A Field Guide
To the Permanent Sample Plots
in the Conventional Logging Blocks 28 & 29
at CIFOR Malinau Research Forest
East Kalimantan
Kuswata Kartawinata
Hari Priyadi
Douglas Sheil
Soedarsono Riswan
Plinio Sist
Machfudh
A Field Guide
To the Permanent Sample Plots
in the Conventional Logging Blocks 28 & 29
at CIFOR Malinau Research Forest
East Kalimantan
Kuswata Kartawinata
Hari Priyadi
Douglas Sheil
Soedarsono Riswan
Plinio Sist
Machfudh
National Library of Indonesia Cataloging-in-Publication Data
Kartawinata, Kuswata et al.
A ield guide to the permanent sample plots in the conventional logging blocks 28 & 29 at CIFOR
Malinau research forest East Kalimantan/by Kuswata Kartawinata, Hari Priyadi, Douglas Sheil,
Soedarsono Riswan, Plinio Sist, Machfudh. Bogor, Indonesia: Center for International Forestry Research
(CIFOR), 2006.
113p.
ISBN: 979-24-4631-1
CAB Int. thesaurus: 1. logging 2. sample plot technique 3. stand density 4. forest trees 5. identiication
6. Indonesia 7. guidelines
© 2006 by CIFOR and ITTO
All rights reserved. Published in 2006
Printed by Intiprima Karya
Cover photos by Hari Priyadi and Ahmad Zakaria
Cover design by Eko Prianto
Layout by Catur Wahyu
Maps by Mohammad Agus Salim
Published by
Center for International Forestry Research
Mailing address: P.O. Box 6596 JKPWB, Jakarta 10065, Indonesia
Ofice address: Jl. CIFOR, Situ Gede, Sindang Barang,
Bogor Barat 16680, Indonesia
Tel.: +62 (251) 622622; Fax: +62 (251) 622100
E-mail: cifor@cgiar.org
Web site: http://www.cifor.cgiar.org
3
Contents
Foreword
4
Introduction
5
The Plot
8
Access to a Plot
9
Plant Identiication
10
Distribution of PSPS
11
Plot# CNV01
12
Plot# CNV02
19
Plot# CNV03
28
Plot# CNV04
35
Plot# CNV05
42
Plot# CNV06
52
Plot# CNV07
60
Plot# CNV08
67
Plot# CNV09
77
Plot# CNV10
87
Plot# CNV11
96
Plot# CNV12
105
4
Foreword
The CIFOR Permanent Sample Plots (PSP) were established in 1998
in conjunction with the Phase 1 of the ITTO Project dealing with
sustainable forest management, integrated land use, reduced-impact
logging, biodiversity in the production forest, and cultural and social
aspects of logging. In terms of technical perspectives, PSP are an
important tool to monitor forest dynamics and changes, long term
growth and yield and to provide critical data for evaluation of ecological
model. For silvicultural purposes PSP supply data on, diameter and
volume increment as well as stand structure dynamics.
I therefore appreciate those who have given strong initiative and
support in the establishment of PSP in Malinau Research Forest and
continuously measure those plots. My utmost appreciation goes also
to our partners: PT INHUTANI II for logging activities, BIOMA, the
Malinau District Ofice for administrative research arrangements and
the Indonesian Institute of Sciences (LIPI) for scientiic identiication
of trees I also wish to express my sincere thanks to Dr. Kuswata
Kartawinata, Dr. Plinio Sist of Cirad Foret, Dr. Kade Sidiyasa of
FORDA and Hari Priyadi who maintain their continuing interest by
revisiting the plots and conirming the accuracy of the identity of trees
found in the plots.
Dr Kartawinata and I take the initiative to publish the guidebook to
satisfy the growing interests and needs of visitors visiting our Seturan
camp. More and more people think highly of the importance of these
plots to understand the secret of forest growth and the diversity of
trees occurring inside the permanent plots.
This initiative is not possible without continuous support of ITTO in
providing fund for both research and publication of this guide. It is
my sincerest hope that this little guide will beneit any one who is
concerned with the future of the tropical forest.
Bogor, May 2006
Dr. Petrus Gunarso
Malinau Research Forest Coordinator
5
Introduction
A set of Permanent Sample Plots (PSP) was established in the lowland
mixed dipetrocarp forest within PT Inhutani II concession at Malinau
(2o45’ – 3o15’N, 116o30’), East Kalimantan (Figure 1). They were
designed for comparative studies of conventional logging (CNV) and
reduced-impact logging (RIL) techniques under the ITTO Project PD
12/97 Rev.1 (F), Forest, Science and Sustainability. Bulungan Model
Forest. The Malinau concession is located about 30 km to the East
of CIFOR’s Bulungan Research Forest in a forest area with elevation
between 100 m to 300 m above sea level. The terrain is undulating
with slopes ranging from 10% to 70%. The mean annual rainfall in the
area is 3.790 mm, with the number of wet months of more than 9
months per year.
This pocket guideline is intended to give visitors to PSPs information
on how to access to the location of plots and distribution of trees with
dbh ≥ 20cm in each plot. The CIFOR’s Seturan station is used as a
starting point to travel to the plots.
6
Figure 1. Malinau Research Forest Map
Figure 2. Location of Permanent Sample Plots
A Field Guide to the Permanent Sample Plots • 7
8
The Plots
The objective of setting up the permanent sample plots is to monitor
long-term impact of logging on forest dynamic, growth and yield.
The plot locations were selected randomly according to the density
of harvestable timber trees (diameter at breast height or dbh ≥ 60cm)
prior to logging using the tree maps and inventory data in the logging
compartments prepared by CIFOR and PT INHUTANI II. A ield
checking was carried out before delimiting each plot to asses the
efective density of harvestable trees. Between June 1998 and September
1998, 12 one-hectare plots were set up in Petak (compartment) 28/29
(Figure 2).
Each one-hectare plot (100 m x 100 m) was horizontally corrected
during its establishment. The plot was divided into 25 subsquares
(20m x 20m), each being divided into 4 elementary squares (a,b,c,d,
in Figure 3 below). At 20 m intervals throughout the plot, PVC piping
was driven into the soil, and a metal tag attached to indicate the sub
square’s coordinates within the plot. A total of 36 PVC stakes were
placed throughout the plot. A wood panel indicating the block and
the number of the plot marks the centre of each. Three control plots
(CNV 04, CNV 08, CNV 12,) were designated and in each one, a 50 m
buffer zone was set up to avoid edge effect caused by logging activities.
After logging, all plots were re-grided (every subsquare 20m x 20 m
materialised by rope) and permanently marked in the ield with iron
wood/ulin poles located at each corner.
9
Access to a Plot
Visitors are advised to refer to the irst map (Figure 4) to get an
overview on how the plots are spatially distributed. The plots are
grouped based on their route. The routes follow the best walking way
to go to each plot, that is the available skidding road in the ield.
To enter a route, visitors should ind direction sign in the ield posted
on the entrance of each route and follow a skidding road directed by
the sign. Zoomed maps of the routes are given in Figures 4.
Figure 3. Permanent Sample Plot structure (PVC piping is located at
each number)
20m
20m
31 0
20m
32
c
b
a
c
b
a
26
25 0
c
a
b
c
a
b
d
c
b
a
d
c
d
c
d
c
b
a
b
a
b
a
b
d
c
d
a
c
b
d
b
d
c
b
a
North
West
East
South
18
c
a
b
a
b
d
c
d
c
d
c
b
a
b
a
b
a
9
10
9
11
3
3 0
11
10
2
2 0
17
d
8
1 0
17
12
4
4 0
5
5 0
20m
a
24
16
16
c
1
23
15
8
a
23
22
d
7 0
b
d
b
20m
b
a
30
22
15
7
d
a
14
a
c
b
21
14
c
d
29
a
21
d
b
29
c
20
c
a
d
20
13
b
28
d
20m
c
13 0
a
c
28
d
19
a
b
36
d
20m
c
19 0
c
27
d
35
d
27
26
20m
34
d
20m
c
25
a
20m
33
d
b
6 0
10
Plant Identiication
Trees with dbh ≥ 20 cm and dbh ≥ 10 cm were identiied in the ield by
botanists of the Herbarium Bogoriense, Puslitbang Biologi-LIPI, Bogor.
Voucher specimens were collected for further identiication and are
stored at the Herbarium Bogorriense. The identiication is not inal,
hence any corrections are welcome and will be very much appreciated.
The nomenclature follows T.C. Whitmore, I.G.M. Tantra and U. Sutisna
(Editor), Tree Flora of Indonesia: Check List for Kalimantan, Agency
for Forestry Research and Development, Ministry of Forestry, Bogor,
and C.G. G.J. Van Steenis, W.J.O. De Wilde, C. Kalkman, etc. (Editor),
Flora Malesiana Serie I, Volumes 1-15, 1948-2001.
Figure 4. Distribution of Permanent Sample Plots in Petak (Compartment) # 28 - 29 (Conventional logging treatment)
Route #1
Route# 2
Route# 3
CNV06, CNV01, CNV04, CNV07
CNV05, CNV11, CNV09, CNV12, CNV03
CNV15, CNV14, CNV02, CNV10, CNV08, CNV13, CNV16
CNV08
an
tur
Se
CNV10
CNV02
CNV11
CNV05
CNV09
CNV03
CNV01
CNV06
Legend
ROAD
RIVER
PSP
BRIDGE
FIRE TOWER
TPN 1
CIFOR RESEARCH STATION
CNV07
CNV04
A Field Guide to the Permanent Sample Plots • 11
CNV12
40
60
80
163
161
160
168
174 175
167
162
194
176
164
165
188
187 189
190
183
180
177
173
169
100
182
181
179
184
136
139
144
148 145
146
147
158
153
193
141
143 142
149
157
192
130
131
150
191
185 186
80
178
151
100
20
159
80
100
0
132
133
128
129
135
85
104
107
103
102
86
95
92
90
88
93
110
94
89
82
121
117 116
125
124
109
111
113
114
112
101
100
123
122
99
81
40
108
87
83
48
80
63
79
72 71
78
74
70
67
68
49
45
53
66
46
44
54
69
41
59
76
65
20
75
73
1
64
57 55
14
5
15
4
23
22 21
24
12
19
40
42
39
18
11
10
35
34
25
27
7
38
28
31
30
17
36
0
0
20
40
60
CNV01
Icacinaceae
FAMILY
Lauraceae
Anacardiaceae
Melastomataceae
Annonaceae
Meliaceae
Apocynaceae
Myristicaceae
Arecaceae
Myrtaceae
Bombacaceae
Oxalidaceae
Burseraceae
Podocarpaceae
Celastraceae
Rosaceae
Clusiaceae
Sapindaceae
Dilleniaceae
Sapotaceae
Dipterocarpaceae
Simarubaceae
Ebenaceae
Sterculiaceae
Euphorbiacea
Theaceae
Fabaceae
Ulmaceae
Fagaceae
Verbenaceae
Flacourtiaceae
0
9
43
56 58
26 60
61
33
13
8
47
52 51
50
77
2
120
119
40
97 98
60
105 106
96
91
20
60
127
84
80
100
12 • A Field Guide to the Permanent Sample Plots
Figure 5. Distribution of Trees in Plot CNV01
A Field Guide to the Permanent Sample Plots • 13
Table 1. List of Trees in Plot CNV01
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea parvifolia Dyer
Shorea fallax Meijer
Unidentiied
Diospyros sp. 1
Parashorea smythiesii Wyat-Smith ex Ashton
Unidentiied
Shorea parvifolia Dyer
Palaquium rostratum (Miq.) Burk.
Shorea paucilora King
Dipterocarpus stellatus Vesque
Shorea parvifolia Dyer
Dacryodes rostrata H.J.Lam forma pubescens
Shorea beccariana Burk.
Lithocarpus gracilis (Korth.) Soepadmo
Syzygium sp.
Dead
Alseodaphne paludosa Gamble
Gluta wallichii (Hk.f.) Ding Hou
Canarium caudatum King
Myristica beccarii Warb
Licania splendens (Korth.) Prance
Shorea sp.
Syzygium sp.
Licania splendens (Korth.) Prance
Sindora sp.
Melanochyla auriculata Hook.f.
Shorea fallax Meijer
Parashorea smythiesii Wyat-Smith ex Ashton
Dead
Diospyros sp.
Neesia cf. altissima Bl.
Lithocarpus nieuwenhuisii (V.Seem) A. Camus
Alseodaphne havilandii (Gamble) Kosterm.
Shorea fallax Meijer
Pternandra rostrata (Cogn.) Ohwi
Lithocarpus blumeanus (Korth.) Rehd.
Dead
14 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Livistona sp.
Livistona sp.
Tetramerista glabra Miq.
Madhuca sp.
Tetramerista glabra Miq.
Pternandra caerulescens Jack.
Lithocarpus benetii (Miq.) Rehd.
Dacryodes rugosa (Bl.) H.J. Lam
Diospyros borneensis Hiern)
Lithocarpus urceolaris (Jack) Merr.
Xylopia altissima Boerl.
Syzygium tawahense (Korth) Merr. & Perry
Shorea parvifolia Dyer ssp.parvifolia
Alseodaphne havilandii (Gamble) Kosterm.
Palaquium rostratum (Miq.) Burk.
Polyalthia sumatrana King
Aglaia cordata Hiern.
Livistona sp.
Parashorea smyethiesi Wyatt-Smith exAshton
Cleistanthes myrianthus Krz.
Vatica rassak (Korth.) Blume
Aporusa falcifera Hk.f.
Shorea ovalis (Korth.) Bl.
Hydnocarpus cf. kunstleri (King) Warb.
Shorea ovalis (Korth.) Bl.
Diospyros ellipticifolia Merr.
Syzygium perpuncticulatum (Merr.) Merr. & Perry
Shorea sp.
Litsea elliptica Bl.
Myristica maxima Warb.
Bhesa paniculata Arn.
Ryparosa acuminata Merr.
Sarcotheca diversifolia (Miq.) Hall.f.
Parashorea smythiesii Wyat-Smith ex Ashton
Heritiera sp.
Artocarpus lanceifolius Roxb.
Garcinia nervosa Miq.
A Field Guide to the Permanent Sample Plots • 15
Table 1. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Syzygium sp.
Actinodaphne glabra Bl.
Stemonurus grandifolius Becc.
Syzygium endertii Merr. & Perry
Artocarpus lanceifolius Roxb.
Diospyros
Irvingia malayana Oliver
Castanopsis philippinensis Vidal
Dillenia exelsa (Jacq.) Gilg.
Parashorea smythiesii Wyat-Smith ex Ashton
Lithocarpus ewyckii (Korth) Rehd.
Nephelium maingayi Hiern.
Dillenia exelsa (Jacq.) Gilg.
Diospyros sp.
Parashorea smythiesii Wyat-Smith ex Ashton
Diospyros sp.
Dehaasia cf. irma Bl.
Podocarpus becarii Parl.
Canarium denticulatum Bl. ssp. denticulatum
Cephalomappa lepidotula Airy Shaw
Tetramerista glabra Miq.
Garcinia maingayi Hook.f.
Popowia pisocarpa (Bl.) Endl.
Castanopsis motleyana King
Shorea ovalis Korth.) Bl.
Cleistanthus myrianthus Krz.
Koompassia malaccensis Maing. ex Benth.
Teijsmaniodendron simplicifolium Merr.
Diospyros macrophylla Bl.
Dyera costulata (Miq.) Hk.f.
Sindora leiocarpa Backer
Teijsmaniodendron simplicifolium Merr.
Syzygium sp.
Shorea agami Ashton
Dipterocarpus humeratus Sloot.
Heritiera simplicifolia (Mart.) Kosterm.
Syzygium tawahense (Korth.) Merr. & Perry
16 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Coccoceras borneense J.J. Smith
Teijsmaniodendron simplicifolium Merr.
Sarcotheca diversifolia (Miq.) Hall.f.
Teijsmaniodendron simplicifolium Merr.
Teijsmaniodendron simplicifolium Merr.
Archidendron fagifolium (Bl. ex Miq.) Nielsen
Dillenia eximia Miq.
Girroniera nervosa Planch.
Dyera costulata (Miq.) Hk.f.
Dacryodes sp.
Scaphium macropodum (Miq.) Beumee ex Heyne
Alseodaphne insignis Gamble
Teijsmaniodendron simplicifolium Merr.
Heritiera simplicifolia (Mart.) Kosterm.
Dead
Palaquium ferox H. J. Lam
Dacryodes sp.
Palaquium calophyllum Pierre ex Burck
Heritiera simplicifolia (Mart.) Kosterm.
Shorea parvifolia Dyer
Teijsmaniodendron simplicifolium Merr.
Tetramerista glabra Miq.
Dead
Palaquium gutta (Hook.f.) Bail
Syzygium chlorantum (Dut.) Merr. & Perry
Baccaurea sumatrana M.A.
Herritiera javanica (Bl.) Kosterm
Cleistanthus myrranthus (Krz.
Dead
Palaquium ferox H.J. Lam
Scaphium macropodum (Miq.) Beumée ex Heyne
Knema conferta (King) Warb
Artocarpus odoratisima Blanco
Oncosperma horridum Scheff
Oncosperma horridum Scheff
Scaphium macropodum (Miq.) Beumée ex Heyne
Canarium denticulatum Bl. ssp. denticulatum
A Field Guide to the Permanent Sample Plots • 17
Table 1. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Santiria oblongifolia Bl.
Syzygium sp.
Shorea ovalis (Korth.) Bl.
Castanopsis hypophoenicea (V.Sleum.) Supadmo
Bhesa paniculata Arn.
Shorea ovalis (Korth.) Bl.
Macaranga pruinosa (Miq.) Muell. Arg
Dead
Artocarpus kemando Miq
Parashorea smythiesii Wyatt-Smith ex Ashton
Shorea parvifolia Dyer
Shorea ovalis (Korth.) Bl.
Baccaurea sumatrana M.A.
Durio sp.
Artocarpus elasticus Reinw.
Licania splendens (Korth.) Prance
Heritiera simplicifolia (Mart.) Kosterm.
Alseodaphne insignis Bamble
Vatica vinosa Ashton
Polyalthia rumphii Merr.
Vatica vinosa Ashton
Dipterocarpus stellatus Vesque
Sindora leiocarpa Backer
Artocarpus kemando Miq.
Durio oxleyanus Griff.
Heritiera simplicifolia (Mart.) Kosterm.
Licania splendens (Korth.) Prance
Myristica beccarii Warb.
Aglaia cordifolia Merr.
Nephelium uncinatum Leenh.
Shorea pinanga Scheff.
Dialium sp.
Dyera costulata (Miq.) Hk.f.
Shorea pinanga Scheff.
Knema cinerea (Poir.) Warb.
Knema cinerea (Poir.) Warb.
Licania splendens (Korth.) Prance
18 • A Field Guide to the Permanent Sample Plots
Table 1. Continued
Tree #
186
187
188
189
190
191
192
193
194
Species
Teijsmaniodendron simplicifolium Merr.
Dehaasia irma Bl.
Canarium denticulatum Bl. ssp. denticulatum
Shorea macroptera Dyer
Alseodaphne sp.
Syzygium tawahense (Korth.) Merr. & Perry
Garcinia sp.
Shorea fallax Meijer
Syzygium tawahense (Korth.) Merr. & Perry
252
254
245
261
240
251
249
271 272
255
253
250
256
262
264
211
220
225
231
222
229
224
60
126
124
131
40
129
203
119
165
164
96
85
98
105
33
34
75
84
80
70
78
36
35
16
31
30
45
46
67
37
29
66
64
60
63 61
44
21
56
53
27
23
24
48
38
25
41 42
40
39
49
52
55
51
0
20
40
60
80
CNV02
Flacourtiaceae
FAMILY
Lauraceae
Anacardiaceae
Meliaceae
Annonaceae
Myristicaceae
Apocynaceae
Myrtaceae
Araucariaceae
Ochnaceae
Arecaceae
Olacaceae
Bombacaceae
Oxalidaceae
Burseraceae
Polygalaceae
Celastraceae
Proteaceae
Clusiaceae
Rosaceae
Crypteroniaceae
Rubiaceae
Dilleniaceae
Sabiaceae
Dipterocarpaceae
Sapindaceae
Ebenaceae
Sapotaceae
Elaeocarpaceae
Sterculiaceae
Euphorbiacea
Thymelaeaceae
Fabaceae
Tiliaceae
Fagaceae
50
0
22
57
0
1
233
24 5
6
3
71
69 68
59
62
28
7
171
72
74
79
90
47
17
77
76
91
18
170
169
81
86
92
99
107
174
172
82
88
104
9
173
166
83
93
108
113 12
13 11 10
191
168
155
154 89
101
112
175
178
156
132
100
117
176
177
190
140
130
121
189
100
A Field Guide to the Permanent Sample Plots • 19
125
116
115
200
207
210
215
208
216 145 147
148
138 134 135
149
144
139
146
150 161 159
143 142
128 133
141
152
136
122
214
180
209
213
217
230
183
184
185
188
192
204
199
218
223
182
187 186
193
198
212
235
20
277
194
197
221
283
285
284
273
270
227 228
232
14
278
269
263
257
195
234
114
286
281
282
274
248
236
100
280
265
239
80
267
266
258
246
247
80
275
100
60
260
80
40
259
60
20
244
243
242
40
0
20
100
Figure 6. Distribution of Trees in Plot CNV02
20 • A Field Guide to the Permanent Sample Plots
Table 2. List of Trees in Plot CNV02
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Anisoptera sp.
Castanopsis oviformis Soepadmo
Castanopsis oviformis Soepadmo
Diospyros sp.
Baccaurea minor Hook.f.
Litsea grandis (Wall. ex Nees) Hook.f.
Baccaurea minor Hook.f.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Pimeleodendron papaveroides J.J.S.
Shorea parvifolia Dyer
Dacryodes rostrata H.J.Lam
Dacryodes rostrata H.J.Lam
Sterculia macrophylla Vent.
Oncosperma horridum Scheff.
Oncosperma horridum Scheff.
Koordersiodendron pinnatum (Blanco) Merr.
Saraca hullettii Prain.
Lithocarpus urceolaris (Jack.) Merr.
Knema latifolia Warb.
Shorea paucilora King
Nephelium uncinatum Leenh.
Knema cinerea (Poir.) Warb.
Dysoxylum sp.
Santiria oblongifolia Bl.
Alseodaphne ceratoxylon Kosterm.
Sarcotheca diversifolia (Miq.) Hall.f.
Dehaasia irma Bl.
Irvingia malayana Oliv.
Melanochyla beccariana Oliv.
Dead
Castanopsis tungurut (Bl.) A.DC.
Sindora sp.
Alseodaphne ceratoxylon Kosterm.
Parashorea smythiesii Wyatt-Smith ex Ashton
Parashorea smythiesii Wyatt-Smith ex Ashton
A Field Guide to the Permanent Sample Plots • 21
Table 2. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Santiria laevigata Bl.
Vatica rassak (Korth.) Blume
Alseodaphne cuneata (Bl.) Kosterm.
Myristica sp.
Polyalthia sp.
Dead
Syzygium sp.
Lithocarpus ulceolaris (Jack) Merr.
Canarium caudatum King
Shorea macroptera Dyer.
Xanthophyllum parvum Chod.
Timonius ovalis Boerl
Shorea parvifolia Dyer
Maccaranga gigantea M.A.
Shorea parvifolia Dyer
Dehaasia irma Bl.
Dead
Syzygium sp.
Polyalthia glauca Boerl.
Shorea seminis Slooten
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Shorea macroptera Dyer
Artocarpus odoratissimus Blanco
Knema intermedia Warb.
Helicia fuscotomentosa Suesseng
Dead
Polyalthia laterilora (Bl.) King
Shorea exelliptica Meijer
Aporusa antennifera (Airy Shaw) Airy Shaw
Santiria laevigata Bl. f. laevigata
Lithocarpus bennetii (Miq.) Rehd.
Dyera costulata (Miq.) Hk.f.
Agathis borneensis Warb.
Dead
Macaranga pruinosa M.A.
22 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Ochanostachys amentacea Mast.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Elaeocarpus griithii A. Gray
Syzygium sp.
Vatica umbonta (Hook.f.) Burk.
Dyera costulata (Miq.) Hk.f.
Diospyros kunstleri King & Gamble
Shorea parvifolia Dyer.
Myristica beccarii Warb.
Baccaurea delexa M.A.
Pometia pinnata Forst.
Dead
Melanochyla beccariana Oliv.
Chisocheton sp.
Syzygium sp.
Shorea parvifolia Dyer
Shorea paucilora King
Shorea parvifolia Dyer
Dead
Shorea parvifolia Dyer
Shorea paucilora King
Unidentiied
Koompassia malaccense Maing.. ex Benth.
Aquilaria malaccensis Lam.
Koompassia excelsa Taub.
Knema glauca (Bl.) Warb.
Dead
Dead
Pentace erectinervia Kosterm.
Dillenia excelsa (Jack.) Gilg.
Albizia sp.
Knema conferta (King) Warb.
Macaranga hypoleuca (Richb. f & Zoll.) M.A.
Uidentiied
Dead
Saraca hulletii Prain
A Field Guide to the Permanent Sample Plots • 23
Table 2. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Syzygium operculata Gamb.
Eugeissona utilis Becc.
Mangifera swintonioides Kosterm.
Saraca hulletii Prain
Shorea parvifolia Dyer
Mangifera swintonioides Kosterm.
Knema cinerea (Poir.) Warb
Livistona sp.
Saraca hulletii Prain
Macaranga hypoleuca (Richb. f & Zoll.) M.A.
Shorea parvifolia Dyer
Unidentiied
Shorea parvifolia Dyer
Diospyros ellipticifolia Merr.
Syzygium sp.
Dead
Cococeras borneense J.J. Smith
Lithocarpus nieuwenhuisii (v. Seem) A. Camus
Knema laurina (Bl.) Warb.
Baccaurea kunstlerii King
Shorea parvifolia Dyer
Koompassia malaccensis Maing.. ex Benth.
Shorea parvifolia Dyer
Xanthophyllum amoenum Chod.
Baccaurea angulata Merr.
Unidentiied
Canarium denticulatum Bl. ssp. denticulatum
Shorea parvifolia Dyer ssp. velutina
Knema cinerea (Poir.) Warb.
Ptychophyxys bacciformis Croiz.
Ptychophyxys bacciformis Croiz.
Koompassia malaccensis (Clarke) Breh.
Macaranga hypoleuca (Reichb.f. & Zoll.) M.A.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Pouteria malaccensis (Clarke) Breh.
Knema sp.
24 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Lythocarpus ewyckii (Korth.) Rehd.
Cephalomappa lepidotula Airy Shaw
Shorea parvifolia Dyer
Koompassia malaccensis Maing.. ex Benth.
Girroniera nervosa Planch.
Shorea paucilora King
Artocarpus kemando Miq.
Artocarpus lanceifolius Roxb.
Unidentiied
Unidentiied
Lythocarpus ewyckii (Korth.) Rehd.
Macaranga hypoleuca (Reichb.f. & Zoll.) M.A.
Artocarpus lanceifolius Roxb.
Dead
Mangifera swintonoides Kosterm.
Crypteronia grifithii Clarke
Payena endertii H.J.Lam
Glochidion macrocarpum Bl.
Dead
Polyalthia sclerophylla Hook.f. ex Th.
Dialium trifoliatum de Wit
Garcinia dioica Bl.
Mangifera swintonioides Kosterm.
Palaquium calophyllum Pierre ex Burck
Lithocarpus ewyckii (Korth.) Rehd.
Shorea parvifolia Dyer
Dacryodes rostrata H.J. Lam
Dacryodes rostrata H.J. Lam
Shorea paucilora King
Durio oxleyanus Griff.
Dead
Shorea exelliptica Meijer
Knema latericia Elmer
Shorea sp.
Buchanania sp.
Shorea exelliptica Meijer
Scaphium macropodum (Miq.) Buemee ex Heyne
A Field Guide to the Permanent Sample Plots • 25
Table 2. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Knema sp.
Dialium trifoliatum De Wit
Atuna excelsa (Jack.) Kosterm.
Dialium trifoliatum De Wit
Koordersiodendron pinnatum (Blanco) Merr.
Dacryodes rostrata H.J. Lam
Drypetes kikir Airy Shaw
Shorea paucilora King
Mangifera swintonioides Kosterm.
Helicia serrata (R.Br.) Bl.
Dead
Baccaurea edulis Merr.
Knema latericia Elmer.
Durio lanceolatus Mast.
Dialium platycephalum Backer
Lithocarpus ewyckii (Korth.) Rehd.
Mangifera swintonioides Kosterm.
Sindora leiocarpa Backer ex Heyne
Oncosperma horridum Scheff.
Canarium denticulatum Bl. ssp. denticulatum
Dipterocarpus sp.
Crypteronia grifithi Clarke
Hydnocarpus anomala (Merr.) Sleumer
Dialium platycephalum Backer
Artocarpus kemando Miq.
Knema furfuracea (Hk.f. ex Th.) Warb.
Cratoxylum formosum Benth. & Hook. f. ex Dyer
Vatica umbonata (Hook.f.) Burk.
Dipterocarpus
Artocarpus lanceifolius Roxb.
Palaquium calophyllum Pierre ex Burck
Shorea parvifolia Dyer
Parashorea amythiesii Wyatt-Smith ex Ashton
Dead
Pouteria malaccensis (Clarke) Breh
Diospyros sp.
Meliosma nitida
26 • A Field Guide to the Permanent Sample Plots
Table 2. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Syzygium sp.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Unidentiied
Mangifera magniica Kochumen
Syzygium sp.
Nephelium eriobetalum Miq.
Polyalthia laterilora (Bl.) King
Bhesa paniculata Arm.
Garcinia dioica Bl.
Vatica micrantha Slooten
Dacryodes incurvata (Engl.) H.J.Lam.
Pometia pinnata Forst.
Dacryodes costata (Benn.) H.J. Lam
Artocarpus integer (Thunb.) Merr.
Vatica micrantha Slooten
Cococeras borneensis M.A.
Dipterocarpus cornutus Dyer
Dead
Dipterocarpus confertus Sloot.
Agathis borneensis Warb.
Cococeras borneensis M.A.
Baccaurea sp.
Beilschmiedia sp.
Diopyros ellipticifolia Merr.
Agathis borneensis Warb.
Diospyros everetii Merr.
Bhesa paniculata Arn.
Elateriospermum tapos Bl
Ochanostachys amentacea Mast.
Cleistanthus myrianthus (Hassk.) Kurz.
Palaquium sp.
Sarcotheca diversifolia (Miq.) Hall.f.
Palaquium ottolanderi K. ex V.
Prunus sp.
Palaquium calophyllum Pierre ex Burck
Dysoxylum sp.
A Field Guide to the Permanent Sample Plots • 27
Table 2. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
Species
Cephalomappa beccariana Baill.
Sindora leiocarpa Backer
Dipterocarpus humeratus Sloot.
Elateriospermum tapos Bl.
Madhuca cf. rupicola H.J.Lam
Dacryodes rugosa H.J.Lam var. rugosa
Xanthophyllum vittelinum (Bl.) Dierr.
Xylopia malayana Hook. f. & Thoms.
Dead
Plectronia glabra Benth. & Hook. ex Kurz
Shorea macroptera Dyer.
Brownlowia sp.
Syzygium sp.
Dacryoides rostrata H.J. Lam
Shorea sp.
Dipterocapus stellatus Vesque
Unidentiied
Shorea macroptera Dyer.
Nauclea subdita Steud.
Dead
Shorea parvifolia Dyer.
Dipterocapus stellatus Vesque
Shorea parvifolia Dyer.
Helicia serrata Bl.
Dipterocapus stellatus Vesque
Gymnacranthera paniculata Warb.
Gymnacranthera paniculata Warb.
40
60
80
100
189
209 211
210
208
198
193
192
183
203
182
80
180
217
205
204
163
165
169
168 167
206
202
185
145
159
158 160
166
181
141
161
144 143
162
170
154
153
172
115
82
89
124
117
108
110
126
80
79
85
54
68
58
63
69
77
71
76
95
51
56
53
70
61
75
60
62
15
26
25
49
44
17
36
9
6
1
3
10
5
11
24
23
18
4
20
14 13
40
38
22
21
37
30
19
39
45
46
0
32
0
12
42
34
28
0
20
40
60
80
CNV03
Lauraceae
FAMILY
Meliaceae
Anacardiaceae
Myristicaceae
Annonaceae
Myrtaceae
Arecaceae
Olacaceae
Bombacaceae
Oleaceae
Burseraceae
Proteaceae
Celastraceae
Rosaceae
Clusiaceae
Rubiaceae
Dilleniaceae
Sapindaceae
Dipterocarpaceae
Sapotaceae
Ebenaceae
Sterculiaceae
Euphorbiacea
Thymelaeaceae
Fabaceae
Tiliaceae
Fagaceae
Ulmaceae
48
47
29
8
7
50
52
59
86
20
55
57
66 64
78
94
87
128
65
73 72
133
135
127
74
84
120
118
83
88
90
129
111
81
130
112
40
60
40
91
93
107
102 101
132
122
114
106
98
149
123
113
116
99
137
148
151
121
139 138
147
131
174
105
150
152
173
103
100
142
140
146
156 157
171
176
178
215
207
201
199
212
80
190
184
200
197
195
194
60
196
191
186
100
20
20
100
0
100
28 • A Field Guide to the Permanent Sample Plots
Figure 7. Distribution of Trees in Plot CNV03
A Field Guide to the Permanent Sample Plots • 29
Table 3. List of Trees in Plot CNV03
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Polyalthia glauca Boerl.
Dead
Croton laevifolim Bl.
Artocarpus kemando Miq.
Eugeissona utilis Becc.
Shorea ovalis (Korth.) Bl.
Myristica maxima Warb.
Coccoceras borneense J.J. Smith
Beilschmiedia bankae Kosterm.
Syzygium splendens (Bl.) Merr. & Perry
Gonystylus brunnescens Airy Shaw
Baccaurea
Vatica rassak (Korth.) Blume
Syzygium splendens (Bl.) Merr. & Perry
Vatica oblongifolia Hook.f.
Unidentiied
Vatica rassak (Korth.) Blume
Syzygium splendens (Bl.) Merr. & Perry
Parashorea smythiesii Wyatt-Smith ex Ashton
Cephalomappa lepidotula Airy Shaw
Cephalomappa lepidotula Airy Shaw
Vatica micrantha Slooten
Coccoceras borneense J.J. Smith
Shorea ovalis (Korth.) Bl.
Dillenia excelsa (Jacq.) Gilg.
Shorea parvifolia Dyer
Dead
Cephalomappa lepidotula Airy Shaw
Dyospiros sp.
Syzygium operculata Gamble
Dead
Parashorea sp.
Dead
Vatica rassak (Korth.) Blume
Turpinia sphaerocarpa Hassk.
Litsea noronhae Bl.
Coccoceras borneense J.J. Smith
30 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Shorea paucilora King
Durio kuteyensis Becc.
Dysoxylum sp.
Pternandra rostrata (Cogn.) Ohwi
Artocarpus lanceifolius Roxb.
Dead
Dysoxylum hexandrum Merr.
Scaphium macropodum (Miq.) Beumée ex Heyne
Lithocarpus ewyckii Korth.) Rehd.
Lithocarpus ewyckii Korth.) Rehd.
Myristica malaccensis Hk.f.
Aglaia cordata Hiern.
Lithocarpus ewyckii (Korth.) Rehd.
Shorea ovalis Korth. Bl.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Gymnacranthera paniculata Warb.
Pentace laxilora Merr.
Bhesa paniculata Arn.
Artocarpus anysophyllus Miq.
Chionanthes cuspidatus Bl.
Coccoceras borneense J.J. Smith
Chionanthus sp.
Parashorea smythiesii Wyatt-Smith ex Ashton
Baccaurea sp.
Parashorea smythiesii Wyatt-Smith ex Ashton
Drypetes longifolia Pax. & Hoffm.
Shorea ovalis (Korth.) Bl.
Syzygium sp.
Dead
Coccoceras borneense J.J. Smith
Baccaurea sp.
Dipterocarpus stellatus Vesque
Diospyros malabarica (Desr.) Kostel.
Pentace discolor Merr.
Herritiera javanica (Bl.) Kosterm.
Coccoceras borneense J.J. Smith
A Field Guide to the Permanent Sample Plots • 31
Table 3. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Coccoceras borneense J.J. Smith
Coccoceras borneense J.J. Smith
Shorea maxwelliana King
Licania spendens (Korth.) Prance
Chionanthus cuspidatus Bl.
Parashorea smythiesii Wyatt-Smith ex Ashton
Coccoceras borneense J.J. Smith
Shorea parvifolia Dyer
Lithocarpus nieuwenhuisii (V.Seem) A.Camus
Shorea parvifolia Dyer
Syzygium sp.
Knema cinerea (Poir) Warb.
Licania splendens (Korth.) Prance
Vatica rassak (Korth.) Blume
Castanopsis hypophoenica (V.Seem) Soepadmo
Diospyros malabarica (Desr.) Kostel.
Vatica rassak (Korth.) Blume
Planchonia valida (Bl.) Bl.
Vatica micrantha Slooten
Shorea fallax Meijer
Shorea paucilora King.
Syzygium splendens (Bl.) Merr. & Perry
Shorea fallax Meijer
Licuala sp.
Licuala sp.
Neonauclea purpurascens Moore
Shorea ovalis (Kort.) Bl.
Knema cinerea (Poir.) Warb.
Cratoxylum sumatranum (Jack.) Bl.
Dead
Aporusa falcifera Hk.f.
Shorea paucilora King
Castanopsis javanica (Bl.) A.DC.
Dipterocarpus stellatus Vesque
Dead
Palaquium sumatranum Burck.
Syzygium sp.
32 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Drypetes longifolia Pax. & Hoffm.
Durio carrinatus Mast.
Syzygium sp.
Dysoxylum hexandrum Merr.
Dehaasia irma Bl.
Parashorea smythiesii Wyatt-Smith ex Ashton
Ochanostachys amentacea Mast.
Dead
Dacryodes rogosa H.J.Lam
Coccoceras borneense J.J. Smith
Knema furfuracea (Hk. f. ex Th.) Warb.
Parashorea smythiesii Wyatt-Smith ex Ashton
Shorea pinanga Scheff.
Dead
Shorea beccarina Burk.
Shorea parvifolia Dyer
Knema intermedia (Bl.) Warb
Shorea ovalis (Kort.) Bl.
Shorea paucilora King
Artocarpus kemando Miq.
Koompassia malaccensis Maing. ex Benth
Dialium patens Backer
Dead
Palaquium rostratum (Miq.) Burk.
Dead
Shorea paucilora King
Parashorea smythiesii Wyatt-Smith ex Ashton
Syzygium splendens (Bl.) Merr. & Perry
Lithocarpus blumeanus (Korth.) Rehd.
Gymnacranthera paniculata Warb.
Knema furfuracea (Hk.f. ex Sinel.) Warb
Knema laurina (Bl.) Warb.
Vatica rassak (Korth.) Blume
Durio oxleyanus Griff.
Knema laurina (Bl.) Warb.
Pimeleodendron papaveroides J.J.S.
Parashorea smythiesii Wyatt-Smith ex Ashton
A Field Guide to the Permanent Sample Plots • 33
Table 3. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea pinanga Scheff.
Syzygium angustifolius (Miq.) Hats.
Drypetes sp.
Pometia pinnata Forst.
Durio testudinarum Becc.
Parashorea smythiesii Wyatt-Smith ex Ashton
Dead
Diospyros sp.
Parashorea smythiesii Wyatt-Smith ex Ashton
Knema glauca (Bl.) Warb.
Garcinia parvifolia Miq.
Knema glauca (Bl.) Warb.
Lithocarpus conocarpus (Oudem) Rehd
Shorea seminis (de Vriese) Slooten
Shorea pinanga Scheff
Dead
Coccoceras borneense J.J. Smith
Shorea rubra Ashton
Coccoceras borneense J.J. Smith
Vatica micrantha Slooten
Baccaurea sp.
Cleisthanthus myrianthus Krz.
Parashorea smythiesii Wyatt-Smith ex Ashton
Shorea rubra Ashton
Pterocymbium tubulatum (Mast.) Pierre
Alseodaphne ceratoxylon Kosterm.
Aporusa falcifera Hk.f.
Shorea parvifolia Dyer
Actinodaphne glabra Bl.
Pterocymbium tubulatum (Mast.) Pierre
Dead
Livistona
Shorea parvifolia Dyer
Payena accuminata (Bl.) Pierre
Palaquium rostratum (Miq.) Burk.
Gymnacranthera paniculata Warb.
Diospyros everetii Merr.
34 • A Field Guide to the Permanent Sample Plots
Table 3. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
Species
Knema furfuracea (Hk.f.ex. Th.) Warb.
Dead
Livistona sp.
Dialium modustum (V. Stenis) Stegaert
Calophyllum soulatri Burm.
Cratoxylum formosum (Jack) Dyer
Syzygium splendens (Bl.) Merr. & Perry
Aglaia cf. pseudolansium Kosterm.
Shorea paucilora King
Mangifera quadriida Jack.
Dialium modestum (V. Steenis) Stegaert.
Pentace discolor Merr.
Syzygium splendens (Bl.) Merr. & Perry
Aporusa falcifera Hk.f.
Cleisthanthus myrianthus Krz.
Castanopsis javanica (Bl.) A.DC.
Celtis nigrecens (Miq.) Planch.
Sindora leiocarpa Backer
Diospyros cf. oblonga Wall. ex. G. Don
Prunus sp.
Coccoceras borneense J.J. Smith
Shorea fallax Meijer
Castanopsis javanica (Bl.) A.DC.
Helicia petiolaris Benn.
Coccoceras borneense J.J. Smith
Myristica sp.
Diospyros ellipticifolia Merr.
Aporusa falcifera Hk.f.
Gardenia tubifera Wall.
Diospyros oblonga Wall. ex. G. Don
Calophyllum soulatri Burm.
Diospyros oblonga Wall. ex. G. Don
20
40
175
174
180
170
197
186
193
194
185
80
168
198
152
157
164
163
165
162
200
215
213
211
140 139
141
212
220
223
219
221
222
153
147
146
149
160
145
144
148
156
138
132
137
150
155
210
201
151
158
159
161
202
195
184
179
169
203
214
183
166
204
199
100
218
217
216
209
208
192
196
178
171
207
206
205
187
177
167
80
191
182
176
172
60
190
189
188
131
136
130
143
129
133
154
128
95
91 96
105 108
107
106
90
98
87
40
88
84
68
67
81
82
75
Melastomataceae
Annonaceae
Meliaceae
Apocynaceae
Myristicaceae
Arecaceae
Myrtaceae
Bombacaceae
Oxalidaceae
Burseraceae
Polygalaceae
Celastraceae
Proteaceae
Clusiaceae
Rhizophoraceae
Dipterocarpaceae
Rosaceae
Ebenaceae
Rubiaceae
Elaeocarpaceae
Sapindaceae
Euphorbiacea
Sapotaceae
Fabaceae
Sterculiaceae
Fagaceae
Theaceae
50
49
20
21
10
47
59
56
12
14
2
3
45
36 37
17
35
32 33
26
18
31
19
20
27
13
38
39
30
29
0
28
0
1
9
41
34
25
15
7 8
46
54
57
42
44
58
22
16
5
43
53
23
11
6
48
52
60
73
0
20
40
60
Lauraceae
Anacardiaceae
61
74
Flacourtiaceae
FAMILY
120
63
62
CNV04
121
71
86
60
125
51
64
76
122
66
70
77
117
124
119
69
78
79
83
100
99
116
115
114 118
101 102
89
123
109
111
112
110
113
80
100
A Field Guide to the Permanent Sample Plots • 35
104
40
103
92
20
60
134
94
100
0
80
100
Figure 8. Distribution of Trees in Plot CNV04
36 • A Field Guide to the Permanent Sample Plots
Table 4. List of Trees on Plot# CNV04
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Aporusa falcifera Hk.f.
Pternandra rostrata (Cogn.) Ohwi
Alstonia angustiloba Miq.
Dead
Memecylon myrsinoides Blume
Syzygium bankense (Hassk.) Merr. & Perry
Dialium trifoliatum de Wit.
Artocarpus kemando Miq.
Shorea pinanga Scheff.
Aglaia cf. gamopetala Merr.
Litsea forstenii (Bl.) Boerl.
Palaquium rostratum (Miq.) Burck.
Dacryodes incurvata (Engl.) H.J.Lam
Sarcotheca diversifolia (Miq.) Hall.f.
Koompassia malaccensis Maing. ex Benth
Syzygium tawahense (Korth.) Merr. & Perry
Knema cinerea (Poir.) Warb.
Baccaurea kunstleri King.
Pometia pinnata Forst.
Dyera costulata (Miq.) Hk.f.
Sindora leiocarpa Backer
Parastemon urophyllus (A.DC.) A.DC.
Ternstroemia aneura Miq.
Dead
Calophyllum lowii Hook.f.
Cephallomappa lepidotula Airy Shaw
Myristica beccarii Warb.
Shorea parvifolia Warb.
Baccaurea bracteata M.A.
Myristica beccarii Warb.
Ternstroemia aneura Miq.
Tetramerista glabra Miq.
Dipterocarpus lowii Hk.f.
Myristica beccarii Warb.
Durio lanceolatus Mast.
Shorea seminis (de Vriese) Slooten
Canarium sp.
A Field Guide to the Permanent Sample Plots • 37
Table 4. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Calophyllum pulcherrimum Wall. ex Planch. & Trian.
Palaquium rostratum (Miq.) Burk.
Dead
Durio carinatus Mast.
Santiria laevigata Bl.f. laevigata
Vatica rassak (Korth.) Blume
Canarium littorale Bl.
Myristica beccarii Warb.
Shorea seminis (de Vriese) Slooten
Parastemon urophyllus (A.DC.) A.DC.
Santiria mollissima Ridl.
Shorea seminis (de Vriese) Slooten
Santiria laevigata Bl.
Syzygium tawahense (Korth.) Merr. & Perry
Xanthophyllum parvum Chod
Dyera costulata (Miq.) Hk.f.
Gymnacranthera contracta Warb.
Dead
Shorea seminis (de Vriese) Slooten
Gymnacranthera contracta Warb.
Horsieldia sp.
Syzygium sp.
Gluta wallichii (Hk.f.) Ding Hou
Dehaasia irma Bl.
Shorea ovalis (Korth.) Bl.
Gymnacranthera paniculata Warb.
Gymnacranthera paniculata Warb.
Dead
Knema sp.
Koompassia malaccensis Maing. ex. Benth.
Gluta wallichii (Hk.f.) Ding Hou
Sterculia oblongata R.Br.
Santiria oblongifolia Bl.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Shorea seminis (de Vriese) Slooten
Dialium indum L.
38 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Baccaurea pendula Merr.
Dyera costulata (Miq.) Hk.f.
Syzygium splendens (Bl.) Merr. & Perry
Xylopia caudata Hook. f. & Thoms.
Xylopia caudata Hook. f. & Thoms.
Dead
Santiria laevigata Bl.
Durio sp.
Helicia petiolaris Benn.
Baccaurea sumatrana M.A.
Dead
Gymnacranthera paniculata Warb.
Lophopetalum subobovatum King.
Syzigium fastigiata (Bl.) Merr. & Perry
Cephalomappa lepidotula Airy Shaw
Endiandra ochracea Kosterm.
Baccaurea sp.
Aporusa falcifera Hk.f.
Mastixia trichotoma Bl.
Dehaasia irma Bl.
Aporusa falcifera Hk.f.
Knema cinerea (Poir.) Warb
Dead
Shorea parvifolia Dyer
Parashorea smythiesii Wyatt-Smith ex Ashton
Ixonanthes petiolaris Bl.
Hydnocarpus woodii Merr.
Canarium litorale Bl.
Pternandra rostrata (Cogn.) Ohwi
Memecylon edule Roxb.
Castanopsis motleyana King
Shorea parvifolia Dyer
Syzygium litorale (Bl.) Merr. & Perry
Shorea parvifolia Dyer
Vatica vinosa Ashton
Shorea maxweliana King
Canarium litorale Bl.
A Field Guide to the Permanent Sample Plots • 39
Table 4. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Shorea parvifolia Dyer
Palaquium sp.
Gymnacranthera contracta War.
Nephelium maingayi Hiern.
Shorea agami Ashton
Gymnacranthera contracta War.
Pimeleodendron papaveroides J.J.S.
Dehaasia irma Bl.
Horsieldia sp.
Tetramerista glabra Miq.
Shorea leprosula Miq.
Shorea agami Ashton
Santiria laevigata Bl.
Xylopia altissima Boerl.
Dead
Madhuca malaccensis (Clarke) H.J.Lam.
Myristica sp.
Syzygium ochneocarpa Merr.
Palaquium rostratum (Miq.) Burk.
Myristica vilosa Warb.
Tetramerista glabra Miq.
Lithocarpus cantleyanus (King ex Hk.f.)Rehd.
Vatica rassak (Korth.) Blume
Dead
Palaquium rostratum (Miq.) Burk.
Diospyros oblonga Wall. ex G. Don
Xylopia altissima Boerl.
Madhuca malaccensis (Clarke) H.J.Lam
Pternandra rostrata (Cogn.) Ohwi
Castanopsis aviformis Soepadmo
Dead
Dacryodes incurvata (Engler) H.J.Lam
Syzygium sp.
Durio carinatus Mast.
Oncosperma horridum Scheff.
Shorea seminis (de Vriese) Slooten
Koordersiodendron pinnatum (Blanco) Merr.
40 • A Field Guide to the Permanent Sample Plots
Table 4. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Sarcotheca diversifolia (Miq.) Hall.f.
Tetramerista glabra Miq.
Ternstroemia aneura Miq.
Baccaurea pendula Merr.
Alseodaphne insignis Gamble
Parashorea smythiesii Wyatt-Smith ex Ashton
Aporusa falcifera Hk.f.
Buchanania arborescens Bl.
Madhuca malaccensis (Clarke) H.J.Lam
Endiandra sp.
Nephelium maingayi Hiern.
Pternandra rostrata (Cogn.) Ohwi
Shorea paucilora King
Pternandra rostrata (Cogn.) Ohwi
Shorea rubra Ashton
Parastemon urophyllus (A.DC.) A.DC.
Lithocarpus pulcher (King.) Markgr.
Dipterocarpus lowii Hk.f.
Sizygium splendens (Bl.) Merr. & Perry
Parishia maingayi Hk.f.
Xylopia malayana Hk.f. ex Th.
Elaeocarpus petiolatus Wall.
Artocarpus anisophyllus Miq.
Shorea parvifolia Dyer
Unidentiied
Xylopia malayana Hk.f. ex Th.
Parastemon urophyllus (A.DC.) A.DC.
Diospyros sp.
Castanopsis motleyana King
Palaquium rostratum (Miq.) Burk.
Madhuca malaccensis (Clarke) H.J.Lam.
Sarcotheca diversifolia (Miq.) Hall.f.
Dead
Myristica vilosa Warb.
Jackia ornata Wall.
Dehaasia irma Bl.
Syzigium chloranthum (Duth.) Merr. & Perry
A Field Guide to the Permanent Sample Plots • 41
Table 4. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
223
Species
Semecarpus glaucus Engl.
Pternandra rostrata (Cogn.) Ohwi
Cleisthantus myrianthus Krz.
Castanopsis motleyana King
Baccaurea racemosa (Rinw.) Muel. Arg.
Madhuca malaccensis (Clarke) H.J.Lam.
Cephalomappa lepidotula Airy Shaw
Vatica rassak (Korth.) Blume
Baccaurea edulis Merr.
Baccaurea edulis Merr.
Tetramerista glabra Miq.
Syzigium chloranthum (Duth.) Merr. & Perry
Baccaurea puberula Merr.
Lithocarpus urceolaris (Jack.) Merr.
Ternstroemia aneura Miq.
Baccaurea puberula Merr.
Sterculia foetida L.
Shorea parvifolia Dyer
Shorea seminis (de Vriese) Slooten
Syzygium bankense (Hassk.) Merr. & Perry
Shorea seminis (de Vriese) Slooten
Gymnacranthera paniculata Warb.
Ficus sp.
Syzigium chloranthum (Duth.) Merr. & Perry
Baccaurea sp.
Carallia brachyata (Lour.) Merr.
Xanthophyllum sp.
Tetramerista glabra Miq.
Knema sp.
Baccaurea puberula Merr.
Syzigium chloranthum (Duth.) Merr. & Perry
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Aporusa falcifera Hk.f.
Syzygium sp.
Baccaurea bracteata M.A.
Pternandra rostrata (Cogn.) Ohwi
Aporusa falcifera Hk.f.
40
60
274
265
264
285
275
272
269
259
296
291
273
261 260
80
279
100
300
298
289
284
302
297 299
283
270
303
258
256
254
292
282
293
257
255
244
242 240
80
253
246
294
306 305
295
200
224
173
239
241
100
20
263
262
304
199
196
194
198
197
195
80
100
0
193
248
192
252
188
227
236
60
60
250 249
123
251
122
145 158
138
148
146
151
149
150
156
152
154 153
157 80
132
40
133
97
114
112
111
110
95
81
82
104
92
20
87
91
93
90
36
162
163
34
18
23
27
28
Arecaceae
Melastomataceae
Bombacaceae
Meliaceae
58
Burseraceae
Myristicaceae
57
Celastraceae
Myrtaceae
55
Clusiaceae
Oxalidaceae
Convolvulaceae
Polygalaceae
Dilleniaceae
Rosaceae
Dipterocarpaceae
Sapindaceae
Ebenaceae
Sapotaceae
Elaeocarpaceae
Sterculiaceae
Euphorbiacea
Theaceae
66
70
65
71
63
46
47
41 45
48
43
51
50
49
53
42
29
19
Lecythidaceae
59
40
31
Lauraceae
Araucariaceae
187
73
30
44
0
0
54
0
20
40
60
Fagaceae
Flacourtiaceae
68
16
6
FAMILY
Annonaceae
165
75
38
Fabaceae
Anacardiaceae
174
76
CNV05
166
164
33
13
25
167
169
84
86
89 88
14
12
161
83
98
100 99
172
136
79 78 77
103
106
102
105
170
160
155
129 131
118
159
141
40
130
140
135
20
137
80
100
42 • A Field Guide to the Permanent Sample Plots
Figure 9. Distribution of Trees in Plot CNV05
A Field Guide to the Permanent Sample Plots • 43
Table 5. List of Trees in Plot CNV05
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea sp.
Swintonia glauca Engl.
Dacryodes rugosa H.J.Lam
Shorea macroptera Dyer
Baccaurea parvilora (M.A.) M.A.
Pouteria malaccensis (Clarke) Baeh.
Myristica beccarii Warb.
Shorea pinanga Scheff.
Agathis borneensis Warb.
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Swintonia glauca Engl.
Pouteria malaccensis (Clarke) Baeh.
Agathis borneensis Warb.
Shorea beccariana Burck.
Lithocarpus sp.
Memecylon myrsinoides Blume
Dead
Shorea ochracea Sym.
Garcinia dioica Bl.
Shorea pinanga Scheff.
Shorea venulosa Wood ex Meijer
Celtis philippensis Blanco
Agathis borneensis Warb.
Dead
Shorea maxwelliana King
Dead
Palaqium calophyllum (T.& B.) Pierre
Garcinia dioica Bl.
Shorea pinanga Scheff.
Scaphium macropodum (Miq.) Beumée ex Heyne
Castanopsis sp.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Memecylon myrsinoides Blume
Dead
Scaphium macropodum (Miq.) Beumee ex Heyne
Dead
44 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Artocarpus anisophyllus Miq.
Shorea parvifolia Dyer
Neesia altissima (Bl.) Bl.
Lophopetalum subobovatum King
Diospyros oblonga Wall ex G. Don
Pternandra rostrata (Cogn.) Ohwi
Licania splendens (Korth.) Prance
Licania splendens (Korth.) Prance
Pouteria malaccensis (Clarke) Baeh.
Sarcotheca diversifolia (Miq.) Hall.f.
Dillenia grandifolia Wall. Ex Hk.f.
Erycibe impressa Hoogl.
Shorea venulosa Wood ex Meijer
Myristica beccarii Warb.
Shorea parvifolia Dyer
Tetramerista glabra Miq.
Dacryodes incurvata (Engler) H.J. Lam
Shorea venulosa Wood ex Meijer
Agathis borneensis Warb.
Hydnocarpus kunstleri (King) Warb.
Swintonia glauca Engl.
Swintonia glauca Engl.
Shorea sp.
Pternandra rostrata (Cogn.) Ohwi
Myristica beccarii Warb.
Castanopsis sp.
Myristica beccarii Warb.
Baccaurea edulis Merr.
Baringtonia sp.
Dead
Hydnocarpus kunstleri (King) Warb.
Dead
Hydnocarpus kunstleri (King) Warb.
Scaphium macropodum (Miq.) Beumée ex Heyne
Dead
Calophyllum venulosum Miq.
Unidentiied
A Field Guide to the Permanent Sample Plots • 45
Table 5. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Syzygium chloranthum (Duth.) Merr. & Perry
Shorea sp.
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus kunstleri (King) Warb.
Shorea pinanga Scheff.
Dipterocarpus lowii Hk.f.
Gymnacranthera paniculata Warb.
Gluta wallichii (Hk.f.) Ding Hou
Swintonia glauca Engl.
Shorea sp.
Agathis borneensis Warb.
Shorea maxwelliana King
Koompassia malaccensis Maing. ex Benth.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Syzygium clavilorum (Roxb.) Wall. ex Cowan
Dipterocapus lowii Hk.f.
Dead
Shorea macroptera Dyer
Dead
Koompassia malaccensis Maing. ex Benth.
Garcinia dioica Bl.
Palaquium rostratum (Miq.) Burk.
Shorea seminis (de Vriese) Slooten
Dead
Shorea pinanga Scheff.
Syzygium bankense (Hassk.) Merr. & Perry
Santiria laevigata Bl. f. laevigata
Koompassia malaccensis Maing. ex Benth.
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Dead
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Agathis borneensis Warb.
Mezzetia parvifolia Becc.
Palaquium cochlarilorum V. Royen
46 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Gymnacranthera paniculata Warb.
Dead
Syzygium bankense (Hassk.) Merr. & Perry
Myristica beccarii Warb.
Shorea pinanga Scheff.
Pternandra rostrata (Cogn.) Ohwi
Shorea rubra Ashton
Horsieldia sp.
Vatica oblongifolia Hook.f.
Xylopia malayana Hook.f. ex Th.
Shorea beccariana Scheff.
Shorea macroptera Dyer
Palaqium calophyllum (T.& B.) Pierre
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Garcinia dioica Bl.
Shorea rubra Ashton
Nephelium uncinatum Leenh
Shorea macroptera Dyer
Shorea rubra Ashton
Shorea rubra Ashton
Dead
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea rubra Ashton
Syzygium sp.
Shorea exelliptica Meijer
Artocarpus anysophyllus Miq.
Shorea maxwelliana King
Shorea parvifolia Dyer
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Diospyros sp.
Syzygium sp.
Dead
Agathis borneensis Warb.
A Field Guide to the Permanent Sample Plots • 47
Table 5. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea beccariana Burk.
Syzygium sp.
Pouteria malaccensis (Clarke) Baeh.
Xanthophyllum sp.
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus woodii Merr.
Gymnacranthera paniculata Warb.
Garcinia nervosa Miq.
Hydnocarpus woodii Merr.
Horsieldia sp.
Shorea rubra Ashton
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Aporusa antennifera (Airy Shaw) Airy Shaw
Nephelium uncinatum Leenh.
Shorea parvifolia Dyer
Quercus sp.
Shorea venulosa Wood ex Meijer
Dipterocapus lowii Hk.f.
Shorea rubra Ashton
Dead
Artocarpus sp.
Shorea rubra Ashton
Agathis borneensis Warb.
Diospyros sp.
Santiria grifithii (Hk.f.) Engl.
Madhuca malaccensis (Clarke) H.J.Lam.
Adinandra borneensis Kob.
Shorea rubra Ashton
Shorea maxwelliana King
Adinandra borneensis Kob.
Sindora sp.
Mangifera swintonioides Kosterm
Dead
Shorea macroptera Dyer.
Hydocarpus sp.
Madhuca malaccensis (Clarke) H.J.Lam.
Lithocarpus ewyckii Rehd.
48 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Pouteria malaccensis (Clarke) Baeh.
Shorea maxwelliana King
Shorea rubra Ashton
Koompassia malaccensis Maing. ex Benth.
Shorea rubra Ashton
Calophyllum pulcherrimum Wall. ex Planch. & Trian
Memecylon myrsinoides Blume
Shorea venulosa Wood ex Meijer
Madhuca malaccensis (Clarke) H.J.Lam.
Lithocarpus ewyckii Rehd.
Lithocarpus ewyckii Rehd.
Artocarpus integer (Thunb.) Merr.
Elaeocarpus petiolatus (Thunb)
Dillenia grandifolia Wall.
Madhuca malaccensis (Clarke) H.J.Lam.
Garcinia dioica Bl.
Myristicaceae
Myristicaceae
Shorea rubra Ashton
Shorea venulosa Wood ex Meijer
Pouteria malaccensis (Clarke) Baeh.
Shorea rubra Ashton
Beilschmeidia sp.
Dipterocarpus lowii Hk.f.
Shorea beccariana Burk.
Garcinia sp.
Dead
Atuna excelsa (Benth.) Kosterm.
Syzygium bankense (Hassk.) Merr & Perry
Dead
Dead
Agathis borneensis Warb.
Dead
Agathis borneensis Warb.
Garcinia sp.
Anisoptera costata Korth.
Shorea venulosa Wood ex Meijer
A Field Guide to the Permanent Sample Plots • 49
Table 5. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Madhuca malaccensis (Clarke) H.J.Lam
Ternstroemia aneura Miq.
Syzygium sp.
Shorea maxwelliana King
Madhuca malaccensis (Clarke) H.J.Lam
Ternstroemia aneura Miq.
Santiria grifithii (Hk.f.) Engl.
Shorea venulosa Wood ex Meijer
Baccaurea sumatrana M.A.
Swintonia glauca Engl.
Baccaurea sumatrana M.A.
Horsieldia sp.
Syzygium jambosoides (Laut.) Merr. & Perry
Shorea exelliptica Meijer
Pouteria malaccensis (Clarke) Baeh.
Garcinia nervosa Miq.
Shorea maxwelliana King
Shorea maxwelliana King
Horsieldia sp.
Shorea rubra Ashton
Semecarpus longifolia Bl.
Shorea maxwelliana King
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea maxwelliana King
Dead
Calophyllum lowii Hook.f.
Shorea beccariana Burk.
Gluta wallichii (Hk.f.) Ding Hou
Durio carinatus Mast.
Lithocarpus ewyckii Rehd.
Madhuca malaccensis (Clarke) H.J.Lam
Syzygium splendens (Bl.) Merr. & Perry
Gluta wallichii (Hk.f.) Ding Hou
Ixonanthes petiolaris Bl.
Dipterocarpus lowii Hk.f.
Lithocarpus ewyckii Rehd.
Shorea venulosa Wood ex Meijer
50 • A Field Guide to the Permanent Sample Plots
Table 5. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea macroptera Dyer
Knema laurina (Bl.) Warb.
Adinandra borneensis Kob.
Shorea macroptera Dyer
Gluta wallichii (Hk.f.) Ding Hou
Helicia petiolaris Benn.
Syzygium jambosoides (Laut.) Merr. & Perry
Litsea irma Bl.
Gymnacranthera paniculata Warb.
Madhuca malaccensis (Clarke) H.J.Lam
Unidentiied
Shorea rubra Ashton
Shorea maxwelliana King
Madhuca malaccensis (Clarke) H.J.Lam.
Shorea maxwelliana King
Calophyllum lowii Hook.f.
Madhuca malaccensis (Clarke) H.J.Lam.
Calophyllum lowii Hook.f.
Shorea venulosa Wood ex Meijer
Dead
Xanthophyllum sp.
Sindora coriacea Maing. ex Prain
Melanochyla beccariana Oliv.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Swintonia glauca Engl.
Dead
Shorea rubra Ashton
Eugeissonia utilis Becc.
Gluta wallichii (Hk.f.) Ding Hou
Anisoptera sp.
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea parvifolia Dyer
Shorea rubra Ashton
Agathis borneensis Warb.
Eugeissonia tristis Griff.
A Field Guide to the Permanent Sample Plots • 51
Table 5. Continued
Tree #
297
298
299
300
301
302
303
304
305
306
Species
Koompassia malaccensis Maing. ex Benth.
Castanopsis sp.
Gymnacranthera paniculata Warb.
Tristaniopsis whiteana (Griff.) Wils. & Waterh.
Dead
Adinandra borneensis Kob.
Tetramerista glabra Miq.
Shorea pinanga Scheff.
Macaranga
Castanopsis
20
40
60
80
74
205
136
100
111
100
100
0
106
15
73
162
71
179
27
23
69
34
37
76
125
22 66 213
80
191
4
126
159
178
133
97
180
60
154
171
115
10
224
177
114
13
104
29
112
19
96
113
157
25
57
32
102
172
7
202
9
187
35
158
33
132
105
217
143
109
67
203
99
60
68
80
220
94
230
155
231
129
40
103
107
40
95
17
186
55
18
50 182
20
195
127
1
108
168
169
20
39
225
30 174
226
119
48
101
193
40
85
38
163
208
89
209
185
47
45
210
41
164
49
222
0
0
43
0
20
40
Gnetaceae
FAMILY
Lauraceae
Anacardiaceae
Melastomataceae
Annonaceae
Meliaceae
Apocynaceae
Myristicaceae
Arecaceae
Myrtaceae
Bombacaceae
Oxalidaceae
Burseraceae
Podocarpaceae
Celastraceae
Rhizophoraceae
Clusiaceae
Rosaceae
Cornaceae
Rubiaceae
Dipterocarpaceae
Sapindaceae
Ebenaceae
Sapotaceae
Elaeocarpaceae
Simarubaceae
Euphorbiacea
Sterculiaceae
Fabaceae
Theaceae
Fagaceae
Ulmaceae
Flacourtiaceae
Violaceae
184
11
20
56
CNV06
60
80
100
52 • A Field Guide to the Permanent Sample Plots
Figure 10. Distribution of Trees in Plot CNV06
A Field Guide to the Permanent Sample Plots • 53
Table 6. List of Trees in Plot CNV06
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Koompassia malaccensis Maing. ex. Benth
Dead
Dipterocarpus lowii Hk.f.
Baccaurea pendula Merr.
Cephalomappa lepidota Airy Shaw
Microcos sp.
Shorea parvifolia Dyer
Dipterocarpus lowii Hk.f.
Knema cinerea (Poir.) Warb.
Licania splendens (Korth.) Prance
Syzygium sp.
Dead
Shorea parvifolia Dyer
Dead
Shorea sp.
Dipterocarpus lowii Hk.f.
Litsea castanea Hook.f.
Vatica rasak (Korth.) Bl.
Vatica rasak (Korth.) Bl.
Dehaasia irma Bl.
Vatica rasak (Korth.) Bl.
Palaquium gutta (Hook.f.) Baill.
Tetramerista glabra Miq.
Neoscortechinia kingii (Hk.f.) Pax & Hoffm.
Tetramerista glabra Miq.
Neoscortechinia kingii (Hk.f.) Pax & Hoffm.
Shorea ovalis (Korth.) Bl.
Palaquium obtusifolium Burck.
Vatica rasak (Korth.) Bl.
Vatica rasak (Korth.) Bl.
Amoora lanceolata Hiem.
Syzygium sp.
Lophopetalum subobovatum King
Gymnacranthera paniculata Warb.
Myristica sp.
Dacryodes incurvata (Engl.) H.J.Lam.
Elaeocarpus polystachys Wall.
54 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Dehaasia irma Bl.
Koompassia malaccensis Maing. ex Benth.
Alseodaphne havilandii (Gamble) Kosterm
Scaphium macropodum (Miq.) Beumée ex Heyne
Unidentiied
Gironiera nervosa Planch.
Lophopetalum subobovatum King
Dehaasia irma Bl.
Dead
Carallia brachyata (Lour.) Merr.
Shorea maxwelliana King
Memecylon myrsinoides Blume
Oncosperma horridum Scheff.
Shorea sp.
Unidentiied
Artocarpus anisophyllus Miq.
Gymnacranthera forbesii (King) Warb
Mangifera magniica Kochumen
Knema furfuracea (Hk.f. ex Th.) Warb.
Knema furfuracea (Hk.f. ex Th.) Warb.
Durio lanceolatus Mast.
Carallia brachyata (Lour.) Merr.
Dehaasia irma Kosterm.
Eugeissona utilis Becc
Lithocarpus sp.
Xylopia altissima Boerl.
Shorea parvifolia Dyer
Parastemon urophyllus DC.
Carallia brachyata (Lour) Merr.
Myristica sp.
Sarcotheca diversifolia (Miq.) Hall.f.
Calophyllum lowii Hook.f
Alstonia angustiloba Miq.
Aporusa falcifera Hk.f.
Shorea parvifolia Dyer.
Vatica rasak (Korth.) Bl.
Vatica rasak (Korth.) Bl.
A Field Guide to the Permanent Sample Plots • 55
Table 6. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Pternandra caerulescens Jack.
Syzygium sp.
Palaquium sumatranum Burck
Dipterocarpus lowii Hk.f.
Lophopetalum subobovatum King
Palaquium sumatranum Burck
Knema furfuracea (Hk.f.ex Th.) Warb.
Diospyros sp.
Polyosma sp.
Garcinia sp.
Eugenia chlorantum (Duth.) Merr. & Perry
Lithocarpus blumeanus (Korth.) Rehd.
Sarcotheca diversifolia (Miq.) Hall.f.
Artocarpus kemando Miq.
Tetramerista glabra Miq.
Tetramerista glabra Miq.
Unidentiied
Dehaasia irma Bl.
Myristica maxima Warb.
Sindora sp.
Prunus spicata Kalkm.
Polyalthia sumatrana King
Calophyllum venulosum Zoll.
Dead
Licania splendens (Korth.) Prance
Madhuca malaccensis Lam
Baccaurea delexa M.A.
Baccaurea angulata Merr.
Gnetum cuspidatum Bl.
Polyalthia rumphii Merr.
Eugenia fastigiata Miq.
Shorea seminis (de Vriese) Slooten
Myristica villosa Warb.
Baccaurea angulata Merr.
Eugeissona utilis Becc.
Baccaurea angulata Merr.
Madhuca sericea Lam
56 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Koompassia malaccensis Maing. ex Benth.
Sageraea elliptica (DC.) Hk.f. ex Th.
Myristica villosa Warb.
Mastixia trichotoma Bl.
Diospyros oblonga Wall. ex G.Don
Shorea ovalis (Korth.) Bl.
Myristica villosa Warb.
Baccaurea kunstleri King
Litsea laevigata Gamble
Dead
Unidentiied
Unidentiied
Dipterocarpus lowii Hk.f.
Dacryodes incurvata (Engler) H.J.Lam.
Hydnocarpus woodii Merr. ex Slooten
Licania splendens (Korth.) Prance
Cratoxylum formosum Benth. & Hook. f. ex Dyer
Myristica villosa Warb.
Artocarpus anisophyllus Miq.
Memecylon myrsinoides Blume
Shorea
Hydnocarpus woodii Merr. ex Slooten
Dead
Koompassia malaccensis Maing. ex Benth.
Dyera costulata Hook.f.
Shorea paucilora King
Helicia petiolaris Benn.
Baccaurea pendulla Merr.
Melanochyla beccariana Oliv.
Dead
Rhodamnia cinerea Jack.
Artocarpus anisophyllus Miq.
Dipterocarpus lowii Hook.f.
Buchanania arborescens (Bl.) Bl.
Teysmaniodendron simplicifolium Merr.
Dead
Sarcotheca diversifolia (Miq.) Hall.f.
A Field Guide to the Permanent Sample Plots • 57
Table 6. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Syzygium chloranthum (Duth.) Merr. & Perry
Shorea parvifolia Dyer
Palaquium obtusifolium Burck.
Vatica rasak (Korth.) Bl.
Ixonanthes petiolaris Bl.
Adinandra borneensis Kob.
Vatica rasak (Korth.) Bl.
Tetramerista glabra Miq.
Melanochyla beccariana Oliv.
Koona ochracea (Elm) Merr.
Parishia maingayi Hook.f
Pternandra rostrata (Cogn.) Ohwi
Dead
Mastixia trichotoma Bl.
Durio dulcis Becc.
Lithocarpus sp.
Shorea sp.
Diospyros sp.
Dead
Durio lanceolatus Mast.
Vatica rasak (Korth.) Bl.
Dead
Myristica sp.
Palaquium rostratum (Miq.) Burck
Pternandra rostrata (Cogn.) Ohwi
Calophyllum lowii Hook.f.
Dead
Hopea sp.
Nephelium uncinatum Leenh.
Pternandra rostrata (Cogn.) Ohwi
Sarcotheca diversifolia (Miq.) Hall.f.
Myristica sp.
Tetramerista glabra Miq.
Palaquium rostratum (Miq.) Burck
Macaranga hypoleuca M.A.
Archidendron sp.
Palaquium sumatranum Burck
58 • A Field Guide to the Permanent Sample Plots
Table 6. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Tetramerista glabra Miq.
Shorea parvifolia Dyer
Pternandra rostrata (Cogn.) Ohwi
Knema furfuracea (Hk.f.et Th.) Warb.
Tetramerista glabra Miq.
Palaquium rostratum (Miq.) Burck
Palaquium rostratum (Miq.) Burck
Podocarpus neriifolius D.Don.
Shorea seminis (de Vriese) Slooten
Aglaia cf. gamopetala Merr.
Celtis philippensis Blanco
Shorea parvifolia Dyer
Pternandra rostrata (Cogn.) Ohwi
Dead
Oncosperma horridum Scheff.
Adinandra borneensis Kob.
Shorea seminis (de Vriese) Slooten
Tetramerista glabra Miq.
Neesia malayana Bakh.
Gardenia tubifera Wall.
Syzygium ochneocarpa Merr.
Shorea seminis (de Vriese) Slooten
Irvingia malayana Oliv.
Shorea seminis (de Vriese) Slooten
Alstonia spathulata Blume
Canarium littorale Bl.
Lithocarpus blumeanus (Korth.) Rehd.
Palaquium rostratum (Miq.) Burck
Licania splendens (Korth.) Prance
Shorea johorensis Foxw.
Dead
Dehaasia irma Bl.
Calophyllum lowii Hook.f.
Vatica sp.
Archidendron sp.
Lophopetalum subobovatum King
Vatica sp.
A Field Guide to the Permanent Sample Plots • 59
Table 6. Continued
Tree #
223
224
225
226
227
228
229
230
231
Species
Dillenia grandifolia Wall. ex Hk.f.
Tetramerista glabra Miq.
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea sp.
Shorea sp.
Gymnacranthera contracta Warb.
Artocarpus kemando Miq.
Castanopsis motleyana King.
Lophopetalum subobovatum King
20
40
60
80
100
189
176
200 201
194
190
168
174
191
183
175
195
184
199
192
80
153
148
166
160
135
144
154 155
165
208
206
133
145 143
146
152
164
80
173
169
204
207
203
172
167
209
202
181
177
100
100
0
134
137 136
128
127
138
156
162
157
151
139
150
161
60
158
141
101
103
100
99
96
113
98
97
115 116
40
66
79
74
78
70
77
75
82
62
71
49
60
42
68
76
84
64
69
73
83
65
72
6
67
12
7
20
9
32
23
13
35
19
16
3
122
0
20
36 37 38
30
17
15
0
Myristicaceae
FAMILY
Myrtaceae
Alangiaceae
Olacaceae
Anacardiaceae
Oleaceae
Annonaceae
Polygalaceae
Apocynaceae
Proteaceae
Arecaceae
Rhizophoraceae
Bombacaceae
Rosaceae
Burseraceae
Rubiaceae
Clusiaceae
Sapindaceae
Dipterocarpaceae
Sapotaceae
Ebenaceae
Sterculiaceae
Euphorbiacea
Theaceae
Fabaceae
Thymelaeaceae
Fagaceae
Tiliaceae
Lauraceae
Verbenaceae
29
4
2
41
18
14
1
CNV07
46
59
21
11
8
22
20
81
40
60
80
Meliaceae
39
0
40
86
80
20
118
112
88
5
120
121
119
110
90
89 95
85
111
109
102
87
108
107
93 94
130
131
142
60
159
91
100
60 • A Field Guide to the Permanent Sample Plots
Figure 11. Distribution of Trees in Plot CNV07
A Field Guide to the Permanent Sample Plots • 61
Table 7. List of Trees in Plot CNV07
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Shorea sp.
Dehaasia irma Bl.
Shorea sp.
Shorea sp.
Beilschmiedia glabra Kosterm
Shorea fallax Meijer
Shorea fallax Meijer
Aglaia harmsiana Park.
Baccaurea edulis Merr.
Dead
Durio grandilorus Kost.
Polyalthia glauca Boerl.
Garcinia maingayi Hook.f.
Spatholobus sp.
Shorea fallax Meijer
Knema furfuracea (Hk.f. ex. Th.) Warb.
Plectronia sp.
Gonystylus keithii Airy Shaw
Elateriospermum tapos Bl.
Parashorea smithiesii Wyatt-Smith ex Ashton
Tabernaemontana sphaerocarpa Bl.
Shorea ovalis (Korth.) Bl.
Artocarpus lanceifolius Roxb.
Beilschmiedia glabra Kosterm.
Polyalthia glauca Boerl.
Dead
Dead
Elateriospermum tapos Bl.
Prunus arborea (Bl.) Kalkm.
Santiria sp.
Mastixia rostrata Bl.
Ochanostachys amentacea Mast.
Aglaia ganggo Miq.
Shorea parvifolia Dyer
Baccaurea pendula Merr.
Polyalthia glauca Boerl.
Teijsmaniodendron simplicifolium Merr.
62 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Parashorea smithiesii Wyatt-Smith ex Ashton
Plectronia sp.
Litsea angulata Bl.
Pentace laxilora Merr.
Chionanthus sp.
Dead
Teijsmaniodendron simplicifolium Merr.
Shorea ochracea Sym
Carallia brachyata (Lour.) Merr.
Shorea seminis (de Vriese) Slooten
Chisocheton divergens Bl.
Heritiera borneensis (Merr.) Kosterm.
Heritiera borneensis (Merr.) Kosterm.
Teijsmaniodendron simplicifolium Merr.
Aquilaria malaccensis Lamk.
Shorea ovalis (Korth.) Bl.
Aporusa falcifera Hk.f.
Syzygium sp.
Palaquium calophyllum Pierre ex Burck
Payena acuminta (Bl.) Pierre
Shorea ovalis (Korth.) Bl.
Shorea ovalis (Korth.) Bl.
Helicia serrata (R.Br.) Bl.
Shorea fallax Meijer
Palaquium dasyphyllum Pierre
Dead
Shorea ovalis (Korth.) Bl.
Baccaurea edulis Merr.
Nephelium uncinatum Leenh
Baccaurea edulis Merr.
Shorea fallax Meijer
Dysoxylum sp.
Shorea fallax Meijer
Polyalthia sumatrana King
Artocarpus lanceifolius Roxb.
Mezzetia parvifolia Becc.
Artocarpus lanceifolius Roxb.
A Field Guide to the Permanent Sample Plots • 63
Table 7. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Shorea ovalis (Korth.) Bl.
Dacryodes rugosa H.J.Lam
Knema laurina (Bl.) Warb.
Knema furfuracea (Hk.f. ex. Th.) Warb.
Knema sp.
Diospyros sp.
Macaranga pruinosa (Miq.) Muell Arg.
Amoora lanceolata Hiern
Shorea fallax Meijer
Diospyros sp.
Shorea ovalis (Korth.) Bl.
Artocarpus lanceifolius Roxb.
Elateriospermum tapos Bl.
Shorea ovalis (Korth.) Bl.
Shorea ovalis (Korth.) Bl.
Shorea fallax Meijer
Knema furfuracea (Hk.f. ex. Th.) Warb.
Knema laurina (Bl.) Warb.
Syzygium splendens (Bl.) Merr. & Perry
Palaquium rostratum Burck
Heritiera javanica (Bl.) Kosterm.
Shorea ovalis (Korth.) Bl.
Parashorea smithiesii Wyatt-Smith ex Ashton
Tetramerista glabra Miq.
Gymnacranthera paniculata Warb
Chisocheton divergens Bl.
Xylopia altissima Boerl.
Lithocarpus sundaicus (Bl.) Rehd.
Diospyros borneensis Hiern
Pometia pinnata Forst.
Shorea ovalis (Korth.) Bl.
Livistona sp.
Shorea leprosula Miq.
Madhuca oblongifolia Merr.
Parishia maingayi Hk.f.
Syzygium splendens (Bl.) Merr. & Perry
Shorea parvifolia Dyer
64 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Polyalthia glauca Boerl.
Shorea fallax Meijer
Dead
Shorea ovalis (Korth.) Bl.
Cleistanthus myrianthus (Hassk.) Krz.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Lithocarpus blumeanus (Korth.) Rehd.
Shorea parvifolia Dyer
Nephelium lappaceum Miq.
Kokoona ohracea (Elmer) Merr.
Palaquium sumatranum Burk.
Shorea ovalis (Korth.) Bl.
Xanthophyllum sp.
Livistona sp.
Koodersiodendron pinnatum (Blanco) Merr.
Adinandra sp.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Pometia pinnata Forst.
Ficus glandulifera Wall.
Lithocarpus sp.
Syzygium sp.
Shorea parvifolia Dyer
Xanthophyllum amoenum Chod.
Polyalthia glauca Boerl.
Aporusa lunata (Miq.) Kurz.
Shorea ovalis (Korth.) Bl.
Myristica sp.
Dehaasia irma Bl.
Knema furfuracea (Hk.f. ex Th.) Warb.
Shorea parvifolia Dyer
Dead
Syzygium laxilorum DC.
A Field Guide to the Permanent Sample Plots • 65
Table 7. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Dead
Pterocymbium tubulatum (Mast.) Pierre
Shorea fallax Meijer
Pentace laxilora Merr.
Helicia petiolaris Benn.
Actinodaphne glomerata (Bl.) Nees
Koompassia malaccensis Maing. ex Benth.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Palaquium dasyphyllum Pierre
Palaquium dasyphyllum Pierre
Cleistanthus myrianthus (Hassk.) Krz.
Diospyros borneensis Hiern
Dehaasia irma Bl.
Dead
Shorea ovalis (Korth.) Bl.
Ficus globosa Bl.
Shorea leprosula Miq.
Diospyros borneensis Hiern
Cleistanthus myrianthus (Hassk.) Krz.
Syzygium sp.
Pentace laxilora Merr.
Macaranga hosei King ex Hook.f.
Lithocarpus blumeanus (Korth.) Rehd.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Knema cinerea (Poir.) Warb.
Aporusa falcifera Hk.f.
Shorea parvifolia Dyer
Livistona sp.
Shorea parvifolia Dyer
Livistona sp.
Dead
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Shorea parvifolia Dye r
66 • A Field Guide to the Permanent Sample Plots
Table 7. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
Species
Pternandra rostrata (Cogn.) Ohwi
Dead
Nepheium lapaceum Miq.
Syzygium sp.
Alangium sp.
Madhuca malaccensis (Clarke) H.J.Lam
Dehaasia irma Bl.
Shorea ovalis (Korth.) Bl.
Ostodes macrophylla B. ex H.
Hopea dryobalanoides Miq.
Bhesa paniculata Arn
Dead
Dead
Chisocheton divergens Bl.
Baccaurea kunstleri King.
Shorea ovalis (Korth.) Bl.
Shorea ovalis (Korth.) Bl.
Polyalthiya laterilora King
Quercus sp.
Macaranga pruinosa (Miq.) Muell Arg.
Dipterocarpus humeratus Sloot.
Calophyllum pulcherrimum Wall.
Parashorea smithiesii Wyatt-Smith ex Ashton
Calophyllum venulosum Miq.
244
243
236
237
239
80
238
240
234
233
216
230
232
228
227
224
296
185 184
191
196
193
301
297
180 177 176 175
179
178
182
174
192
183
190
202
203
205
201 200
60
139
137
141
134
164
162
151
145
140
133
125
147
142
146
163
119
154
130
129
131
121
127
143
128
99
40
115 106
118
85
105
110
90 89
20
13
3 4 5
12
7
15
17
19
25
Arecaceae
Myristicaceae
67
Bombacaceae
Myrtaceae
65
Burseraceae
Polygalaceae
Clusiaceae
Proteaceae
Dilleniaceae
Rosaceae
Dipterocarpaceae
Sapindaceae
Ebenaceae
33
41
Sapotaceae
Elaeocarpaceae
Simarubaceae
Euphorbiacea
Sterculiaceae
Fabaceae
Theaceae
61
54
42
27
51
52
40 39
55
56
57
43 44
30
32
46
31
47
48
0
45
59
0
1
69
64 63
35
36 37
24
28
8 9
66
38
26
11
Meliaceae
81
20
18
Melastomataceae
Araucariaceae
68
34
21
Lauraceae
Apocynaceae
71
79
87 86
88
Icacinaceae
70
77
91
14 16
73 72
0
20
40
60
Flacourtiaceae
Annonaceae
160
157
74
97 93 92
94
109
2
96
156
Fagaceae
FAMILY
Anacardiaceae
161
159
76
82
104 98
112
111
75
83
102
103
149
84
100
155
150
148
117
158
144
20
120
40
122
CNV08
80
100
A Field Guide to the Permanent Sample Plots • 67
138
173
172
166
168 167
169 165
171
170
186
188
189
187
194
206
136
123
295
294
275
199
204
207
226
124
274
198
221 223
225
293
197
212
220
289
286
273
272
209
291
292
288
287
269
271
211
215
231
290
268
270
100
299
285
278
258
256
254
251
80
284
281 280
279
267
266
265
259
283
298
264
252
60
282
262
260
247
249
246
250
245
242 241
235
40
263
261
100
20
248
80
0
60
100
Figure 12. Distribution of Trees in Plot CNV08
68 • A Field Guide to the Permanent Sample Plots
Table 8. List of Trees on Plot# CNV08
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Artocarpus anisophyllus Miq.
Dacryodes rugosa H.J.Lam var. rugosa
Dialium trifolum de Wit
Shorea exelliptica Meijer
Knema glauca (Bl.) Warb.
Xanthophyllum rufum Bth.
Cryptocarya sp.
Knema glauca (Bl.) Warb.
Myristica tomentosa Warb.
Dead
Shorea parvifolia Dyer
Durio carinatus Mast.
Polyalthia laterilora Kurz.
Prunus spicata Kalkm.
Shorea paucilora King
Dipterocarpus confertus Sloot.
Cinnamomum sp.
Irvingia malayana Oliv.
Shorea macroptera Dyer
Shorea agami Ashton
Shorea macroptera Dyer
Dacryodes incurvata (Engler) H.J.Lam
Prunus sp.
Knema cinerea (Poir.) Warb.
Shorea paucilora King
Syzygium tawahense (Korth.) Merr. & Perry
Dipterocarpus confertus Sloot.
Cryptocarya densilora Bl.
Shorea agami Ashton
Diospyros sp.
Madhuca malaccensis (Clarke.) H.J.Lam
Quercus sp.
Myristica beccarii Warb.
Moultonianthus leembruggianus (Boerl. & Koord.) Steenis
Drypetes kikir Airy Shaw
Syzygium bankensis (Hassk.) Merr. & Perry
Licania splendens (Korth.) Prance
A Field Guide to the Permanent Sample Plots • 69
Table 8. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Shorea agami Ashton
Horsieldia sp.
Litsea sp.
Horsieldia sp.
Nephelium uncinatum Radlk. ex Leenh.
Endiandra coriacea Merr.
Horsieldia sp.
Shorea beccariana Burck
Mangifera swintonioides Kosterm
Drypetes kikir Airy Shaw
Shorea rubra Ashton
Shorea rubra Ashton
Horsieldia sp.
Shorea rubra Ashton
Shorea exelliptica Meijer
Dead
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Syzygium clavilora Wall.
Shorea exelliptica Meijer
Dead
Durio lanceolatus Mast.
Shorea maxwelliana King
Shorea maxwelliana King
Dead
Agathis borneensis Warb.
Santiria grifithii (Hk.f.) Engl.
Knema sp.
Gymnacranthera paniculata Warb
Knema sp.
Artocarpus anisophyllus Miq.
Shorea maxwelliana King
Mangifera magniica Kochumen
Memecylon myrsinoides Bl.
Shorea beccariana Burck
Pimeleodendron papaveroides J.J.S.
Scaphium macropodum (Miq.) Beumee ex Heyne
70 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Aporusa sp.
Shorea beccariana Burck
Durio lanceolatus Mast.
Unidentiied
Xylopia malayana Hk. f. Thoms.
Archidendron fagifolium (Bl. ex Miq.) Nielsen
Canarium littorale Bl.
Pimeleodendron papaveroides J.J.S.
Shorea sp.
Dipterocarpus humeratus Sloot.
Sindora beccariana Back.
Nephelium uncinatum Radlk. ex Leenh.
Mesua cornata Stevens
Canarium littorale Bl.
Syzygium sp.
Shorea agami Ashton
Shorea beccariana Burck
Knema sp.
Shorea sp.
Shorea macroptera Dyer
Dead
Quercus subsericea A.Camus
Shorea maxwelliana King
Shorea beccariana Burck
Licania splendens (Korth.) Prance
Neesia malayana Bakh.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum pulcherrimum Wall.
Dacryodes rugosa H.J.Lam
Artocarpus odoratissimus Blanco
Pternandra galeata Ridl.
Dead
Hydnocarpus woodi Merr.
Syzygium sp.
Quercus subsericea A.Camus
A Field Guide to the Permanent Sample Plots • 71
Table 8. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Moultonianthus leembruggianus (Boerl. & Koord.) Steenis
Diospyros sp.
Prunus spicata Kalkm.
Oncosperma horridum Scheff.
Dead
Aporusa falcifera Hk.f.
Myristica sp.
Canarium denticulatum H.J.Lam
Shorea exelliptica Meijer
Oncosperma horridum Scheff.
Shorea rubra Ashton
Shorea exelliptica Meijer
Shorea maxwelliana King
Artocarpus integer (Thunb.) Merr.
Dead
Shorea beccariana Burck
Stemonurus grandifolius Becc.
Shorea rubra Ashton
Shorea maxwelliana King
Melanochyla pulvinervis (Bl.) Ding Hou
Dead
Shorea maxwelliana King
Shorea macroptera Dyer
Dead
Agathis borneensis Warb.
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Lithocarpus lucidus (Roxb.) Rehd.
Dialium maingayi Backer
Shorea beccariana Burck
Elaeocarpus petiolatus Wall.
Knema sp.
Shorea maxwelliana King
Shorea parvifolia Dyer
Cinnamomum iners Reinw.
Dehaasia brachybotrys (Merr.) Kosterm.
Dipterocarpus humeratus Sloot.
72 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Dacryodes incurvata (Engler) H.J.Lam
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Dead
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Syzygium sp.
Gluta wallichii (Hk.f.) Ding Hou
Koompassia malaccensis Maing. ex Benth.
Artocarpus integer (Thunb.) Merr.
Dillenia grandifolia Wall. ex Hk.f.
Shorea maxwelliana King
Shorea rubra Ashton
Shorea exelliptica Meijer
Shorea rubra Ashton
Hydnocarpus woodii Merr.
Madhuca sp.
Quercus sp.
Shorea exelliptica Meijer
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Pouteria malaccensis (Clarke) Baeh.
Knema glauca (Bl.) Warb.
Shorea agami Ashton
Shorea seminis (de Vriese) Slooten
Cryptocarya tomentosa Bl.
Prunus spicata Kalkm.
Madhuca sericea H.J. Lam
Shorea agami Ashton
Oncosperma horridum Scheff.
Palaquium obtusifolium Burck.
Dead
Syzygium splendens (Bl.) Merr. & Perry
Tetramerista glabra Miq.
Neesia malayana Bakh.
Pouteria malaccensis (Clarke) Baeh.
A Field Guide to the Permanent Sample Plots • 73
Table 8. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Tetramerista glabra Miq.
Kokoona ochracea Merr.
Shorea paucilora King
Syzygium splendens (Bl.) Merr. & Perry
Santiria tomentosa Bl.
Shorea beccariana Burck
Hydnocarpus sp.
Disoxylum sp.
Lithocarpus lucidus (Roxb.) Rehd.
Mangifera swintonioides Ashton
Lithocarpus ewyckii Rehd.
Gluta wallichii (Hk.f.) Ding Hou
Shorea beccariana Burck
Helicia petiolaris Benn.
Shorea parvifolia Dyer
Shorea beccariana Burck
Gluta wallichii (Hk.f.) Ding Hou
Horsieldia sp.
Shorea rubra Ashton
Shorea maxwelliana King
Shorea agami Ashton
Syzygium bankensis (Hassk,) Merr. & Perry
Dead
Dyera costulata (Miq.) Hk.f.
Dead
Shorea beccariana Burck
Horsieldia sp.
Dead
Dead
Horsieldia sp.
Diospyros everetii Merr.
Unidentiied
Shorea rubra Ashton
Dead
Vatica umbonata (Hook.f.) Burk
Shorea rubra Ashton
Dead
74 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Shorea maxwelliana King
Madhuca malaccensis (Clarke) H.J.Lam
Sindora leiocarpa Backer
Shorea rubra Ashton
Shorea artinervosa Sym.
Shorea artinervosa Sym.
Glochidion
Dead
Shorea rubra Ashton
Shorea venulosa Wood. ex Meijer
Koompassia malaccensis Maing. ex Benth.
Santiria sp.
Shorea artinervosa Sym.
Dacryodes rogosa H.J.Lam
Horsieldia sp.
Hydnocarpus woodii Merr.
Shorea maxwelliana King
Xanthophyllum
Artocarpus dadah Miq.
Shorea maxwelliana King
Vatica umbonata (Hook.f.) Burk.
Artocarpus anysophyllus Miq.
Semecarpus longifolia Bl.
Durio lanceolatus Mast.
Shorea exelliptica Meijer
Canarium litorale Bl.
Shorea beccariana Burck
Shorea exelliptica Meijer
Memecylon myrsinoides Bl.
Shorea maxwelliana King
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Dead
Shorea maxwelliana King
Dead
Shorea parvifolia Dyer
Calophyllum pulcherrimum Wall.
A Field Guide to the Permanent Sample Plots • 75
Table 8. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Horsieldia sp.
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Prunus sp.
Gluta wallichii (Hk.f.) Ding Hou
Mezzetia parvilora Becc.
Lithocarpus sp.
Lithocarpus lucidus (Roxb.) Rehd.
Syzygium clavilora Wall.
Santiria oblongifolia Bl.
Melanochyla beccariana Oliv.
Shorea macroptera Dyer
Myristica sp.
Lithocarpus sp.
Shorea rubra Ashton
Licania splendens (Korth.) Prance
Dead
Dead
Hydnocarpus woodii Merr.
Shorea beccariana Burck
Hydnocarpus woodii Merr.
Lithocarpus sp.
Ternstroemia aneura Miq.
Mezzetia parvilora Becc.
Tetramerista glabra Miq
Knema cinerea (Poir.) Warb.
Unidentiied
Prunus spicata Kalkm.
Shorea parvifolia Dyer
Knema sp.
Palaquium calophyllum Pierre ex Burck
Tetramerista glabra Miq.
Shorea macroptera Dyer
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Shorea macroptera Dyer
Dipterocarpus lowii Hook.f.
76 • A Field Guide to the Permanent Sample Plots
Table 8. Continued
Tree #
297
298
299
300
301
302
Species
Lithocarpus sp.
Tetramerista glabra Miq.
Nephelium maingayi Hiern.
Tetramerista glabra Miq.
Shorea macroptera Dyer
Unidentiied
40
264
60
276
275
265
282
271
303
307 310
311
309
294
295
257
292
280
237 236
80
256
291
290
279
258
306 312
302
274
273
260
304
296 297
284 285
270
262
100
305
283
269
80
289
238
286
222 221
308
293
205
191
192
234
189
247
239
203
255
207
212
211
158
139
146
144
122
177
152
126 128
129 125
170
127
40
63
91
84
94
79
80
77
70
69
68
75
66
67
37
57
54
26
5
38
25
11
35 34
4
3
71
85
87
100 99 98
7
59
39
40
33
16
41
24
2
23
22
20
46
45
32
31
42
0
13
30
40
49
48
60
53 52
47
56
CNV09
Flacourtiaceae
FAMILY
Lauraceae
Anacardiaceae
Linaceae
Annonaceae
Meliaceae
Apocynaceae
Myristicaceae
Araucariaceae
Myrtaceae
Arecaceae
Polygalaceae
Bombacaceae
Proteaceae
Burseraceae
Rosaceae
Clusiaceae
Rubiaceae
Dilleniaceae
Sapotaceae
Dipterocarpaceae
Sterculiaceae
Euphorbiacea
Theaceae
Fabaceae
Thymelaeaceae
Fagaceae
Ulmaceae
51
44
50
43
0
6
62
60
76
104
171
180
78
107
105
179
73
93
95
20
90
92
108
0
178
172
147
149
109
1
181
140
123
106
190
187
184
60
137
113
210
214
135
202
201 197 199
196
200
198
215 208
227
241
242
204
188
193
80
100
A Field Guide to the Permanent Sample Plots • 77
112
226
195
20
60
246
251
111
110
114
100
20
266
80
0
40
100
Figure 13. Distribution of Trees in Plot CNV09
78 • A Field Guide to the Permanent Sample Plots
Table 9. List of Trees in Plot CNV09
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Shorea rubra Ashton
Dipterocarpus stellatus Vesque
Xanthophyllum parvum Chod.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Dehaasia sp.
Dead
Helicia petiolaris Benn.
Agathis borneensis Warb.
Shorea rubra Ashton
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Shorea rubra Ashton
Syzygium bankense (Hassk.) Merr.& Perry
Syzygium bankense (Hassk.) Merr.& Perry
Gluta wallichii (Hk.f.) Ding Hou
Syzygium bankense (Hassk.) Merr.& Perry
Agathis borneensis Warb.
Shorea exelliptica Meijer
Palaquium rostratum (Miq.) Burck.
Shorea beccariana Burck.
Vatica umbonata (Hook.f.) Burck.
Shorea maxwelliana King
Baccaurea racemosa M.A.
Syzygium bankense (Hassk.) Merr.& Perry
Dipterocarpus stellatus Vesque
Shorea beccariana Burck.
Shorea exelliptica Meijer
Shorea parvifolia Dyer
Palaquium callophyllum (T.& B.) Pierre
Shorea venulosa Wood. ex Meijer
Syzygium sp.
Cleistanthus sumatranus M.A.
Koompassia malaccensis Maing. ex Benth
A Field Guide to the Permanent Sample Plots • 79
Table 9. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Syzygium sp.
Shorea venulosa Wood. ex Meijer
Gonystylus forbesii Gilg.
Koompassia malaccensis Maing. ex Benth
Shorea rubra Ashton
Shorea maxwelliana King
Madhuca erythrophylla (K.ex G.) H.J.Lam
Gluta wallichii (Hk.f.) Ding Hou
Durio lanceolatus Mast.
Gymnacranthera paniculata Warb.
Knema intermedia (Bl.) Warb.
Shorea parvifolia Dyer
Dacryodes incurvata (Engler) H.J.Lam
Shorea macroptera Dyer
Agathis borneensis Warb.
Shorea beccariana Burck.
Dipterocarpus stellatus Vesque
Horsieldia sp.
Shorea parvifolia Dyer
Hydnocarpus kunstleri (King) Warb.
Atuna excelca (Benth.) Kosterm
Dehaasia polyneura (Miq.) Kosterm
Dillenia grandifolia Wall. ex. Hk.f.
Santiria laevigata Bl.
Dacryodes rugosa (Bl.) H.J.Lam
Dipterocarpus stellatus Vesque
Madhuca malacensis (Clarke) H.J.Lam
Syzygium bankense (Hassk.) Merr.& Perry
Spatholobus sp.
Gymnacranthera paniculata Warb.
Artocarpus integer (Thumb.) Merr.
Xanthophyllum amoenum Chod.
Koompassia malaccensis Maing. ex Benth
Garcinia parvifolia Miq.
Horsieldia sp.
Shorea macroptera Dyer
Cleistanthes myrianthus Krz.
80 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Horsieldia glabra (Bl.) Warb,
Shorea beccariana Burck.
Palaquium rostratum (Miq.) Burck
Shorea venulosa Wood ex Meijer
Syzygium sp.
Agathis borneensis Warb.
Dead
Gonystylus forbesii Gilg.
Calophyllum pulcherrimum Wall.
Gluta wallichii (Hk.f.) Ding Hou
Cleistanthus myrianthus Krz.
Agathis borneensis Warb.
Dipterocarpus stellatus Vesque
Agathis borneensis Warb.
Horsieldia sp.
Shorea parvifolia Dyer
Vatica umbonata (Hook.f.) Burck.
Sindora leiocarpa Backer
Vatica umbonata (Hook.f.) Burck.
Shorea parvifolia Dyer
Shorea macroptera Dyer
Dead
Agathis borneensis Warb.
Vatica umbonata (Hook.f.) Burck.
Hydnocarpus woodii Merr.
Dipterocarpus stellatus Vesque
Xanthophyllum
Dipterocarpus stellatus Vesque
Unidentiied
Horsieldia glabra (Bl.) Warb.
Gymnacranthera paniculata Warb.
Neonauclea excelsa (Bl.) Merr.
Shorea maxwelliana King
Madhuca oblongifolia Merr.
Dipterocarpus lowii Hk.f.
Shorea macroptera Dyer
Helicia petiolaris Benn.
A Field Guide to the Permanent Sample Plots • 81
Table 9. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Syzygium bankense (Hassk.) Merr. & Perry
Santiria grifithii (Hk.f.) Engl.
Vatica umbonata (Hook.f.) Burck.
Barringtonia scortechinii King
Mangifera magniica Kochumen
Shorea maxwelliana King
Horsieldia sp.
Dipterocarpus stellatus Vesque
Podocarpus neriifolius D.Don
Agathis borneensis Warb.
Scaphium macropodum (Miq.) Beumee ex Heyne
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus urceolaris (Jack) Merr.
Parastemon urophyllus A.DC.
Licania splendens (Korth.) Prance
Shorea beccariana Burck.
Shorea parvifolia Dyer
Parastemon urophyllus A.DC.
Palaquium cochlearifolium van Royen
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Shorea maxwelliana King
Swintonia glauca Engl.
Agathis borneensis Warb.
Shorea maxwelliana King
Scaphium macropodum (Miq.) Beumee ex Heyne
Santiria grifithii (Hk.f.) Engl.
Hydnocarpus kunstleri (King) Warb
Vatica umbonata (Hook.f.) Burck.
Agathis borneensis Warb.
Madhuca erythrophylla (K.ex G.) H.J.Lam
Calophyllum pulcherrimum Wall.
Syzygium bankense (Hassk.) Merr. & Perry
Mangifera magniica Kochumen
Shorea maxwelliana King
Shorea rubra Ashton
82 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea venulosa Wood ex Meijer
Dead
Memecylon oleaefolium Bl.
Koompassia malaccensis Maing. ex Benth
Horsieldia sp.
Melanochyla beccariana Oliv.
Aglaia grifithii Kurz.
Dead
Hydnocarpus kunstleri (King) Warb
Melanochyla beccariana Oliv.
Carallia brachyata (Lour.) Merr.
Sindora leiocarpa Backer.
Syzygium tawahense (Korth.) Merr. & Perry
Podocarpus neriifolius D.Don
Shorea agami Ashton
Syzygium tawahense (Korth.) Merr. & Perry
Shorea beccariana Burck.
Dead
Dacryodes rugosa (Bl.) H.J.Lam
Baccaurea kunstleri King
Palaquium cochlearifolium van Royen
Durio carinatus Mast.
Dillenia grandifolia Wall. ex Hk.f.
Litsea nidularis Gamble
Shorea beccariana Burck.
Syzygium tawahense (Korth.) Merr. & Perry
Artocarpus integer (Thumb.) Merr.
Dead
Shorea rubra Ashton
Santiria grifithii (Hk.f.) Engl.
Castanopsis ?argentea A.DC
Dipterocarpus stellatus Vesque
Baccaurea sumatrana M.A.
Shorea agami Ashton
Ternstroemia aneura Miq.
Mezzetia parvifolia Becc.
Dead
A Field Guide to the Permanent Sample Plots • 83
Table 9. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dipterocarpus stellatus Vesque
Syzygium sp.
Dipterocarpus stellatus Vesque
Parastemon urophyllus A.DC.
Myristica beccarii Warb.
Swintonia glauca Engl.
Lithocarpus sp.
Canarium litorale Bl.
Dead
Shorea exelliptica Meijer
Santiria grifithii (Hk.f.) Engl.
Shorea venulosa Wood ex Meijer
Palaquium cochlearifolium van Royen
Gymnacranthera contracta Warb.
Dacryodes incurvata (Engler) H.J.Lam
Ternstroemia aneura Miq.
Hydnocarpus kunstleri (King) Warb
Agathis borneensis Warb.
Gluta wallichii (Hk.f.) Ding Hou
Hydnocarpus woodii Merr.
Prunus arborea (Bl.) Kalkm.
Shorea macroptera Dyer
Agathis borneensis Warb.
Dead
Scaphium macropodum (Miq.) Beumée ex Heyne
Madhuca malaccensis (Clarke) H.J.Lam
Prunus arborea (Bl.) Kalkm.
Horsieldia sp.
Hydnocarpus kunstleri (King) Warb
Hydnocarpus kunstleri (King) Warb
Dehaasia incrassata (Jack.)Kosterm.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Litsea irma Bl.
Unidentiied
Koompassia malaccensis Maing. ex Benth
Dipterocarpus lowii Hk.f.
84 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Hydnocarpus kunstleri (King) Warb
Shorea paucilora King
Madhuca oblongifolia Merr.
Madhuca malacensis (Clarke)H.J.Lam
Shorea macroptera Dyer
Agathis borneensis Warb.
Sindora leiocarpa Backer
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Artocarpus odoratissimus Blanco
Endiandra coriacea Merr.
Parastemon urophyllus A.DC.
Ixonanthes petiolaris Bl.
Oncosperma horridum Scheff.
Dehaasia irma Bl.
Diospyros laevigata Bakh.
Shorea rubra Ashton
Shorea beccariana Burck.
Agathis borneensis Warb.
Agathis borneensis Warb.
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Syzygium
Agathis borneensis Warb.
Agathis borneensis Warb.
Agathis borneensis Warb.
Agathis borneensis Warb.
Calophyllum pulcherrimum Wall.
Agathis borneensis Warb.
Dacryodes rostrata H.J.Lam
Dipterocarpus stellatus Vesque
Shorea parvifolia Dyer
Shorea maxwelliana King
Dead
A Field Guide to the Permanent Sample Plots • 85
Table 9. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Dipterocarpus stellatus Vesque
Callophylum venulosum Miq.
Shorea parvifolia Dyer
Dead
Gymnacranthera contracta Warb.
Santiria laevigata Bl.
Dipterocarpus stellatus Vesque
Pimeleodendron papaveroides J.J.S.
Agathis borneensis Warb.
Celtis sp.
Oncosperma horridum Scheff.
Ctenolophon parvifolius Oliv.
Dead
Melanochila elmeri Merr.
Garcinia maingayi Hook.f.
Horsieldia glabra (Bl.) Warb.
Alstonia spathulata Blume
Eugeissona utilis Becc
Madhuca oblongifolia Merr.
Gymnacranthera paniculata Warb.
Madhuca borneensis van Royen
Dead
Shorea macroptera Dyer
Shorea parvifolia Dyer
Adinandra borneensis Kobuski
Garcinia celebica L.
Pimeleodendron papaveroides J.J.S.
Calophyllum venulosum Miq.
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea parvifolia Dyer
Shorea parvifolia Dyer
Dipterocarpus stellatus Vesque
Palaquium calophyllum (T. & B.) Pierre
Dipterocarpus stellatus Vesque
86 • A Field Guide to the Permanent Sample Plots
Table 9. Continued
Tree #
297
298
299
300
301
302
303
304
305
306
307
308
309
310
311
312
Species
Artocarpus integer (Thunb.) Merr.
Shorea maxwelliana King
Baccaurea angulata Merr.
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Agathis borneensis Warb.
Palaquium calophyllum (T. & B.) Pierre
Callophyllum venulosum Miq.
Syzygium bankense (Hassk.) Merr. & Perry.
Calophyllum pulcherrimum Wall.
Durio lanceolatus Mast.
Palaquium gutta (Hook.f.) Bail.
Swintonia glauca Engl.
Swintonia glauca Engl.
Hydnocarpus kunstleri (King) Warb
Lithocarpus urceolaris (Jack) Merr.
A Field Guide to the Permanent Sample Plots • 87
Table 10. List of Trees in Plot CNV10
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Artocarpus anisophyllus Miq.
Microcos cinnamomifolia Burr.
Buchanania arborescens (Bl.) Bl.
Dillenia grandifolia Wall. ex. Hk.f.
Artocarpus anisophyllus Miq.
Mezzetia parvilora Miq.
Girroniera subaequalis Planch.
Koordersiodendron pinnatum (Blanco) Merr.
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Shorea exelliptica Meijer
Gluta wallichii Ding Hou
Agathis borneensis Warb.
Palaquium obtusifolium Burck
Koompassia malaccensis Maing. ex Benth.
Atuna excelsa (Jack) Kosterm.
Dead
Tetramerista glabra Miq.
Koordersiodendron pinnatum (Blanco) Merr.
Shorea parvifolia Dyer
Licania splendens (Korth.) Prance
Shorea macroptera Dyer
Lithocarpus urceolaris (Jack) Merr.
Palaquium hexandrum Engl.
Syzygium sp.
Shorea exelliptica Meijer
Shorea maxwelliana King
Shorea macroptera Dyer
Hydnocarpus kunstleri (Kirng) Warb.
Baccaurea sumatrana M.A.
Shorea maxwelliana King
Baccaurea edulis Merr.
Agathis borneensis Warb.
Helicia fuscotomentosa Suesseng
Shorea beccariana Burck
Shorea exelliptica Meijer
Hydnocarpus kunstleri (Kirng) Warb.
88 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Dead
Shorea exelliptica Meijer
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Dead
Shorea exelliptica Meijer
Melanochyla beccariana Oliv.
Dead
Calophyllum lowii Hook.f.
Myristica sp.
Palaquium cochlearifolium van Royen
Gluta wallichii Ding Hou
Syzygium bankense (Hassk.) Merr. & Perry
Shorea maxwelliana King
Mangifera swintonioides Kosterm.
Shorea maxwelliana King
Erycibe impressa Hoogl.
Shorea maxwelliana King
Agathis borneensis Warb.
Pimeleodendron papaveroides J.J.S.
Lithocarpus sundaicus Rehd.
Gymnacranthera contracta Warb.
Agathis borneensis Warb.
Syzygium operculata (Roxb.) Nied.
Xanthophyllum vitellinum (Bl.) D.Dietr.
Durio lanceolatus Mast.
Dillenia grandifolia Wall. ex Hk.f.
Palaquium calophyllum Pierre ex Burck
Artocarpus anysophyllus Miq.
Bhesa paniculata Arn.
Shorea exelliptica Meijer
Dialium sp.
Madhuca magniica H.J. Lam
Palaquium cochlearifolium van Royen
Shorea exelliptica Meijer
Shorea maxwelliana King
Shorea beccariana Burck
A Field Guide to the Permanent Sample Plots • 89
Table 10. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Unidentiied
Shorea beccariana Burck
Gluta wallichii (Hk.f.) Ding Hou
Shorea beccariana Burck
Lithocarpus sp.
Shorea maxwelliana King
Xanthophyllum sp.
Shorea maxwelliana King
Shorea sp.
Memecylon myrsinoides Blume
Shorea exelliptica Meijer
Swintonia glauca Engl.
Garcinia macroptera Dyer
Litsea sp.
Shorea macroptera Dyer
Myristica sp.
Calophyllum venulosum Miq.
Shorea maxwelliana King
Santiria grifithii (Hk.f.) Engl.
Shorea parvifolia Dyer
Shorea beccariana Burck
Garcinia dioica Bl.
Shorea maxwelliana King
Shorea sp.
Buchanania sessilifolia Bl.
Gluta wallichii (Hk.f.) Ding Hou
Unidentiied
Palaquium cochlearifolium van Royen
Scaphium macropodum (Miq.) Beumee ex Heyne
Shorea macroptera Dyer
Erycibe impressa Hoogl.
Agathis borneensis Warb.
Scaphium macropodum (Miq.) Beumee ex Heyne
Shorea exelliptica Meijer
Syzygium bankense (Hassk.) Merr. & Perry
Shorea maxwelliana King
Shorea pinanga Scheff.
90 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Nephelium maingayi Hiern.
Terminalia sp.
Shorea macroptera Dyer
Shorea exelliptica Meijer
Dead
Xylopia malayana Hook. f. Th.
Garcinia dioica Bl.
Myristica beccarii Warb.
Gymnacranthera contracta Warb.
Anisoptera sp.
Shorea parvifolia Dyer
Licania splendens (Korth.) Prance
Shorea sp.
Gymnacranthera contracta Warb.
Terminalia sp.
Shorea sp.
Shorea parvifolia Dyer
Dead
Dead
Shorea maxwelliana King
Shorea maxwelliana King
Dead
Ixonanthes petiolaris Bl.
Aporusa antennifera (Airy Shaw) Airy Shaw
Palaquium cochlearifolium van Royen
Shorea venulosa Wood ex Meijer
Gluta wallichii Ding Hou
Garcinia dioica Bl.
Garcinia macrophylla Miq.
Licania splendens (Korth.) Prance
Shorea maxwelliana King
Gluta wallichii Ding Hou
Helicia fuscotomentosa Suesseng
Knema latericia Elmer
Shorea maxwelliana King
Palaquium cochlearifolium van Royen
Myristica sp.
A Field Guide to the Permanent Sample Plots • 91
Table 10. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Calophyllum pulcherrimum Wall.
Shorea maxwelliana King
Garcinia dioica Bl.
Shorea maxwelliana King
Palaquium cochlearifolium van Royen
Sarcotheca diversifolia (Miq.) Hall.f.
Palaquium cochlearifolium van Royen
Pternandra rostrata (Cogn.) Ohwi
Shorea beccariana Burck
Dead
Lithocarpus cantleyanus (King ex Hk.f.) Rehd.
Santiria laevigata Bl. f. laevigata
Palaquium quercifolium Burck
Gluta wallichii (Hk.f.) Ding Hou
Agathis borneensis Warb.
Calophyllum lowii Hook.f.
Dead
Shorea parvifolia Dyer
Lithocarpus sp.
Shorea venulosa Wood ex Meijer
Shorea beccariana Burck
Eugeissonia utilis Becc.
Dead
Shorea parvifolia Dyer
Lithocarpus sp.
Girroniera subaequalis Planch.
Stemonurus grandifolius Becc.
Palaquium sp.
Shorea beccariana Burck
Lithocarpus sp.
Tetramerista glabra Miq.
Shorea exelliptica Meijer
Hydnocarpus kunstleri (King) Warb.
Knema furfuracea (Hk.f. ex Th.) Warb.
Dacryodes sp.
Shorea seminis (de Vriese) Slooten
Hydnocarpus gracilis (v. Sl.) Sleum
92 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dead
Macaranga conifera (Zoll.) M.A.
Erythroxylon sp.
Syzygium sp.
Eugeissonia utilis Becc.
Pternandra rostrata (Cogn.) Ohwi
Lithocarpus sp.
Dead
Eugeissonia utilis Becc.
Lithocarpus sp.
Mallotus tiliaefolia M.A.
Adinandra borneensis Kobuski
Koompassia malaccensis Maing. ex Benth.
Unidentiied
Helicia petiolaris Benn.
Gymnacranthera contracta Warb.
Dead
Dead
Hydnocarpus woodii Merr.
Calophyllum lowii Hook.f.
Shorea exelliptica Meijer
Arthocarpus anisophyllus Miq.
Shorea sp.
Palaquium cochlearifolium van Royen
Shorea beccariana Burck
Ctenolophon parvifolius Oliv.
Durio lanceolatus Mast.
Shorea beccariana Burck
Shorea agami Ashton
Lithocarpus ewyckii Rehd.
Shorea maxwelliana King
Myristica sp.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus lowii Hk.f.
Syzygium bankense (Hassk.) Merr. & Perry
Gluta wallichii (Hk.f.) Ding Hou
A Field Guide to the Permanent Sample Plots • 93
Table 10. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Drypetes kikir Airy Shaw
Helicia petiolaris Benn.
Lithocarpus sp.
Lithocarpus sp.
Palaquium cochlearifolium van Royen
Gymnacranthera contracta Warb.
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Gymnacranthera contracta Warb.
Koordersiodendron pinnatum (Blanco) Merr.
Gymnacranthera contracta Warb.
Shorea maxwelliana King
Dead
Scaphium macropodum (Miq.) Beumee ex Heyne
Palaquium cochlearifolium van Royen
Shorea maxwelliana King
Palaquium cochlearifolium van Royen
Shorea maxwelliana King
Gymnacranthera contracta Warb.
Gluta wallichii (Hk.f.) Ding Hou
Erycibe impressa Hoogl.
Hopea bracteata Burck.
Dipterocarpus lowii Hk.f.
Agathis borneensis Warb.
Dead
Lithocarpus ewyckii Rehd.
Atuna excelsa (Jack) Kosterm.
Dead
Shorea venulosa Wood ex Meijer
Palaqium quercifolium Burk.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Litsea irma Hk. f
Gluta wallichii (Hk.f.) Ding Hou
Syzygium sp.
Baccaurea sumatrana M.A.
Shorea venulosa Wood ex Meijer
94 • A Field Guide to the Permanent Sample Plots
Table 10. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
296
Species
Litsea irma Hk. f
Dead
Gymnacranthera contracta Warb.
Shorea macroptera Dyer
Prunus spicata Kalkm.
Helicia petiolaris Benn.
Shorea exelliptica Meijer
Neonauclea calycina (Bart.) Merr.
Shorea parvilora Dyer
Sarcotheca diversifolia (Miq.) Hall.f.
Scaphium macropodum (Miq.) Beumée ex Heyne
Gluta walichii (Hk.f. Ding Hou
Shorea maxwelliana King
Dipterocarpus lowii Hk.f.
Hopea bracteata Burk.
Dipterocarpus lowii Hk.f.
Agathis borneensis Warb.
Lophopetalum beccarianum Pierre
Shorea maxwelliana King
Hydnocarpus kunstleri (King) Warb.
Hydnocarpus kunstleri (King) Warb.
Shorea exelliptica Meijer
Knema glauca (Bl.) Warb.
Dipterocarpus lowii Hk.f.
Hydnocarpus kunstleri (King) Warb.
Cryptocarya crassinervia Miq.
Shorea maxwelliana King
Shorea maxwelliana King
Shorea rubra Ashton
Shorea macroptera Dyer
Syzygium bankense (Hassk.) Merr. & Perry
Myristica sp.
Xanthophyllum sp.
Gluta wallichii (Hk.f.) Ding Hou
Shorea maxwelliana King
Palaquium leiocarpum Boerl.
Shorea parvifolia Dyer
A Field Guide to the Permanent Sample Plots • 95
Table 10. Continued
Tree #
297
298
299
300
301
302
303
304
305
Species
Pternandra rostrata (Cogn.) Ohwi
Gymnacranthera contracta Warb.
Lithocarpus sericobalanus E.F. Warb.
Lithocarpus sp.
Shorea sp.
Unidentiied
Shorea parvifolia Dyer
Lithocarpus pulcher (King) Markgr.
Alseodaphne havilandii (Gamble) Kosterm.
20
40
60
80
100
289
279
100
100
0
290
288
263
253
254
242 236
248
60
159
220
166
176
164
40
128
165
193
202
194
179
180
190
100
115
101 102
116 114
122
123
124
130
108
106
107
99
136
151
8
9
12
11
133
14
13
87
93
16
10
4
3
15
32
19
27
20
44
61
64
62
58
41
28
37
26
40
39
38
53
52
49
24
57
56
54
92
91
90
75
63
74
60
59
72
55
50 51
67
68
0
20
40
60
80
CNV11
Lauraceae
FAMILY
Linaceae
Anacardiaceae
Melastomataceae
Apocynaceae
Meliaceae
Araucariaceae
Myristicaceae
Arecaceae
Myrtaceae
Bombacaceae
Oxalidaceae
Burseraceae
Polygalaceae
Celastraceae
Rhizophoraceae
Clusiaceae
Rosaceae
Dipterocarpaceae
Sapindaceae
Elaeocarpaceae
Sapotaceae
Euphorbiacea
Sterculiaceae
Fabaceae
Theaceae
Fagaceae
Ulmaceae
76
69
Flacourtiaceae
70
0
22
96
77
0
2
88 89
109
35
31 30
17
18
23
117
36
34
79
80
121
120
132
81
82
134
138
201
199
198
98
131
129
139
200
197
167
126 125
127
141
20
195
186
187
189 188
181
173
163
145 144
140
6
185
40
168
143
152
183 182
177
178
172
162
150
184
175
154
149
203
213
169 170
155
146
204
211
214
157
156
205
206
210
217
160
295
294
293
207
208
209
216 215
234
158
292
97
233
247 245 243
246
278
223
222
232
249
273
227 225 224
230
229
237
235
251
252
284
281
226
239
240
238
291
286
285
20
80
261
255
283
266 269 270
271
267 268
272
262
280
80
260
277
60
264
250
287
265
257
100
96 • A Field Guide to the Permanent Sample Plots
Figure 14. Distribution of Trees in Plot CNV11
A Field Guide to the Permanent Sample Plots • 97
Table 11. List of Trees in Plot CNV11
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Dead
Shorea ovalis (Korth.) Bl.
Drypetes kikir Airy Shaw
Shorea fallax Meijer
Dead
Nephelium maingayi Hiern.
Eugeissona utilis Becc
Scaphium macropodum (Miq.) Beumée ex Heyne
Shorea exelliptica Meijer
Palaquium calophyllum Pierre ex Burck
Durio lanceolatus Mast.
Melanochyla beccariana Oliv
Shorea rubra Ashton
Gymnacranthera contracta Warb.
Quercus subsericea A. Camus
Canarium sp.
Lithocarpus sundaicus Rehd.
Vatica umbonata (Hook.f.) Burck.
Santiria grifithii (Hk.f.) Engl.
Koompassia malaccensis Maing.. ex Benth.
Tetramerista glabra Miq.
Shorea macroptera Dyer
Shorea exelliptica Meijer
Lophopetalum subobovatum King
Dead
Shorea macroptera Dyer
Licania splendens (Korth.) Prance
Shorea rubra Ashton
Drypetes kikir Airy Shaw
Shorea maxwelliana King
Koompassia malaccensis Maing.. ex Benth.
Termstroemia coriacea Scheff.
Dead
Knema sp.
Shorea rubra Ashton
Swintonia glauca Engl.
Shorea parvifolia Dyer
98 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
69
70
71
72
73
74
Species
Garcinia nervosa Miq.
Hydnocarpus sp.
Shorea exelliptica Meijer
Shorea parvifolia Dyer
Swintonia glauca Engl.
Dead
Shorea rubra Ashton
Eugenia fastigiata Miq.
Gluta wallichii (Hk.f.) Ding Hou
Santiria laevigata Bl.
Santiria oblongifolia Bl.
Memecylon myrsinoides Blume
Scaphium macropodum (Miq.) Beumée ex Heyne
Melanochyla cf. becariana Oliv
Syzygium tawahense (Korth.) Merr. & Perry
Shorea macroptera Dyer
Hydnocarpus kunstleri (King) Warb.
Lithocarpus sp.
Madhuca malaccensis (Clarke) H.J. Lam
Shorea parvifolia Dyer
Artocarpus integer (Thunb.) Merr.
Eugeissona utilis Becc
Eugeissona utilis Becc
Shorea maxwelliana King
Shorea maxwelliana King
Santiria grifithii (Hk.f.) Engl.
Shorea agami Ashton
Shorea maxwelliana King
Syzygium sp.
Durio lanceolatus Mast.
Syzygium sp.
Shorea rubra Ashton
Dipterocarpus lowii Hk.f.
Diospyros functiculosa Bakh.
Litsea nidularis Gamble
Lophopetalum beccarianum Pierre
Shorea maxwelliana King
A Field Guide to the Permanent Sample Plots • 99
Table 11. Continued
Tree #
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
Species
Gluta wallichii (Hk.f.) Ding Hou
Syzygium sp.
Shorea sp.
Agathis borneensis Warb.
Garcinia macrophylla Miq.
Shorea sp.
Memecylon myrsinoides Blume
Shorea sp.
Dead
Dead
Calophyllum sp.
Gluta wallichii (Hk.f.) Ding Hou
Dipterocarpus lowii Hk.f.
Gluta wallichii (Hk.f.) Ding Hou
Shorea parvifolia Dyer
Vatica umbonata (Hook.f.) Burck.
Quercus subsericea A. Camus
Agathis borneensis Warb.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Dead
Baccaurea simatrana M.A.
Shorea sp.
Shorea maxwelliana King
Santiria grifithii (Hk.f.) Engl.
Shorea maxwelliana King
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea venulosa Wood ex Meijer
Lithocarpus sp.
Dipterocarpus lowii Hk.f.
Palaquium cochlearifolium van Royen
Shorea sp.
Agathis borneensis Warb.
Shorea sp.
100 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
112
113
114
115
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
Species
Gluta wallichii (Hk.f.) Ding Hou
Dead
Shorea sp.
Palaquium cochlearifolium van Royen
Gluta wallichii (Hk.f.) Ding Hou
Calophyllum sp.
Dead
Dead
Shorea sp.
Shorea venulosa Wood ex Meijer
Shorea sp.
Gluta wallichii (Hk.f.) Ding Hou
Xanthophyllum sp.
Anisoptera sp.
Xanthophyllum sp.
Shorea sp.
Vatica umbonata (Hook.f.) Burck.
Swintonia glauca Engl.
Syzygium sp.
Swintonia glauca Engl.
Palaquium cochleariifolium van Royen
Dialium trifoliatum de Wit.
Gluta wallichii (Hk.f.) Ding Hou
Dead
Gymnacranthera contracta Warb.
Dead
Shorea sp.
Shorea exelliptica Meijer
Dacryodes rugosa H.J.Lam.
Durio sp.
Dead
Shorea exelliptica Meijer
Knema cinerea (Poir.) Warb.
Shorea macroptera Dyer
Memecylon myrsinoides Blme
Dead
Durio oxleyanus Griff.
A Field Guide to the Permanent Sample Plots • 101
Table 11. Continued
Tree #
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
Species
Shorea venulosa Wood ex Meijer
Syzygium sp.
Dipterocarpus stellatus Vasque
Shorea sp.
Unidentiied
Syzygium sp.
Artocarpus odoratissima Blanco
Ptychopyxis bacciformis Croiz.
Ptychopyxis bacciformis Croiz.
Gluta wallichii (Hk.f.) Ding Hou
Pternandra rostrata (Cogn.) Ohwi
Rhodamnia cinerea Jack.
Dead
Gluta wallichii (Hk.f.) Ding Hou
Shorea macroptera Dyer
Hydnocarpus woodi Merr.
Ptychopyxis bacciformis Croiz.
Calophyllum lowii Hook.f.
Sarcotheca diversifolia (Miq.) Hall .f.
Swintonia glauca Engl.
Gluta wallichii (Hk.f.) Ding Hou
Gymnacranthera contracta Warb.
Dead
Shorea sp.
Syzygium sp.
Dead
Syzygium sp.
Agathis borneensis Warb.
Agathis borneensis Warb.
Shorea beccariana Burck.
Lophopetalum beccarianum Pierre
Shorea sp.
Syzygium bankense (Hassk.) Merr. & Perry
Shorea parvifolia Dyer
Ixonanthes petiolaris Bl.
Artocarpus anisophyllus \Miq.
Gluta wallichii (Hk.f.) Ding Hou
102 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
Species
Dyera costulata (Miq.) Hk.f.
Melanochyla beccariana Oliv.
Syzygium sp.
Dipterocarpus lowii Hk.f.
Swintonia glauca Engl.
Gluta wallichii (Hk.f.) Ding Hou
Dacryodes rostrata H.J.Lam forma pubescens
Shorea parvifolia Dyer
Drypetes kikir Airy Shaw
Shorea ovalis (Korth.) Bl.
Mangifera swintonioides Kosterm.
Palaquium eriocalyx H.J.Lam
Shorea sp.
Shorea sp.
Shorea sp.
Shorea maxwelliana King
Dipterocarpus lowii Hk.f.
Celtis philippensis Blanco
Santiria oblongifolia Bl.
Dipterocarpus humeratus Slooten
Madhuca oblongifolia Merr.
Calophyllum venullosum Zoll.
Syzygium splendens (Bl.) Merr. & Perry
Quercus sp.
Tetramerista glabra Miq.
Knema furfuracea (Hk.f. ex Th.) Warb.
Dead
Lithocarpus sp.
Dialium platycepalum Backer
Gluta wallichii (Hk.f.) Ding Hou
Shorea rubra Ashton
Madhuca erythrophylla H.J.Lam
Ptychopyxis bacciformis Croiz.
Dead
Shorea paucilora King
Dead
Knema cinerea (Poir.) Warb.
A Field Guide to the Permanent Sample Plots • 103
Table 11. Continued
Tree #
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
255
256
257
258
259
Species
Durio grandilorus Kost.
Shorea maxwelliana King
Heritiera javanica (Bl.) Kosterm.
Semecarpus macrophylla Merr.
Garcinia nervosa Miq.
Dead
Gymnacranthera contracta Warb.
Shorea agamii Ashton
Hydnocarpus sp.
Vatica sp.
Quercus sp.
Koompassia malaccensis Maing. ex Benth.
Shorea parvifolia Dyer
Vatica umbonata (Hook.f.) Burck.
Cleistanthus bakonensis Airy Shaw
Hydnocarpus sp.
Canarium littorale Bl.
Nephelium maingayi Hiern
Syzygium confertum (Korth.) Merr. & Perry
Shorea parvifolius Dyer
Shorea ovalis (Korth.) Bl.
Unidentiied
Shorea ovalis (Korth.) Bl.
Calophyllum venulosum Zoll.
Phoebe laevis Kosterm.
Koompassia malaccensis Maing. ex Benth.
Shorea ovalis (Korth.) Bl.
Syzygium sp.
Dacryodes sp.
Ternstroemia aneura Miq.
Shorea paucilora King
Gluta wallichii (Hk.f.) Ding Hou
Shorea ovalis (Korth.) Bl.
Dead
Tetramerista glabra Miq.
Pterocarpidia polyneura (Miq.) Brem.
Dead
104 • A Field Guide to the Permanent Sample Plots
Table 11. Continued
Tree #
260
261
262
263
264
265
266
267
268
269
270
271
272
273
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
290
291
292
293
294
295
Species
Syzygium splendens (Bl.) Merr. & Perry
Adinandra dumosa Jack.
Vatica vinosa Ashton
Tetramerista glabra Miq.
Dysoxylum
Vatica vinosa Ashton
Lithocarpus sundaicus Rehd.
Hydnocarpus
Gluta wallichii (Hk.f.) Ding Hou
Shorea paucilora King
Cinnamomum javanicum Bl.
Shorea parvifolia Dyer
Dacryodes sp.
Shorea macroptera Dyer
Pimeleodendron papaveroides J.J.S.
Unidentiied
Syzygium sp.
Nephelium uncinatum Leenh.
Vatica cf. granulata Slooten
Cleistanthes myrianthus Krz.
Macaranga pruinosa M.A.
Dacryodes rostrata H.J.Lam
Dead
Carallia brachyata (Lour.) Merr.
Pternandra rostrata (Cogn.) ohwi
Horsieldia sp.
Shorea ochracea Sym.
Pternandra rostrata (Cogn.) ohwi
Tetramerista glabra Miq.
Ternstroemia aneura Miq.
Knema laurina (Bl.) Warb.
Tetramerista glabra Miq.
Shorea ovalis (Korth.) Bl.
Oncosperma horridum Scheff.
Myristica villosa Warb.
Syzygium sp.
0
20
40
60
80
284
271
263
269
268
283
274
272
273
285
286
281
100
296
295
294
298
297
287
282
292
275
261
291
280
279
100
100
Figure 15. Distribution of Trees in Plot CNV12
259
251
240
250
247
245
60
249
248
141
140
133
132
126
127
152
142
147
146
143
233
153
40
115
106
105
108 107
114
117
123
113
164
162
80
174
173
178
85
72
86
91
69
93
95
73
74
83
75
81
20
24
77 78
80
8
2
17
14 15
4
1
9 12 13
5
6
27
22
10
28
20
40
Araucariaceae
Melastomataceae
187
Arecaceae
Meliaceae
Bombacaceae
Myristicaceae
Burseraceae
Myrtaceae
Clusiaceae
Ochnaceae
Crypteroniaceae
Oxalidaceae
Dilleniaceae
Proteaceae
Dipterocarpaceae
Rosaceae
Ebenaceae
Rubiaceae
Elaeocarpaceae
Sapindaceae
Euphorbiacea
Sapotaceae
Fabaceae
Sterculiaceae
Fagaceae
Tiliaceae
Flacourtiaceae
Verbenaceae
49
50
62
42 43
35
44
48
45
37
33
30
36
34
46 47
32
31
0
20
186
52
55 53
54
56
61
65
39 38
29
21
0
Linaceae
41
19
11
Lecythidaceae
60
63
60
80
Junglandaceae
Apocynaceae
51
67
25 79
26
23
59
FAMILY
Lauraceae
184
68
66
98
57
Icacinaceae
Annonaceae
20
16
7
183
58
109
110
70
71
125
124
185
179
CNV12
Anacardiaceae
188
176 175
88
97 96 94
111
190
181
182
161
87
90
99
191
180
89
100
192
163
160
157
101
104
112
171
166
130 131
103
102
205
201
203
202
204
172
165
156
158
159
151
129
199 200
169 170
154
155
149
193
213
208
168
167
145
116
209
196
198
214
207
206
227
150
121
120
122
211
225
119
215
212
224
139
118
195
219
222
223
100
A Field Guide to the Permanent Sample Plots • 105
135
217
221
228
238
134 136
128
220
236
237
241
230
234
235
239
242
244 243
246
231
252
289
216
0
80
229
253
288
277
256
254
278
276
218
60
267
265
232
40
266
260 262
106 • A Field Guide to the Permanent Sample Plots
Table 12. List of Trees in Plot CNV12
Tree #
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
34
35
36
37
Species
Dillenia grandifolia Wall. ex Hk.f.
Teijsmaniodendron simplicifolium Merr.
Litsea grandis (Wall. ex Nees) Hook.f.
Alseodaphne falcata (Bl.) Boerl.
Dacryodes rubiginosa H.J. Lam.
Syzygium sp.
Dillenia excelsa (Jack.) Gilg.
Syzygium sp.
Dialium trifoliatum de Wit
Shorea macroptera Dyer
Knema sp.
Oncosperma horridum
Oncosperma horridum
Canarium littorale Bl.
Shorea parvifolia Dyer
Koompasia malaccensis Maing ex Benth.
Lithocarpus blumeanus Rehd.
Lithocarpus blumeanus Rehd.
Crypteronia grifithii Clarke
Knema glauca (Bl.) Warb.
Sarcotheca diversifolia (Miq.) Hall.f.
Xylopia caudata Hk.f. ex Th.
Gluta wallichii Ding Hou
Dipterocarpus stellatus Vesque
Gonocaryum impressinervium Sleum.
Dialium trifoliatum de Wit
Sindora coriacea Maingayi ex Prain
Hydnocarpus kunstleri King
Baccaurea kunstleri King
Lithocarpus blumeanus Rehd.
Gymnacranthera contracta Warb.
Madhuca malaccensis (Clarke) H.J.Lam
Swintonia glauca Engl.
Koompasia malaccensis Maing ex Benth.
Shorea beccariana Burk.
Knema glauca (Bl.) Warb.
Shorea maxwelliana King
A Field Guide to the Permanent Sample Plots • 107
Table 12. Continued
Tree #
38
39
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
65
66
67
68
69
70
71
72
73
74
75
77
Species
Syzygium chloranthum
Dialium trifoliatum de Wit
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
Anisoptera costata Korth.
Elaeocarpus cf pedunculatus Wall.
Durio carinatus Mast.
Agathis borneensis Warb.
Syzygium fastigiata
Shorea macroptera Dyer
Palaquium calophyllum
Gluta wallichii (Hk.f.) Ding Hou
Lithocarpus sp.
Knema cinerea (Poir) Warb.
Barringtonia scortechinii Kurz.
Knema cinerea (Poir) Warb.
Mangifera swintonioides Kosterm.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Knema laurina (Bl.) Warb.
Dyera costulata (Miq.) Hk.f.
Shorea maxwelliana King
Helicia pescotomentosa Sueneus
Shorea macroptera Dyer
Shorea parvifolia Dyer ssp. Parvifolia
Shorea maxwelliana King
Shorea maxwelliana King
Vatica rassak (Korth.) Blume
Palaquium hexandrum
Shorea excelliptica Meijer
Knema intermedia
Shorea excelliptica Meijer
Shorea macroptera Dyer
Agathis borneensis Warb.
Dacryodes rubiginosa H.J.Lam
Lithocarpus blumeanus (Korth.) Rehd.
Diospyros sp.
108 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
78
79
80
81
83
85
86
87
88
89
90
91
92
93
94
95
96
97
98
99
100
101
102
103
104
105
106
107
108
109
110
111
112
113
114
115
116
Species
Durio lanceolata Mast.
Dipterocarpus stellatus Vesque
Garcinia macrophylla Miq.
Shorea rubra
Shorea paucilora King
Platea sp.
Shorea parvifolia Dyer
Shorea excelliptica Meijer
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Hydnocarpus woodii Merr.
Shorea maxwelliana King
Aporosa falcifera Hk.f.
Shorea maxwelliana King
Shorea rubra
Gymnacranthera contracta Warb.
Palaquium rostratum (Miq.) Burck.
Shorea parvifolia Dyer
Hydnocarpus woodii Merr.
Anisoptera
Syzygium opaca
Shorea parvifolia Dyer ssp. Parvifolia
Hydnocarpus woodii Merr.
Oncosperma horidum
Timonius wallichianus
Payena acuminatissima
Sarcotheca diversifolia (Miq.) Hall.f.
Oncosperma horidum
Oncosperma horidum
Hydnocarpus woodii Merr.
Licania splendens (Korth.) Prance
Parashorea smythiesii Wyatt-Smith ex Ashton
Neonauclea excelsa (Bl.) Merr.
Durio
Knema laurina (Bl.) Warb.
Vatica sp.
Shorea parvifolia Dyer ssp. Parvifolia
A Field Guide to the Permanent Sample Plots • 109
Table 12. Continued
Tree #
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
134
135
136
137
138
139
140
141
142
143
144
145
146
147
149
150
151
152
153
154
Species
Diospyros sp.
Knema laurina (Bl.) Warb.
Ryparosa hulletii King
Parashorea smythiesii Wyatt-Smith ex Ashton
Knema cinerea (Poir.) Warb.
Hydnocarpus woodii Merr.
Pentace discolor Merr.
Teijsmaniodendron simplicifolium Merr.
Nephelium sp.
Moultonianthus leemburgianus
Oncosperma horidum
Lithocarpus sp.
Madhuca oblongifolia
Macaranga gigantea
Madhuca oblongifolia
Lithocarpus urceolatus
Aporosa sp.
Shorea maxwelliana King
Dipterocarpus stellatus Vesque
Shorea maxwelliana King
dead
dead
Hydnocarpus woodii Merr.
Paratocarpus venenosa Becc.
Shorea maxwelliana King
Shorea agamii Ashton
Ctenolophon parvifolius Oliv.
dead
Shorea rubra
Lithocarpus sp.
Shorea maxwelliana King
Koompasia malaccensis Maing ex Benth.
Dacryodes incurvata H.J.Lam.
Dipterocarpus stellatus Vesque
Dipterocarpus stellatus Vesque
Gardenia tubifera Wall.
Shorea ovalis (Korth.) Bl.
110 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
155
156
157
158
159
160
161
162
163
164
165
166
167
168
169
170
171
172
173
174
175
176
178
179
180
181
182
183
184
185
186
187
188
189
190
191
192
Species
Palaquium rostratum (Miq.) Burck.
Shorea beccariana Burk.
Vatica umbonata (Hook.f.) Burk.
Shorea macroptera Dyer
Melanochyla beccariana Oliv.
Dipterocarpus stellatus Vesque
Agathis boneensis Warb.
Shorea macroptera Dyer
Agathis boneensis Warb.
Shorea macroptera Dyer
Gymnacranthera contracta Warb.
Agathis boneensis Warb.
Gymnacranthera contracta Warb.
Shorea macroptera Dyer
Sarcotheca diversifolia (Miq.) Hall.f.
Shorea beccariana Burk.
Sarcotheca diversifolia (Miq.) Hall.f.
Baccaurea sumatrana M.A.
Knema glauca (Bl.) Warb.
Herritiera javanica (Bl.) Kosterm
Calophyllum lowii Hook.f.
Baccaurea sumatrana M.A.
Prunus spicata Kalkm
Pternandra rostrata (Cogn.) Ohwi
Payena endertii H.J.Lam.
Drypetes kikir Airy Shaw
Shorea parvifolia Dyer
Parkia sp.
Gymnacranthera contracta Warb.
Elateriospermum tapos Bl.
Hydnocarpus kunstlerii (King) Warb.
Dialium trifoliatum de Wit
Knema cinerea (Poir.) Warb.
Castanopsis javanica (Bl.) A.Dc.
Elateriospermum tapos Bl.
Shorea agamii Ashton
Shorea parvifolia Dyer ssp. Parvifolia
A Field Guide to the Permanent Sample Plots • 111
Table 12. Continued
Tree #
193
195
196
197
198
199
200
201
202
203
204
205
206
207
208
209
211
212
213
214
215
216
217
218
219
220
221
222
223
224
225
227
228
229
230
231
232
Species
Shorea macroptera Dyer
Artocarpus anisophyllus Miq.
Artocarpus anisophyllus Miq.
Diospyros sp.
Cephalomappa lepidotula Airy Shaw
Knema cinerea (Poir.) Warb.
Artocarpus anisophyllus Miq.
Englhardia serrata Bl.
Shorea parvifolia Dyer ssp. Parvifolia
Syzygium tawahense
Artocarpus integra (Thunb.) Merr.
Artocarpus anisophyllus Miq.
Knema cinerea (Poir.) Warb.
Shorea sp.
Licania splendens (Korth.) Prance
Litsea nidularis Gamble
Cleistanthus myrianthus Krz.
Dyera costulata (Miq.) Hk.f.
Polyanthia glauca Boerl.
Artocarpus anisophyllus Miq.
Shorea rubra
Palquium sumatranum Burck.
Shorea beccariana Burk.
Syzygium sp.
Knema conferta (King.) Warb.
Knema conferta (King.) Warb.
Sarcotheca diversifolia (Miq.) Hall.f.
Syzygium perpuncticulatum
Syzygium perpuncticulatum
Sarcotheca diversifolia (Miq.) Hall.f.
Agathis boneensis Warb.
Oncosperma horridum
Pterocymbium tubulatum (Most.) Pierre
Mangifera swintonioides Kosterm.
Pimeleodendron papaveroides J.J.S.
Lithocarpus blumeanus (Korth.) Rehd.
Lithocarpus blumeanus (Korth.) Rehd.
112 • A Field Guide to the Permanent Sample Plots
Table 12. Continued
Tree #
233
234
235
236
237
238
239
240
241
242
243
244
245
246
247
248
249
250
251
252
253
254
256
259
260
261
262
263
264
265
266
267
268
269
271
272
273
Species
Sindora beccariana Back.
Teijsmaniodendron simplicifolium Merr.
Shorea leprosula Miq.
Ctenolophon parvifolius Oliv.
Palaqium gutta (Hook.f.) Baill.
Garcinia gaudichaudii
Shorea beccariana Burk.
Garcinia macrophylla Miq.
Shorea maxwelliana Miq.
Shorea parvifolia Dyer.
Diospyros sp.
Gluta wallichii (Hk.f.) Ding Hou
Licania splendens (Korth.) Prance
Baccaurea
Palaquium rostratum
Helicia pescotomentosa Sueneus
Madhuca sp.
Garcinia dioica Bl.
Shorea parvifolia Dyer.
Licania splendens (Korth.) Prance
Calophyllum pulcherrianum Wall. ex Choisy
Hydnocarpus kunstleri (King.) Warb.
Syzygium opaca
Santiria tomentosa
Parinari sp.
Sagera elliptica (DC.) Hk.f. ex Th.
Syzygium sp.
Alstonia soathulata Blume
Elateriospermum tapos Bl.
Elateriospermum tapos Bl.
Calophyllum cf. soulatri Burm.f.
Garcinia nervosa Miq.
Dehaasia irma Bl.
Koompasia malaccensis Maing ex Benth.
Parishia maingayi Hk.f.
Gymnacranthera paniculata Warb.
Anisoptera
A Field Guide to the Permanent Sample Plots • 113
Table 12. Continued
Tree #
274
275
276
277
278
279
280
281
282
283
284
285
286
287
288
289
291
292
293
294
295
296
297
298
Species
Vatica rassak (Korth.) Blume
Syzygium endertii
Dacryodes rostrata H.J.Lam.
Baccaurea edulis Merr.
Lithocarpus sp.
Mallotus echinatus
Hydnocarpus woodii Merr.
Dacryodes rostrata H.J.Lam.
Madhuca corthalsii (Pierre) H.J.Lam
Mangifera swintonioides Kosterm.
Baccaurea edulis Merr.
Brackenridgea palustris Bartell.
Baccaurea motleyana M.A.
Cleistanthus myrianthus Krz.
Knema laurina (Bl.) Warb.
Garcinia dioica Bl.
Vatica rassak (Korth.) Blume
Artocarpus anisophyllus Miq.
Pentace laxilora
Sterculia rubiginosa Vent.
Dyera costulata (Miq.) Hk.f.
Licania splendens (Korth.) Prance
Shorea pinanga Scheff.
Calophyllum pulcherrimum Wall. ex Choisy
The Center for International Forestry Research (CIFOR) is a leading international
forestry research organization established in 1993 in response to global concerns about the
social, environmental, and economic consequences of forest loss and degradation. CIFOR
is dedicated to developing policies and technologies for sustainable use and management
of forests, and for enhancing the well-being of people in developing countries who rely on
tropical forests for their livelihoods. CIFOR is one of the 15 Future Harvest centres of the
Consultative Group on International Agricultural Research (CGIAR). With headquarters
in Bogor, Indonesia, CIFOR has regional ofices in Brazil, Burkina Faso, Cameroon and
Zimbabwe, and it works in over 30 other countries around the world.
Donors
The Center for International Forestry Research (CIFOR) receives its major funding from
governments, international development organizations, private foundations and regional
organizations. In 2005, CIFOR received inancial support from Australia,Asian Development
Bank (ADB), Belgium, Brazil, Canada, China, Centre de coopération internationale
en recherche agronomique pour le développement (CIRAD), Cordaid, Conservation
International Foundation (CIF), European Commission, Finland, Food and Agriculture
Organization of the United Nations (FAO), Ford Foundation, France, German Agency for
Technical Cooperation (GTZ), German Federal Ministry for Economic Cooperation and
Development (BMZ), Indonesia, International Development Research Centre (IDRC),
International Fund for Agricultural Development (IFAD), International Tropical Timber
Organization (ITTO), Israel, Italy, The World Conservation Union (IUCN), Japan, Korea,
Netherlands, Norway, Netherlands Development Organization, Overseas Development
Institute (ODI), Peruvian Secretariat for International Cooperation (RSCI), Philippines,
Spain, Sweden, Swedish University of Agricultural Sciences (SLU), Switzerland, Swiss Agency
for the Environment, Forests and Landscape, The Overbrook Foundation, The Nature
Conservancy (TNC), Tropical Forest Foundation, Tropenbos International, United States,
United Kingdom, United Nations Environment Programme (UNEP), World Bank, World
Resources Institute (WRI) and World Wildlife Fund (WWF).
The CIFOR Permanent Sample Plots (PSP) were
established in 1998 in conjunction with the Phase I of the
ITTO Project dealing with sustainable forest management,
integrated land use, reduced-impact logging, biodiversity
in the production forest, and cultural and social aspects
of logging. In terms of technical perspectives, PSP are an
important tool to monitor forest dynamics and changes,
long term growth and yield and to provide critical data for
evaluation of ecological models. For silvicultural purposes
PSP supply data on diameter and volume increment as
well as stand structure dynamics.
In particular, the PSP in the Malinau Research Forest
provides a valuable insight into the diversity of trees found
there and the growth of the forest. This guidebook has
been designed to facilitate the growing interests and needs
of visitors to the Malinau Research Forest (MRF).
PATEN MA LINAU
KAB U
I N TI M U N G