Vascular plants of Greece: An annotated checklist.
Supplement
Authors: Dimopoulos, Panayotis, Raus, Thomas, Bergmeier, Erwin,
Constantinidis, Theophanis, Iatrou, Gregoris, et al.
Source: Willdenowia, 46(3) : 301-347
Published By: Botanic Garden and Botanical Museum Berlin (BGBM)
URL: https://doi.org/10.3372/wi.46.46303
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Willdenowia
Annals of the Botanic Garden and Botanical Museum Berlin-Dahlem
PANAYOTIS DIMOPOULOS1*, THOMAS RAUS2, ERWIN BERGMEIER3, THEOPHANIS CONSTANTINIDIS4,
GREGORIS IATROU1, STELLA KOKKINI5, ARNE STRID6 & DIMITRIOS TZANOUDAKIS1
Vascular plants of Greece: An annotated checklist. Supplement
Version of record first published online on 26 October 2016 ahead of inclusion in December 2016 issue.
Abstract: Supplementary information on taxonomy, nomenclature, distribution within Greece, total range, life form
and ecological traits of vascular plants known to occur in Greece is presented and the revised data are quantitatively
analysed. Floristic discrepancies between Vascular plants of Greece: An annotated checklist (Dimopoulos & al.
2013) and relevant influential datasets (Flora europaea, Med-Checklist, Euro+Med PlantBase, etc.) are explained
and clarified. An additional quantity of synonyms and misapplied names used in previous Greek floristic literature is
presented. Taxonomic and floristic novelties published after 31 October 2013 are not considered.
Key words: vascular plants, Europe, Greece, checklist, distribution, taxonomy, nomenclature
Article history: Received 6 May 2016; peer-review completed 4 July 2016; received in revised form 28 August
2016; accepted for publication 7 September 2016.
Citation: Dimopoulos P., Raus Th., Bergmeier E., Constantinidis Th., Iatrou G., Kokkini S., Strid A. & Tzanoudakis D. 2016: Vascular plants of Greece: An annotated checklist. Supplement. – Willdenowia 46: 301 – 347. doi:
http://dx.doi.org/10.3372/wi.46.46303
Introduction
Vascular plants of Greece: An annotated checklist (Dimopoulos & al. 2013) constitutes the first comprehensive
inventory of the flora of Greece more than a century after
Eugen von Halácsy, a Hungarian-born physician of Vienna, had finalized the most recent complete Flora of the
country (Halácsy 1900 – 1904, with supplements in 1908
and 1912). The Checklist of 2013, eagerly awaited by the
scientific and general public, met with a kind reception
of professional and amateur botanists, as well as national
and regional politicians and administrators responsible
for nature conservation activities in Greece. The quite
active use of the book, which started immediately after
publication, revealed a certain number of alleged and
factual discrepancies in content compared to contemporary influential datasets (Tutin & al. 1968 – 1980, 1993;
Greuter & al. 1984 – 1989; Greuter & Raab-Straube 2008;
Euro+Med 2006+). Considering the significance of the
work as a reference for scientific and political action in
1 University of Patras, Department of Biology, Division of Plant Biology, Panepistimiopolis, GR-26504 Rio, Greece; *e-mail:
pdimopoulos@upatras.gr (author for correspondence); iatrou@upatras.gr; tzanoyd@upatras.gr
2 Botanischer Garten und Botanisches Museum Berlin, Freie Universität Berlin, Königin-Luise-Str. 6–8, D-14195 Berlin, Germany;
e-mail: t.raus@bgbm.org
3 University of Göttingen, Albrecht von Haller Institute of Plant Sciences, Untere Karspüle 2, D-37073 Göttingen, Germany; e-mail:
erwin.bergmeier@bio.uni-goettingen.de
4 National and Kapodistrian University of Athens, Faculty of Biology, Department of Ecology and Systematics, Panepistimiopolis,
GR-15703, Athens, Greece; e-mail: constgr@biol.uoa.gr
5 Aristotle University of Thessaloniki, School of Biology, Department of Botany, GR-54124 Thessaloniki, Greece; e-mail:
kokkini@bio.auth.gr
6 Bakkevej 6, DK-5853 Ørbæk, Denmark; e-mail: arne.strid@youmail.dk
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302
order to safeguard the wealth of the natural heritage of
Greece, mere omission of names of taxa that are disregarded for Greece as being reported in error, non-established aliens, non-stabilized hybrids, etc. is unsatisfactory if the reasons for their exclusion are not explained
and commented on. Therefore, we offer here an array
of comments, synonyms, revised nomenclatural, chorological and ecological data supplementary to the Greek
checklist (Dimopoulos & al. 2013), as far as single items
were helpfully communicated to the team of compilers by
courtesy of a community of attentive users of the book. A
number of species, admittedly exhibiting morphological
variation in different parts of their total range, are now
more precisely allocated to subspecies. As a result of this,
we offer a supplementary quantitative analysis of the vascular flora of Greece, focusing on overall taxonomic diversity of the Greek vascular plants, taxonomic diversity
across the floristic regions of Greece, and endemic and
range-restricted plant diversity in Greece.
A publication of this kind is “a living thing”, and
Costas Thanos, as President of the Hellenic Botanical
Society and head of one of the editing institutions of the
Greek checklist, stated that “certainly, it will have to be
amended” (Dimopoulos & al. 2013: 11). The added, revised or calculated entries are confined to the state of
taxonomic knowledge prior to 31 October 2013, when
the Checklist was sent to the printer (publication was on
26 November 2013). Taxonomic and floristic novelties
published thereafter are not considered here, but will
rather be subject to prospective editorial decision.
Material and methods
For the reader’s convenience, the supplement is presented in the layout of the printed volume of Dimopoulos &
al. (2013).
The distribution data for the vascular plants are coded
using the 13 floristic regions of Greece, as defined for
the Flora Hellenica project (Strid & Tan 1997). Of the
13 floristic regions, seven are continental: Peloponnisos
(Pe), Sterea Ellas (StE), Southern Pindos (SPi), Northern Pindos (NPi), East Central (EC), North Central (NC)
and North East (NE); and six are island regions: Ionian
Islands (IoI), West Aegean islands (including the large
island of Evvoia) (WAe), North Aegean islands (NAe),
Kiklades (Kik), Kriti and Karpathos (including satellite
islands = the Cretan area) (KK) and East Aegean islands
(EAe) (Fig. 1). The symbols for the distribution of the
plants in floristic (phytogeographical) regions are “x” for
presence, “.” for absence and “?” for doubtful presence.
In the Status column (Stat) of the Floristic catalogue,
non-native taxa (aliens, xenophytes), including cultigens,
are denoted with “X”, provided that they are permanently
established somewhere in the country (Dimopoulos & al.
2013). The origin of alien taxa is given in square brackets
“[ ]” in the chorology column (Ch). Native taxa, applying
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Dimopoulos & al.: Vascular plants of Greece: Supplement
the criteria taken from Med-Checklist (see, e.g., Greuter
& Raab-Straube 2008: xi), are not specifically annotated
in the status column of the Floristic catalogue.
The symbol “r” in the Status column (Stat) of the
Floristic catalogue denotes range-restricted taxa, which
are characterized by a restricted distribution and populations occurring along a linear distance not exceeding 500
km, no matter whether the political borders of Greece are
crossed (Dimopoulos & al. 2013). In contrast to endemic
taxa, range-restricted taxa may well be shared by two,
three or more countries. The assignment of any Greek
taxon to the range-restricted category requires good
knowledge of its overall distribution. For the estimation
of linear distance we used Google Earth and its tools
(https://www.google.com/earth/). This distance is not affected by topography, altitude, habitats, bodies of fresh or
sea water, or political borders.
In the Chorology column (Ch) of the Floristic catalogue, the chorological type of each taxon is denoted on
the basis of a new “Greece-centred” system of chorological categories/types, which was established in Dimopoulos & al. (2013) to better reflect and circumscribe
the distribution ranges of the taxa of the Greek vascular
flora, given that Greece is a country of S Europe, of the
Balkan Peninsula and of the Mediterranean basin. Based
on this system, the Greek vascular flora can be assigned
to 21 chorological categories distinguished for native
taxa, and to one group of various chorological categories representing different origins of alien taxa. Descriptors and abbreviations for each chorological category are
given in Dimopoulos & al. (2013: 24 – 25). One of the
chorological categories is Greek endemics, annotated
with a bullet point “” in the chorology column (Ch) of
the Floristic catalogue. The term “Greek endemic” denotes vascular plant taxa with a distribution restricted
to the territory of Greece, i.e. occurring in any or all of
the 13 floristic regions of Greece but not known to occur
outside of Greece.
In the Life-form column (Lf) of the Floristic catalogue, the life-form categories for the terrestrial and
aquatic (hydrophytes) vascular plants of the Greek flora
are coded according to the life-form system of Raunkiaer (1934) and subsequent extensions to Raunkiaer’s
system by Ellenberg & Mueller-Dombois (1967). The
descriptor and the abbreviation for each category are
provided in Dimopoulos & al. (2013: 25 – 26) and are
summarized as follows (categories and their abbreviations in brackets): phanerophytes (P), chamaephytes
(C), hemicryptophytes (H), geophytes (G), therophytes
(T) and aquatics (A).
For the habitat analysis of the total vascular flora
of Greece, eight groups (categories) of habitats were
distinguished; the descriptor and the abbreviation for
each category are provided in Dimopoulos & al. (2013:
26 – 27) and are summarized as follows (categories and
their abbreviations in brackets): freshwater habitats (A),
cliffs, rocks, walls, ravines, boulders (C), submediter-
Willdenowia 46 – 2016
303
ranean/temperate grasslands (G), high-mountain vegetation (H), coastal habitats (M), xeric Mediterranean
phrygana and associated annual-rich grasslands (P), agricultural and ruderal habitats (R), woodlands and scrub
(W).
In the Habitat column (Hab) of the Floristic catalogue,
the habitat or habitats that a taxon prefers are given using the mentioned abbreviations. The range of habitats that
a species occupies falls mostly into one habitat category,
but may comprise two or more categories. Generally a category is given only when it corresponds to a considerable
proportion of the populations of the respective species. If
more than one category applies, the two or more abbreviations are arranged in alphabetical order. The order of habitat symbols does not express prevalence. If one out of two
or more habitat categories clearly prevails – i.e. representing at least about two-thirds of all known populations – the
respective habitat abbreviation is underlined. The degree
of uncertainty in allocating habitats to plant taxa in Greece
is high in many cases, even with only eight coarse categories adopted. Many species have been seen in the wild
by few persons, and some by one or no living person. For
many taxa, hardly any useful, or no, ecological or habitat
statements are available in literature. Habitat descriptions
in taxonomic studies or on herbarium specimen labels are
often short or misleading, or comprehensive works attempt
to enumerate all possible habitat conditions under which a
species might be encountered. In any of these cases the allocation to a predominant habitat category is made difficult
(Dimopoulos & al. 2013).
Results
Supplementary quantitative analysis of the vascular
flora of Greece
In order to analyse the vascular flora of Greece, we used
the following definitions and derived rules for calculations at different taxonomic ranks (counting families,
genera, species, subspecies and taxa).
Species are defined as comprising (1) species that
have no subspecies and (2) species that have one or more
than one subspecies. Subspecies are defined as comprising all subspecies given for Greece, no matter how many
per species. Taxa are defined as comprising (1) subspecies and (2) species that have no subspecies, i.e. when
a species has subspecies then only its subspecies are
counted. Hence, in the case of a species with no subspecies we have one taxon; in the case of a species with one
subspecies in Greece we have one taxon, not two; and
in the case of a species with two or more subspecies in
Greece, then we have two or more taxa.
Overall taxonomic diversity of the vascular flora of
Greece
The vascular flora of Greece comprises 5758 species and
1970 subspecies (native and naturalized), representing
6620 taxa, belonging to 1073 genera and 185 families
(Table 1). The full dataset has a total of 7739 records,
comprising species and subspecies plus ten sections of
Taraxacum and one aggregate (Portulaca oleracea aggr.).
The only species for which the status of
Table 1. Numbers of plant families, genera, species, subspecies and taxa in the three “native but extinct” was confirmed are
main taxonomic groups of the Greek vascular flora.
Staphylea pinnata (NE) and Stratiotes
aloides (NC).
Taxonomic group Families
Genera
Species Subspecies
Taxa
Pteridophytes
Gymnosperms
Angiosperms
16
4
165
29
7
1037
73
25
5660
18
9
1943
80
28
6512
Total
185
1073
5758
1970
6620
Table 2. Numbers (in descending order from the most taxon-rich to the least taxonrich region) of vascular plant families, genera, species, subspecies and taxa in each
of the 13 floristic regions of Greece.
Floristic region
NE
NC
StE
Pe
SPi
NPi
EAe
KK
EC
WAe
IoI
NAe
Kik
Families
Genera
Species
Subspecies
Taxa
164
158
160
159
155
146
151
146
144
146
146
145
136
871
823
860
856
798
744
756
703
713
695
698
678
619
3264
3116
3116
2971
2666
2572
2381
2079
2086
2024
1949
1928
1661
1054
1015
977
919
820
835
660
571
567
582
526
494
458
3531
3356
3318
3171
2793
2715
2520
2214
2102
2084
2003
1932
1750
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Taxonomic diversity across the floristic regions of Greece
When comparing the different floristic
regions of Greece (Table 2), we find that
the most species-rich and taxon-rich region is NE (3264 species, 3531 taxa),
followed by NC, StE and Pe, whereas
the most species- and taxon-poor region
is Kik (1661 species, 1750 taxa) (Table
2, Fig. 1, abbreviations explained in the
latter).
Generally we observe that the
mainland regions of Greece are more
species- and taxon-rich than the island
regions, a trend that might reflect the
different amount of land surface of each
region. The exception is EC, which has
an intermediate position among the
most species- and taxon-poor island
regions of Greece, after EAe and KK
and before WAe, IoI, NAe and Kik, in
304
Dimopoulos & al.: Vascular plants of Greece: Supplement
North East (NE)
3264
North
Central (NC)
3116
North Aegean
Islands (NAe)
1928
Northern
Pindos (NPi)
2572
Southern
Pindos (SPi)
2666
Ionian Islands
(IoI) 1949
East
Central (EC)
2086
Sterea Ellas (StE)
3116
West Aegean
Islands (WAe)
2024
East Aegean Islands
(EAe) 2381
Peloponnisos (Pe)
2971
Kiklades (Kik)
1661
Kriti and Karpathos (KK)
2079
Fig. 1. Vascular plant species in each of the 13 floristic regions of Greece.
descending order. The floristic regions of NPi, KK, WAe
and IoI each have the same number of families (146 families) (Table 2).
Endemic plant diversity within the vascular flora of
Greece
The endemic vascular flora of Greece comprises 1459
taxa (22 % of the total number of taxa in Greece), corresponding to 1274 endemic species (22.1 % of the total
number of Greek species) and 450 endemic subspecies
(22.8 % of the total number of Greek subspecies) (Table
3).
The endemic richness in absolute numbers and the
rate of endemism are not uniformly distributed across the
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floristic regions; as a general pattern S Greece (Pe, KK,
StE) and E Greece are richer in absolute numbers of endemics (Table 3). The highest number of Greek endemic
species and taxa is observed in Pe (464 taxa), while the
second and the third highest numbers are in the regions
KK (392 taxa) and StE (368 taxa). The lowest numbers
are in the regions NAe (57 taxa), IoI (91 taxa) and EC (96
taxa) (Table 3).
While KK is second highest among the floristic regions in its absolute number of Greek endemic taxa, its
endemism rate is the highest (21.1 % for subspecies,
17.1 % for species and 17.7 % for taxa), followed by Pe
(16.4 % for subspecies, 14.4 % for species and 14.6 % for
taxa). The ranking of the regions according to their total
vascular plant diversity is quite similar across taxonomic
Willdenowia 46 – 2016
305
Table 3. Greek endemic species, subspecies and taxa (absolute
numbers) for each of the 13 floristic regions and for Greece as a
whole.
Floristic region
Numbers of endemic
species
subspecies
taxa
Pe
KK
StE
WAe
NC
Kik
SPi
EAe
NPi
NE
EC
IoI
NAe
427
356
340
194
181
146
149
147
144
122
95
86
57
151
120
130
65
71
55
53
46
46
49
36
34
19
464
392
368
202
193
162
156
152
146
129
96
91
57
Greece
1274
450
1459
levels (families, genera, species and taxa; Table 2), but is
very different to the ranking according to the diversity of
endemic species and taxa (Table 3).
Table 4. Range-restricted species, subspecies and taxa (absolute
numbers) for each of the 13 floristic regions and for Greece as
a whole.
Floristic region
Pe
StE
NC
KK
NE
NPi
SPi
EAe
WAe
Kik
EC
IoI
NAe
Greece
Numbers of range-restricted
species
subspecies
taxa
466
429
386
352
323
306
269
253
211
154
149
99
82
173
166
147
120
116
115
100
73
71
56
51
40
27
505
461
414
388
344
319
278
259
217
169
150
107
82
1703
611
1972
the phytogeography of the Greek flora, although they
offer important information on the local character, the
uniqueness and relations of a flora. From the evaluation
of the range-restricted taxa in the vascular flora of Greece
a slightly different picture compared to the endemic taxa
Range-restricted plant diversity within the vascular flois obtained.
ra of Greece
With the current knowledge, the range-restricted vasMost floristic inventories or publications on the phyto- cular flora of Greece consists of 1972 taxa (29.8 % of the
geography of Greece recognize and analyse the endem- total number of taxa in Greece), corresponding to 1703
ic plants of Greece. However, until Dimopoulos & al. species accounting for 29.6 % of the total number of spe(2013), the range-restricted taxa of Greece have rarely cies, and 611 subspecies (31 % of the total number of subbeen mentioned, and never evaluated, in publications on species in Greece (Table 4). Range-restricted species and
taxa, similar to the endemics,
are not uniformly distributed
across the floristic regions
(Table 4). The region Pe is
again the richest floristic region in Greece (505 taxa),
now followed by StE (461
taxa) and NC (414 taxa).
The N Greek floristic
regions, including NE (344
taxa) and NPi (319 taxa),
with their considerable proportions of cross-border endemics, rank much higher
among the range-restricted
taxa than among the withinGreece endemics. The regions NAe and IoI are the
poorest both in range-restricted as in endemic taxa.
Overall, the range-restricted
taxa that are not also Greek
Fig. 2. Rates of vascular plant endemism and range-restrictedness in each of the 13 floristic endemics are mainly located
regions of Greece. The rates are calculated as the percentage of taxa within a floristic region that on the Greek mainland and
are endemic and range-restricted to Greece.
especially in mountain areas.
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306
If we compare the different floristic regions of Greece
taking into account their total plant diversity, i.e. analysing the diversity of range-restricted taxa as a proportion
of the total flora, then KK is the richest in range-restricted taxa with 17.5 %, followed by Pe with 15.9 % (Fig. 2).
Comparing the patterns exhibited when endemism
and range-restrictedness rates across the floristic regions
of Greece are taken into account, the trend is partly similar and partly different: (1) a high endemism rate is combined with a high rate of range-restricted taxa, decreasing
from 17.7 % and 17.5 %, respectively, in KK to 9.2 % and
9.6 % in KiK, through Pe, StE and WAe with intermediate values; (2) the rates of range-restricted taxa exceed
considerably the respective endemism rates, as for NC,
NPi, EAe, SPi and NE (12.3 %, 11.7 %, 10.3 %, 10 %
and 9.7 % range-restricted taxa, respectively) towards the
lowest rate, in NAe (4.2 %) (Fig. 2).
Habitat preferences of Greek plant taxa
Greece is well-known as a country of islands and mountains, but coastal and high-mountain plants together
comprise about 17.2 % of the Greek flora (Fig. 3). Our
evaluation on the habitat preferences of plant taxa reveals
that Greece is in fact rather a country of cultural, i.e. anthropozoogenic, landscapes. Most common are plants of
agricultural and ruderal habitats (18.1 %), followed by
plants of grasslands and dwarf shrublands, with 17.7 %
representing submediterranean/temperate lowland to
montane pastures and meadows, and 15.4 % Mediterranean annual-rich grasslands and phrygana. Plants of
woodlands and shrublands represent only 13.7 %, although these formations are very diverse and widespread
in Greece, and almost all tree and shrub species belong
here. Specialist plants of high mountains (12.6 %), cliffs
(9.0 %), freshwater (8.9 %) and coastal habitats (4.6 %)
are represented by minor proportions but, considering the
small areas occupied by each of these habitat categories,
Dimopoulos & al.: Vascular plants of Greece: Supplement
Table 5. Habitat categories represented among all taxa, endemic
taxa and range-restricted taxa of Greece. Abbreviations of categories as in Fig. 3.
Habitat
category
A
C
G
H
M
P
R
W
All taxa
[%]
Endemic taxa
[%]
Range-restricted
taxa [%]
8.9
9.0
17.7
12.6
4.6
15.4
18.1
13.7
1.7
22.9
15.6
19.9
6.4
19.3
4.7
9.5
2.4
20.6
18.5
21.2
5.0
17.7
5.1
9.6
their floras are remarkably prominent in the Greek vegetation.
Focusing on endemic and range-restricted taxa, the
evaluation reveals that terrestrial habitat categories with a
high proportion of natural and semi-natural open habitats
prevail. A total of about 78 % of all endemic and rangerestricted taxa are associated with cliff, high-altitude,
xeric Mediterranean and submediterranean grassland
habitats (Table 5). Diversity of paleo- and neo-endemics,
as expressed by the number and proportion of endemic
and range-restricted taxa, tends to be most pronounced in
habitat categories with suitable sites that are more or less
isolated, such as cliffs, high mountains and xeric rocky
habitats on islands and peninsulas.
Coastal habitats are represented among endemic
and range-restricted taxa by a similar proportion to that
among all taxa, and woodlands only slightly less. In contrast, ruderal and freshwater habitats are much poorer in
endemic and range-restricted plant taxa than their proportion among the entire flora would suggest. The latter habitat categories occur widespread and with similar
ecologies throughout the Mediterranean and beyond, and
they accommodate chiefly widespread species, with a
higher proportion of non-native taxa than other habitat
categories.
Acknowledgements
Fig. 3. Habitat categories represented in the total vascular flora
of Greece. – A: freshwater; C: cliff; G: submediterranean grassland; H: high mountain; M: coastal/marine; P: Mediterranean
grassland and phrygana; R: ruderal and agricultural; W: woodland (see also column-related explanatory notes above). The
numbers refer to all single and multiple habitat categories assigned to plant taxa.
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The team of compilers gratefully acknowledges critical
input to the present supplement by the following: Ioannis Bazos (Athens), Erwin Bergmeier (Göttingen), Karl
Peter Buttler (Frankfurt/Main), the late Lance Chilton
(Hunstanton), Michael Damanakis (Iraklion), Manfred
A. Fischer (Vienna), Paul Fontaine (Brussels), Christina
Fournaraki (Chania), Günter Gottschlich (Tübingen),
Werner Greuter (Berlin/Palermo), Jaime Guemes (Valencia), Ralf Hand (Berlin), Per Hartvig (Copenhagen),
Gregoris Iatrou (Patras), Ralf Jahn (Großschirma), Stella
Kokkini (Thessaloniki), Katerina Koutsovoulou (Athens), Per Lassen (Lund), Magnus Lidén (Uppsala), Sabine
von Mering (Berlin), Maria Panitsa (Patras), Eckhard von
Raab-Straube (Berlin), Uwe Raabe (Recklinghausen),
Willdenowia 46 – 2016
Thomas Raus (Berlin), Federico Selvi (Firenze), the late
Franz Speta (Linz), Arne Strid (Ørbæk), Kit Tan (Copenhagen), Costas Thanos (Athens), Nicholas Turland (Berlin), Holger Uhlich (Dresden), Robert Ulrich (Tübingen),
Eckhard Willing (Dessau) and Aris Zografidis (Athens).
The authors also wish to thank the two reviewers, Ralf
Jahn and Maria Panitsa, for their constructive comments
and suggestions on an earlier draft of this paper. A special credit is due to Eckhard and Rita Willing (Dessau),
whose abundant and unselfish original registration entries to the Flora Hellenica Database amount to more
than 150 000 records, thus helping to make the picture
of the taxonomic diversity across the floristic regions of
Greece as precise as possible.
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Willdenowia 46 – 2016
311
Floristic catalogue. Supplement
pallidum, Hypericum hyssopifolium, Isoetes echinospora, I. sicula, Juniperus phoenicea, Leontodon crispus
subsp. asper, L. crispus subsp. crispus, L. crispus subsp.
rossianus, Limonium rhodense, Matricaria chamomilla,
Medicago blancheana subsp. blancheana, M. blancheana
subsp. bonarotiana, Myosurus heldreichii, Ophrys cretica
subsp. ariadnae, O. fuciflora, O. fuciflora subsp. andria,
O. fuciflora subsp. candica, O. fuciflora subsp. fuciflora,
Ornithogalum umbellatum, Phelipanche nana, P. ramosa,
Pilosella alpicola, Plantago macrorrhiza, Polygonatum
latifolium, Pseudorchis albida subsp. albida, Schenkia
spicata subsp. spicata, Spergularia salina, Tragopogon
longirostris, Verbascum glandulosum and Viola tricolor.
The arrow symbol “►” refers to a comment under the
same name in Appendix III.
Ch
Lf
Hab
EM
MA
Me
Ct
H
H
H
H
CH
C
C
C
.
x
x
x
x
x
x
x
x
.
.
.
.
Bo
H
C
.
.
.
.
.
.
x
.
.
.
.
.
.
x
.
.
.
.
.
.
.
.
.
x
.
.
Eu
Eu
H
H
C
C
DENNSTAEDTIACEAE
Pteridium aquilinum (L.) Kuhn in Kersten
subsp. brevipes (Tausch) E. Wulff
.
.
.
.
.
.
.
.
.
.
.
x
?
ME
G
W
ISOETACEAE
Isoetes gymnocarpa (Gennari) A. Braun ►
.
.
.
x
.
.
.
.
.
.
x
.
x
Me
A
A
POLYPODIACEAE
Polypodium cambricum L.
subsp. cambricum
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
MA
Me
G
G
CW
CW
THELYPTERIDACEAE
Thelypteris palustris Schott ►
x
x
x
x
x
.
x
x
x
.
.
.
.
Co
G
A
Ch
Lf
Hab
Me
P
W
.
x
x
x
x
x
x
x
x
x
.
x
x
PW
Hab
W
P
Lf
P
MA
Ch
EM
x
Stat
x
x
EAe
x
x
KK
.
x
Kik
x
x
WAe
x
.
NE
x
.
NAe
x
x
EC
x
x
NC
x
x
StE
x
x
Pe
x
.
SPi
x
IoI
.
x
NPi
CUPRESSACEAE
Juniperus oxycedrus L. ►
subsp. deltoides (R.P. Adams) N.G. Passal. in Bernardo,
Passalacqua & Peruzzi
Juniperus turbinata Guss. ►
Stat
GYMNOSPERMS
Stat
KK
EAe
.
x
x
x
EAe
Kik
x
x
x
x
KK
WAe
.
x
x
x
Kik
.
x
x
x
WAe
NE
NAe
.
x
x
x
NE
.
x
x
x
NAe
EC
NC
.
x
x
x
EC
.
.
.
x
NC
Pe
StE
?
.
.
x
StE
SPi
.
x
x
x
Pe
.
.
.
x
SPi
IoI
.
.
.
x
ASPLENIACEAE
Asplenium aegaeum Lovis, Reichst. & Greuter in Reichst. & al.
Asplenium obovatum Viv.
subsp. obovatum
Asplenium ruta-muraria L.
Asplenium septentrionale (L.) Hoffm.
subsp. septentrionale
Asplenium trichomanes L.
subsp. hastatum (H. Christ) S. Jess.
subsp. pachyrachis (H. Christ) Lovis & Reichst. in Greuter
IoI
.
.
.
x
PTERIDOPHYTES
NPi
NPi
The entries in the Floristic catalogue presented here
are additional to or replace entries in Dimopoulos & al.
(2013); the replaced entries are hence obsolete or redundant and are to be moved as synonyms to Appendix II
(q.v.), viz. Aira elegantissima, Ajuga orientalis subsp. aenesia, A. orientalis subsp. orientalis, Ammophila arenaria
subsp. arundinacea, Ballota nigra subsp. uncinata, Carex
pairae, C. sempervirens, C. tomentosa, Centaurea ptarmicoides, Centaurium erythraea subsp. grandiflorum,
Cerastium holosteoides subsp. vulgare, Epipactis persica
subsp. exilis, Euphorbia villosa, Gagea saxatilis, Galium
recurvum, Goniolimon dalmaticum, Helosciadium repens,
Hieracium parnassi subsp. versutum, H. schmidtii subsp.
ACANTHACEAE
Acanthus mollis L.
subsp. mollis
x
.
.
x
x
x
.
.
x
.
.
x
x
X
[W-Med.]
H
R
AGAVACEAE
Agave americana L.
subsp. americana
x
.
.
x
x
.
.
x
x
x
x
x
x
X
[N-Am.]
P
R
ALISMATACEAE
Baldellia ranunculoides (L.) Parl.
subsp. ranunculoides
x
.
x
x
.
.
x
x
.
.
.
.
x
MA
A
A
ALLIACEAE
Allium orestis Kalpoutz., Trigas & Constantin.
Allium ritsi Iatroú & Tzanoud. ►
Allium scorodoprasum L.
.
.
.
.
.
.
.
.
.
x
x
.
.
.
.
.
.
.
.
.
x
.
.
.
.
.
.
x
.
.
.
.
.
.
.
.
.
.
.
Eu
G
G
G
W
P
R
ANGIOSPERMS
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r
r
EC
NC
NE
NAe
WAe
Kik
KK
EAe
Ch
Lf
Hab
x
x
.
x
.
x
x
x
.
x
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
EM
EM
G
G
W
W
.
.
.
.
.
.
.
x
.
.
.
.
x
.
.
.
.
.
.
.
.
x
.
x
.
.
Eu
MS
H
T
W
R
x
.
x
x
x
.
x
x
.
.
x
x
x
ME
TH
R
x
x
x
.
x
x
?
.
.
x
x
?
.
.
x
x
x
x
.
x
x
x
x
.
x
x
.
.
.
x
x
.
.
x
x
x
.
.
.
x
x
.
.
.
.
x
.
.
.
.
x
.
.
.
.
x
.
.
.
.
x
x
x
.
.
Me
Me
Me
Bk
BI
T
H
H
H
G
P
P
P
W
W
x
x
x
.
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
x
x
.
.
x
.
x
x
.
.
x
.
x
Me
EA
Me
H
T
T
R
R
R
.
.
.
.
.
.
.
.
x
.
.
.
.
MS
H
M
x
x
x
x
x
x
x
x
x
x
x
.
x
ME
G
GPW
G
PW
Stat
Pe
ARACEAE
Arum italicum Mill.
subsp. italicum
Biarum tenuifolium (L.) Schott in Schott & Endl.
subsp. idomenaeum P.C. Boyce & Athanasiou
StE
APOCYNACEAE
Trachomitum sarmatiense Woodson
SPi
APIACEAE
Anthriscus nitidus (Wahlenb.) Hazsl. ►
Bupleurum lancifolium Hornem. ►
Daucus carota L
subsp. major (Vis.) Arcang.
Daucus guttatus Sm. in Sibth. & Sm.
subsp. guttatus
Elaeoselinum asclepium (L.) Bertol.
subsp. asclepium
Eryngium palmatum Pančić & Vis. ►
Geocaryum capillifolium (Guss.) Coss.
Smyrnium perfoliatum L.
subsp. rotundifolium (Mill.) Bonnier & Layens ►
Tordylium maximum L.
Torilis pseudonodosa Bianca
IoI
AMARYLLIDACEAE
Galanthus reginae-olgae Orph.
subsp. reginae-olgae
subsp. vernalis Kamari
Dimopoulos & al.: Vascular plants of Greece: Supplement
NPi
312
.
.
.
.
.
.
.
.
.
.
.
x
.
ARALIACEAE
Hedera helix L.
subsp. helix
x
x
x
x
x
x
x
x
x
x
x
x
x
ME
P
W
ASPARAGACEAE
Asparagus acutifolius L. ►
Asparagus officinalis L.
subsp. officinalis
x
x
x
x
.
.
x
.
.
x
x
x
x
.
.
x
x
x
x
x
x
x
x
x
x
.
.
x
.
.
x
.
.
.
.
.
x
.
.
Me
EA
EA
C
C
C
W
GR
GR
x
x
x
.
x
x
x
.
x
x
.
.
x
.
.
x
x
x
.
x
x
x
.
x
x
x
.
?
x
x
.
x
.
.
.
x
.
.
.
.
.
.
.
.
.
.
.
x
.
.
.
x
BI
ES
[E-As.]
EM
T
C
H
T
R
GH
R
GP
.
.
x
.
?
x
?
?
x
.
.
.
.
EA
H
GR
.
x
.
x
x
.
x
x
.
.
.
.
.
Eu
H
G
.
.
.
x
x
x
.
.
.
.
.
.
.
Bk
H
GH
.
x
x
.
.
.
x
.
.
.
.
.
.
BI
H
G
.
.
x
x
x
.
.
.
x
x
.
x
x
.
.
x
.
.
.
.
.
.
.
?
x
.
.
x
.
.
.
.
.
.
.
?
.
.
x
BA
Me
H
H
T
GH
H
R
.
x
x
x
x
.
x
.
.
.
.
x
x
.
.
x
x
x
x
x
x
x
.
x
x
x
x
.
.
.
?
x
.
.
.
?
x
.
.
.
?
x
.
.
.
x
x
.
x
x
x
.
x
x
x
x
.
x
x
x
x
.
x
x
x
EM
Me
Me
EM
EM
CH
CH
H
T
T
R
R
R
MP
P
.
x
x
.
x
.
x
.
.
.
.
.
.
BI
H
H
.
.
.
.
.
x
.
.
.
.
.
.
x
.
.
.
.
.
.
.
.
.
.
.
.
.
Eu
Eu
H
H
CW
CW
x
.
.
.
.
.
.
x
.
.
.
.
x
x
.
.
.
.
.
x
x
.
.
.
.
.
x
x
.
.
.
.
.
x
x
.
x
.
.
.
.
.
.
.
x
x
.
x
x
x
.
.
.
x
.
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.
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.
.
.
.
ME
ME
EA
Eu
Bk
BA
Bk
H
H
H
H
H
H
H
W
W
W
W
W
CHW
CHW
.
.
.
.
x
.
.
.
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.
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.
x
.
x
.
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.
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.
.
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.
.
.
.
.
.
.
.
Bk
Eu
H
H
H
W
W
W
ASTERACEAE
Anthemis segetalis Ten.
Artemisia absinthium L.
Artemisia verlotiorum Lamotte
Carduus argentatus L.
Carduus hamulosus Ehrh.
subsp. hamulosus
Carlina biebersteinii Hornem.
subsp. brevibracteata (Andrae) K. Werner
Carlina frigida Boiss. & Heldr. in Boiss.
subsp. frigida
Centaurea jacea L.
subsp. weldeniana (Rchb.) Greuter
Centaurea pichleri Boiss.
subsp. pichleri
Centaurea ptarmicifolia Halácsy ex Hayek
Cladanthus mixtus (L.) Chevall. ►
Dittrichia viscosa (L.) Greuter
subsp. angustifolia (Bég.) Greuter
subsp. viscosa
Echinops spinosissimus Turra
Filago aegaea Wagenitz
subsp. aristata Wagenitz
Gnaphalium hoppeanum W.D.J. Koch
subsp. magellense (Fiori) Strid in Strid & Tan
Hieracium bifidum Kit. ex Hornem.
subsp. psammogenes (Zahn) Zahn
subsp. pseudopraecox (Zahn) Zahn
Hieracium brevifolium Tausch
subsp. brevifolium
subsp. muraltae (Zahn) Greuter
Hieracium diaphanoides Lindeb
subsp. pseudoumbrosum Zahn in Engl
Hieracium krischtimanum Mattf. & Zahn in Zahn ►
Hieracium lazistanum Arv.-Touv.
subsp. leithneri (Boiss.) Greuter in Greuter & Raab-Straube
Hieracium murorum L.
subsp. platydiaphanoides Rech. f. & Zahn
subsp. ravnanum K. Malý & Zahn
subsp. subbifidiforme Zahn
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Terms of Use: https://bioone.org/terms-of-use
r
X
r
.
r
r
r
BIEBERSTEINIACEAE
Biebersteinia orphanidis Boiss.
BORAGINACEAE
Buglossoides arvensis (L.) I.M. Johnst.
subsp. sibthorpiana (Griseb.) R. Fern. ►
Cynoglossum officinale L. ►
subsp. officinale
Myosotis discolor Pers. in Murray
subsp. discolor
Myosotis ramosissima Rochel in Schult.
subsp. gracillima (Loscos & J. Pardo) Rivas Mart.
Symphytum circinale Runemark
BRASSICACEAE
Alyssum strigosum Banks & Sol. in Russell
subsp. strigosum
Erysimum corinthium (Boiss.) Wettst.
Fibigia clypeata (L.) Medik.
subsp. clypeata ►
Lepidium draba L.
subsp. draba
Lepidium sativum L.
subsp. sativum
Lunaria annua L.
subsp. pachyrhiza (Borbás) Maire & Petitm. ►
Noccaea versicolor (Stoj . & Kitanov) F.K. Mey.
Rorippa sylvestris (L.) Besser ►
CAMPANULACEAE
Campanula drabifolia Sm. in Sibth. & Sm.
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NE
NAe
WAe
Kik
KK
EAe
.
.
.
x
.
x
x
.
.
x
.
.
.
x
x
x
x
.
.
x
.
.
.
x
.
.
.
.
.
x
.
.
.
x
x
x
x
x
.
.
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.
x
x
.
x
x
x
.
.
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.
x
x
.
x
.
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x
x
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x
.
x
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x
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.
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x
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x
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x
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x
.
.
.
x
.
.
.
x
.
.
r
.
.
.
.
.
.
r
Hab
EC
NC
.
x
.
x
x
x
x
.
.
x
Lf
Pe
StE
x
.
x
x
.
x
x
.
.
x
Ch
SPi
.
.
.
.
.
.
.
.
.
.
Stat
IoI
Hieracium murorum [continued]
subsp. subnemorense (Zahn) Zahn
subsp. sylvularum (Boreau) Zahn
subsp. viridicollum (Boreau) Zahn
Hieracium pannosum Boiss.
subsp. friwaldii (Rchb. f.) Freyn ►
Hieracium parnassi Fr.
Hieracium sericophyllum Nejčeff & Zahn in Zahn
subsp. sericophylloides Hayek
subsp. sericophyllum
Hieracium sermenikense Freyn & Sint. ►
subsp. sermenikense
Hieracium transiens (Freyn) Freyn
Hyoseris lucida L.
Hyoseris radiata L. ►
Inula bifrons (L.) L.
Jacobaea erucifolia (L.) G. Gaertn., B. Mey. & Scherb.
subsp. erucifolia
Jacobaea maritima (L.) Pelser & Meijden
subsp. maritima
Jacobaea vulgaris Gaertn. ►
Leontodon biscutellifolius DC.
Leontodon crispus Vill. ►
Leontodon saxatilis Lam.
subsp. rothii Maire in Jahand. & Maire
subsp. saxatilis
Matricaria discoidea DC.
Matricaria recutita L.
Onopordum myriacanthum Boiss.
Pilosella acutifolia (Vill.) Arv.-Touv.
Pilosella bauhini (Schult.) Arv.-Touv.
subsp. magyarica (Peter) S. Bräut. in Greuter & Raus
Pilosella cymosa (L.) F.W. Schultz & Sch. Bip.
subsp. cymosa
subsp. sabina (Sebast.) H.P. Fuchs
Pilosella rhodopaea (Griseb.) Szeląg
Pilosella sphaerocephala (Rchb.) F.W. Schultz & Sch. Bip.
Senecio vulgaris L.
subsp. denticulatus (O.F. Müll.) P.D. Sell
subsp. vulgaris
Taraxacum haussknechtii R. Uechtr. ex Hausskn.
Taraxacum poliochlorum Dahlst.
Tragopogon coelesyriacus Boiss. ►
Willemetia stipitata (Jacq.) Dalla Torre in Sonklar & al.
subsp. albanica (Kümmerle & Jáv.) Kirschnerová
313
NPi
Willdenowia 46 – 2016
Eu
EA
Eu
BA
Bk
BA
Bk
Bk
H
H
H
H
H
H
H
H
H
H
W
W
W
CHW
CH
CH
CG
CG
CG
CG
BA
Me
Me
Me
H
H
H
H
W
M
P
W
.
x
x
.
x
.
x
x
.
.
.
.
.
ES
H
R
x
.
x
x
x
.
.
x
.
x
x
x
x
Me
C
CM R
.
.
x
x
x
x
.
x
x
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.
x
x
x
x
x
.
.
x
x
.
.
x
x
x
x
.
x
.
x
x
.
.
.
x
x
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x
x
.
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x
x
.
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x
x
.
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x
x
x
.
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.
x
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x
x
x
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x
x
x
.
x
x
x
x
.
x
x
x
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x
x
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x
x
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x
x
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x
x
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x
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x
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.
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.
.
.
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.
?
.
?
.
x
.
.
Me
EA
EA
Me
ME
Me
Eu
[N-Am.]
Co
BA
EA
C
H
H
H
HT
HT
HT
T
T
H
H
CMR
GR
GH
GH
R
R
R
R
R
R
W
x
x
.
x
.
.
x
x
.
x
x
.
x
x
x
x
.
.
x
x
.
x
.
x
x
x
.
x
.
.
.
x
.
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.
.
x
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.
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.
x
x
x
x
.
?
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x
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x
.
x
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.
x
.
.
.
.
.
EA
ES
Eu
EA
Bk
Eu
H
H
H
H
H
H
GW
GHW
GW
GHW
H
G
.
x
.
.
.
.
x
x
.
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x
x
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x
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x
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x
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x
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x
x
x
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x
x
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x
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x
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x
.
x
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x
.
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x
.
.
x
x
x
.
.
x
MA
Pt
EM
Me
EM
T
T
H
H
H
R
R
G
H
GR
.
x
.
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Bk
H
H
.
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.
x
.
.
.
.
.
.
.
.
.
EM
H
G
.
.
.
x
?
?
.
.
.
x
?
?
x
?
?
.
.
.
x
x
x
x
x
x
x
.
.
.
.
.
x
.
.
x
.
.
x
.
.
EA
ES
ES
T
H
H
GPR
R
R
x
x
.
.
.
.
x
x
x
x
x
.
x
EA
T
G
.
.
.
.
x
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
x
.
x
x
r
MA
EM
T
H
GP
C
x
x
x
x
x
.
x
x
.
x
x
x
x
x
x
x
x
.
x
x
.
x
x
.
x
x
.
x
x
.
.
.
.
x
x
.
x
x
.
r
MS
MS
T
T
C
PR
PR
C
.
x
x
x
x
.
x
.
.
x
.
.
.
MS
H
CG
x
x
x
x
x
x
x
x
x
x
x
x
x
Co
GH
R
.
.
.
x
.
.
.
.
x
.
.
x
x
IT
T
R
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
.
x
x
x
x
x
.
.
.
.
.
x
.
.
.
x
.
BI
EM
EA
H
H
H
RW
G
A
x
.
.
x
x
.
.
.
.
.
.
.
.
T
P
X
r
r
Pe
StE
EC
NC
NE
NAe
WAe
Kik
KK
EAe
Ch
Lf
Hab
.
x
.
x
.
x
.
x
.
x
.
.
.
.
.
.
.
.
.
x
.
.
.
.
x
.
EM
Me
H
T
CP
R
.
x
.
x
x
.
x
x
x
.
x
x
x
ME
T
A
.
.
.
.
.
.
.
x
.
.
.
.
.
Pt
H
G
x
x
x
x
x
x
x
x
x
x
x
x
x
EA
T
GHR
.
.
x
.
x
.
x
.
.
.
x
.
.
.
x
.
.
.
x
.
.
.
x
.
x
.
x
.
.
x
x
x
.
.
x
.
.
.
x
.
.
.
x
.
.
.
x
.
.
.
x
.
Bk
Bk
EA
HC
H
T
H
GH
G
GP
G
.
.
.
.
.
.
.
x
.
.
.
.
.
Bk
H
GH
.
.
.
.
?
x
x
x
x
x
.
.
x
MA
T
G
.
x
x
.
x
.
.
x
.
x
.
.
x
x
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
Bk
H
HC
HC
CH
C
C
x
.
x
x
x
.
.
x
x
x
x
x
x
Me
T
MP
.
.
.
x
.
.
x
.
x
.
.
.
.
x
.
x
.
.
x
.
.
.
.
.
.
.
Me
Eu
T
H
R
G
.
x
x
.
x
x
x
x
x
x
x
x
x
x
x
x
.
x
x
x
.
x
.
x
.
x
Me
Pt
H
T
GW
M
x
x
x
x
x
x
x
x
x
x
x
x
x
Pt
T
R
.
.
.
.
.
.
.
.
.
.
.
.
x
EA
T
R
Stat
SPi
CARYOPHYLLACEAE
Corrigiola litoralis L.
subsp. litoralis
Dianthus superbus L.
subsp. superbus
Holosteum umbellatum L.
subsp. umbellatum
Minuartia baldaccii (Halácsy) Mattf.
subsp. baldaccii
Minuartia bosniaca (Beck) K. Malý ►
Minuartia hybrida (Vill.) Schischk. in Kom. ►
Minuartia kitanovii Panov ►
Minuartia saxifraga (Friv.) Graebn.
subsp. saxifraga
Moenchia erecta (L.) G. Gaertn., B. Mey. & Scherb.
subsp. erecta
Silene auriculata Sm. in Sibth. & Sm.
subsp. auriculata
Silene cephallenia Heldr.
subsp. cephallenia
Silene colorata Poir.
subsp. colorata
Silene muscipula L.
subsp. muscipula
Silene otites (L.) Wibel
Silene vulgaris (Moench) Garcke
subsp. bosniaca (Beck) Janch. ex Greuter, Burdet & Long
Spergularia marina (L.) Besser
Vaccaria hispanica (Mill.) Rauschert
subsp. hispanica
IoI
Campanula lyrata Lam.
subsp. lyrata
Campanula ramosissima Sm. in Sibth. & Sm.
Dimopoulos & al.: Vascular plants of Greece: Supplement
NPi
314
r
r
r
r
r
r
CHENOPODIACEAE
Atriplex oblongifolia Waldst. & Kit. ►
Camphorosma monspeliaca L.
subsp. monspeliaca
Chenopodium strictum Roth
subsp. strictum
Salicornia procumbens Sm. in Sowerby
subsp. procumbens
.
.
.
x
.
.
.
x
.
.
.
.
x
EA
T
M
.
x
x
.
.
.
.
.
.
?
.
.
x
.
.
.
.
.
.
.
.
.
x
x
x
.
.
.
.
.
.
.
.
.
.
.
.
?
?
Pt
EA
EA
T
T
T
R
M
M
CISTACEAE
Fumana laevis (Cav.) Pau ►
x
.
x
x
x
x
x
x
x
x
x
x
Me
C
P
COLCHICACEAE
Colchicum chalcedonicum Azn.
subsp. chalcedonicum
Colchicum lingulatum Boiss. & Spruner in Boiss.
subsp. lingulatum
.
.
.
.
.
.
x
.
.
.
.
.
x
x
x
x
.
.
x
.
.
x
.
.
.
.
.
x
x
x
.
.
.
.
.
.
.
.
.
EM
BA
G
G
G
G
P
P
CONVALLARIACEAE
Polygonatum hirtum (Poir.) Pursh
.
.
.
.
.
.
x
x
.
.
.
.
.
ME
G
W
x
x
x
.
x
?
x
x
x
x
x
x
x
.
x
.
x
x
x
x
x
x
x
x
x
x
MS
Me
H
C
W
CP
x
.
.
.
.
.
.
.
.
.
.
.
.
X
Me
H
P
x
x
x
.
.
.
x
x
.
x
x
.
.
.
x
.
.
.
x
.
.
.
x
.
.
x
.
.
.
.
.
.
.
.
x
.
.
x
.
X
ST
[Am.]
ST
T
H
H
P
R
A
CRASSULACEAE
Sedum acre L. ►
Sedum dasyphyllum L.
subsp. dasyphyllum
.
.
.
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
.
x
x
.
x
x
.
.
.
.
.
.
.
.
.
ES
ME
ME
C
H
H
GH
CH
CH
CUCURBITACEAE
Ecballium elaterium (L.) A. Rich. in Bory
subsp. elaterium
x
x
x
x
x
x
x
x
x
x
x
x
x
MS
G
R
CYPERACEAE
Carex atrata L.
subsp. atrata
Carex bulgarica (Domin) Lazare
Carex filiformis L.
.
.
.
.
.
.
x
x
x
x
.
x
.
.
.
.
x
x
.
?
.
.
.
.
.
.
x
x
.
.
.
x
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
AA
AA
Bk
ES
HG
HG
H
G
H
H
H
A
CONVOLVULACEAE
Calystegia silvatica (Kit.) Griseb.
subsp. silvatica
Convolvulus oleifolius Desr. in Lam.
Convolvulus sabatius Viv.
subsp. sabatius
Cuscuta approximata Bab.
subsp. macranthera (Boiss.) Feinbrun & Greuter
Dichondra micrantha Urban
Ipomoea sagittata Poir.
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Terms of Use: https://bioone.org/terms-of-use
r
r
SPi
Pe
StE
EC
NC
NE
NAe
WAe
Kik
KK
EAe
Ch
Lf
Hab
x
x
x
x
x
x
x
x
x
x
x
x
x
EA
H
GW
.
x
.
.
x
x
x
x
.
.
.
.
.
EA
G
G
.
x
x
x
x
x
x
x
x
x
x
.
x
ES
H
GW
.
x
x
.
x
.
x
.
.
.
.
.
.
EA
H
A
.
.
x
.
x
x
x
.
x
x
x
.
x
.
x
x
x
.
x
.
.
.
.
.
.
.
ES
Pt
HG
H
W
AR
x
x
x
x
x
x
x
x
x
x
x
x
x
Co
G
A
.
.
x
x
.
x
x
.
x
.
x
x
.
x
x
.
x
.
x
x
x
x
x
x
x
x
.
x
.
x
.
.
.
x
.
x
.
?
x
Co
ST
Pt
G
T
G
A
A
A
DIPSACACEAE
Knautia integrifolia (L.) Bertol.
subsp. integrifolia
x
.
.
x
x
.
.
x
.
.
.
.
.
Me
T
PR
ERICACEAE
Vaccinium vitis-idaea L.
subsp. vitis-idaea
.
.
.
.
.
.
.
x
.
.
.
.
.
Bo
C
HW
EUPHORBIACEAE
Euphorbia exigua L.
subsp. exigua
Euphorbia illirica Lam.
Euphorbia verrucosa L.
x
.
x
x
x
x
x
x
x
x
x
.
x
x
x
x
.
x
x
x
x
x
x
?
x
.
?
x
.
.
x
x
.
x
.
.
x
.
.
ME
ES
ME
T
G
C
PR
A
H
x
.
.
x
x
x
x
x
x
x
x
x
x
.
x
x
x
x
x
.
x
.
x
.
x
.
Me
Bk
T
C
P
H
x
.
x
.
x
.
.
.
.
x
x
x
.
x
.
.
.
x
x
.
x
.
x
x
x
.
.
?
.
x
x
x
x
.
x
.
.
x
x
.
.
.
.
.
x
x
x
.
x
.
x
x
x
x
.
x
.
.
?
.
.
.
x
.
.
?
.
.
.
?
.
x
.
.
.
.
.
.
x
.
.
.
.
?
.
x
x
.
.
x
.
Me
Me
MS
EA
ME
EM
EA
P
T
T
H
H
T
H
W
P
R
GH
A
R
AGH
.
.
x
.
.
.
.
x
x
.
.
x
.
x
x
.
.
x
.
x
x
x
x
x
x
x
x
.
.
x
.
x
x
.
.
x
.
x
x
.
.
x
.
x
x
x
x
x
.
.
.
x
x
x
.
.
.
x
x
x
.
.
.
x
x
.
.
.
.
.
.
?
x
.
.
x
x
x
Me
ME
Ct
Me
Me
MS
TH
T
H
T
T
T
R
G
G
PR
PR
R
x
x
x
x
x
x
.
.
x
x
.
.
x
x
x
x
x
x
x
x
x
x
.
.
x
x
x
x
x
x
x
.
x
x
x
x
x
x
x
x
x
x
x
x
x
x
?
?
x
x
x
x
MA
MA
EM
EM
T
T
T
T
R
R
R
R
x
.
.
.
.
.
x
.
.
.
.
.
x
Me
T
A
x
.
.
.
.
.
x
.
x
.
.
.
.
.
.
.
.
.
.
.
x
.
x
.
x
x
Me
EM
T
T
A
P
.
x
.
.
x
.
.
x
.
.
.
.
.
ME
H
H
x
x
.
.
x
x
x
.
.
.
x
x
.
.
.
x
x
.
.
x
x
x
.
.
x
.
.
.
.
x
.
.
.
.
.
.
.
.
.
x
.
.
.
.
.
x
.
.
.
x
.
.
x
x
.
x
x
?
?
x
x
x
.
.
x
Me
Me
Me
G
G
G
G
G
R
R
MP
MP
P
x
x
x
x
x
.
x
x
x
.
x
x
x
.
?
.
x
x
x
x
x
x
x
.
x
x
?
.
x
.
x
x
.
.
x
x
x
x
.
.
.
x
x
x
.
.
.
x
x
x
.
.
.
.
x
x
.
.
.
.
x
x
.
.
.
.
x
x
.
.
.
.
x
x
.
x
.
.
Me
ME
BI
Me
Me
G
G
G
G
G
G
GP
R
GP
GP
GP
H
Carex halleriana Asso
subsp. halleriana
Carex liparocarpos Gaudin
subsp. liparocarpos
Carex muricata L.
subsp. muricata
Carex paniculata L.
subsp. paniculata
Carex sylvatica Huds.
subsp. sylvatica
Cyperus glomeratus L.
Eleocharis palustris (L.) Roem. & Schult.
subsp. palustris
Eleocharis uniglumis (Link) Schult.
subsp. uniglumis
Fimbristylis bisumbellata (Forssk.) Bubani
Schoenoplectus lacustris (L.) Palla
FABACEAE
Astragalus pelecinus (L.) Barneby
subsp. pelecinus
Astragalus thracicus Griseb. ►
Coronilla valentina L.
subsp. valentina
Hippocrepis unisiliquosa L.
Lathyrus inconspicuus L.
Lotus corniculatus L. ►
Lotus pedunculatus Cav. ►
Medicago bonarotiana Arcang.
Trifolium hybridum L.
Trifolium resupinatum L.
subsp. suaveolens (Willd.) Ponert
Trifolium strictum L.
Vicia cracca L. ►
Vicia monantha Retz.
subsp. monantha
Vicia narbonensis L.
FUMARIACEAE
Fumaria capreolata L.
subsp. capreolata
Fumaria judaica Boiss.
subsp. judaica
GENTIANACEAE
Blackstonia acuminata (W.D.J. Koch & Ziz) Domin
subsp. aestiva (K. Malý) Zeltner
Blackstonia perfoliata (L.) Huds.
subsp. intermedia (Ten.) Zeltner
Centaurium serpentinicola Carlström
Gentianopsis ciliata (L.) Ma
subsp. ciliata
HYACINTHACEAE
Bellevalia dubia (Guss.) Rchb.
subsp. boissieri (Freyn) Feinbrun
Muscari cycladicum P.H. Davis & D.C. Stuart ►
subsp. subsessile (Bentzer) Raus
Muscari parviflorum Desf.
Ornithogalum collinum Guss.
subsp. collinum
Ornithogalum divergens Boreau ►
Ornithogalum exscapum Ten. ►
Ornithogalum gussonei Ten. ►
Ornithogalum immaculatum Speta
Ornithogalum kochii Parl.
Downloaded From: https://bioone.org/journals/Willdenowia on 18 Dec 2023
Terms of Use: https://bioone.org/terms-of-use
Stat
IoI
315
NPi
Willdenowia 46 – 2016
?X
r
r
r
NE
NAe
WAe
Kik
KK
EAe
Ch
Lf
x
?
x
x
x
x
x
x
x
x
x
x
BA
G
R
.
.
.
x
x
x
x
x
.
x
x
x
x
x
x
x
x
.
x
x
x
.
x
x
?
.
?
.
x
.
.
.
.
.
.
.
.
x
.
BA
EA
ME
G
G
G
GH
GR
GH
HYPERICACEAE
Hypericum barbatum Jacq.
Hypericum hircinum L.
subsp. majus (Aiton) N. Robson
Hypericum perforatum L.
subsp. veronense (Schrank) A. Fröhl.
x
x
x
x
x
x
x
x
x
x
.
.
.
Eu
H
GH
.
.
.
.
.
.
.
.
.
.
.
x
x
Me
CP
A
IRIDACEAE
Crocus robertianus C.D. Brickell
x
x
x
x
x
x
x
x
x
x
x
x
x
Pt
H
R
JUNCACEAE
Juncus effusus L.
subsp. effusus
Juncus heldreichianus T. Marsson ex Parl.
subsp. heldreichianus
Luzula alpinopilosa (Chaix) Breistr.
subsp. deflexa (Kožuharov) Kirschner
Luzula campestris (L.) DC. in Lam. & DC.
subsp. campestris
Luzula forsteri (Sm.) DC. in Lam. & DC. ►
Luzula multiflora (Ehrh.) Lej. ►
subsp. multiflora
.
x
x
.
x
.
.
.
.
.
.
.
.
G
W
x
x
x
x
x
x
x
x
x
x
x
x
x
Eu
HG
A
x
.
.
x
x
x
x
x
x
x
x
x
x
EM
H
AM
.
.
.
.
.
.
x
.
.
.
.
.
.
Bk
H
AH
.
x
x
.
x
x
x
.
x
x
x
?
.
x
x
.
x
x
x
?
x
x
x
.
x
x
x
.
x
x
x
x
x
x
x
.
.
x
.
.
.
x
x
.
.
x
.
.
.
x
x
.
EA
Me
Ct
Ct
H
H
H
H
GH
W
GHW
GHW
JUNCAGINACEAE
Triglochin barrelieri Loisel.
LAMIACEAE
Ajuga orientalis L. ►
Ballota nigra L.
subsp. ruderalis (Sw.) Briq.
Betonica alopecuros L.
subsp. alopecuros
Calamintha grandiflora (L.) Moench
subsp. grandiflora
Lavandula pedunculata (Mill.) Cav.
Leonurus cardiaca L.
subsp. cardiaca
Leonurus marrubiastrum L. ►
Mentha suaveolens Ehrh.
subsp. suaveolens
Origanum vulgare L. ►
subsp. hirtum (Link) A. Terracc.
subsp. viridulum (Martrin-Donos) Nyman
subsp. vulgare
Prunella grandiflora (L.) Scholler
subsp. grandiflora
Teucrium montanum L.
subsp. helianthemoides (Adamović) Baden in Strid & Tan
Ziziphora capitata L.
subsp. capitata
x
.
x
x
x
.
x
.
x
x
x
x
x
MA
G
AM
x
x
x
x
x
x
x
x
x
x
x
x
x
EM
H
RW
x
x
x
x
x
x
x
x
.
x
x
x
x
Me
H
R
.
x
x
x
x
x
x
.
.
.
.
.
.
Eu
H
H
x
.
.
.
.
x
.
x
x
x
x
.
x
x
x
x
.
x
x
.
x
.
x
x
.
x
.
.
.
.
x
.
x
x
x
x
.
x
x
x
.
.
?
?
.
x
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
x
.
.
.
ME
MS
ES
ES
ES
H
P
H
H
H
W
P
R
R
R
.
x
x
.
.
x
x
x
x
x
x
x
x
x
.
x
x
x
x
.
x
x
x
x
.
.
x
x
x
.
.
x
x
x
x
x
x
x
x
x
.
x
x
x
.
.
x
x
x
.
.
x
x
.
.
.
x
x
.
.
x
x
x
.
.
Me
EA
EA
EA
EA
H
H
H
H
H
A
GPW
GPW
GP
GW
.
.
.
.
x
.
x
x
.
.
x
.
.
x
x
.
x
x
x
x
x
x
x
.
.
x
.
.
x
x
.
.
.
.
.
.
.
x
.
EA
ME
H
C
C
G
GH
GH
LILIACEAE
Gagea bohemica (Zauschn.) Schult. & Schult. f.
Tulipa orphanidea Boiss. ex Heldr. ►
.
x
x
x
x
x
x
x
x
x
.
.
.
MS
T
GR
x
.
x
.
x
.
x
x
x
x
x
.
x
.
x
.
x
.
x
.
x
.
x
.
x
.
EA
G
G
GP
PR
MALVACEAE
Malope malacoides L.
subsp. malacoides
Malva cretica Cav.
subsp. cretica
x
.
.
x
x
.
x
x
x
x
.
.
x
Me
HC
R
Me
T
P
r
r
r
Hab
EC
NC
Ornithogalum nutans L.
Ornithogalum oligophyllum E.D. Clarke ►
Ornithogalum pyrenaicum L.
Ornithogalum refractum Kit ex Schltdl. ►
Stat
Pe
StE
x
IoI
SPi
Dimopoulos & al.: Vascular plants of Greece: Supplement
NPi
316
x
.
x
x
x
.
x
x
x
x
x
x
x
MYRTACEAE
Eucalyptus camaldulensis Dehnh.
x
.
x
x
x
.
.
.
.
x
x
x
x
X
[Austr.]
P
AR
OLEACEAE
Fraxinus angustifolia Vahl
subsp. syriaca (Boiss.) Yalt.
.
.
.
.
.
.
.
.
.
.
.
.
x
X
MS
P
CW
ONAGRACEAE
Epilobium tournefortii Michalet
x
x
x
x
x
.
.
x
x
x
x
.
x
Bk
H
AR
ORCHIDACEAE
Epipactis persica (Soó) Hausskn. ex Nannf.
subsp. gracilis W. Rossi
Ophrys cretica (Vierh.) E. Nelson
subsp. karpathensis E. Nelson
Ophrys ferrum-equinum Desf.
subsp. ferrum-equinum
.
.
.
x
x
x
.
.
x
x
x
.
.
x
x
x
x
x
x
x
x
x
.
x
x
x
.
.
x
x
x
.
.
x
x
x
.
.
x
x
.
.
.
x
x
.
.
.
x
x
.
x
x
x
x
.
x
x
x
x
.
x
x
x
x
BI
BA
BA
G
G
G
G
G
W
P
P
P
P
Downloaded From: https://bioone.org/journals/Willdenowia on 18 Dec 2023
Terms of Use: https://bioone.org/terms-of-use
r
WAe
Kik
KK
EAe
.
x
x
.
.
.
.
.
x
x
x
.
.
x
x
.
.
.
.
.
x
.
.
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.
x
x
.
.
.
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.
x
x
.
.
.
x
x
.
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.
x
x
.
.
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.
x
.
.
.
.
x
x
.
x
.
.
.
.
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.
.
.
.
.
.
.
.
.
.
.
.
x
.
.
.
ME
G
PW
G
P
EM
G
PW
Me
G
P
x
EM
G
P
.
.
Me
G
PW
.
.
ES
G
W
Stat
Hab
NE
NAe
.
.
Lf
EC
NC
Ophrys holoserica (Burm. f.) Greuter in Greuter & Rech. f. ►
subsp. andria (P. Delforge) Faurh.
subsp. candica (Greuter, Matthäs & Risse) H.A. Pedersen &
Faurh. ►
subsp. holoserica
Ophrys omegaifera H. Fleischm.
subsp. israelitica (H. Baumann & Künkele) G. Morschek &
K. Morschek
Ophrys sphegodes Mill.
subsp. atrata (Rchb. f.) A. Bolòs ►
Pseudorchis albida (L.) Á. Löve & D. Löve
subsp. tricuspis (Beck) E. Klein ►
Serapias neglecta De Not.
subsp. ionica (E. Nelson ex H. Baumann & Künkele)
H. Baumann & R. Lorenz
Serapias vomeracea (Burm. f.) Briq.
Ch
Pe
StE
x
IoI
SPi
317
NPi
Willdenowia 46 – 2016
r
x
.
.
.
.
.
.
.
.
.
.
.
.
Bk
G
P
x
x
x
x
x
x
x
x
x
x
?
.
x
ME
G
PW
OROBANCHACEAE
Euphrasia minima DC. in Lam. & DC.
subsp. minima
Melampyrum barbatum Waldst. & Kit. ex Willd.
Orobanche reticulata Wallr.
Phelipanche dalmatica (Beck) Soják
Phelipanche lavandulacea (Rchb.) Pomel
subsp. lavandulacea
Phelipanche nana (Reut.) Soják ►
Phelipanche olbiensis (Coss.) Carlón & al.
Phelipanche purpurea (Jacq.) Soják
Rhinanthus pumilus (Sterneck) Pau ►
.
.
.
.
.
x
x
x
x
x
x
x
.
x
.
.
.
.
x
.
x
x
.
x
.
.
.
.
x
.
x
x
.
x
x
.
.
.
x
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
?
ME
ME
BC
Eu
ME
T
T
T
T
T
H
H
G
HR
P
x
x
x
x
.
x
x
.
x
x
.
x
.
x
x
x
x
.
x
.
.
x
.
x
.
x
x
.
.
.
x
x
.
x
x
x
x
.
x
.
.
x
.
.
.
x
x
.
.
.
x
x
.
?
.
x
x
.
?
.
.
x
.
?
.
Me
Pt
Me
EA
Me
T
T
T
T
T
R
GP
R
G
H
OXALIDACEAE
Oxalis stricta L.
x
.
.
.
.
.
.
.
x
.
.
.
x
[Am.]
H
R
PAPAVERACEAE
Papaver davisii (Kadereit) M.V. Agab.
Papaver dubium L. ►
.
?
.
?
.
x
x
x
x
x
.
x
.
x
.
x
.
?
.
x
.
x
.
x
?
x
BA
EA
T
T
R
GR
PLANTAGINACEAE
Plantago altissima L. ►
.
.
x
.
.
.
?
x
.
.
.
.
.
Eu
H
AR
PLUMBAGINACEAE
Goniolimon tataricum (L.) Boiss. in A. DC.
.
.
.
.
.
.
x
x
x
.
.
.
.
ME
H
G
x
x
x
x
x
x
x
x
x
x
x
x
x
MS
T
P
x
.
x
x
x
x
x
x
x
x
x
x
x
Me
G
M
x
.
.
x
x
.
?
x
x
x
x
x
x
Me
T
P
.
.
x
x
.
.
x
x
.
.
x
.
x
MS
H
M
x
.
.
.
x
x
x
.
.
.
.
x
.
x
x
x
x
x
x
x
.
.
x
.
x
x
x
x
x
x
x
.
.
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
x
.
.
x
.
x
x
x
x
x
.
.
.
.
x
.
x
x
x
x
x
x
x
.
.
x
x
x
x
x
x
x
x
x
.
.
x
x
x
x
x
x
x
x
x
.
.
x
x
x
x
x
x
x
x
x
.
.
x
x
x
x
x
x
.
.
.
.
x
.
x
?
.
x
x
.
.
.
.
x
x
x
x
x
x
x
x
x
x
.
x
ES
EM
Co
Co
ST
MS
Me
Bo
Bo
Me
G
H
T
T
H
T
H
H
H
T
T
GR
M
R
R
R
MR
GH
G
G
R
R
x
x
x
.
x
.
x
.
x
.
x
x
.
x
.
.
x
.
.
x
x
x
.
x
.
.
x
.
.
x
x
x
.
x
.
.
.
.
x
.
.
x
x
x
.
.
.
x
.
x
x
x
.
x
.
x
.
.
.
.
x
x
.
x
.
.
.
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.
x
x
x
x
x
.
x
.
.
.
x
x
x
x
x
.
x
.
.
.
.
x
x
x
x
.
.
.
.
.
.
x
.
.
.
.
.
.
x
x
.
x
x
x
.
.
.
x
.
x
.
x
x
x
x
.
x
x
x
x
?
Me
MS
Me
MS
MA
EA
Me
MS
Me
Me
H
T
T
T
T
T
T
T
T
T
GP
RW
W
RW
M
A
R
R
M
GR
POLYGONACEAE
Rumex aquaticus L.
.
.
.
.
.
.
x
x
?
.
.
.
.
ES
H
A
POTAMOGETONACEAE
Potamogeton berchtoldii Fieber in Bercht. & Opiz
Potamogeton pusillus L.
Potamogeton schweinfurthii A. Benn. in Oliv.
.
.
x
.
x
.
.
x
.
x
x
.
.
.
.
.
.
.
x
x
.
x
x
.
.
.
.
.
.
.
.
x
.
.
x
x
.
x
.
Co
ST
Me
A
A
A
A
A
A
POACEAE
Aira elegans Willd. ex Roem. & Schult. ►
Ammophila arenaria (L.) Link
subsp. australis H. Lindb.
Bromus fasciculatus C. Presl
subsp. fasciculatus
Elytrigia elongata (Host) Nevski
subsp. elongata
Elytrigia repens (L.) Nevski
subsp. repens
Elytrigia sartorii (Boiss. & Heldr.) Holub ►
Eragrostis minor Host ►
subsp. minor
Hordeum bulbosum L. ►
Hordeum murinum L.
Koeleria lobata (M. Bieb.) Roem. & Schult.
Koeleria macrantha (Ledeb.) Schult.
subsp. macrantha
Lolium scholzii Greuter
Lolium subulatum Vis.
Melica transsilvanica Schur
subsp. klokovii Tzvelev
Milium vernale M. Bieb.
subsp. montianum (Parl.) K. Richt.
subsp. vernale
Parapholis strigosa (Dumort.) C.E. Hubb.
Pholiurus pannonicus (Host) Trin.
Rostraria pubescens (Lam.) Trin.
Rostraria smyrnaea (Trin.) H. Scholz
Triplachne nitens (Guss.) Link
Ventenata dubia (Leers) Coss. in Durieu
Downloaded From: https://bioone.org/journals/Willdenowia on 18 Dec 2023
Terms of Use: https://bioone.org/terms-of-use
X
r
SPi
Pe
StE
EC
NC
NE
NAe
WAe
Kik
KK
EAe
Ch
Lf
Hab
x
x
.
.
.
x
.
x
.
.
.
.
.
x
x
.
.
x
.
.
.
x
.
.
x
x
.
.
x
x
.
.
x
x
x
.
.
x
.
.
.
x
x
.
.
x
.
.
x
x
x
.
ME
Me
Ct
ES
G
G
T
H
GR
AW
A
A
Stat
IoI
RANUNCULACEAE
Ficaria verna Huds.
subsp. calthifolia (Rchb.) Nyman
subsp. ficariiformis (F.W. Schultz ex Rouy & Foucaud) Maire ►
Myosurus sessilis S. Watson
Ranunculus auricomus L. ►
Ranunculus bullatus L.
subsp. cytheraeus (Halácsy) Vierh.
Dimopoulos & al.: Vascular plants of Greece: Supplement
NPi
318
.
.
.
x
.
.
.
.
.
.
.
x
x
EM
H
P
RESEDACEAE
Reseda alba L.
subsp. alba
Reseda phyteuma L. ►
x
x
.
x
x
x
x
x
x
x
x
.
x
x
x
x
x
.
x
x
x
.
x
.
x
.
ME
ME
TH
TH
R
R
ROSACEAE
Aphanes floribunda (Murb.) Rothm.
Pyracantha coccinea M. Roem.
Pyrus pyraster (L.) Burgsd.
Sorbus aria (L.) Crantz
x
x
x
.
.
x
x
x
.
x
x
x
.
x
x
x
.
x
.
x
.
x
x
.
.
x
x
x
.
x
x
x
.
x
.
.
.
.
.
.
.
.
.
.
x
.
.
.
.
x
x
.
Me
ME
ME
ME
T
P
P
P
P
W
W
CW
.
x
.
x
x
.
.
x
.
.
x
.
.
x
.
.
x
.
.
x
.
x
x
x
x
x
x
.
x
x
.
.
x
.
.
x
.
.
x
BA
EA
EM
H
H
T
G
GW
P
.
.
.
.
x
.
x
x
.
.
.
.
x
.
x
x
.
.
.
.
x
.
x
x
.
.
x
.
x
.
x
x
.
.
x
.
x
.
x
x
.
x
.
.
.
.
x
x
.
x
.
.
x
.
x
x
.
x
.
.
x
.
x
x
.
.
x
x
.
.
x
x
.
.
x
.
.
.
x
x
.
.
x
.
.
.
x
x
x
.
.
.
.
.
x
x
x
.
.
.
.
x
x
x
EM
MS
EA
EM
Me
Me
C
H
H
T
H
T
T
T
C
GW
GP
R
A
W
P
P
x
x
.
x
x
x
x
x
x
x
x
.
x
.
x
.
x
.
x
x
x
x
x
x
x
.
MA
Bk
P
H
W
H
SALICACEAE
Salix elaeagnos Scop. ►
x
x
x
x
x
x
x
x
.
.
.
.
.
ME
P
AW
SCROPHULARIACEAE
Scrophularia canina L.
subsp. bicolor (Sm.) Greuter in Greuter & Rech. f.
subsp. canina
Verbascum chaixii Vill.
Verbascum delphicum Boiss. & Heldr. in Boiss.
Verbascum gloeotrichum Hausskn. & Heldr.
Verbascum nigrum L.
x
x
.
x
.
.
.
x
x
x
x
.
.
x
x
x
x
x
.
x
x
x
x
.
.
.
.
x
x
x
.
.
x
.
.
x
x
x
x
.
.
x
x
x
x
x
.
x
x
x
x
.
x
.
x
x
x
x
.
x
.
.
.
x
x
.
.
x
x
.
x
x
.
.
.
.
.
.
.
.
.
.
.
.
x
x
.
.
.
.
.
ME
EM
ME
ME
Bk
EA
H
H
H
H
H
H
H
GHR
GR
GH
GRW
GR
G
RW
SOLANACEAE
Solanum villosum Mill.
x
x
x
.
.
x
.
.
x
x
x
x
x
EA
T
R
TAMARICACEAE
Tamarix tetrandra Pall. ex M. Bieb. ►
?
.
x
x
x
x
x
x
?
?
?
?
?
ME
P
AM
THYMELAEACEAE
Daphne sojakii Halda
.
.
.
.
.
.
x
.
.
.
.
.
.
C
H
ULMACEAE
Ulmus laevis Pall.
.
.
.
.
.
.
x
x
.
x
.
.
.
EA
P
W
URTICACEAE
Urtica dioica L.
subsp. dioica
x
x
x
x
x
x
x
x
x
x
.
.
x
Co
H
R
x
.
x
x
x
x
.
x
x
x
x
x
x
Me
T
P
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GH
RUBIACEAE
Asperula tenella Degen in A. Kern. ►
Cruciata laevipes Opiz
Galium caminianum Schult. & Schult. f. ►
Galium canum Req. ex DC.
subsp. ovatum Ehrend.
Galium exaltatum Krendl
Galium melanantherum Boiss.
Galium nigricans Boiss. ►
Galium palustre L.
Galium peplidifolium Boiss.
Galium setaceum Lam.
subsp. decaisnei (Boiss.) Ehrend.
Rubia peregrina L.
subsp. peregrina
Valantia aprica (Sm.) Tausch ►
VALERIANACEAE
Centranthus calcitrapae (L.) Dufr. ►
Centranthus longiflorus Steven
subsp. junceus (Boiss. & Heldr.) I. Richardson
Valeriana tuberosa L.
Valerianella vesicaria (L.) Moench
VERONICACEAE
Antirrhinum majus L.
subsp. tortuosum (Vent.) Rouy
Kickxia lanigera (Desf.) Hand.-Mazz.
Kickxia spuria (L.) Dumort.
subsp. integrifolia (Brot.) R. Fern.
Veronica panormitana Tineo ex Guss.
subsp. panormitana
VIOLACEAE
Viola macedonica Boiss. & Heldr. in Boiss.
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Willdenowia 46 – 2016
Appendix I: Excluded taxa. Supplement
Taxa disregarded as being reported in error, non-established aliens, non-stabilized hybrids, taxonomically enigmatic, or vanished. The arrow symbol “►” refers to a
comment under the same name in Appendix III.
Taxa listed in Appendix I, given for Greece in different recent basic floras, checklists and databases, have
been correctly (though without comment) excluded from
the flora of Greece by mere omission in Dimopoulos &
al. (2013). However, supplementary comments on that
exclusion are furnished here in Appendices I and III in
order to serve as a correction tool for floristically deviating basic sources.
Atriplex oblongifolia, Eryngium palmatum, Hyoseris
radiata and Leonurus marrubiastrum, previously disregarded by Dimopoulos & al. (2013), now prove to be
members of the Greek flora (see Floristic catalogue and
Appendix III).
ACERACEAE
Acer ×bornmuelleri Borbás ►
ALLIACEAE
Allium sativum L. ►
APIACEAE
Helosciadium repens (Jacq.) W.D.J. Koch ►
Heracleum sphondylium subsp. orsinii (Guss.) H. Neumayer ►
Pastinaca sativa L. subsp. sativa ►
ASPARAGACEAE
Asparagus aphyllus L. subsp. aphyllus ►
ASPLENIACEAE
Asplenium adulterinum Milde subsp. adulterinum ►
ASTERACEAE
Achillea ×tymphaea Hausskn. ►
Anthemis bornmuelleri Stoj. & Acht. ►
Anthemis macrantha Heuff. ►
Anthemis parvifolia Eig ►
Carduus ×intercedens Hausskn. ►
Centaurea phrygia subsp. indurata (Janka) Stoj. & Acht. ►
Centaurea phrygia subsp. razgradensis (Velen.) Greuter ►
Centaurea stereophylla Besser ►
Cichorium endivia L. ►
Cirsium serrulatum (M. Bieb.) Fisch. ►
Gnaphalium hoppeanum W.D.J. Koch subsp. hoppeanum ►
Hieracium bifidum subsp. basicuneatum (Zahn) Zahn ►
Jacobaea aquatica L. ►
Jurinea kilaea Azn. ►
Jurinea polycephala Formánek ►
Osteospermum barberae (Harv.) Norl. ►
Pilosella alpicola (Froel.) F.W. Schulz & Sch. Bip. ►
Pilosella caespitosa (Dumort.) P.D. Sell & C. West ►
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319
BETULACEAE
Corylus maxima Mill. ►
BRASSICACEAE
Alyssum stribrnyi Velen. ►
CAMPANULACEAE
Asyneuma canescens subsp. cordifolium (Bornm.) Damboldt ►
Edraianthus tenuifolius (Waldst. & Kit.) A. DC. ►
CARYOPHYLLACEAE
Arenaria biflora L. ►
Cerastium arvense L. ►
Cerastium brachypetalum Pers. subsp. brachypetalum ►
Cerastium diffusum Pers. ►
Cerastium gracile Dufour ►
Dianthus giganteus subsp. croaticus (Borbás) Tutin ►
Dianthus leptopetalus Willd. ►
Dianthus microlepis Boiss. ►
Dianthus pallidiflorus Ser. ►
Dianthus pancicii Velen. ►
Dianthus petraeus Waldst. & Kit. subsp. petraeus ►
Dianthus roseoluteus Velen. ►
Holosteum umbellatum subsp. glutinosum (M. Bieb.) Nyman ►
Minuartia erythrosepala (Boiss.) Hand.-Mazz. ►
Minuartia graminifolia (Ard.) Jáv. subsp. graminifolia ►
Minuartia rumelica Panov ►
Minuartia setacea (Thuill.) Hayek subsp. setacea ►
Paronychia sintenisii Chaudhri ►
Saponaria sicula Raf. ►
Silene densiflora d’Urv. ►
Silene heldreichii Boiss. ►
Silene nemoralis Waldst. & Kit. ►
Silene portensis ►
Silene vulgaris (Moench) Garcke subsp. vulgaris ►
Spergularia ×hybrida Hausskn. ►
CRASSULACEAE
Sedum rupestre Chaix ►
Sempervivum octopodes Turrill ►
CUPRESSACEAE
Juniperus phoenicea L. ►
CYPERACEAE
Carex atrata subsp. aterrima (Hoppe) Čelak. ►
Carex pairae F.W. Schultz ►
Carex sempervirens L. ►
DRYOPTERIDACEAE
Polystichum ×lonchitiforme (Halácsy) Becherer ►
ERICACEAE
Calluna vulgaris (L.) Hull ►
EUPHORBIACEAE
Euphorbia anacampseros Boiss. ►
320
Euphorbia hierosolymitana Boiss. ►
Euphorbia hirta L. ►
Euphorbia hypericifolia L. ►
Euphorbia lucida Waldst. & Kit. ►
FABACEAE
Astragalus lanatus Labill. ►
Lathyrus vernus (L.) Bernh. ►
Medicago ×blancheana Boiss. ►
FAGACEAE
Quercus ×kanitziana Borbás ►
Quercus ×szechenyana Borbás ►
FUMARIACEAE
Fumaria officinalis subsp. wirtgenii (W.D.J. Koch) Arcang. ►
Fumaria schleicheri Soy.-Will. ►
Pseudofumaria alba (Mill.) Lidén subsp. alba ►
GERANIACEAE
Geranium pratense L. ►
HYACINTHACEAE
Ornithogalum corsicum Jord. & Fourr. ►
Scilla peruviana L. ►
HYPERICACEAE
Hypericum hyssopifolium Chaix ►
Hypericum maculatum Crantz subsp. maculatum ►
Hypericum richeri Vill. ►
LAMIACEAE
Lycopus ×intermedius Hausskn. ►
Marrubium ×paniculatum Desr. ►
Satureja ×boissieri sensu Hayek, vix Briq. ►
Teucrium spinosum L. ►
Thymus kosteleckyanus Opiz ►
ONAGRACEAE
Epilobium ×persicinum Rchb. ►
ORCHIDACEAE
Anacamptis ×eccarii (Biel) H. Kretzschmar & G. Kretzschmar ►
Anacamptis ×gennarii (Rchb. f.) H. Kretzschmar & al. ►
Anacamptis ×lesbiensis (Biel) H. Kretzschmar & al. ►
Anacamptis ×parvifolia nothosubsp. bicknellii (E.G. Camus) H.
Kretzschmar & al. ►
Anacamptis ×sciathia (Biel) H. Kretzschmar & al. ►
Anacamptis ×simorrensis (E.G. Camus) H. Kretzschmar & al.►
Cephalanthera ×majeri W. Zimm. ►
Epipactis leptochila subsp. aspromontana (Bartolo, Pulv. &
Robatsch) Kreutz ►
Epipactis leptochila subsp. neglecta Kümpel ►
Liparis loeselii (L.) Rich. ►
Ophrys ×asterusica C. Alibertis & A. Alibertis ►
Ophrys ×baumanniana Soó nothosubsp. baumanniana ►
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Dimopoulos & al.: Vascular plants of Greece: Supplement
Ophrys ×baumanniana nothosubsp. hierapetrae H. Baumann
& Künkele ►
Ophrys ×burneriana C. Alibertis & A. Alibertis ►
Ophrys ×corcyrensis Renz ►
Ophrys holoserica subsp. bornmuelleri (M. Schulze) H.
Sund. ►
Ophrys holoserica subsp. grandiflora (H. Fleischmann & Soó)
Faurh. ►
Ophrys ×kastelli nothosubsp. antiskariensis C. Alibertis & A.
Alibertis ►
Ophrys ×kastelli E. Klein nothosubsp. kastelli ►
Ophrys ×keramensis E. Klein ►
Ophrys ×lithinensis C. Alibertis & A. Alibertis ►
Ophrys ×maremmae O. Danesch & E. Danesch ►
Ophrys ×pauliana C. Alibertis & A. Alibertis ►
Ophrys ×pezaenensis E. Klein ►
Ophrys ×plorae C. Alibertis & A. Alibertis ►
Ophrys ×pseudoquadriloba Renz ►
Ophrys ×pseudospruneri Soó ►
Ophrys ×rechingeri Soó ►
Ophrys scolopax subsp. nestoris A. Alibertis & Brütsch ►
Ophrys ×sieberi H. Baumann & Künkele ►
Ophrys ×sivana H. Baumann & Künkele ►
Ophrys ×skopelii Renz ►
Ophrys ×sommieri E.G. Camus ex Cortesi ►
Ophrys sphegodes subsp. atrata (Rchb. f.) A. Bolòs ►
Ophrys sphegodes subsp. litigiosa (E.G. Camus) Bech. ►
Ophrys ×varvarae Faller & Kreutz ►
×Orchinea attica (Hausskn.) F.N. Vázquez ►
×Orchinea hermaniana (C. Alibertis & A. Alibertis) J.M.H.
Shaw ►
Orchis ×adriatica Soó ►
Orchis ×bivonae Tod. ►
Orchis ×dicorifiana G. Thiele & W. Thiele ►
Orchis ×kretzschmariorum B. Baumann & H. Baumann ►
Orchis mascula subsp. speciosa (Mutel) Hegi ►
Orchis ×paschae Hauzinger ►
Orchis ×plessidiaca Renz ►
Orchis ×salkowskiana C. Alibertis & A. Alibertis ►
Orchis ×sezikiana B. Baumann & H. Baumann ►
Orchis ×thriftiensis Renz ►
Orchis ×willingiorum B. Baumann & H. Baumann ►
Serapias ×ambigua E.G. Camus nothosubsp. ambigua ►
Serapias ×ambigua nothosubsp. panormosana B. Baumann &
H. Baumann ►
Serapias ×broeckii A. Camus ►
Serapias ×cythereis Renz ►
Serapias ×fallax Soó ►
Serapias ×halacsyana Soó ►
Serapias ×intermedia F.W. Schultz ►
Serapias ×kelleri A. Camus ►
Serapias ×kelleriana Renz ►
Serapias neglecta subsp. apulica (E. Nelson) Landwehr ►
Serapias neglecta De Not. subsp. neglecta ►
Serapias ×semicolumnae E.G. Camus & A. Camus ►
Serapias ×semilingua E.G. Camus ►
Willdenowia 46 – 2016
Serapias ×sooi Renz ►
×Serapicamptis ligustica (Dupuy) J.M.H. Shaw ►
×Serapicamptis rousii (Dupuy) J.M.H. Shaw ►
OROBANCHACEAE
Phelipanche ramosa (L.) Pomel ►
Rhinanthus illyricus (Sterneck) Soó ►
PAEONIACEAE
Paeonia arietina G. Anderson ►
PINACEAE
Cedrus atlantica (Endl.) Carrière ►
Cedrus deodara (D. Don) G. Don ►
Larix decidua Mill. ►
Picea pungens Engelm. ►
Pinus canariensis C. Sm. ►
Pinus pinaster Aiton ►
Pinus ponderosa P. Lawson & C. Lawson ►
Pseudotsuga menziesii (Mirb.) Franco ►
PLANTAGINACEAE
Plantago macrorrhiza Poir. ►
PLUMBAGINACEAE
Goniolimon dalmaticum (C. Presl) Rchb. f. ►
POACEAE
Helictochloa pratensis (L.) Romero Zarco ►
Helictochloa versicolor (Vill.) Romero Zarco ►
Hierochloe australis (Schrad.) Roem. & Schult. ►
Hierochloe odorata (L.) Wahlenb. ►
Melica picta K. Koch ►
Stipa tirsa Steven ►
POLYGONACEAE
Polygonum ×heldreichii Halácsy ►
Polygonum ×pseudobellardii Hausskn. ►
Polygonum ×pseudopulchellum Hausskn. ►
Rumex ×abortivus Ruhmer ►
Rumex ×dimidiatus Hausskn. ►
Rumex ×halacsyi Rech. ►
Rumex ×muretii Hausskn. ►
Rumex ×pratensis Mert. & W.D.J. Koch ►
Rumex ×semigraecus Hausskn. ►
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321
RANUNCULACEAE
Ceratocephala orthoceras DC. ►
Ranunculus acris subsp. friesianus (Jord.) Syme ►
Ranunculus bulbosus subsp. aleae (Willk.) Rouy & Foucaud ►
Ranunculus carinthiacus Hoppe ►
Ranunculus penicillatus (Dumort.) Bab. subsp. penicillatus ►
Ranunculus polyanthemos subsp. nemorosus (DC.) Schübl. &
G. Martens ►
Ranunculus polyanthemos subsp. serpens (Schrank) Baltisb. ►
Ranunculus pseudomontanus Schur ►
RESEDACEAE
Reseda alba subsp. hookeri (Guss.) Arcang. ►
ROSACEAE
Potentilla ×commixta Hausskn. ►
Potentilla ×degenii Th. Wolf ►
Potentilla ×dispersa Hausskn. ►
Potentilla ×dolosa Hausskn. ►
Potentilla ×intercedens Hausskn. ►
Potentilla ×kerneri Borbás ►
Potentilla ×micans Hausskn. ►
Potentilla ×pedatoides Hausskn. ►
Rosa ×guicciardii Burnat & Gremli ►
Rosa ×oetea Burnat & Gremli ►
SAXIFRAGACEAE
Saxifraga juniperifolia Adams ►
SCROPHULARIACEAE
Verbascum ×ambracicum Halácsy ►
Verbascum ovalifolium Sims subsp. ovalifolium ►
Verbascum ×parallelum Hausskn. ►
Verbascum ×petrophilum Halácsy ►
VERBENACEAE
Verbena ×adulterina Hausskn. ►
VERONICACEAE
Kickxia spuria (L.) Dumort. subsp. spuria ►
Veronica alpina L. ►
VIOLACEAE
Viola ×lacmonica Hausskn. ►
Viola tricolor ►
322
Dimopoulos & al.: Vascular plants of Greece: Supplement
Appendix II: Synonyms and misapplied names. Supplement
Synonyms collected here mirror changes in the Floristic catalogue and in Appendix I due to newly detected nomenclatural priority or taxonomic and floristic assessments. Additional synonyms result not only from continued datamining in old and rarely cited primary literature sources, but also include nomenclatural novelties derived from recent
findings in molecular taxonomy; such novelties, however, are not necessarily accepted in Dimopoulos & al. (2013).
Acer monspessulanum subsp. athoum (Bornm. & Sint.) F.K. Mey.
Acis valentina auct. fl. graec., non (Pau) Lledó & al.
Acuston lunarioides (Willd.) Raf.
Agropyrum sartorii (Boiss. & Heldr.) Grecescu
Aira capillaris Host, non Savi
Aira elegans auct. fl. graec., non Gaudin
Aira elegans subsp. ambigua (Asch.) Holub
Aira elegantissima Schur
Aira elegantissima subsp. ambigua (Asch.) Doğan
Ajuga orientalis subsp. aenesia (Heldr.) Phitos & Damboldt
Alectorolophus mediterraneus Sterneck
Alectorolophus pumilus Sterneck
Alkanna bracteolata Greuter
Alkanna lehmannii (Tineo) A. DC.
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Acer monspessulanum L. subsp. monspessulanum
Leucojum ionicum Kit Tan & al.
Fibigia lunarioides (Willd.) Sweet
Elytrigia sartorii (Boiss. & Heldr.) Holub
Aira elegans Roem. & Schult.
Aira elegans Roem. & Schult.
Aira elegans Roem. & Schult.
Aira elegans Roem. & Schult.
Aira elegans Roem. & Schult.
Ajuga orientalis L.
Rhinanthus pumilus (Sterneck) Pau
Rhinanthus pumilus (Sterneck) Pau
Alkanna tinctoria Tausch
Alkanna tinctoria Tausch
Alkanna matthioli Tausch
Alkanna tinctoria subsp. lehmannii (Tineo) Nyman
Alkanna tuberculata Greuter
Alkanna tuberculata (Forssk.) Meikle, non Greuter
Allium scorodoprasum subsp. rotundum (L.) Stearn
Alyssoides utriculata subsp. graeca (Boiss.) Zangh.
Alyssum diffusum auct. fl. graec., non Ten.
Alyssum murale subsp. chalcidicum (Janka) Contandr.
Alyssum murale subsp. chlorocarpum (Hausskn.) Contandr.
Alyssum suffrutescens Halácsy, non Boiss.
Amaranthus graecizans subsp. sylvestris (Vill.) Brenan
Ammophila arenaria subsp. arundinacea H. Lindb., nom. inval.
Ammophila littoralis (P. Beauv.) Rothm.
Amygdalus incana Sm., non Pall.
Anacamptis papilionacea subsp. expansa (Ten.) Amard. & Dusak
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
×Anacamptorchis simorrensis E.G. Camus
→
×Anacamptorchis simorrensis E.G. Camus
→
Anagallis arvensis subsp. foemina (Mill.) Schinz & R. Keller
Androrchis anatolica (Boiss.) D. Tyteca & E. Klein
Androrchis ×dicorifiana (G. Thiele & W. Thiele) W. Foelsche & Jakely
Androrchis ×kretzschmariorum (B. Baumann & H. Baumann) W. Foelsche & Jakely
Androrchis ovalis (F.W Schmidt.) D. Tyteca & E. Klein
Androrchis pallens (L.) D. Tyteca & E. Klein
Androrchis pauciflora (Ten.) D. Tyteca & E. Klein
Androrchis pinetorum (Boiss. & Kotschy) D. Tyteca & E. Klein
Androrchis ×plessidiaca (Renz) W. Foelsche & Jakely
Androrchis provincialis (Lam. & DC.) D. Tyteca & E. Klein
Androrchis quadripunctata (Ten.) D. Tyteca & E. Klein
Androrchis sitiaca (Renz) D. Tyteca & E. Klein
Androrchis spitzelii (W.D.J. Koch) D. Tyteca & E. Klein
Androrchis ×willingiorum (B. Baumann & H. Baumann) W. Foelsche
& Jakely
→
→
→
→
Alkanna tinctoria Tausch
Alkanna tinctoria Tausch
Alkanna tinctoria Tausch
Alkanna tinctoria Tausch
Allium rotundum L.
Alyssoides utriculata (L.) Medik. subsp. utriculata
Alyssum montanum subsp. repens (Baumg.) Schmalh.
Alyssum chalcidicum Janka
Alyssum chalcidicum Janka
Alyssum sibiricum Willd.
Amaranthus graecizans L.
Ammophila arenaria subsp. australis (Mabille) M. Laínz
Ammophila arenaria subsp. australis (Mabille) M. Laínz
Prunus graeca (Lindl.) Steud.
Anacamptis papilionacea subsp. aegaea (P. Delforge) L. Lewis &
Kreutz
Anacamptis ×simorrensis (E.G. Camus) H. Kretzschmar & al. [see
Appendix I]
Anacamptis ×simorrensis (E.G. Camus) H. Kretzschmar & al. [see
Appendix I]
Anagallis foemina Mill.
Orchis anatolica Boiss.
Orchis ×dicorifiana G. Thiele & W. Thiele [see Appendix I]
Orchis ×kretzschmariorum B. Baumann & H. Baumann [see Appendix I]
Orchis mascula subsp. speciosa (Mutel) Hegi [see Appendix I]
Orchis pallens L.
Orchis pauciflora Ten.
Orchis mascula (L.) L. subsp. mascula
Orchis ×plessidiaca Renz [see Appendix I]
Orchis provincialis Lam. & DC.
Orchis quadripunctata Ten.
Orchis sitiaca (Renz) P. Delforge
Orchis spitzelii W.D.J. Koch
Orchis ×willingiorum B. Baumann & H. Baumann [see Appendix I]
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323
Anemone messarensis Coust. & Gand.
Anthemis galilaea auct. fl. graec., non Eig
Anthemis parvifolia auct. fl. graec., non Eig
Anthemis rigescens Willd.
Anthemis tomentosa subsp. peregrina (L.) Hayek
Anthemis visianii E. Weiss
Anthyllis circinnata (L.) D.D. Sokoloff
Apargia aspera Waldst. & Kit.
Apargia saxatilis Ten.
Apium heldreichii (Boiss.) Calest.
Apium meoides (Griseb.) Calest.
Apium rupestre (Boiss. & Heldr.) Calest.
→
→
→
→
→
→
→
→
→
→
→
→
Aquilegia ottonis subsp. australis Quézel & Contandr.
Aquilegia ottonis subsp. meridionalis Quézel & Contandr.
Arabis ochroleuca (Lam.) Lam.
Arabis ochroleuca Boiss. & Heldr., non (Lam.) Lam.
Arachnites fuciflora F.W. Schmidt
Aspidium ×lonchitiforme Halácsy
Asplenium adulterinum auct. fl. graec., non Milde
Asplenium ceterach subsp. mediterraneum Pinter
→
→
→
→
→
→
→
→
Anemone coronaria L.
Anthemis cotula L.
Anthemis pseudocotula Boiss.
Anthemis triumfettii (L.) DC.
Anthemis tomentosa L. subsp. tomentosa
Anthemis chia L.
Hymenocarpos circinnatus (L.) Savi
Leontodon biscutellifolius DC.
Leontodon crispus Vill.
Carum heldreichii Boiss.
Carum graecum Boiss. & Heldr.
Carum meoides (Griseb.) Halácsy 1894, non Halácsy 1901 [see Appendix III]
Aquilegia ottonis Boiss. subsp. ottonis
Aquilegia ottonis Boiss. subsp. ottonis
Arabis turrita L.
Arabis subflava B.M.G. Jones
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Polystichum ×lonchitiforme (Halácsy) Becherer [see Appendix I]
Asplenium trichomanes L. subsp. trichomanes
Asplenium ceterach L.
Astracantha cretica subsp. rumelica (Bunge) Podlech
Astracantha rumelica (Bunge) Reer & Podlech
Astracantha rumelica subsp. taygetica (Širj.) Reer & Podlech
Astracantha thracica subsp. cyllenea (Fisch.) Greuter
Astragalus albanicus Širj.
Astragalus atticus Nyman
Astragalus macedonicus Heldr. & Nadji
Astragalus maniaticus Kit Tan & Strid
Astragalus pamphylicus subsp. argolicus (Hausskn.) Hayek
Astragalus veluchensis Boiss.
Avena heldreichii Parl.
Avenula agropyroides (Boiss.) Holub
Ballota nigra subsp. uncinata (Bég.) Patzak
Blackstonia perfoliata subsp. serotina (Rchb.) Vollm.
Bromus willdenowii Kunth
Bupleurum baldense auct. fl. graec., non Turra
Bupleurum baldense subsp. gussonei auct. fl. graec., non (Arcang.)
Tutin
Bupleurum falcatum subsp. exaltatum (M. Bieb.) Briq.
Bupleurum gramineum auct. fl. graec., non Vill.
Butinia macrocarpa Boiss. & Spruner
Butinia stylosa Boiss.
Calamagrostis arenaria (L.) Roth
Campanula thessala Maire
Campanula topaliana subsp. cordifolia Phitos
Camphorosma nestensis Turrill
Carduus neglectus Ten., non Steud.
Carduus pindicola Hausskn.
Carduus tmoleus subsp. armatus (Boiss. & Heldr.) Franco
Carex caesia Griseb.
Carex flacca subsp. erythrostachys auct. fl. graec., non (Hoppe) K.
Richt.
Carex muricata subsp. pairae (F.W. Schultz) Čelak.
Carex sempervirens auct. fl. graec., non Vill.
Carex soleirolii DC. & Duby
Carex tomentosa L.
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Astragalus rumelicus Bunge
Astragalus rumelicus Bunge
Astragalus rumelicus Bunge
Astragalus thracicus subsp. cylleneus (Fisch.) Strid
Astragalus rumelicus Bunge
Astragalus monspessulanus L. subsp. monspessulanus
Astragalus monspessulanus L. subsp. monspessulanus
Astragalus suberosus subsp. haarbachii (Boiss.) V.A. Matthews
Astragalus suberosus subsp. haarbachii (Boiss.) V.A. Matthews
Astragalus rumelicus Bunge
Helictotrichon convolutum (C. Presl) Henrard
Helictochloa agropyroides (Boiss.) Romero Zarco
Ballota nigra subsp. ruderalis (Sw.) Briq.
Blackstonia acuminata (Koch & Ziz) Domin subsp. acuminata
Bromus catharticus Vahl
Bupleurum veronense Turra
Bupleurum veronense Turra
→
→
→
→
→
→
→
→
→
→
→
→
→
Bupleurum falcatum subsp. cernuum (Ten.) Arcang.
Bupleurum falcatum subsp. cernuum (Ten.) Arcang.
Geocaryum macrocarpum (Boiss. & Spruner) Engstrand
Geocaryum stylosum (Boiss.) Engstrand
Ammophila arenaria (L.) Link
Campanula pelia Bedd.
Campanula topaliana Beauverd subsp. topaliana
Camphorosma monspeliaca L. subsp. monspeliaca
Carduus acicularis Bertol.
Carduus nutans subsp. leiophyllus (Petrović) Stoj. & Stef.
Carduus tmoleus Boiss. subsp. tmoleus
Carex filiformis L.
Carex flacca Schreb. subsp. serrulata (Spreng.) Greuter
→
→
→
→
Carex pairae F.W. Schultz [see Appendix I]
Carex bulgarica (Domin) Lazare
Carex hispida Willd.
Carex filiformis L.
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→ Carthamus lanatus subsp. baeticus (Boiss. & Reut.) Nyman
→ Carum meoides (Griseb.) Halácsy 1894, non Halácsy 1901 [see Appendix III]
Carum grimburgii Halácsy
→ Trinia glauca subsp. pindica Hartvig
Carum macedonicum Quézel & Contandr.
→ Carum meoides (Griseb.) Halácsy 1894, non Halácsy 1901 [see Appendix III]
Carum meoides auct. fl. graec. sensu Halácsy 1901, non (Griseb.) → Carum graecum Boiss. & Heldr.
Halácsy 1894
Carum meoides subsp. heldreichii (Boiss.) Maire & Petitm.
→ Carum heldreichii Boiss.
Carum pachypodium P. Candargy
→ Oenanthe silaifoia M. Bieb.
Carum rupestre Boiss. & Heldr.
→ Carum meoides (Griseb.) Halácsy 1894, non Halácsy 1901 [see Appendix III]
Carum scaligerioides Bornm.
→ Hellenocarum strictum (Griseb.) Hand
Centaurea albanica Bornm.
→ Centaurea alba subsp. albanica (Bornm.) Dostál
Centaurea deustiformis subsp. ptarmicifolia (Hayek) Dostál
→ Centaurea ptarmicifolia Hayek
Centaurea epirotica (Bald.) Halácsy
→ Centaurea ptarmicifolia Hayek
Centaurea paniculata var. macedonica Griseb.
→ Centaurea grisebachii (Nyman) Heldr.
Centaurea ptarmicoides Halácsy nom. inval.
→ Centaurea ptarmicifolia Hayek
Centaurea stenolepis subsp. indurata auct. fl. graec. non (Janka) Stoj. → Centaurea phrygia subsp. stenolepis (A. Kern.) Gugler
& Acht.
Centaurium erythraea subsp. grandiflorum auct. fl. graec., non (Pers.) → Centaurium erythraea subsp. rhodense (Boiss. & Reut.) Melderis
Carthamus lanatus subsp. creticus (L.) Holmboe
Carum adamovicii Halácsy
Melderis
Cephalanthera pallens (Sw.) Rich.
Cephalanthera pallens auct. fl. graec., non (Sw.) Rich.
Cerastium brachypetalum subsp. tauricum (Spreng.) Murb.
Cerastium caespitosum Asch.
Cerastium chassium Formánek
Cerastium corcyrense Möschl
Cerastium decrescens (Lonsing) Greuter & Burdet
Cerastium fontanum subsp. holosteoides (Fr.) Salman & al.
Cerastium fontanum subsp. triviale Jalas
Cerastium fontanum subsp. vulgare (Hartm.) Greuter & Burdet
Cerastium holosteoides subsp. vulgare (Hartm.) Buttler
Cerastium semidecandrum subsp. balearicum (F. Hermann) Litard.
Cerastium triviale Link
Cerastium vulgatum L.
Ceratocephala orthoceras auct. fl. graec., non DC.
Ceratocephala testiculata (Crantz) Roth
Chaiturus marrubiastrum (L.) Rchb.
Chamaenerion dodonaei (Vill.) Schur
Chamomilla recutita (L.) Rauschert
Cheiranthus senoneri Reut.
Chlora intermedia Ten.
Chondrilla acantholepis Boiss.
Cistus albanicus Heywood
Cleome macedonica Heldr. & Nadji
Coronilla argentea L.
Coronilla emerus L.
Corylus tubulosa Willd.
Cota macrantha (Heuff.) Boiss.
Crepis nemausensis Gouan
Crepis turcica subsp. murganica Kamari
Crocus ionicus Herb.
Crocus marathonisius Heldr.
Crocus mazziaricus Herb.
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→ Cephalanthera longifolia (L.) Fritsch
→ Cephalanthera damasonium (Mill.) Druce
→ Cerastium brachypetalum Pers. subsp. brachypetalum [see Appendix I]
→ Cerastium holosteoides Fr.
→ Cerastium brachypetalum subsp. roeseri (Boiss. & Heldr.) Nyman
→ Cerastium brachypetalum subsp. corcyrense (Möschl) P.D. Sell &
Whitehead
→ Cerastium illyricum subsp. brachiatum (Lonsing) Jalas
→ Cerastium holosteoides Fr.
→ Cerastium holosteoides Fr.
→ Cerastium holosteoides Fr.
→ Cerastium holosteoides Fr.
→ Cerastium semidecandrum L.
→ Cerastium holosteoides Fr.
→ Cerastium holosteoides Fr.
→ Ceratocephala falcata (L.) Pers.
→ Ceratocephala orthoceras DC. [see Appendix I]
→ Leonurus marrubiastrum L. [see Appendix I]
→ Epilobium dodonaei Vill.
→ Matricaria recutita L.
→ Erysimum senoneri (Reut.) Wettst.
→ Blackstonia perfoliata subsp. intermedia (Ten.) Zeltner
→ Chondrilla juncea L.
→ Cistus sintenisii Litard.
→ Cleome aurea Čelak.
→ Coronilla valentina subsp. glauca (L.) Batt.
→ Hippocrepis emerus (L.) Lassen
→ Corylus maxima Mill. [see Appendix I]
→ Anthemis macrantha Heuff. [see Appendix I]
→ Crepis sancta (L.) Bornm.
→ Crepis turcica Degen & Bald.
→ Crocus boryi J. Gay
→ Crocus boryi J. Gay
→ Crocus cancellatus subsp. mazziaricus (Herb.) B. Mathew
Willdenowia 46 – 2016
325
Crocus orphanidis Hook. f.
Crocus pholegandrius Orph.
Crocus spruneri Boiss. & Heldr.
Cucumis citrullus (L.) Ser.
→
→
→
→
Cyanus triumfettii (All.) Á. Löve & D. Löve
Cystopteris fragilis subsp. alpina (Lam.) Hartm.
Cystopteris fragilis subsp. dickieana (R. Sim) Hyl.
Dactylis marina Borrill
Daucus mauritanicus L.
Delphinium hybridum Willd., non L.
Dianthus campestris subsp. pallidiflorus (Ser.) Schmalh.
Dianthus campestris subsp. roseoluteus (Velen.) Stoj. & Acht.
Dianthus pancicii auct. fl. graec., non Velen.
Dianthus tristis Velen.
Dianthus tristis auct. fl. graec., non Velen.
Dinacrusa cretica (Cav.) G. Krebs
Dinacrusa hirsuta (L.) G. Krebs
Doronicum austriacum subsp. giganteum (Griseb.) Stoj. & Stef.
Draba bruniifolia subsp. heterocoma auct. fl. graec., non (Fenzl) Coode
& Cullen
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Draba spathulata (Láng) Sadler, non Bergeret nec Spreng.
Edraianthus australis (Wettst.) F.K. Mey.
Edraianthus tenuifolius auct. fl. graec., non (Waldst. & Kit.) A. DC.
Elymus athericus (Link) Kerguélen
Elymus campestris (Godr. & Gren.) Kerguélen
Elymus farctus subsp. bessarabicus (Săvul. & Rayss) Melderis
Elymus farctus subsp. rechingeri (Runemark) Melderis
Elymus rechingeri (Runemark) Runemark
Elytrigia intermedia subsp. barbulata (Schur) Á. Löve
Elytrigia intermedia subsp. graeca (Melderis) Á. Löve
Elytrigia pycnantha (Godr.) Á. Löve
Elytrigia rechingeri (Runemark) Holub
Emerus major Mill.
Emerus major subsp. emeroides (Boiss & Spruner) Soldano & F. Conti
Epilobium rosmarinifolium Haenke
Epilobium tetragonum subsp. tournefortii (Michalet) H. Lév.
Epipactis baumanniorum Ströhle
Epipactis exilis P. Delforge
Epipactis gracilis B. Baumann & H. Baumann, nom. illeg.
Epipactis helleborine subsp. aspromontana (Bartolo, Pulv. & Robatsch) H. Baumann & R. Lorenz
Epipactis helleborine subsp. aspromontana auct. fl. graec., non (Bartolo, Pulv. & Robatsch) H. Baumann & R. Lorenz
Epipactis leptochila subsp. neglecta auct. fl. graec., non Kümpel
Erophila boerhaavii (H.C. Hall) Dumort.
Erophila spathulata Láng
Erophila verna subsp. spathulata (Láng) Vollm.
Ervum monanthos L.
Eryngium serbicum auct. fl. graec., non Pančić
Eucalyptus rostrata Schltdl., non Cav.
Euphorbia lingulata Heuff.
Euphorbia spinosa Sm., non L.
Euphorbia villosa Willd.
Faba vulgaris Moench
Festuca danthonii Asch. & Graebn.
Festuca elatior subsp. pratensis (Huds.) Hack.
→
→
→
→
→
→
→
→
→
→
→
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→
→
→
→
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→
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Crocus tournefortii J. Gay
Crocus tournefortii J. Gay
Crocus cancellatus subsp. mazziaricus (Herb.) B. Mathew
Citrullus lanatus (Thunb.) Matsum. & Nakai [see Appendix I in Dimopoulos & al. 2013: 152]
Centaurea triumfettii All.
Cystopteris alpina (Lam.) Desv.
Cystopteris dickieana R. Sim
Dactylis glomerata subsp. hackelii (Asch. & Graebn.) Cif. & Giacom.
Daucus carota subsp. maximus (Desf.) Ball
Delphinium fissum Waldst. & Kit.
Dianthus pallidiflorus Ser. [see Appendix I]
Dianthus roseoluteus Velen. [see Appendix I]
Dianthus cruentus Griseb.
Dianthus pancicii Velen. [see Appendix I]
Dianthus cruentus Griseb.
Malva cretica L.
Malva setigera K.F. Schimp. & Spenn.
Doronicum austriacum Jacq.
Draba heterocoma Fenzl subsp. archipelagi (O.E. Schulz) Buttler
Draba boerhaavii (H.C. Hall) Raus
Edraianthus graminifolius (L.) A. DC. subsp. graminifolius
Edraianthus graminifolius (L.) A. DC. subsp. graminifolius
Elytrigia atherica (Link) Kerguélen
Elytrigia campestris (Godr. & Gren.) Kerguélen
Elytrigia bessarabica (Săvul. & Rayss) Prokudin
Elytrigia sartorii (Boiss. & Heldr.) Holub
Elytrigia sartorii (Boiss. & Heldr.) Holub
Elytrigia intermedia subsp. trichophora (Link) Á. Löve & D. Löve
Elytrigia obtusiflora subsp. graeca (Melderis) H. Scholz
Elytrigia atherica (Link) Kerguélen
Elytrigia sartorii (Boiss. & Heldr.) Holub
Hippocrepis emerus (L.) Lassen
Hippocrepis emerus subsp. emeroides (Boiss. & Spruner) Lassen
Epilobium dodonaei Vill.
Epilobium tournefortii Michalet
Epipactis persica subsp. gracilis W. Rossi
Epipactis persica subsp. gracilis W. Rossi
Epipactis persica subsp. gracilis W. Rossi
Epipactis leptochila subsp. aspromontana (Bartolo, Pulv. & Robatsch) Kreutz [see Appendix I]
→ Epipactis olympica Robatsch
→
→
→
→
→
→
→
→
→
→
→
→
→
Epipactis leptochila subsp. naousaensis (Robatsch) Kreutz
Draba boerhaavii (H.C. Hall) Raus
Draba boerhaavii (H.C. Hall) Raus
Draba boerhaavii (H.C. Hall) Raus
Vicia articulata Hornem.
Eryngium palmatum Pančić & Vis.
Eucalyptus camaldulensis Dehnh.
Euphorbia epithymoides L.
Euphorbia acanthothamnos Boiss.
Euphorbia illirica Lam.
Vicia faba L.
Vulpia ciliata Dumort.
Festuca pratensis Huds.
326
Dimopoulos & al.: Vascular plants of Greece: Supplement
Festuca fasciculata Asch. & Graebn.
Festuca ligustica (All.) C. Presl
Festuca muralis Kunth
Festuca myuros L.
Festuca vizzavonae auct. fl. graec., non Ronniger
Fibigia clypeata subsp. eriocarpa (DC.) Greuter
Fibigia eriocarpa (DC.) Boiss.
Ficaria chrysocephala (P.D. Sell) Galasso & al.
Ficaria ficariiformis (Rouy & Foucaud) A.W. Hill
Ficaria grandiflora Robert
Ficaria verna subsp. grandiflora (Robert) Hayek
Freyera stylosa (Boiss.) Boiss.
Fritillaria acmopetala Baker, non Boiss.
Fritillaria chia Nyman
Fritillaria citrina Baker
Fritillaria dasyphylla Baker
Fritillaria sieheana Hayek
Fritillaria tulipifolia Bory & Chaub., non M. Bieb.
Fritillaria tulipifolia var. dasyphylla Baker
Fumana glutinosa (L.) Boiss.
Fumana thymifolia subsp. laevis (Cav.) Molero & Rovira
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Vulpia fasciculata (Forssk.) Fritsch
Vulpia ligustica (All.) Link
Vulpia muralis (Kunth) Nees
Vulpia myuros (L.) C.C. Gmel.
Festuca alfrediana Foggi & Signorini
Fibigia clypeata (L.) Medik. subsp. clypeata
Fibigia clypeata (L.) Medik. subsp. clypeata
Ficaria verna subsp. chrysocephala (P.D. Sell) Stace
Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
Geocaryum stylosum (Boiss.) Engstrand
Fritillaria elwesii Boiss.
Fritillaria carica Rix subsp. carica
Fritillaria bithynica Baker
Fritillaria bithynica Baker
Fritillaria elwesii Boiss.
Fritillaria conica Boiss.
Fritillaria carica Rix subsp. carica
Fumana thymifolia (L.) Webb
Fumana laevis (Cav.) Pau
Fumaria schrammii (Asch.) Velen.
Fumaria vaillantii subsp. schrammii (Asch.) Nyman
Gagea bohemica subsp. saxatilis (Mert. & W.D.J. Koch) Pascher
Gagea rhodiaca A. Terracc.
Gagea saxatilis (Mert. & W.D.J. Koch) Schult. & Schult. f.
Galanthus corcyrensis (Beck) Stern
Galanthus olgae Boiss.
Galanthus reginae-olgae subsp. corcyrensis (Beck) Kamari
Galium apricum Sm.
Galium glabrum (L.) A. Kern.
Galium micranthum d’Urv., non Pursh
Galium mollugo subsp. pycnotrichum (Heinr. Braun) O. Schwarz
Galium recurvum DC.
Galium spruneri Jord.
Galium valantia Weber, non G. Gaertn. & al.
Gentianella austriaca auct. fl. graec., non (A. Kern. & Jos. Kern.)
Holub
Gentianella caucasea subsp. lutescens auct. fl. graec., non Velen.
Gentianella lutescens auct. fl. graec., non (Velen.) Holub
Geranium tuberosum subsp. thasium Stoj. & Kitan.
Gladiolus byzantinus Mill.
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Fumaria vaillantii Loisel.
Fumaria vaillantii Loisel.
Gagea bohemica (Zauschn.) Schult. & Schult. f.
Gagea rigida Boiss. & Spruner
Gagea bohemica (Zauschn.) Schult. & Schult. f.
Galanthus reginae-olgae Orph. subsp. reginae-olgae
Galanthus reginae-olgae Orph. subsp. reginae-olgae
Galanthus reginae-olgae Orph. subsp. reginae-olgae
Valantia aprica (Sm.) Tausch
Cruciata verna (Scop.) Gutermann & Ehrend.
Galium caminianum Schult. & Schult. f.
Galium album subsp. pycnotrichum (Heinr. Braun) Krendl
Galium caminianum Schult. & Schult. f.
Galium melanantherum Boiss.
Galium verrucosum Huds.
Gentianella bulgarica (Velen.) Holub
→
→
→
→
Hedypnois pendula Willd.
Helianthemum umbellatum auct. fl. graec., non L.
Helianthemum viride Ten.
Helichrysum sulfureum P. Candargy
Heracleum verticillatum Pančić
Hermione tazetta (L.) Haw.
Hieracium acutifolium Vill.
Hieracium alpicola auct. fl. graec., non Froel.
Hieracium alpicola subsp. petraeum Nägeli & Peter
Hieracium alpicola subsp. rhodopaeum (Griseb.) Zahn
Hieracium brachiatum DC.
Hieracium brachyphyllum Vuk., nom. inval.
Hieracium brevifolium subsp. brachyphyllum Zahn
Hieracium breviscapum Griseb, non DC.
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Gentianella bulgarica (Velen.) Holub
Gentianella bulgarica (Velen.) Holub
Geranium tuberosum L.
Gladiolus communis L. [see Appendix I in Dimopoulos & al. 2013:
153]
Hedypnois rhagadioloides (L.) F.W. Schmidt subsp. rhagadioloides
Halimium voldii Kit Tan & al.
Fumana laevis (Cav.) Pau
Helichrysum stoechas subsp. barrelieri (Ten.) Nyman
Heracleum sphondylium subsp. verticillatum (Pančić) Brummitt
Narcissus tazetta L.
Pilosella acutifolia (Vill.) Arv.-Touv.
Pilosella rhodopea (Griseb.) Szeląg
Pilosella rhodopea (Griseb.) Szeląg
Pilosella rhodopea (Griseb.) Szeląg
Pilosella acutifolia (Vill.) Arv.-Touv.
Hieracium brevifolium subsp. muraltae (Zahn) Greuter
Hieracium brevifolium subsp. muraltae (Zahn) Greuter
Pilosella rhodopea (Griseb.) Szeląg
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Hieracium breviscapum Hayek, non DC. nec Griseb.
Hieracium maranzae (Murr & Zahn) Prain
Hieracium maranzae auct. fl. graec., non (Murr & Zahn) Prain
Hieracium pallidum auct. fl. graec., non Biv.
Hieracium parnassi subsp. versutum (Griseb.) Zahn
Hieracium petraeum Friv., non Bluff & Fingerh.
Hieracium pratense Tausch
Hieracium racemosum subsp. todaroanum Zahn
Hieracium rhodopaeum Griseb.
Hieracium schmidtii subsp. labillardierei auct. fl. graec., non (Arv.Touv.) Greuter
Hieracium schmidtii subsp. pallidum auct. fl. graec., non (Biv.) O.
Bolòs & Vigo
Himantoglossum caprinum subsp. bolleanum (Siehe) H. Baumann &
R. Lorenz
Himantoglossum caprinum subsp. rumelicum H. Baumann & R. Lorenz
Honorius prasandrus (Griseb.) Holub
Hordeum bulbosum subsp. nodosum (L.) B.R. Baum
Hordeum nodosum L.
Hypericum dubium auct. fl. graec., non Leers
Hypericum ferrugineum Boiss. & Heldr., non Pursh
→
→
→
→
→
→
→
→
→
→
Hypericum hyssopifolium auct. fl. graec., non Chaix
Hypericum perforatum subsp. angustifolium (DC.) A. Fröhl., non
Gaudin
Hypericum richeri subsp. grisebachii auct. fl. graec., non (Boiss.) Nyman
Hypericum rochelii subsp. pseudotenellum (Vandas) Jordanov &
Kožuharov
Hypochaeris radicata subsp. heterocarpa (Moris) Arcang.
Hypochaeris radicata subsp. platylepis (Boiss.) Jahand. & Maire
Hypochoeris setosa Formánek
Inula aspera Poir.
Iris falcata Babal. & Papan., non Tausch
Isoetes echinospora auct. fl. graec., non Durieu
Isoetes histrix var. subinermis Motelay & Vendryès
Isoetes setacea auct. fl. graec., non Lam.
Isoetes sicula Tod.
Isoetes subinermis (Gennari) Cesca & Peruzzi
Jovibarba heuffelii subsp. glabra (Beck & Szyszyl.) Holub
Juncus phalereus Gand.
Juniperus phoenicea auct. fl. graec., non L.
Juniperus phoenicea subsp. eumediterranea P. Lebreton & Thivend
Juniperus phoenicea subsp. turbinata (Guss.) Nyman
Jurinea arachnoidea auct. balc., non Bunge
Kali australe (L.) Akhani & Roalson
Kali soda Moench, non (L.) Scop.
Kali turgidum (Dumort.) Gutermann
Kohlrauschia glumacea (Bory & Chaub.) Hayek
Lagoseris nemausensis K. Malý, non Crepis nemausensis Gouan
Lamium ochroleucum Link
Lapsana stellata L.
Leiotulus aureus (Sm.) Pimenov & Ostr.
Leiotulus involucratus (Boiss. & Spruner) Pimenov & Ostr.
Leontodon asper (Waldst. & Kit.) Poir., non Forssk.
Leontodon asper var. haussknechtii (Hausskn.) Halácsy
Leontodon asper var. saxatilis (Ten.) Halácsy
Leontodon asper var. setulosus Halácsy
→ Hypericum tymphresteum Boiss. & Spruner
→ Hypericum perforatum subsp. veronense (Schrank) Ces.
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Hieracium scapigerum Boiss. & al.
Hieracium sabaudum L.
Hieracium neoplatyphyllum Gottschl.
Hieracium schmidtii Tausch
Hieracium parnassi Fr.
Pilosella rhodopea (Griseb.) Szeląg
Pilosella caespitosa (Dumort.) P.D. Sell & C. West [see Appendix I]
Hieracium racemosum subsp. crinitum (Sm.) Rouy
Pilosella rhodopea (Griseb.) Szeląg
Hieracium schmidtii subsp. samothracis (Ade & Schack) Gottschl.
→ Hieracium schmidtii subsp. creticum (Zahn) Greuter
→ Himantoglossum jankae Somlyay & al.
→
→
→
→
→
→
Himantoglossum jankae Somlyay & al.
Ornithogalum nutans L.
Hordeum bulbosum L.
Hordeum bulbosum L.
Hypericum maculatum subsp. immaculatum (Murb.) A. Fröhl.
Hypericum spruneri Boiss.
→ Hypericum barbatum Jacq.
→ Hypericum rochelii Griseb. & Schenk
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Hypochaeris radicata L.
Hypochaeris radicata L.
Leontodon biscutellifolius DC.
Inula salicina subsp. aspera (Poir.) Hayek
Iris attica Boiss. & Heldr.
Isoetes heldreichii Wettst.
Isoetes gymnocarpa (Gennari) A. Braun
Isoetes heldreichii Wettst.
Isoetes gymnocarpa (Gennari) A. Braun
Isoetes gymnocarpa (Gennari) A. Braun
Sempervivum heuffelii Schott
Juncus heldreichianus Parl. subsp. heldreichianus
Juniperus turbinata Guss.
Juniperus phoenicea L. [see Appendix I]
Juniperus turbinata Guss.
Jurinea polycephala Formánek [see Appendix I]
Salsola kali L. [see Appendix I in Dimopoulos & al. 2013: 152]
Salsola kali L. [see Appendix I in Dimopoulos & al. 2013: 152]
Salsola kali L. [see Appendix I in Dimopoulos & al. 2013: 152]
Petrorhagia glumacea (Bory & Chaub.) P.W. Ball & Heywood
Crepis sancta (L.) Bornm.
Lamium moschatum Mill.
Rhagadiolus stellatus (L.) Gaertn.
Malabaila aurea (Sm.) Boiss.
Malabaila involucrata Boiss. & Spruner
Leontodon biscutellifolius DC.
Leontodon biscutellifolius DC.
Leontodon crispus Vill.
Leontodon biscutellifolius DC.
328
Dimopoulos & al.: Vascular plants of Greece: Supplement
Leontodon asper var. typicus Halácsy
Leontodon asperrimus auct. fl. graec., non (Willd.) Endl.
Leontodon crispus subsp. asper (Waldst. & Kit.) Rohlena
Leontodon crispus subsp. asperrimus auct. fl. graec., non (Willd.)
Finch & P.D. Sell
Leontodon crispus subsp. rossianus (Degen & Lengyel) Hayek
Leontodon hastilis L.
Leontodon haussknechtii Hausskn.
Leontodon longirostris (Finch & P.D. Sell) Talavera
Leontodon saxatilis (Ten.) Rchb., non Lam.
Leontodon taraxacoides subsp. longirostris Finch & P.D. Sell
Limonium rhodense M.B. Crespo & Pena-Martín
Linaria parviflora (Jacq.) Halácsy
Lolium arundinaceum subsp. orientale (Hack.) G.H. Loos
Lolium pratense (Huds.) Darbysh.
Loncomelos pyramidale auct. fl. graec., non (L.) Raf.
Lotus graecus L.
Lotus herbaceus (Vill.) Jauzein
Lotus rectus L.
Luzula alpinopilosa subsp. obscura auct. fl. graec., non S.E. Fröhner
Lyonnetia rigida DC.
→
→
→
→
Leontodon crispus Vill.
Leontodon biscutellifolius DC.
Leontodon biscutellifolius DC.
Leontodon biscutellifolius DC.
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Leontodon crispus Vill.
Leontodon hispidus L. subsp. hispidus
Leontodon biscutellifolius DC.
Leontodon saxatilis subsp. rothii Maire
Leontodon crispus Vill.
Leontodon saxatilis subsp. rothii Maire
Limonium ammophilon (Papatsou & Phitos) Domina
Linaria simplex Desf.
Festuca arundinacea subsp. orientalis (Hack.) K. Richt.
Festuca pratensis Huds.
Ornithogalum brevistylum Wolfner
Dorycnium graecum (L.) Ser.
Dorycnium herbaceum Vill.
Dorycnium rectum (L.) Ser.
Luzula alpinopilosa subsp. deflexa (Kožuharov) Kirschner
Anthemis rigida Heldr.
Lysimachia arvensis subsp. latifolia (L.) Peruzzi
Lysimachia arvensis subsp. parviflora (L.) Peruzzi
Majorana cretica Mill.
Majorana leptoclados Rech. f.
→
→
→
→
Majorana sipylea (L.) Kostel.
Malva althaeoides auct. fl. graec., non Cav.
Malva linnaei M.F. Ray
Malva rotundifolia L., nom. rej.
Marrubium ×remotum Schult.
Maruta cotula (L.) DC.
Matricaria chamomilla L.
Matthiola sinuata subsp. glandulosa (Vis.) Vierh.
Medicago blancheana Boiss. subsp. blancheana
Medicago blancheana var. bonarotiana (Arcang.) Arcang.
Medicago rotata subsp. bonarotiana (Arcang.) Ponert
Mentha brachyodonta P. Candargy
Mentha epicaulos P. Candargy
Mentha noeana Boiss.
Mentha spicata subsp. tomentosa (Briq.) Harley
Mentha tomentosa subsp. condensata Briq.
Minuartia diljanae Panov
Minuartia graminifolia subsp. graminifolia auct. fl. graec., non (Ard.)
Jáv.
Minuartia hybrida subsp. tenuifolia (L.) Kerguélen
Minuartia mutabilis subsp. balcanica Panov
Myosurus breviscapus Huth
Myosurus heldreichii H. Lév.
Myosurus minimus subsp. heldreichii (H. Lév.) O. Bolòs & Vigo
Narcissus aphyllus Sieber
Narcissus corcyrensis (Herb.) Nyman
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Anagallis arvensis L.
Anagallis parviflora Hoffmanns. & Link
Origanum onites L.
Origanum ×minoanum P.H. Davis [see Appendix I in Dimopoulos &
al. 2013: 154]
Origanum sipyleum L.
Malva cretica Cav. subsp. cretica
Malva multiflora (Cav.) Soldano & al.
Malva pusilla Sm.
Marrubium ×paniculatum Desr. [see Appendix I]
Anthemis cotula L.
Matricaria recutita L., nom. cons.
Matthiola sinuata (L.) R. Br.
Medicago ×blancheana Boiss. [see Appendix I]
Medicago bonarotiana Arcang.
Medicago bonarotiana Arcang.
Mentha spicata subsp. condensata (Briq.) Greuter & Burdet
Mentha spicata subsp. condensata (Briq.) Greuter & Burdet
Mentha longifolia (L.) Huds. subsp. longifolia
Mentha spicata subsp. condensata (Briq.) Greuter & Burdet
Mentha spicata subsp. condensata (Briq.) Greuter & Burdet
Minuartia rumelica Panov [see Appendix III]
Minuartia graminifolia subsp. brachypetala Kamari
Narcissus serotinus auct. fl. graec., non L.
Narcissus tazetta subsp. corcyrensis (Herb.) Baker
Nectaroscilla hyacinthoides (L.) Parl.
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→
→
→
→
→
→
→
Minuartia hybrida (Vill.) Schischk.
Minuartia bosniaca (Beck) K. Malý
Myosurus sessilis S. Watson
Myosurus sessilis S. Watson
Myosurus sessilis S. Watson
Narcissus obsoletus (Haw.) Steud.
Narcissus ×corcyrensis (Herb.) Nyman [see Appendix I in Dimopoulos & al. 2013: 149]
→ Narcissus obsoletus (Haw.) Steud.
→ Narcissus ×corcyrensis (Herb.) Nyman [see Appendix I in Dimopoulos & al. 2013: 149]
→ Scilla hyacinthoides L.
Willdenowia 46 – 2016
329
→ Nepeta scordotis L.
→ Alyssum murale Waldst. & Kit.
→ Olea europaea L. subsp. europaea [var. sylvestris (Mill.) Lehr, see
Appendix III in Dimopoulos & al. 2013: 286]
Olea europaea subsp. sylvestris (Mill.) Hegi
→ Olea europaea L. subsp. europaea [var. sylvestris (Mill.) Lehr, see
Appendix III in Dimopoulos & al. 2013: 286]
Oloptum miliaceum (L.) Röser & Hamasha
→ Piptatherum miliaceum (L.) Coss.
Oncostema peruviana (L.) Speta
→ Scilla peruviana L. [see Appendix I]
Ophrys aeoli P. Delforge
→ Ophrys holoserica subsp. andria (P. Delforge) Faurh.
Ophrys andria P. Delforge
→ Ophrys holoserica subsp. andria (P. Delforge) Faurh.
Ophrys andria subsp. halkionis (G. Kretzschmar & H. Kretzschmar) → Ophrys holoserica subsp. andria (P. Delforge) Faurh.
Kreutz
Ophrys apifera subsp. arachnites (L.) Hook. f.
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys arachnites (L.) Reichard
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys aranifera subsp. aesculapii (Renz) Soó
→ Ophrys sphegodes subsp. aesculapii (Renz) J.J. Wood
Ophrys aranifera subsp. parnassica Soó
→ Ophrys ferrum-equinum Desf. subsp. ferrum-equinum
Ophrys aranifera subsp. renzii Soó
→ Ophrys sphegodes subsp. aesculapii (Renz) J.J. Wood
Ophrys aranifera subsp. tommasinii (Vis.) E.G. Camus
→ Ophrys sphegodes subsp. litigiosa (E.G. Camus) Bech. [see Appendix I]
Ophrys aranifera var. atrata Rchb. f.
→ Ophrys scolopax subsp. atrata (Rchb. f.) A. Bolòs [see Appendix I]
Ophrys ariadnae Paulus
→ Ophrys cretica subsp. karpathensis E. Nelson
Nepeta tomentosa Benth., non (Gilib.) Vitman
Odontarrhena muralis (Waldst. & Kit.) Endl.
Olea europaea subsp. oleaster (Hoffmanns. & Link) Negodi
Ophrys atrata Lindl., non L.
Ophrys balcanica Soó
Ophrys bilunulata subsp. sancti-isidorii (A. Saliaris & al.) Paulus
Ophrys candica Greuter & al.
→
→
→
→
Ophrys candica (Soó) H. Baumann & Künkele, nom. inval.
→
Ophrys candica subsp. cytherea B. Baumann & H. Baumann
→
Ophrys candica subsp. lacaena (P. Delforge) Kreutz
→
Ophrys candica subsp. minoa C. Alibertis & A. Alibertis
→
Ophrys cephaloniensis Paulus
Ophrys chiosica P. Delforge & al.
Ophrys colossaea P. Delforge
Ophrys cornuta subsp. balcanica Soó
Ophrys cretica subsp. ariadnae (Paulus) H. Kretzschmar
Ophrys cretica subsp. naxia E. Nelson
Ophrys episcopalis Poir.
Ophrys ferrum-equinum subsp. aegaea (Kalteisen & H.R. Reinhard) H.
Baumann & R. Lorenz
Ophrys ferrum-equinum subsp. convexa B. Baumann & H. Baumann
Ophrys forestieri (Rchb. f.) Lojac.
Ophrys fuciflora (Crantz) Rchb. f., non (F.W. Schmidt) Moench
Ophrys fuciflora subsp. andria (Delforge) Faurh.
Ophrys fuciflora subsp. bornmuelleri (M. Schulze) B. Willing & E.
Willing
Ophrys fuciflora subsp. candica Soó, nom. inval.
→
→
→
→
→
→
→
→
Ophrys fuciflora (F.W. Schmidt) Moench subsp. fuciflora
Ophrys fuciflora subsp. grandiflora (H. Fleischm. & Soó) Faurh.
→
→
Ophrys fuciflora subsp. maxima (H. Fleischm.) Soó
Ophrys fusca subsp. fleischmannii (Hayek) Soó
Ophrys fusca subsp. phaseliana (D. Rückbr. & U. Rückbr.) Kreutz
→
→
→
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→
→
→
→
→
→
Ophrys scolopax subsp. atrata (Rchb. f.) A. Bolòs [see Appendix I]
Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys fusca subsp. fusca
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys cretica subsp. karpathensis E. Nelson
Ophrys cretica subsp. karpathensis E. Nelson
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys argolica subsp. aegaea (Kalteisen & H.R. Reinhard) H.A.
Pedersen & Faurh.
Ophrys ferrum-equinum Desf. subsp. ferrum-equinum
Ophrys fusca Link subsp. fusca
Ophrys holoserica (Burm. f.) Greuter
Ophrys holoserica subsp. andria (P. Delforge) Faurh.
Ophrys holoserica subsp. bornmuelleri (M. Schulze) H. Sund. [see
Appendix I]
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica subsp. grandiflora (H. Fleischmann & Soó) Faurh.
[see Appendix I]
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys omegaifera subsp. fleischmannii (Hayek) Del Prete
Ophrys fusca Link subsp. fusca
330
Dimopoulos & al.: Vascular plants of Greece: Supplement
Ophrys grammica subsp. knossia A. Alibertis
→ Ophrys sphegodes subsp. mammosa (Desf.) E. Nelson
Ophrys halia Paulus
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys heldreichii subsp. pusilla B. Baumann & H. Baumann
→ Ophrys scolopax subsp. heldreichii (Schltr.) E. Nelson
Ophrys heldreichii subsp. schlechteriana Soó
→ Ophrys scolopax subsp. heldreichii (Schltr.) E. Nelson
Ophrys helios Kreutz
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys hellenica Devillers & Devillers-Tersch.
→ Ophrys lutea Link subsp. galilaea (H. Fleischm. & Bornm.) Soó
Ophrys herae subsp. janrenzii (M. Hirth) M. Hirth
→ Ophrys sphegodes subsp. mammosa (Desf.) E. Nelson
Ophrys holoserica subsp. aeoli (P. Delforge) Kreutz
→ Ophrys holoserica subsp. andria (P. Delforge) Faurh.
Ophrys holoserica subsp. candica (Soó) Renz & Taubenheim, nom. → Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen
illeg.
Faurh.
Ophrys holoserica subsp. colossaea (P. Delforge) Kreutz
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica subsp. episcopalis (Poir.) Kreutz
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica subsp. graeca B. Baumann & H. Baumann
→ Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen
Faurh.
Ophrys holoserica subsp. halia (Paulus) Kreutz
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica subsp. helios (Kreutz) Kreutz
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica subsp. lacaena (P. Delforge) H. Baumann & R. → Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen
Lorenz
Faurh.
Ophrys holoserica subsp. lyciensis (Paulus & al.) H. Baumann & R. → Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen
Lorenz
Faurh.
Ophrys holoserica subsp. maxima (H. Fleischm.) Greuter
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica subsp. taloniensis Kreutz
Ophrys holosericea Greuter, orth. var.
Ophrys janrenzii M. Hirth
Ophrys knossia (A. Alibertis) P. Delforge
Ophrys kotschyi subsp. ariadnae (Paulus) Faurh.
Ophrys lacaena P. Delforge
→
→
→
→
→
→
Ophrys lyciensis Paulus & al.
→
Ophrys malvasiana S. Hertel & Weyland
Ophrys mammosa subsp. epirotica (Renz) H. Baumann & R. Lorenz
Ophrys mammosa subsp. gortynia (H. Baumann & Künkele) B. Baumann & R. Lorenz
Ophrys maxima (H. Fleischm.) Paulus & Gack
Ophrys mimnolea auct. fl . graec., non O. Schwarz
Ophrys mimnolea O. Schwarz
→
→
→
&
&
&
&
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys holoserica (Burm. f.) Greuter
Ophrys sphegodes subsp. mammosa (Desf.) E. Nelson
Ophrys sphegodes subsp. mammosa (Desf.) E. Nelson
Ophrys cretica subsp. karpathensis E. Nelson
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica subsp. candica (Greuter & al.) H.A. Pedersen &
Faurh.
Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys sphegodes subsp. epirotica (Renz) Gölz & H.R. Reinhard
Ophrys sphegodes subsp. gortynia H. Baumann & Künkele
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
→ Ophrys reinholdii H. Fleischm.
→ Ophrys reinholdii subsp. straussii (H. Fleischm.) E. Nelson [see Appendix I in Dimopoulos & al. 2013: 155, 287]
Ophrys minoa (C. Alibertis & A. Alibertis) P. Delforge
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys naxensis Rech.
→ Ophrys cretica subsp. karpathensis E. Nelson
Ophrys neglecta Parl.
→ Ophrys tenthredinifera Willd.
Ophrys oestrifera auct. fl. graec., non M. Bieb. nec Rchb.
→ Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys oestrifera M. Bieb.
→ Ophrys apifera Huds.
Ophrys oestrifera Rchb., non M. Bieb.
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys oestrifera subsp. lemnosiana B. Baumann & H. Baumann
→ Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys oestrifera subsp. minutula (Gölz & H.R. Reinhard) Kreutz
→ Ophrys scolopax Cav. subsp. cornuta (Steven) E.G. Camus
Ophrys oestrifera subsp. stavri Kalog. & al.
→ Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys penelopeae Paulus
→ Ophrys lutea Link subsp. lutea
Ophrys phaseliana D. Rückbr. & U. Rückbr.
→ Ophrys fusca Link subsp. fusca
Ophrys praemelena S. Hertel & Presser
→ Ophrys lutea subsp. melena Renz
Ophrys renzii Soó
→ Ophrys sphegodes subsp. aesculapii (Renz) J.J. Wood
Ophrys saliarisii Paulus & M. Hirth
→ Ophrys holoserica (Burm. f.) Greuter subsp. holoserica
Ophrys sancti-isidorii (A.Saliaris & al.) P. Delforge
→ Ophrys fusca Link subsp. fusca
Ophrys sitiaca Paulus & al.
→ Ophrys ×brigittae H. Baumann [see Appendix I in Dimopoulos & al.
2013: 286]
Ophrys sphegodes subsp. atrata auct. fl. graec., non (Rchb. f.) A. Bolòs. → Ophrys sphegodes Mill. subsp. sphegodes
Ophrys sphegodes subsp. janrenzii (M. Hirth) Kreutz
→ Ophrys sphegodes subsp. mammosa (Desf.) E. Nelson
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Willdenowia 46 – 2016
Ophrys sphegodes subsp. zeusii (M. Hirth) Kreutz
Ophrys stavri (Kalog. & al.) P. Delforge
Ophrys subfusca (Rchb.f.) Hausskn.
Ophrys subfusca subsp. archimedea (P. Delforge & M. Walravens)
Kreutz
Ophrys subfusca subsp. blithoperta (Paulus & Gack) Kreutz
331
→
→
→
→
Ophrys sphegodes Mill. subsp. sphegodes
Ophrys scolopax subsp. cornuta (Steven) E.G. Camus
Ophrys fusca Link subsp. fusca
Ophrys lutea subsp. galilaea (H. Fleischm. & Bornm.) Soó
→ Ophrys fusca Link subsp. blithoperta (Paulus & Gack) Faurh. & H.A.
Pedersen
Ophrys subfusca subsp. cinereophila (Paulus & Gack) Kreutz
→ Ophrys fusca Link subsp. cinereophila (Paulus & Gack) Faurh.
Ophrys subfusca subsp. persephonae (Paulus) Kreutz
→ Ophrys fusca Link subsp. fusca
Ophrys tenthredinifera subsp. sanctae-marcellae Saliaris & al.
→ Ophrys tenthredinifera Willd.
Ophrys thesei P. Delforge
→ Ophrys holoserica subsp. andria (P. Delforge) Faurh.
×Orchiaceras bivonae (Tod.) Soó
→ Orchis ×bivonae Tod. [see Appendix I]
×Orchiaceras orphanidesii E.G. Camus
→ Orchis ×bivonae Tod. [see Appendix I]
Orchis anatolica subsp. sezikiana (B. Baumann & H. Baumann) Kreutz → Orchis ×sezikiana B. Baumann & H. Baumann [see Appendix I]
Orchis ×attica Hausskn.
→ Orchinea ×attica (Hausskn.) F.N. Vázquez [see Appendix I]
Orchis ×bicknellii E.G. Camus
→ Anacamptis ×parvifolia nothosubsp. bicknellii (E.G. Camus) H.
Kretzschmar & al. [see Appendix I]
Orchis ×corcyrensis Soó
→ Orchinea ×attica (Hausskn.) F.N. Vázquez [see Appendix I]
Orchis fuciflora Crantz
→ Ophrys holoserica (Burm. f.) Greuter
Orchis ×gennarii Rchb. f.
→ Anacamptis ×gennarii (Rchb. f.) H. Kretzschmar & al. [see Appendix I]
→ Anacamptis ×gerakarionis (Faller & Kreutz) H. Kretzschmar & al.
[see Appendix I in Dimopoulos & al. 2013: 154]
Orchis ×hermaniana C. Alibertis & A. Alibertis
→ ×Orchinea hermaniana (C. Alibertis & A. Aliberti) J.M.H. Shaw
Orchis holoserica Burm. f.
→ Ophrys holoserica (Burm. f.) Greuter
Orchis ×ionica Renz
→ Ophrys ×rechingeri Soó [see Appendix I]
Orchis lactea subsp. minuscula A. Alibertis
→ Neotinea lactea (Poir.) R.M. Bateman & al.
Orchis mascula subsp. signifera (Vest) Soó
→ Orchis mascula subsp. speciosa (Mutel) Hegi [see Appendix I]
Orchis morio subsp. boryi (Rchb. f.) Soó
→ Anacamptis boryi (Rchb. f.) R.M. Bateman & al.
Orchis papilionacea subsp. balcanica H. Baumann & R. Lorenz
→ Anacamptis papilionacea (L.) R.M. Bateman & al. subsp. papilionacea
Orchis quadripunctata subsp. sezikiana (B. Baumann & H. Baumann) → Orchis ×sezikiana B. Baumann & H. Baumann [see Appendix I]
H. Baumann & R. Lorenz
Orchis ×sciathia Biel
→ Anacamptis ×sciathia (Biel) H. Kretzschmar & al. [see Appendix I]
Orchis sezikiana B. Baumann & H. Baumann
→ Orchis ×sezikiana B. Baumann & H. Baumann [see Appendix I]
Orchis tridentata subsp. angelica A. Alibertis
→ Neotinea tridentata (Scop.) R.M. Bateman & al. subsp. tridentata
×Orchiserapias ligustica E.G. Camus
→ ×Serapicamptis ligustica (E.G. Camus) J.M.H. Shaw [see Appendix I]
×Orchiserapias purpurea E.G. Camus
→ ×Serapicamptis rousii (Dupuy) J.M.H. Shaw [see Appendix I]
Ornithogalum alatum Turrill
→ Ornithogalum wiedemannii Boiss.
Ornithogalum atticum sensu Fl. Aeg., non Boiss. & Heldr.
→ Ornithogalum montanum Cirillo
Ornithogalum byzantinum Ten., non Azn.
→ Ornithogalum montanum Cirillo
Ornithogalum collinum W.D.J Koch & sensu Fl. Eur., non Guss.
→ Ornithogalum gussonei Ten.
Ornithogalum costatum Zahar.
→ Ornithogalum gussonei Ten.
Ornithogalum cuspidatum Griseb., non Bertol.
→ Ornithogalum montanum Cirillo
Ornithogalum exscapum var. collinum (Guss.) Stearn
→ Ornithogalum collinum Guss. subsp. collinum
Ornithogalum fimbriatum subsp. atticum (Boiss. & Heldr.) Nyman
→ Ornithogalum atticum Boiss. & Heldr.
Ornithogalum graecum Zahar.
→ Ornithogalum collinum Guss.
Ornithogalum graecum subsp. ciliolatum Zahar.
→ Ornithogalum collinum Guss. subsp. collinum
Ornithogalum graecum subsp. ionicum Zahar.
→ Ornithogalum collinum Guss. subsp. collinum
Ornithogalum macedonicum Velen.
→ Ornithogalum armeniacum Baker
Ornithogalum montanum subsp. atticum (Boiss. & Heldr.) Nyman
→ Ornithogalum atticum Boiss. & Heldr.
Ornithogalum nutans subsp. prasandrum (Griseb.) K. Richt.
→ Ornithogalum nutans L.
Ornithogalum pascheanum auct. fl. graec., non Speta
→ Ornithogalum pumilum Zahar.
Ornithogalum prasandrum Griseb.
→ Ornithogalum nutans L.
Ornithogalum sandalioticum (Tornad. & Garbari) Zahar.
→ Ornithogalum corsicum Jord. & Fourr. [see Appendix III]
Ornithogalum tenuifolium auct. fl. graec., non Guss.
→ Ornithogalum collinum Guss.
Ornithogalum tenuifolium Guss., non F. Delaroche
→ Ornithogalum gussonei Ten.
Orobanche chassia Formánek
→ Orobanche alba Willd.
Orchis ×gerakanionis Faller & Kreutz
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Orobanche coelestis auct. fl. graec., non (Reut.) Beck
Orobanche grandiflora Bory & Chaub.
Orobanche hyalina Reut., non Gren. & Godr.
Orobanche ramosa L.
Paeonia mascula subsp. arietina (G. Anderson) Cullen & Heywood
Paeonia mascula subsp. arietina auct. fl. graec., non (G. Anderson)
Cullen & Heywood
Paronychia echinata DC., non Lam.
Pastinaca pimpinellifolia Bory & Chaub.
Peplis borysthenica Schrank
Peplis tubulosa P. Candargy
Persicaria dubia (A. Braun) Fourr.
Persicaria salicifolia (Willd.) Assenov, non Gray
Petrosedum anopetalum (DC.) Grulich
Petrosedum sediforme (Jacq.) Grulich
Pilosella acutifolia auct. fl. graec., non (Vill.) Arv.-Touv.
Pilosella alpicola auct. fl. graec., non (Froel.) F. W. Schultz & Sch. Bip.
Pimpinella depressa (Spreng.) DC.
Pimpinella tragium subsp. lithophila (Schischk.) Tutin
Pimpinella tragium var. typica Halácsy
Pisum graecum Quézel & Contandr.
Plantago lanceolata subsp. lanuginosa (Mert. & W.D.J. Koch) Ces.
Plantago lanceolata subsp. sphaerostachya (Mert. & W.D.J. Koch)
Hayek
Plantago uliginosa F.W. Schmidt
Platanthera holmboei H. Lindb.
Poa annua subsp. exilis (Freyn) Asch. & Graebn.
Poa exilis (Freyn) Murb.
Poa hackelii auct. fl. graec., non Post
Poa sinaica subsp. graeca H. Scholz
Polygonatum latifolium (Jacq.) Desf.
Populus nigra subsp. thevestina (Dode) Maire
Dimopoulos & al.: Vascular plants of Greece: Supplement
→
→
→
→
→
→
Phelipanche mutelii (F.W. Schultz) Pomel
Orobanche crenata Forssk.
Orobanche minor Sm.
Phelipanche ramosa (L.) Pomel [see Appendix I]
Paeonia arietina G. Anderson [see Appendix I]
Paeonia mascula (L.) Mill. subsp. mascula
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Paronychia echinulata Chater
Malabaila aurea (Sm.) Boiss.
Lythrum borysthenicum (Schrank) Litv.
Lythrum borysthenicum (Schrank) Litv.
Persicaria mitis (Schrank) Assenov, nom. cons.
Persicaria decipiens (R. Br.) K.L. Wilson
Sedum ochroleucum Chaix
Sedum sediforme (Jacq.) Pau
Pilosella sphaerocephala (Rchb.) F.W. Schultz & Sch. Bip.
Pilosella rhodopea (Griseb.) Szeląg
Pimpinella tragium subsp. depressa (Spreng.) Tutin
Pimpinella tragium Vill. subsp. tragium
Pimpinella tragium Vill. subsp. tragium
Lathyrus grandiflorus Sm.
→ Plantago lanceolata L.
→ Plantago lanceolata L.
→
→
→
→
→
→
→
→
Plantago major subsp. intermedia (Gilib.) Lange
Platanthera chlorantha subsp. holmboei (H. Lindb.) J.J. Wood
Poa infirma Kunth
Poa infirma Kunth
Poa pelasgis H. Scholz
Poa pelasgis H. Scholz
Polygonatum hirtum (Poir.) Pursh
Populus thevestina Dode [see Appendix I in Dimopoulos & al. 2013:
157]
Potentilla goulandrii Rech. f.
→ Potentilla tridentula Velen.
Potentilla poetarum Boiss. & Spruner
→ Potentilla speciosa Willd. subsp. speciosa
Pseudorchis albida subsp. albida auct. fl. graec., non (L.) Á. Löve & → Pseudorchis albida subsp. tricuspis (Beck) E. Klein
D. Löve
Pseudoturritis turrita (L.) Al-Shehbaz
→ Arabis turrita L.
Quercus ×braunii Borbás
→ Quercus ×szechenyana Borbás [see Appendix I]
Quercus ×topaliae A. Camus
→ Quercus ×szechenyana Borbás [see Appendix I]
Ranunculus binatus Rchb.
→ Ranunculus auricomus L. s.l.
Ranunculus ficaria subsp. bulbilifer Lambinon
→ Ficaria verna Huds. subsp. verna [see Appendix I in Dimopoulos &
al. 2013: 156]
Ranunculus ficaria subsp. ficariiformis Rouy & Foucaud
→ Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
Ranunculus ficaria subsp. grandiflorus (Robert) Cout.
→ Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
Ranunculus ficariiformis (Rouy & Foucaud) Beck
→ Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
Ranunculus ficarioides Bory & Chaub.
→ Ficaria ficarioides (Bory & Chaub.) Halácsy
Ranunculus nemorosus DC.
→ Ranunculus polyanthemos subsp. nemorosus (DC.) Schübl. & G.
Martens [see Appendix I]
Ranunculus parvulus L.
→ Ranunculus sardous Crantz
Ranunculus peloponnesiacus Boiss.
→ Ranunculus gracilis E.D. Clarke
Ranunculus philonotis Ehrh.
→ Ranunculus sardous Crantz
Ranunculus sardous subsp. balcanicus Kümmerle & Jáv.
→ Ranunculus sardous Crantz
Ranunculus serpens auct. fl. graec., non Schrank
→ Ranunculus polyanthemos L.
Ranunculus serpens subsp. nemorosus (DC.) G. López
→ Ranunculus polyanthemos subsp. nemorosus (DC.) Schübl. & G.
Martens [see Appendix I]
Ranunculus testiculatus auct. fl. graec., non Crantz
→ Ceratocephala falcata (L.) Cramer
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Rhinanthus mediterraneus (Sterneck) Sennen
Rubus ionicus Utsch
Rubus paramethystinus Formánek
Sagina melitensis Duthie
Salicornia emerici auct. fl. graec., non Douval-Jouve
Salvia grandiflora subsp. aegaea (Bornm.) Rech. f.
Salvia grandiflora subsp. rotundifolia (Vis.) Rech. f.
Salvia sylvestris L.
Sanguisorba minor subsp. spachiana (Coss.) Cout.
Satureja hispida (Benth.) Nyman, non Ehrh.
Satureja pilosa subsp. origanita Dardioti & Kokkini
Sedum albescens auct. fl. graec., non Haw.
Sempervivum erythraeum Velen.
Sempervivum marmoreum subsp. reginae-amaliae (Baker) Zonn.
Sempervivum marmoreum subsp. reginae-amaliae (Boiss.) Zonn.
Sempervivum marmoreum subsp. reginae-amaliae Maire & Petitm.
Sempervivum reginae-amaliae Baker
Sempervivum reginae-amaliae Halácsy, non Baker nec Boiss.
→ Ceratocephala orthoceras DC. [see Appendix I]
→ Ranunculus polyanthemos subsp. nemorosus (DC.) Schübl. & G.
Martens [see Appendix I]
→ Rhinanthus pumilus (Sterneck) Pau
→ Rubus sanctus Schreb.
→ Rubus sanctus Schreb.
→ Sagina apetala Ard.
→ Salicornia procumbens Sm. subsp. procumbens
→ Salvia tomentosa Mill.
→ Salvia tomentosa Mill.
→ Salvia ×sylvestris L [see Appendix I in Dimopoulos & al. 2013: 154]
→ Sanguisorba verrucosa (G. Don) Ces.
→ Micromeria hispida Benth.
→ Satureja pilosa Velen.
→ Sedum ochroleucum Chaix
→ Sempervivum marmoreum Griseb. subsp. marmoreum
→ Sempervivum heuffelii Schott
→ Sempervivum marmoreum Griseb. subsp. marmoreum
→ Sempervivum marmoreum Griseb. subsp. marmoreum
→ Sempervivum heuffelii Schott
→ Sempervivum marmoreum Griseb. subsp. marmoreum
Sempervivum tectorum subsp. reginae-amaliae Maire & Petitm.
Senecio aquaticus auct. fl. graec., non Hill
Senecio doronicum subsp. transylvanicus (Boiss.) Nyman
Senecio transylvanicus Boiss.
Serapias columnae Aurnier
Serapias columnae (Rchb. f.) H. Fleischm., non Aurnier
Serapias cordigera subsp. cycladum (H. Baumann & Künkele) Kreutz
→
→
→
→
→
→
→
Serapias cycladum H. Baumann & Künkele
→
Serapias ionica H. Baumann & Künkele
→
Serapias laxiflora var. columnae Rchb. f., non S. columnae Aurnier
Serapias longipetala (Ten.) Pollini
Serapias neglecta subsp. apulica auct. fl. graec., non Landwehr
→
→
→
Serapias neglecta subsp. neglecta auct. fl. graec., non De Not.
→
Serapias orientalis subsp. moreana H. Baumann & R. Lorenz
→
Serapias parviflora subsp. columnae (Asch. & Graebn.) Soó
Serapias parviflora var. columnae Asch. & Graebn., non S. columnae
Aurnier nec (Rchb. f.) H. Fleischm.
Serapias parviflora subsp. hellenica (Renz) Soó
Serapias sennii Renz
→
→
Ranunculus testiculatus Crantz
Ranunculus tuberosus Lapeyr.
Sempervivum marmoreum Griseb. subsp. marmoreum
Jacobaea erratica (Bertol.) Fourr.
Senecio doronicum (L.) L.
Senecio doronicum (L.) L.
Serapias parviflora Parl.
Serapias lingua L. subsp. lingua
Serapias orientalis (Greuter) H. Baumann & Künkele subsp. orientalis
Serapias orientalis (Greuter) H. Baumann & Künkele subsp. orientalis
Serapias neglecta subsp. ionica (H. Baumann & Künkele) H. Baumann & R. Lorenz
Serapias lingua L. subsp. lingua
Serapias vomeracea (Burm. f.) Briq.
Serapias neglecta subsp. ionica (H. Baumann & Künkele) H. Baumann & R. Lorenz
Serapias neglecta subsp. ionica (H. Baumann & Künkele) H. Baumann & R. Lorenz
Serapias orientalis (Greuter) H. Baumann & Künkele subsp. orientalis
Serapias bergonii E.G. Camus
Serapias bergonii E.G. Camus
→ Serapias bergonii E.G. Camus
→ Serapias orientalis (Greuter) H. Baumann & Künkele subsp. orientalis
Serapias vomeracea subsp. columnae auct. fl. graec., non (Aurnier) H. → Serapias bergonii E.G. Camus
Sund.
Serapias wettsteinii H. Fleischm.
→ Serapias bergonii E.G. Camus
Seseli gummiferum subsp. aegaeum P.H. Davis
→ Seseli crithmifolium DC.
Sesleria argentea auct. fl. graec., non (Savi) Savi
→ Sesleria anatolica Deyl
Silaus meoides Griseb.
→ Carum meoides (Griseb.) Halácsy 1894, non Halácsy 1901 [see Appendix III]
Silene graefferi Guss.
→ Silene ciliata subsp. graefferi (Guss.) Nyman
Silene italica subsp. nemoralis (Waldst. & Kit.) Nyman
→ Silene nemoralis Waldst. & Kit. [see Appendix I]
Silene roeseri Boiss. & Heldr.
→ Silene ciliata subsp. graefferi (Guss.) Nyman
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334
Silene uniflora auct. fl. graec., non Roth
Silene uniflora subsp. prostrata (Gaudin) Chater & Walters
Silene vulgaris subsp. antelopum (Vest) Hayek
Sison acaule Steud.
Sison alpinum Schult.
Sison sieberianum DC.
Sium nodiflorum L.
Spergula bocconei (Scheele) Pedersen
Spergula diandra (Guss.) Murb.
Spergula media (L.) Bartl.
Spergula rubra (L.) D. Dietr.
Spergula salina (J. Presl & C. Presl) D. Dietr.
Spergularia atheniensis Asch.
Spergularia campestris (Kindb.) Willk.
Spergularia rubra subsp. atheniensis (Asch.) Rouy & Foucaud
Spergularia salina J. Presl & C. Presl
Stachys acutifolia auct. fl. graec., non Bory & Chaub. nec Link
Stachys orientalis L.
Stachys orientalis (Mill.) Vahl, non L.
Sternbergia fischeriana (Herb.) Roem.
Sternbergia minoica Ravenna
Dimopoulos & al.: Vascular plants of Greece: Supplement
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Silene vulgaris (Moench) Garcke
Silene vulgaris subsp. prostrata (Gaudin) Schinz & Thell.
Silene vulgaris subsp. bosniaca (Beck) Greuter & al.
Ormosolenia alpina (Schult.) Pimenov
Ormosolenia alpina (Schult.) Pimenov
Ormosolenia alpina (Schult.) Pimenov
Helosciadium nodiflorum (L.) W.D.J. Koch
Spergularia bocconei (Scheele) Graebn.
Spergularia diandra (Guss.) Heldr.
Spergularia media (L.) C. Presl
Spergularia rubra (L.) J. Presl & C. Presl
Spergularia marina (L.) Besser
Spergularia bocconei (Scheele) Graebn.
Spergularia bocconei (Scheele) Graebn.
Spergularia bocconei (Scheele) Graebn.
Spergularia marina (L.) Besser
Stachys graeca Boiss. & Heldr.
Stachys alpina L. subsp. alpina
Stachys spinulosa Sm.
Sternbergia vernalis (Mill.) Gorer & J.H. Harvey [see Appendix I]
Sternbergia lutea subsp. greuteriana (Kamari & R. Artelari) Strid
Stipella capensis (Thunb.) Röser & Hamasha
→ Stipa capensis Thunb.
Stipella parviflora (Desf.) Röser & Hamasha
→ Stipa parviflora Desf.
Stipellula capensis (Thunb.) Röser & Hamasha
→ Stipa capensis Thunb.
Stipellula parviflora (Desf.) Röser & Hamasha
→ Stipa parviflora Desf.
Thinopyrum bessarabicum (Săvul. & Rayss) Á. Löve
→ Elytrigia bessarabica (Săvul. & Rayss) Prokudin
Thinopyrum elongatum (Host) D.R. Dewey
→ Elytrigia elongata (Host) Nevski
Thinopyrum intermedium (Host) Barkworth & D.R. Dewey
→ Elytrigia intermedia (Host) Nevski
Thinopyrum intermedium subsp. barbulatum (Schur) Barkworth & → Elytrigia intermedia subsp. trichophora (Link) Á. Löve & D. Löve
D.R.Dewey
Thinopyrum junceum (L.) Á. Löve
→ Elytrigia juncea (L.) Nevski
Thinopyrum ponticum (Podp.) Barkworth & D.R. Dewey
→ Elytrigia obtusiflora (DC.) Tzvelev subsp. obtusiflora
Thinopyrum pycnanthum (Godr.) Barkworth
→ Elytrigia atherica (Link) Kerguélen
Thinopyrum sartorii (Boiss. & Heldr.) Á. Löve
→ Elytrigia sartorii (Boiss. & Heldr.) Holub
Thinopyrum scirpeum (C. Presl) D.R. Dewey
→ Elytrigia scirpea (C. Presl) Holub
Thlaspi goesingense auct. fl. graec., non Halácsy
→ Noccaea tymphaea (Hausskn.) F.K. Mey.
Thlaspi ochroleucum Boiss. & Heldr.
→ Noccaea ochroleuca (Boiss. & Heldr.) F.K. Mey. [see Appendix I in
Dimopoulos & al. 2013: 151]
Thlaspi zaffranii (F.K. Mey.) Greuter & Burdet
→ Noccaea zaffranii F.K. Mey.
Thrincia hirta Roth
→ Leontodon saxatilis Lam. subsp. saxatilis
Thrincia oliverii (DC.) Hausskn.
→ Leontodon tuberosus L.
Thymus kosteleckyanus auct. fl. graec., non Opiz
→ Thymus sibthorpii Benth.
Trachomitum venetum subsp. sarmatiense (Woodson) Avet.
→ Trachomitum sarmatiense Woodson
Tragium depressum (Spreng.) DC.
→ Pimpinella tragium subsp. depressa (Spreng.) Tutin
Tragopogon longirostris Sch. Bip.
→ Tragopogon coelesyriacus Boiss.
Tragopogon porrifolius subsp. longirostris (Sch. Bip.) Greuter
→ Tragopogon coelesyriacus Boiss.
Trifolium brachycalycinum (Katzn. & F.H.W. Morley) F.H.W. Morley → Trifolium subterraneum L.
Trifolium clusii auct. fl. graec., non Godr. & Gren.
→ Trifolium tomentosum L.
Trifolium clusii Godr. & Gren.
→ Trifolium resupinatum L. subsp. resupinatum
Trifolium congestum Link, non Guss.
→ Trifolium fragiferum L.
Trifolium creticum L.
→ Melilotus creticus (L.) Desr.
Trifolium glaucescens Hausskn.
→ Trifolium campestre Schreb.
Trifolium intermedium Guss., non Lapeyr.
→ Trifolium infamia-ponertii Greuter
Trifolium meneghinianum auct. fl. graec., non Clementi
→ Trifolium michelianum Savi
Trifolium nervulosum Boiss. & Heldr.
→ Trifolium glanduliferum Boiss.
Trifolium niethammeri Rothm.
→ Trifolium pignantii Fauché & Chaub.
Trifolium nigrotinctum Boiss. & Orph.
→ Trifolium squamosum L.
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Trifolium preslianum Boiss.
Trifolium speciosum Boiss., non Willd.
Trifolium tenoreanum Boiss. & Spruner
Trifolium xerocephalum Fenzl
Triticum ventricosum (Tausch) Ces.
Valantia aristata Boiss. & Heldr.
Valantia humifusa Sieber
Valeriana phitosiana Quézel & Contandr.
Valerianella thelocarpa P. Candargy
Verbascum blattariforme Griseb.
Verbascum glandulosum Delile, non Roem. & Schult.
Verbascum gloeotrichum subsp. doiranense Bornm.
Verbascum meteoricum Hausskn.
Verbascum perviscosum Borbás
Veronica austriaca subsp. teucrium (L.) D.A. Webb
Viola alba subsp. thessala (Boiss. & Spruner) Hayek
Viola pindicola Formánek
Viola tricolor auct. fl. graec., non L.
Viola tricolor subsp. macedonica (Boiss. & Heldr.) A. Schmidt
Viscum laxum subsp. abietis (Wiesb.) O. Schwarz
Visnaga daucoides Gaertn.
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
→
Xanthium orientale subsp. saccharatum (Wallr.) B. Bock
→ Xanthium orientale subsp. italicum (Moretti) Greuter
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Trifolium affine C. Presl
Trifolium boissieri Guss.
Trifolium tenuifolium Ten.
Trifolium argutum Banks & Sol.
Aegilops ventricosa Tausch
Valantia aprica (Sm.) Tausch
Valantia aprica (Sm.) Tausch
Valeriana crinii subsp. epirotica (Phitos) Franzén
Valerianella orientalis (Schlecht.) Boiss. & Balansa
Verbascum blattaria L.
Verbascum gloeotrichum Hausskn. & Heldr.
Verbascum gloeotrichum Hausskn. & Heldr.
Verbascum gloeotrichum Hausskn. & Heldr.
Verbascum gloeotrichum Hausskn. & Heldr.
Veronica teucrium L. [see Appendix I in Dimopoulos & al. 2013: 158]
Viola alba Besser subsp. alba
Viola macedonica Boiss. & Heldr.
Viola macedonica Boiss. & Heldr.
Viola macedonica Boiss. & Heldr.
Viscum album subsp. abietis (Wiesb.) K. Malý
Ammi visnaga (L.) Lam.
336
Appendix III: Comments. Supplement
For authors of plant names, see entries arrowed with
“►” in the Floristic catalogue or Appendices I and II
(above).
► Acer ×bornmuelleri
Represents the hybrid A. campestre × A. monspessulanum, disregarded.
► Achillea ×tymphaea
Represents the hybrid A. coarctata × A. nobilis subsp. neilreichii, disregarded.
► Aira elegans
According to W. Greuter (pers. comm.), A. elegans Gaudin
(Agrost. Helv. 1: 130, 355. 1811) was published as a synonym
instead of an accepted name and is therefore not validly published. Aira elegans Willd. ex Roem. & Schult. (Syst. Veg., ed.
15bis, 2: 682. 1817) is the correct name of this taxon at specific
rank (Wipff 2007: 616), antedating A. elegantissima Schur (in
Verh. Mitth. Siebenbürg. Vereins Naturwiss. Hermannstadt 4:
85. 1853).
► Ajuga orientalis
Infraspecific variation deserves at most varietal rank (Euro+Med
2006+).
► Allium ritsi
Correct orthography of the epithet of this species named after Yannis Ritsos (Euro+Med 2006+; IPNI 2012+), contrary to
what is given in Tan & Iatroú (2001: 428).
► Allium sativum
Cultivated as vegetable crop, not naturalized (Euro+Med
2006+).
► Alyssum stribrnyi
Not known to occur in Greece (Greuter & al. 1986: 49; Tutin
& al. 1993: 366; Strid & Tan 2003). Erroneously reported in
Euro+Med (2006+), based on a mapping mistake by Jalas & al.
(1996: 46), who misplaced a single dot in NW Greece, which
refers to populations in former Jugoslavia just north of the border from Greece (see Vandas 1909: 36, under A. montanum var.
galicicae).
► Anacamptis ×eccarii
Represents the hybrid A. boryi × A. coriophora subsp. fragrans,
disregarded.
► Anacamptis ×gennarii
Represents the hybrid A. morio × A. papilionacea, disregarded.
► Anacamptis ×lesbiensis
Represents the hybrid A. pyramidalis × A. sancta, disregarded.
► Anacamptis palustris subsp. robusta
Absent but reported in error, confined to the W Mediterranean
area (Euro+Med 2006+), records from KK refer to A. palustris
subsp. elegans (Kretzschmar & al. 2007: 104, fig. 104/1).
► Anacamptis ×parvifolia nothosubsp. bicknellii
Represents the hybrid A. coriophora subsp. fragrans × A. laxiflora, disregarded.
► Anacamptis ×sciathia
Represents the hybrid A. boryi × A. morio, disregarded.
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► Anacamptis ×simorrensis
Represents the hybrid A. coriophora × A. pyramidalis, disregarded.
► Anthemis bornmuelleri
A single literature record from EAe, based on Major 927 from
Samos and accepted by Greuter & Raab-Straube (2008: 23),
refers to A. cotula (rev. Grierson & Yavin in Davis 1975: 209).
► Anthemis macrantha
An unsubstantiated literature record from Thessaly (Hayek
1931: 620, under A. triumfettii var. rigescens) was rejected for
Greece in Tutin & al. (1976: 157) and is likely to refer to A.
triumfettii s.str.
► Anthemis parvifolia
Confined to the Levant. Literature records from EAe (Rodos)
refer to A. pseudocotula (Carlström 1987: 90).
► Anthriscus nitidus
Anthriscus nitidus (Wahlenb.) Hazsl. (Éjsz. Magyarh. Vir.: 152.
1864) antedates A. nitidus (Wahlenb.) Garcke (Fl. N. Mitt.Deutschland, ed. 7: 180. 1865).
► Arenaria biflora
Absent from Greece (see Greuter & al. 1984: 161). Greek
records refer to A. rotundifolia (Strid & Tan 1997: 159 – 160).
► Asparagus acutifolius
Absent from the Cretan area (Fielding & Turland 2005: 471),
previous literature records may be referable to misidentified
shade forms of A. aphyllus subsp. orientalis (R. Jahn, pers.
comm.).
► Asparagus aphyllus subsp. aphyllus
Assumed occurrence of this W & C Mediterranean subspecies
in continental Greece (Euro+Med 2006+; derived from Davis
1984: 77) not substantiated by herbarium material, hence disregarded.
► Asperula tenella
All records need confirmation and might refer to forms of A.
aristata (A. Strid).
► Asplenium adulterinum subsp. adulterinum
A single unsubstantiated record from NPi is queried by Strid
(1986: 18) and is likely to refer to misidentified material of A.
trichomanes L. subsp. trichomanes.
► Astragalus lanatus
Confined to Lebanon and SW Syria (Podlech & Zarre 2013:
1166). A record from EAe (Rodos) cited by Rechinger (1944:
318) has not been confirmed and is probably incorrect.
► Astragalus thracicus
Literature records from EAe refer to A. lesbiacus (Chios, Lesvos) and A. condensatus Ledeb. (Samos), respectively (A. Strid,
based on Podlech & Zarre 2013).
► Asyneuma canescens subsp. cordifolium
Erroneously given for Greece in Euro+Med (2006+), the record
incorrectly inferred from Tutin & al. (1972: 277). Endemic to
former Jugoslavia (Greuter & al. 1984: 121).
► Atriplex oblongifolia
Reported in error by Jalas & Suominen (1980: map 518), according to Strid & Tan (1997: 126). However, recently collected
in EAe (Kalimnos, Zervou & al. 2009: 168).
Willdenowia 46 – 2016
► Buglossoides arvensis subsp. sibthorpiana
Subspecific rank inferred from Euro+Med (2006+) only tentatively accepted. Infraspecific variability of B. arvensis seems
to be predominantly ecology-driven and better deserves varietal rank (see, e.g., Halácsy 1902: 349; Hayek 1931: 82; Davis
1978: 316).
► Bupleurum falcatum subsp. falcatum
Erroneously given for Greece in Euro+Med (2006+) inferred
from Tutin & al. (1968: 349), but not substantiated by previous literature sources (see Hayek 1931: 971 – 972) or herbarium
material seen. A single literature report of this subspecies (Parent 2005: 211) from Mt Trapezitsa needs confirmation. Several
other collections from this area (NPi) have been identified as
subsp. cernuum. It seems probable that only subsp. cernuum
occurs in Greece.
► Bupleurum lancifolium
In Greece confined to Pe, Kik and KK (see map in Snogerup
& Snogerup 2001: 220, fig. 5). Records from IoI, NPi, StE,
EC, NE, NAe and EAe refer to B. subovatum, which has been
widely but taxonomically inadequately lumped with B. lancifolium based on Tutin & al. (1968: 427; see also Dimopoulos &
al. 2013: 174).
► Calluna vulgaris
A single unsubstantiated literature record from NE (Zaganiaris
1940: 85), never confirmed later, disregarded and likely to be
based on misidentified material of Bruckenthalia spiculifolia.
► Capsella rubella
Excluded from Greece by Akeroyd (in Greuter & Raus 1986:
417). Plants resembling C. rubella have been recorded from IoI,
StE, KK and EAe, but are considered to belong to one of the biotypes of the C. bursa-pastoris complex (Strid & Tan 2003: 250).
► Carduus ×intercedens
Represents the hybrid C. hamulosus subsp. hamulosus × C. nutans subsp. leiophyllus, disregarded.
► Carex atrata subsp. aterrima
Assumed occurrence of this W & C Mediterranean subspecies (Euro+Med 2006+; derived from Tutin & al. 1980: 317) in
Greece not substantiated by herbarium material so far (Strid &
Tan 1991: 862).
► Carex pairae
Centred in W & C Europe and absent from most of the Balkan
Peninsula (Euro+Med 2006+), previous records from NE, WAe
and Kik refer to C. muricata subsp. muricata (see Strid & Tan
1991: 845).
► Carex sempervirens
Confined to mountains of W & C Europe (Castroviejo &
al. 2007b: 227), records from Greece refer to C. bulgarica
(Euro+Med 2006+).
► Carum meoides
Correct nomenclature emended by Wolff (1927: 154, 156), the
name often misapplied to the similar C. graecum in previous
floristic literature (see Appendix II). Erroneously given for
IoI (Mt Enos, Strid 1986: 702, under C. rupestre), based on
misidentified material of Trinia glauca subsp. pindica (rev. P.
Hartvig).
► Cedrus atlantica, C. deodara
Two species occasionally planted for timber (Tutin & al. 1993:
40), not naturalized.
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337
► Centaurea phrygia subsp. indurata
Literature records from NE (Strid & Papanicolaou 1981) are
incorrect and fall within the variability of C. phrygia subsp. stenolepis (rev. A. Strid).
► Centaurea phrygia subsp. razgradensis
A single unsubstantiated literature record from NE (W Rodopi)
accepted by Eleftheriadou & Raus (1996: 467) is likely to belong to C. phrygia subsp. stenolepis, which is well documented
by herbarium collections from the area (A. Strid).
► Centaurea stereophylla
A Pontic steppe element occurring as far south as Serbia and
Bulgaria (Assyov & Petrova 2006), erroneously given for
Greece in Tutin & al. (1976: 268) by misinterpretation of the
imprecise “Ma” (for Macedonia) and “Thra” (for Thrace) in
Hayek (1931: 744).
► Centranthus calcitrapae
Subspecies in C. calcitrapae (Tutin & al. 1976: 56; Kerguélen
1998 – 2002) considered as taxonomically overrated (Castroviejo & al. 2007a: 231).
► Cephalanthera ×majeri
Represents the hybrid C. damasonium × C. rubra, disregarded.
► Cerastium arvense
Absent from Greece, reported in error based on misidentified
material of C. decalvans (Strid 1986: 114; Jalas & Suominen
1983: 93).
► Cerastium brachypetalum subsp. brachypetalum
Absent from Greece (see Euro+Med 2006+). A single literature record from WAe (Evvia, erroneously accepted in Jalas &
Suominen 1983: 7) refers to C. brachypetalum subsp. roeseri
(Strid & Tan 1997: 210).
► Cerastium diffusum
Confined to W Europe, records from Greece refer to C. glutinosum (Strid & Tan 1997: 213).
► Cerastium gracile
Endemic to Spain and NW Africa, records from Greece refer to
C. ramosissimum (Strid & Tan 1997: 213).
► Cerastium holosteoides
Often considered a subspecies of C. fontanum (see Appendix
II), but species concept in an informal C. fontanum aggr., in
which C. holosteoides Fr. 1817 antedates C. vulgare Hartm.
1820, proves appropriate, advocated by most recent C European floras (Fischer & al. 2008; Jäger 2011).
► Ceratocephala orthoceras
Erroneously given for Greece in Greuter & al. (1989: 441, under Ranunculus testiculatus) and in Euro+Med (2006+) by misinterpretation of the imprecise “Thra” (for Thrace) in Hayek
(1924: 345).
► Cichorium endivia
Widely cultivated as a vegetable crop (Rechinger 1944: 673;
Tutin & al. 1976: 305), believed to have originated in China
(Davis 1975: 628); not naturalized, hence disregarded.
► Cirsium serrulatum
A Pontic steppe element extending to E Romania (Tutin &
al. 1976: 237), erroneously given for Greece in Euro+Med
(2006+).
338
► Cladanthus mixtus
Presence in KK questionable, based on 19th century records
from Kriti (Raulin 1869: 781), not reconfirmed later and probably referring to a lost casual introduction. Not mentioned for
Kriti by Fielding & Turland (2005).
Dimopoulos & al.: Vascular plants of Greece: Supplement
► Edraianthus tenuifolius
Excluded from Greece in Strid & Tan (1991: 396), relevant
material seen from NPi belongs to E. graminifolius (rev. P.
Hartvig).
► Elytrigia sartorii
Elytrigia sartorii (Boiss. & Heldr.) Holub (in Folia Geobot.
Phytotax. 23: 413. 1988) antedates E. sartorii (Boiss. & Heldr.)
H. Scholz (in Ber. Inst. Landschafts Pflanzenökol. Univ. Hohenheim Beih. 16: 46. 2003).
► Corylus maxima
Probably of hybrid origin, derived from C. avellana under cultivation (see Browicz & Zieliński 1982: 29). In Greece only
cultivated and not naturalized (see also Tutin & al. 1993: 71),
hence disregarded in accordance with Boratyński & al. (1992).
Actually given as established only in the Adriatic coastal NW
Balkans by Jalas & Suominen (1976: 65).
► Epilobium ×persicinum
Represents the hybrid E. parviflorum × E. roseum, disregarded.
► Cynoglossum officinale
Erroneously recorded from IoI and SPi, based on misidentified
material of C. columnae (rev. F. Selvi). Questionable records
from NPi, Pe and StE still to be revised accordingly (A. Strid).
► Epipactis leptochila subsp. aspromontana
Endemic to S Italy (Apulia, Calabria), see Euro+Med (2006+).
Records from Greece (Baumann & al. 2006: 81) refer to E. olympica.
► Dianthus giganteus subsp. croaticus
Erroneously given for Greece by Greuter & al. (1984: 194), but
absent (see Jalas & Suominen 1986: 206, map 1481).
► Epipactis leptochila subsp. neglecta
Confined to France, Germany and Italy (Euro+Med 2006+). An
alleged occurrence in Greece (Baumann & al. 2006: 86) refers
to Epipactis leptochila subsp. naousaensis.
► Dianthus leptopetalus
No data available from Greece (Jalas & Suominen 1986: 188 &
map 1436). A single unsubstantiated record for Greece (Greuter & al. 1984: 198) probably based on misinterpretation of the
imprecise “Ma” (for Macedonia) and “Thra” (for Thrace) in
Hayek (1924: 250) ), hence disregarded.
► Eragrostis minor
A single record from Crete is queried by Böhling & Scholz
(2003) because they did not see any corroborating herbarium
specimen. Until further evidence is available, its presence in
KK should be regarded as doubtful (R. Jahn).
► Dianthus microlepis
A single unsubstantiated literature record from NE (Mt Belles)
not confirmed (Strid & Tan 1997: 355), hence disregarded.
► Dianthus pallidiflorus
No material from Greece has been seen, hence disregarded
(not mapped for Greece in Jalas & Suominen 1986: 186).
Questionable records for Greece (Greuter & al. 1984: 198;
Tutin & al. 1993: 244), misleadingly accepted in Euro+Med
(2006+), are based on the imprecise “Ma” (for Macedonia)
and “Thra” (for Thrace) in Hayek (1924: 242, under D. aridus
Janka).
► Dianthus pancicii
No material from Greece has been seen, hence disregarded.
Collections from NC, initially reported by Strid (1986: 199)
as D. tristis, belong to D. cruentus (rev. A. Strid). Unsubstantiated records for Greece (Greuter & al. 1984: 195; Tutin & al.
1993: 236, under D. tristis), accepted in Euro+Med (2006+),
are based on the imprecise “Ma” (for Macedonia) in Hayek
(1924: 234, 236). Dianthus pancicii Velen. 1886 antedates the
conspecific D. tristis Velen. 1890. An alleged earlier homonym, “D. pancicii F. N. Williams” (Tutin & al. 1993: 236),
does not in fact exist, but was erroneously inferred from D.
capitatus var. pancicianus F. N. Williams (in J. Bot. 23: 342.
1885), a synonym of D. capitatus subsp. andrzejowskianus
(see Appendix I).
► Eryngium palmatum
In Greece confined to the Prespa area (Nomos Florina), the S
limit of the total range of this Balkan endemic. Previous records
from elsewhere in Greece (Nomoi Pella, Pieria, Kozani) are
considered incorrect, referring to E. wiegandii (A. Strid).
► Euphorbia anacampseros
Endemic to Turkey (Davis 1982: 612; Greuter & al. 1986: 216).
Given for Greece in Euro+Med (2006+), but probably in error,
the record likely referring to E. myrsinites.
► Euphorbia hierosolymitana
Absent from Greece, confined to SW Anatolia, Cyprus and the
Levant. Erroneously given for Greece in Euro+Med (2006+),
wrongly inferred from Davis (1982: 583). The record essentially refers to E. acanthothamnos (see Rechinger 1944: 113).
► Euphorbia hirta
A tropical annual weed of irrigated fields, reported as introduced in EAe (Euro+Med 2006+), but not established, hence
disregarded.
► Euphorbia hypericifolia
An annual of Central and South America, cultivated for ornament in flowerbeds, reported as introduced in EAe (Euro+Med
2006+), but not established, hence disregarded.
► Dianthus petraeus subsp. petraeus
Not in Greece according to Jalas & Suominen (1986: 172).
Greek records (e.g. Greuter & al. 1984: 201) are likely to refer
to D. petraeus subsp. orbelicus (see Strid & Tan 1997: 349).
► Euphorbia lucida
No material from Greece has been seen, hence disregarded.
Given for Greece in Euro+Med (2006+), inferred from Tutin
& al. (1968: 225) and obviously based on the imprecise “Thra”
(for Thrace) in Hayek (1924: 131).
► Dianthus roseoluteus
Mentioned for Greece by Jalas & Suominen (1986: 189) based
on a single literature record for NE with no exact data available. No material from Greece has been seen, hence disregarded
(Strid & Tan 1997: 372).
► Fibigia clypeata subsp. clypeata
This is represented by var. clypeata (straight stiff hairs in the
central part of the silicula) and var. eriocarpa (DC.) Thiéb.
(long whitish hairs concealing the silicula and giving it a
hirsute-villous appearance). The two varieties are completely
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mixed throughout their total range and cannot be regarded as
geographical races (subspecies). Morphologically and chorologically more distant Anatolian populations, however, call for
recognition at subspecies level (F. clypeata subsp. anatolica A.
Duran & Tuştaş).
► Ficaria verna subsp. ficariiformis
Ficaria verna subsp. ficariiformis (Rouy & Foucaud) Maire
(in Bull. Soc. Hist. Nat. Afrique N. 21: 59. 1930) antedates F.
verna subsp. ficariiformis ([F.W. Schultz ex] Rouy & Foucaud)
B. Walln. (in Ann. Naturhist. Mus. Wien, B, 109: 277. 2008).
► Fumana laevis
Given occurrence in Greece particularly based on herbarium
material revised by J. Guemes. Considered conspecific with F.
thymifolia, or reduced to varietal rank under the latter, in previous floristic literature (Boissier 1867: 449; Hayek 1925: 498;
Rechinger 1944: 252; Tutin & al. 1968: 433).
► Fumaria officinalis subsp. wirtgenii
Occurrence in Greece of this W & C European taxon queried by
M. Lidén (Strid & Tan 2003: 112).
► Fumaria schleicheri
Not in Greece (Strid & Tan 2003: 112), previous records refer
to F. vaillantii.
► Galium caminianum
Galium caminianum Schult. & Schult. f. (Mantissa 3: 186.
1827) antedates Galium recurvum Req. ex DC. (Prodr. 4: 609.
1830).
► Galium nigricans
A record from NAe (Thasos) of this chiefly SW Asian species
needs confirmation. Collections from Samos (EAe), identified
as G. nigricans by Ehrendorfer in 1988, turned out to represent
G. floribundum (rev. A. Strid).
► Geranium pratense
Reported once from NE (Mt Falakro; Goulimis 1956), probably
in error for G. sylvaticum (Strid 1986: 543).
► Gnaphalium hoppeanum subsp. hoppeanum
Not in Greece, as misleadingly suggested in Euro+Med (2006+,
as G. hoppeanum s.str.), replaced there by G. hoppeanum subsp.
magellense (see Greuter & Raab-Straube 2008: 230, under G.
diminutum).
► Goniolimon dalmaticum
Endemic to Croatia, where it grows on saline ground near the
sea, whereas G. tataricum is of Pontic-Mediterranean range
and grows in xerophilous pastures in hilly regions. Both species
have been reported from N Greece, but it seems probable that
all Greek records of the former are incorrect and refer to the
latter (A. Strid).
► Helictochloa pratensis
Old unsubstantiated literature records under the synonyms of
Avena pratensis or Avenula pratensis from Greece (Diapoulis
1939: 196; Kavvadas 1956–1964: 6) are erroneous. The species is absent from the Balkan Peninsula south of Serbia (see
Euro+Med 2006+).
► Helictochloa versicolor
Old unsubstantiated literature records under the synonyms of
Avena versicolor or Avenula versicolor from Greece (Diapoulis
1939: 196; Kavvadas 1956 – 1964: 5 – 6) are erroneous. The species is absent from Greece (see Euro+Med 2006+).
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► Helosciadium repens
Occurrence in Greece not substantiated so far. Given for the
Cretan area in Tutin & al. (1968: 351), although there are no
records as a basis for this (Greuter 1974: 139; Turland & al.
1993: 148). A specimen from NC (Haristos 1161, ATH), preliminarily referred to H. repens, has stems apparently suberect
and its identity needs to be verified (A. Strid).
► Heracleum sphondylium subsp. orsinii
Absent from Greece. The record in Euro+Med (2006+) is merely
calculated based on Tutin & al. (1968: 366) and is not substantiated by herbarium specimens or documented literature sources.
► Hieracium bifidum subsp. basicuneatum
A single unsubstantiated literature record for Greece without
locality (Zahn 1921: 424, followed by Ascherson & Graebner
1935: 645; Hayek 1931: 911; Greuter & Raab-Straube 2008:
260) is not placeable to a relevant Greek floristic region, hence
disregarded unless corroborated by confirmed re-collections
(G. Gottschlich).
► Hieracium krischtimanum
Originally described as H. kritschimanum (Zahn 1928: 384;
see Hayek 1931: 990; Greuter & Raab-Straube 2008: 347).
Gottschlich’s demand (in Greuter & Raus 2011: 314), based on
ICN Art. 60.1 (McNeill & al. 2012), for an orthographical correction of the epithet to krischtimanum, because the species was
described from the Krischtima valley (Bulgarian C Rhodope
Mts), was accepted and implemented in Euro+Med (2006+).
► Hieracium lazistanum subsp. leithneri
Endemic to continental Greece and Bulgaria, possibly extending to Albania (Strid & Tan 1991: 635; Assyov & al. 2006:
206), but, contrary to what is given in Greuter & Raab-Straube
(2008: 365), absent from Anatolia, where it is replaced by subsp. lazistanum (Tutin & al. 1976: 390; Davis 1975: 729). 19th
century records from Crete (see Rechinger 1944: 703, under
H. leithneri) are considered erroneous and likely to refer to H.
schmidtii (Turland & al. 1993: 67).
► Hieracium pannosum subsp. friwaldii
The identity of a single 19th century collection from Crete (see
Rechinger 1944: 704) is uncertain (Buttler in Strid & Tan 1991:
621). Since there are no confirming later collections, Turland &
al. (1993: 67) regarded H. pannosum as absent from KK.
► Hieracium parnassi
Infraspecific taxa of this Balkan endemic are obsolete (see discussion in Strid & Tan 1991: 633). A 19th century record from KK
(Rechinger 1944: 704, as H. parnassi subsp. versutum, followed
by Greuter & Raab-Straube 2008: 407) is considered erroneous
and likely to refer to H. schmidtii (Turland & al. 1993: 67).
► Hieracium sermenikense
Named after its locus classicus Sermeniko (Σερμένικο, the
former Turkish Sirminik, since 1928 called Filakti/Φυλακτή),
a village near Karditsa (SPi). The original spelling “sermonikense” of the epithet in the protologue of 1897 (documented in
IPNI 2012+) is considered an orthographical error according to
ICN Art. 60.1 (McNeill & al. 2012) and has to be corrected to
sermenikense, as already implemented in basic floristic literature five years afterwards (Halácsy 1902) and ever since (Tutin
& al. 1976: 408; Greuter & Raab-Straube 2008: 453).
► Hierochloe australis, H. odorata
Old unsubstantiated literature records of these taxa from Greece
(Diapoulis 1939: 212) are erroneous. Both species are absent
from Greece (see Euro+Med 2006+).
340
► Holosteum umbellatum subsp. glutinosum
Erroneously given for Greece in Euro+Med (2006+) inferred
from Tutin & al. (1993: 164), but absent (see Greuter & al.
1984: 214).
► Hordeum bulbosum
This species comprises diploids in the W & C Mediterranean
area to W Greece and tetraploids from C Greece east to Afghanistan, the border following the Pindos mountains. However, the
two cytotypes cannot be distinguished morphologically and a
taxonomic subdivision of the species is not recommended (Jørgensen 1982).
► Hyoseris radiata
All Aegean records of H. radiata belong to H. lucida (syn. H.
radiata subsp. graeca).
► Hypericum hyssopifolium
Not in Greece, confined to the C & W Mediterranean area (Robson 2010). Greek records are based on misidentified material of
H. tymphresteum.
► Hypericum maculatum subsp. maculatum
Absent from Greece (see Greuter & al. 1986: 269), replaced by
H. maculatum subsp. immaculatum (Strid 1986: 608).
► Hypericum richeri
Not in Greece (Strid 1986: 608; Greuter & al. 1986: 272). Material of H. barbatum from NE (Mt Vrondous, Rechinger 10853,
type of H. aucheri var. punctatofimbriatum Rech. f.) was incorrectly identified by Robson in 1967 as H. richeri subsp. grisebachii (Boiss.) Nyman and is the source for the record of this
subspecies from Greece in Tutin & al. (1968: 267), see Robson
(2012: 77).
► Isoetes gymnocarpa
Only recently reported from Pe (Greuter 2012: 24, as I. sicula)
and known from Kik and EAe as I. histrix var. subinermis
(Milos, Siros, Tinos, Ikaria: all LD online; Rodos, Carlström
1987: 44). Isoetes gymnocarpa (Gennari) A. Braun (in Monatsber. Koenigl. Preuss. Akad. Wiss. Berlin 1863: 555. 1864)
antedates I. sicula Tod. (in Giorn. Sci. Nat. Econ. Palermo 1:
251. 1866).
► Isoetes setacea
Absent but reported in error (Hayek 1924: 12), confined to the
W Mediterranean area. Greek records refer to I. heldreichii.
► Jacobaea aquatica
Absent from Greece. The record in Tutin & al. (1976: 202,
as Senecio aquaticus), accepted by Greuter & Raab-Straube
(2008: 498), is not substantiated by herbarium specimens or
previous literature except for the misapplication of the name
S. aquaticus to material of J. erratica (see Boissier 1875: 392,
under S. erraticus).
► Jacobaea vulgaris
Distribution of different subspecies in Greece unsettled so far.
Plants from NE with basal leaves subentire or showing a large
terminal lobe, instead of deeply pinnatifid as in J. vulgaris
subsp. vulgaris, were identified as J. vulgaris subsp. gotlandica
(Neuman) B. Nord., a steppic element with a Pontic-Pannonian
range extending to the islands of Öland and Gotland in E Sweden. It possibly deserves specific rank due to genetic discontinuities with J. vulgaris subsp. vulgaris (Wysk & al. 2009).
Similar collections have been seen from SPi and NPi (B!, Th.
Raus).
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Dimopoulos & al.: Vascular plants of Greece: Supplement
► Juniperus oxycedrus subsp. oxycedrus
A W Mediterranean taxon absent from Greece, replaced there
by J. oxycedrus subsp. deltoides (Adams 2004, 2011; Bernardo
& al. 2009). Records of the latter from IoI and Kik are erroneous, referring to misidentified material of J. macrocarpa (see
Strid & Tan 1997: 13).
► Juniperus phoenicea, J. turbinata
Previous E. Mediterranean and Greek records of J. phoenicea belong to J. turbinata (Macaronesia to SW Asia), while J.
phoenicea s.str. is confined to Mediterranean France and Spain
(Adams & al. 2013: 203).
► Jurinea kilaea
Euxine element confined to the Black Sea coast of Bulgaria
and Turkey (Davis 1975: 445), erroneously given for Greece
in Euro+Med (2006+) by misinterpretation of the imprecise
“Thra” (for Thrace) in Hayek (1931: 700).
► Jurinea polycephala
Absent from Greece, from where it was erroneously reported
in Greuter & Raab-Straube (2008: 505) by misinterpretation of
the imprecise “Ma” (for Macedonia) in Hayek (1931: 700–701,
under J. arachnoidea).
► Kickxia spuria subsp. spuria
Erroneously given for Greece in Euro+Med (2006+), incorrectly inferred from Tutin & al. (1972: 239). All Greek material
seen belongs to subsp. integrifolia (see also Davis 1978: 677).
► Larix decidua
Occasionally planted for timber in NE (Strid & Tan 1997: 4),
not naturalized.
► Lathyrus vernus
No Greek material of this Euro-Siberian taxon has been seen.
Unsubstantiated literature records (Tutin & al. 1968: 138; Greuter & al. 1989: 125; Euro+Med 2006+) are likely to refer to L.
venetus.
► Leontodon asperrimus
Reported in error from Greece (Euro+Med 2006+). Greek
records refer to L. biscutellifolius (see Strid & Tan 1991: 530,
under L. crispus subsp. asper).
► Leontodon crispus
As advocated in Euro+Med (2006+), specific rank alongside L.
graecus in an informal L. crispus aggr. is appropriate for the two
subspecies of L. crispus traditionally distinguished in Greece
(viz. subsp. asper, subsp. crispus). The two taxa, accordingly
referred to as L. biscutellifolius DC. 1838 (= L. asper (Waldst.
& Kit.) Poir. 1814, non Forssk. 1775) and L. crispus s.str. (L.
crispus subsp. “eu-crispus” sensu Hayek 1931: 813), occupy
extensively vicariant total ranges of different chorotype (Mediterranean-Atlantic vs. subcontinental) but are largely sympatric
in SE Europe. Whether they prefer different ecological niches
where they chorologically co-occur in Greece is unexplored
so far (see, e.g., Karagiannakidou & Raus 1996: 508). Previous literature records of L. crispus from Greece not determined
to subspecies thus regard L. crispus aggr. and may belong to
either taxon. Unfortunately Halácsy, in his Conspectus florae
graecae, applied the name L. asper to L. crispus (s.l.), referring correctly to collections of L. biscutellifolius from NPi and
SPi (see Halácsy 1902: 187, under L. asper var. haussknechtii
and L. asper var. setulosus) but incorrectly to collections of L.
crispus (s.str.) from IoI, NPi and SPi (see Halácsy 1902: 187,
under L. asper var. typicus and L. asper var. saxatilis). Leontodon crispus subsp. rossianus, erroneously mentioned in Tutin
Willdenowia 46 – 2016
& al. (1976: 314) as a third subspecies of L. crispus to occur in
Greece due to a misinterpretation of Hayek (1931: 813), falls
within the range of variation of L. crispus s.str. (Strid & Tan
1991: 529; see Appendix II).
► Leonurus marrubiastrum
Old records (Halácsy 1902: 534; Hayek 1929: 277), previously
considered erroneous, corroborated by several recent collections.
► Liparis loeselii
Given for N Greece without further details by Baumann & al.
(2006: 122), although there are no records as a basis for this (see
Euro+Med 2006+; Hayek 1933: 416; Tutin & al. 1980: 350).
► Lotus corniculatus
There are no verified records from the Aegean islands. Includes
L. corniculatus var. stenodon Boiss. & Heldr., a montane ecotype described from the Greek mountains, sometimes overrated
at specific rank in previous floristic literature (see, e.g., Strid
1986: 519).
► Lotus pedunculatus
Previously reported from W Kriti and N Evvia, probably in
error, confused with L. preslii or L. tenuis. The only verified
Greek records are from wet habitats in Kerkira and the W and
N mainland.
► Lunaria annua subsp. pachyrhiza
Lunaria annua subsp. pachyrhiza (Borbás) Maire & Petitm.
(in Matér. Étude Fl. Géogr. Bot. Orient 4: 30. 1908) antedates
L. annua subsp. pachyrhiza (Borbás) Hayek (in Repert. Spec.
Nov. Regni Veg. Beih. 30(1) [Prodr. Fl. Penins. Balcan. 1]: 424.
1925).
► Luzula forsteri
The occurrence of Luzula forsteri (Sm.) DC. subsp. forsteri in
Greece, although given in Euro+Med (2006+), is queried by
Kaplan (2001: 60).
341
► Minuartia graminifolia subsp. graminifolia
Absent from Greece, confined to Italy. Greek records refer to
M. graminifolia subsp. brachypetala (Strid & Tan 1997: 190).
► Minuartia hybrida
Varietal rank is adopted as appropriate for glabrous and glandular hairy plants in M. hybrida, which are optionally treated
as subsp. tenuifolia (L.) Kerguélen and subsp. hybrida, respectively (see, e.g., Kerguélen 1998 – 2002; Jäger 2011).
► Minuartia kitanovii
Occurrence in Greece (Jalas & Suominen 1983: 46; Greuter &
al. 1984: 225) accidentally omitted in Flora hellenica (Strid &
Tan 1997).
► Minuartia rumelica
Erroneously given for Greece in Euro+Med (2006+), but absent, confined to Bulgaria (Jalas & Suominen 1986: 46, map
764).
► Minuartia setacea subsp. setacea
A record for Greece of this subspecies (Euro+Med 2006+, inferred from Tutin & al. 1993: 156) is not substantiated by material seen, hence disregarded (Strid & Tan 1997: 178 – 179).
► Muscari cycladicum
The record for the Cretan area of M. cycladicum subsp. cycladicum (Euro+Med 2006+) should be regarded as erroneous since
it is not backed by literature sources or herbarium specimens.
Records of M. cycladicum subsp. subsessile for the Cretan area
(Bentzer 1973) are likely to refer to M. spreitzenhoferi or M.
weissii and must be regarded as dubious (A. Strid).
► Ophrys ×asterusica
Represents the hybrid O. omegaifera subsp. fleischmannii × O.
omegaifera subsp. omegaifera, disregarded.
► Ophrys ×baumanniana nothosubsp. baumanniana
Represents the hybrid O. cretica subsp. cretica × O. sphegodes
subsp. gortynia, disregarded.
► Luzula multiflora
Distribution of the two subspecies in Greece only incompletely
known so far (Kirschner 1993: 161, 165). According to Kirschner (1992: 239), the true hexaploid subsp. multiflora seems to
be very rare in the Balkan Peninsula, particularly in its S half.
► Ophrys ×baumanniana nothosubsp. hierapetrae
Represents the hybrid O. cretica subsp. cretica × O. sphegodes
subsp. cretensis, disregarded.
► Lycopus ×intermedius
Represents the hybrid L. europaeus × L. exaltatus, disregarded.
► Ophrys ×burneriana
Represents the hybrid O. sphegodes subsp. cretensis × O. sphegodes subsp. spruneri, disregarded.
► Marrubium ×paniculatum
Represents the hybrid M. peregrinum × M. vulgare, disregarded.
► Medicago ×blancheana
Represents the hybrid M. bonarotiana × M. rotata, not established, hence disregarded (see Lassen 1999; Small 2011: 222).
► Melica picta
Old unsubstantiated literature records from Greece (Diapoulis
1939: 176; Kavvadas 1956 – 1964: 2552) are erroneous. The
species is absent from Greece (see Euro+Med 2006+).
► Minuartia bosniaca
Occurrence in Greece (Jalas & Suominen 1983: 48; Greuter &
al. 1984: 215; Tutin & al. 1993: 156) accidentally omitted in
Flora hellenica (Strid & Tan 1997).
► Minuartia erythrosepala
Absent from Greece, confined to Turkey. Greek records refer to
M. anatolica (Strid & Tan 1997: 182).
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► Ophrys ×corcyrensis
Represents the hybrid O. ferrum-equinum × O. sphegodes, disregarded.
► Ophrys holoserica
Contrary to what has been accepted in Euro+Med (2006+) based
on Pedersen & Faurholdt (2007), O. holoserica is the correct
name for what has been called O. fuciflora in previous floristic
literature (see Greuter 2008 and Appendix II for details). Orthography of the epithet follows Cribb & Wood (1981), Willing &
Willing (1988) and Buttler & al. (2015); see also IPNI (2012+),
which makes Greuter’s (l.c.) etymological and quantitative arguments in favour of the spelling “holosericea” obsolete.
► Ophrys holoserica subsp. bornmuelleri
Absent but reported in error, confined to SW Asia (Rechinger
1944: 814, under O. bornmuelleri; Pedersen & Faurholdt 2007:
228, as O. fuciflora subsp. bornmuelleri); records from EAe
(Rodos) refer to O. ×vicina (disregarded hybrid).
342
► Ophrys holoserica subsp. candica
The name O. fuciflora subsp. candica E. Nelson ex Soó in Bot.
J. Linn. Soc. 76: 368. 1978 is not validly published (McNeill
& al. 2012: Art. 33.1) and consequently the same is true for O.
candica (Soó) H. Baumann & Künkele. The names have to be
replaced by O. candica Greuter & al. (in Willdenowia 15: 53.
1985) and O. holoserica subsp. candica (Greuter & al.) H. A.
Pedersen & Faurh. (in J. Eur. Orch. 37: 288. 2005), respectively
(nomenclatural advice by E. von Raab-Straube).
► Ophrys holoserica subsp. grandiflora
Absent but reported in error, confined to Cyprus and Anatolia
(Pedersen & Faurholdt 2007: 228 – 229, under O. fuciflora subsp.
grandiflora); records from EAe (Rodos) refer to O. ×vicina (disregarded hybrid).
► Ophrys ×kastelli nothosubsp. antiskariensis
Represents the hybrid O. bombyliflora × O. cretica subsp. cretica, disregarded.
► Ophrys ×kastelli nothosubsp. kastelli
Represents the hybrid O. bombyliflora × O. cretica subsp. karpathensis, disregarded.
► Ophrys ×keramensis
Represents the hybrid O. scolopax subsp. heldreichii × O. tenthredinifera, disregarded.
► Ophrys ×lithinensis
Represents the hybrid O. ×brigittae (O. fusca × O. omegaifera)
× O. omegaifera subsp. omegaifera, disregarded.
► Ophrys ×maremmae
Represents the hybrid O. holoserica × O. tenthredinifera, disregarded.
► Ophrys ×pauliana
Represents the hybrid O. ×brigittae (O. fusca × O. omegaifera)
× O. omegaifera subsp. fleischmannii, disregarded.
► Ophrys ×pezaenensis
Represents the hybrid O. bombyliflora × O. scolopax subsp.
heldreichii, disregarded.
► Ophrys ×plorae
Represents the hybrid O. cretica subsp. karpathensis × O. sphegodes subsp. spruneri, disregarded.
► Ophrys ×pseudoquadriloba
Represents the hybrid O. lutea × O. sphegodes subsp. mammosa, disregarded.
► Ophrys ×pseudospruneri
Represents the hybrid O. sphegodes subsp. mammosa × O.
sphegodes subsp. spruneri, disregarded.
► Ophrys ×rechingeri
Represents the hybrid O. ferrum-equinum × O. sphegodes subsp.
mammosa, disregarded.
Dimopoulos & al.: Vascular plants of Greece: Supplement
► Ophrys ×sivana
Represents the hybrid O. holoserica subsp. candica × O. holoserica subsp. holoserica, disregarded.
► Ophrys ×skopelii
Represents the hybrid O. apifera × O. scolopax subsp. cornuta,
disregarded.
► Ophrys ×sommieri
Represents the hybrid O. bombyliflora × O. tenthredinifera, disregarded.
► Ophrys sphegodes subsp. atrata
A W & C Mediterranean element considered absent from
Greece. Its occurrence in IoI, previously supposed by Hölzinger
& al. (1985: 20, 42), has not been confirmed (Kapteyn den Boumeester & Willing 1988; Hirth 2002). Literature records from
mainland Greece and the Aegean area (Euro+Med 2006+, incorrectly inferred from Boissier 1884: 78) are referable to O.
sphegodes subsp. mammosa and O. sphegodes subsp. sphegodes, respectively (see, e.g., Rechinger 1944: 818 – 819).
► Ophrys sphegodes subsp. litigiosa
The periadriatic range of this chiefly C and SW European taxon
does not include Greece (Euro+Med 2006+; Dimopoulos 2013:
155, 287). Supposed occurrences in IoI (Corfu, Keller & Soó
1931: 52, 388, under O. aranifera subsp. tommasinii) and KK
(Crete, Landwehr 1977: 398) have not been confirmed (see also
Baumann & al. 2006: 198, under O. sphegodes subsp. tommasinii).
► Ophrys ×varvarae
Represents the hybrid O. cretica × O. fusca, disregarded.
► ×Orchinea attica
Represents the hybrid Neotinea tridentata × Orchis italica, disregarded.
► Orchis ×adriatica
Represents the hybrid Anacamptis morio subsp. caucasica ×
Orchis quadripunctata, disregarded.
► Orchis ×bivonae
Represents the hybrid O. anthropophora × O. italica, disregarded.
► Orchis ×dicorifiana
Represents the hybrid O. mascula × O. pauciflora, disregarded.
► Orchis ×kretzschmariorum
Represents the hybrid O. anatolica × O. provincialis, disregarded.
► Orchis mascula subsp. speciosa
Given for Greece in Euro+Med (2006+, probably incorrectly
inferred from O. mascula subsp. signifera in Tutin & al. 1980:
341), although there are no records as a basis for this (Hayek
1933: 390; see also Landwehr 1977: 119; Baumann & al. 2006:
228).
► Orchis ×paschae
Represents the hybrid Anacamptis collina × O. spitzelii subsp.
nitidifolia, disregarded.
► Ophrys scolopax subsp. nestoris
Considered to be a hybrid (= nothosubsp. nestoris) of O. scolopax s.l. with unknown parentage, disregarded.
► Orchis ×plessidiaca
Represents the hybrid O. pallens × O. provincialis, disregarded.
► Ophrys ×sieberi
Represents the hybrid O. cretica subsp. cretica × O. sphegodes
subsp. mammosa, disregarded.
► Orchis ×salkowskiana
Represents the hybrid Anacamptis collina × Orchis sitiaca, disregarded.
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Willdenowia 46 – 2016
► Orchis ×sezikiana
Represents the hybrid O. anatolica × O. quadripunctata, disregarded.
► Orchis ×thriftiensis
Represents the hybrid O. anatolica × O. pauciflora, disregarded.
► Orchis ×willingiorum
Represents the hybrid O. provincialis × O. spitzelii, disregarded.
► Origanum vulgare
Previous records of subsp. viridulum from IoI, Kik and EAe,
and of subsp. vulgare from SPi and StE, are erroneous, based
on misidentified material of O. vulgare subsp. hirtum (rev. S.
Kokkini).
► Ornithogalum corsicum
Confined to Corsica and Sardinia (Euro+Med 2006+). A record
from KK (Karpathos) under its synonym O. sandalioticum
(Zahariadi 1982: 145) is erroneous and likely to represent O.
pumilum.
► Ornithogalum divergens, [O. umbellatum]
Ornithogalum umbellatum was typified by Stearn on triploid
plants (2n = 27) (as shown by Speta 2000a) with few large, leafbearing bulbils and a corymbose inflorescence. This is a mainly
C and W European taxon. Its name is inappropriate for Greek
plants of this complex. Landström (1989) accepted another typification on polyploid material from Spain by Raamsdonk, who
found only hexaploid plants at the type locality (but Moret &
al. 1991 found also triploid ones), which is in conflict with the
protologue, which says “Habitat in Germania, Gallia.” Raamsdonk’s typification has not been accepted recently (see, e.g.,
Jarvis 2007: 709). Triploid plants do not appear in the study
of Landström (1989), where only tetra- to hexaploid numbers
have been counted, so they can be regarded as actually unknown from Greece. Ornithogalum umbellatum in the sense of
Landström is at least largely what is called by Martínez-Azorin
O. divergens from the habit of the plants figured by Landström
and from at least the pentaploid and hexaploid plants. It remains
unclear whether the Greek plants belong to O. divergens at all
(Speta restricted the use of O. divergens to W European plants;
see Speta 2000a: 781), especially the tetraploids. As nothing
has been published and as no other name is available, placing
the Greek plants to O. divergens in a broad sense referring to
Martínez-Azorin & al. (2009) best reflects the current state of
knowledge. It makes no sense to place this unclear complex into
two taxa in Greece. On Crete, there are no distinguishable two
members of this complex (R. Jahn).
► Ornithogalum exscapum
A chiefly Italian species with a transadriatic range comprising westernmost Greece from Kerkira and Vikos to Mt Killini. Records from other parts of Greece (Landström (1989:
22, 30; Strid & Tan 1991: 690) refer to O. collinum (Speta
1990a: 116, 157, 162). Conspecificity of both taxa, as erroneously presumed by Landström (1989, followed in Strid &
Tan 1991), is incorrect. Ornithogalum exscapum exhibits an
epigeal cotyledon, O. collinum on the contrary a hypogeal
cotyledon. The former is rare, the latter abundant in Greece
(see Speta 1990a for details).
► Ornithogalum gussonei
Much confused with O. collinum and O. exscapum, in Greece
only known to occur in IoI (Gutermann 1995; Speta 2000b:
383) and in Pe and EAe (Landström 1989: 37; Speta 1990a:
106, fig. 4). Records from other parts of Greece are most likely
to represent O. refractum (see, e.g., Strid & Tan 1991: 691).
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343
► Ornithogalum oligophyllum
Reported from Thasos by Chilton (2010: 30), apparently based
on a field note. It is a mountain species of the Greek mainland
and Peloponnisos; occurrence on Thasos needs confirmation
(A. Strid).
► Ornithogalum refractum
Old records of this species from Thasos and Samothraki (Stojanov & Kitanov 1945: 272; 1944: 422) need confirmation and
may refer to a form of O. divergens. Ornithogalum refractum is
a rare species of the C & N Greek mainland.
► Osteospermum barberae
Listed by Arianoutsou & al. (2010), planted for ornament, not
established. Native to South Africa, where it is accepted under
its basionym Dimorphotheca barberae Harv.
► Paeonia arietina
Erroneously given for Greece in previous floristic literature, at
specific or subspecific rank (P. mascula subsp. arietina), but
absent (see Strid & Tan 2003). Records from KK (Crete) refer
to P. clusii subsp. clusii, records from StE to P. parnassica, and
records from EAe (Samos) to P. mascula subsp. mascula.
► Papaver dubium
Papaver dubium s.str. (P. dubium subsp. dubium) is reported from SPi, Pe, StE, EC, NC, NE, WAe, Kik, KK and EAe
(Eleftheriadou & al. 1995: 223; Strid & Tan 2003: 89; Willing
& Willing 2007: 92, 2008: 121, 2009: 121). Previous literature
records of P. dubium (s.l.) from IoI (Chilton & Allen 1996: 12),
NPi (Chanlidou & Kokkini 1997: 95; Chitos 2009: 34), and
NAe (Stojanov & Kitanov 1944: 429, 1945: 304; Panitsa & al.
2003: 97) need revision and may refer to P. albiflorum (Elkan)
Pasz., P. confine Jord., or P. lecoqii Lamotte, respectively.
► Paronychia sintenisii
Erroneously given for Greece in Davis (1967: 257), but absent,
endemic to NW Anatolia (Greuter & al. 1984: 235; Tutin & al.
1993: 183).
► Pastinaca sativa subsp. sativa
Vegetable crop listed for Greece in Euro+Med (2006+) merely
calculated from Tutin & al. (1968: 364), disregarded as not established.
► Phelipanche nana
Traditional species concept for this taxon follows Euro+Med
(2006+) although, according to H. Uhlich (pers. comm.), the
morphological differences between P. mutelii and P. nana are
too vague and inconsistent for them to be retained as separate
species. To treat the latter as a variety of the former (Phelipanche mutelii var. nana (Reut.) Uhlich & Rätzel) seems more
appropriate. Subspecific rank is unsuitable since they overlap
completely in distribution and ecology.
► Phelipanche ramosa
This species is confined to the W & C Mediterranean area and
is regarded as being absent from Greece. Greek records refer to
P. mutelii (H. Uhlich, pers. comm.).
► Picea pungens
Occasionally planted for timber in NE (Strid & Tan 1997: 4),
not naturalized.
► Pilosella alpicola
Endemic to the Alps and absent from the Balkan Peninsula, in
Greece replaced by P. rhodopaea (Szeląg 2008).
344
► Pilosella caespitosa
Absent from Greece, replaced there by P. onegensis (Greuter &
Raus 2011: 316).
► Pinus canariensis
Occasionally planted for ornament (Strid & Tan 1997: 5), not
naturalized.
► Pinus pinaster
Occasionally planted for timber and shelter (Tutin & al. 1993:
41; Strid & Tan 1997: 5), not naturalized.
► Pinus ponderosa
Occasionally planted for timber (Tutin & al. 1993: 42), not
naturalized.
► Plantago altissima
Only known with certainty from SPi (Arta, Thesprotia) and NE
(Serres). Almost all other reports of this species from Greece
refer to large forms of P. lanceolata, namely P. lanceolata var.
mediterranea (Kerner) Pilg.
► Plantago macrorrhiza
Not in Greece, confined to the W & C Mediterranean region.
Two collections from Skiros under this designation (Snogerup
3883 and Snogerup & Gustafsson 42775, both at LD) have been
redetermined as P. coronopus s. lat. (rev. P. Lassen).
► Polygonum ×heldreichii
Represents the hybrid Persicaria decipiens × P. lapathifolia,
disregarded.
► Polygonum ×pseudobellardii
Represents the hybrid P. arenarium × P. bellardii, disregarded.
► Polygonum ×pseudopulchellum
Represents the hybrid P. arenarium × P. aviculare, disregarded.
► Polystichum ×lonchitiforme
Represents the hybrid P. aculeatum × P. lonchitis, disregarded.
► Potentilla ×commixta
Represents the hybrid P. detommasii × P. recta, disregarded.
► Potentilla ×degenii
Represents the hybrid P. inclinata × P. pedata, disregarded.
► Potentilla ×dispersa
Represents the hybrid P. pedata × P. pindicola, disregarded.
► Potentilla ×dolosa
Represents the hybrid P. argentea × P. pindicola, disregarded.
► Potentilla ×intercedens
Represents the hybrid P. detommasii × P. pedata, disregarded.
► Potentilla ×kerneri
Represents the hybrid P. argentea × P. inclinata, disregarded.
► Potentilla ×micans
Represents the hybrid P. detommasii × P. pindicola, disregarded.
► Potentilla ×pedatoides
Represents the hybrid P. pedata × P. recta, disregarded.
► Pseudofumaria alba subsp. alba
Erroneously given for Greece in Euro+Med (2006+), incor-
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Dimopoulos & al.: Vascular plants of Greece: Supplement
rectly inferred from Greuter & al. (1989: 290) and Tutin & al.
(1993: 305), see also Lidén (1986: 32).
► Pseudorchis albida subsp. tricuspis
Following Euro+Med (2006+), the rank of subspecies is accepted for this taxon, which was previously considered a variety under P. albida subsp. albida (Dimopoulos & al. 2013:
117; see also taxonomic discussion in Tsiftsis & Antonopoulos
2011: 798 – 799).
► Pseudotsuga menziesii
Occasionally planted for timber (Tutin & al. 1993: 38), not
naturalized.
► Quercus ×kanitziana
Represents the hybrid Q. pubescens × Q. robur subsp. pedunculiflora, disregarded.
► Quercus ×szechenyana
Represents the hybrid Q. frainetto × Q. pubescens, disregarded.
► Ranunculus acris subsp. friesianus
Unsubstantiated literature records from Greece have not been
confirmed and are likely to refer to R. acris subsp. acris (Strid
& Tan 2003: 44).
► Ranunculus auricomus (s. lat.)
The name designates a group of apomictic species, spread all
over Europe (Jalas & Suominen 1989: 171), in Greece only
known to be represented by R. binatus Kit. ex Rchb. so far
(Euro+Med 2006+).
► Ranunculus bulbosus subsp. aleae
Absent from Greece (Jalas & Suominen 1989: 140). Greek
records refer to R. neapolitanus (Strid & Tan 2003: 46).
► Ranunculus carinthiacus
Absent from Greece (Greuter & al. 1989: 425; Jalas & Suominen 1989: 132). Misleadingly given in Euro+Med (2006+),
wrongly inferred from R. oreophilus [subsp. carinthiacus] var.
sartorianus (= R. sartorianus) in Hayek (1924: 339 – 340).
► Ranunculus penicillatus subsp. penicillatus
Absent from Greece (Strid & Tan 2003: 68). Erroneously given
in Euro+Med (2006+), inferred from Tutin & al. (1993: 286)
without being backed by Greek collections seen.
► Ranunculus polyanthemos subsp. nemorosus
Given for Greece (Jalas & Suominen 1989: 121), but no confirmed material has been seen, the records referring to R. polyanthemos subsp. polyanthemoides (see Strid & Tan 2003: 42,
under R. polyanthemoides).
► Ranunculus polyanthemos subsp. serpens
Erroneously mentioned for Greece in Euro+Med (2006+, inferred from Tutin & al. 1993: 273), but absent, confined to
C & SW Europe (Castroviejo & al. 1986: 338; Jalas & Suominen 1989: 122; Greuter & al. 1989: 437).
► Ranunculus pseudomontanus
Erroneously mentioned for Greece in Euro+Med (2006+, inferred from Tutin & al. 1993: 275), but absent, confined to the
Carpathians extending south to Bulgaria and former Jugoslavia
(Jalas & Suominen 1989: 133; Greuter & al. 1989: 426).
► Reseda alba subsp. hookeri
Although erroneously given for “Cr” and “Gr” in Euro+Med
(2006+), this W & C Mediterranean taxon is absent from
Greece (Strid & Tan 2003: 299).
Willdenowia 46 – 2016
► Reseda phyteuma
Infraspecific variability, sometimes overrated as subspecies, deserves at most varietal rank (Castroviejo & al. 1993: 471).
► Rhinanthus illyricus
Erroneously given for Greece in Euro+Med (2006+), the unsubstantiated record incorrectly inferred from Tutin & al.
(1972: 277).
► Rhinanthus pumilus
Rhinanthus pumilus (Sterneck) Pau (in Actas Mem. Prim. Congr.
Nat. Esp. Zaragoza: 248. 1909) antedates R. pumilus (Sterneck)
Soldano (in Atti Soc. Ital. Sci. Nat. Mus. Civ. Stor. Nat. Milano 127: 216. 1986). Represented in Greece by its montane
ecotype, in previous floristic literature called R. mediterraneus.
Rhinanthus mediterraneus (Sterneck) Sennen (in Actas Mem.
Prim. Congr. Nat. Esp. Zaragoza: 289. 1909) antedates R. mediterraneus (Sterneck) Adamović (in Rad Jugoslav. Acad. Znan.
1913: 63. 1913).
► Rorippa sylvestris
A single old record from Kriti by Raulin, cited in Rechinger
(1944: 210) and accepted by Greuter & al. (1986: 153) and
Euro+Med (2006+), not confirmed later and probably incorrect
(see Strid & Tan 2003: map 1023).
► Rosa ×guicciardii
Represents the hybrid R. heckeliana × R. pulverulenta, disregarded.
345
► Saxifraga juniperifolia s.str.
Misleadingly given for Greece in Euro+Med (2006+), but confined to Bulgaria and Caucasus (Strid 1986: 375). In Greece
replaced by S. sancta (Jalas & al. 1999: 150).
► Scilla peruviana
Ornamental plant of SW Mediterranean origin, occasional escape from cultivation, not established, hence disregarded.
► Sedum acre
Old records from Kriti, cited in Rechinger (1944: 292) and accepted by Greuter & al. (1986: 19) and Euro+Med (2006+), not
confirmed later and probably incorrect (see Strid & Tan 2003:
316 and map 1265).
► Sedum rupestre
Absent from Greece (see Strid 1986: 344; Strid & Tan 2003:
330). Greek records refer to S. ochroleucum.
► Sempervivum octopodes
Erroneously given for Greece in Euro+Med (2006+) due to
misinterpretation of Jalas & al. (1999: 53, map 2989). Endemic to former Jugoslavia (SW Makedonija; see Tutin & al.
1993: 426).
► Serapias ×ambigua nothosubsp. ambigua
Represents the hybrid S. cordigera subsp. cordigera × S. lingua,
disregarded.
► Rosa ×oetea
Represents the hybrid R. glauca × R. pulverulenta, disregarded.
► Serapias ×ambigua nothosubsp. panormosana
Represents the hybrid S. cordigera subsp. cretica × S. lingua,
disregarded.
► Rumex ×abortivus
Represents the hybrid R. conglomeratus × R. obtusifolius, disregarded.
► Serapias ×broeckii
Represents the hybrid S. parviflora × S. vomeracea, disregarded.
► Rumex ×dimidiatus
Represents the hybrid R. crispus × R. cristatus, disregarded.
► Serapias ×cythereis
Represents the hybrid S. bergonii × S. cordigera, disregarded.
► Rumex ×halacsyi
Represents the hybrid R. palustris × R. pulcher, disregarded.
► Serapias ×fallax
Represents the hybrid S. bergonii × S. vomeracea, disregarded.
► Rumex ×muretii
Represents the hybrid R. conglomeratus × R. pulcher subsp.
woodsii, disregarded.
► Rumex ×pratensis
Represents the hybrid R. crispus × R. obtusifolius, disregarded.
► Rumex ×semigraecus
Represents the hybrid R. conglomeratus × R. cristatus, disregarded.
► Salix elaeagnos
Infraspecific variability, sometimes overrated as subspecies, deserves at most varietal rank (Castroviejo & al. 1993: 507).
► Saponaria sicula
Given for Greece (Euro+Med 2006+), but absent, merely the
name appearing as an informal aggregate designation with
no nomenclatural status in Greuter & al. (1984: 246). Greek
records refer to S. intermedia (Strid & Tan 1997: 330 – 331).
► Satureja ×boissieri
Represents the hybrid Calamintha incana × C. nepeta, disregarded.
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► Serapias ×halacsyana
Represents the hybrid S. bergonii × S. cordigera, disregarded.
► Serapias ×intermedia
Represents the hybrid S. lingua × S. vomeracea, disregarded.
► Serapias ×kelleri
Represents the hybrid S. cordigera × S. vomeracea, disregarded.
► Serapias ×kelleriana
Represents the hybrid S. bergonii × S. lingua, disregarded.
► Serapias neglecta subsp. apulica
Absent from Greece, endemic to SE Italy (Mte Gargano to
Lecce; see Landwehr 1977: 166; Baumann & al. 2006: 290,
under S. orientalis subsp. apulica). Records for Greece (e.g.
Euro+Med 2006+) refer to S. neglecta subsp. ionica (Baumann
& al. 2006: 285).
► Serapias neglecta subsp. neglecta
Absent from Greece, endemic to SE France (incl. Corsica) and
NE Italy (Baumann & al. 2006: 284). Records for Greece (e.g.
Landwehr 1977: 165) refer to S. neglecta subsp. ionica (Baumann & al. 2006: 285).
346
Dimopoulos & al.: Vascular plants of Greece: Supplement
► Serapias ×semicolumnae
Represents the hybrid S. bergonii × S. lingua, disregarded.
ror (see Strid & Tan 1991: 829). Absent from Greece, according
to Hayek (1932 – 1933: 349).
► Serapias ×semilingua
Represents the hybrid S. lingua × S. parviflora, disregarded.
► Tamarix tetrandra
Island records from the Ionian and Aegean Seas are solely
based on material revised by, or determined on advice of, J.
Zieliński, who advocates the controversial synonymization of
T. parviflora with T. tetrandra; they are all likely to refer to T.
parviflora if the two species are kept separate (Th. Raus).
► Serapias ×sooi
Represents the hybrid S. bergonii × S. vomeracea, disregarded.
► ×Serapicamptis ligustica
Represents the hybrid Anacamptis papilionacea × Serapias
vomeracea, disregarded.
► ×Serapicamptis rousii
Represents the hybrid Anacamptis laxiflora × Serapias vomeracea, disregarded.
► Silene densiflora
Absent from Greece, confined to Crimea, W Transcaucasia and
Anatolia (Wrigley 1986). Greek records refer to S. exaltata
(Strid & Tan 1997: 268 – 269).
► Silene heldreichii
Absent from Greece, endemic to Anatolia. Greek records refer
to S. remotiflora (Strid & Tan 1997: 309 – 310).
► Silene nemoralis
Erroneously given for Greece in Euro+Med (2006+) inferred
from Tutin & al. (1993: 197), but absent (Greuter & al. 1984:
261; Jalas & Suominen 1986: 23).
► Silene nutans
Given for Greece by Tutin & al. (1993) and Greuter & al. (1984)
based on an old and probably incorrectly labelled collection by
Chaubard (Halácsy 1900: 182), hence disregarded. The record
for Greece of S. nutans subsp. insubrica (Gaudin) Soldano
(Euro+Med 2006+) is erroneous (Greuter & al. 1986: 267).
► Silene portensis
Absent from Greece, confined to SW Europe and NW Africa
(Greuter & al. 1984: 269; Jalas & Suominen 1986: 95). Greek
records refer to S. corinthiaca (Strid & Tan 1997: 297).
► Silene uniflora
Absent from Greece, confined to Atlantic W & N Europe (Jalas
& Suominen 1986: 60). Merely the name appears in the synonymy of S. vulgaris subsp. prostrata (see Appendix II), and as
an informal aggregate designation with no nomenclatural status
in Greuter & al. (1984: 276).
► Silene vulgaris subsp. vulgaris
The subspecies was given for Greece in Greuter & al. (1984:
279) and Euro+Med (2006+), inferred from Hayek (1924: 257),
but is probably absent. Only material matching other subspecies of S. vulgaris has been seen from Greece (Strid & Tan
1997: 274 – 278).
► Smyrnium perfoliatum subsp. rotundifolium
Smyrnium perfoliatum subsp. rotundifolium (Mill.) Bonnier
& Layens (Tabl. Syn. Pl. Vasc. France: 135. 1894) antedates
S. perfoliatum subsp. rotundifolium (Mill.) Hartvig (in Strid,
Mountain Fl. Greece 1: 672. 1986).
► Spergularia ×hybrida
Represents the hybrid S. bocconei × S. diandra, disregarded.
► Stipa tirsa
Given for Greece in Tutin & al. (1980: 250), but probably in er-
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► Teucrium spinosum
Reported by Candargy (1898) from Lesvos, but never confirmed by subsequent collectors (I. Bazos, pers. comm.), hence
disregarded. The species is widespread in Anatolia and was illustrated on Plate 539 of Flora Graeca in 1825, based on material collected “in arvis inter Smyrnam [Izmir] et Bursam” (Strid
& Strid 2011: 280).
► Thelypteris palustris
Infraspecific variability within the total range of this cosmopolitan fern deserves at most varietal rank (Flora of North America
Editorial Committee 1993: 213).
► Thymus kosteleckyanus
Reported in error for Greece (Greuter & al. 1986: 385), although there are no records as a basis for this (see Hayek 1931:
352 and Tutin & al. 1972: 179, under T. pannonicus).
► Tragopogon coelesyriacus
Tragopogon coelesyriacus Boiss. (Diagn. Pl. Orient. 2: 47.
1849) antedates T. longirostris Sch. Bip. (in Webb & Berthelot,
Hist. Nat. Îles Canaries 3(2,2): 469. 1850), under which name
this taxon was widely treated in previous Greek floristics.
► Triglochin bulbosa
Reported in error (Halácsy 1904: 143; Euro+Med 2006+), but
confined to S Africa, in Greece replaced by T. barrelieri (Köcke
& al. 2010).
► Tulipa orphanidea
Endemic to the S Greek mainland. Alleged presence in KK
(Euro+Med 2006+) is based on the inclusion of T. goulimyi and
T. doerfleri in a broader concept of T. orphanidea by Zonneveld
(2009) and Christenhusz & al. (2013), which is refused here on
morphological, evolutionary, phytogeographical and ecological
grounds (see, e.g., Fielding & Turland 2005: 523).
► Valantia aprica
Valantia aprica (Sm.) Tausch (in Flora 12: 647. 1829) antedates
V. aprica Boiss. & Heldr. (in Boissier, Diagn. Pl. Orient. 2: 72.
1849).
► Verbascum ×ambracicum
Represents the hybrid V. guicciardii × V. sinuatum, disregarded.
► Verbascum ovalifolium subsp. ovalifolium
A Pontic steppe element occurring as far south as E Bulgaria and
European Turkey, misleadingly given for Greece in Euro+Med
(2006+) based on the record in Tutin & al. (1972: 208) by misinterpretation of the imprecise “Ma” (for Macedonia) in Hayek
(1931: 112, under V. crenatifolium). No material has been seen
from Greece so far (see also Murbeck 1933; Davis 1978: 510).
► Verbascum ×parallelum
Represents the hybrid V. blattaria × V. sinuatum, disregarded.
► Verbascum ×petrophilum
Represents the hybrid V. blattaria × V. pulverulentum, disregarded.
Willdenowia 46 – 2016
347
► Verbena ×adulterina
Represents the hybrid V. officinalis × V. supina, disregarded.
► Viola ×lacmonica
Represents the hybrid V. aetolica × V. orphanidis, disregarded.
► Veronica alpina
Erroneously given for Greece in Euro+Med (2006+), but absent, on the Balkan Peninsula extending south to Albania. The
imprecise “Ma” (for Macedonia) in Hayek (1931: 159) refers to
occurrences in SW Bulgaria and S former Jugoslavia.
► Viola tricolor
According to Erben (1985: 651, corroborating Vandas 1909:
56), records of V. tricolor in previous Greek floristic literature
are all referable to V. macedonica.
► Veronica glauca subsp. kavusica
A single literature record of this Cretan endemic from IoI (Mt
Enos, Strid & Tan 1991: 229) refers to dwarf plants of V. glauca
subsp. peloponnesiaca (rev. M. A. Fischer).
Appendix V: Colour plates. Corrections
► Vicia cracca
No confirmed collections from Aegean islands. Literature
records from Kriti, Lesvos, Kos and Evvia may refer to forms
of V. tenuifolia or V. villosa (A. Strid).
► Vicia monantha subsp. calcarata
Absent but reported in error (Tutin & al. 1968: 133), Greek
records refer to V. monantha subsp. monantha.
Two captions of colour plates in Dimopoulos & al. (2013:
334, 342) should be corrected as follows:
Page 334, Plate 9, caption 9, line 1: replace “Crassulaceae”
with “Cupressaceae”.
Page 342, Plate 13, caption 5, line 2: replace “bifolia” with
“nivalis”.
Willdenowia
Open-access online edition www.bioone.org/loi/will
Online ISSN 1868-6397 · Print ISSN 0511-9618 · Impact factor 0.500
Published by the Botanic Garden and Botanical Museum Berlin, Freie Universität Berlin
© 2016 The Authors · This open-access article is distributed under the CC BY 4.0 licence
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