Professional Documents
Culture Documents
of Mexican
and
Central
American
Rubiaceae
David H. Lorence
AUTHOR'S ADDRESS
David H. Lorence
National Tropical Botanical Garden
P.O. Box 340
Lawai, Hawaii 96765
VOLUME EDITOR
Marshall R. Crosby
MANAGING EDITOR
Amy Scheuler McPherson
Introduction
Format
Acknowledgments
References Cited
INTRODUCTION
FORMAT
At the beginning of each generic listing the name of each genus and its type species are always written
in normal Roman type. Typifications for these generic names mostly follow Index Nominwn Generic orum
iPLintarum) (Farr et al., 1979).
For each entry, an infrageneric name written in boldface type is accepted as valid, whereas an italicized
name is considered to be a synonym of the name in boldface type preceded by an em dash (—) at the end
of each entry. Each name is cross-referenced with its basionym, which is also italicized. Blocking names
have been indicated in cases where generic transfers have necessitated new names. Other generic
combinations (homotypic synonyms) that appear in this catalog are indicated by the generic name(s) listed
in italics just preceding the accepted name in boldface at the end of each citation. The following examples
illustrate these points.
(1)The accepted name (boldface type) is followed by the author(s) and publication citation, the basionym
(italics) its publication citation, and type information; this is followed by generic names (also in italics)
under which other combinations (=homotypic synonyms) are listed in this catalog:
Elaeagia myriantha (Standi.) C. M. Taylor & Hammel, Novon 3:490.1993. Sickingia myriantha Standi.,
Publ. Field Columbian Mus., Bot. Ser. 7: 27. 1930. Type: COLOMBIA: Magdalena, Las Nubes road,
region of Santa Marta, 1200 m, 3 Dec 1898, H. H. Smith 1810 (Holotype F 138660; isotypes MO, NY,
n.v., US, n.v., W, n.v., Y, n.v.). Holtonia, Simira.
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIA
(2) A homotypic synonym (italics) is followed by its publication citation, the basionym (italics) and its
publication citation and type information, and another genus under which a homotypic synonym exists
(italics). Finally, this is cross-referenced to the accepted name (boldface):
Holtonia myriantha (Standi.) Standi., Trop. Woods 30: 37. 1932. Sickingia myriantha Standi., Publ. Field
Columbian Mus., Bot. Ser. 7: 27. 1930. Type: COLOMBIA: Magdalena, Las Nubes road, region of
Santa Marta, 1200 m, 3 Dec 1898, H. H. Smith 1810 (Holotype F 138660; isotypes MO, NY, n.v., US,
n.v., W, n.v., Y, n.v.). Simira — Elaeagia myriantha (Standi.) C. M. Taylor & Hammel
ACKNOWLEDGMENTS
I sincerely thank the following individuals and institutions whose assistance has been instrumental in
completing this work: John Dwyer for inspiring this nomenclator and for making available to me notes
on the types he examined at BM and K; C. Dennis Adams (BM) for welcome advice and assistance with
Spermacoceae; Tim McDowell for information regarding Exostema and Syringantha; Robert M. Rhode
for information regarding Chiococca; Marshall R. Crosby, Gerrit Davidse, and Amy McPherson for
helpful discussions, comments, and suggestions; Lennart Andersson and Charlotte Taylor for their detailed
reviews, which substantially improved the manuscript; Janet Leopold and Teri Teasdale for help with
often tedious manuscript preparation; Rick Hanna for locating many obscure references; and Barbara
Mack for updating and formatting the manuscript for publication. I also thank the curators of the herbaria
cited within for loaning specimens and for providing work space and access to their collections. During
my tenure as Investigador at the Herbario Nacional de Mexico (MEXU), the Instituto de Biologia of the
Universidad Nacional Autonoma de Mexico provided financial support, enabling me to study and
photograph types of Mexican and Central American Rubiaceae at the F, MO, P, and US herbaria.
Additional financial support enabling me to study and photograph types at P was provided by a grant from
CONACYT (Mexico). The Missouri Botanical Garden provided facilities and support during my many
visits there. Finally, I am grateful to William L. Theobald and the late William McK. Klein, Jr., former
directors of the National Tropical Botanical Garden, for supporting various phases of work on this project.
My sincere gratitude is extended to all who have helped me.
REFERENCES CITED
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1-128.
Andersson, L. 1992. A Provisional Checklist of Neotropical Rubiaceae. Scripta Botanica Belgica 1:
1-199.
Andersson, L. & C. M. Taylor. 1994. Rubiaceae-Cinchoneae-Coptosapelteae. In G. Harling & L.
Andersson (editors), Flora of Ecuador 50. No. 162 (1-4). Berlings, Arldv, Sweden.
Blackwell, W. FL, Jr. 1968. Revision of Bouvardia (Rubiaceae). Ann. Missouri Bot. Gard. 55: 1-30.
Boom, B. M. 1984. A revision of Isertia (Isertieae: Rubiaceae). Brittonia 36: 425^54.
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Bridson, G D. R. & E. R. Smith. 1991. Botanico-Periodicum-Huntianum/Supplementum. Pittsburgh: Hunt
Brummitt, R. K. & C. E. Powell. 1992. Authors of Plant Names. Kew: Royal Botanic Gardens.
Burger, W. C. & C. M. Taylor. 1993. Family # 202 Rubiaceae. In W. C. Burger (editor), Flora
Costaricensis. Fieldiana, Bot., n.s. 33: 1-333.
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of Mexico and Southwestern United States. Univ. Calif. Publ. Bot. 64: 1-36.
Dempster, L. T. 1973b. Note on the type of Rubia acuminata Mart. & Gal. Brittonia 25: 313.
Dempster, L. T. 1976. Galium mexicanum of Central American and western North America. Madrono 23:
378-386.
Dempster, L. T. 1978. The genus Galium (Rubiaceae) in Mexico and Central America. Univ. Calif. Publ.
Bot. 73: 1-33.
Dwyer, J. D. 1980. Rubiaceae. In R. E. Woodson, Jr., R. W. Schery & collaborators, Flora of Panama-Part
IX. Ann. Missouri Bot. Gard. 67: 1-522.
Elias, T. S. 1976. A monograph of the genus Hamelia (Rubiaceae). Mem. New York Bot. Gard. 26:
81-144.
Farr, E. R., J. A. Leussink & F. A. Stafleu. 1979. Index Nominum Genericorum (Plantarum). 3 vols. The
Hague: Bohn, Scheltema & Holkema.
Fosberg, F. R. 1974. Studies in American Rubiaceae 2. Ayuque, Balmea stormae, an endangered Mexican
species. Sida 5: 268-270.
Fosberg, F. R. & M.-H. Sachet. 1989a. Three cultivated ixoras (Rubiaceae). Baileya 23: 74-85.
Fosberg, F. R. & M.-H. Sachet. 1989b. Lectotypification of Ixora coccinea L. (Rubiaceae). Taxon 38:
McDowell, T. 1996b. Exostema (Rubiaceae): Taxonomic history, nomenclature, position and subgeneric
classification. Opera Bot. Belg. 7: 277-295.
Mena V., P. 1986. A revision of the genus Arcytophyllum (Rubiaceae: Hedyotideae). Mem. New York
Bot. Gard. 60: 1-26.
Natali, A. 1993. In C. E. Jarvis, F. R. Barrie, D. M. Allan & J. L. Reveal, A List of Linnaean Generic
Names and their Types. Regnum Vegetabile 127. [Typification of Sherardia an>ensis p. 88.]
Nicolson, D. 1977. Notes for the flora of Dominica. Spermacoce confusa and Schradera exotica. J. Arnold
Arbor. 58: 445^47. [Typification of Spermacoce confusa.}
Puff, C. 1991. The genus Paederia L.: A multidisciplinary study. Opera Bot. Belg. 3: 1 376.
Rendle, A. B. 1934. Linnaean species of Spermacoce. J. Bot. 72: 329-333.
Rendle, A. B. 1936. Spermacoce remota Lam. J. Bot. 74: 10-12.
Reveal, J. 1993. In C. E. Jarvis, F. R. Barrie, D. M. Allan & J. L. Reveal, A List of Linnaean Generic
Names and their Types. Regnum Vegetabile 127. [Typification of Cephalanthus occidentalism p. 31.]
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mentioned in the first two editions of Linnaeus' Species plantarum. Huntia 7: 209 245.
[Lectotypification of Mitchella repens, p. 215.]
Rogers, G. K. 1987. The genera of Cinchonoideae (Rubiaceae) in the Southeastern United States. J.
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Schumann, K. 1891. Rubiaceae. In A. Engler & K. Prantl, Die natiirlichen Pflanzenfamilien IV (4): 1-156.
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Smith, A. C. 1974. Studies of Pacific Island plants. XXVII. The genus Gardenia (Rubiaceae) in the Fijian
region. Amer. J. Bot. 61: 109-128.
Stafleu, F. & R. Cowan. 1976-1988. Taxonomic Literature. Vols. 1-7. Utrecht: Bonn, Scheltema &
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Standley, P. C. 1926. Rubiaceae. In Trees and Shrubs of Mexico. Contr. U.S. Natl. Herb. 23: 1349-1394.
Standley, P. C. 1934. Rubiaceae. In North American Flora 32(3-4): 159-300.
Standley, P. C. 1938. Rubiaceae. In Flora of Costa Rica. Publ. Field Mus. Nat. Hist., Bot. Ser. 18:
1264-1380.
Standley, P. C. & L. O. Williams. 1975. Rubiaceae. In Flora of Guatemala. Fieldiana: Bot. 24(11): 1-274.
Steyermark, J. A. 1972. Rubiaceae, Part IX. In B. Maguire and collaborators, The Botany of the Guyana
Highland. Mem. New York Bot. Gard. 23: 227-832.
Steyermark, J. A. 1974. Rubiaceae. In Flora de Venezuela 9(1-3): 1-2070.
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Taylor, C. M. 1994. Revision of Hillia (Rubiaceae). Ann. Missouri Bot. Gard. 81: 571-609.
Taylor, C. M. & D. H. Lorence. 1985. Lectotypification of Palicourea galeottiana M. Martens
(Rubiaceae) and a new name for this common species. Taxon 34: 667-669.
Taylor, C. M. & D. H. Lorence. 1993. On the status of Randia armata (Sw.) DC. (Rubiaceae: Gardenieae).
Taxon 42: 865-867.
Terrell, E. E. 1980. Two new species and a new combination in Houstonia (Rubiaceae) from Mexico.
Brittonia 32: 490-494.
Terrell, E. E. 1996. Revision of Houstonia (Rubiaceae-Hedyotideae). Syst. Bot. Monogr. 48: 1-118.
Terrell, E. E. & W. H. Lewis. 1990. Oldenlandiopsis (Rubiaceae), a new genus from the Caribbean Basin,
based on Oldenlandia callitrichoides Grisebach. Brittonia 42: 185-190.
Verdcourt, B. 1975. Studies in the Rubiaceae-Rubioideae for the 'Flora of Tropical East Africa': I. Kew
Bull. 30: 247-326.
Verdcourt, B. 1976. In R. M. Polhill (editor), Flora of Tropical East Africa. Rubiaceae, Part 1: 1^14.
London: Crown Agents for Overseas Governments. [Lectotypifications for Oldenlandia corymbosa L.
on p. 308 and O. lancifolia on p. 292.]
Verdcourt, B. 1983. Notes on Mascarene Rubiaceae. Kew Bull. 37: 521-547. [Spermacoce typifications.]
M > \1 F \ ( LATURALINDEX
Aeginetia Cav., Anal. Ci. Nat. 3: 129. 1801. Alibertia premontana C. M. Taylor, Novon 6:
Type not designated. 215. 1996. Type: COSTA RICA: Alajuela,
Aeginetia longiflora Cav., Anal. Ci. Nat. 3: 130. Reserva Biologica Monteverde, Valle del rio
1801. Type: MEXICO: Between Queretaro Penas Blancas, fila del Toro, serdero Pipilacha
and Guanajuato, Oct, Nee (Holotype y Campo Tres, 18°N, 84°W, 900-1100 m, 21
presumably MA, n.v.).—Bouvardia Jan 1991, W. Haber & E. Cruz 10630
longiflora (Cav.) Kunth (Holotype MO 4234216; Isotype CR, n.v.).
-ieginetici multiflora Cav., Anal. Ci. Nat. 3: 131, Alibertia utleyorum (Dwyer) C. M. Taylor,
t. 28, fig. 2. 1801; Icon. 6: 52, t. 572, fig. 2. Novon 6: 215. 1996. Duroia utleyorum Dwyer
1801. Type: MEXICO: Queretaro, Nee s.n. in W. C. Burger & C. M. Taylor, Fieldiana
(Holotype MA, n.v. (photo MO)). Bot., n.s. 33: 127. 1993. Type: COSTA RICA:
—Bouvardia multiflora (Cav.) Schult. & Puntarenas, adjacent to airfield, Rincon de
Schult. f. Osa, 6-7 Feb 1974, R. Liesner 1774
(Holotype CR, n.v.; Isotype MO 3030387).
Aegiphila Jacq. [Verbenaceae]
K wphila hnjjmumuoides Standi. & Steyerm., s Standi, Ann. Missouri Bot. Gard.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 227. 27: 344. 1940. Type: Allenanthus
1947. Type: GUATEMALA: Huehuetenango, erythrocarpa Standi.
Steyermark 48843 (Holotype F, n.v.). Allenanthus erythrocarpus Standi., Ann.
—Hoffmannia nicotianifolia (M. Martens & Missouri Bot. Gard. 27: 344. 1940. Type:
Galeotti) L. O. Williams PANAMA: Code, Vicinity of El Valle de
Anton, ca. 600 m, 17 Sep 1939, P. H. Allen
Alibertia A. Rich., Mem. Soc. Hist. Nat. Paris 5: 1999 (Holotype MO 1195153 (photos MEXU,
234, t. 21, fig. 1. 1830. Type: Alibertia edulis PTBG); Isotypes F 1015649, GH, n.v., US
(Rich.) A. Rich, ex DC. (photos MEXU, PTBG)).
Alibertia edulis (Rich.) A. Rich, in DC, Prodr. Allenanthus hondurensis Standi., Ceiba 1: 45.
4: 443. 1830. Genipa edulis Rich., Act. Soc. 1950. Type: HONDURAS: Morazan, Faldas
Hist. Nat. Paris 1: 107. 1792. Type: FRENCH del Cerro Majicaran, aguas arriba del Rio de
GUIANA: Guyane, Richard s.n. (Holotype P, la Onlla, 950 m, 27 Nov 1948, A. Molina R.
n.v.; Isotype F 971706 (photo PTBG)). 1750 (Holotype F 152284 (photo PTBG);
Cordiera, Gardenia. Isotypes EAP, n.v., US (photos MEXU,
Alibertia garapatica K. Schum. in Mart., Fl. PTBG)).
Bras. 6(6): 384. 1889. Type not cited. Allenanthus hondurensis var. parvifolia L. O.
Alibertia longistiptdata L. Riley, Kew Bull. Williams, Phytologia 25: 461. 1973. Type:
1927: 122. 1927. Type: PANAMA: Panama, MEXICO: Chiapas, Mun. Venustiano
Old Panama, 16 Jun 1924, L. A. M. Riley 145 Carranza, Rancho Carmen along the road
(Holotype K, n.v.; Isotype US (photos from Acala to Venustiano Carranza, 2600 ft.,
MEXU, PTBG)).—Alibertia edulis (Rich.) 23 Aug 1966, R. M. Lauglin 1647 (Holotype F
A. Rich ex DC. 1662766 (photos MEXU, PTBG)).
Alibertia panamensis L. Riley, Kew Bull. 1927:
123. 1927. Type: PANAMA: Between Alseis Schott in Spreng., Syst. Veg. 4(2): 404.
Panama and Sabana, 14 Jun 1924, L. A. M. 1827. Type: Alseis floribunda Schott
Riley 111 (Holotype K, n.v.; Isotypes US, 2 Alseis blackiana Hemsl., Diag. PI. Mex. 30.
sheets (photos MEXU, PTBG)).—Alibertia 1879. Type: PANAMA: In dense woods,
edulis (Rich.) A. Rich, ex DC. Manma [?] Station, P. R. R., Jun 1862, S.
L NOMENCLATOR OF MEXICAN / ) CENTRAL AMERICAN RUBIACEAE
Hayes 665 (Holotype K, n.v.; Isotype P Agricola and Alto Piedra (above Santa Fe)
(photos MEXU, PTBG)). and Rio Dos Bocas ca. 5-8 km from Escuela,
Alseis dariensis Dwyer, Ann. Missouri Bot. 730-770 m, 26 Jul 1974, T. B. Croat 25937
Gard. 67: 27. 1980. Type: PANAMA: Darien, (Holotype MO 2212775 (photo PTBG);
Rio Tuquesa, middle Tuquesa Mining Co. Isotypes F (photo PTBG), GH, n.v., MO
camp called Charco Peje, 250 m, S. Mori 2212776 (photo PTBG)).
7001 (Holotype MO, not located 6/96).
—Wittmackanthus stanleyanus (Schomb.) Amphidaysa Standi., Field Mus. Nat. Hist., Bot.
Ser. 11: 180. 1936. Type: Amphidaysa
Alseis hondurensis Standi., Trop. Woods 16: ambigua (Standi.) Standi.
48. 1928. Type: HONDURAS: La Ceiba, Dec Amphidaysa ambigua (Standi.) Standi., Publ.
1927, H. Johansen 9 (Holotype F 572615 Field Columbian Mus., Bot. Ser. 11: 181.
(photo PTBG)). 1936. Sabicea ambigua Standi., Publ. Field.
Alseis schippii Lundell, Lloydia 4: 56. 1941. Mus. Nat. Hist., Bot. Ser. 7: 49. 1930. Type:
Type: BRITISH HONDURAS (BELIZE): COLOMBIA: Without precise locality, 1847,
Toledo Distr., Machaca, 50 ft., 4 Nov 1932, Triana s.n. (Holotype K, n.v. (photo F)).
W. A. Schipp 1230 (Holotype TEX-LL Amphidaysa spathulata Dwyer, Ann. Missouri
(photos MEXU, PTBG); Isotype F (photo Bot. Gard. 67: 33. 1980. Type: PANAMA:
PTBG)).—Alseis hondurensis Standi. Darien, Cerro Campamento, south of Cerro
Alseis yucatanensis Standi., Publ. Field Pire, cloud forest, 20-27 Mar 1968, J. A.
Columbian Mus., Bot. Ser. 8: 50. 1930. Type: Duke 15615 (Holotype MO 1964213 (photos
MEXICO: Yucatan, without precise locality, MEXU, PTBG)).
1917-1921, G. F. Gaumer 24247 (Holotype F
552253 (photo PTBG); Isotype US (photos Anisomeris C. Presl, Symb. Bot. 2: 5, t. 54.
MEXU, PTBG)). 1834. Type: Anisomeris spinosa C. Presl.
[Herein treated as a synonym of Chomelia
Amaioua Aubl., Hist. PI. Guiane 13: 1775. Jacq.]
Type: Amaioua guianensis Aubl. Anisomeris brachypoda (Donn. Sm.) Standi.,
Amaioua corymbosa Kunth, Nov. Gen. Sp. 3: Publ. Field Columbian Mus., Bot. Ser. 4: 293.
419, pi. 294. 1820. Type: VENEZUELA: In 1929. Chomelia brachypoda Donn. Sm., Bot.
locis frondosis Novae-Andalusia prope Gaz. (Crawfordsville) 47: 255. 1909. Type:
Cumanacoa, Humboldt & Bonpland s.n. GUATEMALA: Alta Verapaz, Sasis, banks of
(Holotype presumably P-Bonpland, n.v.; Rio Ogeqaj, 900 m, May 1908, H von
Isotype B-W (photo MO)). Tuerckheim 2253 (Holotype US; Isotypes F
Amaioua hirsuta Poepp. & Endl., Nov. Gen. Sp. (photos MEXU, PTBG), US).—Chomelia
PI. 3: 25, pi. 230 1841. Type: Chili, Perou, et brachypoda Donn. Sm.
Fleuve de La Plata, 1836, Poeppig 2823 Anisomeris englesingii (Standi.) Standi., Publ.
(Holotype W, n.v., fragment F; Isotype F Field Columbian Mus., Bot. Ser. 8: 360. 1931.
(photo PTBG)).—Duroia hirsuta (Poepp. & Chomelia englesingii Standi., Trop. Woods
Endl.) K. Schum. 16: 45. 1928. Type: NICARAGUA: Region of
Amaioua magnicarpa Dwyer, Ann. Missouri Braggman's Bluff (Kukalaya River), 200-600
Bot. Gard. 67: 29. 1980. Type: PANAMA: ft., 2 Dec 1927, F. C Englesing 49 (Holotype
Colon, Santa Rita Ridge, 23 Mar 1972, J. D. F 572606 (photo PTBG)).—Chomelia
Dwyer 9552 (Holotype MO 2162079 (photo recordii Jacq.
PTBG); Isotype MO 4199142 (photo PTBG)). Anisomeris microloba (Donn. Sm.) Standi.,
Amaioua pedicellata Dwyer, Ann. Missouri Publ. Field Columbian Mus., Bot. Ser. 4: 293.
Bot. Gard. 67: 30. 1980. Type: PANAMA: 1929. Chomelia microloba Donn. Sm., Bot.
Veraguas, along road between Escuela Gaz. (Crawfordsville) 31: 114. 1901. Type:
COSTA RICA: Forets de Santo Domingo de
Golfo Dulce, Feb 1896, A. Tonduz 9874
(Holotype US; Isotypes F, MO (photos Standley & J. Valerio 49196 (Holotype US
MEXU, PTBG), P (photo MEXU), US, 3 1305902).—Chione sylvicola (Standi.) W. C.
sheets).—Chomelia microloba Donn. Sm.
Anisomeris pringlei (S. Watson) Standi., Publ.
Field Columbian Mus., Bot. Ser. 4: 293. 1929. Anotis DC, Prodr. 4: 431. 1830. Type not
Chomelia pringlei S. Watson, Proc. Amer. designated.
Acad. Sci. 26: 137. 1891. Type: MEXICO: Anotis cervantesii (Kunth) DC, Prodr. 4: 432.
San Luis Potosi, Tamasopo Canyon, 6 Aug 1830. Hedyotis cervantesii Kunth in Humb.,
1890, C. G. Pringle 3209 (Holotype US Bonpl. & Kunth, Nov. Gen. Sp. 3: 390. 1820.
48535; Isotypes F (photos MEXU, PTBG), Type: MEXICO: Without precise locality, A.
MO (photos MEXU, PTBG), P (photos Bonplands.n. (Holotype P-Bonpland, n.v.;
MEXU, PTBG)).—Chomelia pringlei S. Isotype B-W (photo MO)).—Hedyotis
Watson cervantesii Kunth
Anisomerisprotracta (Bartl. ex DC.) Standi., Anotis longiflora Benth., PI. Hartw. 23. 1839.
Publ. Field Columbian Mus., Bot. Ser. 4: 293. Type: MEXICO: Without precise locality,
1920. Guettarda protracta Bartl. ex DC, 1837, T. K. Hartweg 206 (Holotype K, n.v.;
Prodr. 4: 457. 1830. Type: MEXICO: Without Isotype GH, n.v.). Bouvardia.—Bouvardia
precise locality, Haenkes.n. (Holotype multiflora (Cav.) Schult. & Schult. f.
presumably PR, n.v.).—Chomelia protracta
(Bartl. ex DC.) Standi. Antirhea Comm. ex Juss., Gen. PL 204. 1789.
Anisomeris purpusii Brandegee, Univ. Calif. Type: Antirhea borbonica J. F. Gmelin
Publ. Bot. 6: 70. 1914. Type: MEXICO: Antirhea aromatica Cast-Campos & Lorence,
Chiapas, nearTonala, Sep 1913, C. A. Purpus Ann. Missouri Bot. Gard. 72: 268 Jig. I.
6940 (Holotype UC 173028, n.v.; Isotypes F 1985. Type: MEXICO: Veracruz, Mun.
(photo PTBG), US (photo PTBG)). Chomelia. Jalcomulco, ca. 10 km al S de Apazapan,
—Chomelia spinosa Jacq. Barranca de Monterrey, al S de Cuetzalan,
Anisomeris recordii (Standi.) Standi., N. Amer. entre Cuetzalan y Apazapan, 350 m, 17 Sep
Fl. 32: 227. 1934. Chomelia recordii Standi.. 1983, G. Castillo-Campos 2957 (Holotype
Trop. Woods 7: 9. 1926. Type: XAL; Isotypes MEXU, MO, XAL, plus about
GUATEMALA: Izabal, high forest between 50 more distributed to a wide range of
Los Andes and Entre Rios, lower Rio herbaria).
Motagua Valley, Record 8862 (Holotype Y, Antirhea lucida (Sw.) Benth. & Hook, f., Gen.
n.v.; Isotype NY, n.v.).—Chomelia recordii PI. 2: 100. 1873. Laugeria lucida Sw., Prodr.
Standi. Veg. Ind. Occ. 48. 1788. Type: JAMAICA:
Anisomeris rudis Standi., Field Mus. Nat. Hist., Without precise locality, Swartz s.n.
Bot. Ser. 17: 394. 1938. Type: HONDURAS: (Holotype presumably S, n.v.).
Comayagua, summit of ridge above El —Stenostomum lucidum (Sw.) C. F. Gaertn.
Achote, above the plains of Siguatepeque, Antirhea panamensis Standi., N. Amer. Fl. 32:
1800 m, 1 Aug 1936, T. G. Yunker, R. F. ' 264. 1934. Type: PANAMA: San Bias
Dawson & H. R. Youse 6253 (Holotype F District, 21-28 Apr 1933, G. P. Cooper 238
857678 (photo PTBG); Isotype MO (Holotype F 686345 (photo PTBG); Isotype F
1115471).—Chomelia rudis (Standi.) 709955). Chomelia.—Chomelia microloba
Donn. Sm.
Anisomeris sylvicola (Standi.) Standi., N. Amer. Antirhea trichantha (Griseb.) Hemsl., Biol.
Fl. 32: 225. 1934. Chomelia ? sylvicola Centr.-Amer., Bot. 2: 42. 1881. Pittoniotis
Standi., J. Wash. Acad. Sci. 18: 182. 1928. trichantha Griseb., Bonplandia 6: 8. 1858.
A NOMENCLATOR OF MEXICAN AND CENTRAL
Type: PANAMA: Without precise locality, m, 14 Jan 1905, W. R. Maxon & R. Hay 3275
Duchaissing s.n. (Holotype presumably (Holotype US 473244 (photos MEXU,
GOET or P, n.v.).—Pittoniotis trichantha PTBG); Isotype US (photos MEXU, PTBG)).
Griseb. —Rondeletia aetheocalymna Donn. Sm.
Arachnothryx albida (Lundell) Borhidi, Acta
Appunia Hook. f. in Benth. & Hook, f., Gen. PI. Bot. Hung. 35: 309. 1989. Rondeletia albida
2: 120. 1873. Type: Appunia tenuiflora Lundell, Wrightia 5: 323. 1976. Type:
(Benth.) Hook. f. MEXICO: Chiapas, Carelas, near Motozintla,
Appunia guatemalensis Donn. Sm., Bot. Gaz. 2176 m, Apr 1945, E. Matuda 5509 (Holotype
(Crawfordsville) 48: 294. 1909. Type: TEX-LL (photos MEXU, PTBG); Isotypes F,
GUATEMALA: Izabal, silva prope pagam MEXU, TEX-LL).—Rondeletia laniflora
martimam Livingston, Jun 1906, H. von Benth.
Tuerckheim IIf-230 (Holotype US 943501 Arachnothryx aspera (Standi.) Borhidi, Acta
(photos MEXU, PTBG)). Morinda. Bot. Acad. Sci. Hung. 28: 68. 1982.
Appunia parviflora Lundell, Contr. Univ. Rondeletia aspera Standi., N. Amer. Fl. 32:
Michigan Herb. 4: 28. 1940. Type: BRITISH 54. 1918. Type: COSTA RICA: San Jose,
HONDURAS (BELIZE): Stann Creek Distr.. Pont du Rodeo, Pacacol (Villa Colon), Jan
Silk Grass Creek Reserve, 5 Sep 1939, P. H. 1891, H. Pittier 3243 (Holotype US 941408
Gentle 2986 (Holotype MICH, n.v.; Isotypes (photo PTBG)).—Rondeletia aspera Standi.
A, n.v., F (photo PTBG), US (photos MEXU, Arachnothryx belizensis (Standi.) Borhidi, Acta
PTBG)).—Psychotria erecta (Alibi.) Standi. Bot. Acad. Sci. Hung. 28: 70. 1982.
& Steyerm. Rondeletia belizensis Standi., Carnegie Inst.
Appunia seibertii Standi., Ann. Missouri Bot. Wash. Publ. 461: 91. 1935. Type: BRITISH
Gard. 24: 208. 1937. Type: PANAMA: HONDURAS (BELIZE): Jasinto Hills, 300
r Rio ft., 28 Aug 1933, W. A. Schipp 1201
Trinidad, 26-50 m, 7-10 Aug 1935, R. J. (Holotype F 683600 (photos MEXU, PTBG);
Seibert 624 (Holotype MO 1110146 (photos Isotypes GH, n.v., K (photo PTBG), MO).
MEXU, PTBG); Isotypes A, n.v., F). —Rondeletia belizensis Standi.
Arachnothryx belizensis (Standi.) Borhidi var.
Arachnothryx Planch., Fl. Serres 5: tab. 442. longiloba (Lundell) Borhidi, Acta Bot. Hung.
1849. Type: Arachnothryx leucophylla 35: 310. 1989. Rondeletia belizensis Standi,
(Kunth) Planch. Type designated by Borhidi, var. longiloba Lundell, Wrightia 5: 323. 1976.
1982. [Herein treated as a synonym of Type: GUATEMALA: Alta Verapaz,
Rondeletia L.] Rubelsanto, 2.3 km ESE of Balastrera, 20 Jul
Arachnothryx acuminata (Oerst. ex Standi.) 1985, C. L. Lundell & E. Contreras 19545
Borhidi, Acta Bot. Hung. 35: 310. 1989 [as (Holotype TEX-LL (photos MEXU, PTBG);
acumunata]. Sommera acuminata Oerst. ex Isotypes F, MO (photo PTBG)).—Rondeletia
Standi., J. Wash. Acad. Sci. 17: 340. 1927. belizensis Standi.
Type: MEXICO: Oaxaca, Amatlan, F. Arachnothryx bertieroides (Standi.) Borhidi,
Liebmann 11712 (Holotype C (photo MEXU); Acta Bot. Acad. Sci. Hung. 28: 70. 1982.
Isotype US (photos MEXU, PTBG)). Rondeletia bertieroides Standi., Publ. Field
Rondeletia.—Rondeletia scabra Hemsl. Columbian Mus., Bot. Ser. 4: 267. 1929.
Arachnothryx aetheocalymna (Donn. Sm.) Type: PANAMA: Bocas del Toro, Buena
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 70. Vista Camp, Chinqui Trail, 900 m, 1928, G
1982. Rondeletia aetheocalymna Donn. Sm., P. Cooper 598 (Yale 12231) (Holotype F
Bot. Gaz. (Crawfordsville) 42: 298. 1906. 579614 (photo PTBG); Isotype NY, n.v.).
Type: GUATEMALA: Alta Verapaz, trail i bertieroides Standi.
between Sepacuite and Secanquim, 550-1000
Arachnothryx bourgaei (Standi.) Borhidi, Acta Arachnothryx calycosa (Donn. Sm.) Borhidi,
Bot. Acad. Sci. Hung. 28: 69. 1982. Acta Bot. Acad. Sci. Hung. 28: 70. 1982.
Rondeletia bourgaei Standi., N. Amer. Fl. 32: Rondeletia calycosa Donn. Sm., Bot. Gaz.
56. 1918. Type: MEXICO: Veracruz, region (Crawfordsville) 56: 59. 1913. Type: COSTA
de Orizaba, Monte San Cristobal, 8 Jul 1866, RICA: Forets de la Palma, 1460 m, 25 Sep
E. Bourgeau 2618 (Holotype US 49964 1898, A. Tonduz 11633 (Holotype US 941406
(photos MEXU, PTBG); Isotypes P, 4 sheets (photos F, MEXU, PTBG)).—Rondeletia
(photos MEXU, PTBG)).—Rondeletia calycosa Donn. Sm.
laniflora Benth. Arachnothryx capitellata (Hemsl.) Borhidi, Acta
| Standi.) Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
Bot. Hung. 33: 301. 1987. Rondeletia brenesii Rondeletia capitellata Hemsl., Diag. PI. Mex.
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. 28. 1879. Type: MEXICO: Without precise
18: 1370. 1938. Type: COSTA RICA: locality, Jurgensen 355 (Lectotype K, here
Camino de San Ramon a La Calera, 20 Mar designated (photo PTBG); Isolectotypes G-
1929, A. M. Brenes 6757 (Holotype F 854972 DEL (n.v., fragment F), K (photo PTBG)).
(photo PTBG)).—Rondeletia brenesii Standi. Syntypes: MEXICO: Oaxaca, Joya, F.
Arachnothryx brevicorolla (J. H. Kirkbr.) Liebmann 202 (K (photo PTBG); without
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69. precise locality, Jurgensen 3 76 (K (photo
1982 [as brt alk ifolia PTBG)).—Rondeletia capitellata Hemsl.
Dwyer & Hayden subsp. brevicorolla J. H. Arachnothryx chinajensis (Standi. & Steyerm.)
Kirkbr., Ann. Missouri Bot. Gard. 55: 383. Borhidi, Acta Bot. Hung. 33: 301. 1987.
1963. Type: PANAMA: Bocas del Toro, Rondeletia chinajensis Standi. & Steyerm.,
cloud forest above Quebrada Huron on Cerro Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.
Bonyik, 166^00 m, 13 Apr 1968, J. H. 1947. Type: GUATEMALA: Alta Verapaz,
Kirkbride, Jr. &J.A. Duke 597 (Holotype Cerro Chinaja, between Finca Yalpemech and
MO 1946832 (photo PTBG); Isotypes NY, Chinaja, above source of Rio San Diego,
n.v., US (photos MEXU, PTBG)). 150-700 m, 1-2 Apr 1942, J. A. Steyermark
—Rondeletia bertieroides Standi. 45637 (Holotype F 1134210 (photos MEXU,
Arachnothryx buddleioides (Benth.) Planch., Fl. PTBG)).—Rondeletia chinajensis Standi. &
Serres 5: tab. 442. 1849. Rondeletia Steyerm.
buddleioides Benth., PI. Hartweg. 69. 1840. Arachnothryx cooperi (Standi.) Borhidi, Acta
Type: MEXICO: Oaxaca, in planitie Llano Bot. Acad. Sci. Hung. 28: 69. 1982.
Verde, T. K. Hartweg 502 (Holotype K (photo Rondeletia cooperi Standi., Publ. Field Mus.
PTBG); Isotypes NY, n.v. (fragment US Nat. Hist., Bot. Ser. 4: 267. 1929. Type:
(photos MEXU, PTBG), P (photos MEXU, PANAMA: Bocas del Toro, Buena Vista
PTBG)).—Rondeletia buddleioides Benth. Camp, on Chiriqui Trail, 1250 ft., Jan-Mar
Arachnothryx buddleioides (Benth.) Planch, var. 1928, G. P. Cooper 600 (Yale 12233)
aspera (J. H. Kirkbr.) Borhidi, Acta Bot. (Holotype F 579596 (photo PTBG); Isotype
Hung. 33: 301. 1987. Rondeletia buddleioides NY, n.v.).—Rondeletia cooperi Standi.
Benth. var. aspera J. H. Kirkbr., Ann. Arachnothryx cordovana (Standi. & Steyerm.)
Missouri Bot. Gard. 55: 379. 1969. Type: Borhidi, Acta Bot. Hung. 33: 301. 1987.
PANAMA: Bocas del Toro, between Criollo Rondeletia cordovana Standi. & Steyerm.,
(just above Buena Vista) and Quebrada Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 26.
Higueron on Chiriqui Trail, 18 Apr 1968, 1943. Type: GUATEMALA: Zacapa, in
J. H. Kirkbride, Jr. & J. A. Duke 780 ravine bordering Quebrada Alejandria,
(Holotype MO 1942973 (photos MEXU, summit of Sierra de las Minas, vicinity of
PTBG)).—Rondeletia buddleioides Benth. Finca Alejandria, 2500 m, 13 Oct 1939, J. A.
Steyermark 29906 (Holotype F 1042952
A NOMENCLATOR OF MEXICAN AND CENTRAL
867136 (photos MEXU, PTBG); Isotype F Acta Bot. Hung. 33: 302. 1987 [as
(photo PTBG)).—Rondeletia dariensis gonzalaeoides]. Rondeletia gonzaleoides
Standi. Standi., Publ. Field Columbian Mus., Bot. Ser.
Arachnothryx deamii (Donn. Sm.) Borhidi, Acta 8: 160. 1930. Type: MEXICO: Chiapas,
Bot. Acad. Sci. Hung. 28: 68. 1982. Pueblo Nuevo, 2200 m, 1840, J. Linden 149
Bouvardia deamii Donn. Sm., Bot. Gaz. (Holotype LE, n.v. (fragment F, photos
(Crawfordsville) 49: 455. 1910. Type: MEXU, PTBG); probable Isotype P (photos
GUATEMALA: At base of Rillo de las MEXU, PTBG)).—Rondeletia £
Nabajas, about V* mile front side of Fiscal,
3700 ft., 5 Jun 1909, C C. Deam 6190 Arachnothryx gracilis (Hemsl.) Borhidi, Acta
(Holotype US 579579 (photos MEXU, Bot. Acad. Sci. Hung. 28: 69. 1982.
PTBG); Isotypes A, n.v., F (photo PTBG), Rondeletia gracilis Hemsl., Diag. PI. Nov.
GH, n.v., MO (photos MEXU, PTBG), US Mexic. 53. 1880. Type: GUATEMALA: Alta
(photos MEXU, PTBG)).—Rondeletia Verapaz, 3600 ft., H. von Tuerckheim 404
deamii (Donn. Sm.) Standi. (Holotype K (photo PTBG)).—Rondeletia
Arachnothryx edwardsii (Standi.) Borhidi, Acta gracilis Hemsl.
Bot. Hung. 33: 301. 1987. Rondeletia oica (Standi.) Borhidi,
edwardsii Standi., Trop. Woods 37: 31. 1934. Acta Bot. Hung. 33: 302. 1987. Rondeletia
Type: HONDURAS: Comayagua, Pito Solo, gracilispica Standi., Publ. Field Mus. Nat.
Lake Yojoa, 2000 ft, 29 Aug 1932,/ B. Hist., Bot. Ser. 8: 342. 1931. Type: MEXICO:
Edwards P476 (Holotype F 662645 (photos Oaxaca, Sierra San Pedro Nolasco, Talea, etc.,
MEXU, PTBG); Isotype A, n.v.). 1843-33, C. Jurgensen 935 (Holotype K
i Standi. (photo PTBG; fragment F, photos MEXU,
PTBG))—Rondeletia gracilispica Standi.
'Ha (Dwyer & Hay den) Isotypes B (presumably destroyed, photo
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 70. MO), P (photos MEXU, PTBG)).
1982. Rondeletia hameliifolia Dwyer & —Rondeletia laniflora Benth.
Hayden, Ann. Missouri Bot. Gard. 54: 144. Arachnothryx leptodictya (B. L. Rob.) Borhidi,
1967. Type: PANAMA: Veraguas, Santiago, Acta Bot. Acad. Sci. Hung. 28: 68. 1982.
Aug 1961, J. D. Dwyer 1350 (Holotype MO Rondeletia leptodictya B. L. Rob., Proc.
1824657 (photo PTBG); Isotype NY, n.v.). Amer. Acad. Sci. 45: 402. 1910. Type:
—Rondeletia hameliifolia Dwyer & Hayden MEXICO: [Boundary between Michoacan
Arachnothyx heteranthem (Brandegee) and Guerrero] Rives du Rio Petatlan, 500 m,
Borhidi, Acta Bot. Acad. Sci. Hung. 28: 69. 24 Nov 1898, E. Langlasse 666 (Holotype
1982. Rondeletia heteranthera Brandegee, GH, n.v.; Isotypes F (photos MEXU, PTBG),
Univ. Calif. Publ. Bot. 4: 387. 1913. Type: G, n.v., P (2 sheets, photos MEXU, PTBG),
MEXICO: Veracruz, Banos del Carrizal, Aug US (photos MEXU, PTBG)).—Rondeletia
1912, C. A. Purpus 6148 (Holotype UC leucophylla Kunth
155217, n.v.; Isotypes F, MO).—Rondeletia IT leucophylla (Kunth) Planch., Fl.
heteranthera Brandegee Serres 5: tab. 442. 1849. Rondeletia
Irachnothryx izabalensis (Standi. & Steyerm.) leucophylla Kunth in Humb., Bonpl. & Kunth,
Borhidi, Acta Bot. Hung. 33: 302. 1987. Nov. Gen. Sp. 3: 395. t. 290. 1820. Type:
Rondeletia izabalensis Standi. & Steyerm., MEXICO: Inter Alto del Peregrino et Flumen
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 286. Papagallo, Humboldt & Bonpland s.n.
1940. Type: GUATEMALA: Izabal, Escoba, (Holotype P-Bonpland, n.v.; Isotypes B, n.v.,
across the bay (west) from Puerto Barrios, P (photos MEXU, PTBG)).—Rondeletia
near sea level, 3 May 1939, P. C. Standlex leucophylla Kunth
72891 (Holotype F 991611 (photos MEXU, Arachnothyx licbmannii (Standi.) Borhidi,
PTBG)).—Rondeletia izabalensis Standi. & Acta Bot. Hung. 33: 302. 1987. Rondeletia
liebmannii Standi., J. Wash. Acad. Sci. 17:
Arachnothryxjurgensenii (Hemsl.) Borhidi, 336. 1927. Type: MEXICO: Oaxaca, without
Acta Bot. Acad. Sci. Hung. 28: 70. 1982. precise locality, 1841-43, F. Liebmann 11S34
Rondeletia jurgensenii Hemsl., Diag. PI. Nov (Holotype US 1315231 (photos MEXU,
Mexic. 29. 1879. Type: MEXICO: Oaxaca, PTBG); probable Isotype P (photos MEXU,
Sierra de San Pedro Nolasco, Jurgensen 248 PTBG)).—Rondeletia capitellata Hemsl.
(Holotype K (photo PTBG)).—Rondeletia Arachnothryx linguiformis (Hemsl.) Borhidi,
jurgensenii Hemsl. Acta Bot. Acad. Sci. Hung. 28: 70. 1982.
Arachnothryx kirkbridei (Dwyer) Borhidi, Acta Rondeletia linguiformis Hemsl, Diag. PI.
Bot. Acad. Sci. Hung. 28: 70. 1982. Nov. Mexic. 29. 1879. Type: GUATEMALA:
Rondeletia kirkbridei Dwyer, Ann. Missouri Without precise locality, Skinner s.n.
Bot. Gard. 67: 469. 1980. Type: PANAMA: (Holotype K (photo PTBG)).—Rondeletia
Veraguas, 16 km NW of Santa Fe on road to linguiformis Hemsl.
Calovebora (Panama Hwy. 35), 300-500 m Arachnothryx longipetiolata (Lundell) Borhidi,
16 May 1975, S. Mori & J. Kallunki 6112 Acta Bot. Hung. 35: 310. 1989. Rondeletia
(Holotype MO 2300094 (photo PTBG)). longipetolata Lundell, Wrightia 5: 325. 1976.
—Rondeletia bertieroides Standi. Type: MEXICO: Chiapas, Mt. Ovando, near
Arachnothryx laniflora (Benth.) Planch., Fl. Escuintla, 14 Nov 1945, E. Matuda 6146
Serres 5: tab. 442. 1849. Rondeletia laniflora (Holotype TEX-LL; Isotypes MEXU, TEX-
Benth., PI. Hartw. 85. 1841. Type: LL).—Rondeletia buddleioides Benth.
GUATEMALA: In montibus las Nubes, T. Arachnothryx lundelliana (Standi.) Borhidi,
Hartweg 584 (Holotype K (photo PTBG), Acta Bot. Hung. 33: 302. 1987. Rondeletia
fragment US (photos MEXU, PTBG); lundelliana Standi., Contr. Univ. Mich. Herb.
A NOMENCLATOR OF MEXICAN AND CEN
4: 31. 1940. Type: BRITISH HONDURAS Rondeletia myriantha Standi. & Steyerm. var.
(BELIZE): El Cayo Distr., Vaca, 21 Apr armentalis L. O. Williams, Phytologia 26:
1938, P. H. Gentle 2504 (Holotype F 941520 127. 1973. Type: GUATEMALA: Quiche,
(photos MEXU, PTBG); Isotypes A, n.v., Nebaj, 5600 ft., 22 Nov 1934, A. F Skutch
TEX-LL ex MICH (photos MEXU, PTBG), 1776 (Holotype F 815811 (photos MEXU,
US (photos MEXU, PTBG)).—Rondeletia PTBG)).~Rondeletia myriantha Standi. &
stenosiphon Hemsl. Steyerm. var. armentalis L. O. Williams
Arachnothryx macrocalyx (Standi. & Steyerm.) Arachnothryx nebulosa (Standi.) Borhidi, Acta
Borhidi, Acta Bot. Hung. 33: 302. 1987. Bot. Hung. 33: 302. 1987. Rondeletia
Rondeletia macrocalyx Standi. & Steyerm., nebulosa Standi., Trop. Woods 37: 32. 1934.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254. Type: HONDURAS: Tegucigalpa, La Aurora
1947. Type: GUATEMALA: Izabal, Cerro Trail, Rosario, San Juancito, 5400 ft., 2 Apr
San Gil, 300-900 m, 25 Dec 1941, J. A. 1932, J. B. Edwards P10 (Holotype F 662646
Steyermark 41864 (Holotype F 1134224 (photos MEXU, PTBG); Isotype A, n.v.).
(photos MEXU, PTBG); Isotype F (photos —Rondeletia nebulosa Standi.
MEXU, PTBG)).—Rondeletia macrocalyx Arachnothryx nicaraguensis (Oerst.) Borhidi,
Standi. & Steyerm. Acta Bot. Hung. 35: 310. 1989. Rondeletia
Arachnothryx mexiae (Standi.) Borhidi, Acta nicaraguensis Oerst., Vidensk. Meddel.
Bot. Hung. 33: 302. 1987. Rondeletia mexiae Dansk Naturhist. Foren. Kjobenhavn 1852:
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. 43. 1853. Type: NICARAGUA: Matagalpa,
22: 55. 1940. Type: MEXICO: Oaxaca, Distr. Segovia, in monte Pantasmo, Jan 1848,
Choapan, Santa Maria, Montana Santa Maria, Oersted s.n. (Holotype C (photos F, MO, US),
1500 m, 1 Apr 1938, Y. Mexia 9261 fragment US).—Rondeletia nicaraguensis
(Holotype F 982856 (photos MEXU, PTBG); Oerst.
Isotypes GH, n.v., MO (photos MEXU, Arachnothryx nitida (Hemsl.) Borhidi, Acta Bot.
PTBG), US (photos MEXU, PTBG)). Acad. Sci. Hung. 28: 69. 1982. Rondeletia
—Rondeletia scabra Hemsl. nitida Hemsl., Diag. PI. Nov. Mexic. 29.
Arachnothryx minor (Lundell) Borhidi, Acta 1879. Type: MEXICO: Chiapas, Fl. rosatres,
Bot. Hung. 35: 310. 1989. Rondeletia minor forets des pins, Chiapas, in fevr., Linden 1660
Lundell, Wrightia 5: 326. 1976. Type: (Lectotype K, here designated (photo PTBG);
MEXICO: Veracruz, about 10 miles north of Isolectotypes G-DEL, n.v., fragment F, P
Teziutlan, 6 Jan 1964, D. S. Correll & H. B. (photos MEXU, PTBG)). Syntype: MEXICO:
Correll 28736 (Holotype TEX-LL (photos Chiapas, Ghiesbreght 728 (K (photo PTBG),
MEXU, PTBG); Isotype MEXU). GH, n.v.). Linden 1660 was designated as
—Rondeletia gracilispica Standi. lectotype, as it is the better specimen with
Arachnothryx myriantha (Standi. & Steyerm.) more duplicates.—Rondeletia nitida Hemsl.
Borhidi, Acta Bot. Hung. 33: 302. 1987. Arachnothryx oaxacana (Standi.) Borhidi, Acta
Rondeletia myriantha Standi. & Styerm., Bot. Hung. 33: 302. 1987. Rondeletia
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 288. oaxacana Standi., Field Mus. Nat. Hist., Bot.
1940. Type: GUATEMALA: Suchitepequez, Ser. 11: 255. 1936. Nomen superfl., based on
Finca Moca, 3200 ft., 31 Oct 1934, A. F. same type as Rondeletia mexicana (Turcz.)
Skutch 1569 (Holotype F 813463 (photos Standi. Type: MEXICO: Oaxaca, without
MEXU, PTBG); Isotypes A, n.v., US (photos precise locality, 3000 ft., H. Galeotti 2664
MEXU, PTBG)).—Rondeletia myriantha (Holotype P (photos MEXU, PTBG),
Standi. & Steyerm. fragment F (photos MEXU, PTBG)).
Arachnothryx myriantha (Standi. & Steyerm.) —Rondeletia mexicana (Turcz.) Standi.
Borhidi var. armentalis (L.O. Williams) Arachnothryx ovandensis (Lundell) Borhidi,
Borhidi, Acta Bot. Hung. 35: 310. 1989. Acta Bot. Hung. 35: 310. 1989. Rondeletia
ovandensis Lundell, Wrightia 5: 326. 1976. PTBG); Isotypes F, MEXU, TEX-LL).
Type: MEXICO: Chiapas, Mt. Ovando, near —Rondeletia pyramidalis Lundell
Escuintla, 24 Oct 1941, E. Matuda 6067 Arachnothryx rekoi (Standi.) Borhidi, Acta Bot.
(Holotype TEX-LL (photos MEXU, PTBG); Acad. Sci. Hung. 28: 69. 1982. Rondeletia
Isotypes F (photo PTBG), MEXU, TEX- rekoi Standi, Proc. Biol. Soc. Wash. 8: 126.
LL).—Rondeletia ovandensis Lundell 1918. Type: MEXICO: Oaxaca, Las Pilas
Arachnothryxpansamalana (Standi.) Borhidi, (Cerro Espino), 400 m, 10 Oct 1917, B. P.
Acta Bot. Acad. Sci. Hung. 28: 69. 1982. Reko 3490 (Holotype US 867145 (photos
Rondeletia pansamalana Standi., N. Amer. Fl. MEXU, PTBG)).—Rondeletia rekoi Standi.
32: 58. 1918. Type: GUATEMALA: Alta Arachnothryx rubens (L. O. Williams) Borhidi,
Verapaz, Pansamala, 3800 ft., Apr 1886, H. Acta Bot. Hung. 35: 311. 1989. Rondeletia
von Tuerckheim 897 (Holotype US 49985 rubens L. O. Williams, Phytologia 26: 128.
(photos MEXU, PTBG); Isotype GH, n.v.). 1973. Type: GUATEMALA: Quiche, Nebaj,
—Rondeletia pansamalana Standi. 6000 ft, 19 Nov 1934, A. F. Skutch 1725
Arachnothryxphyllocephala (Standi. & L. O. (Holotype F 815804 (photos MEXU, PTBG)).
Williams) Borhidi, Acta Bot. Hung. 33: 302. —Rondeletia rubens L. O. Williams
1987. Rondeletia phyllocephala Standi. & Arachnothryx rufescens (B. L. Rob.) Borhidi,
L. O. Williams, Ceiba 3: 63. 1952. Type: Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
HONDURAS: Cortes, faldas hiimedas de Rondeletia rufescens B. L. Rob, Proc. Amer.
Montana Sta. Ana, matorrales hiimedos de Acad. Arts 45: 402. 1910. Type:
Rio Sta. Ana, 210 m, 6 Dec 1950, A. Molina GUATEMALA: Alta Verapaz, Coban, 4800
R. 3608 (Holotype US 2216049 (photos ft. Mar 1881, H. von Tuerckheim 582
MEXU, PTBG); Isotype F (photos MEXU, (Holotype GH, n.v.; Isotypes P (photos
PTBG))—Rondeletia thiemei Donn. Sm. MEXU, PTBG), US, 2 sheets (photos MEXU,
•\ x platysepala (Standi.) Borhidi, PTBG)).—Rondeletia gracilis Hemsl.
Acta Bot. Acad.'Sci. Hung. 28: 70. 1982. Arachnothryx salicifolia (Dwyer & Hay den)
Rondeletia platysepala Standi., Ann. Missouri Borhidi, Acta Bot. Acad. Sci. Hung. 28: 70.
Bot. Gard. 27: 343. 1940. Type: PANAMA: 1982. Rondeletia salicifolia Dwyer &
Code, El Valle, North Rim, 600-1000 m, 14 Hayden, Phytologia 15: 58. 1967. Type:
May 1939, P. H. Allen 1791 (Holotype F PANAMA: Panama, Cerro Jefe in "Clusia
989484 (photo PTBG); Isotypes F (photos forest" at 2700-3000 ft, 27 Jan 1966, E. L.
MEXU, PTBG), G (n.v.), GH, n.v, MO Tyson, J. D. Dwyer & K. Blum 3319
(photos MEXU, PTBG), NY, n.v.). (Holotype MO 1824503 (photo PTBG);
—Rondeletia bertieroides Standi. Isotypes F, US (photos MEXU, MO)).
Arm hnotkryxpotycephala (Standi.) Borhidi, —Rondeletia bertieroides Standi.
Acta Bot. Hung. 35: 310. 1989. Rondeletia Arachnothryx secunda (Standi.) Borhidi, Acta
polycephala Standi, J. Wash. Acad. Sci. 17: Bot. Acad. Sci. Hung. 28: 70. 1982.
337. 1927. Type: MEXICO: Oaxaca, without Rondeletia secunda Standi, Contr. U.S. Natl.
precise locality, F. Liebtnann 11826 Herb. 18: 141. 1916. Type: PANAMA: San
(Holotype C, fragment US (photos MEXU, Bias, forests around Puerto Obaldia, San Bias
PTBG)).—Rondeletia mexicana (Turcz.) coast, 0-50 m, Aug 1911, H. Pittier 4279
Standi. (Holotype US 679392 (photos MEXU,
Arachnothryxpyramidalis (Lundell) Borhidi, PTBG); Isotype NY, n.v.).—Rondeletia
Acta Bot. Hung. 35: 310. 1989. Rondeletia secunda Standi.
pyramidalis Lundell, Wrightia 5: 327. 1976. Arachnothryx secundiflora (B. L. Rob.) Borhidi,
Type: MEXICO: Chiapas, San Luis, near Acta Bot. Acad. Sci. Hung. 28: 69. 1982.
Siltepec, 1300 m, 24 Jan 1945, E. Matuda Rondeletia secundiflora B. L. Rob, Proc.
5292 (Holotype TEX-LL (photos MEXU, Amer. Acad. Arts 45: 403. 1910. Type:
A NOMENCLATOR OF MEXICAN AND CENTRAL /
Bertiera angustifolia Benth., Bot. Voy. Sulphur um mexicanum Standi.. Publ. Field
103. 1844. Type: COSTA RICA: Cocos Mus. Nat. Hist., Bot. Ser. 22: 108. 1940.
Island, Barclay s.n. (Holotype BM, n.v.). Type: MEXICO: Chiapas, Rvina [Ruina],
Bertiera guianensis Aubl., Hist. PL Guiane 1: Palenque, 10-12 Jul 1939, E. Matuda 3670
180, pi. 69. 1775. Type: FRENCH GUIANA: (Holotype F 1011462 (photo PTBG); Isotypes
"prope Aroura," Aublet s.n. (Holotype A, n.v., TEX-LL (photos MEXU, PTBG), US
presumably BM, n.v.). (photos MEXU, PTBG)).—Blepharidium
Bertiera guianensis var. leiophylla Steyerm., guatemalense Standi.
Mem. New York Bot. Gard. 17: 319. 1967.
Type: PANAMA: Chiriqui, vicinity of San Borojoa Cuatrecasas, Ser. Bot. Aplicada, Serv.
Felix, 0-120 m, 17 Dec 1911, H. Pittier 5178 Devulg., Seer. Agric. Ganad. (Call, Colombia)
(Holotype NY, n.v.; Isotype US (photos 2: 474. 1949. (Preprint of Rev. Acad.
MEXU, PTBG)).—Bertiera guianensis Aubl. Colombia Ci. Exact. 7: 474. 1950.) Type:
Bertiera oligosperma Wernham, J. Bot. 50: 162. Borojoa patinoi Cuatrecasas.
1912. Type: NICARAGUA: Seemann 129 Borojoa atlantica Dwyer, Ann. Missouri Bot.
(Holotype BM, n.v.).—Status uncertain. Gard. 67: 46. 1980. Type: PANAMA: Colon,
Standi., N. Amer. Fl. 32: 2-3 miles up from the Rio Guanche, 10-20 m,
180. 1934. Type: PANAMA: Darien, Cana 19 Jan 1973, H. Kennedy & R. Foster 2197
and vicinity, 2000-5600 ft., 17 Apr-8 Jun (Holotype MO 2163702 (photo PTBG);
1908, R. S. Williams 832 (Holotype US Isotype MO (photo PTBG)).
678312 (photos MEXU, PTBG); Isotype NY, Borojoa panamensis Dwyer, Phytologia 17:
n.v.).—Raritebe palicoureoides Wernham 446. 1968. Type: PANAMA: Code, Cerro
subsp. dwyeranum J. H. Kirkbr. Pilon, El Valle, 4 Jan 1968, J. A. Duke & B. R.
Bertiera tenuis Lundell, Wrightia 4: 49. 1968. Lallathin 15014 (Holotype MO 1968459
Type: GUATEMALA: Alta Verapaz, Sebol, (photo PTBG); Isotype MO).
in high forest about 800 m west of village, Borojoa patinoi Cuatrec, Ser. Bot. Aplicada,
bordering arroyo, 15 Apr 1964, E. Contreras Serv. Devulg., Seer. Agric. Ganad. (Cali,
4318 (Holotype TEX-LL (photos MEXU, Colombia) 2: 474. 1949. (Preprint of Rev.
PTBG); Isotype F).—Bertiera guianensis Acad. Colombia Ci. Exact. 7: 474. 1950.)
Aubl. Type: COLOMBIA: Valle, Rio Calima
Bertiera viburnoides (Standi.) J. H. Kirkbr., (region del Choco); margen derecha, entre
Brittonia 31:311. 1979. Raritebe viburnoides Quebrada de la Brea y La Esperanza, 25 m, 19
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. May 1946, J. Cuatrecasas 21137 (Holotype F
7: 56. 1930. Type: COLOMBIA: "Caucas, La 1251558 (photo PTBG); Isotypes F (3 sheets),
Gallera, Mica Valley," Killip 7934 (Holotype US (photo PTBG)).
US, n.v., fragment F, n.v.; Isotypes G, n.v., Y,
n.v.). Borreria G. Mey., Prim. Fl. Esseq. 79, t. 1. 1818.
Type: Borreria suaveolens G. Mey., excl. ref.
Browne Jam. 141, no. 4, nom. et typ. cons.
[Herein treated as a synonym of Spermacoce
L. In cases where generic transfers have not
Blepharidium guatemalense Standi., J. Wash. yet been published, the correct genus is given
Acad. Sci. 8: 59. 1918. Type: GUATEMALA: in boldface type.]
Alta Verapaz, forest along Saclac River, Borreria arenicola Brandegee, Univ. Calif.
below Secanquim, 300 m, 7 May 1905, H. Publ. Bot. 10: 417. 1924. Type: MEXICO:
Pittier 266 (Holotype US 472925 (photos Chiapas, in sandy arroyo near Jalisco, Aug
MEXU, PTBG); Isotype US (photos MEXU, 1923, C. A. Purpus 8968 (Holotype UC
PTBG)).
220266, n. Kunth in Humb., Bonpl. & Kunth, Nov. Gen.
(DC.) Alain Sp. 3: 344. 1820. Type: MEXICO: Jalisco,
Borreria aspera M. Martens & Galeotti, Bull. Volcan de Jorullo, 3000 ft., Humboldt &
Acad. Roy. Bruxelles 11: 130. 1844. Type: Bonpland s.n. (Holotype P-Bonpland, n.v.
MEXICO: Oaxaca, Juquila, 4000 ft., H. (photo MO)).—Diodia apiculata (Willd. ex
Galeotti 2625 (Holotype BR, n.v.). Roem. & Schult.) K. Schum., vel sp. aff.
—Spermacoce suaveolens (G. May.) Kuntze Borreria exilis L. O. Williams, Phytologia 28:
Borreria asperifolia (M. Martens & Galeotti) 227. 1974. Type: COSTA RICA: Cocos
B. L. Rob., Proc. Amer. Acad. Arts 45: 409. Island, Wafer Bay, 28 June 1932, J. T. Howell
1910. Spermacoce asperifolia M. Martens & 10178 (Holotype F 1031197 (photo PTBG)).
Galeotti, Bull. Acad. Roy. Bruxelles 11: 132. Nom. nov. for Borreria gracilis L. O.
1844. Type: MEXICO: Oaxaca, pres de Williams, Phytologia 26: 487. 1973.
1'Ocean Pacifique, H. Galeotti 2626 —Spermacoce exilis (L. O. Williams) C D.
(Holotype BR, n.v.; Isotype P, n.v. (photo
MO)).—Staelia scabra (C. Presl) Standi. Borreria fruticosa Standi., J. Arnold Arbor. 11:
Borreria assurgens (Ruiz & Pav.) Griseb., Abh. 42. 1930. Type: HONDURAS: Comayagua,
Konigl. Ges. Wissensch. Gottingen 19: 159. Vicinity of Siguatepec, 1080-1400 m., 14-27
1874. Spermacoce assurgens Ruiz & Pav., Feb 1928, P. C. Standley 56240 (Holotype F
Syst. Veg. Fl. Peru 1: 60, t. 92b. 1798. Type: 581434 (photo PTBG); Isotype US (photos
PERU: Habitat m ruderalis Limae Chancay et MEXU, PTBG)).—Spermacoce
Huanuci, Ruiz Lopez & Pavon s.n. (syntype Borreria fruticosa Standi, var. pubescent Standi.
MA, n.v. (photo MO)); Plukenet Aim. 33, tab. & L. O. Williams, Ceiba 1: 165. 1950. Type:
136/4 (syntype, n.v.).—Spermacoce HONDURAS: El Paraiso, 2 miles W of
assurgens Ruiz & Pav. Guinope, 1400 m, 16 Aug 1946, L. O.
Borreria auriculata Brandegee, Univ. Calif. Williams & A. Molina R. 10315 (Holotype US
Publ. Bot. 10: 417. 1924. Type: MEXICO: 1206971 (photos MEXU, PTBG); Isotype F
Chiapas, meadows near Jalisco, Aug 1923, (photo PTBG)).—Spermacoce
C A. Purpus 9218 (Holotype UC, n.v.; Borreria gracilis L. O. Williams, Phytologia 26:
Isotype F (photo PTBG)).—Crusea 487. 1973. Not Borreria gracilis Scheele,
psyllioides (Kunth) W. R. Anderson 1843, nor Miq. ex Hook., 1855. Type:
Borreria bartlingiana DC, Prodr. 4: 544. 1830. COSTA RICA: Cocos Island, Wafer Bay, 28
Type: MEXICO: Without precise locality, Jun 1932, J. T. Howell 10178 (Holotype F
Haenke s.n. (Holotype presumably PR, n.v.). 1031197 (photo PTBG)).—Spermacoce exilis
—Spermacoce latifolia Aubl. (L. O. Williams) C D. Adams
Borreria chiapensis Brandegee, Univ. Calif. Borreria graminifolia M. Martens & Galeotti,
Publ. Bot. 10: 417. 1924. Type: MEXICO: Bull. Acad. Roy. Bruxelles 11: 129. 1844.
Chiapas, Hacienda Monserrate, Sep 1923, Type: MEXICO: El Sabino, pres de Zimapan,
C. A. Purpus 9216 (Holotype UC 223384, 6500 pieds, H. Galeotti 2599 (Holotype BR,
n.v.; Isotypes F (photo PTBG), NY, n.v., US n.v.).—Spermacoce verticillata L.
(photos MEXU, PTBG)).—Crusea setosa Borreria haenkeana DC, Prodr. 4: 547. 1830.
(M. Martens & Galeotti) Standi. & Steyerm. Type: MEXICO: Without precise locality or
Borreria densiflora DC, Prodr. 4: 542. 1830. collector, presumably Haenke (G-DEL, n.v.
Type: JAMAICA: Without precise locality, (photo MO)).—Borreria suaveolens G. Mey.
Swartzs.n. (Holotype G-DC, n.v. (microfiche Borreria heterophylla Brandegee, Univ. Calif.
4. 542.3)).—Spermacoce densiflora (DC) Publ. Bot. 10: 418. 1924. Type: MEXICO:
Chiapas, in sandy arroyo near Jalisco, Aug
1923, C. A. Purpus 8969 (Holotype UC
220267, n.v.).—Spermacoce prostrata Aubl.
L NOMENCLATOR OF MEXICAN AND CENTRAL
Madre, 16 Aug 1897, J. N. Rose 3516 Bouvardia steyermarkii Standi., Publ. Field
(Holotype US 302493 (photos MEXU, Mus. Nat. Hist., Bot. Ser. 22: 383. 1940.
PTBG); Isotype F). Type: GUATEMALA: Zacapa, Sierra de las
Bouvardia rzedowskii Terrell & S. D. Koch, Minas, between Rio Hondo and Finca
Novon4: 179, fig. 1. 1994. Type: MEXICO: Alejandna, 1700-2000 m, 11 Oct 1939, J. A.
Oaxaca, ladera caliza con vegetacion de Steyermark 29671 (Holotype F 1043386
encinar, 2300 m, 6 km al SW de (photos F, PTBG)).—Bouvardia dolichantha
Tamazulapan, sobre la carretera a Chilapa, 12
Aug 1977, J. Rzedowski 35142 (Holotype Bouvardia strigosa Benth., PI. Hartweg. 75.
ENCB; Isotypes ARIZ, n.v., ASU, n.v., 1841. Type: GUATEMALA: Quezaltenango,
MICH, n.v., XAL). [Volcan] Zunil, Nov., T. Hartweg 530
Bouvardia salvadorensis Steyerm., Ceiba 4: (Lectotype K (photo PTBG), here designated;
302. 1955. Type: EL SALVADOR: Isolectotypes G-DEL, n.v. (fragment F), P
Sonsonante, Rio Acachapa near headquarters (photos MEXU, MO, PTBG), W, n.v. (photos
of Hacienda Las Tablas, Balsam Range, 570 F, MO)). Syntype: GUATEMALA: Volcan de
m, May 1941, J. M. Tucker 1368 (Holotype F Fuego, 6000-6400 ft., Salvin s.n. (K (photo
1492726 (photo PTBG); Isotypes MICH, n.v., PTBG),fragment F). Rogiera.—Rondeletia
UC, n.v., US (photos MEXU, PTBG)). strigosa (Benth.) Hemsl.
—Bouvardia multiflora (Cav.) Schult. & Bouvardia subcordata Standi, N. Amer. Fl.
Schult. f. 32: 105. 1921. Type: MEXICO: Sinaloa, near
Bouvardia scabra Hook. & Arn., Bot. Beechey Colomas, 13 Jul 1897,7. N. Rose 1628
Voy. 427. 1840. Type: MEXICO: Nayarit, (Holotype US 300476 (photos MEXU,
between San Bias and Tepic, Sinclair s.n. PTBG); Isotype F).
(Holotype K, n.v.). Bouvardia tenuijlora Schltdl., Linnaea 26: 97.
Bouvardia scabrida M. Martens & Galeotti, 1853. Type: Based on living plants in
Bull. Acad. Roy. Sci. Bruxelles 11: 237. 1844. cultivation (B, presumably destroyed).
Type: MEXICO: Oaxaca, ravin de Yavezia, —Bouvardia ternifolia (Cav.) Schltdl.
cordillere orientale, 6000-7000 pieds, Dec Bouvardia tenuifolia Standi., N. Amer. Fl. 32:
1840, H. Galeotti 2624 (Holotype BR, n.v.; 104. 1921. Type: MEXICO: Jalisco, grassy
Isotype P, n.v. (photo MO)).—Bouvardia hillsides near Guadalajara, Oct 1889, C. G.
ternifolia (Cav.) Schltdl. Pringle 2292 (Holotype US 49988 (photos
Bouvardia schiedeana Schltdl., Linnaea 26: MEXU, PTBG); Isotypes A, n.v., BM, n.v.,
123. 1853. Type: MEXICO: Barranca de BR, n.v., F, G, n.v., LE, n.v., M, n.v., MICH,
Tioselo, Aug, Schiedes.n. (Holotype B, n.v., MO, MSC, n.v., P, n.v., US, n.v., W,
presumably destroyed).—Bouvardia
multiflora (Cav.) Schult. & Schult. f. Bouvardia ternifolia (Cav.) Schltdl., Linnaea
Bouvardia splendens J. Graham, Bot. Mag. pi. 26: 98. 1853. Ixora ternifolia Cav., Ic. 4: 3, /.
3781. 1840. Type: Based on greenhouse- 305. 1797. Lectotype: Protologue illustration
grown plants (Holotype K, n.v.).—Bouvardia of Ixora ternifolia Cav., Ic. 4: 3, /. 305. 1797
ternifolia (Cav.) Schltdl. (based on a plant grown at MA from Mexican
Bouvardia standleyana W. H. Blackw., Ann. seed; designated by Blackwell, 1968).
Missouri Bot. Gard. 55: 13. 1968. Type: Bouvardia ternifolia (Cav.) Schltdl. var.
MEXICO: Michoacan, Coalcoman, S angustifolia (Kunth) B. L. Rob., Proc. Amer.
Naranjillo, 1360 m, 26 Nov 1938, G. B. Acad. Sci. 45: 405. 1910. Bouvardia
Hinton et al. 12677 (Holotype US 2020693 angustifolia Kunth in Humb., Bonpl. &
(photos MEXU, PTBG); Isotypes K, n.v., Kunth, Nov. Gen. Sp. 3: 384. 1819. Type:
MICH, n.v., UC, n.v.). MEXICO: Moran (?), A. Bonpland s.n.
(Holotype P-Bonpland, n.v. (photo MO);
Isotype F (photo PTBG)).—Bouvardia (photo PTBG), GH, n.v.).—Bouvardia
ternifolia (Cav.) Schltdl. xylosteoides Hook. & Arn.
Bouvardia tolucana Hook. & Am., Bot. Bouvardia viminalis Schltdl., Linnaea 26: 120.
Beechey Voy. 427. 1840. Type: MEXICO: 1854. Type: MEXICO: Oaxaca, Monte Alban,
Mexico, Toluca, Andheux 332 (Holotype K, 6000 ft., 4 Sep 1894, C G. Pringle 4888
n.v.; Isotype W, n.v.).—Bouvardia ternifolia (Neotype MO 174074 (photos MEXU,
(Cav.) Schltdl. PTBG); Isoneotypes BM, n.v., BR, n.v., F,
Bouvardia triflora Kunth in Humb., Bonpl. & GH, n.v., M, n.v., MEXU, 2 sheets, MSC,
Kunth, Nov. Gen. Sp. 3: 386. 1820. Type: n.v., P, n.v., UC, n.v., US (photos MEXU,
MEXICO: In temperatis, Humboldt & PTBG), W, n.v.; designated by Blackwell,
Bonplands.n. (Holotype P-Bonpland, n.v. 1968).
(fragment P, n.v.)). Houstonia.—Bouvardia Bouvardia violacea Rzed., Ciencia (Mexico) 19:
multiflora (Cav.) Schult. & Schult. f. 82. 1959. Type: MEXICO: San Luis Potosi, J.
Bouvardia triflora Kunth var. hirsuta M. Rzedowski 7680 (Holotype MEXU; Isotype
Martens & Galeotti, Bull. Acad. Bruxelles 11: ENCB).—Bouvardia rosea Schltdl.
236. 1844. Type: MEXICO: Oaxaca, Bouvardia xestosperma (B. L. Rob. &
cordillere orientale, dans les montagnes du Greenm.) Terrell & S. D. Koch, Novon 4:
Rincon, 5000 pieds, H. Galeotti 2658 bis 181. 1994. Oldenlandia xestosperma B. L.
(Holotype BR, n.v.).—Bouvardia multiflora Rob. & Greenm., Proc. Amer. Acad. Arts 32:
(Cav.) Schult. & Schult. f. 41. 1896. Type: MEXICO: Oaxaca, open
Bouvardia triphylla Salisb., Parad. Lond.,p/. 88. glades, Sierra de San Felipe, 8000 ft., Jun
1807. Norn, superfl., based on Ixora ternifolia 1894, C. G. Pringle 4692 (Lectotype GH,
Cav.—Bouvardia ternifolia (Cav.) Schltdl. n.v.; Isolectotypes BM, n.v., CM, n.v., F, K,
Bouvardia venosissima Lundell, Bull. Torrey n.v., MEXU, 2 sheets, MICH, n.v., MO
Bot. Club 66: 602. 1939. Type: MEXICO: (photos MEXU, PTBG), MSC, n.v., PH, n.v.,
Chiapas, Mt. Tacana, 1400 m, 17-23 Mar US, 2 sheets, VT, n.v.; designated by Terrell,
1939, E. Matuda 2748 (Holotype MICH, n.v.; 1980).
Isotype MEXU).—Bouvardia dictyoneura Bouvardia xylosteoides Hook. & Arn., Bot.
Standi. Beechey Voy. 428. 1840. Type: MEXICO:
Bouvardia versicolor KerGawl., Bot. Reg. 3: pi. Oaxaca, on slightly elevated mountains about
245. 1817. Type: Based on a greenhouse- Mitlam [Mitla], Andrieux 335 (Holotype K,
grown plant (the protologue illustration is n.v.; Isotype G-DEL, n.v., fragment F).
taken as the type; designated by Blackwell,
1968).—Bouvardia multiflora (Cav.) Schult. Buena Cav., Anal. Hist. Nat. (Madrid) 2: 278, /.
& Schult. f. 23. 1800. Type: Buena panamensis Cav. Not
Bouvardia versicolor Ker Gawl. var. graciliflora Buena Pohl, 1826. [Herein treated as a
A. Gray in S. Watson, Proc. Amer. Acad. Arts synonym of Gonzalagunia Ruiz & Pavon.]
22: 416. 1887. Type: MEXICO: Jalisco, Rio Buena panamensis Cav., Anal. Hist. Nat.
Blanco, E. Palmer 154 (Syntype GH, n.v.); (Madrid) 2: 279. 1800. Type: PANAMA:
Tequila, E. Palmer 369 (Syntype GH, n.v.); Without precise locality, Nee s.n. (Holotype
Chapala, E. Palmer 708 (Syntype GH, n.v.). MA, n.v.; Isotype F (photo PTBG)).
—Bouvardia multiflora (Cav.) Schult. & Gonzalea.—Gonzalagunia panamensis
Schult. f. (Cav.) K. Schum.
Bouvardia villosa Standi., N. Amer. Fl. 32: 107. Buena skinneri Oerst., Vidensk. Meddel. Dansk
1921. Type: MEXICO: Oaxaca, Alturas de Naturhist. Foren. Kjobenhavn 1852: 48. 1853.
Matatlan, 1800 m, C. Conzatti I486 TYPE: NICARAGUA: Masaya, toppen af
(Holotype US 764094 (photos MEXU, Vulkanen Masaya, 2000-3000 ft., Oersted
PTBG)); Isotypes G-DEL, n.v., fragment F 11134 (Syntypes C, n.v. (photos A, F), US);
\. NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Cephaelis costaricensis Schltdl., Linnaea 28: Cephaelis gaugeri Dwyer, Ann. Missouri Bot.
546. 1856. Type: COSTA RICA: Type not Gard. 67: 71. 1980. Type: PANAMA:
designated.—Psychotria elata (Sw.) Hammel Panama, Cerro Jefe, near junction of roads to
Cephaelis croatii Dwyer, Ann. Missouri Bot. Cerro Jefe and Altos de Pacora, 18 Dec 1974,
Gard. 67: 64. 1980. Type: PANAMA: Darien, S. Mori & J. Kallunki 3779 (Holotype MO
upper gold mining camp of Tyler Kittredge, 2240077 (photos MEXU, PTBG)).
headwaters of Rio Tuquesa, ca. 2 air km from —Psychotria camponutans (Dwyer &
continental divide, 26 Aug 1974, T. B. Croat Hayden) Hammel
27204 (Holotype MO 2201416 (photos Cephaelis gentryi Dwyer, Selbyana 2: 57. 1977.
MEXU, PTBG); Isotypes F, GH, n.v.). Type: ECUADOR: Los Rios, Rio Palenque
—Psychotria gentryi (Dwyer) C. M. Taylor Science Center, km 56 of Quevedo-Santo
Cephaelis dichroa (Standi.) Standi., Publ. Field Domingo Road, C. Dodson 5169 (Holotype
Columbian Mus., Bot. Ser. 4: 296. 1929. Evea MO; Isotypes RPSC, n.v., SEL, n.v., US,
dichroa Standi., Contr. U.S. Nat. Herb. 18: n.v.).—Psychotria gentryi (Dwyer) C. M.
124. 1916. Type: PANAMA: Chiriqui, humid Taylor
forest of Cuesta de Las Palmas, southern Cephaelis glomerulata Dorm. Sm., Bot. Gaz.
slopes of Cerro de la Horqueta, 1700-2100 m, (Crawfordsville) 16: 12. 1891. Type:
17-19 Mar 1911, H. Pittier 3218 (Holotype GUATEMALA: Dept. Livingston, Rio Dulce,
US 677615 (photos MEXU, PTBG); Isotype Mar 1880, J. D. Smith 1637 (Holotype US
F)).—Psychotria dichroa (Standi.) C. M. 943529 (photos MEXU, PTBG); Isotype US
Taylor (photos MEXU, PTBG)).—Psychotria
Cephaelis dimorphandrioides Dwyer, Ann. glomerulata (Donn. Sm.) Steyerm.
Missouri Bot. Gard. 67: 66. 1980. Type: Cephaelis guapilense (Standi.) Standi., Publ.
PANAMA: Veraguas, Valley of Rio Dos Field Columbian Mus., Bot. Ser. 4: 295. 1929.
Bocas on road between Alto Piedra (above Evea guapilensis Standi., J. Wash. Acad. Arts
Santa Fe) and Calovebora, 350-400 m, 29 Sci. 15: 104. 1925. Type: COSTA RICA:
Aug 1974, T. B. Croat 27470 (Holotype MO Limon, vicinity of Guapiles, 300-500 m, P. C.
2201358 (photos MEXU, PTBG)). Standley 37025 (Holotype US 1153029
—Palicourea dimorphandrioides (Dwyer) (photos MEXU, PTBG)).—Psychotria
C. M. Taylor guapilensis (Standi.) C. W. Ham.
Cephaelis discolor Pol., Linnaea 41: 572. 1877. Cephaelis hirsute M. Martens & Galeorti, Bull.
Type: COSTA RICA: Angustura, 1875, H. Acad. Roy. Bruxelles 11: 135. 1844. Type:
Polakowsky 384 (Holotype B, presumably MEXICO: Oaxaca, la Chinantla, 4000 pieds,
destroyed (photos F, MO, US), fragment F H. Galeotti 7185 (Holotype BR, n.v.).
(photo PTBG)). Not Psychotria discolor —Psychotria poeppigiana Mull. Arg.
Huber, 1901, nor P. discolor (Griseb.) Rolfe, Cephaelis insueta Dwyer, Ann. Missouri Bot.
1893—Psychotria guapilensis (Standi.) Gard. 67: 73. 1980. Type: PANAMA:
Hammel Veraguas, 11 km from Escuela Agricola Alto
Cephaelis dressleri Dwyer, Ann. Missouri Bot. de Piedra, forest along Rio Dos Bocas,
Gard. 67: 68, fig. 15. 1980. Type: PANAMA: tropical wet forest, Atlantic slope, 15 Nov
San Bias, forest south of Puerto Obaldia, 18 1974, S. Mori & J. Kallunki 3108 (Holotype
Aug 1971, T. B. Croat 16801 (Holotype MO MO 2220981 (photo PTBG)).—Psychotria
2138462 (photo MO)).—Psychotria dressleri insueta (Dwyer) C. W. Ham.
(Dwyer) C. W. Ham. Cephaelis ipecacuanha (Brot.) A. Rich., Bull.
Cephaelis elata Sw., Prodr. Veg. Ind. Occ. 45. Fac. Med. 4: 92. 1818. Callicocca
1788. Type: JAMAICA, Swartz s.n. ipecacuanha Brot., Trans. Linn. Soc. London
(presumably S, n.v.).—Psychotria elata 6: 137. 1802. Type: BRAZIL: Without precise
(Sw.) Hammel
locality, Gomez s.n. (presumably LISU, n.v.).
—Psychotria ipecacuanha (Brot.) Stokes
Cephaelis kennedyae Dwyer, Ann. Missouri A. Gentry & S. Mori 13895 (Holotype MO
Bot. Gard. 67: 75. 1980 [as kennedyi}. Type: 2276226 (photos MEXU, PTBG)).
PANAMA: Panama, 6 km from Panamerican —Psychotria dressleri (Dwyer) C. W. Ham.
Highway on road from El Llano to Carti- Cephaelis phoenicia (K. Schum.) Dorm. Sm.,
Tupile, H. Kennedy 1779 (Holotype MO Enum. PI. Guat. 5: 39. 1899. Uragoga
2161974 (photo PTBG); Isotype GH, n.v.). Phoenicia K. Schum. in Engl. & Prantl, Nat.
—Psychotria muscosa (Jacq.) Steyerm. Pflanzenfam. 4, 4: 120. 1891. Type: COSTA
( 'cphcichs lansupulata Standi., J. Wash. Acad. RICA: Suerre, Llanuras de Santa Clara, 300
Sci. 18: 281. 1928. Type: COSTA RICA: m, Feb 1896, J. D. Smith 6584 (Syntype US,
Cartago, vicinity of Orosi, 30 Mar 1924, P. C. n.v.); same locality, J. D. Smith 6586
Standley 39695 (Holotype US 1153168 (Syntype US, n.v.).—Psychotria elata (Sw.)
(photos MEXU, PTBG)). Not Psychotria Hammel
latistipula Benth. in Hook., 1849. Cephaelis pittieri K. Krause, Bot. Jahrb. Syst.
—Psychotria cartagoensis Nepokroeff 54, Beibl. 119: 45. 1916. Type: COSTA
Cephaelis nana (Standi.) Standi., J. Wash. Acad. RICA: Puntarenas, Cafias Gordas, 1100 m,
Sci. 17: 171. 1927. Evea nana Standi., J. Feb 1897, H. Pittier 11162 (Holotype B,
Wash. Acad. Sci. 15: 105. 1925. Type: presumably destroyed (photos F, MO;
PANAMA: Canal Zone, hills north of Fnjoles, fragment F); Isotype US (photos MEXU,
19 Dec 1923, P. C Standley 27550 (Holotype PTBG)). not Psychotria pittieri Standi. 1916.
US 1153871 (photos MEXU, PTBG); Isotype —Psychotria aurantibracteata C. M. Taylor
F). Not Psychotria nana K. Krause 1908. Cephaelis psychotriifolia Seem., Bot. Voy.
—Psychotria guapilensis (Standi.) Hammel Herald 138. 1852 [aspsychotriaefolia]. Type:
Cephaelis nicaraguensis Standi., Trop. Woods PANAMA: Panama, damp forest near Cruces;
16: 46. 1928. Type: NICARAGUA: Region of and Darien, Chirambira, Seemann 482
Braggmann's Bluff, on bank of Kukalaya (Holotype BM, n.v. (photo MO); Isotype K
River near Camp 14, 200 ft., 8 Dec 1927, [as Seemann 482bis]).—Psychotria
F. C. Englesing 58 (Holotype F 572595 psychotriifolia (Seem.) Standi.
(photo PTBG)).—Psychotria guapilensis Cephaelis punicea Vahl, Eclog. Amer. 1: 19.
(Standi.) Hammel 1796. Type: JAMAICA: Without precise
Cephaelis ostreophora Wernham, J. Bot. 55: locality, von Rohr s.n. (n.v.).—Psychotria
284. 1917. Type: COLOMBIA: Bogota, elata (Sw.) Hammel
Servita y Villavicencio, Triana s.n. (Holotype Cephaelis rigidifolia Dwyer & Hayden, Ann.
BM, n.v.; Isotype F).—Psychotria Missouri Bot. Gard. 55: 37. 1968. Type:
ostreophora (Wernham) C. M. Taylor PANAMA: Panama, Cerro Jefe, ca. 2700 ft.,
Cephaelis panamensis Dwyer, Ann. Missouri 20 Aug 1967, J. D. Dwyer 8075A (Holotype
Bot. Gard. 67: 77. 1980. Type: PANAMA: MO 2165097 (photos MEXU, PTBG)).
Colon, Rio Guanche, ca. 2.5 km upriver from Palicourea.—Status uncertain.
bridge on road to Portobelo, along small creek Cephaelis sessilifolia Standi., Publ. Field
to the S of the river, 10-100 m, 14 Dec 1974, Columbian Mus., Bot. Ser. 4: 295. 1929.
S. Mori & J. Kallunki 3714 (Holotype MO Type: MEXICO: Jalisco, Quimixto, trail from
2353037 (photos MEXU, PTBG); Isotypes San Pedro el Tuito, 60 m, 2 Dec 1926, Y.
MO 2353038 (photos MEXU, PTBG), PMA Mexia 1240 (Holotype F 579912 (photo
22701, n.v).—Rudgea panamensis (Dwyer) PTBG); Isotypes A, n.v., MO (photos MEXU,
C. M. Tayor PTBG), US (photos MEXU, PTBG)). Not
Cephaelis panchocoensis Dwyer, Ann. Missouri Psychotria sessilifolia M. Martens & Galeotti,
Bot. Gard. 67: 78. 1980. Type: PANAMA: 1844—Psychotria molinarum Lorence
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Cephaelis tetragona Donn. Sm., Bot. Gaz. •'••us occidentalis I var. salicifolius
(Crawfordsville) 61: 376. 1916. Type: (Humb. & Bonpl.) A. Gray, Syn. PI. 1: 29.
COSTA RICA: Forets de Tuis, Oct 1897, 1878. Cephalanthus salicifolius Humb. &
Tonduz 11354 (Syntype US 943500 (photos Bonpl., PL Aequin. 2: 63. 1809. Type:
MEXU, PTBG)), same locality, Tonduz 12997 MEXICO: [Probably Guerrero], near
(Syntype US). Not Psychotria tetragona Acapulco, Humboldt & Bonpland s.n.
Seem., 1867.—Psychotria chiapensis Standi. (Holotype presumably P-Bonpland, n.v.).
Cephaelis tomentosa (Aubl.) Vahl, Eclog.
Amer. 1: 19. 1796. Tapogomea tomentosa Cephalanthus peroblongus Wernham, J. Bot.
Aubl., Hist. PI. Guiane 1: 160, pi 61. 1775. 55: 176. 1917. Type: MEXICO: Nayarit,
Type: FRENCH GUIANA: In sylvarum locis Tepic, 31 May-3 Jun 1837, G. W. Barclay
apertis, Aublet s.n. (Holotype presumably P, 1193 (Holotype BM, n.v.; Isotype US (photos
n.v.; Isotype B, presumably destroyed (photo MEXU, PTBG)).—Cephalanthus salicifolius
MO)). Not Psychotria tomentosa Mull. Arg. Humb. & Bonpl.
ex Mart., 1881.—Psychotria poeppigiana Cephalanthus salicifolius Humb. & Bonpl., PL
Mull. Arg. Aequin. 2: 63. 1809. Type: MEXICO:
Cephaelis tonduzii K. Krause, Bot. Jahrb. Syst. [Probably Guerrero] near Acapulco,
54, Beibl. 119: 45. 1916. COSTA RICA: Bei Humboldt & Bonpland s.n. (Holotype
Tuis, 650 m, Nov 1897, A. Tonduz 11461 presumably P-Bonpland, n.v.).
(Holotype B, presumably destroyed (photos F,
PTBG; fragment F (photo PTBG)). Not Chimarrhis Jacq., Sel. Stirp. Amer. 61. 1763.
Psychotria tonduzii Standi., 1925. Type: Chimarrhis cymosa Jacq.
—Psychotria guapilensis (Standi.) Hammel Chimarrhis decurrens Steyerm., Ceiba 3:18.
Cephaelis vultusmimi Dwyer, Ann. Missouri 1952. Type: PANAMA: Code, El Valle, 600
Bot. Gard. 67: 81, fig. 17. 1980. Type: m, 28 Jun 1946, P. H. Allen 3566 (Holotype F
PANAMA: Colon, Santa Rita, E. Ridge, 23 1394240 (photo PTBG); Isotypes F, 2 sheets,
Mar 1972, A. Gentry & J. D. Dwyer 4811 MO).—Allenanthus erythrocarpa Standi.
(Holotype MO 2164626 (photo PTBG); Chimarrhis latifolia Standi., Publ. Field
Isotype MO (photo PTBG)).—Psychotria Columbian Mus., Bot. Ser. 4: 265. 1929.
poeppigiana Mull. Arg. Type: PANAMA: Chiriqui, Progreso,
Cephaelis williamsii Standi., Publ. Field Jul-Aug 1927, G. P. Cooper & G. M. Slater
Columbian Mus., Bot. Ser. 8: 185. 1930. 260 (Holotype F 573160 (photo PTBG);
Type: PERU: Loreto, Manfina, on upper Rio Isotype US (photos MEXU, PTBG)).
Nany, 29 Jun 1929, L. Williams 1114 Chimarrhis parviflora Standi., Trop. Woods
(Holotype F 604625). Not Psychotria 11: 26. 1927. Type: PANAMA: Bocas del
williamsii Standi., 1930.—Psychotria Toro, Changuinola Valley, 1927, G. P.
zevallosii C. M. Taylor Cooper & G. M. Slater 120 (Holotype US
1315890 (photos MEXU, PTBG); Isotypes A,
Cephalanthus L., Sp. PI. 1: 95. 1753. Type: n.v., F, NY, n.v.).
Cephalanthus occidentalis L.
Cephalanathus berlandieri Wernham, J. Bot. Chiococca P. Browne, Civ. Nat. Hist. Jamaica
55: 175. 1917. Type: MEXICO: Berlandier 164. 1756. Type: Chiococca alba (L.) Hitchc.
1620 (Holotype BM, n.v.).—Cephalanthus Chiococca alba (L.) Hitchc, Annual Rep.
Missouri Bot. Gard. 4: 94. 1893. Lonicera
Cephalanthus occidentalis L., Sp. PL 1: 95 alba L., Sp. PL 175. 1753. Type: JAMAICA
1753. Type: America septentrionali, Herb. or BARBADOS: According to Howard
Linn. 118.1 (Lectotype LINN, (1989), a specimen in the Sloan herbarium
by Reveal, 1993). (BM) may be an appropriate lectotype.
Chiococca axillares Sesse & Moc., La Chiococca jefensis Dwyer, Ann. Missouri Bot.
Naturaleza ser. 2, 1: 36. 1888. Type: Gard. 67: 88. 1980. Type: PANAMA:
MEXICO: Habitat in Tepetlpae prope Sancti Panama, La Eneida, region of Cerro Jefe, R.
Angeli montibus, Sesse & Mocino s.n. Dressier 3296 (Holotype MO).—Elaeagia
(Holotype presumably MA, n.v.; Fl. Mex. Ic. nitidifolia Dwyer
50 (G-DC, n.v.)).—Status uncertain. Chiococca macrocarpa M. Martens & Galeotti,
Chiococca belizensis Lundell, Amer. Midi. Bull. Acad. Roy. Bruxelles 11: 230. 1844.
Naturalist 29: 492. 1943. Type: BRITISH Type: MEXICO: Veracruz, bois de Mirador,
HONDURAS (BELIZE): Toledo Distr., Cow 3000 pieds, H. Galeotti 7064 (Holotype
Pen, near Monkey River, 2 Sep 1942, P. H. presumably BR, n.v.).—Chiococca alba (L.)
Gentle 4115 (Holotype MICH, n.v.; Isotypes Hitchc.
A, n.v., F (photo F), MO (photos MEXU, Chiococca mexicana Lundell, Wrightia 5: 71.
PTBG), NY, n.v., TEX-LL, PTBG), US 1974. Type: MEXICO: Puebla, Mun. de
(photos MEXU, PTBG)). Zacapoaxtla, Tespilco, 1800 m, 23 Apr 1970,
Chiococca caputensis Lorence & C. M. Taylor, F. Ventura A. 955 (Holotype TEX-LL (photos
Novon 5: 210, fig. IB, C. 1995. Type: MEXU, PTBG); Isotype ENCB).—Chiococca
PANAMA: Panama, Cerro Jefe, NE of pachyphylla Wernham
Panama City, 9°15'N, 79°30'W, 12 Jul 1986, Chiococca nocturna Sesse & Moc., Fl. Mexic,
G. McPherson 9744 (Holotype MO 3624428; ed. 2, 59. 1894. Type: MEXICO: Habitat in
Isotype PMA, n.v.). Tuxtlae montibus, floret Julio, Sesse &
Chiococca coriacea M. Martens & Galeotti, Mocino (Holotype presumably MA, n.v.). Not
Bull. Acad. Roy. Bruxelles 11: 231. 1844. Chiococca nocturna Jacq. 1761.—Status
Type: MEXICO: Veracruz, environs de Vera uncertain, probably Cestrum nocturnum L.
Cruz, J. Linden 423 (Holotype presumably (Solanaceae), as protologue states leaves
BR, n.v.; Isotype MICH, n.v.). alternate, flowers nocturnal, smelling of
Chiococca durifolia Dwyer, Ann. Missouri Bot. jasmine.
Gard. 67: 87. 1980. Type: PANAMA: Code, Chiococca oaxacana Standi., Publ. Field Mus.
woods adjacent to chicken farm, La Mesa, Nat. Hist., Bot. Ser. 22: 383. 1940. Type:
above El Valle, 3 Jan 1974, J. D. Dwyer MEXICO: Oaxaca, Cerro de San Antonio,
11865 (Holotype MO 2188155 (photos 1650 m, 26 Jun 1906, C. Conzatti 1418
MEXU, PTBG)).—Chiococca belizensis (Holotype F 579006 (photos F, PTBG);
Lundell Isotypes GH (fragment), US).
Chiococca filipes Lundell, Contr. Univ. Chiococca pachyphylla Wernham, J. Bot. 51:
Michigan Herb. 7: 53. 1942. Type: MEXICO: 323. 1913. Type: MEXICO: Veracruz, entre
Chiapas, Nuevo Amatenango, 1300 m, 17 Jul Acatlan et Chiconquiaco, Schiede & Deppe
1941, E. Matuda 4783 (Holotype MICH, n.v.; (Holotype K, n.v.).
Isotypes A, n.v., MO (photos MEXU, PTBG), Chiococca paniculata Sesse & M09., Fl. Mexic,
NY, n.v., TEX-LL (photos MEXU, PTBG), ed. 2, 58. 1894. Type: MEXICO: Habitat in
US (photos MEXU, PTBG)).—Chiococca calidis Cordovae montibus, floret Augusto,
alba (L.) Hitchc. Sesse & Mocino (Holotype presumably MA,
Chiococca henricksonii M. C. Johnst., n.v.). Not Chiococca paniculata Hoffmanns,
Madrono 28: 30. 1981. Type: MEXICO: ex Roem. & Schult., 1819.—Status
Coahuila, 1-2 km N. of Puerto Colorado, near
crest at SW end of Sierra de la Fragua, near Chiococca petenensis Lundell, Wrightia 5: 7.
26°45TSf, 102°30*W, 2 Sep 1941, /. M. 1972. Type: GUATEMALA: Peten, Dolores,
Johnston 8738 (Holotype TEX 301584, n.v.; in high forest on Santo Toribio trail, about
Isotypes F (photo PTBG), GH (photo PTBG), 200 m west, 7 Jun 1961, E. Contreras 2439
MICH, n.v.). (Holotype TEX-LL (photo F)).
i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Chiococca petrina Wiggins, Contr. Dudley 265 (Holotype DS 20437, n.v.; Isotype UC
Herb. 3: 76,pi. 20, f. 5-13. 1940. Type: (n.v.)).—Status uncertain, probably
MEXICO: Sonora, gorge in andesitic rock 23 Chiococca pueblensis Lundell
miles S. of Divisadero, 26 Sep 1934, /. L. Chiococca pueblensis Lundell, Wnghtia 5: 7.
Wiggins 7493 (Holotype DS 263330, n.v.; 1972. Nom. nov. for Chiococca pubescens
Isotypes F, TEX-LL, n.v., US (photos MEXU, Standi., not Chiococca pubescens Humb. &
PTBG)). Bonpl. ex Roem. & Schultes, 1819.
Chiococca phaenostemon Schltdl., Linnaea 9: Chiococca racemosa L., Syst. Nat. ed. 10, 2:
594. 1834. Type: MEXICO: Veracruz, "In 917. 1759. Nom. illegit., citing Lonicera alba
sylvis Jalapae, Majo florens, ad Hac. d. 1. L—Chiococca alba (L) Hitchc.
Laguna Jul. fructifer," collector not cited Chiococca racemosa Sesse & Moc., Fl. Mexic,
(Holotype presumably HAL, n.v.). Note: no ed. 2, 58. 1893, nom. illegit. Not Chiococca
actual specimens or collector(s) were cited in racemosa L., 1759, nor C. racemosa Schltdl.
Schlechtendal's protologue (1834), merely "In & Cham., 1830. Type: MEXICO: Sesse &
sylvis Jalapae Majo florens, ad Hac. d. 1. Moqiho (presumably MA, n.v.).—Status
Laguna Jul. fructifer." There is a Schiede 255 uncertain, probably Chiococca alba (L.)
specimen at Paris (with a duplicate at MO) in Hitchc.
Schlechtendal's hand stating only "Mexique." Chiococca racemosa L. var. yucatana Loes.,
Schlechtendal's types are mostly at HAL, so Repert. Sp. Nov. 18: 361. 1922. Type:
the holotype [or possibly syntypes] is MEXICO: Yucatan, im Distrikt Ticul, auf der
presumably deposited there. Schiede is known Hacienda Yaxche, E. Seler & C Seler 5591
to have collected at Hacienda de la Laguna, (Holotype B, presumably destroyed).
Veracruz, and therefore the Schiede 255 —Chiococca alba (L.) Hitchc.
specimens at P & MO are very likely original Chiococca rubriflora Lundell, Wrightia 5: 7.
material, probably isotypes. However, 1972. Type: GUATEMALA: Peten, west of
mention of "Jalapa" suggests that km 165 of Poptun road, 10 Sep 1969, E
Schlechtendal may have been referring to Contreras 9110 (Holotype TEX-LL (photos
another collection (syntype) as well. This is a MEXU, PTBG); Isotype F (photos F, PTBG)).
vining species with a short, internally villous Chiococca semipilosa Standi. & Steyerm., Publ.
corolla rube and exserted stamens with Field Mus. Nat. Hist., Bot. Ser. 22: 279. 1940.
filaments villous in the basal half. Collections Type: GUATEMALA: Chiquimula, Cerro
from southern Mexico and Central America Caracol, 1.5 miles north of Quezaltepeque,
erroneously identified as Chiococca 1200-1400 m, 7 Nov 1939, J. A. Steyermark
phaenostemon have the habit of a small tree, a 31406 (Holotype F 1039351 (photo PTBG)).
glabrous corolla, and actually represent C Chiococca sessilifolia Miranda, Anales Inst.
pachyphylla Wernham. Biol. Mexico 21(2): 306,/ 4. 1950 [1951].
Chiococca pubescens Standi., Contr. U.S. Natl. Type: MEXICO: Chiapas, en borde de bosque
Herb. 20: 209. 1919. Type: MEXICO: Puebla, bajo hacia Las Vistas, al N de Berriozabal, 24
vicinity of San Luis Tultitlanapa, Jul 1908, Sep 1950, F. Miranda 6672 (Holotype
C. A. Purpus 3334 (Holotype US 840975 MEXU).
(photos MEXU, PTBG); Isotypes F, P). Not
Chiococca pubescens Humb. & Bonpl. ex Bull. Acad. Bruxelles 11: 231. 1844. Type:
Roem. & Schultes, 1819.—Chiococca MEXICO: Veracruz, environs de Xalapa,
4000 pieds, / Linden 48 (Holotype BR, n.v).
Chiococca pubescens Standi, var. peninsularis —Status uncertain, probably Chiococca
Wiggins, Contr. Dudley Herb. 4: 24. 1950. phaenostemon Schltdl.
Type: MEXICO: Baja California, Sierra San Chiococca steyermarkii Standi., Publ. Field
Francisquito, 18 Oct 1890, T. S. Brandegee Mus. Nat. Hist., Bot. Ser. 22: 280. 1940.
Type: GUATEMALA: Dept. San Marcos, Rio Chione chiapasensis Standi., Publ. Field Mus.
Vega, near San Rafael and Guatemala-Mexico Nat. Hist, Bot. Ser. 22: 110. 1940. Type:
boundary, Volcan de Tacana, 2500-3000 m, MEXICO: Chiapas, Palenque, Rvina [Ruina],
20 Feb 1940, J. A. Steyermark 36237 river side in virgin forest, Jul 1939, E. Matuda
(Holotype F 1043222 (photos F, PTBG); 3675 (Holotype F 1011456 (photo PTBG);
Isotype F 1043229). Isotypes A, n.v., US (photos MEXU, PTBG),
Chiococca vestita Lundell, Wrightia 5: 8. 1972. MEXU).
Type: GUATEMALA: Dept. Peten, Poptun, 2 Chione costaricensis Standi., Publ. Field Mus.
km east, in hammock in pineland, 13 Jul Nat. Hist., Bot. Ser. 22: 111. 1940. Type:
1959, C L. Lundell 16425 (Holotype TEX- COSTA RICA: Alajuela, Canton de San
LL, n.v. (photo F)).—Chiococca semipilosa Carlos, Villa Quesada, Caribbean rain forest,
Standi. & Steyerm. 825 m, 14 Mar 1939, Austin Smith F1778
Chiococca vestita Lundell var. glaberrima (Holotype F 980765 (photo PTBG); Isotype
Lundell, Wrightia 5: 8. 1972. Type: US (photos MEXU, PTBG)).—Chione
GUATEMALA: Dept. Peten, Poptun, 2 km sylvicoia (Standi.) W. C. Burger
east, in hammock in pineland, 13 Jul 1959, Chione dariensis Dwyer, Ann. Missouri Bot.
C L. Lundell 16425A (Holotype TEX-LL, Gard. 67: 94. 1980. Type: PANAMA: Darien,
n.v. (photo F)).—Chiococca semipilosa Rio Balsa, between Manene and Tusijuanda,
Standi. & Steyerm. 26 Jul 1967, J. Duke 13571 (Holotype MO
1968196 (photos MEXU, PTBG); Isotypes
Chione DC, Prodr. 4: 461. 1830. Type: Chione MO, 2 sheets, US (photo PTBG)).
glabra DC. Chione guatemalensis Standi. & Steyerm.,
Chione allenii L. O. Williams, Phytologia 25: Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 174.
462. 1973. Type: COSTA RICA: Golfo Dulce 1940. Type: GUATEMALA: Dept. Izabal,
area, vicinity of Esquinas Experiment Station, Rio Dulce, between Livingston and 6 miles up
sea level, 8 Jul 1949, P. H. Allen 5321 river, on north side, 1-25 m, 14 Apr 1940,
(Holotype F 1679433; Isotypes MO (photo J. A. Steyermark 39375 (Holotype F 1035017
PTBG), TUL (photo PTBG)).—Chione (photo PTBG)).
sylvicoia (Standi.) W. C. Burger Chione mexicana Standi., J. Wash. Acad. Sci.
Chione buxifolia Dwyer & Hayden, Phytologia 17: 340. 1927. Type: MEXICO: Veracruz,
15: 55. 1967. Type: PANAMA: Panama, Zacuapan, open forests, Jun 1926, C. A.
Cerro Jefe, E. Tyson et al. 3291 (Holotype Purpus 10757 (Holotype US 1266079 (photos
MO 2210380 (photo PTBG); Isotypes F, NY, MEXU, PTBG); Isotypes A, n.v., F, 2 sheets,
n.v.). MO, 2 sheets (photos MEXU, PTBG)).
Chione campanensis Dwyer, Ann. Missouri Chionepanamensis Steyerm., Ceiba 3: 19.
Bot. Gard. 67: 92. 1980. Type: PANAMA: 1952. Type: PANAMA: Chiriqui, Boquete
Cerro Campana, trail to Campana Ridge, 2300 region, Cerro Horqueta, 6500 ft., 1 Jul 1940,
ft., 18 Sep 1968, A R. Wilbur, R. Foster, C. von Hagen & W. von Hagen 2137
H. Kennedy & R. Dressier 1020 (Holotype (Holotype F 1397595 (photo PTBG); Isotype
MO 2224284 (photo PTBG)). MO, n.v.).—Chione sylvicoia (Standi.) W. C.
Chione chambersii Dwyer & Hayden, Ann.
Missouri Bot. Gard. 54: 138. 1967. Type: Chione sylvicoia (Standi.) W. C. Burger,
PANAMA: Comarca de Bani, area near Selbyana 12: 138. 1991. Chomelia ? sylvicoia
Puerto Armuelles, between Canazo and Standi., J. Wash. Acad. Sci. 18: 182. 1928.
Cocos, 17 Jun 1957, W. I. Stern & K L. Type: COSTA RICA: Heredia, Yerba Buena,
Chambers 144 (Holotype MO 1739838; northeast of San Isidro, 2000 m, 22-28 Feb
Isotype GH (photo PTBG)).—Chomelia 1926, P. C. Standley & J. Valerio 49196
i Donn. Sm. (Holotype US 1305902 (photo PTBG)).
i NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Chomelia Jacq., Enum. PI. Carib. 12. 1760. C, n.v. (photo US), fragment US (photo
Type: Chomelia spinosa Jacq. PTBG)).—Chomelia spinosa Jacq.
Chomelia atlantica Dwyer, Ann. Missouri Bot. Chomelia grandicarpa Dwyer, Ann. Missouri
Gard. 67: 96. 1980. Type: PANAMA: Colon, Bot. Gard. 67: 97. 1980. Type: PANAMA:
swamp forest and bordering terra firma, ca. Colon, 2-3 miles up Rio Guanche, lowland
0.5 km from the Rio Buenaventura, near rain forest, 10-20 m, 19 Jan 1973,
Portobelo, 30 Jan 1973, H. Kennedy & S. Gra H. Kennedy & R. Foster 2174 (Holotype MO.
2242 (Holotype MO 2161971 (photo PTBG); 2465399 (photo PTBG); Isotype MO).
Isotype MO)).—Chomelia barbellata Standi. Chomelia leucophylla Dwyer, Ann. Missouri
Chomelia barbata Standi, J. Wash. Acad. Sci. Bot. Gard. 67: 98. 1980. Type: PANAMA:
13: 7. 1923. Type: MEXICO: Oaxaca, Darien, slopes of Cerro Chucula, drainage of
Juquila, vicinity of Chacahua, 5 m, 17 Dec Rio Pavarando, 11 Feb 1972, A. Gentry 4262
1921, C. Conzatti 4475 (Holotype US (Holotype MO; Isotype MO). Not Rondeletia
1110841 (photo US)). leucophylla Kunth, 1820.—Rondeletia
Chomelia barbellata Standi., Field Mus. Nat. dwyeri Lorence
Hist., Bot. Ser. 13(7): 120. 1936. Type: Chomelia microloba Donn. Sm., Bot. Gaz.
PERU: San Martin, Klug 3920 (Holotype F, (Crawfordsville) 31: 114. 1901. Type:
n.v.). COSTA RICA: Santo Domingo de Golfo
Chomelia brachyloba Standi., Contr. U.S. Natl. Duke, Feb 1896, A. Tonduz 7048 (Herb. Nac.
Herb. 18: 136. 1916. Type: PANAMA: Costa Rica 9874) (Holotype US 943460
Panama, Sabana de San Juan Corso, near (photo PTBG); Isotypes A, n.v., MO (photos
Chepo, 60-80 m, Oct 1911, H. Pittier 4673 MEXU, PTBG), P (photos MEXU, PTBG),
(Holotype US 679833 (photo PTBG)). US (3 sheets; photos PTBG)). Anisomeris.
—Chomelia spinosa Jacq. Chomelia panamensis (Standi.) Dwyer, Ann.
Chomelia brachypoda Donn. Sm., Bot. Gaz. Missouri Bot. Gard. 67: 100. 1980. Antirhea
(Crawfordsville) 47: 255. 1909. Type: panamensis Standi., N. Amer. Fl. 32: 264.
GUATEMALA: Alta Verapaz, Sasis, banks of 1934. Type: PANAMA: San Bias District,
Rio Ogewaj, 900 m, May 1908, H. von 21-28 Apr 1933, G. P. Cooper 238 (Holotype
Tuerckheim II2253 (Holotype US 941389 F 686345 (photo PTBG); Isotype F 709955).
(photos PTBG, US); Isotypes F (photos —Chomelia microloba Donn. Sm.
MEXU, PTBG), US (photo PTBG), W). Chomelia peninsularis Dwyer, Ann. Missouri
Anisomeris. Bot. Gard. 67: 101. 1980. Type: PANAMA:
Chomelia coclensis Dwyer, Ann. Missouri Bot. Chiriqui, Burica Peninsula, along Quebrada
Gard. 67: 97. 1980. Type: PANAMA: Code, Punta de Piedra, 2 mi SE of Puerto Armuelles,
La Mesa above El Valle, 800 m, T. B. Croat 0-100 m, 1 Mar 1973, T. B. Croat 22440
25396 (Holotype MO 2210285; Isotype MO (Holotype MO 2785727 (photo PTBG);
2219529).—Malanea colombiana Standi. Isotype F).—Chomelia microloba Donn. Sm.
Chomelia englesingii Standi., Trop. Woods 16: Chomelia pringlei S. Watson, Proc. Amer.
45. 1928. Type: NICARAGUA: Region of Acad. Arts 26: 137. 1891. Type: MEXICO:
Braggman's Bluff, Kukalaya River bank west San Luis Potosi, Tamasopo Canyon, 6 Aug
of Camp 14, 2 Dec 1927, F. C Englesing 49 1890, C. G. Pringle 3209 (Holotype GH, n.v.;
(Yale 1233) (Holotype F 572606 (photo Isotypes A, n.v., F (photos MEXU, PTBG),
PTBG)). Anisomeris.—Chomelia recordii MO (photos MEXU, PTBG), P (photos
Standi. MEXU, PTBG), US). Anisomeris.
Chomelia filipes Benth. in Oerst., Vidensk. Chomelia protracta (Bartl. ex DC.) Standi.,
Meddel Dansk Naturhist. Foren. Kjobenhavn Contr. U.S. Natl. Herb. 23: 1384. 1926.
1852: 41. 1853. Type: NICARAGUA: Guettarda protracta Bartl. ex DC. Prodr. 4:
Granada, Oersted s.n. (Holotype presumably 457. 1830. Type: MEXICO: Without precise
locality, Haenke s.n. (Holotype presumably Chomelia venulosa W. C. Burger & C. M.
PR, n.v.). Anisomeris, Pittoniotis. Taylor, Fieldiana, Bot, n.s. 33: 101. 1993.
Chomelia psilocarpa Dwyer & Hayden, Ann. Type: COSTA RICA: Alajuela, Reserva
Missouri Bot. Gard. 54: 139. 1967. Type: Biologica Monteverde, E. Bello 414
PANAMA: Canal Zone, Barro Colorado (Holotype CR, n.v.; Isotypes F, n.v., MO).
Island, SE shoreline from lab, Aug 1965,
M. V. Hayden 143 (Holotype MO 1831286 Choristes Benfh., PI. Hartw. 63. 1840. Type:
(photo PTBG); Isotypes MO, 3 sheets). Choristes obtusiflora Benth. [Herein treated as
Chomelia purpusii (Brandegee) Rusby, Bull. a synonym of Deppea Cham. & Schltdl.]
Torrey Bot. Club 52: 138. 1925. Anisomeris Choristes cornifolia (Benth.) Benth., PI. Hartw.
purpusii Brandegee, Univ. Calif. Publ. Bot. 6: 64. 1840. Rondeletia cornifolia Benth., PI.
70. 1914. Type: MEXICO: Chiapas, Tonala, Hartw. 39. 1840. Type: MEXICO:
Sep 1913, C. A. Purpus 6940 (Holotype UC, Michoacan, prope Morelia, T. Hartweg 299
n.v.; Isotypes F (photo PTBG), US (photo (Holotype K (photo MEXU); Isotypes BM, P,
PTBG))./ " - US (fragment), W, 2 sheets).—Deppea
cornifolia (Benth.) Benth.
i Standi., Trop. Woods 7: 9. Choristes obtusiflora Benth., PI. Hartw. 63.
1926. Type: GUATEMALA: Izabal, Los 1840. Type: MEXICO: Oaxaca, in plamtie
Andes to Entre Rios, lower Rio Motagua Llano Verde, T. Hartweg 473 (Holotype K;
Valley, 1 Mar 1926, S. J. Record 8862 Isotypes P, F, W, 2 sheets).—Deppea
(Holotype US 1209919 (photo PTBG); obtusiflora (Benth.) Benth.
Isotype NY, n.v.). Anisomeris.
Chomelia recordii Standi, var. paucipubescens Cigarrilla Aiello, J. Arnold Arbor. 60: 109.
Dwyer, Ann. Missouri Bot. Gard. 67: 103. 1979. Type: Cigarilla mexicana (Zucc. &
1980. Type: PANAMA: Canal Zone, near Mart, ex DC.) Aiello. [Herein treated at a
archeological site, edge of Madden Lake, A. synonym of Nernstia Urb.]
Gentry & E. Tyson 5038 (Holotype MO (not Cigarrilla mexicana (Zucc. & Mart, ex DC.)
located 6/96)).—Chomelia recordii Standi. Aiello, J. Arnold Arbor. 60: 110. 1979.
Chomelia rudis (Standi.) Lorence, Novon 7: Coutarea mexicana Zucc. & Mart, ex DC,
48. 1997. Anisomeris rudis Standi., Field Mus Prodr. 4: 350. 1830. Type: MEXICO: Without
Nat. Hist, Bot. Ser. 17: 394. 1938. Type: precise locality, Martius s.n. in 1829
HONDURAS: Comayagua, summit of the (Holotype G-DC, n.v. (microfiche IDC
ridge above El Achote, above the plains of 800.680: II.7)).—Nernstia mexicana (Zucc.
Siguatepeque, 1800 m, 1 Aug 1936, G. T. & Mart, ex DC.) Urb.
Yuncker, R. F. Dawson & H. R. Youse 6253
(Holotype F 857678 (photo PTBG); Isotype Cinchona L., Sp. PI. 1: 172. 1753. Type:
\I< ) 1 Cinchona officinalis L.
Chomelia spinosa Jacq., Enum. PI. Carib 12 Cinchona caribaea Jacq., Enum. PI. Carib. 16.
1760. Type: COLOMBIA: Cartagena, in 1760. Type: CUBA: Thickets near Havana,
sylvaticis ad pendem montis de La Po'pa, Jacquin s.n. (location unknown).—Exostema
Jacquin s.n. (location unknown). Ixora. ' caribaeum (Jacq.) Roem. & Schultes
Chomelia sylvicola Standi., J. Wash Acad Sci Cinchona grandiflora Ruiz & Pav., F. Peruv. 2:
18: 182. 1928. Type: COSTA RICA: Heredia, 54, pi. 89. 1799. Type: PERU: Huanuco,
Yerba Buena, northeast of San Isidro 2000 m Andes de Peru, ad Pozuzo et San Antonii de
22-28 Feb 1926, P. C Standley & J Valeria ' Playa grande, Ruiz Lopez & Pavon s.n.
49196 (Holotype US 1305902 (photo (Holotype presumably MA, n.v.; Isotype B,
PTBG)). Anisomeris.—Chione sylvicola destroyed (photo P)).—Cosmibuena
(Standi.) W. C. Burger grandiflora (Ruiz & Pav.) Rusby
Cinchona macrocarpa Vahl, Schr. Naturh. Ges. Huitamulco et Cuapa, Schiede & Deppe 1273
^Copenhagen 1(1): 20. 1793. Type: (Holotype presumably HAL, n.v.).—Status
COLOMBIA: In regno Santa Fe, Ortega s.n. uncertain, probably Coccocypselum
(Lectotype BM, n.v.; designated by hirsutum Bartl. ex DC.
Andersson in Andersson & Taylor, 1994: 16). Coccocypselum cordifolium Nees & Mart.,
—Ladenbergia macrocarpa (Vahl) Klotzsch Nova Acta Phys.-Med. Acad. Caes. Leop.-
Cinchona officinalis L., Sp. PI. 1: 172. 1753. Carol. Nat. Cur. 12: 14. 1824. Type: BRAZIL:
Type: ECUADOR: Habitat in Loxa Pemviae, Ad viam Felisbertiam, Wied-Neuwied
Quinquina La Condamine, Hist. Acad. Roy. (location unknown).
Sci. Paris 1738: 226, t. 5-6. 1738. (For Coccocypselum glabrum Bartl. ex DC, Prodr. 4:
typification see Andersson & Taylor, 1994: 397. 1830. Type: PANAMA: Without precise
locality, Haenke s.n. (Holotype presumably
Cinchona pubescens Vahl, Skr. Naturhist. PR, n.v., fragment G-DC, n.v. (microfiche 4:
Selsk. Kjoebenhavn 1: 119. 1790. Type: 397.14)). Tontanea.—Coccocypselum
Probably ECUADOR: precise locality hirsutum Bartl. ex DC. var. glabrum (Bartl.
unknown, J. deJussieu s.n. (Holotype C, n.v. ex DC.) L. O. Williams
(photos F, MO); Isotype P-JU 9900, n.v.). Coccocypselum guianense (Aubl.) K. Schum.
Cinchona rosea Ruiz & Pav., Fl. Peruv. 2: 54. in Mart., Fl. Bras. 6(6): 315. 1889. Tontonea
1799. Type: PERU: Huanuco, Pozuzo, Ruiz guianensis Aubl., Hist. PI. Guiane 1: 108, tab.
Lopez & Pavon s.n. (Lectotype MA, n.v.; 42. 1775. Type: FRENCH GUIANA: Sylvis
Isolectotypes BM, n.v., G, n.v., G-DC, n.v., Cayenne, Aublet s.n. (Holotype presumably P
K, n.v., MA, n.v., P, n.v.; designated by or BM, n.v.).
Andersson in Andersson & Taylor, 1994: 12). Coccocypselum herbaceum P. Browne, Civ.
—Macrocnemum roseum (Ruiz & Pav.) Nat. Hist. Jamaica 144, pi. 6 J. 2. 1756. Type:
Wedd. JAMAICA: Cooler mountains of Liguancee
Cinchona succirubra Pav. ex Klotzsch, Abh. and Mount-Diable, P. Browne (LINN, n.v.).
Akad.Berl. 1857: 60. 1858. Type: Tontanea.
ECUADOR: Bolivar, San Antonio on trail to Coccocypselum hirsutum Bartl. ex DC, Prodr.
Guaranda, [Tafalla in] Pavon 464 (Holotype 4: 396. 1830. Type: Precise locality unknown,
B, presumably destroyed (photo F); Isotypes probably Trinidad, Haenke s.n. (Holotype
G, n.v., K, n.v., MA, n.v.).—Cinchona presumably PR, n.v., fragment G-DC, n.v.
pubescens Vahl (microfiche 4: 396.5)). Tontanea.
Coccocypselum hirsutum var. glabrum (Bartl.
Coccocypselum P. Browne, Civ. Nat. Hist. ex DC.) L. O. Williams, Phytologia 25: 462.
Jamaica 144. 1756. [as Coccocipsilum], nom. Coccocypselum glabrum Bartl. ex DC, Prodr.
et orthogr. cons. Type: Coccocypselum repens 4: 397. 1830. Type: PANAMA: Without
Sw., Prodr. Veg. Ind. Occ. 31. 1788, not precise locality, Haenke s.n. (Holotype
Kunth, 1819. presumably PR, n.v., fragment G-DC, n.v.
Coccocypselum canescens Willd. ex Cham. & (microfiche 4: 397.14)).
Schltdl., Linnaea4: 139. 1829. Nom. Meg., Coccocypselum hispidulum (Standi.) Standi.,
not Coccocypselum canescens Willd. ex Publ. Field Columbian Mus., Bot. Ser. 4: 281.
Schult. & Schult., 1827. Type: VENEZUELA: 1929. Tontanea hispidula Standi., N. Amer.
Buenavista prope Caracas, Humboldt s.n. Fl. 32: 147. 1921. Type: PANAMA: Chagres,
(Holotype B-W 2874, n.v). Tontanea. Isthmus of Panama, Jan-Feb 1850, A. Fendler
—Coccocypselum lanceolatum (Ruiz & 179 (Holotype US 42058 (photo MO); Isotype
Pav.) Pers. US).
Coccocypselum ciliatum Cham. & Schltdl.,
Linnaea 6: 414. 1831. Type: MEXICO: Inter
Coccocypselum lanceolatum (Ruiz & Pav.) Trans. Linn. Soc, Ser. 2: 171, tab. 24. 1876.
Pers., Syn. PL 1: 132. 1805. Condalia Type: SIERRA LEONE: Daniell A608
lanceolata Ruiz & Pav., PL Fl. Peruv. 1: 54. (Holotype BM, n.v. (photo MO)).
1798. Type: PERU: In nemoribus Cuchero et Occasionally cultivated in Mexico and Central
Chinchao, Ruiz & Pavon s.n. (Holotype
presumably MA, n.v.). ( offea mexicana DC, Prodr. 4: 501. 1830.
Coccocypselum macropodum (Ruiz & Pav.) Type: MEXICO: Without precise locality,
Britton, Bull. Torrey Bot. Club 18: 107. 1891. Haenke s.n. (Holotype presumably PR, n.v.,
Psychotria macropoda Ruiz & Pav., Fl. fragment G-DC, n.v. (microfiche 4: 501.25)).
Peruv. 2: 63, pi 211, fig. b. 1799. Type: —Rudgea cornifolia (Humb. & Bonpl.)
PERU: Huanuco, nemoribus secus Pillao ad Standi.
Iscutunam, Ruiz & Pavon s.n. (Holotype Coffea obovata Cham. & SchltdL, Linnaea 6:
presumably MA, n.v.).—Geophila 412. 1832. Type: MEXICO: Veracruz, m
macropoda (Ruiz & Pav.) DC. umbrosis prope Misantla, Schiede & Deppe
Coccocypselum pleuropodum (Donn. Sm.) 1268 (Holotype presumably HAL, n.v.).
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. —Status uncertain, probably Hoffmannia
4: 281. 1929. Geophila pleuropoda Donn. sp.
Sm., Bot. Gaz. (Crawfordsville) 52: 50. 1911. Coffea rosea Moc. & Sesse ex DC, Prodr. 4:
Type: GUATEMALA: Alta Verapaz, 499. 1830. Type: MEXICO: Without precise
Secanquim, trail to Sepacuite, 4000 ft., 5 Feb locality, Sesse & Mocino s.n. (presumably G-
1905, G. P. Golls.n. (Holotype US 860647 DC, n.v.; Fl. Mex. Ic. (G-DEL, n.v. (photo
(photos MEXU, PTBG)).—Coccocypselum MO), caique 479 (US)).—Status uncertain,
cordifolium Nees & Mart. probably I
Coccocypselum rothschuhii Loes., Bot. Jahrb.
Syst. 60: 370. 1926. Type: NICARAGUA: Condalia Ruiz & Pav., Prodr. II,pi. 2. 1794.
Matagalpa, Canada Yasica, 950 m, Rothschuh Nomen rejic, not Condalia Cav. (1799)
179 (Holotype B, presumably destroyed). (Rhamnaceae). Type: Condalia repens Ruiz &
—Coccocypselum cordifolium Nees & Mart.
Coccocypselum P. Browne.]
Coffea L., 172. 1753. Type: Coffea Ruiz & Pav., Fl. Peruv.
Chil. 1: 54. 1798. Type: PERU: In nemoribus
Coffea arabica L., Sp. PL 1: 172. 1753. Type: Cuchero et Chinchao, Ruiz Lopez & Pavon
Habitat in Arabia felici, Herb. Clifford 9, s.n. (Holotype presumably MA, n.v.).
Coffea 1 (Lectotype BM, n.v.; designated by —Coccocypselum lanceolatum (Ruiz &
Bridson & Verdcourt, 1988: 713). Widely Pav.) Pers.
cultivated in Mexico and Central America.
Coffea canephora Pierre ex Froehner, Notizbl. Condaminea DC, Prodr. 4: 402. 1830.
Bot. Gard. Berlin 1: 237. 1897. Type: Lectotype: Condaminea corymbosa (Ruiz &
GABON: Gabun, Herb. L. Pierre n. 247 Pav.) DC.
(Holotype presumably P, n.v.). Occasionally Condaminea corymbosa (Ruiz & Pav.) DC,
cultivated in Mexico and Central America. Prodr. 4: 402. 1830. Macrocnemum
Coffea lanceolata Cham. & SchltdL, Linnaea 6: corymbosum Ruiz & Pav., Fl. Peruv. Prodr. 2:
412. 1831. Type: MEXICO: Veracruz, prope 4S, pi. 189. 1799. Type: PERU: In Andium
Jalapam, Schiede & Deppe 1269 (Holotype
presumably HAL, n.v.).—Status uncertain,
possibly Chiococca sp.
Coffea liberica W. Bull ex Hieron., Retail List
New, Beautiful and Rare Plants 97: 4. 1874;
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Condaminea petiolata Dwyer, Ann. Missouri Standi., Publ. Field Mus. Nat. Hist., Bot. Ser.
Bot. Gard. 67: 117. 1980. Type: PANAMA 22: 51. 1940. Type: MEXICO: Chiapas,
Panama, Rio Maje, near new Bayano Lake, Volcan Tacana, 2000^039 m, Aug 1938, E.
<100 m, 5 May 1976, T. B. Croat 34706 Matuda 2327 (Holotype F 980255 (photo F);
(Holotype MO 2398904 (photo PTBG): Isotypes A, n.v., NY, n.v., US (photos
Isotype MO). Rondeletia [H MEXU, PTBG)).
Cosmibuena ovalis Standi., Contr. U.S. Natl.
Cordiera A. Rich., Mem. Soc. Hist. Nat., Paris Herb. 18: 137. 1916. Type: PANAMA: Code,
5: 222, t. 20. 1830. (Herein considered as a vicinity of Ola, 100-350 m, 7-9 Dec 1911,//.
synonym of AI'ibertia A. Rich.) Pittier5074 (Holotype US 715202 (photos F,
Cordiera edulis (Rich). Kuntze, Rev. Gen. PI. 1: MEXU, PTBG); Isotype NY, n.v.).
279. 1891. Genipa edulis Rich., Act. Soc. —Cosmibuena grandiflora (Ruiz & Pav.)
Hist. Nat. Paris 1: 107. 1792. Type: FRENCH Rusby
GUIANA: Guyane, without precise locality, Cosmibuena paludicola Standi., Contr. U.S.
Richard s.n. (Holotype P, n.v.; Isotype F Natl. Herb. 18: 137. 1916. Type: PANAMA:
971706 (photo PTBG)).—Alibertia edulis Colon, vicinity of Viento Frio between Playa
(Rich.) A. Rich. Damas and Viento Frio, along beach and near
sea level, in mangrove formation, 7-8 Aug
Cosmibuena Ruiz & Pav., Fl. Peruv. 3: 2. 1802, 1911, H. Pittier 4107 (Holotype US 679204
nom. cons. Type: Cosmibuena obtusifolia (photos F, MEXU, PTBG); Isotypes GH, n.v.,
Ruiz & Pav., nom. illeg., type. cons. [= NY, n.v.)—Cosmibuena macrocarpa
Cosmibuena grandiflora (Ruiz & Pav.) (Benth.) Klotzsch
Rusby]. Not Cosmibuena Ruiz & Pav., 1794, Cosmibuena skinneri (Oerst.) Hemsl., Biol.
Centr.-Amer., Bot. 2: 12. 1881. Buena
iz & Pav.) Rusby, skinneri Oerst., Vidensk. Meddel. Dansk
Bull. New York Bot. Gard. 4: 368. 1907. Naturhist. Foren Kjobenhavn 1852: 48. 1853.
Cinchona grandiflora Ruiz & Pav., Fl. Peruv. Type: NICARAGUA: In monte ignivomo,
2: 54,/?/. 89. 1799. Type: PERU: Huanuco, Masaya, Jan 1848, Oersted 11134 (Holotype
Andes de Peru, ad Pozuzo et San Antonii de C, n.v. (photos A, F, MO); Isotype US (photos
Playa grande, Ruiz Lopez & Pavon s.n. MEXU, PTBG))—Cosmibuena grandiflora
(Holotype presumably MA, n.v.; Isotype B, (Ruiz & Pav.) Rusby
destroyed (photo F)). Cosmibuena valerii (Standi.) C. M. Taylor,
Cosmibuena holdridgei Monach., Phytologia 3: Ann. Missouri Bot. Gard. 79: 897. 1992.
64. 1949. Type: GUATEMALA: Hillia valerii Standi., J. Wash. Acad. Sci. 18:
Huehuetenango, Barillas to Tokija, Jul 1948, 164. 1928. Type: COSTA RICA: San Jose,
L. R. Holdridge s.n. (Monachino 519) vicinity of La Palma, on the road to La
(Holotype NY 3187, n.v. (photo NY); Isotype Hondura, 1500-1700 m, 17-18 Jul 1923,
NY, n.v. (photo NY)).—Cosmibuena W. R. Maxon & A. D. Harvey 8092 (Holotype
matudae (Standi.) L. O. Williams US 1181776 (photos F, MEXU, PTBG)).
Cosmibuena macrocarpa (Benth.) Klotzsch ex
Walp., Repert. Bot. Syst. 6: 69. 1846. Buena Cosmocalyx Standi., Publ. Field Mus. Nat. Hist.,
macrocarpa Benth., Bot. Voy. Sulphur 104, /. Bot. Ser. 8: 56. 1930. Type: Cosmocalyx
38, f. 1-14. 1845. Not Buena macrocarpa spectabilis Standi.
Wedd. 1869. Type: COLOMBIA: Narino, Cosmocalvx spectabilis Standi., Publ. Field
Island of Gorgona, Hinds 353 (Holotype K, Mus. Nat. Hist., Bot. Ser. 8: 56. 1930. Type:
n.v. (photo NY)). MEXICO: Yucatan, without precise locality,
Cosmibuena matudae (Standi.) L. O. Williams, 1917-1921, G. F. Gaumer 24270 (Holotype F
Fieldiana, Bot. 31: 45. 1965. Hillia matudae
vicinity of Yaviza, 100 ft., 9-11 Jun 1947,
P. H. Allen 4576 (Holotype F 1397594 (photo
PTBG); Isotypes MO, NY, n.v.).—Coussarea
Coussarea Aubl., Hist. PI. Guiane 1: 98. 1775. paniculata (Vahl) Standi.
Type: Coussarea violacea Aubl. Coussarea durifolia Dwyer, Ann. Missouri
Coussarea austin-smithii Standi., Publ. Field Bot. Gard. 67: 124. 1980. Type: PANAMA:
Mus. Nat. Hist, Bot. Ser. 18: 1286. 1938. Panama, along El Llano-Carti Tupile road, 12
Type: COSTA RICA: Region of Zarcero, miles above Panamerican Highway, 200-500
4700 ft., 25 Aug 1937, A. Smith 4226 m, 26-27 Mar 1973, R. Liesner 1180
(Holotype F 905233 (photos MEXU, PTBG)). (Holotype MO 2219526 (photo PTBG)).
Coussarea caroliana Standi., Publ. Field Mus. Coussarea enneantha Standi, J. Wash. Acad.
Nat. Hist., Bot. Ser. 22: 178. 1940. Type: Sci. 18: 282. 1928. Type: PANAMA: Darien,
COSTA RICA: Alajuela, Canton San Carlos, Cana and vicinity, 2000-6500 ft., 17 Apr-8
Suerre, 1000 m, 2 Mar 1939, A. Smith HI645 Jun 1908, R. S. Williams 841 (Holotype US
(Holotype F 981032 (photo PTBG); Isotype 678317 (photos MEXU, PTBG)).
US (photos MEXU, PTBG)). Coussarea euryphylla Standi., Publ. Field Mus.
Coussarea cerroazulensis Dwyer & Hayden, Nat. Hist., Bot. Ser. 4: 294. 1929. Type:
Ann. Missouri Bot. Gard. 53: 374. 1966. PANAMA: Bocas del Toro, Buena Vista
Type: PANAMA: Panama, Cerro Azul, 600 camp on Chiriqui trail, region of Almirante,
m, 6 Aug 1961, J. D. Dwyer 1368 (Holotype Jan-Mar 1928, G. P. Cooper 228 (Holotype F
MO 1810161 (photos MEXU, PTBG)). 579282). Dukea, Raritebe.—Raritebe
Coussarea chiriquiensis (Dwyer) C. M. Taylor palicoureoides Wernham subsp.
in W. C. Burger & C. M. Taylor, Fieldiana, dwyerianum J. H. Kirkbr.
Bot., n.s. 33: 113. 1993. Rudgea chiriquiensis Coussarea garciae Standi., Publ. Field Mus.
Dwyer, Ann. Missouri Bot. Gard. 67: 476. Nat. Hist., Bot. Ser. 22: 180. 1940. Type:
1980. Type: PANAMA: Chiriqui, Cerro COLOMBIA: Choco, Bahia Solano, along
Colorado, along road above San Felix, 29 km Quebrada Jellita, 50-100 m, 22 Feb 1939,
above bridge over Rio San Felix (7.9 km E. P. Killip & H. Garcia 33524 (Holotype F
above turnoff to Escopeta), 1500 m, 14 Jul 999258 (photo PTBG); Isotype US (photo
1976, T. B. Croat 37071 (Holotype MO PTBG)).
2798805 (photo PTBG); Isotype MO). Coussarea hondensis (Standi.) C. M. Taylor &
—Psychotria nebulosa (Dwyer) C. M. Taylor W. C. Burger, Selbyana 12: 138. 1991.
Coussarea colonis Standi., Publ. Field Mus. Nat. Psychotria hondensis Standi., J. Wash. Acad.
Hist., Bot. Ser. 22: 179. 1940. Type: Sci. 18: 183. 1928. Type: COSTA RICA: Bois
PANAMA: Bocas del Toro, Buena Vista, de Rio Hondo, versant atlantique, 100 m, Aug
region of Almirante, Jan-Mar 1928, G. P. 1901,//. Pittier 16161 (Holotype US 764151
Cooper 228 (Holotype F 579282 (photo (photos MEXU, PTBG)).
PTBG)). Norn, superfl. for Coussarea Coussarea imitans L. O. Williams, Phytologia
euryphylla Standi., 1929 —Raritebe 26: 488, fig. 1973. Type: GUATEMALA:
palicoureoides Wernham subsp. Izabal, along trail beginning from mile 33.23
dwyerianum J. H. Kirkbr. between Dartmouth and Moralaes toward
Coussarea curvigemma Dwyer, Phytologia 38- Lake Izabal, Montana de Mico, 30-150 m, 7
215. 1978. Type: PANAMA: Barro Colorado Apr 1940, J. A. Steyermark 39013 (Holotype
Island, Wheeler Trail 850, 1 Jun 1971, T. B. F 1042963 (photos MEXU, PTBG)).
Croat 14863 (Holotype MO 2050013 (photo Coussarea impetiolaris Donn. Sm., Bot. Gaz.
PTBG)). (Crawfordsville) 37: 418. 1904. COSTA
Coussarea dariensis Steyerm., Ceiba 3: 20. RICA: La Colombiana, Llanuras de Santa
1952. Type: PANAMA: Darien, Rio Chico, Clara, 200 m, Jun 1899, H. Pittier 7582
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Crusea Schltdl. & Cham., Linnaea 5: 165. Jan Crusea calocephala DC, Prodr. 4: 567. 1830.
1830. Not Crusea A. Rich., Dec 1830. Type: MEXICO: Without precise locality,
Lectotype: Crusea hispida (Mill.) B. L. Rob. "Pavon " [Sesse, Mocino et al], (Presumably
(Lectotype designated by Schumann, 1891: Herb. Dunat, location unknown (n.v.),
141). fragment P, n.v.; Isotype P, n.v.; see
Crusea allococca Ton. & A. Gray, Proc. Amer. Anderson, 1972: 97).
Acad. Arts 19: 78. 1883, nom. superfl. illeg., Crusea chiapensis Brandegee, Univ. Calif. Publ.
based on Diodia tricocca A. Gray (See Bot. 6: 374. 1917. Type: MEXICO: Chiapas,
Anderson, 1972: 116 for discussion.). C. A. Purpus 7526 (Holotype UC, n.v.).
—Richardia tricocca (Torr. & A. Gray) —Crusea longiflora (Willd. ex Roem. &
Standi, subsp. tetracocca (M. Martens & Schultes) W. R. Anderson
Galeotti) W. H. Lewis & R. L. Oliv. Crusea coccinea DC, Prodr. 4: 567. 1830.
Crusea andersoniorum Lorence, Novon 7: 57, Type: MEXICO: Nova Hispana, without
fig. 6. 1997. Type: MEXICO: Oaxaca, Distr. precise locality, "Pavon 588" [=Sesse, Mocino
de Miahuatlan, crest of ridge and steep SW et al. 588] (Lectotype MA (n.v.), fragment G-
facing slope 90-100 km SE of Sola de Vega DC, n.v.; Isolectotype F 850991 (photo
along road to Puerto Escondido, 1830 m, 25 PTBG); designated by Anderson, 1972: 39).
Oct 1984, D. E. Breedlove 62063 (Holotype Crusea coccinea DC. var. breviloba Loes.,
CAS 727669 (photo PTBG)). Verh. Bot. Vereins Prov. Brandenburg 65:
Crusea aspera Bartl., Ind. Sem. Hort. Gotting 115. 1923. Type: GUATEMALA:
1838. Type: Probably MEXICO: Herb. Huehuetenagno, Jacaltenango, San Martin, 9
Fischer, a garden specimen grown during Sep 1896, E. Seler & C Seler 3113 (Holotype
Bartling's tenure (Lectotype GOET, n.v.; B, presumably destroyed; Isotype GH, n.v.)
designated by Anderson, 1972: 111). Crusea coccinea DC. var. chiriquensis W. R.
—Crusea hispida (Mill.) B. L. Rob. var. Anderson, Mem. New York Bot. Gard. 22: 45.
hispida 1972. Type: PANAMA: Chiriqui, Casita Alta
Crusea brachyphylla Schltdl. & Cham., Linnaea to Cerro Copete, 2300-3300 m, 10 Jul 1040,
5: 165. 1830. Type: MEXICO: Veracruz: Woodson & Schery 358 (Holotype F 1089337
Serro [Cerro] Colorado, Aug 1828, Schiede & (photo PTBG); Isotypes GH, n.v., MO).
Deppe 524b (Holotype presumably HAL, n.v.; Crusea coccinea DC. var. pubescens Greenm.,
Isotype LE, n.v., fragment F) —Crusea Proc. Amer. Acad. Arts 32: 306. 1897. Type:
longiflora (Willd. ex Roem. & Schultes) MEXICO: Oaxaca: NW slope of Mt.
W. R. Anderson Zempoaltepec, 8000-10000 ft., 10 Jul 1894,
Crusea brachyphylla Schltdl. & Cham. f. E. W. Nelson 699 (Lectotype GH, n.v.;
microcephala Loes., Verh. Bot. Vereins Prov. Isolectotype US (photo PTBG); designated by
Brandenburg 65: 115. 1923. Type: MEXICO: Anderson, 1972: 39).—Crusea coccinea DC.
Michoacan, Patzcuaro, Monte Calvario, 1 Nov
1895, E. Seler & C Seler 1211 (Holotype B, Crusea coronata B. L. Rob. & Greenm., Proc.
presumably destroyed; Isotype GH, n.v.). Amer. Acad. Arts 29: 387. 1894. Type:
—Crusea longiflora (Willd. ex Roem. & MEXICO: Jalisco, Tequila, 3 Oct 1893, C G.
Schultes) W. R. Anderson Pringle 5424 (Holotype GH, n.v.; Isotype US
Crusea calicola Greenm., Proc. Amer. Acad. (photos MEXU, PTBG)).
Arts 32: 305. 1897. Type: MEXICO: Oaxaca, Crusea cruciata S. Watson, Proc. Amer. Acad.
dry calcareous hills, Las Sedas, 6000 ft., 8 Sep Arts 25: 152. 1890. Type: MEXICO: Jalisco,
1894, C. G. Pringle 4869 (Holotype GH, n.v.; barranca near Guadalajara, 19 Oct 1889, C. G.
Isotypes BM, n.v., BR, n.v., F, MICH, n.v., Pringle 2969 (Holotype GH, n.v.; Isotype F)).
MO (photos MEXU, PTBG), NY, n.v., UC —Crusea setosa (M. Martens 8c Galeotti)
(photo PTBG), US (photos MEXU, PTBG)). Standi. & Steyerm.
Crusea cruciata S. Watson var. villosior Crusea hispida (Mill.) B. L. Rob. var.
Greenm., Proc. Amer. Acad. Arts 32: 306. grandiflora (Paul G. Wilson) W. R.
1897. Type: MEXICO: Jalisco, Guadalajara, Anderson, Mem. New York Bot. Gard. 22:
Jul-Oct 1886, E. Palmer 901 (Lectotype GH, 115. 1972. Crusea grandiflora Paul G.
n.v.; Isolectotype US; designated by Wilson, Kew. Bull. 13: 161. 1958. Type:
Anderson, 1972: 83).—Crusea setosa (M. MEXICO: Mexico, G. B. Hinton 4848
Martens & Galeotti) Standi. & Steyerm. (Holotype K, n.v.).
Crusea diversifolia (Kunth) W. R. Anderson, Crusea hispidula Bartl., Ind. Sem. Hort.
Mem. New York Bot. Gard. 22: 58. 1972. Gotting. 1839: 3. 1839. Type: Herb. Fischer, a
Spermacoce diversifolia Kunth in Humb., garden specimen grown during Bartling's
Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 341. tenure (Holotype GOET (n.v.)).—Crusea
1818 [1819]. Type: MEXICO: Guanajuato, calocephala DC.
Santa Rosa de la Sierra, Humboldt & Crusea longibracteata Benth. in Oerst,
Bonplands.n. (Holotype P-Bonpland, n.v. Vidensk. Meddel. Dansk. Naturhist. Foren.
(photos F, MO)). Kjoebenhavn 1852: 28. 1853. Type:
Crusea elata Brandegee, Univ. Calif. Publ. Bot. NICARAGUA: Granada, Jun 1846, Oersted
6: 68. 1914. Type: MEXICO: Chiapas, Sierra s.n. (Holotype, K, n.v.; Isotype US)).
de Tonala, Sep 1913, C. A. Purpus 6640 —Crusea setosa (M. Martens & Galeotti)
(Holotype UC 172353 (n.v.); Isotypes BM, Standi. & Steyerm.
n.v., F, GH, n.v., NY, n.v., US (photos Crusea longiflora (Willd. ex Roem. &
MEXU, PTBG)). Listed in synonymy under Schultes) W. R. Anderson, Mem. New York
Crusea calocephala by Anderson (1972), but Bot. Gard. 22: 89. 1972. Spermacoce
it actually represents a valid species (W. R. longiflora Willd. ex Roem. & Schultes, Syst.
Anderson, pers. comm. 1984). Veg. 3: 531. 1818. Type: MEXICO:
Crusea gentryi Standi., Publ. Field Mus. Nat. Michoacan, near Ario [de Rosales], and
Hist, Bot. Ser. 22: 112. 1940. Type: Mexico City, Humboldt & Bonpland s.n.
MEXICO: Sonora: Alamos Distr., Quirocoba, (Holotype B-W, n.v.).
12 Nov 1923, H. S. Gentry 766M (Holotype F Crusea lucida Benth., Bot. Voy. Sulphur 109.
968108 (photo PTBG); Isotype MICH, n.v.). 1845. Type: MEXICO: Without precise
—Crusea lucida Benth. locality, southern Mexico, Sinclair s.n.
Crusea grandiflora Paul G. Wilson, Kew Bull. (Holotype K, n.v. (photo MICH)).
13: 161. 1958. Type: MEXICO: Mexico, Crusea megalocarpa (A. Gray) S. Watson,
Distr. Temascaltepec, Tenayac, 1540 m, 28 Proc. Amer. Acad. Arts 26: 137. 1891.
Sep 1933, G. B. Hinton 4848 (Lectotype K, Spermacoce megalocarpa A. Gray, Proc.
n.v.; designated by Anderson, 1972). Amer. Acad. Arts 21: 381. 1886. Type:
—Crusea hispida (Mill.) B. L. Rob. var. MEXICO: Chihuahua, southwestern
Chihuahua, wet places in the higher
Crusea guatemalensis Gand., Bull. Soc. Bot. mountains above Batopilas, Aug-Nov 1885,
France 65: 34. 1918. Type: GUATEMALA: E. Palmer 185 (Holotype GH, n.v.; Isotypes
Alta Verapaz, H. von Tuerckheim s.n. NY, n.v., US (photos MEXU, PTBG)).
(Holotype presumably P, n.v.).—Crusea Crusea palmeri A. Gray, Proc. Amer. Acad.
calocephala DC. Arts 21: 381. 1886. Type: MEXICO:
Crusea hispida (Mill.) B. L. Rob., Proc. Amer. Chihuahua, southwestern Chihuahua, 25 miles
Acad. Arts 45: 409. 1910. var. hispida. S of Batopilas, Aug-Sep 1885, E. Palmer 70
Crucianella hispida Mill, Gard. Diet., ed. 8, (Holotype GH, n.v.; Isotypes BM, n.v.,
no. 4. 1768. Type: MEXICO: Veracruz, MEXU, NY, n.v., US (photos MEXU,
grown at Chelsea from seed sent by Houstoun, PTBG)).—Crusea psyllioides (Kunth) W. R.
Miller s.n. (Holotype BM, n.v. (photo US)). Anderson
^ NOMENCLATOR OF MEXICAN AND CENTRAL
Crusea parviflora Hook. & Arn., Bot. Beechey n.v.).—Crusea diversifolia (Kunth) W. R.
Voy. 430, pi. 99c. 1840 (text), 1841 (plate). Anderson
Type: MEXICO: Guerrero, Acapulco, Sinclair Crusea subulata (DC.) A. Gray var. leiocarpa
s.n. (Holotype K, n.v.). Greenm., Proc. Amer. Acad. Arts 40: 33.
Crusea psyllioides (Kunth) W. R. Anderson, 1904. Type: MEXICO: San Luis Potosi,
Mem. New York Bot. Gard. 22: 75. 1972. Alvarez, 28 Sep-3 Oct 1902, E. Palmer 147
Spermacoce psyllioides Kunth in Humb., (Holotype GH, n.v.); Isotypes F, MO (photos
Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 346, pi. MEXU, PTBG), NY, n.v., US (photos
278. 1818 [1819]. Type: MEXICO: MEXU, PTBG)).—Crusea diversifolia
Michoacan, near Ario [de Rosales], Humboldt (Kunth) W. R. Anderson
& Bonplands.n. (Holotype P-Bonpland, n.v. Crusea tricocca (Torr. & A. Gray) A. Heller,
(photo MICH)). Contr. Herb. Franklin & Marshall College 1:
Crusea rosea Loes., Repert. Spec. Nov. Regni 97. 1895. Diodia tricocca Torr. & A. Gray, Fl.
Veg. 18: 362. 1922. Type: MEXICO: N. Amer. 2: 30. 1841. Type: U.S.A.: Texas,
Morelos: barranca bei Cuernava Drummond 117 (Holotype F; Isotype G, n.v.).
[Cuernavaca], 14 Oct 1904, E. Seler & C. —Richardia tricocca (Torr. & A. Gray)
Seler 4181 (Lectotype GH, n.v.; designated by Loes. subsp. tricocca
Anderson, 1972: 111).—Crusea hispida Crusea villosa S. Watson, Proc. Amer. Acad.
(Mill.) B. L. Rob. var. hispida Arts 25: 152. 1890. Type: MEXICO: Jalisco,
Crusea rubra (Jacq.) Schltdl. & Cham., Linnaea rocky hillsides near Guadalajara, 9 Oct 1889,
5: 165. 1830. Spermacoce rubra Jacq., PL C. G. Pringle 2448 (Holotype GH, n.v.;
Rar. Hort. Schoenbr. 3: 3,pi. 256. 1798. Isotypes BM, n.v., MEXU, MO (photos
Type: Unknown (see Anderson, 1972: 110). MEXU, PTBG), NY, n.v., US (photos
—Crusea hispida (Mill.) B. L. Rob. var. MEXU, PTBG)).—Crusea psyllioides
(Kunth) W. R. Anderson
Crusea setosa (M. Martens & Galeotti) Standi. Crusea violacea Brongn. ex Neumann, Rev.
& Steyerm., Publ. Field Mus. Nat. Hist., Bot. Hort. II. 4: 368. 1846; 5: 61. 1846. Type:
Ser. 23: 22. 1943. Borreria setosa M. Martens Unknown (see Anderson, 1972: 97).—Crusea
& Galeotti, Bull. Acad. Roy. Sci. Bruxelles calocephala DC.
11:131. 1844. Type: MEXICO: Oaxaca, dans Crusea wrightii A. Gray, PL Wright. 2: 68.
les forets de palmiers et sur les dunes de la 1853. Type: MEXICO: Sonora, prairies near
cote d'Oaxaca, baignee par l'Ocean Pacifique, Santa Cruz, Sep 1851, C. Wright 1121
pres du Rancho du Rio Grande, Sep 1839, H. (Holotype GH, n.v.; Isotypes GH, n.v., US
Galeotti 2627 (Holotype BR, n.v.; Isotype P, (photos MEXU, PTBG)).
Crusea wrightii A. Gray var. angustifolia
Crusea subalata Hook. & Arn., Bot. Beechey W. R. Anderson, Mem. New York Bot. Gard.
Voy. 431. 1840. Type: MEXICO: Nayarit, 22: 108. 1972. Type: MEXICO: Guerrero, 6.3
between San Bias and Tepic, Sinclair s.n. miles east of Chilpancingo on road to Tixtla,
(Holotype K, n.v.).—Crusea parviflora at km 12, 1600 m, W. R. Anderson & C W.
Hook. & Arn. Laskowski 4334 (Holotype MICH, n.v.;
Crusea subulata (DC.) A. Gray, Proc. Amer. Isotype MO (photos MEXU, PTBG)).
Acad. Arts 19: 78. 1883. Borreria subulata
DC, Prodr. 4: 543. 1830. Type: MEXICO: Dasycephala DC, Prodr. 4: 565. 1830. (Diodiae
Without precise locality, Berlandier 826 sect. 2). Type: Not designated.
(Lectotype [annotated by de Candolle but not Dasycephala spicata (Miq.) Benth. in Benth. &
cited in protologue, according to Anderson, Hook, f, Gen. PL 2: 144. 1873. Diodia
1972: 58] G, n.v. (photo F); Isolectotype BM, spicata Miq., Stirp. Surin. Sel. 179, pi 52.
1851. Type: SURINAM: Sylvarum prope
Bergendaal, Oct, Focke s.n. (presumably U, Deppea Cham. & Schltdl., Linnaea 5: 167.
n.v.; this is Miquel's lectotype according to 1830. Type: Deppea erythrorhiza Cham. &
Howard, 1989).—Diodia spicata Miq. Schltdl.
Deppea amaranthina Standi. & Steyerm., Publ.
Declieuxia Kunth in Humb., Bonpl. & Kunth, Field Mus. Nat. Hist., Bot. Ser. 22; 281. 1940.
Nov. Gen. Sp. 352,/?/. 25/. 1818. Type: Type: GUATEMALA: Chiquimula, Cerro
Declieuxia fruticosa (Willd. ex Roem. & Brujo, in vicinity of Rio Negro, below
Schult.) Kuntze. Montana Montenegro, near village of Brujo,
Declieuxia fruticosa (Willd. ex Roem. & 1500-2000 m, 1 Nov 1939, J. A. Steyermark
Schult.) Kuntze, Rev. Gen. 1: 279. 1891. 30926 (Holotype F 1035809 (photos F,
Houstonia fruticosa Willd. ex Roem. & MEXU, PTBG)).
Schult., Syst. Veg. 3: 527. 1818. Type: Deppea anisophylla L. O. Williams, Phytologia
VENEZUELA: Monagas, Caripe, Humboldt 26: 490. 1973. Type: GUATEMALA:
s.n. (presumably B-W, n.v.). Suchitepequez, Finca Moca, 1000 m, 7 Jan
Declieuxia fruticosa (Willd. ex Roem. & 1935, A. F. Skutch 2072 (Holotype F 815818
Schult.) Kuntze var. mexicana (DC.) Standi., (photos MEXU, PTBG); Isotype US).
Field Mus. Nat. Hist, Bot. Ser. 12: 378. 1936. Deppea cornifolia (Benth.) Benth., PI. Hartw.
Declieuxia mexicana DC, Prodr. 4: 479. 349. 1857. Rondeletia cornifolia Benth., PI.
1830. Type: MEXICO: Without precise Hartw. 39. 1840. Type: MEXICO:
locality, Haenke s.n. (Holotype presumably Michoacan, prope Morelia, T. Hartweg 299
PR, n.v.).—Declieuxia fruticosa (Willd. ex (Holotype K (photo MEXU); Isotypes BM, G-
Roem. & Schult.) Kuntze DEL, n.v., fragment F, P (photos MEXU,
Det lieuxia galeottii M. Martens in M. Martens PTBG, US), US, fragment (photos MEXU,
& Galeotti, Bull. Acad. Roy. Soc. Bruxelles PTBG), W, 2 sheets (photos PTBG)).
11: 231. 1844. Type: MEXICO: Oaxaca, Choristes.
Llano Verde. 3000-7000 ft., Apr 1840, H. Deppea costaricensis Pol., Linnaea 41: 566.
Galeotti 2603 (Holotype BR (photo PTBG); 1877. Type: COSTA RICA: Desengano,
Isotypes BR, n.v., F (photos PTBG), US). prope Alajuela, 27 Jun 1875, H. Polakowski
Hedyotis galeottii (M. Martens) Terrell & 148 (Holotype B, presumably destroyed
(photos F, MEXU, US); Isotypes BM, W
Declieuxia mexicana DC, Prodr. 4: 479. 1830. (photo PTBG), fragment F).—Deppea
Type: MEXICO: Without precise locality, grandiflora Schltdl.
He k (Holotype presumably PR, n.v.). Deppea ehrenbergii Standi., Publ. Field
—Declieuxia fruticosa (Willd. ex Roem & Columbian Mus., Bot. Ser. 8: 161. 1930.
Schult.) Kuntze Type: MEXICO: Without precise locality, C.
Ehrenberg 1318 (Holotype B, presumably
Dentella J. R. & G. Forst., Char. Gen. PI. 25, t. destroyed (fragment F (photos MEXU,
13. 1776. Type: Dentella repens (L.) J. R & PTBG)).
G. Forst. Deppea erythrorhiza Cham. & Schltdl.,
Dentella repens (L.) J. R. & G. Forst., Char. Linnaea 5: 168. 1830 (Jan 1830). Type:
Gen. PL 26. 1776. Oldenlandia repens L., MEXICO: Veracruz, in umbrosis prope
Mant. 40. 1767. Type: INDIA: (Lectotype Hacienda de la Laguna, Aug 1829, C. J. W.
LINN 155/2, n.v.; designated by Verdcourt, Schiede & F. Deppe 388 (Holotype HAL
1983). Naturalized in Mexico at least in (photo PTBG); Isotype HAL (photo PTBG)).
Colima and Guerrero; native to Australasia. Deppea excelsa (Kunth) Standi., N. Amer. Fl.
32: 89. 1921. Psychotria (?) excelsa Kunth in
Humb., Bonpl. & Kunth, Nov. Gen. Sp. 3:
355, t. 282. 1819. Type: MEXICO: Veracruz,
A NOMENCLATOR OF MEXICAN AND CENTRAL
Dukea trifoliata Dwyer & Hayden, Ann. (Holotype CR, n.v.; Isotypes MO 3030387
Missouri Bot. Gard. 54: 139. 1967. Type: (photo PTBG), TEX-LL).—Alibertia
PANAMA: Code, El Valle, potato farm utleyorum (Dwyer) C. M. Taylor
above village of El Valle, 12 Jul 1962, J. D.
Dwyer 1933 (Holotype MO 3831231 (photos Edithea Standi., Trop. Woods 34: 1. 1933. Type:
MO, PTBG)). Raritebe.—Raritebe Edithea floribunda Standi. [Herein treated as a
palicoureoides Wernham subsp. synonym of Deppea Cham. & Schltdl.]
dwyerianum J. H. Kirkbr. Edithea floribunda Standi, Trop. Woods 34: 1.
Dukea victoriae Dwyer, Ann. Missouri Bot. 1933. Not Deppea floribunda Hemsl., 1879.
Gard. 53: 364. 1966. Type: PANAMA: Type: MEXICO: Guerrero, Malinantepec,
Darien, Cerro Pirre, cloud forest, Oct 1962, Mar 1930, L. Schultze 363 (Holotype B,
J. A. Duke 6099 (Holotype MO 1840647 presumably destroyed, fragment F (photos
(photo PTBG); Isotype MO 1832075 (photo MEXU, PTBG)).—Deppea schultzei Lorence
PTBG)). Raritebe.—Raritebe palicoureoides
Wernham subsp. dwyerianum J. H. Kirkbr. Eizia Standi., Publ. Field Mus. Nat. Hist., Bot.
Ser. 22: 50. 1940. Type: Eizia mexicana
Duroia L. f, Suppl. PI. 209. 1781. Nomen. cons. Standi.
Type: Duroia eriopila L. f. Eizia mexicana Standi., Publ. Field Mus. Nat.
Duroia costaricensis Standi., Conrr. U.S. Natl. Hist., Bot. Ser. 22: 50. 1940. Type: MEXICO:
Herb. 20: 208. 1919. Type: COSTA RICA: Chiapas, Mt. Tacana, 1000-2000 m, Aug
Marais de Sierpe, Mar 1892, H. Pittier 6803 1938, E. Matuda 2403 [sphalm. as 2402 in
(Holotype US 938658 (photos MEXU, protologue] (Holotype MICH, n.v. (fragment
PTBG)). The name Duroia hirsuta (Poepp. & and photo F); Isotypes A, n.v., TEX-LL
Endl.) K. Schum., an Amazonian species, was (photos MEXU, PTBG), US, 2 sheets (photos
misapplied to this species by Dwyer (1980: MEXU, PTBG)).
155).
Duroia genipifolia Standi. & Steyerm., Publ. Elaeagia Wedd., Monogr. Cinch. 94. 1849.
Field Mus. Nat. Hist., Bot. Ser. 22: 186. 1940. Type: Elaeagia utilis (Goud.) Wedd.
Type: GUATEMALA: Izabal, Rio Dulce, Elaeagia asperula Standi, ex Steyerm., Acta
between Livingston and 6 miles up river, on Biol. Venez. 4: 37. 1964. Type: COLOMBIA:
north side (right side going up river), sea Valle, Rio Calima (region del Choco), La
level, 14 Apr 1940, J. A. Steyermark 39382 Trojita, 5-50 m, 19 Feb-10 Mar 1944, J.
(Holotype F 1035218 (photo PTBG)). Cuatrecasas 16647 (Holotype VEN, n.v.;
—Randia genipifolia (Standi. & Steyerm.) Isotype F (photo PTBG)).
Elaeagia auriculata Hemsl., Diag. PI. Nov.
Duroia panamensis Dwyer in Dwyer & Hayden, Mexic. 32. 1879. Type: COSTA RICA:
Ann. Missouri Bot. Gard. 55: 38. 1968. Type: Without precise locality, alt. 4500 ped.,
PANAMA: Bocas del Toro, Duwebdulup Endres 95 (Holotype K, n.v.)
Peak, N of Rio Teribe across from W. Huron Elaeagia karstenii Standi., Publ. Field
(behind chiefs house), 300-900 ft., 13 Apr Columbian Mus., Bot. Ser. 8: 50. 1930. Type:
1968, J. H. Kirkbride &J.A. Duke 571 VENEZUELA: Near Colonia Tovar,
(Holotype MO 1968346); Isotype MO). 1854-55, A. Fendler 594 (Holotype GH, n.v.,
—Hippotis albiflora H. Karst. fragment F; Isotypes MO, NY, n.v.).
Duroia utleyorum Dwyer in W. Burger & C. M. Elaeagia myriantha (Standi.) C. M. Taylor &
Taylor, Fieldiana, Bot., n.s. 33: 127. 1993. Hammel, No von 3: 490. 1993. Sickingia
Type: COSTA RICA: Puntarenas, Rincon de myriantha Standi., Publ. Field Columbian
Osa, stream and slopes adjacent to Airfield, Mus., Bot. Ser. 7: 27. 1930. Type:
20-300 m, 6-7 Feb 1974, R. L. Liesner 1774 COLOMBIA: Magdalena, Las Nubes road,
region of Santa Marta, 1200 m, 3 Dec 1898, Ernodea Sw., Prodr. 29. 1788. Type: Ernodea
H. H. Smith 1810 (Holotype F 138660; littoralis Sw.
Isotypes MO, NY, n.v., US, n.v., W, n.v., Y, Ernodea littoralis Sw., Prodr. Veg. Ind. Occ.
n.v.). Holtonia, Simira. 29, pi. 223, tab. 4. 1788. Type: JAMAICA:
Elaeagia nitidifolia Dwyer, Ann. Missouri Bot. Not designated; Swartz referred to the Sloan
Gard. 67: 157. 1980. Type: PANAMA: and Browne descriptions. According to
Panama, region of Cerro Jefe, ca. 1000 m, 3 Howard (1989) a lectotype needs to be
Oct 1969, M. D. Correa A., R. L. Dressier, N. designated.
Escobar & W. H. Lewis 1608 (Holotype MO
2424954 (photo PTBG); Isotype MO 2424954 Evea Aubl., Hist. PI. Guian. 1: 100, t. 39. 1775.
(photo PTBG)). nom. rej. vs. Cephaelis Sw. 1788, nom. cons.
Elaeagia uxpanapensis Lorence, Bol. Soc. Bot. Type: Evea guianensis Aubl. [Herein treated
Mexico 45: 66, fig. 1. 1983 (1984). Type: as a synonym of Psychotria L.]
MEXICO: Oaxaca, Mun. de Matias Romero Evea axillaris (Sw.) Standi., Contr. U.S. Natl.
(Zona de Uxpanapa), a 2 km SE del Herb. 18: 123. 1916. Cephaelis axillaris Sw.,
aserradero La Floresta sobre el camino a Rio Prodr. 3: 45. 1788. Not Psychotria axillaris
Verde, a 15 km S de Esmeralda, 250 m, 25 Willd., 1798. Type: WEST INDIES: St.
May 1981, T. Wendt, S. Koch, A. Villalobos & Christopher, Swartz s.n. (Holotype
J. Garcia 3299 (Holotype MEXU; Isotypes presumably S, n.v.). Tapogomea.
CHAPA (33 others were also distributed). —Psychotria aubletiana Steyerm.
Evea chiapensis Standi., Contr. U.S. Natl. Herb.
Ereicoctis Kuntze, Rev. Gen. PI. 1: 281. 1891. 23: 1392. 1926. Not Psychotria chiapensis
Type: Not designated. Standi, 1926. Type: MEXICO: Chiapas,
Ereicoctis acerosa (A. Gray) Kuntze, Rev. Gen Cerro del Boqueron, C. A. Purpus 6928
PI. 1: 281. 1891. Hedyotis acerosa A Gray (Holotype US 567214 (photos MEXU,
PI. Wright. 1: 81. 1852. U.S.A.: Texas, high PTBG); Isotypes F, MO (photos MEXU,
prairies, from Live Oak Creek to the Las PTBG)). Cephaelis, Psychotr ~ ' '
Moras, Jun 1849, C. Wright 237 (Holotype
GH, n.v.; Isotype US).—Hedyotis acerosa A Evea dichroa Standi., Contr. U.S. Natl. Herb.
Gray 18: 124. 1916. Type: PANAMA: Chinqui,
Ereicoctis cervantesii (Kunth) Kuntze, Rev. Cuesta de las Palmas, S slope of Cerro de la
Gen. PI. 281. 1891. Hedyotis cervantesii Horqueta, 1700-2100 m, 17-19 Mar 1911, H.
Kunth in Humb., Bonpl. & Kunth, Nov. Gen Pittier 3218 (Holotype US 677615 (photos
Sp. 3: 390. 1820. Type: MEXICO: A. MEXU, PBTG); Isotype F). Cephaelis.
Bonpland s.n. (presumably P-Bonpland, n.v.; —Psychotria dichroa (Standi.) C. M. Taylor
Isotype B, presumably destroyed (photo Evea elata (Sw.) Standi., Contr. U.S. Natl. Herb.
MO)).—Hedyotis cervantesii Kunth 18: 123. 1916. Cephaelis elata Sw., Prodr.
Veg. Ind. Occ. 45. 1788. Type: JAMAICA:
Enthahs P. Browne, Civ. Nat. Hist. Jamaica Swartz s.n. (presumably S, n.v.).—Psychotria
165, t. 17, fig. 3. 1756. Lectotype: Enthalis elata (Sw.) Hammel
fruticosa L. Evea guapilensis Standi., J. Wash. Acad. Sci.
Erithalis fruticosa L., Syst. Nat., ed 10 2- 930 15: 104. 1925. Type: COSTA RICA: Limon,
1759. Type: JAMAICA: Not designated vicinity of Guapiles, 300-500 m, 12-13 Mar
based on Browne, Civ. Nat. Hist. Jamaica t. 1934, P. C. Standley 37025 (Holotype US
E7,fig. 3. According to Howard (1989), a 1153029 (photos MEXU, PTBG)). Cephaelis.
Browne figure or specimen would be a' —Psychotria guapilensis (Standi.) Hammel
probable lectotype. Evea ipecacuanha (Brot.) Standi., Contr. U.S.
Natl. Herb. 18: 123. 1916. Callicocca
^ NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
ipecacuanha Brot., Trans. Linn. Soc. London designated by McDowell (1996b: 289)).
6: 137. 1802. Type: BRAZIL: Without precise Howard (1989) stated that no type specimens
locality, Gomez s.n. (presumably LISU, n.v.). are known and a neotype needs to be
Cephaelis.—Psychotria ipecacuanha (Brot.) designated. Later, T. McDowell, in his review
Stokes of the genus, selected a lectotype from among
Evea nana Standi., J. Wash. Acad. Sci. 15: 105. original material.
1925. Cephaelis nana (Standi.) Standi. Type: Exostema coulteri Hook. f. ex Hemsl., Diag. PI.
PANAMA: Canal Zone, hills N of Frijoles, 19 Nov. 32. 1879, [as Exostemma]. Type:
Dec 1923, P. C. Standley 27550 (Holotype US MEXICO: Hidalgo, Zimapan, Coulter 209
1153871 (photos MEXU, PTBG); Isotype F). (Holotype K, n.v.; Isotype MO).
Psychotria guapilensis (Standi.) —Syringantha coulteri (Hook. f. ex Hemsl.)
Hammel T. McDowell
Evea tomentosa (Aubl.) Standi, Contr. U.S. bandl., N. Amer. Fl. 32:
Natl. Herb. 18: 123. 1916. Tapogomea 126. 1921. Type: MEXICO: Oaxaca, Villa
tomentosa Aubl., Hist. PI. Guiane 160. 1775. Alta, 3000 ft., Aug, H. Galeotti 2664
Type: FRENCH GUIANA: Aublets.n. (Holotype US 572997; Isotypes P, 2 sheets
(Holotype presumably P or BM, n.v.; Isotype (photos MEXU, PTBG)).—Exostema
B, presumably destroyed (photo MO)). mexicanum A. Gray
Cephaelis.—Psychotria poeppigiana Muell. Exostema longicuspe Oerst., Vidensk. Meddel.
Arg. Dansk Naturhist. Foren. Kjobenhavn 1852:
48. 1853. Type: COSTA RICA: In insula San
Exandra Standi., J. Wash. Acad. Sci. 13: 292. Lucas, Aug 1849, Oersted 11403 (Holotype
1923. Type: Exandra rhodoclada Standi. C, n.v. (photo US)).—Exostema caribaeum
[Herein treated as a synonym of Simira Aubl.] (Jacq.) Roem. & Schult.
Exandra rhodoclada Standi., J. Wash. Acad. Exostema mexicanum A. Gray, Proc. Amer.
Sci. 13: 292. 1923. Type: MEXICO: Oaxaca, Acad. Arts 5: 180. 1861. Type: MEXICO:
between La Venta and Niltepec, 200 ft., 14 Jul [Presumably Veracruz] Prov. Huasteca,
1895, E. W. Nelson 2796 (Holotype US Wartenberg, near Tantoyuca, 1858, L. C
229232 (photos MEXU, PTBG); Isotype F Ervendberg 125 (Holotype GH, n.v.; Isotype
(photos MEXU, PTBG)).—Simira P (photos MEXU, PTBG)).
rhodoclada (Standi.) Steyerm. Exostema occidentale Benth., Bot. Voy. Sulphur
104. 1844. Type: COLOMBIA: Narino, Isla
Exostema (Pers.) Bonpl. in Humb. & Bonpl., PL Gorgona, Barclay 354 or Hinds 354
Aequin. 1: 131. 1807 ("1808"). Lectotype: (Holotype K, n.v.).—Rustia occidentalis
Exostema caribaeum (Jacq.) Roem. & Schult. (Benth.) Hemsl.
(designated by Rogers, 1987: 165).
Exostema canescens Bartl. ex DC, Prodr. 4: Faramea Aubl., Hist. PI. Guian. 1: 192, t. 40.
359. 1830. Type: MEXICO: Without precise 1775. Type: Faramea corymbosa Aubl.
locality, Haenke s.n. (Holotype presumably Faramea altipetens Dwyer, Ann. Missouri Bot.
PR, n.v., fragment G-DC, n.v. (microfiche 4: Gard. 67: 162. 1980. Type: PANAMA:
359.3)).—Status uncertain ("scarcely of this Darien, Cerro Tacarcuna, south face of
genus" according to Standley, 1926). westernmost peak, lower montane wet forest,
Exostema caribaeum (Jacq.) Roem. & Schult., 1500 m, 30 Jan 1975, A. Gentry & S. Mori
Syst. Veg. 5: 19. 1819. Cinchona caribaea 14013 (Holotype MO 2272402 (photo
Jacq., Enum. PI. Carib. 16. 1760. Type: PTBG)).
DOMINICA: Without precise locality, De Faramea belizensis Standi., Carnegie Inst.
Ponthieu s.n. (lectotype BM, n.v.; illustrated Wash. Publ. 461: 90. 1935. Type: BRITISH
in Lambert, Desc. Cinchona 24, t. 4. 1797; HONDUDRAS (BELIZE): Camp 36, British
Honduras-Guatemala survey, 2800 ft., 1 Jul Faramea cobana Donn. Sm., Bot. Gaz.
1934, W. A. Schipp S-721 (Holotype F 733535 (Crawfordsville) 57: 422. 1914. Type:
(photo PTBG); Isotypes A, n.v., F, G, n.v., GUATEMALA: Alta Verapaz, Coban, 1550
MO (photos MEXU, PTBG)).—Faramea m, Jul 1912, H. von Tuerckheim 2474
occidentalis (L.) A. Rich. (Holotype US 943628 (photos MEXU,
Faramea bocataurensis Dwyer, Ann. Missouri PTBG)).
Bot. Gard. 67: 163. 1980. Type: PANAMA: Faramea correae C. M. Taylor, Novon 6: 289,
Bocas del Toro, eastern slope of La Zorra to fig 1C, D. 1996. Type: PANAMA: Panama,
Divide, mossy cloud forest, 19 Apr 1968, Parque Nacional Altos de Campana, sendero
J. H. Kirkbride & J. A. Duke 806 (Holotype de Interpretacion, 1 km al E del campamento
MO 1968230 (photos MEXU, PTBG); Isotype de los guardaparques de INRENARE, 8°40'N,
MO 2165813 (photos MEXU, PTBG)). 79°55'W, 800-900 m, 9 Sep 1993, M. D.
Faramea brachysiphon Standi., Publ. Field Correa A., E. Montenegro & E. Hidalgo 9983
Columbian Mus., Bot. Ser. 8: 62. 1930. Type: (Holotype PMA 38229, n.v.; Isotypes MO
BRITISH HONDURAS (BELIZE): 4641338,4658116).
Middlesex, 200 ft., 17 Sep 1928, W. A. Schipp Faramea eurycarpa Donn. Sm., Bot. Gaz.
345 (Holotype F 603047; Isotypes A, n.v., (Crawfordsville) 44: 113. 1907. Type:
GH, n.v., MO (photos MEXU, PTBG), US COSTA RICA: Forets de Las Vueltas,
(photos MEXU, PTBG)). Tucumque, Dec 1898, A. Tonduz 12880
Faramea bullata Standi., Publ. Field Columbian (Holotype US 943627 (photos MEXU,
Mus., Bot. Ser. 4: 294. 1929. Type: PTBG); Isotypes B, presumably destroyed
PANAMA: Bocas del Toro, region of (fragment F), GH, 2 sheets (photos PTBG), P,
Almirante, Kankintoe 10 miles above 2 sheets (photos MEXU, PTBG)).
Holstein, Jan-Mar 1928, G. P. Cooper 507 Faramea hondurae Standi., J. Wash. Acad. Sci.
(Holotype F 579385 (photo PTBG); Isotype 18: 169. 1928. Type: COSTA RICA: San
NY, n.v.).—Faramea trinervia K. Schum. Jose, La Hondura, 1400 m, 16 Mar 1924,
Faramea capulifolia Dwyer in W. C. Burger & P. C. Standley 37890 (Holotype US 1153105
C. M. Taylor, Fieldiana, Bot, n.s. 33: 132. (photos MEXU, PTBG)).
1993. Type: PANAMA: Darien, Cerro Pirre, Faramea jefensis Dwyer & Hayden, Phytologia
ridgetop at 1200 m, J. P. Folsom 4262 15: 56. 1968. Type: PANAMA: Panama,
(Holotype MO 2620580). Note: the following eastern slope of Cerro Jefe in Clusia forest,
collections were also annotated as "type" by 2700 ft., 8 Feb 1966, E. L. Tyson et al. 3396
J. D. Dwyer but are not types: COSTA RICA: (Holotype MO 1936328 (photo PTBG);
Puntarenas, Monteverde Cloud forest Reserve, Isotypes F, GH, n.v., NY, n.v., US (photo
1500-1600 m, 8 Nov 1986, W. A. Haber ex E. PTBG)).
Bello C. 6235 (MO 36482259, photo PTBG). Faramea killipii Standi., Publ. Field Columbian
PANAMA: Darien, Cerro Pirre, 10-20 Jul Mus., Bot. Ser. 7: 73. 1930. Type:
1977, J. P. Folsom 4435 (MO 2620712 (photo COLOMBIA: Norte de Santander, Killip &
PTBG)). Smith 20569 (Holotype US, n.v.).
Faramea caputanguis Dwyer & Hayden, Ann. Faramea liebmannii Standi., J. Wash. Acad.
Missouri Bot. Gard. 55: 29. 1968. Type: Sci. 17: 343. 1927. Type: MEXICO: Oaxaca,
PANAMA: Panama, Cerro Jefe, 2900 ft., 29 Tepitapa, F. Liebmann 11404 (Liebm.
Jul 1967, J. D. Dwyer & G. Gauger 7375 Rubiaceae Mexic. 105) (Holotype presumably
(Holotype MO 1961869; Isotypes GH, n.v., C, n.v.; Isotype US).
K, n.v., MO (photos MEXU, PTBG), NY, ' Faramea Hesneri Dwyer, Ann. Missouri Bot.
n.v., UC, n.v., US).—Faramea papirifolia Gard. 67: 167. 1980. Type: PANAMA:
(Standi, ex Steyerm.) C. M. Taylor Panama, road to Carti (San Bias), 19 km north
of El Llano, ca. 500 m, 13 Mar 1973, P. Busey
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
913 (Holotype MO 2106781 (photos MEXU, 30-50 m, 18, 22 May 1946, J. Cuatrecasas
PTBG)). 21078 (Holotype US 2825318, n.v.; Isotypes
Faramea loftonii Dwyer & Hayden, Phytologia COL, n.v., VEN, n.v.).
15: 56. 1967. Type: PANAMA: Panama, Faramea papulata Dwyer & Hayden,
Cerro Jefe, in C/usia forest, 2700-3000 ft., 27 Phytologia 15: 57. 1967. Type: PANAMA:
Jan 1966, E. L. Tyson, J. D. Dwyer & K. Blum Panama, Cerro Jefe, in Clusia forest,
3323 (Holotype MO 1824505 (photo PTBG); 2700-3000 ft., 27 Jan 1966, E. L. Tyson, J.
Isotype MO (photo PTBG)).—Coussarea Dwyer & K. Blum 3284 (Holotype MO
loftonii (Dwyer & Hayden) Dwyer 1831279 (photos MEXU, PTBG)).
Faramea luteovirens Standi., Contr. U.S. Natl. Faramea parvibracteata Steyerm., Mem. New
Herb. 18: 138. 1916. Type: PANAMA: Colon, York Bot. Gard. 17: 376. 1967. Type:
Loma de la Gloria, near Fato (Nombre de VENEZUELA: Terr. Fed. Amazonas, Cafio
Dios), 10-104 m, Aug 1911, H. Pittier 4098 Tamatama, Rio Orinoco, 150 m, 12 Sep 1954,
(Holotype US 679194 (photos MEXU, S. Silverio Level 145 (Holotype NY, n.v.).
PTBG)). Faramea pauciflora Dwyer, Ann. Missouri
Faramea multiflora A. Rich., Mem. Soc. Hist. Bot. Gard. 67: 172. 1980. Type: PANAMA:
Nat. Paris 5: 176. 1834. Type: FRENCH Veraguas, Valley of Rio Dos Bocas along
GUIANA: Richard s.n. (Holotype presumably road between Escuela Agricola Alta Piedra
P,n.v.). and Calovebora, 15.6 km northwest of Santa
Faramea myrticifolia Dwyer in W. C. Burger Fe, 450-550 m, 31 Aug 1974, T B. Croat
& C. M. Taylor, Fieldiana: Bot., n.s. 33: 134. 27575 (Holotype MO 2201361 (photo
1993. Type: COSTA RICA: Limon, Las PTBG); Isotype MO (photo PTBG)).
Brisas de Pacuarito, Siquirres, 300 m, 18 Apr Faramea permagnifolia Dwyer ex C. M.
1985, L. D. Gomez, C. Herrera, E. Herrera & Taylor, Novon 6: 298, fig. IB. 1996. Type:
W. Murillo 23401 (Holotype CR, n.v.; Isotype COSTA RICA: Puntarenas, Osa Peninsula, in
MO 3648258 (photo PTBG)). forest ca. 9 km W of Rincon on road to
Faramea occidentalis (L.) A. Rich., Mem. Soc. Rancho Quemado, 8°39'N, 83°32'W, 200 m,
Hist. Nat. Paris 5: 176. 1834. Ixora 26 May 1986, B. Hammel, M. Grayum & G.
occidentalis L., Syst., ed. 10, 2: 893. 1759. de Nevers 15212 (Holotype CR, n.v.; Isotypes
Type: JAMAICA: Based on P. Browne, Hist. MO 4658114, MO 4658115).
Jam. t.6,f.2 (LINN, n.v.), and Plumier, Icon. Faramea quercetorum Standi., J. Wash. Acad.
tab. 156, f. 2 (ed. Burmannus). Steyermark Sci. 18: 168. 1928. Type: COSTA RICA: San
(1974: 947) suggested the type was from Jose, oak forest near Quebradillas, north of
Jamaica, thus selecting the Browne reference Santa Maria de Dota, 1800 m, 24 Dec 1925,
over that of Plumier within the references P. C. Standley 42999 (Holotype US 1253066
given by Linnaeus. (photos MEXU, PTBG)).—Faramea ovalis
Faramea ovalis Standi., Contr. U.S. Natl. Herb. Standi.
18: 138. 1916. Type: PANAMA: Chiriqui, Faramea sanblasensis C. M. Taylor, Novon 6:
forests along Rio Ladrillo and vicinity, above 300, fig. I A. 1996. Type: PANAMA:
El Boquete, 1200-1300 m, W. R. Maxon 5397 Comarca de San Bias, Cerro Brewster,
(Holotype US 675764 (photos MEXU, headwaters of Rio Cangandi, 9°18'N,
PTBG)). 79° 16'W, 630 m, 24 Apr 1985, G. de Nevers,
Faramea papirifolia (Standi, ex Steyerm.) H. Herrera, B. Hammel & S. Charnley 5491
C. M. Taylor, Novon 6: 210. 1996. Cephaelis (Holotype PMA 38230, n.v.; Isotype MO
papirifolia Standi, ex Steyerm., Acta Biol. 4658117).
Venez. 4: 21. 1964. Type: COLOMBIA: Faramea scalaris Standi., Contr. U.S. Natl.
Valle, Rio Calima (region del Choco), margen Herb. 13: 139. 1916. Type: PANAMA:
derecha, lomas frente a quebrada de la Brea, Chiriqui, humid forest between Altos de las
Palmas and top of Cerro de la Horqueta, Faramea talamancarum Standi., Publ. Field
2100-2268 m, 18 Mar 1911, H. Pittier 3265 Columbian Mus., Bot. Ser. 4: 332. 1929.
(Holotype US 677656 (photos MEXU, Type: PANAMA: Talamanca Valley, 3 Feb
PTBG)). 1921, M. A. Carleton 135 (Holotype F 578962
Faramea schultesii Standi., Bot. Mus. Leafl. (photo PTBG); Isotype US (photos MEXU,
Harvard Univ. 9: 194. 1941. Type: MEXICO: PTBG)).—Faramea multiflora A. Rich.
Oaxaca, Distr. Choapam [Choapan], Cerro de Faramea terryae Standi., Publ. Field Mus. Nat.
Lalana, near San Juan Lalana, in forest, long. Hist., Bot. Ser. 22: 189. 1940. Type:
95°45', lat. 17°25\ 1300 m, 11 May 1939, PANAMA: Darien, Chepigana Distr.,
R. E. Schultes & B. P. Reko 855 (Holotype F Cana-Cuasi Trail (Camp 2), 3000 ft., 10 Mar
1016864 (photo PTBG)). 1940, M. E. Terry & R. A. Terry 1479
Faramea sessifolia P. H. Allen, Rain Forests (Holotype F 1034536 (photo PTBG); Isotype
Golfo Dulce 409. 1956. Type: COSTA RICA: MO (photo PTBG)).
Esquinas Forest, area between the Rio Faramea trinervia K. Schum. & Donn. Sm.,
Esquinas and Palmar, 200 ft., 22 May 1950, Bot. Gaz. (Crawfordsville) 31: 115. 1901.
P. H. Allen 5539 (Holotype US 2215891 Type: COSTA RICA: Talamanca, forets de la
(photos MEXU, PTBG); Isotypes F, 2 sheets). Boca de Zhorquin, Mar 1894, A. Tonduz 8571
Faramea standleyana L. O. Williams, (Holotype US 943623 (photos MEXU,
Phytologia 26: 490. 1973. Type: PTBG)).
GUATEMALA: Izabal, Cerro San Gil, rocky Faramea trinervia K. Schum. & Donn. Sm. var.
slopes along Rio Frio, 75 m, 17 Dec 1941, suerrensis Donn. Sm., Bot. Gaz.
/ A. Steyermark 39962 (Holotype F 1134267 (Crawfordsville) 31: 115. 1901. Type:
(photo PTBG)).—Faramea occidentalis (L.) COSTA RICA: Suerre, Llanuras de Santa
A. Rich. Clara, 300 m, Feb 1896, J. D. Smith 6589
Faramea stenophylla Standi., J. Wash. Acad. (Holotype US 943626 (photos MEXU,
Sci. 18: 169. 1928. Type: HONDURAS: PTBG); Isotype US 943624 (photos MEXU,
Cuyamel, on hills, 29 Mar 1924, M. A. PTBG)).—Faramea suerrensis (Donn. Sm.)
Carlcton 592 (Holotype US 1208399 (photos Donn. Sm.
MEXU, PTBG)). Faramea woodsonii Standi., Ann. Missouri
Faramea stenura Standi., Publ. Field Bot. Gard. 28: 470. 1921. Type: PANAMA:
Columbian Mus., Bot. Ser. 4: 331. 1929. Code, North rim of El Valle de Anton, near
Type: HONDURAS: Dept. Atlantida, in wet Cerro Turega, 650-700 m, 30 Jun 1940, R. E.
Woodson, Jr. & R. W. Schery 164 (Holotype F
Lancetilla Valley, 600 m, 13 Mar 1928, P. C. 1085608 (photo PTBG); Isotypes F, GH, n.v.,
Standley 56743 (Holotype F 581582 (photo MO (photo PTBG)).
PTBG); Isotype US 1409383 (photos MEXU, Faramea zetekii Standi., Contr. Arnold Arbor. 5:
PTBG)). 147. 1933. Type: PANAMA: Canal Zone,
Faramea suerrensis (Dorm. Sm.) Donn. Sm., Barro Colorado Island, shore of cove W of
Bot. Gaz. (Crawfordsville) 44: 112. 1907. Fuertes House, 21 Feb 1932, R. H.
Faramea trinenia K. Schum. & Donn. Sm. Woodworth & P. A. Vestal 641 (Holotype A,
var. suerrensis Donn. Sm., Bot. Gaz. n.v.; Isotypes F, MO (photo PTBG)).
(Crawfordsville) 31: 115. 1901. Type: —Faramea occidentalis (L.) A. Rich.
COSTA RICA: Suerre, Llanuras de Santa
Clara, 300 m, Feb 1896, J. D. Smith 6589 Ferdinandusa Pohl, PI. Bras. Icon. Descr. 2: 8.
(Holotype US 943626 (photos MEXU, 1828-1829. (Substitute name for Ferdinandea
PTBG); Isotype US 943624 (photos MEXU Pohl, 1827, not Ferdinanda Lagasca, 1816).
PTBG)). Type: Ferdinandusa speciosa Pohl
L NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
PTBG); Isotypes F, n.v., NY, n.v., US 943469 Bartl. ex DC, Prodr. 4: 436. 1830. Type:
(photos MEXU, PTBG)). Duggena. MEXICO: Without precise locality, Haenke
Gonzalagunia brenesii Standi., Publ. Field s.n. (Holotype presumably PR, n.v., fragment
Mus. Nat. Hist., Bot. Ser. 18: 1302. 1938. G-DC, n.v. (microfiche 4: 436.3). Duggena.
Type: COSTA RICA: Near San Ramon, —Rondeletia buddleioides Benth.
Bosque a orillas del mar de Golfito de Osa, 12 Gonzalagunia ovatifolia (Donn. Sm.) B. L.
Apr 1930, A. Brenes 12323 (Holotype F Rob., Proc. Amer. Acad. Arts 45: 405. 1910.
856730 (photo PTBG); Isotypes GH, n.v., US Gonzalea ovatifolia Dorm. Sm., Bot. Gaz.
(photos MEXU, PTBG)). (Crawfordsville) 27: 336. 1899. Type:
Gonzalagunia chiapasensis (Standi.) Standi. & COSTA RICA: Cartago, Atirro, 600 m, J. D.
Steyerm., Publ. Field Mus. Nat. Hist., Bot. Smith 6599 (Lectotype US 943467, designated
Ser. 22: 283. 1940. Rondeletia chiapasensis here under main entry (photos MEXU,
Standi., Publ. Field Mus. Nat. Hist., Bot. Ser. PTBG); Isolectotype F). Syntypes: Costa
22: 54. 1940. Type: MEXICO: Chiapas, Rica, J. D. Smith 6600 (F, n.v., GH, US);
Cascarada, 31 Dec 1936, E. Matuda 1064 Costa Rica, Pittier & Tonduz 9171 (US, n.v.);
(Holotype MICH, n.v.; Isotypes F (fragment), i, Hart 113 (US,:
TEX-LL (photo PTBG)). Rogiera.
Gonzalagunia exaltata Standi., Publ. Field Mus. Mart., Fl. Bras. 6(6): 292. 1889. Buena
Nat. Hist., Bot. Ser. 22: 385. 1940. Type: panamensis Cav., An. Hist. Nat. 2: 279. 1800.
COSTA RICA: San Jose, basin of El General, Type: PANAMA: Lancon, Nee s.n. (Holotype
beside river, 675-900 m, Mar 1940, A. F. presumably MA, n.v.; possible Isotype F
Skutch 4747 (Holotype F 1044350 (photo (photo PTBG)).
PTBG); Isotypes A, n.v., MO (photos MEXU, >ra Cham. & Schltdl.,
PTBG), US (photos MEXU, PTBG)). Linnaea 6: 414. 1831. Type: MEXICO:
—Rondeletia brenesii Standi. Veracruz, Cuesta Grande de Jalacingo,
Gonzalagunia glabra S. Watson, Proc. Amer. Schiede & Deppe 1275 (Holotype presumably
Acad. Arts 25: 152. 1890. Type: MEXICO: HAL, n.v.; Isotype B, presumably destroyed
Jalisco, in the mountains near Lake Chapala, (photo MO)).—Rondeletia septicidalis B. L.
Dec 1889, C. G. Pringle 2442 (Holotype A, Rob., vel sp. aff.
n.v.; Isotype US (photo PTBG)). Gonzalagunia rojasii Standi., Publ. Field Mus.
—Citharexylum glabrum (S. Watson) Nat. Hist., Bot. Ser. 22: 114. 1940. Type:
Greenm. (Verbenaceae) GUATEMALA: Quezaltenango, El Muro,
Gonzalagunia kallunkiae Dwyer, Ann. below Santa Maria de Jesus, ca. 1200 m, 1
Missouri Bot. Gard. 67: 192. 1980 [as Mar 1938, P. C. Standley 67145 (Holotype F
kallunkii]. Type: PANAMA: Veraguas, NW 995085 (photo PTBG)).
of Santa Fe, 11 km Escuela Agricola Alto de Gonzalagunia rosea Standi., Ann. Missouri
Piedra, on road to Calovebora, near Rio Bot. Gard. 25: 836. 1938. Type: PANAMA:
Coloveborita, 15 Nov 1975, S. Mori & J. Chinqui, valley of the upper Rio Chiriqui
Kallunki 3141 (Holotype MO 2285895 (photo Viejo, 1300-1900 m, Jul-Aug 1937, G. White
PTBG)). & P. White 7 (Holotype F 913064 (photo
Gonzalagunia longithyrsa Fosberg, Sida 2: 387. PTBG); Isotypes A, n.v., MO (photo PTBG).
1966. Type: COSTA RICA: Cartago, Tapanti, NY, n.v.).
at edge of wet forest, 1200 m, 1 Nov 1964, Gonzalagunia rudis (Standi.) Standi., J. Wash.
W. H. Hatheway & R. L. Rodriguez 1241 Acad. Sci. 17: 170. 1927, comb, inval.
(Holotype US 2562636 (photos MEXU, Duggena rudis Standi., Contr. U.S. Natl.
PTBG)).—Gonzalagunia rosea Standi. Herb. 18: 125. 1916. Type: PANAMA:
Gonzalagunia nivea (Bartl. ex DC.) Kuntze, Panama, shady places along Chagras at El
Rev. Gen. PI. 1: 284. 1891. Gonzalea nivea
Vigia, 12 Jan 1911, //. Pittier 2378 (Holotype Duggena.—Gonzalagunia bracteosa (Donn.
US 676602 (photos MEXU, PTBG)). Sm.) B. L. Rob.
Gonzalagunia stenostachya (Standi.) W. C. Gonzalea hayesii Wernham, J. Bot. 51:219.
Burger in W. C. Burger & C. M. Taylor, 1913. Type: PANAMA: In woods near
Fieldiana, Bot. 33: 150. 1993. Rondeletia Panama, Hayes s.n. (Holotype BM, n.v.).
stenostachya Standi., Publ. Field Mus. Nat. Duggena.—Gonzalagunia panamensis
Hist., Bot.Ser. 18: 1372. 1938. Type: COSTA (Cav.) K. Schum.
RICA: La Hondura de San Jose, 1300 m, 15 Gonzalea nivea Bartl. ex DC, Prodr. 4: 436.
Aug 1933, M. Valerio 769 (Holotype F 1830. Type: MEXICO: Without precise
681816 (photos MEXU, PTBG)). locality, Haenke s.n. (Holotype presumably
Gonzalagunia tacanensis Lundell, Lloydia 2: PR, n.v., fragment G-DC, n.v. (microfiche 4:
106. 1939. Type: MEXICO: Chiapas, Mt. 436.3)). Duggena.—Rondeletia buddleioides
Tacana, 1000-2000 m, Aug 1938, E. Matuda Benth.
2462 (Holotype TEX-LL (photos MEXU, Gonzalea ovatifolia Donn. Sm., Bot. Gaz.
PTBG)).—Gonzalagunia thyrsoidea (Donn. (Crawfordsville) 27: 336. 1899. Type:
COSTA RICA: Cartago, Atirro, 600 m, J. D.
Gonzalagunia thyrsoidea (Donn. Sin.) B. L. Smith 6599 (Lectotype US 943467, here
Rob., Proc. Amer. Acad. Arts 45: 405. 1910. designated (photos MEXU, PTBG);
Gonzalea thyrsoidea Donn. Sm., Bot. Gaz. Isolectotype F). Syntypes: COSTA RICA:
(Crawfordsville) 13: 188. 1888. Type: J. D. Smith 6600 (F, n.v., GH, US); Pittier &
GUATEMALA: Aha Verapaz, Berge bei Tonduz 9171 (US, n.v.). PANAMA; Hart 113
Pansamala, 3800 ft., May 1887, H. von (US, n.v.). Duggena.—Gonzalagunia
Tuerckheim 1249 (Holotype US 941409 ovatifolia (Donn. Sm.) B. L. Rob.
(photos MEXU, PTBG); Isotype GH, n.v.). Gonzalea panamensis (Cav.) Spreng., Syst. Veg.
Gonzalagunia tonduzii K. Krause, Bot. Jahrb. 1: 417. 1825. Buena panamensis Cav., Anal.
54, Beibl. 119: 42. 1916. Type: COSTA Hist. Nat. (Madrid) 2: 279. 1800. Type:
RICA: Cerro del Roble, ca. 2700-2900 m, PANAMA: Lancon, Nee s.n. (Holotype
Apr 1898, A. Tonduz 11813 (Holotype B, presumably MA, n.v.; possible Isotype F
presumably destroyed (photos F, MEXU, (photo MO)). Duggena.—Gonzalagunia
MO)).—Rondeletia buddleioides Benth. panamensis (Cav.) K. Schum.
Gonzalagunia veraguensis Dwyer, Ann. Gonzalea parviflora G. Don., Gen. Hist. 3: 537.
Missouri Bot. Gard. 67: 197. 1980. Type: 1834. Type: MEXICO: Oaxaca, 3000 ft., H
PANAMA: Veraguas, NW of Santa Fe, 8.8 Galeotti 7245 (Holotype presumably BM,
km from Escuela Agricola Alto de Piedra, on n.v.; Isotype G-DC, n.v. (photo MO)).
road to Calovebora, 1900 ft., 17 Jun 1975, 5. —Rondeletia capitellata Hemsl., vel. sp. aff;
Mori 6673 (Holotype MO 2353238 (photos perhaps the earliest available epithet for this
PTBG); Isotype MO 2353239 (photo PTBG)).
Gonzalea secunda M. Martens & Galeotti, Bull.
Gonzalea Pers., Syn. PI. 1: 132. 1805. Norn, Acad. Roy. Sci. Bruxelles 11: 235. 1844.
superfl. for Gonzalagunia Ruiz & Pav. 1794 Type: MEXICO: Veracruz, bois humides de
(for which it is herein treated as a synonym). Xalapa, a 4000 pieds, Juin, H. Galeotti 7077
Gonzalea bracteosa Donn. Sm., Bot. Gaz. (Holotype presumably BR, n.v.).—Status
(Crawfordsville) 33: 252. 1902. Type: uncertain.
COSTA RICA: Suerre, Llanuras de Santa Gonzalea thyrsoidea Donn. Sm., Bot. Gaz.
Clara, J. D. Smith 6583 (Holotype US 943468 (Crawfordsville) 13: 188. 1888. Type:
(photos MEXU, PTBG); Isotypes F, n.v., NY, GUATEMALA: Alta Verapaz, Berge bei
n.v., US 943469 (photos MEXU, PTBG)). Pansamala, 3800 ft., May 1887, H. von
Tuerckheim 1249 (Holotype US 941409
A NOMENCLATOR OF MEXICAN A - AMERICAN RUBIACEA
(photos MEXU, PTBG); Isotype GH, n.v.). Guettarda crispiflora Vahl. Eclog. Amer. 1:
Duggena.—Gonzalagunia thyrsoidea (Donn. 36, /. 6. 1796. Type: MONTSERRAT: Versus
Sm.) B. L. Rob. summitates in insula Montserrat, Ryan s.n
(Holotype presumably C, n.v.).
Guettarda L., Sp. PI. 2: 991. 1753. Type: Guettarda dealbata M. Martens & Galeotti,
Guettarda speciosa L. Bull. Acad. Roy. Soc. Bruxelles 11: 232.
Guettarda armata Bartl. ex DC, Prodr. 4: 457. 1844. Type: MEXICO: Oaxaca, dans les
1830. Type: MEXICO: Without precise forets epaisses et humides de Llano Verde et
locality, Haenke s.n. (Holotype presumably de Tanetze, dans le Rincon (cordillere
PR, n.v. (fragment G-DC, n.v. (microfiche 4: orientale d'Oaxaca), 4000-6000 ft., H.
457.19)).—Status uncertain, probably . Galeotti 7131 (Holotype presumably BR,
Chomelia spinosa Jacq. n.v.).—Status uncertain, "scarcely of this
Guettarda brenesii Standi., Publ. Field Mus. genus" according to Standley (1934).
Nat. Hist., Bot. Ser. 18: 1303. 1938. Type: Guettarda deamii Standi., Contr. U.S. Natl.
COSTA RICA: Golfo de Nicoya, Isla de Herb. 20: 209. 1919. Type: GUATEMALA:
Caballo, Jun 1932, A. Brenes 15694 Zacapa, Gualan, 620 ft., 15 Jun 1909, C. C.
(Holotype F 857985 (photo PTBG)). Deam 6271 (Holotype US 796136 (photo
Guettarda chiriquiensis Standi., Ann. Missouri PTBG); Isotype GH, n.v.).
Bot. Gard., 25: 838. 1938. Type: PANAMA: Guettarda dichotoma M. Martens & Galeotti,
Chiriqui, Valley of upper Rio Chiriqui Viejo, Bull. Acad. Roy. Sci. Bruxelles 11: 233. 1844.
1300-1900 m, 27 Jul 1937, P. White & G. Type: MEXICO: Veracruz, savanes de
White 22 (Holotype MO 1177311 (photo Consoquitla, pres la colonie de Zacuapan et
PTBG): Isotypes A, n.v., F).—Guettarda Mirador, 1500-2000 pieds, 1840,//. Galeotti
crispiflora Vahl 2601 (Holotype presumably BR, n.v.; Isotype
Guettarda cobanensis Donn. Sm., Bot. Gaz. P (photos MEXU, PTBG)).—Guettarda
(Crawfordsville) 47. 255. 1909. Type: elliptica Sw.
GUATEMALA: Alta Verapaz, near Coban, Guettarda elliptica Sw., Prodr. Veg. Ind. Occ.
1500 m, H. von Tuerckheim 11.2096 (Holotype 59. 1788. Type: JAMAICA: Campestris
US 941415 (photo PTBG); Isotype US (photo siccis, Swartzs.n. (presumably S, n.v.).
PTBG)). Guettarda filipes Standi., Contr. U.S. Natl.
Guettarda colubrinoides Standi., Publ. Field Herb. 20: 210. 1919. Type: MEXICO:
Columbian Mus., Bot. Ser. 8: 358. 1931. Durango, near Huasemote, 15 Aug 1897,7. N.
Type: MEXICO: Tabasco, Paso del Carrizal, 4 Rose 3498 (Holotype US 302475).
May 1890, J. N. Rovirosa 787 (Holotype K, Guettarda foliacea Standi., Contr. U.S. Natl.
n.v., fragment F)).—Status uncertain, Herb. 18: 139. 1916. Type: PANAMA: Canal
probably Guettarda elliptica Sw. Zone, along the Trinidad River, near sea level,
Guettarda combsii Urb., Symb. Antill. 6: 48. 19-21 Jul 1911, //. Pittier 4031 (Holotype US
1909. Type: CUBA: Distr. Cienfuegos, Prov. 679116; Isotype F).
Santa Clara, prope Cieneguito, 2 Jul 1895, R. Guettarda galeotti Standi., Contr. U.S. Natl.
Combs 270 (Syntype B, destroyed; Lectotype Herb. 23: 1384. 1926. Type: MEXICO:
F 358073 (photo PTBG), here designated; Oaxaca, Prov. Pinotepa, H. Galeotti 2576
Isolectotypes F, US). Syntype: Prov. Pinar del (Holotype US 572992, fragment F (photos
Rio inter Candelaria et Artemisa, Sept., MEXU, PTBG)).—Chomelia protracta
Wilson 1783 (B, destroyed). (Bartl. ex DC.) Standi.
Guettarda conferta Benth., Bot. Voy. Sulphur Guettarda gaumeri Standi., Publ. Field
106. 1845. Type: COSTA RICA: Cocos Columbian Mus., Bot. Ser. 8: 58. 1930. Type:
Island, Barclay s.n. (Holotype BM, n.v.). MEXICO: Yucatan, without precise locality,
1917-1921, G. F. Gaumer 24239 (Holotype F
552244 (photos MEXU, PTBG); Isotypes A, Guettarda protracta Bartl. ex DC. var. brevipes
n.v., GH, n.v., MO (photos MEXU, PTBG)). DC, Prodr. 4: 457. 1830. Type: MEXICO:
(As for other taxa described by Standley in Without precise locality, Haenke s.n.
this publication, the first specimen he cited in (Holotype presumably PR, n.v.). Anisomeris,
the protologue ("Gaumer 24239, Herb. Field Pittonioitis — Chomelia protracta (Bartl. ex
Mus. No. 552,244") was intended to be the DC.) Standi.
holotype, rather than the second one (Lundell Guettarda protracta Bartl. ex DC. var. longipes
352). Consequently lectotypification is DC, Prodr. 4: 457. 1830. Type: MEXICO:
Guerrero, ad Acapulco, Haenke s.n. (Holotype
(iuanmlu insulari.s Brandegee, Univ. Calif. presumably PR, n.v.).—Chomelia protracta
Publ. Bot. 10: 416. 1924. Type: MEXICO: (Bartl. ex DC.) Standi.
Baja California, Socorro Island, Mar-Jun Guettarda ramuliflora Beurl, Kongl. Vetensk.
1897, A. W. Anthony 377 & F. E. Barkelmew Acad. Handl. 132. 1854. Type: PANAMA:
179 (Holotype UC 102472, n.v.; Isotypes F, Colon, near Porto Bello, BUlberg 642
GH, n.v., MO (photos MEXU, PTBG)). (Holotype S, n.v.).
—Guettarda elliptica Sw. Guettarda sageretioides A. R. Molina, Ceiba 3:
Guettarda macrosperma Donn. Sm., Bot. Gaz. 96. 1952. Type: HONDURAS: El Paraiso,
(Crawfordsville) 18: 204. 1893. Type: matorrales hiimedos y rocosos del Rio Lizapa,
GUATEMALA: Santa Rosa, 3000 ft., May 1000 m, 19 Sep 1951, A Molina R. 4125
1892, Heyde & Lux 3160 (Holotype US (Holotype US 2216040 (photo PTBG);
48602 (photo PTBG); Isotype US (photo Isotypes F (photo PTBG), GH, n.v.).
PTBG)). Guettarda sanblasensis Dwyer, Ann. Missouri
Guettarda odorata (Jacq.) Lam., Tabl. Encycl. Bot. Gard. 67: 204. 1980. Type: PANAMA:
Meth. 2: 219. 1819. Laugieria odorata Jacq., San Bias, headwaters of Rio Cuadi, Camp
Enum. PI. Carib. 2: 16. 1760. Type: CUBA: Diablo (Drill site 22), 273 ft., 18 Dec 1967,
Ex Carthagena et Havana, Jacquin s.n. J. A. Duke, A. G. Robyns & S. E. Verhoek
(Lectotype P, n.v.; designated by Steyermark, 3646 (Holotype MO 2183824 (photo PTBG)).
1972: 360). i abra Lam. var. seleriana Loes.,
Guettarda petenensis Lundell, Wrightia 4: 124. Fedde Rep. Sp. Nov. 18: 361. 1922. Type:
1969. Type: GUATEMALA: Peten, La MEXICO: Yucatan, Chiken Itza, im Walde, 2
Cumbre, in high forest, about 6 km from Jun 1914, C. Seler & E. Seler 5574 (Holotype
village, on top of hill bordering Rio Pusila, 18 B, presumably destroyed, fragment F;
May 1967, E. Contreras 6947 (Holotype Isotypes GH, n.v., US (photos MEXU,
TEX-LL (photos MEXU, PTBG); Isotype PTBG)).—Guettarda combsii Urb.
US). Guettarda seleriana (Loes.) Standi., Contr. U.S.
Guettarda poasana Standi., J. Wash. Acad. Sci. Natl. Herb. 23: 1384. 1926. Guettarda scabra
18: 182. 1928. Type: COSTA RICA: Alajuela, Lam. var. seleriana Loes., Fedde Rep. Sp.
Viento Fresco, slopes of Volcan de Poas, 1800 Nov. 18: 361. 1922. Type: MEXICO:
m, 13 Feb 1926, P. C. Standley & Torres Yucatan, Chiken Itza, im Walde, 2 Jun 1914,
47807 (Holotype US 1305283 (photo C. Seler & E. Seler 5574 (Holotype B,
PTBG)). presumably destroyed, fragment F; Isotypes
Guettarda protracta Bartl. ex DC, Prodr. 4: GH, n.v., US (photos MEXU, PTBG)).
457. 1830. Type: MEXICO: Without precise —Guettarda combsii Urb.
locality, Haenke s.n. (Holotype presumably Guettarda tetrandra Sesse & Moc., Fl. Mexic,
PR, n.v., fragment G-DC, n.v. (microfiche 4: ed. 2, 218. 1894. Type: MEXICO: Without
457.18)).—Chomelia protracta (Bartl. ex precise locality, Sesse & Mociiio s
DC.) Standi. (presumably MA, n.v.).—Status i
L NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
231. 1844. Type: MEXICO: Oaxaca, Llano Hedyotis latifolia (M. Martens & Galeotti)
Verde, 3000-7000 ft., Apr 1840, H. Galeotti Walp., Rep. Sp. Nov. 6: 55. 1846.
2603 (Holotype BR (photo PTBG); Isotypes Oldenlandia latifolia M. Martens & Galeotti,
BR, n.v., F, US (photo PTBG)). Bull. Acad. Roy. Sci. Bruxelles 11: 235. 1844.
Hedyotis gracilenta (I. M. Johnst.) W. H. Not Houstonia latifolia Willd. ex Roem. &
Lewis, Rhodora 63: 222. 1961. Houstonia Schult., 1818. Type: MEXICO: Veracruz,
gracilenta I. M. Johnst., Proc. Calif. Acad. Mirador, 3000 pieds, May 1840, H. Galeotti
Sci. 4: 1174. 1824. Type: MEXICO: Baja 2556 (Holotype presumably BR, n.v.; Isotype
California, San Diego Island, rocky island F (photos MEXU, PTBG)).—Hedyotis
slope facing sea, 2 May 1921, /. M. Johnston exigula W. H. Lewis
3927 (Holotype UC, n.v.; Isotype US (photos Hedyotis leucophylla (Kunth) Spreng., Syst. 1:
MEXU, PTBG)). 411. 1826 lla Kunth in
Hedyotis gracilis Brandegee, Zoe 5: 238. 1907. Humb., Bonpl. & Kunth, Nov. Gen. Sp. PI. 3:
Not Hedyotis gracilis DC, 1830. Type: 395, t. 290. 1820. Type: MEXICO: Inter Alto
MEXICO: Veracruz, Orizaba, May 1904, de Peregrino et flumen Papagallo, Humboldt
C. A. Purpus 1251 (Holotype UC, n.v.; & Bonplands.n. (Holotype P-Bonpland, n.v.;
Isotypes F, MO (photos MEXU, Isotypes B (photo MO), P (photos MEXU,
PTBG)).—Hedyotis exigula W. H. Lewis PTBG)).—Rondeletia leucophylla Kunth in
Hedyotis herbacea L., Sp. PI. 102. 1753. Type: Humb., Bonpl. & Kunth
CEYLON: (Presumably LINN, n.v.). Hedyotis longipes (S. Watson) W. H. Lewis,
Oldenlandia. Rhodora 63: 222. 1961. Houstonia longipes S.
Hedyotis humifusa A. Gray, Boston J. Nat. Watson, Proc. Amer. Acad. Arts 18: 97. 1883.
Hist. 6: 216. 1850. Type: U.S.A.: Texas, near Syntypes: MEXICO: Without precise locality,
Fredericksburg, 1847, F. Lindheimer 621 States of Coahuila and Nuevo Leon, Feb-Oct
(Holotype GH, n.v.; Isotypes BM, n.v., K, 1880, E. Palmer 394 (US 47845 (photos
n.v., MICH, n.v., MO, PH, n.v., US, n.v.). MEXU, PTBG)); Nuevo Leon, near
Houstonia, Oldenlandia. Monterrey, 1-26 Feb 1880, E. Palmer 395
Hedyotis intricata Fosberg, Lloydia 4: 290. (US, 2 sheets (photos MEXU, PTBG)).
1941. Based on Houstonia fasciculata A. Hedyotis lutea Sesse & Moc., Fl. Mexic. 22.
Gray, Proc. Amer. Acad. Arts 17: 203. 1882. 1893. Type: Habitat in montibus
Not Hedyotis fasciculata Bertol., 1850. Temascaltepec, floret Julio, Sesse & Mocino
Syntypes: U.S.A.: Texas, Presidio, Bigelow s.n. (presumably MA, n.v.).—Status
s.n. (GH, n.v.); MEXICO: Coahuila, near
Parras and Monclova, Palmer 404 (GH, n.v.), Hedvotis mcvaughii Terrell, Novon 6: 128, fig.
Palmer 406 (GH, n.v.). 1. "l996. Type: MEXICO: Jalisco, Mun. de
Hedyotis kingii (Terrell) G. L. Nesom, Syst. Cabo Corrientes, steep mountain sides 3-10
Bot. 13: 434. 1988. Houstonia kingii Terrell, km generally E on the road to Mina del Cuale
Brittonia 32: 491, fig. 2. 1980. Type: from the junction 5 km NW of El Tuito,
MEXICO: Oaxaca, E-facing slopes, 850-1150 m, 16-19 Feb 1975, R. McVaugh
mountains along route 175, 28 km by road 26426 (Holotype MICH, n.v.).
south of Valle Nacional on way to Ixtlan de Hedyotis mexicana Sesse & Moc., PI. Nov.
Juarez, ca. 2000 m, 30 Jul 1959, R. M. King Hispan. ed. 1,15. 1887. Type: MEXICO:
2107 (Holotype MICH, 2 sheets, n.v.). Habitat in Mexici circuitibus, Sesse & Mocino
Hedyotis lancifolia Schumach. in Schumach. & s.n. (presumably MA, n.v.).—Bouvardia
Thonn., Beskr. Guin. PL 72. 1827. Type: ternifolia (Cav.) Schltdl.
GHANA: Valley of Aquapim, Thonning 210 Hedvotis microtheca (Schltdl. & Cham.)
(Lecototype C, n.v.; Isolectotype S, n.v.; Steud., Nomencl. Bot. ed. 2, 1: 728. 1840.
designated by Verdcourt, 1976: 292). Gerontogea microtheca Schltdl. & Cham.,
Linnaea 5: 169. 1830. Type: MEXICO: Retion, Sierra de la Trinidad, Nov 1902, T. S.
Veracruz, in sylvis Papantlae, Schiede & Brandegee s.n. (Holotype UC, n.v.; Isotypes
Deppe 390 (Holotype presumably HAL, n.v.; GH, n.v., US (photos MEXU, PTBG)).
Isotype MO (photos MEXU, PTBG)). Hedyotis pentandra Schumach. & Thonn.,
Oldenlandia. Kongel. Danske Vidensk. Selsk.
Hedyotis mucronata Benth., Bot. Voy. Sulphur Naturvidensk., Math. Afh. 3: 71. 1827. Type:
19. 1844. Type: MEXICO: Baja California, GUINEA: Aquapim, "S" [Haaslund Smith?]
Magdalena Bay, Barclay s.n. (Holotype BM, (location unknown).—Pentodon pentandrus
n.v.). Houstonia. (Schumach. & Thon.) Vatke
Hedyotis mullerae Fosberg, Lloydia 4: 288,^/g. Hedyotis polypremoides (A. Gray) Shinners,
3. 1941. Type: MEXICO: Coahuila, Mun. de Field & Lab. 17: 168. 1949. Houstonia
Cuatro Cienegas, Canon del Agua, Sierra de la polypremoides A. Gray, Proc. Amer. Acad.
Madera, 9 Sep 1939, C. H. Midler & K. Arts 21: 379. 1886. Type: MEXICO:
Midler 3234 (Holotype US 2133305 (photos Chihuahua, Santa Eulalia Mountains, 26 Sep
MEXU, PTBG); Isotype GH, n.v.). 1885, C. G. Pringle 356 (Lectotype GH, n.v.;
Houstonia. Isolectotypes MO (photos MEXU, PTBG),
Hedyotis mutica Wedd., Chlor. And. 2: 43. US (photos MEXU, US)).
1857. Type: COLOMBIA: Cundimarca, Prov. •o/ypremoides A. Gray var. bigelovii
Bogota, Goudot s.n. (Lectotype P, n.v.; Greenman, Proc. Amer. Acad. Arts 32: 291.
Isolectotype F, n.v.; designated by Mena V., 1897. Type: U.S.A.: Florence Mountains, Jun
1986).—Arcytophyllum muticum (Wedd.) 1852, Bigelow 437 (Holotype GH, n.v.).
Standi. —Hedyotis polypremoides (A. Gray)
Hedyotis nigricans (Lam.) Fosberg, Lloydia 4: Shinners
287. 1941. Gentiana nigricans Lam., Encycl. Hedyotis pringlei (B. L. Rob.) W. H. Lewis,
2: 645. 1788. Type: MEXICO: Locality and Rhodora 63: 222. 1961. Oldenlandia pringlei
collector unknown (Holotype P-JU, n.v.). B. L. Rob., Proc. Amer. Acad. Arts 27: 169.
Houstonia. 1892. Type: MEXICO: San Luis Potosi,
Hedyotis nigricans (Lam.) Fosberg var. scabra alkaline plains, Hacienda de Angostura, 27
(S. Watson) Fosberg, Lloydia 4: 288. 1941. Jun 1891, C. G. Pringle 3758 (Holotype GH,
Houstonia angustifolia Michx. var. scabra S. n.v.; Isotypes F, MO (photo PTBG), US, 2
Watson, Proc. Amer. Acad. Arts 18: 97. 1883. sheets (photos MEXU, PTBG)).
Type: MEXICO: Coahuila, Saltillo, Caracol Hedyotis pygmaea Roem. & Schult., Syst. Veg.
Mountains, 1-15 Apr 1880, E. Palmer 410 3: 526. 1818. Type: MEXICO: Without
(Holotype GH, n.v.; Isotype US (photos precise locality, A. Bonpland s.n. (Holotype P-
MEXU, PTBG)).—Hedyotis nigricans Bonpland, n.v.; Isotype B, presumably
(Lam.) Fosberg destroyed (photo MO)).—Hedyotis wrightii
Hedyotis palmeri (A. Gray) W. H. Lewis, (A. Gray) Fosberg
Rhodora 63: 222. 1961. Houstonia palmeri A. Hedyotis rubra (Cav.) A. Gray, Mem. Amer.
Gray, Proc. Amer. Acad. Arts 17: 202. 1882. Acad. Arts, n.s. 4: 61. 1849. Houstonia rubra
Type: MEXICO: Coahuila, in the mountains Cav., Icon. 5: 48, t. 474, fig. I. 1799. Type:
east and south of Saltillo, Feb-Oct 1880, E. MEXICO: Hidalgo, Ixmiquilpan, Nee s.n.
Palmer 397 (Syntype US 47431 (photos (presumably MA, n.v.). Oldenlandia.
MEXU, PTBG)); E. Palmer 398 (Syntype US Hedyotis serpyllacea Schltdl., Linnaea 9: 599.
70740 (photos MEXU, PTBG)). 1834. Type: MEXICO: Mexico, inter La Joya
Hedyotis peninsularis (Brandegee) W. H. et San Salvador, Jun 1829, Schiede & Deppe
Lewis, Rhodora 63: 222. 1961. Houstonia 265 (Holotype presumably HAL, n.v.;
peninsularis Brandegee, Zoe 5: 160. 1903. Isotypes B (photo MO), F (photo PTBG),
Type: MEXICO: Baja California, Cape MO). Houstonia.
INOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Hedyotis sharpii (Terrell) G. L. Nesom, Syst. B. L. Rob., Proc. Amer. Acad. Arts 45: 401.
Bot. 13: 434. 1988. Houstonia sharpii Terrell, 1910. Type: MEXICO: Nuevo Leon, shaded
Brirtonia 32: 490,/?g. 1. 1980. Type: cliffs and mountains, near Monterrey, 25 Apr
MEXICO: Hidalgo, camino enrre Metepec y 1906, C. G Pringle 13877 (Holotype GH,
Tenango de Doria, entre 16 y 31.5 km al NNE n.v.; Isotypes MO (photos MEXU, PTBG),
de Tulancingo, 18 km NNE de Metepec, 29 US (photos MEXU, PTBG)).
Mar 1974, S. D. Koch 7423 (Holotype US Hedyotis vergrandis W. H. Lewis, Rhodora 63:
2754863 (photos MEXU, PTBG); Isotypes 222. 1961. Nom. nov. for Houstonia prostrata
CHAPA, ENCB, MEXU). Brandegee, Zoe 5: 105. 1901. Not Hedyotis
Hedyotis sinaloeae W. H. Lewis, Rhodora 63: prostrata Blume, 1823, nor Korth., 1851.
222. 1961, nom. nov. for Houstonia parvula Type: MEXICO: Baja California, La Palma,
Brandegee, Zoe 5: 221. 1905. Not Hedyotis Cape Region, 25 sep 1899, T. S. Brandegee
parvula (A. Gray) Fosb., 1943. Type: s.n. (Holotype UC, n.v.; Isotypes GH, n.v.,
MEXICO: Sinaloa, vicinity of Culiacan, 12 US, 2 sheets (photos MEXU, PTBG)).
Oct 1904, T. S Brandegee s.n. (Holotype UC, Hedyotis watsonii W. H. Lewis, Rhodora 63:
n.v.; Isotypes GH, n.v., US (photos MEXU, 222. 1961. Nom. nov. for Oldenlandia ovata
PTBG)). S. Watson, Proc. Amer. Acad. 18: 97. 1883.
Hedyotis spellenbergii G. L. Nesom & Not Hedyotis ovata Thunb. ex Maxim., 1833,
Vorobik, Syst. Bot. 13: 432. 1988. Type: nor Thwaites ex Hook, f., 1892. Type:
MEXICO: Chihuahua, Mun. Ocampo, area of MEXICO: Nuevo Leon, Guajuco, Feb-Oct
Cascada de Basaseachic at the confluence of 1880, E. Palmer 399 (Lecototype GH, n.v.;
Rio Basaseachic and Rio Durazno, ca. 2 mi S Isolectotype US 47919 (photos MEXU,
of village of Basaseachic, 2100-2200 m, 18 PTBG)).
May 1985, J. Grimes, T. Ayers, M. Lavan & Hedyotis wrightii (A. Gray) Fosberg, Lloydia
R. Scott 2855 (Holotype TEX, n.v.; Isotypes 4: 290. 1941. Houstonia wrightii A. Gray,
ARIZ, n.v., CAS, n.v., MEXU, n.v., NMC, Proc. Amer. Acad. Arts 17: 202. 1882. Type:
. IS, MEXICO: Nuevo Leon, shaded cliffs of
Hedyotis spinescens Sesse & Moc., Fl. Mexic. mountains, near Monterrey, 25 Apr 1906,
22. 1893. Type: MEXICO: Habitat in aridis C. G. Pringle 13877 (Holotype GH, n.v.;
Tehuacani circuitibus. Floret Maio, Sesse & Isotypes MO (photos MEXU, PTBG), US
Mocino s.n. (presumably MA, n.v.). (photos MEXU, PTBG)).
—Bouvardia erecta (DC.) Standi. Hedyotis xestosperma (B. L. Rob. & Greenman)
Hedyotis teretifolia (Terrell) G. L. Nesom, W. H. Lewis, Rhodora 63: 222. 1961.
Syst. Bot. 13: 434. 1988. Houstonia teretifolia Oldenlandia xestosperma B. L. Rob. &
Terrell, Brirtonia 31: 166. 1979. Type: Greenman, Proc. Amer. Acad. Arts 32: 41.
MEXICO: Coahuila, 12 km NNE of Las 1896. Type: MEXICO: Oaxaca, open glades,
Margaritas on easternmost ridge of Sierra de Sierra de San Felipe, 8000 ft., Jun 1894, C. G.
las Margaritas, 1300-1400 m, 24 Sep 1972, Pringle 4692 (Lectotype GH, n.v.;
M. C. Johnston, F. Chiang & T. Wendt 9508E Isolectotypes BM, n.v., F, K, n.v., MO
(Holotype TEX, n.v.). (photos MEXU, PTBG), MSC, n.v., PH, n.v.,
Hedyotis terrellii Lorence, Novon 7: 51,fig. 3. US, 2 sheets (photos MEXU, PTBG), VT
1997. Type: MEXICO: Oaxaca, Distr. de n.v.); designated by Terrell, 1980). Houstonia.
Mixe, Mun. de Totontepec, 3 km al SW de —Bouvardia xestosperma (B. L. Rob. &
Totontepec, carretera a Mitla, 1910 m, 6 Sep Greenman) Terrell & S. D. Koch
1986, R. Torres C. & C. Martinez 9067
(Holotype PTBG 12314; Isotype MEXU).
Hedyotis umbratilis (B. L. Rob.) W. H. Lewis,
Rhodora 63: 222. 1961. Houstonia u
(Willd.) K. Schum. [Herein treated as a of Mexican origin cultivated at the Berlin
synonym of Diodia L.] Botanical Garden.—Hoffmannia excelsa
, \ Willd. ex Roem. & (Kunth) K. Schum.
Schult.) K. Schum. in Mart., F. Bras. 6(6): 29. Higginsia psychotriifolia Benth. in Oerst.,
1888 (as ocimitolid) Spenmicoce ocxmifoha Vidensk. Meddel. Dansk Naturhist. Foren.
Willd. ex Roem. & Schult., Syst. Veg. 3: 530. Kjobenhavnl852:50. 1853 [as
1819. Type: WEST INDIES: Without precise psychotriaefolia]. Type: COSTA RICA: In
locality, 1768, Rudolphi s.n. (presumably B- Monte Barba, 6000 ft., May 1847, Oersted
W, n.v.). Diodia.—Spermacoce ocymifolia 11507. (Holotype C, n.v. (photos F, MO)
Willd. ex Roem. & Schult. Isotype US (photos MEXU, PTBG)).
-Hoffmannia psychotriifolia (Benth. in
Oerst.) Griseb.
Higginsia refulgens Hook, f, Bot. Mag. tab.
DC. 5346. 1862. Type: Based on material
Icxascpalum angustifolium Bartl. ex DC, cultivated at Kew thought to be from South
Prodr. 4: 561. 1830. Type: MEXICO: America, but most probably from Chiapas,
Hidalgo, Real del Monte, Haenke s.n. Mexico, or Central America (Holotype
(Holotype PR, n.v., fragment G-DC, n.v. presumably K, n.v.).—Hoffmannia refulgens
(microfiche 4: 561.1); Isotype MO (photos (Hook, f.) Hemsl.
MEXU, PTBG)).—Diodia c Higginsia refulgens Hook. f. var. 2 argyroneura
(Linden) Regel, Gartenfl. 17: 116. 1868.
Campylobotrys argyroneura Linden, Belg.
Higginsia Pers., Syn. PI. 1: 133. 1805. Type: Hortic. 8: 166. 1858. Type: Not designated,
Higginsia obovata Pers.. "Higgensia" orth. based on cultivated material collected by
mut. Steud. Norn., ed. 2, 1: 767. 1840. [Herein Ghiesbreght in Chiapas, Mexico.
treated as a synonym of Hoffmannia Sw.] —Hoffmannia refulgens (Hook, f.) Hemsl.
discolor (Lem.) Planch., Fl. Serres. 5: i fulgens Hook. f. var. 6 ortgiesii
482d. 1849. Campylobotrys discolor Lem., Fl. Regel, Gartenfl. 17: 116. 1868. Type: Not
Serres 3: Misc. 37, fig. 1837. Type: Not designated, based on cultivated material of
designated, based on cultivated material Mexican origin.—Hoffmannia regalis (Hook,
collected by Galeotti in Mexico. f.) Hemsl.
—Hoffmannia discolor (Lem.) Hemsl. Higginsia refulgens Hook. f. var. P pyrophylla
Higginsia ghiesbreghtii (Lem.) Hook., Bot. Regel, Gartenfl. 17: 116. 1868. Type: Not
Mag. tab. 5383. 1863. Campylobotrys designated, based on cultivated material from
ghiesbreghtii Lem., Illust. Hort. 8: pi. 279. Volcan San Martin, Veracruz, Mexico.
1861 [as Gheisbechtii]. Type: Not designated, —Hoffmannia refulgens (Hook, f.) Hemsl.
based on cultivated material from seed Higginsia refulgens Hook. f. var. S roezlii
collected by Ghiesbreght in Chiapas, Mexico. Regel, Gartenfl. 17: 116. 1868. Type: Not
—Hoffmannia ghiesbreghtii (Lem.) Hemsl. designated, based on cultivated material of
Higginsia macrophylla Planch., Fl. Serres 5: Mexican origin.—Hoffmannia regalis (Hook,
482d. 1849. Type: Not designated, described f.) Hemsl.
from cultivated material grown from seeds Higginsia refulgens Hook. f. var. J" smaragdina
collected in Guatemala.—Status uncertain, Regel, Gartenfl. 17: 116. 1868. Type: Not
probably Hoffmannia nicotianifolia (M. designated, based on cultivated material of
Martens & Galeotti) L. O. Williams Mexican origin—Hoffmannia regalis (Hook,
Higginsia mexicana Klotzsch in Link, Klotzsch f.) Hemsl.
& Otto, Ic. PL Rar. Hort. Berol. 1: 57, tab. 23. Higginsia regalis Hook, f., Bot. Mag. tab. 5280.
1841. Type: Not designated, based on material 1861. Type: Based on material cultivated at
^ NOMENCLATOR OF MEXICAN AND CENTRAL
1800-2000 m, Apr 1898, A. Tonduz 12230 A. Gentry & S. Mori 14061 (Holotype MO
(Holotype US 392077 (photos MEXU, 2353753; Isotype MO 2423663).
PTBG); Isotypes P, 2 sheets (photos MEXU, Hoffmannia gesnerioides (Oerst.) Kuntze, Rev.
PTBG)). Gen. PI. 285. 1891. Ophryococcus
Hoffmannia discolor (Lem.) Hemsl., Biol. gesnerioides Oerst., Vidensk. Meddel. Dansk
Centr.-Amer., Bot. 2: 36. 1881. Naturhist. Foren. Kjoebenhavn 1852: 53.
Campylobotrys discolor Lem., Fl. Serres 3: 1853. Type: NICARAGUA: Segovia, Monte
Misc. 37. fig. 1847. Type: Not designated, Pantasmo, Jan 1848, Oersted 11823
based on cultivated plants grown from seed (Holotype C, n.v. (photos F, MO); Isotype F).
collected in Mexico by H. Galeotti. Note: A Hoffmannia ghiesbreghtii (Lem.) Hemsl., Biol.
specimen at P ["en herb. Houtte Juin 1857. Centr.-Amer., Bot. 2: 36. 1881.
Hort. Parisei"] and a specimen at G ["Mexici, Campylobotiys ghiesbreghtii Lem., Illust.
Oaxaca, 3000', fl. albis, Galeotti 7183"] may Hort. S:pl. 279. 1861. Type: Not designated,
correspond with Galeotti's seed collection and based on cultivated material from seed
represent authentic cultivated material. collected by Ghiesbreght, probably in
Higginsia. Chiapas, Mexico. Higginsia.
Hoffmannia dotae Standi., J. Wash. Acad. Sci. Hoffmannia hamelioides Standi., J. Wash.
18: 181. 1928. Type: COSTARICA: San Acad. Sci. 15: 8. 1925. Type: COSTA RICA:
Jose, vicinity of Santa Maria de Dota, San Jose, between Aserri and Tarbaca,
1500-1800 m, 26 Dec 1925-3 Jan 1926, P. C. 1600-1800 m, 12 Feb 1924, P. C. Standley
Standley & J. Valerio 43277 (Holotype US 34149 (Holotype US 1152904 (photos
1253172 (photos MEXU, PTBG)). MEXU, PTBG)).
Hoffmannia eliasii Dwyer, Ann. Missouri Bot. Hoffmannia hammelii C. M. Taylor in W. C.
Gard. 56: 276. 1969. Type: PANAMA: Burger & C. M. Taylor, Fieldiana, Bot., n.s.
Darien, Rio Cuasi mainstream, 2.5 miles S of 33: 174. 1993. Type: COSTA RICA:
Tres Bocas, 28 Apr 1968, J. H. Kirkbride, Jr. Puntarenas, near Sirena, Corcovado National
& J. A. Duke 1129 (Holotype MO 1982892 Park, 0-200 m, 5 Jul 1977, R. L. Liesner 2907
(photos MEXU, PTBG)).—Hoffmannia (Holotype CR, n.v.; Isotype MO).
subauriculata Standi. Hoffmannia haydenii Dwyer, Ann. Missouri
Hoffmannia excelsa (Kunth) K. Schum. in Bot. Gard. 56: 277. 1969. Type: PANAMA:
Mart., Fl. Bras. 6(6): 327. 1889. Psychotria Chiriqui, Cerro Campana, Jul 1965, s. M. V.
excelsa Kunth in Humb., Bonpl. & Kunth, Hayden 117 (Holotype MO 2030831). Note:
Nov. Gen. Sp. 3: 355. 1819. Type: MEXICO: In the protologue the type locality is
Veracruz, Xalapa, A. Bonpland 1445 incorrectly cited as "Finca Collins, ca. 5000
(Holotype presumably P-Bonpland, n.v.; ft." —Hoffmannia subauriculata Standi.
Isotypes B, n.v. (photo MO), P (photos Hoffmannia hondurensis Standi., Publ. Field
MEXU, PTBG)). Deppea. Columbian Mus., Bot. Ser. 4: 329. 1929.
Hoffmannia fimbrianthera Dwyer, Ann. Type: HONDURAS: Atlantida, Lancetilla
Missouri Bot. Gard. 56: 277. 1969. Type: Valley, 200-500 m, 6 Dec 1927-20 Mar
PANAMA: Rio Teribe, just below Puerto 1928, P. C. Standley 52905 (Holotype F
Palenque, 350 m, 12 Apr 1968, J. H. 582885 (photo PTBG); Isotype US (photos
Kirkbride, Jr. & J. A. Duke 556 (Holotype MEXU, PTBG)).
MO 1999724).—Hoffmannia inamoena Hoffmannia huehueteca Standi. & Steyerm.,
Standi. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 249.
Hoffmannia gentryi Dwyer, Ann. Missouri 1947. Type: GUATEMALA: Huehuetenango,
Bot. Gard. 67: 230. 1980. Type: PANAMA: Sierra de los Cuchumatanes, Cerro Negro, 2
Darien, Cerro Tacarcuna, summit camp, along miles east of Las Palmas, 1600-2000 m, 31
stream N of camp, 1550-1650 m, 1 Feb 1975,
Aug 1942, J. A. Steyermark 51697 (Holotype 2992 collection at P was chosen as lectotype,
F 1132825 (photo PTBG)). because it is most representative, with flowers,
Hoffmannia inamoena Standi., J. Wash. Acad. fruits, and full-sized leaves.—Hoffmannia
Sci. 18: 179. 1928. Type: COSTA RICA: nicotianifolia (M. Martens & Galeotti) L. O.
Guanacaste, Los Ayotes, near Tilaran, 600 m, Williams
21 Jan 1926, P. C. Standley & J. Valerio Hoffmannia leucocarpa Standi., J. Wash.
45421 (Holotype US 1254102 (photos Acad. Sci. 15: 9. 1925. Type: COSTA RICA:
MEXU, PTBG)). San Jose, Las Nubes, 1300-1600 m, Mar
Hoffmannia ixtlanensis Lorence, Novon 4: 1924, P. C. Standley 38340 (Holotype US
\2\,fig. 2a, b. 1994. Type: MEXICO: 1153121 (photos MEXU, PTBG)).
Oaxaca, Distr. de Ixtlan, Sierra de Juarez; ruta Hoffmannia lewisiana Dwyer, Ann. Missouri
175 Tuxtepec a Oaxaca, ca. 5 km al N de Bot. Gard. 56: 279. 1969. Type: PANAMA:
Vista Hermosa, 1260 m, 14 Apr 1982, D. H. Chiriqui, vicinity of Cerro Punta, 6800 ft., 31
Lorence, R. Torres C. & R. Cedillo T. 4013 May-1 Jun 1967, Ridgway & Solis 2413
(Holotype MEXU; Isotypes MO, PTBG). (Holotype MO 1966156).
Hoffmannia josefina Standi., J. Wash. Acad. Sci. Hoffmannia liesneriana L. O. Williams,
15: 8. 1925. Type: COSTA RICA: San Jose, Fieldiana, Bot. 36: 54. 1973. Type: COSTA
between Aserri and Tarbaca, 1800 m, 12 Feb RICA: Limon, Guapiles, a 2 millas al norte de
1924, P. C. Standley 34143 (Holotype US Diamantes, 150 m, 9 Aug 1963, A. Jimenez
1152900 (photos MEXU, PTBG)). 1054 M. (Holotype F 1690189 (photo
—Hoffmannia arborescens Donn. Sm. PTBG)).
a kirkbridei Dwyer, Ann. Missouri Hoffmannia lineolata Donn. Sm., Bot. Gaz.
Bot. Gard. 56: 278. 1969. Type: PANAMA: (Crawfordsville) 40: 5. 1905. Type:
Bocas del Toro, Quebrada Huron, ca. GUATEMALA: Alta Verapaz, Cubliquitz,
30(M00 ft., 11 Apr 1968, J. H. Kirkbride, Jr. 350 m, May 1902, H. von Tuerckheim 8227
& J. A. Duke 429 (Holotype MO 1968253). (Holotype US no 576514 (photos MEXU,
—Hoffmannia vesiculifera Standi. PTBG); Isotypes US, 2 sheets (photos MEXU,
Hoffmannia lancistigma Dwyer, Ann. Missouri PTBG)).—Hoffmannia regalis (Hook, f.)
Bot. Gard. 56: 279. 1969. Type: PANAMA: Hemsl.
Chiriqui, Boquete, Cerro Horqueta, Hoffmannia longicalycina Dwyer, Ann.
5000-6000 ft., 8 Aug 1967, J. D. Dwyer & Missouri Bot. Gard. 56: 280. 1969. Type:
S. M. V. Hayden 7735 (Holotype MO PANAMA: Darien, Cuasi-Cana Trail between
2065523). Cerro Campamiento and La Escalera to
Hoffmannia laxa Standi., Publ. Field Mus. Nat. paramo, E of Tres Bocas, 30 Apr 1968, J. H.
Hist., Bot. Ser. 22: 116. 1940. Type: COSTA Kirkbride, Jr. & J. A. Duke 1278 (Holotype
RICA: Alajuela, Canton de San Carlos, Sucre, MO 2165098; Isotype MO 1686749).
1000 m, 30 Mar 1939, A. Smith F1851 —Status uncertain, probably Amphidaysa
(Holotype F 980999 (photo PTBG); Isotype sp.
US (photos MEXU, PTBG)). Hoffmannia longipetiolata Pol., Linnaea 41:
Hoffmannia lenticellata Hemsl., Diag. PI. Nov. 567. 1877. Type: COSTA RICA: In sylvis
Mexic. 30. 1879. Type: MEXICO: Veracruz, primaevis Carpinterae, 24 Jun 1875, H.
region d'Orizaba, Rio Blanco, 29 Aout 1866, Polakowsky 134 (B, presumably destroyed
Bourgeau 2992 (Lectotype P, here designated (photo F), lectotype fragment F, here
(photos MEXU, PTBG); Isolectotype P designated (photo PTBG)). Syntype: COSTA
(photos MEXU, PTBG)). Syntypes: RICA: In sylvis primaevis Carpinterae, 24 Jun
MEXICO: Veracruz, Orizaba, Botteri 974 1875, H. Polakowksy 140 (B, presumably
(Syntype P, photos MEXU, PTBG); Jalapa, destroyed). The fragment of Polakowsky 134
Linden 1 (Syntype K, n.v.). The Bourgeau ; lectotype.
k NOMENCLATOR OF MEXICAN /
Hoffmannia refulgens (Hook, f.) Hemsl., Biol. lectotype as it is most complete, with intact
Centr.-Amer., Bot. 2: 37. 1881. Higginsia leaves, flowers, and inflorescences.
refulgens Hook, f, Bot. Mag. tab. 5346. 1862. Hoffmannia rotundata Standi., Contr. U.S. Natl.
Type: Based on plants grown at Kew from Herb. 20: 204. 1919. Type: MEXICO:
seed thought to be from South America, but Chiapas, Cerro del Boqueron, Jun 1914, C. A.
most probably from Central America or Purpus 7268 in part (Holotype MO 765057).
Chiapas, Mexico (Holotype presumably K, —Hoffmannia nicotianifolia (M. Martens &
n.v.). Galeotti) L. O. Williams
Hoffmannia regalis (Hook, f.) Hemsl., Biol. Hoffmannia rubripigmenta Dywer, Ann.
Centr.-Amer., Bot. 2: 37. 1881. Higginsia Missouri Bot. Gard. 56: 283. 1969. Type:
regalis Hook, f, . Mag. tab. PANAMA: Darien, between upper Rio
Type: Based on ] ts grown at Kew Membrillo and Camp 7 on construction road
presumably fron ;xican seed (Holotype to San Bias, 100-800 m, 26-28 Apr 1967,
presumably K, n.v.). Campylobotrys. J. A. Duke 10870 (Holotype MO 1981851).
Hoffmannia rexmontis Dywer, Ann. Missouri Hoffmannia sessilifolia L. O. Williams,
Bot. Gard. 67: 241. 1980. Type: PANAMA: Phytologia 25: 463. 1973. Type:
Chiriqui, NW side of Cerro Pando, 21 Jul GUATEMALA: Baja Verapaz, Sierra de las
1971, T. Croat 15934 (Holotype MO Minas, above Purulha, 1600 m, 2 Jan 1973,
2692337; Isotypes MO 2692338, MO L. O. Williams, A. Molina R. & T. P. Williams
2150879).—Hoffmannia pallidiflora Standi. 41977 (Holotype F 1716220 (photo PTBG);
Hoffmannia rhizantha Standi., Carnegie Inst. Isotype 1716221).
Wash. Publ. 461: 90. 1935. Type: BRITISH Hoffmannia steinvorthii Fosberg, Sida 2: 388.
HONDURAS (BELIZE): Esperanza Trail, 1966. Type: COSTA RICA: Heredia, south
630 m, 26 Jun 1934, W. A. Schipp S-731 slope of Volcan Barba, 2000 m, 26 Jun 1965,
(Holotype F 729968 (photo PTBG)). W. H. Hatheway & A. Jimenez 1461
Hoffmannia riparia Standi., Field Mus. Nat. (Holotype US 2562634 (photos MEXU,
Hist., Bot. Ser. 17:214. 1937. Type: PTBG); Isotypes F, GH, n.v.).—Hoffmannia
GUATEMALA: Quezaltenango, Colomba, arborescens Donn. Sm.
720 m, 24 Sep 1934, A. F. Skutch 1291 Hoffmannia steyermarkii Standi., Publ. Field
(Holotype A, n.v. (photo and fragment F)). Mus. Nat. Hist., Bot. Ser. 22: 192. 1940.
—Hoffmannia regalis (Hook, f.) Hemsl. Type: GUATEMALA: San Marcos, between
Hoffmannia rosei B. L. Rob., Proc. Amer. Acad. Canjula and La Union, near southeast portion
Arts 45: 407. 1910. Type: MEXICO: Nayarit, of Volcan de Tacana, 2000-3000 m, 22 Feb
Tepic, near Pedro Paulo, 3 Aug 1897, J. N. 1940, J. A. Steyermark 36426 (Holotype F
Rose 1968 (Holotype US 300860 (photos 1035015 (photo PTBG); Isotype A, n.v.).
MEXU, PTBG); Isotype GH, n.v.). Hoffmannia strigillosa Hemsl., Diag. PI. Nov.
31. 1879. Type: MEXICO: Without precise
Hoffmannia rotata Donn. Sm., Bot. Gaz. locality, Salle s.n. (Holotype K, n.v.).
(Crawfordsville) 18: 204. 1893. Type: Hoffmannia subauriculata Standi., J. Wash.
GUATEMALA: Quiche, San Miguel Acad. Sci. 18: 179. 1928. Type: COSTA
Uspantan, 7000 ft., Apr 1892, Heyde & Lux RICA: Cartago, El Mufieco, on the Rio
3169B (Lectotype US 943496 (photos MEXU, Navarro, 1400-1500 m, 6-7 Mar 1926, P. C.
PTBG), here designated). Syntype: Standley & R. Torres R. 50956 (Holotype US
GUATEMALA: Quiche, San Miguel 1306554 (photos MEXU, PTBG)).
Uspantan, 7000 ft., Apr 1892, Heyde & Lux Hoffmannia teruae L. O. Williams. & A. R.
3174 (P (photos MEXU, PTBG)). The US Molina, Fieldiana, Bot. 36: 59. 1973. Type:
sheet of Heyde & Lux 3169B v GUATEMALA: San Marcos, near Aldea
Fraternidad, between San Rafael Pie de la
Cuesta and Palo Gordo, west facing slopes of J. Valerio 57 (Holotype US 1206194 (photos
Sierra Madre Mountains, 1800-2400 m, MEXU, PTBG)).
10-18 Dec 1963, L. O. Williams, A. Molina R. Hoffmannia vesiculifera Standi, Publ. Field
& T P. Williams 26028 (Holotype F 1651553 Columbian Mus., Bot. Ser. 4: 285. 1929.
(photo PTBG); Isotype EAP, n.v.). Type: PANAMA: Bocas del Toro, Buena
Hoffmannia tetrastigma Dorm. Sm., Bot. Gaz. Vista Camp, Chiriqui Trail, 750 m, Jan-Mar
(Crawfordsville) 27: 336. 1899. Type: 1928, G. P. Cooper 231 (Holotype F 579284
GUATEMALA: Santa Rosa, Zamorora, 1800 (photo PTBG)).
m, Apr 1893, Heyde & Lux 4490 (Holotype Hoffmannia viridis Rusby, Descr. N. Sp. S.
US 354989 (photos MEXU, PTBG); Isotypes Amer. PI. 134. 1920. Type: COLOMBIA:
F, US, 2 sheets (photos MEXU, PTBG)). Santa Marta, H. H. Smith 2649 (Holotype NY,
Hoffmannia tonduzii Standi., Contr. U.S. Natl. n.v., photo and fragment F).
Herb. 20: 205. 1919. Type: COSTA RICA: Hoffmannia vulcanicola Standi. & Steyerm.,
Forets de Las Vueltas, Tucurrique, 635-700 Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 251.
m, May 1899, A. Tonduz 13373 (Holotype US 1947. Type: GUATEMALA: Solola, southern
941386 (photos MEXU, PTBG)). slopes of Volcan de Atitlan above Finca
—Hoffmannia longipetiolata Pol. Moca, 1000-1250 m, 20 Jun 1942, J. A.
Hoffmannia trichocalyx Standi., J. Wash. Steyermark 47931 (Holotype F 1132754
Acad. Sci. 18: 181. 1928. Type: COSTA (photo PTBG)).
RICA: Alajuela, vicinity of Fraijanes, Hoffmannia wilsonii Standi., Publ. Field Mus.
1500-1700 m, 12-13 Feb 1926, P. C Nat. Hist., Bot. Ser. 22: 117. 1940. Type:
Standley & R. Torres R. 47690 (Holotype US GUATEMALA: Alta Verapaz, Finca Los
1305242 (photos MEXU, PTBG); Isotype US Alpes, 1200 m, 21 Mar 1939, C. L. Wilson
1305243 (photos MEXU, PTBG)). 344 (Holotype F 968020 (photo PTBG)).
Hoffmannia tuerckheimii Donn. Sm., Bot. Hoffmannia witheringioides Dwyer, Ann.
Gaz. (Crawfordsville) 47: 254. 1909. var. Missouri Bot. Gard. 67: 244. 1980. Type:
tuerckheimii. Type: GUATEMALA: Alta PANAMA: Veraguas, Rio Segundo Braso, 8
Verapaz, bei Coban, 1600 m, Mar 1908, H. km beyond Escuela Agricola Alto Piedra
von Tuerckheim 11.2160 (Holotype US 941383 beyond Santa Fe, 750 m, 24 Jul 1974, T. B.
(photos MEXU, PTBG); Isotype US 941384 Croat 25551 (Holotype MO 2214610; Isotype
(photos MEXU, PTBG)). MO 2207374).
Hoffmannia tuerckheimii Donn. Sm. var. Hoffmannia woodsonii Standi., Ann. Missouri
glabra Standi. & Steyerm., Publ. Field Mus. Bot. Gard. 28: 471. 1941. Type: PANAMA:
Nat. Hist., Bot. Ser. 23: 250. 1947. Type: Chiriqui, vicinity of Callejon Seco, Volcan de
GUATEMALA: El Progreso, Sierra de las Chiriqui, 1700 m, 17 Jul 1940, R. E.
Minas, Montana Canahui, between Finca San Woodson, Jr. & R. W. Schery 503 (Holotype
Miguel and summit of mountain, upper limits MO 1210619 (photo PTBG); Isotypes F
of Finca Caieta, 1600-2300 m, 10 Feb 1942, 1085625 (photo PTBG), GH, n.v.).
J. A. Steyermark 43807 [43806 in protologue] —Hoffmannia longipetiolata Pol.
(Holotype F 1133774 (photo PTBG)).
Hoffmannia uniflora Standi., Contr. U.S. Natl Holtonia Standi., Trop. Woods. 30: 37. 1932.
Herb. 20: 204. 1919. Type: GUATEMALA: Type: Holtonia myriantha (Standi.) Standi.
Alta Verapaz, Coban, 1100 m, Feb 1908, H. Herein treated as a synonym of Elaeagia
von Tuerckheim 11.2107 (Holotype US 941382 Wedd.]
(photos MEXU, PTBG)). Holtonia myriantha (Standi.) Standi., Trop.
Hoffmannia valerii Standi., J. Wash. Acad Sci Woods 30: 37. 1932. Sickingia myriantha
18: 178. 1928. Type: COSTA RICA: Standi., Publ. Field Columbian Mus., Bot. Ser.
Guanacaste, El Arenal, 600 m, 20 Mar 1923 7: 27. 1930. Type: COLOMBIA: Magdalena:
^. NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Las Nubes road, region of Santa Marta, 1200 n.v.).—Hedyotis arenaria (Rose) W. H.
m, 3 Dec 1898, H. H. Smith 1810 (Holotype F
138660; Isotypes MO, NY, n.v., US, n.v., W, Houstonia asperuloides (Benth.) A. Gray, Proc.
n.v., Y, n.v). Simira.—Elaeagia myriantha Amer. Acad. Arts 5: 158. 1860. Hedyotis
(Standi.) C. M. Taylor & Hammel asperuloides Benth., Bot. Voy. Sulphur 19,/?/.
13. 1844. Type: MEXICO: Baja California,
Houstonia L., Sp. PI. 1: 101. 1753. Type: Cabo San Lucas, Barclay s.n. (Holotype BM,
Houstonia caerulea L. [Herein treated as a n.v.).—Hedyotis asperuloides Benth.
synonym of Hedyotis L., although some Houstonia asperuloides (Benth.) A. Gray var.
authors (e.g., Terrell, 1996) recognize brandegeana (Rose) Wiggins in Shreve &
Houstonia as a genus.] Wiggins, Veg. Fl. Sonoran Desert 2: 1399.
Houstonia acerosa (A. Gray) Benth. & Hook, f, 1964. Houstonia brandegeanna Rose, Contr.
Gen. PI. 2: 60. 1873. Hedyotis acerosa A. U.S. Natl. Herb. 1: 83. 1890. Type: MEXICO:
Gray, PI. Wright. 1: 81. 1852. Type: U.S.A.: Baja California, La Paz, 20 Jan-5 Feb 1890,
Texas, high prairies from Live Oak Creek to E. Palmer 31 (Holotype US 47259 (photos
Las Moras, C. Wright 237 (Holotype GH, MEXU, PTBG); Isotypes F, GH, n.v.).
n.v.). Oldenlandia.—Hedyotis acerosa A. —Hedyotis asperuloides Benth. f.
brandegeana (Rose) W. H. Lewis
Houstonia acerosa (A. Gray) Benth. & Hook. f. Houstonia bouvardioides (Seem.) Benth. &
subsp. polypremoides (A. Gray) Terrell, Hook, f, Gen. PI. 2: 60. 1873. Hedyotis
Brittonia31: 168. 1979. Houstonia bouvardioides Seem., Bot. Voy. Herald 2:
polypremoides A. Gray, Proc. Amer. Acad. 296. 1856. Type: MEXICO: Without precise
Arts 21: 379. 1886. Type: MEXICO: locality [Durango ?], Sierra Madre, Seemann
Chihuahua, Santa Eulalia Mountains, 26 Sep s.n. (Holotype BM, n.v.; Isotypes C, n.v.,
1885, C. G. Pringle 356 (Lectotype GH, fragment F (photo PTBG), GH, n.v.).
n.v.; Isolectotypes F, MO (photos MEXU, —Bouvardia bouvardioides (Seem.) Standi.
PTBG), US (photos MEXU, PTBG); Houstonia brandegeana Rose, Contr. U.S. Natl.
designated by Lewis, 1968).—Hedyotis Herb. 1: 83. 1890. Type: MEXICO: Baja
acerosa A. Gray var. polypremoides (A. California, La Paz, 20 Jan-5 Feb 1889, E.
Gray) W. H. Lewis Palmer 31 (Holotype US 47259 (photos
Houstonia angustifolia Michx., Fl. Bor. Amer. MEXU, PTBG); Isotypes F, GH, n.v.).
1: 85. 1803. Type: U.S.A.: Florida, in —Hedyotis asperuloides Benth. f.
submaritimus Floridae, collector not cited brandegeana W. H. Lewis
(Holotype presumably P, n.v.).—Hedyotis Houstonia brevipes Rose, Contr. U.S. Natl.
nigricans (Lam.) Fosberg Herb. 1: 83. 1890. Type: MEXICO: Baja
Houstonia angustifolia Michx. var. scabra S. California, Santa Rosalia, 24 Feb-3 Mar 1889,
Watson, Proc. Amer. Acad. Arts 18: 97. 1883. E. Palmer 202 (Holotype US 47258 (photos
Type: MEXICO: Coahuila, Saltillo, Caracol MEXU, PTBG); Isotypes F, GH, n.v., US
Mountains, 1-15 Apr 1880, E. Palmer 410 (photos MEXU, PTBG)).—Hedyotis
(Holotype GH, n.v.; Isotype US (photos brevipes (Rose) W. H. Lewis
MEXU, PTBG)).—Hedyotis nigricans Houstonia coccinea Andr., Bot. Repos.pl. 106.
(Lam.) Fosberg 1800. Type: No herbarium specimens are
Houstonia arenaria Rose, Contr. U.S. Natl. known to exist, only the drawing pi. 106.
Herb. 1: 70. 1890. Type: MEXICO: Baja —Bouvardia ternifolia (Cav.) Schltdl.
California, La Paz, 20 Jan-5 Feb 1890, E. Houstonia drymarioides Standi., J. Wash. Acad.
Palmer 28 (Holotype US 47260 (photos Sci. 18: 162. 1928. Type: MEXICO:
MEXU, PTBG); Isotypes F, GH, Tamaulipas, uthofVic
1000 m, 9 Apr 1926, R. Runyon 870
(Holotype US 1315865 (photos MEXU, slopes, mountains along route 175, 28 km by
PTBG); Isotype F) —Hedyotis c road south of Valle Nacional on way to Ixtlan
(Standi.) W. H. Lewis de Juarez, ca. 2000 m, 30 Jul 1959, R. M.
Houstonia fasciculata A. Gray, Proc. Amer. King 2107 (Holotype MICH, n.v.; Isotype
Acad. Arts 17: 203. 1882. Syntypes: U.S.A.: MICH, n.v.).—Hedyotis kingii (Terrell) G. L.
Texas, Presidio, Bigelow s.n. (GH); New
Mexico, Organ Mountains, G. R. Vasey s.n. Houstonia latifolia Willd. ex Roem. & Schult.,
(GH, n.v.); MEXICO: Coahuila, near Parras Syst. 527. 1808. Not Hedyotis latifolia (M.
and Monclova, E. Palmer 404 (GH, n.v.), E. Martens & Galeotti) Walp., 1846. Type:
Palmer 406 (GH, n.v.).—Hedyotis intricata America borealis, Muhlenberg s.n.
(presumably B-WI, n.v.).—Hedyotis exigula
Houstonia fruticosa Willd. ex Roem. & Schult., W. H. Lewis
Syst. 3: 527. 1818. Type: VENEZUELA: Houstonia longipes S. Watson, Proc. Amer.
Monagas, Caripe, Humboldt s.n. (B, Acad. Arts 18: 97. 1883. Syntypes: MEXICO:
presumably destroyed).—Declieuxia Without precise locality, States of Coahuila
fruticosa (Willd. ex Roem. & Schult.) Kuntze and Nuevo Leon, Feb-Oct 1880, E. Palmer
Houstonia fruticosa Rose, Contr. U.S. Natl. 394 (GH, n.v.; US 47845 (photos MEXU,
Herb. 1: 132. 1892. Not Hedyotis fruticosa L., PTBG)); Nuevo Leon, Monterrey, Feb 1-26
1753, nor Willd. ex Roem. & Schult., 1818. 1880, E Palmer 395 (GH, n.v., US 70739
Type: MEXICO: Baja California, Carmen (photos MEXU, PTBG), US 1390542 (photos
Island (Isla Carmen), 1-7 Nov 1890, E. MEXU, PTBG)).—Hedyotis palmeri (A.
Palmer 885 (Holotype US 47256 (photos Gray) W. H. Lewis
MEXU, PTBG); Isotypes MO (photos Houstonia mucronata (Benth.) B. L. Rob., Proc.
MEXU, PTBG), US (photos MEXU, PTBG)). Amer. Acad. Arts 45: 401. 1910. Hedyotis
—Hedyotis mucronata Benth. mucronata Benth., Bot. Voy. Sulphur 19.
Houstonia gracilenta I. M. Johnst., Proc. Calif. 1844. Type: MEXICO: Baja California,
Acad. IV, 12: 1174. 1924. Type: MEXICO: Magdalena Bay, Barclay s.n. (Holotype BM,
Baja California, San Diego Island, rocky n.v.).—Hedyotis mucronata Benth.
island slope facing sea, 2 May 1921,1. M. Houstonia mullerae (Fosberg) Terrell, Brittonia
Johnston 3927 (Holotype UC, n.v.; Isotypes 31: 169. 1979. Hedyotis mullerae Fosberg,
GH, n.v., US (photos MEXU, PTBG)). Lloydia4: 288. 1941. Type: MEXICO:
—Hedyotis gracilenta (I. M. Johnst.) W. H. Coahuila, Mun. de Cuatro Cienegas, Canon
del Agua, Sierra de la Madera, 9 Sep 1939,
Houstonia gracilis Brandegee, Zoe 5: 238. C. H. Muller 3234 (Holotype US 2133305 ex
1907. Not Hedyotis gracilis DC, 1830. Type: NA (photos MEXU, PTBG); Isotype GH,
MEXICO: Veracruz, Orizaba, May 1904, n.v).—Hedyotis mullerae Fosberg
C. A. Purpus 1251 (Holotype UC, n.v.; Houstonia nigricans (Lam.) Fernald, Rhodora
Isotypes F, MO (photos MEXU, PTBG)). 42: 299. 1940. Gentiana nigricans Lam.,
—Hedyotis exigula W. H. Lewis Encycl. 2: 645. 1786. Type: MEXICO:
Houstonia humifusa (A. Gray) A. Gray, Proc. Collector unknown, (presumably P-LAM,
Amer. Acad. Arts 4: 314. 1859. Hedyotis n.v.).—Hedyotis nigricans (Lam.) Fosberg
humifusa A. Gray, Boston J. Nat. Hist. 6- 216 Houstonia palmeri A. Gray, Proc. Amer. Acad.
1850. Type: U.S.A.: Texas, near Arts 17: 202. 1882. Syntypes: MEXICO:
Fredrickburg, 1847, F. Lindheimer 621 Coahuila, Sierra Madre, 40 mi. south of
(Holotype GH, n.v.; Isotypes BM, n.v., K, Saltillo, 25-31 Jul 1880, E. Palmer 397 (GH,
n.v., MICH, n.v., MO, PH, n.v., US). ' n.v., US 47431 (photos MEXU, PTBG); E.
Houstonia kingii Terrell, Brittonia 32: 491, fig Palmer 398 (GH, n.v., F (2 sheets), US 70740
2. 1980. Type: MEXICO: Oaxaca, E-facing
i NOMENCLATOR OF MEXICAN AND CENTRAL ;
(photos MEXU, PTBG)).—Hedyotis palmeri Brandegee s.n. (Holotype UC, n.v.; Isotypes
(A. Gray) W. H. Lewis GH, n.v., US (photos MEXU,
Houstonia parvula Brandegee, Zoe' 5: 221. PTBG)).—Hedyotis sinaloae W. H. Lewis
1905. Not Hedyotis parvula (A. Gray) Houstonia rubra Cav., Icon. 5: 48, /. 474, fig. I.
Fosberg, 1943. Type: MEXICO: Sinaloa, 1799. Type: MEXICO: Hidalgo, Ixmiquilpan,
vicinity of Culiacan, 12 Oct 1904, T. S. Nee s.n. (Holotype presumably MA, n.v.).
Brandegee s.n. (Holotype UC, n.v.; Isotypes —Hedyotis rubra (Cav.) A. Gray
GH, n.v., US (photos MEXU, PTBG)). Houstonia serpyllacea (Schltdl.) C. L. Sm. ex
—Hedyotis sinaloae W. H. Lewis Greenman, Proc. Amer. Acad. Arts 32: 284.
Houstonia peninsularis Brandegee, Zoe 5: 160. 1897. Hedyotis serpyllacea Schltdl., Linnaea
1903. Type: MEXICO: Baja California, Cape 9: 599. 1834. Type: MEXICO: Mexico, inter
Region, Sierra de la Trinidad, Nov 1902, T. S. La Joya et San Salvador, Jun 1829, Schiede &
Brandegee s.n. (Holotype UC, n.v.; Isotypes Deppe 265 (Holotype presumably HAL, n.v.;
GH, n.v., US (photos MEXU, PTBG)). Isotypes B (photo MO), F (photo PTBG),
—Hedyotis peninsularis (Brandegee) W. H. MO).—Hedyotis serpyllacea Schltdl.
Houstonia sharpii Terrell, Brittonia 32: 490, fig.
Houstonia polypremoides A. Gray, Proc. Amer. I. 1980. Type: MEXICO: Hidalgo, camino
Acad. Arts 21: 379. 1886. Type: MEXICO: entre Metepec y Tenango de Doria, entre 16 y
Chihuahua, Santa Eulalia Mountains, 26 Sep 31.5 km al NNE de Tulancingo, 18 km N.NE.
1885, C. G Pringle 356 (Lectotype GH; n.v.; de Metepec, 29 Mar 1974, S. D. Koch 7423
Isolectotypes F, MO (photos MEXU, PTBG), (Holotype US 2754863 (photos MEXU,
US (photos MEXU, PTBG); designated by PTBG); Isotypes CHAPA, ENCB, MEXU).
Lewis, 1968). Syntype: MEXICO: Chihuahua, —Hedyotis sharpii (Terrell) G. L. Nesom
Santa Eulalia Mountains, 29 May 1885, C. G Houstonia teretifolia Terrell, Brittonia 31: 166.
Pringle 16 (GH, n.v., US (photos MEXU, 1979. Type: MEXICO: Coahuila, 12 km NNE
PRBG)).—Hedyotis polypremoides (A. of Las Margaritas on easternmost ridge of
Gray) Shinners Sierra de las Margaritas, 1300-1400 m, 24
Houstonia polypremoides var. bigelovii Sep 1972, M. C. Johnston, F. Chiang & T.
Greenman, Proc. Amer. Acad. Arts 32: 291. Wendt 9508E (Holotype TEX, n.v.).
1897. Type: U.S.A.: Florence Mountains, Jun —Hedyotis teretifolia (Terrell) G. L. Nesom
1852, Bigelow 437 (Holotype GH, n.v.). Houstonia triflora (Kunth) A. Gray, Proc. Amer.
—Hedyotis polypremoides (A. Gray) W. H. Acad. Arts 4: 314. 1860. Bouvardia triflora
Kunth in Humb., Bonpl. & Kunth, Nov. Gen.
Houstonia prostrata Brandegee, Zoe 5: 105. Sp. 3: 386. 1820. Type: MEXICO: Without
1901. Not Hedyotis prostrata Blume, 1823 precise locality, Humboldt & Bonplands.n.
norKorth., 1851. Type: MEXICO: Baja (Holotype P-Bonpland, n.v. (photo MO)).
California, La Palma, Cape Region, 25 Sep —Bouvardia multiflora (Cav.) Schult. &
1899, T. S. Brandegee s.n. (Holotype UC, Schult. f.
n.v.; Isotypes GH, n.v., US 382879 (photos Houstonia umbratilis B. L. Rob., Proc. Amer.
MEXU, PTBG), US 735791 (photos MEXU, Acad. Arts 45: 401. 1910. Type: MEXICO:
PTBG)).—Hedyotis vergrandis W. H. Lewis Nuevo Leon, shaded cliffs of mountains, near
Houstonia prostrata Brandegee var. parvula Monterrey, 25 Apr 1906, C. G. Pringle 13877
(Brandegee) Wiggins in Shreve & Wiggins, (Holotype GH, n.v.; Isotypes MO (photos
Veg. Fl. Sonoran Desert 2: 1399. 1964. MEXU, PTBG), US (photos MEXU, PTBG)).
Houstonia parvula Brandegee, Zoe 5: 221. —Hedyotis umbratilis (B. L. Rob.) W. H.
1905. Not Hedyotis parvula (A. Gray)
Fosberg, 1943. Type: MEXICO: Sinaloa, Houstonia wrightii A. Gray, Proc. Amer. Acad.
vicinity of Culiacan, 12 Oct 1904, T. S. Arts 17: 202. 1882. Type: MEXICO: Nuevo
Leon, shaded cliffs of mountains, near H Pittier 3889 (Holotype US 678963 (photo
Monterrey, 25 Apr 1906, C. G. Pringle 13877 PTBG); Isotype US (photo PTBG)).—Isertia
(Holotype GH, n.v.; Isotypes MO (photos laevis (Triana) B. M. Boom
MEXU, PTBG), US (photos MEXU, PTBG)). Isertia scorpioides B. M. Boom, Brittonia 36:
—Hedyotis wrightii (A Gray) Fosberg 435. 1984. Type: PANAMA: Colon, Atlantic
Houstonia xestosperma (B. L. Rob. & side of Isthmus on road to Pifia, 25 Aug 1967,
Greenman) Terrell, Brittonia 32: 492. 1980. J. H Kirkbride, Jr. & S. M. V. Hayden 316
Oldenlandia xestosperma B. L. Rob. & (Holotype NY, n.v.; Isotypes MO (2 sheets;
Greenman, Proc. Amer. Acad. Arts 32: 41. photos PTBG), NY, n.v.).
1896. Type: MEXICO: Oaxaca, Sierra de San
Felipe, open glades, 8000 ft., Jun 1894, C. G. Ixora L., Sp. PI. 1: 110. 1753. Type: Ixora
Pringle 4692 (Lectotype GH, n.v.; coccinea L.
Isolectotypes BM, n.v., F, K, n.v., MO Ixora alba Sesse & Mo?., La Naturaleza, ser. 2,
(photos MEXU, PTBG), MSC, n.v., PH, n.v., 1. app. 16. 1887. Not Ixora alba L., 1753, nor
UC, US, 2 sheets (photos MEXU, PTBG), Roxb., 1814, nor Sus. ex Wight & Arm, 1834.
VT, n.v.; designated by Terrell, 1980). Type: MEXICO: Habitat in India ad montibus
Hedyotis.—Bouvardia xestosperma (B. L. Mazatlan (presumably MA, n.v.).—Status
Rob. & Greenman) Terrell & S. D. Koch
Uacq., Hort. Schonbr. 3:4,/?/.
Isertia Schreb., Gen. PI. 1: 234, n. 602. 1789. 257. 1798. Type: Cultivated at Schobrunn,
Type: Isertia coccinea (Aubl.) Vahl Vienna, said to have been brought from
unii Bartlett, Proc. Amer. Acad. Arts Mexico.—Bouvardia ternifolia (Cav.)
43: 59. 1907. Type: GUATEMALA: Izabal, Schltdl.
Puerto Barrios, 24 Feb 1905, C. C. Deam 48 Ixora aphylla Sesse & Mo?., Pi. Nuev. Hispan.
(Holotype GH, n.v.).—Isertia haenkeana 16. 1887; La Naturaleza, ser. 2, 1. app. 16, ed.
DC. 2, 15. 1887. Type: MEXICO: Habitat prope
Isertia deamii Bartlett var. stenophylla Dorm. Xochistlan, Sesse & Mociiio s.n. (presumably
Sm., Bot. Gaz. (Crawfordsville) 61: 374. MA, n.v.).—Status uncertain.
1916. Type: COSTA RICA: Boca Culebra, Ixora casei Hance in Walp., Ann. Bot. Syst. 2:
plaines des Pacifique, 50 m, 20 Jan 1898, H. 754. 1852. Type: CAROLINE ISLANDS:
Pit tier 11989 (Holotype US 941413 (photo Strong Island, Q. W. Case s.n. (Isotype or
PTBG); Isotypes F (photo PTBG), P, n.v., possible Holotype K, n.v.). For typification
fragment F).—Isertia haenkeana DC. see Fosberg & Sachet (1989a), who note that
Isertia haenkeana DC, Prodr. 4: 437. 1830. var. casei is cultivated in Mesoamerica.
Type: MEXICO: Without precise locality, Ixora coccinea L., Sp. PI. 1: 110. 1753.
Haenke s.n. (Holotype presumably PR, n.v.). Lectotype: Rheede, Hortus malabaricus 2:17,
Isertia laevis (Triana) B. M. Boom, Brittonia /. 13. 1679. Designated by Fosberg & Sachet
36: 433. 1984. Cassupa laevis Triana, Ann. (1989b), who note that Ixora coccinea L. var.
Sci. Nat, Bot., Ser 4, 9: 44. 1858. Type: coccinea and var. intermedia Fosberg &
COLOMBIA: Cundinamarca, Triana 1838 Sachet are cultivated in Mexico and Central
(Lectotype P; Isolectotypes BR, n.v., COL,
n.v., L, n.v., P, n.v., US, n.v.; designated by Ixora coccinea L. var. intermedia Fosberg &
Boom, 1984). Sachet, Baileya 23: 80. 1989. Type: VIRGIN
Isertia panamensis (Standi.) Standi., Publ. Field ISLANDS: St. Croix, Teague Bay, cultivated,
Columbian Mus., Bot. Ser. 8: 346. 1931. F. R., Fosberg 55279 (Holotype US, n.v.)
Cassupa panamensis Standi., Contr. U.S. —Ixora coccinea L.
Natl. Herb. 18: 135. 1916. Type: PANAMA: Ixora finlaysoniana Wall, ex G. Don, Gen.
Colon, along Rio Fato, 10-100 m, 8 Jul 1911, Hist. Dichlam. PI. 3: 572. 1834. Type: EAST
i NOMENCLATOR OF MEXICAN AND CENTRAL
was the Santa Fe in Mexico (L. Andersson, F. Liebmann 11662 (Lectotype C (photos
pers. comm., 1996). No herbarium specimens MEXU, PTBG); Isolectotype US, n.v.;
are known to exist, and a neotype needs to be designated by Hamilton, 1989). Syntypes:
MEXICO: Veracruz, Mirador, Apr 1847, F.
Manettia seleriana Loes., Verhandl. Bot. Verein Liebmann 11577'(C (photos MEXU, PTBG);
Brandenburg 65: 107. 1923. Type: Mirador, F. Liebmann 11614 (C (photos
GUATEMALA: Izabal, Los Amates, 1 Jan MEXU, PTBG)). Psychotria.—Psychotria
1897, E. Seler & C. Seler 3309 (Holotype B, costivenia Griseb. var. costivenia
presumably destroyed (photos F, MO, US; Mapouria obovata Oerst., Amer. Centr. 17, t.
fragment F); Isotype US (photos MEXU, 14, figs. 3, 4. 1863. Not Psychotria obovata
PTBG)).—Manettia reclinata L. Ruiz & Pav., 1799. Type: MEXICO:
Manettia stenophylla Donn. Sm., Bot. Gaz. Veracruz, F. Liebmann s.n. (Holotype
(Crawfordsville) 56: 58. 1913. Type: COSTA presumably C, n.v.). Psychotria.—Psychotria
RICA: Tucurrique, Las Vueltas, 635 m, Jan quinqueradiata Pol.
1899, A. Tonduz 12969 (Holotype US 943614 Mapouria papantlensis, Oerst., Amer. Centr. 17.
(photos F, MEXU, PTBG, US); Isotype B, t. 14, fig. 5. 1863. Type: MEXICO: Veracruz,
destroyed (fragment F)).—Manettia barbata Papantla,/ Liebmann s.n. (Holotype
Oerst. presumably C, n.v.). Uragoga.—Psychotria
Manettia zimapanica Hemsl., Diag. PI. Nov. papantlensis (Oerst.) Hemsl.
Mexic. 30. 1879. Type: MEXICO: Hidalgo, Mapouria remota (Benth.) Muell. Arg., Flora
Zimapan, Coulter 203 (Holotype K, n.v.). 59: 459. 1876. Psychotria remota Benth., J.
—Status u Bot. Hook. 3: 225. 1841. Type: BRAZIL: On
the Rio Negro, Schomburgk 963 (Holotype K,
Mapourea Aubl., Hist. PL Guiane 1: 175. 1775. n.v. (photo NY); Isotypes F, W, n.v.).
Type: Mapouria guianensis Aubl. [Herein —Psychotria remota Benth.
treated as a synonym of Psychotria L.] Mapouria salicifolia Oerst., Amer. Centr. 17, t.
Mapourea chamissoana Loes., Verhandl. Bot. 14, fig. 2. 1863. Not Psychotria salicifolia
Verein Brandenburg 65: 112. 1923. Type: Kunth in Humb., Bonpl. & Kunth, 1819.
MEXICO: Veracruz, Tecolute [Tecolutla], Type: MEXICO: Oaxaca, Lacoba [Lacova],
Schiede 1266 (Holotype B, presumably F. Liebmann s.n. (Holotype presumably C,
destroyed (photo MO)). Psychotria. n.v). Psychotria.—Psychotria papantlensis
—Psychotria erythrocarpa Schltdl. (Oerst.) Hemsl.
Mapourea graciliflora (Benth.) Oerst., Amer. Mapouria stipulata Oerst., Amer. Centr. 17, t.
Centr. 17, t. 14, fig. 6. 1863. Psychotria 14, fig. 7. 1863. Type: MEXICO: Veracruz,
graciliflora Benth. in Oerst., Vidensk. Papantla, F. Liebmann 11615 (Lectotype C,
Meddel. Dansk Naturhist. Foren. Kjobenhavn here designated (photos MEXU, PTBG);
1852: 35. 1853. Type: COSTA RICA: Isolectotype C (photos MEXU, PTBG)).
Alajuela, ad Naranjo, Apr 1847, Oersted Syntype: MEXICO: Veracruz, Papantla, F.
11611 (Holotype C (photos F, MEXU, Liebmann 11636 (C (photos MEXU, PTBG)).
PTBG); Isotypes K, n.v., US (photos MEXU, Liebmann 11615, a well preserved flowering
PTBG)).—Psychotria graciliflora Benth. specimen with a duplicate, is the best choice
Mapouria micrantha (Kunth) Wernham, Kew for lectotype. Psychotria.—Psychotria
Bull. 1914: 69. 1914. Type: PERU (?): A. costivenia Griseb. var. costivenia
Bonplands.n. (Holotype P-Bonpland, n.v. Mapouria tomentosa Oerst., Amer. Centr. 17, t.
(fragment F)).—Psychotria micrantha Kunth 14, fig. 8. 1863. Not Psychotria tomentosa
Mapouria miradorensis Oerst., Amer. Centr. 17, Mtill Arg. ex Mart., 1881, nor Hemsl., 1881.
/. 14, fig. 9. 1863. Type: MEXICO: Veracruz, Type: MEXICO: Veracruz, prope Papantla, F.
Mirador, in quercetum, 3000 ft., 1841-1843, Liebmann s.n. (Holotype presumably C, n.v.).
—Status uncertain, probably Psychotria MEXICO: Baja California, Cape [Cabo] San
micrantha Kunth in Humbl., Bonpl. & Kunth Lucas, Hinds 65 (Holotype BM, n.v.).
Mapouria trichogyne Muell. Arg. in Mart., Fl. Mitracarpus pallidus Hook. & Am., Bot.
Brasil. 6: 426. 188. Type: BRAZIL: Habitat in Beech. Voy. 4: 30. 1840. Type: MEXICO:
prov. de Alto Amazonas, prope Manaos ad Realjo, collector probably Sinclair s.n.
Rio Negro, Spruce s.n. (B, presumably (Holotype presumably K, n.v.).
destroyed). Geophila.—Geophila cordifolia Mitracarpus rhadinophyllus (B. L. Rob.) L. O.
Miq. Williams, Fieldiana, Bot. 29: 371. 1961.
Borreria rhadinophylla B. L. Rob., Proc.
Microsplenium Hook. f. in Benth. & Hook, f., Amer. Acad. 45: 409. 1910. Type: BRITISH
Gen. PI. 2: 4. 1873. Type: Microsplenium HONDURAS (BELIZE): On dry, sandy pine
coulteri Hook. f. [Herein treated as a synonym ridges, 23 Oct 1905, M. E. Peck 180
of Machaonia Humb. & Bonpl] (Holotype GH, n.v.; Isotype NY, n.v.)
Microsplenium coulteri Hook. f. in Benth. & Mitracarpus schizangius DC, Prodr. 4: 572.
Hook, f., Gen. PI. 2: 4. 1873. Type: MEXICO: 1830 (as Mitracarpum schizangium). Type:
Hidalgo, Zimapan, Coulter (Holotype K, MEXICO: Without precise locality, Haenke
n.v.).-^Machaonia coulteri (Hook, f.) Standi. s.n. (Holotype presumably PR, n.v.).
Mitracarpus schizangius DC. var. angx
Mitchella L., Sp. PI. 1: 111. 1753. Type: Oerst., Vidensk. Meddel. Dansk Naturhist.
Mitchella repens L. Foren. Kjobenhavn 1852: 7. 1853. Type:
Mitchella repens L., Sp. PI. 1: 111. 1753. Type: NICARAGUA: Volcan El Viejo, Oersted s.n.
U.S.A.: Habitat in Carolina, Terra Mariana, (Holotype C, n.v.).—Status uncertain.
Virginia, Herb. Linn. 135.1 (Lectotype LINN, Mitracarpus villosus (Sw.) Cham. & SchltdL,
n.v.; designated by Reveal et al., 1987: 215). Linnaea 3: 363. 1828. Spermacoce villosa
Sw., Prodr. Veg. Ind. Occ. 29. 1788. Type:
. [Here JAMAICA: Swartz s.n. (Holotype presumably
BM, n.v.).—Mitracarpus hirtus (L.) DC
Mitracarpus Zucc. ex Schult. & Schult f. in Montamans Dwyer, Ann. Missouri Bot. Gard.
Roem. & Schult., Mant. PI. 3: 210 (as 67: 284. 1980. Type: Montamans panamensis
Mitracarpum), 399. 1827. Type: Mitracarpus Dwyer [Herein treated as a synonym of
scaber Zucc. ex Schult. & Schult. f. Psychotria L.]
Mitracarpus breviflorus A. Gray, PI. Wright. 2: Montamans panamensis Dwyer, Ann. Missouri
68. 1853 (as Mitracarpum brevifiorum). Type: Bot. Gard. 67: 286, fig. 62. 1980. Type:
MEXICO: Sonora, mountain ravines and hill- PANAMA: Chiriqui, Cerro Horqueta, rain
sides near Santa Cruz, Sep 1851, C. Wright forest, ca. 5500 ft., 20 Jul 1968, J. D. Dwyer
1120 (Holotype GH, n.v.; Isotype US (photos & B. Lallathin 8792 (Holotype MO 2163225
MEXU, PTBG)).—Mitracarpus hirtus fL.) (photos MEXU, PTBG)).—Psychotria
DC. aggregata Standi.
Mitracarpus hirtus (L.) DC, Prodr. 4: 572.
1830. Spermacoce hirta L., Sp. PL, ed. 2, 1: Morinda L., Sp. PL 1: 176. 1753. Type: Morinda
148. 1762. Not Spermacoce hirta L., 1767,
nor Sw., 1791. Type: JAMAICA: Browne Morinda asperula Standi., Publ. Field Mus.
'Spermacoce # 3' (LINN 125.4, n.v.). For Nat. Hist., Bot. Ser. 22: 54. 1940. Type:
typification see Verdcourt (1975: 317-322). GUATEMALA: Izabal, hilly pine forest near
Mitracarpus linearis Benth., Bot. Voy. Sulphur Quirigua, 72-150 m, 26-27 Apr 1939, P. C.
20. 1844 (as Mitracarpium linear e). Type: Standley 72368 (Holotype F 991552 (photo
PTBG)).
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Morinda citrifolia L., Sp. PI. 1: 176. 1753. ..,Sp.Pl. 1: 177. 1753. Type:
Type: INDIA: (Holotype LINN 236.1, n.v. i frondosa L.
(microfiche MO)). Mussaenda acutiflora Bartl. ex DC, Prodr. 4:
Morinda guatemalensis (Donn. Sm.) Steyerm., 370. 1830. Type: MEXICO: Without precise
Mem. New York Bot. Gard. 23: 385. 1972. locality, Haenke s.n. (Holotype presumably
Appunia guatemalensis Donn. Sm., Bot. Gaz. PR, n.v., fragment G-DC, n.v. (microfiche 4:
(Crawfordsville) 48: 294. 1909. Type: 370.2)).—Status uncertain.
GUATEMALA: Izabal, Livingston, Jun 1906, Mussaenda candidissima (Vahl) Roem. &
H. von Tuerckheim III-230 (Holotype US Schult., Syst. Veg. 5:251. 1819.
943501 (photos MEXU, PTBG)).—Appunia Macrocncn: .M\, Symb.
guatemalensis Donn. Sm. Bot. 2: 38,/?/. 30. 1791. Type: TRINIDAD:
Morinda laxa Bartl. ex DC, Prodr. 4: 449. Vicinia St. Marthae, von Rohrs.n. (Holotype
1830. Type: MEXICO: Without precise C, n.v).—Calycophyllum candidissumum
locality, Haenke s.n. (Holotype presumably (Vahl) DC.
PR, n.v.).—Status uncertain. Mussaenda coccinea (Vahl) Poir. in Lam.,
Morinda mesochora Standi., Publ. Field Encycl. Meth. 4: 394. 1797. Macrocnemum
Columbian Mus., Bot. Ser. 4: 345. 1929. coccineum Vahl, Symb. Bot. 2: 38. 1791.
Type: BRITISH HONDURAS (BELIZE): Type: TRINIDAD: Without precise locality,
Honey Camp, Orange Walk, Sep 1928, C. L. von Rohrs.n. (Holotype C, n.v.).
Lundell 19 (Holotype F 580825; Isotype US —Warszewiczia coccinea (Vahl) Klotzsch
(photos MEXU, PTBG)). Mussaenda erythrophylla Schumach. &
Morinda muscosa Jacq., Sel. Stirp. Amer. 65. Thonn., Beskr. Guin. PI. 116. 1827. Type:
1763. Type: MARTINIQUE: Jacquin s.n. TROPICAL AFRICA: Guinea, Thonnings.n.
(location unknown).—Psychotria muscosa (Holotype presumably C, n.v.; probable
(Jacq.) Steyerm. Isotype G-DC, n.v. (microfiche 4: 371.8)).
Morinda panamensis Seem., Bot. Voy. Herald Cultivated in Mesoamerica.
136. 1854. Type: PANAMA: Panama, Mussaenda formosa Jacq., Enum. PI. Carib. 16.
Chagres, 10 Feb 1850, Fendler 148 (Holotype 1760. Type: COLOMBIA: Cartagena, Jacquin
BM, n.v.; Isotypes GH, n.v., MO (photos s.n. (presumably G, n.v.). Randia.
MEXU, PTBG), US (photo PTBG)). —Rosenbergiodendron formosum (Jacq.)
Morinda royoc L., Sp. PI. 1: 176. 1753 [as Fagerl.
roioc]. Type: Habitat in America calidiore, Mussaenda glabra Vahl, Symb. Bot. 3: 38.
Herb. Clifford 73, Morinda 1 (Lectotype BM; 1794. Type: Collector and location unknown.
designated by Lorence, 1993: 68). Cultivated in Mesoamerica.
Morinda yucatanensis Greenm., Publ. Field Mussaenda luteola Delile in Caill., Voy. Meroe
Columbian Mus., Bot. Ser. 2: 262. 1907. 4: 65. 1827. Type: ARABIA: Nubia, collector
Type: MEXICO: Yucatan, in brush lands unknown (Holotype presumably MPU, n.v.;
about Izamal, 1895, G F. Gaunter 362 probable isotype G-DC, n.v. (microfiche 4:
(Lectotype F 36165 (photo PTBG), here 371.9)). Cultivated in Mesoamerica.
designated; Isolectotypes GH, n.v., US Mussaenda spinosa Sesse & Moc., La
(photos MEXU, PTBG)). Note: in his Naturaleza, ser. 2, 1: 36. 1888. Type:
protologue Greenman cited duplicates of MEXICO: Habitat in Martinica, Indiarum
Gaunter 362 at both F and GH without Cartagine et calidis Novae Hispaniae
designating a holotype. An additional regionibus, Sesse & Mocirio 1583
duplicate exists at US. I have selected the (presumably MA, n.v. (fragment F)).
excellent flowering and fruiting specimen at F —Randia tetracantha (Cav.) DC.
as lectotype.—Morinda royoc L. Mussaenda tetracantha Cav., Icon. PI. 4: 20,/?/.
435. 1799. Type: MEXICO: [Probably
Guerrero], cerca Acapulco, Nee s.n. however, it does not occur in the Northern
(presumably MA, n.v. (photos F, MEXU, Hemisphere.
MO)).—Randia tetracantha (Cav.) DC. Nertera alsinoides Schltdl. & Cham., Linnaea 6:
413. 1831. Type: MEXICO: Questa Grande
Nauclea L., Sp. PL, ed. 2, 1: 243. 1762. Type: de Jalacingo, Schiede & Deppe 1271
Nauclea orientalis (L.) L., based on (Holotype presumably HAL, n.v.).
Cephalanthus orientalis L. —Didymaea alsinoides (Schltdl. & Cham.)
Nauclea cadamba Roxb., Fl. Ind. 2: 121. 1824. Standi.
Type: INDIA: Rheede, Hort. Ind. Malabar. 3: Nertera granadensis (Mutis ex L. f.) Druce,
tab. ii, Katou-tsjaka (1678-1703). Rep. Bot. Soc. Exch. Club Brit. Isles 1916:
—Neolamarckia cadamba (Roxb.) Bosser 637. 1917. Gomozia granadensis Mutis ex
Nauclea tomentosa Willd. ex Roem. & Schult, L. f., Suppl. PL 129. 1781. Type:
Syst. Veg. 5: 221. 1819. Type: COLOMBIA: COLOMBIA: America meridionali, Mutis s.n.
In ripa flumen Magdalenae prope Naers, (LINN 172.1, n.v. (microfiche)).
Humboldt s.n. (Holotype B-W 3910, n.v.).
—Uncaria tomentosa (Willd. ex Roem. & Nonatelia AubL, Hist. PL Guiane 1: 182, pi.
Schult.) DC. 70-75. Ml 5. Type: Nonatelia racemosa AubL
[Herein treated as a synonym of Psychotria
Neolamarckia Bosser, Bull. Mus. Natl. Hist. L-]
Nat, Paris, Ser. 4, Sect. B, Adansonia 6(3): Nonatelia officinalis Aubl, Hist. PL Guiane 1:
247. 1984. Type: Neolamarckia cadamba 182, tab. 70. 1775. Type: FRENCH
(Roxb.) Bosser GUIANA: Cayenne, Aublet s.n. (Holotype
Neolamarckia cadamba (Roxb.) Bosser, Bull. presumably BM, n.v.).—Psychotria
Mus. Natl. Hist. Nat., Paris, Ser. 4, Sect. B, s (AubL) Raeusch.
Adansonia 6(3): 247. 1984. Nauclea cadamba apanamensis DC, Prodr. 4: 466. 1830.
Roxb., Fl. Ind. 2: 121. 1824. Type: INDIA: Type: PANAMA: Without precise locality,
Rheede, Hort. Ind. Malabar. 3: tab. 33, Haenke s.n. (Holotype presumably PR,
Katou-tsjaka (1678-1703). Cultivated and n.v.).—Status uncertain, probably
naturalized in Costa Rica according to Burger Psychotria racemosa (AubL) Raeusch.
& Taylor (1993). Nonatelia racemosa AubL, Hist. PL Guiane 1:
187, t. 72. Ml 5. Type: FRENCH GUYANA:
In sylvis territorii Orapu, Aublet s.n.
(Holotype P, n.v.).—Psychotria r
Urb. Rich.
Nernstia mexicana (Zucc. & Mart, ex DC.)
Urb., Symb. Antill. 9: 146. 1923. Coutarea Nothophlebia Standi., Contr. U.S. Natl. Herb.
mexicana Zucc. & Mart, ex DC, Prodr. 4: 17: 438. 1914. Type: Nothophlebia
350. 1830. Type: MEXICO: Without precise costancensis Standi. [Herein treated as a
locality, Martius in 1829 (Holotype G-DC, synonym of Pentagonia Benth.]
n.v. (IDC 800.680: II.7)). Cigarilla. Nothophlebia costaricensis Standi., Contr. U.S.
Natl. Herb. 17: 438. 1914. Type: COSTA
Nertera Banks & Sol. ex C. F. Gaertn., Fruct. RICA: Collines de Moin, littoral atlantique,
Sem. PL 1: 124, pi. 26. 1788. Nomen Nov 1899, H. Pittier 16024 (Holotype US
conservandum. Type: Nertera depressa 578472 (photos MEXU, PTBG)).
Gaertn. [=Nertera granadensis (Mutis ex L. f.) —Pentagonia costaricensis (Standi.) W. C
Druce.] Note: Nertera depressa was listed, as Burger & C M. Taylor
a synonym of Gomozia granadensis, by
Standley (1938) as occurring in Costa Rica;
i NOMENCLATOR OF MEXICAN AND CENTRAL AMER
Oldenlandia L., Sp. PI. 1: 119. 1753. Type: Oldenlandia herbacea (L.) Roxb., Hort. Bengal.
Oldenlandia corymbosa L. [Herein treated as a 11. 1814. Hedyotis herbacea L., Sp. PI. 1:
synonym of Hedyotis L.] 102. 1753. Type: CEYLON: (Presumably
Oldenlandia acerosa (A. Gray) A. Gray, PI. LINN, n.v.).—Hedyotis herbacea L.
Wright. 2: 67. 1853. Hedyotis acerosa A. Oldenlandia humifusa (A. Gray) A. Gray,
Gray, PI. Wright. 1:81. 1852. Type: U.S.A.: Smiths. Contr. Knowl. 5: 68. 1853. Hedyotis
Texas, high prairies from Live Oak Creek to humifusa A. Gray, J. Boston Soc. Nat. Hist. 6:
the Las Moras, Jun 1849, C Wright 237 216. 1850. Type: U.S.A.: Texas, near
(Holotype GH, n.v.; Isotype US). Houstonia, Fredericksburg, 1847, F. Lindheimer 621
Mallostoma.—Hedyotis acerosa A. Gray (Holotype GH, n.v.; Isotypes BM, n.v.,
Oldenlandia angustifolia (Michx.) A. Gray, PI. MICH, n.v., MO, PH, n.v., US). Houstonia.
Wright. 2: 68. 1853. Houstonia angustifolia —Hedyotis humifusa A. Gray
Michx., Fl. Bor. Amer. 1: 85. 1803. Type: Oldenlandia lancifolia (Schumach.) DC, Prodr.
U.S.A.: Florida in submantimis, collector not 4: 425. 1830. Hedyotis lancifolia Schumach.
cited (presumably P, n. v.).—Hedyotis in Schumach. & Thonn., Beskr. Guin. PI. 72.
nigricans (Lam.) Fosberg 1827. Type: GHANA: Valley of Aquapim,
Oldenlandia callitrichoides Griseb., Mem. Thonning 210 (Lectotype C, n.v.; Isolectotype
Amer. Acad. Arts, n.s. 8: 506. 1863. Type: S, n.v.; designated by Verdcourt, 1976: 292).
CUBA: Without precise locality, ad ripas —Hedyotis lancifolia Schumach. in
rivulorum, Wright 268 (Lectotype GOET, Schumach. & Thonn.
n.v.; designated by Terrell & Lewis, 1990). Oldenlandia latifolia M. Martens & Galeotti,
Syntype: CUBA: Wright 1640 (GOET, n.v.). Bull. Acad. Roy. Sci. Bruxelles 11: 235. 1844.
Oldenlandiopsis.—Hedvotis callitrichoides Type: MEXICO: Veracruz, Mirador, near
(Griseb) W. H. Lewis Xalapa, 3000 ft., 1840, H Galeotti 2556
Oldenlandia corymbosa L., Sp. PI. 1: 119. 1753. (Holotype BR, n.v.; Isotypes F (photos
Lectotype: "Oldenlandia," Plumier, Nov. PI. MEXU, PTBG), GH).—Hedyotis exigula
Amer. t. 36. 1703; designated by Verdcourt W. H. Lewis
(1976: 308).—Hedyotis corymbosa (L.) Oldenlandia microtheca (Schltdl. & Cham.)
DC, Prodr. 4: 428. 1830. Gerontogea
Oldenlandia deppeana (Schltdl. & Cham.) DC, microtheca Schltdl. & Cham., Linnaea 5: 169.
Prodr. 4: 428. 1830. Gerontogea deppeana 1830. Type: MEXICO: Veracruz, Barranca de
Schltdl. & Cham., Linnaea 5: 169. 1830. Tioselos in umbrosis, Sep, Schiede & Deppe
Type: MEXICO: Veracruz, ad San Andres, 390 (Holotype presumably HAL, n.v.; Isotype
1829, Schiede & Deppe 391 (Holotype MO (photos MEXU, PTBG), label states In
presumably HAL, n.v.; Isotype MO (photos sylvis Papanatlae, Jan., Dec, corolla alba).
MEXU, PTBG)). Hedyotis.—Hedyotis —Hedyotis microtheca (Schltdl. & Cham.)
microtheca (Schltdl. & Cham.) Steud. Steud.
Oldenlandia depressa Sesse & Moc., Fl. Mexic. Oldenlandia ovata S. Watson, Proc. Amer.
27. 1893. Not O. depressa Willd. in L., 1798. Acad. Arts 18: 97. 1883. Not Hedyotis ovata
Type: MEXICO: Habitat in montibus S. Thunb. ex Maxim., 1883. Type: MEXICO:
Augustine et viciniis. Floret Julio, Sesse, Nuevo Leon, Guajuco, 27 miles SE of
Mocino, Castillo & Maldonado 601 (Fl. Mex. Monterrey, 1-8 Mar 1880, E. Palmer 399
Ic 325) (Holotype presumably MA, n.v.; (Lectotype GH, n.v.; Isolectotype F (photos
Isotype F, n.v.) —Richardia tricocca (Torr. MEXU, PTBG), US (photo PTBG);
& A. Gray) Standi, subsp. tetracocca (M. designated by Terrell, 1978, in herb.).
Martens & Galeotti) W. H. Lewis & R. L. Syntype: MEXICO: Ervandberg 200 (GH,
Oliv. n.v.).—Hedyotis watsonii W. H. Lewis
Oldenlandia pringlei B. L. Rob., Proc. Amer.
Acad. Arts 27: 169. 1892. Type: MEXICO:
San Luis Potosi, alkaline plaines, Hacienda de
Angostura, 27 Jun 1891, C. G. Pringle 3758 Omiltemia Standi., J. Wash. Acad. Sci. 8: 427.
(Holotype GH, n.v.; Isotypes F, MO (photo 1918. Type: Omiltemia longipes Standi.
PTBG), US (2 sheets; photos MEXU, Omiltemia filisepala (Standi.) C. V. Morton,
PTBG)).—Hedyotis pringlei (B. L. Rob.) Baileya 16: 98. 1968. Kohleria filisepala
W. H. Lewis Standi, Publ. Field Mus. Nat. Hist., Bot. Ser.
Oldenlandia racemosa Sesse & Moc., Fl. 22: 107. 1940. Type: MEXICO: Chiapas, Mt.
Mexic. 28. 1893. Type: MEXICO: Habitat in Ovando, 16 Dec 1936, E. Matuda 574
Cordovae pratis. Floret Julio, Sesse & Mocino (Holotype F 916483 (photo PTBG); Isotypes
s.n. (presumably MA, n.v.).—Status MEXU, US).
uncertain. Omiltemia floribunda (Standi.) J. H. Kirkbr.,
Oldenlandia repens L., Mant. 40. 1767. Type: Syst. Bot. 9: 413. 1984. Edithea floribunda
INDIA: Without precise locality or collector Standi., Trop. Woods. 34: 2. 1933. Not
(Lectotype LINN 155.2, n.v.; designated by Deppea floribunda Hemsl., 1879. Type:
Verdcourt, 1983).—Dentella repens (L.) J. R. MEXICO: Guerrero, Malinaltepec, Mar 1930,
& G. Forst. L. Schultze 363 (Holotype B, destroyed
Oldenlandia rubra (Cav.) A. Gray, PI. Wright. (fragment F)).—Deppea schultzei Lorence
2: 68. 1853. Houstonia rubra Cav., Icon. 5: Omiltemia longipes Standi., J. Wash. Acad.
48, t. 474, fig. 1. 1799. Type: MEXICO: Sci. 8: 427. 1918. Type: MEXICO: Guerrero,
Hidalgo, Ixmiquilpan, Nee s.n. (Presumably Omilteme, May 1903, E. W. Nelson 7054
MA, n.v.).—Hedyotis rubra (Cav.) A. Gray (Holotype US 1084692 (photos MEXU,
Oldenlandia xestosperma B. L. Rob. & PTBG); Isotypes GH, n.v., US).
Greenm., Proc. Amer. Acad. Arts 32: 41. Omiltemia scoti J. H. Kirkbr., Syst. Bot. 9: 411.
1896. Type: MEXICO: Oaxaca, open glades, 1984. Type: MEXICO: Puebla, above
Sierra de San Felipe, 8000 ft., Jun 1894, C G. Teotitlan del Camino on road to Huautla de
Pringle 4692 (Lectotype GH, n.v.; Jimenez, 2000-3250 m, 3 Aug 1961, C. E.
Isolectotypes BM, n.v., F, K, n.v., MO Smith, Jr., F. A. Peterson & N. Tejada 4137
(photos MEXU, PTBG), MSC, n.v., PH, n.v., (Holotype US 2645608; Isotype GH (photo
US (2 sheets; photos MEXU, PTBG), VT, PTBG)).—Deppea scoti (J. H. Kirkbr.)
n.v.; designated by Terrell, 1980). Hedyotis,
Houstonia.—Bouvardia xestosperma (B. L.
Rob. & Greenm.) Terrell & S. D. Koch Ophiorhiza L., Gen., ed. 5, 74. 1754.
(=Ophiorrhiza L., 1753.)
Oldenlandiopsis Terrell & W. H. Lewis, Ophiorhiza lanceolata Forssk., F. Aegypt.-Arab.
Brittonia42: 185. 1990. Type: 42. 1775. Type: YEMEN: Hadie Mts.,
Oldenlandiopsis callitrichoides (Gnseb.) Forsskals.n. (Holotype C, n.v.; Isotype BM,
Terell & W. H. Lewis. [Herein treated as a n.v.).—Pentas lanceolata (Forssk.) Deflers
synonym of Hedyotis L.)
Oldenlandiopsis callitrichoides (Griseb.) Terrell Ophryococcus Oerst, Vidensk. Meddel. Dansk.
& W. H. Lewis, Brirtonia 42: 185. 1990. Naturhist. Foren. Kjobenhavn 1852: 52. 1853.
Oldenlandia callitrichoides Griseb., Mem. Type: Ophryococcus gesnerioides Oerst.
Amer. Acad. Arts, n.s. 8: 506. 1863. Type: [Herein treated as a synonym of Hoffmannia
CUBA: Ad ripas rivulorum, C Wright 268 Sw.]
(Lectotype GOET, n.v.; designated by Terrell Ophryococcus gesnerioides Oerst., Vidensk.
& Lewis, 1990). Syntype: CUBA: Wright Meddel. Dansk. Naturhist. Foren. Kjobenhavn
1852: 53. 1853. Type: NICARAGUA:
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Segovia, Pantasmo, Jan 1848, Oersted 9332 Haenke s.n. (presumably PR, n.v.; not located
[=Plantae centroamericanae Oersted 9332] by Puff, 1991).
(Holotype C, n.v. (photo F); Isotype F). Paederia pringlei Greenm., Proc. Amer. Acad.
—Hoffmannia gesnerioides (Oerst.) Kuntze Arts 39: 92. 1903. Type: MEXICO: Morelos,
lava fields near Yautepec, 4500 ft., 27 Oct
Oregandra Standi., Publ. Field Columbian Mus., 1902, C G. Pringle 8682 (Holotype GH, n.v.;
Bot. Ser. 4: 265. 1929; Trop. Woods 16: 22. Isotypes MEXU, US (photos MEXU, PTBG)).
1928 (nomen nudum). Type: Oregandra —Paederia ciliata (Bartl. ex DC) Standi.
panamensis Standi. [Herein treated as a
synonym of Chione DC] Pahcourea Aubl., Hist. PI. Guiane 1: 172,/?/. 66.
Oregandra panamensis Standi., Publ. Field Ml5. Type: Palicourea guianensis Aubl.
Columbian Mus., Bot. Ser. 4: 265. 1929. Palicourea adusta Standi., J. Wash. Acad. Sci.
Type: PANAMA: Talamanca Valley, 1927, 18: 279. 1928. Type: COSTA RICA: San
G. P. Cooper & G. M. Slater 144 {Yale Jose, Cerro de las Vueltas, 2700-3000 m, 29
10490) (Holotype F 573817 (photo PTBG); Dec 1925-1 Jan 1926, P. C. Standley & J.
Isotypes US 1316774 (photos MEXU, PTBG), Valeria 43666 (Holotype US 1253350 (photo
US 1316777).—Chione sylvicola (Standi.) PTBG)).
W. C. Burger Palicourea albocaerulea C M. Taylor, Syst.
Bot. Monogr. 26: \5,fig. 3c, d. 1989. Type:
Osa Aiello, J. Arnold Arbor. 60: 115. 1979. COSTA RICA: Puntarenas, Monteverde
Type: Osa pulchra (D. R. Simpson) Aiello Cloud Forest Reserve, Pacific slope of
Osa pulchra (D. R. Simpson) Aiello, J. Arnold Continental Trail, Nuboso Trail, 1540-1620
Arbor. 60: 116. 1979. Hintonia pulchra D. R. m, 16 Sep 1983, Busby 107 (Holotype DUKE,
Simpson, Phytologia 29: 277. 1974. Type: n.v.).
COSTA RICA: Puntarenas, Osa Peninsula, Palicourea angustifolia Kunth in Humb.,
about 5 km W. of Rincon de Osa, W. Burger Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 367.
& R. Liesner 7320 (Holotype F 1729029 1819. Type: VENEZUELA: Amazonas, banks
(photo PTBG); Isotypes MO (photos MEXU, of Orinoco and Rio Negro, Humboldt &
PTBG), US (photos MEXU, PTBG)). Bonplands.n. (Holotype P-Bonpland, n.v.;
Isotype B-WI, n.v. (photo MO)).
Otocalyx Brandegee, Univ. Calif. Publ. Bot. 6: Palicourea austin-smithii Standi., Publ. Field
68. 1914. Type: Otocalyx chiapensis Mus. Nat. Hist., Bot. Ser. 18: 1333. 1938.
Brandegee [Herein treated as a synonym of Type: COSTA RICA: Alajuela, Canton Alfaro
Rondeletia L.] Ruiz, Palmira, 2300 m, 4 Aug 1938, A. Smith
Otocalyx chiapensis Brandegee, Univ. Calif. HI034 (Holotype F 942192 (photo PTBG);
Publ. Bot. 6: 69. 1914. Not Rondeletia Isotypes F, GH, n.v., MO (photos MEXU,
chiapasensis Standi., 1940. Type: MEXICO: PTBG), US (photo PTBG)).—Palicourea
Chiapas, Cerro del Boqueron, 1913, C. A. salicifolia Standi.
Purpus 7041 (Holotype UC 173047 (photos Palicourea bella (Standi.) Dwyer, Ann.
MEXU, PTBG); Isotype GH, n.v.). Missouri Bot. Gard. 67: 299. 1980. Psychotria
—Rondeletia h bella Standi., J. Wash. Acad. Sci. 18: 185.
1928. Type: PANAMA: Chiriqui, between
Paederia L., Syst. Nat., ed. 12, 2: 189. 1767; Alto de Las Palmas and Cerro de la Horqueta,
Mant. PI. 52. 1767. Type: Paederia foetida L. 2100-2268 m, 18 Mar 1911, H Pittier 3250
Paederia ciliata (Bartl. ex DC.) Standi., Publ. (Holotype US 677646 (photo PTBG); Isotype
Field Mus. Nat. Hist., Bot. Ser. 22: 386. 1940. F (photos MEXU, PTBG)).
Lygodysodea ciliata Bartl. ex DC, Prodr. 4: Palicourea bellula C M. Taylor, Syst. Bot.
470. 1830. Type: MEXICO: Ad Acahuitzola, Monogr. 26: 24, fig. 9, a, b, d. 1989. Type:
COSTA RICA: Heredia, Rio Vueltas, upper designated by Standley, 1938).—Palicourea
Rio Patria, eastern slope of Volcan Barba at padifolia (Willd. ex Roem. & Schult.) C. M.
the Continental Divide, ca. 2000 m, 27 Jan Taylor & Lorence
1970, W. C. Burger & R. L. Liesner 7514 Palicourea crocea (Sw.) Roem. & Schult., Syst.
(Holotype F 1833197 (photo PTBG); Isotype Veg. 5: 193. 1819. Psychotria crocea Sw.,
MO (photo PTBG)). Prodr. Veg. Ind. Occ. 44: 1788. Type:
Palicourea brenesii Standi., Publ. Field Mus. JAMAICA: Without precise locality, Swartz
Nat. Hist., Bot. Ser. 18: 1333. 1938. Type: s.n. (Holotype BM, n.v.).
COSTA RICA: Palmira del Naranjo, in forest, Palicourea dimorphandrioides (Dwyer) C. M.
1850 m, Apr 1921, Brenes 3512 (Holotype F Taylor, Novon 6: 211. 1996. Cephaelis
851630 (photo PTBG); Isotype GH, n.v.). dimorphandrioides Dwyer, Ann. Missouri
Palicourea caerulescens Suessenguth, Bot. Bot. Gard. 67: 66. 1980. Type: PANAMA:
Jahrb. Syst. 72: 286. 1942. Type: COSTA Veraguas, valley of Rio Dos Bocas on road
RICA: San Jose-Limon border region, between Alto Piedras (above Santa Fe) and
Chirripo Grande, 3000 m, 30 Apr 1932, Calovebora, 350-400 m, 19 Aug 1974, T. B.
Kupper 1200 (Holotype M, n.v.). Croat 27470 (Holotype MO 2201358 (photos
—Palicourea salicifolia Standi. MEXU, PTBG)).
Palicourea carpirostrata Dwyer, Ann. Missouri Palicourea discolor K. Krause, Bot. Jahrb.
Bot. Gard. 67: 299. 1980. Type: PANAMA: Engl. 54, Beibl. 119: 40. 1916. Type: COSTA
Veraguas, NW of Santa Fe, 2 km from RICA: Canas Gordas, 1100 m, 18 Feb 1897,
Escuela Agricola Alto de Piedra. Cloud forest H. Pittier 11161 (Holotype B, presumably
on ridge top below summit of Cerro Tute, 28 destroyed (photos F, US), fragment F; Isotype
Mar 1975, S. Mori & J. Kallunki 5262 US (photo PTBG)).
(Holotype MO 2385000 (photo PTBG); Palicourea domingensis (Jacq.) DC, Prodr. 4:
Isotype MO 2384999 (photo PTBG)). 529. 1830. Psychotria domingensis Jacq.,
—Palicourea ochnoides Dwyer Enum. PI. Carib. 16. 1760. Type: SANTO
Palicourea chiriquina Standi., Contr. U.S. Natl. DOMINGO: Without precise locality, Jacquin
Herb. 18: 140. 1916. Type: PANAMA: (location unknown).—Psychotria
Chiriqui, Cuesta de las Palmas, southern slope domingensis Jacq.
of Cerro de la Horqueta, 1700-2100 m, 17-19 Palicourea galeottiana M. Martens, Bull. Acad.
Mar 1911, H. Pittier 3211 (Holotype US Roy. Sci. Bruxelles 11: 136. 1844. Type:
677607 (photo PTBG); Isotype US (photo MEXICO: Oaxaca, H. Galeotti 2602
PTBG)). (Lectotype BR (photos MEXU, PTBG);
Palicourea copensis (Dwyer) C. M. Taylor in Isolectotypes K, n.v., P (photos MEXU,
W. C. Burger & C. M. Taylor, Fieldiana, Bot., PTBG); designated by Taylor & Lorence,
n.s. 33: 204. 1993. Psychotria copensis 1985: 669).—Psychotria galeottiana (M.
Dwyer, Ann. Missouri Bot. Gard. 67: 365. Martens) C. M. Taylor & Lorence
1980. Type: PANAMA: Code, El Cope, El Palicourea garciae Standi., Publ. Field Mus.
Potroso, Atlantic slope of Alto Calvario, 26 Nat. Hist., Bot. Ser. 22: 195. 1940. Type:
Feb 1977, J. P. Folsom & R. Lantz 1880 COLOMBIA: El Valle, San Antonio, W of
(Holotype MO 2605731 (photos MEXU, Cali, near summit of Cordillera Occidental,
PTBG)). 1900-2350 m, Feb-Mar 1939, Killip &
Palicourea costaricensis Benth. in Oerst., Garcia 33660 (Holotype F; Isotypes A, n.v.,
Vidensk. Medd. Dansk. Naturhist. Foren. NY, n.v.).
Kjobenhavn 1852: 38. 1853. Type: COSTA Palicourea gardenioides (Scheidw.) Benth. &
RICA: Heredia, Volcan de Barba et Cerros de Hook. f. ex Hemsl., Biol. Centr.-Amer., Bot.
Candelaria, Oersted 11540 (Lectotype C, n.v. 2: 52. 1881. Rhodostoma gardenioides
(photo F); Isolectotypes MICH, n.v., US, n.v.; Schiedw. in Otto & Dietr., Allgem. Gartenz.
^ NOMENCLATOR OF MEXICAN AND CENTRAL J
10: 286. 1842. Type: Based on cultivated Palicourea lanceolata Oerst. ex Pol., Vidensk.
material of unknown origin, presumably from Meddel. Dansk Narurhist. Foren. Kjobenhavn
Veracruz, Mexico; type not designated. 1852: 39. 1853. Type: COSTA RICA: San
—Psychotria gardenioides (Scheidw.) Jose, in monte Candelaria, 2100 m, 1845-48,
Standi. Oersted 11556 (Lectotype C, n.v. (photo &
Palicourea gibbosa Dwyer, Ann. Missouri Bot. fragment F); designated by Standley (1938);
Gard. 67: 304. 1980. Type: PANAMA: see Uragoga carpinterensis O. Kuntze).
Darien, Cerro Tacaracuna, West Ridge trail —Palicourea angustifolia Kunth in Humb.,
from summit camp to waterfall, E of camp, Bonpl. & Kunth
1550-1700 m, 2 Feb 1975, A. Gentry & S. Palicourea lancifera Standi. & L. O. Williams,
Mori 14108 (Holotype MO 2354276; Isotype Ceiba 1: 249. 1951. Type: COSTA RICA:
MO 2354275). Alajuela, Canton Alfaro Ruiz, Zarcero, 1500
Palicourea gomezii C. M. Taylor, Ann. m, 17 Jun 1941, A. Smith 2794 (Holotype US
Missouri Bot. Gard. 77: 215. 1990. Type: 2215921 ex EAP (photo PTBG); Isotype F).
COSTA RICA: Heredia, quebrada Palicourea lasiorrhachis Oerst., Vidensk.
Sangrijuela, 700 m, L. D. Gomez P., Chacon Meddel. Dansk Narurhist. Foren. Kjobenhavn
& Herrera 20896 (Holotype CR, n.v.; Isotype 1852: 39. 1853. Type: COSTA RICA: In
MO, n.v.). monte Candelaria, Feb 1847, Oersted 11557
Palicourea grandibracteata Dwyer, Ann. (Holotype C, n.v. (photos F, NY); Isotype US
Missouri Bot. Gard. 67: 305. 1980. Type: (photo PTBG)).
PANAMA: Veraguas, NW of Santa Fe, 2 km Palicourea leucantha Donn. Sm., Bot. Gaz.
from Escuela Agricola Alto de Piedra, cloud (Crawfordsville) 48: 295. 1909. Type:
forest on ridge top below summit of Cerro GUATEMALA: Alta Verapaz, between
Tute, 28 Mar 1975, S. Mori & J. Kallunki Coban and Tactic, 1800 m, Mar 1903, H. von
5262 (Holotype MO 2384999 (photo PTBG); Tuerckheim 8400 (Holotype US 943517
Isotype MO 2385000 (photo PTBG)). (photo PTBG); Isotype US (photo PTBG)).
—Palicourea ochnoides Dwyer Palicourea longibracteata Bartl. ex DC, Prodr.
Palicourea guianensis Aubl., Hist. PI. Guiane 4: 527. 1830. Type: PANAMA: Without
1: 173, t. 66. 1775. Type: FRENCH precise locality, Haenke s.n. (Holotype
GUIANA: In Guajanae sylvis de Caux, Aublet presumably PR, n.v.).—Palicourea triphylla
s.n. (Holotype BM, n.v.). DC.
Palicourea hammelii C. M. Taylor, Syst. Bot. Palicourea lusinaturalis Dwyer, Ann. Missouri
Monogr. 26: 43, fig. 22 c, d. 1989. Type: Bot. Gard. 53: 106. 1966. Type: PANAMA:
PANAMA: Near top of Continental Divide Colon, NW part of Canal Zone, area W of
ridge between Chiriqui and Bocas del Toro Limon Bay, Gatun Locks and Gatun Lake,
provinces near Cerro Pate Macho NE of Rio Pina-Rio Media divide, Johnston 1807
Boquete, 7100 ft., 15 Mar 1982, B. Hammel, (Holotype MO, not located 6/96).
W. J. Kress, S. Knapp, Trainer & Will 11408 —Psychotria luxurians Rusby
(Holotype DUKE, n.v.; Isotype MO (photo Palicourea macrantha Loes., Verh. Bot.
PTBG)). Vereins Brandenburg 65: 113. 1923. Type:
Palicourea intermedia Oerst. ex Pol., Linnaea GUATEMALA: Huehuetenango, Nenton
41: 571. 1877. Type: COSTA RICA: Forests District, near Yalambohoch, Aug 1896, E.
of La Carpintera, Polakowsky 143 (Holotype Seler & C. Seler 2871 (Holotype B,
B, presumably destroyed (photo F); Isotype presumably destroyed (photos F, MO, NY,
W, n.v. (fragment F)).—Palicourea US); fragment F).
angustifolia Kunth in Humb., Bonpl. & ocalyx Standi., J. Wash. Acad.
Kunth : 278. 1928. Type: COSTA RICA:
a, Cerro de Las Lajas, north of San
Isidro, 2000-2200 m, 7 Mar 1926, P. C. Palicourea ochnoides Dwyer, Ann. Missouri
Standley & J. Valerio 51611 (Holotype US Bot. Gard. 67: 310. 1980. Type: PANAMA:
1306801 (photo PTBG)). Veraguas, trail up E side of Cerro Tute, to
Palicourea macrosepala K. Krause, Bot. Jahrb. 1200 m, 25 Oct 1975, J. T. Witherspoon, F.
Syst. 54, Beibl. 119: 41. 1916. Type: COSTA Witherspoon & R. Dressier 8872 (Holotype
RICA: Forets de Alto del Pito (Ojo de Agua), MO 2302202 (photo PTBG)).
pres de San Marcos, 1000 m, Mar 1893, A. Palicourea orosiana C. M. Taylor, Syst. Bot.
Tonduz 7704 (Holotype B, presumably Monogr. 26: 6\,fig. 33 a, b. 1989. Type:
destroyed (photos F, MO, US; fragment F); COSTA RICA: Cartago, about 10 km south of
Isotype BR, n.v.).—Palicourea discolor K. Tapanti along new road on east slope above
the Rio Grande de Orosi, 1600 m, 14 & 17 Jul
Palicourea mexicana Benth. in Oerst., Vid. 1971, W. C Burger & M. Burger 7574
Meddel. Dansk Naturhist. Foren. Kjobenhavn (Holotype F 1940946 (photo PTBG); Isotype
1852: 37. 1853. Type: COSTA RICA: In MO (photo PTBG)).
monte Barba, Oersted 11559 (Lectotype C, Palicourea padifolia (Willd. ex Roem. &
n.v.; designated by Standley (1938); see Schult.) C. M. Taylor & Lorence, Taxon 34:
Uragoga cordillerae O. Kuntze). 669. 1985. Psychotria padifolia Willd. ex
—Palicourea padifolia (Willd. ex Roem. & Roem. & Schult., Syst. Veg. 5: 189. 1819.
Schult.) C. M. Taylor & Lorence Type: MEXICO: Veracruz, Xalapa, Humboldt
Palicourea molliramis K. Schum. & K. Krause, & Bonpland 4420 (Holotype B-WI (photos
Engl. Bot. Jahrb. 40: 337. 1908. Type: DUKE, F, MO, NY)).
COLOMBIA: Cauca, prope La Junta ad Rio Palicoureapanamensis Standi., Ann. Missouri
d'Agua, 200-500 m, Lehmann 4667 Bot. Gard. 25: 839. 1938. Type: PANAMA:
(Holotype B, presumably destroyed). Chiriqui, Valley of upper Rio Chiriqui Viejo,
/tycAo/r/a.—Psychotria hebeclada DC. 1300-1900 m, Jul-Aug 1937, G. White & P.
Palicourea montensis Dwyer, Ann. Missouri White 14 (Holotype F 913062 (photo PTBG);
Bot. Gard. 67: 309. 1980. Type: PANAMA: Isotypes A, n.v., MO (photo PTBG)).
Chiriqui, Bocas del Toro-Chiriqui border —Palicourea discolor K. Krause
region, on Cerro Colorado copper mine road, Palicourea parvijlora Benth., Bot. Voy. Sulphur
5000-5500 ft., 4 May 1975, S. Mori & J. 107. 1845. Type: PANAMA: Island off the
Kallunki 5880 (Holotype MO).—Psychotria coast of Veraguas, Barclay & Hinds s.n.
croceovenosa Dwyer (which is based on the (Holotype BM, n.v.).—Palicourea triphylla
DC.
placement for this species.) Palicourea pauciflora Standi., J. Wash. Acad.
Palicourea montivaga Standi., J. Wash. Acad. Sci. 18: 278. 1928. Type: PANAMA:
Sci. 18: 279. 1928. Type: COSTA RICA: Chiriqui, between Alto de las Palmas and top
Heredia, Yerba Buena, northeast of San of Cerro de la Horqueta, 2100-2265 m, 18
Isidro, 2000 m, 28 Feb 1926, P. C. Standley & Mar 1911, H Pittier 3237 (Holotype US
J. Valerio 49850 (Holotype US 1306225 677618 (photo PTBG); Isoytpe F (photo
(photo PTBG, fragment F)). PTBG)).
Palicourea nigrescens M. Martens 8c Galeotti, Palicourea pendula C. M. Taylor, Syst. Bot.
Bull. Acad. Roy. Soc. Sci. Bruxelles 11: 136. Monogr. 26: 68, fig. 37 a-d. 1989. Type:
1844. Not Psychotria nigrescens De Wild., PANAMA: Veraguas, ridge of Cordillera de
1924. Type: MEXICO: Veracruz, Cordillera, Tute, trail to Cerro Tute above Escuela
Xalapa, 4000 pieds, Jun-Oct 1840, H. Agricola Alto de Piedra, just W of Santa Fe,
Galeotti 2653 (Holotype BR, n.v. (photo F); 800-1350 m, 5 Jun 1982, S. Knapp & R.
Isotype P (photos MEXU, PTBG)). Dressier 5409 (Holotype MO 3041797 (photo
—Psychotria mexiae Standi. PTBG)).
t NOMENCLATOR OF MEXICAN AND CENTRAL
Palicourea phanaerandra Standi. & Steyerm., Palicourea skotakii C. M. Taylor, Syst. Bot.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 252. Monogr. 26: 76, fig. 41. 1989. Type: COSTA
1947. Type: GUATEMALA: Izabal, Cerro RICA: San Jose, along a road to the S from
San Gil, uppermost ridges and summit, the Panamerican Highway at km 67, about 1
1200-1300 m, 26-27 Dec \9A\,J.A. km along the road, 2700-2800 m, 5 Mar
Steyermark 41952 (Holotype F 1132612 1985, C. M. Taylor & C. Skotak 4756
(photo PTBG); Isotype US).—Psychotria (Holotype DUKE, n.v.; Isotypes F (photo
phanaerandra (Standi. & Steyerm.) Lorence PTBG), MO (photo PTBG)).
Palicourea purpurea C. M. Taylor, Syst. Bot. Palicourea spathacea C. M. Taylor, Syst. Bot.
Monogr. 26: 71, fig. 33 c, d. 1989. Type: 9: 226. 1984. Type: COSTA RICA: Cartago,
PANAMA: Chiriqui, NW of Boquete along along the trail about 6 km beyond the gate to
road to Palo Alto Arriba, from 2 km until road the meteorology station in Alto Patillos,
ends then another 0.5 km by trail, 1400-1600 Tapanti Hydroelectric Project, 7 Apr 1976, J.
m, 13 May 1984, C M. Taylor 3201 Vtley & K. Utley 4538 (Holotype DUKE, n.v.;
(Holotype DUKE, n.v.; Isotypes F (photo Isotypes CAS, n.v., CR, n.v., F (photo PTBG),
PTBG), MO (photo PTBG)). US (photo PTBG)).
Palicouraea purpusii Standi., Publ. Field Palicourea standleyana C. M. Taylor, Syst.
Columbian Mus., Bot. Ser. 8: 67. 1930. Type: Bot. Monogr. 26: 81, fig. 45 d-f. 1989. Norn,
MEXICO: Veracruz, Zacuapam [Zacuapan], nov. for Psychotria brenesii Standi., Publ.
shady forest, May 1928, C. A. Purpus 10110 Field Mus. Nat. Hist., Bot. Ser. 18: 1347.
(Holotype F 602907 (photos MEXU, PTBG); 1938. Not Palicourea brenesii Standi., 1938.
Isotype MO (photos MEXU, PTBG)). Type: COSTA RICA: Alajuela, Los Angeles
—Palicourea macrantha Loes. de San Ramon (Cerros de La Balsa), Mar
Palicourea rigidifolia (Dwyer & Hayden) 1928, Brenes 6050 (Holotype F 854521
Dwyer, Ann. Missouri Bot. Gard. 67: 313. (photos MEXU, PTBG)).
1980. Cephaelis rigidifolia Dwyer & Hayden, Palicourea stevensonii Standi., Trop. Woods.
Ann. Missouri Bot. Gard. 55: 37. 1968. Type: 16: 42. 1928. Type: BRITISH HONDURAS
PANAMA: Panama, Cerro Jefe, 2700 ft., 20 (BELIZE): Middlesex, Jul 1927, N. S.
Aug 1967, J. D. Dwyer 8075A (Holotype MO Stevenson C(Yale 10683) (Holotype F
2165097 (photo PTBG)).—Status uncertain, 572673; Isotype NY, n.v.).—Psychotria
probably Psychotria subg. Heteropsychotria capitata Ruiz & Pav.
Palicourea salicifolia Standi., J. Wash. Acad. Palicourea subrubra Pol., Linnaea 41: 571.
Sci. 18: 280. 1928. Type: COSTA RICA: San 1877. Type: COSTA RICA: San Jose, Cerro
Jose, Laguna de La Chonta, northeast of Santa de La Carpintera, 24 Jun 1875, Polakowsky
Maria de Dota, 2000-2100 m, 18 Dec 1925, 200A (Holotype B, presumably destroyed
P. C. Standley 42174 (Holotype US 1252646 (photo F)).—Palicourea padifolia (Willd. ex
(photo PTBG)). Roem. & Schult.) C. M. Taylor & Lorence
Palicourea seleri Loes., Verh. Bot. Vereins Palicourea talamancana Standi. & L. O.
Prov. Brandenburg 65: 114. 1923. Type: Williams, Ceiba 1: 250. 1951. Type: COSTA
GUATEMALA: Huehuetenango, zwischen RICA: Cartago, Cordillera de Talamanca, near
Trinidad und Rosario, 13 Aug 1896, E. Seler Vuelta Alta, Cerro de las Vueltas, 3200 m, 10
& C. Seler 3049 (Syntype B, destroyed Apr 1949, L. O. Williams 16264 (Holotype
(photos F, GH, MO, NY, US), fragment F, EAP, n.v.; Isotypes F (photo PTBG), GH,
GH, n.v.); Yalambahoch, E. Seler & C. Seler n.v., US (photo PTBG)).—Palicourea
2676 (Syntype B, destroyed).—Psychotria brenesii Standi.
galeottiana (M. Martens) C. M. Taylor & Palicourea tilaranensis C. M. Taylor, Syst.
Bot. Monogr. 26: S4, fig. 3 a, b. 1989. Type:
COSTA RICA: Puntarenas-Alajuela border
region, in and around Monteverde Nature MA, n.v.; probable Isotype F).—Rondeletia
Reserve, mostly on Pacific watershed,
10°18'N, 84°47*N, 1450-1650 m, 29 Jun
1977, R. O. Lawton 1189 (Holotype F Pentagonia Benth., Bot. Voy. Sulphur /. 39.
1885069 (photo PTBG)). 1804; 105. 1845 (nom. cons.). Type:
Palicourea triphylla DC, Prodr. 4: 526. 1830. Pentagonia macrophylla Benth.
Type: FRENCH GUIANA: Guiana Gallica, Pentagonia alba Dwyer, Ann. Missouri Bot.
without precise locality, Patris s.n. (Holotype Gard. 67: 321. 1980. Type: PANAMA:
G-DC, n.v. (photo MO)). Psychotria. Darien, Cerro Pirre, Bristan 1224 (Holotype
Palicourea tubiflora Dwyer, Ann. Missouri MO, not located 6/1996; Isotype NY, n.v.).
Bot. Gard. 67: 316. 1980. Type: PANAMA: Pentagonia alfaroana (Standi.) Standi., J. Wash.
Panama, Cerro Jefe, along trail on ridge Acad. Sci. 17: 171. 1927. Watsonamra
running NE from summit, ca. 1000 m, 11 May alfaroana Standi., J. Wash. Acad. Sci. 15:
1975, S. Mori & J. Kallunki 6082 (Holotype 287. 1925. Type: COSTA RICA: Alajuela,
MO 2271726 (photo PTBG)). vicinity of Capulin, on the Rio Grande de
Palicourea tumidonodosa Dwyer, Ann. Tarcoles, 80 m, 2 Apr 1924, P. C. Standley
Missouri Bot. Gard. 67: 317. 1980. Type: 40194 (Holotype US 1152712 (photos
PANAMA: Panama, Cerro Mali, base camp MEXU, PTBG); Isotype F).—Pentagonia
near Colombian border, lower montane wet tinajita Seem.
forest, 1500 m, 22 Jan 1975, A. Gentry & S. Pentagonia brachyotis (Standi.) Standi., J.
Mori 13787 (Holotype MO 2259310 (photo Wash. Acad. Sci. 17: 171. 1927. Watsonamra
PTBG)). brachyotis Standi., Contr. U.S. Natl. Herb. 17:
Palicourea veraguensis Dwyer, Ann. Missouri 443. 1914. Type: PANAMA: Darien,
Bot. Gard. 67: 318. 1980. Type: PANAMA: Marraganti, 3 Apr 1908, R. S. Williams 999
Chiriqui, Cerro Colorado, 50 km N of San (Holotype US 678351 (photos MEXU,
Felix on the continental divide, 1200-1500 m, PTBG)).
17 Aug 1975, S. Mori & R. Dressier 7796 Pentagonia costaricensis (Standi.) W. C.
(Holotype MO 2777356 (photo PTBG)). Burger & C. M. Taylor, Fieldiana, Bot., n.s.
—Palicourea lasiorrhachis Oerst. 33: 213. 1993. Nothophlebia costaricensis
Palicourea vestita Standi., J. Wash. Acad. Sci. Standi., Contr. U.S. Natl. Herb. 17: 438. 1914.
18: 277. 1928. Type: COSTA RICA: San Type: COSTA RICA: Collines de Moin,
Jose, near Quebradillas, about 7 km north of littoral atlantique, Nov 1899, H. Pittier 16024
Santa Maria de Dota, 1800 m, 24 Dec 1925, (Holotype US 578472 (photos MEXU,
P. C. Standley 42909 (Holotype US 1253015 PTBG)).
(photos F, PTBG)). Pentagonia donnell-smithii (Standi.) Standi., J.
Wash. Acad. Sci. 17: 170. 1927. Watsonamra
Pavetta L., Sp. PL 1: 110. 1753. Type: Pavetta donnell-smithii Standi., Contr. U.S. Natl.
indica L. Herb. 17: 442. 1914. Type: COSTA RICA: La
Pavetta odoratissima Sesse & Moc., Fl. Mexic. Emilia, Llanuras de Santa Clara, 250 m, Apr
20. 1893; La Naturaleza, ser. 2, 2: app. 20. 1896, J. D. Smith 6590 (Holotype US 355176
1893. Type: MEXICO ?: Habitat in Praedio (photos MEXU, PTBG); Isotype F).
Sacarmo de la Langosta, Equitis Gonzalo de Pentagonia gymnopoda (Standi.) Standi., J.
Herrera, floret Agusto, Sesse & Mocino s.n. Wash. Acad. Sci. 17: 171. 1927. Watsonamra
(presumably MA, n.v.).—Status uncertain. gymnopoda Standi., Contr. U.S. Natl. Herb.
Pavetta racemosa Sesse & Mo?., Fl. Mexic. 20. 17: 444. 1914. Type: PANAMA: Colon,
1893; La Naturaleza, ser. 2, 2: app. 2, 18. Loma de la Gloria, near Fato (Nombre de
1893. Type: MEXICO: Without precise Dios), 10-104 m, 8 Jul 1911, H. Pittier 3858
locality, Sesse & Mocino 543 (presumably (Holotype US 678935 (photos MEXU,
L NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
PTBG); Isotype US 678934 (photos MEXU, Pentagonia tinajita Seem., Bot. Voy. Herald
PTBG)). 134, t. 28. 1854. Type: PANAMA: Veraguas,
Pentagonia hirsuta Standi., J. Wash. Acad. Sci. western parts, Seemann 1595 (Holotype BM,
18: 165. 1928. Type: COSTA RICA: n.v. (photo MO)).
Talamanca, forets au-dessus de Tsaki, 500 m, Pentagonia veraguensis Dwyer, Ann. Missouri
Mar 1895, A. Tonduz 9415 (Holotype US Bot. Gard. 67: 328. 1980. Type: PANAMA:
938648 (photos MEXU, PTBG); Isotype F). Veraguas, Valley of Rio Dos Bocas, 11 km
Pentagonia Iobata C. M. Taylor, Novon 5: 202, from Escuela Agricola Alto Piedra (above
fig. ID. 1995. Type: COSTA RICA: Santa Fe) on road to Calovebora, 450 m, 30
Puntarenas. Canton de Golfito, Reserva Aug 1974, T. B. Croat 27504 (Holotype MO
Natural de Vida Silvestre Golfito, final del 2201425 (photo PTBG)).
camino hacia las torres, 8°39*52"N, Pentagonia wendlandii Hook, f, Bot. Mag. pi.
83°10'00"W, 7 Dec 1992, B. Hammel 18619 5230. 1861. Type: CENTRAL AMERICA
(Holotype MO 4355639; Isotypes CR, n.v., F, [probably Costa Rica or Panama]: Described
n.v., MO 4355638).
Pentagonia macrophylla Benth., Bot. Voy.
Sulphur 105, tab. 39. 1845. Type: PANAMA:
Panama, forests between Panama and Cruces, Pentas, Benth. in Hook., Bot. Mag. 4086. 1844.
Barclay s.n. (Holotype BM, n.v.). Type: Pentas carnea Benth.
Watsonamra. Pentas lanceolata (Forssk.) Deflers, Voy.
Pentagonia nuciformis Dwyer, Ann. Missouri Yemen 142. 1889. Ophiorhiza lanceolata
Bot. Gard. 67: 326. 1980. Type: PANAMA: Forssk., F. Aegypt.-Arab. 42. 1775. Type:
Veraguas, 5 miles W of Santa Fe on the road AFRICA: Yemen, Hadie Mts., Forssk&ls.n.
past Escuela Agricola Alto Piedra on Pacific (Holotype C, n.v.; Isotype BM, n.v.). Widely
side of divide, 800-1200 m, 18 Mar 1973, cultivated in Mexico and Mesoamerica.
T. B. Croat 23093 (Holotype MO 2161493
(photo PTBG)). Pentodon Hochst., Flora 27: 552. 1844. Type:
Pentagonia pinnatifida Seem., J. Bot. Hooker Pentodon decumbens Hochst.
7: 566. 1848. Type: PANAMA: Darien, banks Pentodon pentandrus (Schumach. & Thonn.)
of the river Cupica, Seemann 1019 (Holotype Vatke, Osterr. Bot. Zeitschr. 25: 231. 1875.
BM, n.v. (photo MO)). Hedyotispentandra Schumach. & Thonn.,
Pentagonia pittieri (Standi.) Standi., J. Wash. Kongel. Danske Vidensk. Selsk.
Acad. Sci. 17: 170. 1927. Watsonamra pittieri Naturvidensk., Math. Afh. 3: 71. 1827. Type:
Standi., Contr. U.S. Natl. Herb. 17: 443. 1914. GUINEA: Aquapim, "S" [Haaslund Smith?]
Type: PANAMA: San Bias, forests around (location unknown). Naturalized at least in
Puerto Obaldia, San Bias coast, 0-50 m, Aug Mexico (Veracruz), Belize, Nicaragua, and
1911,//. Pittier 4298 (Holotype US 679414 Costa Rica.
(photos MEXU, PTBG); Isotype US 679415
(photos MEXU, PTBG)).—Pentagonia Petesia P. Browne, Civ. Nat. Hist. Jamaica 143,
pinnatifida Seem. t. 2, 3. 1756. Type not designated.
Pentagonia pubescens (Standi.) Standi., J. Petesia grandis Bartl. ex DC, Prodr. 4: 395.
Wash. Acad. Sci. 17: 171. 1927. Watsonamra 1830. Type: MEXICO: Without precise
pubescens Standi., Contr. U.S. Natl. Herb. 17: locality, Haenke s.n. (Holotype presumably
441. 1914. Type: PANAMA: Canal Zone, PR, n.v., fragment G-DC, n.v. (microfiche 4:
around Tabernilla, along the railroad, 20-25 395.1)).—Sommera grandis (Bartl. ex DC.)
m, 6 Jul 1911, //. Pittier 3822 (Holotype US Standi.
678894 (photo PTBG); Isotype NY, n.v.). Petesia minor Griseb., Bonplandia 6: 8. 1858.
—Pentagonia macrophylla Benth. Type: PANAMA: Without precise locality,
. (Holotype presumably BM, Placocarpa Hook. f. in Benth. & Hook, f., Gen.
i donnell-smithii Standi. PI. 2: 107. 1873. Type: Placocarpa mexicana
Petesia repens Sesse & Moc., Fl. Mexic. 21. Hook. f.
1893. Type: MEXICO: Habitat in Ahualuci Placocarpa mexicana Hook. f. in Benth. &
pratis, florit Octobris, Sesse & Mociho s.n. Hook, f., Gen. PI. 2: 108. 1873. Type:
(presumably MA, n.v.).—Status uncertain. MEXICO: Veracruz, Orizaba, Apr 1855,
Petesia stipularis Sesse & M09., Fl. Mexic. 21. Botteri 913 (Lectotype P, (photo PTBG), here
1893, La Naturaleza 2, 2: App. 2, 16. 1893. designated; Isolectotypes F (fragment), K
Not Petesia stipularis L., 1759. Type (listed (photo PTBG). Syntypes: MEXICO:
twice): MEXICO: Habitat in Montibus Veracruz, Orizaba, Botteri 316 (F, K (photo
Mazatlanensibus, floret Maio, habitat in PTBG)); Bourgeau 3022 (K (photo PTBG)).
Acahuitzotlae montibus, floret Julio, Sesse & The sheet of Botteri 913 at P, with duplicates
Mociiio s.n. (presumably MA, n.v.).—Status in two other herbaria and copious flowers and
uncertain, possibly Chomelia or Sommera. fruits, represents the most appropriate choice
Protologue states leaves are prominently for lectotype.
lineolate beneath and inflorescence is once
dichotomous. Plocaniophyllon Brandegee, Univ. Calif. Publ.
Bot. 6: 69. 1914. Type: Plocaniophyllon
Pinarophyllon Brandegee, Univ. Calif. Publ. flavum Brandegee
Bot. 6: 71. 1914. Type: Pinarophyllon flavum Plocaniophyllon flavum Brandegee, Univ.
Brandegee Calif. Publ. Bot. 6: 69. 1914. Type: MEXICO:
Pinarophyllon bullatum Standi., N. Amer. Fl. Chiapas, Finca Mexiquito, Jun 1913, C. A.
32: 87. 1921. Type: GUATEMALA: Alta Purpus 7019 (Holotype UC 173077, n.v.;
Verapaz, between Sepacuite and Secoyocte, Isotypes F (photos MEXU, PTBG), US
1100 m, 24 May 1905, H. Pittier 348 (photos MEXU, PTBG)).
(Holotype US 473013 (photos MEXU, PTBG,
US)). Pogonopus Klotzsch, Monatsber. Konigl.
Pinarophyllon flavum Brandegee, Univ. Calif. Preuss. Acad. Wiss. Berlin 1853: 500. 1853.
Publ. Bot. 6: 71. 1914. Type: MEXICO: Type: Pogonopus ottonis Klotzsch
Chiapas, Finca Mexiquito, Jun 1913, C /f Pogonopus exsertus (Oerst.) Oerst, Amer.
Purpus 6700 (Holotype UC 172447, n.v.; Centr. 17. t. 13. 1863. Macrocnemum
Isotypes F (photo PTBG), GH, n.v., US exsertum Oerst., Vidensk. Meddel. Naturhist.
(photos MEXU, PTBG)). Foren Kjobenhavn 1852: 45. 1853. Type:
COSTA RICA: Jaris, Oersted (Holotype C,
Pittoniotis Griseb., Bonplandia 6: 8. 1858. Type: n.v.). Note: Pogonopus speciosus (Jacq.) K.
Pittoniotis tnchantha Griseb. Schum., a species from northern South
Pittoniotis protracta (Bartl. ex DC.) Griseb., America, was erroneously recorded from
Bonplandia 6: 8. 1858. Guettarda protracta Central America by Standley & Williams
Bartl. ex DC, Prodr. 4: 457. 1830. Type: (1975) and Dwyer (1980).
MEXICO: Without precise locality, Haenke
s.n. (Holotype presumably PR, n.v.). Portlandia P. Browne, Hist. Jamaica 164, t. 11.
Anisomeria.—Chomelia protracta (Bartl. ex 1756. Type: Portlandia grandiflora L.
DC.) Standi. Portlandia ghiesbreghtiana Baill., Adansonia
Pittoniotis trichantha Griseb., Bonplandia 6: 8. 12: 300. 1879. Type: MEXICO: Oaxaca, pres
1858. Type: PANAMA: Without precise de Hacienda de Huijastla au bord des ravine,
locality, Duchassaing s.n. (possibly P, n.v.). 1842-3, Ghiesbreght 27 (Holotype P (photos
Antirhea. MEXU, PTBG); Isotypes F, P (3 sheets;
OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
photos MEXU), US (photos MEXU, PTBG)). 2000-4000 ft., H. Galeotti 1580 (Holotype
—Coutaportla ghiesbreghtiana (Baill.) Urb. BR, n.v.; Isotype P (photos F, MEXU, MO)).
guatemalensis Standi., J. Wash. Posoqueria grandiflora Standi., J. Wash. Acad.
Acad.Sci. 18: 162. 1928. Type: Sci. 18: 166. 1928. Type: COSTA RICA:
GUATEMALA: Alta Verapaz, Quebradas Limon, wet thicket at Finca Hamburg on the
Secas, 2500 ft., 1 Jun 1920, H. Johnson 282 Rio Reventazon below El Cairo, 55 m, 19 Feb
(Holotype US 1081354 (photos MEXU, 1926, P. C. Startdley & J. Valeria 48706
PTBG)).—Coutaportla guatemalensis (Holotype US 1305673 (photos MEXU,
(Standi.) Lorence PTBG)).
Portlandia hexandra Jacq., Enum. PL Carib. 16. Posoqueria latifolia (Rudge) Roem. & Schult.,
1760. Type: COLOMBIA: Bolivar, vicinity of Syst. Veg. 5: 227. 1819. Solena latifolia
Cartagena, Jacquin (location unknown). Rudge, PL Guian. 1: 26. 1805; t. 40. 1806.
—Coutarea hexandra (Jacq.) K. Schum. Type: FRENCH GUIANA: Martin s.n. (BM,
Portlandia lumaeana (Baill.) Baill., Hist. PI. 7:
333. 1880. Coutarea lumaeana Baill., Posoqueria obliquinenia Standi., J. Wash.
Adansonia 12: 301. 1879. Type: Acad. Sci. 18: 167. 1928. Type: COSTA
GUATEMALA: Cote occidentale des Andes, RICA: Forets et bords du Rio Naranjo,
450-500 m, 1878, G. Luma s.n. (Holotype P 200-250 m, Mar 1893, A. Tonduz 7528
(photos MEXU, PTBG)).—Hintonia [published erroneously as 9528} (Holotype
lumaeana (Baill.) Bullock US 764158 (photos MEXU, PTBG)).
Portlandia mexicana (Zucc. & Mart.) Hemsl., —Tocoyena pittieri (Standi.) Standi.
Diagn. PI. Nov. 31. 1879. Coutarea mexicana Posoqueria panamensis (Walp. & Duchass.)
Zucc. & Mart, ex DC, Prodr. 4: 350. 1830. Walp., Ann. Bot. Syst. 2: 797. 1852. Stannia
Type: MEXICO: Without precise locality, panamensis Walp. & Duchass., Linnaea 23:
1829, Martius s.n. (Holotype G-DC, n.v. 755. 1850. Type: PANAMA: Isthmo
(microfiche IDC 800.680. II.7).—Nernstia Panamensi, Duchassaing ? (location
mexicana (DC.) Urb.
Portlandia pterosperma S. Watson, Proc. Amer. Posoqueria pittieri Standi., J. Wash. Acad. Arts
Acad. Arts 24: 52. 1889. Type: MEXICO: Sci. 18: 167. 1928. Type: PANAMA: Canal
Sonora, Guaymas, 1887, E. Palmer 298 Zone, near Hydrographic Station on Trinidad
(Holotype US 47978 (photos MEXU, PTBG); River, 17 May 1914, H. Pittier 6635
Isotype US 47979 (photos MEXU, PTBG)). (Holotype US 716697 (photos MEXU,
Coutarea.—Hintonia latiflora (Sesse & Moc. PTBG); Isotype P, n.v. (fragment F)).
ex DC.) Bullock —Tocoyena pittieri (Standi.) Standi.
Portlandia speciosa (Aubl.) Baill., Hist. PI. 7:
381. 1879. Coutarea speciosa Aubl., Hist. PI.
Guiane 1: 314, t. 122. 1775. Type: FRENCH
GUIANA: Aublet s.n. (presumably P, n.v.).
—Coutarea hexandra (Jacq.) K. Schum. Psychotria acicularis C. M. Taylor in W. C.
Burger & C. M. Taylor, Fieldiana, Bot., n.s.
Posoqueria Aubl., Hist. PL Guiane 1: 133. 1775. 33: 231. 1993. Type: COSTA RICA:
Type: Posoqueria longiflora Aubl. Puntarenas, Rmcon de Osa, region to
Posoqueria coriacea M. Martens & Galeotti, southwest of airstrip, 20-60 m, 20 Jul 1974, J.
Bull. Acad. Roy. Sci. Bruxelles 11: 240. 1844. Utley & K. Utley 1036 (Holotype CR, n.v.;
Type: MEXICO: Oaxaca, Cordillere orientale, Isorypes F, n.v., MO).
forets humides de la Chinantla, pres des Psychotria acuminata Benin., Bot. Voy.
bourgs indiens de Teotalcingo et Choapam, Sulphur 107. 1845. Type: COLOMBIA: Isle
of Gorgona, and Colombian coast, Barclay Psychotria allenii Standi., Ann. Missouri Bot.
s.n. (Holotype BM, n.v.). Gard. 27: 342. 1940. Type: PANAMA: Code,
Psychotria acuminata Benth. subsp. vicinity of El Valle, 600-1000 m, 8 Dec 1938,
boraginoides Dwyer, Ann. Missouri Bot. P. H. Allen 1796 (Holotype F 989006 (photos
Gard. 67: 344. 1980. Type: PANAMA: MEXU, PTBG); Isotypes GH, n.v., NY, n. v.,
Panama, El Llano Carti-Tupile road, 12 mi. US (photos MEXU, PTBG)).
above Panamerican Highway, 26-27 Mar Psychotria altorum Standi. & Steyerm., Publ.
1973, R. L. Liesnerll87 (Holotype MO Field Mus. Nat. Hist., Bot. Ser. 23: 86. 1944.
2164366).—Psychotria acuminata Benth. Type: GUATEMALA: Quezaltenango,
Psychotria aggregata Standi., Contr. U.S. Natl. Montana Chicharro, on lower southeast-facing
Herb. 18: 128. 1916. Type: PANAMA: slopes of Volcan Santa Maria, 2-4 miles south
Clin lqui, between Alto de las Palmas and of Santa Maria de Jesus, 1400-1500 m, 17 Jan
Cerro de la Horqueta, 2100-2268 m, 18 1940, J. A. Steyermark 34302 (Holotype F
March 1911,//. Pittier 3264 (Holotype US 1056806 (photos MEXU, PTBG)).
677655 (photos MEXU, PTBG)). —Psychotria costivenia Griseb. var. altorum
Psychotria aguilarii Standi. & Steyerm., Publ. (Standi. & Steyerm.) C. W. Ham.
Field Mus. Nat. Hist., Bot. Ser. 23: 23. 1943. Psychotria americana Sesse & Moc., Fl. Mexic,
Type: GUATEMALA: Chimaltenango, on ed. 2, 57. 1894. Type: MEXICO: Habitat in
road between Chimaltenango and San Martin montibus calidarum regionum, ut Tenampulco
Jilotepeque, 1500-1700 m, 22 Dec 1940, prope Huastecam et Acauizotla versus
P. C. Standley 80900 (Holotype F 1092416 Acapulcum, Sesse & Mocino s.n. (presumably
(photos MEXU, PTBG)).
Psrchotna alba Ruiz & Pav. var. mexicana Psychotria angustiflora K. Krause, Bot. Jahrb.
Oerst, Vidensk. Meddel. Dansk Naturhist. Syst. 54, Beibl. 119: 43. 1916. Type: COSTA
Foren. Kjobenhavn 1852: 12. 1853. Syntypes: RICA: Sur les collines autour de Las Vueltas,
MEXICO: South Mexico, Jecaltepec, F. Tucurrique, 600-700 m, Jan 1899, A. Tonduz
Liebmann 17 (presumably C, n.v.); Cazadera, 12996 (Holotype B, presumably destroyed
F. Liebmann 18 (presumably C, n.v.); Pital. F (photos F, MO); Isotypes P (photos MEXU,
Liebmann 23 (presumably C, n.v.).—Status PTBG), US (photos MEXU, PTBG)).
Psychotria anomothyrsa K. Schum. & Donn.
Standi., Contr. U.S. Natl. Sm., Bot. Gaz. (Crawfordsville) 35: 3. 1903.
Herb. 18: 131. 1916. Type: PANAMA: Type: GUATEMALA: Alta Verapaz,
Chagres, Isthmus of Panama, Feb-Mar 1850, Cubliquitz, 350 m, Dec 1900, H. von
Fendler 62 (Holotype US 47282 (photos Tuerckheim 7909 (Holotype US 943471
MEXU, PTBG); Isotype MO).—Psychotria (photos MEXU, PTBG); Isotypes GH, n.v.,
capitata Ruiz & Pav. US (photos MEXU, PTBG)).—Psychotria
Psychotria alboviridula K. Krause, Notizbl. macrophylla Ruiz & Pav.
Konigl. Bot. Gard. Berlin 6: 208. 1914. Type: Psychotria arborea Sesse & Moc., Fl. Mexic,
BRAZIL: Im Gebiet des Alto Acre, Seringal, ed. 2, 57. 1894. Type: MEXICO: Habitat in
Sao Francisco, Sep 1911, Ule 9846 (Holotype Ahualuci atque calidis Guatemalae montibus.
B, presumably destroyed (photo F, fragment Floret Maio, Sesse & Mocino s.n. (presumably
F); Isotype US).—Psychotria remota Benth. MA, n.v.).—Status uncertain.
Psychotria alfaroana Standi, J. Wash. Acad. Psychotria aubletiana Steyerm., Mem. New
Sci. 18: 273. 1928. Type: COSTA RICA: York Bot. Gard. 23: 694. 1972. Cephaelis
Guanacaste, El Arenal, 485-600 m, 18-19 Jan axillaris Sw., Prodr. 45. 1788. Not Psychotria
1926, P. C. Standley & J. Valerio 45179 axillaris Willd., 1798, nor Veil., 1827. Type:
(Holotype US 1253966 (photos MEXU, WEST INDIES: St. Christopher, Swartz s.n.
PTBG)). (Holotype presumably S, n.v.).
A NOMENCLATOR OF MEXICAN AND CENTRAL
Maria, 2-4 miles south of Santa Maria de Psychotria deflexa DC, Prodr. 4: 510. 1830.
Jesus, 1400-1500 m, 17 Jan 1940, J. A. Type: FRENCH GUIANA: Without precise
Steyermark 34302 (Holotype F 1056806 locality or collector (Holotype presumably G-
(photos MEXU, PTBG)). DC (n.v.); Isotype G-DEL, n.v. (photo MO)).
Psychotria crebrinervia Standi., Publ. Field Psychotria deneversii C M. Taylor, Novon 6:
Mus. Nat. Hist, Bot. Ser. 4: 343. 1929. Type: 300, fig. 2C, D. 1996. Type: PANAMA:
HONDURAS: Atlantida, Lancetilla valley, Comarca de San Bias, El Llano-Carti road, 18
near Tela, 20-600 m, 6 Dec 1927-20 Mar km from Interamerican Highway, headwaters
1928, P. C. Standley 55286 (Holotype F of Atlantic-draining creeks, 9°19'N, 78°55'W,
584201 (photos MEXU, PTBG)). 300 m, 7 Sep 1984, G. de Nevers 3861
—Psychotria berteriana DC. (Holotype PMA 38224, n.v.; Isotype MO
Psychotria crocea Sw., Prodr. Veg. Ind. Occ. 4658111).
44. 1788. Type: JAMAICA: Without precise Psychotria dichroa (Standi.) C M. Taylor in
locality, Swartz s.n. (Holotype BM, n.v.). W. C. Burger & C M. Taylor, Fieldiana, Bot.,
Uragoga.—Palicourea crocea (Sw.) Roem. n.s. 33: 245. 1993. Evea dichroa Standi.,
& Schult. Contr. U.S. Natl. Herb. 18: 124. 1916. Type:
Psychotria croceovenosa Dwyer, Ann. PANAMA: Chiriqui, humid forest of Cuesta
Missouri Bot. Gard. 67: 367. 1980. Type: de Las Palmas, southern slopes of Cerro de la
PANAMA: Chiriqui, N of San Felix at Horqueta, 1700-2100 m, 17-19 Nov 1911, H.
Chiriqui-Bocas del Toro border on Cerro Pittier 3218 (Holotype US 677615 (photos
Colorado copper mine road along continental MEXU, PTBG); Isotype F). Cephaelis.
divide, 5000-5500 ft., 4 May 1975, S. Mori & Psychotria dispersa Standi, J. Wash. Acad. Sci.
J. Kallunki 5880 (Holotype MO 2579259 18: 184. 1928. Type: COSTA RICA:
(photos MEXU, PTBG)). Based on the same Guanacaste, Los Ayotes, near Tilaran,
type as Palicourea montensis Dwyer, 1980, 600-700 m, 21 Jan 1926, P. C. Standley & J.
which Valerio 45548 (Holotype US 1254170 (photos
MEXU, PTBG)).—Psychotria pittieri Standi.
Psychotria cuneifolia Sesse & Moc,, Fl. Mexic, Psvchotria domingensis Jacq., Enum. PI. Carib.
ed. 2, 58. 1894. Not P. cuneifolia DC, 1830. 16. 1760. Type: SANTO DOMINGO
Type: MEXICO: Habitat in Cordovae vicinis (Hispaniola): Jacquin (no Jacquin material
montibus, floret Augusto, Sesse & Mocino s.n. was located, according to Howard, 1989).
(presumably MA, n.v.).—Status uncertain. Palicourea.
Psychotria cuspidata Bredem. ex Roem. & Psvchotria dosbocensis Dwyer, Ann. Missouri
Schult., Syst. Veg. 5: 192. 1819. Type: Bot. Gard. 67: 370. 1980. Type: PANAMA:
VENEZUELA: Ad Caracas, Bredemeyer s.n. Veraguas, valley of Rio Dos Bocas on road
(Holotype B, presumably destroyed). between Alto Piedra (above Santa Fe) and
—Psychotria acuminata Benth. Calovebora, 350-400 m, 29 Aug 1974, T. B.
Psychotria cyanococca Seem., Fl. Mag. tab. Croat 27427 (Holotype MO 2216296 (photo
479. 1870. Type: NICARAGUA: Chontales, PTBG)).
Seemann s.n. (Holotype presumably BM, Psychotria dressleri (Dwyer) C W. Ham.,
nv.).—Status uncertain. Phytologia 64: 223. 1988. Cephaelis dressleri
Psychotria davidsei Dwyer, Ann. Missouri Bot. Dwyer, Ann. Missouri Bot. Gard. 67: 68.
Gard. 67: 368. 1980. Type: PANAMA: 1980. Type: PANAMA: San Bias, forest
Darien, Cerro Tacarcuna, summit camp along southeast of Puerto Obaldia, 18 Aug 1971,
stream N of camp, 1550-1650 m, A. Gentry & T. B. Croat 16801 (Holotype MO 2138462
S. Mori 14071 (Holotype MO (not located (photo PTBG)).
6/1996)). Psychotria dukei Dwyer, Ann. Missouri Bot.
Gard. 67: 371. 1980. Type: PANAMA:
Darien, Casi-Cana Trail on Cerro Darien, top of W peak of Cerro Tacarcuna
Campamiento, E of Tres Bocas, headwaters of massif, 1800-1850 m, 28 Jan 1975, A. Gentry
Rio Coas, 29 Apr 1968, J. H. Kirkbride, Jr. & & S. Mori 13984 (Holotype MO 2280217
J. A. Duke 1245 (Holotype MO 2162265 (photos MEXU, PTBG)).—Psychotria
(photo PTBG)). goldmanii Standi.
Psychotria durilancifolia Dwyer, Ann. Missouri Psychotria eurycarpa Standi., J. Wash. Acad.
Bot. Gard. 67: 372. 1980. Type: PANAMA: Sci. 18: 275. 1928. Type: COSTA RICA:
Panama, El Llano-Carti Road, Gorgas Lab, Guanacaste, Quebrada Serena, southeast of
Mosquito Project study area, 250 m, 1 Aug Tilaran, 700 m, 27 Jan 1926, P. C. Standley &
1974, T. B. Croat 26028 (Holotype MO J. Valerio 46237 (Holotype US 1254544
2240525 (photo PTBG)).—Psychotria (photos MEXU, PTBG)).
panamensis Standi, var. panamensis Psychotria (?) excelsa Kunth in Humb., Bonpl.
Psychotria dwyeri C. W. Ham., Phytologia 64: & Kunth, Nov. Gen. Sp. 3: 355, tab. 282.
223. 1988. Type: MEXICO: Oaxaca, Distr. de 1819. Type: MEXICO: Veracruz, Xalapa, A.
Choapam [Santiago Choapan], Santa Maria, Bonpland 1445 (Holotype P-Bonpland, n.v.;
Montana Santa Maria, 1500 m, 7 Apr 1938, Y. Isotypes B, n.v. (photo MO), P (photos
Mexia 9265 (Holotype NY, n.v.; Isotypes B, MEXU, PTBG)). Deppea.—Hoffmannia
n.v., F (photo PTBG), GH, n.v., K, n.v., MO excelsa (Kunth) K. Schum.
(photo PTBG), US). Psvchotria faxlucens Lorence & Dwyer, Bol.
Psychotria elata (Sw.) Hammel, Selbyana 12: Soc. Bot. Mexico 47: 50. 1987. Type:
139. 1991. Cephaelis elata Sw., Prodr. 45. MEXICO: Veracruz, Mun. de Catemaco,
1788. Type: JAMAICA: Swartzs.n. Ejido de Coscoapan, 8 km SE de Coyame, 16
(presumably S, n.v.). Abr 1974, F. Ponce & R. Cedillo T. 2
Psychotria elongata Benth. in Oerst., Vidensk. (Holotype MEXU; Isotypes F (photo PTBG),
Meddel. Dansk Naturhist. Foren. Kjobenhavn MO).
1852: 32. 1853. Type: NICARAGUA: Volcan Psychotria fendleri Standi., Contr. U.S. Natl.
de Mombacho, Oersted 11594 (Holotype C Herb. 18: 129. 1916. Type: PANAMA:
(photos MEXU, PTBG), fragment US (photos Chagres, Isthmus of Panama, 25 Feb 1850,
MEXU, PTBG); Isotype K, n.v.). Fendler 59 (Holotype US 48567 (photos
—Psychotria nervosa Sw. var. nervosa MEXU, PTBG); Isotype GH (photos MEXU,
Psychotria emetica L. f., Suppl. PI. 144. 1781. PTBG)).
Type: COLOMBIA: America septentrionalis Psychotria fimbriata Benth., J. Bot. Hook. 3:
provinciae Gironensis, Mutis 1602 (Holotype 226. 1841. Type: GUYANA: Essequibo
LINN, n.v.; probable Isotype US (photo River, Schomburgk 51 (Holotype presumably
PTBG)). Uragoga. K, n.v.). Rudgea.—Rudgea cornifolia
Psychotria erecta (Aubl.) Standi. & Steyerm., (Kunth) Standi.
Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 24. Psychotria flava Oerst. ex Standi., J. Wash.
1943. Ronabea erecta Aubl., Hist. PI. Guian. Acad. Sci. 17: 341. 1927. Type: MEXICO:
1: 156. 1775. Type: FRENCH GUIANA: In Veracruz, Misantla, F. Liebmann 11605
sylvis Oyao, Orapu, Sinemari, Aublets.n. (Holotype C (photos F, MEXU, MO, PTBG)).
(Holotype presumably BM, n.v.). Psychotria fosteri C. W. Ham., Phytologia 64:
Psychotria erythrocarpa Schltdl., Linnaea 9: 224. 1988. Type: PANAMA: Veraguas, Playa
595. 1834. Type: MEXICO: Veracruz, Rosario, northern tip of Coiba Island, 26 Aug
Hacienda de la Laguna, Schiede & Deppe 385 1970, R. Foster 1605 (Holotype DUKE, n.v.;
(Holotype B, presumably destroyed (photo & Isotypes F, GH, MO (photo PTBG)).
fragment F); Isotype K, n.v.). Uragoga. Psychotria fruticetorum Standi., J. Arnold
Psychotria eugenifolia Dwyer, Ann. Missouri Arbor. 11: 42. 1930. Type: HONDURAS:
Bot. Gard. 67: 375. 1980. Type: PANAMA: Comayagua, vicinity of Siguatepeque,
^ NOMENCLATOR OF MEXICAN AND CENTRAL
Psychotria nebulosa (Dwyer) C. M. Taylor, Galeotti 7066 (Holotype BR, n.v.; Isotypes G-
Novon 5: 205. 1995. Coussarea nebulosa DC (photo MO), P (label states "cerca
Dwyer, Ann. Missouri Bot. Gard. 67: 131. Xalapa," photos MEXU, PTBG)).
1980. Type: PANAMA: Chiriqui, Monte Rey —Hoffmannia nicotianifolia (M. Martens &
near Boquete, 20 Jul 1971, T. B. Croat 15868 Galeotti) L. O. Williams
(Holotype MO 2162999 (photo PTBG); / nubiphila Dwyer, Ann. Missouri
Isotype MO 4043108). Bot. Gard. 67: 403. 1980. Type: PANAMA:
Psychotria neillii C. W. Ham. & Dwyer, Chiriqui, NW side of Cerro Pando, T. B.
Phytologia 64: 231. 1988. Type: Croat 15990 (Holotype MO).—Psychotria
NICARAGUA: Rio San Juan, Rio Sabalos, 2 panamensis Standi, var. panamensis
km al W de Santa Eduviges, 80 m, 18 Feb Psvclwtria oaxacam Standi., Contr. U.S. Natl.
1984, P. P. Moreno 23060 (Holotype MO Herb. 23: 1391. 1926. Type: MEXICO:
3192079 (photo PTBG)). Oaxaca, from near Santo Domingo, 1600 ft.,
Psychotria nervosa Sw., Prodr. 43. 1788. Type: 18 Jun 1895, E. W. Nelson 2688 (Holotype
JAMAICA: Without precise locality. Swart: US 574444 (photos MEXU, PTBG)).
s.n. (Holotype S, n.v. (photo A)). —Psychotria gracilifiora Benth. in Oerst.
Psychotria nervosa Sw. var. rufescens (Kunth) Psychotria obovata (Oerst.) Hemsl., Biol.
L. O. Williams, Phytologia 26: 492. 1973. Centr.-Amer., Bot. 2: 50. 1881. Mapouria
Psychotria rufescens Kunth in Humb., Bonpl. obovata Oerst., Amer. Centr. 17, tab. 14, figs.
& Kunth, Nov. Gen. Sp. 3: 364. 1819. Type: 3, 4. 1863. Not Psychotria obovata Ruiz &
VENEZUELA: Caripe, Humboldt & Pav„ 1799. Type: MEXICO: Veracruz, F.
Bonplands.n. (Holotype P-Bonpland, n.v.; Liebmann s.n. (Holotype C, n.v.).
Isotype B, presumably destroyed (photos F, —Psychotria quinqueradiata Pol.
MO))—Psychotria nervosa Sw. Psychotria oerstediana (Kuntze) Standi., Contr.
Psychotria nervosa Sw. subsp. rufescens U.S. Natl. Herb. 23: 1390. 1926. Uragoga
(Kunth) Steyerm., Mem. New York Bot. oerstediana Kuntze, Rev. Gen. PI. 2: 957.
Gard. 23: 480. 1972. Psychotria rufescens 1891. Based on Mapouria obovata Oerst.,
Kunth in Humb., Bonpl. & Kunth, Nov. Gen. Amer. Centr. 17, tab. 14, figs 3, 4. 1863.
Sp. 3: 364. 1819. Type: VENEZUELA: Type: MEXICO: Veracruz, F. Liebmann s.n.
Caripe, Humboldt & Bonplands.n. (Holotype (Holotype C, n.v.).—Psychotria
P-Bonpland, n.v.; Isotype B, presumably quinqueradiata Pol.
destroyed (photos F, MO)).—Psychotria Psvchotria officinalis (Aubl.) Raeusch. ex
nervosa Sw. Sandwith, Kew Bull. 1931: 473. 1931.
Psychotria nicaraguensis Benth. in Oerst., Nonatelea officinalis Aubl., Hist. PI. Guiane
Viddensk. Meddel. Dansk Naturhist. Foren. 1: 182, tab. 70. 1775. Type: FRENCH
Kjobenhavn 1852: 34. 1853. Type: GUIANA: Cayenne, Aublet s.n. (presumably
NICARAGUA: Rio San Juan, Oersted 11634 BM, n.v.).
(Lectotype C, here designated (photos MEXU, Psychotria olgae Dwyer & Hayden, Ann
PTBG)). Syntype: COSTA RICA: Ad Jaris, Missouri Bot. Gard. 55: 42, fig. 3. 1968. Type:
Oersted 11635 (C (photos MEXU, PTBG)). PANAMA: Panama, between Cerro Jefe and
Oersted 11634 is the better choice for Eneida, 2100-2900 ft., 17 Jan 1968, J. D.
lectotype as it is a well developed fruiting Dwvcr. J. Duke & R. Dressier 8193 (Holotype
specimen. Uragoga.—Psychotria marginata MO 1962026 (photos MEXU, PTBG);
Isotypes DUKE, n.v., F, NY, n.v.).
Psychotria nicotianifolia M. Martens & Psvchotria orchidearum Standi.. J. Wash.
Galeotti, Bull. Acad. Roy. Sci. Bruxelles 11: Acad. Sci. 18: 276. 1928. Type: COSTA
229. 1844 [as nicotianaefolia]. Type: RICA: Heredia, Cerros de Zurqui, northeast of
MEXICO: Veracruz, Mirador, 3000 ft., H. San Isidro, 2000-2400 m, 3 Mar 1926, P. C.
Standley & J. Valerio 50863 (Holotype US Villavicencio, Triana s.n. (Holotype BM, n.v.;
1306503 (photos MEXU, PTBG)). Isotype F).
Psychotria orchidearum Standi, subsp. Psychotria pachecoana Standi. & Steyerm.,
persedens Dwyer, Ann. Missouri Bot. Gard. Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 205.
67: 407. 1980. Type: PANAMA: Panama, 1940. Type: GUATEMALA: Baja Verapaz,
Cerro Campana, 2500 ft., 7 Feb 1969, E. L. mountain side of divide north of Santa Rosa,
Tyson 5331 (Holotype MO 1973872). 1650 m, 30 Mar 1939, P. C. Standley 69922
—Status uncertain, probably Psychotria (Holotype F 994220 (photos MEXU, PTBG)).
orchidearum Standi. —Psychotria galeottiana (M. Martens) C. M.
Psychotria oreodoxa L. O. Williams, Taylor & Lorence
Phytologia 28: 230. 1974. Type:
GUATEMALA: Suchitepequez, south-facing Phytologia 64: 232. 1988. Type: PANAMA:
slopes and barrancos of Volcan Santa Clara, Panama, vicinity of Cerro Jefe, Altos de
1 Vz-2 miles west of Finca El Naranjo, 1250 m, Pacora, 2400 ft., 18 Dec 1980, T. Antonio
1 Jun 1942,7. A. Steyermark 46803 (Holotype 3241 (Holotype MO 2892015 (photo PTBG);
F 1133947 (photos MEXU, PTBG)). Isotype ENCB, n.v.).
Psychotria oresbia Dwyer, Ann. Missouri Bot. Psychotria padifolia Willd. ex Roem. & Schult.,
Gard. 67: 408. 1980. Type: PANAMA: Syst. Veg. 5: 189. 1819. Type: MEXICO:
Darien, Cerro Tacarcuna W ridge, trail toward Veracruz, Xalapa, Humboldt & Bonpland
Rio Pucuro just below summit camp, 4420 (Holotype B-WI (photos DUKE, F, MO,
1500-1600 m, 5 Feb 1975, A. Gentry & S. NY)).—Palicourea padifolia (Willd. ex
Mori 14150 (Holotype MO 2280279 (photos Roem. & Schult.) C. M. Taylor & Lorence
MEXU, PTBG)). Psychotria panamensis Standi., Contr. U.S.
Psychotria owgenes L. O. Williams, Phytologia Natl. Herb. 18: 132. 1916. var. panamensis.
28: 231. 1974. Type: GUATEMALA: Baja Type: PANAMA: Chiriqui, humid forest
Verapaz, Sierra de las Minas about 5 km south around Los Siguas Camp, southern slope of
of Purulha, 1600 m, 2 Jan 1973, L. O. Cerro de la Horqueta, 1700 m, 17-19 Mar
Williams. A. Molina R. & T. P. Williams 1911,//. Pittier 3194 (Holotype US 677588
41924 (Holotype F 1722437 (photos MEXU, (photos MEXU, PTBG)).
PTBG)).—Psychotria galeottiana (M. Psychotria panamensis Standi, var.
Martens) C. M. Taylor & Lorence compressicaulis (K. Krause) C. W. Ham.,
Psychotria orosiana Standi., J. Wash. Acad. Phytologia 64: 233. 1988. Psychotria
Sci. 15: 288. 1925. Type: COSTA RICA: compressicaulis K. Krause, Bot. Jahrb. Syst.
Cartago, vicinity of Orosi, 30 Mar 1924, P. C. 54, Beibl. 119: 44. 1916. Type: COSTA
Standley 39803 (Holotype US 1153182 RICA: Cartago, Tuis, 600 m, Jul 1898, H.
(photos MEXU, PTBG)). Pittier 12412 (Holotype B, destroyed (photo
Psychotria ostanrea Dwyer & Hayden, Ann. GH); Isotype US (photos MEXU, PTBG)).
Missouri Bot. Gard. 54: 143. 1967. Type: Psychotria panamensis Standi, var. ixtlanensis
PANAMA: Bocas del Toro, vicinity of C. W. Ham., Phytologia 64: 233. 1988. Type:
Chinqui Lagoon, Water Valley, 1 Nov 1940, MEXICO: Oaxaca, Distr. de Ixtlan, 21.4 km S
H. von Wedel 1461 (Holotype MO 1240228 de Valle Nacional, 1140 m, 28 Nov 1979, T.
(photos MEXU, PTBG); Isotype F). Wendt et al. 2258 (Holotype MEXU 317322;
—Coussarea hondensis (Standi.) C. M. Isotypes CHAPA, ENCB).
Taylor & W. C. Burger Psychotria panamensis Standi, var. magna
Psychotria ostreophora (Wernh.) C. M. (Standi.) C. W. Ham., Phytologia 64: 234.
Taylor, Novon 4: 305. 1994. Cephaelis 1988. Psychotria magna Standi., Contr. U.S.
ostreophora Wernh., J. Bot. 55: 284. 1917. Natl. Herb. 18: 131. 1916. Type: PANAMA:
Type: COLOMBIA: Bogota, Servita y Colon, Loma de la Gloria, near Fato (Nombre
^ NOMENCLATOR OF MEXICAN AND CENTRA!
de Dios), 10-104 m, Aug 1911, H. Pittier 41952 (Holotype F 1132612 (photo PTBG);
4092 (Holotype US 679188 (photos MEXU, Isotype US).
PTBG); Isotype US 693188). Psychotria philacra Dwyer, Ann. Missouri Bot.
Psychotria papantlensis (Oerst.) Hemsl., Biol. Gard. 67: 412. 1980. Type: PANAMA:
Centr.-Amer., Bot. 2: 50. 1881. Mapouria Darien, Cerro Tacarcuna, S slope, ridge top
papantlensis Oerst., Amer. Centr. 17, tab. 14, forest well below summit, 1250-1450 m, 26
fig. 5. 1863. Type: MEXICO: Veracruz, Jan 1975, A. Gentry & S. Mori 13916
Papantla, F. Liebmann s.n. (Holotype (Holotype MO 2280220 (photo PTBG)).
presumably C, n.v.). Uragoga. Psychotria phoeniciana Standi., J. Wash. Acad.
Psychotria parasitica Sw., Prodr. Veg. Ind. Occ. Sci. 16: 17. 1926. Type: MEXICO: Chiapas,
44. 1788, nomen illegit. citing Viscoides mountains near Fenix, May 1925, C. A.
pendulum Jacq. 1763. (see Steyermark, 1972: Purpus 316 (Holotype US 1208247 (photos F,
569 for discussion). Uragoga.—Psychotria MEXU, PTBG)).—Palicourea macrantha
guadalupensis (DC.) Howard Loes.
Psychotria parvifolia Benth. in Oerst., Psychotria pilosa Ruiz & Pav., Fl. Peruv. 2: 60,
Vidensk. Meddel. Naturhist. Foren. pi. 208, fig. 1. 1799. Type: PERU: Ruiz Lopez
Kjobenhavn 1852: 36. 1853. Type: COSTA & Pavon s.n. (Holotype presumably MA, n.v.;
RICA: Ad Naranjo, Apr 1847, Oersted 11637 Isotype B, n.v (photo MO)).
(Holotype C (photos F, MEXU, MO, PTBG); Psychotria pinularis Sesse & Moc., Fl. Mexic,
Isotype US (photos MEXU, PTBG)). ed. 2, 57. 1894. Type: PUERTO RICO:
Psychotria peperomiae Standi., Contr. U.S. Habitat in monticulo de la Aguadilla vicino,
Natl. Herb. 18: 132. 1916. Type: PANAMA: Sesse & Mocifio s.n. (presumably MA, n.v.).
Chiriqui, top of Cerro de la Horqueta, 2265 m, —Psychotria microdon (DC.) Urb.
18 Mar 1911, H. Pittier 3235 (Holotype US Psvchotria pithecobia Standi., Publ. Field
677632, n.v.).—Psychotria guadalupensis Columbian Mus., Bot. Ser. 8: 187. 1930.
(DC.) Howard Type: COSTA RICA: La Palma, 1550 m, 15
Psychotria perparva Dwyer, Ann. Missouri Aug 1898, A. Tonduz 12432 (Holotype B,
Bot. Gard. 67: 411. 1980. Type: PANAMA: presumably destroyed; Isotype US (photos
Panama, Cerro Tacarcuna Massif between MEXU, PTBG)).
summit camp on westernmost ridge and Pico Psvchotria pittieri Standi., Contr. U.S. Natl.
Tacarcuna, mostly elfin forest, 1650-1800 m, Herb. 18: 132. 1916. Type: PANAMA: Canal
22 Jul 1976, A. Gentry, H. Leon & L. Forero Zone, railroad relocation between Gorgona
16921 (Holotype MO 2416828 (photos and Gatun, 10-50 m, 7 Jan 1911, H. Pittier
MEXU, PTBG); Isotype F). 2266 (Holotype US 676523 (photos MEXU,
Psychotria persearum Standi., Ceiba 1: 48. PTBG)).
1950. Type: HONDURAS: Morazan, Cerro de Psychotria pittieri Standi, subsp. oinochrophylla
Uyuca, ca. 1800 m, 2 Mar 1947, P. C. Dwyer, Ann. Missouri Bot. Gard. 67: 416.
Standley 4865 (Holotype F 1256865 (photos 1980. Type: PANAMA: Darien, Cerro Pirre,
MEXU, PTBG); Isotype EAP, n.v.). cloud forest and/or mossy forest, 2500-4000
—Psychotria galeottiana (M. Martens) C. M. ft., 9-10 Aug 1967, J. A. Duke & T. Elias
Taylor & Lorence 13668 (Holotype MO 2138646 (photos
Psychotria phanaerandra (Standi. & Steyerm.) MEXU, PTBG))—Status uncertain,
Lorence, Novon 2: 260. 1992. Palicourea Psvchotria subgen. Heteropsychotria.
phanaerandra Standi. & Steyerm., Publ. Field Psychotria platypoda DC, Prodr. 4: 510. 1830.
Mus. Nat. Hist., Bot. Ser. 23: 252. 1947. Type: GUYANA: Patris s.n. (presumably G-
Type: GUATEMALA: Izabal, Cerro San Gil,
1200 m, 26-27 Dec 1941, J. A. Steyermark
GUATEMALA: Alta Verapaz, Cubliquitz, Psychotria purpureocapitata Dwyer ex C. M.
350 m, Aug 1903, H. von Tuerckheim 8529 Taylor, Novon 6: 301, fig. 2A, B. 1996. Type:
(Lectotype GH, n.v. (fragment F; photos PANAMA: Code, El Cope on Pacific side, lA
MEXU, PTBG); Isolectotype US (photos hour walk from sawmill, 2400 ft. [774 m], 16
MEXU, PTBG); designated by Hamilton, Oct 1979, T. Antonio 2154 (Holotype PMA,
1989). n.v.; Isotype MO 4658112).
Psychotria poeppigiana Mull. Arg., Fl. Bras. Psychotria purpusii Standi., Contr. U.S. Natl.
6(5): 370. 1881. Syntypes: BRAZIL: Herb. 23: 1388. 1926. Type: MEXICO:
Amazonas Habitat in regione superiore Chiapas, Cerro del Boqueron, Aug 1913, C. A.
fluminis Amazonum prope Ega, Poeppig Purpus 7012 (Holotype US 567269 (photos
3065 (presumably M, n.v.), Martius s.n. MEXU, PTBG); Isotypes F (photos MEXU,
(presumably M, n.v.); in sylvis Japurensibus PTBG), GH, n.v., MO (photos MEXU,
ad Maribi, Martius s.n. (presumably M, n.v.); PTBG)).
in regione Rio Negro ad Uanaca et S. Gabriel, Psychotria quadrangulata Dwyer, Ann.
Spruce IS92 (presumably M, n.v.); prope Missouri Bot. Gard. 67: 1980. Type:
Barra do Rio Negro. Martius s.n. (presumably PANAMA: Veraguas, NW. of Santa Fe, 2.7
M, n.v.); in silvis ad Nagueria, Martius s.n. km from Escuela Agricola Alto de Piedras,
(presumably M, n.v.). road to Calovebora, S. Mori & J. Kallunki
Psychotria polyphobia Donn. Sm., Bot. Gaz. 5363 (Holotype MO (not located 6/1996)).
(Crawfordsville) 33: 253. 1902. Type: Psychotria quiinifolia Dwyer, Ann. Missouri
COSTA RICA: Suerre, Llanuras de Santa Bot. Gard. 53: 108. 1966. Type: PANAMA:
Clara, 300 m, Apr 1896, J. D. Smith 6854 Bocas del Toro, Almirante, Changuinola
(Lectotype US 943508 (photos MEXU, Canal, K. Blum 1402 (Holotype MO 2448117
PTBG), here designated; Isolectotype US (photo PTBG)).—Psychotria nervosa Sw.
943509 (photos MEXU, PTBG). Syntype: Psychotria quinqueradiata Pol, Linnaea 41:
COSTA RICA: Talamanca, forets au-dessus 570. 1877. Type: COSTA RICA: San Jose, 24
de Tsaki, 500 m, Apr 1895, A. Tonduz 9577 Jun 1875, H. Polakowski 135 (Lectotype B,
(P (photos MEXU, PTBG)). Smith 6854, with destroyed (photos BH, MO, US); designated
abundant flowers and fruit and a duplicate at by Hamilton, 1989). Uragoga
US, is the better choice for lectotype. Psychotria racemosa Rich., Actes Soc. Hist.
Psychotria psychotriifolia (Seem.) Standi., Nat. Paris 1: 107. 1792. Type: FRENCH
Contr. U.S.Natl. Herb. 18: 133. 1916. GUIANA: L. C. M. Richard s.n. (Neotype P,
Cephaelis psychotriifolia Seem., Bot. Voy. n.v.; designated by Kirkbride, 1997).
Herald 138. 1852 [as psychotriaefolia]. Type: Psychotria racemosa (Aubl.) Raeusch., Nomen.,
PANAMA: Panama, dense forests near ed. 3, 56. 1797, not Rich. 1792. Nonatelia
Cruces; and Darien, Chirambira, Seemann 482 racemosa Aubl., Hist. PI. Guiane 187, t. 72.
(Holotype BM, n.v. (photo MO); Isotype K Ml5. Type: FRENCH GUIANA: In sylvis
[as Seemann 482bis)). territoni Orapu, Aublet s.n. (Holotype P, n.v.).
Psychotria pubescens Sw., Prodr. Veg. Ind. —Psychotria racemosa Rich.
Occ. 44. 1788. Type: JAMAICA: A neotype Psychotria ramonensis Standi., Publ. Field Mus.
needs to be designated according to Howard Nat. Hist., Bot. Ser. 18: 1360. 1938. Type:
(1989). COSTA RICA: La Palma de San Ramon,
Psychotria pumilocarpa Dwyer, Ann. Missouri 1250 m, Nov 1925, A. Brenes 4599 (Holotype
Bot. Gard. 67: 419. 1980. Type: PANAMA: F 852181 (photos MEXU, PTBG)).
Veraguas, Alto de Piedra, 800 m, 7 Sep 1974, —Psychotria hazenii Standi.
P. J. M. Maas & R. L. Dressier 1612 Psychotria remota Benth., J. Bot. Hook. 3:
(Holotype MO 2287628). 225. 1841. Type: BRAZIL: On the Rio Negro,
k. NOMENCLATOR OF MEXICAN AND CENTRAL
Schomburgk 963 (Holotype presumably K, 5000-5500 ft., S. Mori & J. Kallunki 5871
n.v. (photo NY)). Mapouria (Holotype MO 3646500)
Psychotria roseocremea (Dwyer) C. M. Taylor, Psychotria santaritensis Dwyer, Ann. Missouri
Novon 5: 205. 1995. Coussarea roseocremea Bot. Gard. 424. fig. 78. 1980. Type:
Dwyer, Ann. Missouri Bot. Gard. 67: 133. PANAMA: Colon, Santa Rita, near Agua
1980. Type: PANAMA: Panama, El Llano- Clara weather station, 23 Mar 1972, J. D.
Carti highway, 6-10 km N of El Llano, 13 Dwyer & A. Gentry 9565 (Holotype MO
Apr 1973, R. L. Dressier 4334 (Holotype MO 2183828 (photos MEXU, PTBG)).
2126258 (photo PTBG)). Psychotria santarosae Standi., Publ. Field
Psychotria rosulatifolia Dwyer, Ann. Missouri Columbian Mus., Bot. Ser. 7: 113. 1930.
Bot. Gard. 67: 423, fig. 77. 1980. Type: Type: COLOMBIA: El Valle, Santa Rosa,
PANAMA: Darien, Cerro Tacarcuna, trail Dagua Valley, 200-300 m, 22 Sep 1922, E. P.
from Rio Pucuro base camp up west ridge of Killip 11530 (Holotype US 1140714).
Cerro Mali, 640-1000 m, 6 Feb 1975, /i. Psychotria sarapiquensis Standi., Publ. Field
Gentry & S. Mori 14174 (Holotype MO Mus. Nat. Hist., Bot. Ser. 18: 1360. 1938.
2276271 (photos MEXU, PTBG)). Type: COSTA RICA: Vara Blanca de
Psychotria rufescens Kunth in Humb., Bonpl. & Sarapiqui, north slope of Central Cordillera,
Kunth, Nov. Gen. Sp. 3: 364. 1819. Type: 1500 m, Jul-Sep 1937, A. F. Skutch 3330
VENEZUELA: Canpe, Humboldt <£ (Holotype US 1643233 (photos F, MEXU,
Bonpland s.n. (Holotype presumably P- PTBG); Isotypes GH (photo PTBG), MO).
Bonpland, n.v.; Isotype B, presumably Psychotria scabriuscula Bartl. ex DC, Prodr. 4:
destroyed (photos F, MO)). Uragoga. 513. 1830. Type: MEXICO: Guerrero,
—Psychotria nervosa Sw. Acapulco, Haenke s.n. (Holotype presumably
Psychotria rufescens Kunth var. haenkeana PR, n.v.).—Psychotria pubescens Sw.
DC, Prodr. 4: 512. 1930. Type: MEXICO: Psychotria schippii Standi. & Steyerm., Publ.
Without precise locality, Haenke s.n. Field Mus. Nat. Hist., Bot. Ser. 23: 24. 1943.
(Holotype presumably PR, n.v.).—Status Type: BRITISH HONDURAS (BELIZE): El
uncertain, probably Psychotria nervosa Sw. Cayo district, Valentin, in advanced forest,
Psychotria salicifolia (Oerst.) Hemsl., Biol. limestone valley, Jun-Jul 1936, C. L. Lundell
Centr.-Amer., Bot. 2: 51. 1881. Mapouria 6193 (Holotype F 861234 (photos MEXU,
salicifolia Oerst., Amer. Centr. 17, tab. 14, PTBG); Isotypes GH, n.v., NY, n.v., US
fig- 2. 1863. Not Psychotria salicifolia Kunth (photo PTBG)).—Psychotria mexiae Standi.
in Humb., Bonpl. & Kunth, 1819. Type: Psychotria sessilifolia M. Martens & Galeotti,
MEXICO: Oaxaca, Lacoba [Lacova], F. Bull. Acad. Roy. Sci. Bruxelles 11: 228. 1844.
Liebmann (Holotype presumably C, n.v.). Type: MEXICO: Veracruz, Miradoret
—Psychotria papantlensis (Oerst.) Hemsl. Zacuapan [label on isotype states "Xalapa"],
Psychotria saltatrix C. M. Taylor, Novon 5: 3000 ft., H. Galeotti 7078 (Holotype
205, fig. 2. 1995. Type: COSTA RICA: presumably BR, n.v.; Isotypes G-DEL, n.v.
Cartago, Refugio Nacional Tapanti, por la (photo MO), P (photos MEXU, PTBG)).
orilla del Rio Grande de Orosi, 09°45'N, Uragoga.—Psychotria tenuifolia Sw.
83°47'W, 14 Aug 1991, B. Hammel et al. Psychotria siccorubra Dwyer, Ann. Missouri
18328 (Holotype CR, n.v.; Isotypes F, n.v., Bot. Gard. 67: 428. 1980. Type: PANAMA:
MO 4619859). Veraguas, Cerro Tute, ca. 10 km NW of Santa
Psychotria sanfelicensis Dwyer, Ann. Missouri Fe, on ridgetop in cloud forest, above 1000 m,
Bot. Gard. 67: 424. 1980. Type: PANAMA: 19Jun 1975, S. Mori 6781 (Holotype MO
Chiriqui, N of San Felix on Cerro Colorado 2272240 (photos MEXU, PTBG)).
copper mine road, along continental divide, —Psychotria ostreophora (Werhnam) C. M.
Taylor
Psychotria siggersiana Standi., J. Wash. Acad. between Santa Maria de Jesus and
Sci. 15: 289. 1925. Type: COSTA RICA: Calahuache, along great barranco between
Limon, vicinity of Guapiles, 300-500 m, Mar Finca Pireneos and San Juan Patzulin,
1924, P. C. Standley 37210 (Holotype US 1300-1400 m, 6 Jan 1940, J. A. Steyermark
1153058 (photos MEXU, PTBG)). 33700 (Holotype F 1043219 (photos MEXU,
Psychotria simiarum Standi., Publ. Field PTBG)).
Columbian Mus., Bot. Ser. 4: 344. 1929. Psychotria stipulata (Oerst.) Hemsl., Biol.
Type: HONDURAS: Atlantida, Lancetilla Centr.-Amer., Bot. 2: 51. 1881. Mapouria
Valley, near Tela, 20-600 m, 6 Dec 1927-20 stipulata Oerst., Amer. Cent. 17, tab. 14, fig.
Mar 1928, P. C. Standlev 56772 (Holotype F 7. 1863. Type: MEXICO: Veracruz, Papantla,
581799 (photos MEXU, PTBG); Isotype US Liebmann 11615 (Lectotype C, here
(photos MEXU, PTBG)). designated (photos MEXU, PTBG);
Psychotria sixaolensis C. W. Ham., Phytologia Isolectotype C (photos MEXU, PTBG)).
64: 234. 1988. Type: COSTA RICA: Limon, Syntype: MEXICO: Veracruz, Papantla,
along road between Limon and Shiroles, Liebmann 11636 (C (photos MEXU, PTBG)).
along Rio Sixaola, 0.9 miles SW of Bambu, Liebmann 11615, a flowering specimen with a
6.5 miles SW of Bribni, less than 50 m alt., 12 duplicate, is the more suitable choice for
Aug 1977, T. B. Croat 43303 (Holotype MO lectotype.—Psychotria costivenia Griseb.
2587929 (photo PTBG)).
Psychotria skutchii Standi, Field Mus. Nat. Psychotria stockwellii C. W. Ham., Phytologia
Hist., Bot. Ser. 17:283. 1937. Type: 64: 235. 1988. Type: COSTA RICA: Alajuela,
GUATEMALA: San Juan Ixcoy, oak woods, region of Zarcero, 1800 m, 18 Oct 1937, A.
8000 ft., 22 Aug 1924, A. F. Skutch 1074 Smith 543 (Holotype US; Isotype F).
(Holotype F 813469 (photos MEXU, PTBG); Psychotria subsessilis Benth. in Oerst., Vidensk.
Isotype A, n.v.).—Psychotria galeottiana Meddel. Dansk Naturhist. Foren. Kjobenhavn
(M. Martens) C. M. Taylor & Lorence 1852: 32. 1853. Type: COSTA RICA: Ad
Psychotria solitudinum Standi., Publ. Field Turrialba, Oersted 11613 (Holotype C (photos
Mus. Nat. Hist., Bot. Ser. 22: 207. 1940. MEXU, PTBG): Isotype US (photos MEXU,
Type: COSTA RICA: San Jose, Valley of El PTBG)).—Psychotria costivenia Griseb. var.
General, 700 m, Jun 1939, A. F. Skutch 4347
(Holotype F 974021 (photos MEXU, PTBG); Psychotria suerrensis Dorm. Sm., Bot. Gaz.
Isotype US (photos MEXU, PTBG)). (Crawfordsville) 27: 337. 1899. Type:
Psychotria sousae Lorence & Dwyer, Bol. Soc. COSTA RICA: Suerre, Llanuras de Santa
Bot. Mexico 47: 59. 1987. Type: MEXICO: Clara, 300 m, Feb-Apr 1896, J. D. Smith
Oaxaca, Mun. de Matias Romero, zona de 6580 (Syntype US 943504 (photos MEXU,
Uxpanapa, lomas al E de Arroyo Hamaca, 9.5 PTBG), US 943405 (photos MEXU, PTBG));
km al SE de La Floresta, a 21.5 km S de Tonduz 8683 (Syntype US, n.v.).
Esmeralda, 400 m, 22 May 1981, T. Wendt, A. Psychotria sylvivaga Standi., J. Wash. Acad.
Villalobos, I. Navarrete & J. Anguiano 3282 Sci. 18: 274. 1928. Type: COSTA RICA:
(Holotype MEXU; Isotypes CHAPA, PTBG). Heredia, Yerba Buena, northeast of San
Psychotria stenostachya Standi., Publ. Field Isidro, ca. 2000 m, 28 Feb 1926, P. C.
Columbian Mus., Bot. Ser. 8: 207. 1930. Standley & J. Valerio 49989 (Holotype US
Type: PERU: Loreto, Killip & Smith 29294 1306274 (photos MEXU, PTBG)).
(Holotype F, n.v.). Psychotria tacarcunensis Dwyer, Ann.
Psychotria steyermarkii Standi., Publ. Field Missouri Bot. Gard. 67: 431. 1980. Type:
Mus. Nat. Hist., Bot. Ser. 22: 387. 1940. PANAMA: Darien, elfin forest at top of W
Type: GUATEMALA: Quezaltenango, lower peak of Cerro Tacarcuna massif, 1800-1850
south-facing slopes of Volcan Santa Maria, m, 28 Jan 1975, A. Gentry & S. Mori 13974
L NOMENCLATOR OF AND CENTRAL AMERICAN RUBIACE
8: 185. 1930. Kot Psychotria williamsii Randia armata (Sw.) DC, Prodr. 4: 387. 1830.
Standi., 1930. Type: PERU: Loreto, Manfina, subsp. armata. Gardenia armata Sw., Prodr.
on upper Rio Nanay, 29 Jun 1929, L. Williams Fl. Ind. Occ. 51. 1788. Type: COLOMBIA:
1114 (HolotypeF 604625). Cartagena, Jacquin s.n. (Lectotype: Jacquin's
plate; designated by Taylor & Lorence, 1993).
Rachicallis DC, Prodr. 4: 433. 1830. Type: Randia armata (Sw.) DC. subsp. panamensis
Rachicallis mpestris (Sw.) DC. (Standi.) Lorence, Novon 8: 250. 1998.
Rachicallis americana (Jacq.) A. S. Hitchc, Randia panamensis Standi., Publ. Field
Missouri Bot. Gard. Ann. Rep. 4: 92. 1893. Columbian Mus., Bot. Ser. 4: 288. 1929.
Hedyotis americana Jacq., Enum. PI. Carib. Type: PANAMA: Prov. Bocas del Toro,
12. 1760. Type: CUBA: India Occidentalis, region of Almirante, Flat Rock, Jan-Mar
Swartzs.n. (locatic 1928, G. Proctor Cooper 213 (Holotype F
579675 (photos MEXU, PTBG)).
Randia L., Sp. PL 2: 1192. 1753. Type: Randia Randia blepharodes Standi., Contr. U.S. Natl.
mitis L. (=Randia aculeata L.). Herb. 23: 1378. 1926. Type: MEXICO:
Randia aculeata L., Sp. PL 2: 1192. 1753. var. Guerrero, Mexcala, versant oriental, terre
aculeata. Type: JAMAICA: (LINN 214.1, chaude, 500 m, 23 May 1899, E. Langlasse
n.v.; for typification, see Howard, 1989: 452). 1033 (Holotype US 386343 (photos MEXU,
Randia aculeata L. var. chiapasina Loes., PTBG); Isotypes B, presumably destroyed
Verhandl. Bot. Ver. Brandenburg 65: 109. (fragment F), F, GH, n.v., K, P (2 sheets;
1923. Type: MEXICO: Chiapas, Comitan, photos MEXU, PTBG)).^Randia thurberi
near Bahucuc, E. Seler & C. Seler 2203 S. Watson
(Holotype B, presumably destroyed).—Status Randia blepharophylla Standi., Contr. U.S.
, probably Randia aculeata L. var. Natl. Herb. 23: 1375. 1926. Type: MEXICO:
Nayarit, foothills of Sierra Madre, Tepic,
Randia aculeata L. var. dasyclada Steyerm., between Aguacates and Dolores, J. N. Rose
Mem. New York Bot. Gard. 23: 341. 1972. 2028 (Holotype US 300921 (photos MEXU,
Based on Randia obovata Kunth in Humb., PTBG); Isotypes F (photos MEXU, PTBG),
Bonpl. & Kunth, Nov. Gen. Sp. PL 3: 409. GH, n.v.).—Glossostipula blepharophylla
1820. Not Randia obovata Ruiz & Pav., 1799. (Standi.) Lorence
Type: COLOMBIA: Dept. Cordoba, ad ostia Randia brenesii Standi., Publ. Field Mus. Nat.
flumiinis Sinu, A. Bonpland s.n. (Holotype P- Hist., Bot. Ser. 18: 1365. 1938. Type: COSTA
Bonpland, n.v. (photo F)). RICA: La Palma de San Ramon, 10 Mar
Randia albonervia Brandegee, Zoe 5: 257. 1932, Brenes 15048 (Holotype F 857063
1908. Type: MEXICO: Veracruz, vicinity of (photos MEXU, PTBG)).
Zacuapan, Corral de las Piedras, May 1907, Randia calycosa Standi., Contr. U.S. Natl.
C. A. Purpus 2424 (Holotype UC, n.v.; Herb. 20: 201. 1919. Type: PANAMA:
Isotypes BM, F (photos MEXU, PTBG), GH, Chiriqui, Los Siguas Camp, southern slope of
n.v., US (photos MEXU, PTBG)).—Randia Cerro de la Horqueta, 1700 m, 17-19 Mar
monantha Benth. 1911,//. Pittier 3198 (Holotype US 677593
Randia altiscandens (Ducke) C. M. Taylor in (photos MEXU, PTBG); Isotype F (photos
W. C. Burger & C. M. Taylor, Fieldiana, Bot., MEXU, PTBG)).
n.s. 33: 282. 1993. Basanacantha altiscandens Randia canescens Greenm., Proc. Amer. Acad.
Ducke, Archiv. Inst. Biol. Veg. Rio Janeiro 2: Arts 34: 573. 1899. Type: MEXICO: Morelos,
172. 1935. Type: VENEZUELA: Amazonas, barrancas near Cuemavaca, 5000 ft., 28 May
Ducke s.n. (Holotype RB 24415, n.v.; Isotype 1898 C G. Pringle 6863 (Holotype GH, n.v.;
Isotypes F (photos MEXU, PTBG), MEXU (2
sheets), MO (photos MEXU, PTBG), P (2
sheets; photos MEXU, PTBG), US (2 sheets; • i.) K. Schum. in Mart., Fl.
photos MEXU, PTBG)). Bras. 6(6): 342. 1889. Mussaenda formosa
Randia capitata DC, Prodr. 4: 387. 1830. Jacq., Enum. PI Carib. 16. 1760. Type:
Type: MEXICO: Mexico ad Anasteca COLOMBIA: Cartagena, Jacquin s.n.
[Huasteca], May 1827, Berlandier 513 (Holotype G, n.v).—Rosenbergiodendron
(Holotype G-DC, n.v. (photos F, MEXU)). formosum (Jacq.) F. Fagerl
Randia chiapensis Standi, Contr. U.S. Natl. Randia gaumeri Greenm. & C H. Thomps.,
Herb. 23: 1377. 1926. Type: MEXICO: Ann. Missouri Bot. Gard. 1: 410. 1915. Type:
Chiapas, Canjob, 27 Mar 1904, E. A. MEXICO: Yucatan, Izamal, 1895, G. F.
Goldman 797 (Holotype US 470602 (photos Gaumer 589 (Holotype MO 951695 (photos
K, MEXU, PTBG)). MEXU, PTBG); Isotypes F (photos MEXU,
Randia cinerea (Femald) Standi., Contr. U.S. PTBG), MO (photos MEXU, PTBG), US
Natl. Herb. 20: 201. 1919. Genipa cinerea (photos MEXU, PTBG)).—Randia
Femald, Proc. Amer. Acad. Arts. 33: 93. obcordata S. Watson
1897. Type: MEXICO: Guerrero, Acapulco Randia genipifolia (Standi. & Steyerm.)
and vicinity, Oct 1894-Mar 1895, E. Palmer Lorence, Novon 8: 249. 1998. Duroia
348 (Holotype GH, n.v.; Isotype US (photos genipifolia Standi. & Steyerm., Publ. Field
MEXU, PTBG)). Mus. Nat. Hist., Bot. Ser. 22: 186. 1940.
Randia concinna Standi., Publ. Field Columbian Type: GUATEMALA: Izabal, Rio Dulce,
Mus., Bot. Ser. 4: 288. 1929. Type: MEXICO: between Livingston and 6 miles up river, on
Jalisco, San Sebastian, Hacienda del Ocotal, north side (right side going up river), sea
1500 m, Y. Mexia 1690 (Holotype F 579911 level, 14 Apr 1940, J. A. Steyermark 39382
(photos MEXU, PTBG); Isotypes A, n.v., (Holotype F 1035218 (photo PTBG)).
BM, GH, n.v., MO (photos MEXU, PTBG), Randia genipoides Dwyer in W. C Burger &
US (photos MEXU, PTBG)) — Glossostipula C M. Taylor, Fieldiana, Bot., n.s. 33: 284.
concinna (Standi.) Lorence 1993. Type: COSTA RICA: Alajuela, Upala,
Randia cookii Standi., Contr. U.S. Natl. Herb. G. Herrera 1129 (Holotype CR, n.v.; Isotypes
32: 176. 1934. Type: GUATEMALA: Baja F, n.v., MO).
Verapaz, Santa Rosa, June 1904, O. F. Cook Randia gentlei Lundell, Wrightia 4: 125. 1969.
249 (Holotype US 860099 (photos MEXU, Type: BRITISH HONDURAS (BELIZE):
PTBG), fragment F). Toledo Distr., Edwards Rd. beyond Columbia,
Randia crescentioides Standi., Publ. Field Mus. 24 May 1951, P. H. Gentle 7343 (Holotype
Nat. Hist., Bot. Ser. 23: 88. 1944. Type: TEX-LL (photos MEXU, PTBG); Isotypes F,
COSTA RICA: Port Parker, 4 Jul 1932, J. T. TEX-LL (2 sheets), US (photos MEXU,
Howell s.n. (Holotype CAS, n.v., fragment F PTBG)).—Randia genipifolia (Standi. &
(photos MEXU, PTBG)).—Randia thurberi Steyerm.) Lorence
S. Watson Randia gentryi Dwyer, Ann. Missouri Bot.
Randia echinocarpa Sesse & Moc. ex DC, Gard. 67: 447. 1980. Type: PANAMA: Prov.
Prodr. 4: 385. 1830. Type: MEXICO: Fl. Darien, top of Cerro Mali, 1300-1420 m,
Mexic. Ic. 469 (caique presumbably MA, n.v., A. H. Gentry & S. Mori 13629 (Holotype MO
G-DC, n.v. (copies MEXU, US)). 2258315 (photos MEXU, PTBG); Isotype MO
Basanacantha, Genipa. 2578524 (photo PTBG)).
Randia ehrenbergii Standi, Contr. U.S. Natl. Randia grandifolia (Donn. Sm.) Standi, J.
Herb. 32: 167. 1934. Type: MEXICO: Wash. Acad. Sci. 18: 166. 1928.
Veracruz, Plan de Amilpas, Mar 1831, C Basanacantha grandifolia Donn. Sm., Bot.
Ehrenberg 575 (Holotype B, presumably Gaz. (Crawfordsville) 55: 436. 1913. Type:
destroyed).—Randia capitata DC. COSTA RICA: Forets de Santo Domingo de
Golfo Dulce, Feb 1896, A. Tonduz 9878
OF MEXICAN AND CENTRAL
destroyed (photo MO), BM, P [label states armata (Sw.) DC. subsp. panamensis
"Mexique"] photos MEXU, PTBG)). (Standi.) Lorence
Randia montana L. O. Williams, Phytologia 24: Randia pepoformis Dwyer, Ann. Missouri Bot.
160. 1972. Type: HONDURAS, Intibuca, Gard. 67: 451. 1980. Type: PANAMA:
Cordillera Opalaca, between Calaveras and El Darien, trail from Tacarcuna Village on Rio
Duraznillo, 1800 m, 12 Mar 1970, A. Molina Tacarcuna to Cerro Mali, 800-1300 m, 16 Jar
R. & A. R. Molina 25550 (Holotype F 1975, A. H. Gentry & S. Mori 13614
1695297 (photos MEXU, PTBG); Isotype (Holotype MO 2248019 (photos MEXU,
EAP, n.v.).—Randia aculeata L. var. PTBG); Isotypes F (photos MEXU, PTBG),
aculeata MO (photos MEXU, PTBG)).—Randia
Randia nelsonii Greenm., Proc. Amer. Acad. altiscandens (Ducke) C. M. Taylor
Arts 34: 574. 1899. Type: MEXICO: Oaxaca, Randia petenensis Lundell, Wrightia 4: 127.
Juchitan to Chivela, 150-900 ft., 1895, E. W. 1969. Type: GUATEMALA: Peten, La
Nelson 2635 (Holotype GH, n.v.; Isotype US Cumbre, about 5 km from the village, on
569319 (photos MEXU, PTBG)).—Randia Pusila River Road, 17 May 1967, E.
thurberi S. Watson Contreras 6941 (Holotype TEX-LL (photos
Randia nicaraguensis Lorence & Dwyer, MEXU, PTBG); Isotypes F, US (photos
Novon 8: 247, fig. 1. 1998. Type: MEXU, PTBG)).
NICARAGUA: Esteli, 4.9-7.6 km NE of Randia pittieri (Standi.) Standi., Contr. U.S.
Hwy. 1 at Esteli along road to Yali, ca. Natl. Herb. 20: 201. 1919. Basanacantha
13°08-09', 86°19-20'W, 1100 m, 14 Nov pittieri Standi., Contr. U.S. Natl. Herb. 18:
1979, W. D. Stevens & A. Grijalva 15553 134. 1916. Type: COSTA RICA: Gent
(Holotype MO 2872306; Isotypes F, HNMN, Junction, pres Matina, Feb 1900, H. Pither
MEXU, MO). 16036 (Holotype US 578473 (photos K,
Randia oaxacana Standi., Contr. U.S. Natl. MEXU, PTBG); Isotype P (fragment F,
Herb. 32(3): 165. 1934. Type: MEXICO: photos MEXU, PTBG)).
Oaxaca, Choapam [=Santiago Choapan], Randia pleiomeris Standi., Contr. U.S. Natl.
2800-1500 ft., 28-29 Jul 1894, E W. Nelson Herb. 20: 202. 1919. Type: GUATEMALA:
842 (Holotype F 599712 (photos MEXU, Santa Rosa, Santa Rosa, 3000 ft., May 1892,
PTBG)). Heyde & Lux 3166 (Lectotype US 888490,
Randia obcordata S. Watson, Proc. Amer. here designated [left- and right-hand
Acad. Arts 24: 53. 1889. Type: MEXICO: collections only; the center collection is R.
Sonora, Guaymas, 1887, E. Palmer 648 cookii]; Isolectotype GH, n.v.).—Randia
(Holotype US 47988 (photos MEXU, PTBG); thurberi S. Watson
Isotype BM). Randia pringlei (S. Watson) A. Gray, Proc.
Randia obovata Kunth in Humb., Bonpl. & Amer. Acad. Arts 21: 379. 1886.
Kunth, Nov. Gen. Sp. 3: 409. 1820. Not Basanacantha pringlei S. Watson, Proc.
Randia obovata Ruiz & Pav., 1799. Type: Amer. Acad. Arts 18: 98. 1883. Type:
COLOMBIA: Ad ostia fluminis Sinu, MEXICO: Coahuila, Mountains, Jimulco, 9
A. Bonplands.n. (Holotype P-Bonpland, Apr 1885, C. G. Pringle 142 (Holotype GH,
n.v.).—Randia aculeata L. var. dasyclada n.v.; Isotypes A, n.v., BM, F (photos MEXU,
Steyerm. PTBG), MEXU, P (photos MEXU, PTBG),
Randia panamensis Standi., Publ. Field US (photos MEXU, PTBG)).
Columbian Mus., Bot. Ser. 4: 288. 1929. Randia pterocarpa Lorence & Dwyer, Bol.
Type: PANAMA: Prov. Bocas del Toro, Soc. Bot. Mexico 47: 43, fig. 2. 1987. Type:
region of Almirante, Flat Rock, Jan-Mar MEXICO: Veracruz, Mun. San Andres
1928, Q. Proctor Cooper 213 (Holotype F Tuxtla, Estacion de Biologia Tropical Los
579675 (photos MEXU, PTBG)).—Randia Tuxtlas, jardin botanico, 95°04-09'W,
18°34-36*N, 160 m, 6 Dec 1983, G. Ibarra subcordata Standi, Contr. U.S. Natl. Herb.
M. 1109 (Holotype MEXU; Isotypes BM, F, 18: 133. 1916. Type: COSTA RICA: Buissons
MO, XAL). et bois des chemins a Nicoya, May 1900, A.
Randia purpusii Greenm. & C. H. Thomps., Tonduz 13934 (Holotype US 861251 (photos
Ann. Missouri Bot. Gard. 1: 410. 1914. Type: MEXU, PTBG); Isotypes B, destroyed
MEXICO: San Luis Potosi, Minas de San (fragment F, photos MEXU, PTBG), GH, n.v.,
Rafael, May \9\\,CA. Purpus 5208 LE, n.v. (fragment F, photo PTBG), P (photos
(Holotype MO 4186182 (photos MEXU, MEXU, PTBG), fragment F (photos MEXU,
PTBG); Isotypes F, (photos MEXU, PTBG), PTBG)).—Randia monantha Benth.
GH, n.v., MO (photos MEXU, PTBG), US Randia tetracantha (Cav.) DC, Prodr. 4: 387.
(photos MEXU, PTBG)).—Randia capitata 1830. Mussaenda tetracantha Cav., Icon. PI.
DC. 4: 20,/?/. 435. 1799. Type: MEXICO:
Randia retroflexa Lorence & M. Nee, Brittonia [Probably Guerrero], cerca Acapulco, Nee s.n.
39: 371, fig. 1. 1987. Type: MEXICO: (Holotype MA, n.v. (photos F, MEXU, MO)).
Veracruz, Mun. de Hidalgotitlan,
Campamento Hermanos Cedillo a 4 km de Randia tetracantha Sesse & Moc., PI. Nov.
Pancho Villa, 150 m, 25 Mar 1975, B. Ortiz & Hispan. 33. 1888 (La Naruraleza ser. 2, 1:
J. Martiniano A. 56 (Holotype MEXU; app. 33. 1888). Not Randia tetracantha (Cav.)
Isotypes BM, BR, ENCB, F (photo PTBG), DC, 1830. Type: MEXICO: Habitat in
MO (photo PTBG), NY, UC, XAL). montibus Haereditatis-Ahuesuyo, fl. Dec,
Randia rhagocarpa Standi., Contr. U.S. Natl. Sesse & Mocino s.n. (presumably MA,
Herb. 23: 1376. 1926. Type: MEXICO:
Tamaulipas, vicinity of Victoria, 320 m, 1
Feb-9 Apr 1907, E. Palmer 38 (Holotype US
572266 (photos MEXU, PTBG); Isotypes F Yucatan, Xkombec and Uxmal, E. Seler & C
(photos MEXU, PTBG), GH, n.v., MO Seler 4035 (Syntype B, presumably destroyed,
(photos MEXU, PTBG)).—Randia GH, n.v.); E. Seler & C. Seler 5559a (Syntype
obcordata S. Watson B, presumably destroyed).—Randia truncata
Randia rosei Standi., Contr. U.S. Natl. Herb. 20: Greenm. & C H. Thomps.
203. 1919. Type: MEXICO: Sinaloa, Rosario, Randia texensis Lundell, Field & Lab. 13: 22.
7 Jul 1897, J. N. Rose 1551 (Holotype US 1945. Type: U.S.A.: Texas, Cameron County,
300395 (photos MEXU, PTBG); Isotype F about 5 miles north of Brownsville off Point
(photos MEXU, PTBG), GH, n.v.).—Randia Isabel road, 21 Sep 1943, C L. Lundell &
thurberi S. Watson A. A. Lundell 12456 (Holotype SMU,
Randia sonorensis Wiggins, Contr. Dudley n.v.).—Randia obcordata S. Watson
Herb. 3: 75,pi. 19, figs. 4-6. 1940. Type: Randia thurberi S. Watson, Proc. Amer. Acad.
MEXICO: Sonora, granitic hillside 12 miles E Arts 24: 53. 1889. Type: MEXICO: Sonora,
of Ures on road to Babiacora, 2760 ft., 20 Sep Guayamas, 1887, Thurber in E. Palmer 229
1934, /. L. Wiggins 7360 (Holotype DS (Holotype US 1393134 (photos MEXU,
263358, n.v.; Isotype US (photos MEXU, PTBG); Isotypes BM, US).
PTBG)). Randia tomatillo Loes., Rep. Spec. Nov. 18:
Randia standleyana L. O. Williams, Phytologia 360. 1922. Type: MEXICO: Veracruz, Puerto
24: 162. 1972. Type: GUATEMALA: Peten, de Alvarado, and Isla de Sacrificios, E. Seler
La Libertad, 29 May 1933, C. L. Lundell 3474 & C. Seler 4498 (Lectotype US 1205748, here
(Holotype F 692287 (photos MEXU, PTBG); designated; Isolectotypes B, destroyed; GH,
Isotype MICH, n.v.). n.v.). Same locality, E. Seler & C. Seler 5433
Randia subcordata (Standi.) Standi., Contr. U.S. (Syntype B, destroyed). The US collection of
Natl. Herb. 20: 201. 1919. Basanacantha Seler & Seler 4498 is a good fruiting
specimen and the best choice for 1911,//. Pittier 4223 (Holotype MO 1227470
lectotype.—Randia aculeata L. var. (photo PTBG)).—Raritebe palicoureoides
dasyclada Steyerm. Wernham subsp. dwyerianum J. H. Kirkbr.
Randia tomentosa S. Watson, Proc. Amer. Raritebe dariensis (Dwyer) Dwyer, Ann.
Acad. Arts 25: 152. 1890. Not Randia Missouri Bot. Gard. 67: 453. 1980. Dukea
tomentosa Wight & Arn., 1834, nor R. dariensis Dwyer, Ann. Missouri Bot. Gard.
tomentosa Hook., 1880. Type: MEXICO: 53: 365. 1966. Type: PANAMA: Darien,
Nuevo Leon, Sierra de la Silla, near Chepigana District, Cana-Cuasi Trail (Camp
Monterrey, Aug 1889, C. G Pringle 2865 2), 10 Mar 1940, M. E. Terry & R. A. Terry
(Holotype GH, n. v.).—Randia capitata DC. 14871 (Holotype MO 1196543 (photo
Randia truncata Greenm. & C. H. Thomps., PTBG)).—Raritebe palicoureoides
Ann. Missouri Bot. Gard. 1: 411. 1914. Type: Wernham subsp. dwyerianum J. H. Kirkbr.
MEXICO: Yucatan, vicinity of Izamal, 1895, Raritebe euryphyllum (Standi.) Dwyer, Ann.
G F. Gaumer 713 (Holotype MO 46009 Missouri Bot. Gard. 67: 453. 1980. Coussarea
(photos MEXU, PTBG); Isotypes F (photos euryphylla Standi., Publ. Field Columbian
MEXU, PTBG), MO (photos MEXU, PTBG), Mus., Bot. Ser. 4: 294. 1929. Type:
US (photos MEXU, PTBG)). PANAMA: Bocas del Toro, Buena Vista
Randia vazquezii Lorence & Dwyer, Bol. Soc. Camp on Chiriqui Trail, 375 m, 1928, G P.
Bot. Mexico 47: 46,/?g. 3. 1987. Type: Cooper 228 (Holotype F 579282, n.v.).
MEXICO: Veracruz, Mun. de Jesus Carranza, —Raritebe palicoureoides Wernham subsp.
Zona de Uxpanapa, 2 km N del Poblado 2, dwyerianum J. H. Kirkbr.
Ejido F. J. Mina, 17° 16'N, 94°40'W, 120 m, Raritebe palicoureoides Wernham subsp.
23 May 1982, M. Vazquez T. & I. Navarrete dwyerianum J. H. Kirkbr., Brittonia 31: 304.
2520 (Holotype MEXU; Isotypes CHAPA, 1979. Type: PANAMA: Panama, on trail to
PTBG). Cerro Campana, 23 Aug 1967, J. H.
Randia watsonii B. L. Rob., Proc. Amer. Acad. Kirkbride, Jr. & S. M. V. Hay den 273
Arts 29: 317. 1894 [as watsoni}. Based on (Holotype MO 1917794 (photos MO, PTBG);
Randia tomentosa S. Watson, Proc. Amer. Isotypes F, MO (photo PTBG)).
Acad. Arts 25: 152. 1890. Not Randia Raritebe panamensis (Dwyer) Dwyer, Ann.
tomentosa Wight & Arn., 1834, nor R. Missouri Bot. Gard. 67: 454. 1980. Dukea
tomentosa Hook., 1880. Type: MEXICO: panamensis Dwyer, Ann. Missouri Bot. Gard.
Nuevo Leon, Sierra de la Silla, near 53: 364. 1966. Type: PANAMA: Panama,
Monterrey, Aug 1889, C. G. Pringle 2865 Cerro Jefe in Clusia rain forest, 3000 ft., E. L.
(Holotype GH, n.v.).—Randia capitata DC. Tyson, J. Dwyer & K. Blum 3326 (Holotype
Randia xalapensis M. Martens & Galeotti, MO 2084467 (photo PTBG)).—Raritebe
Bull. Acad. Roy. Sci. Bruxelles 11: 239. 1844. palicoureoides Wernham subsp.
Type: MEXICO: Veracruz, Xalapa, dwyerianum J. H. Kirkbr.
Cordillera, Jun-Oct 1840, H. Galeotti 7116 Raritebe trifoliatum (Dwyer & Hay den) Dwyer,
(Holotype BR, n.v.; Isotypes GH (label states Ann. Missouri Bot. Gard. 67: 455. 1980.
"Galeotti s.n"), P (photos MEXU, PTBG)). Dukea trifoliata Dwyer & Hayden, Ann.
Missouri Bot. Gard. 54: 139. 1967. Type:
Raritebe Wernham, J. Bot. 55: 336. 1917. Type: PANAMA: Code, El Valle, potato farm
Raritebe palicoureoides Wernham above village of El Valle, 12 Jul 1962, J. D.
Raritebe blumii (Dwyer) Dwyer, Ann. Missouri Dwyer 1933 (Holotype MO 3831231 (photo
Bot. Gard. 67: 452. 1980. Dukea blumii PTBG)).—Raritebe palicoureoides
Dwyer, Ann. Missouri Bot. Gard. 53: 365. Wernham subsp. dwyerianum J. H. Kirkbr.
1966. Type: PANAMA: Colon, around Dos Raritebe victoriae (Dwyer) Dwyer, Ann.
Bocas, Rio Fato valley, 40-80 m, 16 Aug Missouri Bot. Gard. 67: 456. 1980. Dukea
victoriae Dwyer, Ann. Missouri Bot. Gard. Relbunium Hook. f. in Benth. & Hook, f., Gen.
53: 364. 1966. Type: PANAMA: Darien, PI. 2: 149. 1873. Type: Not designated.
cloud forest, Cerro Pirre, Oct 1962, J. A. Duke [Herein treated as a synonym of Galium L.]
6099 (Holotype MO 1832075 (photo PTBG); • aschenbornii (S. Schauer) Hemsl.,
Isotype MO (photo PTBG)).—Raritebe Biol. Centr.-Amer., Bot. 2: 62. 1881. Galium
palicoureoides Wernham subsp. aschenbornii S. Schauer, Linnaea 20: 701.
dwyerianum J. H. Kirkbr. 1847. Type: MEXICO: Zimapan in Mexico,
Raritebe viburnioides Standi., Publ. Field Aschenborn 92 (Holotype presumably BR or
Columbian Mus., Bot. Ser. 7: 56. 1930. Type: K, n.v.; probable fragment JE, n.v.).—Galium
COLOMBIA: Cauca, La Gallera, Micay
Valley, 1 Jul 1922, Killip 7934 (Holotype US, urn (L.) Hemsl., Biol.
n.v.; fragment F, n.v.; Isotypes G, n.v., NY, Centr.-Amer., Bot. 2: 63. 1881. Valantia
n.v.).—Bertiera viburnoides (Standi.) J. H. hypocarpia L., Syst. Nat., ed. 10, 1307. 1759.
Kirkbr. Type: JAMAICA: Rubia I, Browne, Hist.
Jamaica 141 (Holotype LINN, n.v.).
Ravnia Oerst., Vidensk. Meddel. Dansk —Galium hypocarpium (L.) Endl. ex Griseb.
Naturhist. Foren. Kjobenhavn 1852: 49. 1853. (DC.) Hemsl., Biol.
Type: Ravnia triflora Oerst. [Herein treated as Centr.-Amer., Bot. 2: 63. 1881. Rubia
a synonym of Hillia Jacq. subg. Ravnia laevigata DC, Prodr. 4: 591. 1830. Type:
(Oerst.) C. M. Taylor.] MEXICO: Without precise locality, Haenke
Ravnia longifilamentosa Steyerm., Ceiba 3: 21. s.n. (Holotype presumbly PR, n.v.).—Galium
1952. Type: PANAMA: Chiriqui, upper microphyllum A. Gray
forested southern slopes of Quebrada Velo, mm Greenm., Proc. Amer.
vicinity of Finca Lerida, 5000 ft., 25 Jul 1947, Acad. Arts 41: 250. 1905. Type: MEXICO:
P. H. Allen 4744 (Holotype F 1394241 (photo Hidalgo, moist banks, barranca below
PTBG); Isotype MO, n.v.).—Hillia Trinidad Iron Works, 5500 ft., 6 May 1904,
longifilamentosa (Steyerm.) C. M. Taylor C. G. Pringle 8834 (Holotype GH, n.v.;
Ravnia panamensis Steyerm., Ceiba 3: 22. 1952. Isotypes MEXU, UC, n.v., US (photos
Not Hillia panamensis Standi., 1921. Type: MEXU, PTBG)).—Galium sphagnophilum
PANAMA: Code, North Hills, El Valle de (Greenm.) Dempster var. mazocarpum
Anton, 29 Jun 1946, P. H. Allen 3558 (Greenm.) Dempster
[collector erroneously omitted in protologue] Relbunium microphyllum (A. Gray) Hemsl.,
(Holotype F 1394242 (photo PTBG); Isotype Biol. Centr.-Amer., Bot. 2: 63. 1881. Galium
MO, n.v.).—Hillia allenii C. M. Taylor microphyllum A. Gray, PI. Wright. 1: 80.
Ravnia pittieri Standi., N. Amer. Fl. 32: 114. 1852. Type: U.S.A.: New Mexico, from
1921. Type: COSTA RICA: Santa Clara, Jul western Texas to El Paso, New Mexico,
1899, H. Pittier 1346J (Holotype NY, n.v.). May-Oct 1849, C. Wright 234 (Holotype GH,
—Hillia triflora (Oerst.) C. M. Taylor var. n.v.; Isotype US (photo PTBG)).—Galium
pittieri (Standi.) C. M. Taylor microphyllum A. Gray
Ravnia triflora Oerst., Vidensk. Meddel. Dansk Relbunium microphyllum (A. Gray) Hemsl. var.
Naturhist. Foren. Kjobenhavn 1852: 50. 1853. hispidulum Greenm., Proc. Amer. Acad. Arts
Type: COSTA RICA: Near Cartago and 33: 470. 1898. Syntypes: MEXICO: San Luis
Candelaria, 6000-8000 ft., Oersted 11696 Potosi, valley of San Luis Potosi, Schaffner
(Lectotype C, n.v. (photos F, MO, NY); 842 (GH, n.v.). Region of San Luis Potosi,
designated by Standley, 1938).—Hillia 1850-2460 m, Perry & Palmer 308 (GH,
triflora (Oerst.) C. M. Taylor var. triflora n.v).—Galium microphyllum A. Gray
Relbunium microphyllum (A. Gray) Hemsl. var.
setulosum Greenm., Proc. Amer. Acad. Arts
A NOMENCLATOR OF MEXICAN / D CENTRAL AMERICAN RUBIACEAE
33: 470. 1898. Syntypes: MEXICO: Hidalgo, terrenos arenosos, 23 Jun 1954, J. Rzedowsh
Zimapan, Coulter 186 (GH, n.v.); San Luis 3168 (Holotype ENCB).
Potosi, region of San Luis Potosi, C. C. Perry Richardia haenkeana (DC.) Steud., Nom. Bot.,
& E. Palmer 309 (GH, n.v.), C. C. Pern' & E. ed. 2, 459. 1841. Richardsonia haenkeana
Palmer 310 (GH, n.v.).~Galium DC, Prodr. 4: 569. 1830. Type: MEXICO:
microphyllum A. Gray Without precise locality, Haenke s.n.
Relbunium polyplocum Hemsl., Diag. PI. Nov. (Holotype presumably PR, n.v.).—Richardia
Mexic. 53. 1880. Type: MEXICO: San Luis humistrata (Cham. & Schltdl.) Steud.
Potosi, without precise locality, 1878, C. C. Richardia humistrata (Cham. & Schltdl.)
Parry & E. Palmer 308 (Syntype US 19271 Steud., Nom. Bot., ed. 2, 459. 1841.
(photos MEXU, PTBG), K, n.v.). MEXICO: Richardsonia humistrata Cham. & Schltdl.,
Veracruz, Mirador, Linden 516 (Syntype K, Linnaea 3: 353. 1828. Type: URUGUAY:
n.v.), Liebmann 243 (Syntype K, n.v.), Montevideo, in Brasihae meridionalis
provincia Montevideo, Sellow s.n. (Holotype
microphyllum A. Gray B, presumably destroyed (fragment F)).
m sphagnophilum Greenm., Proc. Richardia procumbens Sesse & Moc., Fl.
Amer. Acad. Arts 33: 468. 1898. Type: Mexic, ed. 2, 83. 1894. Type: MEXICO:
MEXICO: Oaxaca, Sierra de Clavellinas, Habitat in Ahualulci apricis, Sesse & Mocino
2270 m, 17 Oct 1894, C. G. Pringle 5812 s.n. (presumably MA, n.v.).—Richardia
(Holotype GH, n.v.).—Galium scabra L.
sphagnophilum (Greenm.) Dempster var. Richardia scabra L., Sp. PI. 1: 330. 1753.
sphagnophilum Type: MEXICO: Veracruz (Lectotype LINN
451.1 (n.v.); for typification see Lewis &
Rhodostoma Scheidw., Allg. Gartenzeitung 10: Oliver, 1974).
286. 1842. Type: Rhodostoma gardenioides Richardia tricocca (Torr. & A. Gray) Standi.,
Scheidw. Publ. Field Columbian Mus., Bot. Ser. 8: 386.
Rhodostoma gardenioides Scheidw., Allgem. 1931. subsp. tricocca. Diodia tricocca Torr.
Gartenzeitung 10: 286. 1842. Type: Based on & A. Gray, Fl. N. Amer. 2: 30. 1841. Type:
cultivated material of uncertain origin, U.S.A.: Texas, Drummond 117 (Holotype
presumably Veracruz, Mexico (type not GH, n.v.; Isotype G, n.v.).
designated). Palicourea.—Psychotria Richardia tricocca (Torr. & A. Gray) Standi,
gardenioides (Scheidw.) Standi. subsp. tetracocca (M. Martens & Galeotti)
W. H. Lewis & R. L. Oliv., Bnttonia 26: 287.
Richardia L., Sp. PI. 1: 330. 1753. Type: 1974. Spermacoce tetracocca M. Martens &
Richardia scabra L. Galeotti, Bull. Acad. Roy. Soc. Bruxelles 11:
Richardia adscendens (Pav. ex DC.) Steud., 132. 1844. Type: MEXICO: Michoacan,
Norn. Bot., ed. 2, 458. 1841. Richardsonia Santa Maria, near Morelia, H. Galeotti 2587
adscendens Pav. ex DC, Prodr. 4: 569. 1830. (Holotype BR, n.v.; Isotypes K, n.v., P
Type: MEXICO: Without precise locality, (photos MEXU, PTBG)).
Moricands.n. (Holotype G-DC, n.v. (photo
MO)).—Richardia brasiliensis Gomes Richardsonia Kunth in Humb., Bonpl. & Kunth,
Richardia brasiliensis Gomes, Mem. Nov. Gen. Sp. PI. 3: 350. 1819. [Herein
Ipecacuanha Bras. 31, t. 2. 1801. Type: treated as a synonym of Richardia L.]
BRAZIL: Without precise locality, Gomes s.n. Richardsonia adscendens Pav. ex DC, Prodr. 4:
(Holotype LISU, n.v.). 569. 1830. Type: MEXICO: Without precise
Richardia gandarae Rzed., Bol. Soc. Bot. locality, Moricands.n. (Holotype G-DC, n.v.
Mexico 44: 11, fig. 2. 1983. Type: MEXICO: (photo MO)). Richardia.—Richardia
San Luis Potosi, Santa Maria del Rio, 1750 m, brasiliensis Gomes
Richardsonia brasiliensis (Gomes) Hayne, Getr. Rogiera amoena Planch., Fl. Serr. Jard. 5: tab.
Darstell. Gew. 14: tab. 21. 1846. Richardia 442, figs. 1, 2. 1849. Type: Based on plants
brasUiensis Gomes, Mem. Ipecacuanha Bras. cultivated in Belgium, presumably from
31, tab. 2. 1801. Type: BRAZIL: Without Guatemalan seed (type not designated).
precise locality, Gomes s.n. (LISU, n.v.). —Rondeletia amoena (Planch.) Hemsl.
—Richardia brasiliensis Gomes Rogiera aprica (Lundell) Borhidi, Acta Bot.
Richardsonia haenkeana DC., Prodr. 4: 569. Hung. 33: 301. 1987. Rondeletia aprica
1830. Type: MEXICO: Without precise Lundell, Contrib. Univ. Michigan Herb. 7: 54.
locality, Haenke s.n. (Holotype presumably 1942. Type: MEXICO: Chiapas, Saxchanal,
PR, n.v). Richardia.—Richardia humistrata Sierra Madre, 1-5 Jul 1941, E. Matuda 4314
(Cham. & Schltdl.) Steudel (Holotype TEX-LL (photos MEXU, MO,
Richardsonia humistrata Cham. & Schltdl., PTBG); Isotypes A, n.v., F, MO, US (photos
Linnaea 3: 353. 1828. Type: URAGUAY: MEXU, PTBG))—Rondeletia cordata
Montevideo, in Brasiliae meridionalis Benth.
provincia Montevideo, Sellow s.n. (Holotype Rogiera backhousii (Hook, f.) Borhidi, Acta
B, presumably destroyed (fragment F)). Bot. Hung. 33: 301. 1987. Rondeletia
—Richardia humistrata (Cham.) Schltdl.) backhousii Hook, f., Bot. Mag. 103: tab.
Steud. 6290. 1877. Type: Based on material of
Richardsonia scabra (L.) A. St.-Hil., PI. Us. unknown origin cultivated at Kew, obtained
Bras. 8: 1.1. 8. 1824, p.p. Richardia scabra L., from the Backhouse establishment in York
Sp. PL 1: 330. 1753. Type: MEXICO: (type not designated).—Rondeletia
Veracruz, (Holotype LINN 451.1 (n.v.)). backhousii Hook. f.
—Richardia scabra L. Rogiera brachistantha (Standi. & Steyerm.)
Richardsonia tetracocca (M. Martens & Borhidi, Acta Bot. Hung. 33: 301. 1987.
Galeotti) K. Schum. ex Loes., Repert. Sp. Rondeletia brachistantha Standi. & Steyerm.,
Nov. 18: 362. 1922. Spermacoce tetracocca Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 25.
M. Martens & Galeotti, Bull. Acad. Roy. Soc. 1943. Type: GUATEMALA: San Marcos, Rio
Bruxelles 11: 132. 1844. Type: MEXICO: Vega, near San Rafael and Guatemala-Mexico
Michoacan, Santa Maria, near Morelia, H. boundary, Volcan Tacana, 2500-3000 m, 20
Galeotti 2587 (Holotype BR, n.v.; Isotypes K, Feb 1940, J. A. Steyermark 36286 (Holotype
n.v., P (photos MEXU, PTBG)). Diodia. F 1050045 (photos MEXU, PTBG); Isotype
—Richardia tricocca (Torr. & A. Gray) US (photos MEXU, PTBG)).—Rondeletia
Standi, subsp. tetracocca (M. Martens & cordata Benth.
Galeotti) W. H. Lewis & R. L. Oliv. Rogiera chiapasensis (Standi.) Borhidi, Acta
Richardsonia tetracocca var. calvescens Loes., Bot. Hung. 33: 301. 1987. Rondeletia
Repert. Sp. Nov. 18: 362. 1922. Syntypes: chiapasensis Standi., Publ. Field Mus. Nat.
MEXICO: Without precise locality, Schaffner Hist., Bot. Ser. 22: 54. 1940. Type: MEXICO:
281 (K, n.v.); Schaffner 282 (K, n.v.). Chiapas, Cascarada, 31 Dec 1936, E. Matuda
—Richardia tricocca (Torr. & A. Gray) 1064 (Holotype MICH, n.v. (fragment F);
Standi, subsp. tetracocca (M. Martens & Isotype A, n.v).—Gonzalagunia
Galeotti) W. H. Lewis & R. L. Oliv. chiapasensis (Standi.) Standi. & Steyerm.
Rogiera cordata (Benth.) Planch., Fl. Serr. Jard.
Rogiera Planch., Fl. Serr. Jard. 5: tab. 442. 5: tab. 442b. 1849. Rondeletia cordata Benth.,
1849. Lectotype: Rogiera amoena Planch, PI. Hartweg. 85. 1841. Type: GUATEMALA:
(designated by Borhidi, Acta Bot. Acad. S< In praeruptis prope Guatemala [Guatemala
Hung. 28: 67. 1982). [Herein treated as a City], Jan 1841, T. Hartweg 585 (Holotype K
synonym of Rondeletia L.] (photo PTBG); Isotype G-DEL, n.v. (fragment
F)).—Rondeletia cordata Benth.
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
(K). The K specimen of Sumichrast sub Rondeletia belizensis Standi, var. longiloba
Botteri 1017 is the best choice for lectotype Lundell, Wrightia 5: 323. 1976. Type:
and has duplicates in two other herbaria. GUATEMALA: Alta Verapaz, Rubelsanto,
—Rondeletia buddleioides Benth. 2.3 km SEE [=ESE] of Balastrera, 20 Jul
Rondeletia albida Lundell, Wrightia 5: 323. 1975, C. L. Lundell & E Contreras 19545
1976. Type: MEXICO: Chiapas, Carelas, near (Holotype TEX-LL (photos MEXU, PTBG);
Motozintla, 2176, Apr 1945, E. Matuda 5509 Isotypes F, MO (photo PTBG)).—Rondeletia
(Holotype TEX-LL (photos MEXU, PTBG); $ Standi.
Isotypes F, MEXU, TEX-LL).—Rondeletia i bertieroides Standi., Publ. Field
laniflora Benth. Columbian Mus., Bot. Ser. 4: 267. 1929.
Rondeletia amoena (Planch.) Hemsl., Diag. PI. Type: PANAMA: Bocas del Toro, Buena
Mex. 26. 1879. Rogiera amoena Planch., Fl. Vista Camp, Chiriqui Trail, 900 m, 1928,
Serr. Jard. 5: tab. 442, figs 1, 2. 1849. Type: G. P. Cooper 598 (Yak 12231) (Holotype F
Based on plants cultivated in Belgium, 579614 (photo PTBG); Isotype NY, n.v.).
presumably from Guatemalan seed (type not Arachnothryx.
designated). Rondeletia bourgaei Standi., N. Amer. Fl. 32:
Rondeletia aprica Lundell, Contrib. Univ. 56. 1918. Type: MEXICO: Veracruz, Region
Michigan Herb. 7: 54. 1942. Type: MEXICO: de Orizaba, Monte San Cristobal, 8 Jul 1866,
Chiapas, Saxchanal, Sierra Madre, in E. Bourgeau 2618 (Holotype US 49964
advanced forest, 1-5 Jul 1941, E. Matuda (photos MEXU, PTBG); Isotypes P (4 sheets,
4314 (Holotype TEX-LL (photos MEXU, photos MEXU, PTBG)). Arachnothryx.
MO, PTBG); Isotypes A, n.v., F, MO, US —Rondeletia laniflora Benth.
(photos MEXU, PTBG)). Rogiera. Rondeletia brachistantha Standi. & Steyerm.,
—Rondeletia cordata Benth. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 25.
Rondeletia aspera Standi., N. Amer. Fl. 32: 54. 1943. Type: GUATEMALA: San Marcos, Rio
1918. Type: COSTA RICA: San Jose, Pont du Vega, near San Rafael and Guatemala-Mexico
Rodeo, Pacacol (Villa Colon), Jan 1891, f/. boundary, Volcan Tacana, 2500-3000 m, 20
Pittier 3243 (Holotype US 941408 (photo Feb 1940, J. A. Steyermark 36286 (Holotype
PTBG)). Arachnothryx. F 1050045 (photos MEXU, PTBG); Isotype
Rondeletia atravesadensis Lorence, Novon 1: US (photos MEXU, PTBG)). Rogiera.
137,/jg. 1A, B. 1991. Type: MEXICO: —Rondeletia cordata Benth.
Oaxaca, Cerro Atravesada-Cerro Azul, Rondeletia brandegeeana Lorence, Novon 1:
Arroyo Rana, 6 Dec 1956, T. MacDougall s.n. 138. 1991. Based on Otocalyx chiapensis
(Holotype MEXU 253128 (photo PTBG)). Brandegee, Univ. Calif Publ. Bot. 6: 69.
Rondeletia backhousii Hook.f, Bot. Mag. 103: 1914. Not Rondeletia chiapasensis Standi..
tab. 6290. 1877. Based on material of 1940. Type: MEXICO: Chiapas, Cerro del
unknown origin cultivated at Kew, obtained Boqueron, middle region, moist valley slopes,
from the Backhouse establishment in York Aug 1913, C. A. Purpus 7041 (Holotype UC
(type not designated). Rogiera.—Status 173047 (photo PTBG); Isotypes GH, US
uncertain, resembles Rondeletia panamensis (fragment)).
DC. but is nearly glabrous. Rondeletia breedlovei Lorence. Novon 4: 129,
Rondeletia belizensis Standi., Carnegie Inst. fig 6. 1994. Type: MEXICO: Chiapas, Mun.
Wash. Publ. 461: 91. 1935. Type: BRITISH of Tenejapa, paraje of Mahben Chauk, on
HONDURAS (BELIZE): Jacinto Hills, 300 slopes along the Tana Te' River near Shal
ft., 28 Aug 1933, W. A. Schipp 1201 K'esh, 2900 ft., 27 Nov 1964, D. E Breedlove
(Holotype F 683600 (photos MEXU, PTBG); 7682 (Holotype F 1617735 (photo PTBG);
Isotypes GH, n.v., K (photo PTBG), MO). Isotype CAS (photo PTBG)).
Arachnothryx.
Rondeletia brenesii Standi., Publ. Field Mus. 1993. Type: COSTA RICA: Heredia,
Nat. Hist., Bot. Ser. 18: 1370. 1938. Type: Magsasay, pasturas, bosque socolado, finca de
COSTA RICA: Camino de San Ramon a La Bernardo Gomez, 700 m, 1 Jan 1983, /. C.
Calera, 20 Mar 1929, A. M. Brenes 6757 Chacon 177 (Holotype CR 92850; Isotype
(Holotype F 854972 (photo PTBG)). MO).
Arachnothryx. Rondeletia chiapasensis Standi., Publ. Field
Rondeletia buddleioides Benth., PI. Hartweg. Mus. Nat. Hist., Bot. Ser. 22: 54. 1940. Type:
69. 1840. Type: MEXICO: Oaxaca, in planitie MEXICO: Chiapas, Cascarada, 31 Dec 1936,
Llano Verde, T. Hartweg 502 (Holotype K E. Matuda 1064 (Holotype MICH, n.v.
(photo PTBG)); Isotypes B (destroyed, photo (fragment F); Isotype A, n.v.). Rogiera.
MO, fragment G-DEL), K (photo PTBG), —Gonzalagunia chiapasensis (Standi.)
NY, n.v. (fragment US, photos MEXU, Standi. & Steyerm.
PTBG), P (photos MEXU, PTBG)). Rondeletia chinajensis Standi. & Steyerm.,
Arachnothryx. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.
Rondeletia buddleioides Benth. var. aspera J. H. 1947. Type: GUATEMALA: Alta Verapaz,
Kirkbr., Ann. Missouri Bot. Gard. 55: 379. Cerro Chinaja, between Finca Yalpemech and
1968. Type: PANAMA: Bocas del Toro, Chinaja, above source of Rio San Diego,
between Criollo (just above Buena Vista) and 150-700 m, 1-2 Apr 1942, J. A Si
Quebrada Higueron on Chiriqui Trail, 18 Apr 45637 (Holotype F 1134210 (photos MEXU,
1968, J. H. Kirkbride, Jr. &J.A. Duke 780 PTBG)). Arachnothryx.
(Holotype MO 1942973 (photos MEXU, Rondeletia chiriquiana Lorence, Novon 4:
PTBG); Isotype US). Arachnothryx. 129, fig. 7. 1994. Type: PANAMA: Chiriqui,
—Rondeletia buddleioides Benth. Fortuna Dam area, along shores of Rio
Rondeletia calycosa Donn. Sm., Bot. Gaz. Chiriqui where it enters reservoir, 1000 m,
(Crawfordsville) 56: 59. 1913. Type: COSTA 8°45'N, 82°13'W, 2 Aug 1984, H. W.
RICA: San Jose, forets de La Palma, 1460 m, Churchill 5924 (Holotype MO 3595277
25 Sep 1898, A. Tonduz 11633 (Holotype US (photo PTBG)).
no 941406 (photos F, MEXU, PTBG)). Rondeletia cooperi Standi., Publ. Field
Arachnothryx. Columbian Mus., Bot. Ser. 4: 267. 1929.
Rondeletia capitellata Hemsl., Diag. PI. Mex. Type: PANAMA: Bocas del Toro, Buena
28. 1879. Type: MEXICO: Without precise Vista Camp, on Chiriqui Trail, 1250 ft.,
locality, Jurgensen 355 (Lectotype K, here Jan-Mar 1928, G. P. Cooper 600 (Yale
designated (photo PTBG); Isolectotypes G- 12233) (Holotype F 579596 (photo PTBG);
DEL, n.v. (fragment F), K (photo PTBG). Isotype NY, n.v.). Arachnothryx.
Syntypes: MEXICO: Oaxaca, Joya, F. Rondeletia cordata Benth., PI. Harrweg. 85.
Liebmann 202 (K (photo PTBG)). MEXICO: 1841. Type: GUATEMALA: In praeruptis
Without precise locality, Jurgensen 376 (K prope Guatemala [Guatemala City], Jan 1841,
(photo PTBG)). Arachnothryx. The K sheet of T. Hartweg 585 (Holotype K (photo PTBG);
Jurgensen 355 is very floriferous and Isotype G-DEL, n.v. (fragment F)). Rogiera.
represents the best choice for lectotype. Note: Rondeletia cordovana Standi. & Steyerm., Publ.
Rondeletia capitellata and R. liebmannii Field Mus. Nat. Hist., Bot. Ser. 23: 26. 1943.
Standi, are synonymous. Material from Type: GUATEMALA: Zacapa, in ravine
Veracruz and Hidalgo erroneously identified bordering Quebrada Alejandria, summit of
as R. capitellata in many herbaria actually Sierra de las Minas, vicinity of Finca
represents a distinct species, R. pringlei Alejandra, 2500 m, 13 Oct 1939, J. A.
Steyermark 29906 (Holotype F 1042952
(photos MEXU, PTBG); Isotype US (photos
A NOMENCLATOR OF MEXICAN AND CENTRAL /
Sierra de San Pedro Nolasco, Jurgensen 798 presumably destroyed (fragment F), G-DEL,
(K (photo PTBG)). A probable isolectotype at n.v. (fragment F), K (photo PTBG), P (2
P [Linden s.n. (fragment F)] annotated "fleurs sheets; photos MEXU, PTBG)). Rogiera.
rosatres" has corolla tubes 6-7 mm long and —Rondeletia amoena (Planch.) Hemsl.
is likely a duplicate of Linden 1661 at K. A Rondeletia laniflora Benth., PL Hartw. 85.
probable syntype at P annotated "fleurs 1841. Type: GUATEMALA: Mountains of
rouges" has corolla tubes 5-7 mm long and Las Nubes, March 1841, T Hartweg 584
larger leaves, and is likely a duplicate of (Holotype K (photo PTBG), fragment US
Linden 1659 at K. The Linden 1661 specimen (photos MEXU, PTBG); Isotypes B,
at K is selected as lectotype, because it has presumably destroyed (photos F, MO, US), G-
more leaves, inflorescences, and young fruit. DEL, n.v. (fragment F), P (photos MEXU,
Hemsley noted that this species was PTBG), W, n.v. (photo F)). Arachnothryx.
intermediate between R. amoena and R. Rondeletia latifolia Oerst., Vidensk Meddel.
cordata, but the original material has sparsely Dansk Naturhist. Foren. Kjobenhavn 1852:
pubescent lower leaf surfaces and is clearly 43. 1853. Type: NICARAGUA: In monte
synonymous with R. cordata. In the Flora of Pantasmo in Segovia, Jan 1848, Oersted 76
Guatemala Rubiaceae treatment Standley & [=Plantae centroamericanae Oersted 11678]
Williams (1975) also treated this species as a (Holotype C, n.v.; Isotypes K (photo PTBG),
synonym of R. cordata Benth. Rogiera. US (photo PTBG)). Rogiera.—Rondeletia
amoena (Planch.) Hemsl.
a izabalensis Standi. & Steyerm., Rondeletia leptodicfta B. L. Rob., Proc. Amer.
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 286. Acad. Arts 45: 402. 1910. Type: MEXICO:
1940. Type: GUATEMALA: Izabal, Escoba, [Boundary between Michoacan and
across the bay (west) from Puerto Barrios, Guerrero], Rives du Rio Petatlan, 500 m, 24
near sea level, 3 May 1939, P. C. Standley Nov 1898, E Langlasse 666 (Holotype GH,
72891 (Holotype F 991611 (photos MEXU, n.v.; Isotypes F (photos MEXU, PTBG), G-
PTBG)). Arachnothryx. DEL, n.v. (fragment F), K (photo PTBG), P (2
Rondeletia jurgensenii Hemsl., Diag. PL Nov. sheets; photos MEXU, PTBG; fragment F),
Mexic. 29. 1879. Type: MEXICO: Without US (photos MEXU, PTBG)). Arachnothryx.
precise locality [protologue states "Oaxaca, —Rondeletia leucophylla Kunth in Humb.,
Sierra de San Pedro Nolasco"], Jurgensen 248 Bonpl. & Kunth
(Holotype K (photo PTBG); Isotype G-DEL, Rondeletia leucophylla Kunth in Humb.,
n.v. (fragment F, label states "Trapiche de Bonpl. & Kunth, Nov. Gen. Sp. PI. 3: 395, t.
Concepcion and Chicahuastla, Jun 1845")). 290. 1820. Type: MEXICO: Guerrero, inter
Arachnothryx. Alto del Peregrino et flumen Papagallo,
Rondeletia kirkbridei Dwyer, Ann. Missouri Humboldt & Bonpland s.n. (Holotype P-
Bot. Gard. 67: 469. 1980. Type: PANAMA: Bonpland, n.v.; Isotypes B, presumably
Veraguas, 16 km NW of Santa Fe, on road to destroyed (photos F, MO), P (photos MEXU,
Calovebora (Panama Hwy. 35), 300-500 m, PTBG)). Arachnothryx.
16 May 1975, S. Mori & J. Kallunki 6112 Rondeletia leucophylla Kunth in Humb., Bonpl.
(Holotype MO 2300094 (photo PTBG)). & Kunth var. calycosa Greenm., Proc. Amer.
Arachnothryx.—Rondletia bertieroides Acad. Arts 39: 92. 1903. Type: MEXICO:
Standi. Oaxaca, 1650 m, Dec 1900, C. Conzatti & V.
Rondeletia langlassei Standi., N. Amer. Fl. 32: Gonzalez 1127 (Syntype GH, n.v.); Oaxaca,
53. 1918. Type: MEXICO: [Michoacan or city of Oaxaca, 1550 m, C. Conzatti 970
Guerrero] Sierra Madre, 1750 m, 26 Jan 1899, (Syntype GH, n.v.).—Rondeletia leucophylla
E Langlasse 797 (Holotype US 386194 Kunth in Humb., Bonpl. 8c Kunth
(photos MEXU, PTBG); Isotypes B,
Rondeletia liebmannii Standi., J. Wash. Acad.
Sci. 17: 336. 1927. Type: MEXICO: Oaxaca,
without precise locality, 1841^3, F. Rondeletia macdougallii Lorence, Novon 1:
Liebmann J1834 (Holotype US 1315231 144, Jig. 3A, B. 1991. Type: MEXICO:
(photos MEXU, PTBG); Isotypes C, n.v. Oaxaca, Cerro Azul (top), N of Niltepec, 7000
(photo F), P (photos MEXU, PTBG)). ft., 7 Mar 1956, T. MacDougalls.n. (Holotype
Arachnothryx.—Rondeletia capitellata MEXU 253131 (photo PTBG)).
Hemsl. Rondeletia macrocalyx Standi. & Steyerm.,
Rondeletia ligustroides Hemsl., Diag. PI. Nov. Publ. Field Mus. Nat. Hist., Bot. Ser. 23: 254.
Mexic. 26. 1879. Type: MEXICO: Veracruz, 1947. Type: GUATEMALA: Izabal, Cerro
Orizaba, Botteri 971 (Lectotype K, here San Gil, 300-900 m, 25 Dec 1941, J. A.
designated (photo PTBG); Isolectotypes P (2 Steyermark 41864 (Holotype F 1134224
sheets, photos MEXU, PTBG). Syntypes: (photos MEXU, PTBG); Isotype F (photos
MEXICO: Veracruz, Orizaba, Bourgeau 2518 MEXU, PTBG), US). Arachnothryx.
(K (photo PTBG), P (3 sheets; photos MEXU, Rondeletia manantlanensis Lorence, Novon 1:
PTBG)). Region d'Orizaba, Bourgeau 2622 145, fig. 4A, B. 1991. Type: MEXICO:
(K (photo PTBG), B (n.v.; fragment F), F, K Jalisco, Mun. de Autlan, Canada del Laurelito,
(photo PTBG), P (4 sheets; photos MEXU, Las Joyas, 1850 m, 7 Dec 1985, A. Vazquez
PTBG), US (2 sheets)). Orizaba, Botteri 474 3750 (Holotype WIS (photo PTBG); Isotype
(K (2 sheets [label on first states "coll. Salle"; IBUG, n.v.).
second states "Sumichrast coll."] photos Rondeletia megalantha Lorence, Novon 4:
PTBG)), P (photos MEXU, PTBG). Orizaba, Uljig. 8. 1994. Type: HONDURAS:
Botteri 1951 (K, n.v., P (2 sheets; photos Cuyamel, 23 Aug 1924, M. A. Carleton 653
MEXU, PTBG). Orizaba, Muller 969 (K, n.v., (Holotype US 1208409; Isotype US
P (photos MEXU, PTBG). Orizaba, Muller 1208410).
1308 (K, n.v.). Botteri 971 at K is selected as Rondeletia mexiae Standi., Publ. Field Mus.
lectotype because it has three leafy twigs with Nat. Hist., Bot. Ser. 22: 55. 1940. Type:
an abundance of flowers at anthesis, and three MEXICO: Oaxaca, Distr. Choapan, Santa
duplicates exist at P. Rogiera. Maria, Montana Santa Maria, 1500 m, 1 Apr
Rondeletia linguiformis Hemsl., Diag. PL Nov. 1938, Y. Mexia 9261 (Holotype F 982856
Mexic. 29. 1879. Type: GUATEMALA: (photos MEXU, PTBG); Isotypes GH, n.v.,
Without precise locality, Skinner s.n. in 1857 MO (photos MEXU, PTBG), UC (photo
(Holotype K (photo PTBG)). Arachnothryx. PTBG), US (photos MEXU, PTBG)).
Rondeletia longipetiolata Lundell, Wrightia 5: Arachnothryx.—Rondeletia scabra Hemsl.
325. 1976. Type: MEXICO: Chiapas, Mt. Rondeletia mexicana (Turcz.) Standi., Publ.
Ovando, near Escuintla, 14 Nov 1945, E. Field Mus. Nat. Hist., Bot. Ser. 22: 388. 1940.
Matuda 6146 (Holotype TEX-LL; Isotypes Siphonandra mexicana Turcz., Bull. Soc Imp.
MEXU, TEX-LL). Arachnothryx. Naturalistes Moscou 21: 581. 1848. Type:
—Rondeletia buddleioides Benth. MEXICO: Oaxaca, without precise locality,
Rondeletia lundelliana Standi., Contr. Univ. 3000 ft., Oct, H. Galeotti 2664bis (Holotype
Michigan Herb. 4: 31. 1940. Type: BRITISH G-DEL, n.v. (photos F, MO, US); Isotype P
HONDURAS (BELIZE): El Cayo District, (photos MEXU, PTBG; fragment F, photos
Vaca, 21 Apr 1938, P. H. Gentle 2504 MEXU, PTBG)).
(Holotype F 941520 (photos MEXU, PTBG); Rondeletia microdon DC, Prodr. 4: 408. 1830.
Isotypes A, n.v., K (photo PTBG), TEX-LL Type: CUBA: Havana, de la Ossa s.n.
ex MICH (photos MEXU, PTBG), US (2 (Holotype G-DC, n.v. (microfiche IDC
sheets; photos MEXU, PTBG)). 800.691:111.7,8)).—Psychotria r
(DC.) Urb.
i NOMENCLATOR OF MEXICAN AND CENTRAL
Rondeletia minor Lundell, Wrightia 5: 326. lectotype, because it has abundant flowers,
1976. Type: MEXICO: Veracruz, about 10 and duplicates exist in two other herbaria.
miles north of Teziutlan, 6 Jan 1964, D. S. Arachnothryx.
Correll & H. B. Correll 28736 (Holotype Rondeletia oaxacana Standi., Field Mus. Nat.
TEX-LL (photos MEXU, PTBG); Isotype Hist., Bot. Ser. 11:255. 1936. Type:
MEXU). Arachnothryx.—Rondeletia MEXICO: Oaxaca, without precise locality,
gracilispica Standi. 3000 ft., Oct, H. Galeotti 2664bis (Holotype P
Rondeletia monteverdensis Lorence, Novon 1: (photos MEXU, PTBG), fragment F (photos
147, fig. 5A, B. 1991. Type: COSTA RICA: MEXU, PTBG)). Norn, superfl. for and based
Puntarenas, Monteverde, road to TV tower, on same type as Rondeletia mexicana (Turcz.)
1700 m, 24 Aug 1985, W. A. Haber 2340 Standi. Arachnothryx.—Rondeletia mexicana
(Holotype MO 3595758 (photo PTBG)). (Turcz.) Standi.
Rondeletia myriantha Standi. & Steyerm., Rondeletia odorata Jacq., Enum. PI. 16. 1760.
Publ. Field Mus. Nat. Hist., Bot. Ser. 22: 288. Type: CUBA: Havanna, coastal thickets,
1940. var. myriantha. Type: GUATEMALA: Jacquin s.n. (location unknown). Cultivated in
Suchitepequez, Finca Moca, 3200 ft., 31 Oct
1934, A. F. Skutch 1569 (Holotype F 813463 Rondeletia odorata Jacq. var. breviflora Hook,
(photos MEXU, PTBG); Isotypes A, n.v., US f, Curtis Bot. Mag. 104, tab. 6350. 1878.
(photos MEXU, PTBG)). Arachnothryx. Type: CUBA: Collector and locality unknown
Rondeletia myriantha Standi. & Steyerm. var. (presumably K, n.v.). Reported by Kirkhnde
armentalis L. O. Williams, Phytologia 26: (1969) to occur in Panama.—Rondeletia
127. 1973. Type: GUATEMALA: Quiche, odorata Jacq.
Nebaj, 5600 ft., 22 Nov 1934, A. F. Skutch Rondeletia ovandensis Lundell. Wrightia .v
1776 (Holotype F 815811 (photos MEXU, 326. 1976. Type: MEXICO: Chiapas, Mt.
PTBG)). Arachnothryx. Ovando, near Escuintla, 24 Oct 1941, E.
Rondeletia nebulosa Standi., Trop. Woods 37: Matuda 6067 (Holotype TEX-LL (photos
32. 1934. Type: HONDURAS: Tegucigalpa, MEXU, PTBG); Isotypes F (photo PTBG),
La Aurora Trail, Rosario, San Juancito, 5400 MEXU, TEX-LL, n.v.). Arachnothryx.
ft., 2 Apr 1932, J. B. Edwards PI0 (Holotype Rondeletia panamensis DC, Prodr. 4: 408.
F 662646 (photos MEXU, PTBG); Isotype A, 1830. Type: PANAMA: Without precise
n.v.). Arachnothryx. locality, Haenke s.n. (Holotype PR, n.v.,
Rondeletia nicaraguensis Oerst., Vidensk. fragment G-DC, n.v. (microfiche 4: 408, 8)).
Meddel. Dansk Naturhist. Foren. Kjobenhavn Rondeletia pansamalana Standi., N. Amer. Fl.
1852: 43. 1853. Type: NICARAGUA: 32- 58. 1918. Type: GUATEMALA: Alta
Matagalpa, Segovia, in monte Pantasmo, Jan Verapaz, Pansamala, 3800 ft., Apr 1886, H.
1848, Oersted 11825 (Holotype C, n.v. von Tuerckheim 897 (Holotype US 49985
(photos F, MEXU, PTBG); Isotype GH, n.v.).
(photos F, MO, US; fragment US)).
Arachnothryx. Arachnothryx.
Rondeletia phvllocephala Standi. & L. O.
Rondeletia nitida Hemsl., Diag. PI. Nov. Williams, Ceiba 3: 63. 1952. Type:
Mexic. 29. 1879. Type: MEXICO: Chiapas, fl.
HONDURAS: Cortes, faldas humedas de
rosatres, forets des pins, en fevr, Linden 1660
Montana Sta. Ana, matorrales humedos de
(Lectotype K, here designated (photo PTBG): Rio Sta. Ana, 210 m. 6 Dec 1950, A. Molina
Isolectotypes G-DEL (n.v.; fragment F), P R. 3608 (Holotype US 2216049 (photos
(unnumbered Linden collection, almost MEXU, PTBG); Isotypes F (phot<~5 MEXU,
**cv
certainly a duplicate; photos MEXU, PTBG)). PTBG, US), MO)—Rondeletia t
Syntype: MEXICO: Chiapas, Ghiesbreght 728
(K (photo PTBG), GH, n.v.). Arachnothryx. Donn. Sm.
The Linden 1660 collection at K is designated
Rondeletia pittieri K. Schum. & K. Krause ex K. Rondeletia rekoi Standi., Proc. Biol. Soc.
Krause, Bot. Jahrb. Syst. 40: 316. 1908. Type: Wash. 8: 126. 1918. Type: MEXICO: Oaxaca,
COSTA RICA: Bords du Rio Poros, 2000 m, Las Pilas (Cerro Espino), 400 m, 10 Oct 1917,
10 Jan 1890, H. Pittier & T Duranci 1729 B. P. Reko 3490 (Holotype US 867145
(Holotype B, presumably destroyed (photos F, (photos MEXU, PTBG)). Arachnothryx.
MO); Isotype US (photos MEXU, PTBG)). Rondeletia repens L., Syst., ed. 10, 2: 928.
Rogiera.—Rondeletia amoena (Planch.) 1759. Type: JAMAICA: Sloanes.n.
Hemsl. (Lectotype: Sloane herb, iv.l 11, n.v.;
Rondeletia platysepala Standi., Ann. Missouri designated by Howard, 1989).—Geophila
Bot. Gard. 27: 343. 1940. Type: PANAMA: repens (L.) I. M. Johnst.
Code, El Valle, North Rim, 600-1000 m, 14 Rondeletia ricoi Lorence, Novon 1: 152, fig.
May 1939, P. H. Allen 1791 (Holotype F 8E, F, 9A, B. 1991. Type: MEXICO: Oaxaca,
989484 (photo PTBG); Isotypes F (photos Distr. de Putla de Guerrero, a 10 km al NE de
MEXU, PTBG), G, n.v., GH, n.v., MO Putla, 950 m, 9 Dec 1982, L. Rico A., E. M.
(photos MEXU, PTBG), NY, n.v.). Martinez & O. Tellez V. 566 (Holotype
Arachnothryx.—Rondeletia bertieroides MEXU (photo PTBG); Isotypes BM, F, MO,
Standi. PTBG).
Rondeletia polycephala Standi., J. Wash. Acad. Rondeletia roezlii (Planch.) Hemsl., Diag. PI.
Sci. 17: 337. 1927. Type: MEXICO: Oaxaca, Nov. Mexic. 25. 1879. Rogiera roezlii
without precise locality, F. Liebmann 11826 Planch., Fl. Serr. Jard. 5: tab. 442. 1849.
(Holotype C, n.v. (fragment US; photos Type: Based on cultivated material grown in
MEXU, PTBG)). Arachnothryx.—Rondeletia Belgium, probably of Guatemalan origin (type
mexicana (Turcz.) Standi. not designated).—Rondeletia amoena
Rondeletia povedae Lorence in W. C. Burger (Planch.) Hemsl.
& C. M. Taylor, Fieldiana Bot., n.s. 33: 299. Rondeletia rothschuhii Loes., Engl. Bot. Jahrb.
1993. Type: COSTA RICA: San Jose, Santa 60: 369. 1926. Type: NICARAGUA:
Rosa de Puriscal, 17 Jan 1973, L. J. Poveda Matagalpa, Canada Yucul, 1000 m, Rothschuh
441 (Holotype CR 51260; Isotype MO 592 (B, presumably destroyed (photos F, MO;
2211249). fragment F)).—Rondeletia buddleioides
Rondeletia pringlei Lorence, Novon 7: 54, fig. Benth.
5. 1997. Type: MEXICO: Veracruz, hills near Rondeletia rubens L. O. Williams, Phytologia
Orizaba, 5000 ft., 9 Feb 1895, C G. Pringle 26: 128. 1973. Type: GUATEMALA: Quiche,
6135 (Holotype GH; Isotypes K, MEXU, MO, Nebaj, 6000 ft., 19 Nov 1934, A. F. Skutch
WU). 1725 (Holotype F 815804 (photos MEXU,
Rondeletia purpurea Lorence, Novon 1: 148, PTBG)). Arachnothryx.
fig. 6A-F, 7A, B. 1991. Type: MEXICO: Rondeletia rufescens B. L. Rob., Proc. Amer.
Oaxaca, Distr. de Ixtlan, Sierra de Juarez, Acad. Arts 45: 402. 1910. Type:
Ruta 175 Tuxtepec a Oaxaca, a 5 km al N. de GUATEMALA: Alta Verapaz, Coban, 4800
Vista Hermosa, 1175 m, 14 Apr 1982, D. H. ft., Mar 1881,//. von Tuerckheim 582
Lorence & R. Torres C. 4009 (Holotype (Holotype GH, n.v.; Isotypes F, K (photo
MEXU (photo PTBG); Isotypes BM, F, MO, PTBG), P (photos MEXU, PTBG), US (2
PTBG). sheets; photos MEXU, PTBG)).
Rondeletia pyramidalis Lundell, Wrightia 5: Arachnothryx.—Rondeletia gracilis Hemsl.
327. 1976. Type: MEXICO: Chiapas, San Rondeletia rufescens B. L. Rob. var. ovata B. L.
Luis, near Siltepec, 1300 m, 24 Jan 1945, E. Rob., Proc. Amer. Acad. Arts 45: 403. 1910.
Matuda 5292 (Holotype TEX-LL (photos Type: GUATEMALA: Alta Verapaz, Tactic,
MEXU, PTBG); Isotypes F, MEXU, TEX-LL, 1800 m, Mar 1903, H. von Tuerckheim 8401
n.v.). Arachnothryx. (Holotype GH, n.v.; Isotypes F (2 sheets), US
^ NOMENCLATOR OF MEXICAN AND CENTRAL
Chagres, Jan-Mar 1850, A. Fendler 180 (F, Sickingia calderoniana Standi., Publ. Field Mus.
(photo PTBG), BM, n.v., W, n.v., fragment Nat. Hist., Bot. Ser. 22: 214. 1940. Type: EL
F); same locality, Hayes s.n. (n.v.), same SALVADOR: Jayaque, finca del Dr. Zarate, 9
locality, Seemann 1073 (n.v.). COSTA RICA: Jan 1939, M. L. Van Severen 66 (Holotype F
Limon, Kuntze 1995 (n.v.); Punta Arena, 1033327 (photo PTBG); Isotype F).—Simira
Tonduz 6712 (n.v.), Tonduz 9955 (n.v.); six calderoniana (Standi.) Steyerm.
additional syntypes are cited from St. Thomas, f ) Standi., Trop. Woods
Trinidad, and Brazil. 22: 10. 1930. Calderonia klugei Standi., J.
Wash. Acad. Sci. 15:6. 1924. Type:
Schradera Vahl, Eclog. Amer. 1: 35. 1796, nom. PANAMA: Panama, near Chepo, 1924, H. C.
cons. Type: Schradera capitata Vahl, nom. Kluge 19 (Yale 7132) (Holotype US 1206424
illeg. {=Schradera invohtcrata (Sw.) K. (photos MEXU, PTBG); Isotype F (plus
Schum.) another probable isotype at F, a fruiting
Schradera blumii Dwyer & Hayden, collection from type tree, Kluge s.n.. also Yale
Phytologia 15: 59. 1967. Type: PANAMA: 7132)).—Simira klugei (Standi.) Steyerm.
Panama, Cerro Jefe in Clusia forest at Sickingia lancifoliu Lundell. Wrightia4: 50.
2700-3000 ft., 27 Jan 1966, E. L. Tyson, J. 1968. Type: GUATEMALA: Peten, Laguna
Dwyer & K. Blum 3218 (Holotype MO Petexbatun, in high forest bordering lake, on
1831023 (photo PTBG)). Escarbadero, 2 Apr 1964, C. L. Lundell 18224
Schradera costaricensis Dwyer in W. C. (Holotype TEX-LL, n.v.; Isotype US (photos
Burger & C. M. Taylor, Fieldiana, Bot., n.s. MEXU, PTBG)).—Simira salvadorensis
33: 309. 1993. Type: COSTA RICA: Alajuela, (Standi.) Steyerm.
Bijagua, G. Herrera 2004 (Holotype CR, n.v.; ', ;,. Lundell var. puberula
Isotypes F, n.v., MO). Lundell, Wrightia 4: 50. 1968. Type:
GUATEMALA: Peten, Lacandon, in bajo,
Serissa Comm. ex Juss., Gen. 209. 1789. Type: about 1 km, 700 m south. 12 Mar 1962, E.
Serissa foetida Comm. ex Juss. Contreras 3514 (Holotype TEX-LL, n.v.).
Serissa japonica (Thunb.) Thunb., Nov. Gen. —Simira salavadorensis (Standi.) Steyerm.
PI. 132, pi. 9. 1798. Type: JAPAN: Collector Sickingia maxonii (Standi.) Standi., Trop.
unknown (presumably UPS, n.v.). Lycium Woods 14: 30. 1928. Genipa maxonii Standi.,
japonicum Thunb., Nov. Act. Roy. Soc. J. Wash. Acad. Sci. 8: 642. 1918. Type:
Upsal. 3: 207. 1780. Cultivated at least in PANAMA: Panama, Canal Zone, forest along
Costa Rica and Panama. the Rio Indio de Gatiin, near sea level, 17 Feb
1911, W. R. Maxon 4848 (Holotype US
Sherardia L., Sp. PI. 1: 102. 1753. Type: 675223 (photos MEXU, PTBG)).—Simira
Sherardia arvensis L. maxonii (Standi.) Steyerm.
Sherardia arvensis L., Sp. PI. 1: 102. 1753. dlock in Hook. Icon., ser.
Type: Habitat in arvensis Scaniae, Germaniae, 4, 4: tab. 3321. 1936. Type: MEXICO:
Angliae, (Lectotype Sherard, Herb. Clifford Mexico, Distr. Temascaltepec, Luvianos, 26
33: Sherardia 1 (BM, n.v.); designated by Sep 1933, G. B. Hinton 4818 (Holotype K,
Natali in Jarvis et al., 1993: 88). Naturalized n.v.; Isotype US (photos MEXU, PTBG)).
at least in Veracruz and Costa Rica. —Simira mexicana (Bullock) Steyerm.
Sickingia mollis Lundell, Contr. Univ. Michigan
Sickingia Willd., Ges. Naturf. Freunde Berlin Herb. 4: 32. 1940. Not Sickingia mollis
Neue Schrift. 3: 445. 1801. Type not Standi., 1936. Type: MEXICO: Tabasco,
designated. [Herein treated as a synonym of Reforma, on the Rio San Pedro de Martir,
Simira Aubl.l above Balancan, 22-26 May 1939, E. Matuda
3161 (Holotype MICH, n.v.; Isotypes A, n.v.,
TEX-LL (photos MEXU, PTBG), US (photo Siderodendron Cothen., Disp. 6. 1790.
PTBG)).—Simira salvadorensis (Standi.) (=Sideroxyloides Jacq., 1763.) Type:
Steyerm. Siderodendron ferreum (Jacq.) J. F. Gmel.
Sickingia multiflora Lundell, Wrightia 5: 258. [Herein treated as a synonym of Ixora L.]
1976. Type: GUATEMALA: Peten, Tikal Siderodendi Rich, in Sagra,
National Park, Tikal, in ramonal covering the Hist. Cuba 11: 24. 1850. Type: CUBA:
ruins, on Temple IV, 11 Jul 1960, E. Without precise locality, A. Richard s.n.
Contreras 1239 (Holotype TEX-LL (photos (Holotype presumably P, n.v.).—Ixora
MEXU, PTBG); Isotype US (photos MEXU, floribunda (A. Rich.) Griseb.
PTBG)).—Simira salvadorensis (Standi.)
Steyerm. Simira AubL, Hist. PL Guiane 1: 170, t. 65.
Sickingia myriantha Standi., Pub. Field 1775, not Simira Raf, 1836. Type: Simira
Columbian Mus., Bot. Ser. 7: 27. 1930. Type: tinctoria Aubl.
COLOMBIA: Magdalena, Las Nubes road, Simira calderoniana (Standi.) Steyerm., Mem.
region of Santa Marta, 1200 m, 3 Dec 1898, New York Bot. Gard. 23: 306. 1972. Sickingia
H. H. Smith 1810 (Holotype F 138660; calderoniana Standi., Publ. Field Mus. Nat.
Isotypes MO, NY, n.v., US, n.v., W, n.v., Y, Hist., Bot. Ser. 22: 214. 1940. Type: EL
n.v.). Holtonia, Simira.—Elaeagia SALVADOR: Jayaque, finca del Dr. Zarate, 9
myriantha (Standi.) C. M. Taylor & Hammel Jan 1939, M. L. Van Severen 66 (Holotype F
Sickingia rhodoclada (Standi.) Standi., Publ. 1033327 (photo PTBG)).
Field Mus. Nat. Hist., Bot. Ser. 22: 215. 1940. Simira daricncns > IKvyer, Ann. Missouri Bot.
Exandra rhodoclada Standi., J. Wash. Acad. Gard. 67: 493. 1980. Nom. superfl. for
Sci. 13: 292. 1923. Type: MEXICO: Oaxaca, Bathysa panamensis Dwyer, Ann. Missouri
between La Venta and Niltepec, 200 ft., 14 Jul Bot. Gard. 55: 34. 1968 (based on same type).
1895, E. W. Nelson 2796 (Holotype US Type: PANAMA: Darien, Rio Ucurganti, 7
229232 (photos MEXU, PTBG); Isotype F Jul 1967, N. Bristan 1187 (Holotype MO
(photos MEXU, PTBG)).—Simira 1968219 (photo PTBG); Isotype GH, n.v.).
rhodoclada (Standi.) Steyerm.
Sickingia salvadorensis (Standi.) Standi., Field Simira klugei (Standi.) Steyerm., Mem. New
Mus. Nat. Hist., Bot. Ser. 12: 390. 1936. York Bot. Gard. 23: 302. 1972. Calderonia
Calderonia salvadorensis, J. Wash. Acad. Sci. klugei Standi., J. Wash. Acad. Sci. 15:6.
13: 290. 1923. Type: EL SALVADOR: San 1924. Type: PANAMA: Panama, near Chepo,
Salvador, Jul 1922, S. Calderon 761 1924, H. C. Kluge 19 [-Yale 7132] (Holotype
(Holotype US 1151718 (photos MEXU, US 1206424 (photos MEXU, PTBG); Isotype
PTBG)).—Simira salvadorensis (Standi.) F (photo PTBG), (plus another probable
Steyerm. isotype at F, a fruiting collection from type
Sickingia vestita Lundell, Wrightia 4: 51. 1968. tree, Kluge s.n., also Yale 7132)).
Nom. no\ Lundell, Simira maxonii (Standi.) Steyerm., Mem. New
Contrib. Univ. Mich. Herb. 4: 32. 1940. Not York Bot. Gard. 23: 306. 1972. Genipa
Sickingia vestita Standi, 1936. Type: maxonii Standi., J. Wash. Acad. Sci. 8: 642.
MEXICO: Tabasco, Reforma, on the Rio San 1918. Type: PANAMA: Panama, Canal Zone,
Pedro de Martir, above Balacan, 22-26 May forest along the Rio Indio de Ganin, near sea
1939, E. Matuda 3161 (Holotype MICH, n.v.; level, 17 Feb 1911, W. R. Maxon 4848
Isotypes A, n.v., TEX-LL (photos MEXU, (Holotype US 675223 (photos MEXU,
PTBG), US (photos MEXU, PTBG)). PTBG)).
—Simira salvadorensis (Standi.) Steyerm. Simira mexicana (Bullock) Steyerm., Mem.
New York Bot. Gard. 23: 307. 1972. Sickingia
mexicana Bullock in Hook. Icon., ser. 4, 4:
OF MEXICAN AND CENTRAL AN
tab. 3321. 1936. Type: MEXICO: Mexico, Turcz. [Herein treated as a synonym of
Distr. Temascaltepec, Luvianos, 26 Sep 1933, Rondeletia L.]
G. B. Hinton 4818 (Holotype K, n.v.; Isotype Siphonandra mexicana Turcz., Bull. Soc. Imp.
US (photos MEXU, PTBG)). Sickingia. Naruralistes Moscou 21: 581. 1848. Type:
Simira myrkmtha (Standi.) Steyerm., Mem. MEXICO: Oaxaca, without precise locality,
New York Bot. Gard. 23: 306. 1972. Sickingia 3000 ft.. H Galeotti 266-1 hi.s (Holotype
myriantha Standi., Publ. Field Columbian presumably LE, n.v.; Isotype G-DEL. n.v.
Mus., Bot. Ser. 7: 27. 1930. Type: (photos A, MO)).—Rondeletia mexicana
COLOMBIA: Magdalena, Las Nubes road, (Turcz.) Standi.
region of Santa Marta, 1200 m, 3 Dec 1898,
H. H. Smith 1810 (Holotype F 138660; Siphoma Benth., PI. Hartweg. 84. 1841. Not
Isotypes MO, NY, n.v., US, n.v., W, n.v., Y, Siphonia D. Richard ex Schreber, 1791, nor
n.v). Holtonia.—Elaeagia mvriantha Siphoma E. M. Fries, 1820. Type: Siphonia
(Standi.) C. M. Taylor & Hammel nvalis Benth. (=Lindenia Benth., 1841). See
Simira rhodoclada (Standi.) Steyerm., Mem. Darwin (1976: 427) for discussions regarding
New York Bot. Gard. 23: 307. 1972. Exandra publication of the generic names Lindcma vs.
rhodoclada Standi., J. Wash. Acad. Sci. 13: Siphonia. [Herein treated as a synonym of
292. 1923. Type: MEXICO: Oaxaca, between Augusta Pohl.]
La Venta and Niltepec, 200 ft., 14 Jul 1895, Siphonia rivalis Benth., Pi. Hartweg. 84. 1841.
E. W. Nelson 2796 (Holotype US 229232 Type: GUATEMALA: In rivis, la Vera Paz, T.
(photos MEXU, PTBG); Isotype F (photos Hartweg 581 (Holotype K, n.v. (photo MO)).
MEXU, PTBG)). Simira. —Augusta rivalis (Benth.) J. H. Kirkbr.
Simira salvadorensis (Standi.) Steyerm., Mem. Note: according to Farr et al. (1979: 1620),
New York Bot. Gard. 23: 307. 1972. although the name Siphoma was actually
Calderonia salvadorensis Standi., J. Wash. printed, it was crossed out in ink and the name
Acad. Sci. 13: 290. 1923. Type: EL Lindenia substituted, "
SALVADOR: San Salvador, Jul 1922, S copies before publication,
Calderdn 761 (Holotype US 1151718 (photos published on p. 351.
MEXU, PTBG)). Sickingia.
Solena Willd., Sp. PL 1: 961. 1798. Not Solena
Sipanea Aubl., Hist. PI. Guiane 1: 147, t. 56. Loureiro, 1790. Type: Solena longiflora
1775. Type: Sipanea pratensis Aubl. (Aubl.) Willd. (= Posoquena longiflora Aubl.)
Sipanea biflora (L. f.) Cham. & Schltdl., [Herein treated as a synonym of Posoqueria
Linnaea4: 168. 1829. Virecta biflora L. f, Aubl.]
Suppl. 17, 134. 1781. Type: SURINAM: Solena latifolia Rudge, PI. Guian. 26. 1805.
Without precise locality, Dalberg s.n. Type: FRENCH GUIANA: Martin s.n. (BM,
(presumably LINN, n.v.). n.v).—Posoqueria latifolia (Rudge) Roem.
Sipanea palustris Seem., Bot. Voy. Herald 136. & Schult.
1854. Type: PANAMA: Panama, in swamps
near Panama, Seemann 347 (Holotype BM, in Sommera Schltdl., Linnaea 9: 602. 1835. Type:
part [excluding specimen of Microlicia Sommera arborescens Schltdl.
(Melastomataceae)], n.v. (photo MO)). Sommera acuminata Oerst. ex Standi.. J. Wash.
Limnosipanea. Acad. Sci. 17: 340. 1927. Type: MEXICO:
Oaxaca, Amatlan, F. Liebmann 11712
Siphonandra Turcz., Bull. Soc. Imp. Naruralistes (Holotype C, n.v. (photo MEXU); Isotype
Moscou 21: 581. 1848. Not Siphonandra US). Arachnothryx, Rondeletia.—Rondeletia
Klotzsch 1851. Type: Siphonandra mexicana scabra Hemsl.
$ Schltdl., Linnaea 9: 602. Sommera montana L. O. Williams, Phytologia
1835. Type: MEXICO: Veracruz, Hacienda de 26: 124. 1973. Type: HONDURAS: Morazan,
la Laguna, Schiede 272 (Holotype presumably margen de la Quebrada de La Paz, en medio
HAL, n.v.; Isotypes F (photos MEXU, de la floresta de pino-liquidambar de la
PTBG), LE, n.v.(fragment F), MO (photos Montana Zanquin, 1600 m, 14 May 1950, A.
MEXU, PTBG), US (photos MEXU, PTBG)). Molina R. 2829 (Holotype F 1389080 (photo
Sommera chiapensis Brandegee, Univ. Calif. PTBG); Isotypes EAP, n.v., US (photos
Publ. Bot. 6: 196. 1915. Type: MEXICO: MEXU, PTBG)).
Chiapas, Finca Irlanda, Jun 1914, C. A. Sommera parva Lorence, Novon4: 134, fig. 9.
Purpus 7263 (Holotype UC 174914. n.v.; 1994. Type: MEXICO: Chiapas, Mun. de
Isotypes F (photos MEXU, PTBG), MO, 2 Tila, Pie de Cerro K'abana, 900 m, 1 Jul
sheets (photos MEXU, PTBG), US (photos 1982, A. Mendez Ton {A. Shilom Ton) 4367
MEXU, PTBG)). (Holotype MEXU; Isotypes BM, F, MO
Sommera donnell-smithii Standi., Contr. U.S. (photo PTBG), NY, PTBG, US, XAL).
Natl. Herb. 17: 436. 1914. Type: COSTA Sommera rivularis L. O. Williams, Phytologia
RICA: San Jose, Alajuehta, 3000 ft., Mar 26: 125. 1973. Type: COSTA RICA:
1894, J. D. Smith 6592 (Holotype US 245836 Puntarenas, Rio Esquinas, near kilometer 42,
(photos MEXU, PTBG)). 30 m, 8 Dec 1949, P. H. Allen 5431 (Holotype
Sommera dunlapii L. O. Williams, Phytologia F 1679274 (photo PTBG); Isotypes EAP, n.v.,
26: 122. 1973. Type: PANAMA: Changuinola F, US (photos MEXU, PTBG)).-
Valley, 17 Jan 1924, V. C. Dunlap 351
(Holotype F 635718 (photo PTBG); Isotype ubcordata Standi., N. Amer. Fl. 32:
US (photos MEXU, PTBG)).—Sommera 144. 1921. Type: MEXICO: Sinaloa,
donnell-smithii Standi. Colomas, 1897, J. N. Rose s.n. (Holotype US
Sommera fusca Oerst. ex Standi., J. Wash. 866799 (photos MEXU, PTBG)).—Sommera
Acad. Sci. 17: 339. 1927. Type: MEXICO: grandis (Bartl. ex DC.) Standi.
[Presumably Oaxaca] Jocoaltepec, Jun 1842, Sommera zygocalyx L. O. Williams, Phytologia
F. Liebmann 11720 [=Liebmann Rubiaceae 26: 126. 1973. Type: MEXICO: Oaxaca,
36] (Holotype C, n.v.; Isotype US (photos Distr. Choapam [=Choapan], Yaveo, Arroyo
MEXU, PTBG)). de Culebras, 430 m, 21 Mar 1938, Y. Mexia
Sommera grandis (Bartl. ex DC.) Standi., N. 9191 (Holotype F 982858 (photos MEXU,
Amer. Fl. 32: 145. 1921. Petesia grandis PTBG); Isotypes UC (photo PTBG), US
Bartl. ex DC, Prodr. 4: 395. 1830. Type: (photos MEXU, PTBG)).—Sommera
MEXICO: Without precise locality, Haenke arborescens Schltdl.
s.n. (Holotype presumably PR, n.v., fragment
G-DC, n.v. (microfiche 4: 395.1)). Spermacoce L., Sp. PI. 1: 102. 1753. Type:
Sommera guatemalensis Standi., Contr. U.S.
Natl. Herb. 17: 436. 1914. Type: Spermacoce ambigua Brandegee, Zoe 5: 180.
GUATEMALA: Alta Verapaz, Cubliquitz, 1904. Type: MEXICO: Veracruz, Cordoba,
350 m, May 1902, H. von Tuerckheim 8225 Jan 1904, C. A. Purpus 417 (Holotype UC,
(Holotype US 398487 (photos MEXU, n.v.; Isotypes MO (photo PTBG), US (photos
PTBG)). MEXU, PTBG)).—Diodia brasiliensis
Sommera mesochora Standi., Contr. U.S. Natl. Spreng.
Herb. 17: 439. 1914. Type: PANAMA: Spermacoce apiculata Willd. ex Roem. &
Chiriqui, vicinity of El Boquete, 1000-1300 Schult., Syst. Veg. 3: 531. 1818. Type:
m, 2-8 Mar 1911, W. R. Maxon 4941 BRAZIL: Without precise locality,
(Holotype US 675326 (photos MEXU, Hoffmannsegg s.n. (B-WI 2626,
PTBG)).—S
k. NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
Galeotti 2608 (Holotype BR, n.v.; Isotype K, Crusea.—Crusea hispida (Mill.) B. L. Rob.
n.v.).—Spermacoce prostrata Aubl.
Spennacoce polycephala Bartl. ex DC., Prodr. Spermacoce scabiosoides (Cham. & Schltdl.)
4: 558. 1830. Type: MEXICO: Without Kuntze, Revis. Gen. PI. 3: 123. 1898. Borreria
precise locality, Haenke s.n. (Holotype scabiosoides Cham. & Schltdl., Linnaea 3:
presumably PR, n.v., fragment G-DC, n.v. 318. 1828. Type: BRAZIL: Sellow s.n.
(microfiche 4: 558.54)).—Status uncertain. (presumably B, n.v.).
Spermacocepringlei S. Watson, Proc. Amer. Spermacoce serrulata P. Beauv., Fl. Oware
Acad. Arts 25: 152. 1890. Type: MEXICO: Benin 1: 39. tab. 23. 1805. Type: NIGERIA:
Jalisco, shaded hillsides near Guadalajara, 27 Palisot de Beauvois s.n. (presumably P,
Sep 1889, C. G. Pringle 2464 (Holotype GH, n.v.).—Diodia serrulata (P. Beauv.) G.
n.v.; Isotypes MO (photos MEXU, PTBG), Taylor
US (photos MEXU, PTBG)).—Spermacoce Spermacoce stricta Sesse & Mo?., Fl. Mexic.
ovalifolia (M. Martens & Galeotti) Hemsl. 24. 1893. Type: MEXICO: Habitat in calidis,
Spermacoce prostrata Aubl., Hist. PI. Guiane NIL, Sesse & Mocino s.n. (presumably MA,
Franc. 1: 58, tab. 20, fig. 3. 1775. Type:
FRENCH GUIANA: Aublet s.n. (Holotype P. iveolens (G. Mey.) Kuntze,
Revis. Gen. PI. 3: 123. 1898. Borreria
Spermacoce psyllioides Kunth in Humb., Bonpl. suaveolens G. Mey., Prim. Fl. Esseq. 81, pi. 1.
& Kunth, Nov. Gen. Sp. 3: 346. 1819. Type: 1818. Type: SURINAM: In arenosis
MEXICO: Without precise locality, Humboldt continentis circa Arouabischkrek, G. Meyer
& Bonplands.n. (Holotype P-Bonpland, n.v. s.n. (Holotype GOET, n.v.).
(photo MO)). Crusea psyllioides (Kunth) Spermacoce tenui Sesse & Moc., Fl. Mexic. 25.
W. R. Anderson 1893 (LaNaturaleza2,l. suppl. 25. 1897).
Spermacoce reclinata Nees, Horae Phys. Berol. Type: MEXICO: Habitat Patzcuan. Floret
50. 1820. Type: MEXICO: In regno Augusto, Sesse & Mocino s.n. (presumably
Mexicano, without precise locality or MA, n.v.).—Status uncertain.
collector (location unknown).—Spermacoce Spermacoce tenuiflora Sesse & Mo?., Fl. Mexic.
24. 1893 (La Naturaleza 2.1. suppl. 24. 1897).
Spermacoce rigida Willd. ex Roem. & Schult., Type: Habitat in Zitacuan. Floret Augusto,
Syst.Veg.3:531. 1818. Type: Sesse & Mocino s.n. (presumably MA, n.v.).
VENEZUELA: Silla de Caracas, Humboldt & —Crusea longifiora (Roem. & Schult.) W. R.
Bonpland s.n. (Holotype B-WI 2628, n.v.; Anderson
Isotype G-DC, n.v. (microfiche Spermacoce tenuifolia Sesse & Moc., PI. Nov.
IDC.7440.174:111,4)). Diodia.—Diodia Hispan. 17. 1897 (La Naturaleza 2, 1. suppl.
apiculata (Willd. ex Roem. & Schult.) K. 17. 1897). Type: MEXICO: Habitat Patzquari.
Sesse & Mocino s.n. (presumably MA, n.v.).
Spermacoce riparia Cham. & Schltdl., Linnaea
3: 355. 1828. Type: BRAZIL: Rio Negro, Spermacoce tenuior L., Sp. PI. 1: 102. 1753.
(Rincon de Gallinas), Sellow s.n. (Holotype Lectotype: Habitat in Carolina, Spermacoce
presumably B or HAL, verticillis tenuioribus Dillenius, Hortus
Elthamensis 370, tab. 277, fig. 359. 1732.
Spermacoce rubra Jacq., PI. Rar. Hort. Typotype in Herb. Sherard (OXF, n.v.), drawn
Schoenbr. 3: 3,pi. 256. 1798. Type: Cuba [but from specimens grown at Eltham; designated
probably from MEXICO], Jacquin s.n. by Rendle (1934: 333). Typification
(probable type W, n.v., fragment F). For confirmed by Verdcourt (1983).
discussion, see Anderson (1972: 110-111). Spermacoce tenuior Sesse & Mocino, Fl. Mexic.
24. 1893. Not Spermacoce tenuior L., 1753.
Staelia scabra (C. Presl) Standi., Publ. Field
Columbian Mus., Bot. Ser. 8: 385. 1931.
MA, n.v.).—St Diphragmus scaber C. Presl, Bot. Bermerk.
Spermacoce tetracocca M. Martens & Galeotti, 81. 1844. Type: MEXICO: Habitat in Mexico,
Bull. Acad. Roy. Soc. Bruxelles 112: 132. precise locality or collector not cited.
1844. Type: MEXICO: Michoacan, Santa
Maria, near Morelia, H. Galeotti 2587 Stannia H. Karst., Auswahl Gewachse
(Holotype presumably BR, n.v.; Isotype K, Venezuela 27. 1848. Type: Stannia formosa
n.v., P (photos MEXU, PTBG)).—Richardia H. Karst.
tricocca (Torr. & A. Gray) Standi, subsp. Stannia panamensis Walp. & Duchass., Linnaea
tetracocca (M. Martens & Galeotti) W. H. 23: 755. 1850. Type: PANAMA: Isthmo
Lewis & R. L. Oliv. Panamensis, Duchassaing ? (location
Spermacoce tetraquetra A. Rich, in Sagra, unknown).—Posoqueria panamensis (Walp.
Hist. Fis. Cub. Bot. 11: 29. 1850. Type: & Duchass.) Walp.
CUBA: Without precise locality, R. de la
Sagra s.n. (Holotype presumably P, n.v.; Stenostomum C. F. Gaertn., Fruct. Sem. PI. 3:
probable Isotype F (photo PTBG)). 69. 1806. Type: Stenostomum lucidum (Sw.)
Spermacoce vegeta (Standi. & Steyerm.) C. D. C. F. Gaertn.
Adams in W. C. Burger & C. M. Taylor, Stenostomum lucidum (Sw.) C. F. Gaertn.,
Fieldiana, Bot., n.s. 30: 320. 1993. Borreria Fruct. Sem. PI. 3: 69. 1806. Laugeria lucida
vegeta Standi. & Steyerm., Publ. Field Mus. Sw., Prodr. Veg. Ind. Occ. 48. 1788. Type:
Nat. Hist., Bot. Ser. 23: 21. 1943. Type: JAMAICA: Without precise locality, Swartz
GUATEMALA: Dept. Jutiapa, between s.n. (presumably S, n.v.).
Jutiapa and La Burrera, NE of Jutiapa,
800-850 m, 1 Nov 1940, P. C. Standley Steyermarkia Standi., Publ. Field Mus. Nat.
76007 (Holotype F 1091300 (photos MEXU, Hist, Bot. Ser. 22: 216. 1940. Type:
PTBG)). Steyermarkia guatemalensis Standi.
Spermacoce verticillata L., Sp. PI. 1: 102. Steyermarkia guatemalensis Standi., Publ.
1753. Type: JAMAICA: Hort. Cliff. 33 (BM, Field Mus. Nat. Hist., Bot. Ser. 22: 216. 1940.
n.v.). Borreria. For typification see Rendle Type: GUATEMALA: Izabal, Rio Dulce, 2^
(1934:331). miles west of Livingston, on south side (left.-
Spermacoce villosa Sw., Prodr. Veg. Ind. Occ. hand side going up river), alt. 1-3 m, 16 Apr
29. 1788. Type: JAMAICA: Swartzs.n. 1940, J. A. Steyermark 39520 (Holotype F
(Holotype presumably BM, n.v.). 1035008 (photo PTBG); Isotypes F 1035009,
—Mitracarpus hirtus (L.) DC. MO (photos MEXU, PTBG)).
Stachyarrhena Hook, f., Hook. Icon. PI. 1068. Stomandra Standi., Publ. Field Mus. Nat. Hist.,
1870. Type: Stachyarrhena spicata Hook. f. Bot. Ser. 23: 247. 1947. Type: Stomandra
Stachyarrhena heterochroa Standi., Contr. costaricensis Standi. [Herein treated as a
U.S. Natl. Herb. 18: 142. 1916. Type: synonym of Rustia Klotzsch.]
PANAMA: Colon, Loma de la Gloria, near Stomandra costaricensis Standi., Publ. Field
Fato (Nombre de Dios), 10-104 m, Jul-Aug Mus. Nat. Hist., Bot. Ser. 23: 247. 1947.
1911,//. Pittier 4078 (Holotype US 679174 Type: COSTA RICA: San Jose, basin of El
(photos MEXU, PTBG); Isotype F). General, 675-900 m, Mar 1940, A. F. Skutch
4777 (Holotype F 1044344 (photo PTBG);
Staelia Cham. & Schltdl., Linnaea 3: 364, pi. 3, Isotypes MO (photo PTBG), US).—Rustia
fig. 3. 1828. Type: Staelia thymoides Cham. costaricensis (Standi.) Lorence
& Schltdl.
^ NOMENCLATOR OF MEXICAN AND CENTRAL AME
Uncaria tomentosa (Willd. ex Roem. & Monte Verde, Jan-Jul 1859, Wright 242
Schult.) DC, Prodr. 4: 349. 1830. Nauclea (Lectotype GH, n.v.; Isolectotype BR, n.v.;
tomentosa Willd. ex Roem. & Schult., Syst. designated by Hamilton, 1989).—Psychotria
Veg. 5: 221. 1819. Type: COLOMBIA: In costivenia Griseb.
ripa flumen Magdalenae prope Naers, Uragoga crocea (Sw.) Kuntze, Rev. Gen. PI. 1:
Humboldt s.n. (Holotype B-WI 3910, n.v.). 299. 1891. Psychotria crocea Sw., Prodr.
Veg. Ind. Occ. 44. 1788. Type: JAMAICA:
Uragoga Baill., Adansonia 12: 323. 1879. Type: Swartzs.n. (Holotype BM, n.v.).—Palicourea
Not designated. crocea (Sw.) Roem. & Schult.
Uragoga carpinterensis Kuntze, Rev. Gen. PI. Uragoga emetica (L. f) Baill., Hist. PI. 7: 371.
2: 955. 1891. Based on Palicourea lanceolata 1880. Psychotria emetica L. f, Suppl. PI. 144.
Oerst. ex Pol., Vidensk. Meddel. Dansk 1781. Type: COLOMBIA: America
Naturhist. Foren. Kjobenhavn 1852: 39. 1853, septentrionalis provinciae Gironensis, Mutis
pro syn. Type: COSTA RICA: San Jose, in 1602 (Holotype LINN, n.v.; probable Isotype
Monte Candelaria, 2100 m, Oersted 11556 US (photo PTBG)).—Psychotria emetica L.
(Holotype C, n.v. (photo and fragment F)). f.
—Palicourea angustifolia Kunth Uragoga erythrocarpa (Schltdl.) Kuntze. Rev.
Uragoga carthagenensis (Jacq.) Kuntze, Rev. Gen. PI. 2: 960. 1891. Psychotria
Gen. PI. 2: 959. 1891. Psychotria erythrocarpa Schltdl., Linnaea 9: 595. 1834.
carthagenensis Jacq., Enum. Syst. PI. Carib. Type: MEXICO: Veracruz, Hacienda de la
16. 1760. Type: COLOMBIA: Bolivar, Laguna, Schiede & Deppe 385 (Holotype B,
vicinity of Cartagena, Jacquin s.n. (location presumably destroyed (photo & fragment F);
unknown).—Psychotria carthagenensis Isotype K, n.v )—Psychotria erythrocarpa
Schltdl.
Uragoga cordillerae Kuntze, Rev. Gen. PI. 2: Uragoga galeottiana (M. Martens) Kuntze, Rev.
955. 1891. Based on Palicourea mexicana Gen. PI. 2: 960. 1891. Palicourea galeottiana
Benth. in Oerst., Vidensk. Dansk Meddel. M. Martens, Bull. Acad. Roy. Sci. Bruxelles
Naturhist. Foren. Kjobenhavn 1852: 37. 1853. 14: 136. 1844. Type: MEXICO: Oaxaca,
Not Uragoga mexicana (Willd ex Roem. & Cordillera, 6500 ft., 1840, H. Galeotti 2602
Schult.) Kuntze, 1891. Type: COSTA RICA: (Lectotype BR (photos MEXU, PTBG):
In monte Barba, Oersted 11559 (Lectotype C, Isolectotypes K, n.v., P (photos MEXU,
n.v.; designated by Standley 1938). PTBG); designated by Taylor & Lorence,
—Palicourea padifolia (Willd. ex Roem. & 14S5).
Schult.) C. M. Taylor & Lorence Martens) C. M. Taylor & Lorence
Uragoga costaricensis (Benth.) Kuntze, Rev. Uragoga graciliflora (Benth.) Kuntze, Rev.
Gen. PI. 2: 960. 1891. Palicourea Gen. PI. 2: 960. 1891. Psychotria graciliflora
costaricensis Benth. in Oerst., Vidensk. Benth. in Oerst., Vidensk. Meddel. Dansk
Meddel. Dansk Naturhist. Foren. Kjobenhavn Naturhist. Foren. Kjobenhavn 1852: 35. 1853.
1852: 38. 1853. Type: COSTA RICA: Type: COSTA RICA: Alajuela, ad Naranjo,
Heredia, Volcan de Barba and Cerros de Apr 1847, Oersted 11611 (Holotype C
Candelaria, Oersted 11540 (Holotype C, n.v. (photos F, MEXU, PTBG); Isotype US
(photo F); Isotypes MICH, n.v., US, n.v.). (photos MEXU, PTBG)). Mapouria.
—Palicourea padifolia (Willd. ex Roem. & —Psychotria graciliflora Benth. in Oerst.
Schult.) C. M. Taylor & Lorence Uragoga granadensis (Benth.) Kuntze, Rev.
Uragoga costivenia (Griseb.) Kuntze, Rev. Gen. Gen. PI. 2: 960. 1891. Psychotria granadensis
PI. 2: 960. 1891. Psychotria costivenia Benth. in Oerst., Vidensk. Meddel. Dansk
Griseb., PI. Wright. Cuba Orient. 2: 508. Naturhist. Foren. Kjobenhavn 1852: 34. 1853.
1862. Type: CUBA: Cuba Orientali, prope Type: NICARAGUA: Granada, Oersted
11612 (Holotype C (photos MEXU, C (photos MEXU, PTBG)).—Psychotria
PTBG)).—Psychotria nervosa Sw. marginata Sw.
Uragoga grandis (Sw.) Kuntze, Rev. Gen. Pi. 2: Uragoga mexicana (Willd. ex Roem. & Schult.)
960. 1891. Psychotria grandis Sw., Prodr. Kuntze, Rev. Gen. PL 2: 961. 1891.
Veg. Ind. Occ. 43. 1788. Type: JAMAICA: Psychotria mexicana Willd. ex Roem. &
Interioris occidentalis, Swartz s.n. (Holotype Schult., Syst. Veg. 5: 189. 1819. Type:
S, n.v. (photo A)).—Psychotria grandis Sw. MEXICO: Without precise locality, Humboldt
Uragoga horizontalis (Sw.) Kuntze, Rev. Gen. & Bonplands.n. (Holotype B-WI, n.v.).
PL 1: 300. 189\. Psychotria horizontalis Sw., —Palicourea padifolia (Willd. ex Roem. &
Prodr. Veg. Ind. Occ. 44. 1788. Type: Schult.) C. M. Taylor & Lorence
SANTO DOMINGO: Without precise Uragoga micrantha (Kunth) Kuntze, Rev. Gen.
locality, Swartz s.n. (Holotype S, n.v. (photo PL 2: 961. 1891. Psychotria micrantha Kimtli
A)).—Psychotria horizontalis Sw. in Humb., Bonpl. & Kunth, Nov. Gen. Sp. PL
Uragoga ipecacuanha (Brot.) Baill., Hist. PI. 7: 3: 363, pi. 284. 1819. Type: PERU?: Without
370. 1880. Calicocca ipecacuanha Brot., precise locality, A. Bonplands.n. (Holotype P,
Trans. Linn. Soc. 6: 137. 1802. Type: n.v. (fragment F)). Mapouria.—Psychotria
BRAZIL: Without precise locality, Gomes s.n. micrantha Kunth in Humb., Bonpl. & Kunth
(presumably LISU, n.v.). Cephaelia, Evea. Uragoga microdesmia (Oerst.) Kuntze, Rev.
—Psychotria ipecacuanha (Brot.) Stokes Gen.PL2:961.1891.Psyc/^na
Uragoga justicioides (Schltdl.) Kuntze, Rev. microdesmia Oerst., Vidensk. Meddel. Dansk
Gen. PL 1: 300. 1891. Psychotria justicioides Naturhist. Foren. Kjobenhavn 1852: 36. 1853.
Schltdl., Linnaea 9: 596. 1834. Type: Type: COSTA RICA: Ad Jaris, Oersted
MEXICO: Barranca de Tioselo, Schiede & 11610 (Holotype C (photos MEXU, MO,
Deppe 259 (Holotype presumably HAL, n.v.; PTBG); Isotypes C (photos MEXU, PTBG),
Isotypes B, presumably destroyed (photo US (photo PTBG)).—Psychotria viridis Ruiz
MO), P (photos MEXU, PTBG)). & Pavon
—Psychotria hebeclada DC. Uragoga miradorensis (Oerst.) Kuntze, Rev.
Uragoga longicollis (Benth.) Kuntze, Rev. Gen. Gen. PL 2: 961. 1891. Mapouria miradorensis
PI. 2: 961. 1891. Psychotria longicollis Benth. Oerst., Amer. Centr. 17, tab. 14, fig- 9. 1863.
in Oerst., Vidensk. Meddel. Dansk Naturhist. Type: MEXICO: Veracruz, Mirador, in
Foren. Kjobenhavn 1852: 33. 1853. Type: quercetum, 3000 ft., F. Liebmann 11662
COSTA RICA: Insula San Lucas, Oersted (Lectotype C (photos MEXU, PTBG);
11621 (Lectotype C, here designated (photos Isolectotype US, n.v.; designated by
MEXU, PTBG); Isolectotype C, lower right- Hamilton, 1989).—Psychotria costivenia
hand collection, mounted on same sheet as Griseb. var. costivenia
syntype Oersted 11622). Syntypes: COSTA Uragoga morae (Pol.) Kuntze, Rev. Gen. PL 2:
RICA: In monte Aguacate, Nov 1846, Oersted 961. 1891. Psychotria morae Pol., Linnaea
11622 (C (photos MEXU, PTBG)); Aguacate, 41: 570. 1877. Type: COSTA RICA: San
Oersted 11629 (C (photos MEXU, PTBG)). Jose, Jun 1875, H. Polakowski 171 (Holotype
Oersted 11621, with two leafy fruiting B, presumably destroyed (photos GH, US)).
branches, is in excellent condition and —Psychotria quinqueradiata Pol.
represents the best choice for lectotype. Uragoga nicaraguensis (Benth.) Kuntze, Rev.
—Psychotria horizontalis Sw. Gen. PL 2: 957. 1891. Psychotria
Uragoga marginata (Sw.) Kuntze, Rev. Gen. PL nicaraguensis Benth. in Oerst., Viddensk.
2: 961. 1891. Psychotria marginata Sw., Meddel. Dansk Naturhist. Foren. Kjobenhavn
Prodr. Veg. Ind. Occ. 43. 1788. Type: 1852: 34. 1853. Type: NICARAGUA: Rio
JAMAICA: In sylvestribus, Swartz s.n. San Juan, Oersted 11634 (Lectotype C, here
(Holotype S, n.v. (photo A); probable Isotype designated (photos MEXU, PTBG)). Syntype:
A NOMENCLATOR OF MEXICAN AND CENTRAL AMERICAN RUBIACEAE
(Holotype US 678894 (photo PTBG); Isotype Xerococcus Oerst., Vidensk. Meddel. Dansk.
NY, n.v.).—Pentagonia macrophylla Benth. Naturhist. Foren. Kjobenhavn 1852: 52. 1853.
Watsonamra wendlandii (Hook.) Kuntze, Rev. Type: Xerococcus congestus Oerst. [Herein
Gen. 302. 1891. Pentagonia wendlandii treated as a synonym of Hoffmannia Sw.]
Hook., Bot. Mag./?/. 5230. 1861. Type: Xerococcus congestus Oerst., Vidensk. Meddel.
Described from cultivated material, probably Dansk. Naturhist. Foren. Kjobenhavn 1852:
from Costa Rica or Panama. Type not 52. 1853. Type: COSTA RICA: Turrialba,
designated.—Pentagonia wendlandii Hook. 3000 ft., Oersted 11776 (Holotype C, n.v.
(photos F, MO); Isotype K, n.v.).
Wittmackanthus Kuntze, Rev. Gen. PI. 1: 302. —Hoffmannia congesta (Oerst.) Dwyer
1891. Type: Wittmackanthus stanleyanus
(R. H. Schomb.) Kuntze (Basionym:
Calycophyllum Stanleyanum R. H. Schomb.)
Wittmackanthus stanleyanus (R. H. Schomb.)
Kuntze, Rev. Gen. PI. 1:302. 1891.
vanum R. H. Schomb., J.
Bot. Hook. 3: 622. 1844. Type: GUYANA:
Without precise locality, Schomburgk 356/411
B (Holotype K, n.v. (photo MO); Isotype NY,