Bolivia Checklist Bromelias

Selbyana 20(2): 201-223. 1999.
CHECKLIST OF BOLIVIAN BROMELIACEAE WITH NOTES ON SPECIES

DISTRIBUTION AND LEVELS OF ENDEMISM
THORSTEN KROMER AND MICHAEL KESSLER
I
Albrecht-von-Haller Institut ffir Pflanzenwissenschaften der Universitat Gottingen,
Abteilung Systematische Botanik, Untere Karsptile 2, Gottingen, D-37073.
E-mail forMK:106606.464@ucompuserve.com
BRUCE K. HOLST* AND HARRY E. LUTHER
The Marie Selby Botanical Gardens, 811 South Palm Ave., Sarasota, FL 34236 USA.
E-mail for BKH: bholst@virtu.sar.usf.edu
ERIC J. GOUDA
University Botanic Gardens, P.O. Box 80.162, NL-3508 TD Utrecht, The Netherlands
E-mail: E.J.Gouda@bio.uu.nl
PIERRE L. I1nsCH
Fundaci6n Amigos de la NaturalezaIBotanical Institute of the University of Bonn, Germany,
Casilla 2241, Santa Cruz, Bolivia. E-mail: pibisch@fan-bo.org
WALTER TILL
Botanical Institute of the University of Vienna, Rennweg 14, A-1030, Wien, Austria
E-mail: walter.till@univie.ac.at
ROBERTO V A.SQUEZ
Sociedad Boliviana de Botanica, Casilla 3822, Santa Cruz, Bolivia
E-mail: r.vasquez@daitec.scz.com
ABSTRACT. A discussion of Bromeliaceae diversity in Bolivia, and a checklist of the 21 genera and 281
species occuring there is presented. Each species entry in the checklist includes accepted name, compre-
hensive synonymy for all species described based on Bolivian types, additional pertinent synonymy, ele-
vation range above sea level, distribution by department, and an indication of which species are endemic
to Bolivia.
RESUMEN. Aqui se describe la diversidad de la familia Bromeliaceae en Bolivia. Se incluye un listado de
los 21 generos y 281 especies que occurren ahi. Cada especie en ellistado contiene inforrnaci6n del nombre
aceptado, sinonimia comprensiva para cada especie descrita en base a muestras Bolivianas, sinonimia
pertinente adicional, rango de elevaci6n sobre el nivel del mar, distribuci6n por departamento, y se indica
caules especies son endemicas a Bolivia.
Key words: Bromeliaceae, Bolivia, checklist, endemic
INTRODUCTION
The checklist presented here provides updated
information on the diversity and distribution of
Bromeliaceae in Bolivia. Numerous collections
in the last two decades, especially from field-
work by S.G. Beck, H. & L. Hromadnik, M.
Kessler, P. Ibisch, R. V iisquez, and T. Kromer,
have resulted in the addition of many new re-
cords for the flora since the last summary of
* Corresponding author.
Bolivian bromeliads (Smith 1969). These in-
clude seven genera (Araeococcus, Catopsis,
Deinacanthon, Greigia, Mezobromelia, Pepinia,
and Werauhia) and 125 species. Nevertheless, it
is clear that the record is incomplete, and further
surveys are needed to clarify taxonomic prob-
lems and fragmentary distribution records.
METHODS
The following herbaria were surveyed for col-
lections from Bolivia: Herbario Nacional de Bo-
201
202 SELBYANA Volume 20(2) 1999
livia, La paz (LPB), Museo de Historia Natural
Noel Kempff, Santa Cruz (USZ), The Marie
Selby Botanical Gardens, Sarasota (SEL), the
Missouri Botanical Garden, St. Louis (MO), In-
stitute of Botany, University of Vienna (WU),
Utrecht Univerisity, Utrecht (U), and the person-
al Herbarium of R. Vasquez, Santa Cruz
(V ASQ). Principal literature references used
were the Flora Neotropica Monograph Brome-
liaceae treatments (Smith & Downs 1974, 1977,
1979), a preliminary country checklist by R.
Foster (1958), a country-level family treatment
by L.B. Smith (1969), and a modem epiphyte
survey by P. Ibisch (1996). While an attempt
was made to verify specimens of all taxa in-
cluded in the list, the occurrence of some species
in Bolivia is only known to us from literature
sources or from identifications made by other
botanists. For these, an indication of the citation
or determiner source is made in the list.
The checklist was compiled from a database
prepared at the Marie Selby Botanical Gardens
which includes approximately 2250 herbarium
records. Author abbreviations follow Brummitt
and Powell (1992), abbreviations ofliterature ci-
tations follow Stafteu and Cowan (1976-1988)
for books, and Lawrence et al. (1968) for jour-
nals. Herbarium acronyms follow Holmgren et
al. (1990). All basionyms are included, jiS well
as all species-level names based on types from
Bolivia and pertinent recent synonymy. Distri-
bution at the department level in Bolivia and the
elevational range where the species occur are
listed after each species, as well as the type col-
lection, country where the type was collected,
and where possible, an additional voucher spec-
imen. The word "Endemic" is added if the spe-
cies is wholly confined to the political bound-
aries of Bolivia.
Nomenclature used in the checklist follows
that used in An Alphabetic list of Bromeliad Bi-
nomials (Luther & Sieff 1998) which recognizes
three recently created or resurrected genera: Ra-
cinaea (Spencer & Smith 1993), Werauhia
(Grant 1995), and Pepinia (Varadarajan & Gil-
martin 1988). Species which formerly belonged
to Streptocalyx can now be found in Aechmea
(Smith & Spencer 1992).
RESULTS AND DISCUSSION
Although Bolivia's species totals still rank be-
low those of other Andean countries (see Holst
1994), the number of genera (21, see TABLE 1)
is second only to Venezuela. This high number
reflects the unique biogeographical position of
Bolivia at the meeting point of the humid north-
ern and central Andes, the dry southern Andes,
TABLE 1. Species numbers of Bromeliaceae in Boliv-
ia, arranged by subfamily and genus.
Subfamily/Genus
PITCAIRNIOIDEAE
Deuterocohnia
Dyckia
Fosterella
Pepinia
Pitcairnia
Puya
Subtotal:
BROMELIOIDEAE
Aechmea
Ananas
Araeococcus
Billbergia
Bromelia
Deinacanthon
Greigia
Pseudananas
Subtotal:
TILLANDSIOIDEAE
Catopsis
Guzmania
Mezobromelia
Racinaea
Tillandsia
Vriesea
Werauhia
Subtotal:
Family total:
No. species
in Bolivia
11
6
20
1
15
57
110
10
2
1
8
4
1
3
1
30
1
17
2
12
97
9
3
141
281
No. species
endemic to
Bolivia
5
2
15
8
47
77
3
3
7
2
31
1
35
119
Amazonia, the Gran Chaco, and the Brazilian
Shield (FIGURE 1).
The 281 recorded species from the country
(TABLE 1; APPENDIX) is still well below the num-
ber to be expected. Based on a species/area re-
lationship in comparison to Peru, Ibisch (1996)
estimated a total 300-350 bromeliad species for
Bolivia. Considering the increase in species
numbers in the last decades (on average, about
four new species have been found per year since
1969), this estimate seems realistic. In 1997,
seven new species were described from Bolivia
(Till & Hromadnik 1997, Luther 1997, Ibisch et
al. 1997, Gross 1997, Ehlers 1997, Ehlers &
Hromadnik 1997, Till 1997); in 1998, six new
taxa were described (Luther 1998a, 1998b),
among them, three Greigia species; and in 1999
at least 8 new species have been published
(lbisch et al. 1999, Luther 1999, Ibisch et al.
1999, Kessler et al. 1999). Among the two latter
papers are five new Fosterella species.
Almost 43% of the Bolivian bromeliads are
endemic to the country (TABLE 1). This high
number is due in large part to terrestrial taxa,
KROMER ET AL.: BOLIVIAN BROMELIACEAE 203
BRAZIL
PERU
CHILE
250 KM
ARGENTINA
< 500 m
500 -1500 m co = COCHABAMBA
1500 - 3000 m CH = CHUQUISACA
> 3000 m TA = TARIJA
~ ~ ~ - Departmental boundaries
FIGURE 1. Bolivia. Political subdivsions and broad
elevational zones.
particularly those in the subfamily Pitcairnioi-
deae which has three genera (Deuterocohnia,
Fosterella, Puya) with centers of diversity in
Bolivia (Ibisch 1996). Eleven out of 15 Deute-
rocohnia species, 20 of approximately 25 Fos-
terella species occur in Bolivia, and 47 of the
nearly 200 species of Puya are endemic. Some-
what surprisingly, the number of Pitcairnia spe-
cies (15) is low compared to other Andean coun-
tries. This may be the result of inadequate sam-
pIing or the fact that some morphologically and
ecologically variable species (e.g., P. cardenasii,
P. crassa, P. trianae) may include more than
one species.
Another striking feature of the Bolivian bro-
meliad flora is the low number of large, green,
tank bromeliads in the Tillandsioideae and Bro-
melioideae in the genera Guzmania, Tillandsia,
Vriesea, and Aechmea. While this is in part due
to inadequate sampling, these groups have their
centers of diversity elsewhere; southeastern Bra-
zil for Aechmea and Vriesea, and the northern
Andes for Guzmania and Tillandsia.
Three departments contain more than 100 spe-
cies each, La Paz, Cochabamba, and Santa Cruz
(TABLE 2). While these appear to be the best
sampled (ca. 1700 ofthe 2250 collection records
in our database come from there), they are also
the most varied topographically. At the other
end of the sampling spectrum, our database con-
tains only two Bromeliaceae collections from
Oruro and 10 from Pando. While nearly all lo-
calities merit further collection for new Brome-
liaceae, priorities include the humid mid-eleva-
tion forests (1500-2500 m) in La Paz, semi-hu-
mid forests at the transition of dry, rainshad-
owed valleys to humid montane forest in almost
any valley system in La Paz and adjacent Co-
chabamba, and the interior portions of Ambor6
National Park in Santa Cruz. Although brome-
liads occur at low densities in the Amazonian
lowlands, a thorough search in Pando and the
Amazonian part of Beni should yield elements
of the Brazilian bromeliad flora which are not
yet known from Bolivia. Likewise, the dry Cha-
co lowland and foothill forests of Chuquisaca
and Tarija are likely to yield new discoveries. In
most need of study taxonomically are the terres-
trial and saxicolous genera, particularly Puya
and Pitcairnia, and the largely epiphytic genus
Tillandsia.
TABLE 2. Bromeliaceae species diversity, number of department- and country-level political endemic species,
and number of collection records in database used.
No. BoHvian
No. species wholly endemic species
endemic to each f'Ound in each No. collection
Department N o ~ species department department records in database
La Paz 154 23 45 714
Santa Cruz 119 20 37 575
Cochabamba 115 14 35 417
Chuquisaca 65 8 24 220
Tarija 63 12 18 146
PotOSI 30 3 12 78
Beni 28 2 67
Pando 6 10
Oruro 2 2
The database resulting from this project is
available to all those who want to contribute fur-
ther records or use it for purposes of research or
conservation. Please contact B. Holst or M.
Kessler for more information.
ACKNOWLEDGMENTS
We thank the following herbaria for providing
specimens and data used in this work: LPB, MO,
SCZ, SEL, U, and WU. The studies of P. lbisch
and R. Vasquez partially were conducted under
the Research and Development program of the
Science Department of Fundaci6n Amigos de la
Naturaleza Noel Kempff, Bolivia as part of the
Noel Kempff Climate Action Project.
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of Plant Names. Royal Botanic Gardens, Kew,
London.
Ehlers, R. 1997. Tillandsia colganii, a new species. J.
Brome1iad Soc. 47: 103-106.
Ehlers, R. and L. Hromadnik. 1997. Tillandsia hasei
R. Ehlers & L. Hromadnik spec. nov. Die Bro-
melie 1996(3): [74-75, figs.] 76-78 [backcover].
Foster, R.C. 1958. A catalogue of the ferns and flow-
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vard Univ. 184: 1-223.
Grant, J.R. 1995. The resurrection of Alcantarea and
Werauhia, a new genus. Trop. Subtrop. Pflanzen-
welt 91: 16-17.
Gross, E. 1997. Bromelienstudien. 1. Neue und wenig
bekannte Arten aus Peru und anderen Liindern
(23. Mitteilung). Trop. Subtrop. Pflanzenwelt 95:
20-21,39.
Holmgren, P.K., N.H. Holmgren and L.c. Barnett.
1990. Index Herbariorium. Part I: The Herbaria of
the World, ed. 8. New York Botanical Garden,
Bronx, New York.
Holst, B.K. 1994. Checklist of Venezuelan Bromelia-
ceae with notes on species distribution by state
and levels of endemism. Selbyana 15: 132-149.
Ibisch, P.L. 1996. Neotropische Epiphytendiversitiit-
das Beispiel Bolivien. Archiv naturwissenschaft-
licher Dissertationen 1: I-XIV, 1-357 + 107 pp.,
Martina Galunder-Verlag, Wiehl.
!bisch, P.L., E. Gross, G. Rauer & D. Rudolph 1997.
On the diversity and biogeography of the genus
Fosterella L.B. Sm., Bromeliaceae, with the de-
scription of a new species from eastern Bolivia. J.
Bromeliad Soc. 47: 211-217.
!bisch, P.L., R. Vasquez Ch. and E. Gross. 1999. New
species of Puya and Pitcairnia (Bromeliaceae)
from the Ambor6 National Park and vicinity (San-
ta Cruz, Bolivia)-a neglected centre of diversity
and endemism of Pitcairnoideae. J. Bromeliad
Soc. 49: 124-135.
!bisch, P.L., R. Vasquez and E. Gross. 1999. More
novelties of Fosterella L.B. Smith (Bromeliaceae,
Pitcairnioideae) from Bolivia. Rev. Soc. Bo1iviana
Bot. 2(2): 117-132.
Kessler, M., P.L. Ibisch and E. Gross. 1999. Fosterella
cotacajensis, una nueva especie de Bromeliaceae
de los valles secos andinos de Bolivia. Rev. Soc.
Boliviana Bot. 2(2): 111-116.
Lawrence, G.H.M., A.EG. Buchheim, G.S. Daniels
and H. Dolezal, eds. 1968. Botanico-Periodicum-
Huntianum. Hunt Botanical Library, Pittsburgh.
Luther, H.E. 1997. A showy new Fosterella from Bo-
livia. J. Bromeliad Soc. 47: 118-119.
---. 1998a. Miscellaneous new taxa of Bromelia-
ceae (XII). Selbyana 19(1): 83-90.
---. 1998b. Miscellaneous new taxa of Brome1ia-
ceae (XIII). Se1byana 19(2): 218-226.
---. 1999. Miscellaneous new taxa of Bromelia-
ceae (XIV). Selby ana 20(1): 10-15.
Luther, H.E. and E. Sieff. 1998. An Alphabetic List of
Bromeliad Binomials, 5th ed. The So-
ciety, International, Newberg.
Ribera, M.O., M. Libermann, S. Beck & M. Moraes.
1996. Vegetaci6n de Bolivia. Pp. 169-222 in K.
Mihotek, ed. Comunidades, Territorios Indfgenas
y Biodiversidad en Bolivia. CIMAR-U.A.G.R.M.,
Santa Cruz, Bolivia.
Smith, L.B. 1969. The Bromeliaceae of Bolivia. Rho-
dora 71: 35-57,220-279.
Smith, L.B. and R.J. Downs. 1974. Pitcairnioideae
(Bromeliaceae). Fl. Neotrop. Monogr. 14(1): 1-
660. Hafner Press, New York.
---. 1977. Tillandsioideae (Brome1iaceae). FI.
Neotrop. Monogr. 14(2): 663-1492. Hafner Press,
New York.
---.1979. Bromelioideae (Bromeliaceae). Fl. Neo-
trop. Monogr. 14(3): 1493-2142. New York Bo-
tanical Garden, New York.
Smith, L.B. and M.A. Spencer. 1992. Reduction of
Streptocalyx (Bromeliaceae: Brome1ioideae).
Phytologia 72: 96-98.
Spencer, M.A. and L.B. Smith. 1993. Racinaea, a new
genus of Bromeliaceae (Tillandsioideae). Phyto-
logia 74: 151-160.
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Literature: A Selective Guide to Botanical Publi-
cations and Collections with Dates, Commentaries
and Types, ed. 2. 7 volumes. Utrecht, Antwerpen,
The Hague, and Boston.
Till, W. 1997. Tillandsia samaipatensis, eine iiberse-
hene Art aus Zentralbolivien. Die Bromelie 21
1996: 36-38.
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Incomum de Deuterocohnia da Bolfvia. An un-
usual new Deuterocohnia from Bolivia. Bromelia
4(1): 19-22.
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ic realignments within the subfamily Pitcairnioi-
deae (Bromeliaceae). Syst. Bot. 13: 294-299.
ApPENDIX
Checklist of Bolivian Bromeliaceae
Department abbreviations. BE-Beni; CH-Chuquisaca; CO-Cochabamba; LP-La paz;
OR-Oruro; PA-Pando; PO-Potosi; TA-Tarija.
Type designations. HT-Holotype; IT-Isotype; LT-Lectotype; ILT-Isolectotype; CT-Clon-
otype.
Endemic = confined to the political boundaries of Bolivia.
AECHMEA
Aechmea angustifolia Poepp. & Endl., Nov. gen.
sp. PI. 2: 43. 1838.
Synonym(s): Aechmea boliviana Rusby, Bull. New
York Bot. Gard. 4: 456. 1907; Aechmea incon-
spicua Harms, Notizbl. Bot. Gart. Berlin-Dah-
lem 10: 786. 1929.
Dept.: BE, CO, LP, PA, SC.
Elevation: 150-1300 m.
Type: Poeppig 1823. HT: W (lost). Peru.
Voucher: M. Kessler et al. 8457 (LPB, SEL).
Aechmea bromel;ifolia (Rudge) Baker, in Benth. &
Hook. f., Gen. pI. 3: 664. 1883. Basionym: Til-
landsia bromeliaefolia Rudge, PI. Guian. 32, t.
50. 1807.
Synonym(s): Aechmea ellipsoidea Rusby, Mem.
New York Bot. Gard. 7: 212. 1927.
Dept.: BE, CH, LP, SC.
Type: Martin S.n. HT: BM. French Guiana.
Voucher: J. Solomon 6179 (SEL).
Aechmea castelnavii Baker, Handb. Bromel. 39.
1889.
Dept.: SC.
Type: Castelnau s.n. HT: P. Bolivia.
Voucher: In Bolivia, only known from type collec-
tion.
Aechmea distichantha Lem., Jard. Fleur. 3: t. 269.
1853.
Synonym(s): Platyaechmea distichantha (Lem.)
L.B. Sm. & Kress, Phytologia 69: 272. 1990.
Dept.: CH, CO, LP, SC, TA.
Type: Libon s.n. HT: BR? Brazil.
Voucher: B. Holst et al. 4948 (USZ, SEL).
Aechmea fuerstenbergii E. Morren & Wittm., Bel-
gique Hort. 29: 42. 1879.
Synonym(s): StreptocalyxJuerstenbergii (E. Morren
& Wittm.) E. Morren, Belgique Hort. 33: 16.
1883.
Dept.: BE, LP.
Elevation: 300-850(-1050?) m.
Type: Porte S.n. HT: LG. Brazil.
Voucher: T. Kromer (GOET, LPB, SEL).
Aechmea kuntzeana Mez, in C. DC., Monogr.
phan. 9: 208. 1896.
Dept.: BE, LP, SC.
Type: Kuntze s.n. HT: NY; IT: US. Bolivia.
Voucher: M. Kessler 3791 (LPB, SEL).
Endemic.
Aechmea longifolia (Rudge) L.B. Sm. & M.A.
Spencer, Phytologia 72: 97. 1992. Basionym:
Bromelia longifolia Rudge, Guyan. I: 31, pI.
49. (1805) 1806.
Synonym(s): Streptocalyx longifolius (Rudge) Bak-
er, Handb. Bromel. 31. 1889.
Dept.: CO.
Type: Martin s.n. HT: BM. French Guiana.
Aechmea mertensii (G. Mey.) Schult. & Schult.f.,
in Roem. & Schult., Syst. veg. 7(2): 1272.
1830. Basionyrn: Bromelia mertensii Mey.,
Prim. fl. esseq. 144. 1818.
Dept.: BE, PA.
Elevation: ca. 200 m.
Type: E.K. Rodschied S.n. HT: GOET. Guyana.
Voucher: S. Beck et al. 19320 (SEL).
Aechmea setigera Mart. ex Schult. & Schult.f., in
Roem. & Schult., Syst. veg. 7(2): 1273. 1830.
Dept.: BE, SC.
Type: C. Martius s.n. HT: M. Brazil.
Voucher: S. Beck et al. 5978 (LPB, SEL).
Aechmea tocantina Baker, Handb. Bromel. 39.
1889.
Dept.: BE, SC.
Type: Weddell 2365. HT: P. Brazil.
Voucher: R. Vasquez et al. 1813 (VASQ).
ANANAS
Ananas ananassoides (Baker) L.B. Sm., Bot. Mus.
Leafl. 7: 70. 1939. Basionym: Acanthostachys
ananassoides Baker, Handb. Bromel. 25. 1889.
Dept.: BE, LP, PA, Sc.
Type: LT: Regnell (Ill) 1261 in part. HT: P. Brazil.
Voucher: M. Moriies et al. 1633 (SEL).
Ananas comosus (L.) Merr., Interpr. Herb. amboin.
133. 1917. Basionym: Bromelia comosa L.,
Herb. ambo 21. 1754.
Dept.: Widely cultivated.
Elevation: range unknown.
Type: Type not yet selected.
Voucher: M. Lewis 37568 (SEL).
ARAEOCOCCUS
Araeococcus ftagellifolius Harms, Notizbl. Bot.
Gart. Berlin-Dahlem 10: 784. 1929.
Dept.: SC.
Type: Huebner 58. HT: B. Brazil.
Voucher: A. Jardim et at. 53 (SEL).
BILLBERGIA
Billbergia brasilieusis L.B. Sm., Arq. Bot. Estado
Sao Paulo II. 1: 105. 1943. Basionym: Helico-
dea leopoldii Versch. Hort ex Lem., Ill. Hort.
11: sub pi. 421. 1864.
Synonym(s): Billbergia leopoldii (Versch. Hort. ex
Lem.) Linden ex Houllet, Rev. Hort. 41: 87.
1869; non K. Koch 1857.
Dept.: Unknown.
Elevation: Unknown.
Type: De Vos in Verschaffett Hortus s.n. Descrip-
tion and illustration. Brazil.
Voucher: D. Beadle SU 126 (SEL).
Billbergia cardenasii L.B. Sm., Phytologia 4: 382.
1953.
Dept.: CO?
Elevation: Unknown.
Type: Cardenas 4907. HT: US. Bolivia.
Voucher: D. Beadle SU 029 (SEL).
Endemic.
Billbergia decora Poepp. & EndL, Nov. gen. sp. pI.
2: 42, t. 157. 1838.
Synonym(s): Billbergia boliviensis Baker, Handb.
Bromel. 81. 1889.
Dept.: LP.
Elevation: 2400 m.
Type: Poeppig s.n. HT: W (lost). Peru.
Voucher: H. Rusby 2853 (NY). Cited for Bolivia in:
Fi. Neotrop. Monogr. 14(3): 2021. 1979.
Billbergia meyeri Mez, Bot. Jahrb. Syst. 30: 148.
1902.
Dept.: Sc.
Type: Pilger 705. HT: B. Brazil.
Voucher: R. Guillen 1910 (SEL).
Billbergia microlepis L.B. Sm., Phytologia 8: 504.
1963.
Dept.: CO, LP.
Type: B. Krukoff 11158. HT: GH. Bolivia.
Voucher: A. Gentry et al. 44301 (SEL).
Endemic.
Billbergia robert-readii E. Gross & Rauh, Trop.
SUbtrop. Pflanzenwelt 60: 89. 1987.
Dept.:BE,LP.
Type: Rauh 26163. lIT: HEID. Peru.
Voucher: T. Kromer 137 (GOET, LPB, SEL).
BiUbergia velascana M. cardenas, Bull. Bromeliad
Soc. 7: 35. 1957.
Dept.: SC.
Type: Father Lorenzo Hammerschmidt 5486. HT:
Cardenas Herbarium (LlL ?). Bolivia.
Voucher: D. Beadle SU 029A (SEL).
Endemic.
BiUbergia violacea Beer, Fam. Bromel. 116. 1856.
Dept.: CO, SC.
Type: Appun S.n. HT: B? "Brazil" Guyana.
Voucher: R. Vasquez & Ric 1845 (VASQ).
Bromelia balansae Mez, in Mart., FI. bras. 3(3):
191. 1891.
Dept.: BE, Sc.
Type: Balansa 608. HT: G. Paraguay.
Voucher: S. Beck 20689 (LPB, SEL).
BROMELIA
Bromelia hieronymi Mez, in Mart., Fl. bras. 3(3):
199. 1891.
Dept.: SC, TA.
Type: Hieronymus & Lorentz s.n. HT: B; IT: US.
Argentina.
Voucher: M. Nee & G. Coimbra S. 39933 (LPB).
Bromelia serra Griseb., Abh. Konigi. Ges. Wiss.
Gottingen 24: 328. 1879.
Dept.: BE, LP, SC.
Type: Lorentz & Hieronymus s.n. LT: GOET. Ar-
gentina.
Voucher: M. Moriies et at. 1631 (LPB, SEL).
Bromelia vilIosa Mez, Bot. Jahrb. Syst. 30 (BeibI.
67): 3. 1901.
Dept.: SC.
Elevation: 400 m.
Type: Glaziou 22191. HT: B. Brazil.
Voucher: Fuentes 1148 (USZ).
CATOPSIS
Catopsis sessilillora (Ruiz & Pav.) Mez, in C. DC.,
Monogr. phan. 9: 625. 1896. Basionym: Til-
landsia sessiliflora Ruiz & Pav., Fl. peruv. 3:
42, pl. 271, fig. b. 1802.
Dept.: LP.
Elevation: 1250-1300 In.
Type: Ruiz & Pavon S.n. HT: M; IT: B. Peru.
DEINACANTHON
Deinacantbon urbanianum (Mez) Mez, in C. DC.,
Monogr. phan. 9: 13. 1896. Basionym: Rhodos-
tachys urbaniana Mez, in Mart., Fl. bras. 3(3):
182. 1891.
Dept.: sc.
Elevation: 400 m.
Type: Hieronymus s.n. HT: G; IT: B, CORD, GH,
US. Argentina.
Voucher: G. Navarro 2221 (USZ). Det. G. Navarro.
DEUTEROCOHNIA
Deuterocohnia bracteosa W. Till & L. Hrom., Bro-
melia 4(1): 19. 1997.
Dept.: CH.
Elevation: 3000 m.
Type: L. Hromadnik 19029. HT: WU. Bolivia.
Voucher: Only known from type collection.
Endemic.
Deuterocohnia brevifolia (Griseb.) M.A. Spencer
& L.B. Sm., Bradea 6: 144. 1993. Basionym:
Navia brevifolia Griseb., Abh. Konigl. Ges.
Wiss. Gottingen 24: 332. 1879.
Synonym(s): Abromeitiella brevifolia (Griseb.) A.
Cast., Anales Mus. Nac. Hist. Nat. Buenos Ai-
res 36: 371. 1931; Abromeitiella pulvinata
Mez, Bot. Archiv. 19: 460. 1927.
Dept.: TA.
Type: Lorentz & Hieronymus 947 (in part). HT:
GOET; IT: CORD. Argentina.
Voucher: S. Beck 14276 (SEL).
Deuterocohnia brevispicata Rauh & L. Hrom.,
Trop. SUbtrop. Pflanzenwelt 65: 5. 1988.
Dept.: CH.
Elevation: 1200-1300(-2050) m.
Type: Hromadnik 5213. HT: HElD. Bolivia.
Voucher: D. Cathcart B52 (SEL).
Endemic.
Deuterocohnia digitata L.B. Sm., Phytologia 18:
137. 1969.
Dept.: CO, PO, SC, TA.
Type: Castellanos s.n. HT: BA; IT: US. Argentina.
Voucher: 1. Vargas 3185 (SEL).
Deuterocohnia glandulosa E. Gross, Trop. Subtrop.
Pflanzenwelt 75: 5. 1990.
Dept.: TA.
Elevation: 1200 m.
Endemic.
Deuterocohnia longipetala (Baker) Mez, in Mart.,
Fl. bras. 3(3): 506. 1894. Basionym: Dyckia
longipetala Baker, Handb. Bromel. 135. 1889.
Dept.: CO, SC, TA.
Type: Humboldt & Bonpland 3595. HT: B. Peru.
Voucher: J. Balcazar 100 (USZ, SEL).
Deuterocohnia lorentziana (Mez) M.A. Spencer &
L.B. Sm., Bradea 6: 144. 1993. Basionym: Pit-
caimia iorentzina Mez, in C. DC., Monogr.
phan. 9: 974. 1896.
Synonym(s): Abromeitiella lorentziana (Mez) A.
Cast., Lilloa 10: 459. 1944.
Dept.: PO, TA.
Type: Lorentz s.n. HT: B. Argentina.
Voucher: W. Till 62a (WU).
Deuterocohnia lotteae (Rauh) M.A. Spencer &
L.B. Sm., Bradea 6: 145. 1993. Basionym:
Abromeitiella lotteae Rauh, Trop. Subtrop.
Dept.: TA.
Elevation: 2700 m.
Voucher: W. Till 62 A (WU).
Endemic.
Deuterocohnia meziana Kuntze ex Mez, in C. DC.,
Monogr. phan. 9: 465. 1896.
Dept.: SC.
Type: O. Kuntze s.n. HT: NY. Brazil.
Voucher: M. Nee 35601 (LPB).
Deuterocohnia scapigera (Rauh & L. Hrom.) M.A.
Spencer & L.B. Sm., Bradea 6: 141-145. 1993.
Basionym: Abromeitiella scapigera Rauh & L.
Hrom., Trop. SUbtrop. Pflanzenwelt 60: 5.
1987.
Dept.: CH, PO.
Voucher: M. Kessler et at. 5085 (LPB, SEL).
Endemic.
Deuterocohnia strobilifera Mez, Repert. Spec.
Nov. Regni Veg. 3: 15, 1906.
Dept.: CH, PO, TA.
Type: Fiebrig 2933. HT: B. Bolivia.
Voucher: S. Beck 14122 (LPB). Det. G. Varadarajan.
DYCKIA
Dyckia crassifolia Rauh, Trop. Subtrop. Pflanzen-
welt 79: 10. 1991.
Dept.: SC.
Elevation: 1200 m.
Type: W. Rauh 40670. HT: HElD. Bolivia.
Endemic.
Dyckia ferox Mez, in C. DC., Monogr. phan. 9:
511. 1896.
Synonym(s): Dyckia meziana 0. Kuntze, Revis.
gen. pI. 3(2): 302. 1898.
Dept.: SC.
Elevation: 1200 m.
Type: Kuntze s.n. HT: NY. Paraguay.
Voucher: M. Cardenas 2679 (GH). Cited for Bolivia
in: Fl. Neotrop. Monogr. 14(1): 543. 1974.
Dyckia ferrnginea Mez, in C. DC., Monogr. phan.
9: 533. 1896.
Dept.: Sc.
Elevation: 300 m.
Type: Kuntze S.n. HT: NY. Brazil.
Voucher: P. lbisch et ai. 93.0614 (FR, LPB, SEL).
Dyckia gracilis Mez, in C. DC., Monogr. phan. 9:
516. 1896.
Dept.: Sc.
Type: D'Orbigny 1006. HT: P; IT: G. Bolivia.
Voucher: A. Fuentes 1151 (LPB, USZ).
Dyckia leptostachya Baker, Gard. Chron. 1884(2):
198. 1884.
Synonym(s): Dyckia boliviensis Mez in C. DC.,
Monogr. phan. 9: 524. 1896.
Dept.: CH, sc.
Elevation: 350-500(-1850?) m.
Type: Kew Hortus. HT: K. Brazil.
Voucher: T. Killeen et al. 7030 (SEL).
Dyckia pulquinensis Wittm., Rijks-Herb. 29: 88.
1916.
Dept.: SC.
Elevation: 1500 m.
Type: Herzog 1849. HT: L; IT: B. Bolivia.
Voucher: G. Balcazar 35 (USZ).
Endemic.
FOSTERELLA
Fosterella albicans (Griseb.) L.B. Sm., Phytologia
7: 171. 1960. Basionym: Cottendorfia albicans
Griseb., Abh. Konigl. Ges. Wiss. Gottingen 24:
330. 1879.
Synonym(s): Lindmania alibcans -(Griseb.) Mez in
C. DC., Monogr. phan. 9: 537. 1896.
Dept.: CH, CO, LP, SC.
Type: Lorentz & Hieronymus 502. HT: GOET; IT:
B, CORD. Argentina.
Fosterella cauIescens Rauh, Trop. SUbtrop. Pflan-
zenwelt 31: 23. 1979.
Dept.: LP.
Elevation: 1200 m.
Type: W. Rauh 40579A. HT: HEID. Bolivia.
Endemic.
Fosterella chaparensis Ibisch, Vasquez & Gross,
Rev. Soc. Boliviana Bot. 2(2): 118. 1999.
Dept.: CO.
Elevation: 600 m.
Type: R. Vasquez & M.l. Vasquez 2792. HT: LPB;
IT: FR, USZ. Bolivia.
Endemic.
Fosterella chiquitana Ibisch, Vasquez & Gross,
Dept.: Sc.
Elevation: 500 m.
Type: P. Ibisch 98.0125. HT: LPB; IT: USZ. Boliv-
ia.
Endemic.
Fosterella cotacajensis M. Kessler, P. Ibisch & E.
Gross. Rev. Soc. Boliviana Bot. 2(2): 111.
1999.
Dept.: CO, LP.
Type: Kessler et al. 9620. HT: LPB; IT: SEL. Bo-
livia.
Voucher: M. Kessler et al. 5531 (LPB).
Endemic.
Fosterella elata H. Luther, Selbyana 5: 310. 1981.
Dept.: LP.
Type: Luer et al. s.n. HT: SEL; IT: US. Bolivia.
Endemic.
Fosterella floridensis Ibisch, Vasquez & Gross,
Dept.: SC.
Type: 1bisch & Ibisch 97.83. HT: LPB; IT: FR, USZ.
Bolivia.
Voucher: Vasquez 1808 (JBSCZ).
Endemic.
Fosterella gracilis (Rusby) L.B. Sm., Phytologia 7:
171. 1960. Basionym: Catopsis gracilis Rusby,
Bull. New York Bot. Gard. 6: 489. 1910.
Synonym(s): Lindmania gracilis (Rusby) L.B. Sm.,
Contr. Gray Herb 104: 78. 1934.
Dept.: LP.
Elevation: 500 m.
Type: R.S. Williams 738. HT: NY. Bolivia.
Voucher: White 1086 (NY). Cited in FI. Neotrop.
Monogr. 14(1): 209. 1974.
Endemic.
Fosterella graminea (L.B. Sm.) L.B. Sm., Phyto-
logia 7: 171. 1960. Basionym: Lindmania gra-
minea L.B. Sm., Lilloa 14: 93. 1948.
Dept.: LP.
Elevation: 7oo-900(-1450?) m.
Type: Buchtien 417. HT: US. Bolivia.
Endemic.
Fosterella heterophylla Rauh, Trop. Subtrop.
Dept.: LP.
Elevation: 1000 m.
Type: W. Rauh 40583A. HT: HEID. Bolivia.
Endemic.
Fosterella latifolia Ibisch, Vasquez & Gross, Rev.
Soc. Bol. Bot. 2(2): 123. 1999.
Dept.: SC.
Type: P. Ibisch 98.0098. HT: LPB; IT: FR, USZ.
Bolivia.
Voucher: Ibisch 98.0173 (USZ).
Endemic.
Fosterella pearcei (Baker) L.B. Sm., Phytologia 7:
172. 1960. Basionym: Cottendorfia pearcei
Baker, Handb. Bromel. 128. 1889.
Synonym(s): Lindmania pearcei (Baker) Mez in C.
DC., Monogr. phan. 9: 537. 1896.
Dept.: LP.
Type: Pearce s.n. HT: BM. Bolivia.
Voucher: B. Krukoff 10328 (GH, LPB, NY). Cited
for Bolivia in Fl. Neotr. 14(1): 202. 1974.
Endemic.
Fosterella penduliflora (C.H. Wright) L.B. Sm.,
Phytologia 7: 172. 1960. Basionym: Catopsis
pendulifiora C.H. Wright, Bull. Misc. Inform.
1910: 197. 1910.
Synonym(s): Lindmania pendulifiora (C.H. Wright)
Stapf, Bot. Mag. 150: pi. 9029. 1924.
Dept.: CH, LP, SC, TA.
Type: Forget S.n. HT: K. Peru.
Fosterella rojasii (LB. Sm.) LB. Sm., Revista Ar-
gent. Agron. 7: 162. 1940. Basionym: Lind-
mania rojasii LB. Sm., Phytologia 7: 172.
1960.
Dept.: Sc.
Type: Rojas 6771. HT: GH. Paraguay.
Voucher: H. Amerhauser 5 (WU).
Fosterella rusbyi (Mez) LB. Sm., Phytoiogia 7:
172. 1960. Basionym: Lindmania rushyi Mez,
Bot. J ahrb. Syst. 30 (BeibL 67): 6. 1901.
Dept.: LP.
Elevation: 850-1500(-1800?) m.
Type: Bang 2571. HT: B; IT: BM, F, G, GH, K, NY,
US. Bolivia.
Voucher: M. Lewis 36936 (MO, SEL). "cf.".
Fosterella schidosperma (Baker) LB. Sm., Phy-
tologia 8: 500. 1963. Basionym: Chlorophytum
schidospermum Baker, J. Linn. Soc., Bot. 15:
326. 1876.
Synonym(s): Cottendorfia rushyi Baker, BulL Tor-
rey Bot. Club 29: 697. 1902, non Lindmania
rusbyi Mez, 1901; Lindmania schidosperma
(Baker) P. Ravenna, Wrightia 7(3): 253. 1983.
Dept.: CO, LP.
Type: Lechler 2382. HT: CGE?; IT: G? Peru.
Voucher: M. Kessler et at. 7786 (LPB, SEL). "cf.".
Fosterella spectabilis H. Luther, J. Bromeliad Soc.
47: 118. 1997.
Dept.: Sc.
Elevation: 1700 m.
Type: Cathcart B 17. HT: SEL; IT: LPB, MO, NY,
US, WU. Bolivia.
Endemic.
Fosterella vasquezii E. Gross & P. Ibisch, J. Bro-
meliad Soc. 47: 212. 1997.
Dept.: Sc.
Type: Ibisch et at. 930652. HT: HElD; IT: LPB.
Bolivia.
Voucher: F. Mamani & A. Jardim 126J(USZ, SEL).
Endemic.
Fosterella villosula (Harms) LB. Sm., Phytologia
7: 172. 1960. Basionym: Lindmania villosula
Harms, NotizbL Bot. Gart. Berlin-Dahlem 10:
794. 1929.
Dept.: CO, LP, Sc.
Type: Werdermann 2120. HT: B. Bolivia.
Endemic.
FostereHa weddelliana (Brongn. ex Baker) LB.
Sm., Phytologia 7: 172. 1960. Basionym: Cot-
tendorfia weddelliana Brongn. ex Baker,
Handb. BrameL 129. 1889.
Synonym(s): Lindmania weddelliana (Brongn. ex
Baker) Mez in C. DC., Monogr. phan. 9: 538.
1896.
Dept.: LP.
Type: Weddell 4233. HT: P. Bolivia.
Voucher: M. Lewis 40425 (LPB, MO, SEL).
Endemic.
GREIGIA
Greigia atrocastanea H. Luther, Selbyana 19: 218.
1998 [1999].
Dept.: LP.
Elevation: (2500?-)3300 m.
Type: E. Garcia 1977. HT: SEL; IT: LPB. Bolivia.
Endemic.
Greigia cochabambae H. Luther, Selbyana 19: 83.
1998.
Dept.: CO, LP.
Type: M. Kessler et al. 7111. HT: SEL; IT: LPB.
Bolivia.
Endemic.
Greigia kessleri H. Luther, Selbyana 19: 83. 1998.
Dept.: CO, LP.
Type: M. Kessler et al. 9067. HT: SEL; IT: LPB.
Bolivia.
Endemic.
GUZMANIA
Guzmania besseae H. Luther, Selbyana 5: 310.
1981.
Dept.: CO, LP, Sc.
Type: L. Besse et at. 669. HT: SEL; IT: US. Bolivia.
Voucher: P. lbisch & c. 1bisch 93.0111 (LPB, SEL).
Guzmania bipartita L.B. Sm., Phytologia 6: 437.
1959.
Dept.: LP, Sc.
Type: Asplund 19474. HT: S. Ecuador.
Guzmania calothyrsus Mez, in C. DC., Monogr.
phan. 9: 910. 1896.
Dept.: BE, CO, LP, SC.
Type: Poeppig 1224. HT: W (lost). Peru.
Voucher: T. Kriimer et at. 57 (LPB, SEL).
Guzmania danielii L.B. Sm., Phytologia 4: 360.
1953.
Dept.: CO, LP?
Type: Daniel 3350. HT: OH; IT: US. Colombia.
Guzmania gioriosa (Andre) Andre ex Mez, in C.
DC., Monogr. phan. 9: 922. 1896. Basionym:
Caraguata gloriosa Andre, Enum. BromeI. 5.
13 Dec. 1888; Rev. Hort. 60: 565. 16 Dec.
1888.
Dept.: CO, LP.
Type: Andre 3791. HT: K. Ecuador.
Guzmania killipiana L.B. Sm., Contr. Gray Herb.
98: 29. 1932.
Dept.: CO, LP.
Type: Killip & Smith 25630. HT: US. Peru.
Guzmania Jingulata (L.) Mez, in C. DC., Monogr.
phan. 9: 899. 1896. Basionym: Tillandsia lin-
gulata L., Sp. pI. 1: 286. 1753.
Dept.: LP, SC.
Type: Plumier s.n. HT: P. West Indies.
Voucher: M. Saldias & L. Arroyo 3572 (SEL).
Guzmania marantoidea (Rusby) H. Lutber, Sel-
byana 19: 86. 1998. Basionym: Tillandsia mar-
antoidea Rusby, Descr. S. Amer. pI. 4. 1920.
Synonym(s): Guzmania rio-nievensis Rauh, Trop.
Subtrop. Pflanzenwelt 50: 22. 1984.
Dept.: CO, LP.
Type: Bang 2570. HT: NY; IT: US. Bolivia.
Guzmania melinonis Regel, Gartenfiora 34: 116.
1885.
Dept.: CO, LP, Sc.
Type: Linden in Petrograd Hortus s.n. HT: LE.
French Guiana.
Guzmania monostachia (L.) Rusby ex Mez, in C.
DC., Monogr. phan. 9: 905. 1896. Basionym:
Renealmia monostachia L., Sp. pI. 1: 287.
1753.
Dept.: LP.
Elevation: 1300 m.
Type: Plumier S.n. HT: P. West Indies.
Guzmania morreniana (Linden Hortus) Mez, in C.
DC., Monogr. phan. 9: 932. 1896. Basio-
nym: Massangea morreniana Linden Hortus,
Cat. Expos. Brux. 1880.
Dept.: CO, LP.
Type: E. Morren Hortus s.n. CT: LG. Soutb Amer-
ica.
Guzmania remediosensis E. Gross, Trop. SUbtrop.
Dept.: LP.
Elevation: Unknown.
Type: M. Miyagawa s.n. HT: HElD. Bolivia.
Endemic.
Guzmania retusa L.B. Sm., Fieldiana, Bot. 28(1):
143. 1951.
Dept.: CO, LP.
Type: J. Steyermark 60677. HT: E Venezuela.
Guzmania roezlii (E. Morren) Mez, in C. DC.,
Monogr. phan. 9: 948. 1896. Basionym:
Schlumbergeria roezli E. Morren, Belgique
Hort. 28: 311. 1878.
Dept.: BE, CO, LP.
Type: Roezl in Morren Hortus S.n. CT: LG. Peru.
Voucher: T. Kromer 185 (LPB, SEL).
Guzmania sphaeroidea (Andre) Andre ex Mez, in
C. DC., Monogr. phan. 9: 942. 1896. Basio-
nym: Caraguata sphaeroidea Andre, Enum.
Bromel. 6. 13 Dec. 1888; Rev. Hort. 60: 566.
16 Dec. 1888.
Dept.: CO, LP.
Type: Andre 2685. HT: K. Colombia.
Guzmania squarrosa (Mez & Sodiro) L.B. Sm. &
Pittendr., J. Wash. Acad. Sci. 43: 403. 1953
[1954]. Basionym: Thecophyllum squarrosum
Mez & Sodiro, Bull. Herb. Boissier, ser. 2, 4:
877. 1904.
Dept.: CO, LP.
Type: Sodiro 47. HT: B. Ecuador.
Guzmania tarapotina Ule, Verh. Bot. Vereins Prov.
Brandenburg 48: 147. 1907.
Dept.: LP.
Elevation: 600(-2350) m.
Type: Ule 6683. HT: B. Peru.
MEZOBROMELIA
Mezobromelia capituIigera (Griseb.) J.R. Grant,
Phytologia 74: 428. 1993. Basionym: Tillandsia
capituligera Griseb., Cat. pI. Cub. 254. 1866.
Dept.: CO, LP.
Type: Wright 3275. HT: GOET; IT: GH. Cuba.
Mezobromelia pleiosticha (Griseb.) Utley & H.
Lutber, Ann. Missouri Bot. Gard. 78: 270.
1991. Basionym: Tillandsia pleiosticha Griseb.,
Nachr. Konigl. Ges. Wiss. Georg-Augusts-
Univ. [1]: 19. 1864.
Dept.: LP.
Elevation: 1300 m.
Type: A. Fendler 1514. HT: GOET. Venezuela.
PF;PINIA
Pepinia caricifolia (Mart. ex Schult. f.) G.S. Varad.
& Gilmartin, Syst. Bot. 13: 297. 1988. Basion-
KROMER ET AL: BOLIVIAN BROMELIACEAE 211
ym: Pitcairnia caricifolia Mart. ex Schult. f., in
Roem. & Schult., Syst. Veg. 7(2): 1242. 1830.
Dept.: PA, Sc.
Voucher: T. Killeen et al. 5448 (SEL).
PITCAIRNIA
Pitcairnia amboroensis Ibisch, Vasquez, Grass &
Kessler, J. Bromeliad Soc. 49: 129. 1999.
Dept.: Sc.
Type: P. lbisch & c. lbisch 97.0004A. HT: LPB.
Bolivia.
Voucher: M. Kessler et al. 8660 (GOET, LPB, SEL).
Endemic.
Pitcairnia aurea Rusby ex L.B. Sm., Phytologia
19: 286. 1970.
Dept.: LP.
Elevation: 900 m.
Type: H. Rusby 665. HT: NY. Bolivia.
Endemic.
Pitcairnia brittoniana Mez, in C. DC., Managr.
phan. 9: 451. 1896.
Synonym(s): Hepetis brittoniana (Mez) Mez in C.
DC., Monogr. phan. 9: 973. 1896.
Dept.: CO(?), LP.
Type: Bang 592. HT: US; IT: NY. Bolivia.
Voucher: J. Solomon 9281 (LPB, SEL).
Pitcairnia cardenasii L.B. Sm., Phytologia 8: 505.
1963.
Dept.: CH, LP, SC.
Voucher: Guardia et al. 230 (SEL).
Endemic.
Pitcairnia crassa L.B. Sm., Lilloa 14: 94. 1948.
Dept.: LP, SC.
Type: Cardenas 3690. HT: GH. Bolivia.
Voucher: T. Kromer et al. 126 (LPB, SEL).
Endemic.
Pitcairnia divaricata Wittm., Rijks-Herb. 29: 81.
1916.
Dept.: LP, SC, TA?
Elevation: 900-1300(-1900) m.
Voucher: S. Beck 1765 (LPB).
Endemic.
Pitcairnia inermis (E. Mey.) E. Mey. ex Schult .. &
Schult. f., in Roem. & Schult., Syst. veg. 7(2):
1238. 1830. Basionym: Pourretia inermis E.
Mey., in C. Pres!, Re1iq. haenk. 1(2): 123. 1827.
Dept.: Unknown.
Elevation: Unknown.
Type: Haenke S.n. HT: PRo Peru.
Voucher: Pearce 521 (K). Cited for Bolivia in Fl.
Neotrap. Monogr. 14(1): 326. 1974.
Pitcairnia lanuginosa Ruiz & Pav., Fl. peruv. 3:
35. 1802; Phytologia 15: 195. 1967.
Synonym(s): Orthophytum lanuginosum (Ruiz &
Pav.) Beer, Fam. Bramel. 70. 1856; Pitcairnia
sessiliflora Rusby, Bull. New York Bot. Gard.
4: 457. 1907.
Dept.: CO, LP, SC.
Type: Ruiz & Pavon S.n. HT: MA. Peru.
Voucher:J. Solomon 14173 (LPB, SEL).
Pitcairnia longissimiflora Ibisch, Vasquez, &
Grass, J. Brameliad Soc. 49: 127. 1999.
Dept.: Sc.
Elevation: 1000 m.
Type: P. lbisch & c. lbisch 97.0097. HT: LPB. Bo-
livia.
Voucher: P. lbisch & c. Ibisch 98.0225 (USZ).
Endemic.
Pitcairnia multiramosa Mez, in C. DC., Monogr.
phan. 9: 419. 1896.
Synonym(s): Hepetis multiramosa (Mez) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: CH.
Elevation: 1900 m.
Voucher: M. Cardenas 5732 (US). Cited in FI. Neo-
trap. Monogr. 14(1): 306. 1974.
Endemic.
Pitcairnia odontopoda Baker, Handb. Bramel. 93.
1889.
Synonym(s): Hepetis odontopoda (Baker) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: LP.
Type: Pearce s.n. HT: K. Bolivia.
Voucher: Eyerdam 25357 (F, UC). Cited in Fl. Neo-
trap. Monogr. 14(1): 319. 1974.
Pitcairnia paniculata (Ruiz & Pav.) Ruiz & Pav.,
Syst. veg. fl. peruv. chi!. 3: 36. 1802. Basion-
ym: Pourretia paniculata Ruiz & Pav., Syst.
veg. fl. peruv. chi!. 1: 81. 1798.
Synonym(s): Pitcairnia biattenuata Rusby, Bull.
New York Bot. Gard. 4: 457. 1907.
Dept.: CH, CO?, LP, Sc.
Elevation: (500)1750-2000(-2550) m.
Type: Ruiz & Pavon s.n. Herbarium of deposit un-
known. Peru.
Pitcairnia platystemon Mez, in C. DC., Monogr.
phan. 9: 421. 1896.
Synonym(s): Hepetis platystemon (Mez) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: Sc.
Type: D'Orbigny 937. HT: P. Bolivia.
Voucher: P. Ibisch & Nowicki 990124 (USZ).
Endemic.
Pitcairnia riparia Mez, Repert. Spec. Nov. Regni
Veg. 12: 416. 1913.
Dept.: LP.
Elevation: 1300 m.
Type: Weberbauer 6156. HT: B. Peru.
Voucher: M. Kessler 9806 (GOET, LPB).
Pitcairnia trianae Andre, Enum. Bromel. 4. 13 Dec
1888; Rev. Hort. 60: 564. 16 Dec 1888.
Synonym(s): Hepetis trianae (Andre) Mez in C.
DC., Monogr. phan. 9: 974. 1896.
Dept.: CO, LP.
Voucher: T. Kromer 134 (LPB, SEL).
PSEUDANANAS
Pseudananas sagenarius (Arruda) Camargo, Revis-
ta Agric. (Piracicaba) 14(7, 8): reprint p. 4.
1939. Basionym: Bromelia sagenaria Arruda,
Diss. pI. Brazil 13. 1810.
Dept.: BE, LP, SC.
Type: Original description.
PuYA
Puya alata L.B. Sm., Phytologia 7: 422. 1961.
Dept.: PO.
Elevation: 1000 m.
Type: Cardenas 2851. HT: F. Bolivia.
Endemic.
Puya alba L.B. Sm., Phytologia 5: 49. 1954.
Dept.: TA.
Elevation: 800 m.
Type: Cardenas 4952a. HT: US. Bolivia.
Endemic.
Puya atra L.B. Sm., Contr. U.S. Natl. Herb. 29:
536. 1954.
Dept.: CO, LP, Sc.
Type: M.B. Foster 2551. HT: US. Bolivia.
Endemic.
Puya brittoniana Baker, Handb. Bromel. 124.
1889.
Dept.: LP.
Voucher: C. Varadarajan 1466 (LPB). Del. G. Var-
adarajan.
Endemic.
Puya cardenasii L.B. Sm., Lilloa 14: 94. 1948.
Dept.: CO.
Type: Cardenas 4082. HT: US; IT: GH. Bolivia.
Voucher: C. Varadarajan 1442 (LPB). Det. G. Var-
adarajan.
Endemic.
Puya claudiae Ibisch, Vasquez & Gross, J. Bro-
meliad Soc. 49: 132. 1999.
Dept.: Sc.
Elevation: 2000 m.
Type: P. lhisch & c. 1hisch 980040. HT: LPB. Bo-
livia.
Endemic.
Puya cristata L.B. Sm., Lilloa 14: 95. 1948.
Dept.: CO.
Elevation: 3000 m.
Endemic.
Puya ctenorhyncha L.B. Sm., Phytologia 5: 49.
1954.
Dept.: LP.
Voucher: L. Besse et ai. 6]8 (SEL).
Endemic.
Puya dyckioides (Baker) Mez, in C. DC., Monogr.
phan. 9: 486. 1896. Basionym: Pitcairnia dy-
ckioides Baker, Handb. Bromel. 118. 1889.
Dept.: CH, SC, TA.
Elevation: 2250-2400(-3750?) m.
Voucher: P. lbisch & c. lbisch 93.1026 (SEL).
"cf." .
Puya entre-riosensis Ibisch & Gross, J. Bromeliad
Soc. 48: 217. 1998.
Dept.: TA.
Elevation: 1800 m.
Type: P. Ibisch & c. Ibisch 93.1282. HT: LPB; IT:
HEID. Bolivia.
Endemic.
Puya ferruginea (Ruiz & Pav.) L.B. Sm., Phyto-
logia 16: 461. 1968. Basionym: Pitcairniafer-
ruginea Ruiz & Pav., Fl. peruv. 3: 36. 1802.
Synonym(s): Pitcairnia consimilis Baker, J. Bot. 19:
226, 266. 1881.
Dept.: CO, LP, Sc.
Type: Ruiz & Pavon s.n. HT: BM. Peru.
Puya fiebrigii Mez, Repert. Spec. Nov. Regni Veg.
3: 9. 1906.
Dept.: TA?
Elevation: Unknown.
Type: Fiebrig 3210. LT: B. Bolivia.
Endemic.
Puya fosteriana L.B. Sm., J. Wash. Acad. Sci. 40:
216. 1950.
Dept.: LP.
Voucher: J. Solomon 18299 (MO, SEL).
Endemic.
Puya glabrescens L.B. Sm., Contr. U.S. Natl. Herb.
29: 537. 1954.
Dept.: CO, PO.
Voucher: L. Besse et al. 502 (SEL).
Endemic.
Puya giareosa L.B. Sm., Lilloa 14: 96. 1948.
Dept.: LP.
Elevation: 3600 m.
Endemic.
Puya herzogii Wittm., Rijks-Herb. 29: 86. 1916.
Dept.: CO, LP.
Voucher: M. Kessler et al. 6581 (SEL).
Endemic.
Puya hofstenii Mez, Repert. Spec. Nov. Regni Veg.
3: 8. 1906.
Synonym(s): Puya hauthalii Mez, Repert. Spec.
Nov. Regni Veg. 16: 65. 1919.
Dept.: CH, PO.
Type: Hofsten 1710. HT: B. Argentina.
Voucher: Hromadnik 19017a (WU).
Puya hromadnikii Rauh. Trop. Subtrop. Pflanzen-
welt 41: 5. 1983.
Dept.: TA.
Voucher: W. Till 60a (WU).
Endemic.
Puya humilis Mez, in C. DC., Monogr. phan. 9:
498. 1896.
Synonym(s): Puya werdermannii Harms, Notizbl.
Bot. Gart. Berlin-Dahlem 10: 793. 1929.
Dept.: CH, CO, PO.
Elevation: (2100-)2800-4200 m.
Type: Kuntze s.n. HT: NY. Bolivia.
Voucher: C. Varadarajan et al. 1451 (LPB, SEL).
Det. G. Varadarajan.
Endemic.
Puya kuntzeana Mez. in C. DC., Monogr. phan. 9:
490. 1896.
Dept.: CO.
Elevation: 1300 m.
Endemic.
Puya lasiopoda L.B. Sm., Proc. Amer. Acad. Arts
70: 153. 1935.
Dept.: BE, LP?
Elevation: 500(- J 850?) m.
Voucher: J. Solomon 10862 (MO). "cf.".
Puya laxa L.B. Sm., Phytologia 6: 195. 1958.
Dept.: Sc.
Voucher: S. Beck 7100 (LPB). Det. G. Varadarajan.
Endemic.
Puya leptostachya L.B. Sm., Lilloa 14: 97. 1948.
Dept.: CO.
Voucher: C. Varadarajan et al. 1445 (LPB). Det.
G. Varadarajan.
Endemic.
Puya meziana Wittm., Rijks-Herb. 29: 85. 1916.
Dept.: CO(?), LP.
Elevation: (2200?-)3000-3600 m.
Type: Herzog 2487. HT: L. Bolivia.
Endemic.
Puya micrantha Mez, Repert. Spec. Nov. Regni
Veg. 3: 8. 1906.
Dept.: TAIArgentina border.
Elevation: 1700 m.
Type: Fiebrig 2244. HT: B; IT: BM, GH. Argentina.
Voucher: No known collections from Bolivia.
Puya minima L.B. Sm., Phytologia 7: 423. 1961.
Synonym(s): Puya pusilla H. Luther, Bromelia 4(4):
30 Nov. 1997 [1998].
Dept.: TA.
Type: West 8345. HT: Uc. Bolivia.
Voucher: S. Beck & M. Libermann 14310 (LPB,
SEL).
Endemic.
Puya mirabilis (Mez) L.B. Sm., Phytologia 16:
461. 1968. Basionym: Pitcairnia mirabilis
Mez, Repert. Spec. Nov. Regni Veg. 3: 6. 1906.
Dept.: CO, TA.
Voucher: P. lbisch & Rojas 415 (LPB). Det. G. Var-
adarajan.
Puya mollis Baker ex Mez, in C. DC., Monogr.
phan. 9: 488. 1896.
Dept.: LP.
Type: Bang 1629. LT: K; ILT: BM, GH, US. Bolivia.
Voucher: (Syntype) C. Mandan 1190 (BR, GH,
MANCH, P, PL, STU, WU, Z). Cited in FI.
Neotrop. Monogr. 14(1): 123. 1974.
Endemic.
Puya nana Wittm., Rijks-Herb. 29: 85. 1916.
Synonym(s): Pitcairnia nana (Wittm.) L.B. Sm.,
Phytologia 7: 5. 1959; 10: 30. 1964.
Dept.: Sc.
Elevation: (1350-)1700-2100 m.
Type: Herzog 1856a. HT: L. Bolivia.
Endemic.
Puya olivacea Wittm., Rijks-Herb. 29: 83. 1916;
emend. L.B. Sm., Contr. U.S. Nat. Herb. 29:
538. 1954.
Synonym(s): Pitcairnia olivacea (Wittm.) Mez in
Engler, Pfianzenr. IV, 32: 247. 1935.
Dept.: Sc.
Type: Herzog 1216. HT: L. Bolivia.
Voucher: J. Solomon & M. Nee 17936 (MO, SEL).
Endemic.
Puya paupera Mez, Repert. Spec. Nov. Regni Veg.
3: 14. 1906.
Dept.: TA.
Elevation: 2600 m.
Endemic.
Puya pearcei (Baker) Mez, in C. DC., Monogr.
phan. 9: 480. 1896. Basionym: Pitcairnia pear-
cei Baker, Handb. Bromel. 120. 1889.
Dept.: CO, LP, SC?
Type: Pearce s.n. HT: K. Bolivia.
Voucher: A. Gentry et al. 44254 (LPB, SEL).
Endemic.
Puya pendulifiora L.B. Sm., Contr. Gray Herb. 98:
12. 1932.
Dept.: LP.
Elevation: 1400 m.
Endemic.
Puya potosina L.B. Sm., Phytologia 7: 423. 1961.
Dept.: CR, PO.
Elevation: (2050-)3400 m.
Endemic.
Puya prosanae Ibisch & E. Gross, 1. Bromeliad
Soc. 43: 212. 1993.
Dept.: CO.
Type: P. Ibisch & c. Ibisch 625A. HT: HElD; IT:
LPB. Bolivia.
Endemic.
Puya raimondii Harms, Notizbl. Bot. Gart. Berlin-
Dahlem 10: 213. 1928.
Dept.: CO, LP.
Elevation: (2400-)3000-4200 m.
Voucher: G. & D. Schmitt (LPB, SEL).
Puya reducta L.B. Sm., Phytoiogia 10: 472. 1964.
Dept.: LP.
Elevation: 2600 m.
Endemic.
Puya riparia L.B. Sm., Phytologia 7: 424. 1961.
Dept.: LP.
Elevation: 1350(-3950) m.
Voucher: S. Beck 11277 (LPB). Det. A. Gilmartin.
Endemic.
Puya rusbyi (Baker) Mez, in C. DC., Monogr. phan.
9: 482. 1896. Basionym: Pitcaimia rusbyi Bak-
er, Handb. Bromel. 122. 1889.
Dept.: LP.
Voucher: West 6402 (UC). Cited in Fl. Neotrop.
Monogr. 14(1): 89. 1974.
Endemic.
Puya sanctae-crucis (Baker) L.B. Sm., Proc. Amer.
Acad. Arts 70: 154. 1935. Basionym: Pitcaimia
sanctae-crucis Baker, Handb. Bromel. 120.
1889.
Synonym(s): Pitcaimia robusta Rusby, Bull. New
York Bot. Gard. 6: 488. 1910.
Dept.: LP, SC.
Type: Castelnau s.n. HT: P. Bolivia.
Voucher: L. Besse et al. 636 (SEL).
Endemic.
Puya secunda L.B. Sm., Phytologia 7: 424. 1961.
Dept.: CO.
Voucher: P. lbisch 14074 (LPB).
Endemic.
Puya serranoensis Raub, Trop. SUbtrop. Pflanzen-
welt 41: 14. 1983.
Dept.: SC.
Voucher: l. Vargas & Fuentes 3046 (SEL).
Endemic.
Puya solomonii Varad., J. Bromeliad Soc. 39: 12l.
1989.
Dept.: LP.
Elevation: 3050 m.
Type: G. Varadarajan, U. Varadarajan & J. Solo-
mon 1471. HT: GH; IT: LPB, MO, US, WS.
Bolivia.
Endemic.
Puya spathacea (Griseb.) Mez, in C. DC., Monogr.
phan. 9: 481. 1896. Basionym: Pitcaimia spa-
thacea Griseb., Abh. KOnig!. Ges. Wiss. GOt-
tingen 24: 329. 1879.
Dept.: TA.
Elevation: 2050 m.
Type: Hieronymus 175. HT: GOET; IT: CORD. Ar-
gentina.
Voucher: E. Bastian 414 (LPB).
Puya stenothyrsa (Baker) Mez. in C. DC., Monogr.
phan. 9: 481. 1896. Basionym: Pitcaimia sten-
othyrsa Baker, Handb. Bromel. 122. 1889.
Synonym(s): Pitcaimia bangii Baker, Mem. Torrey
Bot. Club 6: 124. 1896.
Dept.: CO, LP, SC, TA.
Elevation: (1350-)2700-3800 m.
Type: Mandon 1189. HT: K; IT: GH, PI, US. Boliv-
ia.
Voucher: L. Hromadnik 19026 (WU).
Endemic.
Puya tristis L.B. Sm., Phytologia 10: 475. 1964.
Dept.: CO.
Elevation: 3900 m.
Endemic.
Puya trollii L.B. Sm., Phytologia 16: 459. 1968.
Dept.: PO.
Elevation: 3350 m.
Type: Troll 661. HT: M. Bolivia.
Voucher: S. Beck et al. 26787 (LPB).
Endemic.
Puya tuberosa Mez, in C. DC., Monogr. phan. 9:
483. 1896; emend. L.B. Sm., Contr. U.S. Natl.
Herb. 29: 539. 1954.
Dept.: SC, TA.
Type: Haenke S.n. HT: M. Bolivia.
Voucher: Cardenas 4647 (US). Cited in Fl. Neotrop.
Monogr. 14(1): 120. 1974.
Endemic.
Puya tunariensis Mez, in C. DC., Monogr. phan.
9: 498. 1896.
Dept.: CO.
Elevation: (2000-)3150-3800 m.
Endemic.
Puya ugentiana L.B. Sm., Phytologia 13: 147.
1966.
Dept.: CH.
Type: Ugent 4998. HT: US; IT: WIS. Bolivia.
Voucher: L. Hromadnik 19066 (WU).
Endemic.
Puya ultima L.B. Sm., Contr. U.S. Natl. Herb. 29:
540. 1954.
Dept.: LP.
Voucher: M. Kessler et al. 10574 (SEL).
Endemic.
Puya valida L.B. Sm., Phytologia 10: 474. 1964.
Dept.: CH.
Elevation: 2500 m.
Endemic.
Puya vallo-grandensis Rauh, Trop. Subtrop. Pflan-
zenwelt 41: 9. 1983.
Dept.: SC.
Voucher: P. 1bisch & c. 1bisch 93.0861 (SEL).
Endemic.
Puya vasquezii P. Ibisch & E. Gross, Rev. Soc. Bo-
liviana Bot. 2(1): 26. 1998.
Dept.: SC.
Elevation: 600 m.
Type: R. Vasquez et al. 2025. HT: LPB; IT: NY,
USZ. Bolivia.
Endemic.
Puya weberbaueri Mez, Bull. Herb. Boissier, ser.
2,4: 633. 1904.
Dept.: LP.
Elevation: Unknown.
Voucher: Cardenas 5534 (US). Cited for Bolivia in
Fl. Neotrop. Monogr. 14(1): 148. 1974.
Puya weddelliana (Baker) Mez, in C. DC., Monogr.
phan. 9: 475. 1896. Basionym: Pitcairnia wed-
delliana Baker, Handb. Bromel. 12. 1889.
Dept.: CH, TA.
Voucher: L. Hromadnik 19046a (WU).
Endemic."
Puya sp. 1.
Dept.: LP.
Elevation: 1450 m.
RACINAEA
Racinaea cr. riocreuxii (Andre) M.A. Spencer &
L.B. Sm., Phytologia'74: 156. 1993. Basionym:
Tillandsia riocreuxii Andre, Enum. Bromel. 7.
13 Dec. 1888; Rev. Hort. 60: 567. 16 Dec.
1888.
Dept.: LP.
Elevation: 3250 m.
Type: Andre 4408. HT: K. Ecuador.
Voucher: B. Herzog 368 (LPB).
Racinaea flexuosa (Baker) M.A. Spencer & L.B.
Sm., Phytologia 74: 154. 1993. Basionym: Ca-
topsisjlexuosa Baker, J. Bot. 25: 175. 1887.
Synonym(s): Tillandsia bakeri L.B. Sm., Contr.
Gray Herb. 95: 45. 1931.
Dept.: CO, LP.
Type: Mandon 1172. HT: K. Bolivia.
Racinaea kessleri H. Luther, SeJbyana 19: 86.
1998.
Dept.: CO.
Type: M. Kessler et at. 8380. HT: SEL; IT: LPB.
Bolivia.
Endemic.
Racinaea monticola (Mez & Sodiro) M.A. Spencer
& L.B. Sm., Phyto10gia 74: 155. 1993. Basion-
ym: Tillandsia monticola Mez & Sodiro, Bull.
Herb. Boissier, ser. 2, 4: 1135. 1904.
Dept.: LP.
Type: Sodiro 171/33. HT: B; IT: F, GH. Ecuador.
Voucher: O. Buchtien 1266 (US). Cited for Bolivia
in FI. Neotrop. Monogr. 14(2): 1038. 1977.
Racinaea nervibractea (Gilmartin & H. Luther)
J.R. Grant, Phyto10gia 74: 429. 1993. Basion-
ym: Tillandsia nervibractea Gilmartin & H.
Luther, J. Bromeliad Soc. 40(6): 250. 1990.
Dept.: CO, SC.
Type: Kalmbacher S.n. HT: US. Ecuador.
Voucher: P. 1bisch 93.1537 (LPB, SEL).
Racinaea pardina (L.B. Sm.) M.A. Spencer & L.B.
Sm., Phytologia 74: 155. 1993. Basionym: TU-
landsia pardina L.B. Sm., Lilloa 14: 98. 1948.
Dept.: LP, Sc.
Endemic.
Racinaea parvifiora (Ruiz & Pav.) M.A. Spencer
& L.B. Sm., Phytologia 74: 156. 1993. Basion-
ym: Tillandsia parvijlora Ruiz & Pav., FL pe-
ruv. 3: 41. 1802.
Dept.: CO, LP, Sc.
Elevation: (500-)950-2500 m.
Type: Ruiz & Pavon s.n. HT: BM. Peru.
Racinaea pendulispica (Mez) M.A. Spencer &
L.B. Sm., Phytologia 74: 156. 1993. Basionym:
Tillandsia pendulispica Mez, in C. DC., Mon-
ogr. phan. 9: 745. 1896.
Synonym(s): Tillandsia triangularis Rusby, Descr.
S. Amer. pI. 3. 1920.
Dept.: CO.
Elevation: Unknown (> 2500 m?).
Type: Poeppig 1348. HT: G; IT: W (lost). Peru.
Voucher: M. Bang s.n. (NY). Cited for Bolivia in
FL Neotop. Monogr., 14(2): 1040. 1977.
Racinaea schumanniana (Wittm.) J.R. Grant, No-
von 4: 363. 1994. Basionym: Catopsis schu-
manniana Wittm., Bot. lahrb. Syst. 11: 70.
1890.
Dept.: CO, LP.
Type: Lehmann 22. HT: G. Colombia.
Voucher: 1. Solomon 9189 (SEL).
Racinaea seemannii (Baker) M.A. Spencer & L.B.
Sm., Phytologia 74: 157. 1993. Basionym: Guz-
mania seemannii Baker, Handb. Bramel. 153.
1889.
Dept.: CO, LP.
Type: Seemann 898. HT: K. Ecuador.
Racinaea spiculosa (Griseb.) M.A. Spencer & L.B.
Sm., Phytologia 74: 157. 1993. Basionym: Til-
landsia spiculosa Griseb., Nachr. Konig!. Ges.
Wiss. Georg-Augusts-Univ. [1]: 17. 1864.
Synonym(s): Tillandsia brittoniana Baker, Handb.
Bramel. 195. 1889; Tillandsia micrantha Bak-
er, Bull. Torrey Bot. Club 29: 698. 1902.
Dept.: BE, CO, SC, LP.
Racinaea tetrantha (Ruiz & Pav.) M.A. Spencer &
L.B. Sm., Phytologia 74: 158. 1993. Basionym:
Tillandsia tetrantha Ruiz & Pav., F!. peruv. 3:
39, pi 265. 1802.
Dept.: CO, LP.
Type: Ruiz & Pavon s.n. HT: MA. Peru.
Voucher: M. Kessler et al.10421 (SEL).
TILLANDSIA
Tillandsia acuminata L.B. Sm., Contr. U.S. Nat!.
Herb. 29: 434. 1951.
Dept.: CO, LP.
Type: M.E. Foster & L.E. Smith 1402. HT: GH. Co-
lombia.
Tillandsia adpressifiora Mez, in C. DC., Monogr.
phan. 9: 661.1896.
Dept.: Sc.
Type: Wullschlaegel 1335. HT: BM. Suriname.
Voucher: S. Halloy et al. 4406 (SEL).
Tillandsia aizoides Mez, in C. DC,. Monogr. phan.
9: 866.1896.
Dept.: PO, TA.
Type: Hieronymus & Niederlein 850. HT: B; IT:
CORD. Argentina.
Voucher: H. Hromadnik & L. Hromadnik 5086
(WU).
TiUandsia argentina C.H. Wright, Bull. Misc. In-
form. Kew 60. 1907.
Dept.: CH, TA.
Type: Stuckert s.n. HT: K. Argentina.
SEL).
Tillandsia asplundii L.B. Sm., Phytologia 6: 441.
1959.
Dept.: CO, LP.
Type: Asplund 19279. HT: S; IT: US. Ecuador.
Tillandsia australis Mez, Repert. Spec. Nov. Regni
Veg. 16: 75. 1919.
Synonym(s): Tillandsia maxima Lillo & Hauman,
Anales Mus. Nac. Hist. Nat. Buenos Aires 29:
245,416. 1917, non Strangeways, 1882.
Elevation: 800-2100(-3100) m.
Type: Lorentz & Hieronymus 287. HT: LIL; IT: B,
CORD, US. Argentina.
Tillandsia bandensis Baker, J. Bot. 25: 234. 1887.
Synonym(s): Tillandsia pilosa L.B. Sm., Phytologia
7: 175. 1960.
Dept.: CH, CO, SC, TA.
Elevation: (600-)1200-2200 m.
Type: Tweedie S.n. HT: K. Uruguay.
Tillandsia bermejoensis H. Hromadnik, Trap. Sub-
trap. Pflanzenwelt 31: 9. 1979.
Dept.: Sc.
Voucher: D. Cathcart 16 (SEL).
Endemic.
Tillandsia biflora Ruiz & Pav., FL peruv. 3: 41.
1802.
Dept.: CO, LP.
Tillandsia boliviana Mez, Bull. Herb. Boissier, ser.
2, 4: 1l30. 1904.
Dept.: CH7, LP7.
Elevation: 15007-2730? m.
Type: Bang 2202. HT: B; IT: BM, F, G, GH, K, LE,
M, MICH, MO, 0, US, WU. Bolivia.
Voucher: M. Kessler et al. 4535 (LPB, SEL). "cf.".
Endemic.
Tillandsia boliviensis Baker, Mem. Torrey Bot.
Club 4(3): 267. 1895; emend. Contr. Gray Herb.
98: 15. 1932.
Dept.: LP.
Type: Bang 159 A. HT: NY. Bolivia.
Voucher: Buchtien 808 (US). Cited in Fl. Neotrop.
Monogr. 14(2): 769. 1977.
Endemic.
Tillandsia cr. buseri Mez, Bull. Herb. Boissier, ser.
2, 3: 145. 1903.
Dept.: LP.
Elevation: 2750 m.
Type: Langlasse 102-A. HT: G; IT: K. Colombia.
Tillandsia caliginosa W. Till, PI. Syst. Evol. 147:
282. 1984. Basionym: Tillandsia crocata (E.
Morren) Baker var. tristis Rauh, Trop. Subtrop.
Pflanzenwelt 43: 17. 1983, non T. tristis Mez
1896.
Dept.: CH, CO. PO, TA.
Type: D. Muhr s.n. HT: HEID. Argentina.
Voucher: P. 1bisch & c. 1bisch 93.0329 (LPB, SEL).
Tillandsia calochlamys Ehlers & L. Hrom., Die
Bromelie 2/1998: 38. 1998.
Dept.: CO.
Type: R. Ehlers EB952403. HT: LPB. Bolivia.
Voucher: W. Till 158 (WU).
Endemic.
Tillandsia camargoensis L. Hrom., Die Bromelie
111991: 9. 1991.
Dept.: CH, PO.
Type: Hromadnik 5073 A. HT: WU. Bolivia.
Voucher: E. Haugg 7023 (WU). in part.
Endemic.
Tillandsia capillaris Ruiz & Pav., FI. peruv. 3: 42.
1802.
Dept.: CH, CO, LP, OR, PO, SC, TA.
Type: Ruiz & Pavon S.n. LT: BM; ILT: BM, MA.
Peru.
Tillandsia cardenasii L.B. Sm., Proc. Amer. Acad.
Arts 70: 154. 1935.
Dept.: CH, CO, PO, TA.
Endemic.
Tillandsia carnosa L.B. Sm., Phytologia 9: 251.
1963.
Dept.: CO, LP.
Type: J. Wurdack 603. HT: US; IT: USM. Peru.
Tillandsia caulescens Brongn. ex Baker, Handb.
Bromel. 168. 1889.
Dept.: LP.
Elevation: 850 - J 550 m.
Type: Gay 1186. HT: P. Peru.
Tillandsia cochabambae E. Gross & Rauh, Die
Bromelie 2/86: 14. 1986.
Dept.: CO.
Type: W. Rauh 40609. HT: HElD. Bolivia.
Voucher: M. Kessler et al. (LPB, SEL).
Endemic.
Tillandsia colganii Ehlers, J. Bromeliad Soc. 47:
103. 1997.
Dept.: SC.
Elevation: 2700-3200 ill.
Type: R. Ehlers EB 951002. HT: WU. Bolivia.
Voucher: R. Ehlers 951101 (WU). Det. R. Ehlers.
Endemic.
Tillandsia comarapaensis H. Luther, J. Bromeliad
Soc. 34: 54. 1984.
Dept.: CH, CO, SC.
Elevation: (400-)1550-2600 m.
Type: L. Besse et al. 1713. HT: SEL. Bolivia.
Endemic.
Tillandsia compacta Griseb., Nachr. Konigl. Ges.
Wiss. Georg-Augusts-Univ. [1]: 18. 1864.
Dept.: CO.
Elevation: 2000 m.
Type: A. Fendler 1508. HT: GOET; IT: K. Vene-
zuela.
Voucher: L. Dorr 7052 (LPB, SEL).
Tillandsia compianata Benth., Bot. voy. Sulphur
173. 1846.
Dept.: CO, LP, Sc.
Elevation: (300-)1050-2900 m.
Type: Sinclair s.n. HT: K. Ecuador.
Tillandsia confinis L.B. Sm., Phytologia 4: 218.
1953.
Synonym(s): Tillandsia subtropicalis L.B. Sm.,
Phytologia 8: 505. 1963.
Dept.: CO, LP, Sc.
Type: Ferreyra 1686. HT: USM. Peru.
Voucher: J. Solomon 9253 (LPB, MO, SEL).
Tiliandsia cotagaitensis L. Hrom., PI. Syst. Evo!.
147: 284, 1984.
Dept.: PO.
Elevation: 3000 m.
Type: Hromadnik 5093. HT: WU; IT: LI, NY, WU.
Bolivia.
Endemic.
Tillandsia denudata Andre, Enum. Bromel. 8. 13
Dec. 1888; Rev. Hort. 60: 568. 16 Dec. 1888.
Dept.: LP.
Elevation: 1250 m.
Type: Andre 1358. HT: HElD. Colombia.
Tillandsia didisticha (E. Morren) Baker, J. Bot. 26:
16. 1888. Basionym: Anoplophytum didisti-
chum E. Morren, Belgique Hort. 31: 164. 1881.
Synonym(s): Guzmania complanata Wittm., Meded.
Rijks-Herb. 29: 92. 1916.
Dept.: BE, CH, CO, LP, SC, TA.
Type: Jacob Matoy Hortus s.n. HT: LG. Brazil.
Tillandsia duratii Visiani, Nuovi. Saggi r. Accad.
Sci. Padova 5: 271. 1840.
Synonym(s): Tillandsia weddellii Baker, Handb.
Bromel. 181. 1889.
Dept.: CH, CO, LP, PO, SC, TA.
Type: Padua Hortus s.n. HT: PAD.
Tillandsia edithae Rauh, Trop. Subtrop. Pflanzen-
welt 8: 19. 1974.
Dept.: CH, LP, SC.
Type: Krahn K 295. HT: HElD; IT: US, HAL. Bo-
livia.
Endemic.
Tillandsia engleriana Wittm., Bot. Jahrb. Syst. 11:
63. 1890.
Dept.: CO, LP, SC.
Type: Lehmann 520. HT: G-B. Colombia.
Voucher: D. Smith 13442 (LPB, MO).
Tillandsia erici Ehlers, Die Bromelie 111998: 20.
1998.
Dept.: TA.
Type: E. Haugg & E. Heger 11247. HT: WU; IT:
LPB. Bolivia.
Endemic.
Tillandsia fendleri Griseb., Nachr. Konigi. Ges.
Wiss. Georg-Augusts-Univ. [l): 17. 1864.
Dept.: BE, CO, LP, Sc.
Elevation: 700-2200(-3000) m.
Tillandsia funebris A. Cast., Anales Mus. Nac.
Hist. Nat. Buenos Aires 37: 502. 1933.
Dept.: CH, CO, PO, SC, TA.
Type: Schreiter 1689. HT: BA; IT: B, GH, LIL. Ar-
gentina.
Voucher: M. Kessler et al. 4710 (LPB, SEL, WU).
Tillandsia gerdae Ehlers, Die Bromelie 2/87: 32.
1987.
Dept.: CH, CO.
Elevation: (1700-)2000-2600 m.
Type: E. Haugg 7163. HT: WU; IT: WU. Bolivia.
Voucher: E. Haugg 8283 (WU).
Endemic.
Tillandsia gilliesii Baker, J. Bot. 16: 240. 1878.
Type: Gillies s.n. HT: K; IT: CGE, GH. Argentina.
Voucher: M. Cardenas 2418 (GH). Cited for Bolivia
in FI. Neotop. Monogr. 14(2): 885. 1977.
Tillandsia basei Ehlers & L. Hrom., Die Bromelie
311996: 176. July 1997.
Dept.: CH.
Elevation: 3100 m.
Type: F. Hase et al. EB 951501. HT: WU. Bolivia.
Voucher: L Hromadnik 19046 (WU).
Endemic.
Tillandsia hegeri Ehlers, Die Bromelie 2/1994: 33.
1994.
Dept.: CH, TA.
Type: E. Heger s.n. HT: WU. Bolivia.
Endemic.
Tillandsia helmutii L. Hrom., Die Bromelie 3/
1990: 63. Jan. 1990 [1991).
Dept.: CH.
Type: L. & H. Hromadnik 13009. HT: WU; IT: WU.
Bolivia.
Voucher: L & H. Hromadnik 5028 (WU).
Endemic.
Tillandsia hirta w. Till & L. Hrom., PI. Syst. Evoi.
147: 285. 1984.
Type: L & R. Hromadnik HR 5001. HT: WU; IT:
WU. Bolivia.
Tillandsia imporaensis R. Ehlers, Die Bromelie 3/
1998: 76. 1998.
Dept.: CH, CO.
Elevation: 3050 m.
Type: E. Haugg 7078. HT: WU. Bolivia.
Voucher: G. Kohres & L. KiJhres s.n. (WU).
Endemic.
Tillandsia ionochroma Andre ex Mez, in C. DC.,
Monogr. phan. 9: 801. 1896.
Dept.: CO, LP, SC.
Type: Andre 2746. HT: K; IT: NY. Ecuador.
Tillandsia ixioides Griseb., Abh. Konigl. Ges.
Wiss. Gottingen 24: 333. 1879.
Dept.: CH, TA.
Type: Lorentz 1196. HT: B; IT: BM, G-B, GOET,
K, M, US. Argentina.
Voucher: M. Cardenas 4093 (GH, US). Cited for
Bolivia in Fl. Neotrop. Monogr. 14(2): 821.
1977.
Tillandsia jucunda A. Cast., Anales Mus. Nac.
Hist. Nat. Buenos Aires 36: 56. 1929.
Dept.: CH.
Type: Schreiter 26/2367. HT: BA. Argentina.
Tillandsia juncea (Ruiz & Pav.) Poir., Encycl.
suppl. 5: 309. 1817. Basionym: Bonapartea
juncea Ruiz & Pav., FI. peruv. 3: 38. 1802.
Dept.: BE, CO, LP, PA, SC.
Elevation: 800-1400(-2000) m.
Tillandsia kessleri H. Luther, Selbyana 20: 10.
1999.
Dept.: CO.
Type: M. Kessler et at. 9557. HT: SEL; IT: LPB.
Bolivia.
Endemic.
Tillandsia koehresiana Ehlers, Die Bromelie 3/
1990: 59. 1990 [Jan. 1991].
Dept.: CO, PO.
Type: K6hres S.n. HT: WU; IT: WU. Bolivia.
Voucher: P. lbisch 216 (LPB, SEL).
Endemic.
Tillandsia krukoffiana L.B. Sm., Contr. Gray Herb.
154: 36. 1945.
Dept.: BE, CO, LP, SC?
Elevation: 850-2200(-2700) m.
Type: B. Krukoff 10503. HT: GH;. IT: K, MICH, NY,
S. Bolivia.
Endemic.
Tillandsia kuehhasii W. Till, Die Bromelie 2/1995:
33. 1995.
Dept.: CH, PO?
Elevation: 3600 m.
Type: E. Kirschnek & F. Kuehhas S.n. HT: WU; IT:
WU. Bolivia.
Voucher: R. Ehlers 950601 (WU).
Endemic.
Tillandsia kuntzeana Mez, in C. DC., Monogr.
phan. 9: 790. J 896.
Dept.: CO, LP?
Elevation: 1300-2100(-3050) m.
Endemic.
Tillandsia laxissima Mez, BulL Herb. Boissier, ser.
2, 5: 108. 1905,
Dept.: SC, LP.
Type: Bang 2301 (in part). HT: B; IT: G, GH, LE,
M, MO, NY, PH, US, W, WU. Bolivia.
Voucher: M. Kessler 4064 (LPB).
Endemic.
Tillandsia loliacea Mart. ex Schult. & Schult.f., in
Roem. & Schult., Syst. veg. 7(2): 1204. 1830.
Elevation: 200-1400(-2200) m.
Tillandsia longifolia Baker, Handb. Bromel. 185.
1889.
Dept.: CO, SC?
Elevation: 2000-2100(-2600) m.
Type: A. Fendler 1522. HT: K; IT: GH. Venezuela.
Voucher: P. 1bisch & c. 1bisch 93.1628 (SEL).
Tillandsia lorentziana Griseb., Abh. Konigl. Ges.
Wiss. Gottingen 19: 271.1874.
Synonym(s): Tillandsia lorentziana f. simplex 0.
Kuntze, Revis. gen. pl. 3(2): 304. 1898.
Type: Lorentz 69. HT: B; IT: CORD. Argentina.
Voucher: P. 1bisch & c. 1bisch 93.0208 (LPB, SEL).
Tillandsia lotteae H. Hrom., J. Bromeliad Soc. 28:
263. 1978.
Dept.: CH, CO, PO, SC.
Elevation: (900-)1550-2900 m.
Type: H. Hromadnik PB 77. HT: HElD; IT: HAL.
Bolivia.
Endemic.
Tillandsia mandonii E. Morren ex Mez, in C. DC.,
Monogr. phan. 9: 871. 1896.
Dept.: LP, TA.
Type: Mandon 1180. LT: not selected; ILT:
BM,GH,K,PI,S,W. Bolivia.
Tillandsia moms H. Hrom. & W. Till, PI. Syst.
Evol. 142: 123. 1983.
Dept.: TA.
Elevation: 2700 m.
Type: H. Hromadnik 9088. HT: WU; IT: B, HAL,
K, LI, NY, W, WU. Bolivia.
Voucher: E. Heger s.n. (WU).
Endemic.
Tillandsia muhriae W. Weber, Feddes Repert. 97:
101. 1986.
Synonyrn(s): Tillandsia alberi Ehlers, Die Bromelie
4/87: 55. 1987.
Dept.: PO.
Type: Muhr s.n. HT: HAL. Argentina.
Voucher: E. Haugg 7029 (WU).
Tillandsia myosura Griseb. ex Baker, J. Bot. 16:
240. 1878.
Type: Lorentz 122. HT: BM; IT: B, NY. Argentina.
SEL).
Tillandsia nana Baker, Handb. Bromel. 172. 1889.
Synonym(s): Tillandsia calocephala Wittm., Med-
ed. Rijks-Herb. 29: 90. 1916.
Dept.: CO, LP.
Elevation: 3150 m.
Type: Gay s.n. HT: P. Peru.
Voucher: Shepard 234 (GH, US). cited for Bolivia
in FI. Neotrop. Monogr. 14(2): 807. 1977.
Tillandsia oropezana L. Hrom., Die Bromelie 1/
89: 14. 1989.
Dept.: CH.
Elevation: 1800 m.
Type: H. Hromadnik 9012. HT: WU; IT: WU. Bo-
livia.
Endemic.
Tillandsia paleacea C. PresJ, Re1iq haenk. 1(2):
125. 1827.
Synonym(s): Tillandsia scalarifolia Baker, I. Bot.
25: 235. 1887.
Dept.: LP.
Elevation: 2600 m.
Type: Haenke S.n. HT: PR. Chile.
Voucher: Julio 158 (BM, US). Cited for Bolivia in:
FI. Neotrop. Monogr. 14(2): 863. 1977.
Tillandsia paraensis Mez, in Mart., FI. bras. 3(3):
586. 1894.
Dept.: LP, PA, Sc.
Type: Sieber 68. HT: BM. Brazil.
Voucher: T. Kromer & A. Acebey 187 (LPB, SEL).
Tillandsia pedicellata (Mez) A. Cast., Lilloa 11:
144. 1945. Basionym: Tillandsia coarctata Gil-
lies ex Baker vaT. pedicellata Mez, in C. DC.,
Monoge phan. 9: 865. 1896.
Dept.: CH, LP, TA.
Type: Hieronymus 795. LT: B; ILT: BM, CORD,
GH, GOET. Argentina.
Voucher: B. Holst et ai. 4958 (USZ).
Tillandsia platyrachis Mez, in C. DC., Monoge
phan. 9: 848. 1896.
Dept.: LP.
Elevation: 900 m.
Type: Kalbreyer 1328. HT: K. Colombia.
Voucher: T. Kromer et ai. 199 (GOET, LPB, SEL).
Tillandsia pohliana Mez, in Mart., FI. bras. 3(3):
597. 1894.
Dept.: BE, CH, LP, Sc.
Type: Pohl 3658. HT: W (lost). Brazil.
Voucher: M. Kessler et ai. 4845 (LPB, SEL).
Tillandsia polystachia (L.) L., Sp. pI. (ed. 2) 1:
410. 1762. Basionym: Renealmia polystachia
L., Sp. pI. 1: 286. 1753.
Dept.: BE, LP, Sc.
Type: Plumier S.n. HT: P. West Indies.
Tillandsia pseudocardenasii W. Weber, Feddes Re-
pert. 97: 103. 1986.
Dept.: LP, Sc.
Elevation: 2300 m.
Type: H. Hromadnik & L. Hromadnik 5323. HT:
HAL; IT: WU. Bolivia.
Voucher: K Uhlig S.n. (WU).
Endemic.
Tillandsia pyramidata Andre, Bromel. Andr. 86.
1889; emend. Rauh, Trop. Subtrop. 3: 15. 1973.
Dept.: LP.
Elevation: 2400 m.
Voucher: S. Beck 3142 (LPB). Del. L.B. Sm.
Tillandsia rauschii Rauh & Lehmann, Trop. Sub-
trop. Pflanzenwelt 21: 41. 1977.
Dept.: CH?
Elevation: 2500? m.
Type: Bot. Garten Heidelberg 31623 ( = Rausch
s.n.). HT: HElD; IT: LPB, WU. Bolivia.
Voucher: Only known from type collection, without
specific locality.
Endemic.
Tillandsia rectangula Baker, J. Bot. 16: 238. 1878.
Dept.: TA.
Type: Lorentz 127. HT: K. Argentina.
Voucher: E. Bastian 1295 (LPB).
Tillandsia recurvata (L.) L., Sp. pI. (ed. 2) 1: 410.
1762. Basionym: Renealmia recurvata L., Sp.
pI. 1: 287. 1753.
Synonym(s): Tillandsia recurvata f. brevifolia An-
dre, Bromel. Andr. 65. 1889.
Type: Sloane s.n. HT: BM; IT: B, GOET, M, P. Ja-
maica.
Tillandsia recurvifolia Hook., Bot. Mag. 87: pI.
5246. 1861.
Synonym(s): Tillandsia meridionalis auct. non Bak-
er: sensu Smith & Downs, FI. Neotrop. Monoge
14(2): 820. 1977.
Dept.: CH, SC, TA.
Type: Original description and plate.
Tillandsia recurvispica L. Hrom. & P. Schneid.,
Die Bromelie 1/87: L 1987.
Dept.: CO.
Type: H. Hromadnik & L. Hromadnik 3007. HT:
WU; IT: LPB, WU. Bolivia.
Voucher: H. Hromadnik & L. Hromadnik 5330
(WU).
Endemic.
Tillandsia reichenbachii Baker, Handb. BromeL
166. 1889.
Synonym(s): Tillandsia herzogii Wittm., Meded.
Rijks-Herb. 29: 89. 1916.
Dept.: CH, CO, SC, TA.
Type: Reichenbach icon; with no specific locality.
(K).
Voucher: R. Martinez 591 (LPB, SEL).
Tillandsia rhomboidea Andre, Enum, Bromel. 6.
13 Dec. 1888; Rev. Hort. 60: 566. 16 Dec.
1888.
Dept.: BE.
Type: Andre 2745. LT: K; IT: NY. Colombia.
Voucher: S. Beck 1732 (US).
Tillandsia rubella Baker, J. Bot. 26: 44. 1888.
Dept.: CO, LP, Sc.
Type: Mandon 1187. HT: K; IT: GH, P. Bolivia.
Tillandsia rusbyi Baker, Handb. Bromel. 176.
1889.
Dept.: LP, Sc.
Voucher: T. Kromer & A. Acebey 183 (LPB, SEL).
Tillandsia samaipatensis W. Till, Die Bromelie 21
1996: 36. Feb. 1997.
Dept.: LP, Sc.
Type: W. Till 149/2. HT: LPB; IT: SEL, WU. Bo-
livia.
Endemic.
Tillandsia schreiteri Lillo & A. Cast., Anales Mus.
Nac. Hist. Nat. Buenos Aires 36: 56. 1929.
Dept.: TA.
Elevation: 1800 m.
Type: Schreiter S.n. HT: BA. Argentina.
Voucher: H. Hromadnik 9099 (WU).
Tillandsia sphaerocephala Baker, J. Bot. 26: 141.
1888.
Type: Mandon 1188. HT: K; IT: G. Bolivia.
Tiliandsia spiralipetala Gouda, J. Bromeliad Soc.
36: 165. 1986. Basionym: Tillandsia tricholepis
Baker var. macrophylla L.B. Sm., Lilloa 14: 98.
1948.
Dept.: CH, LP, Sc.
Elevation: 350-1450(-2300) m.
Type: Buchtien 1270. HT: US; IT: E. Bolivia.
Tillandsia stenoura Harms, NotizbL Bot. Gart.
Berlin-Dahlem 12: 537. 1935.
Dept.: CO.
Type: Diels 558. HT: B. Ecuador.
Voucher: P. Ibisch & c. lbisch 93.0098 (SEL).
Tillandsia streptocarpa Baker, J. Bot. 25: 241.
1887.
Dept.: CO, CH, LP, PO, Sc.
Type: Balansa 615. HT: K; IT: P, S. Paraguay.
Tillandsia superba Mez & Sodiro, Bull. Herb.
Boissier, ser. 2, 4: 1132. 1904.
Dept.: CO, LP.
Type: Sodiro 37-D. HT: B. Ecuador.
Voucher: R. Seidel 7015 (LPB, SEL).
Tillandsia tenuifolia L., Sp. pI. 1: 286. 1753.
Type: Royen s.n. HT: L. Country unknown.
Voucher: R. Seidel 7011 (LPB, SEL).
Tillandsia tovarensis Mez, in C. DC., Monogr.
phan. 9: 769. 1896.
Dept.: CO, LP.
Tillandsia tricholepis Baker, J. Bot. 16: 237. 1878.
Type: Mandon 1179 in part. HT: K; IT: BM, NY, P,
S. Bolivia.
Voucher: S. Beck 3200 (LPB, SEL). Det. L.B. Sm.
Tillandsia usneoides (L.) L., Sp. pI. (ed. 2) 1: 411.
1762. Basionym: Renealmia usneoides L., Sp.
pI. 1: 287. 1753.
Type: Anonymous in Clifford Herbarium s.n. LT:
BM. Jamaica.
Tillandsia variabilis SchltdL, Linnaea 18: 418.
1844.
Synonym(s): Tillandsia valenzuelana A. Rich. in
Sagra, Hist. fis. Cuba, Bot. 11: 267. 1850.
Dept.: CO, LP, Sc.
Type: Schiede s.n. LT: HAL; ILT: HAL. Mexico.
Tillandsia vernicosa Baker, J. Bot. 25: 241. 1887.
Type: Christie s.n. HT: K. Argentina.
Tillandsia violascens Mez, in C. DC., Monogr.
phan. 9: 797. 1896.
Dept.: CO, LP, SC.
Type: Mandon 1174 in part. HT: P; IT: BM. Bolivia.
Tillandsia virescens Ruiz & Pav., FI. Peruv. 3: 43.
1802.
Synonym(s): Tillandsia capillaris f. virescens (Ruiz
& Pavon) L.B. Sm., Proc. Amer. Acad. Arts 70:
212. 1935.
Type: Ruiz & Pavon S.n. LT: BM. Peru.
Tillandsia walter-richteri W .. Weber, J. Bromeliad
Soc. 34: 102. 1984.
Dept.: TA.
Elevation: 1800 m.
Type: H. Hromadnik & E. Hromadnik 5142. HT:
HAL; IT: WU. Bolivia.
Endemic.
Tillandsia walteri Mez, Repert. Spec. Nov. Regni
Veg. 3: 43. 1906.
Dept.: CO, LP.
Tillandsia xiphloides Ker-Gawler, Bot. Reg. 2: pl.
105. 1816.
Dept.: CH, CO, LP, PO, TA.
Type: Original description and plate.
Voucher: M. Cardenas 93 (GH, SIlo
Tillandsia yuncharaensis W. Till, J. Bromeliad
Soc. 40: 214. 1990.
Dept.:TA.
Elevation: 3000 m.
Type: w: Till et al. sub WT 55. HT: WU. Bolivia.
Endemic.
Tillandsia sp. 1.
Synonym(s): Tillandsia bryoides auct. non Griseb.
ex Baker: Smith and Downs, Fl. Neotrop. Mon-
ogr. 14: 879. 1977; Brako and Zarucchi, Mon
ogr. Syst. Bot. Missouri Bot. Gard. 45: 250.
1993; Zuloaga and Morrone, Monogr. Syst.
Bot. Missouri Bot. Gard. 60: 114. 1996.
Tillandsia bryoides Grisebach, Abh. Konig!. Ges.
Wiss. Gottingen 24: 334. 1879, in part; Tilland-
sia coarctata Gillies ex Mez, in C. DC., Mon-
ogr. phan. 9: 865. 1896, in part.
Dept.: CR, CO, LP, OR, PO, SC, TA.
Elevation: (400-)1200-3000 m.
Voucher: M. Cardenas 3309 (GH).
Note: This taxon has often been identified as Til-
landsia bryoides, but appears to be a fairly
widespread, undescribed species.
VRIESEA
Vriesea cf. chrysostachys E. Morren, Belgique
Hort. 31: 87. 1881.
Dept.: LP.
Elevation: 900 m.
Type: Davis in Hort. Veitch s.n. HT: LG. Peru.
"cf.".
Vriesea chontalensis (Baker) L.B. Sm., Contr. U.S.
Nat!. Herb. 29: 518. 1951. Basionym: Tilland-
sia chontalensis Baker, J. Bot. 25: 237. 1887.
Dept.: LP.
Elevation: 1300 m.
Type: R. Tate 413. HT: BM. Nicaragua.
Vriesea friburgensis Mez, in Mart., FI. bras. 3(3):
537. 1894.
Dept.: CH, SC, TA.
Type: Glaziou 16467. HT: K; IT: C, LE, P. Brazil.
Voucher: B. Holst et al. 4887 (SEL, USZ).
Vriesea heliconioides (Kunth) Hook. ex WaIp.,
Ann. Bot. Syst. 3: 623. 1852. Basionym: Til-
landsia heliconioides Kunth, Nov. gen. sp. 1:
293. 1816.
Dept.: BE, LP, SC.
Type: Humboldt & Bonpland s.n. HT: P. Colombia.
Vriesea heterandra (Andre) L.B. Sm., Contr. U.S.
Nat!. Herb. 29: 443. 1951. Basionym: Tilland-
sia heterandra Andre, Enum. Bromel. 7. 13
Dec. 1888; Rev. Hort. 60: 567. 16 Dec. 1888.
Dept.: LP, SC, TA.
Vriesea incurva (Griseb.) Read, Phytoiogia 16:
458. 1968. Basionym: Tillandsia incurva Gri-
seb., Nachr. Konigl. Ges. Wiss. Georg-Augusts-
Univ. [II: 15 .. 1864.
Dept.:BE?,LP.
Type: A. Fendler 1524. HT: GOET; IT: GH. Vene-
zuela.
Voucher: Buchtien 373 (US). Cited for Bolivia in
Fl. Neotrop. Monogr. 14 (2): 1251. 1977.
Vriesea maxoniana (L.B. Sm.) L.B. Sm., Phytolo-
gia 6: 441. 1959. Basionym: Tillandsia max-
oniana L.B. Sm., Contr. Gray Herb. 124: 11.
1939.
Vriesea platynema Gaudich., Voy. Bonite, Bot. pI.
66. 1843.
Dept.: LP.
Elevation: 1050 m.
Type: Gaudichaud s.n. HT: P. Brazil.
Vriesea rubra (Ruiz & Pav.) Beer, Fam. BromeI.
98. 1856. Basionym: Tillandsia rubra Ruiz &
Pav., Fl. peruv. 3: 40, pI. 266. 1802.
Dept.: BE, LP.
WERAUHlA
Werauhia boliviana H. Luther, Selbyana 19: 218.
1998 [1999].
Dept.: LP.
Elevation: 950 m.
Type: M. Kessler 3907. HT: SEL; IT: LPB. Bolivia.
Endemic.
Werauhia gladioliflora (H. Wend!.) J.R. Grant,
Trop. Subtrop. Pflanzenwelt 91: 31. 1995. Bas-
ionym: Tillandsia gladioli flora Wendland,
Hamburger Garten-Blumenzeitung 19: 31.
1863.
Dept.: CO, LP.
Elevation: 350-1300 Ill.
Type: Wendland in Herrenhausen Hortus s.n. HT:
HBG? Costa Rica.
Vouchet: M. Kessler 9874 et al. (LPB, SEL).
Werauhia sanguinolenta (Linden ex Cogn. & Mar-
chat) l.R. Grant, Trop. Subtrop. Pflanzenwelt
91: 35. 1995. Basionym: Vriesea sanguinolenta
Linden ex Cogn. & Marchal, Pl. ornem. 2: pi.
52. 1874.
Dept.: SC.
Type: E. Morren in Liege Hortus s.n. CT?: LG. Cos-
ta Rica.

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Selbyana 20(2): 201-223. 1999.

CHECKLIST OF BOLIVIAN BROMELIACEAE WITH NOTES ON SPECIES

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