Proc. Amer. Acad. Galium denticulatum Bartl. ex DC, Prodr. 4:
Arts 11: 74. 1876. Type: MEXICO: Baja 612. 1830. Type: MEXICO: Hidalgo, Real del
California, Guadalupe Island, 1875, E. Palmer Monte, Haenke (Holotype presumably PR,
36 (Holotype GH, n.v.; Isotypes MO, 2 sheets n.v.; Isotypes GOET, n.v., W, n.v.).
(photos MEXU, PTBG), NY, n.v., UC, n.v.). Galium diabolense Dempster, Univ. Calif.
Galium angustifolium Nutt. in Torr. & A. Publ. Bot. 64: 28. 1973. Type: MEXICO: Baja
Gray, Fl. N. Amer. 2: 22. 1841. Type: U.S.A.: California, Canon del Diablo, N and W of
California, San Francisco (probably from Picacho del Diablo, E flank of Sierra San
Pedro Martir, 4000 ft., 14 Jun 1954,
 Chambers 571 (Holotype UC M034766, n.v.; Galium glaberrimum Hemsl., Diag. PI. Nov.
Mexic. 3: 53. 1878. Type: MEXICO: Without
Galium fendleri A. Gray, PI. Fendl. 60. 1849. precise locality, Rubiaceae Mexic. Liebmann
Type: U.S.A.: New Mexico, Santa Fe Co., 220 (= Liebmann PI. Mex. 11426) (Holotype
valley of Santa Fe Creek, Fendler 288 K, n.v.; Isotype US (photos MEXU, PTBG)).
(Holotype GH, n.v.; Isotypes DS, n.v., UC, Galium hypadenium S. Schauer, Linnaea 20:
700. 1847. Type: MEXICO: Without precise
Galium fuscum M. Martens & Galeotti, Bull. locality, Aschenborn 396 (Holotype B, n.v.,
Acad. Bruxelles 11: 126. 1844. Type: presumably destroyed). Neotype: MEXICO:
MEXICO: Oaxaca, cordillera, H. Galeotti Puebla, dry limestone ledges near Tehuacan,
2633 (Holotype BR, n.v.; Isotype K, n.v.). 5000 ft., 27 Aug 1897, C. G. Pringle 6694
Galium fuscum M. Martens & Galeotti subsp. (UC 102526, n.v.; Isoneotypes BM, n.v., F,
altiplanicum Dempster, Univ. Calif. Publ. GH, n.v., K, n.v., MEXU, MICH, n.v., MO
Bot. 64: 16. 1973. Type: MEXCICO: Jalisco, (photos MEXU, PTBG), NY, n.v., US;
near Guadalajara, C. G. Pringle 5046 designated by Dempster, 1973a).—Galium
(Holotype GH, n.v.; Isotype MEXU). fuscum M. Martens & Galeotti subsp.
Galium fuscum M. Martens & Galeotti subsp. hypadenium (S. Schauer) Dempster
guerrericum Dempster, Univ. Calif. Publ. Galium hypocarpium (L.) Endl. ex Griseb., Fl.
Bot. 64: 19. 1973. Type: MEXICO: Guerrero, Brit. W. I. 4: 351. 1861. Valantia hypocarpia
between Chilapa and Tixtla, 5200-7000 ft., L., Syst. Nat., ed. 10, 1307. 1759. Type:
E. W. Nelson 2166 (Holotype US, n.v.; JAMAICA: Rubia I, Browne, Hist. Jamaica
Isotype GH, n.v.). 141 (LINN, n.v.). Galium hypocarpium (L.)
Galium fuscum M. Martens & Galeotti subsp. Fosberg, Sida 2: 386. 1966, redundant
hypadenium (S. Schauer) Dempster, Univ. combination. Relbunium.
Calif. Publ. Bot. 64: 16. 1973. Galium Galium hystricocarpum Greenm., Proc. Amer.
hypadenium S. Schauer, Linnaea 20: 700. Acad. Arts 33: 464. 1898. Type: MEXICO:
1847. Type: MEXICO: Puebla, dry limestone Chihuahua, cool cliffs and ledges, mountains
ledges near Tehuacan, 5000 ft., 27 Aug 1897, near Chihuahua, 23 Oct 1886, C. G. Pringle
C. G. Pringle 6694 (Neotype UC 102526, 741 (Holotype US 19277 (photos MEXU,
n.v.; Isoneotypes BM, n.v., F, GH, n.v., K, PTBG); Isotypes BM, n.v., GH, n.v., K, n.v.,
n.v., MEXU, MICH, n.v., MO (photos MEXU, MO (photos MEXU, PTBG), NY,
MEXU, PTBG), NY, n.v., US; designated by n.v., RSA, n.v., TEX-LL, n.v., UC, n.v., US, 2
Dempster, 1973a). sheeets (photos MEXU, PTBG)).
Galium galeottianum Walp., Rep. Bot. Syst. 6: Galium iltisii Dempster, Univ. Calif. Publ. Bot.
17. Sept. 1846, nom. superfl. Based on Rubia 73: S,fig. 1. 1978. Type: MEXICO: Guerrero,
( M. Martens & Galeotti, Bull. Sierra Madre del Sur, along the Milpillas-
Acad. Roy. Belg. 11: 127. 1844. Not Galium Atoyac road, ca. 13 mi NE of Puerto del
acuminatum Ball, 1873. Type: MEXICO: Gallo, J. Reveal et al. 4306 (Holotype UC
Puebla, Tehuacan, 5000 pieds, H. Galeotti 1415881, n.v.).
2362 (Lectotype K, n.v.; Isolectotype P, n.v. Galium jalapense Schltdl., Linnaea 9: 591.
(photo MO); designated by Dempster, 1834. Type: MEXICO: Veracruz, in sylvis
1973a).—Galium aschenbornii S. Schauer umbrosis Jalapae, Schiede & Deppe 247
Galium geminiflorum M. Martens & Galeotti, (Holotype HAL, n.v.; Isotypes BM, n.v., MO
Bull. Acad. Bruxelles 11: 126. 1844. Type: (photos MEXU, PTBG)).—Galium triflorum
MEXICO: Veracruz, regiones du pic
d'Orizaba, vers 9500 pieds, H. Galeotti 2604 Galium juniperinum Standi., Publ. Field Mus.
(Holotype BR, n.v.; Isotype K, n.v.). Nat. Hist., Bot. Ser. 22: 114. 1940. Type:
—Galium aschenbornii S. Schauer MEXICO: Nuevo Leon, Mun. Galeana, within
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

1 mi of Galeana, earth wall of canyon, 5400 l Kunth subsp. flexicum

ft., 4 Aug 1939, V. H. Chase 7772 (Holotype Dempster, Madrono 23: 384. 1976. Type:
F 1002599 (photo PTBG); Isotypes GH, n.v., U.S.A.: Texas, Brewster Co., Chisos
MO (photos MEXU, PTBG), NY, n.v., TEX- Mountains, C. H. Mueller 8281 (Holotype
LL, n.v., US (photos MEXU, PTBG)). TEX, n.v.; Isotypes GH, n.v., MICH, n.v.,
(,uli l lacriforme Dempster, Madrono 23: NY,:
108. 1975. Type: MEXICO: Nuevo Leon, in Galium mexicanum Kunth var. glabratum
oak-pine forest 8 mi east of Dulces Nombres, Greenm., Contr. Gray Herb., n.s. 14: 458.
1550 m, 18 Jun 1948, F G. Meyer & D. J. 1898. Type: MEXICO: Oaxaca, west slope of
Rogers 2586 (Holotype MO 1597282 (photos Mt. Zempoaltepec, 7700-8000 ft., 5-13 Jul
MEXU, PTBG); Isotypes BM, n.v., BR, n.v., 1894, E. W. Nelson 583 (Holotype US 569280
GH. (photo PTBG); Isotypes GH, n.v., MO (photos
Galium leucotrichum Hemsl., Diagn. PL Nov. MEXU, PTBG)).—Galium mexicanum
Mexic. 53. 1880. Syntypes: MEXICO:
Veracruz, region of Orizaba, Muller J 74 (K, Galium mexicanum Kunth var. platyphyllum
n.v., GH, n.v.), Bourgeau 3125 (K, n.v., P, Greenm., Contr. Gray Herb., n.s. 14: 458.
n.v., fragment F), Sumichrast 514 (K, n.v.); 1898. Type: MEXICO: Chiapas, near San
Puebla, Tehuacan, F. Liebmann 238 (K, n.v.). Cristobal, 7000-8800 ft., 18 Sep 1895, E. W.
—Galium fuscum M. Martens & Galeotti Nelson 3165 (Lectotype US 233084 (photo
subsp. hypadenium (S. Schauer) Dempster PTBG); Isolectotype MO (photos MEXU,
Galium martirense Dempster & Stebbins, PTBG)); designated by Dempster, 1976).
Univ. Calif. Publ. Bot., 46: 50. 1968. Type: —Galium mexicanum Kunth subsp.
MEXICO: Baja California, the Oaks Pasture,
NE Meling Ranch (Rancho San Jose), east of Galium microphyllum A. Gray, PI. Wright. 1:
San Telmo, 5600 ft., 3 Jun 1964, L. T. 80. 1852. Type: U.S.A.: Texas, Jeff David
Dempster & L. Cory 3987 (Holotype UC, n.v.; Co., Mountains at the pass of the Limpia...and
Isotype F (photo PTBG)). in the valley of the Limpia, Aug 1849, C.
Galium mechudoense Dempster, Univ. Calif. Wright 234 (Holotype GH, n.v.; Isotype US
Publ. Bot. 73: 15. 1978. Type: MEXICO: Baja (photo PTBG)). Relbunium.
California del Sur, vicinity of Cerro Mechudo, Galium moranii Dempster, Brittonia 22: 190.
on talus and steep eastern escarpment south of 1970. subsp. moranii. Type: MEXICO: Baja
Portezuelo de San Antonio, 1 Nov 1971, R. California del Sur, Sierra de la Giganta,
Moran 18852 (Holotype UC 1432117, n.v.; Arroyo Hondo, north side of Cerro Giganta,
Isotype US (photos MEXU, PTBG)). 840 m, 14 Oct 1966, A. Carter & M. Sousa S.
Galium mexicanum Kunth in Humb., Bonpl. & 5172 (Holotype UC, n.v.; Isotype MO (photos
Kunth, Nov. Gen. Sp. 3: 337. 1818. subsp. MEXU, PTBG)).
mexicanum. Neotype: MEXICO: Guanajuato, Galium moranii Dempster subsp. aculeolatum
18 miles east of Guanajuato on road to (Dempster) Dempster, Univ. Calif. Publ. Bot.
Dolores, Hidalgo, O. Solbrig & R. Ornduff 73: 19. 1978. Galium aculeolatum Dempster,
4540 (UC, n.v.; designated by Dempster, Univ. Calif. Publ. Bot. 64: 27. 1973. Type:
1976). MEXICO: Baja California del Sur, north side
Galium mexicanum Kunth subsp. asperrimum of Cerro Azufre, R. Moran 18755 (Holotype
(A. Gray) Dempster, Madrono 23: 384. 1976. UC, n.v.; Isotype SD, n.v.).
Galium asperrimum A. Gray, PI. Fendl. 60. Galium nelsonii Greenm., Proc. Amer. Acad.
1879. Type: U.S.A.: New Mexico, Santa Fe, Arts 33: 460. 1898. Type: MEXICO: Oaxaca,
Fendler 289 (Holotype GH, n.v.; Isotypes summit of Mt. Zempoaltepec, 11400 ft., 9 Jul
MO, UC, n.v.). 1894, E. W. Nelson 628 (Holotype US 569281
 (photos MEXU, PTBG); Isotype GH, n.v.). Galium pennellii Dempster, Univ. Calif. Publ.
Bot. 73: 31. 1978. Type: MEXICO: San Luis
Galium nuttalii A. Gray, PL Wright. 1: 80. Potosi, Sierra Madre Oriental, Alvarez, Sierra
1852. Based on Galium suffruticosum Torr. & de Alvarez, rocky crevices of limestone,
A. Gray, Fl. N. Am. 2: 21. 1841. Not Galium 2200-2400 m, 30-31 Jul 1934, W. Pennell
suffruticosum Hook. & Am., 1833. Type: 17853 (Holotype PH, n.v.; Isotypes F (photo
U.S.A.: California, San Diego, Nuttall s.n. PTBG), MEXU, MICH, n.v., NY, n.v.).
(Holotype GH, n.v.). Galium porrigens Dempster, Madrono 22: 313.
I kdium nuttattii subsp. ovalifolium (Dempster) 1974. Based on Galium nuttallii A. Gray var.
Dempster & Stebbins, Univ. Calif. Publ. Bot. ovalifolium Dempster, Brittonia 10: 183.
46: 37. 1968. Based on Galium nuttallii A. 1958. Not Galium ovalifolium Schott ex
Gray var. ovalifolium Dempster, Brittonia 10: Schult. & Schult f., 1827. Type: U.S.A.:
183. 1958. Not Galium ovalifolium Schott f. California, Humboldt Co., Blue Slide, Van
ex Schult., 1827. Type: U.S.A.: California, Duzen River, Tracy 68 J 6 (Holotype JEPS,
Humboldt Co., Blue Slide, Van Duzen River,
Tracy 6816 (Holotype JEPS, n.v.).—Galium Galium praetermissum Greenm., Proc. Amer.
porrigens Dempster Acad. Arts 32: 306. 1897. Type: MEXICO:
Galium obstipum SchltdL, Linnaea 9: 592. 1834. Distr. Federal, pine woods, base of Sierra de
Type: MEXICO: Veracruz, in sylvis umbrosis Ajusco, 8000 ft., 19 Sep 1896, C. G. Pringle
Jalapae, May 1829, Schiede & Deppe 248 6596 (Holotype US 287866 (photos MEXU,
(Holotype HAL (n.v.); Isotypes MO (photos PTBG); Isotypes CAS, n.v., ENCB, F, GH,
MEXU, PTBG), NY, n.v., P, n.v. (photo MO), n.v., M, n.v., MEXU, MO (photos MEXU,
fragment F, W, n.v.).—Galium uncinulatum PTBG), NY, n.v., UC, n.v., US, 2 sheets).
DC. Galium pringlei Greenm., Proc. Amer. Acad.
Galium oresbium Greenm., Proc. Amer. Acad. Arts 33: 458. 1898. Type: MEXICO: Nuevo
Arts 33: 462. 1898. Type: MEXICO: Nuevo Leon, shaded ledges of the Sierra Madre near
Leon, Sierra de la Silla, dry mountain walls, Monterrey, 9 Jul 1888, C. G. Pringle 1884
16 Jul 1889, C G. Pringle 2542 (Holotype US (Holotype US 19276 (photos MEXU, PTBG);
19288 (photos MEXU, PTBG); Isotypes BR, Isotypes BM, n.v., BR, n.v., GH, n.v., K, n.v.,
n.v., F, GH, n.v., M, n.v., MEXU, MO M, n.v., MO (photos MEXU, PTBG), NY,
(photos MEXU, PTBG), NY, n.v., RSA, n v n.v., RSA, n.v., UC, n.v.).
US, n.v.). Galium proliferum A. Gray, PL Wright. 2: 67.
Galium orizabense HemsL, Diag. PL Nov. 1853. Based on Galium virgatum var.
Mexic. 3: 54. 1878. Type: MEXICO: diffusum A. Gray, PL Wright. 1: 80. 1852.
Veracruz, Orizaba, Botteri 838 (Lectotype K, Not Galium diffusum Clas in Gay, 1848.
n.v; Isolectorypes BM, n.v., P, n.v., fragment Type: U.S.A.: Texas, high, rocky hills of the
F; designated by Dempster, 1978.) Pecos, Jun 1851, C. Wright 1112 (Holotype
Galium pendulum Greenm., Proc. Amer. Acad. GH, n.v.; Isotypes GH, n.v., MO (photos
Arts 41: 249. 1905. Type: MEXICO: Hidalgo, MEXU, PTBG), UC, n.v., US (photo PTBG)).
hanging (5-8 ft.) from wet cliffs and banks, Galium quichense Dempster, Phytologia 26:
barranca below Trinidad Iron Works, 5200 ft., 12. 1973. Type: GUATEMALA: Quiche, near
6 May 1904, C. G. Pringle 8985 (Holotype junction of Nebaj road with road between
US 461457; Isotypes BM, n.v., CAS, n.v., Sacapulas and Cunen, Proctor 25139
GH, n.v., K, n.v., M, n.v., MEXU, MO (Holotype TEX-LL, n.v.; Isotypes IJ, n.v.,
(photos MEXU, PTBG), NY, n.v PH nv UC,n.v.).
POM, n.v., UC, n.v, US 1133652 (photos " Galium repens Sesse & Mo?., Fl. Mexic. 24.
MEXU, PTBG)). 1893. Type: MEXICO: Habitat in pratis
arenosis de Calpulalpam, Sesse & Mocino s.n.
A NOMENCLATOR OF MEXICAN AND CENTRAL

(presumably MA, n.v.).—Si Galium triflorum Michx., Fl. 1: 80. 1803.

probably Galium microphyllum A. Gray Type: CANADA: Location and collector
Galium rothrockii A. Gray, Proc. Amer. Acad. unknown (presumably P, n.v.).
Arts 17: 203. 1882. Type: U.S.A.: Arizona, Galium uncinulatum DC, Prodr. 4: 600. 1830.
Camp Crittenden, Rothrock 675 (Holotype Type: MEXICO: Between Tampico and Real
GH, n.v.; Isotype NY, n.v.).—Galium del Monte, Berlandier 342 (Holotype G-DC,
wrightii A. Gray n.v. (microfiche 4: 600.55); Isotype W, n.v.).
Galium rzedowskii Dempster, Bol. Soc. Bot. ( nilium uin iimhitum DC. var. nanum Greenm.,
Mexico 35: 13. 1975. Type: MEXICO: San Proc. Amer. Acad. Sci. 33: 459. 1898. Type:
Luis Potosi, Aguaje de Garcias, 10 km al MEXICO: Veracruz, Orizaba, Muller 1523
oeste de Guadalcazar, J. Rzedowski 4693 (Holotype NY, n.v.; Isotype GH, n.v.).
(Holotype ENCB; Isotype MEXU). —Galium orizabense Hemsl.
Galium seatonii Greenm., Proc. Amer. Acad. • cntr.-Amer..
Arts 33: 460. 1898. Type: MEXICO: Distr. Bot. 2: 66. 1881, nom. superfl. Based on
Federal, Serrania de Ajusco, 10000 ft., 6 Sep Ritbia acuminata M. Martens & Galeotti,
1896, C. G. Pringle 6461 (Lectotype US Bull. Acad. Roy. Sci. Bruxelles 11: 127. 1844.
96421 (photos MEXU, PTBG); Isolectotypes Not Galium acuminatum Ball, 1873. Type:
B, n.v., GH, n.v., MEXU, MO (photos MEXICO: Puebla, Tehuacan, 5000 ft., H.
MEXU, PTBG), NY, n.v., PH, n.v., UC, n.v.; Galeotti 2362 (Lectotype K, n.v.; Isolectotype
designated by Dempster, 1978). P, n.v. (photo MO); designated by Dempster,
Galium sphagnophilum (Greenm.) Dempster, 1973b).—Galium aschenbornii S. Schauer
Phytologia 26: 221. 1973. Relbunium Galium volcanense Dempster, Univ. Calif.
sphagnophilum Greenm., Proc. Amer. Acad. Publ. Bot. 64: 27. 1973. Type: MEXICO: Baja
Arts 33: 468. 1898. Type: MEXICO: Oaxaca, California del Sur, north ridge of Cerro
Sierra de Clavellinas, C. G. Pringle 5812 Azufre, R. Moran 18753 (Holotype UC
(Holotype GH, n.v.). 1380662, n.v.; Isotype GH, n.v.).
Galium sphagnophilum (Greenm.) Dempster Galium wigginsii Dempster, Brittonia 22: 188.
1970. Type: MEXICO: Baja California,
Phytologia 26: 222. 1973. Relbunium meadow at La Encantada, Sierra San Pedro
mazocarpum Greenm., Proc. Amer. Acad. Martir, 2200 m, 18 Sep 1930,1. L. Wiggins &
Arts 41: 250. 1905. Type: MEXICO: Hidalgo, D. Demaree 4879 (Holotype UC, n.v.;
moist banks, barranca below Trinidad Iron Isotypes DS, n.v., F (photo PTBG), GH, n.v.,
Works, 5500 ft., 6 May 1904, C G. Pringle NY, n.v., US (photos MEXU, PTBG)).
8834 (Holotype GH, n.v.; Isotypes MEXU, Galiium wrightii A. Gray, PI. Wright. 1: 80.
UC, n.v., US (photos MEXU, PTBG)). 1852. Type: U.S.A.: Texas, Jeff Davis Co.,
Galium stellatum Kellogg, Proc. Calif. Acad. pass of the Limpia, Aug 1849, C. Wright 235
2: 97, fig. 26. 1861. [1863]. Type: MEXICO: (Holotype GH, n.v.; Isotype NY, n.v.).
Baja California, Cedros Island, Veatch s.n. Galium wrightii A. Gray var. latifolium B. L.
(Holotype CAS, n.v.). Rob. & Femald, Proc. Amer. Acad. Arts 30:
Galium suffruticosum Nutt. in Torr. & A. Gray, 116. 1894. Type: MEXICO: Sonora, Sierra de
Fl. N. Amer. 2: 21. 1841. Horn. Illeg. Not las Gronillas, 5800 ft., 2 Oct 1890, C. V.
Galium suffruticosum Hook. & Arn., 1833. Hartman 58 (Holotype GH, n.v.; Isotypes
Type: U.S.A.: California, San Diego, Nuttall BM, n.v., F, n.v., NY, n.v., UC, n.v., US
s.n. (Holotype GH, n.v.).—Galium nuttallii (photos MEXU, PTBG)).—Galium wrightii
A. Gray subsp. nuttallii. A. Gray
Galium trifidum L., Sp. PI. 1: 105. 1753. Type: Galium wrightii A. Gray var. rothrockii (A.
CANADA: Kalm (presumably LINN, n.v.). Gray) Ehrend., Contr. Dudley Herb. 5: 99.
1958. Galium rothrockii A. Gray, Proc. Amer.
 Acad. Arts 17: 203. 1882. Type: U.S.A.: uncertain, probably Randia sp. (protologue
Arizona, Camp Crittenden, Rothrock 675 states twigs have 4 spines).
(Holotype GH, n.v.; Isotype NY, n.v.). Gardenia schiedeana Schltdl., Linnaea 9: 601.
—Galium wrightii A. Gray 1834. Type: MEXICO: Veracruz, prope
Actopan, Schiede s.n. (presumably HAL,
Garapatica H. Karat, Linnaea 30: 143. 1859. n.v.).—Status uncertain, possibly
Type: Garapatica edulis H. Karst. synonymous with Randia monantha Benth.
Garapatica edulis (L. C. Rich.) H. Karst., Fl. (1841); perhaps the oldest available epithet for
Columb. 1: 57, /. 28. 1860. Genipa edulis this species.
L. C. Rich., Actes Soc. Hist. Nat. Paris 1: 107.
1792. Type: FRENCH GUIANA: Guyana, Genipa L., Syst. Nat., ed. 10, 2: 931. 1759.
Richard s.n. (Holotype P, n.v.; Isotype F Type: Genipa americana L.
971706 (photo PTBG)).—Alibertia edulis (L. Genipa americana L., Syst. Nat., ed. 10, 2:
C.Rich.) A. Rich. 931. 1759. Lectotype: BRAZIL: Janipaba Piso
& Marcgraf, Hist. Nat. Bras. 52, Ic. 1648.
Gardenia J. Ellis, Phil. Trans. 51: 935, t. 23. (Lectotype designated by Howard, 1989.)
1761 (nom. cons.). Type: Gardenia Genipa americana L. var. caruto (Kunth) K.
jasminoides J. Ellis Schum. in Mart., Fl. Bras. 6(6): 352. 1889.
Gardenia armata Sw., Prodr. Fl. Veg. Ind. Occ. Genipa caruto Kunth in Humb., Bonpl. &
51. 1788. Type: COLOMBIA: Cartagena, Kunth, Nov. Gen. Sp. 3: 407. 1820. Type: In
Jacquin s.n. (Lectotype: Swartz's plate; riparia Orinoci et Fluminis Nigri, prope
designated by Taylor & Lorence, 1993). Caracas [VENEZUELA] et Carthagena
—Randia armata (Sw.) DC. [COLOMBIA], Humboldt & Bonpland s.n.
Gardenia augusta (L.) Merr., Interpret. (Holotype P-Bonpland, n.v.). For typification
Rumph. Herb. Amboin. 485. 1917. Vameria see Steyermark, 1972: 349.—Genipa
augusta L., Amoen. Acad. 4: 136. 1759. americana L.
Type: Not cited; the description and Genipa caruto Kunth in Humb., Bonpl. &
illustration may be taken as the type according Kunth, Nov. Gen. Sp. 3: 407. 1820. Type: In
to Smith (1974). Frequently cultivated in riparia Orinoci et Fluminis Nigri, prope
Mexico and Mesoamerica. Caracas [VENEZUELA] et Carthagena
( hirdenia edulis (Rich.) Poir. in Lam., Encycl. [COLOMBIA], Humboldt & Bonpland s.n.
Meth. Bot. Suppl. 2: 708. 1812. Genipa edulis (Holotype P-Bonpland, n.v.). For typification
Rich., Act. Soc. Hist. Nat. Pans 1: 107. see Steyermark, 1972: 349.—Genipa
17792. Type: FRENCH GUIANA: Guyane, americana L.
without precise locality, Richard s.n. Genipa cinerea Fernald, Proc. Amer. Acad. Arts
(Holotype P, n.v.; Isotype F 971706 (photo 33: 93. 1897. Type: MEXICO: Guerrero,
PTBG)).—Alibertia edulis (Rich.) A. Rich. Acapulco and vicinity, Oct 1894-Mar 1895,
Gardenia laccifera Sesse & Moc., Fl. Mexic, E. Palmer 348 (Holotype GH, n.v.; Isotype
ed. 2, 54. 1894. Type: MEXICO: Habitat in US (photos MEXU, PTBG)).—Randia
Ayecapixtla haud procul ab Eclesia. Floret cinerea (Fernald) Standi.
Aprili, Sesse & Mociiio s.n. (presumably MA, Genipa codonocalyx Standi., Contr. U.S. Natl.
n.v.).—Status uncertain, probably Randia Herb. 17: 446. 1914. Type: COSTA RICA:
sp. (protologue states twigs have paired Boca Matapalo, cordon littoral du Pacifique,
10 Apr 1898, H. Pittier 12085 (Holotype US
Gardenia mexicana Sesse & Moc., Fl. Mexic, 577536 (photos MEXU, PTBG); Isotype US).
ed. 2, 54. 1894. Type: MEXICO: Habitat in —Genipa americana L.
Ayacapixtla. Floret Aprili, Sesse & Mocino Genipa curviflora Dwyer, Ann. Missouri Bot.
s.n. (presumably MA, n.v.).—Status Gard. 64: 184. 1980. Type: PANAMA:
 Panama, along El Llano-Carti Road, 8 km
north of Pan American Highway at El Llano,
450 m, 5 Mar 1974, M. Nee & R. Warmbrodt
10347 (Holotype MO 2579400 (photo Geocardia Standi., Contr. U.S. Natl. Herb. 17:
PTBG); Isotype MO (photo PTBG)). 444. 1914. Published as a substitute name for
Genipa echinocarpa (Moc. & Sesse ex DC.) A. Geophila D. Don, 1825 (nom. cons.), non
Gray, Proc. Amer. Acad. Arts 21: 380. 1886. Bergeret, 1803 (nom. rej.). [Herein treated as
Randia echinocarpa Moc. & Sesse ex DC., a synonym of Geophila D. Don.]
Prodr. 4: 385. 1830. Type: MEXICO: Fl. Geocardia pleuropoda (Donn. Sm.) Standi.,
Mexic. Ic. t. 469 (caique G-DC, n.v.; copies Contr. U.S. Natl. Herb. 17: 445. 1914.
MEXU, US). Basanacantha.—Randia Geophila pleuropoda Donn. Sm., Bot. Gaz.
echinocarpa Moc. & Sesse ex DC. (Crawfordsville) 52: 50. 1911. Type:
Genipa edulis Rich., Act. Soc. Hist. Nat. Paris 1: GUATEMALA: Alta Verapaz, Secanquim
107. 1792. Type: FRENCH GUYANA: trail to Sepaquite, 4000 ft., 5 Feb 1904, G. P.
Guyane, without precise locality, Richard s.n. Golls.n. (Holotype US 860647 (photos
(Holotype presumably P, n.v.; Isotype F MEXU, PTBG)).—Coccocypselum
971706 (photo PTBG)).—Alibertia edulis i Nees & Mart.
(Rich.) A. Rich, ex DC.
Genipa maxonii Standi., J. Wash. Acad. Sci. 18: Geophila D. Don, Prodr. Fl. Nepal. 136. 1825.
642. 1918. Type: PANAMA: Panama, forest Nom. cons. Not Geophila Bergeret, 1803.
along the Rio Indio de Gatun, Canal Zone, Type: Geophila reniformis D. Don, nom. illeg.
near sea level, 17 Feb 1911, W. R. Maxon (= Geophila herbacea (Jacq.) K. Schum.).
4848 (Holotype US 675223 (photos MEXU, Geophila cordifolia Miq., Stirp. Sunn. Sel. 176.
PTBG)). Sickingia.—Simira maxonii 1850. Type: SURINAM: Without precise
(Standi.) Steyerm. locality, Kappler 1896 (location unknown).
Genipa venosa Standi, J. Wash. Acad. Sci. 18: Geophila croatii Steyerm., Phytologia 35: 401.
168. 1928. Type: COSTA RICA: Guanacaste, 1977. Type: PANAMA: Canal Zone, Barro
El Arenal, wet forest, 485-600 m, 18-19 Jan Colorado Island, Zetek Trail, 3 Oct 1968,
1926, P. C. Standley & J. Valerio 45269 T. B. Croat 6647 (Holotype MO 1969871
(Holotype US 1254013 (photos MEXU, (photo PTBG)).—Geophila gracilis (Ruiz &
PTBG); Isotype US).—Genipa americana L. Pav.) DC.
Genipa vulcanicola Standi., Field Mus. Nat. Geophila gracilis (Ruiz & Pav.) DC, Prodr. 4:
Hist., Bot. Ser. 17: 213. 1937. Type: 537. 1830. Psychotria gracilis Ruiz & Pav.,
GUATEMALA: Suchitepequez, Volcan de Fl. Peruv. 2: 63, t. 211, fig- C. 1799. Type:
Atitlan, 7700 ft., 24 Oct 1934, A. F. Skutch PERU: Huanuco, Pillao et Pueblo Nuevo,
1512 (Holotype F 813461 (photos MEXU, Ruiz Lopez & Pavon s.n. (Presumably MA,
PTBG); Isotypes A, n.v., US (photos MEXU, n.v.).
PTBG)).—Glossostipula concinna (Standi.) Geophila herbacea (Jacq.) K. Schum. in Engl.
& Prantl, Nat. Pflanzenfam. 4, 4: 119. 1891.
Genipa williamsii Standi., J. Wash. Acad. Sci. Psychotria herbacea Jacq., Enum. Pi. Carib.
8: 642. 1918. Type: PANAMA: Cana and 16. 1760. Type: MARTINIQUE: Without
vicinity, 17 Apr-8 Jun 1908, R. S. Williams precise locality, Jacquin (location unknown).
814 (Holotype US 678301 (photos MEXU, —Geophila repens (L.) I. M. Johnst.
PTBG)). Geophila macropoda (Ruiz & Pav.) DC,
Prodr. 4: 537. 1830. Psychotria macropoda
Gentiana L. [Gem Ruiz & Pav., Fl. Peruv. 2: 63, pi. 211, fig. b.
Gentiana nigricans Lam., Encycl. 2: 645. 1788. 1799. Type: PERU: Huanuco, Nemonbus
Type: MEXICO: Collector and locality secus Pillao ad Iscutunam, Ruiz Lopez &
 Pavon s.n. (Presumably MA, n.v.). Nayarit, foothills of Sierra Madre, Tepic,
Coccocypselum. between Aguacate & Dolores, 6 Aug 1897,
Geophila pleuropoda Donn. Sm., Bot. Gaz. J. N. Rose 2028 (Holotype US 300921 (photos
(Crawfordsville) 52: 50. 1911. Type: MEXU, PTBG); Isotype F (photos MEXU,
GUATEMALA: Alta Verapaz, Secanquim, PTBG)).
trail to Sepacuite, 4000 ft., 5 Feb 1905, G. P Glossostipula concinna (Standi.) Lorence,
Golls.n. (Holotype US 860647 (photos Candollea 41: 458. 1986. Randia concinna
MEXU, PTBG)). Coccocypselum, Geocardia. Standi., Publ. Field Columbian Mus., Bot. Ser.
—Coccocypselum cordifolium Nees & Mart. 4: 288. 1929. Type: MEXICO: Jalisco, San
Geophila repens (L.) I. M. Johnst., Sargentia 8: Sebastian, Hacienda del Ototal, Sierra Madre
281. 1949. Rondeletia repens L., Syst. Nat., Occidental, 1500 m, 15 Feb 1937, Y. Mexia
ed. 10, 2: 928. 1759. Type: JAMAICA: 1690 (Holotype F 579911 (photos MEXU,
Shane s.n. (Lectotype: Sloane Herb, iv.iii, PTBG); Isotypes MO (photos MEXU, PTBG),
n.v.; designated by Howard, 1989). US (photos MO, US)).
Geophila trichogyne (Muell. Arg.) Standi., Publ. Glossostipula strigosa Lorence, Novon 7: 46,
Field Columbian Mus., Bot. Ser. 7: 423. 1931. fig. 1. 1997. Type: HONDURAS: Dept.
Mapouria trichogyne Muell. Arg. in Mart., Fl. Atlantida, base of N slope of Pico Bonito, E of
Bras. 6: 426. 1881. Type: BRAZIL: Habitat in new CURLA (Centre Universitario Regional
prov. do Alto Amazonas, prope Manaos ad del Litoral Atlantico) camp building on
Rio Negro, Spruce s.n. (B, presumably Quebrada Grande, ca. 1/3 km above its
destroyed).—Geophila cordifolia Miq. confluence with the Rio Bonito, ca. 10 km
SW of La Ceiba, Parque Nacional Pico
Gerontogea Cham. & Schltdl., Linnaea 4: 1154. Bonito, 15°42'N, 86°51W, 140 m, 8 May
1829. Type: Not designated. [Herein treated 1993, R. Evans 1559 (Holotype MO
as a synonym of Hedyotis L.] 04585952 (photo PTBG); Isotype PTBG
Gerontogea deppeana Cham. & Schltdl., 16114).
Linnaea 5: 169. 1830. Type: MEXICO:
Veracruz, ad San Andres, 1829, Schiede & Gomozia Mutis ex L. f, Suppl. Pi. 17, 129.
Deppe 391 (Holotype presumably HAL, n.v.; 1781. Norn. rej. vs. Nertera J. Gaertn., 1788.
Isotype MO (photos MEXU, PTBG)). Type: Gomozia granadensis L. f. [Herein
Hedyotis.—Hedyotis microtheca (Schltdl. & treated as a synonym of Nertera Banks & Sol.
Cham.) Steud. ex C. F. Gaertn. according to the
Gerontogea microtheca Cham. & Schltdl., specifications of the rejection.]
Linnaea 5: 169. 1830. Type: MEXICO: Gomozia granadensis L. f, Suppl. PI. 129.
Veracruz, Barranca de Tioselos, Schiede & 1781. Type: America meridionali, Mutis s.n.
Deppe 390 (Holotype presumably HAL, n.v.; (LINN 172.1, n.v.)—Nertera granadensis
Isotype MO (photos MEXU, PTBG) [isotype (L. f.) Druce
label states in sylvis Papantla]). Oldenlandia.
—Hedyotis microtheca (Schltdl. & Cham.) Gonzalagunia Ruiz & Pav., Fl. Peruv. Prodr. 12,
Steud. t. 3. 1794. Type: Gonzalagunia dependens
Ruiz & Pav.
Glossostipula Lorence, Candollea 41: 454. Gonzalagunia bracteosa (Donn. Sm.) B. L.
1986. Type: Glossostipula concinna (Standi.) Rob., Proc. Amer. Acad. Sci. 45: 405. 1910.
Gonzalea bracteosa Donn. Sm., Bot. Gaz.
Glossostipula blepharophylla (Standi.) (Crawfordsville) 33: 252. 1902. Type:
Lorence, Candollea 41: 457. 1986. Randia COSTA RICA: Suerre, Llanuras de Santa
blepharophylla Standi., Contr. U.S. Natl. Clara, 300 m, Apr 1896, J. D. Smith 6583
Herb. 23: 1375. 1926. Type: MEXICO: (Holotype US 943468 (photos MEXU,
 <m Mi AH AN AND CENTRA

PTBG); Isotypes F, n.v., NY, n.v., US 943469 Bartl. ex DC, Prodr. 4: 436. 1830. Type:
(photos MEXU, PTBG)). Duggena. MEXICO: Without precise locality, Haenke
Gonzalagunia brenesii Standi., Publ. Field s.n. (Holotype presumably PR, n.v., fragment
Mus. Nat. Hist., Bot. Ser. 18: 1302. 1938. G-DC, n.v. (microfiche 4: 436.3). Duggena.
Type: COSTA RICA: Near San Ramon, —Rondeletia buddleioides Benth.
Bosque a orillas del mar de Golfito de Osa, 12 Gonzalagunia ovatifolia (Donn. Sm.) B. L.
Apr 1930, A. Brenes 12323 (Holotype F Rob., Proc. Amer. Acad. Arts 45: 405. 1910.
856730 (photo PTBG); Isotypes GH, n.v., US Gonzalea ovatifolia Dorm. Sm., Bot. Gaz.
(photos MEXU, PTBG)). (Crawfordsville) 27: 336. 1899. Type:
Gonzalagunia chiapasensis (Standi.) Standi. & COSTA RICA: Cartago, Atirro, 600 m, J. D.
Steyerm., Publ. Field Mus. Nat. Hist., Bot. Smith 6599 (Lectotype US 943467, designated
Ser. 22: 283. 1940. Rondeletia chiapasensis here under main entry (photos MEXU,
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. PTBG); Isolectotype F). Syntypes: Costa
22: 54. 1940. Type: MEXICO: Chiapas, Rica, J. D. Smith 6600 (F, n.v., GH, US);
Cascarada, 31 Dec 1936, E. Matuda 1064 Costa Rica, Pittier & Tonduz 9171 (US, n.v.);
(Holotype MICH, n.v.; Isotypes F (fragment), i, Hart 113 (US,:
TEX-LL (photo PTBG)). Rogiera.
Gonzalagunia exaltata Standi., Publ. Field Mus. Mart., Fl. Bras. 6(6): 292. 1889. Buena
Nat. Hist., Bot. Ser. 22: 385. 1940. Type: panamensis Cav., An. Hist. Nat. 2: 279. 1800.
COSTA RICA: San Jose, basin of El General, Type: PANAMA: Lancon, Nee s.n. (Holotype
beside river, 675-900 m, Mar 1940, A. F. presumably MA, n.v.; possible Isotype F
Skutch 4747 (Holotype F 1044350 (photo (photo PTBG)).
PTBG); Isotypes A, n.v., MO (photos MEXU, >ra Cham. & Schltdl.,
PTBG), US (photos MEXU, PTBG)). Linnaea 6: 414. 1831. Type: MEXICO:
—Rondeletia brenesii Standi. Veracruz, Cuesta Grande de Jalacingo,
Gonzalagunia glabra S. Watson, Proc. Amer. Schiede & Deppe 1275 (Holotype presumably
Acad. Arts 25: 152. 1890. Type: MEXICO: HAL, n.v.; Isotype B, presumably destroyed
Jalisco, in the mountains near Lake Chapala, (photo MO)).—Rondeletia septicidalis B. L.
Dec 1889, C. G. Pringle 2442 (Holotype A, Rob., vel sp. aff.
n.v.; Isotype US (photo PTBG)). Gonzalagunia rojasii Standi., Publ. Field Mus.
—Citharexylum glabrum (S. Watson) Nat. Hist., Bot. Ser. 22: 114. 1940. Type:
Greenm. (Verbenaceae) GUATEMALA: Quezaltenango, El Muro,
Gonzalagunia kallunkiae Dwyer, Ann. below Santa Maria de Jesus, ca. 1200 m, 1
Missouri Bot. Gard. 67: 192. 1980 [as Mar 1938, P. C. Standley 67145 (Holotype F
kallunkii]. Type: PANAMA: Veraguas, NW 995085 (photo PTBG)).
of Santa Fe, 11 km Escuela Agricola Alto de Gonzalagunia rosea Standi., Ann. Missouri
Piedra, on road to Calovebora, near Rio Bot. Gard. 25: 836. 1938. Type: PANAMA:
Coloveborita, 15 Nov 1975, S. Mori & J. Chinqui, valley of the upper Rio Chiriqui
Kallunki 3141 (Holotype MO 2285895 (photo Viejo, 1300-1900 m, Jul-Aug 1937, G. White
PTBG)). & P. White 7 (Holotype F 913064 (photo
Gonzalagunia longithyrsa Fosberg, Sida 2: 387. PTBG); Isotypes A, n.v., MO (photo PTBG).
1966. Type: COSTA RICA: Cartago, Tapanti, NY, n.v.).
at edge of wet forest, 1200 m, 1 Nov 1964, Gonzalagunia rudis (Standi.) Standi., J. Wash.
W. H. Hatheway & R. L. Rodriguez 1241 Acad. Sci. 17: 170. 1927, comb, inval.
(Holotype US 2562636 (photos MEXU, Duggena rudis Standi., Contr. U.S. Natl.
PTBG)).—Gonzalagunia rosea Standi. Herb. 18: 125. 1916. Type: PANAMA:
Gonzalagunia nivea (Bartl. ex DC.) Kuntze, Panama, shady places along Chagras at El
Rev. Gen. PI. 1: 284. 1891. Gonzalea nivea
 Vigia, 12 Jan 1911, //. Pittier 2378 (Holotype Duggena.—Gonzalagunia bracteosa (Donn.
US 676602 (photos MEXU, PTBG)). Sm.) B. L. Rob.
Gonzalagunia stenostachya (Standi.) W. C. Gonzalea hayesii Wernham, J. Bot. 51:219.
Burger in W. C. Burger & C. M. Taylor, 1913. Type: PANAMA: In woods near
Fieldiana, Bot. 33: 150. 1993. Rondeletia Panama, Hayes s.n. (Holotype BM, n.v.).
stenostachya Standi., Publ. Field Mus. Nat. Duggena.—Gonzalagunia panamensis
Hist., Bot.Ser. 18: 1372. 1938. Type: COSTA (Cav.) K. Schum.
RICA: La Hondura de San Jose, 1300 m, 15 Gonzalea nivea Bartl. ex DC, Prodr. 4: 436.
Aug 1933, M. Valerio 769 (Holotype F 1830. Type: MEXICO: Without precise
681816 (photos MEXU, PTBG)). locality, Haenke s.n. (Holotype presumably
Gonzalagunia tacanensis Lundell, Lloydia 2: PR, n.v., fragment G-DC, n.v. (microfiche 4:
106. 1939. Type: MEXICO: Chiapas, Mt. 436.3)). Duggena.—Rondeletia buddleioides
Tacana, 1000-2000 m, Aug 1938, E. Matuda Benth.
2462 (Holotype TEX-LL (photos MEXU, Gonzalea ovatifolia Donn. Sm., Bot. Gaz.
PTBG)).—Gonzalagunia thyrsoidea (Donn. (Crawfordsville) 27: 336. 1899. Type:
COSTA RICA: Cartago, Atirro, 600 m, J. D.
Gonzalagunia thyrsoidea (Donn. Sin.) B. L. Smith 6599 (Lectotype US 943467, here
Rob., Proc. Amer. Acad. Arts 45: 405. 1910. designated (photos MEXU, PTBG);
Gonzalea thyrsoidea Donn. Sm., Bot. Gaz. Isolectotype F). Syntypes: COSTA RICA:
(Crawfordsville) 13: 188. 1888. Type: J. D. Smith 6600 (F, n.v., GH, US); Pittier &
GUATEMALA: Aha Verapaz, Berge bei Tonduz 9171 (US, n.v.). PANAMA; Hart 113
Pansamala, 3800 ft., May 1887, H. von (US, n.v.). Duggena.—Gonzalagunia
Tuerckheim 1249 (Holotype US 941409 ovatifolia (Donn. Sm.) B. L. Rob.
(photos MEXU, PTBG); Isotype GH, n.v.). Gonzalea panamensis (Cav.) Spreng., Syst. Veg.
Gonzalagunia tonduzii K. Krause, Bot. Jahrb. 1: 417. 1825. Buena panamensis Cav., Anal.
54, Beibl. 119: 42. 1916. Type: COSTA Hist. Nat. (Madrid) 2: 279. 1800. Type:
RICA: Cerro del Roble, ca. 2700-2900 m, PANAMA: Lancon, Nee s.n. (Holotype
Apr 1898, A. Tonduz 11813 (Holotype B, presumably MA, n.v.; possible Isotype F
presumably destroyed (photos F, MEXU, (photo MO)). Duggena.—Gonzalagunia
MO)).—Rondeletia buddleioides Benth. panamensis (Cav.) K. Schum.
Gonzalagunia veraguensis Dwyer, Ann. Gonzalea parviflora G. Don., Gen. Hist. 3: 537.
Missouri Bot. Gard. 67: 197. 1980. Type: 1834. Type: MEXICO: Oaxaca, 3000 ft., H
PANAMA: Veraguas, NW of Santa Fe, 8.8 Galeotti 7245 (Holotype presumably BM,
km from Escuela Agricola Alto de Piedra, on n.v.; Isotype G-DC, n.v. (photo MO)).
road to Calovebora, 1900 ft., 17 Jun 1975, 5. —Rondeletia capitellata Hemsl., vel. sp. aff;
Mori 6673 (Holotype MO 2353238 (photos perhaps the earliest available epithet for this
PTBG); Isotype MO 2353239 (photo PTBG)).
Gonzalea secunda M. Martens & Galeotti, Bull.
Gonzalea Pers., Syn. PI. 1: 132. 1805. Norn, Acad. Roy. Sci. Bruxelles 11: 235. 1844.
superfl. for Gonzalagunia Ruiz & Pav. 1794 Type: MEXICO: Veracruz, bois humides de
(for which it is herein treated as a synonym). Xalapa, a 4000 pieds, Juin, H. Galeotti 7077
Gonzalea bracteosa Donn. Sm., Bot. Gaz. (Holotype presumably BR, n.v.).—Status
(Crawfordsville) 33: 252. 1902. Type: uncertain.
COSTA RICA: Suerre, Llanuras de Santa Gonzalea thyrsoidea Donn. Sm., Bot. Gaz.
Clara, J. D. Smith 6583 (Holotype US 943468 (Crawfordsville) 13: 188. 1888. Type:
(photos MEXU, PTBG); Isotypes F, n.v., NY, GUATEMALA: Alta Verapaz, Berge bei
n.v., US 943469 (photos MEXU, PTBG)). Pansamala, 3800 ft., May 1887, H. von
Tuerckheim 1249 (Holotype US 941409
A NOMENCLATOR OF MEXICAN A - AMERICAN RUBIACEA

(photos MEXU, PTBG); Isotype GH, n.v.). Guettarda crispiflora Vahl. Eclog. Amer. 1:
Duggena.—Gonzalagunia thyrsoidea (Donn. 36, /. 6. 1796. Type: MONTSERRAT: Versus
Sm.) B. L. Rob. summitates in insula Montserrat, Ryan s.n
(Holotype presumably C, n.v.).
Guettarda L., Sp. PI. 2: 991. 1753. Type: Guettarda dealbata M. Martens & Galeotti,
Guettarda speciosa L. Bull. Acad. Roy. Soc. Bruxelles 11: 232.
Guettarda armata Bartl. ex DC, Prodr. 4: 457. 1844. Type: MEXICO: Oaxaca, dans les
1830. Type: MEXICO: Without precise forets epaisses et humides de Llano Verde et
locality, Haenke s.n. (Holotype presumably de Tanetze, dans le Rincon (cordillere
PR, n.v. (fragment G-DC, n.v. (microfiche 4: orientale d'Oaxaca), 4000-6000 ft., H.
457.19)).—Status uncertain, probably . Galeotti 7131 (Holotype presumably BR,
Chomelia spinosa Jacq. n.v.).—Status uncertain, "scarcely of this
Guettarda brenesii Standi., Publ. Field Mus. genus" according to Standley (1934).
Nat. Hist., Bot. Ser. 18: 1303. 1938. Type: Guettarda deamii Standi., Contr. U.S. Natl.
COSTA RICA: Golfo de Nicoya, Isla de Herb. 20: 209. 1919. Type: GUATEMALA:
Caballo, Jun 1932, A. Brenes 15694 Zacapa, Gualan, 620 ft., 15 Jun 1909, C. C.
(Holotype F 857985 (photo PTBG)). Deam 6271 (Holotype US 796136 (photo
Guettarda chiriquiensis Standi., Ann. Missouri PTBG); Isotype GH, n.v.).
Bot. Gard., 25: 838. 1938. Type: PANAMA: Guettarda dichotoma M. Martens & Galeotti,
Chiriqui, Valley of upper Rio Chiriqui Viejo, Bull. Acad. Roy. Sci. Bruxelles 11: 233. 1844.
1300-1900 m, 27 Jul 1937, P. White & G. Type: MEXICO: Veracruz, savanes de
White 22 (Holotype MO 1177311 (photo Consoquitla, pres la colonie de Zacuapan et
PTBG): Isotypes A, n.v., F).—Guettarda Mirador, 1500-2000 pieds, 1840,//. Galeotti
crispiflora Vahl 2601 (Holotype presumably BR, n.v.; Isotype
Guettarda cobanensis Donn. Sm., Bot. Gaz. P (photos MEXU, PTBG)).—Guettarda
(Crawfordsville) 47. 255. 1909. Type: elliptica Sw.
GUATEMALA: Alta Verapaz, near Coban, Guettarda elliptica Sw., Prodr. Veg. Ind. Occ.
1500 m, H. von Tuerckheim 11.2096 (Holotype 59. 1788. Type: JAMAICA: Campestris
US 941415 (photo PTBG); Isotype US (photo siccis, Swartzs.n. (presumably S, n.v.).
PTBG)). Guettarda filipes Standi., Contr. U.S. Natl.
Guettarda colubrinoides Standi., Publ. Field Herb. 20: 210. 1919. Type: MEXICO:
Columbian Mus., Bot. Ser. 8: 358. 1931. Durango, near Huasemote, 15 Aug 1897,7. N.
Type: MEXICO: Tabasco, Paso del Carrizal, 4 Rose 3498 (Holotype US 302475).
May 1890, J. N. Rovirosa 787 (Holotype K, Guettarda foliacea Standi., Contr. U.S. Natl.
n.v., fragment F)).—Status uncertain, Herb. 18: 139. 1916. Type: PANAMA: Canal
probably Guettarda elliptica Sw. Zone, along the Trinidad River, near sea level,
Guettarda combsii Urb., Symb. Antill. 6: 48. 19-21 Jul 1911, //. Pittier 4031 (Holotype US
1909. Type: CUBA: Distr. Cienfuegos, Prov. 679116; Isotype F).
Santa Clara, prope Cieneguito, 2 Jul 1895, R. Guettarda galeotti Standi., Contr. U.S. Natl.
Combs 270 (Syntype B, destroyed; Lectotype Herb. 23: 1384. 1926. Type: MEXICO:
F 358073 (photo PTBG), here designated; Oaxaca, Prov. Pinotepa, H. Galeotti 2576
Isolectotypes F, US). Syntype: Prov. Pinar del (Holotype US 572992, fragment F (photos
Rio inter Candelaria et Artemisa, Sept., MEXU, PTBG)).—Chomelia protracta
Wilson 1783 (B, destroyed). (Bartl. ex DC.) Standi.
Guettarda conferta Benth., Bot. Voy. Sulphur Guettarda gaumeri Standi., Publ. Field
106. 1845. Type: COSTA RICA: Cocos Columbian Mus., Bot. Ser. 8: 58. 1930. Type:
Island, Barclay s.n. (Holotype BM, n.v.). MEXICO: Yucatan, without precise locality,
1917-1921, G. F. Gaumer 24239 (Holotype F
 552244 (photos MEXU, PTBG); Isotypes A, Guettarda protracta Bartl. ex DC. var. brevipes
n.v., GH, n.v., MO (photos MEXU, PTBG)). DC, Prodr. 4: 457. 1830. Type: MEXICO:
(As for other taxa described by Standley in Without precise locality, Haenke s.n.
this publication, the first specimen he cited in (Holotype presumably PR, n.v.). Anisomeris,
the protologue ("Gaumer 24239, Herb. Field Pittonioitis — Chomelia protracta (Bartl. ex
Mus. No. 552,244") was intended to be the DC.) Standi.
holotype, rather than the second one (Lundell Guettarda protracta Bartl. ex DC. var. longipes
352). Consequently lectotypification is DC, Prodr. 4: 457. 1830. Type: MEXICO:
Guerrero, ad Acapulco, Haenke s.n. (Holotype
(iuanmlu insulari.s Brandegee, Univ. Calif. presumably PR, n.v.).—Chomelia protracta
Publ. Bot. 10: 416. 1924. Type: MEXICO: (Bartl. ex DC.) Standi.
Baja California, Socorro Island, Mar-Jun Guettarda ramuliflora Beurl, Kongl. Vetensk.
1897, A. W. Anthony 377 & F. E. Barkelmew Acad. Handl. 132. 1854. Type: PANAMA:
179 (Holotype UC 102472, n.v.; Isotypes F, Colon, near Porto Bello, BUlberg 642
GH, n.v., MO (photos MEXU, PTBG)). (Holotype S, n.v.).
—Guettarda elliptica Sw. Guettarda sageretioides A. R. Molina, Ceiba 3:
Guettarda macrosperma Donn. Sm., Bot. Gaz. 96. 1952. Type: HONDURAS: El Paraiso,
(Crawfordsville) 18: 204. 1893. Type: matorrales hiimedos y rocosos del Rio Lizapa,
GUATEMALA: Santa Rosa, 3000 ft., May 1000 m, 19 Sep 1951, A Molina R. 4125
1892, Heyde & Lux 3160 (Holotype US (Holotype US 2216040 (photo PTBG);
48602 (photo PTBG); Isotype US (photo Isotypes F (photo PTBG), GH, n.v.).
PTBG)). Guettarda sanblasensis Dwyer, Ann. Missouri
Guettarda odorata (Jacq.) Lam., Tabl. Encycl. Bot. Gard. 67: 204. 1980. Type: PANAMA:
Meth. 2: 219. 1819. Laugieria odorata Jacq., San Bias, headwaters of Rio Cuadi, Camp
Enum. PI. Carib. 2: 16. 1760. Type: CUBA: Diablo (Drill site 22), 273 ft., 18 Dec 1967,
Ex Carthagena et Havana, Jacquin s.n. J. A. Duke, A. G. Robyns & S. E. Verhoek
(Lectotype P, n.v.; designated by Steyermark, 3646 (Holotype MO 2183824 (photo PTBG)).
1972: 360). i abra Lam. var. seleriana Loes.,
Guettarda petenensis Lundell, Wrightia 4: 124. Fedde Rep. Sp. Nov. 18: 361. 1922. Type:
1969. Type: GUATEMALA: Peten, La MEXICO: Yucatan, Chiken Itza, im Walde, 2
Cumbre, in high forest, about 6 km from Jun 1914, C. Seler & E. Seler 5574 (Holotype
village, on top of hill bordering Rio Pusila, 18 B, presumably destroyed, fragment F;
May 1967, E. Contreras 6947 (Holotype Isotypes GH, n.v., US (photos MEXU,
TEX-LL (photos MEXU, PTBG); Isotype PTBG)).—Guettarda combsii Urb.
US). Guettarda seleriana (Loes.) Standi., Contr. U.S.
Guettarda poasana Standi., J. Wash. Acad. Sci. Natl. Herb. 23: 1384. 1926. Guettarda scabra
18: 182. 1928. Type: COSTA RICA: Alajuela, Lam. var. seleriana Loes., Fedde Rep. Sp.
Viento Fresco, slopes of Volcan de Poas, 1800 Nov. 18: 361. 1922. Type: MEXICO:
m, 13 Feb 1926, P. C. Standley & Torres Yucatan, Chiken Itza, im Walde, 2 Jun 1914,
47807 (Holotype US 1305283 (photo C. Seler & E. Seler 5574 (Holotype B,
PTBG)). presumably destroyed, fragment F; Isotypes
Guettarda protracta Bartl. ex DC, Prodr. 4: GH, n.v., US (photos MEXU, PTBG)).
457. 1830. Type: MEXICO: Without precise —Guettarda combsii Urb.
locality, Haenke s.n. (Holotype presumably Guettarda tetrandra Sesse & Moc., Fl. Mexic,
PR, n.v., fragment G-DC, n.v. (microfiche 4: ed. 2, 218. 1894. Type: MEXICO: Without
457.18)).—Chomelia protracta (Bartl. ex precise locality, Sesse & Mociiio s
DC.) Standi. (presumably MA, n.v.).—Status i
 L NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Guettarda tikalana Lundell, Wrightia 2: 63. GUATEMALA: Escuintla, Finca Monterrey,

1960. Type: GUATEMALA: Peten, Tikal, on S slope of Volcan de Fuego, 1140-1260 m, 5
Uaxactun trail, in center of escobal, 18 Jul Feb 1939, P. C. Standley 64569 (Holotype F
1959, C. L. Lundell 16519 (Holotype TEX-LL 982608 (photo PTBG); Isotypes A, n.v., NY,
(photos MEXU, PTBG); Isotypes F, US). n.v., US (photos MEXU, PTBG)).
Guettarda tournefortiopsis Standi., Publ. Field Hamelia calycosa Donn. Sm., Bot. Gaz.
Columbian Mus., Bot. Ser. 7: 293. 1931. (Crawfordsville) 12: 132. 1887. Type:
Based on Tournefortiopsis reticulata Rusby, GUATEMALA: Alta Verapaz, Pansamala,
Bull. New York Bot. Gard. 4: 369. 1907. Not 3600 ft., Jun 1885, H. von Tuerckheim 454
Guettarda reticulata Griseb., 1863. Type: (Holotype US 943617 (photos MEXU,
BOLIVIA: Without precise locality, Bang PTBG); Isotypes GH, n.v., K, n.v., NY, n.v..
2230 (Holotype US, n.v.). US (photos MEXU, PTBG)).
Guettarda turrialbana N. Zamora & Poveda, andegee, Univ. Calif.
Ann. Missouri Bot. Gard. 75: 1151, fig. 1. Publ. Bot. 6: 71. 1914. Type: MEXICO:
1988. Type: COSTA RICA: Cartago, Chiapas, Finca Mexiquito, Jul 1913, C. A.
Turrialba, CATIE, Bosque de Los Espabeles, Purpus 7030 (Holotype UC 173058, n.v.;
600 m, 26 Jun 1986, N. Zamora, M. M. Isotypes F (photos MEXU, PTBG), GH, n.v.,
Chavarria & H. Gomez 1263 (Holotype CR, MO (photos MEXU, PTBG), NY (n.v.), US
n.v.; Isotypes F (photo PTBG), K, n.v., MO (photos MEXU, PTBG)).—Hamelia calycosa
(photo PTBG), NY, n.v.). Donn. Sm.
Hamelia costaricensis Standi., Contr. U.S. Natl.
Habroneuron Standi., J. Wash. Acad. Sci. 17: Herb. 20: 207. 1919. Type: COSTA RICA:
338. 1927. Type: Habroneuron mexicanum Surulres [?] pres San Mateo, Aug 1890, P.
Standi. Biolley 2656 (Holotype US 764417 (photos
Habroneuron mexicanum Standi., J. Wash. MEXU, PTBG)).—Hamelia xerocarpa
Acad. Sci. 17: 339. 1927. Type: MEXICO: Kuntze var. costaricensis (Standi.) Elias
Oaxaca, Tintalcingo, F. Liebmann 11527 Hamelia corymbosa Sesse & Moc., La
(Holotype C, n.v.; Isotype C, n.v., fragment Naturaleza, ser. 2, 1: 37. 1888 (as
US (photos MEXU, PTBG)).—Habroneuron "Hamellia"). Type: MEXICO: Habitat in
radicans (Wernham) S. P. Darwin montibus Haereditatis-Ahuesuyo, Sesse &
Habroneuron radicans (Wernham) S. P. Mocino s.n. (presumably MA, n.v.).
Darwin, Brittonia 32: 343. 1980. Lindenia —Hamelia patens Jacq. var. patens
radicans Wernham, J. Bot. 52: 226. 1914. Hamelia declinata Sesse & Moc., Fl. Mexic,
Type: MEXICO: Oaxaca, shaded places on ed. 2, 60. 1894. Type: MEXICO: Sesse &
rocks at crossing of the river near Mocino s.n. (presumably MA, n.v.).
Tocotepeque, Jul 1839, T. Hartwegs.n. —Hamelia axillaris Sw.
(Holotype K, n.v. (photo MO)). Hamelia erecta Jacq., Enum. Syst. PI. Carib. 16.
1760. Type: COLOMBIA: Bolivar, vicinity of
Hamelia Jacq., Enum. Syst. PI. 2: 16. 1760. Cartagena, Jacquin s.n. (location unknown).
Type: Hamelia erecta Jacq. (Lectotype —Hamelia patens Jacq. var. patens
designated by Britton & Millsp., Bahama Fl. Hamelia hypomalaca B. L. Rob., Proc. Amer.
441. 1920). Acad. Arts 45: 406. 1910. Type: MEXICO:
Hamelia axillaris Sw., Prodr. Veg. Ind. Occ. Durango, without precise locality, Aug 1897,
46. 1788. Type: JAMAICA: Without precise J. N. Rose 2304 (Holotype US 301215 (photos
locality, Swartz s.n. (Lectotype S, n.v.; MEXU, PTBG); Isotypes GH, n.v., MO
designated by Elias, 1976). (photos MEXU, PTBG)).—Hamelia
Hamelia barbata Standi., Publ. Field Mus. Nat. xorullensis Kunth in Humb., Bonpl. & Kunth
Hist., Bot. Ser. 22: 115. 1940. Type:
 . Martens & Galeotti, Hamelia patens Jacq., Enum. Syst. PI. Carib.
Bull. Acad. Roy. Sci. Bruxelles 11: 233. 1844. 16. 1760. var. patens. Type: SANTO
Type: MEXICO: Veracruz, Mirador, Xalapa, DOMINGO (DOMINICAN REPUBLIC):
1841, H. Galeotti 2615 (Holotype BR, n.v.; Jacquin s.n. (location unknown). Dunhamelia.
Isotypes MO (photos MEXU, PTBG), P Hamelia patens Jacq. var. coronata Dorm. Sm.,
(photos MEXU, PTBG)).—Hamelia patens Bot. Gaz. (Crawfordsville) 40: 4. 1905. Type:
Jacq. var. patens GUATEMALA: Alta Verapaz, Cubliquitz,
Hamelia longipes Standi., Proc. Biol. Soc. 350 m, Aug 1903, H. von Tuerckheim 8532
Wash. 37: 53. 1924. Type: GUATAMALA: (Holotype US 943525 (photos MEXU,
Izabal, vicinity of Escoba, on the bay opposite PTBG); Isotypes F, GH, n.v., NY, n.v., US
Puerto Barrios, 150 m or less, 2 Jun 1922, (photos MEXU, PTBG)).—Hamelia
P. C Standley 24829 (Holotype US 1150447 rovirosae Wernh.
(photos MEXU, PTBG); Isotype GH, n.v.). Hamelia patens Jacq. var. glabra Oerst.,
Hamelia lutea Rohr ex Rees, Cyclop. 17. 1811. Vidensk. Meddel. Dansk Naturhist. Foren.
Type: Based on manuscript by von Rohr and Kjobenhavn 1852: 42. 1853. Type:
Jaquin's (Collectanea, Ic. Rar. tab 335) NICARAGUA: San Juan, Oersted s.n.
description and illustration of//, chrysantha. (Holotype C, n.v.).
—Hamelia axillaris Sw. Hamelia pauciflora Standi., Contr. U.S. Natl.
Hamelia macrantha Little, Carib. Forester 9: Herb. 18: 139. 1916. Type: PANAMA:
274. 1948. Type: ECUADOR: Chimborazo, Panama, forests on dry limestone around
El Oro, Spruce 6193 (Lectotype K, n.v. Alhajuela, Chagres Valley, 30-100 m, 12-15
(photos F, MO); designated by Elias, 1976). May 1911,//. Pittier 3469 (Holotype US
Hamelia magnifolia Wernham, J. Bot. 49: 210. 678513 (photos MEXU, PTBG)). Not H.
1911. Type: COSTA RICA: Puntarenas, bords pauciflora Willd., 1825.—Hamelia
du Rio Corozal, pres Santo Domingo de Golfo sanguinea Elias
Dulce, Mar 1896, H. Pittier 10091 Hamelia purpurascens S. F. Blake, Contr. Gray
(Lecototype BM, n.v.; Isolectotypes F, Herb. II, 52: 105. 1917. Type: BRITISH
fragment n.v., GH, n.v., P (photos MEXU, HONDURAS (BELIZE): On pine ridge near
PTBG), US (photos MEXU, PTBG); Manatee Lagoon, 6 Aug 1904, M. E. Peck 104
designated by Elias, 1976). (Holotype GH, n.v.).—Hamelia rovirosae
Hamelia magniloba Wernham, J. Bot. 49: 209. Wernham
1911. Type: NICARAGUA: Chontales, Tate Hamelia rostrata Bartl. ex DC, Prodr. 4: 442.
200 (Holotype K, n.v.).—Hamelia xerocarpa 1830. Type: MEXICO: Guerrero, Acapulco,
Kuntze var. xerocarpa Haenke s.n. (Holotype presumably PR (n.v.)).
Hamelia nodosa M. Martens & Galeotti, Bull. According to Elias (1976: 136), "This may be
Acad. Roy. Sci. Bruxelles 11: 234. 1844. the oldest valid name for H. versicolor, but
Type: MEXICO: Veracruz, pres Mirador, the description is too brief for accurate
3000 pieds, H. Galeotti 2581 (Holotype determination."—Status uncertain, probably
presumably BR, n.v.; Isotype P, n.v.). Hamelia versicolor A. Gray
—Hamelia patens Jacq. var. glabra Oerst. Hamelia rovirosae Wernham, J. Bot. 49: 211.
Hamelia panamensis Standi., Contr. U.S. Natl. 1911. Type: MEXICO: Tabasco, marshes on
Herb. 20: 208. 1919. Type: PANAMA: Above banks of Rio Sebastian, 11 Jun 1889, J. N.
Paca, 16 Apr 1908, R. S. Williams 744 Rovirosa 499 (Holotype K, n.v.; Isotypes NY,
(Holotype US 678258 (photos MEXU, n.v., US (photos MEXU, PTBG)).
PTBG); Isotype NY, n.v.).—Hamelia Hamelia rowleei Standi., J. Wash. Acad. Sci. 15:
xerocarpa Kuntze var. costaricensis (Standi.) 7. 1925. Type: COSTA RICA: Livingston on
Elias the Reventazon River, Jul-Aug 1920, W. W.
Rowlee & H. E. Stork 751 (Holotype US
 L NOMENCLATOR OF MEXICAN A ) CENTRAL AMERICAN RUBIACEAE

1080249 (photos MEXU, PTBG)).—Hamelia no type status. A syntype may be designated

xerocarpa Kuntze var. xerocarpa
Hamelia sanguinea Elias, Mem. New York Hamelia xerocarpa Kuntze var. costaricensis
Bot. Gard. 26: 116. 1976. Norn. nov. for (Standi.) Elias, Mem. New York Bot. Gard.
Hamelia pauciflora Standi., 1916. Not 26: 119. 1976. Hamelia costaricensis Standi.,
Hamelia pauciflora Willd. in M. Martens & Contr. U.S. Natl. Herb. 20: 207. 1919. Type:
Galeotti, 1844. Type: PANAMA: Panama, COSTA RICA: Alajuela, Surulres [?] pres San
forests on dry limestone around Alhajuela, Mateo, Aug 1890, P. Biolley 2656 (Holotype
Chagres Valley, 30-100 m, 12-15 May 1911, US 764417 (photos MEXU, PTBG)).
H. Pittier 3469 (Holotype US 678513 (photos Hamelia xorullensis Kunth in Humb., Bonpl. &
MEXU, PTBG); Isotype US (photos MEXU, Kunth, Nov. Gen. Sp. 3: 414. 1818; folio ed.
PTBG)). 323. 1818. Type: MEXICO: Michoacan, El
Hamelia storkii Standi., J. Wash. Acad. Sci. 15: Jorullo [=Xorullo], 1200 m, A. Bonpland s.n.
7. 1925. Type: PANAMA: Western Panama, (Holotype P-Bonpland., n.v.; Isotypes P, 2
without precise locality, 1 Aug 1923, H. E. sheets (photos MEXU, PTBG)).
Stork 38 (Holotype US 1166466 (photos
MEXU, PTBG)).—Hamelia xerocarpa Hedyotis L., Sp. PI. 1: 101. 1753. Type:
Kuntze var. costaricensis (Standi.) Elias Hedyotis fruticosa L. Nom. et typ. cons. prop,
Hamelia suaveolens Kunth in Humb., Bonpl. & (vide Jarvis in Taxon 41: 564. 1992). (As
Kunth, Nov. Gen. Sp. 3: 414. 1818. Type: herein treated, Hedyotis includes Carterella
COLOMBIA: (Garapata), Rio Magdalena, A. Terrell, Houstonia L., Oldenlandia L., and
Bonpland 1637 (Holotype P-Bonpland Oldenlandiopsis Terrell & W. H. Lewis.)
(photos MEXU, PTBG); Isotype P (photos Hedyotis acerosa A. Gray, PI. Wright. 1:81.
MEXU, PTBG)).—Hamelia patens Jacq. var. 1852. var. acerosa. Type: U.S.A.: Texas, high
glabra Oerst. prairies, from Live Oak Creek to the Las
Hamelia versicolor A. Gray, Proc. Amer. Acad. Moras, Jun 1849, C. Wright 237 (Holotype,
Arts 21: 416. 1886. Type: MEXICO: Jalisco, GH n.v.; Isotype US). Ereicoctis, Houstonia,
barranca near Guadalajara, Jun 1886, E. Mallostoma, Oldenlandia.
Palmer 125 (Holotype GH, n.v.; Isotypes MO Hedyotis acerosa A. Gray var. polypremoides
(photos MEXU, PTBG), P (photos MEXU, (A. Gray) W. H. Lewis, Ann. Missouri Bot.
PTBG), US (photos MEXU, PTBG)). Gard. 55: 31. 1968. Houstonia polypremoides
Hamelia viridifolia Wernham, J. Bot. 49: 213. A. Gray, Proc. Amer. Acad. Arts 21: 379.
1911. Type: COSTA RICA: Broussailles a 1886. Type: MEXICO: Chihuahua, Santa
Nicoya, Mar 1900, A. Tonduz 13867 Eulaha Mountains, 26 Sep 1885, C G.
(Holotype K, n.v.; Isotypes GH, n.v., P Pringle 356 (Lectotype GH, n.v.;
(photos MEXU, PTBG), US (photos MEXU, Isolectotypes MO (photos MEXU, PTBG),
PTBG))—Hamelia patens Jacq. var. glabra US (photos MEXU, PTBG); designated by
Oerst. Lewis, 1968).
Hamelia xerocarpa Kuntze, Rev. Gen. PI. 1: Hedyotis alexandrae (A. Carter) W. H. Lewis,
284. 1891. var. xerocarpa. Type: COSTA Ann. Missouri Bot. Gard. 55: 31. 1968.
RICA: Guanacaste?, Baguar, 600 m, Tate 222 Bouvardia alexandrae A. Carter, Madrono 13:
(Syntype B, presumably destroyed, K, n.v.). 142 Jigs. 1. 2. 1955. Type: MEXICO: Baja
NICARAGUA: Seemann 120 (Syntype B, California, Distr. del Sur, Arroyo del Salto,
presumably destroyed, BM, n.v.). Elias (1976: east of La Paz, 30 Mar 1949, A. Carter 2577
118) wrongly cited Kuntze 2028 (NY) as the (Holotype UC 985926, n.v.; Isotypes F,
"holotype," even though this specimen was MEXU, US (photos MEXU, PTBG)).
not cited in Kuntze's protologue and has Carterella.
Hedyotis arenaria (Rose) W. H. Lewis, (Holotype P-Bonpland, n.v.; Isotype B,
Rhodora 63: 211. 1961. Houstonia arenaria presumably destroyed (photo MO)). Anotis,
Rose, Contr. U.S. Natl. Herb. 1: 70. 1890. Ereicoctis.—Hedyotis wrightii (A. Gray)
Type: MEXICO: Baja California, La Paz, 20
Jan-5 Feb 1890, E. Palmer 28 (Lectotype US Hedyotis corymbosa (L.) Lam., Tabl. Encycl.
47260 (photos MEXU, PTBG)). 1: 272. 1792. Oldenlandia corymbosa L., Sp.
Hedyotis asperuloides Benth., Bot. Voy. PI. 1: 119. 1753. Type: America Meridionali
Sulphur 19,/?/. 13. 1844. f. asperuloides. (Holotype LINN 155.9, n.v.).
Type: MEXICO: Baja California, Cabo San Hedyotis deppeana (Schltdl. & Cham.) Steud.,
Lucas, Barclay s.n. (Holotype BM, n.v.). Nomencl. Bot., ed. 2, 1: 727. 1840.
Gerontogea deppeana Schltdl. & Cham.,
Hedyotis asperuloides Benth. i Linnaea 5: 169. 1830. Type: MEXICO:
(Rose) W. H. Lewis, Rhodora 63: 221. 1961. Veracruz, San Andres, 1829, Schiede &
Houstonia brandegeana Rose, Contr. U.S. Deppe 391 (Holotype presumably HAL, n.v.;
Natl. Herb. 1: 70. 1890. Type: MEXICO: Baja Isotype MO (photos MEXU, PTBG)).
California, La Paz, 20 Jan-5 Feb 1989, E. Oldenlandia—Hedyotis microtheca
Palmer 31 (Lectotype US 47259 (photos (Schltdl. & Cham.) Steud.
MEXU, PTBG); Isolectotypes F, GH, n.v.). Hedyotis dichotoma Sesse & Moc., Fl. Mexic.
Hedyotis bouvardioides Seem., Bot. Voy. 22. 1893. Not Hedyotis dichotoma Cav. 1801.
Herald 296. 1856. Type: MEXICO: Type: Habitat in montibus Sancti Angeli et
Durango ?, Sierra Madre, Seemann s.n. Sancti Augustini. Floret Junio, Sesse &
(Holotype BM, n.v.; Isotypes C, n.v., Mocifio s.n. (presumably MA, n.v.).
fragment F (photo PTBG), GH, n.v.). —Hedyotis exigula W. H. Lewis
Houstonia.—Bouvardia bouvardioides Hedyotis drymarioides (Standi.) W. H. Lewis,
(Seem.) Standi. Rhodora 63: 221. 1961. Houstonia
Hedyotis brevipes (Rose) W. H. Lewis, drymarioides Standi., J. Wash. Acad. Sci. 18:
Rhodora 63: 221. 1961. Houstonia brevipes 162. 1928. Type: MEXICO: Tamaulipas, on
Rose, Contr. U.S. Natl. Herb. 1: 83. 1890. mountains south of Victoria, 1000 m, 9 Apr
Type: MEXICO: Baja California, Santa 1926, R. Runyon 870 (Holotype US 1315865
Rosalia, 24 Feb-3 Mar 1889, E. Palmer 202 (photos MEXU, PTBG)).
(Lectotype US 47258 (photos MEXU, Hedyotis exigula W. H. Lewis, Rhodora 63:
PTBG); Isolectotype US (photos MEXU, 221. 1961. Nom. nov. fox Houstonia gracilis
PTBG)). Brandegee, Zoe 5: 238. 1907. Not Hedyotis
Hedyotis callitrichoides (Griseb.) W. H. Lewis, gracilis DC, 1830. Type: MEXICO:
Rhodora 63: 222. 1961. Oldenlandia Veracruz, Orizaba, May 1904, C. A. Purpus
callitrichoides Griseb. Mem. Amer. Acad. 1251 (Holotype UC, n.v.; Isotypes F, MO
Arts, n.s. 8: 506. 1863. Type: CUBA: Without (photos MEXU, PTBG)).
precise locality, ad ripas rivulorum, C. Wright Hedyotis fruticosa Sesse & Moc., Fl. Mexic. 22.
268 (Lectotype GOET, n.v.; designated by 1893. Not Houstonia fruticosa Willd. ex
Lewis, 1961). Syntype: CUBA: C. Wright Roem. & Schult., 1818. Type: MEXICO:
1640 (GOET, n.v.). Oldenlandiopsis. Habitat m Mexici circuitibus, Sesse & Mocifio
Hedyotis cervantesii Kunth in Humb., Bonpl. & 553 (Presumably MA, n.v.; Isotype F (photo
Kunth, Nov. Gen. Sp. 3: 390. 1820. Norn, MO)).—Bouvardia obovata Kunth in
superfl. Based on same type as Hedyotis Humb., Bonpl. & Kunth,
pygmaea Roem. & Schult., Syst. Veg. 3: 526. Hedyotis galeottii (M. Martens) Terrell &
1818. Not Houstonia pygmaea C. H. Mueller Lorence, Phytologia 66: 1. 1989. Declieuxia
& M. T. Muller, 1936. Type: MEXICO: galeottii M. Martens in M. Martens &
Without precise locality, A. Bonpland s.n. Galeotti, Bull. Acad. Roy. Sci. Bruxelles 11:
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

231. 1844. Type: MEXICO: Oaxaca, Llano Hedyotis latifolia (M. Martens & Galeotti)
Verde, 3000-7000 ft., Apr 1840, H. Galeotti Walp., Rep. Sp. Nov. 6: 55. 1846.
2603 (Holotype BR (photo PTBG); Isotypes Oldenlandia latifolia M. Martens & Galeotti,
BR, n.v., F, US (photo PTBG)). Bull. Acad. Roy. Sci. Bruxelles 11: 235. 1844.
Hedyotis gracilenta (I. M. Johnst.) W. H. Not Houstonia latifolia Willd. ex Roem. &
Lewis, Rhodora 63: 222. 1961. Houstonia Schult., 1818. Type: MEXICO: Veracruz,
gracilenta I. M. Johnst., Proc. Calif. Acad. Mirador, 3000 pieds, May 1840, H. Galeotti
Sci. 4: 1174. 1824. Type: MEXICO: Baja 2556 (Holotype presumably BR, n.v.; Isotype
California, San Diego Island, rocky island F (photos MEXU, PTBG)).—Hedyotis
slope facing sea, 2 May 1921, /. M. Johnston exigula W. H. Lewis
3927 (Holotype UC, n.v.; Isotype US (photos Hedyotis leucophylla (Kunth) Spreng., Syst. 1:
MEXU, PTBG)). 411. 1826 lla Kunth in
Hedyotis gracilis Brandegee, Zoe 5: 238. 1907. Humb., Bonpl. & Kunth, Nov. Gen. Sp. PI. 3:
Not Hedyotis gracilis DC, 1830. Type: 395, t. 290. 1820. Type: MEXICO: Inter Alto
MEXICO: Veracruz, Orizaba, May 1904, de Peregrino et flumen Papagallo, Humboldt
C. A. Purpus 1251 (Holotype UC, n.v.; & Bonplands.n. (Holotype P-Bonpland, n.v.;
Isotypes F, MO (photos MEXU, Isotypes B (photo MO), P (photos MEXU,
PTBG)).—Hedyotis exigula W. H. Lewis PTBG)).—Rondeletia leucophylla Kunth in
Hedyotis herbacea L., Sp. PI. 102. 1753. Type: Humb., Bonpl. & Kunth
CEYLON: (Presumably LINN, n.v.). Hedyotis longipes (S. Watson) W. H. Lewis,
Oldenlandia. Rhodora 63: 222. 1961. Houstonia longipes S.
Hedyotis humifusa A. Gray, Boston J. Nat. Watson, Proc. Amer. Acad. Arts 18: 97. 1883.
Hist. 6: 216. 1850. Type: U.S.A.: Texas, near Syntypes: MEXICO: Without precise locality,
Fredericksburg, 1847, F. Lindheimer 621 States of Coahuila and Nuevo Leon, Feb-Oct
(Holotype GH, n.v.; Isotypes BM, n.v., K, 1880, E. Palmer 394 (US 47845 (photos
n.v., MICH, n.v., MO, PH, n.v., US, n.v.). MEXU, PTBG)); Nuevo Leon, near
Houstonia, Oldenlandia. Monterrey, 1-26 Feb 1880, E. Palmer 395
Hedyotis intricata Fosberg, Lloydia 4: 290. (US, 2 sheets (photos MEXU, PTBG)).
1941. Based on Houstonia fasciculata A. Hedyotis lutea Sesse & Moc., Fl. Mexic. 22.
Gray, Proc. Amer. Acad. Arts 17: 203. 1882. 1893. Type: Habitat in montibus
Not Hedyotis fasciculata Bertol., 1850. Temascaltepec, floret Julio, Sesse & Mocino
Syntypes: U.S.A.: Texas, Presidio, Bigelow s.n. (presumably MA, n.v.).—Status
s.n. (GH, n.v.); MEXICO: Coahuila, near
Parras and Monclova, Palmer 404 (GH, n.v.), Hedvotis mcvaughii Terrell, Novon 6: 128, fig.
Palmer 406 (GH, n.v.). 1. "l996. Type: MEXICO: Jalisco, Mun. de
Hedyotis kingii (Terrell) G. L. Nesom, Syst. Cabo Corrientes, steep mountain sides 3-10
Bot. 13: 434. 1988. Houstonia kingii Terrell, km generally E on the road to Mina del Cuale
Brittonia 32: 491, fig. 2. 1980. Type: from the junction 5 km NW of El Tuito,
MEXICO: Oaxaca, E-facing slopes, 850-1150 m, 16-19 Feb 1975, R. McVaugh
mountains along route 175, 28 km by road 26426 (Holotype MICH, n.v.).
south of Valle Nacional on way to Ixtlan de Hedyotis mexicana Sesse & Moc., PI. Nov.
Juarez, ca. 2000 m, 30 Jul 1959, R. M. King Hispan. ed. 1,15. 1887. Type: MEXICO:
2107 (Holotype MICH, 2 sheets, n.v.). Habitat in Mexici circuitibus, Sesse & Mocino
Hedyotis lancifolia Schumach. in Schumach. & s.n. (presumably MA, n.v.).—Bouvardia
Thonn., Beskr. Guin. PL 72. 1827. Type: ternifolia (Cav.) Schltdl.
GHANA: Valley of Aquapim, Thonning 210 Hedvotis microtheca (Schltdl. & Cham.)
(Lecototype C, n.v.; Isolectotype S, n.v.; Steud., Nomencl. Bot. ed. 2, 1: 728. 1840.
designated by Verdcourt, 1976: 292). Gerontogea microtheca Schltdl. & Cham.,
 Linnaea 5: 169. 1830. Type: MEXICO: Retion, Sierra de la Trinidad, Nov 1902, T. S.
Veracruz, in sylvis Papantlae, Schiede & Brandegee s.n. (Holotype UC, n.v.; Isotypes
Deppe 390 (Holotype presumably HAL, n.v.; GH, n.v., US (photos MEXU, PTBG)).
Isotype MO (photos MEXU, PTBG)). Hedyotis pentandra Schumach. & Thonn.,
Oldenlandia. Kongel. Danske Vidensk. Selsk.
Hedyotis mucronata Benth., Bot. Voy. Sulphur Naturvidensk., Math. Afh. 3: 71. 1827. Type:
19. 1844. Type: MEXICO: Baja California, GUINEA: Aquapim, "S" [Haaslund Smith?]
Magdalena Bay, Barclay s.n. (Holotype BM, (location unknown).—Pentodon pentandrus
n.v.). Houstonia. (Schumach. & Thon.) Vatke
Hedyotis mullerae Fosberg, Lloydia 4: 288,^/g. Hedyotis polypremoides (A. Gray) Shinners,
3. 1941. Type: MEXICO: Coahuila, Mun. de Field & Lab. 17: 168. 1949. Houstonia
Cuatro Cienegas, Canon del Agua, Sierra de la polypremoides A. Gray, Proc. Amer. Acad.
Madera, 9 Sep 1939, C. H. Midler & K. Arts 21: 379. 1886. Type: MEXICO:
Midler 3234 (Holotype US 2133305 (photos Chihuahua, Santa Eulalia Mountains, 26 Sep
MEXU, PTBG); Isotype GH, n.v.). 1885, C. G. Pringle 356 (Lectotype GH, n.v.;
Houstonia. Isolectotypes MO (photos MEXU, PTBG),
Hedyotis mutica Wedd., Chlor. And. 2: 43. US (photos MEXU, US)).
1857. Type: COLOMBIA: Cundimarca, Prov. •o/ypremoides A. Gray var. bigelovii
Bogota, Goudot s.n. (Lectotype P, n.v.; Greenman, Proc. Amer. Acad. Arts 32: 291.
Isolectotype F, n.v.; designated by Mena V., 1897. Type: U.S.A.: Florence Mountains, Jun
1986).—Arcytophyllum muticum (Wedd.) 1852, Bigelow 437 (Holotype GH, n.v.).
Standi. —Hedyotis polypremoides (A. Gray)
Hedyotis nigricans (Lam.) Fosberg, Lloydia 4: Shinners
287. 1941. Gentiana nigricans Lam., Encycl. Hedyotis pringlei (B. L. Rob.) W. H. Lewis,
2: 645. 1788. Type: MEXICO: Locality and Rhodora 63: 222. 1961. Oldenlandia pringlei
collector unknown (Holotype P-JU, n.v.). B. L. Rob., Proc. Amer. Acad. Arts 27: 169.
Houstonia. 1892. Type: MEXICO: San Luis Potosi,
Hedyotis nigricans (Lam.) Fosberg var. scabra alkaline plains, Hacienda de Angostura, 27
(S. Watson) Fosberg, Lloydia 4: 288. 1941. Jun 1891, C. G. Pringle 3758 (Holotype GH,
Houstonia angustifolia Michx. var. scabra S. n.v.; Isotypes F, MO (photo PTBG), US, 2
Watson, Proc. Amer. Acad. Arts 18: 97. 1883. sheets (photos MEXU, PTBG)).
Type: MEXICO: Coahuila, Saltillo, Caracol Hedyotis pygmaea Roem. & Schult., Syst. Veg.
Mountains, 1-15 Apr 1880, E. Palmer 410 3: 526. 1818. Type: MEXICO: Without
(Holotype GH, n.v.; Isotype US (photos precise locality, A. Bonpland s.n. (Holotype P-
MEXU, PTBG)).—Hedyotis nigricans Bonpland, n.v.; Isotype B, presumably
(Lam.) Fosberg destroyed (photo MO)).—Hedyotis wrightii
Hedyotis palmeri (A. Gray) W. H. Lewis, (A. Gray) Fosberg
Rhodora 63: 222. 1961. Houstonia palmeri A. Hedyotis rubra (Cav.) A. Gray, Mem. Amer.
Gray, Proc. Amer. Acad. Arts 17: 202. 1882. Acad. Arts, n.s. 4: 61. 1849. Houstonia rubra
Type: MEXICO: Coahuila, in the mountains Cav., Icon. 5: 48, t. 474, fig. I. 1799. Type:
east and south of Saltillo, Feb-Oct 1880, E. MEXICO: Hidalgo, Ixmiquilpan, Nee s.n.
Palmer 397 (Syntype US 47431 (photos (presumably MA, n.v.). Oldenlandia.
MEXU, PTBG)); E. Palmer 398 (Syntype US Hedyotis serpyllacea Schltdl., Linnaea 9: 599.
70740 (photos MEXU, PTBG)). 1834. Type: MEXICO: Mexico, inter La Joya
Hedyotis peninsularis (Brandegee) W. H. et San Salvador, Jun 1829, Schiede & Deppe
Lewis, Rhodora 63: 222. 1961. Houstonia 265 (Holotype presumably HAL, n.v.;
peninsularis Brandegee, Zoe 5: 160. 1903. Isotypes B (photo MO), F (photo PTBG),
Type: MEXICO: Baja California, Cape MO). Houstonia.
 INOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Hedyotis sharpii (Terrell) G. L. Nesom, Syst. B. L. Rob., Proc. Amer. Acad. Arts 45: 401.
Bot. 13: 434. 1988. Houstonia sharpii Terrell, 1910. Type: MEXICO: Nuevo Leon, shaded
Brirtonia 32: 490,/?g. 1. 1980. Type: cliffs and mountains, near Monterrey, 25 Apr
MEXICO: Hidalgo, camino enrre Metepec y 1906, C. G Pringle 13877 (Holotype GH,
Tenango de Doria, entre 16 y 31.5 km al NNE n.v.; Isotypes MO (photos MEXU, PTBG),
de Tulancingo, 18 km NNE de Metepec, 29 US (photos MEXU, PTBG)).
Mar 1974, S. D. Koch 7423 (Holotype US Hedyotis vergrandis W. H. Lewis, Rhodora 63:
2754863 (photos MEXU, PTBG); Isotypes 222. 1961. Nom. nov. for Houstonia prostrata
CHAPA, ENCB, MEXU). Brandegee, Zoe 5: 105. 1901. Not Hedyotis
Hedyotis sinaloeae W. H. Lewis, Rhodora 63: prostrata Blume, 1823, nor Korth., 1851.
222. 1961, nom. nov. for Houstonia parvula Type: MEXICO: Baja California, La Palma,
Brandegee, Zoe 5: 221. 1905. Not Hedyotis Cape Region, 25 sep 1899, T. S. Brandegee
parvula (A. Gray) Fosb., 1943. Type: s.n. (Holotype UC, n.v.; Isotypes GH, n.v.,
MEXICO: Sinaloa, vicinity of Culiacan, 12 US, 2 sheets (photos MEXU, PTBG)).
Oct 1904, T. S Brandegee s.n. (Holotype UC, Hedyotis watsonii W. H. Lewis, Rhodora 63:
n.v.; Isotypes GH, n.v., US (photos MEXU, 222. 1961. Nom. nov. for Oldenlandia ovata
PTBG)). S. Watson, Proc. Amer. Acad. 18: 97. 1883.
Hedyotis spellenbergii G. L. Nesom & Not Hedyotis ovata Thunb. ex Maxim., 1833,
Vorobik, Syst. Bot. 13: 432. 1988. Type: nor Thwaites ex Hook, f., 1892. Type:
MEXICO: Chihuahua, Mun. Ocampo, area of MEXICO: Nuevo Leon, Guajuco, Feb-Oct
Cascada de Basaseachic at the confluence of 1880, E. Palmer 399 (Lecototype GH, n.v.;
Rio Basaseachic and Rio Durazno, ca. 2 mi S Isolectotype US 47919 (photos MEXU,
of village of Basaseachic, 2100-2200 m, 18 PTBG)).
May 1985, J. Grimes, T. Ayers, M. Lavan & Hedyotis wrightii (A. Gray) Fosberg, Lloydia
R. Scott 2855 (Holotype TEX, n.v.; Isotypes 4: 290. 1941. Houstonia wrightii A. Gray,
ARIZ, n.v., CAS, n.v., MEXU, n.v., NMC, Proc. Amer. Acad. Arts 17: 202. 1882. Type:
. IS, MEXICO: Nuevo Leon, shaded cliffs of
Hedyotis spinescens Sesse & Moc., Fl. Mexic. mountains, near Monterrey, 25 Apr 1906,
22. 1893. Type: MEXICO: Habitat in aridis C. G. Pringle 13877 (Holotype GH, n.v.;
Tehuacani circuitibus. Floret Maio, Sesse & Isotypes MO (photos MEXU, PTBG), US
Mocino s.n. (presumably MA, n.v.). (photos MEXU, PTBG)).
—Bouvardia erecta (DC.) Standi. Hedyotis xestosperma (B. L. Rob. & Greenman)
Hedyotis teretifolia (Terrell) G. L. Nesom, W. H. Lewis, Rhodora 63: 222. 1961.
Syst. Bot. 13: 434. 1988. Houstonia teretifolia Oldenlandia xestosperma B. L. Rob. &
Terrell, Brirtonia 31: 166. 1979. Type: Greenman, Proc. Amer. Acad. Arts 32: 41.
MEXICO: Coahuila, 12 km NNE of Las 1896. Type: MEXICO: Oaxaca, open glades,
Margaritas on easternmost ridge of Sierra de Sierra de San Felipe, 8000 ft., Jun 1894, C. G.
las Margaritas, 1300-1400 m, 24 Sep 1972, Pringle 4692 (Lectotype GH, n.v.;
M. C. Johnston, F. Chiang & T. Wendt 9508E Isolectotypes BM, n.v., F, K, n.v., MO
(Holotype TEX, n.v.). (photos MEXU, PTBG), MSC, n.v., PH, n.v.,
Hedyotis terrellii Lorence, Novon 7: 51,fig. 3. US, 2 sheets (photos MEXU, PTBG), VT
1997. Type: MEXICO: Oaxaca, Distr. de n.v.); designated by Terrell, 1980). Houstonia.
Mixe, Mun. de Totontepec, 3 km al SW de —Bouvardia xestosperma (B. L. Rob. &
Totontepec, carretera a Mitla, 1910 m, 6 Sep Greenman) Terrell & S. D. Koch
1986, R. Torres C. & C. Martinez 9067
(Holotype PTBG 12314; Isotype MEXU).
Hedyotis umbratilis (B. L. Rob.) W. H. Lewis,
Rhodora 63: 222. 1961. Houstonia u
 (Willd.) K. Schum. [Herein treated as a of Mexican origin cultivated at the Berlin
synonym of Diodia L.] Botanical Garden.—Hoffmannia excelsa
, \ Willd. ex Roem. & (Kunth) K. Schum.
Schult.) K. Schum. in Mart., F. Bras. 6(6): 29. Higginsia psychotriifolia Benth. in Oerst.,
1888 (as ocimitolid) Spenmicoce ocxmifoha Vidensk. Meddel. Dansk Naturhist. Foren.
Willd. ex Roem. & Schult., Syst. Veg. 3: 530. Kjobenhavnl852:50. 1853 [as
1819. Type: WEST INDIES: Without precise psychotriaefolia]. Type: COSTA RICA: In
locality, 1768, Rudolphi s.n. (presumably B- Monte Barba, 6000 ft., May 1847, Oersted
W, n.v.). Diodia.—Spermacoce ocymifolia 11507. (Holotype C, n.v. (photos F, MO)
Willd. ex Roem. & Schult. Isotype US (photos MEXU, PTBG)).
-Hoffmannia psychotriifolia (Benth. in
Oerst.) Griseb.
Higginsia refulgens Hook, f, Bot. Mag. tab.
DC. 5346. 1862. Type: Based on material
Icxascpalum angustifolium Bartl. ex DC, cultivated at Kew thought to be from South
Prodr. 4: 561. 1830. Type: MEXICO: America, but most probably from Chiapas,
Hidalgo, Real del Monte, Haenke s.n. Mexico, or Central America (Holotype
(Holotype PR, n.v., fragment G-DC, n.v. presumably K, n.v.).—Hoffmannia refulgens
(microfiche 4: 561.1); Isotype MO (photos (Hook, f.) Hemsl.
MEXU, PTBG)).—Diodia c Higginsia refulgens Hook. f. var. 2 argyroneura
(Linden) Regel, Gartenfl. 17: 116. 1868.
Campylobotrys argyroneura Linden, Belg.
Higginsia Pers., Syn. PI. 1: 133. 1805. Type: Hortic. 8: 166. 1858. Type: Not designated,
Higginsia obovata Pers.. "Higgensia" orth. based on cultivated material collected by
mut. Steud. Norn., ed. 2, 1: 767. 1840. [Herein Ghiesbreght in Chiapas, Mexico.
treated as a synonym of Hoffmannia Sw.] —Hoffmannia refulgens (Hook, f.) Hemsl.
discolor (Lem.) Planch., Fl. Serres. 5: i fulgens Hook. f. var. 6 ortgiesii
482d. 1849. Campylobotrys discolor Lem., Fl. Regel, Gartenfl. 17: 116. 1868. Type: Not
Serres 3: Misc. 37, fig. 1837. Type: Not designated, based on cultivated material of
designated, based on cultivated material Mexican origin.—Hoffmannia regalis (Hook,
collected by Galeotti in Mexico. f.) Hemsl.
—Hoffmannia discolor (Lem.) Hemsl. Higginsia refulgens Hook. f. var. P pyrophylla
Higginsia ghiesbreghtii (Lem.) Hook., Bot. Regel, Gartenfl. 17: 116. 1868. Type: Not
Mag. tab. 5383. 1863. Campylobotrys designated, based on cultivated material from
ghiesbreghtii Lem., Illust. Hort. 8: pi. 279. Volcan San Martin, Veracruz, Mexico.
1861 [as Gheisbechtii]. Type: Not designated, —Hoffmannia refulgens (Hook, f.) Hemsl.
based on cultivated material from seed Higginsia refulgens Hook. f. var. S roezlii
collected by Ghiesbreght in Chiapas, Mexico. Regel, Gartenfl. 17: 116. 1868. Type: Not
—Hoffmannia ghiesbreghtii (Lem.) Hemsl. designated, based on cultivated material of
Higginsia macrophylla Planch., Fl. Serres 5: Mexican origin.—Hoffmannia regalis (Hook,
482d. 1849. Type: Not designated, described f.) Hemsl.
from cultivated material grown from seeds Higginsia refulgens Hook. f. var. J" smaragdina
collected in Guatemala.—Status uncertain, Regel, Gartenfl. 17: 116. 1868. Type: Not
probably Hoffmannia nicotianifolia (M. designated, based on cultivated material of
Martens & Galeotti) L. O. Williams Mexican origin—Hoffmannia regalis (Hook,
Higginsia mexicana Klotzsch in Link, Klotzsch f.) Hemsl.
& Otto, Ic. PL Rar. Hort. Berol. 1: 57, tab. 23. Higginsia regalis Hook, f., Bot. Mag. tab. 5280.
1841. Type: Not designated, based on material 1861. Type: Based on material cultivated at
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

Kew of unknown origin (Holotype Hatheway 1371 (Holotype US (photos

presumably K, n.v.).—Hoffmannia regalis MEXU, PTBG); Isotypes F, GH, n.v.).
(Hook, f.) Hemsl. —Hillia panamensis Standi.
Hillia ligulifolia Dwyer, Ann. Missouri Bot.
Hillia Jacq., Enum. PI. Carib. 3. 1760. Type: Gard. 67: 218. 1980. Type: PANAMA:
Hillia parasitica Jacq. Veraguas, lower montane wet forest 6-7 km
Hillia allenii C. M. Taylor, Selbyana 11: 32. W of Santa Fe on new road past agriculture
1989. Based on Ravnia panamensis Steyerm., school, 2900 ft., 17 Feb 1974, M. Nee 9812
Ceiba 3: 22. 1952. Not Hillia panamensis (Holotype MO 2244248 (photo PTBG);
Standi, 1921. Type: PANAMA: Code, North Isotype MO 2244247 (photo PTBG)).
Hills, El Valle de Anton, 29 Jun 1946, P. H —Cosmibuena valerii (Standi.) C. M. Taylor
Allen 3558 [collector's name and number Hillia longifilamentosa (Steyerm.) C. M.
erroneously omitted in the protologue] Taylor, Selbyana 11: 32. 1989. Ravnia
(Holotype F 1394242 (Photo PTBG); Isotype longifilamentosa Steyerm., Ceiba 3(1): 21.
MO, n.v.). 1952. Type: PANAMA: Chiriqui, upper
Hillia chiapensis Standi., J. Wash. Acad. Sci. southern slopes of Quebrada Velo, vicinity of
16: 16. 1926. Type: MEXICO: Chiapas, damp Finca Lerida, 5000 ft., 25 Jul 1947, P. H
forest in mountains near Fenix, Apr 1925, Allen 4744 (Holotype F 1394241 (photo
C. A. Purpus 262 (Holotype US 1208244 PTBG); Isotype MO, n.v.).
(photos F, MEXU, PTBG)).—Hillia Hillia loranthoides Standi., J. Wash. Acad. Sci.
panamensis Standi. 18: 165. 1928. Type: COSTA RICA:
Hillia chiapensis Standi, var. grandifolia Guanacaste, Quebrada Serena, southeast of
Dwyer, Ann. Missouri Bot. Gard. 67: 216. Tilaran, 700 m, 27 Jan 1926, P. C. Standley &
1980. Type: PANAMA: Veraguas, lower J. Valerio 46152 (Holotype US 1254482
montane wet forest 6-7 km W of Santa Fe on (photos MEXU, PTBG)).
new road past agricultural school, 2900 ft., M. Hillia macrocarpa Standi. & Steyerm., Publ.
Nee 9698 (Holotype MO 2413929).—Hillia FieldMus. Nat. Hist., Bot. Ser. 23: 23. 1943.
palmana Standi. Type: GUATEMALA: Quezaltenango, lower
iquiensis Dwyer, Ann. Missouri Bot. south-facing slopes of Volcan de Santa Maria,
Gard. 67. 216. 1980. Type: PANAMA: between Santa Maria de Jesus and
Chiriqui, N of San Felix at Chriqui-Bocas del Calahuache, along the great barranca between
Toro border, on Cerro Colorado copper mine Finca Pinneos and Patzulin, 1300-1500 m,
road along continental divide, 5000-5500 ft., Jan 1940, J. A. Steyermark 33667 (Holotype
3 May 1975, 5. Mori & J. Kallunki 5810 F, n.v.).
(Holotype MO 2259027 (photo PTBG); Hillia macrophylla Standi., Publ. Field
Isotype MO 2259026 (photo PTBG)). Columbian Mus., Bot. Ser. 7: 201. 1931.
—Cosmibuena valerii (Standi.) C. M. Taylor Type: ECUADOR: Chimborazo, Spruce 6186
Hillia gravumii C. M. Taylor, Selbyana 12: (Holotype K, n.v.).
137, fig. 3. 1991. Type: COSTA RICA: Hillia matudae Standi., Publ. Field Mus. Nat.
Heredia, Finca La Selva, the OTS field station Hist., Bot. Ser. 22: 51. 1940. Type: MEXICO:
near Puerto Viejo de Sarapiqui, near the Chiapas, Mt. Tacana, 2000-4040 m, Aug
junction of the Rios Puerto Viejo and 1938, E. Matuda 2327 (Holotype F; Isotypes
Sarapiqui, 100 m, 5 May 1980, M. H Grayum GH, n.v., MICH, n.v.).—Cosmibuena
2793 (Holotype DUKE, n.v.; Isotype DUKE, matudae (Standi.) L. O. Williams
n.v.). Hillia maxonii Standi., J. Wash. Acad. Sci. 18:
Hillia hathewayi Fosberg, Sida 2: 387. 1966. 163. 1928. Type: NICARAGUA: Las Nubes
Type: COSTA RICA: Heredia, south slope of and vicinity, south of Managua, 800-900 m,
Volcan Barba, 1950 m, 26 May 1965, W. H 28 Jan 1923, W. R. Maxon, A. D. Harvey &
 A. T. Valentine 7501 (Holotype US 1181212 Hondura, 1500-1700 m, 17-18 Jul 1923,
(photos MEXU, PTBG)). W. R. Maxon & A. D. Harvey 8092 (Holotype
Hillia oaxacana C. M. Taylor, Ann. Missouri US 1181776 (photos MEXU, PTBG)).
Bot. Gard. 78: 523, fig. 2. 1991. Type: —Cosmibuena valerii (Standi.) C. M. Taylor
MEXICO: Oaxaca, Distr. de Putla, Arroyo de
San Isidro, 3 km al este de Concepcion, Hintonia Bullock, Hook. Icon. Pi. 33, tab. 3295:
Buenavista, 10 Jun 1985, A. Garcia M. & 1. 1935. Type: Hintonia latiflora (Sesse &
R. Torres C. 1523 (Holotype MO; Isotype Mo?, ex DC.) Bullock
MEXU). Hintonia latiflora (Sesse & Mo?, ex DC.)
Hillia palmana Standi., J. Wash. Acad. Sci. 18: Bullock, Hook. Icon. PI. 33, tab. 3295: 4.
164. 1928. Type: COSTA RICA: Heredia, 1935. Coutarea latiflora Sesse & Mo?, ex
vicinity of La Palma, on the road to La DC, Prodr. 4: 350. 1830. Type: Not
Hondura, 1500-1700 m, 17-18 lul 1923, designated, based on an unpublished
W. R. Maxon & A. D. Harvey 8045 (Holotype illustration, Fl. Mexic. Ic. (caique presumably
US 1181721 (photos MEXU, PTBG)). MA, n.v., G-DC, n.v.).
Hillia panamensis Standi., N. Amer. Fl 32 Hintonia latiflora (Sesse & Mo?, ex DC.)
117. 1921. Type: PANAMA: Chinqui, humid Bullock var. leiantha Bullock, Hook. Icon. PI.
forest around Los Siguas Camp, southern 33, tab. 3295: 1. 1935. Type: MEXICO:
slope of Cerro de la Horqueta, 1700 m, 17-19 Mexico, Temascaltepec, Ixtapan, 1000 m,
Mar 1911, H. Pittier 3190 (Holotype US Aug 1932, G. B. Hinton 1258 (Holotype K,
677584 (photos MEXU, PTBG); Isotype US n.v.).—Hintonia latiflora (Sesse & Mo?, ex
(photos MEXU, PTBG)). DC.) Bullock
Hillia tetrandra Sw., Prodr. Veg. Ind Occ 58 Hintonia lumaeana (Baill.) Bullock, Hook.
1788. Type: JAMAICA: Swartzs.n. Icon. PI. 33, tab. 3295: 5. 1935. Coutarea
(presumably S, n.v.). lumaeana Baill., Adansoma 12: 301. 1879.
Hillia tetrandra Sesse & Mo?., Fl. Mexic, ed 2 Type: GUATEMALA: Cote occidentale des
90. 1894. Type: MEXICO: Habitat parasitica ' Andes, 450-500 metres, 14° lat. N, 1878, G.
supra proceriorum arborum axillas in Luma s.n. (Holotype P (photos MEXU,
Tuxtlensibus sylvis, Sesse & Mocino s.n. PTBG)). Portlandia.
(presumably MA, n.v.).—Hillia tetrandra Hintonia octomera (Hemsl.) Bullock, Hook.
Icon. PI. 3, tab. 3295: 6. 1935. Coutarea
Hillia triflora (Oerst.) C. M. Taylor, Selbyana octomera Hemsl., Biol. Centr.-Amer., Bot. 4:
11: 30. 1989. var. triflora. Ravnia triflora 101. 1887. Type: MEXICO: Yucatan,
Oerst., Vidensk. Meddel. Dansk Naturhist. Cozumel Island, Apr 1895, G. F. Gaumer 148
Foren. Kjobenhavn 1852: 50. 1853. Type- (Holotype K, n.v.).
COSTA RICA: Cartago, near Cartago and Hintonia pulchra D. R. Simpson, Phytologia 29:
Candelana, 1800-2400 m, Oersted 11696 277. 1974. Type: COSTA RICA: Osa
(Lectotype C, n.v. (photos F, MO, NY); Peninsula, about 5 km W of Rincon de Osa,
designated by Standley, 1938) 50-200 m, 9-12 Jan 1970, W. C Burger &
Hillia triflora (Oerst.) C. M. Taylor var. pittieri R. L. Liesner 7320 (Holotype F 1729029;
(Standi.) C. M. Taylor, Selbyana 11: 31. 1989 Isotypes MO (photos MEXU, PTBG), US
Ravnia pittieri Standi., N. Amer Fl 32- 114 (photos MEXU, PTBG)).—Osa pulchra
1921. Type: COSTA RICA: Alajuela, Santa' (D. R. Simpson) Aiello
Clara, H. Pittier 13461 (Holotype NY n v Hintonia standleyana Bullock, Hook. Icon. PI.
(photo F)). 3, tab. 3295: 6. 1935. Syntypes: MEXICO:
Hillia valerii Standi., J. Wash. Acad. Sci 18- Chihuahua, E. Palmer 16(K, n.v.); Guerrero,
164. 1928. Type: COSTA RICA: San Jose El Limon, 3 Apr 1903, E. W. Nelson 6985
Vicinity of La Palma, on the road to La (US 399334 (photos MEXU, PTBG)),
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

Langlasse 234 (K, n.v.); Nayarit, Tepic, (Holotype MO 2470655; Isotype MO

Lumholtz s.n. (US); Puebla, Andrieux 337 (K, 2470654).
n.v.), Pringle 8583 (K, n.v.), Seler 849 (US), Hoffmannia arborescens Donn. Sm., Bot. Gaz.
Rose & Hay 5932 (US), Rose, Painter & Rose (Crawfordsville) 37: 417. 1904. Type:
10123 (US); Oaxaca, Purpus 3279 (US), Reko COSTA RICA: Forets de Santa Rosa del
4253 (US), Pringle 7473 (US), Pringle & Copey, 1800 m, Apr 1898, A. Tonduz 8121
Conzatti 27 (US). C. V. Morton annotated the [Herb. Inst. Fysico-geogr. nat. costaricensis
Nelson 6985 specimen at US as lectotype, but 12230] (Holotype US 943523 (photos MEXU,
this choice was not published.—Hintonia PTBG)).
latiflora (Sesse & Moc. ex DC.) Bullock Hoffmannia areolata Standi., Publ. Field Mus.
Nat. Hist., Bot. Ser. 22: 52. 1940. Type:
Hippotis Ruiz & Pavon, Fl. Peruv. Chil. 33 PANAMA: Chiriqui, Boquete District, near
1794. Type: Hippotis trifolia Ruiz & Pavon Bajo Chorro, Jan 1938, M. E. Davidson 40
Hippotis albiflora H. Karat, Fl. Columb. 1: 3 (Holotype F 915621 (photo PTBG); Isotypes
1860. Type: COLOMBIA: Mountains E of GH, n.v., MO).
Bogota, Karsten s.n. (presumably W, n.v.) Hoffmannia asclepiadea Standi., J. Wash.
Acad. Sci. 15: 7. 1925. Type: COSTA RICA:
Hoffmannia Sw., Prodr. 30. 1788. Type: San Jose, Las Nubes, 1500-1900 m, 20-25
Hoffmannia pedunculata Sw. Mar 1924, P. C. Standley 38349 (Holotype
Hoffmannia aeruginosa Standi., Publ. Field US 1153122 (photos MEXU, PTBG)).
Mus. Nat. Hist., Bot. Ser. 18: 1313. 1938. Hoffmannia bullata L. O. Williams, Fieldiana,
Type: COSTA RICA: Zarcero, 1500 m, 10 Bot. 36: 52. 1973. Type: GUATEMALA:
Aug 1937, Austin Smith A86 (Holotype F Izabal, between Virginia and Lago Izabal,
941349 (photo PTBG)). Montana del Mico, 50-100 m, 4 Apr 1940,
nia affinis Hemsl., Diag. PI. Nov. J. A. Steyermark 38719 (Holotype F).
Mexic. 31. 1879. Type: COSTA RICA: —Hoffmannia discolor (Lem.) Hemsl.
Without precise locality, Endres 150 Hoffmannia calycosa Donn. Sm., Bot. Gaz.
(Holotype K, n.v.; Isotype W (photo PTBG)). (Crawfordsville) 40: 4. 1905. Type:
—Hoffmannia nicotianifolia (M. Martens & GUATEMALA: Alta Verapaz, Cubliquitz,
Galeotti) L. O. Williams 350 m., May 1901 H. von Tuerckheim 7912
Hoffmannia amplexifolia Standi., J. Wash. (Holotype US 576489 (photos MEXU,
Acad. Sci. 15: 8. 1925. Type: COSTA RICA: PTBG); Isotypes F (photos MEXU, PTBG),
Cartago, vicinity of Orosi, moist forest, 30 US, 2 sheets (photos MEXU, PTBG)).
Mar 1924, P. C. Standley 39869 (Holotype Hoffmannia capillacea Dwyer, Ann. Missouri
US 1153192 (photos MEXU, PTBG)). Bot. Gard. 56: 274. 1969. Type: PANAMA:
Hoffmannia angustifolia Standi., Contr. U.S. Code, mountains N of El Valle, 2500-3000
Natl. Herb. 20: 206. 1919. Type: ft., 28 May 1967, Lewis, MacBryde. Oliver &
GUATEMALA: Zacatepequez, Acatepeque, Ridgwav 1783 (Holotype MO; Isotypes MO
1290 m, Mar 1892, J. D. Smith 2747 2214603, 2163471). Note: Between the time
(Holotype US 888492 (photos MEXU, Hoffmannia capillacea Dwyer was published
PTBG); Isotype GH, n.v.).—Hoffmannia and the Rubiaceae treatment for the Flora of
nicotianifolia (M. Martens & Galeotti) L. O. Panama was published (Dwyer, 1980), the
Williams wrong labels (T. Elias 1783) were
inadvertently mounted on the type collection.
Missouri Bot. Gard. 67: 226. 1980. Type: Dwyer's protologue lists the type as being
PANAMA: Danen, Cerro Mali on Colomt Lewis et al. 1783. The three collections in the
border, S slope just below summit, 1400 n type folder at MO correspond with the
Feb 1975, A. Gentry & S. Mori 14166 description of H. capillacea but bear Elias
 1783 labels. A reexamination of the Lewis (Holotype US 888491 (photos MEXU,
and Elias field books by C. M. Taylor and me PTBG); Isotype GH, n.v.).
in June 1996 revealed that Elias 1783 was ioffmannia congesta (Oerst.) Dwyer in W. C.
collected in San Bias, Panama, reputedly as a Burger & C. M. Taylor, Fieldiana, Bot., n.s.
species of Annonaceae: "headquarters of Rio 33: 171. 1993. Xerococcus congestus Oerst.,
Mulatupo, tree 5 m, fruits green ripening Vidensk. Meddel. Dansk. Naturhist. Foren
yellow." The Lewis et al. 1783 collection was Kjobenhavn 1852: 52. 1853. Type: COSTA
said to be "Rubiaceae, shrub to 2.2 m, calyx RICA: Turrialba, 3000 ft., Oersted 11776
tube red, flowers white, fruit red." Therefore, (Holotype C, n.v. (photos F, MO); Isotype F,
the Lewis et al. 1783 label seems to be the n.v.).
Hoffmannia conzattii B. L. Rob., Proc. Amer.
Hoffmannia carlsoniae Standi. & L. O. Williams Acad. Arts 45: 406. 1910. Based on Coffea
ex A. R. Molina, Ceiba 14: 3. 1968. Type: obovata Schlecht. & Cham., Linnaea 6: 412.
MEXICO: Chiapas, Los Lagos, 3 mi. NW of 1*31 !//-••/ An, ,(Ru
Rancho San Jose, which is 34 mi. SE of Pavon) Standi., 1930 (based on Ohigginsia
Comitan, 5,000 ft., 15-20 Apr 1949, M. C. obovata Ruiz & Pavon 1798). Type:
Carlson 1874 (Holotype presumably EAP, MEXICO: Veracruz, Canton de Cordoba,
n.v.; Isotype F 1412975).—Hoffmannia Colonia Melchor Ocampo, 1200 m, 19 Jun
huehueteca Standi. & Steyerm. 1896, C. Conzatti 168 (Holotype GH, n.v.).
Hoffmannia carpinterae Standi., N. Amer. Fl. Hoffmannia cryptoneura Standi., Publ. Field
32: 199. 1934. Based on Hoffmannia Mus. Nat. Hist., Bot. Ser. 22: 53. 1940. Type:
macrophylla Standi. 1925, not H. macrophylla MEXICO: Chiapas, Mt. Ovando, 4 Apr 1936,
Hemsl., 1881. Type: COSTA RICA: Cartago, E. Matuda 944 (Holotype F 898899 (photo
Cerro de la Carpintera, 1700 m, P. C. Standley PTBG); Isotypes MICH, n.v., MO (photos
35636 (Holotype US 1152958 (photos MEXU, PTBG), TEX-LL (photos MEXU,
MEXU, PTBG); Isotype US 1152957 (photos PTBG)).
MEXU, PTBG)).—Hoffmannia leucocarpa Hoffmannia culminicola Standi. & L. O.
Standi. Williams, Ceiba 1: 166. 1950. Type:
Hoffmannia cauliflora Hemsl., Diag. PI. HONDURAS: Morazan, Cerro de Uyuca, ca.
Mexic. 30. 1879. Type: GUATEMALA: 1600 m, 2 Mar 1947, P. C. Standley 4797
Without precise locality, Skinner s.n. (Holotype F 1256864 (photos MEXU,
(Holotype K, n.v.). PTBG); Isotype EAP, n.v.).
Hoffmannia cercidifolia Dwyer, Ann. Missouri Hoffmannia cuneatissima B. L. Rob., Proc.
Bot. Gard. 56: 275. 1969. Type: PANAMA: Amer. Acad. Arts 45: 407. 1910. Type:
Darien, Cerro Campamento, S of Cerro Pirre, MEXICO: Morelos, mountain canyon near
20-22 Mar 1968, J. A. Duke 15597 (Holotype Cuernavaca, 6000 ft., 29 May 1898, C. G.
MO 1956874 (photo PTBG)). Pringle 7662 (Holotype GH, n.v.; Isotype US
Hoffmannia chiapensis Standi., Contr. U.S. 334830 (photos MEXU, PTBG)).
Natl. Herb. 20: 206. 1919. Type: MEXICO: Hoffmannia davidsoniae Standi., Publ. Field
Chiapas, Cerro del Boqueron, Jun 1914, C. A. Mus. Nat. Hist., Bot. Ser. 22: 53. 1940. Type:
Purpus 7268, in part (Holotype US 567526 PANAMA: Chiriqui, Boquete District, Bajo
(photos MEXU, PTBG); Isotype F (photos Chorro, 6000 ft., 26 Jan 1938, M. E. Davidson
MEXU, PTBG)).—Hoffmannia 216 (Holotype F 915570 (photo PTBG);
psychotriifolia (Benth in Oerst.) Griseb. Isotypes A, n.v., MO, US (photos MEXU,
Hoffmannia confertiflora Standi., Contr. U.S. PTBG)).
Natl. Herb. 20: 206. 1919. Type: Hoffmannia decurrens Standi., Contr. U.S.
GUATEMALA: Quiche, San Miguel Natl. Herb. 20: 205. 1919. Type: COSTA
Uspantan, 2100 m, 1892, Heyde & Lux 3169 RICA: Forets de Santa Rosa del Copey,
 AND CENTRAL AMERICAN RUBIACEAE

1800-2000 m, Apr 1898, A. Tonduz 12230 A. Gentry & S. Mori 14061 (Holotype MO
(Holotype US 392077 (photos MEXU, 2353753; Isotype MO 2423663).
PTBG); Isotypes P, 2 sheets (photos MEXU, Hoffmannia gesnerioides (Oerst.) Kuntze, Rev.
PTBG)). Gen. PI. 285. 1891. Ophryococcus
Hoffmannia discolor (Lem.) Hemsl., Biol. gesnerioides Oerst., Vidensk. Meddel. Dansk
Centr.-Amer., Bot. 2: 36. 1881. Naturhist. Foren. Kjoebenhavn 1852: 53.
Campylobotrys discolor Lem., Fl. Serres 3: 1853. Type: NICARAGUA: Segovia, Monte
Misc. 37. fig. 1847. Type: Not designated, Pantasmo, Jan 1848, Oersted 11823
based on cultivated plants grown from seed (Holotype C, n.v. (photos F, MO); Isotype F).
collected in Mexico by H. Galeotti. Note: A Hoffmannia ghiesbreghtii (Lem.) Hemsl., Biol.
specimen at P ["en herb. Houtte Juin 1857. Centr.-Amer., Bot. 2: 36. 1881.
Hort. Parisei"] and a specimen at G ["Mexici, Campylobotiys ghiesbreghtii Lem., Illust.
Oaxaca, 3000', fl. albis, Galeotti 7183"] may Hort. S:pl. 279. 1861. Type: Not designated,
correspond with Galeotti's seed collection and based on cultivated material from seed
represent authentic cultivated material. collected by Ghiesbreght, probably in
Higginsia. Chiapas, Mexico. Higginsia.
Hoffmannia dotae Standi., J. Wash. Acad. Sci. Hoffmannia hamelioides Standi., J. Wash.
18: 181. 1928. Type: COSTARICA: San Acad. Sci. 15: 8. 1925. Type: COSTA RICA:
Jose, vicinity of Santa Maria de Dota, San Jose, between Aserri and Tarbaca,
1500-1800 m, 26 Dec 1925-3 Jan 1926, P. C. 1600-1800 m, 12 Feb 1924, P. C. Standley
Standley & J. Valerio 43277 (Holotype US 34149 (Holotype US 1152904 (photos
1253172 (photos MEXU, PTBG)). MEXU, PTBG)).
Hoffmannia eliasii Dwyer, Ann. Missouri Bot. Hoffmannia hammelii C. M. Taylor in W. C.
Gard. 56: 276. 1969. Type: PANAMA: Burger & C. M. Taylor, Fieldiana, Bot., n.s.
Darien, Rio Cuasi mainstream, 2.5 miles S of 33: 174. 1993. Type: COSTA RICA:
Tres Bocas, 28 Apr 1968, J. H. Kirkbride, Jr. Puntarenas, near Sirena, Corcovado National
& J. A. Duke 1129 (Holotype MO 1982892 Park, 0-200 m, 5 Jul 1977, R. L. Liesner 2907
(photos MEXU, PTBG)).—Hoffmannia (Holotype CR, n.v.; Isotype MO).
subauriculata Standi. Hoffmannia haydenii Dwyer, Ann. Missouri
Hoffmannia excelsa (Kunth) K. Schum. in Bot. Gard. 56: 277. 1969. Type: PANAMA:
Mart., Fl. Bras. 6(6): 327. 1889. Psychotria Chiriqui, Cerro Campana, Jul 1965, s. M. V.
excelsa Kunth in Humb., Bonpl. & Kunth, Hayden 117 (Holotype MO 2030831). Note:
Nov. Gen. Sp. 3: 355. 1819. Type: MEXICO: In the protologue the type locality is
Veracruz, Xalapa, A. Bonpland 1445 incorrectly cited as "Finca Collins, ca. 5000
(Holotype presumably P-Bonpland, n.v.; ft." —Hoffmannia subauriculata Standi.
Isotypes B, n.v. (photo MO), P (photos Hoffmannia hondurensis Standi., Publ. Field
MEXU, PTBG)). Deppea. Columbian Mus., Bot. Ser. 4: 329. 1929.
Hoffmannia fimbrianthera Dwyer, Ann. Type: HONDURAS: Atlantida, Lancetilla
Missouri Bot. Gard. 56: 277. 1969. Type: Valley, 200-500 m, 6 Dec 1927-20 Mar
PANAMA: Rio Teribe, just below Puerto 1928, P. C. Standley 52905 (Holotype F
Palenque, 350 m, 12 Apr 1968, J. H. 582885 (photo PTBG); Isotype US (photos
Kirkbride, Jr. & J. A. Duke 556 (Holotype MEXU, PTBG)).
MO 1999724).—Hoffmannia inamoena Hoffmannia huehueteca Standi. & Steyerm.,
Standi. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 249.
Hoffmannia gentryi Dwyer, Ann. Missouri 1947. Type: GUATEMALA: Huehuetenango,
Bot. Gard. 67: 230. 1980. Type: PANAMA: Sierra de los Cuchumatanes, Cerro Negro, 2
Darien, Cerro Tacarcuna, summit camp, along miles east of Las Palmas, 1600-2000 m, 31
stream N of camp, 1550-1650 m, 1 Feb 1975,
 Aug 1942, J. A. Steyermark 51697 (Holotype 2992 collection at P was chosen as lectotype,
F 1132825 (photo PTBG)). because it is most representative, with flowers,
Hoffmannia inamoena Standi., J. Wash. Acad. fruits, and full-sized leaves.—Hoffmannia
Sci. 18: 179. 1928. Type: COSTA RICA: nicotianifolia (M. Martens & Galeotti) L. O.
Guanacaste, Los Ayotes, near Tilaran, 600 m, Williams
21 Jan 1926, P. C. Standley & J. Valerio Hoffmannia leucocarpa Standi., J. Wash.
45421 (Holotype US 1254102 (photos Acad. Sci. 15: 9. 1925. Type: COSTA RICA:
MEXU, PTBG)). San Jose, Las Nubes, 1300-1600 m, Mar
Hoffmannia ixtlanensis Lorence, Novon 4: 1924, P. C. Standley 38340 (Holotype US
\2\,fig. 2a, b. 1994. Type: MEXICO: 1153121 (photos MEXU, PTBG)).
Oaxaca, Distr. de Ixtlan, Sierra de Juarez; ruta Hoffmannia lewisiana Dwyer, Ann. Missouri
175 Tuxtepec a Oaxaca, ca. 5 km al N de Bot. Gard. 56: 279. 1969. Type: PANAMA:
Vista Hermosa, 1260 m, 14 Apr 1982, D. H. Chiriqui, vicinity of Cerro Punta, 6800 ft., 31
Lorence, R. Torres C. & R. Cedillo T. 4013 May-1 Jun 1967, Ridgway & Solis 2413
(Holotype MEXU; Isotypes MO, PTBG). (Holotype MO 1966156).
Hoffmannia josefina Standi., J. Wash. Acad. Sci. Hoffmannia liesneriana L. O. Williams,
15: 8. 1925. Type: COSTA RICA: San Jose, Fieldiana, Bot. 36: 54. 1973. Type: COSTA
between Aserri and Tarbaca, 1800 m, 12 Feb RICA: Limon, Guapiles, a 2 millas al norte de
1924, P. C. Standley 34143 (Holotype US Diamantes, 150 m, 9 Aug 1963, A. Jimenez
1152900 (photos MEXU, PTBG)). 1054 M. (Holotype F 1690189 (photo
—Hoffmannia arborescens Donn. Sm. PTBG)).
a kirkbridei Dwyer, Ann. Missouri Hoffmannia lineolata Donn. Sm., Bot. Gaz.
Bot. Gard. 56: 278. 1969. Type: PANAMA: (Crawfordsville) 40: 5. 1905. Type:
Bocas del Toro, Quebrada Huron, ca. GUATEMALA: Alta Verapaz, Cubliquitz,
30(M00 ft., 11 Apr 1968, J. H. Kirkbride, Jr. 350 m, May 1902, H. von Tuerckheim 8227
& J. A. Duke 429 (Holotype MO 1968253). (Holotype US no 576514 (photos MEXU,
—Hoffmannia vesiculifera Standi. PTBG); Isotypes US, 2 sheets (photos MEXU,
Hoffmannia lancistigma Dwyer, Ann. Missouri PTBG)).—Hoffmannia regalis (Hook, f.)
Bot. Gard. 56: 279. 1969. Type: PANAMA: Hemsl.
Chiriqui, Boquete, Cerro Horqueta, Hoffmannia longicalycina Dwyer, Ann.
5000-6000 ft., 8 Aug 1967, J. D. Dwyer & Missouri Bot. Gard. 56: 280. 1969. Type:
S. M. V. Hayden 7735 (Holotype MO PANAMA: Darien, Cuasi-Cana Trail between
2065523). Cerro Campamiento and La Escalera to
Hoffmannia laxa Standi., Publ. Field Mus. Nat. paramo, E of Tres Bocas, 30 Apr 1968, J. H.
Hist., Bot. Ser. 22: 116. 1940. Type: COSTA Kirkbride, Jr. & J. A. Duke 1278 (Holotype
RICA: Alajuela, Canton de San Carlos, Sucre, MO 2165098; Isotype MO 1686749).
1000 m, 30 Mar 1939, A. Smith F1851 —Status uncertain, probably Amphidaysa
(Holotype F 980999 (photo PTBG); Isotype sp.
US (photos MEXU, PTBG)). Hoffmannia longipetiolata Pol., Linnaea 41:
Hoffmannia lenticellata Hemsl., Diag. PI. Nov. 567. 1877. Type: COSTA RICA: In sylvis
Mexic. 30. 1879. Type: MEXICO: Veracruz, primaevis Carpinterae, 24 Jun 1875, H.
region d'Orizaba, Rio Blanco, 29 Aout 1866, Polakowsky 134 (B, presumably destroyed
Bourgeau 2992 (Lectotype P, here designated (photo F), lectotype fragment F, here
(photos MEXU, PTBG); Isolectotype P designated (photo PTBG)). Syntype: COSTA
(photos MEXU, PTBG)). Syntypes: RICA: In sylvis primaevis Carpinterae, 24 Jun
MEXICO: Veracruz, Orizaba, Botteri 974 1875, H. Polakowksy 140 (B, presumably
(Syntype P, photos MEXU, PTBG); Jalapa, destroyed). The fragment of Polakowsky 134
Linden 1 (Syntype K, n.v.). The Bourgeau ; lectotype.
 k NOMENCLATOR OF MEXICAN /

Hoffmannia longipetiolata Pol. var. minor Pol., Hoffmannia montana L. O. Williams,

Linnaea41: 568. 1877. Type: COSTA RICA: Fieldiana, Bot. 36: 55. 1973. Type:
In sylvis primaevis Carpinterae, Jun 1875, H. GUATEMALA: Quezaltenango, Volcan
Polakowsky 144 (Holotype B, presumably Zunil, 8000 ft., 4 Aug 1934, A. F. Skutch 936
destroyed).—Hoffmannia longipetiolata Pol. (Holotype F 934132 (photos MEXU, PTBG);
Hoffmannia macrophylla (Planch.) Hemsl., Biol. Isotype A (photo PTBG)).
Centr.-Amer., Bot. 2: 37. \U\. Higginsia Hoff nam a morn Dwyer, Ann. Missouri Bot.
macrophylla Planch., Fl. Serres 5: 482d. 1849. Gard. 67: 236. 1980. Type: PANAMA:
Type: Not designated, based on cultivated Chiriqui, N of San Felix at Chiriqui-Bocas del
material grown from seeds collected in Toro border, on Cerro Colorado copper mine
Guatemala.—Hoffmannia nicotianifolia (M. road along continental divide, 5000-5500 ft.,
Martens & Galeotti) L. O. Williams 5 May 1975, S. Mori & J. Kallunki 5915
Hoffmannia macrophylla Standi., J. Wash. (Holotype MO 2259023).—Hoffmannia
Acad. Sci. 15: 9. 1925. Not//, macrophylla vesiculifera Standi.
(Planch.) Hemsl., 1881. Type: COSTA RICA: Hoffmannia nebulosa Standi, in Yunck., Field
Cartago, Cerro de La Carpintera, 1500-1850 Mus. Nat. Hist., Bot. Ser. 17: 396. 1938.
m, Feb 1924, P. C. Standley 35636 (Holotype Type: HONDURAS: Comayagua, above the
US 1152957 (photos MEXU, PTBG); Isotype plains of Siguatepeque, near summit of range
US 1152958 (photos MEXU, PTBG)). above El Achote, 1850 m, 1 Aug 1936, T. G
—Hoffmannia leucocarpa Standi. Yuncker, R. F. Dawson & H. R. Youse 6252
Hoffmannia macrosiphon, Standi., Field Mus. (Holotype F 857687 (photo PTBG)).
Nat. Hist., Bot. ser. 17: 276. 1937. Type: Hoffmannia nesiota Donn. Sm., Bot. Gaz.
GUATEMALA: Quezaltenango, Volcan de (Crawfordsville) 61: 374. 1916. Type:
Zunil, 8000 ft., 4 Aug 1934, A. F. Skutch 937 COSTA RICA: Cocos Island, Wafer Bay, 18
(Holotype F 813468 (photo PTBG); Isotype Jun 1898, H. Pittier 12387 (Holotype US
941381 (photos MEXU, PTBG)).
Hoffmannia manussatani Dwyer, Ann. Hoffmannia nicotianifolia (M. Martens &
Missouri Bot. Gard. 67: 234. 1980. Type: Galeotti) L. O. Williams, Fieldiana, Bot. 36:
PANAMA: Bocas del Toro, Robalo Trail, N 56. 1973. Psychotria nicotianifolia M.
slopes of Cerro Horqueta, 6000-7000 ft., 5-7 Martens & Galeotti, Bull. Acad. Roy. Sci.
Aug 1947, Allen 4978 (Holotype MO Bruxelles 11: 229. 1844 [as nicotianaefolia].
1636310). Type: MEXICO: Veracruz, Cordillera pres
Hoffmannia mexicana (Link, Klotzsch & Otto) Xalapa, 3000 pieds, 1840, H. Galeotti 7066
Hemsl., Biol. Centr.-Amer., Bot. 2: 37. 1881. (Holotype presumably BR, n.v.; Isotype P
Higginsia mexicana Link, Klotzsch & Otto, (photos MEXU, PTBG)).
Ic. PI. Rar. Hort. Ber. 1: 57, t. 23. 1841. Type: Hoffmannia nubium Standi. & L. O. Williams,
Not designated, based on material of Mexican Ceiba 1: 167. 1950. Type: HONDURAS:
origin cultivated at the Berlin Botanical Morazan, drainage of the Rio Yeguare, at
Garden.—Hoffmannia excelsa (Kunth) K. about long. 87°W, lat. 14°N, 6000 ft., 20 Jul
Schum. 1948, S. F. Glassman 1996 (Holotype US
Hoffmannia minuticarpa Dwyer & Lorence, 2215976 (photos MEXU, PTBG); Isotypes F,
Novon4: 122, fig. 2c, d. 1994. Type: GH (photo PTBG), MEXU, WIS).
MEXICO: Veracruz, Mun. de Soteapan, —Hoffmannia cryptoneura Standi.
Cumbres de Bastonal, ca. 10-11 km al SE de Hoffmannia oreogena L. O. Williams,
Tebanca, 1200 m, 26 Apr 1982, R. Cedillo T.. Fieldiana, Bot. 36: 57. 1973. Type:
D. Lorence & G. Ibarra M. 1280 (Holotype HONDURAS: La Paz, Cordillera Guajiquiro,
MEXU; Isotype MO). Montana Verde, aldea Las Marias, 2100 m, 23
May 1964, A. Molina R. & A. R. Molina
 14052 (Holotype F 1616225 (photo PTBG); (Holotype US 579835 (photos MEXU,
Isotype EAP, n.v.). PTBG); Isotype GH (photo PTBG)).
Hoffmannia oreophila L. O. Williams, Hoffmannia pittieri Standi, Contr. U.S. Natl
Fieldiana, Bot. 36: 57. 1973. Type: Herb. 18: 140. 1916. Type: PANAMA:
NICARAGUA: Matagalpa, Cordillera Central Chiriqui, between Alto de las Palmas and top
de Nicaragua between Matagalpa and of Cerro de la Horqueta, 2100-2268 m, 18
Jinotega, Disparate de Potter, near Santa Mar 1911, H Pittier 3247 (Holotype US
Maria de Ostuma, 1500 m, 20-24 Feb 1963, 677642 (photos MEXU, PTBG)).
L. O. Williams, A. Molina R. & T. Williams Hoffmannia psychotriifolia (Benth. in Oerst.)
24709 (Holotype F 1606614 (photo PTBG); Griseb., Fl Brit. W. Ind. 321. 1861. Higginsia
Isotype EAP, n.v.).—Hoffmannia psychotriifolia Benth. in Oerst., Vidensk.
nicotianifolia (M. Martens & Galeotti) L. O. Meddel Dansk Naturhist. Foren. Kjobenhavn
Williams 1852: 50. 1853 [aspsychotriaefolia]. Type:
Hoffmannia orizabensis Standi., Contr. U.S. COSTA RICA: In monte Barba (Volcan de
Natl. Herb. 20: 205. 1919. Type: MEXICO: Barba), 6000 ft., May 1847, Oersted 11507
Veracruz, region of Orizaba, 1855, Mueller (Holotype C, n.v. (photos F, MO); Isotype US
1359 (Holotype GH, n.v.). (photos MEXU, PTBG)).
Hoffmannia ostaurea Dwyer, Ann. Missouri Hoffmannia pusilla L. O. Williams, Fieldiana,
Bot. Gard. 56: 280. 1969. Type: PANAMA: Bot. 34: 117. 1972. Type: GUATEMALA:
Bocas del Toro, Rio Terrible between Alta Verapaz, between Sepacuite and
Quebrada Huron and Puerto Palenque, Secoyocte, 1100 m, 24 May 1905, H. Pittier
120-140 m, 12 Apr 1968, J. H. Kirkbride, Jr. 348 (Holotype US 1083004 (photos MEXU,
& J. A. Duke 501 (Holotype MO 1982893). PTBG)).—Pinarophyllon bullatum Standi.
Hoffmannia pallidiflora Standi., J. Wash. Acad. Hoffmannia pustulata Dwyer, Ann. Missouri
Sci. 15: 9. 1925. Type: COSTA RICA: San Bot. Gard. 67: 240. 1980. Type: PANAMA:
Jose, La Hondura, 1300-1700 m, 16 Mar Colon, Rio Guanche, pressed from cultivation,
1924, P. C. Standley 37877 (Holotype US May-Jun 1972, R. L. Dressier s.n. (Holotype
1153101 (photos MEXU, PTBG)). MO 2178490 (photo PTBG)).
—Hoffmannia viridis Rusby Hoffmannia quadrifolia Standi. & Steyerm.,
Hoffmannia panamensis Standi., Contr. U.S. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 250.
Natl. Herb. 20: 204. 1919. Type: PANAMA: 1947. GUATEMALA: Zacapa, Sierra de las
Chiriqui, above El Boquete, forests along the Minas, around the summit of Monte Virgen,
Rio Ladrillo and vicinity, 1200-1300 m, 6 ca. 2400 m, 12-13 Jan 1942, J. A. Steyermark
Mar 1911, H Pittier 3056 (Holotype US 42600 (Holotype F 1133818 (photo PTBG)).
677443 (photos MEXU, PTBG)). Hoffmannia racemifera Standi. & Steyerm.,
—Hoffmannia psychotriifolia (Benth. in Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 249.
Oerst.) Griseb. 1947. Type: GUATEMALA: Izabal, Cerro
Hoffmannia phoenicopoda K. Schum., San Gil, 300-900 m, 25 Dec \94\,J.A.
Notizbl. Bot. Gart. Berlin 2: 276. 1899. Type: Steyermark 41910 (Holotype F 1134162
Not designated, based on material cultivated (photos PTBG); Isotypes F 1134161 (photos
in the Berlin Botanical Garden of uncertain PTBG), US (photos MEXU, PTBG)).
origin, probably from Mexico or Central Hoffmannia ramonensis Standi, J. Wash. Acad.
America. Sci. 18: 180. 1928. Type: COSTA RICA: Rio
Hoffmannia piratarum Standi, J. Wash. Acad. Barranca a San Juan, pres de San Ramon,
Sci. 18: 180. 1928. Type: COSTA RICA: 1300-1400 m, 25 Apr 1913, A Tonduz 17812
Cocos Island, Forets humides de la vallee de (Holotype US 861910 (photos MEXU,
Wafer, 10-200 m, Jan 1902, H. Pittier 16259 PTBG)).—Hoffmannia dotae Standi.
A NOMENCLATOR OF MEXICAN AND CENTRAL

Hoffmannia refulgens (Hook, f.) Hemsl., Biol. lectotype as it is most complete, with intact
Centr.-Amer., Bot. 2: 37. 1881. Higginsia leaves, flowers, and inflorescences.
refulgens Hook, f, Bot. Mag. tab. 5346. 1862. Hoffmannia rotundata Standi., Contr. U.S. Natl.
Type: Based on plants grown at Kew from Herb. 20: 204. 1919. Type: MEXICO:
seed thought to be from South America, but Chiapas, Cerro del Boqueron, Jun 1914, C. A.
most probably from Central America or Purpus 7268 in part (Holotype MO 765057).
Chiapas, Mexico (Holotype presumably K, —Hoffmannia nicotianifolia (M. Martens &
n.v.). Galeotti) L. O. Williams
Hoffmannia regalis (Hook, f.) Hemsl., Biol. Hoffmannia rubripigmenta Dywer, Ann.
Centr.-Amer., Bot. 2: 37. 1881. Higginsia Missouri Bot. Gard. 56: 283. 1969. Type:
regalis Hook, f, . Mag. tab. PANAMA: Darien, between upper Rio
Type: Based on ] ts grown at Kew Membrillo and Camp 7 on construction road
presumably fron ;xican seed (Holotype to San Bias, 100-800 m, 26-28 Apr 1967,
presumably K, n.v.). Campylobotrys. J. A. Duke 10870 (Holotype MO 1981851).
Hoffmannia rexmontis Dywer, Ann. Missouri Hoffmannia sessilifolia L. O. Williams,
Bot. Gard. 67: 241. 1980. Type: PANAMA: Phytologia 25: 463. 1973. Type:
Chiriqui, NW side of Cerro Pando, 21 Jul GUATEMALA: Baja Verapaz, Sierra de las
1971, T. Croat 15934 (Holotype MO Minas, above Purulha, 1600 m, 2 Jan 1973,
2692337; Isotypes MO 2692338, MO L. O. Williams, A. Molina R. & T. P. Williams
2150879).—Hoffmannia pallidiflora Standi. 41977 (Holotype F 1716220 (photo PTBG);
Hoffmannia rhizantha Standi., Carnegie Inst. Isotype 1716221).
Wash. Publ. 461: 90. 1935. Type: BRITISH Hoffmannia steinvorthii Fosberg, Sida 2: 388.
HONDURAS (BELIZE): Esperanza Trail, 1966. Type: COSTA RICA: Heredia, south
630 m, 26 Jun 1934, W. A. Schipp S-731 slope of Volcan Barba, 2000 m, 26 Jun 1965,
(Holotype F 729968 (photo PTBG)). W. H. Hatheway & A. Jimenez 1461
Hoffmannia riparia Standi., Field Mus. Nat. (Holotype US 2562634 (photos MEXU,
Hist., Bot. Ser. 17:214. 1937. Type: PTBG); Isotypes F, GH, n.v.).—Hoffmannia
GUATEMALA: Quezaltenango, Colomba, arborescens Donn. Sm.
720 m, 24 Sep 1934, A. F. Skutch 1291 Hoffmannia steyermarkii Standi., Publ. Field
(Holotype A, n.v. (photo and fragment F)). Mus. Nat. Hist., Bot. Ser. 22: 192. 1940.
—Hoffmannia regalis (Hook, f.) Hemsl. Type: GUATEMALA: San Marcos, between
Hoffmannia rosei B. L. Rob., Proc. Amer. Acad. Canjula and La Union, near southeast portion
Arts 45: 407. 1910. Type: MEXICO: Nayarit, of Volcan de Tacana, 2000-3000 m, 22 Feb
Tepic, near Pedro Paulo, 3 Aug 1897, J. N. 1940, J. A. Steyermark 36426 (Holotype F
Rose 1968 (Holotype US 300860 (photos 1035015 (photo PTBG); Isotype A, n.v.).
MEXU, PTBG); Isotype GH, n.v.). Hoffmannia strigillosa Hemsl., Diag. PI. Nov.
31. 1879. Type: MEXICO: Without precise
Hoffmannia rotata Donn. Sm., Bot. Gaz. locality, Salle s.n. (Holotype K, n.v.).
(Crawfordsville) 18: 204. 1893. Type: Hoffmannia subauriculata Standi., J. Wash.
GUATEMALA: Quiche, San Miguel Acad. Sci. 18: 179. 1928. Type: COSTA
Uspantan, 7000 ft., Apr 1892, Heyde & Lux RICA: Cartago, El Mufieco, on the Rio
3169B (Lectotype US 943496 (photos MEXU, Navarro, 1400-1500 m, 6-7 Mar 1926, P. C.
PTBG), here designated). Syntype: Standley & R. Torres R. 50956 (Holotype US
GUATEMALA: Quiche, San Miguel 1306554 (photos MEXU, PTBG)).
Uspantan, 7000 ft., Apr 1892, Heyde & Lux Hoffmannia teruae L. O. Williams. & A. R.
3174 (P (photos MEXU, PTBG)). The US Molina, Fieldiana, Bot. 36: 59. 1973. Type:
sheet of Heyde & Lux 3169B v GUATEMALA: San Marcos, near Aldea
Fraternidad, between San Rafael Pie de la
 Cuesta and Palo Gordo, west facing slopes of J. Valerio 57 (Holotype US 1206194 (photos
Sierra Madre Mountains, 1800-2400 m, MEXU, PTBG)).
10-18 Dec 1963, L. O. Williams, A. Molina R. Hoffmannia vesiculifera Standi, Publ. Field
& T P. Williams 26028 (Holotype F 1651553 Columbian Mus., Bot. Ser. 4: 285. 1929.
(photo PTBG); Isotype EAP, n.v.). Type: PANAMA: Bocas del Toro, Buena
Hoffmannia tetrastigma Dorm. Sm., Bot. Gaz. Vista Camp, Chiriqui Trail, 750 m, Jan-Mar
(Crawfordsville) 27: 336. 1899. Type: 1928, G. P. Cooper 231 (Holotype F 579284
GUATEMALA: Santa Rosa, Zamorora, 1800 (photo PTBG)).
m, Apr 1893, Heyde & Lux 4490 (Holotype Hoffmannia viridis Rusby, Descr. N. Sp. S.
US 354989 (photos MEXU, PTBG); Isotypes Amer. PI. 134. 1920. Type: COLOMBIA:
F, US, 2 sheets (photos MEXU, PTBG)). Santa Marta, H. H. Smith 2649 (Holotype NY,
Hoffmannia tonduzii Standi., Contr. U.S. Natl. n.v., photo and fragment F).
Herb. 20: 205. 1919. Type: COSTA RICA: Hoffmannia vulcanicola Standi. & Steyerm.,
Forets de Las Vueltas, Tucurrique, 635-700 Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 251.
m, May 1899, A. Tonduz 13373 (Holotype US 1947. Type: GUATEMALA: Solola, southern
941386 (photos MEXU, PTBG)). slopes of Volcan de Atitlan above Finca
—Hoffmannia longipetiolata Pol. Moca, 1000-1250 m, 20 Jun 1942, J. A.
Hoffmannia trichocalyx Standi., J. Wash. Steyermark 47931 (Holotype F 1132754
Acad. Sci. 18: 181. 1928. Type: COSTA (photo PTBG)).
RICA: Alajuela, vicinity of Fraijanes, Hoffmannia wilsonii Standi., Publ. Field Mus.
1500-1700 m, 12-13 Feb 1926, P. C Nat. Hist., Bot. Ser. 22: 117. 1940. Type:
Standley & R. Torres R. 47690 (Holotype US GUATEMALA: Alta Verapaz, Finca Los
1305242 (photos MEXU, PTBG); Isotype US Alpes, 1200 m, 21 Mar 1939, C. L. Wilson
1305243 (photos MEXU, PTBG)). 344 (Holotype F 968020 (photo PTBG)).
Hoffmannia tuerckheimii Donn. Sm., Bot. Hoffmannia witheringioides Dwyer, Ann.
Gaz. (Crawfordsville) 47: 254. 1909. var. Missouri Bot. Gard. 67: 244. 1980. Type:
tuerckheimii. Type: GUATEMALA: Alta PANAMA: Veraguas, Rio Segundo Braso, 8
Verapaz, bei Coban, 1600 m, Mar 1908, H. km beyond Escuela Agricola Alto Piedra
von Tuerckheim 11.2160 (Holotype US 941383 beyond Santa Fe, 750 m, 24 Jul 1974, T. B.
(photos MEXU, PTBG); Isotype US 941384 Croat 25551 (Holotype MO 2214610; Isotype
(photos MEXU, PTBG)). MO 2207374).
Hoffmannia tuerckheimii Donn. Sm. var. Hoffmannia woodsonii Standi., Ann. Missouri
glabra Standi. & Steyerm., Publ. Field Mus. Bot. Gard. 28: 471. 1941. Type: PANAMA:
Nat. Hist., Bot. Ser. 23: 250. 1947. Type: Chiriqui, vicinity of Callejon Seco, Volcan de
GUATEMALA: El Progreso, Sierra de las Chiriqui, 1700 m, 17 Jul 1940, R. E.
Minas, Montana Canahui, between Finca San Woodson, Jr. & R. W. Schery 503 (Holotype
Miguel and summit of mountain, upper limits MO 1210619 (photo PTBG); Isotypes F
of Finca Caieta, 1600-2300 m, 10 Feb 1942, 1085625 (photo PTBG), GH, n.v.).
J. A. Steyermark 43807 [43806 in protologue] —Hoffmannia longipetiolata Pol.
(Holotype F 1133774 (photo PTBG)).
Hoffmannia uniflora Standi., Contr. U.S. Natl Holtonia Standi., Trop. Woods. 30: 37. 1932.
Herb. 20: 204. 1919. Type: GUATEMALA: Type: Holtonia myriantha (Standi.) Standi.
Alta Verapaz, Coban, 1100 m, Feb 1908, H. Herein treated as a synonym of Elaeagia
von Tuerckheim 11.2107 (Holotype US 941382 Wedd.]
(photos MEXU, PTBG)). Holtonia myriantha (Standi.) Standi., Trop.
Hoffmannia valerii Standi., J. Wash. Acad Sci Woods 30: 37. 1932. Sickingia myriantha
18: 178. 1928. Type: COSTA RICA: Standi., Publ. Field Columbian Mus., Bot. Ser.
Guanacaste, El Arenal, 600 m, 20 Mar 1923 7: 27. 1930. Type: COLOMBIA: Magdalena:
 ^. NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Las Nubes road, region of Santa Marta, 1200 n.v.).—Hedyotis arenaria (Rose) W. H.
m, 3 Dec 1898, H. H. Smith 1810 (Holotype F
138660; Isotypes MO, NY, n.v., US, n.v., W, Houstonia asperuloides (Benth.) A. Gray, Proc.
n.v., Y, n.v). Simira.—Elaeagia myriantha Amer. Acad. Arts 5: 158. 1860. Hedyotis
(Standi.) C. M. Taylor & Hammel asperuloides Benth., Bot. Voy. Sulphur 19,/?/.
13. 1844. Type: MEXICO: Baja California,
Houstonia L., Sp. PI. 1: 101. 1753. Type: Cabo San Lucas, Barclay s.n. (Holotype BM,
Houstonia caerulea L. [Herein treated as a n.v.).—Hedyotis asperuloides Benth.
synonym of Hedyotis L., although some Houstonia asperuloides (Benth.) A. Gray var.
authors (e.g., Terrell, 1996) recognize brandegeana (Rose) Wiggins in Shreve &
Houstonia as a genus.] Wiggins, Veg. Fl. Sonoran Desert 2: 1399.
Houstonia acerosa (A. Gray) Benth. & Hook, f, 1964. Houstonia brandegeanna Rose, Contr.
Gen. PI. 2: 60. 1873. Hedyotis acerosa A. U.S. Natl. Herb. 1: 83. 1890. Type: MEXICO:
Gray, PI. Wright. 1: 81. 1852. Type: U.S.A.: Baja California, La Paz, 20 Jan-5 Feb 1890,
Texas, high prairies from Live Oak Creek to E. Palmer 31 (Holotype US 47259 (photos
Las Moras, C. Wright 237 (Holotype GH, MEXU, PTBG); Isotypes F, GH, n.v.).
n.v.). Oldenlandia.—Hedyotis acerosa A. —Hedyotis asperuloides Benth. f.
brandegeana (Rose) W. H. Lewis
Houstonia acerosa (A. Gray) Benth. & Hook. f. Houstonia bouvardioides (Seem.) Benth. &
subsp. polypremoides (A. Gray) Terrell, Hook, f, Gen. PI. 2: 60. 1873. Hedyotis
Brittonia31: 168. 1979. Houstonia bouvardioides Seem., Bot. Voy. Herald 2:
polypremoides A. Gray, Proc. Amer. Acad. 296. 1856. Type: MEXICO: Without precise
Arts 21: 379. 1886. Type: MEXICO: locality [Durango ?], Sierra Madre, Seemann
Chihuahua, Santa Eulalia Mountains, 26 Sep s.n. (Holotype BM, n.v.; Isotypes C, n.v.,
1885, C. G. Pringle 356 (Lectotype GH, fragment F (photo PTBG), GH, n.v.).
n.v.; Isolectotypes F, MO (photos MEXU, —Bouvardia bouvardioides (Seem.) Standi.
PTBG), US (photos MEXU, PTBG); Houstonia brandegeana Rose, Contr. U.S. Natl.
designated by Lewis, 1968).—Hedyotis Herb. 1: 83. 1890. Type: MEXICO: Baja
acerosa A. Gray var. polypremoides (A. California, La Paz, 20 Jan-5 Feb 1889, E.
Gray) W. H. Lewis Palmer 31 (Holotype US 47259 (photos
Houstonia angustifolia Michx., Fl. Bor. Amer. MEXU, PTBG); Isotypes F, GH, n.v.).
1: 85. 1803. Type: U.S.A.: Florida, in —Hedyotis asperuloides Benth. f.
submaritimus Floridae, collector not cited brandegeana W. H. Lewis
(Holotype presumably P, n.v.).—Hedyotis Houstonia brevipes Rose, Contr. U.S. Natl.
nigricans (Lam.) Fosberg Herb. 1: 83. 1890. Type: MEXICO: Baja
Houstonia angustifolia Michx. var. scabra S. California, Santa Rosalia, 24 Feb-3 Mar 1889,
Watson, Proc. Amer. Acad. Arts 18: 97. 1883. E. Palmer 202 (Holotype US 47258 (photos
Type: MEXICO: Coahuila, Saltillo, Caracol MEXU, PTBG); Isotypes F, GH, n.v., US
Mountains, 1-15 Apr 1880, E. Palmer 410 (photos MEXU, PTBG)).—Hedyotis
(Holotype GH, n.v.; Isotype US (photos brevipes (Rose) W. H. Lewis
MEXU, PTBG)).—Hedyotis nigricans Houstonia coccinea Andr., Bot. Repos.pl. 106.
(Lam.) Fosberg 1800. Type: No herbarium specimens are
Houstonia arenaria Rose, Contr. U.S. Natl. known to exist, only the drawing pi. 106.
Herb. 1: 70. 1890. Type: MEXICO: Baja —Bouvardia ternifolia (Cav.) Schltdl.
California, La Paz, 20 Jan-5 Feb 1890, E. Houstonia drymarioides Standi., J. Wash. Acad.
Palmer 28 (Holotype US 47260 (photos Sci. 18: 162. 1928. Type: MEXICO:
MEXU, PTBG); Isotypes F, GH, Tamaulipas, uthofVic
1000 m, 9 Apr 1926, R. Runyon 870
 (Holotype US 1315865 (photos MEXU, slopes, mountains along route 175, 28 km by
PTBG); Isotype F) —Hedyotis c road south of Valle Nacional on way to Ixtlan
(Standi.) W. H. Lewis de Juarez, ca. 2000 m, 30 Jul 1959, R. M.
Houstonia fasciculata A. Gray, Proc. Amer. King 2107 (Holotype MICH, n.v.; Isotype
Acad. Arts 17: 203. 1882. Syntypes: U.S.A.: MICH, n.v.).—Hedyotis kingii (Terrell) G. L.
Texas, Presidio, Bigelow s.n. (GH); New
Mexico, Organ Mountains, G. R. Vasey s.n. Houstonia latifolia Willd. ex Roem. & Schult.,
(GH, n.v.); MEXICO: Coahuila, near Parras Syst. 527. 1808. Not Hedyotis latifolia (M.
and Monclova, E. Palmer 404 (GH, n.v.), E. Martens & Galeotti) Walp., 1846. Type:
Palmer 406 (GH, n.v.).—Hedyotis intricata America borealis, Muhlenberg s.n.
(presumably B-WI, n.v.).—Hedyotis exigula
Houstonia fruticosa Willd. ex Roem. & Schult., W. H. Lewis
Syst. 3: 527. 1818. Type: VENEZUELA: Houstonia longipes S. Watson, Proc. Amer.
Monagas, Caripe, Humboldt s.n. (B, Acad. Arts 18: 97. 1883. Syntypes: MEXICO:
presumably destroyed).—Declieuxia Without precise locality, States of Coahuila
fruticosa (Willd. ex Roem. & Schult.) Kuntze and Nuevo Leon, Feb-Oct 1880, E. Palmer
Houstonia fruticosa Rose, Contr. U.S. Natl. 394 (GH, n.v.; US 47845 (photos MEXU,
Herb. 1: 132. 1892. Not Hedyotis fruticosa L., PTBG)); Nuevo Leon, Monterrey, Feb 1-26
1753, nor Willd. ex Roem. & Schult., 1818. 1880, E Palmer 395 (GH, n.v., US 70739
Type: MEXICO: Baja California, Carmen (photos MEXU, PTBG), US 1390542 (photos
Island (Isla Carmen), 1-7 Nov 1890, E. MEXU, PTBG)).—Hedyotis palmeri (A.
Palmer 885 (Holotype US 47256 (photos Gray) W. H. Lewis
MEXU, PTBG); Isotypes MO (photos Houstonia mucronata (Benth.) B. L. Rob., Proc.
MEXU, PTBG), US (photos MEXU, PTBG)). Amer. Acad. Arts 45: 401. 1910. Hedyotis
—Hedyotis mucronata Benth. mucronata Benth., Bot. Voy. Sulphur 19.
Houstonia gracilenta I. M. Johnst., Proc. Calif. 1844. Type: MEXICO: Baja California,
Acad. IV, 12: 1174. 1924. Type: MEXICO: Magdalena Bay, Barclay s.n. (Holotype BM,
Baja California, San Diego Island, rocky n.v.).—Hedyotis mucronata Benth.
island slope facing sea, 2 May 1921,1. M. Houstonia mullerae (Fosberg) Terrell, Brittonia
Johnston 3927 (Holotype UC, n.v.; Isotypes 31: 169. 1979. Hedyotis mullerae Fosberg,
GH, n.v., US (photos MEXU, PTBG)). Lloydia4: 288. 1941. Type: MEXICO:
—Hedyotis gracilenta (I. M. Johnst.) W. H. Coahuila, Mun. de Cuatro Cienegas, Canon
del Agua, Sierra de la Madera, 9 Sep 1939,
Houstonia gracilis Brandegee, Zoe 5: 238. C. H. Muller 3234 (Holotype US 2133305 ex
1907. Not Hedyotis gracilis DC, 1830. Type: NA (photos MEXU, PTBG); Isotype GH,
MEXICO: Veracruz, Orizaba, May 1904, n.v).—Hedyotis mullerae Fosberg
C. A. Purpus 1251 (Holotype UC, n.v.; Houstonia nigricans (Lam.) Fernald, Rhodora
Isotypes F, MO (photos MEXU, PTBG)). 42: 299. 1940. Gentiana nigricans Lam.,
—Hedyotis exigula W. H. Lewis Encycl. 2: 645. 1786. Type: MEXICO:
Houstonia humifusa (A. Gray) A. Gray, Proc. Collector unknown, (presumably P-LAM,
Amer. Acad. Arts 4: 314. 1859. Hedyotis n.v.).—Hedyotis nigricans (Lam.) Fosberg
humifusa A. Gray, Boston J. Nat. Hist. 6- 216 Houstonia palmeri A. Gray, Proc. Amer. Acad.
1850. Type: U.S.A.: Texas, near Arts 17: 202. 1882. Syntypes: MEXICO:
Fredrickburg, 1847, F. Lindheimer 621 Coahuila, Sierra Madre, 40 mi. south of
(Holotype GH, n.v.; Isotypes BM, n.v., K, Saltillo, 25-31 Jul 1880, E. Palmer 397 (GH,
n.v., MICH, n.v., MO, PH, n.v., US). ' n.v., US 47431 (photos MEXU, PTBG); E.
Houstonia kingii Terrell, Brittonia 32: 491, fig Palmer 398 (GH, n.v., F (2 sheets), US 70740
2. 1980. Type: MEXICO: Oaxaca, E-facing
 i NOMENCLATOR OF MEXICAN AND CENTRAL ;

(photos MEXU, PTBG)).—Hedyotis palmeri Brandegee s.n. (Holotype UC, n.v.; Isotypes
(A. Gray) W. H. Lewis GH, n.v., US (photos MEXU,
Houstonia parvula Brandegee, Zoe' 5: 221. PTBG)).—Hedyotis sinaloae W. H. Lewis
1905. Not Hedyotis parvula (A. Gray) Houstonia rubra Cav., Icon. 5: 48, /. 474, fig. I.
Fosberg, 1943. Type: MEXICO: Sinaloa, 1799. Type: MEXICO: Hidalgo, Ixmiquilpan,
vicinity of Culiacan, 12 Oct 1904, T. S. Nee s.n. (Holotype presumably MA, n.v.).
Brandegee s.n. (Holotype UC, n.v.; Isotypes —Hedyotis rubra (Cav.) A. Gray
GH, n.v., US (photos MEXU, PTBG)). Houstonia serpyllacea (Schltdl.) C. L. Sm. ex
—Hedyotis sinaloae W. H. Lewis Greenman, Proc. Amer. Acad. Arts 32: 284.
Houstonia peninsularis Brandegee, Zoe 5: 160. 1897. Hedyotis serpyllacea Schltdl., Linnaea
1903. Type: MEXICO: Baja California, Cape 9: 599. 1834. Type: MEXICO: Mexico, inter
Region, Sierra de la Trinidad, Nov 1902, T. S. La Joya et San Salvador, Jun 1829, Schiede &
Brandegee s.n. (Holotype UC, n.v.; Isotypes Deppe 265 (Holotype presumably HAL, n.v.;
GH, n.v., US (photos MEXU, PTBG)). Isotypes B (photo MO), F (photo PTBG),
—Hedyotis peninsularis (Brandegee) W. H. MO).—Hedyotis serpyllacea Schltdl.
Houstonia sharpii Terrell, Brittonia 32: 490, fig.
Houstonia polypremoides A. Gray, Proc. Amer. I. 1980. Type: MEXICO: Hidalgo, camino
Acad. Arts 21: 379. 1886. Type: MEXICO: entre Metepec y Tenango de Doria, entre 16 y
Chihuahua, Santa Eulalia Mountains, 26 Sep 31.5 km al NNE de Tulancingo, 18 km N.NE.
1885, C. G Pringle 356 (Lectotype GH; n.v.; de Metepec, 29 Mar 1974, S. D. Koch 7423
Isolectotypes F, MO (photos MEXU, PTBG), (Holotype US 2754863 (photos MEXU,
US (photos MEXU, PTBG); designated by PTBG); Isotypes CHAPA, ENCB, MEXU).
Lewis, 1968). Syntype: MEXICO: Chihuahua, —Hedyotis sharpii (Terrell) G. L. Nesom
Santa Eulalia Mountains, 29 May 1885, C. G Houstonia teretifolia Terrell, Brittonia 31: 166.
Pringle 16 (GH, n.v., US (photos MEXU, 1979. Type: MEXICO: Coahuila, 12 km NNE
PRBG)).—Hedyotis polypremoides (A. of Las Margaritas on easternmost ridge of
Gray) Shinners Sierra de las Margaritas, 1300-1400 m, 24
Houstonia polypremoides var. bigelovii Sep 1972, M. C. Johnston, F. Chiang & T.
Greenman, Proc. Amer. Acad. Arts 32: 291. Wendt 9508E (Holotype TEX, n.v.).
1897. Type: U.S.A.: Florence Mountains, Jun —Hedyotis teretifolia (Terrell) G. L. Nesom
1852, Bigelow 437 (Holotype GH, n.v.). Houstonia triflora (Kunth) A. Gray, Proc. Amer.
—Hedyotis polypremoides (A. Gray) W. H. Acad. Arts 4: 314. 1860. Bouvardia triflora
Kunth in Humb., Bonpl. & Kunth, Nov. Gen.
Houstonia prostrata Brandegee, Zoe 5: 105. Sp. 3: 386. 1820. Type: MEXICO: Without
1901. Not Hedyotis prostrata Blume, 1823 precise locality, Humboldt & Bonplands.n.
norKorth., 1851. Type: MEXICO: Baja (Holotype P-Bonpland, n.v. (photo MO)).
California, La Palma, Cape Region, 25 Sep —Bouvardia multiflora (Cav.) Schult. &
1899, T. S. Brandegee s.n. (Holotype UC, Schult. f.
n.v.; Isotypes GH, n.v., US 382879 (photos Houstonia umbratilis B. L. Rob., Proc. Amer.
MEXU, PTBG), US 735791 (photos MEXU, Acad. Arts 45: 401. 1910. Type: MEXICO:
PTBG)).—Hedyotis vergrandis W. H. Lewis Nuevo Leon, shaded cliffs of mountains, near
Houstonia prostrata Brandegee var. parvula Monterrey, 25 Apr 1906, C. G. Pringle 13877
(Brandegee) Wiggins in Shreve & Wiggins, (Holotype GH, n.v.; Isotypes MO (photos
Veg. Fl. Sonoran Desert 2: 1399. 1964. MEXU, PTBG), US (photos MEXU, PTBG)).
Houstonia parvula Brandegee, Zoe 5: 221. —Hedyotis umbratilis (B. L. Rob.) W. H.
1905. Not Hedyotis parvula (A. Gray)
Fosberg, 1943. Type: MEXICO: Sinaloa, Houstonia wrightii A. Gray, Proc. Amer. Acad.
vicinity of Culiacan, 12 Oct 1904, T. S. Arts 17: 202. 1882. Type: MEXICO: Nuevo
 Leon, shaded cliffs of mountains, near H Pittier 3889 (Holotype US 678963 (photo
Monterrey, 25 Apr 1906, C. G. Pringle 13877 PTBG); Isotype US (photo PTBG)).—Isertia
(Holotype GH, n.v.; Isotypes MO (photos laevis (Triana) B. M. Boom
MEXU, PTBG), US (photos MEXU, PTBG)). Isertia scorpioides B. M. Boom, Brittonia 36:
—Hedyotis wrightii (A Gray) Fosberg 435. 1984. Type: PANAMA: Colon, Atlantic
Houstonia xestosperma (B. L. Rob. & side of Isthmus on road to Pifia, 25 Aug 1967,
Greenman) Terrell, Brittonia 32: 492. 1980. J. H Kirkbride, Jr. & S. M. V. Hayden 316
Oldenlandia xestosperma B. L. Rob. & (Holotype NY, n.v.; Isotypes MO (2 sheets;
Greenman, Proc. Amer. Acad. Arts 32: 41. photos PTBG), NY, n.v.).
1896. Type: MEXICO: Oaxaca, Sierra de San
Felipe, open glades, 8000 ft., Jun 1894, C. G. Ixora L., Sp. PI. 1: 110. 1753. Type: Ixora
Pringle 4692 (Lectotype GH, n.v.; coccinea L.
Isolectotypes BM, n.v., F, K, n.v., MO Ixora alba Sesse & Mo?., La Naturaleza, ser. 2,
(photos MEXU, PTBG), MSC, n.v., PH, n.v., 1. app. 16. 1887. Not Ixora alba L., 1753, nor
UC, US, 2 sheets (photos MEXU, PTBG), Roxb., 1814, nor Sus. ex Wight & Arm, 1834.
VT, n.v.; designated by Terrell, 1980). Type: MEXICO: Habitat in India ad montibus
Hedyotis.—Bouvardia xestosperma (B. L. Mazatlan (presumably MA, n.v.).—Status
Rob. & Greenman) Terrell & S. D. Koch
Uacq., Hort. Schonbr. 3:4,/?/.
Isertia Schreb., Gen. PI. 1: 234, n. 602. 1789. 257. 1798. Type: Cultivated at Schobrunn,
Type: Isertia coccinea (Aubl.) Vahl Vienna, said to have been brought from
unii Bartlett, Proc. Amer. Acad. Arts Mexico.—Bouvardia ternifolia (Cav.)
43: 59. 1907. Type: GUATEMALA: Izabal, Schltdl.
Puerto Barrios, 24 Feb 1905, C. C. Deam 48 Ixora aphylla Sesse & Mo?., Pi. Nuev. Hispan.
(Holotype GH, n.v.).—Isertia haenkeana 16. 1887; La Naturaleza, ser. 2, 1. app. 16, ed.
DC. 2, 15. 1887. Type: MEXICO: Habitat prope
Isertia deamii Bartlett var. stenophylla Dorm. Xochistlan, Sesse & Mociiio s.n. (presumably
Sm., Bot. Gaz. (Crawfordsville) 61: 374. MA, n.v.).—Status uncertain.
1916. Type: COSTA RICA: Boca Culebra, Ixora casei Hance in Walp., Ann. Bot. Syst. 2:
plaines des Pacifique, 50 m, 20 Jan 1898, H. 754. 1852. Type: CAROLINE ISLANDS:
Pit tier 11989 (Holotype US 941413 (photo Strong Island, Q. W. Case s.n. (Isotype or
PTBG); Isotypes F (photo PTBG), P, n.v., possible Holotype K, n.v.). For typification
fragment F).—Isertia haenkeana DC. see Fosberg & Sachet (1989a), who note that
Isertia haenkeana DC, Prodr. 4: 437. 1830. var. casei is cultivated in Mesoamerica.
Type: MEXICO: Without precise locality, Ixora coccinea L., Sp. PI. 1: 110. 1753.
Haenke s.n. (Holotype presumably PR, n.v.). Lectotype: Rheede, Hortus malabaricus 2:17,
Isertia laevis (Triana) B. M. Boom, Brittonia /. 13. 1679. Designated by Fosberg & Sachet
36: 433. 1984. Cassupa laevis Triana, Ann. (1989b), who note that Ixora coccinea L. var.
Sci. Nat, Bot., Ser 4, 9: 44. 1858. Type: coccinea and var. intermedia Fosberg &
COLOMBIA: Cundinamarca, Triana 1838 Sachet are cultivated in Mexico and Central
(Lectotype P; Isolectotypes BR, n.v., COL,
n.v., L, n.v., P, n.v., US, n.v.; designated by Ixora coccinea L. var. intermedia Fosberg &
Boom, 1984). Sachet, Baileya 23: 80. 1989. Type: VIRGIN
Isertia panamensis (Standi.) Standi., Publ. Field ISLANDS: St. Croix, Teague Bay, cultivated,
Columbian Mus., Bot. Ser. 8: 346. 1931. F. R., Fosberg 55279 (Holotype US, n.v.)
Cassupa panamensis Standi., Contr. U.S. —Ixora coccinea L.
Natl. Herb. 18: 135. 1916. Type: PANAMA: Ixora finlaysoniana Wall, ex G. Don, Gen.
Colon, along Rio Fato, 10-100 m, 8 Jul 1911, Hist. Dichlam. PI. 3: 572. 1834. Type: EAST
 i NOMENCLATOR OF MEXICAN AND CENTRAL

INDIES: Wallich Catalog 6166 (Holotype K, Komlodi. [Herein treated as a synonym of

n.v. (microfiche MO)). Cultivated in Mexico Rondeletia L.]
and Central America. Javorkaea hondurensis (Donn. Sm.) Borhidi &
Ixora floribunda (A. Rich.) Griseb., Cat. PI. J. Komlodi, Acta Bot. Hung. 29: 16. 1983.
Cub. 134. 1866. Siderodendron floribundum
A. Rich, in Sagra, Hist. Cuba 11: 24. 1850.
Type: CUBA: Without precise locality, A. HONDURAS: Santa Barbara, Rio
Richard s.n. (Holotype presumably P, n.v.). Chamelecon, 300 m, Dec 1888, C. Thieme
Ixora nicaraguensis Wernham, J. Bot. 50: 243. 5267 (Holotype US 943502 (photos MEXU,
1912. Type: NICARAGUA: Chontales, PTBG); Isotypes F (photos MEXU, PTBG),
Seemann 117 (Holotype BM, n.v.). US (2 sheets, photos MEXU, PTBG)).
Ixora occidentalis L., Syst., ed. 10, 2: 893.
1759. Type: JAMAICA: Based on P. Browne,
Hist. Jam. t. 6, fig. 2 (LINN, n.v.), and Joosia H. Karst., FL Columb. 1: 9, tab. 5. 1858.
Plumier, Icon. tab. 156, f. 2 (ed. Burmannus). Type: Joosia umbellifera H. Karst.
—Faramea occidentalis (L.) A. Rich. Joosia panamensis Dwyer, Ann. Missouri Bot.
Ixora pulcherrima Sesse & Moc., La Naturaleza, Gard. 67: 262. 1980. Type: PANAMA:
ser. 2, 1. app. 16. 1887. Type: MEXICO: Veraguas, road between Escuela Agricola
Habitat in montibus Mazatlan, Sesse & Alto de Piedra above Santa Fe and Rio Dos
Mocino s.n. (presumably MA, n.v.).—Status bocas, 730-770 m, 26 Jul 1974, T. B. Croat
uncertain. 25932 (Holotype MO 2248047 (photo
Ixora rauwolfwides Standi., Trop. Woods 11: PTBG)).—Joosia umbellifera H. Karst.
25. 1927. Type: PANAMA: Changuinola Joosia umbellifera H. Karst., Fl. Columb. 1: 9,
Valley, 1927, G P. Cooper & G. M. Slater 93 tab. 5. 1858. Type: COLOMBIA: Meta,
[Yale 10274] (Holotype US 1315888 (photo Servita, Karsten & Triana s.n. (Lectotype W
PTBG); Isotype F).—Ixora nicaraguensis 134852, n.v.; Isolectotype NY, n. v.;
Wernham designated by Andersson in Brittonia 49: 32.
Ixora spinosa (Jacq.) Lam., Encycl. Meth. Bot. 1997).
3: 344. 1789. Chomelia spinosa Jacq., Enum.
PL Carib. 12. 1760. Type: COLOMBIA: Near Kohleria Regel. [Ge<
Cartagena, Jacquin s.n. (location unknown). Kohleriafdisepala Standi., Publ. Field Mus.
—Chomelia spinosa Jacq. Nat. Hist., Bot. Ser. 22: 107. 1940. Type:
Ixora ternifolia Cav., Icon. 4: 3, t. 305. 1797. MEXICO: Chiapas, Mt. Ovando, 16 Dec
Type: Based on cultivated material grown at 1936, E. Matuda 574 (Holotype F 916483
Madrid from seed collected in Mexico (photo PTBG); Isotypes MEXU, US).
(Lectotype is tab. 305; designated by —Omiltemia filisepala (Standi.) C. V.
Blackwell, 1968).—Bouvardia ternifolia
(Cav.) Schltdl.
Ixora uniflora Sesse & Moc., La Naturaleza, ser. Ladenbergia Klotzsch in Hayne, Getreue
2, 1. app. 16. 1887. Type: MEXICO: Habitat Darstell. Gew. 14: pi. 15. 1846. Lectotype:
Michaelapoli, Sesse & Mocino s.n. Ladenbergia moritziana Klotzsch (Designated
(presumably MA, n.v.; Fl. Mex. Ic. 26 (G- by Andersson & Taylor, 1994: 17.)
DEL, n.v.).—Status i Ladenbergia brenesii Standi., Publ. Field Mus.
Nat. Hist., Bot. Ser. 18: 1323. 1938. Type:
Javorkaea Bohridi & J. Komlodi, Acta B< COSTA RICA: San Ramon, San Miguel, La
Hung. 29: 16. 1983. Type: Javorkaea Palma, San Pedro, 20-25 Mar 1932, A. M.
hondurensis (Donn. Sm.) Borhidi & J. Brenes 15146 (Holotype F 856805 (photo
PTBG)).
Ladenbergia bullata (Wedd.) Standi., Trop. Cerro Jefe, 25 Jul 1968, R. L. Dressier 3560
Woods 34: 41. 1933. Cascarilla bullata (Holotype MO 2267568 (photo PTBG);
Wedd., Ann. Sci. Nat., Bot., ser. 3, 10: 11. Isotypes MEXU, US (photos MEXU, PTBG)).
1848. Type: PERU: Puuo, Prov. de Carabaya,
Jan-Feb 1847, Weddell 4358 (Holotype P-JU, Lasionema D. Don, Lond. & Edinb. Phil. Mag.
n.v.; Isotypes G, n.v., K, n.v.). Note: Although 2: 377. 1833. Type: Lasionema roseum (Ruiz
Dwyer (1980) erroneously recorded this & Pav.) D. Don (Cinchona rosea Ruiz &
species from Panama, it is endemic to the Pav.). (Here treated as a synonym of
sourthern Andes (L. Andersson, pers. comm. Macrocnemum R. Br.)
1996). Lasionema glabrescens Benth., Bot. Voy.
Ladenbergia laurifolia Dwyer, Ann. Missouri Sulphur 105. 1845. Type: COSTA RICA:
Bot. Gard. 67: 265. 1980. Type: PANAMA: Lagartos Bay (Carno Bay), Gulf of Nicoya,
Veraguas, beyond Rio Tres Brazos, 9 km 12-16 Feb 1839, G. W. Barclay 2780
from Santa Fe, T. B. Croat 25586 (Holotype (Holotype BM, n.v.; Isotype F (photo
MO, n.v.). PTBG)). Macrocnemum.—Macrocnemum
Ladenbergia macrocarpa (Vahl) Klotzsch in roseum (Ruiz & Pav.) Wedd.
Hayne, Getreue Darstell. Gew. 14: tab. 15.
1846. Cinchona macrocarpa Vahl, Schr. Laugeria Vahl ex Hook. f. in Benth. & Hook, f,
naturh. Ges. Kopenhagen 1(1): 20. 1793. Gen. 2: 101. 1873. Not Laugeria L., 1764.
Type: COLOMBIA: In regno Santa Fe, Lectotype: Laugeria resinosa Vahl (Laugeria
Ortega s.n. (Lectotype BM, n.v.; designated Vahl, Eclog. 1: 26. 1796, lacked a generic
by Andersson in Andersson & Taylor, 1994: description. For discussion see Little, 1959:
16). 27.)
Laugeria lucida Sw., Prodr. Veg. Ind. Occ. 48.
1788. Type: JAMAICA: Without precise
Type: COSTA RICA: Two miles southeast of localiity, Swartz s.n. (presumably S, n.v.).
San Isidro del General, Aug 1936, R. E. Antirhea, Stenostomum.—Stenostomum
Danforth 32 (Holotype F; Isotype GH (photo lucidum (Sw.) C. F. Gaertn.
PTBG)).
Ladenbergia valerii Standi., Publ. Field Mus. Laugieria Jacq., Enum. PI. Carib. 2: 16. 1760.
Nat. Hist, Bot. Ser. 18: 1324. 1938. Type: Type: Laugieria odorata Jacq. [Herein treated
COSTA RICA: La Paz de San Ramon, 1225 as a synonym of Guettarda L.]
m, Jun 1925, Brenes 4296 (Holotype F). Laugieria odorata Jacq., Enum. PI. Carib. 2: 16.
1760. Type: CUBA: Ex Carthagena et
Lagenanthus Gilg [Gentianaceae] Havana, Jacquin s.n. (Lectotype P, n.v.;
Lagenanthus pannflorus Ewan, Mutisia 4: 5. designated by Steyermark, 1972: 360).
1952. Type: PANAMA: Code, region north —Guettarda odorata (Jacq.) Lam.
of El Valle of Anton, 1000 m, P. H. Allen
3601 (Holotype MO, n.v.).—Hillia triflora Limnosipanea Hook. f. in Hook., Icon. PI. 11:
var. pittieri (Standi.) C. M. Taylor 38, t. 1050. 1868. Type: Limnosipanea
spruceana Hook. f.
Lasianthus Jack, Trans. Linn. Soc. 14: 125. Limnosipanea palustris (Seem.) Hook. f. in
1823. Type: Lasianthus cyanocarpus Jack Hook., Icon. PI. 11: 38, t. 1050. 1868. Sipanea
Lasianthus panamensis (Dwyer) Robbr., Ann. palustris Seem., Bot. Voy. Herald 136. 1853.
Missouri Bot. Gard. 69: 427. 1982. Type: PANAMA: In a swamp near Panama
Dressleriopsis panamensis Dwyer, Ann. City, Seemann 347 (Holotype BM, in part
Missouri Bot. Gard. 67: 154. 1980. Type: [excluding specimen of Microlicia,
PANAMA: Panama, i r La Eneida, beyond
 INOMENCLATOR OF MEXICAN AND CENTRAL

Lindenia Benth., PL Hartw. 84. 1841. Type: Loranthus Jacq. [Loranthaceae]

Lindenia rivalis Benth. (See Darwin (1976: Loranthus guadalupensis DC, Prodr. 4: 294.
427) and Farr et al. (1979: 1620) for 1830. Type: GUADALOUPE: Without
s regarding publication of the precise locality, Bertero s.n. (Holotype G-DC,
: names Lindenia vs. Siphonia.) [Herein n.v. (microfiche IDC 800.666.II.2)).
as a synonym of Augusta Pohl. See —Psychotria guadalupensis (DC.) Howard
jitfta by Kirkbride, 1997.]
Lindenia acutiflora Benth. in Hooker, Icon. PL Lycium L. [Solanaceae]
5: tab. 475. 1842. Type: MEXICO: Veracruz, Lycium japonicum Thunb., Nov. Gen. PL 132,
Puente Nacional, 1838, Linden 358 (Holotype pi. 9. 1798. Type: JAPAN: Collector and
K, n.v. (photo MO)).—Augusta rivalis locality unknown (presumably UPS, n.v.).
(Benth.) J. H. Kirkbr. —Serissa japonica (Thunb.) Thunb.
Lindenia radiums Wernham, J. Bot. 52. 226.
1914. Type: MEXICO: Oaxaca, shaded places Lygodysodea Ruiz & Pav., Fl. Peruv. Chil. 32, i
on rocks at crossing of the river near 5. 1794. Type: Lygodysodea fetida Ruiz &
Tocotepeque, Jul 1839, T. Hartwegs.n. Pav. [Herein treated as a synonym of
(Holotype K, n.v. (photo MO)). Paederia L.]
—Habroneuron radicans (Wernham) S. P. Lvgodvsodea ciliata Bartl. ex DC, Prodr. 4:
Darwin 470. 1830. Type: MEXICO: Ad Acahuitzola,
Lindenia rivalis Benth., PL Hartweg. 84. 1841. Haenke s.n. (Holotype presumably PR, n.v.;
Type: GUATEMALA: In nvis, la Vera Paz, T. not located by Puff, 1991).—Paederia ciliata
Hartweg 581 (Holotype K, n.v. (photo MO)). (Bartl. ex DC.) Standi.
Siphonia rivalis Benth., PL Hartweg. 84.
1841. Note: According to Farr et al. (1979:
1620) the name Siphonia was actually printed,
but was crossed out in ink and the name
Lindenia substituted in manuscript in all Machaonia acuminata Humb. & BonpL, PL
copies before distribution. The correction is Aequm. 1:101, A 29. 1806. Type:
printed on p. 351.—Augusta rivalis (Benth.) ECUADOR: In ipsa urbe Guyaquil, A.
J. H. Kirkbr. Bonpland3828 (Holotype P-Bonpland, n.v.;
Lindenia rivalis Benth. f. glabra S. P. Darwin, J. Isotype P (photos MEXU, PTBG)).
Arnold Arbor. 57: 439. 1976. Type: Machaonia coulteri (Hook, f.) Standi., Contr.
MEXICO: Chiapas, Chicomucelo [as U.S. Natl. Herb. 20: 209. 1919.
"Chicomuselo"], 800 m, 14 Jul 1941, E. Microsplenium coulteri Hook. f. in Benth. &
Matuda 4449 (Holotype US 2087017 (photos Hook, f, Gen. PL 2: 4. 1873. Type:
MEXU, PTBG); Isotypes A, n.v., F, MO MEXICO: Hidalgo: Zimapan, Coulter s.n.
(photos MEXU, PTBG), NY, n.v.). (Holotype K, n.v.).
—Lindenia rivalis Benth. Machaoniafasciculata A. Gray, Proc. Amer.
Acad. Arts 19: 77. 1883. Type: MEXICO:
Lonicera L. [Caprifoliaceae] Without precise locality, Coulter 1167
Lonicera alba L., Sp. PL 175. 1753. Type: (Holotype GH, n.v.).—Machaonia coulteri
JAMAICA or BARBADOS: Material (Hook, f.) Standi.
cultivated in the Clifford garden (possibly Machaonia floribunda Greenm., Proc. Amer.
BM, n.v.). According to Howard (1989) a Acad. Arts 33: 488. 1898. Type: MEXICO:
.pecii be. Tamaulipas, hills, Tampico, 16 Jun 1897,
 C. G Pringle 6641 (Holotype GH, n.v.; Machaonia pringlei A. Gray, Proc. Amer.
Isotypes A, n.v., MO, P (2 sheets; photos Acad. Arts 21: 380. 1886. Type: MEXICO:
MEXU, PTBG), US (photos MEXU, PTBG)). Coahuila, limestone ledges, mountains,
—Machaonia acuminata Humb. & Bonpl. Jimulco, 27 Apr 1885, C. G. Pringle 146
Machaonia galeottiana Baill., Bull. Mens. Soc. (Holotype GH, n.v.; Isotypes A, n.v., F
Linn. Paris 1: 204. 1879. Type: CUBA: Le (photos MEXU, PTBG), P (photos MEXU,
Havane, H. Galeottis.n. (Holotype P (photo PTBG), US (photos MEXU, PTBG)).
MO)). On the holotype label the word Machaonia rotundata Griseb. var. dodgei
"Veracruz" is crossed out and "Le Havane" Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.
written in. As noted by Standley (1934: 222) 18: 1326. 1938. Type: COSTA RICA: Beach
the collection resembles no Mexican species. between Rio Sandoval and River Tigre, 1 May
—Machaonia havanensis (Jacq.) Alain 1930, C. W. Dodge 7752 (Holotype F 890888
Machaonia hahniana Baill., Bull. Mens. Soc. (photo PTBG); Isotype MO (photo PTBG)).
Linn. Paris 1: 204. 1879. Type: MEXICO: —Machaonia martinicensis (DC.) Standi.
Puebla, Acatlan, Juillet 1865, L Hahn s.n. Machaonia tysonii Dwyer, Ann. Missouri Bot.
(Holotype P (photos MEXU, MO, PTBG)). Gard. 53: 106. 1966. Type: PANAMA. Los
Machaonia havanensis (Jacq. ex J. F. Gmel.) Santos, thicket near man-made dam, main
Alain, Phytologia 8: 370. 1962. Spermacoce highway, Monagre Beach, J. D. Dwyer 5067
havanensis Jacq. in J. F. Gmel., Syst. Nat. 2: (MO, not located in 6/1996). Note: John
234. 1791. Type: Based on Spermacoce Dwyer (pers. coram. 24 March 1993) believes
fruticosa Jacq., ined.? (type not located). citation of this collection was a typographical
Machaonia lindeniana Baill., Bull. Mens. Soc. error and the following collection actually
Linn. Paris 1: 204. 1879. Type: MEXICO: represents the holotype: PANAMA: Along
Campeche, Campeche, fl. en Juin, (Yucatan), Quebrada Vital off Rio Chucanagre, km 16 de
Linden s.n. (Holotype P (photos MEXU, MO, Yavita, 4 Jul 1962, J. A. Duke 5067 (Holotype
PTBG)). MO 3725554 (photo PTBG)). In the Flora of
Machaonia martinezorum Lorence, Novon 4: Panama Rubiaceae treatment, Dwyer (1980)
123, fig. 3. 1994. Type: MEXICO: Chiapas, erroneously cited Dwyer 5061 as the holotype;
Mun. de Ocosingo, a 50 km al S de Boca the latter specimen was not located at MO in
Lacantum, 130 m, 24 Jun 1986, E. Martinez S. 3/1993 or 6/1996—Machanonia acuminata
19016 (Holotype PTBG 7356; Isotypes Humb. & Bonpl.
MEXU, MO). Machaonia velutina M. Martens & Galeotti,
Machaonia martinicensis (DC.) Standi., Publ. Bull. Acad. Roy. Sci. Bruxelles 11: 134. 1844.
Field Mus. Nat. Hist., Bot. Ser. 22: 193. 1940. Type: MEXICO: Veracruz, ravines de Puente
Tertrea martinicensis DC, Prodr. 4: 481. Nacional, pres Veracruz, H. Galeotti 7104
1830. Type: MARTINIQUE: Richard s.n. (Holotype BR, n.v.; Isotype P (photos MEXU,
(Holotype P, n.v.). MO, PTBG)).—Machaonia acuminata
Machaonia martinicensis (DC.) Standi, var. Humb. & Bonpl.
dodgei (Standi.) Standi., Publ. Field Mus. Nat. Machaonia veracruzeana Baill., Bull. Mens.
Hist., Bot. Ser. 22: 193. 1940. Machaonia Soc. Linn. Paris 1: 204. 1879. Norn, superfl.,
rotundata Griseb. var. dodgei Standi., Publ. based on Machaonia velutina M. Martens &
Field Mus. Nat. Hist., Bot. Ser. 18: 1326. Galeotti, 1844— Machaonia acuminata
1938. Type: COSTA RICA: Beach between Humb. & Bonpl.
Rio Sandio and Rio Tigre, 1 May 1930, C. W.
Dodge 7752 (Holotype F 890888 (photo Macrocnemum P. Browne, Hist. Jam. 165.
PTBG); Isotype MO (photo PTBG)). 1756. Type: Macrocnemum jamaicense L.
—Machaonia martinicensis (DC.) Standi. Macrocnemum candidissimum Vahl, Symb. 2:
38,/?/. 30. 1791. Type: TRINIDAD: Vicinia
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

St. Marthae, von Rohrs.n. (Holotype C, —Pogonopus speciosus (Jacq.) K. Schum.

Mussaenda.—Calycophvllu Note: Standley & Williams (1975) and Dwyer
(Vahl) DC. (1980) incorrectly referred collections from
Macrocnemum coccineum Vahl, Symb. Bot. 2: Central America to this South American
38. 1791. Type: TRINIDAD: Without precise species; the Central American species is
locality, von Rohrs.n. (C, n.v.). Mussaenda. Pogonopus exsertus (Oerst.) Oerst.
—Warszewiczia coccinea (Vahl) Klotzsch
Macrocnemum corymbosum Ruiz & Pav., Fl. Malanea Aubl., Hist. PI. Guiane 1: 106, t. 41.
Peruv. Prodr. 2: 48,/?/. 189. 1799. Type: 1775. Type: Malanea sarmentosa Aubl.
PERU: In Andium collibus et runcationibus Malanea colombiana Standi., Publ. Field
calidis versus Chinchao, Acomayo, Pillao, Columbian Mus., Bot. Ser. 7: 66. 1930. Type:
Muiia, Ruiz Lopez & Pavon s.n. (Presumably COLOMBIA: El Valle, Pennell & Killip 5987
MA, n.v.).—Condaminea corymbosa (Ruiz (Holotype US, n.v.: Isotypes GH, n.v., NY,
& Pav.) DC. n.v.).
Macrocnemum exsertum Oerst., Vidensk. Malanea erecta Seem., Bot. Voy. Herald 136.
Meddel. Dansk Naturhist. Foren. Kjobenhavn 1854. Type: PANAMA: Island of Taboga,
1852: 45. 1853. Type: COSTA RICA: Jarvis, Bay of Panama, Seemann s.n. (Holotype
3000 ft., Oersted s.n. (Holotype presumably presumably BM, n.v.).
C, n.v.).—Pogonopus exsertus (Oerst.) Oerst.
Macrocnemum glabrescens (Benth.) Wedd., i G. Karst., Fl. Columb. 2: 9, t. 105.
Ann. Sci. Nat., Bot., Ser. 4, 1: 76. 1854. 1862. Type: Mallostoma cachirense (G.
Lasionema glabrescens Benth., Bot. Voy. Karst.) G. Karst. Based on Hedyotis
Sulphur 105. 1844. Type: COSTA RICA: cachirensis G. Karst. (=Arcytophyllum
Lagartos Bay (Carno Bay), Gulf of Nicoya, cachirense (G. Karst.) K. Schum.)
12-16 Feb 1839, G. W. Barclay 2780 Mallostoma acerosa (A. Gray) Hemsl., Biol.
(Holotype BM, n.v.; Isotype F (photo Centr.-Amer., Bot. 2: 31. 1881. Hedyotis
PTBG)).—Macrocnemum roseum (Ruiz & acerosa A. Gray, PI. Wright. 1: 81. 1852.
Pav.) Wedd. Note: Macrocneum glabrescens Type: U.S.A.: Texas, high prairies, from Live
is regarded as a taxonomic synonym of M. Oak Creek to the Las Moras, Jun 1849, C.
roseum by Andersson in Andersson & Taylor Wright 237 (Holotype GH, n.v.; Isotype US).
(1994). Ereicoctis, Houstonia, Oldenlandia.
Macrocnemum pastoense G. Karst, Linnaea —Hedyotis acerosa A. Gray
30: 150. 1859. Type: COLOMBIA: Ad littora Mallostoma lavarum Donn. Sm., Enum. PI.
fluminis Mayo et Juanambu provinciae Pasto, Guatemal. 5: 36. 1899. Type: PANAMA:
Karsten (presumably W, n.v.). Chiriqui, summit of Chiriqui Volcano, 3374
Macrocnemum roseum (Ruiz & Pav.) Wedd., m, 12 Mar 1911, W. R. Maxon 5349
Ann. Sci. Nat., Bot., Ser. 4, 1: 76. 1854. (Holotype B, presumably destroyed (photo
Cinchona rosea Ruiz & Pav., Fl. Peruv. 2: 54. MO); Isotype US (photos MEXU, PTBG)).
1799. Type: PERU: Huanuco, Pozuzo, Ruiz —Arcytophyllum lavarum (Donn. Sm.) K.
Lopez & Pavon s.n. (Lectotype MA, n.v.; Schum. ex Standi.
Isolectotypes BM, n.v., G, n.v., G-DC, n.v., Mallostoma shannonii Donn. Sm., Bot. Gaz.
K, n.v., MA, n.v., P, n.v.; designated by (Crawfordsville) 18: 203. 1893. Type:
Andersson in Andersson & Taylor, 1994: 12). GUATEMALA: Chimaltenango, Chicoy,
Macrocnemum speciosus Jacq., Hort. Schoenbr. 9000 ft., Mar 1892, W. C. Shannon s.n.
1: 19. 1797. Type: VENEZUELA: Based on (Holotype US 48603 (photos MEXU, PTBG);
material cultivated at Schonbrunn, Vienna, Isotype US 943475).—Hedyotis serpyllacea
said to have been brought from vicinity of Schltdl.
Caracas, Venezuela (not located).
Manettia Mutis ex L., Mant. PL: 553, 558. 1771, Galeotti 2635 (Holotype BR, r
nom. cons. Type: Manettia reclinata L.
Manettia arboricola Lorence, Novon 3: 59. Manettia hydrophila Dwyer, Ann. Missouri Bot.
1993. Type: PANAMA: Comarca de San Gard. 67: 280. 1980. Type: PANAMA:
Bias, El Llano-Carti Rd., km 16.7, 350 m, trail Veraguas, mouth of Rio Concepcion, in
west to waterfall 5 km from road, 16 Jun swamp, 4 Dec 1967, W. H. Lewis, T. B. Croat
1985, G. de Nevers & S. Charnley 5907 & J. L. Hawker 2822 (Holotype MO
(Holotype MO 3623655 (photo PTBG)). 2268375).—Sipanea biflora (L. f.) Cham. &
Manettia barbata Oerst., Vidensk. Meddel. Schltdl.
Dansk Naturhist. Foren. Kjobenhavn 1852: Manettia liebmannii Standi., J. Wash. Acad. Sci.
47. 1853. Type: COSTA RICA: In Monte 17: 337. 1927. Type: MEXICO: Oaxaca,
Aguacate, 2000 ft., Feb 1846, Oersted 11521 Pelado [probably Cerro Pelon], Liebmann
(Holotype C, n.v. (photos MO, US); Isotypes 11485 [=Liebmann Rubiaceae 147] (Holotype
K, n.v., US (2 sheets; photos MEXU, PTBG)). C, n.v. (fragment US; photos MEXU,
Manettia hocataurensis Dwyer, Ann. Missouri PTBG)).—Hedyotis galeottii (M. Martens)
Bot. Gard. 67: 278. 1980. Type: PANAMA: Terrell & Lorence
Bocas del Toro, along RR (railroad) near Manettia longicalycina Dwyer & Lorence,
station at Milla 10, 27 Jul 1971, T. B. Croat & Novon 3: 61. 1993. Type: PANAMA: Code,
D. M. Porter 16334 (Holotype MO 2166138 Alto Calvario, 800-900 m, 20 Apr 1977, J. P.
(photo PTBG)).—Hedyotis lancifolia K. Folsom & Jaslon 2665 (Holotype MO
Schum. 2608777 (photo PTBG)).
Manettia costaricensis Wernham, J. Bot. 41 Manettia longipedicellata C. M. Taylor, Novon
(suppl.): 38. 1919. Syntypes: COSTA RICA: 5: 202, fig. 1A. 1995. Type: COSTA RICA:
Tucurrique, deboisements de Las Vueltas, Limon, Canton de Matina, Cordillera
650-700 m, Dec 1898, A. Tonduz 12873 (B, Talamanca, 200 m, aguas abajo de la
presumably destroyed (photos F, MO; confluencia de la quebrada Canabral con el no
fragment F), BM, n.v., US 1390734 (photos Barbilla, 10°00'10"N, 83°25'30"W, 5 Nov
MEXU, PTBG), US 354369). COSTA RICA: 1988, G. Herrera 2290 (Holotype CR, n.v.;
Without precise locality, 3500 ft., Endres 2 Isotype MO 3649361).
(W, n.v. (fragment F)).—Manettia reclinata Manettia microphylla Lorence & Dwyer,
L. Novon 3: 60. 1993. Type: PANAMA: Darien,
Manettia estrellae Standi., J. Wash. Acad. Sci. Pirre Massif, Alturas de Nique, above Cana
15: 6. 1925. Type: COSTA RICA: Cartago, Mine, 850-1150 m, 2 Mar 1988, G.
La Estrella, 26-27 Mar 1924, P. C Standley McPherson 12189 (Holotype MO 3648261
39224 (Holotype US 1153146 (photos (photo PTBG); Isotypes PTBG, SCZ).
MEXU, PTBG))—Manettia flexilis Manettia orbifera Wernham, J. Bot. 57 (suppl.):
Brandegee 41. 1919. Type: GUATEMALA: Retalhuleu,
Manettia flexilis Brandegee, Univ. Calif. Publ. Bernoulli & Cairo 1649 (Holotype K, n.v.).
Bot. 6: 196. 1915. Type: MEXICO: Chiapas, —Manettia reclinata L.
Cerro del Boqueron, Jun 1914, C. A. Purpus Manettia panamensis Duchass. & Walp.,
7118 (Holotype UC 174907, n.v.; Isotypes F, Linnaea 23: 753. 1850. Type: PANAMA:
GH, n.v., MO (photos MEXU, PTBG), US Duchassaing s.n. (presumably GOET, n.v.).
(photos MEXU, PTBG)).
Manettia hirtella M. Martens & Galeotti, Bull. lata L., Mant. PI. 2: 558. 1771.
Acad. Roy. Sci. Bruxelles 11: 238. 1844. Type: Not designated. Note: this species is
Type: MEXICO: Oaxaca, cordillere orientale, actually based on Colombian material seen b)
au bords des rivieres, dans les forets de Mutis, not Mexican material. Apparently
Tonaguia, 4000-5000 pieds, Jun 1840, H. Linnaeus believed that Santa Fe de Bogota
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

was the Santa Fe in Mexico (L. Andersson, F. Liebmann 11662 (Lectotype C (photos
pers. comm., 1996). No herbarium specimens MEXU, PTBG); Isolectotype US, n.v.;
are known to exist, and a neotype needs to be designated by Hamilton, 1989). Syntypes:
MEXICO: Veracruz, Mirador, Apr 1847, F.
Manettia seleriana Loes., Verhandl. Bot. Verein Liebmann 11577'(C (photos MEXU, PTBG);
Brandenburg 65: 107. 1923. Type: Mirador, F. Liebmann 11614 (C (photos
GUATEMALA: Izabal, Los Amates, 1 Jan MEXU, PTBG)). Psychotria.—Psychotria
1897, E. Seler & C. Seler 3309 (Holotype B, costivenia Griseb. var. costivenia
presumably destroyed (photos F, MO, US; Mapouria obovata Oerst., Amer. Centr. 17, t.
fragment F); Isotype US (photos MEXU, 14, figs. 3, 4. 1863. Not Psychotria obovata
PTBG)).—Manettia reclinata L. Ruiz & Pav., 1799. Type: MEXICO:
Manettia stenophylla Donn. Sm., Bot. Gaz. Veracruz, F. Liebmann s.n. (Holotype
(Crawfordsville) 56: 58. 1913. Type: COSTA presumably C, n.v.). Psychotria.—Psychotria
RICA: Tucurrique, Las Vueltas, 635 m, Jan quinqueradiata Pol.
1899, A. Tonduz 12969 (Holotype US 943614 Mapouria papantlensis, Oerst., Amer. Centr. 17.
(photos F, MEXU, PTBG, US); Isotype B, t. 14, fig. 5. 1863. Type: MEXICO: Veracruz,
destroyed (fragment F)).—Manettia barbata Papantla,/ Liebmann s.n. (Holotype
Oerst. presumably C, n.v.). Uragoga.—Psychotria
Manettia zimapanica Hemsl., Diag. PI. Nov. papantlensis (Oerst.) Hemsl.
Mexic. 30. 1879. Type: MEXICO: Hidalgo, Mapouria remota (Benth.) Muell. Arg., Flora
Zimapan, Coulter 203 (Holotype K, n.v.). 59: 459. 1876. Psychotria remota Benth., J.
—Status u Bot. Hook. 3: 225. 1841. Type: BRAZIL: On
the Rio Negro, Schomburgk 963 (Holotype K,
Mapourea Aubl., Hist. PL Guiane 1: 175. 1775. n.v. (photo NY); Isotypes F, W, n.v.).
Type: Mapouria guianensis Aubl. [Herein —Psychotria remota Benth.
treated as a synonym of Psychotria L.] Mapouria salicifolia Oerst., Amer. Centr. 17, t.
Mapourea chamissoana Loes., Verhandl. Bot. 14, fig. 2. 1863. Not Psychotria salicifolia
Verein Brandenburg 65: 112. 1923. Type: Kunth in Humb., Bonpl. & Kunth, 1819.
MEXICO: Veracruz, Tecolute [Tecolutla], Type: MEXICO: Oaxaca, Lacoba [Lacova],
Schiede 1266 (Holotype B, presumably F. Liebmann s.n. (Holotype presumably C,
destroyed (photo MO)). Psychotria. n.v). Psychotria.—Psychotria papantlensis
—Psychotria erythrocarpa Schltdl. (Oerst.) Hemsl.
Mapourea graciliflora (Benth.) Oerst., Amer. Mapouria stipulata Oerst., Amer. Centr. 17, t.
Centr. 17, t. 14, fig. 6. 1863. Psychotria 14, fig. 7. 1863. Type: MEXICO: Veracruz,
graciliflora Benth. in Oerst., Vidensk. Papantla, F. Liebmann 11615 (Lectotype C,
Meddel. Dansk Naturhist. Foren. Kjobenhavn here designated (photos MEXU, PTBG);
1852: 35. 1853. Type: COSTA RICA: Isolectotype C (photos MEXU, PTBG)).
Alajuela, ad Naranjo, Apr 1847, Oersted Syntype: MEXICO: Veracruz, Papantla, F.
11611 (Holotype C (photos F, MEXU, Liebmann 11636 (C (photos MEXU, PTBG)).
PTBG); Isotypes K, n.v., US (photos MEXU, Liebmann 11615, a well preserved flowering
PTBG)).—Psychotria graciliflora Benth. specimen with a duplicate, is the best choice
Mapouria micrantha (Kunth) Wernham, Kew for lectotype. Psychotria.—Psychotria
Bull. 1914: 69. 1914. Type: PERU (?): A. costivenia Griseb. var. costivenia
Bonplands.n. (Holotype P-Bonpland, n.v. Mapouria tomentosa Oerst., Amer. Centr. 17, t.
(fragment F)).—Psychotria micrantha Kunth 14, fig. 8. 1863. Not Psychotria tomentosa
Mapouria miradorensis Oerst., Amer. Centr. 17, Mtill Arg. ex Mart., 1881, nor Hemsl., 1881.
/. 14, fig. 9. 1863. Type: MEXICO: Veracruz, Type: MEXICO: Veracruz, prope Papantla, F.
Mirador, in quercetum, 3000 ft., 1841-1843, Liebmann s.n. (Holotype presumably C, n.v.).
 —Status uncertain, probably Psychotria MEXICO: Baja California, Cape [Cabo] San
micrantha Kunth in Humbl., Bonpl. & Kunth Lucas, Hinds 65 (Holotype BM, n.v.).
Mapouria trichogyne Muell. Arg. in Mart., Fl. Mitracarpus pallidus Hook. & Am., Bot.
Brasil. 6: 426. 188. Type: BRAZIL: Habitat in Beech. Voy. 4: 30. 1840. Type: MEXICO:
prov. de Alto Amazonas, prope Manaos ad Realjo, collector probably Sinclair s.n.
Rio Negro, Spruce s.n. (B, presumably (Holotype presumably K, n.v.).
destroyed). Geophila.—Geophila cordifolia Mitracarpus rhadinophyllus (B. L. Rob.) L. O.
Miq. Williams, Fieldiana, Bot. 29: 371. 1961.
Borreria rhadinophylla B. L. Rob., Proc.
Microsplenium Hook. f. in Benth. & Hook, f., Amer. Acad. 45: 409. 1910. Type: BRITISH
Gen. PI. 2: 4. 1873. Type: Microsplenium HONDURAS (BELIZE): On dry, sandy pine
coulteri Hook. f. [Herein treated as a synonym ridges, 23 Oct 1905, M. E. Peck 180
of Machaonia Humb. & Bonpl] (Holotype GH, n.v.; Isotype NY, n.v.)
Microsplenium coulteri Hook. f. in Benth. & Mitracarpus schizangius DC, Prodr. 4: 572.
Hook, f., Gen. PI. 2: 4. 1873. Type: MEXICO: 1830 (as Mitracarpum schizangium). Type:
Hidalgo, Zimapan, Coulter (Holotype K, MEXICO: Without precise locality, Haenke
n.v.).-^Machaonia coulteri (Hook, f.) Standi. s.n. (Holotype presumably PR, n.v.).
Mitracarpus schizangius DC. var. angx
Mitchella L., Sp. PI. 1: 111. 1753. Type: Oerst., Vidensk. Meddel. Dansk Naturhist.
Mitchella repens L. Foren. Kjobenhavn 1852: 7. 1853. Type:
Mitchella repens L., Sp. PI. 1: 111. 1753. Type: NICARAGUA: Volcan El Viejo, Oersted s.n.
U.S.A.: Habitat in Carolina, Terra Mariana, (Holotype C, n.v.).—Status uncertain.
Virginia, Herb. Linn. 135.1 (Lectotype LINN, Mitracarpus villosus (Sw.) Cham. & SchltdL,
n.v.; designated by Reveal et al., 1987: 215). Linnaea 3: 363. 1828. Spermacoce villosa
Sw., Prodr. Veg. Ind. Occ. 29. 1788. Type:
. [Here JAMAICA: Swartz s.n. (Holotype presumably
BM, n.v.).—Mitracarpus hirtus (L.) DC

Mitracarpus Zucc. ex Schult. & Schult f. in Montamans Dwyer, Ann. Missouri Bot. Gard.
Roem. & Schult., Mant. PI. 3: 210 (as 67: 284. 1980. Type: Montamans panamensis
Mitracarpum), 399. 1827. Type: Mitracarpus Dwyer [Herein treated as a synonym of
scaber Zucc. ex Schult. & Schult. f. Psychotria L.]
Mitracarpus breviflorus A. Gray, PI. Wright. 2: Montamans panamensis Dwyer, Ann. Missouri
68. 1853 (as Mitracarpum brevifiorum). Type: Bot. Gard. 67: 286, fig. 62. 1980. Type:
MEXICO: Sonora, mountain ravines and hill- PANAMA: Chiriqui, Cerro Horqueta, rain
sides near Santa Cruz, Sep 1851, C. Wright forest, ca. 5500 ft., 20 Jul 1968, J. D. Dwyer
1120 (Holotype GH, n.v.; Isotype US (photos & B. Lallathin 8792 (Holotype MO 2163225
MEXU, PTBG)).—Mitracarpus hirtus fL.) (photos MEXU, PTBG)).—Psychotria
DC. aggregata Standi.
Mitracarpus hirtus (L.) DC, Prodr. 4: 572.
1830. Spermacoce hirta L., Sp. PL, ed. 2, 1: Morinda L., Sp. PL 1: 176. 1753. Type: Morinda
148. 1762. Not Spermacoce hirta L., 1767,
nor Sw., 1791. Type: JAMAICA: Browne Morinda asperula Standi., Publ. Field Mus.
'Spermacoce # 3' (LINN 125.4, n.v.). For Nat. Hist., Bot. Ser. 22: 54. 1940. Type:
typification see Verdcourt (1975: 317-322). GUATEMALA: Izabal, hilly pine forest near
Mitracarpus linearis Benth., Bot. Voy. Sulphur Quirigua, 72-150 m, 26-27 Apr 1939, P. C.
20. 1844 (as Mitracarpium linear e). Type: Standley 72368 (Holotype F 991552 (photo
PTBG)).
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Morinda citrifolia L., Sp. PI. 1: 176. 1753. ..,Sp.Pl. 1: 177. 1753. Type:
Type: INDIA: (Holotype LINN 236.1, n.v. i frondosa L.
(microfiche MO)). Mussaenda acutiflora Bartl. ex DC, Prodr. 4:
Morinda guatemalensis (Donn. Sm.) Steyerm., 370. 1830. Type: MEXICO: Without precise
Mem. New York Bot. Gard. 23: 385. 1972. locality, Haenke s.n. (Holotype presumably
Appunia guatemalensis Donn. Sm., Bot. Gaz. PR, n.v., fragment G-DC, n.v. (microfiche 4:
(Crawfordsville) 48: 294. 1909. Type: 370.2)).—Status uncertain.
GUATEMALA: Izabal, Livingston, Jun 1906, Mussaenda candidissima (Vahl) Roem. &
H. von Tuerckheim III-230 (Holotype US Schult., Syst. Veg. 5:251. 1819.
943501 (photos MEXU, PTBG)).—Appunia Macrocncn: .M\, Symb.
guatemalensis Donn. Sm. Bot. 2: 38,/?/. 30. 1791. Type: TRINIDAD:
Morinda laxa Bartl. ex DC, Prodr. 4: 449. Vicinia St. Marthae, von Rohrs.n. (Holotype
1830. Type: MEXICO: Without precise C, n.v).—Calycophyllum candidissumum
locality, Haenke s.n. (Holotype presumably (Vahl) DC.
PR, n.v.).—Status uncertain. Mussaenda coccinea (Vahl) Poir. in Lam.,
Morinda mesochora Standi., Publ. Field Encycl. Meth. 4: 394. 1797. Macrocnemum
Columbian Mus., Bot. Ser. 4: 345. 1929. coccineum Vahl, Symb. Bot. 2: 38. 1791.
Type: BRITISH HONDURAS (BELIZE): Type: TRINIDAD: Without precise locality,
Honey Camp, Orange Walk, Sep 1928, C. L. von Rohrs.n. (Holotype C, n.v.).
Lundell 19 (Holotype F 580825; Isotype US —Warszewiczia coccinea (Vahl) Klotzsch
(photos MEXU, PTBG)). Mussaenda erythrophylla Schumach. &
Morinda muscosa Jacq., Sel. Stirp. Amer. 65. Thonn., Beskr. Guin. PI. 116. 1827. Type:
1763. Type: MARTINIQUE: Jacquin s.n. TROPICAL AFRICA: Guinea, Thonnings.n.
(location unknown).—Psychotria muscosa (Holotype presumably C, n.v.; probable
(Jacq.) Steyerm. Isotype G-DC, n.v. (microfiche 4: 371.8)).
Morinda panamensis Seem., Bot. Voy. Herald Cultivated in Mesoamerica.
136. 1854. Type: PANAMA: Panama, Mussaenda formosa Jacq., Enum. PI. Carib. 16.
Chagres, 10 Feb 1850, Fendler 148 (Holotype 1760. Type: COLOMBIA: Cartagena, Jacquin
BM, n.v.; Isotypes GH, n.v., MO (photos s.n. (presumably G, n.v.). Randia.
MEXU, PTBG), US (photo PTBG)). —Rosenbergiodendron formosum (Jacq.)
Morinda royoc L., Sp. PI. 1: 176. 1753 [as Fagerl.
roioc]. Type: Habitat in America calidiore, Mussaenda glabra Vahl, Symb. Bot. 3: 38.
Herb. Clifford 73, Morinda 1 (Lectotype BM; 1794. Type: Collector and location unknown.
designated by Lorence, 1993: 68). Cultivated in Mesoamerica.
Morinda yucatanensis Greenm., Publ. Field Mussaenda luteola Delile in Caill., Voy. Meroe
Columbian Mus., Bot. Ser. 2: 262. 1907. 4: 65. 1827. Type: ARABIA: Nubia, collector
Type: MEXICO: Yucatan, in brush lands unknown (Holotype presumably MPU, n.v.;
about Izamal, 1895, G F. Gaunter 362 probable isotype G-DC, n.v. (microfiche 4:
(Lectotype F 36165 (photo PTBG), here 371.9)). Cultivated in Mesoamerica.
designated; Isolectotypes GH, n.v., US Mussaenda spinosa Sesse & Moc., La
(photos MEXU, PTBG)). Note: in his Naturaleza, ser. 2, 1: 36. 1888. Type:
protologue Greenman cited duplicates of MEXICO: Habitat in Martinica, Indiarum
Gaunter 362 at both F and GH without Cartagine et calidis Novae Hispaniae
designating a holotype. An additional regionibus, Sesse & Mocirio 1583
duplicate exists at US. I have selected the (presumably MA, n.v. (fragment F)).
excellent flowering and fruiting specimen at F —Randia tetracantha (Cav.) DC.
as lectotype.—Morinda royoc L. Mussaenda tetracantha Cav., Icon. PI. 4: 20,/?/.
435. 1799. Type: MEXICO: [Probably
 Guerrero], cerca Acapulco, Nee s.n. however, it does not occur in the Northern
(presumably MA, n.v. (photos F, MEXU, Hemisphere.
MO)).—Randia tetracantha (Cav.) DC. Nertera alsinoides Schltdl. & Cham., Linnaea 6:
413. 1831. Type: MEXICO: Questa Grande
Nauclea L., Sp. PL, ed. 2, 1: 243. 1762. Type: de Jalacingo, Schiede & Deppe 1271
Nauclea orientalis (L.) L., based on (Holotype presumably HAL, n.v.).
Cephalanthus orientalis L. —Didymaea alsinoides (Schltdl. & Cham.)
Nauclea cadamba Roxb., Fl. Ind. 2: 121. 1824. Standi.
Type: INDIA: Rheede, Hort. Ind. Malabar. 3: Nertera granadensis (Mutis ex L. f.) Druce,
tab. ii, Katou-tsjaka (1678-1703). Rep. Bot. Soc. Exch. Club Brit. Isles 1916:
—Neolamarckia cadamba (Roxb.) Bosser 637. 1917. Gomozia granadensis Mutis ex
Nauclea tomentosa Willd. ex Roem. & Schult, L. f., Suppl. PL 129. 1781. Type:
Syst. Veg. 5: 221. 1819. Type: COLOMBIA: COLOMBIA: America meridionali, Mutis s.n.
In ripa flumen Magdalenae prope Naers, (LINN 172.1, n.v. (microfiche)).
Humboldt s.n. (Holotype B-W 3910, n.v.).
—Uncaria tomentosa (Willd. ex Roem. & Nonatelia AubL, Hist. PL Guiane 1: 182, pi.
Schult.) DC. 70-75. Ml 5. Type: Nonatelia racemosa AubL
[Herein treated as a synonym of Psychotria
Neolamarckia Bosser, Bull. Mus. Natl. Hist. L-]
Nat, Paris, Ser. 4, Sect. B, Adansonia 6(3): Nonatelia officinalis Aubl, Hist. PL Guiane 1:
247. 1984. Type: Neolamarckia cadamba 182, tab. 70. 1775. Type: FRENCH
(Roxb.) Bosser GUIANA: Cayenne, Aublet s.n. (Holotype
Neolamarckia cadamba (Roxb.) Bosser, Bull. presumably BM, n.v.).—Psychotria
Mus. Natl. Hist. Nat., Paris, Ser. 4, Sect. B, s (AubL) Raeusch.
Adansonia 6(3): 247. 1984. Nauclea cadamba apanamensis DC, Prodr. 4: 466. 1830.
Roxb., Fl. Ind. 2: 121. 1824. Type: INDIA: Type: PANAMA: Without precise locality,
Rheede, Hort. Ind. Malabar. 3: tab. 33, Haenke s.n. (Holotype presumably PR,
Katou-tsjaka (1678-1703). Cultivated and n.v.).—Status uncertain, probably
naturalized in Costa Rica according to Burger Psychotria racemosa (AubL) Raeusch.
& Taylor (1993). Nonatelia racemosa AubL, Hist. PL Guiane 1:
187, t. 72. Ml 5. Type: FRENCH GUYANA:
In sylvis territorii Orapu, Aublet s.n.
(Holotype P, n.v.).—Psychotria r
Urb. Rich.
Nernstia mexicana (Zucc. & Mart, ex DC.)
Urb., Symb. Antill. 9: 146. 1923. Coutarea Nothophlebia Standi., Contr. U.S. Natl. Herb.
mexicana Zucc. & Mart, ex DC, Prodr. 4: 17: 438. 1914. Type: Nothophlebia
350. 1830. Type: MEXICO: Without precise costancensis Standi. [Herein treated as a
locality, Martius in 1829 (Holotype G-DC, synonym of Pentagonia Benth.]
n.v. (IDC 800.680: II.7)). Cigarilla. Nothophlebia costaricensis Standi., Contr. U.S.
Natl. Herb. 17: 438. 1914. Type: COSTA
Nertera Banks & Sol. ex C. F. Gaertn., Fruct. RICA: Collines de Moin, littoral atlantique,
Sem. PL 1: 124, pi. 26. 1788. Nomen Nov 1899, H. Pittier 16024 (Holotype US
conservandum. Type: Nertera depressa 578472 (photos MEXU, PTBG)).
Gaertn. [=Nertera granadensis (Mutis ex L. f.) —Pentagonia costaricensis (Standi.) W. C
Druce.] Note: Nertera depressa was listed, as Burger & C M. Taylor
a synonym of Gomozia granadensis, by
Standley (1938) as occurring in Costa Rica;
 i NOMENCLATOR OF MEXICAN AND CENTRAL AMER

Oldenlandia L., Sp. PI. 1: 119. 1753. Type: Oldenlandia herbacea (L.) Roxb., Hort. Bengal.
Oldenlandia corymbosa L. [Herein treated as a 11. 1814. Hedyotis herbacea L., Sp. PI. 1:
synonym of Hedyotis L.] 102. 1753. Type: CEYLON: (Presumably
Oldenlandia acerosa (A. Gray) A. Gray, PI. LINN, n.v.).—Hedyotis herbacea L.
Wright. 2: 67. 1853. Hedyotis acerosa A. Oldenlandia humifusa (A. Gray) A. Gray,
Gray, PI. Wright. 1:81. 1852. Type: U.S.A.: Smiths. Contr. Knowl. 5: 68. 1853. Hedyotis
Texas, high prairies from Live Oak Creek to humifusa A. Gray, J. Boston Soc. Nat. Hist. 6:
the Las Moras, Jun 1849, C Wright 237 216. 1850. Type: U.S.A.: Texas, near
(Holotype GH, n.v.; Isotype US). Houstonia, Fredericksburg, 1847, F. Lindheimer 621
Mallostoma.—Hedyotis acerosa A. Gray (Holotype GH, n.v.; Isotypes BM, n.v.,
Oldenlandia angustifolia (Michx.) A. Gray, PI. MICH, n.v., MO, PH, n.v., US). Houstonia.
Wright. 2: 68. 1853. Houstonia angustifolia —Hedyotis humifusa A. Gray
Michx., Fl. Bor. Amer. 1: 85. 1803. Type: Oldenlandia lancifolia (Schumach.) DC, Prodr.
U.S.A.: Florida in submantimis, collector not 4: 425. 1830. Hedyotis lancifolia Schumach.
cited (presumably P, n. v.).—Hedyotis in Schumach. & Thonn., Beskr. Guin. PI. 72.
nigricans (Lam.) Fosberg 1827. Type: GHANA: Valley of Aquapim,
Oldenlandia callitrichoides Griseb., Mem. Thonning 210 (Lectotype C, n.v.; Isolectotype
Amer. Acad. Arts, n.s. 8: 506. 1863. Type: S, n.v.; designated by Verdcourt, 1976: 292).
CUBA: Without precise locality, ad ripas —Hedyotis lancifolia Schumach. in
rivulorum, Wright 268 (Lectotype GOET, Schumach. & Thonn.
n.v.; designated by Terrell & Lewis, 1990). Oldenlandia latifolia M. Martens & Galeotti,
Syntype: CUBA: Wright 1640 (GOET, n.v.). Bull. Acad. Roy. Sci. Bruxelles 11: 235. 1844.
Oldenlandiopsis.—Hedvotis callitrichoides Type: MEXICO: Veracruz, Mirador, near
(Griseb) W. H. Lewis Xalapa, 3000 ft., 1840, H Galeotti 2556
Oldenlandia corymbosa L., Sp. PI. 1: 119. 1753. (Holotype BR, n.v.; Isotypes F (photos
Lectotype: "Oldenlandia," Plumier, Nov. PI. MEXU, PTBG), GH).—Hedyotis exigula
Amer. t. 36. 1703; designated by Verdcourt W. H. Lewis
(1976: 308).—Hedyotis corymbosa (L.) Oldenlandia microtheca (Schltdl. & Cham.)
DC, Prodr. 4: 428. 1830. Gerontogea
Oldenlandia deppeana (Schltdl. & Cham.) DC, microtheca Schltdl. & Cham., Linnaea 5: 169.
Prodr. 4: 428. 1830. Gerontogea deppeana 1830. Type: MEXICO: Veracruz, Barranca de
Schltdl. & Cham., Linnaea 5: 169. 1830. Tioselos in umbrosis, Sep, Schiede & Deppe
Type: MEXICO: Veracruz, ad San Andres, 390 (Holotype presumably HAL, n.v.; Isotype
1829, Schiede & Deppe 391 (Holotype MO (photos MEXU, PTBG), label states In
presumably HAL, n.v.; Isotype MO (photos sylvis Papanatlae, Jan., Dec, corolla alba).
MEXU, PTBG)). Hedyotis.—Hedyotis —Hedyotis microtheca (Schltdl. & Cham.)
microtheca (Schltdl. & Cham.) Steud. Steud.
Oldenlandia depressa Sesse & Moc., Fl. Mexic. Oldenlandia ovata S. Watson, Proc. Amer.
27. 1893. Not O. depressa Willd. in L., 1798. Acad. Arts 18: 97. 1883. Not Hedyotis ovata
Type: MEXICO: Habitat in montibus S. Thunb. ex Maxim., 1883. Type: MEXICO:
Augustine et viciniis. Floret Julio, Sesse, Nuevo Leon, Guajuco, 27 miles SE of
Mocino, Castillo & Maldonado 601 (Fl. Mex. Monterrey, 1-8 Mar 1880, E. Palmer 399
Ic 325) (Holotype presumably MA, n.v.; (Lectotype GH, n.v.; Isolectotype F (photos
Isotype F, n.v.) —Richardia tricocca (Torr. MEXU, PTBG), US (photo PTBG);
& A. Gray) Standi, subsp. tetracocca (M. designated by Terrell, 1978, in herb.).
Martens & Galeotti) W. H. Lewis & R. L. Syntype: MEXICO: Ervandberg 200 (GH,
Oliv. n.v.).—Hedyotis watsonii W. H. Lewis
Oldenlandia pringlei B. L. Rob., Proc. Amer.
Acad. Arts 27: 169. 1892. Type: MEXICO:
San Luis Potosi, alkaline plaines, Hacienda de
Angostura, 27 Jun 1891, C. G. Pringle 3758 Omiltemia Standi., J. Wash. Acad. Sci. 8: 427.
(Holotype GH, n.v.; Isotypes F, MO (photo 1918. Type: Omiltemia longipes Standi.
PTBG), US (2 sheets; photos MEXU, Omiltemia filisepala (Standi.) C. V. Morton,
PTBG)).—Hedyotis pringlei (B. L. Rob.) Baileya 16: 98. 1968. Kohleria filisepala
W. H. Lewis Standi, Publ. Field Mus. Nat. Hist., Bot. Ser.
Oldenlandia racemosa Sesse & Moc., Fl. 22: 107. 1940. Type: MEXICO: Chiapas, Mt.
Mexic. 28. 1893. Type: MEXICO: Habitat in Ovando, 16 Dec 1936, E. Matuda 574
Cordovae pratis. Floret Julio, Sesse & Mocino (Holotype F 916483 (photo PTBG); Isotypes
s.n. (presumably MA, n.v.).—Status MEXU, US).
uncertain. Omiltemia floribunda (Standi.) J. H. Kirkbr.,
Oldenlandia repens L., Mant. 40. 1767. Type: Syst. Bot. 9: 413. 1984. Edithea floribunda
INDIA: Without precise locality or collector Standi., Trop. Woods. 34: 2. 1933. Not
(Lectotype LINN 155.2, n.v.; designated by Deppea floribunda Hemsl., 1879. Type:
Verdcourt, 1983).—Dentella repens (L.) J. R. MEXICO: Guerrero, Malinaltepec, Mar 1930,
& G. Forst. L. Schultze 363 (Holotype B, destroyed
Oldenlandia rubra (Cav.) A. Gray, PI. Wright. (fragment F)).—Deppea schultzei Lorence
2: 68. 1853. Houstonia rubra Cav., Icon. 5: Omiltemia longipes Standi., J. Wash. Acad.
48, t. 474, fig. 1. 1799. Type: MEXICO: Sci. 8: 427. 1918. Type: MEXICO: Guerrero,
Hidalgo, Ixmiquilpan, Nee s.n. (Presumably Omilteme, May 1903, E. W. Nelson 7054
MA, n.v.).—Hedyotis rubra (Cav.) A. Gray (Holotype US 1084692 (photos MEXU,
Oldenlandia xestosperma B. L. Rob. & PTBG); Isotypes GH, n.v., US).
Greenm., Proc. Amer. Acad. Arts 32: 41. Omiltemia scoti J. H. Kirkbr., Syst. Bot. 9: 411.
1896. Type: MEXICO: Oaxaca, open glades, 1984. Type: MEXICO: Puebla, above
Sierra de San Felipe, 8000 ft., Jun 1894, C G. Teotitlan del Camino on road to Huautla de
Pringle 4692 (Lectotype GH, n.v.; Jimenez, 2000-3250 m, 3 Aug 1961, C. E.
Isolectotypes BM, n.v., F, K, n.v., MO Smith, Jr., F. A. Peterson & N. Tejada 4137
(photos MEXU, PTBG), MSC, n.v., PH, n.v., (Holotype US 2645608; Isotype GH (photo
US (2 sheets; photos MEXU, PTBG), VT, PTBG)).—Deppea scoti (J. H. Kirkbr.)
n.v.; designated by Terrell, 1980). Hedyotis,
Houstonia.—Bouvardia xestosperma (B. L.
Rob. & Greenm.) Terrell & S. D. Koch Ophiorhiza L., Gen., ed. 5, 74. 1754.
(=Ophiorrhiza L., 1753.)
Oldenlandiopsis Terrell & W. H. Lewis, Ophiorhiza lanceolata Forssk., F. Aegypt.-Arab.
Brittonia42: 185. 1990. Type: 42. 1775. Type: YEMEN: Hadie Mts.,
Oldenlandiopsis callitrichoides (Gnseb.) Forsskals.n. (Holotype C, n.v.; Isotype BM,
Terell & W. H. Lewis. [Herein treated as a n.v.).—Pentas lanceolata (Forssk.) Deflers
synonym of Hedyotis L.)
Oldenlandiopsis callitrichoides (Griseb.) Terrell Ophryococcus Oerst, Vidensk. Meddel. Dansk.
& W. H. Lewis, Brirtonia 42: 185. 1990. Naturhist. Foren. Kjobenhavn 1852: 52. 1853.
Oldenlandia callitrichoides Griseb., Mem. Type: Ophryococcus gesnerioides Oerst.
Amer. Acad. Arts, n.s. 8: 506. 1863. Type: [Herein treated as a synonym of Hoffmannia
CUBA: Ad ripas rivulorum, C Wright 268 Sw.]
(Lectotype GOET, n.v.; designated by Terrell Ophryococcus gesnerioides Oerst., Vidensk.
& Lewis, 1990). Syntype: CUBA: Wright Meddel. Dansk. Naturhist. Foren. Kjobenhavn
1852: 53. 1853. Type: NICARAGUA:
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Segovia, Pantasmo, Jan 1848, Oersted 9332 Haenke s.n. (presumably PR, n.v.; not located
[=Plantae centroamericanae Oersted 9332] by Puff, 1991).
(Holotype C, n.v. (photo F); Isotype F). Paederia pringlei Greenm., Proc. Amer. Acad.
—Hoffmannia gesnerioides (Oerst.) Kuntze Arts 39: 92. 1903. Type: MEXICO: Morelos,
lava fields near Yautepec, 4500 ft., 27 Oct
Oregandra Standi., Publ. Field Columbian Mus., 1902, C G. Pringle 8682 (Holotype GH, n.v.;
Bot. Ser. 4: 265. 1929; Trop. Woods 16: 22. Isotypes MEXU, US (photos MEXU, PTBG)).
1928 (nomen nudum). Type: Oregandra —Paederia ciliata (Bartl. ex DC) Standi.
panamensis Standi. [Herein treated as a
synonym of Chione DC] Pahcourea Aubl., Hist. PI. Guiane 1: 172,/?/. 66.
Oregandra panamensis Standi., Publ. Field Ml5. Type: Palicourea guianensis Aubl.
Columbian Mus., Bot. Ser. 4: 265. 1929. Palicourea adusta Standi., J. Wash. Acad. Sci.
Type: PANAMA: Talamanca Valley, 1927, 18: 279. 1928. Type: COSTA RICA: San
G. P. Cooper & G. M. Slater 144 {Yale Jose, Cerro de las Vueltas, 2700-3000 m, 29
10490) (Holotype F 573817 (photo PTBG); Dec 1925-1 Jan 1926, P. C. Standley & J.
Isotypes US 1316774 (photos MEXU, PTBG), Valeria 43666 (Holotype US 1253350 (photo
US 1316777).—Chione sylvicola (Standi.) PTBG)).
W. C. Burger Palicourea albocaerulea C M. Taylor, Syst.
Bot. Monogr. 26: \5,fig. 3c, d. 1989. Type:
Osa Aiello, J. Arnold Arbor. 60: 115. 1979. COSTA RICA: Puntarenas, Monteverde
Type: Osa pulchra (D. R. Simpson) Aiello Cloud Forest Reserve, Pacific slope of
Osa pulchra (D. R. Simpson) Aiello, J. Arnold Continental Trail, Nuboso Trail, 1540-1620
Arbor. 60: 116. 1979. Hintonia pulchra D. R. m, 16 Sep 1983, Busby 107 (Holotype DUKE,
Simpson, Phytologia 29: 277. 1974. Type: n.v.).
COSTA RICA: Puntarenas, Osa Peninsula, Palicourea angustifolia Kunth in Humb.,
about 5 km W. of Rincon de Osa, W. Burger Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 367.
& R. Liesner 7320 (Holotype F 1729029 1819. Type: VENEZUELA: Amazonas, banks
(photo PTBG); Isotypes MO (photos MEXU, of Orinoco and Rio Negro, Humboldt &
PTBG), US (photos MEXU, PTBG)). Bonplands.n. (Holotype P-Bonpland, n.v.;
Isotype B-WI, n.v. (photo MO)).
Otocalyx Brandegee, Univ. Calif. Publ. Bot. 6: Palicourea austin-smithii Standi., Publ. Field
68. 1914. Type: Otocalyx chiapensis Mus. Nat. Hist., Bot. Ser. 18: 1333. 1938.
Brandegee [Herein treated as a synonym of Type: COSTA RICA: Alajuela, Canton Alfaro
Rondeletia L.] Ruiz, Palmira, 2300 m, 4 Aug 1938, A. Smith
Otocalyx chiapensis Brandegee, Univ. Calif. HI034 (Holotype F 942192 (photo PTBG);
Publ. Bot. 6: 69. 1914. Not Rondeletia Isotypes F, GH, n.v., MO (photos MEXU,
chiapasensis Standi., 1940. Type: MEXICO: PTBG), US (photo PTBG)).—Palicourea
Chiapas, Cerro del Boqueron, 1913, C. A. salicifolia Standi.
Purpus 7041 (Holotype UC 173047 (photos Palicourea bella (Standi.) Dwyer, Ann.
MEXU, PTBG); Isotype GH, n.v.). Missouri Bot. Gard. 67: 299. 1980. Psychotria
—Rondeletia h bella Standi., J. Wash. Acad. Sci. 18: 185.
1928. Type: PANAMA: Chiriqui, between
Paederia L., Syst. Nat., ed. 12, 2: 189. 1767; Alto de Las Palmas and Cerro de la Horqueta,
Mant. PI. 52. 1767. Type: Paederia foetida L. 2100-2268 m, 18 Mar 1911, H Pittier 3250
Paederia ciliata (Bartl. ex DC.) Standi., Publ. (Holotype US 677646 (photo PTBG); Isotype
Field Mus. Nat. Hist., Bot. Ser. 22: 386. 1940. F (photos MEXU, PTBG)).
Lygodysodea ciliata Bartl. ex DC, Prodr. 4: Palicourea bellula C M. Taylor, Syst. Bot.
470. 1830. Type: MEXICO: Ad Acahuitzola, Monogr. 26: 24, fig. 9, a, b, d. 1989. Type:
 COSTA RICA: Heredia, Rio Vueltas, upper designated by Standley, 1938).—Palicourea
Rio Patria, eastern slope of Volcan Barba at padifolia (Willd. ex Roem. & Schult.) C. M.
the Continental Divide, ca. 2000 m, 27 Jan Taylor & Lorence
1970, W. C. Burger & R. L. Liesner 7514 Palicourea crocea (Sw.) Roem. & Schult., Syst.
(Holotype F 1833197 (photo PTBG); Isotype Veg. 5: 193. 1819. Psychotria crocea Sw.,
MO (photo PTBG)). Prodr. Veg. Ind. Occ. 44: 1788. Type:
Palicourea brenesii Standi., Publ. Field Mus. JAMAICA: Without precise locality, Swartz
Nat. Hist., Bot. Ser. 18: 1333. 1938. Type: s.n. (Holotype BM, n.v.).
COSTA RICA: Palmira del Naranjo, in forest, Palicourea dimorphandrioides (Dwyer) C. M.
1850 m, Apr 1921, Brenes 3512 (Holotype F Taylor, Novon 6: 211. 1996. Cephaelis
851630 (photo PTBG); Isotype GH, n.v.). dimorphandrioides Dwyer, Ann. Missouri
Palicourea caerulescens Suessenguth, Bot. Bot. Gard. 67: 66. 1980. Type: PANAMA:
Jahrb. Syst. 72: 286. 1942. Type: COSTA Veraguas, valley of Rio Dos Bocas on road
RICA: San Jose-Limon border region, between Alto Piedras (above Santa Fe) and
Chirripo Grande, 3000 m, 30 Apr 1932, Calovebora, 350-400 m, 19 Aug 1974, T. B.
Kupper 1200 (Holotype M, n.v.). Croat 27470 (Holotype MO 2201358 (photos
—Palicourea salicifolia Standi. MEXU, PTBG)).
Palicourea carpirostrata Dwyer, Ann. Missouri Palicourea discolor K. Krause, Bot. Jahrb.
Bot. Gard. 67: 299. 1980. Type: PANAMA: Engl. 54, Beibl. 119: 40. 1916. Type: COSTA
Veraguas, NW of Santa Fe, 2 km from RICA: Canas Gordas, 1100 m, 18 Feb 1897,
Escuela Agricola Alto de Piedra. Cloud forest H. Pittier 11161 (Holotype B, presumably
on ridge top below summit of Cerro Tute, 28 destroyed (photos F, US), fragment F; Isotype
Mar 1975, S. Mori & J. Kallunki 5262 US (photo PTBG)).
(Holotype MO 2385000 (photo PTBG); Palicourea domingensis (Jacq.) DC, Prodr. 4:
Isotype MO 2384999 (photo PTBG)). 529. 1830. Psychotria domingensis Jacq.,
—Palicourea ochnoides Dwyer Enum. PI. Carib. 16. 1760. Type: SANTO
Palicourea chiriquina Standi., Contr. U.S. Natl. DOMINGO: Without precise locality, Jacquin
Herb. 18: 140. 1916. Type: PANAMA: (location unknown).—Psychotria
Chiriqui, Cuesta de las Palmas, southern slope domingensis Jacq.
of Cerro de la Horqueta, 1700-2100 m, 17-19 Palicourea galeottiana M. Martens, Bull. Acad.
Mar 1911, H. Pittier 3211 (Holotype US Roy. Sci. Bruxelles 11: 136. 1844. Type:
677607 (photo PTBG); Isotype US (photo MEXICO: Oaxaca, H. Galeotti 2602
PTBG)). (Lectotype BR (photos MEXU, PTBG);
Palicourea copensis (Dwyer) C. M. Taylor in Isolectotypes K, n.v., P (photos MEXU,
W. C. Burger & C. M. Taylor, Fieldiana, Bot., PTBG); designated by Taylor & Lorence,
n.s. 33: 204. 1993. Psychotria copensis 1985: 669).—Psychotria galeottiana (M.
Dwyer, Ann. Missouri Bot. Gard. 67: 365. Martens) C. M. Taylor & Lorence
1980. Type: PANAMA: Code, El Cope, El Palicourea garciae Standi., Publ. Field Mus.
Potroso, Atlantic slope of Alto Calvario, 26 Nat. Hist., Bot. Ser. 22: 195. 1940. Type:
Feb 1977, J. P. Folsom & R. Lantz 1880 COLOMBIA: El Valle, San Antonio, W of
(Holotype MO 2605731 (photos MEXU, Cali, near summit of Cordillera Occidental,
PTBG)). 1900-2350 m, Feb-Mar 1939, Killip &
Palicourea costaricensis Benth. in Oerst., Garcia 33660 (Holotype F; Isotypes A, n.v.,
Vidensk. Medd. Dansk. Naturhist. Foren. NY, n.v.).
Kjobenhavn 1852: 38. 1853. Type: COSTA Palicourea gardenioides (Scheidw.) Benth. &
RICA: Heredia, Volcan de Barba et Cerros de Hook. f. ex Hemsl., Biol. Centr.-Amer., Bot.
Candelaria, Oersted 11540 (Lectotype C, n.v. 2: 52. 1881. Rhodostoma gardenioides
(photo F); Isolectotypes MICH, n.v., US, n.v.; Schiedw. in Otto & Dietr., Allgem. Gartenz.
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL J

10: 286. 1842. Type: Based on cultivated Palicourea lanceolata Oerst. ex Pol., Vidensk.
material of unknown origin, presumably from Meddel. Dansk Narurhist. Foren. Kjobenhavn
Veracruz, Mexico; type not designated. 1852: 39. 1853. Type: COSTA RICA: San
—Psychotria gardenioides (Scheidw.) Jose, in monte Candelaria, 2100 m, 1845-48,
Standi. Oersted 11556 (Lectotype C, n.v. (photo &
Palicourea gibbosa Dwyer, Ann. Missouri Bot. fragment F); designated by Standley (1938);
Gard. 67: 304. 1980. Type: PANAMA: see Uragoga carpinterensis O. Kuntze).
Darien, Cerro Tacaracuna, West Ridge trail —Palicourea angustifolia Kunth in Humb.,
from summit camp to waterfall, E of camp, Bonpl. & Kunth
1550-1700 m, 2 Feb 1975, A. Gentry & S. Palicourea lancifera Standi. & L. O. Williams,
Mori 14108 (Holotype MO 2354276; Isotype Ceiba 1: 249. 1951. Type: COSTA RICA:
MO 2354275). Alajuela, Canton Alfaro Ruiz, Zarcero, 1500
Palicourea gomezii C. M. Taylor, Ann. m, 17 Jun 1941, A. Smith 2794 (Holotype US
Missouri Bot. Gard. 77: 215. 1990. Type: 2215921 ex EAP (photo PTBG); Isotype F).
COSTA RICA: Heredia, quebrada Palicourea lasiorrhachis Oerst., Vidensk.
Sangrijuela, 700 m, L. D. Gomez P., Chacon Meddel. Dansk Narurhist. Foren. Kjobenhavn
& Herrera 20896 (Holotype CR, n.v.; Isotype 1852: 39. 1853. Type: COSTA RICA: In
MO, n.v.). monte Candelaria, Feb 1847, Oersted 11557
Palicourea grandibracteata Dwyer, Ann. (Holotype C, n.v. (photos F, NY); Isotype US
Missouri Bot. Gard. 67: 305. 1980. Type: (photo PTBG)).
PANAMA: Veraguas, NW of Santa Fe, 2 km Palicourea leucantha Donn. Sm., Bot. Gaz.
from Escuela Agricola Alto de Piedra, cloud (Crawfordsville) 48: 295. 1909. Type:
forest on ridge top below summit of Cerro GUATEMALA: Alta Verapaz, between
Tute, 28 Mar 1975, S. Mori & J. Kallunki Coban and Tactic, 1800 m, Mar 1903, H. von
5262 (Holotype MO 2384999 (photo PTBG); Tuerckheim 8400 (Holotype US 943517
Isotype MO 2385000 (photo PTBG)). (photo PTBG); Isotype US (photo PTBG)).
—Palicourea ochnoides Dwyer Palicourea longibracteata Bartl. ex DC, Prodr.
Palicourea guianensis Aubl., Hist. PI. Guiane 4: 527. 1830. Type: PANAMA: Without
1: 173, t. 66. 1775. Type: FRENCH precise locality, Haenke s.n. (Holotype
GUIANA: In Guajanae sylvis de Caux, Aublet presumably PR, n.v.).—Palicourea triphylla
s.n. (Holotype BM, n.v.). DC.
Palicourea hammelii C. M. Taylor, Syst. Bot. Palicourea lusinaturalis Dwyer, Ann. Missouri
Monogr. 26: 43, fig. 22 c, d. 1989. Type: Bot. Gard. 53: 106. 1966. Type: PANAMA:
PANAMA: Near top of Continental Divide Colon, NW part of Canal Zone, area W of
ridge between Chiriqui and Bocas del Toro Limon Bay, Gatun Locks and Gatun Lake,
provinces near Cerro Pate Macho NE of Rio Pina-Rio Media divide, Johnston 1807
Boquete, 7100 ft., 15 Mar 1982, B. Hammel, (Holotype MO, not located 6/96).
W. J. Kress, S. Knapp, Trainer & Will 11408 —Psychotria luxurians Rusby
(Holotype DUKE, n.v.; Isotype MO (photo Palicourea macrantha Loes., Verh. Bot.
PTBG)). Vereins Brandenburg 65: 113. 1923. Type:
Palicourea intermedia Oerst. ex Pol., Linnaea GUATEMALA: Huehuetenango, Nenton
41: 571. 1877. Type: COSTA RICA: Forests District, near Yalambohoch, Aug 1896, E.
of La Carpintera, Polakowsky 143 (Holotype Seler & C. Seler 2871 (Holotype B,
B, presumably destroyed (photo F); Isotype presumably destroyed (photos F, MO, NY,
W, n.v. (fragment F)).—Palicourea US); fragment F).
angustifolia Kunth in Humb., Bonpl. & ocalyx Standi., J. Wash. Acad.
Kunth : 278. 1928. Type: COSTA RICA:
a, Cerro de Las Lajas, north of San
 Isidro, 2000-2200 m, 7 Mar 1926, P. C. Palicourea ochnoides Dwyer, Ann. Missouri
Standley & J. Valerio 51611 (Holotype US Bot. Gard. 67: 310. 1980. Type: PANAMA:
1306801 (photo PTBG)). Veraguas, trail up E side of Cerro Tute, to
Palicourea macrosepala K. Krause, Bot. Jahrb. 1200 m, 25 Oct 1975, J. T. Witherspoon, F.
Syst. 54, Beibl. 119: 41. 1916. Type: COSTA Witherspoon & R. Dressier 8872 (Holotype
RICA: Forets de Alto del Pito (Ojo de Agua), MO 2302202 (photo PTBG)).
pres de San Marcos, 1000 m, Mar 1893, A. Palicourea orosiana C. M. Taylor, Syst. Bot.
Tonduz 7704 (Holotype B, presumably Monogr. 26: 6\,fig. 33 a, b. 1989. Type:
destroyed (photos F, MO, US; fragment F); COSTA RICA: Cartago, about 10 km south of
Isotype BR, n.v.).—Palicourea discolor K. Tapanti along new road on east slope above
the Rio Grande de Orosi, 1600 m, 14 & 17 Jul
Palicourea mexicana Benth. in Oerst., Vid. 1971, W. C Burger & M. Burger 7574
Meddel. Dansk Naturhist. Foren. Kjobenhavn (Holotype F 1940946 (photo PTBG); Isotype
1852: 37. 1853. Type: COSTA RICA: In MO (photo PTBG)).
monte Barba, Oersted 11559 (Lectotype C, Palicourea padifolia (Willd. ex Roem. &
n.v.; designated by Standley (1938); see Schult.) C. M. Taylor & Lorence, Taxon 34:
Uragoga cordillerae O. Kuntze). 669. 1985. Psychotria padifolia Willd. ex
—Palicourea padifolia (Willd. ex Roem. & Roem. & Schult., Syst. Veg. 5: 189. 1819.
Schult.) C. M. Taylor & Lorence Type: MEXICO: Veracruz, Xalapa, Humboldt
Palicourea molliramis K. Schum. & K. Krause, & Bonpland 4420 (Holotype B-WI (photos
Engl. Bot. Jahrb. 40: 337. 1908. Type: DUKE, F, MO, NY)).
COLOMBIA: Cauca, prope La Junta ad Rio Palicoureapanamensis Standi., Ann. Missouri
d'Agua, 200-500 m, Lehmann 4667 Bot. Gard. 25: 839. 1938. Type: PANAMA:
(Holotype B, presumably destroyed). Chiriqui, Valley of upper Rio Chiriqui Viejo,
/tycAo/r/a.—Psychotria hebeclada DC. 1300-1900 m, Jul-Aug 1937, G. White & P.
Palicourea montensis Dwyer, Ann. Missouri White 14 (Holotype F 913062 (photo PTBG);
Bot. Gard. 67: 309. 1980. Type: PANAMA: Isotypes A, n.v., MO (photo PTBG)).
Chiriqui, Bocas del Toro-Chiriqui border —Palicourea discolor K. Krause
region, on Cerro Colorado copper mine road, Palicourea parvijlora Benth., Bot. Voy. Sulphur
5000-5500 ft., 4 May 1975, S. Mori & J. 107. 1845. Type: PANAMA: Island off the
Kallunki 5880 (Holotype MO).—Psychotria coast of Veraguas, Barclay & Hinds s.n.
croceovenosa Dwyer (which is based on the (Holotype BM, n.v.).—Palicourea triphylla
DC.
placement for this species.) Palicourea pauciflora Standi., J. Wash. Acad.
Palicourea montivaga Standi., J. Wash. Acad. Sci. 18: 278. 1928. Type: PANAMA:
Sci. 18: 279. 1928. Type: COSTA RICA: Chiriqui, between Alto de las Palmas and top
Heredia, Yerba Buena, northeast of San of Cerro de la Horqueta, 2100-2265 m, 18
Isidro, 2000 m, 28 Feb 1926, P. C. Standley & Mar 1911, H Pittier 3237 (Holotype US
J. Valerio 49850 (Holotype US 1306225 677618 (photo PTBG); Isoytpe F (photo
(photo PTBG, fragment F)). PTBG)).
Palicourea nigrescens M. Martens 8c Galeotti, Palicourea pendula C. M. Taylor, Syst. Bot.
Bull. Acad. Roy. Soc. Sci. Bruxelles 11: 136. Monogr. 26: 68, fig. 37 a-d. 1989. Type:
1844. Not Psychotria nigrescens De Wild., PANAMA: Veraguas, ridge of Cordillera de
1924. Type: MEXICO: Veracruz, Cordillera, Tute, trail to Cerro Tute above Escuela
Xalapa, 4000 pieds, Jun-Oct 1840, H. Agricola Alto de Piedra, just W of Santa Fe,
Galeotti 2653 (Holotype BR, n.v. (photo F); 800-1350 m, 5 Jun 1982, S. Knapp & R.
Isotype P (photos MEXU, PTBG)). Dressier 5409 (Holotype MO 3041797 (photo
—Psychotria mexiae Standi. PTBG)).
 t NOMENCLATOR OF MEXICAN AND CENTRAL

Palicourea phanaerandra Standi. & Steyerm., Palicourea skotakii C. M. Taylor, Syst. Bot.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 252. Monogr. 26: 76, fig. 41. 1989. Type: COSTA
1947. Type: GUATEMALA: Izabal, Cerro RICA: San Jose, along a road to the S from
San Gil, uppermost ridges and summit, the Panamerican Highway at km 67, about 1
1200-1300 m, 26-27 Dec \9A\,J.A. km along the road, 2700-2800 m, 5 Mar
Steyermark 41952 (Holotype F 1132612 1985, C. M. Taylor & C. Skotak 4756
(photo PTBG); Isotype US).—Psychotria (Holotype DUKE, n.v.; Isotypes F (photo
phanaerandra (Standi. & Steyerm.) Lorence PTBG), MO (photo PTBG)).
Palicourea purpurea C. M. Taylor, Syst. Bot. Palicourea spathacea C. M. Taylor, Syst. Bot.
Monogr. 26: 71, fig. 33 c, d. 1989. Type: 9: 226. 1984. Type: COSTA RICA: Cartago,
PANAMA: Chiriqui, NW of Boquete along along the trail about 6 km beyond the gate to
road to Palo Alto Arriba, from 2 km until road the meteorology station in Alto Patillos,
ends then another 0.5 km by trail, 1400-1600 Tapanti Hydroelectric Project, 7 Apr 1976, J.
m, 13 May 1984, C M. Taylor 3201 Vtley & K. Utley 4538 (Holotype DUKE, n.v.;
(Holotype DUKE, n.v.; Isotypes F (photo Isotypes CAS, n.v., CR, n.v., F (photo PTBG),
PTBG), MO (photo PTBG)). US (photo PTBG)).
Palicouraea purpusii Standi., Publ. Field Palicourea standleyana C. M. Taylor, Syst.
Columbian Mus., Bot. Ser. 8: 67. 1930. Type: Bot. Monogr. 26: 81, fig. 45 d-f. 1989. Norn,
MEXICO: Veracruz, Zacuapam [Zacuapan], nov. for Psychotria brenesii Standi., Publ.
shady forest, May 1928, C. A. Purpus 10110 Field Mus. Nat. Hist., Bot. Ser. 18: 1347.
(Holotype F 602907 (photos MEXU, PTBG); 1938. Not Palicourea brenesii Standi., 1938.
Isotype MO (photos MEXU, PTBG)). Type: COSTA RICA: Alajuela, Los Angeles
—Palicourea macrantha Loes. de San Ramon (Cerros de La Balsa), Mar
Palicourea rigidifolia (Dwyer & Hayden) 1928, Brenes 6050 (Holotype F 854521
Dwyer, Ann. Missouri Bot. Gard. 67: 313. (photos MEXU, PTBG)).
1980. Cephaelis rigidifolia Dwyer & Hayden, Palicourea stevensonii Standi., Trop. Woods.
Ann. Missouri Bot. Gard. 55: 37. 1968. Type: 16: 42. 1928. Type: BRITISH HONDURAS
PANAMA: Panama, Cerro Jefe, 2700 ft., 20 (BELIZE): Middlesex, Jul 1927, N. S.
Aug 1967, J. D. Dwyer 8075A (Holotype MO Stevenson C(Yale 10683) (Holotype F
2165097 (photo PTBG)).—Status uncertain, 572673; Isotype NY, n.v.).—Psychotria
probably Psychotria subg. Heteropsychotria capitata Ruiz & Pav.
Palicourea salicifolia Standi., J. Wash. Acad. Palicourea subrubra Pol., Linnaea 41: 571.
Sci. 18: 280. 1928. Type: COSTA RICA: San 1877. Type: COSTA RICA: San Jose, Cerro
Jose, Laguna de La Chonta, northeast of Santa de La Carpintera, 24 Jun 1875, Polakowsky
Maria de Dota, 2000-2100 m, 18 Dec 1925, 200A (Holotype B, presumably destroyed
P. C. Standley 42174 (Holotype US 1252646 (photo F)).—Palicourea padifolia (Willd. ex
(photo PTBG)). Roem. & Schult.) C. M. Taylor & Lorence
Palicourea seleri Loes., Verh. Bot. Vereins Palicourea talamancana Standi. & L. O.
Prov. Brandenburg 65: 114. 1923. Type: Williams, Ceiba 1: 250. 1951. Type: COSTA
GUATEMALA: Huehuetenango, zwischen RICA: Cartago, Cordillera de Talamanca, near
Trinidad und Rosario, 13 Aug 1896, E. Seler Vuelta Alta, Cerro de las Vueltas, 3200 m, 10
& C. Seler 3049 (Syntype B, destroyed Apr 1949, L. O. Williams 16264 (Holotype
(photos F, GH, MO, NY, US), fragment F, EAP, n.v.; Isotypes F (photo PTBG), GH,
GH, n.v.); Yalambahoch, E. Seler & C. Seler n.v., US (photo PTBG)).—Palicourea
2676 (Syntype B, destroyed).—Psychotria brenesii Standi.
galeottiana (M. Martens) C. M. Taylor & Palicourea tilaranensis C. M. Taylor, Syst.
Bot. Monogr. 26: S4, fig. 3 a, b. 1989. Type:
COSTA RICA: Puntarenas-Alajuela border
 region, in and around Monteverde Nature MA, n.v.; probable Isotype F).—Rondeletia
Reserve, mostly on Pacific watershed,
10°18'N, 84°47*N, 1450-1650 m, 29 Jun
1977, R. O. Lawton 1189 (Holotype F Pentagonia Benth., Bot. Voy. Sulphur /. 39.
1885069 (photo PTBG)). 1804; 105. 1845 (nom. cons.). Type:
Palicourea triphylla DC, Prodr. 4: 526. 1830. Pentagonia macrophylla Benth.
Type: FRENCH GUIANA: Guiana Gallica, Pentagonia alba Dwyer, Ann. Missouri Bot.
without precise locality, Patris s.n. (Holotype Gard. 67: 321. 1980. Type: PANAMA:
G-DC, n.v. (photo MO)). Psychotria. Darien, Cerro Pirre, Bristan 1224 (Holotype
Palicourea tubiflora Dwyer, Ann. Missouri MO, not located 6/1996; Isotype NY, n.v.).
Bot. Gard. 67: 316. 1980. Type: PANAMA: Pentagonia alfaroana (Standi.) Standi., J. Wash.
Panama, Cerro Jefe, along trail on ridge Acad. Sci. 17: 171. 1927. Watsonamra
running NE from summit, ca. 1000 m, 11 May alfaroana Standi., J. Wash. Acad. Sci. 15:
1975, S. Mori & J. Kallunki 6082 (Holotype 287. 1925. Type: COSTA RICA: Alajuela,
MO 2271726 (photo PTBG)). vicinity of Capulin, on the Rio Grande de
Palicourea tumidonodosa Dwyer, Ann. Tarcoles, 80 m, 2 Apr 1924, P. C. Standley
Missouri Bot. Gard. 67: 317. 1980. Type: 40194 (Holotype US 1152712 (photos
PANAMA: Panama, Cerro Mali, base camp MEXU, PTBG); Isotype F).—Pentagonia
near Colombian border, lower montane wet tinajita Seem.
forest, 1500 m, 22 Jan 1975, A. Gentry & S. Pentagonia brachyotis (Standi.) Standi., J.
Mori 13787 (Holotype MO 2259310 (photo Wash. Acad. Sci. 17: 171. 1927. Watsonamra
PTBG)). brachyotis Standi., Contr. U.S. Natl. Herb. 17:
Palicourea veraguensis Dwyer, Ann. Missouri 443. 1914. Type: PANAMA: Darien,
Bot. Gard. 67: 318. 1980. Type: PANAMA: Marraganti, 3 Apr 1908, R. S. Williams 999
Chiriqui, Cerro Colorado, 50 km N of San (Holotype US 678351 (photos MEXU,
Felix on the continental divide, 1200-1500 m, PTBG)).
17 Aug 1975, S. Mori & R. Dressier 7796 Pentagonia costaricensis (Standi.) W. C.
(Holotype MO 2777356 (photo PTBG)). Burger & C. M. Taylor, Fieldiana, Bot., n.s.
—Palicourea lasiorrhachis Oerst. 33: 213. 1993. Nothophlebia costaricensis
Palicourea vestita Standi., J. Wash. Acad. Sci. Standi., Contr. U.S. Natl. Herb. 17: 438. 1914.
18: 277. 1928. Type: COSTA RICA: San Type: COSTA RICA: Collines de Moin,
Jose, near Quebradillas, about 7 km north of littoral atlantique, Nov 1899, H. Pittier 16024
Santa Maria de Dota, 1800 m, 24 Dec 1925, (Holotype US 578472 (photos MEXU,
P. C. Standley 42909 (Holotype US 1253015 PTBG)).
(photos F, PTBG)). Pentagonia donnell-smithii (Standi.) Standi., J.
Wash. Acad. Sci. 17: 170. 1927. Watsonamra
Pavetta L., Sp. PL 1: 110. 1753. Type: Pavetta donnell-smithii Standi., Contr. U.S. Natl.
indica L. Herb. 17: 442. 1914. Type: COSTA RICA: La
Pavetta odoratissima Sesse & Moc., Fl. Mexic. Emilia, Llanuras de Santa Clara, 250 m, Apr
20. 1893; La Naturaleza, ser. 2, 2: app. 20. 1896, J. D. Smith 6590 (Holotype US 355176
1893. Type: MEXICO ?: Habitat in Praedio (photos MEXU, PTBG); Isotype F).
Sacarmo de la Langosta, Equitis Gonzalo de Pentagonia gymnopoda (Standi.) Standi., J.
Herrera, floret Agusto, Sesse & Mocino s.n. Wash. Acad. Sci. 17: 171. 1927. Watsonamra
(presumably MA, n.v.).—Status uncertain. gymnopoda Standi., Contr. U.S. Natl. Herb.
Pavetta racemosa Sesse & Mo?., Fl. Mexic. 20. 17: 444. 1914. Type: PANAMA: Colon,
1893; La Naturaleza, ser. 2, 2: app. 2, 18. Loma de la Gloria, near Fato (Nombre de
1893. Type: MEXICO: Without precise Dios), 10-104 m, 8 Jul 1911, H. Pittier 3858
locality, Sesse & Mocino 543 (presumably (Holotype US 678935 (photos MEXU,
 L NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

PTBG); Isotype US 678934 (photos MEXU, Pentagonia tinajita Seem., Bot. Voy. Herald
PTBG)). 134, t. 28. 1854. Type: PANAMA: Veraguas,
Pentagonia hirsuta Standi., J. Wash. Acad. Sci. western parts, Seemann 1595 (Holotype BM,
18: 165. 1928. Type: COSTA RICA: n.v. (photo MO)).
Talamanca, forets au-dessus de Tsaki, 500 m, Pentagonia veraguensis Dwyer, Ann. Missouri
Mar 1895, A. Tonduz 9415 (Holotype US Bot. Gard. 67: 328. 1980. Type: PANAMA:
938648 (photos MEXU, PTBG); Isotype F). Veraguas, Valley of Rio Dos Bocas, 11 km
Pentagonia Iobata C. M. Taylor, Novon 5: 202, from Escuela Agricola Alto Piedra (above
fig. ID. 1995. Type: COSTA RICA: Santa Fe) on road to Calovebora, 450 m, 30
Puntarenas. Canton de Golfito, Reserva Aug 1974, T. B. Croat 27504 (Holotype MO
Natural de Vida Silvestre Golfito, final del 2201425 (photo PTBG)).
camino hacia las torres, 8°39*52"N, Pentagonia wendlandii Hook, f, Bot. Mag. pi.
83°10'00"W, 7 Dec 1992, B. Hammel 18619 5230. 1861. Type: CENTRAL AMERICA
(Holotype MO 4355639; Isotypes CR, n.v., F, [probably Costa Rica or Panama]: Described
n.v., MO 4355638).
Pentagonia macrophylla Benth., Bot. Voy.
Sulphur 105, tab. 39. 1845. Type: PANAMA:
Panama, forests between Panama and Cruces, Pentas, Benth. in Hook., Bot. Mag. 4086. 1844.
Barclay s.n. (Holotype BM, n.v.). Type: Pentas carnea Benth.
Watsonamra. Pentas lanceolata (Forssk.) Deflers, Voy.
Pentagonia nuciformis Dwyer, Ann. Missouri Yemen 142. 1889. Ophiorhiza lanceolata
Bot. Gard. 67: 326. 1980. Type: PANAMA: Forssk., F. Aegypt.-Arab. 42. 1775. Type:
Veraguas, 5 miles W of Santa Fe on the road AFRICA: Yemen, Hadie Mts., Forssk&ls.n.
past Escuela Agricola Alto Piedra on Pacific (Holotype C, n.v.; Isotype BM, n.v.). Widely
side of divide, 800-1200 m, 18 Mar 1973, cultivated in Mexico and Mesoamerica.
T. B. Croat 23093 (Holotype MO 2161493
(photo PTBG)). Pentodon Hochst., Flora 27: 552. 1844. Type:
Pentagonia pinnatifida Seem., J. Bot. Hooker Pentodon decumbens Hochst.
7: 566. 1848. Type: PANAMA: Darien, banks Pentodon pentandrus (Schumach. & Thonn.)
of the river Cupica, Seemann 1019 (Holotype Vatke, Osterr. Bot. Zeitschr. 25: 231. 1875.
BM, n.v. (photo MO)). Hedyotispentandra Schumach. & Thonn.,
Pentagonia pittieri (Standi.) Standi., J. Wash. Kongel. Danske Vidensk. Selsk.
Acad. Sci. 17: 170. 1927. Watsonamra pittieri Naturvidensk., Math. Afh. 3: 71. 1827. Type:
Standi., Contr. U.S. Natl. Herb. 17: 443. 1914. GUINEA: Aquapim, "S" [Haaslund Smith?]
Type: PANAMA: San Bias, forests around (location unknown). Naturalized at least in
Puerto Obaldia, San Bias coast, 0-50 m, Aug Mexico (Veracruz), Belize, Nicaragua, and
1911,//. Pittier 4298 (Holotype US 679414 Costa Rica.
(photos MEXU, PTBG); Isotype US 679415
(photos MEXU, PTBG)).—Pentagonia Petesia P. Browne, Civ. Nat. Hist. Jamaica 143,
pinnatifida Seem. t. 2, 3. 1756. Type not designated.
Pentagonia pubescens (Standi.) Standi., J. Petesia grandis Bartl. ex DC, Prodr. 4: 395.
Wash. Acad. Sci. 17: 171. 1927. Watsonamra 1830. Type: MEXICO: Without precise
pubescens Standi., Contr. U.S. Natl. Herb. 17: locality, Haenke s.n. (Holotype presumably
441. 1914. Type: PANAMA: Canal Zone, PR, n.v., fragment G-DC, n.v. (microfiche 4:
around Tabernilla, along the railroad, 20-25 395.1)).—Sommera grandis (Bartl. ex DC.)
m, 6 Jul 1911, //. Pittier 3822 (Holotype US Standi.
678894 (photo PTBG); Isotype NY, n.v.). Petesia minor Griseb., Bonplandia 6: 8. 1858.
—Pentagonia macrophylla Benth. Type: PANAMA: Without precise locality,
 . (Holotype presumably BM, Placocarpa Hook. f. in Benth. & Hook, f., Gen.
i donnell-smithii Standi. PI. 2: 107. 1873. Type: Placocarpa mexicana
Petesia repens Sesse & Moc., Fl. Mexic. 21. Hook. f.
1893. Type: MEXICO: Habitat in Ahualuci Placocarpa mexicana Hook. f. in Benth. &
pratis, florit Octobris, Sesse & Mociho s.n. Hook, f., Gen. PI. 2: 108. 1873. Type:
(presumably MA, n.v.).—Status uncertain. MEXICO: Veracruz, Orizaba, Apr 1855,
Petesia stipularis Sesse & M09., Fl. Mexic. 21. Botteri 913 (Lectotype P, (photo PTBG), here
1893, La Naturaleza 2, 2: App. 2, 16. 1893. designated; Isolectotypes F (fragment), K
Not Petesia stipularis L., 1759. Type (listed (photo PTBG). Syntypes: MEXICO:
twice): MEXICO: Habitat in Montibus Veracruz, Orizaba, Botteri 316 (F, K (photo
Mazatlanensibus, floret Maio, habitat in PTBG)); Bourgeau 3022 (K (photo PTBG)).
Acahuitzotlae montibus, floret Julio, Sesse & The sheet of Botteri 913 at P, with duplicates
Mociiio s.n. (presumably MA, n.v.).—Status in two other herbaria and copious flowers and
uncertain, possibly Chomelia or Sommera. fruits, represents the most appropriate choice
Protologue states leaves are prominently for lectotype.
lineolate beneath and inflorescence is once
dichotomous. Plocaniophyllon Brandegee, Univ. Calif. Publ.
Bot. 6: 69. 1914. Type: Plocaniophyllon
Pinarophyllon Brandegee, Univ. Calif. Publ. flavum Brandegee
Bot. 6: 71. 1914. Type: Pinarophyllon flavum Plocaniophyllon flavum Brandegee, Univ.
Brandegee Calif. Publ. Bot. 6: 69. 1914. Type: MEXICO:
Pinarophyllon bullatum Standi., N. Amer. Fl. Chiapas, Finca Mexiquito, Jun 1913, C. A.
32: 87. 1921. Type: GUATEMALA: Alta Purpus 7019 (Holotype UC 173077, n.v.;
Verapaz, between Sepacuite and Secoyocte, Isotypes F (photos MEXU, PTBG), US
1100 m, 24 May 1905, H. Pittier 348 (photos MEXU, PTBG)).
(Holotype US 473013 (photos MEXU, PTBG,
US)). Pogonopus Klotzsch, Monatsber. Konigl.
Pinarophyllon flavum Brandegee, Univ. Calif. Preuss. Acad. Wiss. Berlin 1853: 500. 1853.
Publ. Bot. 6: 71. 1914. Type: MEXICO: Type: Pogonopus ottonis Klotzsch
Chiapas, Finca Mexiquito, Jun 1913, C /f Pogonopus exsertus (Oerst.) Oerst, Amer.
Purpus 6700 (Holotype UC 172447, n.v.; Centr. 17. t. 13. 1863. Macrocnemum
Isotypes F (photo PTBG), GH, n.v., US exsertum Oerst., Vidensk. Meddel. Naturhist.
(photos MEXU, PTBG)). Foren Kjobenhavn 1852: 45. 1853. Type:
COSTA RICA: Jaris, Oersted (Holotype C,
Pittoniotis Griseb., Bonplandia 6: 8. 1858. Type: n.v.). Note: Pogonopus speciosus (Jacq.) K.
Pittoniotis tnchantha Griseb. Schum., a species from northern South
Pittoniotis protracta (Bartl. ex DC.) Griseb., America, was erroneously recorded from
Bonplandia 6: 8. 1858. Guettarda protracta Central America by Standley & Williams
Bartl. ex DC, Prodr. 4: 457. 1830. Type: (1975) and Dwyer (1980).
MEXICO: Without precise locality, Haenke
s.n. (Holotype presumably PR, n.v.). Portlandia P. Browne, Hist. Jamaica 164, t. 11.
Anisomeria.—Chomelia protracta (Bartl. ex 1756. Type: Portlandia grandiflora L.
DC.) Standi. Portlandia ghiesbreghtiana Baill., Adansonia
Pittoniotis trichantha Griseb., Bonplandia 6: 8. 12: 300. 1879. Type: MEXICO: Oaxaca, pres
1858. Type: PANAMA: Without precise de Hacienda de Huijastla au bord des ravine,
locality, Duchassaing s.n. (possibly P, n.v.). 1842-3, Ghiesbreght 27 (Holotype P (photos
Antirhea. MEXU, PTBG); Isotypes F, P (3 sheets;
 OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

photos MEXU), US (photos MEXU, PTBG)). 2000-4000 ft., H. Galeotti 1580 (Holotype
—Coutaportla ghiesbreghtiana (Baill.) Urb. BR, n.v.; Isotype P (photos F, MEXU, MO)).
guatemalensis Standi., J. Wash. Posoqueria grandiflora Standi., J. Wash. Acad.
Acad.Sci. 18: 162. 1928. Type: Sci. 18: 166. 1928. Type: COSTA RICA:
GUATEMALA: Alta Verapaz, Quebradas Limon, wet thicket at Finca Hamburg on the
Secas, 2500 ft., 1 Jun 1920, H. Johnson 282 Rio Reventazon below El Cairo, 55 m, 19 Feb
(Holotype US 1081354 (photos MEXU, 1926, P. C. Startdley & J. Valeria 48706
PTBG)).—Coutaportla guatemalensis (Holotype US 1305673 (photos MEXU,
(Standi.) Lorence PTBG)).
Portlandia hexandra Jacq., Enum. PL Carib. 16. Posoqueria latifolia (Rudge) Roem. & Schult.,
1760. Type: COLOMBIA: Bolivar, vicinity of Syst. Veg. 5: 227. 1819. Solena latifolia
Cartagena, Jacquin (location unknown). Rudge, PL Guian. 1: 26. 1805; t. 40. 1806.
—Coutarea hexandra (Jacq.) K. Schum. Type: FRENCH GUIANA: Martin s.n. (BM,
Portlandia lumaeana (Baill.) Baill., Hist. PI. 7:
333. 1880. Coutarea lumaeana Baill., Posoqueria obliquinenia Standi., J. Wash.
Adansonia 12: 301. 1879. Type: Acad. Sci. 18: 167. 1928. Type: COSTA
GUATEMALA: Cote occidentale des Andes, RICA: Forets et bords du Rio Naranjo,
450-500 m, 1878, G. Luma s.n. (Holotype P 200-250 m, Mar 1893, A. Tonduz 7528
(photos MEXU, PTBG)).—Hintonia [published erroneously as 9528} (Holotype
lumaeana (Baill.) Bullock US 764158 (photos MEXU, PTBG)).
Portlandia mexicana (Zucc. & Mart.) Hemsl., —Tocoyena pittieri (Standi.) Standi.
Diagn. PI. Nov. 31. 1879. Coutarea mexicana Posoqueria panamensis (Walp. & Duchass.)
Zucc. & Mart, ex DC, Prodr. 4: 350. 1830. Walp., Ann. Bot. Syst. 2: 797. 1852. Stannia
Type: MEXICO: Without precise locality, panamensis Walp. & Duchass., Linnaea 23:
1829, Martius s.n. (Holotype G-DC, n.v. 755. 1850. Type: PANAMA: Isthmo
(microfiche IDC 800.680. II.7).—Nernstia Panamensi, Duchassaing ? (location
mexicana (DC.) Urb.
Portlandia pterosperma S. Watson, Proc. Amer. Posoqueria pittieri Standi., J. Wash. Acad. Arts
Acad. Arts 24: 52. 1889. Type: MEXICO: Sci. 18: 167. 1928. Type: PANAMA: Canal
Sonora, Guaymas, 1887, E. Palmer 298 Zone, near Hydrographic Station on Trinidad
(Holotype US 47978 (photos MEXU, PTBG); River, 17 May 1914, H. Pittier 6635
Isotype US 47979 (photos MEXU, PTBG)). (Holotype US 716697 (photos MEXU,
Coutarea.—Hintonia latiflora (Sesse & Moc. PTBG); Isotype P, n.v. (fragment F)).
ex DC.) Bullock —Tocoyena pittieri (Standi.) Standi.
Portlandia speciosa (Aubl.) Baill., Hist. PI. 7:
381. 1879. Coutarea speciosa Aubl., Hist. PI.
Guiane 1: 314, t. 122. 1775. Type: FRENCH
GUIANA: Aublet s.n. (presumably P, n.v.).
—Coutarea hexandra (Jacq.) K. Schum. Psychotria acicularis C. M. Taylor in W. C.
Burger & C. M. Taylor, Fieldiana, Bot., n.s.
Posoqueria Aubl., Hist. PL Guiane 1: 133. 1775. 33: 231. 1993. Type: COSTA RICA:
Type: Posoqueria longiflora Aubl. Puntarenas, Rmcon de Osa, region to
Posoqueria coriacea M. Martens & Galeotti, southwest of airstrip, 20-60 m, 20 Jul 1974, J.
Bull. Acad. Roy. Sci. Bruxelles 11: 240. 1844. Utley & K. Utley 1036 (Holotype CR, n.v.;
Type: MEXICO: Oaxaca, Cordillere orientale, Isorypes F, n.v., MO).
forets humides de la Chinantla, pres des Psychotria acuminata Benin., Bot. Voy.
bourgs indiens de Teotalcingo et Choapam, Sulphur 107. 1845. Type: COLOMBIA: Isle
 of Gorgona, and Colombian coast, Barclay Psychotria allenii Standi., Ann. Missouri Bot.
s.n. (Holotype BM, n.v.). Gard. 27: 342. 1940. Type: PANAMA: Code,
Psychotria acuminata Benth. subsp. vicinity of El Valle, 600-1000 m, 8 Dec 1938,
boraginoides Dwyer, Ann. Missouri Bot. P. H. Allen 1796 (Holotype F 989006 (photos
Gard. 67: 344. 1980. Type: PANAMA: MEXU, PTBG); Isotypes GH, n.v., NY, n. v.,
Panama, El Llano Carti-Tupile road, 12 mi. US (photos MEXU, PTBG)).
above Panamerican Highway, 26-27 Mar Psychotria altorum Standi. & Steyerm., Publ.
1973, R. L. Liesnerll87 (Holotype MO Field Mus. Nat. Hist., Bot. Ser. 23: 86. 1944.
2164366).—Psychotria acuminata Benth. Type: GUATEMALA: Quezaltenango,
Psychotria aggregata Standi., Contr. U.S. Natl. Montana Chicharro, on lower southeast-facing
Herb. 18: 128. 1916. Type: PANAMA: slopes of Volcan Santa Maria, 2-4 miles south
Clin lqui, between Alto de las Palmas and of Santa Maria de Jesus, 1400-1500 m, 17 Jan
Cerro de la Horqueta, 2100-2268 m, 18 1940, J. A. Steyermark 34302 (Holotype F
March 1911,//. Pittier 3264 (Holotype US 1056806 (photos MEXU, PTBG)).
677655 (photos MEXU, PTBG)). —Psychotria costivenia Griseb. var. altorum
Psychotria aguilarii Standi. & Steyerm., Publ. (Standi. & Steyerm.) C. W. Ham.
Field Mus. Nat. Hist., Bot. Ser. 23: 23. 1943. Psychotria americana Sesse & Moc., Fl. Mexic,
Type: GUATEMALA: Chimaltenango, on ed. 2, 57. 1894. Type: MEXICO: Habitat in
road between Chimaltenango and San Martin montibus calidarum regionum, ut Tenampulco
Jilotepeque, 1500-1700 m, 22 Dec 1940, prope Huastecam et Acauizotla versus
P. C. Standley 80900 (Holotype F 1092416 Acapulcum, Sesse & Mocino s.n. (presumably
(photos MEXU, PTBG)).
Psrchotna alba Ruiz & Pav. var. mexicana Psychotria angustiflora K. Krause, Bot. Jahrb.
Oerst, Vidensk. Meddel. Dansk Naturhist. Syst. 54, Beibl. 119: 43. 1916. Type: COSTA
Foren. Kjobenhavn 1852: 12. 1853. Syntypes: RICA: Sur les collines autour de Las Vueltas,
MEXICO: South Mexico, Jecaltepec, F. Tucurrique, 600-700 m, Jan 1899, A. Tonduz
Liebmann 17 (presumably C, n.v.); Cazadera, 12996 (Holotype B, presumably destroyed
F. Liebmann 18 (presumably C, n.v.); Pital. F (photos F, MO); Isotypes P (photos MEXU,
Liebmann 23 (presumably C, n.v.).—Status PTBG), US (photos MEXU, PTBG)).
Psychotria anomothyrsa K. Schum. & Donn.
Standi., Contr. U.S. Natl. Sm., Bot. Gaz. (Crawfordsville) 35: 3. 1903.
Herb. 18: 131. 1916. Type: PANAMA: Type: GUATEMALA: Alta Verapaz,
Chagres, Isthmus of Panama, Feb-Mar 1850, Cubliquitz, 350 m, Dec 1900, H. von
Fendler 62 (Holotype US 47282 (photos Tuerckheim 7909 (Holotype US 943471
MEXU, PTBG); Isotype MO).—Psychotria (photos MEXU, PTBG); Isotypes GH, n.v.,
capitata Ruiz & Pav. US (photos MEXU, PTBG)).—Psychotria
Psychotria alboviridula K. Krause, Notizbl. macrophylla Ruiz & Pav.
Konigl. Bot. Gard. Berlin 6: 208. 1914. Type: Psychotria arborea Sesse & Moc., Fl. Mexic,
BRAZIL: Im Gebiet des Alto Acre, Seringal, ed. 2, 57. 1894. Type: MEXICO: Habitat in
Sao Francisco, Sep 1911, Ule 9846 (Holotype Ahualuci atque calidis Guatemalae montibus.
B, presumably destroyed (photo F, fragment Floret Maio, Sesse & Mocino s.n. (presumably
F); Isotype US).—Psychotria remota Benth. MA, n.v.).—Status uncertain.
Psychotria alfaroana Standi, J. Wash. Acad. Psychotria aubletiana Steyerm., Mem. New
Sci. 18: 273. 1928. Type: COSTA RICA: York Bot. Gard. 23: 694. 1972. Cephaelis
Guanacaste, El Arenal, 485-600 m, 18-19 Jan axillaris Sw., Prodr. 45. 1788. Not Psychotria
1926, P. C. Standley & J. Valerio 45179 axillaris Willd., 1798, nor Veil., 1827. Type:
(Holotype US 1253966 (photos MEXU, WEST INDIES: St. Christopher, Swartz s.n.
PTBG)). (Holotype presumably S, n.v.).
A NOMENCLATOR OF MEXICAN AND CENTRAL

I aubletiana Steyerm. var. centro- PTBG); Isotype F (photos MEXU, PTBG)).

americana Steyerm., Mem. New York Bot. —Palicourea bella (Standi.) Dwyer
Gard. 23: 694. 1972. Type: HONDURAS: Psychotria berteriana DC, Prodr. 4: 515.
Morazan, La Tigra, entre El Piliguin y Prado 1830. Type: SANTO DOMINGO:
de Fatima, 1800 m, 30 Die 1962, A. Molina R. Dominique, without precise locality, Bertero
11237 (Holotype NY, n.v.; Isotype F (photos s.n. (a specimen at G-DC, n.v. (photo MO)
MEXU, PTBG)).—Psvchotria aubletiana likely represents authentic material).
Steyerm. Psychotria berteriana DC. subsp. luxurious
Psychotria aurantibractea C. M. Taylor in (Rusby) Steyerm., Mem. New York Bot.
W. C. Burger & C. M. Taylor, Fieldiana, Bot., Gard. 3: 534. 1972. Type: BOLIVIA:
n. s. 33: 235. 1993. CephaelispittieriK. Between Tipuani and Guanai, Dec 1892, Bang
Krause, Bot. Jahrb. Syst. 54, Beibl. 119: 45. 1741 (Holotype US, n.v.).—Psychotria
1916. Not Psychotria pittieri Standi., 1916. luxurians Rusby
Type: COSTA RICA: Puntarenas, Cafias Psyt hotria biaristata Bartl. ex DC, Prodr. 4:
Gordas, 1100 m, Feb 1897, H. Pittier 11162 513. 1830. Type: MEXICO: Without precise
(Holotype B, destroyed (photos F, MO, locality, Haenke s.n. (Holotype presumably
fragment F); Isotype US (photos MEXU, PR, n.v.).—Psychotria graciliflora Benth. ex
PTBG)). Oerst.
Psychotria aureola Bartl. ex DC, Prodr. 4: 513. Psychotria bimea L. Riley, Kew Bull. 1927:
1830. Type: MEXICO: Without precise 124. 1927. Type: PANAMA: Between
locality, Haenkes.n. (Holotype presumably Panama & Sabanas, 14 Jun 1924, L.A.M.
PR, n.v.).—Status uncertain, probably Riley 112 (Holotype K, n.v.; Isotypes MO (2
Psychotria hebeclada DC. or P. pubescens sheets; photos MEXU, PTBG), US (photo
Sw. PTBG)).—Psychotria horizontalis Sw.
Psychotria bakeri Dwyer, Ann. Missouri Bot. Psychotria boquetensis Dwyer, Ann. Missouri
Gard. 67: 347. 1980. Type: PANAMA: Bot. Gard. 67: 349. 1980. Type: PANAMA:
Chiriqui, Los Santos, Loma Prieta, Cerro Chiriqui, Boquete, 3800 ft., 25 May 1938,
Grande, 2400-2800 ft, 8 Jun 1967, W. H. M. E. Davidson 709 (Holotype MO 1172505
Lewis, R. K. Baker, B. MacBrytfe & R. L. (photos MEXU, PTBG); Isotypes F, US).
Oliver 2249 (Holotype MO 2219527 (photos Psychotria borucana (A. R. Molina) C. M.
MEXU, PTBG); Isotypes F, GH, n.v.). Taylor & W. C Burger, Fieldiana: Bot., n.s.
Psychotria balancanensis C. W. Ham., 33: 236. 1993. Cephaelis borucana A. R.
Phytologia 64: 219. 1988. Type: MEXICO: Molina, Ceiba 4: 31. 1951. Based on
Tabasco, Balancan, La Palma, 1-6 Jul 1939, Cephaelis affinis Standi., Publ. Field
E. Matuda 3286 (Holotype F 1026360 (photo Columbian Mus., Bot. Ser. 8: 184. 1930, not
PTBG); Isotypes A, n.v., MEXU, NY, n.v.). P. ajfinis Baker, 1877. Type: COSTA RICA:
Psychotria barbiflora DC, Prodr. 4: 509. 1830. Puntarenas, forest near Esquinas Experimental
Type: BRAZIL: Bahia, Salzmann 1 (Holotype Station, area between Rio Esquinas and Rio
G-DC, n.v. (photo MO)). According to Palmar Sur de Osa, 200 ft., 22 May 1950,
Steyermark (1972) this species occurs in P. H. Allen 5540 (Holotype US 2215926 ex
Central America, where it has been called EAP (photos MEXU, PTBG); Isotype US
Psychotria involucrata of authors, not Sw. 2828131 (photos MEXU, PTBG)). Note:
Psychotria bella Standi., J. Wash. Acad. Sci. 18: A. R. Molina annotated as "typo" the sheet
185. 1928. Type: PANAMA: Chiriqui, US 2215926 and as "isotype" US 2828131.
between Alto de las Palmas and Cerro Psychotria brachiata Sw., Prodr. Veg. Ind.
Horqueta, 2100-2268 m, 18 Mar 1911, H. Occ. 45. 1788. Type: JAMAICA: Swartzs.n.
Pittier 3250 (Holotype US 677646 (photo (presumably S, n.v.).
Psychotria brachybotrya Mull. Arg. in Mart., Type: PERU: Loreto, Williams 1226
Fl. Brasil. 6(5): 327. 1881. Syntypes: (Holotype F, n.v.).
BRAZIL: Amazonas, habitat in regione Rio Psychotria calophylla Standi., Contr. U.S. Natl.
Negro prope S. Gabriel, Spruce 2190 (B, Herb. 18: 129. 1916. Type: PANAMA:
presumably destroyed), Martius s.n. (B, Chagres, Isthmus of Panama, 15 Mar 1830, A.
presumably destroyed). Fendler 60 (Holotype US 48568 (photos
Psychotria bracteolata M. Martens & Galeotti, MEXU, PTBG); Isotypes GH (photo PTBG),
Bull. Acad. Roy. Sci. Bruxelles 11: 228. 1844. MO (photo PTBG)).
Type: MEXICO: Oaxaca, Sierra de Psychotria calopogon L. O. Williams,
Calpulalpan, 5000-7000 ft., H. Galeotti 7226 Phytologia 28: 227. 1974. Type:
(Holotype presumably BR, n.v.; Isotypes K, GUATEMALA: Huehuetenango, between
n.v., P (photos MEXU, MO, PTBG)). Ixcan and Rio Ixcan, Sierra de los
—Psychotria hebeclada DC. Cuchumatanes, 150-200 m, 23 Jul 1942, J. A.
Psychotria breedlovei Lorence, Novon 4: 125, Steyermark 49308 (Holotype F 1133859
fig. 4. 1994. Type: MEXICO: Chiapas, Mun. (photos MEXU, PTBG)).
La Trinitaria, a 1 km al E de Tziscao 6 a 11 Psychotria camponutans (Dwyer & Hayden)
km al E de la entrada al Parque Nacional Hammel, Selbyana 12: 139. 1991. Cephaelis
Lagos de Montebello, 16°05'N, 91°39'0, camponutans Dwyer & Hayden, Ann.
1330 m, 9 Ago 1985, T. Chehaibar 170 Missouri Bot. Gard. 55: 35. 1968. Type:
(Holotype PTBG 9595; Isotypes MEXU, PANAMA: Panama, Cerro Jefe, Dwyer &
UAMIZ). Gauger 7346 (Holotype MO (not located
Psychotria brenesii Standi., Publ. Field Mus. 6/1996)).
Nat. Hist., Bot. Ser. 18: 1347. 1938. Type: Psychotria capacifolia Dwyer, Ann. Missouri
COSTA RICA: Los Angeles de San Ramon, Bot. Gard. 67: 353. 1980. Type: PANAMA:
Cerros de La Balsa, 16 Mar 1928, A. M. Bocas del Toro, above Quebrada Huron on
Brenes 6050 (Holotype F 854521 (photos Cerro Bonyik, 500-1200 ft., 13 Apr 1968, J.
MEXU, PTBG)).—Palicourea standlevana Kirkbride, Jr. & J. Duke 601 (Holotype MO
C. M. Taylor 2623364 (photos MEXU, PTBG)).
Psychotria burgeri C. M. Taylor, Novon 5: Psychotria capitata Ruiz & Pav., Fl. Peruv. 2:
204,y?g. IB. 1995. Type: COSTA RICA: 59,/?/. 206, fig. a. 1799. Type: PERU:
Puntarenas, Canton de Puntarenas, Huanuco, Chinchao, Ruiz Lopez & Pavon s.n.
Monteverde, along Quebrada Maquina, (presumably MA, n.v.; for typification see
10°18'N, 84°48'W, 12 Mar 1990, W. Haber Steyermark, 1972: 618).
9778 (Holotype CR, n.v.; Isotypes F, n.v., MO Psychotria capitata Ruiz & Pav. subsp.
4619858, US, n.v.). fissistipularis Dwyer, Ann. Missouri Bot.
Psychotria caerulea Ruiz & Pav., Fl. Peruv. 2: Gard. 67: 355. 1980. Type: PANAMA: Colon,
62. pi. 213, fig. b. 1799. Type: PERU: Andes Miguel de la Borda, edge of forest near beach,
of Peru, Huanuco, ad Vitoc, Ruiz Lopez & T. B. Croat 10037 (Holotype MO 2050309).
Pavon s.n. (Holotype presumably MA, n.v.). —Psychotria capitata Ruiz & Pav.
Note: this species occurs in Panama according Psychotria capitata Ruiz & Pav. subsp. rubra
toSteyermark(1972). Dwyer, Ann. Missouri Bot. Gard. 67: 356.
Psychotria calimensis Standi, ex Steyerm., 1980. Type: PANAMA: Panama, El Llano to
Acta Biol. Venez. 4: 96. 1964. Type: Carti road, 8 km N of Panamerican Highway,
COLOMBIA: El Valle, Cuatrecasas 16298 6 Apr 1977, J. P. Folsom & L. Skog 2454
(Holotype F, n.v.; Isotypes US, n.v., VEN, (Holotype MO 2606808 (photos MEXU,
PTBG)).—Psychotria capitata Ruiz & Pav.
Psychotria carnosocarpa Dwyer & Hayden,
Ann. Missouri Bot. Gard. 55: 41. 1968. Type:
 i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

PANAMA: Chiriqui, Frances Arriba School, 1266 (Holotype B, presumably destroyed

ca. 14 miles N of David, 1200 ft., 14 Dec (photos F, MO)).—Psychotria erythrocarpa
1966, W. H. Lewis, D. Burch, J. D. Dwyer, Schltdl.
T. S. Elias, N. Escobar, R. L. Oliver & K. R. Psychotria chiapensis Standi., Contr. U.S. Natl.
Robertson 964 (Holotype MO 1966154 Herb. 23: 1390. 1926. Based on Cephaelis
(photos MEXU, PTBG)). tetragona Donn. Sm., Bot. Gaz.
Psychotria cartagoensis Nepokroeff in W. C. (Crawfordsville) 61: 376. 1916, not
Burger & C. M. Taylor, Fieldiana, Bot, n.s. Psychotria tetragona Seem., 1867. Type:
33: 239. 1993. Nom. nov. for Cephaelis MEXICO: Chiapas, Finca Mexiquito, Jul
latistipula Standi., J. Wash. Acad. Sci. 18: 1913, C. A. Purpus 6963 (Holotype US
281. 1928, not Psvchotria latistipula Benth. in 567237 (photos MEXU, PTBG); Isotype GH,
Hook., 1849. Type: COSTA RICA: Cartago, n.v.). Cephaelis, Evea.
vicinity of Orosi, 30 Mar 1924, P. C. Standley Psvchotria chiriquiensis (Standi.) C. M. Taylor
39695 (Holotype US 1153168 (photos in W. C. Burger & C. M. Taylor, Fieldiana,
MEXU, PTBG)). Bot., n. s. 33: 241. 1993. Cephaelis
Psychotria carthaginensis Jacq., Enum. Syst. chiriquiensis Standi., Ann. Missouri Bot.
PI. Carib. 16. 1760. Type: COLOMBIA: Gard. 28: 469. 1941. Type: PANAMA:
Bolivar, vicinity of Cartagena, Jacquin s.n. Chiriqui, Bajo Chorro, 1900 m, 20-22 Jul
(location unknown). Uragoga. 1940, R. E. Woodson, Jr. & R. W. Scheiy 610
Psychotria cascajalensis C. W. Ham., (Holotype MO 1209028 (photos MEXU,
Phytologia 64: 220. 1988. Type: PANAMA: PTBG); Isotypes F, GH, n.v.).
Code, N of Penonome, between Llano Psychotria chiriquina Standi., Contr. U.S. Natl.
Grande and Cascajal, rd. to Coclesito, 480 m, Herb. 18: 129. 1916. Type: PANAMA:
2 May 1979, B. Hammel 7227 (Holotype MO Chiriqui, Alto de la Cuesta, around Camp
2901097; Isotypes ENCB, n.v., MO). Aguacatal, eastern slope of Chiriqui Volcano,
Psychotria catillicalyx Dwyer, Ann. Missouri 2100-2200 m, 10-13 Mar 1911, H. Pittier
Bot. Gard. 67: 358. 1980. Type: PANAMA: 3118 (Holotype US 677509 (photos MEXU,
Veraguas, Valley of Rio Dos Bocas, 11 km PTBG); Isotype F).
from Escuela Agricola Alto Piedra on road to Psychotria chitariana Dwyer & C. W. Ham.,
Calovebora, 450 m, 30 Aug 1974, T. B. Croat Phytologia 64: 221. 1988. Type: COSTA
27548 (Holotype MO 2272494).—Psychotria RICA: Cartago, 13 km E of Turrialba on the
panamensis Standi, var. panamensis highway to Limon; canyon of Rio Chitaria,
Psychotria cerroazulensis Dwyer, Ann. 750-800 m, 10 May 1983, R. Liesner, E.
Missouri Bot. Gard. 53: 107. 1966. Type: Judciewicz & B. Perez G. 15400 (Holotype
PANAMA: Panama, Cerro Azul, Goofy Lake MO 3235522 (photo PTBG); Isotype CR,
area, 600 m, 1 Sep 1963, J. D. Dwyer 2875
(Holotype MO 1800011 (photo PTBG)). Psvchotria chlorobotrya Standi., J. Wash.
—Psychotria calophylla Standi. Acad. Arts 16: 17. 1926. Type: MEXICO:
Psychotria chagrensis Standi., J. Wash. Acad. Chiapas, damp forest, mountains near Fenix,
Sci. 15: 105. 1925. Type: PANAMA: Canal May 1925, C A. Purpus 104 (Holotype US
Zone, Barro Colorado Island, Gatun Lake, 17 1208242 (photos MEXU, PTBG)).
Jan 1924, P. C Standley 31373 (Holotype US Psychotria chlorotica Mull. Arg., Flora 59:
1215962 (photos MEXU, PTBG)). 545. 1876. Type: BRAZIL: Bahia, Blanchet
Psychotria chamissoana (Loes.) Standi., J. 908 (Lectotype presumably G, n.v.;
Wash. Acad. Sci. 17: 341. 1927. Mapouria designated by Dwyer, 1980: 362).
nma L. O. Williams,
chamissoana Loes., Verh. Bot. Ver.
Brandenburg 65: 112. 1923. Type: MEXICO: Phytologia 28: 228. 1974. Type:
Veracruz, Tecolute [Tecolutla], Jan, Schiede GUATEMALA: El Progreso, hills north of
 Finca Piamonte, between Finca Piamonte and MEXU, PTBG)).—Palicourea copensis
summit of Volcan Santa Luisa, 2400-3333 m, (Dwyer) C. M. Taylor
5 Feb 1942, J. A. Steyermark 43518 (Holotype Psychotria copeyana Standi. & L. O. Williams,
F 1133849 (photos MEXU, PTBG)). Ceiba 1: 251. 1951. Type: COSTA RICA:
—Psychotria purpusii Standi. Cartago, near El Copey, Cordillera de
Psychotria cincta Standi., Publ. Field Talamanca, 1800 m, 23 Apr 1949, L. O.
Columbian Mus. Nat. Hist., Bot. Ser. 7: 90. Williams & P. H. Allen 16482 (Holotype F
1930. Type: COLOMBIA: El Valle, Cordoba, 1388092 (photos MEXU, PTBG); Isotype US
Dagua Valley, 6-8 May 1922, E. P. Killip (photos MEXU, PTBG)).—Palicourea
5140 (Holotype US 1141369 (photo PTBG) garciae Standi.
Isotype G, n.v.). Psychotria cornifolia Humb. & Bonpl. ex
Psychotria clivorum Standi. & Steyerm., Publ. Roem. 8c Schult., Syst. Veg. 5: 191. 1819.
Field Mus. Nat. Hist., Bot. Ser. 23: 87. 1944. Type: VENEZUELA: Near Maypures, on the
Based on Psychotria limonensis var. Orinoco, Humboldt & Bonpland 893
angustifolia Standi., not Psychotria (Holotype P-Bonpland, n.v.; Isotype B,
angustifolia Poir. in Lam., 1804. Type: presumably destroyed (photo MO)).
GUATEMALA: Suchitepequez, Finca Moca, —Rudgea cornifolia (Humb. & Bonpl.)
3400 ft., 7 Jan 1935, A. F. Skutch 2073 Standi.
(Holotype F 934091 (photos MEXU, PTBG); Psvchotria correae (Dwyer & Hayden) C. M.
Taylor in W. C. Burger & C. M. Taylor,
Psychotria cocosensis C. W. Ham., Phytologia Fieldiana, Bot., n.s. 33: 244. 1993. Cephaelis
64: 222. 1988. Type: COSTA RICA: Cocos correae Dwyer & Hayden, Ann. Missouri Bot.
Island, 18 Jun 1898, H. Pittier 12375 Gard. 55: 36. 1968. Type: PANAMA: Code,
(Holotype US, n.v.). mountains N of El Valle de Anton,
Psychotria compressicaulis K. Krause, Bot. 2500-3000 ft., 28 May 1967, W. H. Lewis, B.
Jahrb. Syst. 54, Beibl. 119: 44. 1916. Type: McBryde, R. L. Oliver & J. E. Ridgway 1759
COSTA RICA: Forets de Tuis, 600 m, Jul (Holotype MO 1887361 (photos MEXU,
1898, H. Pittier 12412 (Holotype B, PTBG); Isotypes GH, n.v., UC, n.v., US, n.v.).
presumably destroyed; Isotype US (photos Psychotria costaricensis Pol., Linnaea 41: 571.
MEXU, PTBG)).—Psychotria panamensis 1877. Type: COSTA RICA: Angostura, Nov
Standi, var. compressicaulis (K. Schum.) 1875, H. Polakowsky 384A (Holotype B,
C. W. Ham. presumably destroyed)).—Psychotria pilosa
Psychotria concolor Benth. in Oerst., Vidensk. Ruiz & Pav.
Meddel. Dansk Naturhist. Foren. Kjobenhavn Psychotria costivenia Griseb., Plantae
1852: 31. 1853. Type: NICARAGUA: Volcan Wrightianae e Cuba Onentale, pars II: 508.
de Mombacho, Oersteds.n. (Holotype 1862; Mem. Am. Acad. Arts, n.s. 8: 508.
presumably K, n.v.).—Status uncertain. 1863. var. costivenia. Type: CUBA: Cuba
Psychotria cooperi Standi., Publ. Field Mus. Orientali, prope Monte Verde, Jan-Jul 1859,
Wright 242 (Lectotype GH, n.v.; Isolectotype
BR, n.v.; designated by Hamilton, 1989).
Camp on Chinqui trail. 1250 ft., Jan-Mar Uragoga.
1928, P. Cooper 577 (Holotype F 579633 Psychotria costivenia Griseb. var. altorum
(photos MEXU, PTBG); Isotype NY, n.v.). (Standi. & Steyerm.) C. W. Ham., Phytologia
Psychotria copensis Dwyer, Ann. Missouri Bot. 64: 223. 1988. Psychotria altorum Standi. &
Gard. 67: 365. 1980. Type: PANAMA: Code, Steyerm., Publ. Field Mus. Nat. Hist., Bot.
El Cope, El Potroso, Atlantic slope of Alto Ser. 23: 86. 1944. Type: GUATEMALA:
Calvario, 26 Feb 1977, J. P. Folsom & R. Quezaltenango, Montana Chicharro, on lower
Lantz 1880 (Holotype MO 2605731 (photos southeast-facing slopes of Volcan Santa
A NOMENCLATOR OF MEXICAN AND CENTRAL

Maria, 2-4 miles south of Santa Maria de Psychotria deflexa DC, Prodr. 4: 510. 1830.
Jesus, 1400-1500 m, 17 Jan 1940, J. A. Type: FRENCH GUIANA: Without precise
Steyermark 34302 (Holotype F 1056806 locality or collector (Holotype presumably G-
(photos MEXU, PTBG)). DC (n.v.); Isotype G-DEL, n.v. (photo MO)).
Psychotria crebrinervia Standi., Publ. Field Psychotria deneversii C M. Taylor, Novon 6:
Mus. Nat. Hist, Bot. Ser. 4: 343. 1929. Type: 300, fig. 2C, D. 1996. Type: PANAMA:
HONDURAS: Atlantida, Lancetilla valley, Comarca de San Bias, El Llano-Carti road, 18
near Tela, 20-600 m, 6 Dec 1927-20 Mar km from Interamerican Highway, headwaters
1928, P. C. Standley 55286 (Holotype F of Atlantic-draining creeks, 9°19'N, 78°55'W,
584201 (photos MEXU, PTBG)). 300 m, 7 Sep 1984, G. de Nevers 3861
—Psychotria berteriana DC. (Holotype PMA 38224, n.v.; Isotype MO
Psychotria crocea Sw., Prodr. Veg. Ind. Occ. 4658111).
44. 1788. Type: JAMAICA: Without precise Psychotria dichroa (Standi.) C M. Taylor in
locality, Swartz s.n. (Holotype BM, n.v.). W. C. Burger & C M. Taylor, Fieldiana, Bot.,
Uragoga.—Palicourea crocea (Sw.) Roem. n.s. 33: 245. 1993. Evea dichroa Standi.,
& Schult. Contr. U.S. Natl. Herb. 18: 124. 1916. Type:
Psychotria croceovenosa Dwyer, Ann. PANAMA: Chiriqui, humid forest of Cuesta
Missouri Bot. Gard. 67: 367. 1980. Type: de Las Palmas, southern slopes of Cerro de la
PANAMA: Chiriqui, N of San Felix at Horqueta, 1700-2100 m, 17-19 Nov 1911, H.
Chiriqui-Bocas del Toro border on Cerro Pittier 3218 (Holotype US 677615 (photos
Colorado copper mine road along continental MEXU, PTBG); Isotype F). Cephaelis.
divide, 5000-5500 ft., 4 May 1975, S. Mori & Psychotria dispersa Standi, J. Wash. Acad. Sci.
J. Kallunki 5880 (Holotype MO 2579259 18: 184. 1928. Type: COSTA RICA:
(photos MEXU, PTBG)). Based on the same Guanacaste, Los Ayotes, near Tilaran,
type as Palicourea montensis Dwyer, 1980, 600-700 m, 21 Jan 1926, P. C. Standley & J.
which Valerio 45548 (Holotype US 1254170 (photos
MEXU, PTBG)).—Psychotria pittieri Standi.
Psychotria cuneifolia Sesse & Moc,, Fl. Mexic, Psvchotria domingensis Jacq., Enum. PI. Carib.
ed. 2, 58. 1894. Not P. cuneifolia DC, 1830. 16. 1760. Type: SANTO DOMINGO
Type: MEXICO: Habitat in Cordovae vicinis (Hispaniola): Jacquin (no Jacquin material
montibus, floret Augusto, Sesse & Mocino s.n. was located, according to Howard, 1989).
(presumably MA, n.v.).—Status uncertain. Palicourea.
Psychotria cuspidata Bredem. ex Roem. & Psvchotria dosbocensis Dwyer, Ann. Missouri
Schult., Syst. Veg. 5: 192. 1819. Type: Bot. Gard. 67: 370. 1980. Type: PANAMA:
VENEZUELA: Ad Caracas, Bredemeyer s.n. Veraguas, valley of Rio Dos Bocas on road
(Holotype B, presumably destroyed). between Alto Piedra (above Santa Fe) and
—Psychotria acuminata Benth. Calovebora, 350-400 m, 29 Aug 1974, T. B.
Psychotria cyanococca Seem., Fl. Mag. tab. Croat 27427 (Holotype MO 2216296 (photo
479. 1870. Type: NICARAGUA: Chontales, PTBG)).
Seemann s.n. (Holotype presumably BM, Psychotria dressleri (Dwyer) C W. Ham.,
nv.).—Status uncertain. Phytologia 64: 223. 1988. Cephaelis dressleri
Psychotria davidsei Dwyer, Ann. Missouri Bot. Dwyer, Ann. Missouri Bot. Gard. 67: 68.
Gard. 67: 368. 1980. Type: PANAMA: 1980. Type: PANAMA: San Bias, forest
Darien, Cerro Tacarcuna, summit camp along southeast of Puerto Obaldia, 18 Aug 1971,
stream N of camp, 1550-1650 m, A. Gentry & T. B. Croat 16801 (Holotype MO 2138462
S. Mori 14071 (Holotype MO (not located (photo PTBG)).
6/1996)). Psychotria dukei Dwyer, Ann. Missouri Bot.
Gard. 67: 371. 1980. Type: PANAMA:
 Darien, Casi-Cana Trail on Cerro Darien, top of W peak of Cerro Tacarcuna
Campamiento, E of Tres Bocas, headwaters of massif, 1800-1850 m, 28 Jan 1975, A. Gentry
Rio Coas, 29 Apr 1968, J. H. Kirkbride, Jr. & & S. Mori 13984 (Holotype MO 2280217
J. A. Duke 1245 (Holotype MO 2162265 (photos MEXU, PTBG)).—Psychotria
(photo PTBG)). goldmanii Standi.
Psychotria durilancifolia Dwyer, Ann. Missouri Psychotria eurycarpa Standi., J. Wash. Acad.
Bot. Gard. 67: 372. 1980. Type: PANAMA: Sci. 18: 275. 1928. Type: COSTA RICA:
Panama, El Llano-Carti Road, Gorgas Lab, Guanacaste, Quebrada Serena, southeast of
Mosquito Project study area, 250 m, 1 Aug Tilaran, 700 m, 27 Jan 1926, P. C. Standley &
1974, T. B. Croat 26028 (Holotype MO J. Valerio 46237 (Holotype US 1254544
2240525 (photo PTBG)).—Psychotria (photos MEXU, PTBG)).
panamensis Standi, var. panamensis Psychotria (?) excelsa Kunth in Humb., Bonpl.
Psychotria dwyeri C. W. Ham., Phytologia 64: & Kunth, Nov. Gen. Sp. 3: 355, tab. 282.
223. 1988. Type: MEXICO: Oaxaca, Distr. de 1819. Type: MEXICO: Veracruz, Xalapa, A.
Choapam [Santiago Choapan], Santa Maria, Bonpland 1445 (Holotype P-Bonpland, n.v.;
Montana Santa Maria, 1500 m, 7 Apr 1938, Y. Isotypes B, n.v. (photo MO), P (photos
Mexia 9265 (Holotype NY, n.v.; Isotypes B, MEXU, PTBG)). Deppea.—Hoffmannia
n.v., F (photo PTBG), GH, n.v., K, n.v., MO excelsa (Kunth) K. Schum.
(photo PTBG), US). Psvchotria faxlucens Lorence & Dwyer, Bol.
Psychotria elata (Sw.) Hammel, Selbyana 12: Soc. Bot. Mexico 47: 50. 1987. Type:
139. 1991. Cephaelis elata Sw., Prodr. 45. MEXICO: Veracruz, Mun. de Catemaco,
1788. Type: JAMAICA: Swartzs.n. Ejido de Coscoapan, 8 km SE de Coyame, 16
(presumably S, n.v.). Abr 1974, F. Ponce & R. Cedillo T. 2
Psychotria elongata Benth. in Oerst., Vidensk. (Holotype MEXU; Isotypes F (photo PTBG),
Meddel. Dansk Naturhist. Foren. Kjobenhavn MO).
1852: 32. 1853. Type: NICARAGUA: Volcan Psychotria fendleri Standi., Contr. U.S. Natl.
de Mombacho, Oersted 11594 (Holotype C Herb. 18: 129. 1916. Type: PANAMA:
(photos MEXU, PTBG), fragment US (photos Chagres, Isthmus of Panama, 25 Feb 1850,
MEXU, PTBG); Isotype K, n.v.). Fendler 59 (Holotype US 48567 (photos
—Psychotria nervosa Sw. var. nervosa MEXU, PTBG); Isotype GH (photos MEXU,
Psychotria emetica L. f., Suppl. PI. 144. 1781. PTBG)).
Type: COLOMBIA: America septentrionalis Psychotria fimbriata Benth., J. Bot. Hook. 3:
provinciae Gironensis, Mutis 1602 (Holotype 226. 1841. Type: GUYANA: Essequibo
LINN, n.v.; probable Isotype US (photo River, Schomburgk 51 (Holotype presumably
PTBG)). Uragoga. K, n.v.). Rudgea.—Rudgea cornifolia
Psychotria erecta (Aubl.) Standi. & Steyerm., (Kunth) Standi.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 24. Psychotria flava Oerst. ex Standi., J. Wash.
1943. Ronabea erecta Aubl., Hist. PI. Guian. Acad. Sci. 17: 341. 1927. Type: MEXICO:
1: 156. 1775. Type: FRENCH GUIANA: In Veracruz, Misantla, F. Liebmann 11605
sylvis Oyao, Orapu, Sinemari, Aublets.n. (Holotype C (photos F, MEXU, MO, PTBG)).
(Holotype presumably BM, n.v.). Psychotria fosteri C. W. Ham., Phytologia 64:
Psychotria erythrocarpa Schltdl., Linnaea 9: 224. 1988. Type: PANAMA: Veraguas, Playa
595. 1834. Type: MEXICO: Veracruz, Rosario, northern tip of Coiba Island, 26 Aug
Hacienda de la Laguna, Schiede & Deppe 385 1970, R. Foster 1605 (Holotype DUKE, n.v.;
(Holotype B, presumably destroyed (photo & Isotypes F, GH, MO (photo PTBG)).
fragment F); Isotype K, n.v.). Uragoga. Psychotria fruticetorum Standi., J. Arnold
Psychotria eugenifolia Dwyer, Ann. Missouri Arbor. 11: 42. 1930. Type: HONDURAS:
Bot. Gard. 67: 375. 1980. Type: PANAMA: Comayagua, vicinity of Siguatepeque,
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

1080-1400 m, 14-27 Feb 1928, P. C. 943529 (photos MEXU, PTBG); Isotype US

Standley 56197 (Holotype F 581450 (photos (photos MEXU, PTBG)).
MEXU, PTBG); Isotype A, n.v.). Psychotria goldmanii Standi., Contr. U.S. Natl.
Psychotria furcata DC, Prodr. 4: 512. 1830. Herb. 18: 130. 1916. Type: PANAMA:
Type: PANAMA: Without precise locality, Panama, Mont Pirre, head of Rio Limon, 1500
Haenke s.n. (Holotype presumably PR, n.v.). m, 1 Mar 1911, E. A. Goldman 1883
Psychotria galeottiana (M. Martens) C. M. (Holotype US 716141 (photos MEXU,
Taylor & Lorence, Taxon 34: 669. 1985. PTBG)).
Palicourea galeottiana M. Martens, Bull. Psychotria graciliflora Benth. in Oerst.,
Acad. Roy. Sci. Bruxelles 14: 136. 1844. Vidensk. Meddel. Dansk Naturhist. Foren.
Type: MEXICO: Oaxaca, Cordillera, 6500 ft., Kjobenhavn 1852: 35. 1853. Type: COSTA
1840, H. Galeotti 2602 (Lectotype BR (photos RICA: Alajuela, ad Naranjo, Apr 1847,
MEXU, PTBG); Isolectotypes K, n.v., P Oersted 11611 (Holotype C (photos F,
(photos MEXU, PTBG); designated by Taylor MEXU, PTBG); Isotype US (photos MEXU,
& Lorence, 1985). Uragoga. PTBG)). Mapouria, Uragoga.
Psychotria gardenioides (Scheidw.) Standi., J. Psychotria gracilis Ruiz & Pav., Fl. Peruv. 2:
Wash. Acad. Sci. 17: 342. 1927. Rhodostoma 63, t. 211, fig. C. 1799. Type: PERU:
gardenioides Scheidw., Allg. Gartenzeitung Huanuco, Pillao et Pueblo Nuevo, Ruiz Ldpez
10: 286. 1842. Type: Based on cultivated & Pavon s.n. (presumably MA, n.v.).
material of uncertain origin, presumably from —Geophila gracilis (Ruiz & Pav.) DC.
Veracruz, Mexico. Type not designated. Psychotria granadensis Benth. in Oerst.,
Palicourea. Vidensk. Meddel. Dansk Naturhist. Foren.
Psychotria gatunensis Dwyer & Hayden, Ann. Kjobenhavn 1852: 34. 1853. Type:
Missouri Bot. Gard. 54: 142. 1976. Type: NICARAGUA: Granada, Oersted 11612
PANAMA: Canal Zone, W of Limon Bay, (Holotype C (photos MEXU, PTBG)).
Ganin Locks and Gatun Lake, 5 Aug 1955, Uragoga.—Psychotria nervosa Sw.
Johnston 1554 (Holotype MO 1709657; Psychotria grandicarpa Dwyer & Hayden, Ann.
Isotypes A (2 sheets)).—Psychotria Missouri Bot. Gard. 55: 42. 1968. Type:
calophylla Standi. PANAMA: Panama, Cerro Jefe, 12 Mar 1967,
Psychotria gentryi (Dwyer) C. M. Taylor, J. D. Dwyer, Baker & Gauger 7244
Novon4: 305. 1994. Cephaelis gentryi (Lectotype MO 2002447; Isolectotypes MO
Dwyer, Selbyana 2: 57. 1977. Type: 2002694, MO 2002695; designated by
ECUADOR: Los Rios, Rio Palenque Science Hamilton, 1989).—Psychotria calophylla
Center, km 56 of Quevedo-Santo Domingo Standi.
road, 150-220 m, 19 Sep 1972, C. Dodson Psychotria grandis Sw., Prodr. Veg. Ind. Occ.
5169 (Holotype MO 2281982; Isotypes 43. 1788. Type: JAMAICA: Interioris
RPSC, n.v., SEL, n.v.). occidentalis, Swartz s.n. (Holotype S, n.v.
Psychotria glauca Pol., Linnaea 41: 569. 1877. (photo A)). Uragoga.
Syntypes: COSTA RICA: San Jose, H. Psychotria grandistipula Standi., J. Wash. Acad.
Polakowski 377 (B, presumably destroyed); Sci 18: 276. 1928. Not Psychotria
H. Polakowski 378 (B, presumably grandistipula Merrill, 1923. Type: COSTA
destroyed)—Psychotria pubescens Sw. RICA: San Jose, vicinity of Santa Maria de
Psychotria glomerulata (Donn. Sm.) Steyerm., Dota, 1500-1800 m, 26 Dec 1925-3 Jan
Mem. New York Bot. Gard. 23: 670. 1972. 1926, P. C. Standley & J. Valerio 43268
Cephaelis glomerulata Donn. Sm., Bot. Gaz. (Holotype US 1253164 (photos MEXU,
(Crawfordsville) 16: 12, pi. 1. 1891. Type: PTBG)).—Psychotria panamensis Standi.
GUATEMALA: Dept. Livingston, Rio Dulce,
Mar 1880, J. D. Smith 1637 (Holotype US
Psychotria guadalupensis (DC.) Howard, J. Psychotria hirsuta (M. Martens & Galeotti)
Arnold Arbor. 47: 139. 1966. Loranthus Mflll. Arg. in Mart., Fl. Bras. 6(5): 370. 1881.
guadalupensis DC, Prodr. 4: 294. 1830. Cephaelis hirsuta M. Martens & Galeotti,
Type: GUADALOUPE: Bertero s.n. Bull. Acad. Roy. Bruxelles 11: 35. 1844.
(Holotype G-DC, n.v. (microfiche IDC Type: MEXICO: Oaxaca, La Chinantla, 4000
800.666: II.2)). pieds, H. Galeotti 7185 (Holotype BR, n.v.).
Psychotria guapilensis (Standi.) Hammel, —Psychotria poeppigiana Mull. Arg.
Selbyana 12: 139. 1991. Evea guapilensis Psychotria hoffmannseggiana (Willd. ex
Standi., J. Wash. Acad. Sci. 15: 104. 1925. Roem. & Schult.) Mull. Arg. in Mart., Fl.
Type: COSTA RICA: Limon, vicinity of Bras. 6(5): 336. 1881. Cephaelis
Guapiles, 300-500 m, P. C Standley 37025 hoffmannseggiana Willd. ex Roem. & Schult.,
(Holotype US 1153029 (photos MEXU, Syst. Veg. 5: 214. 1819. Type: BRAZIL: Para,
PTBG)). Cephaelis. Sieber in Hoffmanns egg s.n. (location
Psychotria haematocarpa Standi., J. Wash. unknown).
Acad. Sci. 18: 274. 1928. Type: COSTA Psychotria hondensis Standi., J. Wash. Acad.
RICA: Guanacaste, Naranjos Agrios, near Sci. 18: 183. 1928. Type: COSTA RICA: Bois
Tilaran, 600-700 m, 29 Jan 1926, P. C de Rio Hondo, versant atlantique, 100 m, Aug
Standley & J. Valerio 46407 (Holotype US 1901, H. Pittier 16161 (Holotype US 764151
1254627 (photos MEXU, PTBG)). (photos MEXU, PTBG)).—Coussarea
Psychotria hammelii Dwyer, Ann. Missouri hondensis (Standi.) C M. Taylor & W. C
Bot. Gard. 67: 383. 1980. Type: PANAMA:
Veraguas, forested mountains W of Alto de Psychotria horizontalis Sw., Prodr. Veg. Ind.
Piedras, W of Santa Fe, 3200-5600 ft., 8 Sep Occ. 44. 1788. Type: SANTO DOMINGO:
1978, B. Hammel 4634 (Holotype MO Swartz s.n. (presumably S, n.v. (photo A)).
2680614 (photos MEXU, PTBG); Isotype Uragoga.
MO). Psychotria horizontalis Sw. subsp. basicorda
Psychotria hazenii Standi., Publ. Field Dwyer, Ann. Missouri Bot. Gard. 55: 42.
Columbian Mus., Bot. Ser. 7: 96. 1930. Type- 1968. Type: PANAMA: Veraguas, Isla de
COLOMBIA: El Valle, Killip 5573 (Holotype Coiba (Penal Colony), Camp Aguja, 8 Jul
US, n.v.; Isotypes GH, n.v., NY, n.v.). 1962, J. D. Dwyer 2384A (Holotype MO
Psychotria hebeclada DC, Prodr. 4: 513. 1830. 2165270 (photos MEXU, PTBG); Isotype
Type: MEXICO: Without precise locality, MO).—Psychotria horizontalis Sw.
Haenke s.n. (Holotype presumably PR, n.v.). Psychotria hornitensis Dwyer & C W. Ham.,
Psychotria herbacea Jacq., Enum. PI. Carib. 16 Phytologia 64: 225. 1988. Type: PANAMA:
1760. Type: MARTINIQUE: Jacquin Chiriqui, ridge and summit of Cerro Hornito,
(location unknown). Geophila.—Geophila above Los Planes de Hornito, 2100 m, 14 Mar
gracilis (Ruiz & Pav.) DC 1982, S. Knapp et al. 4198 (Holotype MO
Psychotria herbacea Sesse & Moc., Fl. Mexic, (not located 6/1996)).
ed. 2, 58. 1894, not Psychotria herbacea Psychotris impatiens Dwyer, Ann. Missouri
Jacq., 1760. Type: MEXICO: Habitat in Bot. Gard. 67: 385. 1980. Type: PANAMA:
umbrosis montibus prope Oppidium de la Darien, north slopes of Cerro Pirre, 700-950
Aguadilla, floret Maio, Sesse & Mocifio s.n. m, 6 Apr 1975, S. Mori & J. Kallunki 5446
(presumably MA, n.v.).—Status uncertain. (Holotype MO 2272508 (photos MEXU,
Psychotria heydei Standi., J. Wash. Acad. Sci PTBG)).
18: 184. 1928. Type: GUATEMALA: Quiche Psychotria incurvata Sesse & Mo?., Fl. Mexic,
Chiul, 2600 m, Apr 1892, Heyde & Lux 3173' ed. 2, 58. 1894. Type: MEXICO: Habitat in
(Holotype US 939642 (photos MEXU, Tuxtlae sylvis. Floret Julio, Sesse & Mocino
PTBG); Isotype F (photos MEXU, PTBG)).
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

s.n. (presumably MA, n.v.).—Status Psychotria jinotegensis C. Nelson, A. R.

uncertain. Molina & Standi., Phytologia 50: 1. 1981. var.
Psychotria indigotica Dwyer, Ann. Missouri jinotegensis. Type: NICARAGUA: Jinotega,
Bot. Gard. 67: 386. 1980. Type: PANAMA: region of La Montanita and Las Mesitas in
Canal Zone, forest of Gatun, 15 Sep 1967, A. sierra W of Jinotega, ca. 1100-1400 m, 29 Jun
Hladik511 (Holotype MO 2165124). 1947, P. C Standley 10314 (Holotype EAP,
—Rudgea isthmensis Standi. n.v.; Isotype F).
Psychotria insignis Standi., Contr. U.S. Natl. Psychotria jinotegensis C. Nelson, A. R.
Herb. 18: 130. 1916. Type: PANAMA: San Molina & Standi, var. morazanensis C. W.
Bias, hills of Sperdi, near Puerto Obaldia, San Ham., Ann. Missouri Bot. Gard. 76: 408.
Bias coast, 20-200 m, Sep 1911, H. Pittier 1989. Type: EL SALVADOR: Morazan,
4410 (Holotype US 679567 (photos MEXU, easternmost peak, Monte de Cascaquatique,
PTBG); Isotype F (photos MEXU, PTBG)). near summit on N side, 1500 m, 25 Dec 1941,
Psychotria insueta (Dwyer) C. W. Ham., J. M. Tucker 610 (Holotype US, n.v.; Isotypes
Phytologia 64: 226. 1988. Cephaelis insueta F (photo PTBG), K, n.v., NY, n.v., P (photo
Dwyer, Ann. Missouri Bot. Gard. 67: 73. PTBG)).
1980. Type: PANAMA: Veraguas, 11 km Psychotria juarezana C. M. Taylor & Lorence,
from Escuela Agricola Alto de Piedra, in Novon 2: 264, fig 2. 1992. Type: MEXICO:
forest along Rio Dos Bocas, tropical wet Oaxaca: Distr. de Ixtlan, Sierra de Juarez, ruta
forest, Atlantic slope, 15 Nov 1974, S. Mori & 175, Tuxtepec a Oaxaca ca. 4 km al SO de La
J. Kallunki 3108 (Holotype MO 2220981 Esperanza, 1,700 m, 4 Apr 1983, D. H.
(photo PTBG)). Lorence & R. Cedillo T. 4266 (Holotype MO
Psychotria ipecacuanha (Brot.) Stokes, Bot. 3750634; Isotype MEXU).
Mat. Med. 1: 365. 1812. Callicocca Psychotria justicioides Schltdl., Linnaea 9: 596.
ipecacuanha Brot., Trans. Linn. Soc. London 1834. Type: MEXICO: Barranca de Tioselo,
6: 137. 1802. Type: BRAZIL: Gomez s.n. Schiede & Deppe 259 (Holotype presumably
(presumably LISU, n.v.). Cephaelis. Evea, HAL, n.v.; Isotypes B, presumably destroyed
Uragoga. (photo MO), P (photos MEXU, PTBG)).
Psychotria izabalensis L. O. Williams, Uragoga.—Psychotria hebeclada DC.
Phytologia 28: 229. 1974. Type: Psychotria laevis DC. var. angustifolia Oerst.,
GUATEMALA: Izabal, vicinity Lago Izabal, Vidensk. Meddel. Dansk Naturhist. Foren.
600 m, 22 Apr 1966, G. C. Jones, G. R. Kjobenhavn 1852: 15. 1853. Type:
Proctor & L. Facey 3024 (Holotype F NICARAGUA: Monte Pantasmo, Jan 1848,
1665554 (photos MEXU, PTBG)). Oersted 11652 (Holotype C (photos MEXU,
Ps>c i 1 O. PTBG)).—Psychotria uliginosa Sw.
oaxacana Lorence, Bol. Soc. Bot. Mexico 47: Psychotria lamarinensis C. W. Ham.,
53. 1987. Type: MEXICO: Oaxaca, Distr. de Phytologia 64: 227. 1988. Type: COSTA
Ixtlan, Sierra de Juarez, Ruta 175 Tuxtepec a RICA: Alajuela, east of Rio Rafael and south
Oaxaca, 2.5 km NE de Puerto Eligio, 800 m, 1 of the hot springs west of La Marina, 500 m,
Jun 1983, D. Lorence & R. Cedilla T. 4250 19 May 1968, W. C Burger & R. G. Stoltze
(Holotype MEXU; Isotypes BM, BR, CAS, 5062 (Holotype NY, n.v.; Isotype MO (photo
ENCB, F, MO, NY, UC, US, XAL). PTBG)).
Psychotria jimenezii Standi., J. Wash. Acad. Psvchotria laselvensis C. W. Ham., Phytologia
Sci. 15: 288. 1925. Type: COSTA RICA: San 64: 228. 1988. Type: COSTA RICA: Heredia,
Jose, La Hondura, 1300-1700 m, 16 Mar Finca La Selva, OTS field station, Rio Puerto
1924, P. C. Standley 37892 (Holotype US Viejo just E of its junction with Rio Sarapiqui,
1153107 (photos MEXU, PTBG)). 100 m, 19 May 1980, B. Hammel 8706
(Holotype DUKE, n.v.; Isotype CR, n.v.).
Psychotria liesneri Dwyer, Ann. Missouri Bot. lectotype. Uragoga.—Psycho
Gard. 67: 387. 1980. Type: PANAMA:
Panama, NE of town of Cerro Azul, 20 km by Psychotr
road from Inter-American Hwy., 10 Dec Missoi
1974, S. Mori & J. Kallunki 3658 (Holotype Psychotria longipedunculata (Gardn.) Muell.
MO 2257865 (photos MEXU, PTBG)). Arg. in Mart., 1881, nor Elmer, 1911. Type:
Psychotria lilacina Standi. & Steyerm., Publ. PANAMA: Veraguas, valley of Rio Dos
Field Mus. Nat. Hist., Bot. Ser. 23: 252. 1947. Bocas, 15.6 km NW of Santa Fe, road
Type: GUATEMALA: Huehuetenango, Cerro between Escuela Agricola Alto Piedra and
Huitz, between Mimahuitz and Yulhuitz, Calovebora, 450-550 m, 31 Aug 1974, T. B.
Sierra de los Cuchumatanes, 1500-2600 m, 14 Croat 27621 (Holotype MO 2203532).
Jul 1942, J. A. Steyermark 48564 (Holotype F —Psychotria longipedunculoides C. M.
1133819 (photos MEXU, PTBG)). Taylor
—Psychotria minarum Standi. & Steyerm. Psychotria longipedunculoides C. M. Taylor in
Psychotria limonensis K. Krause, Bot. Jahrb. W. C. Burger & C. M. Taylor, Fieldiana, Bot.,
Syst. 54, Beibl. 119: 43. 1916. Type: COSTA n.s. 33: 257. 1993. Norn. nov. {ox Psychotria
RICA: Limon, Hot de la Uvita, Puerto Limon, longipedunculata Dwyer, 1980, not P.
Jul 1898, H. Pittier 12681 (Holotype B, longipedunculata (Gardn.) Muell. Arg. in
presumably destroyed (photo and fragment F); Mart., 1881. Type: PANAMA: Veraguas,
Isotype US (photos MEXU, PTBG)). valley of Rio Dos Bocas, 15.6 km NW of
Psychotria limonensis K. Krause var. Santa Fe, road between Escuela Agricola Alto
angustifolia Standi., Publ. Field Mus. Nat. Piedra and Calovebora, 450-550 m, 31 Aug
Hist., Bot. Ser. 17: 282. 1937. Type: 1974, T. B. Croat 27621 (Holotype MO
GUATEMALA: Suchitepequez, Finca Moca, 2203532).
1020 m, 7 Jan 1935, A. F. Skutch 2073 Psychotria lundellii Standi, in Lundell, Contr.
(Holotype F 934091 (photos MEXU, PTBG); Univ. Michigan Herb. 4: 29. 1940. Type:
Isotype A, n.v.).—Psychotria clivorum BRITISH HONDURAS (BELIZE): El Cayo
Standi. & Steyerm. District, Valentin, in advanced forest,
Psychotria limonensis K. Krause var. laxinervia limestone valley, 26 Jun 1936, C L. Lundell
Loes., Repert. Sp. Nov. 18: 361. 1922. Type: 6260 (Holotype F 861235 (photos MEXU,
MEXICO: Chiapas, ruinas del Palenque, 10 PTBG); Isotypes GH, n.v., MICH, n.v., NY,
Mar 1911, E. Seler & C. Seler 5463 (384) n.v., TEX-LL (photos MEXU, PTBG), US
(Holotype B, presumably destroyed (photo (photo PTBG)).
US; fragment F); Isotype GH (photo PTBG)). Psychotria luteotuba Lorence, Bol. Soc. Bot.
—Psychotria limonensis K. Krause Mexico 47: 55. 1987. Type: MEXICO:
Psychotria longicollis Benth. in Oerst., Vidensk. Veracruz, Mun. de Hidalgotitlan, zona de
Meddel. Dansk Naturhist. Foren. Kjobenhavn Uxpanapa, Rio Solosuchil a 5-6 km ESE de
1852: 33. 1853. Type: COSTA RICA: Insula Hermanos Cedillo, 100-200 m, 24 Mar 1982,
San Lucas, Oersted 11621 (Lectotype C D. Lorence, T. Wendt, R. Riviere, M. Vazquez
(photos MEXU, PTBG), here designated; T., A. S. Ton & I. Navarrete 3910 (Holotype
Isolectotype C (lower right-hand collection MEXU; Isotypes BM, BR, CHAPA, ENCB,
numbered 11621, mounted on same sheet with F, MO, UC, XAL).—Psychotria
Syntype Oersted 11622). Syntypes: COSTA phanaerandra (Standi. & Steyerm.) Lorence
RICA: In monte Aguacate, Nov 1846, Oersted Psychotria luxurians Rusby, Mem. Torrey Bot.
11622 (C (photos MEXU, PTBG)); Aguacate, Club 6: 50. 1896. Type: BOLIVIA: Between
Oersted 11629 (C (photos MEXU, PTBG)). Tipuani and Guanai, Dec 1892, Bang 1741
Oersted 11621, a flowering specimen with a (Lectotype US 1393044 (photos PTBG), here
duplicate, is the most appropriate choice for
A NOMENCLATOR OF MEXICAN AND CENTRAL

designated; Isolectotypes US (2 sheets; photos Type: MEXICO: Jalisco, Santa Cruz de

PTBG)). Vallarta, 300 m, 9 Dec 1926, Y. Mexia 1262
Psychotria macrophylla Ruiz & Pav., Fl. (Holotype F 579924 (photos MEXU, PTBG);
Peruv. 2: 56, pi. 202, fig. a. 1799. Type: Isotypes A, n.v., C (photos MEXU, PTBG).
PERU: Huanuco, Andes of Peru, nemonbus GH (photo PTBG), MO (photos MEXU,
versus Pillao ad Iscutunam, Ruiz Lopez & PTBG), US (2 sheets; photos MEXU, PTBG;
Pavon s.n. (presumably MA, n.v.). fragment F)).
Psychotria macrophylla Ruiz & Pav. var. Psychotria mexicana Willd. ex Roem. &
angustissitna Standi., Publ. Field Mus. Nat. Schult., Syst. Veg. 5: 189. 1819. Type:
Hist., Bot. Ser. 22: 203. 1940. Type: MEXICO: Without precise locality, Humboldt
PANAMA: Darien, Cana-Cuasi Trail, north & Bonplands.n. (presumably B-WI, n.v.).
slope, 5500 ft., 16 Mar 1940, M. E. Terry & Uragoga.—Palicourea padifolia (Willd. ex
R. A. Terry 1597 (Holotype F 1034533 Roem. & Schult.) C. M. Taylor & Lorence
(photos MEXU, PTBG)).—Psychotria Psychotria micrantha Kunth in Humb., Bonpl.
macrophylla Ruiz & Pav. & Kunth, Nov. Gen. Sp. PI. 3: 363, pi. 284.
Psychotria macropoda Ruiz & Pav., Fl. Peruv. 1819. Type: PERU: A. Bonpland s.n.
2: 63, pi. 21 J, fig. b. 1799. Type: PERU: (Holotype P-Bonpland, n.v. (fragment F)).
Huanuco, nemoribus secus Pillao ad Mapouria, Uragoga.
Iscutuvam, Ruiz Lopez & Pavon s.n. Psychotria microbotrys Ruiz ex Standi., Publ.
(presumably MA, n.v.).—Geophila Field Mus. Nat. Hist., Bot. Ser. 8: 204. 1930.
macropoda (Ruiz & Pav.) DC. Type: PERU: Loreto, Williams 2983
Psychotria magna Standi., Contr. U.S. Natl. (Holotype F, n.v.).
Herb. 18: 131. 1916. Type: PANAMA: Colon, Psychotria microdesmia Oerst., Vidensk.
Loma de la Gloria, near Fato (Nombre de Meddel. Dansk Naturhist. Foren. Kjobenhavn
Dios), 10-104 m, Aug 1911, H. Pittier 4092 1852: 36. 1853. Type: COSTA RICA: Ad
(Holotype US 679188 (photos MEXU, Jaris, Oersted 11610 (Holotype C (photos
PTBG)).—Psychotria panamensis Standi, MEXU, MO, PTBG); Isotypes C (photos
var. compressicaulis (K. Schum.) C. W. Ham. MEXU, PTBG), US (photo PTBG)).
Psychotria marginata Sw., Prodr. Veg. Ind. Uragoga.—Psychotria viridis Ruiz & Pavon
Occ. 43. 1788. Type: JAMAICA: In Psychotria microdon (DC.) Urb., Symb. Antill.
sylvestribus, Swartz s.n. (Holotype S, n.v. 9: 539. 1928. Rondeletia microdon DC,
(photo A); possible Isotype C (photos MEXU, Prodr. 4: 408. 1830. Type: CUBA: Havana, de
PTBG)). Uragoga. la Ossa s.n. (Holotype G-DC, n.v. (microfiche
Psychotria maxonii Standi., Proc. Biol. Soc. IDC 800.691:111.7,8)).
Wash. 37: 53. 1924. Type: COSTA RICA: Psychotria mima Standi., Publ. Field Mus. Nat.
Cartago, Santa Clara de Cartago, 20 Jul 1903, Hist., Bot. Ser. 22: 386. 31 Oct 1940. Type:
W. R. Maxon & A. D. Harvey 8250 (Holotype COSTA RICA: Cartago, vicinity of Pejivalle,
US 1180093 (photos MEXU, PTBG)). 600-850 m, Jan 1940, A. F. Skutch 4589
Psychotria megalantha Lorence, Bol. Soc. Bot. (Holotype F 1044330 (photos MEXU,
Mexico 47: 58. 1987. Type: MEXICO: PTBG); Isotypes GH, n.v., MO (photos
Oaxaca, Distr. de Ixtlan, Sierra de Juarez, MEXU, PTBG), US (photos MEXU, PTBG).
Ruta 175 Tuxtepec a Oaxaca, 5 km NE de Nomen. illegit., not Psychotria mima Standi.
Vista Hermosa, 1300 m, 28 May 1983, D. 10 Sep 1940 [Type from Brazil].—Psychotria
Lorence & R. Cedillo T. 4190 (Holotype angustiflora K. Krause
MEXU; Isotypes BM, BR, CAS, ENCB, F, K, Psvchotria minarum Standi. & Steyerm., Publ.
MEXU, MO, NY, UC, US, WIS, TEX, XAL). Field Mus. Nat. Hist., Bot. Ser. 23: 253. 1947.
Psychotria mexiae Standi., Publ. Field Type: GUATEMALA: El Progreso (Sierra de
Columbian Mus., Bot. Ser. 4: 296. 1929. las Minas), between Calera and summit of
 Volcan Siglo, 2000-3000 m, 21 Jan 1942, (Holotype B, presumably destroyed).
J. A. Steyermark 43106 (Holotype F 1133839 —Psychotria hebeclada DC.
(photos MEXU, PTBG); Isotype US (photos Psychotria mollis Poir. in Lam., Encycl. Meth.
MEXU, PTBG)). Bot. 5: 701. 1804. Type: MEXICO: Without
Psychotria miradorensis (Oerst.) Hemsl., Biol. precise locality, Haenke s.n. (Holotype P-LA
Centr.-Amer., Bot. 2: 50. 1881. Mapouria (photos MEXU, MO, PTBG)).—Psychotria
miradorensis Oerst., Amir. Centr. 17, tab. 14,
fig. 9. 1863. Type: MEXICO: Veracruz, Psychotria mombachensis Standi., Publ. Field
Mirador, in quercetum, 3000 ft., F. Liebmann Mus. Nat. Hist., Bot. Ser. 8: 188. 1930. Type:
11662 (Lectotype C (photos MEXU, PTBG); NICARAGUA: Mombacho Volcano,
Isolectotype US, n.v.; designated by 600-750 m, 5 Jul 1923, W. R. Maxon, A. D.
Hamilton, 1989). Syntypes: MEXICO: Harvey & A. T. Valentine 7818 (Holotype F
Veracruz, Mirador, Apr 1847, F. Liebmann 600068 (photos MEXU, PTBG); Isotype US
11577 (C (photos MEXU, PTBG)); Mirador, (photos MEXU, PTBG)).—Psychotria
F Liebmann 11614 (C (photos MEXU, domingensis Jacq.
PTBG)). Psychotria, Uragoga.—Psychotria Psychotria monteverdensis Dwyer & C. W.
costivenia Griseb. var. costivenia Ham., Phytologia 64: 230. 1988. Type:
Psychotria mirandae C. W. Ham., Phytologia COSTA RICA: Guanacaste-Puntarenas border
64: 229. 1988. Type: MEXICO: Chiapas, region, Cordillera de Tilaran, Monteverde, en
Berriozabal a las Vistas, 17 Jul 1949, F. cumbre de fila al oeste de Reserva,
Miranda 5395 (Holotype US, n.v.; Isotype 1520-1590 m, 24 Jun 1977, V. J. Dryer 1531
MEXU). (Holotype MO 3235523 (photo PTBG);
Psychotria molinae Standi., Ceiba 1: 46. 1950. Isotype CR).
Type: HONDURAS: Morazan, drainage of Psychotria morae Pol., Linnaea 41: 570. 1877.
the Rio Yeguare, at about longitude 87 °W and Type: COSTA RICA: San Jose, Jun 1875, H.
latitude 14°N, in cloud forest on Mt. Uyuca, Polakowski 171 (Holotype B, presumably
2000 m, 20 Aug 1946, L. O. Williams & A. destroyed (photos GH, US)). Uragoga.
Molina R. 10390 (Holotype F 1264188 —Psychotria quinqueradiata Pol.
(photos MEXU, PTBG); Isotypes EAP, n.v., morii Dwyer, Ann. Missouri Bot.
GH, n.v., MO (2 sheets; photos MEXU, Gard. 67: 400. 1980. Type: PANAMA:
PTBG), US (photos MEXU, PTBG)). Veraguas, 11 km from Escuela Agricola Alto
—Psychotria panamensis Standi, var. de Piedra, along Rio Dos Bocas, Atlantic
panamensis slope, 15 Nov 1974, S. Mori & J. Kallunki
Psychotria molinarum Lorence, Novon 7: 51. 3139 (Holotype MO 2932202 (photo PTBG)).
1997. Based on Evea chiapensis Standi., —Psychotria siggersiana Standi.
Contr. U.S. Natl. Herb. 23: 1392. 1926. Not Psychotria mortoniana Standi., Publ. Field
Psychotria chiapensis Standi., 1926. Type: Mus. Nat. Hist., Bot. Ser. 18: 355. 1938.
MEXICO: Chiapas, Cerro del Boqueron, C. A. Type: COSTA RICA: San Jose, vicinity of El
Purpus 6928 (Holotype US 567214 (photos General, 950 m, Jul 1936, A. F. Skutch 2697
MEXU, PTBG); Isotypes F, MO (photos (Holotype US 1642538 (photos F, MEXU,
MEXU, PTBG)). PTBG; fragment F); Isotypes GH, n.v., MO
Psychotria molliramis (K. Schum. & K. Krause) (photos MEXU, PTBG)).
Steyerm., Mem. New York Bot. Gard. 23: Psychotria muscosa (Jacq.) Steyerm, Mem.
529. 1972. Palicourea molliramis K. Schum. New York Bot. Gard. 23: 671. 1972. Morinda
& K. Krause, Engl. Bot. Jahrb. 40: 337. 1908. muscosa Jacq., Sel. Stirp. Am. 65. 1763.
Type: COLOMBIA: Cauca, prope La Junta ad Type: MARTINIQUE: Jacquin s.n. (Location
Rio d'Agua, 200-500 m, Lehmann 4667
 L NOMENCLATOR OF MEXICAN AND CENTRAL

Psychotria nebulosa (Dwyer) C. M. Taylor, Galeotti 7066 (Holotype BR, n.v.; Isotypes G-
Novon 5: 205. 1995. Coussarea nebulosa DC (photo MO), P (label states "cerca
Dwyer, Ann. Missouri Bot. Gard. 67: 131. Xalapa," photos MEXU, PTBG)).
1980. Type: PANAMA: Chiriqui, Monte Rey —Hoffmannia nicotianifolia (M. Martens &
near Boquete, 20 Jul 1971, T. B. Croat 15868 Galeotti) L. O. Williams
(Holotype MO 2162999 (photo PTBG); / nubiphila Dwyer, Ann. Missouri
Isotype MO 4043108). Bot. Gard. 67: 403. 1980. Type: PANAMA:
Psychotria neillii C. W. Ham. & Dwyer, Chiriqui, NW side of Cerro Pando, T. B.
Phytologia 64: 231. 1988. Type: Croat 15990 (Holotype MO).—Psychotria
NICARAGUA: Rio San Juan, Rio Sabalos, 2 panamensis Standi, var. panamensis
km al W de Santa Eduviges, 80 m, 18 Feb Psvclwtria oaxacam Standi., Contr. U.S. Natl.
1984, P. P. Moreno 23060 (Holotype MO Herb. 23: 1391. 1926. Type: MEXICO:
3192079 (photo PTBG)). Oaxaca, from near Santo Domingo, 1600 ft.,
Psychotria nervosa Sw., Prodr. 43. 1788. Type: 18 Jun 1895, E. W. Nelson 2688 (Holotype
JAMAICA: Without precise locality. Swart: US 574444 (photos MEXU, PTBG)).
s.n. (Holotype S, n.v. (photo A)). —Psychotria gracilifiora Benth. in Oerst.
Psychotria nervosa Sw. var. rufescens (Kunth) Psychotria obovata (Oerst.) Hemsl., Biol.
L. O. Williams, Phytologia 26: 492. 1973. Centr.-Amer., Bot. 2: 50. 1881. Mapouria
Psychotria rufescens Kunth in Humb., Bonpl. obovata Oerst., Amer. Centr. 17, tab. 14, figs.
& Kunth, Nov. Gen. Sp. 3: 364. 1819. Type: 3, 4. 1863. Not Psychotria obovata Ruiz &
VENEZUELA: Caripe, Humboldt & Pav„ 1799. Type: MEXICO: Veracruz, F.
Bonplands.n. (Holotype P-Bonpland, n.v.; Liebmann s.n. (Holotype C, n.v.).
Isotype B, presumably destroyed (photos F, —Psychotria quinqueradiata Pol.
MO))—Psychotria nervosa Sw. Psychotria oerstediana (Kuntze) Standi., Contr.
Psychotria nervosa Sw. subsp. rufescens U.S. Natl. Herb. 23: 1390. 1926. Uragoga
(Kunth) Steyerm., Mem. New York Bot. oerstediana Kuntze, Rev. Gen. PI. 2: 957.
Gard. 23: 480. 1972. Psychotria rufescens 1891. Based on Mapouria obovata Oerst.,
Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Amer. Centr. 17, tab. 14, figs 3, 4. 1863.
Sp. 3: 364. 1819. Type: VENEZUELA: Type: MEXICO: Veracruz, F. Liebmann s.n.
Caripe, Humboldt & Bonplands.n. (Holotype (Holotype C, n.v.).—Psychotria
P-Bonpland, n.v.; Isotype B, presumably quinqueradiata Pol.
destroyed (photos F, MO)).—Psychotria Psvchotria officinalis (Aubl.) Raeusch. ex
nervosa Sw. Sandwith, Kew Bull. 1931: 473. 1931.
Psychotria nicaraguensis Benth. in Oerst., Nonatelea officinalis Aubl., Hist. PI. Guiane
Viddensk. Meddel. Dansk Naturhist. Foren. 1: 182, tab. 70. 1775. Type: FRENCH
Kjobenhavn 1852: 34. 1853. Type: GUIANA: Cayenne, Aublet s.n. (presumably
NICARAGUA: Rio San Juan, Oersted 11634 BM, n.v.).
(Lectotype C, here designated (photos MEXU, Psychotria olgae Dwyer & Hayden, Ann
PTBG)). Syntype: COSTA RICA: Ad Jaris, Missouri Bot. Gard. 55: 42, fig. 3. 1968. Type:
Oersted 11635 (C (photos MEXU, PTBG)). PANAMA: Panama, between Cerro Jefe and
Oersted 11634 is the better choice for Eneida, 2100-2900 ft., 17 Jan 1968, J. D.
lectotype as it is a well developed fruiting Dwvcr. J. Duke & R. Dressier 8193 (Holotype
specimen. Uragoga.—Psychotria marginata MO 1962026 (photos MEXU, PTBG);
Isotypes DUKE, n.v., F, NY, n.v.).
Psychotria nicotianifolia M. Martens & Psvchotria orchidearum Standi.. J. Wash.
Galeotti, Bull. Acad. Roy. Sci. Bruxelles 11: Acad. Sci. 18: 276. 1928. Type: COSTA
229. 1844 [as nicotianaefolia]. Type: RICA: Heredia, Cerros de Zurqui, northeast of
MEXICO: Veracruz, Mirador, 3000 ft., H. San Isidro, 2000-2400 m, 3 Mar 1926, P. C.
 Standley & J. Valerio 50863 (Holotype US Villavicencio, Triana s.n. (Holotype BM, n.v.;
1306503 (photos MEXU, PTBG)). Isotype F).
Psychotria orchidearum Standi, subsp. Psychotria pachecoana Standi. & Steyerm.,
persedens Dwyer, Ann. Missouri Bot. Gard. Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 205.
67: 407. 1980. Type: PANAMA: Panama, 1940. Type: GUATEMALA: Baja Verapaz,
Cerro Campana, 2500 ft., 7 Feb 1969, E. L. mountain side of divide north of Santa Rosa,
Tyson 5331 (Holotype MO 1973872). 1650 m, 30 Mar 1939, P. C. Standley 69922
—Status uncertain, probably Psychotria (Holotype F 994220 (photos MEXU, PTBG)).
orchidearum Standi. —Psychotria galeottiana (M. Martens) C. M.
Psychotria oreodoxa L. O. Williams, Taylor & Lorence
Phytologia 28: 230. 1974. Type:
GUATEMALA: Suchitepequez, south-facing Phytologia 64: 232. 1988. Type: PANAMA:
slopes and barrancos of Volcan Santa Clara, Panama, vicinity of Cerro Jefe, Altos de
1 Vz-2 miles west of Finca El Naranjo, 1250 m, Pacora, 2400 ft., 18 Dec 1980, T. Antonio
1 Jun 1942,7. A. Steyermark 46803 (Holotype 3241 (Holotype MO 2892015 (photo PTBG);
F 1133947 (photos MEXU, PTBG)). Isotype ENCB, n.v.).
Psychotria oresbia Dwyer, Ann. Missouri Bot. Psychotria padifolia Willd. ex Roem. & Schult.,
Gard. 67: 408. 1980. Type: PANAMA: Syst. Veg. 5: 189. 1819. Type: MEXICO:
Darien, Cerro Tacarcuna W ridge, trail toward Veracruz, Xalapa, Humboldt & Bonpland
Rio Pucuro just below summit camp, 4420 (Holotype B-WI (photos DUKE, F, MO,
1500-1600 m, 5 Feb 1975, A. Gentry & S. NY)).—Palicourea padifolia (Willd. ex
Mori 14150 (Holotype MO 2280279 (photos Roem. & Schult.) C. M. Taylor & Lorence
MEXU, PTBG)). Psychotria panamensis Standi., Contr. U.S.
Psychotria owgenes L. O. Williams, Phytologia Natl. Herb. 18: 132. 1916. var. panamensis.
28: 231. 1974. Type: GUATEMALA: Baja Type: PANAMA: Chiriqui, humid forest
Verapaz, Sierra de las Minas about 5 km south around Los Siguas Camp, southern slope of
of Purulha, 1600 m, 2 Jan 1973, L. O. Cerro de la Horqueta, 1700 m, 17-19 Mar
Williams. A. Molina R. & T. P. Williams 1911,//. Pittier 3194 (Holotype US 677588
41924 (Holotype F 1722437 (photos MEXU, (photos MEXU, PTBG)).
PTBG)).—Psychotria galeottiana (M. Psychotria panamensis Standi, var.
Martens) C. M. Taylor & Lorence compressicaulis (K. Krause) C. W. Ham.,
Psychotria orosiana Standi., J. Wash. Acad. Phytologia 64: 233. 1988. Psychotria
Sci. 15: 288. 1925. Type: COSTA RICA: compressicaulis K. Krause, Bot. Jahrb. Syst.
Cartago, vicinity of Orosi, 30 Mar 1924, P. C. 54, Beibl. 119: 44. 1916. Type: COSTA
Standley 39803 (Holotype US 1153182 RICA: Cartago, Tuis, 600 m, Jul 1898, H.
(photos MEXU, PTBG)). Pittier 12412 (Holotype B, destroyed (photo
Psychotria ostanrea Dwyer & Hayden, Ann. GH); Isotype US (photos MEXU, PTBG)).
Missouri Bot. Gard. 54: 143. 1967. Type: Psychotria panamensis Standi, var. ixtlanensis
PANAMA: Bocas del Toro, vicinity of C. W. Ham., Phytologia 64: 233. 1988. Type:
Chinqui Lagoon, Water Valley, 1 Nov 1940, MEXICO: Oaxaca, Distr. de Ixtlan, 21.4 km S
H. von Wedel 1461 (Holotype MO 1240228 de Valle Nacional, 1140 m, 28 Nov 1979, T.
(photos MEXU, PTBG); Isotype F). Wendt et al. 2258 (Holotype MEXU 317322;
—Coussarea hondensis (Standi.) C. M. Isotypes CHAPA, ENCB).
Taylor & W. C. Burger Psychotria panamensis Standi, var. magna
Psychotria ostreophora (Wernh.) C. M. (Standi.) C. W. Ham., Phytologia 64: 234.
Taylor, Novon 4: 305. 1994. Cephaelis 1988. Psychotria magna Standi., Contr. U.S.
ostreophora Wernh., J. Bot. 55: 284. 1917. Natl. Herb. 18: 131. 1916. Type: PANAMA:
Type: COLOMBIA: Bogota, Servita y Colon, Loma de la Gloria, near Fato (Nombre
 ^ NOMENCLATOR OF MEXICAN AND CENTRA!

de Dios), 10-104 m, Aug 1911, H. Pittier 41952 (Holotype F 1132612 (photo PTBG);
4092 (Holotype US 679188 (photos MEXU, Isotype US).
PTBG); Isotype US 693188). Psychotria philacra Dwyer, Ann. Missouri Bot.
Psychotria papantlensis (Oerst.) Hemsl., Biol. Gard. 67: 412. 1980. Type: PANAMA:
Centr.-Amer., Bot. 2: 50. 1881. Mapouria Darien, Cerro Tacarcuna, S slope, ridge top
papantlensis Oerst., Amer. Centr. 17, tab. 14, forest well below summit, 1250-1450 m, 26
fig. 5. 1863. Type: MEXICO: Veracruz, Jan 1975, A. Gentry & S. Mori 13916
Papantla, F. Liebmann s.n. (Holotype (Holotype MO 2280220 (photo PTBG)).
presumably C, n.v.). Uragoga. Psychotria phoeniciana Standi., J. Wash. Acad.
Psychotria parasitica Sw., Prodr. Veg. Ind. Occ. Sci. 16: 17. 1926. Type: MEXICO: Chiapas,
44. 1788, nomen illegit. citing Viscoides mountains near Fenix, May 1925, C. A.
pendulum Jacq. 1763. (see Steyermark, 1972: Purpus 316 (Holotype US 1208247 (photos F,
569 for discussion). Uragoga.—Psychotria MEXU, PTBG)).—Palicourea macrantha
guadalupensis (DC.) Howard Loes.
Psychotria parvifolia Benth. in Oerst., Psychotria pilosa Ruiz & Pav., Fl. Peruv. 2: 60,
Vidensk. Meddel. Naturhist. Foren. pi. 208, fig. 1. 1799. Type: PERU: Ruiz Lopez
Kjobenhavn 1852: 36. 1853. Type: COSTA & Pavon s.n. (Holotype presumably MA, n.v.;
RICA: Ad Naranjo, Apr 1847, Oersted 11637 Isotype B, n.v (photo MO)).
(Holotype C (photos F, MEXU, MO, PTBG); Psychotria pinularis Sesse & Moc., Fl. Mexic,
Isotype US (photos MEXU, PTBG)). ed. 2, 57. 1894. Type: PUERTO RICO:
Psychotria peperomiae Standi., Contr. U.S. Habitat in monticulo de la Aguadilla vicino,
Natl. Herb. 18: 132. 1916. Type: PANAMA: Sesse & Mocifio s.n. (presumably MA, n.v.).
Chiriqui, top of Cerro de la Horqueta, 2265 m, —Psychotria microdon (DC.) Urb.
18 Mar 1911, H. Pittier 3235 (Holotype US Psvchotria pithecobia Standi., Publ. Field
677632, n.v.).—Psychotria guadalupensis Columbian Mus., Bot. Ser. 8: 187. 1930.
(DC.) Howard Type: COSTA RICA: La Palma, 1550 m, 15
Psychotria perparva Dwyer, Ann. Missouri Aug 1898, A. Tonduz 12432 (Holotype B,
Bot. Gard. 67: 411. 1980. Type: PANAMA: presumably destroyed; Isotype US (photos
Panama, Cerro Tacarcuna Massif between MEXU, PTBG)).
summit camp on westernmost ridge and Pico Psvchotria pittieri Standi., Contr. U.S. Natl.
Tacarcuna, mostly elfin forest, 1650-1800 m, Herb. 18: 132. 1916. Type: PANAMA: Canal
22 Jul 1976, A. Gentry, H. Leon & L. Forero Zone, railroad relocation between Gorgona
16921 (Holotype MO 2416828 (photos and Gatun, 10-50 m, 7 Jan 1911, H. Pittier
MEXU, PTBG); Isotype F). 2266 (Holotype US 676523 (photos MEXU,
Psychotria persearum Standi., Ceiba 1: 48. PTBG)).
1950. Type: HONDURAS: Morazan, Cerro de Psychotria pittieri Standi, subsp. oinochrophylla
Uyuca, ca. 1800 m, 2 Mar 1947, P. C. Dwyer, Ann. Missouri Bot. Gard. 67: 416.
Standley 4865 (Holotype F 1256865 (photos 1980. Type: PANAMA: Darien, Cerro Pirre,
MEXU, PTBG); Isotype EAP, n.v.). cloud forest and/or mossy forest, 2500-4000
—Psychotria galeottiana (M. Martens) C. M. ft., 9-10 Aug 1967, J. A. Duke & T. Elias
Taylor & Lorence 13668 (Holotype MO 2138646 (photos
Psychotria phanaerandra (Standi. & Steyerm.) MEXU, PTBG))—Status uncertain,
Lorence, Novon 2: 260. 1992. Palicourea Psvchotria subgen. Heteropsychotria.
phanaerandra Standi. & Steyerm., Publ. Field Psychotria platypoda DC, Prodr. 4: 510. 1830.
Mus. Nat. Hist., Bot. Ser. 23: 252. 1947. Type: GUYANA: Patris s.n. (presumably G-
Type: GUATEMALA: Izabal, Cerro San Gil,
1200 m, 26-27 Dec 1941, J. A. Steyermark
 GUATEMALA: Alta Verapaz, Cubliquitz, Psychotria purpureocapitata Dwyer ex C. M.
350 m, Aug 1903, H. von Tuerckheim 8529 Taylor, Novon 6: 301, fig. 2A, B. 1996. Type:
(Lectotype GH, n.v. (fragment F; photos PANAMA: Code, El Cope on Pacific side, lA
MEXU, PTBG); Isolectotype US (photos hour walk from sawmill, 2400 ft. [774 m], 16
MEXU, PTBG); designated by Hamilton, Oct 1979, T. Antonio 2154 (Holotype PMA,
1989). n.v.; Isotype MO 4658112).
Psychotria poeppigiana Mull. Arg., Fl. Bras. Psychotria purpusii Standi., Contr. U.S. Natl.
6(5): 370. 1881. Syntypes: BRAZIL: Herb. 23: 1388. 1926. Type: MEXICO:
Amazonas Habitat in regione superiore Chiapas, Cerro del Boqueron, Aug 1913, C. A.
fluminis Amazonum prope Ega, Poeppig Purpus 7012 (Holotype US 567269 (photos
3065 (presumably M, n.v.), Martius s.n. MEXU, PTBG); Isotypes F (photos MEXU,
(presumably M, n.v.); in sylvis Japurensibus PTBG), GH, n.v., MO (photos MEXU,
ad Maribi, Martius s.n. (presumably M, n.v.); PTBG)).
in regione Rio Negro ad Uanaca et S. Gabriel, Psychotria quadrangulata Dwyer, Ann.
Spruce IS92 (presumably M, n.v.); prope Missouri Bot. Gard. 67: 1980. Type:
Barra do Rio Negro. Martius s.n. (presumably PANAMA: Veraguas, NW. of Santa Fe, 2.7
M, n.v.); in silvis ad Nagueria, Martius s.n. km from Escuela Agricola Alto de Piedras,
(presumably M, n.v.). road to Calovebora, S. Mori & J. Kallunki
Psychotria polyphobia Donn. Sm., Bot. Gaz. 5363 (Holotype MO (not located 6/1996)).
(Crawfordsville) 33: 253. 1902. Type: Psychotria quiinifolia Dwyer, Ann. Missouri
COSTA RICA: Suerre, Llanuras de Santa Bot. Gard. 53: 108. 1966. Type: PANAMA:
Clara, 300 m, Apr 1896, J. D. Smith 6854 Bocas del Toro, Almirante, Changuinola
(Lectotype US 943508 (photos MEXU, Canal, K. Blum 1402 (Holotype MO 2448117
PTBG), here designated; Isolectotype US (photo PTBG)).—Psychotria nervosa Sw.
943509 (photos MEXU, PTBG). Syntype: Psychotria quinqueradiata Pol, Linnaea 41:
COSTA RICA: Talamanca, forets au-dessus 570. 1877. Type: COSTA RICA: San Jose, 24
de Tsaki, 500 m, Apr 1895, A. Tonduz 9577 Jun 1875, H. Polakowski 135 (Lectotype B,
(P (photos MEXU, PTBG)). Smith 6854, with destroyed (photos BH, MO, US); designated
abundant flowers and fruit and a duplicate at by Hamilton, 1989). Uragoga
US, is the better choice for lectotype. Psychotria racemosa Rich., Actes Soc. Hist.
Psychotria psychotriifolia (Seem.) Standi., Nat. Paris 1: 107. 1792. Type: FRENCH
Contr. U.S.Natl. Herb. 18: 133. 1916. GUIANA: L. C. M. Richard s.n. (Neotype P,
Cephaelis psychotriifolia Seem., Bot. Voy. n.v.; designated by Kirkbride, 1997).
Herald 138. 1852 [as psychotriaefolia]. Type: Psychotria racemosa (Aubl.) Raeusch., Nomen.,
PANAMA: Panama, dense forests near ed. 3, 56. 1797, not Rich. 1792. Nonatelia
Cruces; and Darien, Chirambira, Seemann 482 racemosa Aubl., Hist. PI. Guiane 187, t. 72.
(Holotype BM, n.v. (photo MO); Isotype K Ml5. Type: FRENCH GUIANA: In sylvis
[as Seemann 482bis)). territoni Orapu, Aublet s.n. (Holotype P, n.v.).
Psychotria pubescens Sw., Prodr. Veg. Ind. —Psychotria racemosa Rich.
Occ. 44. 1788. Type: JAMAICA: A neotype Psychotria ramonensis Standi., Publ. Field Mus.
needs to be designated according to Howard Nat. Hist., Bot. Ser. 18: 1360. 1938. Type:
(1989). COSTA RICA: La Palma de San Ramon,
Psychotria pumilocarpa Dwyer, Ann. Missouri 1250 m, Nov 1925, A. Brenes 4599 (Holotype
Bot. Gard. 67: 419. 1980. Type: PANAMA: F 852181 (photos MEXU, PTBG)).
Veraguas, Alto de Piedra, 800 m, 7 Sep 1974, —Psychotria hazenii Standi.
P. J. M. Maas & R. L. Dressier 1612 Psychotria remota Benth., J. Bot. Hook. 3:
(Holotype MO 2287628). 225. 1841. Type: BRAZIL: On the Rio Negro,
 k. NOMENCLATOR OF MEXICAN AND CENTRAL

Schomburgk 963 (Holotype presumably K, 5000-5500 ft., S. Mori & J. Kallunki 5871
n.v. (photo NY)). Mapouria (Holotype MO 3646500)
Psychotria roseocremea (Dwyer) C. M. Taylor, Psychotria santaritensis Dwyer, Ann. Missouri
Novon 5: 205. 1995. Coussarea roseocremea Bot. Gard. 424. fig. 78. 1980. Type:
Dwyer, Ann. Missouri Bot. Gard. 67: 133. PANAMA: Colon, Santa Rita, near Agua
1980. Type: PANAMA: Panama, El Llano- Clara weather station, 23 Mar 1972, J. D.
Carti highway, 6-10 km N of El Llano, 13 Dwyer & A. Gentry 9565 (Holotype MO
Apr 1973, R. L. Dressier 4334 (Holotype MO 2183828 (photos MEXU, PTBG)).
2126258 (photo PTBG)). Psychotria santarosae Standi., Publ. Field
Psychotria rosulatifolia Dwyer, Ann. Missouri Columbian Mus., Bot. Ser. 7: 113. 1930.
Bot. Gard. 67: 423, fig. 77. 1980. Type: Type: COLOMBIA: El Valle, Santa Rosa,
PANAMA: Darien, Cerro Tacarcuna, trail Dagua Valley, 200-300 m, 22 Sep 1922, E. P.
from Rio Pucuro base camp up west ridge of Killip 11530 (Holotype US 1140714).
Cerro Mali, 640-1000 m, 6 Feb 1975, /i. Psychotria sarapiquensis Standi., Publ. Field
Gentry & S. Mori 14174 (Holotype MO Mus. Nat. Hist., Bot. Ser. 18: 1360. 1938.
2276271 (photos MEXU, PTBG)). Type: COSTA RICA: Vara Blanca de
Psychotria rufescens Kunth in Humb., Bonpl. & Sarapiqui, north slope of Central Cordillera,
Kunth, Nov. Gen. Sp. 3: 364. 1819. Type: 1500 m, Jul-Sep 1937, A. F. Skutch 3330
VENEZUELA: Canpe, Humboldt <£ (Holotype US 1643233 (photos F, MEXU,
Bonpland s.n. (Holotype presumably P- PTBG); Isotypes GH (photo PTBG), MO).
Bonpland, n.v.; Isotype B, presumably Psychotria scabriuscula Bartl. ex DC, Prodr. 4:
destroyed (photos F, MO)). Uragoga. 513. 1830. Type: MEXICO: Guerrero,
—Psychotria nervosa Sw. Acapulco, Haenke s.n. (Holotype presumably
Psychotria rufescens Kunth var. haenkeana PR, n.v.).—Psychotria pubescens Sw.
DC, Prodr. 4: 512. 1930. Type: MEXICO: Psychotria schippii Standi. & Steyerm., Publ.
Without precise locality, Haenke s.n. Field Mus. Nat. Hist., Bot. Ser. 23: 24. 1943.
(Holotype presumably PR, n.v.).—Status Type: BRITISH HONDURAS (BELIZE): El
uncertain, probably Psychotria nervosa Sw. Cayo district, Valentin, in advanced forest,
Psychotria salicifolia (Oerst.) Hemsl., Biol. limestone valley, Jun-Jul 1936, C. L. Lundell
Centr.-Amer., Bot. 2: 51. 1881. Mapouria 6193 (Holotype F 861234 (photos MEXU,
salicifolia Oerst., Amer. Centr. 17, tab. 14, PTBG); Isotypes GH, n.v., NY, n.v., US
fig- 2. 1863. Not Psychotria salicifolia Kunth (photo PTBG)).—Psychotria mexiae Standi.
in Humb., Bonpl. & Kunth, 1819. Type: Psychotria sessilifolia M. Martens & Galeotti,
MEXICO: Oaxaca, Lacoba [Lacova], F. Bull. Acad. Roy. Sci. Bruxelles 11: 228. 1844.
Liebmann (Holotype presumably C, n.v.). Type: MEXICO: Veracruz, Miradoret
—Psychotria papantlensis (Oerst.) Hemsl. Zacuapan [label on isotype states "Xalapa"],
Psychotria saltatrix C. M. Taylor, Novon 5: 3000 ft., H. Galeotti 7078 (Holotype
205, fig. 2. 1995. Type: COSTA RICA: presumably BR, n.v.; Isotypes G-DEL, n.v.
Cartago, Refugio Nacional Tapanti, por la (photo MO), P (photos MEXU, PTBG)).
orilla del Rio Grande de Orosi, 09°45'N, Uragoga.—Psychotria tenuifolia Sw.
83°47'W, 14 Aug 1991, B. Hammel et al. Psychotria siccorubra Dwyer, Ann. Missouri
18328 (Holotype CR, n.v.; Isotypes F, n.v., Bot. Gard. 67: 428. 1980. Type: PANAMA:
MO 4619859). Veraguas, Cerro Tute, ca. 10 km NW of Santa
Psychotria sanfelicensis Dwyer, Ann. Missouri Fe, on ridgetop in cloud forest, above 1000 m,
Bot. Gard. 67: 424. 1980. Type: PANAMA: 19Jun 1975, S. Mori 6781 (Holotype MO
Chiriqui, N of San Felix on Cerro Colorado 2272240 (photos MEXU, PTBG)).
copper mine road, along continental divide, —Psychotria ostreophora (Werhnam) C. M.
Taylor
Psychotria siggersiana Standi., J. Wash. Acad. between Santa Maria de Jesus and
Sci. 15: 289. 1925. Type: COSTA RICA: Calahuache, along great barranco between
Limon, vicinity of Guapiles, 300-500 m, Mar Finca Pireneos and San Juan Patzulin,
1924, P. C. Standley 37210 (Holotype US 1300-1400 m, 6 Jan 1940, J. A. Steyermark
1153058 (photos MEXU, PTBG)). 33700 (Holotype F 1043219 (photos MEXU,
Psychotria simiarum Standi., Publ. Field PTBG)).
Columbian Mus., Bot. Ser. 4: 344. 1929. Psychotria stipulata (Oerst.) Hemsl., Biol.
Type: HONDURAS: Atlantida, Lancetilla Centr.-Amer., Bot. 2: 51. 1881. Mapouria
Valley, near Tela, 20-600 m, 6 Dec 1927-20 stipulata Oerst., Amer. Cent. 17, tab. 14, fig.
Mar 1928, P. C. Standlev 56772 (Holotype F 7. 1863. Type: MEXICO: Veracruz, Papantla,
581799 (photos MEXU, PTBG); Isotype US Liebmann 11615 (Lectotype C, here
(photos MEXU, PTBG)). designated (photos MEXU, PTBG);
Psychotria sixaolensis C. W. Ham., Phytologia Isolectotype C (photos MEXU, PTBG)).
64: 234. 1988. Type: COSTA RICA: Limon, Syntype: MEXICO: Veracruz, Papantla,
along road between Limon and Shiroles, Liebmann 11636 (C (photos MEXU, PTBG)).
along Rio Sixaola, 0.9 miles SW of Bambu, Liebmann 11615, a flowering specimen with a
6.5 miles SW of Bribni, less than 50 m alt., 12 duplicate, is the more suitable choice for
Aug 1977, T. B. Croat 43303 (Holotype MO lectotype.—Psychotria costivenia Griseb.
2587929 (photo PTBG)).
Psychotria skutchii Standi, Field Mus. Nat. Psychotria stockwellii C. W. Ham., Phytologia
Hist., Bot. Ser. 17:283. 1937. Type: 64: 235. 1988. Type: COSTA RICA: Alajuela,
GUATEMALA: San Juan Ixcoy, oak woods, region of Zarcero, 1800 m, 18 Oct 1937, A.
8000 ft., 22 Aug 1924, A. F. Skutch 1074 Smith 543 (Holotype US; Isotype F).
(Holotype F 813469 (photos MEXU, PTBG); Psychotria subsessilis Benth. in Oerst., Vidensk.
Isotype A, n.v.).—Psychotria galeottiana Meddel. Dansk Naturhist. Foren. Kjobenhavn
(M. Martens) C. M. Taylor & Lorence 1852: 32. 1853. Type: COSTA RICA: Ad
Psychotria solitudinum Standi., Publ. Field Turrialba, Oersted 11613 (Holotype C (photos
Mus. Nat. Hist., Bot. Ser. 22: 207. 1940. MEXU, PTBG): Isotype US (photos MEXU,
Type: COSTA RICA: San Jose, Valley of El PTBG)).—Psychotria costivenia Griseb. var.
General, 700 m, Jun 1939, A. F. Skutch 4347
(Holotype F 974021 (photos MEXU, PTBG); Psychotria suerrensis Dorm. Sm., Bot. Gaz.
Isotype US (photos MEXU, PTBG)). (Crawfordsville) 27: 337. 1899. Type:
Psychotria sousae Lorence & Dwyer, Bol. Soc. COSTA RICA: Suerre, Llanuras de Santa
Bot. Mexico 47: 59. 1987. Type: MEXICO: Clara, 300 m, Feb-Apr 1896, J. D. Smith
Oaxaca, Mun. de Matias Romero, zona de 6580 (Syntype US 943504 (photos MEXU,
Uxpanapa, lomas al E de Arroyo Hamaca, 9.5 PTBG), US 943405 (photos MEXU, PTBG));
km al SE de La Floresta, a 21.5 km S de Tonduz 8683 (Syntype US, n.v.).
Esmeralda, 400 m, 22 May 1981, T. Wendt, A. Psychotria sylvivaga Standi., J. Wash. Acad.
Villalobos, I. Navarrete & J. Anguiano 3282 Sci. 18: 274. 1928. Type: COSTA RICA:
(Holotype MEXU; Isotypes CHAPA, PTBG). Heredia, Yerba Buena, northeast of San
Psychotria stenostachya Standi., Publ. Field Isidro, ca. 2000 m, 28 Feb 1926, P. C.
Columbian Mus., Bot. Ser. 8: 207. 1930. Standley & J. Valerio 49989 (Holotype US
Type: PERU: Loreto, Killip & Smith 29294 1306274 (photos MEXU, PTBG)).
(Holotype F, n.v.). Psychotria tacarcunensis Dwyer, Ann.
Psychotria steyermarkii Standi., Publ. Field Missouri Bot. Gard. 67: 431. 1980. Type:
Mus. Nat. Hist., Bot. Ser. 22: 387. 1940. PANAMA: Darien, elfin forest at top of W
Type: GUATEMALA: Quezaltenango, lower peak of Cerro Tacarcuna massif, 1800-1850
south-facing slopes of Volcan Santa Maria, m, 28 Jan 1975, A. Gentry & S. Mori 13974
 L NOMENCLATOR OF AND CENTRAL AMERICAN RUBIACE

(Holotype MO 2242943 (photos MEXU, (photos MEXU, PTBG)).—Psychotria

PTBG)). aggregata Standi.
Psychotria tapantiensis C. M. Taylor in W. C. PsYchotna torrcsiami Standi., J. Wash. Acad.
Burger & C. M. Taylor, Fieldiana, Bot., n.s. Sci. 15: 288. 1925. Type: COSTA RICA:
33: 275. 1993. Type: COSTA RICA: Cartago, Cartago, vicinity of Orosi, 30 Mar 1924, P. C.
Reserva Tapanti, 11 km above the Tapanti Standley 39769 (Holotype US 1153178
bridge, 10 Aug 1980, R. L. Wilbur 30741 (photos MEXU, PTBG)).—Psychotria
(Holotype DUKE, n.v.). goldmanii Standi.
Psychotria taurina Dwyer, Ann. Missouri Bot. Psychotria trichotoma M. Martens & Galeotti,
Gard. 67: 432. 1980. Type: PANAMA: Bocas Bull. Acad. Roy. Sci. Bruxelles 11: 227. 1844.
del Toro, Fish Creek Mountains, vicinity of Type: MEXICO: Veracruz, Xalapa and
Chiriqui Lagoon, 16 Apr 1941, H. von Wedel Mirador, 4000 ft., H. Galeotti 7092 (Holotype
2240 (Holotype MO 1245978 (photos MEXU, presumably BR, n.v.; Isotype P (photos
PTBG)). MEXU, PTBG)). Uragoga.
Psychotria tenuifolia Sw., Prodr. Veg. Ind. Psychotria tricostata Standi., Ceiba 1: 48. 1950.
Occ. 43. 1788. Type: SANTO DOMINGO Type: HONDURAS: Francisco Morazan,
(Hispaniola): A neotype needs to be drainage of the Rio Yeguare; canon de la Mt.
designated according to Howard (1989). Uyuca, 2000 m, 29 May 1948, A. Molino R.
Uragoga. 919 (Holotype F 1352078 (photos MEXU,
Psychotria thesceloantha Steyerm., Ann. PTBG); Isotypes EAP, n.v., GH, n.v.).
Missouri Bot. Gard. 75: 347. 1988. Type: —Psychotria galeottiana (M. Martens) C. M.
VENEZUELA: Amazonas, R. Liesner 16963 Taylor & Lorence
(Holotype MO; Isotype VEN, n.v.). Psychotria triphvlla (DC.) Mull. Arg. in Mart.,
Psychotria thornei Lorence, Novon 4: 127, fig. Fl. Bras. 6(5): 233. 1881. Palicourec
5. 1994. Type: MEXICO: Chiapas, Selva DC, Prodr. 4: 526. 1830. Type: FRENCH
Negra, ca. 15 km N of Pueblo Nuevo GUIANA: Guiana Gallica, without precise
Solistahuacan; canyon with montane rain locality, Paths s.n. (Holotype G-DC, n.v.
forest of Selva Negra on slopes, 5700 ft., 24 (photo MO)).—Palicourea triphvlla DC.
Jun 1970, R. Thome & E. Lathrop 40389b Psvchotria turrubarensis W. C. Burger & Q.
(Holotype CAS (photo PTBG)). Jimenez, Novon 4: 206. 1994 [cited as nom.
Psychotria thyrsoidea Sesse & Mo?., Fl. Mexic, nud. in W. C. Burger & C. M. Taylor,
ed. 2, 58. 1894. Type: MEXICO: Habitat Fieldiana: Bot., n.s. 33: 277. 1993]. Type:
prope Cordovam, Sesse & Mocino s.n. COSTA RICA: San Jose, faldas del Cerro
(presumably MA, n.v.).—Status uncertain. Bares, Zona Protectora Cerros de Turrubares,
Psychotria tomentosa (Oerst.) Hemsl., Biol. 9°47,30"N, 84°28'30"W, 1600 m, 6 Nov
Centr.-Amer., Bot. 2: 51. 1881. Mapouria 1990, Q. Jimenez & G Varela 935 (Holotype
tomentosa Oerst., Amer. Centr. 17, tab. 14, CR, n.v.; Isotypes F, n.v., MO).
fig- 8. 1863. Not Psychotria tomentosa Mull. Psychotria tutensis Dwyer, Ann. Missouri Bot.
Arg., 1881, nor Hemsl., 1881. Type: Gard. 67: 434. 1980. Type: PANAMA:
MEXICO: Veracruz, prope Papantla, F. Veraguas, Cerro Tute, ca. 10 km NW of Santa
Liebmann (presumably C, n.v.).—Status Fe, on ridgetop in cloud forest, above 1000 m,
uncertain, probably Psychotria micrantha 19'jun 1975, S. Mori 6783 (Holotype MO
Kunth in Humb., Bonpl. & Kunth. 2338505 (photos MEXU, PTBG)).
Psychotria tonduzii Standi., J. Wash. Acad. Sci. Coussarea austin-smithii Standi.
15: 287. 1925. Type: COSTA RICA : Psvchotria uliginosa Sw., Prodr. Veg. Ind. Occ.
Cartago, La Estrella, 26-27 Mar 1924, P. C. 43. 1788. Type: JAMAICA: A neotype needs
Standley 39461 (Holotype US 1153160 to be designated according to Howard (1989).
Uragoga.
Psychotria umbelliformis Dwyer & Hayden, Psychotria victoriae Dwyer, Ann. Missouri Bot.
Ann. Missouri Bot. Gard. 55: 44. 1968. Type: Gard. 53: 108. 1966. Not P. victoriae Standi.,
PANAMA: Panama, between peaks of Cerro 1930. Type: PANAMA: Chiriqui, woods at
Trinidad, saddle on SE slope, 5 May 1968, Finca Collins, Boquete, 5000 ft., M. V.
J. A. Duke & J. Kirkbride, Jr. 1641 (Holotype Hayden 164 (Holotype MO (not located
MO 1968366). 6/1996)).—Status uncertain, Psychotria
Psychotria uyucana Standi., Ceiba 1: 48. 1950. macrophylla Ruiz & Pav., vel sp. aff.
Type: HONDURAS: Francisco Morazan, Psychotria viridis Ruiz & Pav., Fl. Peruv. 2:
drainage of Rio Yeguare, Mt. Uyuca, 2000 m, 61, tab. 210 Jig. b. 1799. Type: PERU: Andes
29 May 1948, A. Molina R. 919 (Holotype F; de Peru, Chinchao et Macora, Jul 1794, Ruiz
Lopez & Pavon s.n. (Holotype MA, n.v.
(M. Martens) CM. Taylor & Lorence (photo MO)). Uragoga.
Psychotria Valeriana Standi., Publ. Field Mus. Psychotria wendlandiana Oerst. ex Standi., J.
Nat. Hist., Bot. Ser. 18: 1363. 1938. Type: Wash. Acad. Sci. 18: 9. 1928. Type: COSTA
COSTA RICA: Cerros de San Pedro de San RICA: San Miguel [de Naranjo?], 13 May
Ramon, 1075 m, Jun 1926, A. Brenes 4886 1857, Wendland 781 (Holotype C (photos
(Holotype F 853470 (photos MEXU, PTBG)). MEXU, MO, PTBG); Isotype US (photos
Psychotria vallensis Dwyer, Ann. Missouri Bot. MEXU, PTBG)).—Psychotria jimenezii
Gard. 67: 438. 1980. Type: PANAMA: Code, Standi.
woods adjacent to chicken farm, La Mesa, Psychotria wilburiana Dwyer, Ann. Missouri
above El Valle, 3 Jan 1974, J. D. Dwyer Bot. Gard. 67: 440. 1980. Type: PANAMA:
11867 (Lectotype MO 2163005A (photos Veraguas, valley of the Rio Dos Bocas, 11-13
MEXU, PTBG); Isolectotypes BM, n.v., F km beyond Agricultural School at Santa Fe,
(photos MEXU, PTBG), GH, n.v., PMA, 350-500 m, T. B. Croat 25745 (Holotype MO
n.v.). The lectotype was designated by 2232437 (photos MEXU, PTBG)).
Hamilton (1989), selecting one of three sheets Psychotria xerococcoides Dwyer, Ann.
annotated by J. D. Dwyer as "type," and the Missouri Bot. Gard. 67: 441. 1980. Type:
two isolectotypes at MO were subsequently PANAMA: Darien, Cerro Tacarcuna massif
distributed to BM and PMA.—Psychotria
gracilinora Benth. in Oerst. and Pico Tacarcuna, mostly elfin forest,
Psychotria veracruzensis Lorence & Dwyer, 1650-1800 m, 22 Jul 1976, A. Gentry, H.
Bol. Soc. Bot. Mexico 47: 61. 1987. Type: Leon & L. Forero 16924 (Holotype MO
MEXICO: Veracruz, Mun. de San Andres 2465400 (photo PTBG); Isotype MO (photo
Tuxtla, estacion de biologia tropical Los PTBG)).
Tuxtlas, entre Sontecomapan y Montepio; Psychotria yunckeri Standi., Field Mus. Nat.
vereda al Cerro Vigia, 150-250 m, 18 Apr Hist., Bot. Ser. 17: 397. 1938. Type:
1981, D. H. Lorence &T.P.I HONDURAS: Comayagua, in forest near the
3313 (Holotype MEXU; Isotypes E summit of the range above El Achote, above
(photo PTBG), MEXU, MO (photo PTBG) the plains of Siguatepeque 1850 m, Jun-Aug
UC, XAL). 1936, T. G. Yuncker, R. F. Dawson & H. R-
Psychotria veraguensis Dwyer, Ann. Missouri Youse 6013 (Holotype F 857686 (photos
Bot. Gard. 67: 438. 1980. Type: PANAMA: MEXU, PTBG); Isotypes GH, n.v., MO
Veraguas, along road between Escuela (photos MEXU, PTBG)).-
Agricola Alto Piedra (above Santa Fe) and
Rio Dos Bocas, ca. 10 km from the Escuela, Ps\ch. zevallosii C. M. Taylor, Monogr.
530-620 m, 26 Jul 1974, T. B. Croat 25871 Syst. Bot. Missouri Bot. Gard. 45: 1258.
(Holotype MO 2204045 (photos MEXU, 1993. Nom. nov. for Cephaehs wilhamsn
PTBG)). Standi., Publ. Field Columbian Mus., Bot. S
 \.N0MENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

8: 185. 1930. Kot Psychotria williamsii Randia armata (Sw.) DC, Prodr. 4: 387. 1830.
Standi., 1930. Type: PERU: Loreto, Manfina, subsp. armata. Gardenia armata Sw., Prodr.
on upper Rio Nanay, 29 Jun 1929, L. Williams Fl. Ind. Occ. 51. 1788. Type: COLOMBIA:
1114 (HolotypeF 604625). Cartagena, Jacquin s.n. (Lectotype: Jacquin's
plate; designated by Taylor & Lorence, 1993).
Rachicallis DC, Prodr. 4: 433. 1830. Type: Randia armata (Sw.) DC. subsp. panamensis
Rachicallis mpestris (Sw.) DC. (Standi.) Lorence, Novon 8: 250. 1998.
Rachicallis americana (Jacq.) A. S. Hitchc, Randia panamensis Standi., Publ. Field
Missouri Bot. Gard. Ann. Rep. 4: 92. 1893. Columbian Mus., Bot. Ser. 4: 288. 1929.
Hedyotis americana Jacq., Enum. PI. Carib. Type: PANAMA: Prov. Bocas del Toro,
12. 1760. Type: CUBA: India Occidentalis, region of Almirante, Flat Rock, Jan-Mar
Swartzs.n. (locatic 1928, G. Proctor Cooper 213 (Holotype F
579675 (photos MEXU, PTBG)).
Randia L., Sp. PL 2: 1192. 1753. Type: Randia Randia blepharodes Standi., Contr. U.S. Natl.
mitis L. (=Randia aculeata L.). Herb. 23: 1378. 1926. Type: MEXICO:
Randia aculeata L., Sp. PL 2: 1192. 1753. var. Guerrero, Mexcala, versant oriental, terre
aculeata. Type: JAMAICA: (LINN 214.1, chaude, 500 m, 23 May 1899, E. Langlasse
n.v.; for typification, see Howard, 1989: 452). 1033 (Holotype US 386343 (photos MEXU,
Randia aculeata L. var. chiapasina Loes., PTBG); Isotypes B, presumably destroyed
Verhandl. Bot. Ver. Brandenburg 65: 109. (fragment F), F, GH, n.v., K, P (2 sheets;
1923. Type: MEXICO: Chiapas, Comitan, photos MEXU, PTBG)).^Randia thurberi
near Bahucuc, E. Seler & C. Seler 2203 S. Watson
(Holotype B, presumably destroyed).—Status Randia blepharophylla Standi., Contr. U.S.
, probably Randia aculeata L. var. Natl. Herb. 23: 1375. 1926. Type: MEXICO:
Nayarit, foothills of Sierra Madre, Tepic,
Randia aculeata L. var. dasyclada Steyerm., between Aguacates and Dolores, J. N. Rose
Mem. New York Bot. Gard. 23: 341. 1972. 2028 (Holotype US 300921 (photos MEXU,
Based on Randia obovata Kunth in Humb., PTBG); Isotypes F (photos MEXU, PTBG),
Bonpl. & Kunth, Nov. Gen. Sp. PL 3: 409. GH, n.v.).—Glossostipula blepharophylla
1820. Not Randia obovata Ruiz & Pav., 1799. (Standi.) Lorence
Type: COLOMBIA: Dept. Cordoba, ad ostia Randia brenesii Standi., Publ. Field Mus. Nat.
flumiinis Sinu, A. Bonpland s.n. (Holotype P- Hist., Bot. Ser. 18: 1365. 1938. Type: COSTA
Bonpland, n.v. (photo F)). RICA: La Palma de San Ramon, 10 Mar
Randia albonervia Brandegee, Zoe 5: 257. 1932, Brenes 15048 (Holotype F 857063
1908. Type: MEXICO: Veracruz, vicinity of (photos MEXU, PTBG)).
Zacuapan, Corral de las Piedras, May 1907, Randia calycosa Standi., Contr. U.S. Natl.
C. A. Purpus 2424 (Holotype UC, n.v.; Herb. 20: 201. 1919. Type: PANAMA:
Isotypes BM, F (photos MEXU, PTBG), GH, Chiriqui, Los Siguas Camp, southern slope of
n.v., US (photos MEXU, PTBG)).—Randia Cerro de la Horqueta, 1700 m, 17-19 Mar
monantha Benth. 1911,//. Pittier 3198 (Holotype US 677593
Randia altiscandens (Ducke) C. M. Taylor in (photos MEXU, PTBG); Isotype F (photos
W. C. Burger & C. M. Taylor, Fieldiana, Bot., MEXU, PTBG)).
n.s. 33: 282. 1993. Basanacantha altiscandens Randia canescens Greenm., Proc. Amer. Acad.
Ducke, Archiv. Inst. Biol. Veg. Rio Janeiro 2: Arts 34: 573. 1899. Type: MEXICO: Morelos,
172. 1935. Type: VENEZUELA: Amazonas, barrancas near Cuemavaca, 5000 ft., 28 May
Ducke s.n. (Holotype RB 24415, n.v.; Isotype 1898 C G. Pringle 6863 (Holotype GH, n.v.;
Isotypes F (photos MEXU, PTBG), MEXU (2
sheets), MO (photos MEXU, PTBG), P (2
 sheets; photos MEXU, PTBG), US (2 sheets; • i.) K. Schum. in Mart., Fl.
photos MEXU, PTBG)). Bras. 6(6): 342. 1889. Mussaenda formosa
Randia capitata DC, Prodr. 4: 387. 1830. Jacq., Enum. PI Carib. 16. 1760. Type:
Type: MEXICO: Mexico ad Anasteca COLOMBIA: Cartagena, Jacquin s.n.
[Huasteca], May 1827, Berlandier 513 (Holotype G, n.v).—Rosenbergiodendron
(Holotype G-DC, n.v. (photos F, MEXU)). formosum (Jacq.) F. Fagerl
Randia chiapensis Standi, Contr. U.S. Natl. Randia gaumeri Greenm. & C H. Thomps.,
Herb. 23: 1377. 1926. Type: MEXICO: Ann. Missouri Bot. Gard. 1: 410. 1915. Type:
Chiapas, Canjob, 27 Mar 1904, E. A. MEXICO: Yucatan, Izamal, 1895, G. F.
Goldman 797 (Holotype US 470602 (photos Gaumer 589 (Holotype MO 951695 (photos
K, MEXU, PTBG)). MEXU, PTBG); Isotypes F (photos MEXU,
Randia cinerea (Femald) Standi., Contr. U.S. PTBG), MO (photos MEXU, PTBG), US
Natl. Herb. 20: 201. 1919. Genipa cinerea (photos MEXU, PTBG)).—Randia
Femald, Proc. Amer. Acad. Arts. 33: 93. obcordata S. Watson
1897. Type: MEXICO: Guerrero, Acapulco Randia genipifolia (Standi. & Steyerm.)
and vicinity, Oct 1894-Mar 1895, E. Palmer Lorence, Novon 8: 249. 1998. Duroia
348 (Holotype GH, n.v.; Isotype US (photos genipifolia Standi. & Steyerm., Publ. Field
MEXU, PTBG)). Mus. Nat. Hist., Bot. Ser. 22: 186. 1940.
Randia concinna Standi., Publ. Field Columbian Type: GUATEMALA: Izabal, Rio Dulce,
Mus., Bot. Ser. 4: 288. 1929. Type: MEXICO: between Livingston and 6 miles up river, on
Jalisco, San Sebastian, Hacienda del Ocotal, north side (right side going up river), sea
1500 m, Y. Mexia 1690 (Holotype F 579911 level, 14 Apr 1940, J. A. Steyermark 39382
(photos MEXU, PTBG); Isotypes A, n.v., (Holotype F 1035218 (photo PTBG)).
BM, GH, n.v., MO (photos MEXU, PTBG), Randia genipoides Dwyer in W. C Burger &
US (photos MEXU, PTBG)) — Glossostipula C M. Taylor, Fieldiana, Bot., n.s. 33: 284.
concinna (Standi.) Lorence 1993. Type: COSTA RICA: Alajuela, Upala,
Randia cookii Standi., Contr. U.S. Natl. Herb. G. Herrera 1129 (Holotype CR, n.v.; Isotypes
32: 176. 1934. Type: GUATEMALA: Baja F, n.v., MO).
Verapaz, Santa Rosa, June 1904, O. F. Cook Randia gentlei Lundell, Wrightia 4: 125. 1969.
249 (Holotype US 860099 (photos MEXU, Type: BRITISH HONDURAS (BELIZE):
PTBG), fragment F). Toledo Distr., Edwards Rd. beyond Columbia,
Randia crescentioides Standi., Publ. Field Mus. 24 May 1951, P. H. Gentle 7343 (Holotype
Nat. Hist., Bot. Ser. 23: 88. 1944. Type: TEX-LL (photos MEXU, PTBG); Isotypes F,
COSTA RICA: Port Parker, 4 Jul 1932, J. T. TEX-LL (2 sheets), US (photos MEXU,
Howell s.n. (Holotype CAS, n.v., fragment F PTBG)).—Randia genipifolia (Standi. &
(photos MEXU, PTBG)).—Randia thurberi Steyerm.) Lorence
S. Watson Randia gentryi Dwyer, Ann. Missouri Bot.
Randia echinocarpa Sesse & Moc. ex DC, Gard. 67: 447. 1980. Type: PANAMA: Prov.
Prodr. 4: 385. 1830. Type: MEXICO: Fl. Darien, top of Cerro Mali, 1300-1420 m,
Mexic. Ic. 469 (caique presumbably MA, n.v., A. H. Gentry & S. Mori 13629 (Holotype MO
G-DC, n.v. (copies MEXU, US)). 2258315 (photos MEXU, PTBG); Isotype MO
Basanacantha, Genipa. 2578524 (photo PTBG)).
Randia ehrenbergii Standi, Contr. U.S. Natl. Randia grandifolia (Donn. Sm.) Standi, J.
Herb. 32: 167. 1934. Type: MEXICO: Wash. Acad. Sci. 18: 166. 1928.
Veracruz, Plan de Amilpas, Mar 1831, C Basanacantha grandifolia Donn. Sm., Bot.
Ehrenberg 575 (Holotype B, presumably Gaz. (Crawfordsville) 55: 436. 1913. Type:
destroyed).—Randia capitata DC. COSTA RICA: Forets de Santo Domingo de
Golfo Dulce, Feb 1896, A. Tonduz 9878
 OF MEXICAN AND CENTRAL

(Lectotype US 941393, here designated Magdalena, 260 m, H. Karsten s.n. (Holotype

(photos MEXU, PTBG); Isolectotypes F B, presumably destroyed).
(photos MEXU, PTBG), US 577282). Randia induta Standi., Contr. U.S. Natl. Herb.
Syntype: COSTA RICA: Bords du Rio 23: 1377. 1926. Type: MEXICO: Guerrero,
Corozal, pres Santo Domingo de Golfo Dulce, Cayacol, 12 Apr 1903, E. W. Nelson 7018
Apr 1896, A. Tonduz 9982 (US, F (photos (Holotype US 399362 (photos MEXU,
MEXU, PTBG)). Tonduz 9878, with more PTBG)).
abundant flowers and two duplicates, Randia inermis Sesse & Moc., Fl. Mexic. 54.
represents the better choice for lectotype. 1891. Not Randia inermis (Ducke) Ducke,
Randia grayumii Dwyer & Lorence in W. C. 1932. Type: MEXICO: Habitat in Amatitani
Burger & C. M. Taylor, Fieldiana, Bot., n.s. clivo, ubi Testiculus Taurinus ob Fructus.
33: 285. 1993. Type: COSTA RICA: Limon, Figuram vulgo adpellatur. Florit Maio, Sesse
behind NAISA house, Gandoca, 9°36'N, & Mocino s.n. (presumably MA, n.v.).
82°37'W, 3 m, 4 May 1985, M. H. Grayum & —Status uncertain.
G. Schatz 5287 (Holotype MO 3648263 Randia karstenii Pol., Linnaea 41: 568. 1877.
(photo PTBG); Isotype ?CR, n.v.). Syntypes: COSTA RICA: In sepibus vivis
Randia guatemalensis Standi., Contr. U.S. Natl. prope San Jose, Jun & Sep 1875, H. Karsten
Herb. 20: 202. 1919. Type: GUATEMALA: 74 (B, presumably destroyed); Karsten 302
Alta Verapaz, Secanquim, 550 m, 6 May (B, presumably destroyed, BM, n.v.).
1905, H. Pittier271 (Holotype US 472930 —Randia aculeata L. var. aculeata
(photos MEXU, PTBG)).—Randia aculeata Randia laetevirens Standi.. Contr. U.S. Natl.
L. var. aculeata Herb. 23: 1377. 1926. Type: MEXICO:
Randia guerrerensis Lorence & Rodr. Acosta, Nuevo Leon, Monterrey, Cerro la Silla, 20
Biotica 11: 195,.% /. 1986. Type: MEXICO: Mar 1902, E. W. Nelson 6083 (Holotype US
Guerrero, Mun. de Coyuca de Benitez, 347312 (photos MEXU, PTBG); Isotype F
camino de El Zapote a Hacienda de Cabanas, (photos MEXU, PTBG)).
a 10 km al SO de la alberca El Dorado, cerca Randia laevigata Standi., Contr. U.S. Natl.
de la Laguna de Mitla, 7 m, 21 Jul 1984, M. Herb. 20: 201. 1919. Type: MEXICO: Sonora,
Rodriguez Acosta sub Lorence 4584 Sierra de Alamos, 18 Mar 1910, J. N. Rose,
(Holotype MEXU; Isotypes ENCB, MO P. C. Standley & P. G. Russell 13051
(photo PTBG)). (Holotype US 635870 (photos MEXU,
Randia habrophlebia Standi. & Steyerm., Publ. PTBG)).
Field Mus. Nat. Hist., Bot. Ser. 23: 25. 1943. Randia lasiantha (Standi.) Standi., Contr. U.S.
Type: GUATEMALA: Jutiapa, Valley of Natl. Herb. 20: 201. 1919. Basanacantha
Jutiapa, 850 m, 24 Oct-6 Nov 1940, P. C. lasiantha Standi., Contr. U.S. Natl. Herb. 18:
Standley 76321 (Holotype F 1091237 (photos 134. 1916. Type: PANAMA: Panama,
MEXU, PTBG)).—Randia cinerea (Femald) Alajuela, Chagres Valley, 30-100 m, H.
Standi. Pittier 3473 (Holotype US 678517 (photos
Randia hidalgensis Lorence, Bol. Soc. Bot. MEXU, PTBG)).
Mexico 47: 38,%. la, b. 1987. Type: Randia latifolia Lam. var. micrantha Schltdl.,
MEXICO: Queretaro, Mun. de Arroyo Seco, Linnaea 6: 723. 1831. Type: MEXICO:
12 km NW de El Trapiche, camino a Rio Veracruz, prope Jalapam, Schiede & Deppe
Verde, a 1 km SE de Puerto de Ayutla, 1200 276 (Holotype presumably HAL, n.v.).
m, 29 Abr 1982, P. Tenorio & R. Hernandez —Randia aculeata L. var. aculeata
M. 297 (Holotype MEXU; Isotype MO (photo Randia letreroana Lundell, Wrightia 4: 126.
PTBG)). 1969. Type: MEXICO: Chiapas, Letrero, near
Randia hondensis Karsten, Linnaea 30: 154. Siltepec, 2000 m, 6 Jul 1941, E. Matuda 4358
1859. Type: COLOMBIA: Valley of Rio (Holotype TEX-LL, n.v.; Isotypes F (photo
 PTBG), MEXU, MO (photo PTBG)). Comondu, 2 Feb 1889, T. S. Brandegee s.n.
—Randia thurberi S. Watson (Holotype UC, n.v.).—Randia capitata DC.
Randia longiloba Hemsl., Biol. Centr.-Amer., Randia megalocarpa Dwyer, Ann. Missouri
Bot. 4: 101. 1887. Type: MEXICO: Yucatan, Bot. Gard. 67: 449. 1980. Type: PANAMA:
Cozumel Island, Gaumer s.n. (Holotype Panama, Cerro Penon, 3 km S of Alcade Diaz,
presumably K, n.v.). vicinity of television tower on Continental
Randia loniceroides Dwyer & Lorence, Bol. Divide, 350-440 m, M. Nee 8865 (Holotype
Soc. Bot. Mexico 47: 40, fig. lc, d. 1987 [as MO 2208620 (photo PTBG); Isotype MO
lonicerioides]. Type: MEXICO: Veracruz, 2208621 (photo PTBG)).—Randia Iasiantha
Mun. de Minatitlan, Zona de Uxpanapa, 13.7 (Standi.) Standi.
km E. de La Laguna sobre camino a Randia microcarpa Sesse & Mo?., PI. Nov.
Uxpanapa, luego 8 km N sobre camino a Hispan. 33. 1888 (La Naturaleza ser. 2, 1:
Belisario Dominguez, 17°21'N, 94°23'W, 130 app. 33. 1888) [as microcarpus]. Type:
m, 19 Apr 1981, T. Wendt & E. Lott 3207 MEXICO: Habitat in montibus Haereditatis-
(Holotype MEXU; Isotypes CHAPA, ENCB). Ahuesuyo, Sesse & Mocino s.n. (presumably
nidi, in Lundell, Contr. MA, n.v.).—Status uncertain.
Univ. Michigan Herb. 4: 30. 1940. Type: Randia millspaughiana S. F. Blake, Proc. Biol.
BRITISH HONDURAS (BELIZE): El Cayo Soc. Wash. 34: 45. 1921. Type: MEXICO:
Distrist, Vaca, 26 Apr 1938, P. H. Gentle Yucatan, Maxacanu, Mar 1916, G. F. Gaumer
2524 (Holotype F 941519 (photo PTBG); etal. 23260 (Holotype F 446691 (photos
Isotype TEX-LL).—Randia monantha MEXU, PTBG); Isotypes GH, n.v., US
Benth. (photos MEXU, PTBG)).—Randia longiloba
Randia malacocarpa Standi., Contr. U.S. Natl. Hemsl.
Herb. 20: 202. 1919. Type: MEXICO: Randia mira Dwyer, Ann. Missouri Bot. Gard.
Nayarit, near Acaponeta, Tepic, 30 Jul 1897, 67: 450. 1980. Type: PANAMA: Panama,
J. N. Rose 3298 (Holotype US 302274 (photos 5-10 km NE of Altos de Pacora at end of
MEXU, PTBG, US)). road, 6 Mar 1975, S. Mori & J. Kallunki 4953
Randia matudae Lorence & Dwyer, Bol. Soc. (Holotype MO 2247822 (photos MEXU,
Bot. Mexico 47: 42. 1987. Based on Casasia PTBG); Isotype MO (photos MEXU, PTBG)).
chiapensis Miranda, Ceiba 4: 142, fig. p. 144. Randia mitis L., Sp. PL 2: 1192. 1753. Type:
1954. Not Randia chiapensis Standi., 1926. Habitat in Jamaica. Lectotype: Lycium majus
Type: MEXICO: Chiapas, Rancho el Ocote, americanum
30 km NW de Ocozocuautla, 650 m, 21 May lucitus, Plukenet, Phytographica, t. 97, fig- 6-
1950, F. Miranda 7734 (Lectotype MEXU; 1691. (designated by Lorence in Jarvis et al.,
designated by Lorence & Dwyer, 1987). 1993: 80).—Randia aculeata L. var.
Syntype: MEXICO: Chiapas, same locality, F.
Miranda 6224 (F (photo PTBG)). Randia mollifolia Standi., Field Mus. Nat.
Randia mayana Lundell, Wrightia 5: 322. Hist., Bot. Ser. 17: 219. 1937. Type:
1976. Type: GUATEMALA: Baja Verapaz, MEXICO: Sonora, Saguaribo, Rio Mayo, 3
Nino Perdido, in high forest on top of hill, E. Nov 1935, H. S. Gentry 2139 (Holotype F
of km 150/151, 30 Aug 1975, C. L. Lundell & 809526 (photos MEXU, PTBG); Isotypes GH,
E. Contreras 19758 (Holotype TEX-LL n.v., MEXU, US (photos MEXU, PTBG),
(photos MEXU, PTBG); Isotypes F, MO WIS).
(photo PTBG), TEX-LL (photos MEXU, Randia monantha Benth., PI. Hartweg. 84.
PTBG)). 1841. Type: GUATEMALA: Rio de los
Randia megacarpa Brandegee, Zoe 5: 257. Esclavos, T. Hartweg 582 (Holotype K
1908. Type: MEXICO: Baja California, (photos F, MEXU); Isotypes B, presumably
A NOMENCLATOR OF MEXICAN AND CENTRAL /

destroyed (photo MO), BM, P [label states armata (Sw.) DC. subsp. panamensis
"Mexique"] photos MEXU, PTBG)). (Standi.) Lorence
Randia montana L. O. Williams, Phytologia 24: Randia pepoformis Dwyer, Ann. Missouri Bot.
160. 1972. Type: HONDURAS, Intibuca, Gard. 67: 451. 1980. Type: PANAMA:
Cordillera Opalaca, between Calaveras and El Darien, trail from Tacarcuna Village on Rio
Duraznillo, 1800 m, 12 Mar 1970, A. Molina Tacarcuna to Cerro Mali, 800-1300 m, 16 Jar
R. & A. R. Molina 25550 (Holotype F 1975, A. H. Gentry & S. Mori 13614
1695297 (photos MEXU, PTBG); Isotype (Holotype MO 2248019 (photos MEXU,
EAP, n.v.).—Randia aculeata L. var. PTBG); Isotypes F (photos MEXU, PTBG),
aculeata MO (photos MEXU, PTBG)).—Randia
Randia nelsonii Greenm., Proc. Amer. Acad. altiscandens (Ducke) C. M. Taylor
Arts 34: 574. 1899. Type: MEXICO: Oaxaca, Randia petenensis Lundell, Wrightia 4: 127.
Juchitan to Chivela, 150-900 ft., 1895, E. W. 1969. Type: GUATEMALA: Peten, La
Nelson 2635 (Holotype GH, n.v.; Isotype US Cumbre, about 5 km from the village, on
569319 (photos MEXU, PTBG)).—Randia Pusila River Road, 17 May 1967, E.
thurberi S. Watson Contreras 6941 (Holotype TEX-LL (photos
Randia nicaraguensis Lorence & Dwyer, MEXU, PTBG); Isotypes F, US (photos
Novon 8: 247, fig. 1. 1998. Type: MEXU, PTBG)).
NICARAGUA: Esteli, 4.9-7.6 km NE of Randia pittieri (Standi.) Standi., Contr. U.S.
Hwy. 1 at Esteli along road to Yali, ca. Natl. Herb. 20: 201. 1919. Basanacantha
13°08-09', 86°19-20'W, 1100 m, 14 Nov pittieri Standi., Contr. U.S. Natl. Herb. 18:
1979, W. D. Stevens & A. Grijalva 15553 134. 1916. Type: COSTA RICA: Gent
(Holotype MO 2872306; Isotypes F, HNMN, Junction, pres Matina, Feb 1900, H. Pither
MEXU, MO). 16036 (Holotype US 578473 (photos K,
Randia oaxacana Standi., Contr. U.S. Natl. MEXU, PTBG); Isotype P (fragment F,
Herb. 32(3): 165. 1934. Type: MEXICO: photos MEXU, PTBG)).
Oaxaca, Choapam [=Santiago Choapan], Randia pleiomeris Standi., Contr. U.S. Natl.
2800-1500 ft., 28-29 Jul 1894, E W. Nelson Herb. 20: 202. 1919. Type: GUATEMALA:
842 (Holotype F 599712 (photos MEXU, Santa Rosa, Santa Rosa, 3000 ft., May 1892,
PTBG)). Heyde & Lux 3166 (Lectotype US 888490,
Randia obcordata S. Watson, Proc. Amer. here designated [left- and right-hand
Acad. Arts 24: 53. 1889. Type: MEXICO: collections only; the center collection is R.
Sonora, Guaymas, 1887, E. Palmer 648 cookii]; Isolectotype GH, n.v.).—Randia
(Holotype US 47988 (photos MEXU, PTBG); thurberi S. Watson
Isotype BM). Randia pringlei (S. Watson) A. Gray, Proc.
Randia obovata Kunth in Humb., Bonpl. & Amer. Acad. Arts 21: 379. 1886.
Kunth, Nov. Gen. Sp. 3: 409. 1820. Not Basanacantha pringlei S. Watson, Proc.
Randia obovata Ruiz & Pav., 1799. Type: Amer. Acad. Arts 18: 98. 1883. Type:
COLOMBIA: Ad ostia fluminis Sinu, MEXICO: Coahuila, Mountains, Jimulco, 9
A. Bonplands.n. (Holotype P-Bonpland, Apr 1885, C. G. Pringle 142 (Holotype GH,
n.v.).—Randia aculeata L. var. dasyclada n.v.; Isotypes A, n.v., BM, F (photos MEXU,
Steyerm. PTBG), MEXU, P (photos MEXU, PTBG),
Randia panamensis Standi., Publ. Field US (photos MEXU, PTBG)).
Columbian Mus., Bot. Ser. 4: 288. 1929. Randia pterocarpa Lorence & Dwyer, Bol.
Type: PANAMA: Prov. Bocas del Toro, Soc. Bot. Mexico 47: 43, fig. 2. 1987. Type:
region of Almirante, Flat Rock, Jan-Mar MEXICO: Veracruz, Mun. San Andres
1928, Q. Proctor Cooper 213 (Holotype F Tuxtla, Estacion de Biologia Tropical Los
579675 (photos MEXU, PTBG)).—Randia Tuxtlas, jardin botanico, 95°04-09'W,
 18°34-36*N, 160 m, 6 Dec 1983, G. Ibarra subcordata Standi, Contr. U.S. Natl. Herb.
M. 1109 (Holotype MEXU; Isotypes BM, F, 18: 133. 1916. Type: COSTA RICA: Buissons
MO, XAL). et bois des chemins a Nicoya, May 1900, A.
Randia purpusii Greenm. & C. H. Thomps., Tonduz 13934 (Holotype US 861251 (photos
Ann. Missouri Bot. Gard. 1: 410. 1914. Type: MEXU, PTBG); Isotypes B, destroyed
MEXICO: San Luis Potosi, Minas de San (fragment F, photos MEXU, PTBG), GH, n.v.,
Rafael, May \9\\,CA. Purpus 5208 LE, n.v. (fragment F, photo PTBG), P (photos
(Holotype MO 4186182 (photos MEXU, MEXU, PTBG), fragment F (photos MEXU,
PTBG); Isotypes F, (photos MEXU, PTBG), PTBG)).—Randia monantha Benth.
GH, n.v., MO (photos MEXU, PTBG), US Randia tetracantha (Cav.) DC, Prodr. 4: 387.
(photos MEXU, PTBG)).—Randia capitata 1830. Mussaenda tetracantha Cav., Icon. PI.
DC. 4: 20,/?/. 435. 1799. Type: MEXICO:
Randia retroflexa Lorence & M. Nee, Brittonia [Probably Guerrero], cerca Acapulco, Nee s.n.
39: 371, fig. 1. 1987. Type: MEXICO: (Holotype MA, n.v. (photos F, MEXU, MO)).
Veracruz, Mun. de Hidalgotitlan,
Campamento Hermanos Cedillo a 4 km de Randia tetracantha Sesse & Moc., PI. Nov.
Pancho Villa, 150 m, 25 Mar 1975, B. Ortiz & Hispan. 33. 1888 (La Naruraleza ser. 2, 1:
J. Martiniano A. 56 (Holotype MEXU; app. 33. 1888). Not Randia tetracantha (Cav.)
Isotypes BM, BR, ENCB, F (photo PTBG), DC, 1830. Type: MEXICO: Habitat in
MO (photo PTBG), NY, UC, XAL). montibus Haereditatis-Ahuesuyo, fl. Dec,
Randia rhagocarpa Standi., Contr. U.S. Natl. Sesse & Mocino s.n. (presumably MA,
Herb. 23: 1376. 1926. Type: MEXICO:
Tamaulipas, vicinity of Victoria, 320 m, 1
Feb-9 Apr 1907, E. Palmer 38 (Holotype US
572266 (photos MEXU, PTBG); Isotypes F Yucatan, Xkombec and Uxmal, E. Seler & C
(photos MEXU, PTBG), GH, n.v., MO Seler 4035 (Syntype B, presumably destroyed,
(photos MEXU, PTBG)).—Randia GH, n.v.); E. Seler & C. Seler 5559a (Syntype
obcordata S. Watson B, presumably destroyed).—Randia truncata
Randia rosei Standi., Contr. U.S. Natl. Herb. 20: Greenm. & C H. Thomps.
203. 1919. Type: MEXICO: Sinaloa, Rosario, Randia texensis Lundell, Field & Lab. 13: 22.
7 Jul 1897, J. N. Rose 1551 (Holotype US 1945. Type: U.S.A.: Texas, Cameron County,
300395 (photos MEXU, PTBG); Isotype F about 5 miles north of Brownsville off Point
(photos MEXU, PTBG), GH, n.v.).—Randia Isabel road, 21 Sep 1943, C L. Lundell &
thurberi S. Watson A. A. Lundell 12456 (Holotype SMU,
Randia sonorensis Wiggins, Contr. Dudley n.v.).—Randia obcordata S. Watson
Herb. 3: 75,pi. 19, figs. 4-6. 1940. Type: Randia thurberi S. Watson, Proc. Amer. Acad.
MEXICO: Sonora, granitic hillside 12 miles E Arts 24: 53. 1889. Type: MEXICO: Sonora,
of Ures on road to Babiacora, 2760 ft., 20 Sep Guayamas, 1887, Thurber in E. Palmer 229
1934, /. L. Wiggins 7360 (Holotype DS (Holotype US 1393134 (photos MEXU,
263358, n.v.; Isotype US (photos MEXU, PTBG); Isotypes BM, US).
PTBG)). Randia tomatillo Loes., Rep. Spec. Nov. 18:
Randia standleyana L. O. Williams, Phytologia 360. 1922. Type: MEXICO: Veracruz, Puerto
24: 162. 1972. Type: GUATEMALA: Peten, de Alvarado, and Isla de Sacrificios, E. Seler
La Libertad, 29 May 1933, C. L. Lundell 3474 & C. Seler 4498 (Lectotype US 1205748, here
(Holotype F 692287 (photos MEXU, PTBG); designated; Isolectotypes B, destroyed; GH,
Isotype MICH, n.v.). n.v.). Same locality, E. Seler & C. Seler 5433
Randia subcordata (Standi.) Standi., Contr. U.S. (Syntype B, destroyed). The US collection of
Natl. Herb. 20: 201. 1919. Basanacantha Seler & Seler 4498 is a good fruiting
 specimen and the best choice for 1911,//. Pittier 4223 (Holotype MO 1227470
lectotype.—Randia aculeata L. var. (photo PTBG)).—Raritebe palicoureoides
dasyclada Steyerm. Wernham subsp. dwyerianum J. H. Kirkbr.
Randia tomentosa S. Watson, Proc. Amer. Raritebe dariensis (Dwyer) Dwyer, Ann.
Acad. Arts 25: 152. 1890. Not Randia Missouri Bot. Gard. 67: 453. 1980. Dukea
tomentosa Wight & Arn., 1834, nor R. dariensis Dwyer, Ann. Missouri Bot. Gard.
tomentosa Hook., 1880. Type: MEXICO: 53: 365. 1966. Type: PANAMA: Darien,
Nuevo Leon, Sierra de la Silla, near Chepigana District, Cana-Cuasi Trail (Camp
Monterrey, Aug 1889, C. G Pringle 2865 2), 10 Mar 1940, M. E. Terry & R. A. Terry
(Holotype GH, n. v.).—Randia capitata DC. 14871 (Holotype MO 1196543 (photo
Randia truncata Greenm. & C. H. Thomps., PTBG)).—Raritebe palicoureoides
Ann. Missouri Bot. Gard. 1: 411. 1914. Type: Wernham subsp. dwyerianum J. H. Kirkbr.
MEXICO: Yucatan, vicinity of Izamal, 1895, Raritebe euryphyllum (Standi.) Dwyer, Ann.
G F. Gaumer 713 (Holotype MO 46009 Missouri Bot. Gard. 67: 453. 1980. Coussarea
(photos MEXU, PTBG); Isotypes F (photos euryphylla Standi., Publ. Field Columbian
MEXU, PTBG), MO (photos MEXU, PTBG), Mus., Bot. Ser. 4: 294. 1929. Type:
US (photos MEXU, PTBG)). PANAMA: Bocas del Toro, Buena Vista
Randia vazquezii Lorence & Dwyer, Bol. Soc. Camp on Chiriqui Trail, 375 m, 1928, G P.
Bot. Mexico 47: 46,/?g. 3. 1987. Type: Cooper 228 (Holotype F 579282, n.v.).
MEXICO: Veracruz, Mun. de Jesus Carranza, —Raritebe palicoureoides Wernham subsp.
Zona de Uxpanapa, 2 km N del Poblado 2, dwyerianum J. H. Kirkbr.
Ejido F. J. Mina, 17° 16'N, 94°40'W, 120 m, Raritebe palicoureoides Wernham subsp.
23 May 1982, M. Vazquez T. & I. Navarrete dwyerianum J. H. Kirkbr., Brittonia 31: 304.
2520 (Holotype MEXU; Isotypes CHAPA, 1979. Type: PANAMA: Panama, on trail to
PTBG). Cerro Campana, 23 Aug 1967, J. H.
Randia watsonii B. L. Rob., Proc. Amer. Acad. Kirkbride, Jr. & S. M. V. Hay den 273
Arts 29: 317. 1894 [as watsoni}. Based on (Holotype MO 1917794 (photos MO, PTBG);
Randia tomentosa S. Watson, Proc. Amer. Isotypes F, MO (photo PTBG)).
Acad. Arts 25: 152. 1890. Not Randia Raritebe panamensis (Dwyer) Dwyer, Ann.
tomentosa Wight & Arn., 1834, nor R. Missouri Bot. Gard. 67: 454. 1980. Dukea
tomentosa Hook., 1880. Type: MEXICO: panamensis Dwyer, Ann. Missouri Bot. Gard.
Nuevo Leon, Sierra de la Silla, near 53: 364. 1966. Type: PANAMA: Panama,
Monterrey, Aug 1889, C. G. Pringle 2865 Cerro Jefe in Clusia rain forest, 3000 ft., E. L.
(Holotype GH, n.v.).—Randia capitata DC. Tyson, J. Dwyer & K. Blum 3326 (Holotype
Randia xalapensis M. Martens & Galeotti, MO 2084467 (photo PTBG)).—Raritebe
Bull. Acad. Roy. Sci. Bruxelles 11: 239. 1844. palicoureoides Wernham subsp.
Type: MEXICO: Veracruz, Xalapa, dwyerianum J. H. Kirkbr.
Cordillera, Jun-Oct 1840, H. Galeotti 7116 Raritebe trifoliatum (Dwyer & Hay den) Dwyer,
(Holotype BR, n.v.; Isotypes GH (label states Ann. Missouri Bot. Gard. 67: 455. 1980.
"Galeotti s.n"), P (photos MEXU, PTBG)). Dukea trifoliata Dwyer & Hayden, Ann.
Missouri Bot. Gard. 54: 139. 1967. Type:
Raritebe Wernham, J. Bot. 55: 336. 1917. Type: PANAMA: Code, El Valle, potato farm
Raritebe palicoureoides Wernham above village of El Valle, 12 Jul 1962, J. D.
Raritebe blumii (Dwyer) Dwyer, Ann. Missouri Dwyer 1933 (Holotype MO 3831231 (photo
Bot. Gard. 67: 452. 1980. Dukea blumii PTBG)).—Raritebe palicoureoides
Dwyer, Ann. Missouri Bot. Gard. 53: 365. Wernham subsp. dwyerianum J. H. Kirkbr.
1966. Type: PANAMA: Colon, around Dos Raritebe victoriae (Dwyer) Dwyer, Ann.
Bocas, Rio Fato valley, 40-80 m, 16 Aug Missouri Bot. Gard. 67: 456. 1980. Dukea
 victoriae Dwyer, Ann. Missouri Bot. Gard. Relbunium Hook. f. in Benth. & Hook, f., Gen.
53: 364. 1966. Type: PANAMA: Darien, PI. 2: 149. 1873. Type: Not designated.
cloud forest, Cerro Pirre, Oct 1962, J. A. Duke [Herein treated as a synonym of Galium L.]
6099 (Holotype MO 1832075 (photo PTBG); • aschenbornii (S. Schauer) Hemsl.,
Isotype MO (photo PTBG)).—Raritebe Biol. Centr.-Amer., Bot. 2: 62. 1881. Galium
palicoureoides Wernham subsp. aschenbornii S. Schauer, Linnaea 20: 701.
dwyerianum J. H. Kirkbr. 1847. Type: MEXICO: Zimapan in Mexico,
Raritebe viburnioides Standi., Publ. Field Aschenborn 92 (Holotype presumably BR or
Columbian Mus., Bot. Ser. 7: 56. 1930. Type: K, n.v.; probable fragment JE, n.v.).—Galium
COLOMBIA: Cauca, La Gallera, Micay
Valley, 1 Jul 1922, Killip 7934 (Holotype US, urn (L.) Hemsl., Biol.
n.v.; fragment F, n.v.; Isotypes G, n.v., NY, Centr.-Amer., Bot. 2: 63. 1881. Valantia
n.v.).—Bertiera viburnoides (Standi.) J. H. hypocarpia L., Syst. Nat., ed. 10, 1307. 1759.
Kirkbr. Type: JAMAICA: Rubia I, Browne, Hist.
Jamaica 141 (Holotype LINN, n.v.).
Ravnia Oerst., Vidensk. Meddel. Dansk —Galium hypocarpium (L.) Endl. ex Griseb.
Naturhist. Foren. Kjobenhavn 1852: 49. 1853. (DC.) Hemsl., Biol.
Type: Ravnia triflora Oerst. [Herein treated as Centr.-Amer., Bot. 2: 63. 1881. Rubia
a synonym of Hillia Jacq. subg. Ravnia laevigata DC, Prodr. 4: 591. 1830. Type:
(Oerst.) C. M. Taylor.] MEXICO: Without precise locality, Haenke
Ravnia longifilamentosa Steyerm., Ceiba 3: 21. s.n. (Holotype presumbly PR, n.v.).—Galium
1952. Type: PANAMA: Chiriqui, upper microphyllum A. Gray
forested southern slopes of Quebrada Velo, mm Greenm., Proc. Amer.
vicinity of Finca Lerida, 5000 ft., 25 Jul 1947, Acad. Arts 41: 250. 1905. Type: MEXICO:
P. H. Allen 4744 (Holotype F 1394241 (photo Hidalgo, moist banks, barranca below
PTBG); Isotype MO, n.v.).—Hillia Trinidad Iron Works, 5500 ft., 6 May 1904,
longifilamentosa (Steyerm.) C. M. Taylor C. G. Pringle 8834 (Holotype GH, n.v.;
Ravnia panamensis Steyerm., Ceiba 3: 22. 1952. Isotypes MEXU, UC, n.v., US (photos
Not Hillia panamensis Standi., 1921. Type: MEXU, PTBG)).—Galium sphagnophilum
PANAMA: Code, North Hills, El Valle de (Greenm.) Dempster var. mazocarpum
Anton, 29 Jun 1946, P. H. Allen 3558 (Greenm.) Dempster
[collector erroneously omitted in protologue] Relbunium microphyllum (A. Gray) Hemsl.,
(Holotype F 1394242 (photo PTBG); Isotype Biol. Centr.-Amer., Bot. 2: 63. 1881. Galium
MO, n.v.).—Hillia allenii C. M. Taylor microphyllum A. Gray, PI. Wright. 1: 80.
Ravnia pittieri Standi., N. Amer. Fl. 32: 114. 1852. Type: U.S.A.: New Mexico, from
1921. Type: COSTA RICA: Santa Clara, Jul western Texas to El Paso, New Mexico,
1899, H. Pittier 1346J (Holotype NY, n.v.). May-Oct 1849, C. Wright 234 (Holotype GH,
—Hillia triflora (Oerst.) C. M. Taylor var. n.v.; Isotype US (photo PTBG)).—Galium
pittieri (Standi.) C. M. Taylor microphyllum A. Gray
Ravnia triflora Oerst., Vidensk. Meddel. Dansk Relbunium microphyllum (A. Gray) Hemsl. var.
Naturhist. Foren. Kjobenhavn 1852: 50. 1853. hispidulum Greenm., Proc. Amer. Acad. Arts
Type: COSTA RICA: Near Cartago and 33: 470. 1898. Syntypes: MEXICO: San Luis
Candelaria, 6000-8000 ft., Oersted 11696 Potosi, valley of San Luis Potosi, Schaffner
(Lectotype C, n.v. (photos F, MO, NY); 842 (GH, n.v.). Region of San Luis Potosi,
designated by Standley, 1938).—Hillia 1850-2460 m, Perry & Palmer 308 (GH,
triflora (Oerst.) C. M. Taylor var. triflora n.v).—Galium microphyllum A. Gray
Relbunium microphyllum (A. Gray) Hemsl. var.
setulosum Greenm., Proc. Amer. Acad. Arts
A NOMENCLATOR OF MEXICAN / D CENTRAL AMERICAN RUBIACEAE

33: 470. 1898. Syntypes: MEXICO: Hidalgo, terrenos arenosos, 23 Jun 1954, J. Rzedowsh
Zimapan, Coulter 186 (GH, n.v.); San Luis 3168 (Holotype ENCB).
Potosi, region of San Luis Potosi, C. C. Perry Richardia haenkeana (DC.) Steud., Nom. Bot.,
& E. Palmer 309 (GH, n.v.), C. C. Pern' & E. ed. 2, 459. 1841. Richardsonia haenkeana
Palmer 310 (GH, n.v.).~Galium DC, Prodr. 4: 569. 1830. Type: MEXICO:
microphyllum A. Gray Without precise locality, Haenke s.n.
Relbunium polyplocum Hemsl., Diag. PI. Nov. (Holotype presumably PR, n.v.).—Richardia
Mexic. 53. 1880. Type: MEXICO: San Luis humistrata (Cham. & Schltdl.) Steud.
Potosi, without precise locality, 1878, C. C. Richardia humistrata (Cham. & Schltdl.)
Parry & E. Palmer 308 (Syntype US 19271 Steud., Nom. Bot., ed. 2, 459. 1841.
(photos MEXU, PTBG), K, n.v.). MEXICO: Richardsonia humistrata Cham. & Schltdl.,
Veracruz, Mirador, Linden 516 (Syntype K, Linnaea 3: 353. 1828. Type: URUGUAY:
n.v.), Liebmann 243 (Syntype K, n.v.), Montevideo, in Brasihae meridionalis
provincia Montevideo, Sellow s.n. (Holotype
microphyllum A. Gray B, presumably destroyed (fragment F)).
m sphagnophilum Greenm., Proc. Richardia procumbens Sesse & Moc., Fl.
Amer. Acad. Arts 33: 468. 1898. Type: Mexic, ed. 2, 83. 1894. Type: MEXICO:
MEXICO: Oaxaca, Sierra de Clavellinas, Habitat in Ahualulci apricis, Sesse & Mocino
2270 m, 17 Oct 1894, C. G. Pringle 5812 s.n. (presumably MA, n.v.).—Richardia
(Holotype GH, n.v.).—Galium scabra L.
sphagnophilum (Greenm.) Dempster var. Richardia scabra L., Sp. PI. 1: 330. 1753.
sphagnophilum Type: MEXICO: Veracruz (Lectotype LINN
451.1 (n.v.); for typification see Lewis &
Rhodostoma Scheidw., Allg. Gartenzeitung 10: Oliver, 1974).
286. 1842. Type: Rhodostoma gardenioides Richardia tricocca (Torr. & A. Gray) Standi.,
Scheidw. Publ. Field Columbian Mus., Bot. Ser. 8: 386.
Rhodostoma gardenioides Scheidw., Allgem. 1931. subsp. tricocca. Diodia tricocca Torr.
Gartenzeitung 10: 286. 1842. Type: Based on & A. Gray, Fl. N. Amer. 2: 30. 1841. Type:
cultivated material of uncertain origin, U.S.A.: Texas, Drummond 117 (Holotype
presumably Veracruz, Mexico (type not GH, n.v.; Isotype G, n.v.).
designated). Palicourea.—Psychotria Richardia tricocca (Torr. & A. Gray) Standi,
gardenioides (Scheidw.) Standi. subsp. tetracocca (M. Martens & Galeotti)
W. H. Lewis & R. L. Oliv., Bnttonia 26: 287.
Richardia L., Sp. PI. 1: 330. 1753. Type: 1974. Spermacoce tetracocca M. Martens &
Richardia scabra L. Galeotti, Bull. Acad. Roy. Soc. Bruxelles 11:
Richardia adscendens (Pav. ex DC.) Steud., 132. 1844. Type: MEXICO: Michoacan,
Norn. Bot., ed. 2, 458. 1841. Richardsonia Santa Maria, near Morelia, H. Galeotti 2587
adscendens Pav. ex DC, Prodr. 4: 569. 1830. (Holotype BR, n.v.; Isotypes K, n.v., P
Type: MEXICO: Without precise locality, (photos MEXU, PTBG)).
Moricands.n. (Holotype G-DC, n.v. (photo
MO)).—Richardia brasiliensis Gomes Richardsonia Kunth in Humb., Bonpl. & Kunth,
Richardia brasiliensis Gomes, Mem. Nov. Gen. Sp. PI. 3: 350. 1819. [Herein
Ipecacuanha Bras. 31, t. 2. 1801. Type: treated as a synonym of Richardia L.]
BRAZIL: Without precise locality, Gomes s.n. Richardsonia adscendens Pav. ex DC, Prodr. 4:
(Holotype LISU, n.v.). 569. 1830. Type: MEXICO: Without precise
Richardia gandarae Rzed., Bol. Soc. Bot. locality, Moricands.n. (Holotype G-DC, n.v.
Mexico 44: 11, fig. 2. 1983. Type: MEXICO: (photo MO)). Richardia.—Richardia
San Luis Potosi, Santa Maria del Rio, 1750 m, brasiliensis Gomes
Richardsonia brasiliensis (Gomes) Hayne, Getr. Rogiera amoena Planch., Fl. Serr. Jard. 5: tab.
Darstell. Gew. 14: tab. 21. 1846. Richardia 442, figs. 1, 2. 1849. Type: Based on plants
brasUiensis Gomes, Mem. Ipecacuanha Bras. cultivated in Belgium, presumably from
31, tab. 2. 1801. Type: BRAZIL: Without Guatemalan seed (type not designated).
precise locality, Gomes s.n. (LISU, n.v.). —Rondeletia amoena (Planch.) Hemsl.
—Richardia brasiliensis Gomes Rogiera aprica (Lundell) Borhidi, Acta Bot.
Richardsonia haenkeana DC., Prodr. 4: 569. Hung. 33: 301. 1987. Rondeletia aprica
1830. Type: MEXICO: Without precise Lundell, Contrib. Univ. Michigan Herb. 7: 54.
locality, Haenke s.n. (Holotype presumably 1942. Type: MEXICO: Chiapas, Saxchanal,
PR, n.v). Richardia.—Richardia humistrata Sierra Madre, 1-5 Jul 1941, E. Matuda 4314
(Cham. & Schltdl.) Steudel (Holotype TEX-LL (photos MEXU, MO,
Richardsonia humistrata Cham. & Schltdl., PTBG); Isotypes A, n.v., F, MO, US (photos
Linnaea 3: 353. 1828. Type: URAGUAY: MEXU, PTBG))—Rondeletia cordata
Montevideo, in Brasiliae meridionalis Benth.
provincia Montevideo, Sellow s.n. (Holotype Rogiera backhousii (Hook, f.) Borhidi, Acta
B, presumably destroyed (fragment F)). Bot. Hung. 33: 301. 1987. Rondeletia
—Richardia humistrata (Cham.) Schltdl.) backhousii Hook, f., Bot. Mag. 103: tab.
Steud. 6290. 1877. Type: Based on material of
Richardsonia scabra (L.) A. St.-Hil., PI. Us. unknown origin cultivated at Kew, obtained
Bras. 8: 1.1. 8. 1824, p.p. Richardia scabra L., from the Backhouse establishment in York
Sp. PL 1: 330. 1753. Type: MEXICO: (type not designated).—Rondeletia
Veracruz, (Holotype LINN 451.1 (n.v.)). backhousii Hook. f.
—Richardia scabra L. Rogiera brachistantha (Standi. & Steyerm.)
Richardsonia tetracocca (M. Martens & Borhidi, Acta Bot. Hung. 33: 301. 1987.
Galeotti) K. Schum. ex Loes., Repert. Sp. Rondeletia brachistantha Standi. & Steyerm.,
Nov. 18: 362. 1922. Spermacoce tetracocca Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 25.
M. Martens & Galeotti, Bull. Acad. Roy. Soc. 1943. Type: GUATEMALA: San Marcos, Rio
Bruxelles 11: 132. 1844. Type: MEXICO: Vega, near San Rafael and Guatemala-Mexico
Michoacan, Santa Maria, near Morelia, H. boundary, Volcan Tacana, 2500-3000 m, 20
Galeotti 2587 (Holotype BR, n.v.; Isotypes K, Feb 1940, J. A. Steyermark 36286 (Holotype
n.v., P (photos MEXU, PTBG)). Diodia. F 1050045 (photos MEXU, PTBG); Isotype
—Richardia tricocca (Torr. & A. Gray) US (photos MEXU, PTBG)).—Rondeletia
Standi, subsp. tetracocca (M. Martens & cordata Benth.
Galeotti) W. H. Lewis & R. L. Oliv. Rogiera chiapasensis (Standi.) Borhidi, Acta
Richardsonia tetracocca var. calvescens Loes., Bot. Hung. 33: 301. 1987. Rondeletia
Repert. Sp. Nov. 18: 362. 1922. Syntypes: chiapasensis Standi., Publ. Field Mus. Nat.
MEXICO: Without precise locality, Schaffner Hist., Bot. Ser. 22: 54. 1940. Type: MEXICO:
281 (K, n.v.); Schaffner 282 (K, n.v.). Chiapas, Cascarada, 31 Dec 1936, E. Matuda
—Richardia tricocca (Torr. & A. Gray) 1064 (Holotype MICH, n.v. (fragment F);
Standi, subsp. tetracocca (M. Martens & Isotype A, n.v).—Gonzalagunia
Galeotti) W. H. Lewis & R. L. Oliv. chiapasensis (Standi.) Standi. & Steyerm.
Rogiera cordata (Benth.) Planch., Fl. Serr. Jard.
Rogiera Planch., Fl. Serr. Jard. 5: tab. 442. 5: tab. 442b. 1849. Rondeletia cordata Benth.,
1849. Lectotype: Rogiera amoena Planch, PI. Hartweg. 85. 1841. Type: GUATEMALA:
(designated by Borhidi, Acta Bot. Acad. S< In praeruptis prope Guatemala [Guatemala
Hung. 28: 67. 1982). [Herein treated as a City], Jan 1841, T. Hartweg 585 (Holotype K
synonym of Rondeletia L.] (photo PTBG); Isotype G-DEL, n.v. (fragment
F)).—Rondeletia cordata Benth.
 A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Rogiera ehrenbergii (K. Schum. ex Standi.) Centroamericanae Oersted 11678] (Holotype

Borhidi, Acta Bot. Acad. Sci. Hung. 28: 67. presumably C, n.v.; Isotypes K (photo PTBG),
1982. Rondeletia ehrenbergii K. Schum. ex US (photo PTBG)).—Rondeletia amoena
Standi., N. Amer. Fl. 32: 50. 1918. Type: (Planch.) Hemsl.
GUATEMALA: Totonocapan, Ehrenberg Rogiera ligustroides (Hemsl.) Borhidi, Acta
1033 (Holotype US 594426 (photos F, Bot. Acad. Sci. Hung. 28: 67. 1982.
MEXU, PTBG); Isotype B, presumably Rondeletia ligustroides Hemsl., Diag. PI. Nov.
destroyed (photos F, MO, US)).—Rondeletia Mexic. 26. 1879. Type: MEXICO: Veracruz,
gratissima Planch. & Linden Orizaba, Botteri 971 (Lectotype K, here
Rogiera elegans Planch., Fl. Serr. Jard. 5: tab. designated (photo PTBG); Isolectotypes P (2
442. 1849. Type: Based on cultivated material sheets, photos MEXU, PTBG)). Syntypes:
grown in Belgium of Guatemalan origin (type MEXICO: Veracruz, Orizaba, Bourgeau 2518
not designated).—Rondeletia amoena (K (photo PTBG), P (3 sheets; photos MEXU,
(Planch.) Hemsl. PTBG)). Region d'Orizaba, Bourgeau 2622
Rogiera gratissima Planch. & Linden, Fl. Serr. (K (photo PTBG), B (n.v.; fragment F), F, K
Jard. 15: 133, and unnumbered figure. 1864. (photo PTBG), P (4 sheets; photos MEXU,
Type: Based on cultivated material grown in PTBG), US (2 sheets)). Orizaba, Botteri 474
Belgium brought from Chiapas, Mexico, by (K, 2 sheets: label on first states "coll. Salle,"
Ghiesbreght (type not designated). Note: two second states "Sumichrast coll."; photos
collections at K ["From the garden of Sir PTBG), P (photos MEXU, PTBG). Orizaba,
C. W. Strickland, Reed. Apl. 1897," and Botteri 1951 (K, n.v., P (2 sheets; photos
"Hort. Kew 1879"] are not types. MEXU, PTBG). Orizaba, Muller 969 (K, n.v.,
P (photos MEXU, PTBG). Orizaba, Muller
1308 (K, n.v.). The K sheet of Botteri 971 is
Rogiera intermedia (Hemsl.) Borhidi, Acta Bot. selected as lectotype because it has three leafy
Acad. Sci. Hung. 28: 67. 1982. Rondeletia twigs, an abundance of flowers at anthesis,
intermedia Hemsl., Diag. PI. Nov. Mexic. 26. and two duplicates at P.—Rondeletia
1879. Type: MEXICO: Chiapas, Linden 1661 ligustroides Hemsl.
(Lectotype K (photo PTBG), here designated; Rogiera menechma Planch., Fl. Serr. Jard. 5:
Isolectotype P (photos MEXU, PTBG)). tab. 442. 1849. Type: Based on cultivated
—Rondeletia cordata Benth. material grown in Belgium, of Guatemalan
Rogiera langlassei (Standi.) Borhidi, Acta Bot. origin (type not designated).
Acad. Sci. Hung. 28: 67. 1982. Rondeletia Rogiera pittieri (K. Schum. & K. Krause)
langlassei Standi., N. Amer. Fl. 32: 53. 1918. Borhidi, Acta Bot. Acad. Sci. Hung. 67. 1982.
Type: MEXICO: [Michoacan or Guerrero], Rondeletia pittieri K. Schum. & K. Krause ex
Sierra Madre, 1750 m, 26 Jan 1899, E. K. Schum., Bot. Jahrb. 40: 316. 1908. Type:
Langlasse 797 (Holotype US 386194 (photos COSTA RICA: Bords du Rio Poros, 2000 m,
MEXU, PTBG); Isotypes B, presumably 10 Jan 1890, H. Pittier & T. Durand 1729
destroyed (fragment F), G-DEL, n.v. (Holotype B, presumably destroyed (photos F,
(fragment F), K (photo PTBG), P (2 sheets; MO); Isotype US (photos MEXU, PTBG)).
photos MEXU, PTBG)).—Rondeletia —Rondeletia amoena (Planch.) Hemsl.
amoena (Planch.) Hemsl. Rogiera roezlii Planch., Fl. Serr. Jard. 5: tab.
Rogiera latifolia (Oerst.) Decne., Rev. Hortic, 442. 1849. Type: Based on cultivated material
ser. 4, 2: 121. 1853. Rondeletia latifolia grown in Belgium, probably of Guatemalan
Oerst., Vidensk Meddel. Dansk Naturhist. origin (type not designated).—Rondeletia
Foren. Kjobenhavn 1852: 43. 1853. Type: amoena (Planch.) Hemsl.
NICARAGUA: In monte Pantasmo, in Rogiera seleriana (Loes.) Borhidi, Acta Bot.
Segovia, Jan 1848, Oersted 76 [=Plantae Hung. 35: 311. 1989. Rondeletia seleriana
 Loes., Verh. Bot. Ver. Brandenb. 65: 105. Rogiera versicolor (J. Sm.) Lindl. & Paxton,
1923. Type: GUATEMALA: Huehuetenango, Paxton Fl. Gard. 2: 69. July 1851. Rondeletia
Nenton Distr., near Chacula, E. Seler & C. versicolor J. Sm., Bot. Mag. 7: tab. 4579.
Seler 3015 (Holotype B, presumably 1851. Type: Based on cultivated material
destroyed (photos F, MO, US), fragment F). grown at Kew sent by Seemann from
—Rondeletia gratissima (Planch. & Linden) Boquete, Veraguas, Panama (type not
Hemsl. designated).—Rondeletia amoena (Planch.)
Rogiera stenosiphon (Hemsl.) Borhidi, Acta Hemsl.
Bot. Acad. Sci. Hung. 28: 67. 1982.
Rondeletia stenosiphon Hemsl., Diag. PI. Ronabea Aubl., Hist. PI. Guiane 1: 154; 2
Nov. Mexic. 26. 1879. Type: MEXICO: (Index): 26, t. 59. 1775. Type: Ronabea
Yucatan and Tabasco, without precise latifolia Aubl. [Herein treated as a synonym of
locality, E. P. Johnson s.n. (Holotype K Psychotria L.]
(photo PTBG)).—Rondeletia stenosiphon Ronabea erecta Aubl, Hist. PI. Guiane 1: 156.
Hemsl. 1775. Type: FRENCH GUIANA: In sylvis
Rogiera strigosa (Benth.) Borhidi, Acta Bot. Oyao, Orapu, Sinemari, Aublet s.n. (Holotype
Acad. Sci. Hung. 28: 67. 1982. Bouvardia presumably BM, n.v.).—Psychotria erecta
strigosa Benth., PI. Hartweg. 75. 1841. Type: (Aubl.) Standi. & Steyerm.
GUATEMALA: Quezaltenango, [Volcan]
Zunil, Nov., T. Hartweg 530 (Lectotype K Rondeletia L., Sp. PI. 1: 172. 1753. Lectotype:
(photo PTBG), here designated; Isolectotypes Rondeletia americana L. (Designated by
G-DEL, n.v. (fragment F), P (photos F, Howard, 1989.)
MEXU, MO, PTBG), W, n.v. (photos F, Rondeletia acuminata (Oerst. ex Standi.)
MO)). Syntype: GUATEMALA: Volcan de Lorence & Cast.-Campos, Biotica 13: 147.
Fuego, 6000-6400 ft., Salvin s.n. (K (photo 1988. Sommera acuminata Oerst. ex Standi.,
PTBG, fragment F)). The K specimen of J. Wash. Acad. Sci. 17: 340. 1927. Type:
Hartweg 530 is the better choice for lectotype, MEXICO: Oaxaca, Amatlan, F. Liebmann
with duplicates in three other herbaria. 11712 (Holotype C, n.v. (photo MEXU);
—Rondeletia strigosa (Benth.) Hemsl. Isotype US (photos MEXU, PTBG)).
Rogiera subscandens (Lundell) Borhidi, Acta Arachnothryx.— Rondeletia scabra Hemsl.
Bot. Hung. 35: 311. 1989. Rondeletia Rondeletia aetheocalymna Donn. Sm., Bot.
subscandens Lundell, Wrightia 5: 328. 1976. Gaz. (Crawfordsville) 42: 298. 1906. Type:
Type: GUATEMALA: Peten, La Cumbre, GUATEMALA: Alta Verapaz, trail between
about 500 m west of km 142/143 of the Sepacuite and Secanquim, 550-1000 m, 14
Cadenas Road, 10 Sep 1976, C. L. Lundell & Jan 1905, W. R. Maxon & R. Hay 3275
E. Contreras 19830 (Holotype TEX-LL (Holotype US 473244 (photos MEXU,
(photos MEXU, PTBG); Isotypes F (photo PTBG); Isotype US (photos MEXU, PTBG)).
PTGB), K (photo PTBG)).—Rondeletia Arachnothryx.
subscandens Lundell Rondeletia affinis Hemsl., Diag. PI. Mex. 28.
Rogiera suffrutescens (Brandegee) Borhidi, Acta 1879. Type: MEXICO: Veracruz, Zongolica
Bot. Acad. Sci. Hung. 28: 67. 1982. [Sierra de Zongolica], Jun 1866, Sumichrast
Rondeletia suffrutescens Brandegee, Univ. [sub Botteri 1017] (Lectotype K, here
Calif. Publ. Bot. 6: 70. 1914. Type: MEXICO: designated; Isolectotypes K (photo PTBG), P
Chiapas, Cerro del Boqueron, Aug 1913, C. A. (2 sheets, photos MEXU, MO, PTBG)).
Purpus 6704 (Holotype UC 172451, n.v.; Syntypes: MEXICO: Chiapas, Ghiesbreght
Isotypes F (photos MEXU, PTBG), MO, US 686 (K (photo PTBG), P (photos MEXU,
(photos MEXU, PTBG)).—Rondeletia PTBG), MO). COSTA RICA: Endres 226
suffrutescens Brandegee (K). PANAMA: Veraguas, Seemann 1594
 A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

(K). The K specimen of Sumichrast sub Rondeletia belizensis Standi, var. longiloba
Botteri 1017 is the best choice for lectotype Lundell, Wrightia 5: 323. 1976. Type:
and has duplicates in two other herbaria. GUATEMALA: Alta Verapaz, Rubelsanto,
—Rondeletia buddleioides Benth. 2.3 km SEE [=ESE] of Balastrera, 20 Jul
Rondeletia albida Lundell, Wrightia 5: 323. 1975, C. L. Lundell & E Contreras 19545
1976. Type: MEXICO: Chiapas, Carelas, near (Holotype TEX-LL (photos MEXU, PTBG);
Motozintla, 2176, Apr 1945, E. Matuda 5509 Isotypes F, MO (photo PTBG)).—Rondeletia
(Holotype TEX-LL (photos MEXU, PTBG); $ Standi.
Isotypes F, MEXU, TEX-LL).—Rondeletia i bertieroides Standi., Publ. Field
laniflora Benth. Columbian Mus., Bot. Ser. 4: 267. 1929.
Rondeletia amoena (Planch.) Hemsl., Diag. PI. Type: PANAMA: Bocas del Toro, Buena
Mex. 26. 1879. Rogiera amoena Planch., Fl. Vista Camp, Chiriqui Trail, 900 m, 1928,
Serr. Jard. 5: tab. 442, figs 1, 2. 1849. Type: G. P. Cooper 598 (Yak 12231) (Holotype F
Based on plants cultivated in Belgium, 579614 (photo PTBG); Isotype NY, n.v.).
presumably from Guatemalan seed (type not Arachnothryx.
designated). Rondeletia bourgaei Standi., N. Amer. Fl. 32:
Rondeletia aprica Lundell, Contrib. Univ. 56. 1918. Type: MEXICO: Veracruz, Region
Michigan Herb. 7: 54. 1942. Type: MEXICO: de Orizaba, Monte San Cristobal, 8 Jul 1866,
Chiapas, Saxchanal, Sierra Madre, in E. Bourgeau 2618 (Holotype US 49964
advanced forest, 1-5 Jul 1941, E. Matuda (photos MEXU, PTBG); Isotypes P (4 sheets,
4314 (Holotype TEX-LL (photos MEXU, photos MEXU, PTBG)). Arachnothryx.
MO, PTBG); Isotypes A, n.v., F, MO, US —Rondeletia laniflora Benth.
(photos MEXU, PTBG)). Rogiera. Rondeletia brachistantha Standi. & Steyerm.,
—Rondeletia cordata Benth. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 25.
Rondeletia aspera Standi., N. Amer. Fl. 32: 54. 1943. Type: GUATEMALA: San Marcos, Rio
1918. Type: COSTA RICA: San Jose, Pont du Vega, near San Rafael and Guatemala-Mexico
Rodeo, Pacacol (Villa Colon), Jan 1891, f/. boundary, Volcan Tacana, 2500-3000 m, 20
Pittier 3243 (Holotype US 941408 (photo Feb 1940, J. A. Steyermark 36286 (Holotype
PTBG)). Arachnothryx. F 1050045 (photos MEXU, PTBG); Isotype
Rondeletia atravesadensis Lorence, Novon 1: US (photos MEXU, PTBG)). Rogiera.
137,/jg. 1A, B. 1991. Type: MEXICO: —Rondeletia cordata Benth.
Oaxaca, Cerro Atravesada-Cerro Azul, Rondeletia brandegeeana Lorence, Novon 1:
Arroyo Rana, 6 Dec 1956, T. MacDougall s.n. 138. 1991. Based on Otocalyx chiapensis
(Holotype MEXU 253128 (photo PTBG)). Brandegee, Univ. Calif Publ. Bot. 6: 69.
Rondeletia backhousii Hook.f, Bot. Mag. 103: 1914. Not Rondeletia chiapasensis Standi..
tab. 6290. 1877. Based on material of 1940. Type: MEXICO: Chiapas, Cerro del
unknown origin cultivated at Kew, obtained Boqueron, middle region, moist valley slopes,
from the Backhouse establishment in York Aug 1913, C. A. Purpus 7041 (Holotype UC
(type not designated). Rogiera.—Status 173047 (photo PTBG); Isotypes GH, US
uncertain, resembles Rondeletia panamensis (fragment)).
DC. but is nearly glabrous. Rondeletia breedlovei Lorence. Novon 4: 129,
Rondeletia belizensis Standi., Carnegie Inst. fig 6. 1994. Type: MEXICO: Chiapas, Mun.
Wash. Publ. 461: 91. 1935. Type: BRITISH of Tenejapa, paraje of Mahben Chauk, on
HONDURAS (BELIZE): Jacinto Hills, 300 slopes along the Tana Te' River near Shal
ft., 28 Aug 1933, W. A. Schipp 1201 K'esh, 2900 ft., 27 Nov 1964, D. E Breedlove
(Holotype F 683600 (photos MEXU, PTBG); 7682 (Holotype F 1617735 (photo PTBG);
Isotypes GH, n.v., K (photo PTBG), MO). Isotype CAS (photo PTBG)).
Arachnothryx.
Rondeletia brenesii Standi., Publ. Field Mus. 1993. Type: COSTA RICA: Heredia,
Nat. Hist., Bot. Ser. 18: 1370. 1938. Type: Magsasay, pasturas, bosque socolado, finca de
COSTA RICA: Camino de San Ramon a La Bernardo Gomez, 700 m, 1 Jan 1983, /. C.
Calera, 20 Mar 1929, A. M. Brenes 6757 Chacon 177 (Holotype CR 92850; Isotype
(Holotype F 854972 (photo PTBG)). MO).
Arachnothryx. Rondeletia chiapasensis Standi., Publ. Field
Rondeletia buddleioides Benth., PI. Hartweg. Mus. Nat. Hist., Bot. Ser. 22: 54. 1940. Type:
69. 1840. Type: MEXICO: Oaxaca, in planitie MEXICO: Chiapas, Cascarada, 31 Dec 1936,
Llano Verde, T. Hartweg 502 (Holotype K E. Matuda 1064 (Holotype MICH, n.v.
(photo PTBG)); Isotypes B (destroyed, photo (fragment F); Isotype A, n.v.). Rogiera.
MO, fragment G-DEL), K (photo PTBG), —Gonzalagunia chiapasensis (Standi.)
NY, n.v. (fragment US, photos MEXU, Standi. & Steyerm.
PTBG), P (photos MEXU, PTBG)). Rondeletia chinajensis Standi. & Steyerm.,
Arachnothryx. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.
Rondeletia buddleioides Benth. var. aspera J. H. 1947. Type: GUATEMALA: Alta Verapaz,
Kirkbr., Ann. Missouri Bot. Gard. 55: 379. Cerro Chinaja, between Finca Yalpemech and
1968. Type: PANAMA: Bocas del Toro, Chinaja, above source of Rio San Diego,
between Criollo (just above Buena Vista) and 150-700 m, 1-2 Apr 1942, J. A Si
Quebrada Higueron on Chiriqui Trail, 18 Apr 45637 (Holotype F 1134210 (photos MEXU,
1968, J. H. Kirkbride, Jr. &J.A. Duke 780 PTBG)). Arachnothryx.
(Holotype MO 1942973 (photos MEXU, Rondeletia chiriquiana Lorence, Novon 4:
PTBG); Isotype US). Arachnothryx. 129, fig. 7. 1994. Type: PANAMA: Chiriqui,
—Rondeletia buddleioides Benth. Fortuna Dam area, along shores of Rio
Rondeletia calycosa Donn. Sm., Bot. Gaz. Chiriqui where it enters reservoir, 1000 m,
(Crawfordsville) 56: 59. 1913. Type: COSTA 8°45'N, 82°13'W, 2 Aug 1984, H. W.
RICA: San Jose, forets de La Palma, 1460 m, Churchill 5924 (Holotype MO 3595277
25 Sep 1898, A. Tonduz 11633 (Holotype US (photo PTBG)).
no 941406 (photos F, MEXU, PTBG)). Rondeletia cooperi Standi., Publ. Field
Arachnothryx. Columbian Mus., Bot. Ser. 4: 267. 1929.
Rondeletia capitellata Hemsl., Diag. PI. Mex. Type: PANAMA: Bocas del Toro, Buena
28. 1879. Type: MEXICO: Without precise Vista Camp, on Chiriqui Trail, 1250 ft.,
locality, Jurgensen 355 (Lectotype K, here Jan-Mar 1928, G. P. Cooper 600 (Yale
designated (photo PTBG); Isolectotypes G- 12233) (Holotype F 579596 (photo PTBG);
DEL, n.v. (fragment F), K (photo PTBG). Isotype NY, n.v.). Arachnothryx.
Syntypes: MEXICO: Oaxaca, Joya, F. Rondeletia cordata Benth., PI. Harrweg. 85.
Liebmann 202 (K (photo PTBG)). MEXICO: 1841. Type: GUATEMALA: In praeruptis
Without precise locality, Jurgensen 376 (K prope Guatemala [Guatemala City], Jan 1841,
(photo PTBG)). Arachnothryx. The K sheet of T. Hartweg 585 (Holotype K (photo PTBG);
Jurgensen 355 is very floriferous and Isotype G-DEL, n.v. (fragment F)). Rogiera.
represents the best choice for lectotype. Note: Rondeletia cordovana Standi. & Steyerm., Publ.
Rondeletia capitellata and R. liebmannii Field Mus. Nat. Hist., Bot. Ser. 23: 26. 1943.
Standi, are synonymous. Material from Type: GUATEMALA: Zacapa, in ravine
Veracruz and Hidalgo erroneously identified bordering Quebrada Alejandria, summit of
as R. capitellata in many herbaria actually Sierra de las Minas, vicinity of Finca
represents a distinct species, R. pringlei Alejandra, 2500 m, 13 Oct 1939, J. A.
Steyermark 29906 (Holotype F 1042952
(photos MEXU, PTBG); Isotype US (photos
 A NOMENCLATOR OF MEXICAN AND CENTRAL /

MEXU, PTBG)). Arachnothryx.—Rondeletia (photo PTBG); Isolectotypes G-DEL, n.v.

brandegeeana Lorence (fragment F), P, (photos MEXU, PTBG), W,
Rondeletia cornifolia Benth., PL Hartweg. 39. n.v. (photos F, MO)). Syntype: MEXICO:
1840. Type: MEXICO: Michoacan, prope Sierra Madre, N. W. of Mexico, Seemann
Morelia, T. Hartweg 299 (Holotype K (photo 2057 (K (photo PTBG)). The K sheet of
MEXU); Isotypes BM, G-DEL, n.v. (fragment Andrieux 334 is richly flowering and has
F), P (photos MEXU, MO, PTBG), US duplicates in three other herbaria.
(fragment, photos MEXU, PTBG), W (2 —Rondeletia leucophylla Kunth in Humb.,
sheets, photos PTBG)). Choristes.—Deppea Bonpl. & Kunth
cornifolia (Benth.) Benth. Rondeletia dukei Dwyer & Hayden, Ann.
Rondeletia costaricensis Standi., N. Amer. Fl. Missouri Bot. Gard. 54: 144. 1967. Type:
32: 61. 1918. Type: COSTA RICA: Buena PANAMA: Darien, Rio Pirre, 24 May 1966,
Vista, San Carlos, 900 m, Apr 1903, H. Pittier J. A. Duke 8329 (Holotype MO, unnumbered
16696 (Holotype US 764141 (photos MEXU, sheet (photo PTBG)).—Wittmackanthus
PTBG); Isotypes B, presumably destroyed stanleyanus (R. H. Schomb.) Kuntze
(fragment F), US). Arachnothryx.
Rondeletia darcyi Dwyer, Ann. Missouri Bot. 1991. Based on Chomelia leucophylla Dwyer,
Gard. 67: 467. 1980. Type: PANAMA: Colon, Ann. Missouri Bot. Gard. 67: 98. Jig. 21.
road bank just W of Portobelo, 12 Apr 1970, 1980. Not Rondeletia leucophylla Kunth in
W. G.D'Arcy 4079 (Holotype MO 2238438 Humb., Bonpl. & Kunth, 1820. Type:
(photo PTBG); Isotype MO (photo PTBG)). PANAMA: Darien, slopes of Cerro Chucula,
Arachnothryx.—Rondeletia secunda Standi. drainage of Rio Pavarando, 11 Feb 1972, A.
Rondeletia dariensis Standi., N. Amer. Fl. 32: Gentry 4262 (Holotype MO 2097508; Isotype
53. 1918. Type: PANAMA: Darien, Boca de MO 2161959).
Paurando, on Sambii River, southern Darien, Rondeletia edwardsii Standi., Trop. Woods 37:
20 m, Feb 1912, H. Pittier 5684 (Holotype US 31. 1934. Type: HONDURAS: Comayagua,
867136 (photos F, MEXU, PTBG, US); Pito Solo, Lake Yojoa, 2000 ft., 29 Aug 1932,
Isotypes F (photo PTBG), US (2 sheets)). / B. Edwards P476 (Holotype F 662645
Arachnothryx. (photos MEXU, PTBG); Isotypes A, n.v.,
Rondeletia deamii (Donn. Sm.) Standi., N. US). Arachnothryx.
Amer. Fl. 32: 60. 1918. Bouvardia deamii Rondeletia eherenbergii K. Schum. ex Standi.,
Donn. Sm., Bot. Gaz. (Crawfordsville) 49: N. Amer. Fl. 32: 50. 1918. Type:
455. 1910. Type: GUATEMALA: Guatemala, GUATEMALA: Totonocapan, Ehrenberg
at base of Rillo de las Nabajas, about V* mile 1033 (Holotype US 594426 (photos F,
front side of Fiscal, 3700 ft., 5 Jun 1909, C. C MEXU, PTBG); Isotype B, presumably
Deam 6190 (Holotype US 579579 (photos destroyed (photos F, MO, US)). Rogiera.
MEXU, PTBG); Isotypes A, n.v., F (photo —Rondeletia gratissima Planch. & Linden
PTBG), GH, n.v., MO (photos MEXU, Rondeletia elongata Bartl. ex DC, Prodr. 4:
PTBG), US (photos MEXU, PTBG)). 409. 1830. Type: MEXICO: Guerrero, ad
Arachnothryx. Acapulco, Haenke s.n. (Holotype presumably
Rondeletia dubia Hemsl., Diag. PL Mex. 28. PR, n.v.; Isotype W, n.v. (photos F, MO)).
1879. Based on Bouvardia discolor Hook. & Arachnothryx.—Rondeletia leucophylla
Am., Bot. Beechy Voy. 428. 1840. Not Kunth in Humb., Bonpl. & Kunth
Rondeletia discolor Kunth in Humb.. Bonpl. Rondeletia evansii Lorence, Novon 7: 52, fig.
& Kunth, 1820. Type: MEXICO: Oaxaca, 4. 1997. Type: HONDURAS: Dept. Lempira,
Inter Tehuantepec et Voca [Boca] del Monte, Parque Nacional de Celaque, sendero entre la
in dit. Oaxaca, maxime [?] australi, Sept., G.
Andrieux 334 (Lectotype K, here designated >DonTomas, 14°36'N, 88°40'W, 1800 m,
 28 Jan 1992, P. House 1163 (Holotype PTBG Rondeletia gratissima (Planch. & Linden)
16112; Isotypes EAP, n.v., HEH, n.v., MO, Hemsl., Diag. PI. Nov. Mexic. 25. 1879.
TEFH, n.v.). Rogiera gratissima Planch. & Linden, Fl.
Rondeletiafalciformis Lundell, Wrightia 5: 324. Serr. Jard. 15: 133, and unnumbered figure.
1976. Type: HIDALGO: Km 332 of Mexico 1864. Type: Based on cultivated material
City-Laredo highway, above Chapulhuacan, grown in Belgium brought from Chiapas,
3800 ft., 13 Jul 1943, C. L. Lundell 12234 Mexico by Ghiesbreght (type not designated).
(Holotype TEX-LL (photos MEXU, PTBG); Note: two collections at K ["From the garden
Isotypes F (photo PTBG), K (photo PTBG)). of Sir C. W. Strickland, Reed. Apl. 1897," and
Arachnothryx.—Rondeletia heteranthera "Hort. Kew 1879"] are not types.
Brandegee Rondeletia guerrerensis Lorence, Novon 1:
Rondeletia galeottii Standi., N. Amer. Fl. 32: 143, fig. 8C, D. 1991. Type: MEXICO:
59. 1918. Type: MEXICO: Without precise Guerrero, Mun. de Atoyac de Alvarez, a 23.5
locality, H. Galeotti s.n. (Holotype US km al NE de El Paraiso, camino Atoyac-
454830 (photos MEXU, PTBG); probable Puerto del Gallo, 1700 m, 22 Nov 1983, E.
Isotypes [labels state "Chinantla, 1844, fl. Martinez S. & F. Barrie 5596 (Holotype
Juillet, Galeotti 2684C'] F, P, n.v. (fragment MEXU (photo PTBG); Isotypes BM, CAS,
F)). Arachnothryx. CHAPA, F, MO, PTBG).
Rondeletia ginetteae Lorence, Novon 1: 141, Rondeletia hameliifolia Dwyer & Hayden,
fig. 2A-G. 1991. Type: MEXICO: Oaxaca, Ann. Missouri Bot. Gard. 54: 144. 1967 [as
Distr. de Tuxtepec, Ruta 175 Tuxtepec to hamelifolia]. Type: PANAMA: Veraguas,
Oaxaca, ca. 10 km S of Valle Nacional, Sierra Santiago, Aug 1961, J. D. Dwyer 1350
de Juarez, 750 m, 6 Dec 1980, D. H. Lorence (Holotype MO 1824657 (photo PTBG);
& R. Cedillo T. 2973 (Holotype MEXU Isotype NY, n.v.). Arachnothryx.
(photo PTBG); Isotypes F (photo PTBG), Rondeletia heteranthera Brandegee, Univ.
MO). Calif. Publ. Bot. 4: 387. 1913. Type:
Rondeletia gonzaleoides Standi., Publ. Field MEXICO: Veracruz, Banos del Carrizal, Aug
Columbian Mus., Bot. Ser. 8: 160. 1930. 1912, C. A. Purpus 6184 [as 6148 in
Type: MEXICO: Chiapas, Pueblo Nuevo protologue] (Holotype UC 155217, n.v.;
[Solistahuacan], 2200 m, 1840, J. Linden 149 Isotypes F, MO). Arachnothryx.
(Holotype LE, n.v. (fragment F, photos Rondeletia hondurensis Donn. Sm., Bot. Gaz.
MEXU, PTBG); Isotype K [label states (Crawfordsville) 27: 335. 1899. Type:
"forets humides de Sulusuchiapas"] (photo HONDURAS: Santa Barbara, Rio
PTBG, fragment F), probable isotype P Chamelecon, 300 m, Dec 1888, C. Thieme
(photos MEXU, PTBG)). Arachnothryx. 5267 (Holotype US 943502 (photos MEXU,
Rondeletia gracilis Hemsl., Diag. PI. Nov. PTBG); Isotypes F (photos MEXU, PTBG),
Mexic. 53. 1880. Type: GUATEMALA: Alta US (2 sheets; photos MEXU, PTBG)).
Vera Paz, 3600 ft., March 1879, H. von Javorkaea.
Tuerckheim 404 (Holotype K (photo PTBG)). Rondeletia intermedia Hemsl., Diag. Pi. Nov.
Arachnothryx. Mexic. 26. 1879. Type: MEXICO: Chiapas, a
Rondeletia gracilispica Standi., Publ. Field fleurs rouges, forets de pins de Chiapas,
Columbian Mus., Bot. Ser. 8: 342. 1931. Linden 1661 (Lectotype K, here designated;
Type: MEXICO: Oaxaca, Sierra San Pedro Isolectotype P (photos MEXU, PTBG;
Nolasco, Talea, etc., 1843-44, C. Jurgensen fragment F). Syntypes: MEXICO: Chiapas,
935 (Holotype K (photos F, PTBG, US), Pueblo Nuevo, fl. blanches-rouges avec
fragment F (photos MEXU, PTBG)). extremites, fevrier 1840, Linden 1659 (K), G-
Arachnothryx. DEL, n.v. (fragment F), P (photos MEXU,
PTBG; fragment F)). MEXICO: Oaxaca,
 V NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

Sierra de San Pedro Nolasco, Jurgensen 798 presumably destroyed (fragment F), G-DEL,
(K (photo PTBG)). A probable isolectotype at n.v. (fragment F), K (photo PTBG), P (2
P [Linden s.n. (fragment F)] annotated "fleurs sheets; photos MEXU, PTBG)). Rogiera.
rosatres" has corolla tubes 6-7 mm long and —Rondeletia amoena (Planch.) Hemsl.
is likely a duplicate of Linden 1661 at K. A Rondeletia laniflora Benth., PL Hartw. 85.
probable syntype at P annotated "fleurs 1841. Type: GUATEMALA: Mountains of
rouges" has corolla tubes 5-7 mm long and Las Nubes, March 1841, T Hartweg 584
larger leaves, and is likely a duplicate of (Holotype K (photo PTBG), fragment US
Linden 1659 at K. The Linden 1661 specimen (photos MEXU, PTBG); Isotypes B,
at K is selected as lectotype, because it has presumably destroyed (photos F, MO, US), G-
more leaves, inflorescences, and young fruit. DEL, n.v. (fragment F), P (photos MEXU,
Hemsley noted that this species was PTBG), W, n.v. (photo F)). Arachnothryx.
intermediate between R. amoena and R. Rondeletia latifolia Oerst., Vidensk Meddel.
cordata, but the original material has sparsely Dansk Naturhist. Foren. Kjobenhavn 1852:
pubescent lower leaf surfaces and is clearly 43. 1853. Type: NICARAGUA: In monte
synonymous with R. cordata. In the Flora of Pantasmo in Segovia, Jan 1848, Oersted 76
Guatemala Rubiaceae treatment Standley & [=Plantae centroamericanae Oersted 11678]
Williams (1975) also treated this species as a (Holotype C, n.v.; Isotypes K (photo PTBG),
synonym of R. cordata Benth. Rogiera. US (photo PTBG)). Rogiera.—Rondeletia
amoena (Planch.) Hemsl.
a izabalensis Standi. & Steyerm., Rondeletia leptodicfta B. L. Rob., Proc. Amer.
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 286. Acad. Arts 45: 402. 1910. Type: MEXICO:
1940. Type: GUATEMALA: Izabal, Escoba, [Boundary between Michoacan and
across the bay (west) from Puerto Barrios, Guerrero], Rives du Rio Petatlan, 500 m, 24
near sea level, 3 May 1939, P. C. Standley Nov 1898, E Langlasse 666 (Holotype GH,
72891 (Holotype F 991611 (photos MEXU, n.v.; Isotypes F (photos MEXU, PTBG), G-
PTBG)). Arachnothryx. DEL, n.v. (fragment F), K (photo PTBG), P (2
Rondeletia jurgensenii Hemsl., Diag. PL Nov. sheets; photos MEXU, PTBG; fragment F),
Mexic. 29. 1879. Type: MEXICO: Without US (photos MEXU, PTBG)). Arachnothryx.
precise locality [protologue states "Oaxaca, —Rondeletia leucophylla Kunth in Humb.,
Sierra de San Pedro Nolasco"], Jurgensen 248 Bonpl. & Kunth
(Holotype K (photo PTBG); Isotype G-DEL, Rondeletia leucophylla Kunth in Humb.,
n.v. (fragment F, label states "Trapiche de Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 395, t.
Concepcion and Chicahuastla, Jun 1845")). 290. 1820. Type: MEXICO: Guerrero, inter
Arachnothryx. Alto del Peregrino et flumen Papagallo,
Rondeletia kirkbridei Dwyer, Ann. Missouri Humboldt & Bonpland s.n. (Holotype P-
Bot. Gard. 67: 469. 1980. Type: PANAMA: Bonpland, n.v.; Isotypes B, presumably
Veraguas, 16 km NW of Santa Fe, on road to destroyed (photos F, MO), P (photos MEXU,
Calovebora (Panama Hwy. 35), 300-500 m, PTBG)). Arachnothryx.
16 May 1975, S. Mori & J. Kallunki 6112 Rondeletia leucophylla Kunth in Humb., Bonpl.
(Holotype MO 2300094 (photo PTBG)). & Kunth var. calycosa Greenm., Proc. Amer.
Arachnothryx.—Rondletia bertieroides Acad. Arts 39: 92. 1903. Type: MEXICO:
Standi. Oaxaca, 1650 m, Dec 1900, C. Conzatti & V.
Rondeletia langlassei Standi., N. Amer. Fl. 32: Gonzalez 1127 (Syntype GH, n.v.); Oaxaca,
53. 1918. Type: MEXICO: [Michoacan or city of Oaxaca, 1550 m, C. Conzatti 970
Guerrero] Sierra Madre, 1750 m, 26 Jan 1899, (Syntype GH, n.v.).—Rondeletia leucophylla
E Langlasse 797 (Holotype US 386194 Kunth in Humb., Bonpl. 8c Kunth
(photos MEXU, PTBG); Isotypes B,
Rondeletia liebmannii Standi., J. Wash. Acad.
Sci. 17: 336. 1927. Type: MEXICO: Oaxaca,
without precise locality, 1841^3, F. Rondeletia macdougallii Lorence, Novon 1:
Liebmann J1834 (Holotype US 1315231 144, Jig. 3A, B. 1991. Type: MEXICO:
(photos MEXU, PTBG); Isotypes C, n.v. Oaxaca, Cerro Azul (top), N of Niltepec, 7000
(photo F), P (photos MEXU, PTBG)). ft., 7 Mar 1956, T. MacDougalls.n. (Holotype
Arachnothryx.—Rondeletia capitellata MEXU 253131 (photo PTBG)).
Hemsl. Rondeletia macrocalyx Standi. & Steyerm.,
Rondeletia ligustroides Hemsl., Diag. PI. Nov. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.
Mexic. 26. 1879. Type: MEXICO: Veracruz, 1947. Type: GUATEMALA: Izabal, Cerro
Orizaba, Botteri 971 (Lectotype K, here San Gil, 300-900 m, 25 Dec 1941, J. A.
designated (photo PTBG); Isolectotypes P (2 Steyermark 41864 (Holotype F 1134224
sheets, photos MEXU, PTBG). Syntypes: (photos MEXU, PTBG); Isotype F (photos
MEXICO: Veracruz, Orizaba, Bourgeau 2518 MEXU, PTBG), US). Arachnothryx.
(K (photo PTBG), P (3 sheets; photos MEXU, Rondeletia manantlanensis Lorence, Novon 1:
PTBG)). Region d'Orizaba, Bourgeau 2622 145, fig. 4A, B. 1991. Type: MEXICO:
(K (photo PTBG), B (n.v.; fragment F), F, K Jalisco, Mun. de Autlan, Canada del Laurelito,
(photo PTBG), P (4 sheets; photos MEXU, Las Joyas, 1850 m, 7 Dec 1985, A. Vazquez
PTBG), US (2 sheets)). Orizaba, Botteri 474 3750 (Holotype WIS (photo PTBG); Isotype
(K (2 sheets [label on first states "coll. Salle"; IBUG, n.v.).
second states "Sumichrast coll."] photos Rondeletia megalantha Lorence, Novon 4:
PTBG)), P (photos MEXU, PTBG). Orizaba, Uljig. 8. 1994. Type: HONDURAS:
Botteri 1951 (K, n.v., P (2 sheets; photos Cuyamel, 23 Aug 1924, M. A. Carleton 653
MEXU, PTBG). Orizaba, Muller 969 (K, n.v., (Holotype US 1208409; Isotype US
P (photos MEXU, PTBG). Orizaba, Muller 1208410).
1308 (K, n.v.). Botteri 971 at K is selected as Rondeletia mexiae Standi., Publ. Field Mus.
lectotype because it has three leafy twigs with Nat. Hist., Bot. Ser. 22: 55. 1940. Type:
an abundance of flowers at anthesis, and three MEXICO: Oaxaca, Distr. Choapan, Santa
duplicates exist at P. Rogiera. Maria, Montana Santa Maria, 1500 m, 1 Apr
Rondeletia linguiformis Hemsl., Diag. PL Nov. 1938, Y. Mexia 9261 (Holotype F 982856
Mexic. 29. 1879. Type: GUATEMALA: (photos MEXU, PTBG); Isotypes GH, n.v.,
Without precise locality, Skinner s.n. in 1857 MO (photos MEXU, PTBG), UC (photo
(Holotype K (photo PTBG)). Arachnothryx. PTBG), US (photos MEXU, PTBG)).
Rondeletia longipetiolata Lundell, Wrightia 5: Arachnothryx.—Rondeletia scabra Hemsl.
325. 1976. Type: MEXICO: Chiapas, Mt. Rondeletia mexicana (Turcz.) Standi., Publ.
Ovando, near Escuintla, 14 Nov 1945, E. Field Mus. Nat. Hist., Bot. Ser. 22: 388. 1940.
Matuda 6146 (Holotype TEX-LL; Isotypes Siphonandra mexicana Turcz., Bull. Soc Imp.
MEXU, TEX-LL). Arachnothryx. Naturalistes Moscou 21: 581. 1848. Type:
—Rondeletia buddleioides Benth. MEXICO: Oaxaca, without precise locality,
Rondeletia lundelliana Standi., Contr. Univ. 3000 ft., Oct, H. Galeotti 2664bis (Holotype
Michigan Herb. 4: 31. 1940. Type: BRITISH G-DEL, n.v. (photos F, MO, US); Isotype P
HONDURAS (BELIZE): El Cayo District, (photos MEXU, PTBG; fragment F, photos
Vaca, 21 Apr 1938, P. H. Gentle 2504 MEXU, PTBG)).
(Holotype F 941520 (photos MEXU, PTBG); Rondeletia microdon DC, Prodr. 4: 408. 1830.
Isotypes A, n.v., K (photo PTBG), TEX-LL Type: CUBA: Havana, de la Ossa s.n.
ex MICH (photos MEXU, PTBG), US (2 (Holotype G-DC, n.v. (microfiche IDC
sheets; photos MEXU, PTBG)). 800.691:111.7,8)).—Psychotria r
(DC.) Urb.
 i NOMENCLATOR OF MEXICAN AND CENTRAL

Rondeletia minor Lundell, Wrightia 5: 326. lectotype, because it has abundant flowers,
1976. Type: MEXICO: Veracruz, about 10 and duplicates exist in two other herbaria.
miles north of Teziutlan, 6 Jan 1964, D. S. Arachnothryx.
Correll & H. B. Correll 28736 (Holotype Rondeletia oaxacana Standi., Field Mus. Nat.
TEX-LL (photos MEXU, PTBG); Isotype Hist., Bot. Ser. 11:255. 1936. Type:
MEXU). Arachnothryx.—Rondeletia MEXICO: Oaxaca, without precise locality,
gracilispica Standi. 3000 ft., Oct, H. Galeotti 2664bis (Holotype P
Rondeletia monteverdensis Lorence, Novon 1: (photos MEXU, PTBG), fragment F (photos
147, fig. 5A, B. 1991. Type: COSTA RICA: MEXU, PTBG)). Norn, superfl. for and based
Puntarenas, Monteverde, road to TV tower, on same type as Rondeletia mexicana (Turcz.)
1700 m, 24 Aug 1985, W. A. Haber 2340 Standi. Arachnothryx.—Rondeletia mexicana
(Holotype MO 3595758 (photo PTBG)). (Turcz.) Standi.
Rondeletia myriantha Standi. & Steyerm., Rondeletia odorata Jacq., Enum. PI. 16. 1760.
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 288. Type: CUBA: Havanna, coastal thickets,
1940. var. myriantha. Type: GUATEMALA: Jacquin s.n. (location unknown). Cultivated in
Suchitepequez, Finca Moca, 3200 ft., 31 Oct
1934, A. F. Skutch 1569 (Holotype F 813463 Rondeletia odorata Jacq. var. breviflora Hook,
(photos MEXU, PTBG); Isotypes A, n.v., US f, Curtis Bot. Mag. 104, tab. 6350. 1878.
(photos MEXU, PTBG)). Arachnothryx. Type: CUBA: Collector and locality unknown
Rondeletia myriantha Standi. & Steyerm. var. (presumably K, n.v.). Reported by Kirkhnde
armentalis L. O. Williams, Phytologia 26: (1969) to occur in Panama.—Rondeletia
127. 1973. Type: GUATEMALA: Quiche, odorata Jacq.
Nebaj, 5600 ft., 22 Nov 1934, A. F. Skutch Rondeletia ovandensis Lundell. Wrightia .v
1776 (Holotype F 815811 (photos MEXU, 326. 1976. Type: MEXICO: Chiapas, Mt.
PTBG)). Arachnothryx. Ovando, near Escuintla, 24 Oct 1941, E.
Rondeletia nebulosa Standi., Trop. Woods 37: Matuda 6067 (Holotype TEX-LL (photos
32. 1934. Type: HONDURAS: Tegucigalpa, MEXU, PTBG); Isotypes F (photo PTBG),
La Aurora Trail, Rosario, San Juancito, 5400 MEXU, TEX-LL, n.v.). Arachnothryx.
ft., 2 Apr 1932, J. B. Edwards PI0 (Holotype Rondeletia panamensis DC, Prodr. 4: 408.
F 662646 (photos MEXU, PTBG); Isotype A, 1830. Type: PANAMA: Without precise
n.v.). Arachnothryx. locality, Haenke s.n. (Holotype PR, n.v.,
Rondeletia nicaraguensis Oerst., Vidensk. fragment G-DC, n.v. (microfiche 4: 408, 8)).
Meddel. Dansk Naturhist. Foren. Kjobenhavn Rondeletia pansamalana Standi., N. Amer. Fl.
1852: 43. 1853. Type: NICARAGUA: 32- 58. 1918. Type: GUATEMALA: Alta
Matagalpa, Segovia, in monte Pantasmo, Jan Verapaz, Pansamala, 3800 ft., Apr 1886, H.
1848, Oersted 11825 (Holotype C, n.v. von Tuerckheim 897 (Holotype US 49985
(photos F, MEXU, PTBG); Isotype GH, n.v.).
(photos F, MO, US; fragment US)).
Arachnothryx. Arachnothryx.
Rondeletia phvllocephala Standi. & L. O.
Rondeletia nitida Hemsl., Diag. PI. Nov. Williams, Ceiba 3: 63. 1952. Type:
Mexic. 29. 1879. Type: MEXICO: Chiapas, fl.
HONDURAS: Cortes, faldas humedas de
rosatres, forets des pins, en fevr, Linden 1660
Montana Sta. Ana, matorrales humedos de
(Lectotype K, here designated (photo PTBG): Rio Sta. Ana, 210 m. 6 Dec 1950, A. Molina
Isolectotypes G-DEL (n.v.; fragment F), P R. 3608 (Holotype US 2216049 (photos
(unnumbered Linden collection, almost MEXU, PTBG); Isotypes F (phot<~5 MEXU,
**cv
certainly a duplicate; photos MEXU, PTBG)). PTBG, US), MO)—Rondeletia t
Syntype: MEXICO: Chiapas, Ghiesbreght 728
(K (photo PTBG), GH, n.v.). Arachnothryx. Donn. Sm.
The Linden 1660 collection at K is designated
Rondeletia pittieri K. Schum. & K. Krause ex K. Rondeletia rekoi Standi., Proc. Biol. Soc.
Krause, Bot. Jahrb. Syst. 40: 316. 1908. Type: Wash. 8: 126. 1918. Type: MEXICO: Oaxaca,
COSTA RICA: Bords du Rio Poros, 2000 m, Las Pilas (Cerro Espino), 400 m, 10 Oct 1917,
10 Jan 1890, H. Pittier & T Duranci 1729 B. P. Reko 3490 (Holotype US 867145
(Holotype B, presumably destroyed (photos F, (photos MEXU, PTBG)). Arachnothryx.
MO); Isotype US (photos MEXU, PTBG)). Rondeletia repens L., Syst., ed. 10, 2: 928.
Rogiera.—Rondeletia amoena (Planch.) 1759. Type: JAMAICA: Sloanes.n.
Hemsl. (Lectotype: Sloane herb, iv.l 11, n.v.;
Rondeletia platysepala Standi., Ann. Missouri designated by Howard, 1989).—Geophila
Bot. Gard. 27: 343. 1940. Type: PANAMA: repens (L.) I. M. Johnst.
Code, El Valle, North Rim, 600-1000 m, 14 Rondeletia ricoi Lorence, Novon 1: 152, fig.
May 1939, P. H. Allen 1791 (Holotype F 8E, F, 9A, B. 1991. Type: MEXICO: Oaxaca,
989484 (photo PTBG); Isotypes F (photos Distr. de Putla de Guerrero, a 10 km al NE de
MEXU, PTBG), G, n.v., GH, n.v., MO Putla, 950 m, 9 Dec 1982, L. Rico A., E. M.
(photos MEXU, PTBG), NY, n.v.). Martinez & O. Tellez V. 566 (Holotype
Arachnothryx.—Rondeletia bertieroides MEXU (photo PTBG); Isotypes BM, F, MO,
Standi. PTBG).
Rondeletia polycephala Standi., J. Wash. Acad. Rondeletia roezlii (Planch.) Hemsl., Diag. PI.
Sci. 17: 337. 1927. Type: MEXICO: Oaxaca, Nov. Mexic. 25. 1879. Rogiera roezlii
without precise locality, F. Liebmann 11826 Planch., Fl. Serr. Jard. 5: tab. 442. 1849.
(Holotype C, n.v. (fragment US; photos Type: Based on cultivated material grown in
MEXU, PTBG)). Arachnothryx.—Rondeletia Belgium, probably of Guatemalan origin (type
mexicana (Turcz.) Standi. not designated).—Rondeletia amoena
Rondeletia povedae Lorence in W. C. Burger (Planch.) Hemsl.
& C. M. Taylor, Fieldiana Bot., n.s. 33: 299. Rondeletia rothschuhii Loes., Engl. Bot. Jahrb.
1993. Type: COSTA RICA: San Jose, Santa 60: 369. 1926. Type: NICARAGUA:
Rosa de Puriscal, 17 Jan 1973, L. J. Poveda Matagalpa, Canada Yucul, 1000 m, Rothschuh
441 (Holotype CR 51260; Isotype MO 592 (B, presumably destroyed (photos F, MO;
2211249). fragment F)).—Rondeletia buddleioides
Rondeletia pringlei Lorence, Novon 7: 54, fig. Benth.
5. 1997. Type: MEXICO: Veracruz, hills near Rondeletia rubens L. O. Williams, Phytologia
Orizaba, 5000 ft., 9 Feb 1895, C G. Pringle 26: 128. 1973. Type: GUATEMALA: Quiche,
6135 (Holotype GH; Isotypes K, MEXU, MO, Nebaj, 6000 ft., 19 Nov 1934, A. F. Skutch
WU). 1725 (Holotype F 815804 (photos MEXU,
Rondeletia purpurea Lorence, Novon 1: 148, PTBG)). Arachnothryx.
fig. 6A-F, 7A, B. 1991. Type: MEXICO: Rondeletia rufescens B. L. Rob., Proc. Amer.
Oaxaca, Distr. de Ixtlan, Sierra de Juarez, Acad. Arts 45: 402. 1910. Type:
Ruta 175 Tuxtepec a Oaxaca, a 5 km al N. de GUATEMALA: Alta Verapaz, Coban, 4800
Vista Hermosa, 1175 m, 14 Apr 1982, D. H. ft., Mar 1881,//. von Tuerckheim 582
Lorence & R. Torres C. 4009 (Holotype (Holotype GH, n.v.; Isotypes F, K (photo
MEXU (photo PTBG); Isotypes BM, F, MO, PTBG), P (photos MEXU, PTBG), US (2
PTBG). sheets; photos MEXU, PTBG)).
Rondeletia pyramidalis Lundell, Wrightia 5: Arachnothryx.—Rondeletia gracilis Hemsl.
327. 1976. Type: MEXICO: Chiapas, San Rondeletia rufescens B. L. Rob. var. ovata B. L.
Luis, near Siltepec, 1300 m, 24 Jan 1945, E. Rob., Proc. Amer. Acad. Arts 45: 403. 1910.
Matuda 5292 (Holotype TEX-LL (photos Type: GUATEMALA: Alta Verapaz, Tactic,
MEXU, PTBG); Isotypes F, MEXU, TEX-LL, 1800 m, Mar 1903, H. von Tuerckheim 8401
n.v.). Arachnothryx. (Holotype GH, n.v.; Isotypes F (2 sheets), US
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL

(photos MEXU, PTBG)).—Rondeletia Rondeletia s

gracilis Hemsl. Herb. 18: 1
Rondeletia rugosa Benth. in Oerst., Vidensk. Bias, forests around Puerto Obaldia, San Bias
Meddel. Dansk Naturhist. Foren. Kjobenhavn Coast, 0-50 m, Aug 1911, H. Pittier 4279
1852: 43. 1853. Type: COSTA RICA: (Holotype US 679392 (photos F, MEXU,
Cartago, 5600 ft., Oersteds.n. [=Plantae PTBG); Isotype NY, n.v.). Arachnothryx.
I Oersted 11828] (Holotype Rondeletia secundiflora B. L. Rob., Proc.
; Isotype US (photo PTBG)). Amer. Acad. Arts 45: 403. 1910. Type:
a (Planch.) Hemsl. GUATEMALA: In woods along road from
Rondeletia rzedovvskii Lorence, Novon 1: 154, Patin to Escuintla, 21 Jul 1860, S Hayes s.n
fig. 10. 1991. Type: MEXICO: Guerrero, (Holotype GH, n.v.; Isotype K (photo
Mun. de Atoyac, Puerto de la Piedra PTBG)). Arachnothryx.
Acanalada, 1930 m, 6 Jan 1984, Laboratorio Rondeletia seleriana Loes., Verh. Bot. Ver.
de Biogeografia 867 (Holotype MEXU Brandenb. 65: 105. 1923. Type:
364217 (photo PTBG); Isotypes FCME, GUATEMALA: Huehuetenango, Nenton
MEXU (photo PTBG)). Distr., near Chacula, E. Seler & C. Seler 3015
Rondeletia salicifolia Dwyer & Hayden, (Holotype B, presumably destroyed (photos F,
Phytologia 15: 58. 1967. Type: PANAMA: MO, US), fragment F). Rogiera.—Rondeletia
Panama, Cerro Jefe in Clusia forest at gratissima (Planch. & Limit.) Hemsl.
2700-3000 ft., 27 Jan 1966, E. L. Tyson, J. D. Rondeletia septicidalis B. L. Rob., Proc. Amer.
Dwyer & K. Blum 3319 (Holotype MO Acad. Arts 45: 403. 1910. Type: MEXICO:
1824503 (photo PTBG); Isotypes F, US Chiapas, Chicharras, 3000-6000 ft., 6 Feb
(photos MEXU, MO)). Arachnothryx. 1896, E. W. Nelson 3755 (Holotype US
^Rondeletia bertieroides Standi. 256722 (photos MEXU, PTBG); Isotype GH,
Rondeletia salicifolia Dwyer & Hayden subsp. n.v.). Arachnothryx.
brevicorolla J. H. Kirkbr., Ann. Missouri Bot. Rondeletia silvicola L. O. Williams, Phytologia
Card. 55: 383. 1968. Type: PANAMA: Bocas 26: 128. 1973. Type: HONDURAS: Atlantida,
del Toro, cloud forest above Quebrada Huron Lancetilla Valley, near Tela, 20-600 m, 6 Dec
on Cerro Bonyik, 166-400 m, 13 Apr 1968, 1927-20 Mar 1928, P. C. Standley 56885
/ H. Kirkbride, Jr. & J. A. Duke 597 (Holotype F 581955 (photos MEXU, PTBG)).
(Holotype MO 1946832 (photo PTBG); Arachnothnx.
Isotypes NY, n.v., US (2 sheets; photos Rondeletia skutchii Standi. & Steyerm., Publ.
MEXU, PTBG)). Arachnothryx.—Rondeletia Field Mus. Nat. Hist., Bot. Ser. 22: 289. 1940.
bertieroides Standi. Type: GUATEMALA: Suchitepequez, Finca
Rondeletia scabra Hemsl., Diag. PI. Nov. Moca, 4200 ft., 10 Jan 1935, A. F. Skutch
Mexic. 29. 1879. Type: MEXICO: Oaxaca, 2110 (Holotype F 934088 (photos MEXU,
Sierra San Pedro Nolasco, Talea etc., C. PTBG); Isotypes A, n.v., US (photos MEXU,
Jurgensen 926 (Holotype K (photo PTBG)). PTBG)). Arachnothryx.
Ri
tlti chumanniana K. Krause, Bot. Rondeletia spinosa K. Schum. ex Loes., Bull.
Jahrb. 40: 315. 1908. Type: PANAMA: Herb. Boissier 3: 620. 1895. Type: MEXICO:
Veraguas, Mar 1906, J. von Warszewicz 1168 Hidalgo, prope Huejutla, Apr, E. Seler & C.
(B, presumably destroyed (photo F; fragment Seler668 (Holotype B, presumably
F)).—Rondeletia amoena (Planch.) Hemsl. destroyed).—Status uncertain, possibly
Rondeletia scoti Lorence, Novon 1: 155, fig. Randia sp. The protologue diagnosis states
8A. B. 1991. Type: MEXICO: Oaxaca, La branches armed with paired spines and
Glorieta-Rio Grande (N slope), 12 Mar 1953, flowers solitary or fascicled.
a stachvoidea Dorm. Sm.. Bot. Gaz.
T. MacDougall s.n. (Holotype MEXU (photo
sville) 42: 298. 1906. Type:
PTBG); Isotypes MO, PTBG).
 GUATEMALA: Alta Verapaz,! Isotypes F (photos MEXU, PTBG), MO, US
600-900 m, 25 Apr 1905, R. Hay s.n. (photos MEXU, PTBG)). Rogiera.
(Holotype US 473726 (photos MEXU, Rondeletia tacanensis Lundell, Bull. Torrey
PTBG)). Arachnothryx. Bot. Club 66: 603. 1939. Type: MEXICO:
Rondeletia standleyana A. R. Molina, Ceiba 1: Chiapas, Volcan Tacana West, 2800 m, 30
262. 1951. Type: HONDURAS: Morazan, in Mar 1939, E. Matuda 2928 (Holotype TEX-
pine forest above Zambrano, 1200 m, 20 Jul LL (photos MEXU, PTBG); Isotypes A, n.v.,
1949, L. O. Williams & A. Molina R. 14417 F (photos MEXU, PTBG), GH, n.v., K (photo
(Holotype US 2216018 (photos MEXU, PTBG), MEXU, US (2 sheets; photos MEXU,
PTBG); Isotypes F (photos MEXU, PTBG, PTBG)). Arachnothryx.
US), US). Arachnothryx. Rondeletia taylorae Lorence in W. C. Burger
Rondeletia stenosiphon Hemsl., Diag. PI. Nov. & C. M. Taylor, Fieldiana, Bot., n.s. 33: 300.
Mexic. 26. 1879. Type: MEXICO: Without 1993. Type: COSTA RICA: Limon, Canon de
precise locality, Yucatan & Tabasco [sic], Limon, El Progreso. Fila Matama, Valle de la
E. P. Johnson s.n. (Holotype K (photo Estrella, 9°47T8"N, 83°08'45"W, 1350 m, 19
PTBG)). Rogiera. Apr 1989, G. Herrera & A. Chacon 2656
Rondeletia stenostachxa Standi., Publ. Field (Holotype CR; Isotypes MO, PTBG).
Mus. Nat. Hist., Bot. Ser. 18: 1372. 1938. Rondeletia tenorioi Lorence, Novon 1: 156, fig.
Type: COSTA RICA: La Hondura de San 11 A, B. 1991. Type: MEXICO: Oaxaca, Distr.
Jose, 1300 m, 15 Aug 1933, M. Valerio 769 de Juxtlahuaca, 19 km al SW de Juxtlahuaca y
(Holotype F 681816 (photos MEXU, PTBG)). a 5 km en la misma direccion de Sta. Ma.
Arachnothryx.—Gonzalagunia stenostachya Asuncion, 1900 m, 4 Apr 1982, P. Tenorio &
(Standi.) W. C. Burger R. Torres C. 184 (Holotype MEXU (photo
Rondeletia strigosa (Benth.) Hemsl., Diag. PI. PTBG); Isotypes BM, F, MO).
Nov. Mexic. 27. 1879. , Bot. Gaz.
Benth., PI. Hartweg. 75. 1841. Type: (Crawfordsville) 42: 299. 1906. Type:
GUATEMALA: [Volcan] Zunil, Nov, T. HONDURAS: Santa Barbara, Rio
Hartweg 530 (Lectotype K, here designated Chamelecon, 400 m, 16 Dec 1888, C. Thieme
(photo PTBG); Isolectotypes G-DEL, n.v. 5276 (Holotype US 941403 (photos MEXU,
(fragment F), P (photos F, MEXU, MO, PTBG)). Arachnothryx.
PTBG), W, n.v. (photos F, MO)). Syntype: Rondeletia torresii Standi., J. Wash. Acad. Sci.
GUATEMALA: Volcan de Fuego, 18: 163. 1928. Type: COSTA RICA: Alajuela,
6000-6400 ft., Salvia s.n. (K (photo PTBG; Viento Fresco, 1600-1900 m, 13 Feb 1926,
fragment F)). Rogiera. The K collection of P. C. Standley & R. Torres R. 47839
Hartweg 530 is a good, representative choice (Holotype US 1305295 (photos MEXU,
for lectotype and in addition has duplicates in PTBG)). Arachnothryx.
three herbaria. Rondeletia tuxtlensis Lorence & Cast.-
Rondeletia subscandens Lundell, Wnghtia 5: Campos, Biotica 13: 148, fig. 1. 1988. Type:
328. 1976. Type: GUATEMALA: Peten, La MEXICO: Veracruz, Mun. Mecayapan,
Cumbre, about 500 m west of km 142/143 of Volcan San Martin Pajapan al sur del Ejido La
the Cadenas Road, 10 Sep 1976, C. L. Lundell Valentina, 700 m, 8 May 1985, J. I. Calzada
& E. Contreras 19830 (Holotype TEX-LL 11183 (Holotype XAL; Isotypes F, MEXU,
(photos MEXU, PTBG); Isotypes F (2 sheets; MO (photo PTBG)). Arachnothryx.
photos MO), K (photo PTBG)). Rogiera. Rondeletia urophylla Standi. & L. O.
Rondeletia suffrutescens Brandegee, Univ. Williams, Phytologia 26: 129. 1973. Type:
Calif Publ. Bot. 6: 70. 1914. Type: MEXICO: COSTA RICA: Puntarenas, Esquinas forest,
Chiapas, Cerro del Boqueron, Aug 1913, C. A. 30 m, 3 Aug 1951, P. H. Allen 6291
Purpus 6704 (Holotype UC 172451, n.v.;
^ NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEA

(Holotype F 1562934 (photos MEXU, Jesus and Calahuache, 1300-1500 m. 31 Dec

PTBG); Isotype EAP, n.v.). 1939, J A. Steyermark 33220 (Holotype F
Rondeletia uxpanapensis Lorence & Cast.- 1041786 (photos MEXU, PTBG)).
Campos, Biotica 13: 150, fig. 2. 1988. Type: Arachnothn v.—Rondeletia secundiflora
MEXICO: Veracruz, Mun. Hidalgotitlan, zona B. L. Rob.
de Uxpanapa; Rio Solosuchil a 5-6 km ESE Rondeletia wendtii Lorence & Cast-Campos,
de Hermanos Cedillo, 100-200 m, 24 Mar Biotica 13: 154, fig. 3. 1988. Type: MEXICO:
1982, D. H. Lorence et ah 3909 (Holotype Oaxaca, Distr. Juchitan, Mun. de Santa Maria
MEXU (photo PTBG); Isotypes BM, ENCB, Chimalapa, Arroyo Chocolin, Poblacion de
CHAPA, MO, NY, PTBG, UC, XAL). Nicolas Bravo, cerca del Rancho de Agustin
Arachnothryx. Montero, 3-A km al sur de la Poblacion de
Rondeletia versicolor J. E. Sm., Bot. Mag. 7: Rio Alegre (Veracruz), 150 m, 21 Oct 1983,
tab. 4579. 1851. Type: Based on cultivated
material grown at Kew, sent by Seemann from (Holotype MEXU (photo PTBG); Isotypes
Boquete, Veraguas, Panama (type not CHAPA, PTBG). Arachnothryx.
designated). Rogiera.—Rondeletia amoena Rondeletia vueatanensis Lundell, Wnghtia 5:
(Planch.) Hemsl. 329 1976. Type: MEXICO: Yucatan,
Rondeletia villosa Hemsl., Diag. PI. Nov. Chichen Itza, 28 Apr 1950,/ T. Balthvin. Jr
Mexic. 27. 1879. Type: MEXICO: Oaxaca, 14230 (Holotype TEX-LL). Arachnothryx.
San Pedro Nolasco, etc., Jurgensen 806 Note: the type was likely collected from a
(Lectotype K, here designated (photo PTBG); cultivated plant outside its normal range, as
this species is commonly cultivated in many
Isolectotypes G-DEL, n.v. (fragment F), K
parts of Mexico for its nocturnally fragrant
(photo PTBG), P (photos MEXU, PTBG)).
flowers—Rondeletia leucophylla Kunth in
Syntypes: MEXICO: Oaxaca, Trapiche de
Concepcion, F. Liebmann 70 (K (photo Humb., Bonpl. & Kunth.
Rondeletia zollenana Standi. & Steyerm., Publ.
PTBG), G-DEL, n.v. (fragment F)). Tabasco, Field Mus. Nat. Hist., Bot. Ser. 22: 292. 1940.
arbuste a fleurs blanches en novembre et
Type: GUATEMALA: San Marcos, between
Xbre., environs de Teapa (Tabasco), 1839,
Finca El Porvemr and Loma Corona, 9 mi.
Linden 145 (K (photo PTBG); probable
NW of El Porvemr. southwest-facing slopes
syntypes G-DEL, n.v. (fragment F), P (photos
of Volcan Tajumulco, 1300-2000 m. 14 Mar
MEXU, PTBG)). Arachnothryx- The K sheet
1940, J. A. Steyermark 37738 (Holotype F
of Jurgensen 806 is designated as lectotype,
1039668 (photos MEXU. PTBG I).
because it is representative and has flowers
Arachnothnx. Rondeletia brandegeeana
and fruits, with duplicates in three other

Rondeletia volubilis Sesse & Moc., PI. Nov.

Rosenbereiodendron FagerL Svensk Bot.
Hisp. 36. 1887. Type: MEXICO: Habit in
Tidskrift.42: 150. 1948. Type:
Acahuizotla, Sesse & Mocino 1393 (Holotype
Rosenbergiodendron formosum (Jacq.)
MA, n.v. (photo WU); Isotype F, fragment,
n.v.). Caique of Fl. Mexic. Ic. 275 (G-DC. Fagerl.
Rosenbergiodendron formosum, (Jacq.)
n.v. (photo MO)). For typification, see Puff
Faeerl Svensk Bot. Tidsknft. 42: 152. 1948.
(1991: 326).—Paederia ciliata (Bartl. ex A/^Wa/o^^Jacq..Enum.PlCanb.
DC.) Standi. 16 1760. Tvpe. COLOMBIA: Bolivar.
Rondeletia vulcanicola Standi. & Steyerm.. Cartagena. Jacquin sn. (Holotype G. n .v.).
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 291.
1940. Type: GUATEMALA: Quezaltenango. i. Type: Rubia
Finca Pireneos, lower south-facing slopes of
Volcan Santa Maria, between Santa Maria de
 xta \l Martens & Galeotti, Bull. San Felix, 29 km above bridge over Rio San
Acad. Roy. Sci. Bruxelles 11: 127. 1844. Felix (7.9 km above tumoff to Escopeta),
Type: MEXICO: Puebla, Tehuacan, 5000 ft., 1500 m, 14 Jul 1976, T. B. Croat 37071
H. Galeotti 2362 (Lectotype K, n.v. (photo (Holotype MO 2798805 (photo PTBG);
MO); designated by Dempster, 1973b). Isotype MO).—Psychotria nebulosa (Dwyer)
—Galium aschenbornii S. Schauer C M. Taylor
Rubia dipkylla Sesse & Moc., Fl. Mexic. 23. Rudgea cornifolia (Humb. & Bonpl. ex Roem.
1893. Type: Habitat in anfractibus Santi & Schult.) Standi., Publ. Field Columbian
Angeli et Hortis Sancti Augustini. Floret Julio, Mus., Bot. Ser. 7: 432. 1931. Psychotria
Sesse' <£ Mocirio s.n. (presumably MA, n.v.). cornifolia Humb. & Bonpl. ex Roem. &
—Status uncertain, probably Didymaea sp. Schult., Syst. Veg. 5: 191. 1819; Kunth in
Rubia hypocarpia (L.) DC, Prodr. 4: 591. 1830. Humb., Bonpl. & Kunth, Nov. Gen. Sp. PI. 3:
Valantia hypocarpia L., Syst. Nat., ed. 10, 362. 1819. Type: VENEZUELA: Near
1307. 1759. Type: JAMAICA: Rubia I, Maipures, on the Orinoco, Humboldt &
Browne, Hist. Jamaica 141 (Holotype LINN, Bonpland 893 (Holotype B-WI, n.v. (photo
MO)).
(L.) Endl. ex Griseb. Rudgea coronicarpa Dwyer, Ann. Missouri
Rubia incana Kunth in Humb., Bonpl. & Kunth, Bot. Gard. 67: 478. 1980. Type: PANAMA:
Nov. Gen. Sp. 3: 338. 1820. Type: MEXICO: Code, Cerro Pilon, 2700 ft., 9 Jan 1972, A.
Without precise locality, Humboldt & Gentry & J. D. Dwyer 3647 (Holotype MO
Bonpland s.n. (Holotype presumably B, n.v.; 2176970 (photo PTBG); Isotype MO).
Isotype P, n.v. (photo MO)).—Galium Rudgea fimbriata (Benth.) Standi, in Standi. &
hypocarpium (L.) Endl. ex Griseb. Calderon, Lista Prelim. PL El Salvador 274.
Rubia laevigata DC, Prodr. 4: 591. 1830. Type: 1925. Psychotria fimbriata Benth., J. Bot.
MEXICO: Without precise locality, Haenke Hook. 3: 226. 1841. Type: GUYANA:
s.n. (Holotype presumably PR, n.v.). Essequibo River, Schomburgk 51 (Holotype
Relbumum.—Galium microphyllum A. Gray presumably K, n.v).—Rudgea cornifolia
(Kunth) Standi.
Rudgea Salisb., Trans. Linn. Soc. London 8: Rudgea hemisphaerica Dwyer ex C. M.
327. 1807. Type: Rudgea lanceaefolia Salisb. Taylor, Novon 6: 102, fig. 3A, B. 1996. Type:
Rudgea amplexicaulis Dwyer, Ann. Missouri PANAMA: El Llano-Carti Road, 19.1 km
Bot. Gard. 67: 475. 1980. Type: PANAMA: from Interamerican Highway, 9°19'N,
Chiriqui, above San Felix along mining road 78°55'W, 350 m, 5 Mar 1985, G. de Nevers,
18-27 mi. off Pan-Am. Highway (above H. Herrera & S. Charnley 4962 (Holotype
Chame or tumoff to Escopeta), 1200-1500 m, PMA 38235, n.v.; Isotype MO 4658113).
12 Mar 1976, T. B. Croat 33133 (Holotype Rudgea horquetensis Dwyer & Hayden, Ann.
MO 2399080 (photo PTBG); Isotype MO). Missouri Bot. Gard. 54: 145. 1967. Type:
Rudgea ceratopetala Donn. Sm., Bot. Gaz. PANAMA: Chiriqui, Cerro Horqueta, 6500
(Crawfordsville) 35: 3. 1903. Type: i Hagen & von Hagen 2156 (Holotype
GUATEMALA: Alta Verapaz, Cubliquitz, NY,:
340 m. 1901, H. von Tuerckheim 7904 Rudgea isthmensis Standi, Ann. Missouri Bot.
(Holotype US 943480 (photos MEXU, Gard. 26: 313. 1929. Type: PANAMA: Canal
PTBG); Isotypes F, n.v., US (2 sheets; photos Zone, vicinity of Salamanca Hydrographic
MEXU, PTBG)).—Rudgea cornifolia Station, Rio Pequeni, 80 m, 28-29 Jul 1938,
(Kunth) Standi. R. E. Woodson, Jr., P. H. Allen & R. J- Seibert
Rudgea chiriquiensis Dwyer, Ann. Missouri 1618 (Holotype MO 1169548 (photo PTBG);
Bot. Gard. 67: 476. 1980. Type: PANAMA: Isotypes A, n.v., NY, n.v., US (photos
Chiriqui, Cerro Colorado, along road above MEXU, PTBG)).
 ' MEXICAN AND CENTRAL J

Rudgea laevis C. M. Taylor, Novon 5: 207, fig. Rudgea raveniana W. C. Burger in W. C.

IF, G. 1995. Type: COSTA RICA: Burger & C. M. Taylor, Fieldiana, Bot., n.s.
Puntarenas, Monteverde, Bajos de La Lindora, 33: 304. 1993. Type: COSTA RICA: Rincon
Rio Guacimal, 10°18'N, 84°48'W, 19 Feb de Osa, W. Burger & R. Liesner 7251
1988, W. Haber & E. Bello 8271 (Holotype (Holotype CR, n.v.; Isotype F, n.v.).
CR, n.v.; Isotype MO no 3646480). Rudgea reducticalvx Dwyer in W. C. Burger &
Rudgea magna Standi., Publ. Field Mus. Nat. C. M. Taylor, Fieldiana, Bot., n.s. 33: 304.
Hist., Bot. Ser. 22: 56. 1940. Type: 1993. Type: COSTA RICA: Alajuela, along
PANAMA: San Bias Coast, hills of Sperdi, road between Canas and Upala, 4 km NNE of
near Puerto Obaldia, 20-200 m, Sep 1911, H. Bijagua on slopes leading into Rio Zapote,
Pittier 4350 (Holotype F 600258 (photo 400 m, 24 Jun 1976, T. B. Croat 36330
PTBG)). (Holotype CR, n.v.; Isotype MO 2481680
Rudgea mandevilliifolia Dwyer ex C. M. (photo PTBG)).
Taylor, Novon 6: 304, fig. 3Q D. 1996. Type: Rudgea sanblasensis C. M. Taylor, Novon 6:
PANAMA: Chiriqui, vicinity of Planes de 220. 1996. Type: PANAMA: San Bias. Rio
Hornito beyond Gualaca, ca. 23 km E of Finca Playon Chico, Dukandi, vecinidad de la Isla
Linares, 1400-1900 m, 28 Nov 1979, T. B. Playon Chico, 09° 15'N, 78° 15'W, 30 m, 6
Croat 48863 (Holotype MO 3615290). Sep 1993, H. Herrera et al 1358 (Holotype
Rudgea mcphersonii Dwyer ex C. M. Taylor, MO 4671394; Isotype PMA, n.v.).
Novon 6: 304, fig. 3E, F. 1996. Type: Rudgea .simiarum Standi. & Steyerm., Publ.
PANAMA: Colon, Santa Rita ridge, SE of Field Mus. Nat. Hist., Bot. Ser. 22: 389. 1940.
Colon, 10-11 road-miles from trans-isthmian Type: GUATEMALA: Izabal. between
highway, 9°25'N, 79°40'W, 450 m, 18 Sep Virginia and Lago de Izabal, Montana del
1987, G. McPherson 11771 (Holotype PMA, Mico, 50-500 m, 5 Apr 1940, J. A.
n.v.; Isotype MO 3646497). Stevermark 38839 (Holotype F 1042936
Rudgea monofructus Gomez-Laur. & Dwyer, (photo PTBG)).—Psychotria cf. eurycarpa
Novon 1: 50, fig. I. 1991. Type: COSTA Standi.
RICA: Alajuela, Reserva Forestal de San Rudgea skutchii Standi., Publ. Field Mus. Nat.
Ramon, Campamento Rio San Lorencito, Hist, Bot. Ser. 18: 1374. 1938. Type: COSTA
900-1100 m, 25 Sep 1989, J. Gomez-launto RICA: San Jose, vicinity of El General, 915
& G. Vargas 11822 (Holotype UCR. n.v.: m, Aug 1936, A. F. Shack 2836 (Holotype
Isotype MO). US 1642550 (photos MEXU. PTBG);
Rudgea panamensis (Dwyer) C. M. Taylor, Isotypes GH, n.v., MO (photos MEXU,
Novon 6: 305. 1996. Cephaelispanamensis PTBG)).
Dwyer, Ann. Missouri Bot. Gard. 67: 77. Rudgea th\rsiflora Donn. Sm., Bot. Gaz.
1980. Type: PANAMA: Colon, Rio Guanche, (Crawfordsville) 61: 375. 1916. Type:
ca. 2.5 km upriver from bridge on road to COSTA RICA: Talamanca, Forets de Tsaki,
Portobelo, 10-100 m, 14 Dec 1974, S. Mori & 200 m, Apr 1895, A. Tonduz 9579 (Holotype
J. Kallunki 3714 (Holotype MO 2353037 US 943477 (photos MEXU, PTBG); Isotype
(photos MEXU, PTBG); Isotypes MO BR, n.v).—Psychotria stenostachya Standi.
2353038 (photos MEXU, PTBG), PMA Rudgea trifurcata Gomez-Laur.. Brenesia 33:
22701, n.v.). 139. 1990 (1991). Type: COSTA RICA:
Rudgea pittieri Standi., Publ. Field Mus. Nat. Limon, Talamanca, 3 km al oeste de Volio,
Hist., Bot. Ser. 22: 211. 1940. Type: 9°37,40N, 82°53'50W, 100 m../. Gomez-
PANAMA: Colon, Loma de la Gloria, near Laurito & G. Vargas 11957 (Holotype CR,
Fato (Nombre de Dios), 10-100 m, Jul-Aug n.v.; Isotypes F, n.v., MO, US, n.v.))
1911,//. Pittier 3855 (Holotype F 636043
(photo PTBG); Isotypes F, US).
Rustia Klotzsch in Hayne, Getr. Darstell. Gew. Buenos Aires, buissons, 300 m, Jan 1893, H.
14: sub tab. 14, 15. 1846. Lectotype: Rustia Pittier 6712 [on label "H. Pittier'7 is crossed
formosa (Cham. & Schltdl.) Klotzsch out and "Tonduz" is written in] (US 1393299
(=Exostenia formosum Cham. & Schltdl.). (photos MEXU, PTBG)). On the Savannah,
Rustia costaricensis (Standi.) Lorence in W. C. H. Pittier 4025 (BM, n.v.; Isosyntypes K, n.v.,
Burger & C. M. Taylor, Fieldiana, Bot, n.s. BR, n.v., G, n.v).—Sabicea panamensis
33: 306. 1993. Stomandra costaricensis Wernham
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. Sabicea flagenioides Wernham, Monogr.
23: 247. 1947. Type: COSTA RICA: San Sabicea 57. 1914. Type: MEXICO: Yucatan,
Jose, basin of El General, 675-900 m, Mar Chichankanab, G. F. Gaunter 1432 (Holotype
1940, A. F. Skutch 4777 (Holotype F 1044344 F, n.v.; Isotype B, presumably destroyed).
(photo PTBG); Isotypes MO (photo PTBG), —Status uncertain.
US). Sabicea hirsuta Kunth in Humb., Bonpl. &
Rustia dressleri Delprete, Novon 5: 137. 1995. Kunth var. adpressa Wernham, Monogr.
Type: PANAMA: Panama, El Llano-Carti Sabicea 55. 1914. Syntypes: PANAMA:
highway, 8-10 km N of El Llano, 31 Aug Isthmus of Panama, Chagres, Jan-Mar 1850,
1974, R. L. Dressier 4703 (Holotype MO A. Fendler 180 (F, (photo PTBG), BM, n.v.,
2268783). W, n.v., fragment F); same locality, Hayes s.n.
Rustia ferruginea Standi., Contr. U.S. Natl. (n.v.), same locality, Seemann 1073 (n.v.).
Herb. 18: 142. 1916. Type: PANAMA: Colon, COSTA RICA: Limon, Kuntze 1995 (n.v.);
along Rio Fato, 10-100 m, Aug 1911, M Punta Arena, Tonduz 6712 (n.v.), Tonduz
Pittier 4201 (Holotype US 679302 (photos 9955 (n.v.); six additional syntypes are cited
MEXU, PTBG)). from St. Thomas, Trinidad, and Brazil.
Rustia occidentalis (Benth.) Hemsl., Biol. Sabicea mexicana Wernham, Monogr. Sabicea
Centr.-Amer., Bot. 2: 14. 1881. Exostemma 41. 1914. Syntypes: MEXICO: Oaxaca,
occidental Benth., Bot. Voy. Sulphur 104. without precise locality, 3000 ft., H. Galeotti
1844. Type: PANAMA: Without precise 2662 (K, n.v. (photo MO), B, n.v., BR, n.v.,
locality, Barclay s.n. or Hinds s.n. (Holotype G, n.v., P, n.v., W, n.v.). Without precise
K, n.v.). locality, F. Liebmann 43 (K, n.v., BR, n.v.).
Rustia panamensis Dwyer, Ann. Missouri Bot. Sabicea panamensis Wernham, Monogr.
Gard. 67: 483. 1980. Type: PANAMA: Sabicea 30. 1914. Type: PANAMA: Chagres,
Veraguas, Valley of Rio Dos Bocas, 11 km Isthmus of Panama, Mar 1850, A. Fendler 181
from Escuela Agricola Alto Piedra (above (Holotype K, n.v.; Isotypes US, 2 sheets
Santa Fe), on road to Calovebora, 450 m, 30 (photos MEXU, PTBG)).
Aug 1974, T. B. Croat 27508 (Holotype MO Sabicea stellaris Dwyer, Ann. Missouri Bot.
2214602; Isotypes F (photo PTBG), GH, Gard. 67: 489. 1980. Type: PANAMA:
n.v).—Rustia costaricensis (Standi.) Panama, Rio Pequeni, 10-15 minutes
upstream from hydrographic station by motor,
11 Dec 1974, R. L. Dressier 4887 (Holotype
Sabicea Aubl., Hist. PI. Guiane 192. 1775. Type MO 2353169 (photo PTBG)).
not designated. Sabicea villosa Willd. ex Roem. & Schult.,
Sabicea ambigua Standi., Publ. Field Syst. Veg. 5: 265. 1819. var. villosa. Type:
Columbian Mus., Bot. Ser. 7: 49. 1930. Type: VENEZUELA: Orinoco River, A. Bonpland
COLOMBIA: Without precise locality, 1847, s.n. (Holotype presumably B-WI, n.v.).
Triana s.n. (Holotype K, n.v. (photo F)). Sabicea villosa Willd. ex Roem. & Schult. var.
—Amphidasya ambigua (Standi.) Standi. adpressa (Wernham) Standi., Publ. Field
Sabicea costaricensis Wernham, Monogr. Columbian Mus., Bot. Ser. 7: 52. 1930.
Sabicea 31. 1914. Syntypes: COSTA RICA: Syntypes: PANAMA: Isthmus of Panama,
 CLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACE

Chagres, Jan-Mar 1850, A. Fendler 180 (F, Sickingia calderoniana Standi., Publ. Field Mus.
(photo PTBG), BM, n.v., W, n.v., fragment Nat. Hist., Bot. Ser. 22: 214. 1940. Type: EL
F); same locality, Hayes s.n. (n.v.), same SALVADOR: Jayaque, finca del Dr. Zarate, 9
locality, Seemann 1073 (n.v.). COSTA RICA: Jan 1939, M. L. Van Severen 66 (Holotype F
Limon, Kuntze 1995 (n.v.); Punta Arena, 1033327 (photo PTBG); Isotype F).—Simira
Tonduz 6712 (n.v.), Tonduz 9955 (n.v.); six calderoniana (Standi.) Steyerm.
additional syntypes are cited from St. Thomas, f ) Standi., Trop. Woods
Trinidad, and Brazil. 22: 10. 1930. Calderonia klugei Standi., J.
Wash. Acad. Sci. 15:6. 1924. Type:
Schradera Vahl, Eclog. Amer. 1: 35. 1796, nom. PANAMA: Panama, near Chepo, 1924, H. C.
cons. Type: Schradera capitata Vahl, nom. Kluge 19 (Yale 7132) (Holotype US 1206424
illeg. {=Schradera invohtcrata (Sw.) K. (photos MEXU, PTBG); Isotype F (plus
Schum.) another probable isotype at F, a fruiting
Schradera blumii Dwyer & Hayden, collection from type tree, Kluge s.n.. also Yale
Phytologia 15: 59. 1967. Type: PANAMA: 7132)).—Simira klugei (Standi.) Steyerm.
Panama, Cerro Jefe in Clusia forest at Sickingia lancifoliu Lundell. Wrightia4: 50.
2700-3000 ft., 27 Jan 1966, E. L. Tyson, J. 1968. Type: GUATEMALA: Peten, Laguna
Dwyer & K. Blum 3218 (Holotype MO Petexbatun, in high forest bordering lake, on
1831023 (photo PTBG)). Escarbadero, 2 Apr 1964, C. L. Lundell 18224
Schradera costaricensis Dwyer in W. C. (Holotype TEX-LL, n.v.; Isotype US (photos
Burger & C. M. Taylor, Fieldiana, Bot., n.s. MEXU, PTBG)).—Simira salvadorensis
33: 309. 1993. Type: COSTA RICA: Alajuela, (Standi.) Steyerm.
Bijagua, G. Herrera 2004 (Holotype CR, n.v.; ', ;,. Lundell var. puberula
Isotypes F, n.v., MO). Lundell, Wrightia 4: 50. 1968. Type:
GUATEMALA: Peten, Lacandon, in bajo,
Serissa Comm. ex Juss., Gen. 209. 1789. Type: about 1 km, 700 m south. 12 Mar 1962, E.
Serissa foetida Comm. ex Juss. Contreras 3514 (Holotype TEX-LL, n.v.).
Serissa japonica (Thunb.) Thunb., Nov. Gen. —Simira salavadorensis (Standi.) Steyerm.
PI. 132, pi. 9. 1798. Type: JAPAN: Collector Sickingia maxonii (Standi.) Standi., Trop.
unknown (presumably UPS, n.v.). Lycium Woods 14: 30. 1928. Genipa maxonii Standi.,
japonicum Thunb., Nov. Act. Roy. Soc. J. Wash. Acad. Sci. 8: 642. 1918. Type:
Upsal. 3: 207. 1780. Cultivated at least in PANAMA: Panama, Canal Zone, forest along
Costa Rica and Panama. the Rio Indio de Gatiin, near sea level, 17 Feb
1911, W. R. Maxon 4848 (Holotype US
Sherardia L., Sp. PI. 1: 102. 1753. Type: 675223 (photos MEXU, PTBG)).—Simira
Sherardia arvensis L. maxonii (Standi.) Steyerm.
Sherardia arvensis L., Sp. PI. 1: 102. 1753. dlock in Hook. Icon., ser.
Type: Habitat in arvensis Scaniae, Germaniae, 4, 4: tab. 3321. 1936. Type: MEXICO:
Angliae, (Lectotype Sherard, Herb. Clifford Mexico, Distr. Temascaltepec, Luvianos, 26
33: Sherardia 1 (BM, n.v.); designated by Sep 1933, G. B. Hinton 4818 (Holotype K,
Natali in Jarvis et al., 1993: 88). Naturalized n.v.; Isotype US (photos MEXU, PTBG)).
at least in Veracruz and Costa Rica. —Simira mexicana (Bullock) Steyerm.
Sickingia mollis Lundell, Contr. Univ. Michigan
Sickingia Willd., Ges. Naturf. Freunde Berlin Herb. 4: 32. 1940. Not Sickingia mollis
Neue Schrift. 3: 445. 1801. Type not Standi., 1936. Type: MEXICO: Tabasco,
designated. [Herein treated as a synonym of Reforma, on the Rio San Pedro de Martir,
Simira Aubl.l above Balancan, 22-26 May 1939, E. Matuda
3161 (Holotype MICH, n.v.; Isotypes A, n.v.,
 TEX-LL (photos MEXU, PTBG), US (photo Siderodendron Cothen., Disp. 6. 1790.
PTBG)).—Simira salvadorensis (Standi.) (=Sideroxyloides Jacq., 1763.) Type:
Steyerm. Siderodendron ferreum (Jacq.) J. F. Gmel.
Sickingia multiflora Lundell, Wrightia 5: 258. [Herein treated as a synonym of Ixora L.]
1976. Type: GUATEMALA: Peten, Tikal Siderodendi Rich, in Sagra,
National Park, Tikal, in ramonal covering the Hist. Cuba 11: 24. 1850. Type: CUBA:
ruins, on Temple IV, 11 Jul 1960, E. Without precise locality, A. Richard s.n.
Contreras 1239 (Holotype TEX-LL (photos (Holotype presumably P, n.v.).—Ixora
MEXU, PTBG); Isotype US (photos MEXU, floribunda (A. Rich.) Griseb.
PTBG)).—Simira salvadorensis (Standi.)
Steyerm. Simira AubL, Hist. PL Guiane 1: 170, t. 65.
Sickingia myriantha Standi., Pub. Field 1775, not Simira Raf, 1836. Type: Simira
Columbian Mus., Bot. Ser. 7: 27. 1930. Type: tinctoria Aubl.
COLOMBIA: Magdalena, Las Nubes road, Simira calderoniana (Standi.) Steyerm., Mem.
region of Santa Marta, 1200 m, 3 Dec 1898, New York Bot. Gard. 23: 306. 1972. Sickingia
H. H. Smith 1810 (Holotype F 138660; calderoniana Standi., Publ. Field Mus. Nat.
Isotypes MO, NY, n.v., US, n.v., W, n.v., Y, Hist., Bot. Ser. 22: 214. 1940. Type: EL
n.v.). Holtonia, Simira.—Elaeagia SALVADOR: Jayaque, finca del Dr. Zarate, 9
myriantha (Standi.) C. M. Taylor & Hammel Jan 1939, M. L. Van Severen 66 (Holotype F
Sickingia rhodoclada (Standi.) Standi., Publ. 1033327 (photo PTBG)).
Field Mus. Nat. Hist., Bot. Ser. 22: 215. 1940. Simira daricncns > IKvyer, Ann. Missouri Bot.
Exandra rhodoclada Standi., J. Wash. Acad. Gard. 67: 493. 1980. Nom. superfl. for
Sci. 13: 292. 1923. Type: MEXICO: Oaxaca, Bathysa panamensis Dwyer, Ann. Missouri
between La Venta and Niltepec, 200 ft., 14 Jul Bot. Gard. 55: 34. 1968 (based on same type).
1895, E. W. Nelson 2796 (Holotype US Type: PANAMA: Darien, Rio Ucurganti, 7
229232 (photos MEXU, PTBG); Isotype F Jul 1967, N. Bristan 1187 (Holotype MO
(photos MEXU, PTBG)).—Simira 1968219 (photo PTBG); Isotype GH, n.v.).
rhodoclada (Standi.) Steyerm.
Sickingia salvadorensis (Standi.) Standi., Field Simira klugei (Standi.) Steyerm., Mem. New
Mus. Nat. Hist., Bot. Ser. 12: 390. 1936. York Bot. Gard. 23: 302. 1972. Calderonia
Calderonia salvadorensis, J. Wash. Acad. Sci. klugei Standi., J. Wash. Acad. Sci. 15:6.
13: 290. 1923. Type: EL SALVADOR: San 1924. Type: PANAMA: Panama, near Chepo,
Salvador, Jul 1922, S. Calderon 761 1924, H. C. Kluge 19 [-Yale 7132] (Holotype
(Holotype US 1151718 (photos MEXU, US 1206424 (photos MEXU, PTBG); Isotype
PTBG)).—Simira salvadorensis (Standi.) F (photo PTBG), (plus another probable
Steyerm. isotype at F, a fruiting collection from type
Sickingia vestita Lundell, Wrightia 4: 51. 1968. tree, Kluge s.n., also Yale 7132)).
Nom. no\ Lundell, Simira maxonii (Standi.) Steyerm., Mem. New
Contrib. Univ. Mich. Herb. 4: 32. 1940. Not York Bot. Gard. 23: 306. 1972. Genipa
Sickingia vestita Standi, 1936. Type: maxonii Standi., J. Wash. Acad. Sci. 8: 642.
MEXICO: Tabasco, Reforma, on the Rio San 1918. Type: PANAMA: Panama, Canal Zone,
Pedro de Martir, above Balacan, 22-26 May forest along the Rio Indio de Ganin, near sea
1939, E. Matuda 3161 (Holotype MICH, n.v.; level, 17 Feb 1911, W. R. Maxon 4848
Isotypes A, n.v., TEX-LL (photos MEXU, (Holotype US 675223 (photos MEXU,
PTBG), US (photos MEXU, PTBG)). PTBG)).
—Simira salvadorensis (Standi.) Steyerm. Simira mexicana (Bullock) Steyerm., Mem.
New York Bot. Gard. 23: 307. 1972. Sickingia
mexicana Bullock in Hook. Icon., ser. 4, 4:
 OF MEXICAN AND CENTRAL AN

tab. 3321. 1936. Type: MEXICO: Mexico, Turcz. [Herein treated as a synonym of
Distr. Temascaltepec, Luvianos, 26 Sep 1933, Rondeletia L.]
G. B. Hinton 4818 (Holotype K, n.v.; Isotype Siphonandra mexicana Turcz., Bull. Soc. Imp.
US (photos MEXU, PTBG)). Sickingia. Naruralistes Moscou 21: 581. 1848. Type:
Simira myrkmtha (Standi.) Steyerm., Mem. MEXICO: Oaxaca, without precise locality,
New York Bot. Gard. 23: 306. 1972. Sickingia 3000 ft.. H Galeotti 266-1 hi.s (Holotype
myriantha Standi., Publ. Field Columbian presumably LE, n.v.; Isotype G-DEL. n.v.
Mus., Bot. Ser. 7: 27. 1930. Type: (photos A, MO)).—Rondeletia mexicana
COLOMBIA: Magdalena, Las Nubes road, (Turcz.) Standi.
region of Santa Marta, 1200 m, 3 Dec 1898,
H. H. Smith 1810 (Holotype F 138660; Siphoma Benth., PI. Hartweg. 84. 1841. Not
Isotypes MO, NY, n.v., US, n.v., W, n.v., Y, Siphonia D. Richard ex Schreber, 1791, nor
n.v). Holtonia.—Elaeagia mvriantha Siphoma E. M. Fries, 1820. Type: Siphonia
(Standi.) C. M. Taylor & Hammel nvalis Benth. (=Lindenia Benth., 1841). See
Simira rhodoclada (Standi.) Steyerm., Mem. Darwin (1976: 427) for discussions regarding
New York Bot. Gard. 23: 307. 1972. Exandra publication of the generic names Lindcma vs.
rhodoclada Standi., J. Wash. Acad. Sci. 13: Siphonia. [Herein treated as a synonym of
292. 1923. Type: MEXICO: Oaxaca, between Augusta Pohl.]
La Venta and Niltepec, 200 ft., 14 Jul 1895, Siphonia rivalis Benth., Pi. Hartweg. 84. 1841.
E. W. Nelson 2796 (Holotype US 229232 Type: GUATEMALA: In rivis, la Vera Paz, T.
(photos MEXU, PTBG); Isotype F (photos Hartweg 581 (Holotype K, n.v. (photo MO)).
MEXU, PTBG)). Simira. —Augusta rivalis (Benth.) J. H. Kirkbr.
Simira salvadorensis (Standi.) Steyerm., Mem. Note: according to Farr et al. (1979: 1620),
New York Bot. Gard. 23: 307. 1972. although the name Siphoma was actually
Calderonia salvadorensis Standi., J. Wash. printed, it was crossed out in ink and the name
Acad. Sci. 13: 290. 1923. Type: EL Lindenia substituted, "
SALVADOR: San Salvador, Jul 1922, S copies before publication,
Calderdn 761 (Holotype US 1151718 (photos published on p. 351.
MEXU, PTBG)). Sickingia.
Solena Willd., Sp. PL 1: 961. 1798. Not Solena
Sipanea Aubl., Hist. PI. Guiane 1: 147, t. 56. Loureiro, 1790. Type: Solena longiflora
1775. Type: Sipanea pratensis Aubl. (Aubl.) Willd. (= Posoquena longiflora Aubl.)
Sipanea biflora (L. f.) Cham. & Schltdl., [Herein treated as a synonym of Posoqueria
Linnaea4: 168. 1829. Virecta biflora L. f, Aubl.]
Suppl. 17, 134. 1781. Type: SURINAM: Solena latifolia Rudge, PI. Guian. 26. 1805.
Without precise locality, Dalberg s.n. Type: FRENCH GUIANA: Martin s.n. (BM,
(presumably LINN, n.v.). n.v).—Posoqueria latifolia (Rudge) Roem.
Sipanea palustris Seem., Bot. Voy. Herald 136. & Schult.
1854. Type: PANAMA: Panama, in swamps
near Panama, Seemann 347 (Holotype BM, in Sommera Schltdl., Linnaea 9: 602. 1835. Type:
part [excluding specimen of Microlicia Sommera arborescens Schltdl.
(Melastomataceae)], n.v. (photo MO)). Sommera acuminata Oerst. ex Standi.. J. Wash.
Limnosipanea. Acad. Sci. 17: 340. 1927. Type: MEXICO:
Oaxaca, Amatlan, F. Liebmann 11712
Siphonandra Turcz., Bull. Soc. Imp. Naruralistes (Holotype C, n.v. (photo MEXU); Isotype
Moscou 21: 581. 1848. Not Siphonandra US). Arachnothryx, Rondeletia.—Rondeletia
Klotzsch 1851. Type: Siphonandra mexicana scabra Hemsl.
 $ Schltdl., Linnaea 9: 602. Sommera montana L. O. Williams, Phytologia
1835. Type: MEXICO: Veracruz, Hacienda de 26: 124. 1973. Type: HONDURAS: Morazan,
la Laguna, Schiede 272 (Holotype presumably margen de la Quebrada de La Paz, en medio
HAL, n.v.; Isotypes F (photos MEXU, de la floresta de pino-liquidambar de la
PTBG), LE, n.v.(fragment F), MO (photos Montana Zanquin, 1600 m, 14 May 1950, A.
MEXU, PTBG), US (photos MEXU, PTBG)). Molina R. 2829 (Holotype F 1389080 (photo
Sommera chiapensis Brandegee, Univ. Calif. PTBG); Isotypes EAP, n.v., US (photos
Publ. Bot. 6: 196. 1915. Type: MEXICO: MEXU, PTBG)).
Chiapas, Finca Irlanda, Jun 1914, C. A. Sommera parva Lorence, Novon4: 134, fig. 9.
Purpus 7263 (Holotype UC 174914. n.v.; 1994. Type: MEXICO: Chiapas, Mun. de
Isotypes F (photos MEXU, PTBG), MO, 2 Tila, Pie de Cerro K'abana, 900 m, 1 Jul
sheets (photos MEXU, PTBG), US (photos 1982, A. Mendez Ton {A. Shilom Ton) 4367
MEXU, PTBG)). (Holotype MEXU; Isotypes BM, F, MO
Sommera donnell-smithii Standi., Contr. U.S. (photo PTBG), NY, PTBG, US, XAL).
Natl. Herb. 17: 436. 1914. Type: COSTA Sommera rivularis L. O. Williams, Phytologia
RICA: San Jose, Alajuehta, 3000 ft., Mar 26: 125. 1973. Type: COSTA RICA:
1894, J. D. Smith 6592 (Holotype US 245836 Puntarenas, Rio Esquinas, near kilometer 42,
(photos MEXU, PTBG)). 30 m, 8 Dec 1949, P. H. Allen 5431 (Holotype
Sommera dunlapii L. O. Williams, Phytologia F 1679274 (photo PTBG); Isotypes EAP, n.v.,
26: 122. 1973. Type: PANAMA: Changuinola F, US (photos MEXU, PTBG)).-
Valley, 17 Jan 1924, V. C. Dunlap 351
(Holotype F 635718 (photo PTBG); Isotype ubcordata Standi., N. Amer. Fl. 32:
US (photos MEXU, PTBG)).—Sommera 144. 1921. Type: MEXICO: Sinaloa,
donnell-smithii Standi. Colomas, 1897, J. N. Rose s.n. (Holotype US
Sommera fusca Oerst. ex Standi., J. Wash. 866799 (photos MEXU, PTBG)).—Sommera
Acad. Sci. 17: 339. 1927. Type: MEXICO: grandis (Bartl. ex DC.) Standi.
[Presumably Oaxaca] Jocoaltepec, Jun 1842, Sommera zygocalyx L. O. Williams, Phytologia
F. Liebmann 11720 [=Liebmann Rubiaceae 26: 126. 1973. Type: MEXICO: Oaxaca,
36] (Holotype C, n.v.; Isotype US (photos Distr. Choapam [=Choapan], Yaveo, Arroyo
MEXU, PTBG)). de Culebras, 430 m, 21 Mar 1938, Y. Mexia
Sommera grandis (Bartl. ex DC.) Standi., N. 9191 (Holotype F 982858 (photos MEXU,
Amer. Fl. 32: 145. 1921. Petesia grandis PTBG); Isotypes UC (photo PTBG), US
Bartl. ex DC, Prodr. 4: 395. 1830. Type: (photos MEXU, PTBG)).—Sommera
MEXICO: Without precise locality, Haenke arborescens Schltdl.
s.n. (Holotype presumably PR, n.v., fragment
G-DC, n.v. (microfiche 4: 395.1)). Spermacoce L., Sp. PI. 1: 102. 1753. Type:
Sommera guatemalensis Standi., Contr. U.S.
Natl. Herb. 17: 436. 1914. Type: Spermacoce ambigua Brandegee, Zoe 5: 180.
GUATEMALA: Alta Verapaz, Cubliquitz, 1904. Type: MEXICO: Veracruz, Cordoba,
350 m, May 1902, H. von Tuerckheim 8225 Jan 1904, C. A. Purpus 417 (Holotype UC,
(Holotype US 398487 (photos MEXU, n.v.; Isotypes MO (photo PTBG), US (photos
PTBG)). MEXU, PTBG)).—Diodia brasiliensis
Sommera mesochora Standi., Contr. U.S. Natl. Spreng.
Herb. 17: 439. 1914. Type: PANAMA: Spermacoce apiculata Willd. ex Roem. &
Chiriqui, vicinity of El Boquete, 1000-1300 Schult., Syst. Veg. 3: 531. 1818. Type:
m, 2-8 Mar 1911, W. R. Maxon 4941 BRAZIL: Without precise locality,
(Holotype US 675326 (photos MEXU, Hoffmannsegg s.n. (B-WI 2626,
PTBG)).—S
 k. NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

n.v.).—Diodia apiculata (Willd. ex Roem. &

Schult.) K. Schum. t Galeotti. Howard
Spermacoce ascendens Sesse & Mo?., Fl. (1989: 464) incorrectly c
Mexic. 25. 1893. Type: MEXICO: Habitat in
Ahualulci depressis, ac humidioribus. Floret
Octobri, Sesse & Mocino s.n. (presumably
MA, n.v.).—Status uncertain. 10.;4:
Spermacoce aspera (M. Martens & Galeotti) 330-331).
Hemsl., Biol. Centr.-Amer., Bot. 2: 58. 1881. Spermacoce densiflora (DC.) Alain, Phytologia
Borreria aspera M. Martens & Galeotti, Bull. 54: 113. 1983. Borreria densiflora DC,
Acad. Roy. Sci. Bruxelles 11: 130. 1844. Prodr. 4: 542. 1830. Type: JAMAICA: Swart:
Type: MEXICO: Oaxaca, Juquila, pres de la s.n. (Holotype G-DC, n.v. (microfiche 4:
cote d'Oaxaca baignee par l'Ocean Pacifique, 542.3)). For typification see Steyermark
a 4000 pieds, H. Galeotti 2625 (Holotype (1972).
presumably BR, n.v.).—Spermacoce Spermacoce depressa Sesse & Moc., PI. Nov.
suaveolens (G. Mey.) Kuntze Hispan. 17. 1887 (La Naturaleza 2, 1. suppl.
Spermacoce asperifolia M. Martens & Galeotti, 17. 1887). Type: MEXICO: Habitat Patzquari.
Bull. Acad. Roy. Sci. Bruxelles 11: 132. 1844. Sesse & Mocino s.n. (presumably MA, n.v.).
Type: MEXICO: Oaxaca, pres de l'Ocean —Status uncertain.
Pacifique, H. Galeotti 2626 (Holotype Spermacoce distans Kunth in Humb., Bonpl. &
presumably BR, n.v.; Isotype P, n.v. (photo Kunth, Nov. Gen. Sp. 3: 344. 1820. Type:
MO)). Borreria.Stadia scabra (C. Presl) MEXICO: Jalisco, Volcan de Jorullo, 3000 ft.,
Standi. Humboldt & Bonpland s.n. (Holotype P-
Spermacoce assurgens Ruiz & Pav., Syst. Veg. Bonpland, n.v. (photo MO)). Borreria.
Fl. Peruv. 1: 60, t. 92. 1798. Type: PERU: —Diodia cf. apiculata (Willd. ex Roem. &
Habitat in ruderalis Limae Chancay et Schult.) K. Schum.
Huanuci, Ruiz Lopez & Pavon s.n. (Syntype Spermacoce diversifolia Kunth in Humb.,
MA, n.v. (photo MO); Plukenet Aim. 33, tab. Bonpl. & Kunth, Nov. Gen. Sp. 3: 341. 1820.
136/4 (Syntype, n.v.). Borreria. For Type: MEXICO: Without precise locality,
typification, see Verdcourt, 1983. Humboldt & Bonpland s.n. (Holotype P-
Spermacoce capitata Sesse & Moc., PI. Nov. Bonpland, n.v. (photo MO)).—Crusea
Hispan. 17. 1887 (La Naturaleza 2, 1. suppl. diversifolia (Kunth) W. R. Anderson
17. 1887). Not Spermacoce capitata Ruiz & Spermacoce echioides Kunth in Humb., Bonpl.
Pav., 1798. Type: MEXICO: Habitat & Kunth, Nov. Gen. Sp. 3: 344. 1820. Type:
Patzquari, Sesse & Mocino 633 (Holotype MEXICO: Campeche, Humboldt & Bonpland
presumably MA, n.v.; probable Isotype F s.n. (Holotype presumably P-Bonpland, n.v.).
(photo PTBG)).—Crusea hispida (Mill.) —Spermacoce assurgens Ruiz & Pav.
B. L. Rob. var. hispida Spermacoce exilis (L. O. Williams) C. D.
Spermacoce coccinea Sesse & Moc., Fl. Mexic. Adams in W. C. Burger & C. M. Taylor,
24. 1893. Not Spermacoce coccinea Pav., Fieldiana: Bot., n.s. 33: 316. 1993. Based on
1798. Type: Habitat in montibus Malacatepec Borreria exilis L. O. Williams, Phytologia 28:
ad Zitacuarum interiectis. Floret Augusto, 227. 1974. Not Spermacoce gracilis Ruiz &
Sesse & Mocino s.n. (presumably MA, n.v.). Pav. 1798. Type: COSTA RICA: Cocos
—Status uncertain, possibly Crusea Island, Wafer Bay, 28 Jun 1932, J. T. Howell
coccinea DC. 10178 (Holotype F 1031197 (photo PTBG)).
Spermacoce confusa Rendle, J. Bot. 74: 12. Spermacoce glabra Sesse & Moc., PI. Nov.
1936. Type: MEXICO: Zacuapan, Mirador, Hispan. 17. 1887 (La Naturaleza 2, 1. suppl.
H. Galeotti 2611 (Holotype K, n.v.; Isotype 17. 1887). Not Spermacoce glabra Michx.,
 1803. Type: MEXICO: Habitat Patzquari, Larranaga, 1923. Type: MAURITIUS: Sieber
floret Augusto, Sesse & Mocino s.n. Fl. Maunt. II, 144 (Holotype G, n.v.; Isotypes
(presumably MA, n.v.).—Status uncertain. P). See Verdcourt (1983) for discussion of
Spermacoce haenkeana (DC.) Hemsl., Biol. typification.—Spermacoce exilis (L. O.
Centr.-Amer., Bot. 2: 59. 1881. Borreria Williams) C D. Adams
haenkeana DC, Prodr. 4: 547. 1830. Type: Spermacoce megalocarpa A. Gray, Proc. Amer.
MEXICO or PERU: Without precise locality Acad. Arts 21: 381. 1886. Type: MEXICO:
or collector (G-DEL, n.v. (photo MO)). Chihuahua, south-western Chihuahua, wet
—Spermacoce suaveolens (G. Mey.) Kuntze places in the higher mountains above
Spermacoce jaliscensis M. E. Jones, Extracts Batopilas, Aug-Nov 1885, E. Palmer 185
Contr. Western Bot. 18: 67. 1933. Type: (Holotype GH, n.v.; Isotypes NY, n.v., US
MEXICO: Jalisco, Orendain, 30 Nov 1930, (photos MEXU, PTBG)).—Crusea
Vones 27118 (location unknown).—Status megalocarpa (A. Gray) S. Watson
uncertain. Spermacoce nesiotica (B. L. Rob.) G. A. Levin
Spermacoce laevis Lam., Tabl. Encycl. Meth. in G. A. Levin & R. Moran, Vase. Fl. Isla
Bot. 1: 273, tab. 1435. 1791. Type: SANTO Socorro, Mexico (Mem. San Diego Soc. Nat.
DOMINGO: Without precise locality, Martin Hist. 16): 51. 1989. Borreria nesiotica B. L.
s.n. (Holotype P-LA, n.v.). Rob., Proc. Amer. Acad. Arts 45: 409. 1910.
Borreria.—Spermacoce tenuior L. For a Type: MEXICO: Soccoro Island
discussion of synonymy and misapplication of (Revillagigedo Group), 1897, A. W. Anthony
this name see Verdcourt (1983: 545). s.n. (Holotype GH, n.v.; Isotype US (photos
Spermacoce latifolia Aubl., Hist. PI. Guiane 1: MEXU, PTBG)).
55, /. 19, fig. 1. 1775. Type: FRENCH Spermacoce ocymifolia Willd. ex Roem. &
GUIANA: Cayenne, Aublet s.n. (Holotype P, Schult., Syst. Veg. 3: 530. 1819 [as
n.v.; Isotype MA, n.v. (photo MO)). Borreria. ocimifolia]. Type: WEST INDIES: Without
Spermacoce linearis Sesse & Mo?., Fl. Mexic. precise locality, Rudolphi s.n. in 1768
25. 1893 (La Naturaleza 2, 1: App. 25. 1893). (presumably B-WI, n.v.).
Type: MEXICO: Habitat in montibus Xorullo Spermacoce ocymoides Burm. f, Fl. Ind. 34.
vicinis. Floret Septembri, Sesse & Mocino s.n. 1768. Type: Tropical Asia, collector and
(presumably MA, n.v.).—Status uncertain. precise locality unknown (presumably G,
Spermacoce longiseta (M. Martens & Galeotti) n.v.). Note: Spermacoce ocymoides is not
Hemsl., Biol. Centr.-Amer., Bot. 2: 59. 1882. known from the Neotropics; however, this
Borreria longiseta M. Martens & Galeotti, name has been applied incorrectly by many
Bull. Acad. Roy. Sci. Bruxelles 11: 128. 1844. authors to tropical American plants
Type: MEXICO: Veracruz, Zacuapan et representing S. exilis, S. ovalifolia, and S.
Mirador [label on Isotype at P states
"cordilleres pres de Vera Cruz, 1000 pieds"], e ovalifolia (M. Martens & Galeotti)
H. Galeotti 2611 (Holotype BR, n.v.; Isotypes Hemsl., Biol. Centr.-Amer., Bot. 2: 59. 1881.
K, n.v., P, n.v. (photo MO)). Also see Borreria ovalifolia M. Martens & Galeotti,
comments under Borreria longiseta. The Bull. Acad. Roy. Bruxelles 11: 129. 1844.
description does not apply to the same plant as Type: MEXICO. Veracruz, Mirador, 5000 ft.,
represented by the type (C. D. Adams, pers. H. Galeotti 2609 (Holotype BR, n.v.).
comm. 1996).—Status uncertain. Spermacoce patula (M. Martens & Galeotti)
Spermacoce mauritiana Gideon in Verdcourt, Hemsl., Biol. Centr.-Amer., Bot. 2: 59. 1881.
Kew Bull. 37: 547. 1983. Norn. nov. for Borreria patula M. Martens & Galeotti, Bull.
Borreria repens DC, Prodr. 4: 544. 1830. Not Acad. Roy. Bruxelles 11: 128. 1844. Type:
Spermacoce repens Willd. ex Cham. & MEXICO: Veracruz, Mirador, 3000 ft., H.
Schltdl., 1828, nor Sesse & Moc,, 1893, nor
 ^ NOMENCLATOR OF &

Galeotti 2608 (Holotype BR, n.v.; Isotype K, Crusea.—Crusea hispida (Mill.) B. L. Rob.
n.v.).—Spermacoce prostrata Aubl.
Spennacoce polycephala Bartl. ex DC., Prodr. Spermacoce scabiosoides (Cham. & Schltdl.)
4: 558. 1830. Type: MEXICO: Without Kuntze, Revis. Gen. PI. 3: 123. 1898. Borreria
precise locality, Haenke s.n. (Holotype scabiosoides Cham. & Schltdl., Linnaea 3:
presumably PR, n.v., fragment G-DC, n.v. 318. 1828. Type: BRAZIL: Sellow s.n.
(microfiche 4: 558.54)).—Status uncertain. (presumably B, n.v.).
Spermacocepringlei S. Watson, Proc. Amer. Spermacoce serrulata P. Beauv., Fl. Oware
Acad. Arts 25: 152. 1890. Type: MEXICO: Benin 1: 39. tab. 23. 1805. Type: NIGERIA:
Jalisco, shaded hillsides near Guadalajara, 27 Palisot de Beauvois s.n. (presumably P,
Sep 1889, C. G. Pringle 2464 (Holotype GH, n.v.).—Diodia serrulata (P. Beauv.) G.
n.v.; Isotypes MO (photos MEXU, PTBG), Taylor
US (photos MEXU, PTBG)).—Spermacoce Spermacoce stricta Sesse & Mo?., Fl. Mexic.
ovalifolia (M. Martens & Galeotti) Hemsl. 24. 1893. Type: MEXICO: Habitat in calidis,
Spermacoce prostrata Aubl., Hist. PI. Guiane NIL, Sesse & Mocino s.n. (presumably MA,
Franc. 1: 58, tab. 20, fig. 3. 1775. Type:
FRENCH GUIANA: Aublet s.n. (Holotype P. iveolens (G. Mey.) Kuntze,
Revis. Gen. PI. 3: 123. 1898. Borreria
Spermacoce psyllioides Kunth in Humb., Bonpl. suaveolens G. Mey., Prim. Fl. Esseq. 81, pi. 1.
& Kunth, Nov. Gen. Sp. 3: 346. 1819. Type: 1818. Type: SURINAM: In arenosis
MEXICO: Without precise locality, Humboldt continentis circa Arouabischkrek, G. Meyer
& Bonplands.n. (Holotype P-Bonpland, n.v. s.n. (Holotype GOET, n.v.).
(photo MO)). Crusea psyllioides (Kunth) Spermacoce tenui Sesse & Moc., Fl. Mexic. 25.
W. R. Anderson 1893 (LaNaturaleza2,l. suppl. 25. 1897).
Spermacoce reclinata Nees, Horae Phys. Berol. Type: MEXICO: Habitat Patzcuan. Floret
50. 1820. Type: MEXICO: In regno Augusto, Sesse & Mocino s.n. (presumably
Mexicano, without precise locality or MA, n.v.).—Status uncertain.
collector (location unknown).—Spermacoce Spermacoce tenuiflora Sesse & Mo?., Fl. Mexic.
24. 1893 (La Naturaleza 2.1. suppl. 24. 1897).
Spermacoce rigida Willd. ex Roem. & Schult., Type: Habitat in Zitacuan. Floret Augusto,
Syst.Veg.3:531. 1818. Type: Sesse & Mocino s.n. (presumably MA, n.v.).
VENEZUELA: Silla de Caracas, Humboldt & —Crusea longifiora (Roem. & Schult.) W. R.
Bonpland s.n. (Holotype B-WI 2628, n.v.; Anderson
Isotype G-DC, n.v. (microfiche Spermacoce tenuifolia Sesse & Moc., PI. Nov.
IDC.7440.174:111,4)). Diodia.—Diodia Hispan. 17. 1897 (La Naturaleza 2, 1. suppl.
apiculata (Willd. ex Roem. & Schult.) K. 17. 1897). Type: MEXICO: Habitat Patzquari.
Sesse & Mocino s.n. (presumably MA, n.v.).
Spermacoce riparia Cham. & Schltdl., Linnaea
3: 355. 1828. Type: BRAZIL: Rio Negro, Spermacoce tenuior L., Sp. PI. 1: 102. 1753.
(Rincon de Gallinas), Sellow s.n. (Holotype Lectotype: Habitat in Carolina, Spermacoce
presumably B or HAL, verticillis tenuioribus Dillenius, Hortus
Elthamensis 370, tab. 277, fig. 359. 1732.
Spermacoce rubra Jacq., PI. Rar. Hort. Typotype in Herb. Sherard (OXF, n.v.), drawn
Schoenbr. 3: 3,pi. 256. 1798. Type: Cuba [but from specimens grown at Eltham; designated
probably from MEXICO], Jacquin s.n. by Rendle (1934: 333). Typification
(probable type W, n.v., fragment F). For confirmed by Verdcourt (1983).
discussion, see Anderson (1972: 110-111). Spermacoce tenuior Sesse & Mocino, Fl. Mexic.
24. 1893. Not Spermacoce tenuior L., 1753.
 Staelia scabra (C. Presl) Standi., Publ. Field
Columbian Mus., Bot. Ser. 8: 385. 1931.
MA, n.v.).—St Diphragmus scaber C. Presl, Bot. Bermerk.
Spermacoce tetracocca M. Martens & Galeotti, 81. 1844. Type: MEXICO: Habitat in Mexico,
Bull. Acad. Roy. Soc. Bruxelles 112: 132. precise locality or collector not cited.
1844. Type: MEXICO: Michoacan, Santa
Maria, near Morelia, H. Galeotti 2587 Stannia H. Karst., Auswahl Gewachse
(Holotype presumably BR, n.v.; Isotype K, Venezuela 27. 1848. Type: Stannia formosa
n.v., P (photos MEXU, PTBG)).—Richardia H. Karst.
tricocca (Torr. & A. Gray) Standi, subsp. Stannia panamensis Walp. & Duchass., Linnaea
tetracocca (M. Martens & Galeotti) W. H. 23: 755. 1850. Type: PANAMA: Isthmo
Lewis & R. L. Oliv. Panamensis, Duchassaing ? (location
Spermacoce tetraquetra A. Rich, in Sagra, unknown).—Posoqueria panamensis (Walp.
Hist. Fis. Cub. Bot. 11: 29. 1850. Type: & Duchass.) Walp.
CUBA: Without precise locality, R. de la
Sagra s.n. (Holotype presumably P, n.v.; Stenostomum C. F. Gaertn., Fruct. Sem. PI. 3:
probable Isotype F (photo PTBG)). 69. 1806. Type: Stenostomum lucidum (Sw.)
Spermacoce vegeta (Standi. & Steyerm.) C. D. C. F. Gaertn.
Adams in W. C. Burger & C. M. Taylor, Stenostomum lucidum (Sw.) C. F. Gaertn.,
Fieldiana, Bot., n.s. 30: 320. 1993. Borreria Fruct. Sem. PI. 3: 69. 1806. Laugeria lucida
vegeta Standi. & Steyerm., Publ. Field Mus. Sw., Prodr. Veg. Ind. Occ. 48. 1788. Type:
Nat. Hist., Bot. Ser. 23: 21. 1943. Type: JAMAICA: Without precise locality, Swartz
GUATEMALA: Dept. Jutiapa, between s.n. (presumably S, n.v.).
Jutiapa and La Burrera, NE of Jutiapa,
800-850 m, 1 Nov 1940, P. C. Standley Steyermarkia Standi., Publ. Field Mus. Nat.
76007 (Holotype F 1091300 (photos MEXU, Hist, Bot. Ser. 22: 216. 1940. Type:
PTBG)). Steyermarkia guatemalensis Standi.
Spermacoce verticillata L., Sp. PI. 1: 102. Steyermarkia guatemalensis Standi., Publ.
1753. Type: JAMAICA: Hort. Cliff. 33 (BM, Field Mus. Nat. Hist., Bot. Ser. 22: 216. 1940.
n.v.). Borreria. For typification see Rendle Type: GUATEMALA: Izabal, Rio Dulce, 2^
(1934:331). miles west of Livingston, on south side (left.-
Spermacoce villosa Sw., Prodr. Veg. Ind. Occ. hand side going up river), alt. 1-3 m, 16 Apr
29. 1788. Type: JAMAICA: Swartzs.n. 1940, J. A. Steyermark 39520 (Holotype F
(Holotype presumably BM, n.v.). 1035008 (photo PTBG); Isotypes F 1035009,
—Mitracarpus hirtus (L.) DC. MO (photos MEXU, PTBG)).

Stachyarrhena Hook, f., Hook. Icon. PI. 1068. Stomandra Standi., Publ. Field Mus. Nat. Hist.,
1870. Type: Stachyarrhena spicata Hook. f. Bot. Ser. 23: 247. 1947. Type: Stomandra
Stachyarrhena heterochroa Standi., Contr. costaricensis Standi. [Herein treated as a
U.S. Natl. Herb. 18: 142. 1916. Type: synonym of Rustia Klotzsch.]
PANAMA: Colon, Loma de la Gloria, near Stomandra costaricensis Standi., Publ. Field
Fato (Nombre de Dios), 10-104 m, Jul-Aug Mus. Nat. Hist., Bot. Ser. 23: 247. 1947.
1911,//. Pittier 4078 (Holotype US 679174 Type: COSTA RICA: San Jose, basin of El
(photos MEXU, PTBG); Isotype F). General, 675-900 m, Mar 1940, A. F. Skutch
4777 (Holotype F 1044344 (photo PTBG);
Staelia Cham. & Schltdl., Linnaea 3: 364, pi. 3, Isotypes MO (photo PTBG), US).—Rustia
fig. 3. 1828. Type: Staelia thymoides Cham. costaricensis (Standi.) Lorence
& Schltdl.
 ^ NOMENCLATOR OF MEXICAN AND CENTRAL AME

Strumpfia Jacq., Enum. Syst. PI. 28. 1760.

Type: Strumpfia maritima Jacq.
Strumpfia maritima Jacq., Enum. Syst. PI. 28.
1760. Type: Curacao, in rupestribus Tapogomea Aubl., Hist. PI. Guiane 157. 1775.
maritimus, Jacquin s.n. (location unknown). Type: Tapogomea violacea Aubl. Norn. rej.
Designation of a neotype may be necessary vs. Cephaelis Sw. 1788 (nom. cons.). [Herein
according to Howard (1989: 469). treated as a synonym of Psychotria L.]
Tapogomea axillaris (Sw.) Poir. in Lam.,
Stylosiphonia Brandegee, Univ. Calif. Publ. Bot. Encycl. Meth. Bot. 7: 587. 1806. Cephaelis
6: 70. 1914. Type: Stylosiphonia glabra axillaris Sw., Prodr. 45. 1788. Type: ST.
Brandegee CHRISTOPHER [=ST. KITTS]: Without
Stylosiphonia glabra Brandegee, Univ. Calif. precise locality, Swartzs.n. (presumably S,
Publ. Bot. 6: 70. 1914. Type: MEXICO: n.v.).—PsNihotri.i aubktianj Steyerm.
Chiapas, without precise locality, 1913, C A. Tapogomea elata (Sw.) Poir.in Lam., Encycl.
Purpus 7057 (Holotype UC 173437; Isotypes Meth. Bot. 7: 585. 1806. Cephaelis elata Sw.,
F (photo PTBG), GH, n.v., MO (photos Prodr. 45. 1788. Type: JAMAICA: Without
MEXU, PTBG), US). precise locality. Swartz s.n. (presumably S,
nia Salvadorensis Standi., J. Wash. n.v.).—Psychotria elata (Sw.) Hammel
Acad. Sci. 14: 246. 1924. Type: EL Tapogomea tomentosa Aubl., Hist. PI. Guiane
SALVADOR: San Vicente, Volcan de San 160, pi. 61. 1775. Type: FRENCH GUIANA:
Vicente, 1200-1500 m, 7-8 Mar 1922, P. C In sylvarum locis apertis, Aublet s.n.
Standley 21559 (Holotype US 1137349 (Holotype presumably P, n.v.; Isotype B,
(photos MEXU, PTBG); Isotype GH, n.v.). presumably destroyed (photo MO)).
—Rondeletia jurgensenii Hemsl. Cephaelis. Evea. Uragoga.—Psychotria
" ". Arg.
Syringantha Standi., Publ. Field Columbian
Mus., Bot. Ser. 8: 153. 1930. Type: Tertrea DC, Prodr. 4: 481. 1830. Type: Tertrea
Syringantha loranthoides Standi. martinicensis DC. [Herein treated as a
Syringantha coulteri (Hook. f. ex Hemsl.) T. synonym of Machaonia Humb. & Bonpl.]
McDowell, Novon 6: 277. 1996. Exostema Tertrea martinicensis DC, Prodr. 4: 481. 1830.
coulteri Hook. f. in Hemsl., Diag. PI. Nov. 32. Type: MARTINIQUE: Richard s.n. (Holotype
1879 [as Exostemma]. Type: MEXICO: P, n.v.).—Machaonia martinicensis (DC)
Hidalgo, Zimapan, Coulter 209 (Holotype K,
n.v.; Isotype MO).
ha loranthoides Standi., Publ. Field Tobagoa Urb., Repert. Spec. Nov. Regni Veg.
Columbian Mus., Bot. Ser. 8: 153. 1930. 14: 341. 1916. Type: Tobagoa maleolens Urb.
Lectotype: MEXICO: [Presumably Hidalgo], Tobagoa maleolens Urb., Repert. Spec. Nov.
entre San Diego y la Hacienda de Regni Veg. 14: 343. 1916. Type: TOBAGO:
Santiaguillo, Jul 1842, Karwinsky 308 [upper In sylvis ad Gr. Dog River, Nov 1889, Eggers
specimen] (LE, n.v., fragment F (photos 5783 (Holotype B, presumably destroyed;
MEXU, PTBG); Isolectotype MW, n.v.; Isotypes K, n.v., US (photo PTBG)).
designated by McDowell 1996a: 277).
Syntypes: MEXICO: Santiaguillo, Jul 1842, Tocoyena Aubl.. Hist. PI. Guiane 1: 131, tab.
Karwinsky 308 [lower specimen] (LE, n.v., 50. 1775. Type: Tocoyena longiflora Aubl.
fragment F); El Pantario et in Santiaguillo, Tocoyena obliquinervia (Standi.) Standi., Contr.
Apr 1843, Karwinsky 308b (LE, n.v. Arnold Arbor. 5: 152. 1933. Posoqueria
(fragment F), MW, n.v.); Santiaguillo, Oct obliquinervia Standi., J. Wash. Acad. Sci. 18:
1842, Karwinsky 308c [original 31] (LE, n.v., 167. 1928. Type: COSTA RICA: Forets et
 bords du Rio Naranjo, 200-250 m, Mar 1893, ^rbacea (P. Browne) Standi., N.
A. Tonduz 7528 [erroneously published as Amer. Fl. 32: 147. 1921. Coccocypselum
9528] (Holotype US 764158 (photos MEXU, herbaceum P. Browne, Civ. Nat. Hist. Jam.
PTBG)).—Tocoyena pittieri (Standi.) Standi. 144, pi 6,f. 2. 1756. Type: JAMAICA:
Tocoyena pittieri (Standi.) Standi., Contr. Cooler mountains of Liguancee and Mont-
Arnold Arbor. 5: 151. 1933. Posoqueha Diable, P. Browne s.n. (LINN, n.v.).
pittieri Standi., J. Wash. Acad. Sci. 18: 167. —Coccocypselum herbaceum P. Browne
1928. Type: PANAMA: Near Hydrographic Tontanea hirsuta (Bartl. ex DC.) Standi., N.
Station on Trinidad River, 17 May 1914, H. Amer. Fl. 32: 147. 1921. Coccocypselum
Pittier 6635 (Holotype US 716697 (photos hirsutum Bartl. ex DC, Prodr. 4: 396. 1830.
MEXU, PTBG)). Type: Precise locality unknown [probably
Tocoyena tabascensis Standi., Field Mus. Nat. Trinidad], Haenke s.n. (Holotype presumably
Hist., Bot. Ser. 11: 273. 1936. Type: PR, n.v.. fragment G-DC, n.v. (microfiche 4:
MEXICO: Tabasco, Paso de Espejo, 25 Nov 396.5)).—Coccocypselum hirsutum Bartl. ex
1888, J. N. Rovirosa 301 (Holotype K (photos DC. var. hirsutum
F, MEXU, PTBG)).—Blepharidium Tontanea hispidula Standi., N. Amer. Fl. 32:
guatemalense Standi. 147. 1921. Type: PANAMA: Chagres,
Isthmus of Panama, Jan-Feb 1850, A. Fendler
Tontanea Aubl., Hist. PI. Guiane 1: 108. 1775. 179 (Holotype US 42058 (photo MO); Isotype
Type: Tontanea guianensis Aubl. [Herein US). Coccocypselum.—Coccocypselum
treated as a synonym of Coccocypselum P. herbaceum P. Browne
Browne.]
Tontanea canescens (Willd. ex Cham. & Tournefortiopsis Rusby, Bull. New York Bot.
Schltdl.) Standi., N. Amer. Fl. 32: 146. 1921. Gard. 4: 369. 1907. Type: Tournefortiopsis
Coccocypselum canescens Willd. ex Cham. & reticulata Rusby. [Herein treated as a
Schltdl., Linnaea 4: 139. 1829. Not C. synonym of Guettarda L.]
canescens Willd. ex Schult. & Schult., 1827. lata Rusby, Bull. New
Type: VENEZUELA: Buenavista prope York Bot. Gard. 4: 369. 1907. Not Guettarda
Caracas, Humboldt & Bonpland s.n. reticulata Griseb., 1863. Type: BOLIVIA:
(Holotype B-WI 2874, n.v.). Without precise locality, Bang 2230
—Coccocypselum lanceolatum (Ruiz & (Holotype US, n.v.).—Guettarda
Pav.) Pers. tournefortiopsis Standi.
Tontanea glabra (Bartl. ex DC.) Standi., J.
Wash. Acad. Sci. 15:104. 1925. Triodon A. DC, Prodr. 4: 566. 1830. Not
Coccocypselum glabrum Bartl. ex DC, Prodr. Triodon Baumgartner, 1816. Type: Not
4: 397. 1830. Type: PANAMA: Without designated. [Herein treated as a synonym of
precise locality, Haenke s.n. (Holotype Galianthe Griseb.]
presumably PR, n.v., fragment G-DC, n.v. Triodon angulation Benth., PI. Hartweg. 70.
(microfiche 4: 397.14)).—Coccocypselum 1840. Type: MEXICO: Oaxaca, Teotolcingo
hirsutum Bartl. ex DC. var. glabrum (Bartl. in montibus Chinantla, T. Hartweg 507
ex DC.) L. O. Williams (Holotype K, n.v.). Diodia.—Galianthe
Totanea guianensis Aubl., Hist. PI. Guiane 1: brasiliensis (Spreng.) E. L. Cabral &
108, tab. 42. 1775. Type: FRENCH Bacigalupo subsp. angulata (Benth.) E. L.
GUIANA: In sylvis Cayenne, Aublet s.n. Cabral & Bacigalupo
(Holotype presumably BM or P,
n.v).—Coccocypselum guianense (Aubl.) K. Uncana Schreb., Gen. 125. 1789. Norn. cons.
Schum. in Mart. Type: Uncaria guianensis (Aubl.) J. F. Gmel.
(Basionym: Ourouparia guianensis Aubl.)
 i NOMENCLATOR OF MEXICAN AND CENTRAL

Uncaria tomentosa (Willd. ex Roem. & Monte Verde, Jan-Jul 1859, Wright 242
Schult.) DC, Prodr. 4: 349. 1830. Nauclea (Lectotype GH, n.v.; Isolectotype BR, n.v.;
tomentosa Willd. ex Roem. & Schult., Syst. designated by Hamilton, 1989).—Psychotria
Veg. 5: 221. 1819. Type: COLOMBIA: In costivenia Griseb.
ripa flumen Magdalenae prope Naers, Uragoga crocea (Sw.) Kuntze, Rev. Gen. PI. 1:
Humboldt s.n. (Holotype B-WI 3910, n.v.). 299. 1891. Psychotria crocea Sw., Prodr.
Veg. Ind. Occ. 44. 1788. Type: JAMAICA:
Uragoga Baill., Adansonia 12: 323. 1879. Type: Swartzs.n. (Holotype BM, n.v.).—Palicourea
Not designated. crocea (Sw.) Roem. & Schult.
Uragoga carpinterensis Kuntze, Rev. Gen. PI. Uragoga emetica (L. f) Baill., Hist. PI. 7: 371.
2: 955. 1891. Based on Palicourea lanceolata 1880. Psychotria emetica L. f, Suppl. PI. 144.
Oerst. ex Pol., Vidensk. Meddel. Dansk 1781. Type: COLOMBIA: America
Naturhist. Foren. Kjobenhavn 1852: 39. 1853, septentrionalis provinciae Gironensis, Mutis
pro syn. Type: COSTA RICA: San Jose, in 1602 (Holotype LINN, n.v.; probable Isotype
Monte Candelaria, 2100 m, Oersted 11556 US (photo PTBG)).—Psychotria emetica L.
(Holotype C, n.v. (photo and fragment F)). f.
—Palicourea angustifolia Kunth Uragoga erythrocarpa (Schltdl.) Kuntze. Rev.
Uragoga carthagenensis (Jacq.) Kuntze, Rev. Gen. PI. 2: 960. 1891. Psychotria
Gen. PI. 2: 959. 1891. Psychotria erythrocarpa Schltdl., Linnaea 9: 595. 1834.
carthagenensis Jacq., Enum. Syst. PI. Carib. Type: MEXICO: Veracruz, Hacienda de la
16. 1760. Type: COLOMBIA: Bolivar, Laguna, Schiede & Deppe 385 (Holotype B,
vicinity of Cartagena, Jacquin s.n. (location presumably destroyed (photo & fragment F);
unknown).—Psychotria carthagenensis Isotype K, n.v )—Psychotria erythrocarpa
Schltdl.
Uragoga cordillerae Kuntze, Rev. Gen. PI. 2: Uragoga galeottiana (M. Martens) Kuntze, Rev.
955. 1891. Based on Palicourea mexicana Gen. PI. 2: 960. 1891. Palicourea galeottiana
Benth. in Oerst., Vidensk. Dansk Meddel. M. Martens, Bull. Acad. Roy. Sci. Bruxelles
Naturhist. Foren. Kjobenhavn 1852: 37. 1853. 14: 136. 1844. Type: MEXICO: Oaxaca,
Not Uragoga mexicana (Willd ex Roem. & Cordillera, 6500 ft., 1840, H. Galeotti 2602
Schult.) Kuntze, 1891. Type: COSTA RICA: (Lectotype BR (photos MEXU, PTBG):
In monte Barba, Oersted 11559 (Lectotype C, Isolectotypes K, n.v., P (photos MEXU,
n.v.; designated by Standley 1938). PTBG); designated by Taylor & Lorence,
—Palicourea padifolia (Willd. ex Roem. & 14S5).
Schult.) C. M. Taylor & Lorence Martens) C. M. Taylor & Lorence
Uragoga costaricensis (Benth.) Kuntze, Rev. Uragoga graciliflora (Benth.) Kuntze, Rev.
Gen. PI. 2: 960. 1891. Palicourea Gen. PI. 2: 960. 1891. Psychotria graciliflora
costaricensis Benth. in Oerst., Vidensk. Benth. in Oerst., Vidensk. Meddel. Dansk
Meddel. Dansk Naturhist. Foren. Kjobenhavn Naturhist. Foren. Kjobenhavn 1852: 35. 1853.
1852: 38. 1853. Type: COSTA RICA: Type: COSTA RICA: Alajuela, ad Naranjo,
Heredia, Volcan de Barba and Cerros de Apr 1847, Oersted 11611 (Holotype C
Candelaria, Oersted 11540 (Holotype C, n.v. (photos F, MEXU, PTBG); Isotype US
(photo F); Isotypes MICH, n.v., US, n.v.). (photos MEXU, PTBG)). Mapouria.
—Palicourea padifolia (Willd. ex Roem. & —Psychotria graciliflora Benth. in Oerst.
Schult.) C. M. Taylor & Lorence Uragoga granadensis (Benth.) Kuntze, Rev.
Uragoga costivenia (Griseb.) Kuntze, Rev. Gen. Gen. PI. 2: 960. 1891. Psychotria granadensis
PI. 2: 960. 1891. Psychotria costivenia Benth. in Oerst., Vidensk. Meddel. Dansk
Griseb., PI. Wright. Cuba Orient. 2: 508. Naturhist. Foren. Kjobenhavn 1852: 34. 1853.
1862. Type: CUBA: Cuba Orientali, prope Type: NICARAGUA: Granada, Oersted
 11612 (Holotype C (photos MEXU, C (photos MEXU, PTBG)).—Psychotria
PTBG)).—Psychotria nervosa Sw. marginata Sw.
Uragoga grandis (Sw.) Kuntze, Rev. Gen. Pi. 2: Uragoga mexicana (Willd. ex Roem. & Schult.)
960. 1891. Psychotria grandis Sw., Prodr. Kuntze, Rev. Gen. PL 2: 961. 1891.
Veg. Ind. Occ. 43. 1788. Type: JAMAICA: Psychotria mexicana Willd. ex Roem. &
Interioris occidentalis, Swartz s.n. (Holotype Schult., Syst. Veg. 5: 189. 1819. Type:
S, n.v. (photo A)).—Psychotria grandis Sw. MEXICO: Without precise locality, Humboldt
Uragoga horizontalis (Sw.) Kuntze, Rev. Gen. & Bonplands.n. (Holotype B-WI, n.v.).
PL 1: 300. 189\. Psychotria horizontalis Sw., —Palicourea padifolia (Willd. ex Roem. &
Prodr. Veg. Ind. Occ. 44. 1788. Type: Schult.) C. M. Taylor & Lorence
SANTO DOMINGO: Without precise Uragoga micrantha (Kunth) Kuntze, Rev. Gen.
locality, Swartz s.n. (Holotype S, n.v. (photo PL 2: 961. 1891. Psychotria micrantha Kimtli
A)).—Psychotria horizontalis Sw. in Humb., Bonpl. & Kunth, Nov. Gen. Sp. PL
Uragoga ipecacuanha (Brot.) Baill., Hist. PI. 7: 3: 363, pi. 284. 1819. Type: PERU?: Without
370. 1880. Calicocca ipecacuanha Brot., precise locality, A. Bonplands.n. (Holotype P,
Trans. Linn. Soc. 6: 137. 1802. Type: n.v. (fragment F)). Mapouria.—Psychotria
BRAZIL: Without precise locality, Gomes s.n. micrantha Kunth in Humb., Bonpl. & Kunth
(presumably LISU, n.v.). Cephaelia, Evea. Uragoga microdesmia (Oerst.) Kuntze, Rev.
—Psychotria ipecacuanha (Brot.) Stokes Gen.PL2:961.1891.Psyc/^na
Uragoga justicioides (Schltdl.) Kuntze, Rev. microdesmia Oerst., Vidensk. Meddel. Dansk
Gen. PL 1: 300. 1891. Psychotria justicioides Naturhist. Foren. Kjobenhavn 1852: 36. 1853.
Schltdl., Linnaea 9: 596. 1834. Type: Type: COSTA RICA: Ad Jaris, Oersted
MEXICO: Barranca de Tioselo, Schiede & 11610 (Holotype C (photos MEXU, MO,
Deppe 259 (Holotype presumably HAL, n.v.; PTBG); Isotypes C (photos MEXU, PTBG),
Isotypes B, presumably destroyed (photo US (photo PTBG)).—Psychotria viridis Ruiz
MO), P (photos MEXU, PTBG)). & Pavon
—Psychotria hebeclada DC. Uragoga miradorensis (Oerst.) Kuntze, Rev.
Uragoga longicollis (Benth.) Kuntze, Rev. Gen. Gen. PL 2: 961. 1891. Mapouria miradorensis
PI. 2: 961. 1891. Psychotria longicollis Benth. Oerst., Amer. Centr. 17, tab. 14, fig- 9. 1863.
in Oerst., Vidensk. Meddel. Dansk Naturhist. Type: MEXICO: Veracruz, Mirador, in
Foren. Kjobenhavn 1852: 33. 1853. Type: quercetum, 3000 ft., F. Liebmann 11662
COSTA RICA: Insula San Lucas, Oersted (Lectotype C (photos MEXU, PTBG);
11621 (Lectotype C, here designated (photos Isolectotype US, n.v.; designated by
MEXU, PTBG); Isolectotype C, lower right- Hamilton, 1989).—Psychotria costivenia
hand collection, mounted on same sheet as Griseb. var. costivenia
syntype Oersted 11622). Syntypes: COSTA Uragoga morae (Pol.) Kuntze, Rev. Gen. PL 2:
RICA: In monte Aguacate, Nov 1846, Oersted 961. 1891. Psychotria morae Pol., Linnaea
11622 (C (photos MEXU, PTBG)); Aguacate, 41: 570. 1877. Type: COSTA RICA: San
Oersted 11629 (C (photos MEXU, PTBG)). Jose, Jun 1875, H. Polakowski 171 (Holotype
Oersted 11621, with two leafy fruiting B, presumably destroyed (photos GH, US)).
branches, is in excellent condition and —Psychotria quinqueradiata Pol.
represents the best choice for lectotype. Uragoga nicaraguensis (Benth.) Kuntze, Rev.
—Psychotria horizontalis Sw. Gen. PL 2: 957. 1891. Psychotria
Uragoga marginata (Sw.) Kuntze, Rev. Gen. PL nicaraguensis Benth. in Oerst., Viddensk.
2: 961. 1891. Psychotria marginata Sw., Meddel. Dansk Naturhist. Foren. Kjobenhavn
Prodr. Veg. Ind. Occ. 43. 1788. Type: 1852: 34. 1853. Type: NICARAGUA: Rio
JAMAICA: In sylvestribus, Swartz s.n. San Juan, Oersted 11634 (Lectotype C, here
(Holotype S, n.v. (photo A); probable Isotype designated (photos MEXU, PTBG)). Syntype:
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE

COSTA RICA: Ad Jaris, Oersted 11635 (C "Xalapa," photos MEXU, PTBG)).

(photos MEXU, PTBG)). Oersted 11634 is —Psychotria tenuifolia Sw.
selected as lectotype as it is the most complete Uragoga tenuifolia (Sw.) Kuntze, Rev. Gen. PI.
collection, with leaves, stipules, and a mature 2: 963. 1891. Psychotria tenuifolia Sw.,
infructescence.—Psychotria marginata Sw. Prodr. Veg. Ind. Occ. 43. 1788. Type:
Uragoga oerstediana Kuntze, Rev. Gen. PI. 2: SANTO DOMINGO: According to Howard
957. 1891, nomen superfl Based on (1989), a neotype needs to be designated.
Mapouria obovata Oerst., Amer. Centr. 17, —Psychotria tenuifolia Sw.
tab. 14, figs. 3,4. 1863. Not Psychotria Uragoga tomentosa (Aubl.) K. Schum., Nat.
obovata Ruiz & Pav., 1799. Type: MEXICO: Pflanzenfam. 4(4): 120. 1891. Tapogomea
Veracruz, F. Liebmann s.n. (Holotype C, tomentosa Aubl., Hist. PI. Guian. 160,/?/. 61.
n.v.). Psychotria.—Psychotria 1775. Type: FRENCH GUIANA: In sylvarum
quinqueradiata Pol. locis apertis, Aublet s.n. (presumably P,
Uragoga papantlensis (Oerst.) Kuntze, Rev. n.v).—Psychotria poeppigiana Mull. Arg.
Gen. PI. 2: 962. 1891. Mapouria papantlensis Uragoga trichotoma (M. Martens & Galeotti)
Oerst., Amer. Centr. 17, tab. 14, fig. 5. 1863. Kuntze, Rev. Gen. PI. 2: 963. 1891.
Type: MEXICO: Veracruz, Papantla, F. Psychotria trichotoma M. Martens & Galeotti,
Liebmann s.n. (Holotype presumably C, n.v.). Bull. Acad. Roy. Sci. Bruxelles 11: 227. 1844.
—Psychotria papantlensis (Oerst.) Hemsl. Type: MEXICO: Veracruz, Xalapa et
Uragoga parasitica (Sw.) M. Gomez, Anal. Mirador, 4000 pieds, H. Galeotti 7092
Soc. Esp. Hist. Nat. 3: 293. 1894. Psychotria (Holotype presumably BR, n.v.; Isotype P
parasitica Sw., nom. illegit. For discussion, (photos MEXU, PTBG)).—Psychotria
see Steyermark, 1972: 569.—Psychotria trichotoma M. Martens & Galeotti
guadalupensis (DC.) Howard Uginosa (Sw.) Kuntze, Rev. Gen. PI.
Uragoga phoeniciana K. Schum. in Engl. & 2: 963. 1891. Psychotria uliginosa Sw., Prodr.
Prantl, Naturl. Pflanzenfam. 4(4): 120. 1891. Veg. Ind. Occ. 43. 1788. Type: JAMAICA.
Type: COSTA RICA: Suerre, Llanuras de According to Howard (1989), a neotype needs
Santa Clara, 300 m, Feb 1896, J. Donnell to be designated.—Psychotria uliginosa Sw.
Smith 6584 (Syntype US, n.v.); same locality, Uragoga viridis (Ruiz & Pav.) Kuntze, Rev.
J. Donnell Smith 6586 (Syntype US, n.v.) Gen. PI. 2: 963. 1891. Psychotria viridis Rui/
Cephaelis.—Status uncertain, probably & Pav., Fl. Peruv. 2: 61. tab. 210, fig. b. 1799.
Psychotria sp. Type: PERU: Andes de Peru, Chinchao et
Uragoga quinqueradiata (Pol.) Kuntze, Rev. Macora, Jul 1894, Ruiz Lopez & Pavdn s.n.
Gen. PI. 2: 962. 1891. Psychotria (Holotype MA, n.v. (photo MO)).
quinqueradiata Pol., Linnaea 41: 570. 1877. —Psychotria viridis Ruiz & Pav.
Type: COSTA RICA: San Jose, 24 Jun 1875,
H. Polakowski 135 (Lectotype B, presumably Valantia L., Sp. PI. 2: 1051. 1753. Lectotype:
destroyed (photos BH, MO, US); designated Valantia muralis L.
by Hamilton, 1989).—Psychotria Wdantia hypocarpia L . Syst. Nat., ed. 10, 1307.
quinqueradiata Pol. 1759. Type: JAMAICA: Rubia I, Browne,
Uragoga sessilifolia (M. Martens & Galeotti) Hist. Jamaica 141 (Holotype LINN, n.v.).
Kuntze, Rev. Gen. PI. 2: 962. 1891.
Psychotria sessilifolia M. Martens & Galeotti, Endl. ex Gnseb.
Bull. Acad. Roy. Sci. Bruxelles 11: 228. 1844.
Type: MEXICO: Veracruz, Mirador et Vangueria A. L. Juss., Gen. PI. 206. 1789. Type:
Zacuapan, 3000 pieds, H. Galeotti 7078 Vangueria madagascariensis J. F. Gmel.
(Holotype presumably BR, n.v.; Isotypes G- Vangueria madagascariensis J. F. Gmel.. Syst.
DEL, n.v. (photo MO), P (label states Nat., ed. 13,2:367. 1791. Type:
 MASCARENE ISLANDS: Mauritius et Watsonamra alfaroana Standi., J. Wash. Acad.
Bourbon, 1770-1780, P. Commerson s.n. Sci. 15: 287. 1925. Type: COSTA RICA:
(Holotype P-JU, n.v.). Occasionally cultivated Alajuela, vicinity of Capulin, on the Rio
in tropical America and sparingly naturalized Grande de Tarcoles, 80 m, 2 Apr 1924, P. C.
at least in Costa Rica. The name Vangueria Standley 40194 (Holotype US 1152712
edulis Lam., a synonym, has sometimes been (photos MEXU, PTBG)). Pentagonia.
applied to collections from the Neotropics. —Pentagonia tinajita Seem.
Watsonamra brachyotis Stand., Contr. U.S.
Varneria L. Type: Varneria augusta L., Amoen. Natl. Herb. 17: 443. 1914. Type: PANAMA:
Acad. 4: 136. 1759. [Herein treated as a Danen, Marraganti, 3 Apr 1908, R. S.
synonym of Gardenia L.] Williams 999 (Holotype US 678351 (photos
Varneria augusta L., Amoen. Acad. 4: 136. MEXU, PTBG)).—Pentagonia brachyotis
1759. Type: Not cited. According to Smith (Standi.) Standi.
(1974), the description and illustration may be Watsonamra donnell-smithii Standi., Contr. U.S.
taken as the type.—Gardenia augusta (L.) Natl. Herb. 17: 442. 1914. Type: COSTA
RICA: La Emilia, Llanuras de Santa Clara,
250 m, Apr 1896, J. D. Smith 6590 (Holotype
Virecta L. f, Suppl. 17, 134. 1781. Type: US 355178 (photos MEXU, PTBG)).
Virecta biflora L. f. [Herein treated as a i (Standi.)
synonym of Sipanea Aubl.]
Virecta biflora L. f, Suppl. 17, 134. 1781. Watsonamra gymnopoda Standi., Contr. U.S.
Type: SURINAM: Without precise locality, Natl. Herb. 17: 444. 1914. Type: PANAMA:
Dalbergs.n. (presumably LINN, n.v.). Colon, Loma de la Gloria, near Fato (Nombre
—Sipanea biflora (L. f) Cham. & Schltdl. de Dios), 10-104 m, 8 Jul 1911, H. Pittier
3858 (Holotype US 678935 (photos MEXU,
Warszewiczia Klotzsch, Flora 36: 716. 1853; PTBG); Isotype US 678934 (photos MEXU,
Monatsber. Konigl. Preuss. Akad. Wissensch. PTBG)).—Pentagonia gymnopoda (Standi.)
Berlin 1853: 496. 1853. Type: Warszewiczia Standi.
coccinea (A. DC.) Klotzsch (Basionym: Watsonamra macrophylla (Benth.) Kuntze, Rev.
i Vahl.) Gen. PI. 1: 302. 1891. Pentagonia
i (Vahl) Klotzsch, macrophylla Benth., Bot. Voy. Sulphur 105,
Monatsber. Konigl. Preuss. Akad. Wissensch. tab. 39. 1845. Type: PANAMA: Panama,
Berlin 1853: 497. 1853. Macrocnemum forests between Panama and Cruces, Barclay
coccineum Vahl, Symb. Bot. 2: 38. 1791. s.n. (Holotype BM, n.v.).—Pentagonia
Type: TRINIDAD: Without precise locality, macrophylla Benth.
von Rohr (presumably C, n.v.). Watsonamra pittieri Standi., Contr. U.S. Natl.
Warszewiczia pulcherrima Klotzsch, Monatsber. Herb. 17: 443. 1914. Type: PANAMA: San
Konigl. Preuss. Akad. Wissensch. Berlin Bias, forests around Puerto Obladia, San Bias
1853: 497. 1853. Type: PANAMA: Veraguas, coast, 0-50 m, Aug 1911, H. Pittier 4298
von Warszewicz (Holotype B, presumably (Holotype US 679414 (photos MEXU,
destroyed).—Warszewiczia coccinea (Vahl) PTBG); Isotype US 679415 (photos MEXU,
Klotzsch PTBG)). Pentagonia.—Pentagonia
pinnatifida Seem.
Watsonamra Kuntze, Rev. Gen. PI. 1: 302. Watsonamra pubescens Standi., Contr. U.S.
1891. Type: Watsonamra macrophylla Natl. Herb. 17: 441. 1914. Type: PANAMA:
(Benth.) Kuntze [Herein treated as a synonym Canal Zone, around Tabernilla, along the
of Pentagonia Benth.] railroad, 20-25 m, 6 Jul 1911, H. Pittier 3822
 L NOMENCLATOR OF MEXICA

(Holotype US 678894 (photo PTBG); Isotype Xerococcus Oerst., Vidensk. Meddel. Dansk.
NY, n.v.).—Pentagonia macrophylla Benth. Naturhist. Foren. Kjobenhavn 1852: 52. 1853.
Watsonamra wendlandii (Hook.) Kuntze, Rev. Type: Xerococcus congestus Oerst. [Herein
Gen. 302. 1891. Pentagonia wendlandii treated as a synonym of Hoffmannia Sw.]
Hook., Bot. Mag./?/. 5230. 1861. Type: Xerococcus congestus Oerst., Vidensk. Meddel.
Described from cultivated material, probably Dansk. Naturhist. Foren. Kjobenhavn 1852:
from Costa Rica or Panama. Type not 52. 1853. Type: COSTA RICA: Turrialba,
designated.—Pentagonia wendlandii Hook. 3000 ft., Oersted 11776 (Holotype C, n.v.
(photos F, MO); Isotype K, n.v.).
Wittmackanthus Kuntze, Rev. Gen. PI. 1: 302. —Hoffmannia congesta (Oerst.) Dwyer
1891. Type: Wittmackanthus stanleyanus
(R. H. Schomb.) Kuntze (Basionym:
Calycophyllum Stanleyanum R. H. Schomb.)
Wittmackanthus stanleyanus (R. H. Schomb.)
Kuntze, Rev. Gen. PI. 1:302. 1891.
vanum R. H. Schomb., J.
Bot. Hook. 3: 622. 1844. Type: GUYANA:
Without precise locality, Schomburgk 356/411
B (Holotype K, n.v. (photo MO); Isotype NY,