Pictorial guide Foliage of Hoya Species By Dale Kloppenburg & Dr. Monina Siar

Transcription

1 Hoya Foliage A Pictorial guide Foliage of Hoya Species By Dale Kloppenburg & Dr. Monina Siar Dale Kloppenburg 6427 N. Fruit Ave, Fresno, California All Rights Reserved 1

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3 Hoya foliage, their leaves Hoya leaves are highly variable in size, shape, thickness, texture and venation. They may be glabrous or hairy. The leaf is made up of a petiole and a blade (see labeling on previous page). The blade may have hairs on both surfaces or on only one surface. These hairs are simple or uniseriate. They are thin-walled, very fine, and usually white and vary in length and dispersion. The texture varies from thin e.g. Hoya microphylla, to very thick and succulent e.g. Hoya australis ssp. rupicola, depending on the species and also its habitat. There is usually one or a group of nectariferous glands at the junction of the petiole with the leaf blade. The leaf surface is covered with a relatively thick wax layer (cuticle) above the single celled epidermis. Below the epidermal layer are palisade cells and below this the spongy parenchyma, throughout which is the scattered vein system. Below then the lower epidermis. In Hoya species the mesophyll varies from undifferentiated to strongly differentiated. There is usually a lot of sclerenchymatous mesophyll fibers, like a mat of tangled spider webs. This later also varies in its presence and thickness among the different species. It should be mentioned that leaf size shape are the most variable structure of a hoya plant due to environmental factors of light, nutrition, genetics and other factors even in the wild and more so under domestication. Leaves of hoya may have various distinctive markings on the upper leaf surface. This can be spotting, streaks and blotching. This arises from irregular concentrations of chromoplasts (colored cell inclusions) in cells immediately below the upper leaf surface. Veins give a very characteristic appearance to the leaves and is of two primary types: pinnate and palmate. In pinnate venation the veins branch off the midrib laterally along nearly the entire length. These veins (also referred to as nerves) may be at acute angles to the midrib or in some species perpendicular to the midrib. Most of these veins form loops along and near the margins. Extensive looping connections is termed anastomosing venation. Veins of this type can also be so extensive as to be netted. Veins that arise near the base of the blade in three, five or even seven (including the midrib) primary veins are termed palmate. There are variations as to the positioning of these paired veins off the midrib base, giving rise to many distinct types. The venation of hoya leaves is complex and make an interesting presentation. In this study we have tried to add the angle of the veins to the midrib, primarily the second vein pair from the leaf base. The angle is standard within a range of variability for each species. They do vary, however even within a single leaf and more so under different environments. The data should, however provide some significance in species determinations. Petioles arise from the stem at various angles and are usually slightly twisted. As mentioned before they may be circular but many are grooved on the surface toward the blade. Mostly the petioles are held at about a right angle to the stem. The blade normally then turns another 90º. This creates a anchoring structure, so the vine can entrap itself in surrounding vegetation, thus is an aid in climbing to reach more favorable conditions. 1

4 Leaves may vary in size in species from smaller than a dime to more than 12 long and 8 wide. Even the leaves of one species may be variable. Leaves of wild plants are usually thicker and smaller then the same plant grown in cultivated more ideal conditions. A plants leaves in the wild may look nothing like the foliage that develops in a greenhouse. They not only will show a different size or thickness but even the shape may be drastically different. GROWING HOYAS FROM LEAVES In 1856, Dr. Attilo Tassi, Professor of Botany & Agriculture, Director of the I. & R. Botanical Garden of Lucca presented a paper to the Royal Academy of Lucca, Italy, regarding a seed pod that had matured on a 20 year old Hoya carnosa R. Br. plant. In his lecture he mentioned the methods of propagation for hoyas as being cuttings and from leaves "si moltiplica po per talea, o medianthe le foglie caldo e sotto compana come 1' Aloe, le Glxine, le Calanchoe ecc.". We could add to this that hoyas are also grown from seed. In the past many of us have stuck a leaf in some potting mix, it rooted readily, but then just sat there, not dying, but also never developing into a plant. Most hoya growers have given up on this method. There are, however reasons for developing this means of propagation since there are some very beautiful and exotic foliaged hoyas and leaves which are much easier to mail and transport. This is not a fool-proof method and further study should be conducted on why it succeeds sometimes and not others. A number of years back, Chanin Thorut of Thailand was kind enough to send me 10 leaves and associated flowers from a collecting trip he made into Southern Thailand. These were beautiful leaves, so after making measurements and photographing them, I placed each in its own 4" square dark green plastic pot full of potting mix. These were inserted at a 45 angle and the potting mix firmed up around the leaf peduncle, buried clear up to the leaf base. As usual they rooted quickly in the greenhouse (it was summertime). In the succeeding months 8 of these leaves developed shoots and are now beautiful vigorous healthy plants. Subsequent to this I received a number of exotic and beautiful leaves and flowers collected in Borneo by Ted Green of Kaaawa, Hawaii. I was also able to develop these into plants. The majority of these are Hoya finlaysonii type species along with a couple of Hoya ridleyi types. It seems to me that leaves from plants grown in the tropics will send up shoots much more readily than plants I grow here in Fresno, California, USA. I assume the vigor of the plant, the intensity or type of sunlight or nutrition it has received are factors. It is well established that plants used for tissue culture, form shoots more readily when the derived tissue is from outdoorsvigorous plants. There are some tricks we can use to enhance our success with leaf propagation. The petiole of the leaf calluses over and also produces a mass of roots. The trick is to get this callous to produce meristematic cells and develop shoots. The idea is to stimulate it either by mechanical or chemical means (providing it does not act spontaneously). Sometimes, you can raise the root mass slightly making small cuts into the petiole to mechanically irritate the tissue. This alone will provide the necessary stimulation. 2

5 Chemical stimulation is by means of hormones such as adenine sulfate, kinetin, 2-4 D, gibberelic acid, etc. Since I had a number of these chemicals left over from tissue culture work, I have tried all of the above. 2-4 D is a weed killer and needs to be used in an extremely weak solution-say a drop in a gallon of water (the actual upper limit is 3 parts per million (ppm). I use it with an eyedropper and let the solution run down the center of the leaf on to the root mass and petiole. Gibberelic acid, I have found, produces a weak spindly growth that is unsatisfactory. Lately, I have also used indole-3-buturic acid; only a small crystal in a glass of water and painted it on the leaf-base and petiole after they have rooted. I have not tried these on small or thin leaves. I have had mixed success with Hoya carnosa, pallida, kerrii, diversifolia and finlaysonii. I am sure the process is possible but needs further research. It is worth pursuing and perfecting. Other aspects of the Hoya Plant Leaf petiole, base of leaf blade, usually twisted, round and often grooved above. Node (usually a pair of leaves arise here, however occasionally one will fall. Sometimes 2-3 pairs of leaves will appear to arise from a single node). Leaf blade. Internode (stem between nodes). A typical section of a hoya stem (reduced). Leaf base, here obtuse (rounded). Leaf apex (tip), here acute. 3

6 Terms Associated With Hoya Leaves Not a complete coverage of terms Forms and shapes Definitions from Botanical Latin by W. T. Stern Apex blunt (obtuse) Shape (oblanceolate) Wider toward the apex Base (cuneate) Obtuse (Blunt) terminating gradually in a rounded end. Oblanceolate..reversed lanceolate. Cuneate..wedge shaped. Apex (obtuse) Shape (ovate) Base (obtuse) Ovate broadest at the lower end. (egg like) Apex (acuminate) Shape (elliptic-lanceolate) Base (obtuse) Acuminate Tapered point. Elliptic...(oval) acute at both ends. Lanceolate narrowly elliptic, tapering to each end. 4

7 Apex (acute). Shape (elliptic) Base (cordate) Acute sharp pointed; without tapering in any degree. Cordate...having two rounded lobes at the base. Shape (orbicular) Orbicular perfect circular. Shape(suborbicular) or sub-rotund. Sub nearly or almost. Rotund..a little inclined to be oblong, broadest in the middle. Apex (apiculate) Shape Elliptic-ovate.(oval) Base sub-truncate to obtuse Apiculate terminating abruptly in a little point. Truncate.as if cut off straight across. 5

8 Shape (linear) or (subulate) or (fasciculate) Linear narrow, line like. Subulate linear, very narrow, tapering to a fine point. Fassciculate..narrow, very long. Apex (truncate) Shape (cuneiform) Truncate terminating very abruptly. Cuneiform...wedge shaped. Edges (undulant) Undulant..wavy. Shape (oblong) Oblong elliptic, obtuse at each end. 6

9 Nervation (veins) types. Pinnately veined leaf. With nerves on opposite sides of the midrib. A Palmately veined leaf. 3-5 nerves from the leaf base. Trinerved, Quinquinervis, Septinervis all defining the number of nerves arising from the leaf base. Hoya pottsii is a Trinerved example. 7

10 Another Palmate veined leaf. Many leaves that are palmate as the one above have cordate or obtuse bases but some like this one have narrow bases, nearly cuneate. This one is 3 palmate. A Triplinerved species. With the two lateral nerves emerge from the midrib a little above its base. If Quintuplinerved (with 5 nerves) etc. Triplinerved hoya species. Hoya samoensis, Hoya upoluensis and Hoya pycnophylla are all quintuplinerved species. Also Hoya aurantiaca and Hoya siarae. 8

11 Key Type Nerves: P...pinnate T...triplinerved Pl...palmate O...obscure 5...quintuplinerved R...reticulate...perpendicular Shape: V...narrow linear thick O...ovate C...cordate E...elliptic Ob...obovate Oc...obcordate Obl...oblong L...lanceolate Apex/Base: Cu...cuneate Ob...obtuse Cd...cordate Ap...apiculate A...acute Ac...acuminate At...attenuate Ø...glabrous P...pubescent All measurements converted and presented in Centimeters, cm. Species Type Apex/ Measurements nerve base Shape cm. Hoya acicularis Green & Klopp P A A terete 3 x Hoya acuminata Bentham ex Hook P/O Ac Ø/P Obl x Hoya acuta Haworth 1821 T/P O/A P L/Ac 5-10 x 2.5 Hoya aeschynanthoides Schlechter 1908 P O/A O E x Hoya africana Decaisne 1844 Ob O/Obl Hoya alagensis Kloppenburg 1990 P/R O/A Ø L 11 x Hoya alata K. Hill 1986 PL Cd A Ø O 3.7 x Hoya albiflora Zipp. ex Blume 1848 P Ob A Ø O x Hoya aldrichii Hemsley 1890 T ObAc Ø O 8-12 x Hoya alexicaca Moon 1824 Cd A Ø Hoya amboinensis Warburg 1907 T Ob A Ø Ob 6-7 x Hoya ambyolepis Schlechter P ObOb Ø Ob Hoya amoena Bakhuizen 1950 T Cu A Obl 8-11 x Hoya angustifolia Elmer ex Burton P CuOb ØV 3-12 x Hoya angustifolia Traill 1830 P CuOb Ø L x 1.25 Hoya angustisepala Burton 1987 P Ob A Ø Obl 15 x 6 Hoya andalensis Kloppenburg 2005 P Ob ØO 5 x 2.2 Hoya anncajanoae Klopp. & Siar 2007 P CuAc Ø LE 5-9 x Hoya anulata Schlechter 1905 PL CuAc OObl 6-8 x Hoya apiculata Scheffler 1876 P CdAp ØOrb 7.5 x 5 Hoya apoda S. Moore 1916 Hoya apoensis Kloppenburg & Siar 2010 P Cu Ac Ø E x Hoya archboldiana C. Norman 1937 P Cd A Ø Obl 13 x 7 9

12 Hoya arnottiana Wight 1834 P ObAc Ø EL Hoya attenuata Christophersen 1934 P ObAt Ø EL 5-8 x Hoya aurantiaca Klopp. & Siar C- Ac Ø Ob x Hoya australis R. Br P Ob Ac ØO Ob 2-8 Hoya aust. subsp. nathaleae 2010 P C Ap P O 6.5 x 3.6 Hoya bandaensis Schlechter 1908 P ObAc Ø E O x 5-7 Hoya bella Hooker 1848 P ObAc Ø O 2 Hoya benguetensis Schlechter 1906 T ObAc Ø EO 6-10 x Hoya benvergarai Klopp. & Siar ObAc Ø EL 9-13 x 2-3 Hoya betchei Schlter. ex Whistler 1913 P ObAp Ø LE x Hoya bhutanica Grierson & Long 1979 T P Ob A Ø E 9-15 x Hoya bicarinata A. Gray 1861 P O Ac P O Hoya bicknellii Kloppenburg 1999 P CuAp Ø O 9-12 x Hoya bicolensis Kloppenburg 1994 T ObAc Ø OL x 0.4 Hoya bicolor Kloppenburg 2003 P O Ap Ø OCd Hoya billardieri Decaisne 1844 CdAc Ø O Hoya bilobata Schlechter 1906 P ObOb ØE OB x Hoya blashernaezii Kloppenburg 1999 T CuA Ø EL Hoya blumeana Schlechter 1913 Hoya bonii Costantin 1912 P ObA Ø O 8-10 x Hoya bordenii Schlechter 1906 T ObAc Ø LObl x Hoya brevialata Kleijn &Donkelaar 2001 P Ob Ø ObE x Hoya brittonii Kloppenburg 1992 P Cu Ø 8 x 3 Hoya brooksii Ridley 1925 P Cu A Ø E 5 x 2 Hoya browniana Koorders 1911 T CuAc Ø Obl 20 x Hoya bruensis Miquel 1869 P OB A Ø OE Hoya brunioniana Wight 1834 P ObAc ØObl+O x 1.75 Hoya bulusanensis Elmer 1920 see H. panchoi T P ObAc + O Ø Ob + O Hoya burmanica Rolfe 1920 P Ac Ø O L 5-8 x Hoya buotii Kloppenburg 2002 P ObAc ØEL 10 x 2.5 Hoya burtoniae Kloppenburg 1990 P O A PO 3 x diam x Hoya cagayanensis (Sch.ex Elmer) P Ob A Ø OL x Burton1987 Hoya callistophylla Green 2000 P CuAt ØEL x 6-10 Hoya calycina Schlechter 1908 P CdAp PO x 9-12 Hoya campanulata Blume 1826 P ObAc Ø OL x Hoya camphorifolia Warburg 1904 T oba Ø OE 6-9 x Hoya cardiophylla Merrill 1920 P CdAc Ø C Hoya carnosa R. Brown 1802 P ObAc ØOObl x 3-4 Hoya caudata Hooker 1883 P O Ac ØO x Hoya cebuensis Klopp. & Siar ObA ØOL 9 x 4 Hoya cembra Kloppenburg 1990 P Ob A ØE 8-10 x 2-3 Hoya chinensis Traill 1830 H. carnosa Hoya chlorantha Rechinger 1908 P CuAc Ø L 10 x 2 10

13 Hoya chlorantha var. tutuilensis 1935 P O A Ø O x 6-8 Hoya chloroleuca Schlechter 1913 P Cu A Ø LE 6-8 x Hoya chunii P. T. Li Hoya cinnamomifolia Hooker 1848 PL ObAc Ø O 12.5 x 6.2 Hoya citrina Ridley 1922 PL Ob A Ø C x 7.5 Hoya clandestina Blume 1848 P ObAc Ø EObl Hoya collettii Schlechter 1913 P ObA Ø LObl Hoya collina Schlechter 1913 P CuOb Ø EO x Hoya cominsii Hemsley 1890 PlT CdA Ø E O x 6.3 Hoya commutate Gilbert & Li 1995 Hoya compacta Burton 1990 P Cd Ac Ø x 3 Hoya cordata Li & Huang 1985 P Cd A Ø 5-9 x Hoya coriacea Blume 1826 P ObAc ØE-Ob Hoya coriacea Lindley 1839 H. multiflora Hoya costantinii P. T. Li 1984 Hoya crassicaulis Elm. ex Klopp P ObA ØE 8-17 x 5-8 Hoya crassifolia Haworth 1811 P ObAC ØOb 10 Hoya crassifolia Ridley 1912 P ObAc ØOb 15 x 6.3 Hoya crassior Hochreutiner 1935 P CuAt ØL 4-6 x Hoya crassipes Turczaninow 1948 P ObAc Ø EOb 7-13 x Hoya cumingiana Decaisne 1844 P ObA PC Hoya curtisii King & Gamble 1903 P OBA ØO 1.5 x 1.5 Hoya cystiantha Schlechter 1913 P CuAc ØO-L x Hoya dalrympleana F. Muell 1861 Hoya darwinii Loher 1910 ObA ØEObl x 5-6 Hoya dasyantha Tsiang 1936 P ObA ØO x Hoya davidcummingii Klopp P CuOb Ø LE x Hoya dennisii Forster et D. Liddle 1993 P ObAc ØLO 7.2 x 3.0 Hoya densifolia Turczaninow 1848 P ObOb Ø OblE x 2-4 Hoya dickasoniana P. T. Li Cu-Ob Ob Ø E O x Hoya dictyoneura K. Schumann ObA ØObl x Hoya dimorpha F. M. Bailey 1898 P ObAc ØObl x 5 Hoya diptera Seemann 1861 P ObA ØOE x Hoya dischorensis Schlechter 1913 P ObA ØO x Hoya diversifolia Blume 1826 P ObOb ØOE x Hoya dodecatheiflora Fosberg 1940 P CuAc ØEL 8 x 12 Hoya dolichosparte Schlechter Ø x Hoya edanoi C. M. Burton 1991 P Cu Ob Ø R 3-7 x 3-5 Hoya edeni King ex Hooker 1883 P CuAc ØObE x Hoya eitapensis Schlechter 1909 P CuA ØEO 3-5 x Hoya elliptica Hooker 1883 P ObOb ØE x Hoya elmeri Merrill 1929 P CuAc ØE x Hoya el-nidicus Kloppenburg 1991 P ObAp ØO x Hoya endauensis Kiew 1989 P ObOb ØOCd x

14 Hoya engleriana Hosseus 1907 P ObA PEL 1.5 x 0.4 Hoya epedunculata Schlechter 1913 P CuAc ØE x Hoya erythrina Rintz 1978 PL CdA ØObl 15 x 5 Hoya erythrostemma Kerr 1939 P CuAc ØE 10 x 4 Hoya esculenta Tsiang 1936 P Hoya excavata Teijsm. & Binn P ObAp ØOObl 15 x 9 Hoya exilis Schlechter 1913 P ObAc ØEL 5-10 x Hoya fauensis Klopp. & Siar 2008 P ObAp ØOE 4-5 x 2.2 Hoya fetuana Kloppenburg 2003 P ObAt ØE Hoya filiformis Rechinger 1908 P ObAc ØLO 6 x 2 Hoya finlaysonii Wight 1834 P ObAc ØOblL x Hoya fischeriana Warburg 1904 T CuA ØL x 4-7 Hoya fitchii Klopp. & Siar P ObAp ØO Hoya flagellata Kerr 1940 P CdA ØL x Hoya flavescens Schlechter 1913 P CuAc ØLObl x Hoya flavida Forster et D. Liddle 1993 P CuA ØEO 1.4 x 4.5 Hoya forbesii King & Gamble 1903 P ObAc ØEObl x Hoya formosana Yamazaki ObAt ØO 7-9 x 4-5 Hoya fraterna Blume 1849 P CdA ØOObl x Hoya fungii Merrill 1934 P ObA ØEObl 8-14 x Hoya fusca Wallich ObAc ØLObl x Hoya fuscomarginata N. E. Brown 1910 P CuAp ØE x Hoya gigantanganensis Klopp P CuAp ØE x Hoya gildingii Kloppenburg 2001 P CuAp ØL Hoya glabra Schlechter 1908 PL ObAc ØO x Hoya globiflora Ridley 1915 H. acuta PL-T Ob-A ØO 6.5 x 3.5 Hoya globulifera Blume 1849 P ObA ØOE Hoya globulosa Hooker 1882 P ObAc PEObl 7-8 x Hoya golamcoana Kloppenburg 1991 P Ob ØOc 3 x 2.5 Hoya gonoloboides Regel 1883 P CdAc PCdO Hoya gracilipes Schlechter 1905 P CuAc ØEL x 2-3 Hoya gracilis Schlechter 1908 P CuA ØEL x Hoya grandiflora Sprengel 1825 P ObAc ØO-L Hoya graveolens Kerr 1939 P CuAc ØOblE x 2-3 Hoya greenii Kloppenburg 1995 P ObAc Ø E x 3-4 Hoya griffithii Decais.1883 H. acuminata Hoya griffithii Hooker 1883 P ObAc ØObl L Hoya gymnanthera Wight 1834 leafless Hoya hainanensis Merrill 1923 P A Ac ØOE 6 x Hoya halconensis Kloppenburg 1990 P ObA ØOE x 4-6 Hoya halophila Schlechter 1913 P ObA ØO x Hoya hasseltii Miquel 1856 P ObA ØO 5-7 Hoya hellwigiana Warburg 1907 P CuA ØLObl 9-15 Hoya hellwigii Warburg 1901 ms Hoya heuschkeliana Kloppenburg 1989 P CuOb ØOb 2.5 x

15 Hoya hirsuta Moon 1824 Tylophora Hoya hookeriana Wight 1834 PL ObA ØOblL x Hoya hypolasia Schlechter 1913 P CdAc Ø/PL x Hoya imbricata Decaisne 1844 P ObOb ØO 6-12 round Hoya imperialis Lindley 1846 P CdA ØObl x Hoya inconspicua Hemsley 1894 P ØL -10 Hoya incrassata Warburg 1904 P ObAc ØOObl 9-14 x 5-6 Hoya incurvula Schlechter 1916 P ObOb ØOE x Hoya intermedia Schlechter ms name Hoya intermedia A. C. Smith 1942 P ObA ØObl 8-12 x Hoya ischnopus Schlechter 1913 P CdOb ØO 9-11 x 5-6 Hoya juannguoana Kloppenburg 2002 P CuAp ØOE x Hoya kanyakumariana Henry & P ObOb ØObObc x Swam.1978 Hoya kastbergii Kloppenburg 2003 P ObOb ØO 6 x 4.5 Hoya kenejiana Schlechter 1913 P ObA Ø Obl 6-10 x Hoya kentiana Burton 1991 P Cu A Ø E 3-12 x Hoya kerrii Craib 1911 P CuCd ØOb 4-12 x Hoya keysii Bailey 1884 P CdA POCd x Hoya kingdonwardii P. T. Li 1994 Cu A Ø E x Hoya klosii Moore 1906 P ObOb ØO 4-6 x Hoya krohnian Kloppenburg & Siar 2010 P C Ap Ø O Hoya kuhlii Koorders 1012 P CuAc ØL 5-15 x Hoya kwangsiensis Tsiang et Li 1974 P ObA ØL 8-17 x 2-4 Hoya lactea S. Moore 1914 P ObOb ØObO 9-12 x Hoya lacuna Wight 1834 P ObAc Ø-P x Hoya lacunosa Blume 1826 P ObA ØO-OE Hoya lamingtoniae Bailey 1898 T CuAc ØL x 2.5 Hoya lanceolaria Moore 1916 P ObAt ØOObl x Hoya lanceolata Lindley 1826 = acuta Hoya lanceolata Wallich ex Don 1825 P CuAc ØL Hoya lancilimba Merrill 1932 P CuA ØL x Hoya landgrantensis Klopp. & Siar 2008 T Ob Ac Ø L E 8.8 x 3 Hoya lantsangensis Tsiang et P. T. Li P CuCd ØOb 2-3 x Not a Hoya Hoya lasiantha Blume ex Korthals 1848 P ObAc ØE L x Hoya lasigynostegia P. T. Li 1984 P ObA ØOL x Hoya latifolia G. Don 1838 PL CdAc ØO x Hoya laurifolia Decaisne 1838 P CuA ØLObl 6-10 x Hoya laurifolia Miquel 1863 P ObA ØOObl Hoya laurifoliopsis Hochreutiner 1936 P CuAc ØE 6-10 x Hoya lazeroi Kloppenburg & Siar 2007 P Cu Ac Ø E 11 x 2.8 Hoya leembruggeniana Koorders 1911 P CuA ØEObl x Hoya leucantha S. Moore 1916 P ObA ØLAc x Hoya leucorhoda Schlechter 1913 ObAc ØO 7-11 x

16 Hoya leytensis Elmer ex Burton 1938 P Ob ØObl R x Hoya liangii Tsiang 1936 P CdOb ØObObl x Hoya limonica S. Moore ObAt ØO 7-8 x Hoya linavergariae Klopp. & Siar 2006 P ObA ØEL 4-5 x Hoya lindaueana Koorders 1911 T CuAc ØE x 5-10 Hoya lindleyana F. Vill Hoya linearis Wallich 1834 P CuA PL Hoya lipoensis P. T. Li 1985 P Cucaud. ØE 9-15 x 3-5 Hoya litoralis Schlechter 1905 P CuAc ØL x Hoya lobbii Hooker 1883 P CdA ØE-O Hoya loheri Kloppenburg 1991 P CuA Ø Ob x Hoya longifolia Wallich 1834 P ØL Hoya longifolia (Bl.) Miquel 1857 P CuAc ØL x Hoya lucardenasiana Klopp. & Siar 2008 P Ø O 5.5 x 3.8 Hoya lucyae Klopp. & Siar 2006 P CuA Ø E L 6-10 x Hoya luzonica Schlechter 1904 P ObA ØObl 9-13 Hoya lyi Leveille 1907 P ObOb POblL x Hoya macgillivrayi Bailey 1914 P CdAc ØEO 10 x 5 Hoya macgregorii Schlechter 1906 P CuAc ØLE 7-11 x Hoya macrophylla Blume 1826 T ObAc ØOObl Hoya macrophylla Wight 1834 = latifolia PL Ob-Cd Ac Ø O x Hoya magnifica Forster & Liddle 1995 P CdAAc PEO 19 x 10 Hoya magniflora P. T. Li 1984 not hoya Hoya maingayi Hooker 1883 P AcOb ØELOb 8-20 x Hoya manipurensis Deb 1954 non Hoya P CuObCd Ø 2-3 x Hoya marginata Schlechter CdAc ØEO x Hoya matavanuensis Klop. & Siar 2008 P ObA ØEAt x Hoya maxima Teijsm. & Binnen ObAc2 ØEObl 20 x 10 Hoya maxima Warburg 1907 Hoya megalantha Turrill 1915 P ObAt ØEO 6.5 x 3 Hoya megalaster Warburg 1901 P CdAp ØO 8-14 x 5-9 Hoya mekongensis Gilbert & Li 1995 P Ob Ac P Ob L x 4-5 Hoya meliflua Blanco ex Merrill 1837 P ObA ØObl 9-13 x 5-6 Hoya membranifolia Costantin 1912 P ObAc ØL 7-12 x Hoya memoria Kloppenburg 2004 P ObA ØL Hoya mengtzeensis Tsiang & Li 1974 P ObAc ØL 6-11 x Hoya meredithii Green 1989 P ObAc ØO x Hoya merrillii Schlechter ObAc ØO 6-9 x Hoya micrantha Hooker 1883 P CuA ØOE 5-10 x Hoya microphylla Schlechter 1913 P ObA ØO x Hoya microstemma Schlechter 1913 P CuA ØE x Hoya minahassae Schlechter 1916 P CuA ØLE 4-7 x Hoya mindanaensis Elmer 1938 P ObA ØEL 15 x 6 Hoya mindorensis Schlechter 1906 P ObAc ØOblE 9-12 x

17 Hoya mindorensis ssp. superba Klop.& P ObA Ø E 9-12 x Siar Hoya minima Costantin 1912 P CuA ØL x Hoya mitrata Kerr 1940 P ObAc ØOblE x Hoya montana Schlechter 1913 P ObAc ØOblE x Hoya motoskei Teijsm. & Binn P OBA PO Hoya mucronulata Warburg 1901 P ObOb ØObl 6-12 x 3-5 Hoya multiflora Blume 1823 P CuAc ØOblL x Hoya myanmarica P. T. Li 1994 P Cu-R Ac Ø Ob x Hoya nabawanensis Klopp& Siar2008 P ObA Ø OL 6-10 x Hoya naumanii Schlechter 1908 P CdAp ØE 10 x 7.5 Hoya neocaledonica Schlter PL ObAc ØOObl 5-9 x 3-5 Hoya neoebudica Guillaumin CuAAc ØL 7-9 x Hoya nervosa Tsiang et Li 1974 P ObA ØOObl x Hoya nicholsoniae Mueller ObAc ØO x Hoya nicobarica Br. ex Traill1830 P ObA ØO 5-10 x Hoya nyhuusiae Kloppenburg 2003 P Ob A ØL Hoya nummularia Deca. ex P ObOb ØO 1.9 round Hook.1883 Hoya nummularioides Costantin 1912 P ObAp ØO x Hoya nuuliensis Klopp. & Siar 2008 T CuA ØE 6.8 x 2 Hoya obcordata Hooker 1883 P ObA ØO 1.2 x 1.2 Hoya obcordata Teijs. & Binn.1866 Hoya oblanceolata Hooker 1883 P CuAc ØObl Hoya oblongacutifolia Cost P CuAc ØOblL 5-8 x Hoya obovata Decaisne 1844 P ObOb ØOb 5-11 x Hoya obreniformis King 1901 P ObCd ØOc x Hoya obscure Elmer ex Burton 1986 P CuAp ØE-Ob 2-10 x 4 Hoya obscurinervia Merrill CuA ØE 10 x Hoya obtusifolia Wight 1834 P CdAp ØObl x Hoya occlusa Ridley 1912 P ObAc ØO 10 x 5 Hoya odetteae Kloppenburg 1998 P CuA ØLE 2-7 x 1 Hoya odorata Schlechter 1906 P CuAc ØE-L x Hoya odoratissima Lind.ex Regal 1884 Hoya oleoides Schlechter 1913 P OBAc ØOObt x Hoya oligantha Schlechter 1913 P ObAp ØL x Hoya oligotricha K. Hill 1986 P Ob A ØEO 4-12 x 3-8 Hoya onychoides P. I. Forster 1995 P CdAAp ØLO 12 x 5.5 Hoya opposita G. Don 1834Sperlingia P ObAc ØEO Hoya orbiculata Wall. ex Wight 1834 P ObA Ø 1.3 x 1.3+ Hoya oreogena Kerr 1939 P Cu -Ob ØE 5-7 x Hoya oreostemma Schlechter 1913 P ObAc ØLE x

18 Hoya orientalis Li 1984 H. reticulate Merrill Hoya ottolanderi Koorders CuAc ØL x Hoya ovalifolia Wight & Ar P CuAAc ØE 5-7 Hoya ovalifolia Wallich revoluta 1834 Cu Ac Ø O Hoya oxycoccoides S. Moore 1916 P CuOb ØOOrb x Hoya pachyclada Kerr CuAc ØOb x Hoya pachyphylla Lauter.& P ObAp ØEObt 7-11 x Schu.1907 Hoya pachypus Moore 1914 P ObObA ØOObl x 4-5 Hoya padangensis Schlechter 1916 P CuAAc ØE 7-10 x Hoya palawanica Kloppenburg 1990 P EL x Hoya pallida Lindley 1826 acuta P ObAc ØOL 5-10 x 2.5 Hoya pallida Dalz. & Gibs 1861 Hoya panayensis Klopp. & Siar P ObAp E x Hoya panchoi Kloppenburg 1988 P ObOb Ø O Hoya pandurata Tsiang 1939 P ObAc ØObl 4-7 x 1 Hoya paniculata Sprngl Tylophora Hoya papillantha K. Schumann 1898 P ObA ØObl 11 x 7 Hoya papuana Schlechter 1913 P CuAc ØLE 6-8 x Hoya parasitica Wallich 1830 acuta T/P O/A Ø L/Ac 5-10 x 2.5 Hoya parviflora Wight 1834 P Ac ØL x Hoya parvifolia Schlechter 1908 P CuA ØL 5-10 x Hoya patella Schlechter 1913 P ObAc POblE x Hoya pauciflora Wight 1848 P ObOb ØLObl x 0.6 Hoya paxtonii Nichols 1852 bella? Ac Hoya paziae Kloppenburg 1990 P ObAc Ø E 6-7 x Hoya pedunculata Schlechter 1913 P ObOb ØObl-E x Hoya pendula Wight & Arnott 1834 P ObAc ØOblO Hoya pendula Wight 1918 P ObOb ØEO-L 3.8 x 6.3 Hoya pentaphlebia Merrill ObA ØO-E 8-16 x 6-9 Hoya perakensis Ridley ObA ØO 11.3 x 7.5 Hoya persicinicoronis He, Li et al. P Cu Ob Ø E O 6-10 x 4-5 Hoya philippinensis Li Hoya phyllura O. Schwartz 1931 P CuAc ØOblE 8-16 x Hoya picta Siebold 1848 P ObAt ØOL Hoya picta Blume 1848 P ObA ØO Hoya picta Miquel 1856 P ObA ØO x 4.4 Hoya piestolepis Schlechter 1913 P CdAc ØOblE x Hoya pilosa Seemann 1861 P CdA Ø /PE Hoya planiflora Wall. ex Wight 1861 P CdAc PO

19 Hoya plicata King &. Gamble 1908 P CuAc ØOL 7.5 x 2.5 Hoya piminteliana Klopp P CuAc P L 6-10 x Hoya polyneura Hooker 1883 P ObAc ØOL x Hoya polystachya Blume ObAc ØOObl Hoya poolei White & Francis 1828 P AcCu ØLO 6-9 x 2-3 Hoya pottsii Traill 1830 PL CdAp Ø O Hoya praetorii Miquel 1856 P ObAc ØObl-O Hoya pruinosa Miquel 1856 P ObOb ØEOb -1.3 Hoya pseudolanceolata Cost.1912 P CuAt ØL 4-9 x Hoya pseudolittoralis Norman 1937 P ObA ØO 7-9 x Hoya pseudomaxima Koorders 1919 imbricate Hoya pseudovalifolia Costantin1912 P ObAc ØO x Hoya pubens Costantin 1912 T ObAp ØoObl x Hoya puber Blume 1826 P ObA PEObl Hoya pubescens Reinecke 1898 P ObAc POOb 6-9 x 4-6 Hoya pubicalyx Merrill 1918 P ObAc ØOblO x 3-5 Hoya pubifera Elmer 1938 P ObOb POblL 12 x 3 Hoya pulchella Schlechter 1913 P ObA ØOOb x Hoya purpurascens Teij.& Bin.1863 P CdA ØO 8.5 x 5 Hoya purpureofusca Hooker ObtAc ØO Hoya pusilla Rintz 1987 P ObAc ØOL 8 x 3 Hoya pusilliflora S. Moore 1916 P ObOb ØL-O 6-7 x Hoya pycnophylla Rechinger CuA ØL 9-10 x 2-3 Hoya querinoensis Klopp.& Siar ObAp Ø EL 16.5 x 5.5 Hoya quinquenervia Warburg 1904 PL ObAc ØO 6.5 x 4 Hoya quisumbingii Klopp P CuAp ØL x Hoya radicalis Tsiang et Li1974 P CuAc ØObl x 3-4 Hoya ramosii Klopp. & Siar 2007 P Cu Ac Ø L x 0.6 Hoya rosarioae Klopp. & Siar 2009 P Cu Ap Ø Ob L x Hoya recurvifolia Zipp.ex Blume1849 Hoya recurvula Kloppenburg 2000 P ObA Ø L 8-12 x 3.5 Hoya reflexa Bent. & Hooker 1880 Hoya reticulata Costantin 1912 P CuAc ØObl x Hoya reticulata Schlechter 1913 P ObOb ØE x 3-5 Hoya reticulata Merrill 1920 P Obac ØOObl 8-12 x 4-7 Hoya retusa Dalzell 1852 ObOb Ølinear x Hoya retusa Warburg 1907 P ObAc POb x 7-10 Hoya revolubilis Tsiang et Li 1974 P CuOb ØE 6-9 x 2-3 Hoya revoluta Wight 1883 P CuOb ØOL x Hoya rhodostele Ridley 1923 P Cu A ØO-L x Hoya rhodostemma Schlechter 1913 P CuAc ØL 9-14 x

20 Hoya ridleyi King & Gamble 1908 P CuA ØOObl x Hoya rigida Kerr Cuac ØO-E x Hoya rizaliana Kloppenburg 1991 P CuA ØOE 3.6 x 1.2 Hoya rosea Schumann 1905 P CuAc ØL x Hoya rotundifolia Siebold 1840 P ObA ØEO x 10 Hoya rubida Schlechter 1901 P ObAc ØO-E 4-6 x 2-3 Hoya rufolanata Ridley 1923 P ObA PO x 7.5 Hoya rumphii Blume 1826 P ObAc ØObl L x 5 Hoya rupicola K. Hill 1988 P O A Ø O 14 x Hoya ruscifolia Decaisne 1844 P A ØO Hoya salweenica Tsiang et Li 1974 P ObOb ØEObl x Hoya samoensis Seemann 1866 T ObAc ØOE 7-10 x Hoya sana Bailey 1897 P ObA PObL Hoya sarcophylla Ridley 1917 ObA ØL 8.5 x 3 Hoya schlechteriana Moore 1916 P OB ØObt x Hoya schneei Schlechter 1921 P CuAc ØE 8-11 x Hoya scortechinii King & Gamble 1908 P CdAc ØO-L Hoya serpens Hooker 1883 P ObOb POE Hoya shepherdii Short ex Hooker 1861 P CuA Ø V 5-15 x 1.3 Hoya siamaca Craib 1911 P ObAc ØLObl x Hoya siarae Kloppenburg 2002 T ObA Ø L 14 x 3.5 Hoya sikkimensis Graf 1992 Hoya silvatica Tsiang et P. T. Li 1974 P CuAc PEObl 8-11 x Hoya sipitangensis Klopp.&Wiberg 2002 P CuA Ø L Hoya skinnerae Klopp. & Siar P ObA Ø Hoya smithii Klopp. & Siar P ObA Ø L x 8-12 Hoya sogeriensis Moore 1913 T or PL ObOb ØEObl x 3-4 Hoya solaniflora Schlechter 1913 P ObAc ØL x Hoya soligamiana Klopp. & Siar 2008 T Ob Ac ØEL 9-13 x 2-3 Hoya sororia Schumann 1905 ObAc 5-6 x 1 Hoya spartioides (Kuntz)Klopp T ObOb Ø OE 1 x..5 Hoya splendens Maingay 1837 Hoya stenophylla Schlechter 1913 P ObAp PV 6-8 x Hoya stoneana Kloppenburg 2006 P ObA POL x Hoya suaveolens Miquel 1856 P ObAc ØOE Hoya subglabra Schlechter 1913 P Ac ØE x Hoya subquaterna Miquel 1856 T Ac ØO Hoya subquintuplinervis Miquel ObAp ØO x Hoya tamaleaaensis Klopp. & Siar 2009 Ob At Ø OE x Hoya tauensis Klopp. & Siar ObAt ØOE x Hoya tenggerensis Bakhuizen 1950 T ObAc ØOObl x Hoya teretifolia Griffith ex Hooker 1883 P CuA ØV

21 Hoya teysmanniana Miquel 1856 P ObOb ØLObl x Hoya thomsonii Hooker 1883 P ObAc POObl x Hoya tiapeansis Hochreutiner 1938 P CuAc ØE x Hoya tjadasmalangensis Bakhuizen 1950 P Ob Ac ØO 19.5 x 12 Hoya tjampeaensis Hochreutiner 1936 P CuAp ØE x Hoya tomatoensis Green & Kloppenburg P ObAp ØOE 10.7 x 3.8 Hoya torricellensis Schlechter 1913 P CuAc ØE 9-14 x Hoya treubiana Schlechter 1908 P ObOb ØOblO x Hoya trigonolobos Schlechter 1905 P Ob A ØOObl 14 x 7 Hoya trinervia (Hort.) Mach.ex Regal 3-5 ObA ØO x Hoya trinervis Trail 1830 PL ObAc ØOObl Hoya trukensis Hosakawa 1937 P Ob Ac ØEO x Hoya tsangii Burton ex Klopp PO CuAc Ø V 5-12 x Hoya tsiangiana P. T. Li 1984 Ø Hoya tsoi Merrill 1934 P Ob Ac ØOE 6-8 x Hoya tutuilensis Klopp. & Siar ObA Ø OL x Hoya uncinata Teijsm. & Binnen P ObAc ØL 9 x 3.5 Hoya upoluensis Reinecke CuAc ØEO 7-15 x 5 Hoya vacciniiflora Schwartz 1931 P CuA ØOblL Hoya vaccinioides Hooker 1883 P Ob A ØE 1.25 Hoya variegata Siebold 1846 P CuA ObA ØO + EL Hoya variegata De Vriese 1853 ObAc ØOEObl 2-5 Hoya variifolia Ridley 1926 P ObA CuAc ØEOb + L 1.5 x x 0.2 Hoya velutina Griffith 1854 P ObA ØOL 3.8 Hoya venusta Schlechter 1913 P ObAc ØL -E 3-5 x Hoya verticillata (Vahl) G. Don 1837 P ObAp ØObl 7.5 x 3.1 Hoya villosa Constantin 1912 P ObOb POblE 7-11 x Hoya vitelina Blume 1849 P CuAc ØOblL 8-22 x Hoya vitellinoides Brink f P ObAc ØObl x Hoya vitensis Turrill 1915 P Ob At ØEObl 10 x 7 Hoya volubilis Griffith 1835 T ObA ØOL Hoya warburgii Schlechter Unpub Ø Hoya wariana Schlechter 1913 P CuAp ØV x Hoya walliniana Klopp. & Nyhuus 2003 P Ob A Ø O Hoya wayetii Kloppenburg 1993 P CuA Ø V x Hoya waymaniae Kloppenburg 1995 P P 5-9 x 3-5 Hoya weebella Klopp. & Siar 2005 P ObOb Ø Ob 1.8 x 0.9 Hoya whistlerii Kloppenburg 2002 P ObAc Ø L x 3.8 Hoya wibergiae Kloppenburg 2001 T ObAp Ø E x

22 Hoya wightiana Thwates 1864 P CuAc ØL x Hoya wightii Hooker f P CuAc ØE 5-10 Hoya wrayi King & Gamble 1908 P CuAp ØLO x Hoya yuennanensis Kand_Mass 1936 P Ob Ap Ø

23 Leaf Data from Hoya Type descriptions Hoya acicularis Green & Kloppenburg semi-terete leaves (not linear-lanceolate.. bifoliate at the node, that may be straight to curved and twisted; petiole curved. Hoya acuminata Bentham ex Hooker leaves opposite, oblong, acuminate, fleshy. Hoya acuta Haworth leaves ovate lanceolate acuminate. Hoya aeschynanthoides Schlechter. densely leaved; with leaves shortly petioled, broad ovate-elliptic, acute, pilose (felt like) on both sides, cm. long, nearly cm. wide, petiole hispid cm. long. Hoya africana Decaisne leaves ovate or ovate-oblong acuminate, bases rounded or somewhat attenuate, glabrous on both sides, midrib above at the limb base glandular. Hoya alagensis (Shltr.) Kloppenburg 1990 leaves outspread ovate-lanceolate, with the base rounded, apex acute and beaked, 11 cm long by cm. wide, both surfaces glabrous, not fleshy, veins on both sides 4-6 ascending, reticulate, costa (midrib) thick prominently raised below. Hoya alata K. Hill Leaves fleshy to coriaceous, ovate or rhomboid, acute, tapering to a narrowly chordate base, pale green, pink or pale bronze when growing in strong light. 3-7 cm long, cm wide; margins and apex recurved; venation obscure, sub-parallel or somewhat palmate, petioles 3-8 mm long, 2-4 mm diam. Hoya alba Kosteletsky None Hoya albiflora Zippelius ex Blume leaves elliptic oblong cuspidate.. Leaf inches (long), 1 and 2/3-2.5 inches wide, elliptic-oblong or oblong, with a constricted pointed apex, base somewhat rounded, texture very fleshy, with the lateral veins for the most part not visible, the a strong rounded prominent middle nerve on the back, above channeled and at the base warty. Hoya aldrichii Hemsley leaves with 5 nerves. 21

24 Hoya alexicaca Moon ex Trimen glabrous; l in., ovate or oblong-ovate, usually cordate at base, with rounded lobes, very acute at apex, glabrous, thin, petiole long, slender. Hoya amblyolepis Schlechter, Unpublished, Type #4545 (B) Hoya amboinensis Warburg leaves oblong and broad with a rounded base, acute apex 6-7 cm. long cm. wide, above clear, at the base of the costa somewhat granulose, veins about 4 outspread, curving and curving and running together prior to reaching the margines, not at all distinct, reticulations barely thickened. Hoya amoena Bakhuizen leaves oblong-lanceolate, sometimes almost rhomboid or ovate with the base acute or wedge-shaped, rarely obtuse, acute or briefly acute acuminate, when dried thin but strongly leathery somewhat uninerved or distinctly trinerved at the base, lateral nerves and transverse nerves above fixed at the base which are barely different, glabrous 8-11 cm. long, cm. wide, petiole thin, glabrous 8-15 mm. long. Hoya andalensis Kloppenburg Leaves opposite small, cupped below, ovate 5.0 x 2.2 cm. mostly uniform, medium flat green color above, below much lighter, most leaf pairs turned to one side of the stem. Hoya anncajanoae Kloppenburg & Siar Leaves: lanceolate- elliptic 5-9 cm. x cm. thick and fleshy, base subcuneate apex sub-acute. new growth with more pubescent than older foliage, juvenile lower surface with fine pubescence to finely papilate, edges rounded, upper surface with fewer but more uniform distribution. Mature foliage some hair cells below, upper with fine pustulate. Nerves obsolete but pinnate, 10 or so nerves on each side of midrib anastomosing. Hoya angustifolia Traill Leaves narrowly-lanceolate, Narrow leaved Hoya. Leaves membranaceous. Hoya angustisepala Schlechter 1830 Hoya angustisepala Burton leaves very leathery, glabrous, oblong nearly oblong-ovate, as long as 15 cm., 6 cm. wide, base broadly rotund or obscurely emarginate, apex acute nearly subacuminate, nerves on both sides 3-5, nerves thin; petiole thick nearly 2 cm. long, glabrous deeply grooved. Hoya anulata Schlechter loosely leaved; leaves flat outspread elliptic or oblong elliptic, ovate, sometimes obtuse acuminate, both sides glabrous, nerves barely conspicuous, of thick leathery texture, 6-8 cm. long, near the middle cm. wide. 22

25 Hoya apiculata Scheffler Leaves glabrous at the base almost cordate, somewhat orbicular, tips apiculate, one nerved, with gland inserted at the base. Hoya apoensis Kloppenburg & Siar Leaves glabrous, oblanceolate, base cuneate, apex sub-acute, veins pinnate 7 +/- pairs, at 45 angle to the midrib. Blade cm. long, cm. at the widest. Petiole terete, glabrous, not grooved, cm. long 0.05 cm. in diameter. Hoya archboldiana C. Norman Leaves thick, large, pinnately veined, up to around 13 x 7 cm., oblong, tips abruptly acute, when dry the base lightly cordate with the sinus closed. Hoya ariadna Decaisne 1844 Leaves elliptic, base and apex rounded or above nearly acuminate somewhat distinctive 4.4 inches ling 2 wide, fleshy almost veinless, margins reflexed below concave boat-shaped both surfaces shinny glabrous. Hoya arnottiana Wight leaves fleshy, broadly ovate acuminate leaves fleshy, broadly ovate acuminate. Hoya attenuata Christophersen blades 5 x x x 2.8 cm. long and broad; petiole grooved, glabrous, 8-11 mm. long. Hoya aurantiaca Kloppenburg & Siar Foliage oblanceolate, apex shortly acute, base narrowing subcordate, surface dull green with occasional dull purplish staining. Nerves quintuplinerved of a lighter color distinct above to very distinct even raised, invisible below, first pair arise slightly above base, ending in looping secondary nerves below middle, second pair above base, looping with side reticulations most not reaching the apex cm. long and cm. wide Petioles extremely variable, most thick, curved as large as stem or larger, some extremely long. Hoya australis R. Br. ex Trail Leaves on short petioles, ovate obovate or nearly orbicular, obtuse or shortly acuminate, rounded or rarely almost cordate at the base, thick and fleshy, 2 to 3 in. long or rarely more. Hoya australis subsp. orimicola: subsp. rupicola; subsp sanae; subsp. tenuipes. Hoya australis subsp. nathaliae Kloppenburg & Siar Leaves ovate, cordate, short apiculate apex, 6.5 cm. long 3.6 cm. at the widest, dorsal surface stiff short pubescence, ventral side more dense indumentum. Peduncle 23

26 densely pubescent, 4 triangular glands at the attachment point above, terete, curved 1 cm. long 0.10 cm. in diameter. Hoya balansae Constantine Leaves fleshy glabrous, grayish-green, at times with spots of dark green, oval rounded at the base, tip acuminate, nerves not very apparent 3; lateral nerves alternating, 2 to 4 on each side, meeting at the edge with the others; length 4.5 to 6.5 cm. by 2.5 to 3.5 cm.; petiole thick, to 5 mm. long. Hoya bandaensis Schlechter loosely leaved; leaves petiolate elliptic acute or briefly acuminate, both sides glabrous, with fleshy texture, cm. long. in the middle 5-7 cm. wide, petiole fleshy about 1.5 cm. long. Hoya barbata Sprengel leaves ovate acute very glabrous. Hoya barracki Horne ex Backer. None Hoya bella Hooker with copious foliage, leaves small ovate acute shortly petioled one nerved above intense green below pale, lateral. Hoya benguetensis Schlechter loosely leaved, with the leaves elliptical or ovate elliptical, acuminate, glabrous, with thick coriaceous texture, with 5 primary nerves very conspicuous, 6-10 cm. long in the middle or below cm. wide. Hoya benvergarai Kloppenburg & Siar Blades 9-13 cm. long x 2-3 cm. at the widest. Petiole round cm. long. Hoya betchei (Schlechter) Whistler loosely leaved, leaves outspread petiolate lanceolate-elliptic, with the base rounded, glabrous, texture leathery, cm. long, in the middle cm., petiole fleshy about 1 cm. long; cymes slenderly pedunculate shaped like umbels, of few flowers, peduncle round, glabrous, 5-6 cm. long. Hoya bhutanica Grierson & Long Leaves elliptic and not fleshy, corolla at least twice wider inside. Creeping shrub stem narrow 3-5 meters, rooting. Leaves elliptic (6.5) 9-15 x (2.75) cm., apex acute base rounded, trinerved, lateral veins produced ½ above, slightly leathery, glabrous, margined entire. Hoya bicarinata A. Gray

27 leaves glabrous somewhat fleshy flat obscurely penninerved oval obovate or ovate very briefly and abruptly acuminate bases rounded almost cordate, blade above petiole hairy glandular. Hoya bicknellii Kloppenburg Leaf: Opposite and whorled. Petiole terete, curved, 1.5 cm. long, nearly the same diameter of the stem from which it arises, glabrous, green. Blade broadly ovate to ovate elliptic, flat to sub curved with some undulations cm. long x cm. wide in the center. Base cuneate, edges cupped upward slightly, pinnate anastomosing veining, visible above as is the midrib but obscure below. Apex shortly apiculate to slightly acute. Glossy green above duller below, glabrous. Hoya bicolensis Kloppenburg & Siar Foliage ovate to lanceolate apiculate cm. long and cm. widest, triplinerved, prominent above obscure or less below. A basal pair plus midrib and a pair above, looping and anastomosing. Dull to dark green above paler below. Petiole round curved to twisted cm. long, larger in diameter than the stem cm. in diameter. Hoya bicolor Kloppenburg Foliage opposite, blades ovate with a cordate base and extremely short acute apiculate apex, slightly furrowed at the midrib on the dorsal surface, otherwise horizontal, edges entire, pinnately at about a 40 degree angle, ca. 4 pairs on either side, looping in before reaching the margins and somewhat anastomosing. Lower surface along the midrib with scattered hair cells. Petiole long twister ca. 3 cm. long 0.4 cm. in diameter, slightly grooved near the blade otherwise terete; two small semicircular raised dark glands at attachment above. Internodes variable in length 5 cm cm., puberulous, with nodes enlarged, leaves leaving circular scars. Hoya billardieri Decaisne leaves oval by no means or rarely bases emarginate or somewhat cordate apexes very shortly acuminate with both sides glabrous, limb above petiole glandular, with three 3 obtuse glands. Hoya bilobata Schlechter well leaved; leaves briefly petioled broadly elliptical or somewhat orbicular, obtuse, glabrous, leathery textured, cm. long, cm. long in the middle, petiole about 0.3 cm. long. Hoya blashernaezii Kloppenburg Leaf: petiole terete curved cm. long cm. At the widest, blade elongate elliptic- lanceolate with a slight curve inward apex tapering to a very acute apex, base cuneate or occasionally slightly rounded; venation prominently triplinerved; deep green and glossy above, dull and lighter green below, midrib obscure. Hoya blumeana Schlechter None 25

28 Hoya bonii Costantin Leaves fleshy, opposite, round or heart shaped at the base end with the apex pointed, occasionally oval or even oblong and pointed at apex. Some hairs below, short and with intervals between the hairs visible, often seen with magnifying glass only; hairless and smooth above. Midrib very projecting below and hairy along its sides; secondary nerves 5 to 6 pairs, slanting at 45º for some distance towards the edge. Leaf 8 to 10 cm. long. by 3.8 to 7 cm. wide; petiole to 1 cm., 4mm. thick. Hoya bordenii Schlechter loosely leaved; with leaves outspread lanceolate-oblong or narrowly elliptical, acuminate, glabrous, texture leathery, cm. long, cm. wide at the middle or below the middle, with the petiole short, fleshy cm. long. Hoya brevialata Kleijn & Donkelaar Leaves suborbicular to elliptic, light-green, yellowish to bright red when very exposed. Petioles cm long. Hoya brittonii Kloppenburg, The leaves are noted to be thick and on drying are pinnately veined, 3-4 looping nerves on either side of a somewhat obscure midrib. They appear also to be reticulately nerved Leaf Blade: variable but generally obovate 8 cm. long, 3 cm. wide above the middle, thick, glabrous on both surfaces, with a cuneate base leading into the petiole; pinnately nerved, probably obsolete in the living state. Hoya brooksii Ridley leaves fleshy, elliptic, somewhat acute, bases cuneate, nerves invisible, midrib even obscure, 5 cm. long, 2 cm. wide, petioles 5 mm. long; peduncles somewhat robust 3 cm. long. Hoya browniana Koorders leaves opposite (but not always, rarely one aborts becoming pseudo-alternate), oblong, glabrous, fleshy, dried led-gray, with the base acute, with the apex acuteacuminate, distinctly triple-nerved, cm. long and 6-7½ cm. wide. Petioles 2-2½ cm. long, thickened, glabrous. Hoya brunioniana Wight leaves fleshy, oval-oblong acuminate. Hoya bulusanensis Elmer See under Hoya panchoi Kloppenburg. Leaves very thick, pale or yellowish green beneath, the upper surface paler green, normally opposite along the slender branches, more or less clustered toward the base or upon very short and thick seemingly abnormal branchlets, subdeciduous, glabrous but minutely punctate on both faces, the dry blades wrinkled on the upper but perfectly smooth on the lower side, flat, edges straight and entire, curing yellowish brown, of two distinct types, the orbicular to rotund shape and the elliptic to the obovately oblong type, the latter ones 2 cm wide above the middle and 4 cm in length, obtusely ro6nded at the 26

29 apex, broadly cuneate at the base, the smaller kind 1.5 cm wide across the middle; petiole very short and very thick, curved and expanded toward and at the base, dull brown, rugose on my specimen, leaving large circular scars after falling; midrib scarcely evident or entirely obsolete in the smaller type of leaves, in the middle; petiole very short and very thick, curved and expanded toward and at the base, dull brown, rugose on my specimen, leaving large circular scars after falling; midrib scarcely evident or entirely obsolete in the smaller type of leaves, in the middle region longitudinally striate below especially so of the larger blades, lateral nerves none. Hoya buotii Kloppenburg The typical leaf is elliptic-lanceolate with a long acuminate apex, base narrowly rounded. Blade is 10 cm. long x 2.5 cm. wide near the base. The petiole is grooved above, 1 cm. long x 0.09 cm. in diameter. Nerves are pinnate 7 or so on each side, at from a 30º to nearly perpendicular angle to the midrib; anastomosing. Hoya burmanica Rolfe Leaves somewhat pendulous, shortly petiolate, narrowly ovate-lanceolate, acuminate, somewhat concave, slightly thick, glabrous, glaucous-green 5 to 8 cm. long, 1.3 to 2 cm. wide; petiole 5 mm. long. Hoya burtoniae Kloppenburg The leaves of this species are more regularly ovate-elliptic, rather than long and narrowly lanceolate. Hoya burtoniae contains a lot of anthocyanin pigmentation, which is visible throughout the plant, especially manifest on the lower leaf surfaces Leaves beautifully colored, opposite, far apart, thick or leathery, with revolute margins, narrowly oblong or oblanceolate, rigid, above covered with closely spaced velvety granules, beneath less pubescent and granular, 2.5 to 12 cm. long and above middle 3 cm. broad, bases attenuate, at apexes obtuse or somewhat acute; midrib not prominent, side nerves not at all prominent; petiole velvety 0.8 to 2.5 cm long. Hoya buruensis Miquel Leaves shortly petioled with rounded bases, slightly emarginate, above the petiole minutely callused, (a little) ovate-elliptic or (more or less) elliptic-oblong with the tip acute or obtuse-attenuate, fleshy, glabrous, below the midrib covered over, on both sides about 5 (from the very base) erect and outspread curved almost united. Hoya cagayanensis Burton Leaves elliptic, lightly leathery, penninerved, both sides glabrous, 6-8 cm. by cm., petiole 1 com (cm.) plus or minus long. Hoya callistophylla Green. leaves opposite, elliptic to lanceolate with attenuate tips, cuneate bases, prominent, pinnate, dark green venation, substance heavy and hard, with a matte surface. Hoya calycina Schlechter

30 Leaves spread or outspread, petiolate, broadly oval, apiculate, with base somewhat cordate, leathery, above sparsely puberulous, below shortly tomentose, petiole tomentose above slightly grooved. Hoya campanulata Blume leaves leathery almost veinless oblong, acuminate glabrous. Hoya camphorifolia Warburg Leaf blade glabrous on both sides, parchment like, ovate elliptic or elliptic, 6 to 9 cm. long, in the middle 2.5 to 3.5 cm. wide, apex acuminate acute often curled back, often convex on underside, bases narrowly rounded. Veins 2-3 on both surfaces ascending, not all reaching the apex, reticulate, distinct but not prominent. Peduncles long and filamentous, straight, glabrous, 8-9 cm. long, 0.1 cm. diam. Hoya cardiophylla Merrill Leaves heart-shaped, apparently fleshy when fresh, rather flaccid when dry. 5 to 7 cm. long. 4.5 to 5 cm wide, pale when dry, glabrous or the younger ones slightly ciliate near the basal margins, the base broadly rounded and distinctly cordate, the sinus acute, the basal lobes broadly rounded, the apex shortly and sharply acuminate; petioles 5 to 7 mm long; lateral nerves about 4 on each side of the midrib, somewhat spreading, anastomosing, the reticulations lax and distinct on both surfaces. Hoya carnosa R. Brown Leaves oval-oblong fleshy, corolla hair. Hoya carrii Forster & Liddle 1992 Hoya caudata J. D. Hooker Leaves 2-3 in. diam., sparsely papillose on both surfaces, midrib and arching nerves faint: petiole ¼ in., so thick as to appear globose. Hoya cavaleriei Leveille Leaves leathery, nerves conspicuous, middle nerve elevated. Hoya cebuensis Kloppenburg & Siar leaves are ovate lanceolate with an obtuse base versus linear-elliptic with a cuneate leaf base. Hoya cembra Kloppenburg leaves erect- spreading briefly petiolate, narrow elliptical beaked, glabrous, shinny, somewhat leathery, 8-10 cm. long, below the middle 2-3 cm. wide, petioles 0.5 cm. long. Hoya chinensis Traill None 28

31 Hoya chinghungensis (Tsiang & P. T. Li) M. G. Gilbert, P. T. Li & W. D. Stevens. leaf blade broadly ovate, x cm, glabrous, base rounded to truncate, apex acute to obtuse; lateral veins obsolete. Hoya chlorantha Rechinger Leaves leathery, when dry shiny green, nerves not prominent on either side, briefly petiolate, narrowly acuminate at both ends, penninerved. Hoya chlorantha var. tutuilensis Christophersen Leaves leathery, when dry shiny green, nerves not prominent on either side, briefly petiolate, narrowly acuminate at both ends, penninerved. Hoya chloroleuca Schlechter Leaves spreading or outspread, petiolate, lanceolate or elliptic- lanceolate, acute or sub acute, thinly coriaceous, both sides glabrous, petioles short, glabrous, above slightly grooved. Hoya chunii P. T. Li None Hoya cinnamomifolia Hooker leaves fleshy-leathery, thick ovate or ovate oblong acuminate, 5 nerved, nerves above prominent, above at the petiole callused glandular; peduncle short. Hoya citrina Ridley Leaves thick, fleshy stiffly coriaceous ovate, base broad, round, subcordate; nerves three conspicuous when dry with about 4 pairs of lateral nerves from the central nerve broken up into reticulations large and lax. 4 to 4.5 in. long, 3 in wide; petiole very thick.75 in long. Hoya clandestina Blume not from type description (none) leaves fleshy elliptic-oblong acuminate with the base rounded, three nerved between the nerves obscure veined; Umbels with many flowered long peduncles the inside of the corolla silky lobes ovate acute revolute. Hoya clemensiorum Green. leaves opposite, thick and rigid, petiole 7 mm x 2 cm, curved, blade cm wide x cm long, with raised, prominent, pinnate venation, ovate-lanceolate with attenuate tip. Hoya cochinchinensis Loureiro leaves opposite, ovate. Hoya collettii Schlechter

32 Leaves petiolate fleshy, linear-oblong or linear-lanceolate 3-4½ inches long, somewhat obtuse, base elongated, rounded, above shining, above paler midrib thick elevated; petioles thick 3-4 lines long. Hoya collina Schlechter Leaves erect spreading or widely outspread, shortly petioled, elliptic or ovateelliptic, obtuse, wedge shaped base, fleshy, both sides glabrous, petiole fleshy, above lightly furrowed (grooved). Hoya cominsii Hemsley Leaves fleshy, glabrous, wide ovate, with petiole thick, at least 5 inches long, obtuse or somewhat acute, palmately 5 to 7 nerved. Hoya commutate Gilbert & Li Leaves not known. Hoya compacta Burton 1990 Caulis puberulis, brunnei, albis pilis, nodis brevis 1 1/2-2 cm.. Folia 5 1/2-6 cm. longa X 3 cm. lata, obsolete plicata, inferiora exposita, superiora persipe occulta complanata, (basi cordatis, apices acuminatis), ocula nuda vel negligentius glaberrima sed reversis pilis minutis dispersis, ocula armata aspectabilibus utrinque conspersis. Petioli ± 1 cm. longi, supra non canaliculate. Hoya coriacea Blume leaves a little nerved oval acute or acuminate coriaceous, glabrous. Hoya costantinii P. T. Li Hoya reticulata Cost. in Lec., None Hoya cordata Li & Huang. caule terete cortice cineraceo-flavo 5 mm diametro. Folia opposita succulenta vel exsiccations crasse coriacea, ovate vel long ovate 5 9 cm longa cm lata, apice acute vel obtuse basi cordata, supra glabra et subtus ad costam mediam puberula, nervis lateralibus utrinsecus 6 paribus utrinque leviter elevatis, petiolis robustis 5 mm longis pubescentibus, apice glandulis perpaucis 3 5-fascicularibus instructis. Hoya crassicaulis Elmer ex Kloppenburg leaves leathery elliptic or oblong-acute, shiny, 8-17 cm. more or less large, in the center nearly 5-8 cm. wide, base rounded or a little somewhat acuminate and recurved, margins somewhat revolute, midrib prominent below, veins on both sides about 4, spreading curving confluent to the margins. Hoya crassifolia Haworth ex Traill The leaves are obovate, very bluntly acuminate, upwards of four inches long, without veins, of a darkish green colour above, pale beneath. The petioles are thick and fleshy, of a purplish green colour, and much flattened above. 30

33 Hoya crassifolia Ridleyi 1912 Leaves very coriaceous ovate acuminate slightly cordate at the base, margins undulate and minutely irregular dentate, nerves very inconspicuous 3-4 pairs at an acute angle, midrib inconspicuous 6 inches long 2½ inches wide, petiole very short and thick. Hoya crassior Hochreutiner Leaves opposite, midsize, petioles short, glabrous, more or less fleshy, channeled above and where united with the blade somewhat glandular; blade very glabrous, very thick and fleshy, the midrib it seems depressed on but sides, otherwise lanceolate, apex a little attenuate-acuminate, base acute, margines entire, secondary nerves inconspicuous, or in juvenile leaflet blade about 3, both sides arching and meeting and with reticulate nerves. Hoya crassipes Turczaninow leaves petiolate elliptic or obovate, the base attenuate, briefly acuminate, fleshy veinless glabrous. Hoya cumingiana Decaisne leaves ovate-cordate obtuse somewhat fleshy above glabrous below conspicuously veined papilose velvety with short petioles. Hoya curtisii King & Gamble Leaves sessile, very thick; orbicular, abruptly triangular-acuminate at apex, rounded or sometimes orbicular very slightly attenuate at the base; both surfaces scarious-puncticulate when dry; margine thickened;.4 to.7 in. long,.4 to.6 in broad; midrib when dry faint, other nerves invisible; petiole none. Hoya cystiantha Schlechter Leaves at bases generally acute, obverse oblong or lanceolate acuminate, leathery veined, 3-4 ½ inches long. Hoya dalrympleana F. Muell = H. australis R. Br. ex Trail. None Hoya darwinii Loher Normal leaves elliptic oblong, cm. long, 5-6 cm. wide, glabrous, leathery, with inconspicuous nerves, briefly petiolate; other leaves bullate, a globose pouch of 4-6 cavities, well closed. Hoya dasyantha Tsiang Leaves chartaceous, broad ovate, cm. Long, cm. wide, shortly acuminate, rounded at the base; nerves slightly elevated on both surfaces, lateral nerves 4-5 pairs, veinlets transverse, conspicuous; glands 4 to 6 in a group above the petiole; petiole 5 mm. long. 31

34 Hoya davidcummingii Kloppenburg Leaf Blade: spreading lanceolate-elliptic, upper surface deep green even olivaceous, glabrous with fine granulation s, edges rounded and slightly thickened, lighter green below: apex mostly acute turning down, base cuneate, overall thickened and coriaceous, veins not visible but pinnate, cm. long, widest about the middle cm. Hoya dennisii P. I. Forster et D. J. Liddle Leaves petiolate; lamina lanceolate-ovate, coriaceous, up to 72 mm long and 30 mm wide, discolorous, glabrous, venation obscure; upper surface dark green lower surface pale green; tip acuminate; base rounded to cordate petiole 5-7 mm long and c. 0.8 mm diameter, with scattered trichomes colleters 2, at lamina base. Hoya densifolia Turczaninow leaves somewhat sessile membranaceous densely imbricate elliptic, with short cordate sinus, slightly acute penninerved reticulate glabrous. Hoya deykeae Green Leaf blade: 5-12cm long x 5-8cm wide x 2-2.5mm thick, matte surface. Petiole 3-6 mm diameter x mm long, rough surfaced, tan. Hoya dickasoniana P. T. Li leaves opposite even 3 or 4, elliptic or ovate, 9 19 mm. x 5 10 mm, base broad cuneate or rounded, apex obtuse, glabrous; petiole 1 mm. long. Hoya dictyoneura K. Schumann leaves briefly petiolate oblong or elliptic or somewhat orbicular acute with the base rounded five nerved both sides glabrous. Hoya dimorpha F. M. Bailey Leaves thick-coriaceous, oblong-acuminate, slightly cordate at the base; those on the thick portion of the stem exceeding 5 in. in length and 2 in. in breadth; those on the slender ends often cordate and under 2 in. long, glabrous, undersides pale, the gland on the face indicating the attachment of the petiole, beneath often a broad ciliate disk at times mote or less umbonate. Petiole stout, usually less then ½ in. long. Hoya diptera Seemann leaves ovate-elliptic, acuminate with the base ovate, fleshy, penninerved. Hoya dischorensis Schlechter Leaves spreading or outspread, shortly petioled, oblong, somewhat obtuse, acuminate, base obtuse, a little fleshy, both sides glabrous, petiole short, slightly grooved above. Hoya diversifolia Blume Leaves veinless oval acute orbicular fleshy glabrous. 32

35 Hoya dodecatheiflora Fosberg leaves elliptic-lanceolate, thick, unequal, up to 8 cm. long, umbels axillary, many flowered, pedicles slender, 1 cm. long. Hoya dolichosparte Schlechter loosely leaved, glabrous with numerous wart like lenticels; leaves outspread, petiolate, broadly oval, obtuse, with 5 prominent primary nerves above, both sides glabrous cm. long, almost cm. wide at the middle, petiole fleshy cm. long. Hoya edanoi C. M. Burton Leaves thick leathery, round or oblong, cordate or nearly (no such Latin word) peltate? (shield shaped), with the tip round or mucronate 3-7 cm. long x 3-5 cm. wide. Hoya edenii King ex Hooker Leaves 3-4 by 1-1 ¾ in., elliptic or obovate-lanceolate, obtusely caudateacuminate, base acute, thinly coriaceous, puberulous or glabrate beneath; petiole 1/12 in. pubescent. Hoya eitapensis Schlechter Leaves erect, spreading, elliptic, somewhat obtuse, with wedge shaped base, fleshy, glabrous with very short petioles, almost sessile. Hoya elegans Kosteletsky None Hoya elliptica Hooker f leaves 1 ½ - 2 ½ in. elliptic obtuse et both ends thin in texture nerves very prominent, peduncle very short. Hoya elmeri Merrill leaves opposite, fleshy, when dry leathery, penninerved, somewhat elliptic, 11 to 15 cm long, 5 to 6.5 cm wide, when dry pale, both sides the same color, sparsely covered with small wort-like growths, both ends equally narrow, base acute, apex acute to briefly and obscurely acuminate, primary nerves on both sides of the costa 7, slender, distinct or sometimes somewhat indistinct, curved anastomosing, loosely reticulated; petiole thick, about 1 cm long; fleshy, glabrous 1.3 mm. long. Hoya elmeri Schlechter # 8896 (B). Not published Hoya endauensis Kiew Foliis glabris ovatis-cordatis apicibus acutis 2 ad 2.5 cm longis et 2 ad 2.5 cm latis a congeneribus. 33

36 Hoya engleriana Hosseus leaves short or shortly petiolate, shortly lanceolate or narrowly ovate-lanceolate, margins curled inward leaves of the species accordingly at first appearing chiefly turned in the direction of the inflorescence, fleshy, white pubescent, nerves above inconspicuously marked, 1.5 cm long, 0.4 cm wide, narrow mobile. Hoya epedunculata Schlechter well leaved, round, glabrous. Leaves spreading or outspread, petiolate, elliptic, acuminate, somewhat leathery, both sides glabrous, with the peduncle glabrous, moderately thick slightly grooved above. Hoya erythrina Rintz Leaves coriaceous-fleshy, oblong with strongly corrugate margins and shallowly cordate basis; up to 15 cm long by 5 cm wide; 5-7 veined with a pair of veins parallel to the midrib glossy green or dull red above, red below. Hoya erythrostemma Kerr Leaves elliptic, base acute, apex mildly acuminate, up to 10 cm. long, 4 cm. wide, when dry papery, glabrous, midrib with the lateral and transverse nerves on both sides especially above, lateral nerves 5-6 pairs at 60 from the midrib joining the marginal nerves 3-6 mm. from the margins; petioles moderately robust, glabrous, 3-6 mm. long, above narrowly wedge shaped. Hoya esculenta Tsiang (H. liangii) have penninerved leaves; the leaves are of the same shape but much smaller. Hoya esquirolii Leveille None Hoya excavata Teijsmann & Binnendijk leaves somewhat fleshy, oval-oblong, base rounded, apex briefly apiculate..leaves opposite 0.15 long 0.09 wide, ovate-oblong, base rounded, or obtuse, apexes rounded, briefly aciculate, somewhat fleshy, above shinny, both sides with 6 nerves, with the petiole callous-glandulous. Hoya exilis Schlechter Leaves spreading or outspread, elliptic-lanceolate, obtuse, long-acuminate, texture for the genus thin, both sides glabrous, petiole glabrous, above slightly grooved. Hoya faoensis Kloppenburg & Siar Leaves ovate-elliptic shortly apiculate, nerves obscure but pinnate, anastomosing 4-5 cm. long x 2.2 cm. at the widest in the middle, Petiole cm. long, filiform, same color as the stem. Hoya fetuana Kloppenburg

37 Leaves short elliptic attenuate cm. long, thin but possibly a little fleshy, apex subacute, base rounded cm. at the widest near the middle; midrib narrow, visible below, pinnate netted nerves all the way to the apex. Petioles fine 0.5 cm. long, grooved above. Hoya filiformis Rechinger Leaves thin visibly reticulate veined, very glabrous bright Leaf texture thin, leaf exclusive of the petiole 6 cm long, 2 cm wide, 1 cm near the tapered point turns down. Petioles 8 mm long. Leaves are delicately feather nerved, nevertheless the nerves are distinct on the upper surface and visible on the lower surface, first of all from the median of the leaf bases extend outward in pairs almost to the edge of the leaf. Noteworthy through the long twisted apex of the leaf blade and the fine thin texture. Hoya finlaysonii Wight leaves oblong-lanceolate obtuse acuminate above (dried) reticulate, below somewhat veinless (5-7 inches long 1 ½ -2 wide) petioles short. Hoya fischeriana Warburg Leaves thick leathery broadly lanceolate cm. long 1-7 cm. wide base and apex acute glabrous 5-nerved, nerves inside almost reaching the apexes, nerve reticulations distinct, petiole cm. long 2-3 mm. thick; peduncles about 5 cm. long 2 cm. thick, upper half cylindrical 3 mm. thick. Hoya fitchii Kloppenburg & Siar Foliage is quintuplinerved, with the second pair of nerves nearly reaching the apex, outer connecting nerves anastomosing Nervation white or lighter green than the leaf blade color. Hoya flagellata Kerr Leaves ovate or lanceolate, base rounded and minutely eared, with the apex acute, margine more or less revolute, cm. long, cm. wide, leathery, when dry pale brown, glabrous except that above it is sometimes provided with minute sparse hairs, midrib scarcely visible above, lateral nerves obscure on both sides; petioles thick, wrinkled, glabrous, 3-5 mm. long. Hoya flavescens Schlechter loosely leaved, round, glabrous. Leaves spreading or outspread, oblong or lanceolate-oblong acuminate, base somewhat cuneate rounded, fleshy, glabrous, petiole short, somewhat fleshy above lightly grooved. Hoya flavida P.I. Forster et D. J. Liddle Leaves glabrous, petiolate; lamina elliptic-ovate, up to 140 mm long and 45 rum wide, discolorous, secondary veins 4 per side of midrib just visible on both surfaces, tertiary venation obscure; upper surface dark green; lower surface pale green; tip acute to short acuminate; base cuneate; petiole grooved along top, mm long and c.1.5 mm diameter. 35

38 Hoya flexuosa Spreng leaves cordate oblong veined. Hoya forbesii King & Gamble Leaves coriaceous when dry; elliptic-oblong or obovate, shortly and bluntly acuminate at apex, rounded at bass; upper surface glabrous, shining; lower surface minutely papillose; both pale brown when dry; margins somewhat recurved; 4 to 7 in. long, 2.5 to 4 in. broad; midrib strong, much raised beneath, bearing on the upper surface a gland just above the petiole; main nerves about 7 pairs, irregular but usually at 40 with the midrib, more or less curved and branching; secondary nerves frequent, especially near the top between the main nerves; reticulations transverse, irregular, clearly visible only on the upper surfaces; petiole very thick, bent, articulate in the middle, about 5 in. long. Hoya formosana Yamazaki Leaves fleshy broadly ovate 7-9cm long 4-5cm wide apex abruptly attenuate base rounded when dry flat very glabrous base near the midrib sparsely rough below glabrous with nerves roughly punctate, primary nerves few 2-3 pairs conspicuously curved, petioles thick 2-3cm long above the beginning of the young shoot roughly punctate otherwise glabrous. Hoya fraterna Blume leaves leathery ovate-oblong elliptic acuminate base rounded somewhat veined above the petiole callous-glandulose. Hoya fungii Merrill leaves leathery, penninerved, elliptic to oblong, 8 to 14 cm. long, 4.5 to 8 cm. wide, when dry pale, opaque, both sides glabris uniform color or below a little paler, apexes acute and shortly acuminate, base rounded, midrib thick, below strongly elevated, lateral nerves on both sides about 7, not at all evident, arched meeting at the margins, about 30 angle with the midrib, reticulations loose, obscure; petiole 1.5 to 3 cm. long, glabrous. Hoya fusca Wallich leaves oblong, acuminate. Leaves approximately opposite, very outspread, strongly thick and leathery, firm, flat, oblong narrowly acuminate ½ inch terminal, margins parallel convex, base obtuse, above purple veined, shinny on both sides of the petiole insertion a gland withered (at this place a solitary cilia) below flat, somewhat green, midrib very much elevated, various transverse nerves, capillaries near the margins anastomosing, 6 to 7 inches. Hoya fuscomarginata N. E. Brown Leaves thick fleshy; blade cm. long, 7-8 cm. wide. ovate-lanceolate, sometimes slightly oblique, acute or gradually somewhat acuminate, obtuse or somewhat wedge-shaped, 3-5 nerves, glabrous, green, margins dark brown; petioles cm. long, 7 mm. thick. 36

39 Hoya gigantangensis Kloppenburg Leaf Blade: cm. x cm., pinnate venation, both surfaces glabrous. Petiole: heavy, curved cm. long, probably grooved above. Hoya gildingii Kloppenburg. linear lanceolate opposite leaves, lax or with the petioles upright along the stem. Hoya glabra Schlechter loosely leaved; leaves outspread ovate acuminate, base rounded, glabrous, fleshy texture, cm long in the middle cm wide, 3 primary nerves especially conspicuous, petiole thick cm long. Hoya globiflora Ridley Leaves ovate subacute with rounded bases 6.5 cm. long, 3.5 cm. wide, nerves 3 pairs and nerrules few visible when dry, one pair from the base ascending, the others short horizontal soon broken up, petiole very thick, 6 mm. long. Hoya globosa LeConte None Hoya globulifera Blume leaves almost sessile and ovate elliptic acute with the base rounded fleshy flat lightly veined above toward the base large white spots below punctate. Hoya globulosa Hooker leaves 5-7 in. long-petioled elliptic or oblong cuspidate or acuminate base rounded midrib very stout, nerves distinct, peduncle long. Hoya golamcoana Kloppenburg The leaves are 5.3 cm. long and 2.7 cm. wide at the widest portion, enervis, although in dried specimens the midrib and five pairs of pinnate nerves are visible. It is a densely leaved hanging vine with almost sessile petioles, 0.3 cm. long. Hoya gonoloboides Regel Leaves membranaceous, penninerved, reticulately veined, cordate ovate, acuminate, petiolate, both sides with long ascending hairs, hairs below on the nerves. Petioles and pedicels also with long hispid hairs. Hoya gracilipes Schlechter leaves spreading outspread, lanceolate or elliptic-lanceolate acuminate, both sides glabrous, texture leathery, shortly petiolate, cm. long, below the middle 2-3 cm. wide. petiole cm. long. Hoya gracilis Schlechter

40 loosely leaved, rooting; leaves shortly petioled spreading, lanceolate-elliptical acute or somewhat obtuse, base cuneate, glabrous, lucid, texture thick leathery, cm long, in the middle almost cm. wide. Petiole somewhat thicker cm. Hoya grandiflora Blume ex Decaisne leaves oblong or elliptic-oblong base rounded apexes abruptly and shortly acuminate pubescent above somewhat soft and scattered below densely tomentose velvety reddish (When dried?). Hoya grandiflora Sprengel Hoya leaves cordate-ovate acute. Hoya graveolens Kerr Leaves oblong-elliptic or oblong oblanceolate, base broad wedge shaped or rounded, apex briefly acuminate, margins flat, 5 to 8.3 cm. long, 2 to 4.8 cm. wide, leathery, glabrous, midrib above sunken below almost flat, lateral nerves 4 to 5 pairs often ascending with an angle of 60 difficult to observe; petioles 4 to 6 mm. long, thick, above concave, glabrous. Hoya greenii Kloppenburg the veins of the leaves of this species are perpendicular to the midrib and finely reticulate, whereas those of Hoya darwinii Loher are at 45 degrees to the midrib, venation in neither is prominent. Leaf blade: cm. long x 3-4 cm. wide, narrowly elliptic, both surfaces glabrous, dull, very dark green, above slightly channeled along the midrib area, lighter green below. 10 +/- pairs of veins perpendicular to the midrib, somewhat obscure, with fine secondary reticulate venation with the base tapered acuminate, outer apex curved under acuminate, rigid. Hoya griffithii Hooker Leaves distant, 1-2 in. diam., midrib stout, nerves quite obscure in some, in others distinct and forming large areoles; petiole ½ - 1 in.. very thick. Hoya gymnanthera Wight The specimen of this plant which I have seen are leafless. Hoya hainanensis Merrill leaves when dry papery, ovate-elliptic, penninerved, opposite, 6 to 8 cm long, 2.5 to 4 cm broad, both ends somewhat equally narrow, apex acuminate, base acute, nerves about 4 on both sides slender, obscure, almost oblique, reticulations loosely arranged, inconspicuous; petiole 8 to 10 mm long. Hoya halconensis Kloppenburg Foliage cm. x 4-6 cm. with most small, spreading, glabrous, base broadly rounded, rather rectangular tapering slightly to the mucronate apex. Midrib visible above and below, lateral nerves obscure. 38

41 Hoya halophila Schlechter Leaves spreading or outspread, petiolate, broadly ovate or somewhat round-ovate, briefly acuminate, base very obtuse, both sides glabrous, lucid above, opaque below, fleshy. Hoya hasseltii Miquel leaves ovate acute or acuminate base somewhat cordate margines somewhat wavy glabris below white-punctate. Hoya hellwigiana Warburg leaves cuneate-lanceolate or oblong 9 to 15 cm long, apex and base acute thickly leathery, veins on both sides about 2 ascending almost entirely inconspicuous. Hoya heuschkeliana Kloppenburg Leaves without nerves, petiolate, ovate-elliptic obtuse, base wedge-shaped, fleshy, on the back puberulous both sides punctate (pustulous or granulate) 2 to 5 cm. long, in the middle 0.5 to 2 cm. wide, above convex. Petioles curved about 0.2 cm. long. Hoya heuschkeliana subsp. cajanoae Kloppenburg & Siar Foliage: opposite, ovate x cm. glabrous, flat with edges slightly rolled under, lower surface finely circular granulate (pustulate). Petiole curved 0.8 cm. long, terete, glabrous with a grove above near leaf blade. Hoya hookeriana Wight leaves fleshy oblong-lanceolate 2 nerved (6 to 10 inches long 1 to 1 ½ wide). Hoya hypolasia Schlechter Leaves spreading or outspread, petiolate, lanceolate, acuminate, base somewhat cordate, leathery texture, glabrous above, lucid, shortly and densely puberulous below, petiole grooved above, very shortly and softly puberulous. Hoya imbricata Decaisne leaves aborted alternately unilaterally, imbricate orbicular above convex veinless below veined with reflexed margins. Hoya imperialis Lindley leaves leathery narrowly oblong, ciliate, glabrous, without veins, apiculate, recurved apex with the base rounded or slightly cordate, peduncles tomentose, leaves hanging and small. Hoya inconspicua Hemsley Leaves of primary flowering branches often reduced to filiform rootlike filaments; secondary leaf branchlets strongly abbreviated often erupt from the axils (?) bearing a single pair of leaves, one nearly reduced, briefly petiolate, thickly leathery or somewhat fleshy narrowly lanceolate. 39

42 Hoya incrassata Warburg leaves thick and coriaceous, ovate to oblong, 9-14 cm. long, 5-6 cm. wide, bases rounded or a little subcordate, apexes acuminate, acute, glabrous and shiny above, costa beneath prominent, veins on both sides, 4-5 spreading, curving and meeting before reaching the margins. Hoya incurvula Schlechter loosely leaved; leaves outspread petiolate, ovate elliptic, obtuse, fleshy, glabrous, margins thickened below, cm. long, above the middle cm. wide, petiole fleshy, thick 3-5 mm. long. Hoya inflata (P. I. Forst., Liddle & I. M. Liddle) fleshy leaves. Hoya intermedia Schlechter not published Leaf blades: thin many in close clusters 6-10 cm. mostly 7.5 cm. long. glabrous, narrowly elliptic, thick base narrowing into petiole, apex acute long tapering to sub rounded. Waxy rounded leaf edge. Petiole cm long, thin. Hoya intermedia A. C. Smith leaves; petioles thick (about 3 mm. In diameter) when dry very rugose 10 to 20 mm Long; blades fleshy oblong, 8 to 12 cm. long 2 to 4.5 cm. wide, base obtuse, apex somewhat acute, penninerved, midrib above lightly impressed below prominent, secondary nerves on both sides 5 to 7 ascending often sometimes immersed, both sides prominent, vein sometimes prominent on both sides. Hoya ischnopus Schlechter None. Hoya juannguoana Kloppenburg leaves, obovate to ovate-lanceolate, base cuneate, edges entire, with apiculate acute apex. Blades usually somewhat cupped, waxy medium green upper surface, lighter beneath. Pinnate venation 3 veins on either side of midrib not readily visible (in contrast to those of H. finlaysonii Wight) 4 ½ 6 long 2-2 ½ widest near the middle or beyond. Nodal length variable, stems corky, tan colored, new growth rose bronze, the whole plant glabrous. Hoya kanyakumariana Henry & Swaminathan Leaves x cm, obovate to obcordate, entire, fleshy, glabrous, rounded or emarginate at apex, tapering at base; mid and lateral nerves obscure; petioles 2-6 mm long, thick. Hoya kastbergii Kloppenburg Leaves: nearly round 6 cm. long x 4.5 cm. wide, with variation, nerves obscure, glabrous, medium green. Hoya kenejiana Schlechter

43 loosely leaved, round, glabrous. Leaves spreading or outspread, petioled, narrowly oblong, somewhat obtuse, base rounded, fleshy, both sides glabrous, with a short petiole, lightly grooved above. Hoya kentiana Burton Folia 3-12 cm. longs X cm. late, utrinque acuti; petioli ½ -1 cm longi. Hoya kerrii Craib Leaves opposite, obovate or orbicular, apex bilobed lobes rounded, base broadly wedge-shaped or somewhat rounded 4 to 12 cm. long, 5 to 9.5 cm wide, thickly fleshy, both surfaces glabrous, margins turner under; petiole 5 to 15 mm. long, 5 mm. thick. Hoya keysii Bailey Leaves ovate-cordate 1 to l 1/2 inches long, 3/4 to 1 inch wide, thick, fleshy, prominently penninerved; conspicuously anastomosing within the margin. Hoya kingdonwardii P. T. Li leaves opposite, succulent, elliptic, cm x cm, both ends acute, glabrous; petiole 1 mm. long. Hoya kloppenburgii Green opposite, lanceolate leaves, matte, dark green upper surface with occasional flecking, erose, heavy, thickened edge to blade, underside dull purple with 2-4 mm green edge, cm long and 4-5 cm wide, apex acute base cuneate. Pinnately veined, with secondary veins at degrees to midvein, nearly obscure. Petiole curved, 0.7 cm long and 0.4 cm in diameter. Hoya klossii Moore Leaves with short petioles, ovate, fairly long cordate-acuminate, apex obtuse, somewhat rounded, margines recurved, leathery, glabrous, surface skin shiny and (especially the upper surface) with prominent reticulate nerves, 4 to 6 by 1.5 to 2.5 cm. Hoya krohniana Kloppenburg Small cordate deep green opposite foliage, glabrous. With pinnate nervation, apex apiculate. Hoya kuhlii Koorders Leaves lanceolate acuminate, base narrow: Hoya kwangsiensis Tsiang et P. T. Li leaves long lanceolate up to 17 cm long and flowers much larger 3 cm in diameter. Leaves fleshy and leathery long lanceolate or lanceolate-oblong 8 to 17 cm long and 2 to 4 cm wide, margines slightly recurved, midrib on both sides nearly free of nerves and at other times not at all obscure, petioles strong grooved above 1 to 2 cm long apex with small blackish gland. Hoya lactea S. Moore

44 leaves petiolate oblong-ovate with the apex rounded, sometimes itself briefly acute and cuspidate with a rounded base very slightly cordate thick leathery both sides glabrous. Hoya lacuna Wight ex Don foliis coriacco-membrannaccis cordatis ovatisve acuminatis supra pubescentibus subtis velutinis. Hoya lacunosa Blume leaves veinless ovate acuminate glabrous laciness above beneath convex. Hoya lambii Green leaves opposite, elliptic with apiculate tips and prominent palmate venation, substance papery, with a matte surface.. Leaf blade: Elliptic, 10-15cm long x 6-10cm wide x 1mm thick, matte surface, light green. Leaf venation: Palmate Petiole: 4-5mm diameter x 10-15mm long, rough surfaced, tan and green. Hoya lamingtoniae Bailey Leaves lanceolate-acuminate, 2 to 3 in. long, scarcely exceeding 1 in. wide, of rather pale-green, the gland on the face indicating the position of the attachment of the petiole somewhat raised; lateral nerves few, very oblique, triplinerved at the base, glabrous on both sides, the underside more or less speckled with purple. Petiole 3 or 4 lines long. Hoya lanceolata Wallich ex Don leaves lanceolate acuminate glabrous. Hoya lanceolaria Moore Leaves shortly petioled, ovate-oblong, above ending in a tale or attenuate, apex obtuse, base obtuse briefly rounded, decidedly nerved, fleshy-leathery, both surfaces glabrous, 5.5 to 7.5 by 1.3 to 2.9 cm. peduncles shorter than the leaves. Hoya lancilimba Merrill leaves lanceolate to oblong-lanceolate, flashy, leathery 11 to 14 cm. long 2.5 to 3.5 cm. wide, both ends somewhat equally narrow, base acute, apex acute acuminate, in the dry state pale, wrinkled, with few primary nerves on either side, spreading, obscure arching and anastomosing; petioles thick, 2 to 3 cm. long. Hoya landgrantensis Kloppenburg & Siar 2008 the foliage is lanceolate-elliptic 8.8 cm. long, 3 cm. at widest near middle apex acuminate, base narrowly obtuse; triplinerved 2 near base and 2 further up midrib, angle with midrib 15 first pair 17 second pair. Upper side convex, surface finely granulate, dull, lower surface concave, edges sharp, surface finely punctate slightly lighter color than the dorsal. Hoya lantsangensis Tsiang et P. T. Li (This is not a hoya) 42

45 Hoya lasiantha Blume ex Korthals Leaves ovate briefly cuspidate with obscure veins. Hoya latifolia G. Don leaves broad-ovate, acuminated, obtuse or sub-cordate at the base, 3-nerved, fleshy, glabrous. Hoya latifolia subsp. kinabaluensis Burton 1991 Hoya lasigynostegia P. T. Li Leaves fleshy when dry, thick paper like, ovate lanceolate cm. long cm. wide, apex caudate (tail like) base rounded, above both sides flattened with lateral nerves 4-5 pairs curved ascending anastomosing far from the margines, petiole fleshy 2-4 mm long puberulous becoming glabrous, apex provided with a small brownish gland. Hoya laurifolia Decaisne leaves lanceolate oblong lanceolate, above petiole glandular margins somewhat reflexed undulate leathery very glabrous. Hoya laurifolia Miquel leaves lanceolate oblong lanceolate, above petiole glandular margins somewhat reflexed undulate leathery very glabrous. Hoya laurifoliopsis Hochreutiner Leaves not very fleshy, but nevertheless more or less coriaceous, opposite; petioles glabrous, lenticellate, more or less laterally compressed; leaves very glabrous, elliptic, margins entire and a very little revolute, apexes long acuminate, acute, bases long attenuate and toward ( the margins revolute acuminate. At the leaf base where the petiole meets it, probably glandular. Dried midrib and secondary nerves (about 5 on both sides) more or less conspicuous. Hoya lazaroii Kloppenburg & Siar Foliage: Glabrous, narrowly elliptic, apex acute, base cuneate to sub-rounded, enervis length 11.0 cm. and 2.8 cm. wide. Hoya leembruggeniana Koorders & Schumacher Leaves elliptic-oblong, obovate-oblong or rhomboid-oblong, fleshy, glabrous, base wedge shaped, apex acute or somewhat acuminate, base one nerved, secondary nerves very obscure, 4 to 6 ½ cm wide. Petiole thickened, ½ to 1 cm long, glabrous. Hoya leucantha S. Moore Leaves among the smallest, shortly petioled, lanceolate, acuminate, base obtuse slightly rounded margines recurved, leathery, both sides glabrous, 3.5 to 4.5 cm. long, near the base 8 to 10 mm. broad. 43

46 Hoya leucorhoda Schlechter Leaves spreading or outspread, petiolate, ovate acuminate, base rounded, texture leathery, both sides glabrous. Hoya leytensis Elmer ex Burton. Leaves coriaceous and subglaucous, more or less persistent, glabrous, opposite or in subwhorles at the ends of abortive branchlets, quite variable in size and shape, entire, narrowly oblong to rotund, most of them are from cm. long and from 6-10 mm. wide across the middle or widest portion, the rotund ones measure 10 by 12 mm., the normal or the oblong blades equally tapering toward both ends, obtuse to subacute or even rounded for the smaller rotund to elliptic blades which are in the minority, flat, grayish to brown on my dry specimen; midrib quite fine or obscure, plainer on the lower surface, lateral nerves none, only the upper side marked with faint transverse lines; petioles 3-5 mm. long or much shorter, dark brown, crinkle rugose when dry, leaving orbicular flat scares upon falling Hoya liangii Tsiang Leaves fleshy obovate or obovate-oblong 4.5 to 7.5 cm. long 3 to 4.5 cm. wide, apex rounded or obtuse, base somewhat cordate or rounded, above shiny green below dull when dry pale yellow, lateral nerves 4 in pairs, veins faint, above the petiole 2 to 5 glands, petiole thick 1.5 cm. long. Hoya lii Burton Hoya limoniaca S. Moore leaves far apart, moderately thick; leaves 7 to 8 by 3 to 3.5 cm., petiolate (petiole all 15 mm. long) ovate, attenuate towards the apex the apex itself obtuse, base rounded-obtuse, 5 nerved, moderately thick, leathery. Hoya linavergarae Kloppenburg & Siar The growth of this plant is similar to Hoya cembra Kloppenburg, Hoya odorata Schlechter and Hoya paziae Kloppenburg except here the leaves are more undulant and rigid reminding me of a holly leaf. Foliage opposite, glabrous blade 4-5 x cm. Hoya lindaueana Koorders Leaves opposite, elliptic, when dried lead gray, bases acute, apexes shortly-somewhat-minutely-acuminate, 3-plinerved (or indistinctly 5-plinerved) cm long and 5-10 cm wide, glabrous. Petiole cm long, glabrous. Hoya linearis Wallich leaves lanceolate obtuse fleshy both sides stem-like filiform densely pilose. Three times smaller than above (H. lanceolata) and more slender. Hoya lipoensis P. T. Li

47 Leaves long elliptic, 9 to 15 cm long by 3 to 5 cm wide, apex caudate, cauda 1.5 cm long curved, base wedge-shaped asymmetrical, lateral nerves parallel 10 to 12 pairs, prominent above, obscure below, anastomosing near the margins. Petiole 1 cm long. Hoya litoralis Schlechter loosely leaved, with aerial roots; leaves outspread, spreading, elliptic-lanceolate, acuminate, with the base generally narrowing to the petiole, both surfaces glabrous, texture thickly leathery, cm. long, near the middle cm. wide, petiole about 0.7 cm long. Hoya lobii Hooker leaves 2-3 in. subsessile elliptic or ovate acute or ovate acuminate very fleshy, nerves 6-8 pairs arched spreading. Leaves 1-2 in. diam., base narrowly cordate; midrib rather slender. Hoya loheri Kloppenburg Leaf blade: cm. long, subovate, enervis, glabrous 1.2 cm. at the widest. Petiole: c. 0.5 cm. long 0.15 cm. in diameter, grooved above, bare, rugose. Hoya longifolia Miquel leaves at the base somewhat wedge-shaped otherwise obtuse oblong lanceolate, acuminate, fleshy glabrous 4 to 2 inches long, peduncles somewhat germinal multiflowered. Hoya loyceandrewsiana Green loosely leafed with flat to conduplicate leaf blades, cm wide, cm long; leaf petiole 1 cm thick x 1x 8 cm long with a thick callus on the dorsal side at the blade attachment; leaf blade thick, 2 to 5 mm, ovate to widely cordate with acute tip and obtuse to cordate base; palmately veined with 3 major veins, 2 secondary palmate veins and network of tertiary veins. Hoya lucardenasiana Kloppenburg & Siar 2008 Leaves: ovate 5.5 cm. long and 3.8 cm. at the widest, glabrous, pinnately nerves +/- 7 pairs on either side of midrib, not visible normally, surface cupped, above smooth, underside finely punctate and lighter color. Petiole curved 1.3 cm. long 0.25 cm. in diameter, hirsute, deeply grooved above narrowly round point at leaf base and one semicircular gland. Hoya lucyae Kloppenburg & Siar Leaves narrowly elliptic to oblanceolate, margins entire not revolute, somewhat concave above, Enervis, base cuneate, widest toward the outer apex that is acute, fine scattered hair cells especially on new growth on blade, stems, and petioles, all surfaces. Blades 6-10 cm. long and at the widest. Glossy deep waxy green above, lighter and more dull on the under surface. Petioles: straight or curved, terete, 0.25 cm. in diameter, cm. long, with small gland at blade juncture or none, matt finish. 45

48 Hoya lutea Decaisne leaves elliptic, base and apex round or at apex acuminated, briefly terminated, fleshy, lightly veined and cupped below, glabrous as in the preceding (ariadna) but a little smaller and rounder, pale green. Hoya luzonica Schlechter leaves far apart; with the leaves outspread oblong obtuse or acute; both surfaces glabrous, leathery-fleshy about 9-13 cm. long, in the middle almost 5-6 cm. wide, petiole somewhat fleshy, glabrous cm. long. Hoya lyi Leveille opposite leaves, leathery on both sides corrugated and pubescent, broad green, margins entire, margins revolute, ovate-lanceolate, one nerved, at the apex and base obtuse shortly petioled. Hoya macgillivrayi Bailey Leaves subcoriaceous, about 4 in. long, 2 in. broad across the blunt cordate base, from which it tapers to an acuminate apex; margins recurved; nervation seen under a lens quite dense, the nerves alone prominent and only a few of these conspicuous. Petioles under 1 in. long, minutely glandular at its junction with the lamina. Hoya macgregorii Schlechter Leaves widely spread, lanceolate-elliptic, acuminate, glabrous coriaceous; 7-11 cm. long cm. wide in the middle, triplinerved. Petiole fleshy, almost terete, glabrous, cm. long. Hoya macrophylla Blume leaves trinerved ovate or ovate-oblong acuminate fleshy. Hoya macrophylla Wight leaves broad ovate acuminate base obtuse somewhat cordate 3 nerved fleshy glabrous (7 to 10 inches long 4 to 10 wide). Hoya magniflora P. T. Li = Hoya grandiflora Blume Not a hoya Hoya maingayi Hooker Leaves very thick, fleshy, coriaceous when dry; elliptic-lanceolate or oblanceolate, shortly acuminate at apex, long acuminately narrowed at base; both surfaces glabrous, dull when dry; margins recurved; 7 to 9 in. long, 2.5 to 3.5 in. broad; midrib slender but prominent when dry, with a gland or enlargement at the base above the petiole; main nerves about 5 pairs, prominent when dry, especially above, at about 60 to 70 with the midrib, nearly straight at first, then branching and forming a rough intramarginal nerve; secondary nerves few; reticulations irregular, distant; petiole thick,.25 to.5 in. long. 46

49 Hoya manipurensis Deb Leaves simple, opposite, decussate, thick, coriaceous, puberulous, exstipulate, petiolate, obovate, truncate, pale gray in colour; veins obscure 3-5, at first parallel then diverging, apex slightly depressed and mucronate; base acute attenuated; margin entire; petioles pubescent, 2-4 mm. long, cm broad at the broadest region. Hoya marginata Schlechter loosely leafed, rooting; leaves spreading, elliptic-ovate, acuminate, both sides glabrous, thick leathery, 5-7 nerved, with nerves almost fan shaped, ornate, 13-17cm long, in the middle cm broad, petiole thick, up to 4cm long. Hoya matavanuensis Kloppenburg & Siar 2009 Leaves glabrous, elliptic, attenuate apex cm. long, cm. wide; nerves obscurely pinnate, finely netted, whole plant here fine and delicate with relatively long peduncles & large flowers. Petioles glabrous, cm. long ca 0.1 cm. in diameter, grooved above. Hoya maxima (Karsten) Koorders leaf-base is very united fused folded together or ± fragile and sometimes, profoundly (heart-shaped). Hoya maxima Teijsmann & Binnendijk leaves thick fleshy, elliptic-oblong, shortly acuminate, bases rounded, both sides glabrous, margins reflexed, petiole thick. Hoya megalantha Turrill Leaves elliptic-ovate, apexes attenuate acuminate, bases rounded, up to 6.5 cm. long and 3 cm. wide, entirely, glabrous, somewhat leathery, not very fleshy, midrib impressed above, prominent below, about 5 lateral nerves on both sides, somewhat prominent on both surfaces, as are the transverse reticulations; petiole about 1 cm. long, round glabrous. Hoya megalaster Warburg None Hoya mekongensis M. G. Gilbert & P. T. Li. leaf blade oblong-oblanceolate, x 4-5 cm, minutely hairy, base rounded, apex acuminate to caudate-acuminate; lateral veins ca. 7 pairs, obscure. Hoya meliflua Blanco ex Merrill Opposite leaved, oblong, round, somewhat fleshy, hairless without veins with a gland at the base. Hoya meliflua subsp. fraterna Green Leaves may attain 40 cm in length and 15 cm, in width. 47

50 Hoya memoria Kloppenburg sp. nova leaves shortly petioled, elliptic-lanceolate, base cuneate glabrous. Hoya membranifolia Costantin Leaves lanceolees, attenuated in corner or a little rounded toward the basis, acuminate, very sharp to the top, very thin, translucent on the edges, glabrous; prominent median rib underneath; nerves secondary 5 pairs, inserted perpendicularly from the median rib but immediately arching themselves toward the top and coming 1 mm. from the sides of the leaf; limb 7-12 cm. long on mm. glabrous lateral, stalked Hoya mengtzeensis Tsiang & Li Foliage thinner than general, lanceolate, apexes acuminate, base attenuate and running down into the petiole 6-11 cm long, cm broad, midrib impressed above, nerves and veinlets when dried rugose, petioles 1-2 cm long provided with 2-3 small bunched glands at apex. Hoya meredithii Green loosely, appositely leaved (sometimes lacking the twin of a pair); leaf open, with undulate margins, ovate with acuminate tip, obtuse base, subauriculate, with one large gland, hard and rigid blade; 7.5 to 18 cm. wide by 12 to 30 cm. long; petiole hard and thick, 8 mm. in diameter by cm long, twisted; leaf venation pinnate, with 8-12 secondary veins ending in reticulations, showy, all veins dark green against a light blade. Hoya merrillii Schlechter leaves far apart, fleshy; leaves outspread, ovate acuminate, fleshy, both sides glabrous, with 5 prominent nerves when dry, 6-9 cm. long, below the middle 5-6 cm. wide; petiole short fleshy, cm. long. Hoya micrantha Hooker f leaves ovate or elliptic lanceolate acute thick and fleshy, margins flat. Hoya microphylla Schlechter loosely leafed, round, sparsely puberulous, erect, spreading, ovate, somewhat obtuse, margins recurved, reticulated nerves, leathery, on both sides glabrous; petiole puberulous above lightly furrowed. Hoya microstemma Schlechter Leaves spreading or outspread, shortly petioled, elliptic, obtuse, wedge shaped base, fleshy, both sides glabrous. Hoya minahassae Schlechter loosely leafed; leaves petiolate, spreading, elliptic or lanceolate-elliptic, obtuse or almost obtuse, bases wedge-shaped fleshy, both aides glabrous, 4-7 cm long, in the middle almost 1.5 to 2.5 cm wide, petiole thick 7-10 mm long. Hoya mindanaensis Elmer

51 Leaves opposite, persistent, scattered, mainly descending, thickly coriaceous, yellowish green beneath, tips recurred, otherwise nearly flat or only the entire margins slightly imbricate, also glabrous and smooth, curing subolivaceus on both sides, oblong or more often ovately oblong, base broadly rounded and sometimes obscurely emarginate, the upper portion of the laminae gradually tapering into the acute to subacuminate apex, the average blades 15 cm long by 6 cm wide across the middle or a trifle below it, persistent; midrib thick and pronounced beneath clear into the apex and olivaceus in color, on the upper face deeply and narrowly grooved, totally glabrous on both surfaces; nerves faint, 3 to 5 on each side of the midrib, slightly ascending and straight, forked from above the middle and reticulately united, sometimes with secondary nerves in between, reticulations coarse and only visible from beneath; petioles up to 2 cm in length, blackish brown on my dried specimen, also thick, rugose, glabrous, calculate along the upper side, persistent, leaving a triangularly shaped scar after falling. Hoya mindorensis Schlechter loosely leaved, with the leaves outspread or spreading, oblanceolate-elliptic, shortly acuminate, glabrous, leathery textured 9-12 cm. long, above the middle cm. wide, with the petiole thick, somewhat terete (circular), cm. long. Hoya mindorensis subsp. superba Kloppenburg None. Hoya minima Costantin Leave glabrous, opposite, lanceolate, very small, thin, attenuate at the base, acuminate at the apex, wrinkled, reticulate in dry state; nerves not apparent except when light shines through; secondary nerves not visible, very oblique, towards the top, 4 to 5.5 cm long by 7 to 10 mm wide; petiole 5 to 5 mm, glabrous. Hoya mitrata Kerr Leaves oblanceolate or elliptic, base rounded and lightly eared, apexes briefly acuminate, margins flat, 6.5 to l2 cm long, 4.7 to 6 cm wide, leathery, dried gray, glabrous, above shining, beneath opaque, costa above concave, below prominent, lateral nerves 5-6 pairs towards margins very arching and 5-6 mm from margins anastomosing with network veining prominent on both sides; petioles 6-8 mm long, above concave, glabrous. Hoya monetteae Green Allied to Hoya diversifolia Blume but differing in that the foliage larger 20 cm. x 10 cm versus 5-10 cm. x cm.and flowers larger 2cm in diameter verses 1.3cm, and with clear sap versus milky. Hoya montana Schlechter Leaves erect-spreading or outspread, shortly petiolate, oblong-elliptic, shortly and somewhat obtusely acuminate, bases obtuse, somewhat fleshy, both sides glabrous, petioles very short, above lightly grooved. Hoya motoskei Teijsmann & Binnendijk (not from Type description). 49

52 leaves fleshy, ovate-rounded, acute, base slightly cordate, margins reflexed, glabrous above, pubescent below, with long petioles. Hoya mucronulata Warburg (not from Type description). leaves leathery oblong 6 to 12 cm, long, 3 to 5 cm. wide rounded on both ends, apex sometimes briefly, obliquely mucronulate, glabrous on both sides, above on midrib minutely glandular, veins on both sides 7 to oblique long, meeting and joining at the margins. Hoya multiflora Blume leaves somewhat leathery veined wedge-shaped oblong or oblong-lanceolate glabrous below paler, stiff hairs at base. Hoya myanmarica P. T. Li leaves opposite, succulent, oblong cm. x cm. base cuneate or rounded, apex acuminate; petioles 2 3 mm. long, apex with 2 or 3 glands, umbel terminal, 6-8 cm. long. Hoya myrmecopa Kleijn & van Donkelaar Leaves cm long, ovate to elliptic, acute to acuminate, venation obscure. Petioles cm long. Hoya nabawanensis Kloppenburg & Wiberg Opposite leaved, fleshy, petioles curved to slightly bent, terete, 5-6 times shorter than the leaf blade, 1 cm. long, glabrous. Blades ovate to ovate- lanceolate apex acute to somewhat apiculate, flat to slightly curved down at the apex, deep green, under-sides slightly lighter in color; glabrous 6-10 cm. long by cm. wide, pinnately veined about 4 on each side of the midrib at 45 degrees to it, midrib slightly raised on the upper side, edges entire, texture thick and leathery, sap milky white. Hoya naumanii Schlechter loosely leaved; leaves petiolate spreading outspread, broadly elliptic, widely apiculate, with the base somewhat cordate-rounded both sides glabrous, about 10 cm. long, near the middle about 7.5 cm. wide, petiole somewhat thickened, about 2 cm. long. Hoya neocaledonica Schlechter loosely leaved, with the leaves erect-spreading very outspread, ovate or ovateoblong acuminate, both sides glabrous, texture fleshy, 3 primary nerves very conspicuous above on the herbarium specimen (dried), petiolate, blade 5-9 cm long, in the middle and below the middle 3-5 cm. wide, petiole fleshy, round 1-2 cm. long. Hoya neoebudica Guillaumin leaves 3 to 4 together, lanceolate (7 to 9 cm. by cm.), base wedge-shaped, apexes acute acuminate, glabrous, fleshy, 5 ply nerves, petiole 1 to 1.5 cm. long. Hoya nervosa Tsiang et P. T. Li

53 Leaves large ovate or ovate-oblong 8.5 to 17 cm long 4 to 6.3 cm wide, apexes acute or shortly acuminate and bases rounded, costa nerves and veins very conspicuous; lateral nerves 5 to 6 pairs arching-ascending anastomosing next to margins veins reticulate, petiole strong 1 to 2 cm long with a few glands at apex, 2 to 3 together. Hoya nicholsoniae Mueller leaves ovate acuminate, somewhat fleshy, flat, 5-nerved, reticulate-veined; petiole often longer. Hoya nicobarica R. Brown ex Traill (I do not have original text) leaves ovate-lanceolate. Hoya nummularia Decaisne ex Hooker leaves very small orbicular shining. Hoya nummularioides Costantin Leaves fleshy, small, rounded or oval, but with the apex obtuse and apiculate, very pubescent on both surfaces; nerves not visible; blades 9-16 mm., by mm. long; petiole pubescent, 2-5 mm. long. Hoya nuuuliensis Kloppenburg & Siar Leaves, glabrous, thick wrinkled, elliptic, trinerved indistinct 6.8 cm. long, ca. 2 cm. wide, apex acute, base cuneate. Petiole glabrous, thick 1 cm. long (cannot see a groove above). Hoya nyhuusiae Kloppenburg None. Hoya obcordata Hooker leaves obcordate or obreniform, Leaves ½ in. long, sometimes broader than long, lobes rounded, sinus acute, base rounded or subacute, rather thin, opaque, nerves obscurely reticulate; petiole 1/16-1/15 in. Hoya oblanceolata Hooker leaves 4-6 in. narrowly oblanceolate acuminate very fleshy. Hoya oblongacutifolia Costantin Leaves glabrous, ovate-oblong, or lanceolate, pointed at base, acute or acuminate at apex, as a rule not very fleshy but nevertheless sometimes thick; secondary nerves 4-6 pairs, prominent, at 80 arching and running together as 1 nerve at margins. Leaves 5-8 cm by cm; petiole 4-8 mm long. Hoya obovata Decaisne leaves obovate apexes rounded or rarely emarginate, bases thick attenuate, towards the petioles a callused gland arising from the limb, margins reflexed, both sides glabrous. 51

54 Hoya obreniformis King leaves obcordate or obreniform, sometimes broader than long, lobes rounded, sinus acute, base rounded or subacute, rather thin, opaque, nerves obscurely reticulate, faintly visible above not evident beneath, bright-green above, paler beneath; length cm., width cm. Hoya obscura Elmer ex Merrill ex Burton Folia oppositae, glabra, laevis, stature variabilis, plane elliptica ad obovata-oblonga, in folia longiores, folia minores subelliptica 2 X 3 cm., aliae usque 10 cm. longa et fere 4 cm. late ad medium vel parve supra, gradatim contracta ad acuta-subacuainata apex, folia minores late obtuse ad basim, folia magnae cuneatae, interdum graciliter cuneatae omnes gradatum inter; costa non promines, laterales nervi 3-5 utrinque costa, admodum obliquus praecipue nervi basim, obscure furcati et plus minusve anastomosans ad margine; petioli 5-10 mm longi vel etiam usque 1.5 cm long), glabri, supra canneculeti. Hoya obscurinervia Merrill leaves opposite, fleshy, when dry leathery, pale, elliptic to oblong-elliptic, somewhat thin acuminate, bases for the most part acute, obscurely 3 or 5 plinerved, about 10 cm long, 4 to 4.5 cm wide, nerves thin, reticulations irregular; petiole thick, 2 to 2.5 cm long. Hoya obtusifolia Wight leaves fleshy oval-oblong bases of the apex obtuse cuspidate, above shining below transversely obscurely veined (dried). Hoya occlusa Ridley Leaves ovate acuminate in pairs, base obtuse glabrous above shining greenish when dry, below brownish, coriaceous, nerves 5 pairs rising at an acute angle, slender, elevated beneath, 4 inches long 2 inches wide, petiole 1 inch long. Hoya odetteae Kloppenburg Leaf Blade: l x 2-7 cm. Lanceolate. Matte midrib prominent, secondary veins obscure. Petiole:.25-1 cm. long. Hoya odorata Schlechter well leaved; leaves erect-spreading, short petioled, elliptic or lanceolate-elliptic, acuminate, glabrous, lucid, texture thinly leathery cm. long, below the middle cm. wide. Hoya oleoides Schlechter Leaves erect spreading or outspread, petiolate, elliptic obtusely acuminate, leathery, both sides glabrous, petiole short, somewhat thick, above lightly grooved.. Hoya oligantha Schlechter

55 Leaves erect-spreading or outspread, shortly petioled, narrowly lanceolate, minutely apiculate or somewhat acute, base rounded, leathery texture, both sides glabrous, petiole lightly grooved above, glabrous, very short. Hoya oligotricha K. Hill Leaves fleshy, broadly pinnately veined, elliptical, ovate or orbicular, 4-12 cm long, 3.0-8:0 cm wide; petioles cm long, 3-7 mm diam. Hoya onychoides P. I. Forster & Liddle Leaves petiolate; lamina lanceolate-ovate, up to 120 mm long and 55 mm wide, succulent, discolorous glabrous, with venation obscure, upper surface dark green; lower surface pale green; tip acute to shortly acuminate; base cordate; petiole mm long, 4-5 mm diameter, grooved on upper surface; colleters 3 or 4 at lamina base, often coalesced. Hoya opposita G. Don opposite leaves, the narrower petioles, which are convex beneath, and concave above. Hoya orbiculata Wallich ex Wight leaves fleshy orbicular glabrous. Hoya oreogena Kerr Leaves elliptic, base wedge-shaped or narrow slightly ear-shaped, apex somewhat obtuse margines plus or minus reflexed, 5 to 7 cm. long, 1.8 to 2.9 cm. wide, leathery, glabrous above, below with moderate copious short tawny pilose, when dry midrib obscure above bellow slightly prominent at the base narrowly winged. Lateral nerves obscure on both sides; petiole thick, warty, pilose, 5 to 8 mm. long. Inflorescence lateral, umbel-like, many-flowered: peduncles moderately robust, pilose 5 to 11 mm long. Hoya oreostemma Schlechter Leaves spreading or outspread, petiolate. lanceolate elliptic or elliptic, acuminate, base rounded, leathery texture, both sides glabrous, lucid above (shiny), below opaque, petiole glabrous, above slightly grooved. Hoya orientalis P. T. Li =Hoya reticulata Merrill Leaves fleshy when fresh, coriaceous when dry, ovate to oblong-ovate, shining, 8 to 12 cm long, 4 to 7 cm wide, the base broadly rounded, the apex slenderly and sharply acuminate, the epidermis on both surfaces coarsely reticulate-wrinkled when dry; petioles 1 to 2.5 cm long; lateral nerves about 4 on each side of the midrib, somewhat spreading. Hoya ottolanderi Koorders Leaves opposite very thick, 15 to 28 cm. long and 4 to 7.5 cm wide, glabrous distinct 5 nerved below above about 3 plinerved, lanceolate base acute, apex acuteacuminate. Petiole 1.5 to 3.5 cm long. 53

56 Hoya ovalifolia Wight & Arnott leaves fleshy ovate, acuminate at both ends, peduncles shorter than the leaves. Hoya oxycoccoides S. Moore Leaves bright green and faintly shining above when dry, opaque and pale below; petioles 3-5 mm. long, puberulous. Hoya pachyclada Kerr Leaves obovate, bases cuneate or somewhat rounded apexes very shortly and obtusely acuminate 7.8 to 10.5 cm. long, 5.2 to 6 cm. wide, thickly leathery, both surfaces having a few sprinkled short hairs, dried costa and nerves on both sides obscure but seen with transmitted light, lateral nerves 2 pairs and basal (or almost at the base) arising acutely, arching and ascending almost reaching apex; petioles thick, above concave, minutely puberulous, 6-12 mm. Hoya pachyphylla K. Lauterbach & K. Schumann leaves with short petioles elliptic, obtuse and apiculate with base rounded fleshy shiny both sides very glabrous. Hoya pachypus S. Moore. leaves petiolate broadly ovate-oblong sometimes almost square both ends rounded apex often mucronate thick leathery surface on both sides glabrous. Hoya padangensis Schlechter leaves spreading and widely spaced, narrowly elliptic-lanceolate, acute or acuminate, 7 to 10 cm long, in the middle almost 1.3 to 2 cm wide, fleshy, glabrous, petiole fleshy about 5 mm, long. Hoya palawanica Kloppenburg This species has a very distinctive leaf with venation similar to Hoya finlaysonii Wight with looping inner pairs anastomosing. Leaves spreading or outspread, petiolate lanceolate acuminate, texture leathery, with both surfaces glabrous cm. long, in the middle cm. wide, deciduous with 5 nerves, inner ones nearly reaching the apex with distinct reticulations. Petiole about 1.8 cm. long, 0.6 cm. in diameter, glabrous, grooved above. (from Schlechter). Hoya pallida Lindley leaves ovate-lanceolate acuminate fleshy veined. Hoya pallilimba Kleijn & van Donkelaar Leaves 3--8 cm long, ovate, acute, without obvious venation, margins on basal half of the leaves thin, pale green. Hoya panayensis Kloppenburg & Siar 2010 Leaves glabrous, elliptic-lanceolate, apex acute, pinnate nerved, base sub-obtuse to ovate. 54

57 Hoya panchoi Kloppenburg Leaves very thick, glabrous, with both surfaces minutely punctate, subdeciduous, edges straight and entire; of two distinct types, some round 1.5 cm. in diameter, others obovately oblong 2 cm. wide above the middle and 4 cm. in length, obtusely rounded at the apex, with a broad cuneate base. Petiole very short, thick, curved and expanded toward the base, leaving circular scars after falling. Hoya pandurata Tsiang Leaves erect spreading, a little fleshy, glabrous pandurate or narrowly oblong, margins involute, apex acuminate base rounded and in China very slenderly 2-eared 4 to 7 cm. long and 1 cm. wide (Wong's #74427, 10 cm. by 2 cm ), nerves on both sides not conspicuous, petiole glabrous about 4 mm. long apex glandular 2 to 4 fasicled. Hoya paniculata Sprengel Hoya leaves somewhat cordate ovate acuminate very glabrous. Hoya papillantha K. Schumann leaves petiolate, petioles thick round, leaves oblong acute bases somewhat rounded fleshy both sides very glabrous; umbels outside the axils on short peduncles. Hoya papuana Schlechter loosely leafed; leaves lanceolate-elliptic, long acuminate, both sides glabrous, texture thinly leathery, 6 to 8 cm long, below the middle 1 to 2.3 cm wide, petiole short, glabrous, cm long; rachis very short, about same length as petiole, cylindric. Hoya parasitica Wallich Leaves ovate-lanceolate, fleshy, drooping Leaves, opposite, petioled, retrofracted, ovate-lanceolate, acute, of a firm fleshy texture, and smooth on both sides, veins scarcely conspicuous above, and invisible underneath; from two to four inches long, and about one broad. Petiole short, round, ash coloured, in fact, more like a part of a branch than a petiole. Hoya parviflora Wight leaves fleshy lanceolate base and apex attenuate. Hoya parvifolia Schlechter loosely leaved, having aerial roots; with the leaves shortly petiolate oblong or oblong-elliptic, very obtuse, both surfaces densely and shortly hispid, cm. long, cm. wide, petiole hispid about 0.3 cm. long. Hoya patella Schlechter loosely leaved, round, minutely and sparsely pubescent. Leaves spreading or outspread, briefly petiolate, oblong-elliptic, somewhat obtusely acuminate, below pubescent, petiole pubescent, above slightly grooved. 55

58 Hoya pauciflora Wight leaves approximated, fleshy, glabrous, narrow lanceolate, blunt pointed; flowers few, generally paired, from a short thick peduncle. Hoya paxtonii Nichols Closely resembles the H. belle in habit and appearance, but has less fleshy, more acuminate and lighter green leaves. Hoya paziae Kloppenburg Foliage: held flat (surface to stem) at on twisted ½ turn petioles which are grooved above, also pale green 1 cm. long, 0.18 cm-0.20 cm. in diameter. a divided leaf gland at the base of the leaf tan in color, Midrib a little sunken an the upper surface, blade slightly concave on top edges slightly undulant, apex attenuate, narrowly acute and slightly turned under. Blade 6-7 cm. long and cm. at the widest below the middle. Pinnately veined, 5 pairs not prominent, surface glabrous, waxy, not punctate, very smooth above, granular below, midrib extending below, channeled above. New growth pink-bronze. Hoya pedunculata Schlechter In the leaf form it is midway between Hoya halophila Schlechter and Hoya litoralis Schlechter. Hoya pendula Wight & Arnott leaves fleshy glabrous from oblong-oval acute to broad ovate acuminate, margins revolute. Hoya pentaphlebia Merrill Leaves thickly coriaceous, pale and shining when dry, broadly ovate to elliptic, 8 to 16 cm long, 6 to 9 cm wide, the apex broadly and rather prominently acuminate, base rounded, prominently 5-nerved, the interior pair of nerves reaching the apex, the reticulations very lax, distinct on both surfaces, the margins somewhat recurved; petioles stout, 2 to 5 cm long. Hoya perakensis Ridley Leaves ovate acute coriaceous glabrous base broad truncate rounded 4 ½ inch long 3 inches wide, nerves from the base 5, with few arched secondary nerves, drying brown with recurved edges, petiole thick ¼ inch long. Hoya persicinicoronis He & Li 2008 Leaves alternate, opposite, whorled, leaflet pattern simple, terete, fleshy, blade similar to flat surface, elliptic or obovate, 6-10 cm. long, 4-5 cm. wide 0.03 cm. depth, reticulate, midrib and lateral vein evident or indistinct, lateral vein 3-5 pairs, ventral deep green, dorsal vivid greenish, base rounded, cuneate, apex rounded, cuspidate or retuse. Hoya phyllura O. Schwartz

59 Leaves opposite, blade oblong-elliptic or very slightly ovate-oblong, bases abruptly narrowed, apex long and acute caudate-acuminate, glabrous in the dry state coriaceous shining, nerves very prominent, nervation pinnate veins 3 to 4 on each side disappearing very near apex, curving and anastomosing, nerves narrowly reticulate. Leaf margins entire and very lightly revolute, 8 to 16 cm long, 3.2 to 5.2 cm wide, acumen 1.5 to 2 cm long, in the living state lighter green beneath than above. Petioles robust 1 to 1.7 cm long. Lateral inflorescence umbellate, supported by robust short peduncles 2.3 to 3 cm 2.3 to 3 cm long. Hoya picta Siebold leaves ovate-lanceolate, apex attenuate acute, fleshy, arched in the middle, yellowish and when young purplish or azure-rose and margines green variegate. Hoya picta Blume leaves (small) slightly round to ovate, sharply pointed; both sides purple-stained and glandular; below convex. Hoya picta Miquel Leaves on short petioles, broadly ovate mucronate-acute, both sides purple-dyed and glandular, in the dried state almost three nerved, beneath convex, 4-5 lines long. Hoya piestolepis Schlechter Leaves spreading or outspread, petiolate oblong or elliptic, acuminate, base briefly cordate, texture leathery, both sides glabrous. Hoya pilosa Seemann the leaves more hairy below. Hoya pimenteliana Kloppenburg foliage is opposite, linearly lanceolate with an acute apex and cuneate to slightly rounded base, where a distinct gland is present. The leaves are very distinctive, of a deep dull (flat color) iridescent green, due to an extremely fine velvety upper surface. Venation is somewhat obscure but pinnate with veins at 45 to the midrib (noted when dry). Leaf edges are entire but with slight undulations, the edges being slightly turned under. The foliage is profuse Leaf: 6-10 cm long; cm wide near the middle, apex acute to slightly apiculate, base cuneate to slightly rounded, gland at base, pinnate venation obscure on either surface, Upper surface very finely velvety and deep green, lower surface lighter green, dull with numerous lighter punctations. Leaves rather thick, but not succulent, somewhat leathery. Petiole: short, thick, glabrous, 0.08 cm. long, curved, terete, 0.05 cm. in diameter, not grooved. Hoya plicata King and Gamble Leaves fleshy coriaceous when dry; ovate-lanceolate, acuminate at apex, acute at base; smooth and glabrous on both surfaces; margins slightly recurved; 3 in. long, 1 in. broad; midrib very faint and then only when dry; remaining nerves invisible; petiole stout,.25 in. long, flattened. 57

60 Hoya polyneura Hooker leaves 3-4 in. subsessile rhombic-ovate or lanceolate acuminate very fleshy, nerves very many oblique straight parallel. Hoya polystachya Blume leaves broadly-ovate or oblong-ovate, acuminate, with base rounded or somewhat cordate; fleshy, thick 3-5 nerves, margins a little recurved. Hoya poolei White & Francis Leaf blades fleshy, lanceolate or elliptical, acuminate, venation indistinct in dried specimens, 6-9 cm. long, about thrice as long as broad. Hoya pottsii Traill The leaves are cordate, sharply, acuminate, having occasionally a rusty stairs partly spread over them, beneath of a pale whitish green without veins, above light yellowish green, with three distinct veins of ratter lighter colour than the leaf, from which smaller veins occasionally branch off. The petioles are not particularly thick, and less than half an inch long; they are of the same colour as the stems. Hoya praetorii Miquel leaves oblong or ovate-oblong or somewhat lanceolate acuminate, midrib on both sides with 5 outspread ascending veins prominent beneath, above somewhat grooved, 6 to 3 inches long. Hoya pruinosa Miquel Leaves (small) somewhat round above convex granular- punctate, beneath flat and glaucous-pruinose. Hoya pseudolanceolata Const, Leaves lanceolate attenuate, glabrous, 3-4 pairs of secondary nerves. 4-9 cm. long cm wide. Hoya pseudolittoralis Norman Leaves somewhat fleshy in those collected, red-brown ovate acute without nerves, 7 times 3 to 9 by 3 to 4.5 cm, with petiole very short and narrow. Hoya pseudomaxima Koorders foliage is profoundly cordate and from Hoya maxima Koorders leaves above glabrous and at the apex not at all or rarely emarginate. Leaves round-circular, at the base to 3 cm deeply-heart-shaped, on the top entirely glabrous (not only glabrous, but also without Cutiular-protruberances and polished, undersides glabrous and also without Cutiular-protruberances, to 8 cm long and 9 cm broad. Hoya pseudovalifolia Costantin

61 Leaves ovate-acuminate, fleshy; 4.2 to 6 cm long and 2.4 to 3 cm wide; petioles up to 3 mm long. Hoya pubens Costantin Leaves not Isis fleshy as the preceding species (parasitica) oval-oblong, rounded or often pointed at base shortly apiculate at the apex; veins 2 lateral, in addition to pairs departing at the base and joining the others, reappearing near apex, very strong and prominent on both sides. Three rib-net-work very apparent; leaf 9.5 to 13 cm long by 3.6 to 6.8 cm.; petiole very fleshy, thick, 2 cm long by 3 mm. Hoya puber Blume leaves (small) veinless oval-oblong acute fleshy puberulous. Hoya pubescens Reinecke leaves fleshy-coriaceous, flat, penninerved, oval or ovate, very shortly and abruptly acuminate, bases rounded, or somewhat cordate, petiole long, pubescent, blades on both sides shortly lanate. Hoya pubicalyx Merrill Leaves opposite, fleshy, when dry coriaceous, pale and shining on both surfaces and more or less wrinkled, oblong to oblong-obovate, 10 to 14 cm long, 3 to 6 cm wide, base obtuse, apex distinctly acuminate; lateral nerves pinnately arranged, indistinct, about 6 on each side of the midrib, ascending, scarcely more evident than are the lax, indistinct reticulations; petioles about 1 cm long. Hoya pubifera Elmer Leaves opposite, widely scattered, thick or leathery, curing grayish to yellowish brown, entire, the edges sometimes pressed with involute margins, narrowly oblong or oblanceolate to narrowly spathulate, velutinous beneath as well as densely or numerously papillose, the upper surface very similar but only slightly pubescent, the average blades 12 cm long by nearly 3 cm wide at or above the middle, the base attenuate, apex obtuse to subacute, a few of my detached leaves are only 6 cm long by 2 cm wide and with both ends obtusely rounded; midrib not conspicuous, lateral nerves none; petioles 1 to 3 cm long, dark yellowish brown, leaving circular scars upon falling. Peduncle stout, erect or nearly so, 4 cm long, similarly papillose and of a rich yellowish brown tomentum on my specimen. Hoya pulchella Schlechter loosely leaved, round, minutely pubescent. Leaves erect, spreading or outspread, petiolate, small, ovate, obtuse, slightly leathery, both sides glabrous, petiole minutely pubescent, above slightly grooved. Hoya purpurascens Teijsmann & Binnendijk leaves ovate, acuminate, base rounded, three nerved, above almost lacunose, reddish or purplish, margins revolute. 59

62 Hoya purpureofusca Hooker Leaves opposite, on very thick, brownish petioles, four or five inches long, exactly ovate, acute, or shortly acuminate, thick, fleshy, five-nerved, the nerves all diverging from the base, and having a gland at the base where set on to the petiole. Hoya pusilla Rintz Leaves very similar to those of H. lacunosa; fleshy, oblanceolate with acuminate bases and thickened margins; up to 8cm long by 3cm wide. Peduncle reflexed, rigid, up to 5cm long. Hoya pusilliflora S. Moore Leaves drying pale greenish grey, nervation well marked on the upper side, less clear on the on the lower; petioles 2-3 mm. long, stoutish, channeled above. Hoya pycnophylla Rechinger Leaves thick leathery, margines (when dry) not at all or barely revolute, opaque conspicuously 5 nerved, nerves prominently visible on both sides, shortly petioled. Petioles 1 cm long. Hoya querinoensis Kloppenburg & Siar Leaf blades: elliptic acuminate, quintuplinerved, glabrous cm. long x cm. at the widest in the middle. Petiole: cm. long curved, glabrous. Hoya quinquenervia Warburg coriaceous foliage, standing out, broadly ovate 5.6 cm. long by 4 cm. wide, apex shortly acuminate acute, base rounded, 5 nerved, the innermost about reaching the apex, reticulations subdistinct, edges revolute, equally green on each side. Petiole 1.5 cm. long, 0.3 cm. in diameter, glabrous. Petiole: curved to straight mostly cm. long, round not grooved above and merging with the leaf base. On the new growth there is a fine puberulous indumentum not present on older growth. Hoya quisumbingii Kloppenburg Leaf Blade: 8.f ctn. long; cm. at the widest, near the middle, tapering to a narrow base and with a long drawn out sharp somewhat rigid apex, which in many instances is curved a little downward; with pinnate venation 6-8 pairs on either side of the midrib area, which is more evident above, but also prominent below and on the Side with a few stiff hairs along its straight length from the petiole to the apex. Hoya radicalis Tsiang et P. T. Li Leaves fleshy, spatulate or oblanceolate 10 to 21 cm long 3 to 4 cm wide, apexes shortly acuminate or acute, bases cuneate running into the petioles margins noticeably narrowing all the way down, nervation on both sides conspicuous, nerves on both sides about 10 or more reticulate veins, petiole, when dried, almost flattened to 3 to 6 cm long, with obsolete glands. 60

63 Hoya ramosii Kloppenburg & Siar leaf blade is narrow not elliptic-lanceolate. Hoya recurvula Kloppenburg well leaved with pairs of lanceolate foliage, cm. long by ca. 3.5 cm. wide held mostly upright along the stem; flat to slightly curved, apex acute. Petiole, terete, at first bronze-green becoming corky with age, terete, curved, no groove, about 5 mm. in diameter. Hoya rasarioae Kloppenburg & Siar Foliage ovate apiculate to oblanceolate acute, base mostly cuneate, glabrous cm. long by cm. wide, nerve angle to midrib 44. Hoya reticulata Costantin Leaves oblong, in corner or rarely a little rounded at the base, attenuate progressively from the third lower portion, very sharp points at the apex (acuminate of 5-6 mm.), glabrous, thin, transparent; secondary nerves 9-10 pairs, prominent, almost winged underneath, at about 40, right, divided in regular dichotomies, coufluentes in arches of 4-5 mm. from the edges, very visible on the faces; visible tertiary and quaternary networks; limbe length of cm. by cm.; petiole length of cm. on 2 mm. Hoya reticulata Schlechter Leaves spreading or outspread, shortly petioled, elliptic, somewhat obtuse, fleshy coriaceous, both sides glabrous, reticulately nerved below, petiole short, lightly grooved above. Hoya reticulata Merrill Leaves fleshy when fresh, coriaceous when dry, ovate to oblong-ovate, shining, 8 to 12 cm long, 4 to 7 cm wide, the base broadly rounded, the apex slenderly and sharply acuminate, the epidermis on both surfaces coarsely reticulate-wrinkled when dry; petioles 1 to 2.5 cm long; lateral nerves about 4 on each side of the midrib, somewhat spreading. Hoya retusa Dalzell leaves shortly petioled, linear, three angled, apexes retuse, fleshy, pale Leaves 3-4 mm. long, 6-7 mm. wide, surface channeled above. Hoya retusa Warburg petioles 2 to 2.5 cm long, 2 mm thick, when young densely puberulous becoming glabrous, leaves leathery obovate 10 to 15 cm long, 7 to 10 cm wide bases rounded apexes retuse in sinus shortly apiculate, both sides glabrous, above on juvenile leaves pubescent, veins on both sides 5 to 6, spreading strait and long connected before reaching the margine indistinct, reticulations of nerves somewhat distinct. Hoya revolubilis Tsiang et P. T. Li

64 Leaves with similar features to H. liangii Tsang which differs by the rounded exterior angle of the corona; in this species the leaves are thick fleshy elliptic-oblong 6 9 cm long 2 3 cm wide with the apex obtuse base wedge-shaped or rounded with margines revolute above shiny green and below papilose when dry pale yellowish, midrib above flattened and below prominent nerve with veinlets vanishing, petiole cm long above on the petiole 2-3 glands. Hoya revoluta Wight ex Hooker Leaves 1 ½ - 3 in., midrib and nerves not visible, base acute; petiole 1/8-1/6 in., very thick. Hoya rhodostele Ridley Leaves thick; lanceolate acute, base cuneate: nerves invisible, 3 in. long, 1.5 in. wide; petioles thick.5 in. long. Peduncle axillary 4 in. long. Hoya rhodostemma Schlechter Leaves erect outspread, broadly elliptic, acuminate, base wedge shaped, leathery, both sides glabrous, above lucid, below opaque, petiole glabrous, above slightly grooved. Hoya ridleyi King & Gamble Leaves fleshy, coriaceous when dry, variable; ovate or ovate-oblong or oblong-lanceolate, acute at apex, rounded or narrowed or sometimes cuneate at base; both surfaces g1abrous; margins recurved; 3 to 5 in. long, 1.25 to 1.75 in. broad; midrib slender. Faint, as are the nerves generally, even when dry; main nerves about 5 to 6 pairs, the lowest pair starting from the base and curving along the margin, the other nearly at right angles, quickly branching and joining the marginal one; reticulations irregular netted; petiole very thick,.25 to.75 in. long, broad at the junction of the blade. Hoya rigida Kerr Leaves ovate or ovate-elliptic, bases rounded or broadly cuneate, apex long acutely acuminate, margins narrowly revolute, 12 to 14 cm. long, 4.5 to 6.3 cm. wide, when dry stiffly leathery, very hard, above black-brown, beneath dark, both sides opaque from the base 5-pli-nerved, nerves on both sides flat-convex, transverse nerves between prominent; petioles very thick, above flat-convex, 1.2 to 2 cm. long. Hoya rizaliana Kloppenburg The foliage is small, essentially ovoid, glabrous and enervis with a short petiole. Leaf Blade: 3.6 cm. x 1.2 cm., ovate tapering more toward the petiole, enervis, glabrous. Petiole: cm. long, round, glabrous. Hoya rosea Schumann leaves shortly petiolate, lanceolate, acuminate, bases acute, dried leathery, rugose, both sides glabrous..the petiole is 1 to 1.5 mm long; the leaf blade is 2.5 to 5 cm long and is 8 to 15 mm wide in the middle with veins that are clearly perceivable neither 62

65 on the upper or under side; when dried it is leather yellow bordering on greenish or brownish. Hoya rotundifolia Siebold elliptic-round leaves, those above are elusively suggestively-plain stained. They are thick, succulent, approximately l0 centimeters broad and long. Hoya rubida Schlechter loosely leaved; with leaves generally spreading, ovate or ovate-elliptic, shortly acuminate, both sides glabrous, texture thickly puberulous *,4-6 cm. long, 2-3 cm. wide below the middle, petiole glabrous above slightly grooved, about 1 cm long. Hoya rufolanata Ridley Leaves coriaceous; elliptic-ovate or obovate, acuminate at apex, cuneate or attenuate or rounded at base; upper surface shining, glabrous or slightly villous, pubescent on the midrib and main nerves; lower surface ferruginous-villous, especially on the nerves, dull; margins strongly recurred; 4 to 6 in. long, 2 to 3 in broad; midrib stout, impressed above, as are the 7 to 9 pairs of main nerves which, leaving the midrib at an angle of about 50, are at first straight and then curve upwards to join a rather obscure intramarginal looped nerve, also impressed; secondary nerves few; reticulations transverse, irregular; petiole stout,.75 to 1 in. long, densely ferruginous-tomentose. Hoya rumphii Blume leaves oblong-lanceolate acuminate, with the base very obtuse glabrous, below flat and pale. Hoya rupicola K. Hill Leaves extremely fleshy, obscurely broadly pinnately veined, narrow-ovate to ovate, acute, basally tapered, pale gray-green, 14 cm long, cm wide; petioles cm long, 6-10 mm diem. Hoya ruscifolia Decaisne leaves ovate mucronate mucrone very acute somewhat sessile without veins very glabrous, peduncles short receptacle ovate reduced. Hoya salweenica Tsang & Li Leaves fleshy elliptic or nearly oblong, above opaque green below more pale in the dried state pale yellowish 1 to 12.5 cm. long 3.5 to 4.5 mm wide, apex very obtuse or nearly round, base rounded or broadly wedge shaped, nerves 6 to 8 pairs, small veins disappearing, an inconspicuous gland above on the petiole, petiole thick to 1 cm. long. Hoya samoensis Seemann leaves ovate-elliptic acuminate 5-tuplinerved, nerves prominent on both sides. Hoya sana Bailey

66 Leaves oblong to oblong-lanceolate, from 1 ½ to 3 ¼ in. long, thick, pale in colour, only showing a rather broad midrib, more or less covered on the under side with white hairs, often aciculate, petioles seldom exceeding ½ in. Hoya sarcophylla Ridley Leaves thick leathery lanceolate with the base obtuse, nerves invisible, midrib on the upper surface at the base obscure 8.5 cm. long 3 cm. wide, petioles thick 8 mm. long. Hoya schallertiae Burton (Not a hoya) Leaves sessile (or nearly so), in opposite pairs, bases cordate or truncate, apexes rounded, broader than long ( ¾" X 1" ± ), margins erose-wavy, convex above, and lumpy, puberulous, scabrous beneath. Leaves flattened and oppressed against tree trunk, covering numerous roots. Hoya schneei Schlechter loosely leaved, round, glabrous, leaves spreading or erect outspread, petiolate, elliptic, acuminate, with a wedge shaped base, leathery glabrous, above lucid. Hoya scortechinii King & Gamble Leaves thick, fleshy, drying coriaceous, ovate or ovate-lanceolate, acuminate at apex, rounded or cordate at base, both surfaces glabrous, lower minutely glandular; Margins recurved, 2 to 3.5 in. long,.75 to 1.25 in. broad; nerves only visible when dry; midrib thick; main nerves 5 to 6 pairs at an angle of 50 to 60 with the midrib, branching early and anastomosing near the margin; reticulations transverse; petiole stout. Hoya serpens Hooker very small suborbicular opaque papillose on both surfaces.. Leaves ½ - ¾ in. diam., ovate or very broadly elliptic, base rounded or subcordate, nerves obscure; petiole very short, 1/16-1/10 in. Hoya shepherdii Short ex Hooker leaves 6 inches long and linear-lanceolate, curved downward, thickly-fleshy above dark green channeled, below more pale and semi-rounded, veins and petioles bent abruptly, petioles short round. Hoya siamaca Craib Leaves lanceolate, oblong-lanceolate or ovate lanceolate, apexes somewhat acute, bases obtuse, 3.7 to 9.5 cm. long, by 1.5 to 22 cm. wide, both sides glabrous, margins lightly recurved, when dried leathery, nerves entirely obscure; petioles 0.5 to 1.5 cm. long, glabrous. Hoya siariae Kloppenburg opposite lanceolate glabrous, deep green foliage ca. 14 cm. long x 3.5 cm. wide, tri-ply parallel veined, distinct on the upper surface less so below. Petioles heavy and long, 3.0 cm., diameter 0.09 cm. with deep groove on upper side. 64

67 Hoya silvatica Tsiang et P. T. Li Leaves papery to fleshy elliptic or elliptic-oblong above almost glabrous and beneath pubescent, dense near to the nerves, 8 to 11 cm long by 2.5 to 4 cm wide, apexes shortly acuminate bases cuneate, costa below lightly elevate, lateral nerves 7 to 10 pairs with veins conspicuously reticulate 2 or 3 glands above petiole, petiole 1 to 3 cm long, pubescent. Hoya sipitangensis Kloppenburg & Wiberg Leaves opposite, green with a few silver spots.. Leaves and flowers larger than Hoya lacunosa Blume. Hoya smithii Kloppenburg & Siar 2011 blades fleshy oblong, 8 to 12 cm. long 2 to 4.5 cm. wide, base obtuse, apex somewhat acute, penninerved, midrib above lightly impressed below prominent, secondary nerves on both sides 5 to 7 ascending often sometimes immersed, both sides prominent, vein sometimes prominent on both sides Hoya sogeriensis Moore leaves relatively large elliptic or elliptic-oblong shortly acuminate apex obtuse bases obtuse 5-nerved fleshy-leathery glabrous setting upon moderately long thickened petioles, peduncles strong much shorter than the leaves. Hoya solaniflora Schlechter Leaves spreading or outspread, petiolate, lanceolate, acuminate, texture thinly coriaceous, both sides glabrous, petiole glabrous, slightly grooved above. Hoya soligamiana Kloppenburg & Siar A vine with waxy, opposite elliptic-lanceolate leaves, prominently quintuplinerved, acuminate apex. Blades 9-13 cm. long x 2-3 cm. at the widest. Petiole round cm. long. Hoya sororia Schumann (1 cm. long) petioled leaves of 5 to 6 cm long and 1 cm wide. Hoya spartioides (Kuntz) Kloppenburg The reduced foliage (many times absent) and its prodigious production of green (chlorophyll) peduncles set this species apart as a fringe member of the genus. Hoya speciosa Decaisne leaves very broad ovate or ovate-elliptic with the base rounded apex shortly acuminate margins narrowly revolute both sides glabrous petiolate. Hoya stenophylla Schlechter Leaves spread or outspread, briefly petioled, narrowly linear, base rounded, margins commonly recurved, leathery, both sides glabrous, with the petiole pubescent, short, above slightly grooved. 65

68 Hoya stoneana Kloppenburg & Siar obtuse leaf bases not attenuata, and being pubescent not glabrous.. Foliage: linear lanceolate cm. long and cm. at the widest, waxy dark green, on both surfaces, lighter on underside. Dorsal surface with a few scattered hair cells and more along the edges, ventral surfaces covered with clear curved hair sells. Apex acute to apiculate base mostly obtuse. Petiole green colored pubescent mostly 1 cm. long and grooved above. Hoya subcalva Burkill Leaves ovate-elliptic, very shortly acuminate, bases rounded, 4 to 4.5 inches long 2 to 2.5 inches wide, glabrous, nerves on both sides moderately conspicuous; petioles 6 to 10 lines long. Hoya subglabra Schlechter Leaves spreading or outspread, petiolate, elliptic, acuminate, leathery textured, both sides glabrous, petiole glabrous, lightly grooved above. Hoya subquaterna Miquel leaves distinctly petiolate, opposite here and there almost shield shaped, rhomboid-ovate attenuate-acuminate, fleshy, veinless, petiole almost peltately inserted, 7-4 lines long. Hoya subquintuplinervis Miquel Leaves elliptic-rounded apex here obsoletely somewhat apiculate, fleshy, glabrous, midrib almost 5-plinerved on both sides traversed 2 almost rising out of the base on the other side ½ of the second. Hoya suregaoensis Kloppenburg & Nyhuus Hoya tamaleaaae Kloppenburg & Siar 2010 Leaves broadly ovate-elliptic, shortly attenuate a little rounded, base obtuse, venation pinnate 5 pairs anastomosing-reticulate cm. long, cm. at the widest near the middle, glabrous, midrib inconspicuous. Petiole 2.7 cm. long, terete, glabrous, definitely grooved above, glabrous. Hoya tauensis Kloppenburg & Siar Leaves glabrous, broadly ovate elliptic, edges undulate, glabrous, apex attenuate to apiculate acute, base obtuse, nerves 5 palmate visible on both surfaces central ones extend to near the apex, anastomosing, opposite; cm. long cm. widest near the middle. Petiole long medium heavy, glabrous, cm. long ca 0.15 cm, in diameter, same color (buff) as the stem. Hoya telosmoides Omlar Hoya tenggerensis Bakhuizen

69 leaves ovate-oblong base rounded apex short acuminate glabrous 3 to 5 nerves, within a limited area, middle nerves with 2 basal precursor nerves laterals many, with thickness and thicker nerves similarly at the base, when dry manifestly thick leathery above, below veins not as manifestly coarse, above at the base of the midrib provided with thick kidney shaped cells ( lacking? trachoma base) 10 to 12 long, petiole thick glabrous 1 cm long. Peduncles glabrous 1.5 to 4 cm long. Hoya tenuipes (see subsp of oligotricha and subsp of australis). Leaves fleshy or coriaceous, broadly pinnately veined, elliptical, narrow-ovate or narrow-obovate, 6-14 cm. long, cm. wide. Hoya teretifolia Griffith ex Hooker leaves 5-7 in. very slender cylindric quite terete. Hoya teysmanniana Miquel leaves at bases obtuse-rounded, somewhat obverse lanceolate-oblong (upwards a little wider) at the apex rounded or truncate or lightly emarginate, costa almost extended into a mucrone, thick leathery, veinless 5 3 inches long. Hoya thomsonii Hooker leaves beneath sparsely pilose, leaves 2-3 in. oblong or obovate-oblong acuminate very thick Leaves not very fleshy*, 2/3-1 ½ in. broad, midrib obscure, base obtuse or rounded, nerves very faint, arched; petiole ¼ - ½ in. Hoya tjadasmalangensis Bakhuizen leaves broadly ovate, base rounded, briefly acute and acuminate, glabrous. Not nerved above large warts when dry thickly leathery 19.5 cm long 22 cm wide. Hoya tjampeaensis Hochreutiner blades elliptic, apexes acuminate and acute, base attenuate acute, rarely a little rounded, margins entire, leathery but nevertheless nerves more or less prominent on both sides and conspicuous, middle nerves below very prominent, secondary nerves at the bottom small, oblique, meeting at the margins, over ¼ of blade long inconspicuous. Secondary nerves other than the principle about 6 to 7 on both sides, very irregular, more or less oblique or perpendicular and in this manner arching at an angle and noticeably irregularly anastomosing, i.e. more or less near margins or at a distance from the margins. Blade apparently glabrous, nevertheless very minute pilose very rarely adorned on both sides, mainly on the middle nerve. Hoya tomataensis Green and Kloppenburg foliage is thick, lanceolate ovate, obscure nerved, base narrow. opposite leaves and clear sap. Leaves are glossy deep green with a waxy surface, lanceolate-elliptic with a short apiculate apex, and hang essentially flat. The nerves are somewhat obscure but pinnate anastomosing the leaf base is rather long angustate. Typical leaf is 10.7 cm. long x 3.8 cm. at the widest near the center. The peduncles are curved, relatively long, terete and not grooved above 2.5 cm in length and 0.36 cm. in diameter. 67

70 Hoya torricellensis Schlechter Leaves spreading or outspread, elliptic, acuminate, leathery, glabrous, petiole glabrous, slightly grooved. Hoya treubiana Schlechter loosely leaved; with the leaves petiolate, oblong obtuse, glabrous, thickly leathery, nerves not at all conspicuous, with the margins revolute, cm. long, in the middle nearly cm. wide, petiole fleshy, cm. long. Hoya trigonolobos Schlechter loosely leaved, producing aerial roots; leaves spreading ovate or ovate-oblong, acute or briefly acuminate, both sides glabrous, texture fleshy-leathery, about 14 cm long, in the middle or below the middle about 7 cm. wide, petiole thick, short, cm. long; peduncle slender, round, glabrous about 6.5 cm. long. Hoya trinervis Trail leaves variable in size, oblong, sharply acuminate, pale green beneath, and mottled with small brown spots, light yellowish green above, with 3 prominent veins. Hoya trinervia (Hort.) Mach. ex Regal Smaller 3 5 nerved, but not veined leaves,.. Those of a short thick stalk support thick and leathery leaves, oval, before the pointed end, of 3 or 5 passed through under the long leaf apex anastomosing, the above side bright green, underside pale, glabrous throughout 3-4 inches long and 1 ½ 2 inches broad. Hoya trukensis Hosakawa Leaves outspread or erect-spreading, petioles about 1 cm long glabrous ovate, apexes acuminate or caudate-acuminate, bases rounded, papery, both sides glabrous, above lucid, 4 to 5.5 cm long by 2 to 2.5 cm wide. Hoya tsangii Burton ex Kloppenburg Leaves narrowly elliptic acute, 5-12 cm. long often 5 cm., near the middle cm. wide, on both sides with out nerves; petiole short, glabrous 1 cm. long +/- above lightly grooved. Hoya tsiangiana P. T. L Same as nom. nov. Hoya retusa Warb. Hoya tutuilensis Kloppenburg & Siar 2009 Leaves glabrous, broadly ovate-lanceolate, shortly attenuate, apex acute, base rounded but not cordate, cm. long cm. widest nearer the base, nervation triplinerved 5 distinct on both surfaces, same color as the leaf, secondary nerves some looping near the margins, primary nerves extend to near the apex. Petiole glabrous, not thin 1.5 cm. long probably grooved above, same color as the stem, circular gland at the attachment above. 68

71 Hoya tsoi Merrill leaves oblong or narrowly oblong-elliptic, leathery, when alive fleshy, 6 to 8 cm long, 1.5 to 2 cm wide, pale, on both sides somewhat narrow, base acute, apex mostly acute acuminate, primary nerves on both sides 4 to 5, spreading, obscure, bending, anastomosing; petioles thick 8 to 15 mm long. Hoya uncinata Teijsmann & Binnendijk leaves acuminate, with the base rounded, now and then narrowed, acute, fleshy.. Leaves fleshy 0.09 long wide, acuminate, with the base cuneate, above somewhat lacunose veined, below veinless, both sides with white dots. Leaves (sometimes with the exception of the thinly pubescent midrib) glabrous or subglabrous on both sides. Hoya upoluensis Reinecke leaves leathery, when dry papery, briefly petiolate, often oblique, ellipticlanceolate, distantly acuminate, narrow based, definitely quintuple nerved, nerves prominent on both sides; inflorescence umbel-shaped, peduncles many times longer than the petiole; pedicels slender, shorter than the peduncles. Hoya vacciniiflora Schwartz Leaves oblong-lanceolate both ends acute very glabrous leathery when living probably thick, margins entire one-nerved, nerve impressed above, prominent below, veins throughout frequently uniting. Leaves shortly petioled petioles thick, channeled above, 0.5 to 1 cm long, blade 5.5- to 8.5 cm long and cm wide. Hoya vaccinioides Hooker leaves ½ in. elliptic subacute fleshy nerves obsolete.. leaves diverse when young ovate-orbiculatis, apex briefly acute, later elliptic lanceolate acute, base margine entire, other abbreviated, thick variegation. Hoya variegata Siebold ex Morren leaves diverse, juvenile ones ovate-orbicular, apex short acute, becoming elliptic lanceolate acute, base margine one, the other abbreviated, thick variegated. Hoya variegata De Vriese leaves white-variegated, margins often rosy fleshy, oval, elliptic oblong, somewhat acuminate, petioles reddish, 2 to 5 cm. long. Hoya variifolia Ridley leaves fleshy-leathery dimorphic elliptic obtuse, bases narrow, 1.5 cm. long, 5 mm. wide, and linear-lanceolate, both ends acuminate 4 cm. long, 2 mm. wide, petiole scarcely distinct; umbels 18-flowered; peduncles slender 2 cm. long. Hoya venusta Schlechter

72 long. Leaves 3-5 cm. long, below the middle cm. wide, petioles cm. Hoya verticillata (Vahl) G. Don leaves 4 in a whorl, petiolate, oblong, acuminated, acute, fleshy, glabrous, veinless. Hoya villosa Constantin Leaves oblong, elliptic, rounded at the basis and at the summit sometimes very slightly echancrees or very weekly mucronate, densely villose, yellow or rose colored underneath; a little villose on top; nerves four pairs of laterals, oblique, arched toward the edge, invisible underneath; limbe length of 7-11 cm. x cm.; petiole villous or glabrous, length of cm., 4 mm thick. Hoya vitellina Blume leaves oval oblong acute with base rounded or obtuse fleshy above veined, below no veins. Hoya vitellinoides Bakhuizen Brink f leaves oblong base shortly obtuse and acutely acuminate on both surfaces very sparsely and very shortly pilose thickly coriaceous, penninerved, lateral nerves 4 to 6 parts on both surfaces conspicuously transverse all over by irregular veins without (?) trichomes 14 to 16 cm long, 7 to 7.5 cm wide, petiole thick 7.5 to 10 mm long. Hoya vitiensis Turrill Leaves elliptic-oblong or elliptic-ovate, with apex attenuate acuminate, base rounded or almost cordate, up to 10 cm. long and 7 cm. wide, entire, glabrous lateral nerves on both sides about 7 almost to the margins anastomosing when dry with the midrib flat prominent on both sides. Reticulations prominent in a transverse manner; petiole 0.5 to 1.5 cm. long, thick, glabrous. Hoya volubilis Griffith leaves ovate-lanceolate acute leathery glaucous at the base one callus, indistinctly 3 nerved. Hoya wallichiana Decaisne leaves broad ovate oval or obovate with rounded base or obsoletely emarginate apex short acuminate, with the limb arising from transverse veins transversely thickened and somewhat fleshy, below velvety with middle nerve very prominent glabrous, petiole scarcely 1 inch, thick. Hoya walliniana Kloppenburg & Nyhuus None Hoya warburgii Schlechter. Unpublished but herbarium sheet and drawing. 70

73 Hoya wariana Schlechter Leaves spreading or somewhat outspread, shortly petioled, strap shaped, linear, acute or apiculate, base obtuse, thin leathery texture, both sides glabrous. Hoya wayetii Kloppenburg Leaf Blade: cm. long x cm. wide, thick, succulent, convex to nearly flat on the upper glabrous surface, triangular below and lighter green covered with granulose lighter colored punctations. Upper surface deep green with deeper green edging especially in younger leaves. Some rusty tones may be exhibited in newly formed leaves. Blades tapering to both ends, especially narrow toward the petiole which merges with the blade, leaf gland very small almost non-existent. Hoya waymaniae Kloppenburg Leaf Blade: thick, rigid, succulent, both surfaces finely pubescent especially so with the lower surface. 5-9 cm. long, margins very undulant 3-5 cm wide, enervis, edges rough with small bumpy thickened protrusions, much pinkish pigmentation. with a linear imbedded leaf gland present. Hoya weebella Kloppenburg Leaves: opposite, midrib prominent below, stem rusty, apparently no leaf gland, a few trichomes along the midrib, few hairs on lower midrib end of blade, upper surface rugose, base rounded (obtuse) tapering (lanceolate) to a narrowly rounded apex cm. long and widest near the base 0.09 cm. apex with small apiculation. Blade dark green underside glabrous, edges slightly recurved, thin. Above medium green, slight groove down the center above midrib, edges entire, below granulose punctate, glabrous. Petiole: curved, hirsute, hairs pointing toward the leaf blade, ca cm. long, clean 0.30 cm. in diameter. Hoya whistlerii Kloppenburg opposite, thick lanceolate leaves. The longest leaves are cm. long, mostly with acuminate apices. Hoya wibergiae Kloppenburg leaves opposite tend to be long narrowly elliptic curved along length apex acute to apiculate 6.5 cm cm. long x ca. 4.2 cm wide, but some smaller and broader, triplinerved with faint anastomosing, only primary veins and midrib visible below surfaces glabrous, margins sharp and slightly curved under, blade not flat. Leaf upper surface with splotchy silvering, petiole terete, corky. Hoya wightiana Thwaites leaves glabrous, fleshy, linear-lanceolate, acuminate, obtuse, with narrow base. Hoya wightii Hooker Leaves 3-4 long, long petioled elliptic or elliptic-lanceolate rarely orbicular acute at both ends thick, margins recurved, nerved distinct horizontal or nearly so Leaves broad, margins often revolute, tip obtuse often produced, nervs loosely netted; petiole

74 Hoya wightii ssp. plainensis Matthew Leaves broad-elliptic x cm., margins faintly reflexed; lateral nerves at an acute angle 9c. 45 ) to the midrib Hoya wrayi King & Gamble Leaves fleshy, thinly coriaceous when dry; lanceolate, obtuse and glandular-apiculate at apex, long-attenuate at base; upper surface glabrous smooth, lower minutely papillose, both grayish when dry; margins recurved; 3.5 to 4 in. long, 1 to 1.25 in. broad; midrib slender, broadened at base; main nerves very inconspicuous even when dry, about 4 pairs, at about 30 with the midrib; reticulations not visible; petiole very thick, rugose, jointed at the junction with the blade. Hoya yuennanensis Hand- Mazzetti Leaves almost circular continuously elliptic, 6 to 11 cm long, somewhat continuously longitudinal (rarely) doubly narrow.. briefly and obtusely apiculate, base broadly rounder truncate, very thick, flat, dry light green, both sides with twisted hairs, below when young almost velvety, below 5 to 7 nerves diverging at about 60 from the midrib straight to near the margins barely curved connecting loosely with the reticulate veins when dry prominent on both sides, at the base of the midrib is a yellowish gland; petioles long 3 to 7 mm, 2 to 3 mm wide densely hairy. Peduncles axillary, 2 to 3.5 cm long. Hoya zollingeriana Miquel Leaves with short petioles and bases obtuse, on the upper dorsal side of the petiole somewhat drawn out, elliptic or largely obovate-oblong, or obtuse-apiculate, thick leathery, beneath on both sides 4-5 obsolete veins (when dry, somewhat prominent) in pairs, inches long. 72

75 Hoya Leaves from Holotype Sheets Taken from Herbarium Type sheets (or from the Type plant). Copied and cropped to isolate one or a few leaves. Species and author, date of publication and type number, Collection location, followed by leaf shape, size and nerve angle with the midrib. Hoya acicularis Green & Kloppenburg, Type, Danum Valley, Sabah, Eastern Malaysia; Green No , (BISH) 1/24/91 from Tenom Orchid Center. needle-like,, 0.3 cm. x cm., 1. Hoya aeschynanthoides Schlechter. Borneo: on trees at Long Dett in the state of Koetei. Densely leaved; with leaves shortly petioled, broad ovate-elliptic, acute, pilose (felt like) on both sides, cm. long, nearly cm. wide, petiole hispid cm. long. Hoya alagensis (Schlechter) Kloppenburg 1990, Type 5524 (B). Elevation 150m. Collected by E D. Merrill November 1906 on the Alag River, Mindanao, Philippines. E Ac, 10 x ,

76 Hoya aldrichii Hemsley No type listed. Picture from #94 (SING) leaves being 3 to 5 nerved longitudinally, all of them natives of the Malayan region. Captain Maclear collected the same plant, but without flowers. O Ac, 8-12 cm. long, cm. wide, 32. Hoya amblyoepsis Schlechter (unp.) Type 4545 (B). Collected on Palawan, Philippines May 1906 by H. M. Curran. Ob E Ob, x 2.5-5, 22. Hoya amoena Bakhuizen Typus 20341b (L). Collected 19 Oct from Besoeki, Java, Indonesia. Elevation 10m. E Ac, 8-11 x ,

77 Hoya andalensis Kloppenburg 2005, Typus #20051 (UC). Typus #20051 (UC). Collected by E. de Vogel? J. Vernaden at Padang, Air Sirah in a young secondary forest, Sumatra. Leaves opposite small, cupped below, ob-ovate 5.0 x 2.2 cm. Base cuneate. Hoya angustisepala Schltr. Ex Elmer) ex Burton 1987, Type (B) a synonym of H. coriacea Blume 1826, Collected by A. E. D. Elmer in June 1909 on Mt. Apo, Mindanao, Philippines. O E Ac, Also at (US), Anastomosing. Hoya anncajanoae Kloppenburg & Siar. Typus (CAHUP) collected May 1991 Elevation m Bicol National Park, Luzon, Philippines. R Ob, x

78 Hoya angustisepala Schlechter Type of Hoya mindorensis Elmer. Isotype (L). Collected June 1909 on Mt. Apo, Mindanao, Philippine. Later described by Burton 1987 and subsequently placed into Synonymy as H. coriacea Blume 2001 by Kloppenburg. E Ac, x 4-6, 57. Hoya anulata Schlechter 1902, Type (B). Collected 4 February 1902 by Schlechter at Ramu, New Guinea. Elevation 400m. E Ob, x ,

79 Hoya archboldiana C. Norman 1937 Type 3621 (NY). Collected by L. J. Brass April 1933, Rona, New Guinea. Elevation 450m. EC Ap, x , 60. Hoya attenuata Christophersen Type 2164 Christophersen UC). Savaii: forests above Letui, altitude 1000 meters, flowers, September 27, 1929, Christophersen no x x x 2.8 cm. 40?. Hoya aurantiaca Kloppenburg & Siar 2009, Type (CAHUP).Oblanceolate acute, triplinerved, x 3-5, 10. Hoya australis subsp nathalieae Kloppenburg & Siar, Type (CAHUP). Collected by Nathalie Evelensdaughter at the Morsby Botanical Garden, New Guinea in R C Ap, 6.5 x

80 Hoya bandanensis Schlechter 1908, Type (B). Collected 8 October 1901 at Banda-Archipel, Molucca. E Ac, x 5-7. Hoya benguetensis Schlechter 1906, Type 5979 (B). Collected by A. E. D. Elmer March 1904 at Baguio, Benguit, Luzon, Philippines. O E Ac, 6-10 x 2.5-4,

81 Hoya benvergarai Kloppenburg & Siar 2008 Typus (CAHUP). Collected by Siar & Cajano. Elevation 30m. E L Ac, 9-13 x 2-3, 15. Anastomosing, quintuplinerved. Hoya betchei (Schlechter) Whistler, Type s. n. (B). Collected by E Betche 11 December E Ac, x , 45. Hoya bhutanica Grieson & Long Collected at S. Bhutan: Hastisar, 460 m, In dense forest. Ludlow, Sherriff & Hicks (holo. BM). Leaves elliptic (6.5) 9-15 x (2.75) cm., apex acute base rounded, trinerved, lateral veins produced ½ above, slightly leathery, glabrous, margined entire. 79

82 Hoya bicknellii Kloppenburg, Type 1470 (UC), ex hort. Collected by David Bicknell at Matutinao, (Kawasan) Bidian Cebu, Philippines. O Ap, 9-12 x 4.5-6, 35. Hoya bicolor Kloppenburg 2003, Typus (CAHUP) collected by Renaldo Pimentel from Mt. Banahaw, Quezon Province, Luzon, Philippines. E Ac, x , 42. Hoya bilobata Schlechter Isotype 420 (L) collected by Copeland on Mindanao, Davao Dist., Philippines, March R, 1.5 x 1.5, 21. Type at (B) 80

83 Hoya bordenii Schlechter 1906, Type 1213 (B). Collected by T. E. Borden June 1904 at Lamao river Mt. Merivales, Bataan, Luzon, Philippines. Elevation 650m. L Ac, x , 15. Hoya brevialata Kleijn & van Donkelaar Type IPPS 8836 (L), Indonesia, Central Sulawesi. Province: Tentena, in clove plantation. O Ac, cm. x cm, pinnate,

84 Hoya brittonii Kloppenburg Type (PNH). Collected by B. B. Britton 5 May 1953 at Badoc, Calapan, Mindoro, Philippines. Elevation 200m, Ob L Ac, x , 24. Hoya buotii Kloppenburg, Typus (CAHUP). Collected from Mt. Banahaw, Quezon Province, Philippines. E Ro, x 2-3, 52. Hoya burtoniae Kloppenburg Type s. n. (UC). Collected by Loher in Rizal Prov, Luzon. Philippines Sept O, 3-4 x

85 Hoya buruensis Miquel Type s. n. (L). Collected Moluccas, Indonesia. E Ro, 8-11 x 4-5.2, 25. Hoya cagayanensis (Schlechter) Burton 1987, Type 7374 (B). Collected by M. Ramos at Cagayan, Luzon, Philippines March 1909.E Ac, 8 x 3, 30. Also at (US) 34. Hoya callistophylla Green Type 201 ex hort. (BISH). Collected From Nabawan, Sabah, Eastern Malaysia. Elliptic to lanceolate, 10-20cm long x 6-l0cm wide x l mm thick, matte surface, light green with dark green veins, lateral veins approx.60 to midvein; edge hard, irregular and sharp. Pinnate

86 Hoya calycina Schlechter 1908, Type (B). Collected by Schlechter 26 March 1905 Kani Mts. At 1100m elevation.. O C Ac, 9-11 x 3-3.5, 60. Hoya campanulata Blume. Type s. n. (L). Collected at Burangarang, Indonesia. E Ro, x 3-5, 45. Hoya cardiophylla Merrill 1920, type Dinagat, Bur. Sci Ramos & Pascasio, May 13, 1919, on tree trunks at low altitudes. 5 to 7 cm. long. 4.5 to 5 cm wide, pale when dry, glabrous or the younger ones slightly ciliate near the basal margins, the base broadly rounded and distinctly cordate

87 Hoya carrii Forster & Liddle. Type (NY) Collected by C.E. Carr in Paua New Guinea. Altitude 1376m x 2-3 mm. EL Ac º. Hoya caudata Hooker f Isotype 1128 (L). Collected in Malaya by Maingay. Distributed to Kew O Ob Ac, 9-13 x 5-7, 42. Hoya cebuensis Kloppenburg & Siar, Type XXXX (CAHUP). Collected by David Bicknell at Matutenao, Bidian, Cebu, Philippines. Elevation 10m. O L Ac, 9 x 4. 85

88 Hoya cembra (H. leucantha Schltr. Unpublished) Kloppenburg 1990, Type 5650 (B). Collected by E. D. Merrill on the Alag River, Mindoro, Philippines Nov Elevation 300m. L Ro, 8-10 x 2-3, 73. Also at (US), 63. Hoya chlorantha Rechinger Holotype # 1011 (BISH). Collected on the Island of Upolu, in the forest near Tiavi climbing high up in the trees. May 1905 Elliptic acute. 10 cm x 2 cm., 40. Hoya chlorantha var. tutuilensis Christophersen type1011, type in B. P. Bishop Museum; scrub forest, top of Le Pioa, altitude 500 meters. Elliptic lanceolate acute, agreeing in characters of the leaf with the type of the variety

89 Hoya chloroleuca Schlechter 1913, Type (B). Collected 22 September 1909 by Schlechter in Torricelli Mts. NE New Guinea. Elevation 800m. E Ac, 6-8 x 2-2.3, 32. Hoya clandestina Blume (Under the H. macrophylla heading.) except for the corona structure, easily confused with a species hear-to-for unpublished, H. (Physostemma) clandestina, which I found in each mountain locality in Java and possessed in like manner with similar foliage nerves x cm. L Ac, 23º. Hoya collina Schlechter 1913, Type (B). Collected by Schlechter in August 1908 In the Finisterre Mts., New Guinea. Elevation 400m. E OE Ob, x Hoya cumingiana Decaisne Type 1480 (L) collected by H. Cuming in Philippines. R E Ac, x

90 Hoya coriacea Blume 1826 Type s. n. (L) Labeled first as Hoya fraterna Bl. Herb. Lugd. Batav. E Ac, x , 40. Hoya crassicaulis Elmer ex Kloppenburg. Isotype (UC). Collected by A. E. D. Elmer at Mt. Buluson, Sorsegon, Luzon, Philippines, October Elevation 76m. E Ac,

91 Hoya crassipes Turczaninow Type Zoll. coll. n From the Island of Java. Twining, leaves petiolate elliptic or obovate, the base attenuate, briefly acuminate, fleshy veinless glabrous. RO Ap, x , 42. Hoya darwinii Loher Holotypus (UC) collected by A. Loher Sept 1909 at Rizal, Luzon, Philippines. Ob L Ac, x 2-3, 72. Hoya davidcummingii Kloppenburg 1995, Type: #9 (BISH); isotype (UC), discovered by David Cumming 1.5 Km. from Lake Bulusan turn off towards the lake at the forest edge, when coming from Irosin. Elevation ca Elliptic acute, veins not visible but pinnate, cm. long, widest about the middle cm. 89

92 Hoya deykei Green Type 9912 ex hort (BISH). Collected from Atjeh Barat, Sumatra, Indonesia. truncate to emarginate leaves and ovate. 5-12cm long x 5-8cm wide x 2-2.5mm thick, matte surface. 45. Hoya diptera Seemann 1861, type 320 Viti Levu and Taviuni, on trees, Fiji. Ovate-elliptic acute cm. long, (1-) cm. broad Hoya dischorensis Schlechter 1913, Type (B). Collected by Schlechter in the Dischore Mts., Wari region, NE New Guinea in June Elevation 900m. O E Ac, x 2.8-5,

93 Hoya diversifolia Blume Type s. n. (L) Collected in Java. E Ap, 7-8 x 4.3-5, 45. Hoya diversifolia Blume Type s. n. (L) Collected in Java. This is listed in Leiden under Hoya kuhlii Koorders in The original label says Hoya diversifolia Blume by Blume from Java. There are 4 stems on sheet and it seems to me 2 species. I have shown the 2 leaf types here. E Ac, x , 45. Hoya dolichosparte Schlechter 1916, Type (B). Collected by Schlechter January 1910 in Sulawesi near Tolitoli. Elevation 10m. O R Ob, x 6.5-9,

94 Hoya edanoi C. M. Burton 1990 type (A). rotund obtuse. 40. Hoya eitapensis Schlechter 1909, Type (B). Collected by Schlechter 11 August 1909 at Eitape, NE New Guinea. Elevation 20m.. E Ob, 3-5 x 1.2-2, 31. Hoya elmeri Schlechter unpublished Type (B). collected by A. E. D. Elmer at Benguet, Baguio, Luzon, Philippines March O E Ac, 9 x 3.5, 31. Hoya epedunculata Schlechter 1913, Type (B) Collected by Schlechter 8 Sept in Torricelli Mts. NE New Guinea. Elevation 900m. E Ap, x ,

95 Hoya erythrina Rintz Type 117 (L). Collected by Rintz 27 Sept at Sungai, Teranum, Pahang, Malaysia. Elevation 610m. L Ac, x 2.3-4, 12. Hoya exilis Schlechter 1913, Type (B). Collected by Schlechter in June 1907 at Albo. NE New Guinea. Elevation 300m. E L Ac (Ro), 5-10 x 1.5 3, 56. Hoya faoensis Kloppenburg & Siar, Type 5714 (HAW). Collected by Art Whistler on Mt. Fao, Upolu, Samoa. Elevation 500m. O E Ap, 4-5 x 2.2. anastomosing,

96 Hoya fetuana Kloppenburg, Type W2765 (HAW). Collected by Dr. Art Whistler 5 July 1975at Tutuila, Samoa. Elevation 500m. E At, x 1.2-2, 43. Hoya fitchii Kloppenburg & Siar collected by Charles Marsden Fitch in 1988, Luzon, Philippines. Foliage is quintuplinerved, with the second pair of nerves nearly reaching the apex, outer connecting nerves anastomosing Nervation white or lighter green than the leaf blade color. 30. Hoya flavescens Schlechter 1913, Type (B) Collected by Schlechter 28 April 1908 at Kani Mts, NE New Guinea. Elevation 800m. L Ob Ac, x , 40. Hoya forbesii King & Gamble 1903, Type 2896 a (K). Collected by H. O. Forbes at Kaba Volcano, Sumatra in Elevation 1524m. E Ac, x ,

97 Hoya fraterna Blume 1849 twining, glabrous; leaves leathery ovate-oblong elliptic acuminate base rounded somewhat veined above the petiole callous-glandulose. Notation: The drawings in both Botanical Magazine and Fleur des Serres are not of Hoya Fraterna. A mistake due to Lobb's deception of collection locations. There is no type mentioned nor designated in the Type description. The leaf here is taken from a 1910 sheet s.n. (L) designated as the Type.? 10.5 x cm. 42. Hoya fungii Merrill leaves leathery, penninerved, elliptic to oblong, 8 to 14 cm. long, 4.5 to 8 cm. wide, when dry pale, opaque, both sides glabrous uniform color or below a little paler, apexes acute and shortly acuminate, base rounded, midrib thick, below strongly elevated, lateral nerves on both sides about 7, not at all evident, arched meeting at the margins, about 30 angle with the midrib, reticulations loose, obscure; petiole 1.5 to 3 cm. long, glabrous. Hoya gigantangensis Kloppenburg 1992, Type #36787 PNH collected by Yoshio Kondo & G. Edano 26 March 1957, at Gigantangan, Leyte, Philippines. Elliptic acute cm. x cm., pinnate venation, both surfaces glabrous

98 Hoya gildingii Kloppenburg 2001, type Typus ex hort. Eva Karina Wiberg, Borlänge, Sweden, ex hort (BISH), From Mt. Kota Kinabalu, Sabah Malaysia (Bukit Tapai Trail) Lanceolate elliptic, 85. Hoya glabra Schlechter 1908, Type (B). Collected by Schlechter in August 1901 at Long-Wahau, Borneo. O Ac, x ,

99 Hoya gracilipis Schlechter 1908 (not type) (B) Collected by Schlechter in Torricelli Mts., NE New Guinea. Elevation 1100m E L Ac, x 2-3, 67. Hoya gracilis Schlechter 1908 Type s.n. (B). Collected by Sammlers (a native) in June 1886 at Talisso, Celebes (Sulawesi). L E Ac, x , 18. Hoya greenii Kloppenburg Holotype BISH. #7. collected Mt. Apo, Mindanao, Philippines at 4000 elevation, on trail to hot springs lake cm. long x 3-4 cm. wide, narrowly elliptic

100 Hoya hainanensis Merrill Typus 9750 (UC). Collected by McClure May 1922, Hainan, China. RE Ac, 31. Hoya halconensis (Schltr.) Kloppenburg 1990 Type 5674 (B). Collected by E. D. Merrill on Mt. Halcon, Mindoro, Philippines. Elevation 900m, E Ac, 8-12 x , 35. Hoya halophila Schlechter 1913, Type (B). Collected by Schlechter in August 1909 on the beach at Eitape, NE New Guinea. Elevation 10m. R O Ap, x ,

101 Hoya heuschkeliana Kloppenburg 1989, Type 2175 (CAHUP) collected Luzon (northern portion of Lake Bulusan, Sorsogon, Philippines), epiphytic on trees at ca. 50 m alt. ovate-elliptic obtuse, base wedge-shaped. 2.5 cm. long x cm. wide at the middle, 56. Hoya heuschkeliana subsp. cajanoae Kloppenburg & Siar Type (CAHUP), ovate x cm. glabrous. 56. Hoya hyploasia Schlechter 1913, Type (B). Collected by Schlechter in July 1908 at Wabe river in the Finisterre Mts. N. New Guinea. Elevation 400m. L Ac x , 55. Hoya incurvula Schlechter 1916, Type (B). Collected by Schlechter in Minahassa, Sulawesi in November Elevation 800m. R E Ob, x

102 Hoya intermedia A. C. Smith. Type 399 (UC). Collected in Thankaunirove, Fiji 12 Nov Elevation 10m. OE Ac, Also at (US) Hoya intermedia Schlechter an unpublished species. Leaf blades: thin many in close clusters 6-10 cm. mostly 7.5 cm. long. glabrous, narrowly elliptic, thick base narrowing into petiole, apex acute long tapering to sub rounded. Waxy rounded leaf edge.petiole cm long, thin. There are sheets at (BO, UC, PNH) #75500 labeled as this sp but they are not the same as the type (misidentified). Hoya ischnopus Schlechter 1913 Type (B). Collected by Schlechter in August 1909 at Bulu NE New Guinea. Elevation 10m. O E Ac, 9-11 x 5-6,

103 Hoya juannguoana Kloppenburg 2002, type #2001 ex hort. Fresno, CA., USA. (UC). Collected in 1994 at St. Paul s Underground River, (Sabang Bay area), Palawan, Philippines x5-6.5 cm. Elliptic apiculate, 43. Hoya kastbergii Kloppenburg 2003,. Holotypus (UC), collected at Bau, south West from Kuching, Sarawak, Borneo, elevation of about 300 meters Rotund obtuse. Hoya kenejiana Schlechter 1913, Type (B). Collected by Schlechter at Kenejia, NE New Guinea, on October Elevation 150m. O E Ob, 6-10 x Hoya krohniana Kloppenburg & Siar Type (CAHUP) Leaf from type plant. R O C Ap, 1.5 x 1,

104 Hoya kuhlii Koorders (also labeled Hoya rumphii var. Collected in Java (Herb. Lugd. Batav.) First as Dischidia longifolia (crossed out). Type s.n. (L). L Ac, x 2-4.2, Hoya kuhlii Koorders s. n. (L) First labeled Acanthostemma Blume by Wight from Java. Later as Hoya rumphii Blume β kuhlii and in 1947 as H. kuhlii by Backhuizon. It should be (Bl,) Koorders. L Ac, 12 x 3.3,

105 Hoya lacunosa Blume Type s. n. (L) collected in Java Herb. Ludg. Batav. O Ac, x 1.5-2, Hoya lambii Green Holotype: Bishop Museum, Ted Green No 9905 ex hort. Ted Green of Kaaawa, Hawaii. From Sabah, Eastern Malaysia, Elliptic, 10-15cm long x 6-10cm wide. 50 looping. Hoya lancilimba Merrill Collected at Hainan, Hung Mo Shan, Ip Yulc Shing 18261, August 23, 1929, on the north slope of the mountain. leaves lanceolate to oblong-lanceolate, flashy, leathery 11 to 14 cm. long 2.5 to 3.5 cm. wide, both ends somewhat equally narrow, base acute, apex acute acuminate, in the dry state pale, wrinkled, with few primary nerves on either side, spreading, obscure arching and anastomosing; petioles thick, 2 to 3 cm. long. 103

106 Hoya latifolia G. Don 1838, type 138a Native of Penang. Leaves 7-10 in, long, and 4-10 broad. Broad-leaved Hoya. 33. Hoya leucorhoda Schlechter Type (UC). 14 June 1908 In Keiser Wilhamsland, New Guinea. Elevation 1200m OS Ro, 40. Holotype at (B). Hoya liangii Tsiang Collected at Hainan: Nagi Yuen, H. Y Liang (Type), Sept. 3, 1933; Lingshui, C. F. How 73935m Oct. 21, Leaves fleshy obovate or obovate-oblong 4.5 to 7.5 cm. long 3 to 4.5 cm. wide, apex rounded or obtuse, base somewhat cordate or rounded, above shiny green below dull when dry pale yellow, lateral nerves 4 in pairs, veins faint, above the petiole 2 to 5 glands, petiole thick 1.5 cm. long

107 Hoya linavergarae Kloppenburg & Siar 2006, Type (CAHUP). Collected in Isabella Province, Luzon, Philippines. Foliage opposite, glabrous blade 4-5 x cm. Hoya littoralis Schlechter 1905, Type (B). Collected by Schlechter in October 1901 at Keiser Wilhelmsland, New Guinea. Elevation low. E L Ac, x Hoya loheri Kloppenburg 1991, Type s. n. (UC) Collected by A Loher March 1916 in Paningtingan, Rizal, Luzon, Philippines. E Ob. Hoya loyceandrewsiana Green 1994.Type 9301, (BISH), cm long cm wide, triplinerved,

108 Hoya luzonica Schlechter 1904, Type (B). Collected by Dr. Warburg on Lamo River, Mt. Mariveles, Luzon, Philippines. O OB-Ac, 9-13 x 5-6, 47. Same as H. meliflua. Tsing x cm. Hoya lyi Leveille Collected at Koay ToHedu; environs de Gan-Pin 20 Septembre Lion Matin et J. Seguin No Tsien-Sen-kiao Novembre 1904 No (Sean Ly). No type is specifically designated. The sheet at (NY) is Y. Hoya macgregorii Schlechter 1906, Type 191 (B). Collected by McGregor April- May 1905 on Baco River, Mindanao, Philippines, L E Ac,.5-3.5,

109 Hoya macrophylla Blume Type s.n. (L). Also labeled Hoya clandestina Blume. LE Ac, x , 18. Hoya marginata Schlechter 1905, type (B). Oct elevation 200m Neu-Pommern Ovate acuminate, 5-7 nerved 13-17cm long, in the middle cm. broad,

110 Hoya matavanuensis Kloppenburg & Siar 2008, Type 9539 (HAW). Collected by Art Whistler 9 May 1994 On Savaii, Western Samoa. Elevation 650m. E Ac-Ro, x 1-1.5, 48. Hoya mcgregorii Schlechter 1906, Type 191 (B), April- May 1905, Baco River, Mindoro, Philippines. Leaves lanceolate-elliptic, acuminate, glabrous coriaceous; 7-11 cm. long cm. wide in the middle, triplinerved, 20. Hoya memoria Kloppenburg 2004, Type (UC). Collected from Memorial Gdns. Manila, Philippines, From they type plant. O L Ac. Hoya mengtzeensis Mengtze, Taitang, in N. Y. Bot. Gard., isotype Tsiang & Li Collected at Yunnan: elevated-over rocks, A. Henry Type Arn. Arb. and merotype Kwangtung Bot. 108

111 Inst. Foliage thinner than general, lanceolate, apexes acuminate, base attenuate and running down into the petiole 6-11 cm long, cm broad, midrib impressed above, nerves and veinlets when dried rugose, petioles 1-2 cm long provided with 2-3 small bunched glands at apex. 23. Hoya meredithii Green 1989, Type Meredith (BISH), iso (NY) The foliage is rather rigid, the edge undulant, 3 to 7 inches wide and nearly 12 inches long. The pinnate reticulate venation is deep dark green. 36. Hoya merrillii Schlechter 1904, Type 2218 (B). Collected BY E. D. Merrill at Polo, Mindro, Philippines in May 1903 (Type sheet says May 1898). O Ac, 6-9 x 5-6.5, 34. Hoya microphylla Schlechter 1913, Type (B). Collected in November 1908 in the Bismark Mts. Elevation 1100M NE New Guinea. E O Ob, x Hoya microstemma Schlechter 1913, Type (B). Collected by Schlechter in the Torricelli Mts. NE New Guinea, elevation 800m, in September L E Ob, x ,

112 Hoya minahassae Schlechter 1916, Type (B). Collected by Schlechter near Tomohon, Sulawesi, elevation 800m. in November L Ac, 3.5-7,5 x Hoya mindorensis Schlechter 1906, Type 332 (B). Collected by R. C. McGregor April-may 1905 in Mindoro, Baco River, Philippines. L Ap, 9-12 x , 33. Hoya mitrata Kerr 1940, Type Kerr Thailand (Siam). Surat, Ban Tong Tao, ca. 10 m., in evergreen forest, Leaves oblanceolate or elliptic, base rounded and lightly eared, apexes briefly acuminate, margins flat, 6.5 to l2 cm long, 4.7 to 6 cm wide,

113 Hoya montana Schlechter 1913, Type (B). Collected by Schlechter in The Kani Mts. NE New Guinea, June 1908, elevation 600m. OB E Ac x 3.5-5, 48. Hoya multiflora Blume Type 354 (L) 2 sheets with same number. Collected in Tjibodas, Indonesis. Herb. Lugd. Batav. L Ac, 7 x 3, LE Ro, x ,

114 Hoya nabawanensis Kloppenburg & Wiberg, Type 2002 (UC), here from type plant. Collected by Wiberg et al at Nabawan, Sabah, Malaysia. O L Ac, 6-10 x Hoya naumanii Schlechter 1908, Type 1896 (B). Collected by Naumann on Bougainville, Solomon Islands, (Now New Guinea) 26 August 1875, E Ap, 10 x 7.5, 46. Hoya neo-caledonica Schlechter, 1906 Type (B). Collected by Schlechter in December 1902 in the forest of Berge near Oubatche, New Caledonia. Elevation 600m. O Ac, 5-9 x 3-5,

115 Hoya nervosa Tsiang et P. T. Li Collected at Yunnan: Szemao, large climber on tree, fls. white, A. Henry (Typus! in N. Y. Bot. Gard. and merotype in Kwangtung Bot. Inst.). Leaves large ovate or ovate-oblong 8.5 to 17 cm long 4 to 6.3 cm wide, apexes acute or shortly acuminate and bases rounded, costa nerves and veins very conspicuous; lateral nerves 5 to 6 pairs arching-ascending anastomosing next to margins veins reticulate, petiole strong 1 to 2 cm long with a few glands at apex, 2 to 3 together. 58. Hoya nuuulensis Kloppenburg & Siar, Type 8505 (HAW). Collected by Dr. Art Whistler at Nu uuli, Tutuila, Samoa 16 April Elevation 1m. L Acute, 6.8 x Hoya nyhuusiae Kloppenburg 2003, type 2003 ex hort. (UC), collected by Torill Nyhuus at Mt. Kinabalu, Sabah, Malaya. Lanceolate acute. Hoya obscura Elmer ex Merrill ex Burton Type (L) collected by A.E. D. Elmer July 1916 at Trosin, Mt. Bulusan, Sorsegon, Luzon, Philippines. Ob Ac, 7 x Also at (UC). 113

116 Hoya obscurinervia Merrill Type 8919 (UC). Collected by McClure 1919 in Hainan, China. E Ac. 25. Hoya obtusifolia Wight Lectotype HB 1170 (U). Also labeled Hoya teysmanniana Miquel Collected in Sumatra, Indonesis (Sitabah). E Ob, x 3-5,

117 Hoya odetteae Kloppenburg 1998, Type 5604 (BISH); isotype (UC); (PNH). Collected by Odette Cumming at Patin-ay 1 Bridge, Mindanao, Philippines. : l x 2-7 cm. Lanceolate. Matte midrib prominent, secondary veins obscure. Hoya odorata Schlechter 1906, Type 3202 (B). Collected by A. E. D. Elmer at Mt. Meraveles, Baatan, Luzon, Philippines in January Elevation 1300m. L E Ac, x , 68. Hoya oligantha Schlechter 1913, Type (B). Collected by Schlechter in the Torricelli Mts. NE New Guinea in September Elevation 800m. E L Ap- sub Ac, x Hoya oleoides Schlechter 1913, Type (B). Collected by Schlechter in Dischore Mts. NE New Guinea in June Elevation 1300m. E Ac, x , 51. Hoya opposita G. Don 1837 collected in the East Indies by Vahl Type 72 (C), elliptic-lanceolate apex rostrate, base somewhat cuneate, pinnate anastomosing, typically 9.5 cm. long and 3.5 cm. at the widest near the center. 7 side nerves netted and looping, midribs not prominent. Glabrous all over

118 Hoya oreostemma Schlechter 1913, Type (B). Collected By Schlechter At Keiser Willhemsland NE New Guinea in November Elevation 1300m.( says 1000m on Type sheet) L E AC, x , 54. Hoya ovalifolia Wight & Arnott Wight. catalogue #1522. glabrous, leaves fleshy ovate, acuminate at both ends. LE Ac, x , 36º. Hoya padangensis Schlechter 1916, Type (B). Collected by Schlechter at Padang, Pandjang, Sumatra, Indonesia January 1907, Elevation 900m. L Ac. 7-8 x 1.6-2,

119 Hoya palawanica (Schlechter) Kloppenburg Type 834 (B). Collected by F, W. Foxworthy in Mangrove swamps, Palawan, Philippines L Ac, 14 x 3.5-4, 29, anastomosing. Hoya pallilimba Kleijn & van Donkelaar, 1999, Type IPPS 8864 (L ), collected by Ted Green, Morowali Reserve, near mouth of Ranu River, Sulawesi. Leaves 3--8 cm long, ovate, acute, ca. 54. Hoya panayensis Kloppenburg & Siar 2010, Type (BO) isotype (UC). Collected by McGregor in May, Antigue Prov., Panay, Philippines. L Ac ( E Ac),

120 Hoya panchoi Kloppenburg Type (L) labeled Hoya diversifolia Elmer Isotype of Hoya bulusanensis Elmer Collected on Mt. Bulusan, Sorsogon, Luzon, Philippines May Elevation 475m. R, 1.3 x 1.1, & O Ac, 2.5 x leaf types. Hoya pandurata Tsiang Collected at Yunnan: Sze-Mao, A. Henry l2258 (Type in Herb. N. Y. Bot. Gar., Merotype and Analytical Drawing in Herb. Suny. Univ.. Leaves erect spreading, a little fleshy, glabrous pandurate or narrowly oblong, margins involute, apex acuminate base rounded and in China very slenderly 2-eared 4 to 7 cm. long and 1 cm. wide. L Ro, x 1.3, Sheets at (BO, UC. PNH). Hoya papuana Schlechter 1913, Type (B). Collected by Schlechter in the Toricelli Mts. NE New Guinea on 24 April Elevation m. 6-8 x 1-2.3, L E Ac. Hoya parvifolia Schlechter 1908, Type (B). Collected by Schlechter 9 June 1901, at Danau Kotta, Sumatra, Indonesia, O E Ob, x Hoya patella Schlechter 1913, Type (UC). Collected by Schlechter August 1907 at Wobbe, New Guinea. Elevation 300m. LE Ac. Also at (B) x ,

121 Hoya paziae Kloppenburg 1990, Type 5618 (B), L Ac, x , 900m, 50. Also at (US), 63. Hoya pentaphlebia Merrill Typus (L). Collected March 1914 Samar, Philippines by M Ramos. O E Ac, x Hoya piestolepis Schlechter 1913, Type (B). Collected by Schlechter on Gomadjidji in the Waria Valley NE New Guinea 8 May Elevation 450m. Ob-E Ac, x 5.5-7,

122 Hoya pimenteliana Kloppenburg sp. Nov. Typus #96034 UC. Type material from original collected plant. Original plant collected by Blas Hernaez at Casiguran, Quezon Province, Luzon, Philippines. E Ac : 6-10 cm. long; cm wide; 30. Hoya polystachya Blume Type s.n. (L). Collected by Blume in Java, Indonesia. O C Ac, x 6-8,

123 Hoya poolei White & Francis 1928, Type Plate V., fig.2.). Leaf blades fleshy, lanceolate or elliptical, acuminate, venation indistinct in dried specimens, 6-9 cm. long, about thrice as long as broad. Hoya phyllura Swartz Type 339 (K). Collected by Hans Winkler 24 November 1924 In Hara, West Borneo. Elevation 150m. E Ro, 52. Hoya pseudolittoralis Norman 1937, type 5990, collected Feb. 23, 1934, at Dagwa, Oriomo River, Western Division, British New Guinea, alt. 40 m. Type in British Museum (Natural History); duplicate in Britton Herbarium. New York Botanical Garden. Leaves ovata acuta sine nervis, 7 x 3-9 x 4.5 cm.,

124 Hoya pubifera Elmer1938. Isotype (L). Collected by Elmer August 1916, Mt. Bulusan, Sorsogon, Luzon, Philippines.LE Ac, x 2-3, 42. Also at (US). Hoya pulchella Schlechter 1913, Type (B). Collected by Schlechter 8 September 1909 in the Toricelli Mts. NE New Guinea. Elevation 900m. O Ob, x , 43. Hoya purpureofusca Hooker 1850, type Tab. 4520, four or five inches long, exactly ovate, acute, or shortly acuminate, thick, fleshy, five-nerved, the nerves all diverging from the base,

125 Hoya pusilla Rintz Lectotype 55 (L). Collected by Rintz 4 June 1976 at Sangai Tahan, Malaysia. Elevation 500m. L E Ac, x 1-2.2, 41. Hoya querinoensis Kloppenburg & Siar Typus (CAHUP) collected by Maximo Wayet, Baguio Village, Querino 15 Sept Elevation 150m. L Ac, x ,

126 Hoya quisumbingii Kloppenburg Typus (PNH). Co0llected by Quisumbing at Itbayat Is., Philippines. L, x 1.8-2, 35. Hoya ramosii Kloppenburg & Siar. Type 7550 (UC) as Hoya intermedia Schlechter Collected by Ramos & Edano in Catanduanes, Luzon, Philippines. l Ac. 124

127 Hoya recurvula Kloppenburg 2000, Type #81009 (UC) ex hort. Leaves lanceolate, cm. long by ca. 3.5 cm. wide. 39. Hoya retusa Dalzell 1852, type not indicated. Leaves 3-4 mm. long, 6-7 mm. wide, surface channeled above 125

128 Hoya revoluta Wight ex Hooker 1883, type Kew Distrib. 1127, Collected at Malacca, Maingay. Leaves 1 ½ - 3 in. ovate or ovatelanceolate. 60. Hoya rhodostele Ridley 1923, Type not mentioned.. Leaves thick; lanceolate acute, base cuneate: nerves invisible, 3 in. long, 1.5 in. wide. 42. Hoya rhodostemma Schlechter 1913, Type Collected by Schlechter on 30 May 1909 in the Dischore Mts, NE New Guinea. Elevation 1200m. E Ac, x , 46. glabrous. Hoya rizaliana Kloppenburg 1991, type sheet s.n. (UC). A. Loher collected at Montalban, Rizal Province, Luzon, 3.6 cm. x 1.2 cm., ovate tapering more toward the petiole, enervis, 126

129 Hoya rosarioae Kloppenburg & Siar, 2008, Type 5268 (CAHUP). Collected by Mazimo Wyatt, at Baguio Villiage, Benguet Prov., Luzon, Philippines. O Ap, x 2-3.4, 44. Hoya rubida Schlechter 1901, Type (B). Collected on 5 November 1901 between Massawa and Cape Lambert, New Pommerania. Elevation Om. O-OE Ac, 4-6 x 2-3, 36. Hoya salweenica Tsang & Li. 1974, type Yü (Typus! In Herb. Inst. Bot. Kwangtung. Serv. Leaves elliptic or nearly oblong 1 to 12.5 cm. long 3.5 to 4.5 mm wide. 37. Hoya schneei Schlechter 1921 s. n. (B). Collected by Dr. Schnee, January 1909 in Ponapae. L Ac, x 3 5, 35. Hoya serpens Hooker 1883, type not mentioned, Leaves ½ - ¾ in. diam., ovate or very broadly elliptic, base rounded or subcordate, nerves obscure. 127

130 Hoya shepherdii Short ex Hooker 1861, type Tab. 5148, gathered by Drs. Hooker and Thomson in Sikkim-Himalaya, at elevations of 3000 to 4000 feet, and in Khasya. Leaves from two to (mostly) six inches long, not more than four lines wide. Hoya siariae Kloppenburg 2002 Typus (CAHUP). Collected by Reymold Pimentel from Tayabas, Quezon, Luzon, Philippines., 15 March E Ac, x ,

131 Hoya sigillatis Green 2004, type (BISH) ex hort, collected Tenom Agricultural Park, Tenom, Sabah, Malaysia. Leaves elliptic with obtuse to cuneate base and acuminate tip cm long by cm wide. Hoya silvatica Tsiang et P. T. Li Collected at Yunnan: Gung-Shan, alt m. C. W. Wang (Typus! in Herb. Inst. Bot. Kwangtung, servatus). Also (A, GRAY) Leaves papery to fleshy elliptic or elliptic-oblong above almost glabrous and beneath pubescent, dense near to the nerves, 8 to 11 cm long by 2.5 to 4 cm wide, apexes shortly acuminate bases cuneate, costa below lightly elevate, lateral nerves 7 to 10 pairs with veins conspicuously reticulate 2 or 3 glands above petiole, petiole 1 to 3 cm long, pubescent. 40. Hoya solaniflora Schlechter 1913 Type (UC). Collected by Schlechter in 6 September 1908 In Finisterre Mountains, New Guinea. Elevation 1200m. L Ac, x Also at (B),

132 Hoya soligamiana Kloppenburg & Siar holotypus #69916 (CAHUP). Collected at Atimonian, Quezon, Luzon Philippines (at the foot of Mt. Banahaw).Ovato-ellipticus or ellipticus 9 13 cm. x 2 3 cm., apice acuminate, glabrous. NE New Guinea, l Ap, x 0.4. Hoya stenophylla Schlechter 1913, Type (B). Elevation 800m. Collected by Schlechter in 6 September 1909 at Toricelli. Mts, Hoya stoneae Kloppenburg & Siar ex Hort. Type (CAHUP). Leaf from Type plant. Linear lanceolate Ac, x Hoya subglabra Schlechter 1913, Type (B). Collected by Schlechter July 1908, upper Maijen NE New Guinea. Elevation 450m. E Ac, x ,

133 Hoya subquintuplinervis Miquel 1869, type not designated. Collected Siam, prope Radboeri, ubi Tjang-Kemoi Leaves 3 3/4-3 inches long, 3-2 inches wide. 38. Hoya tamaleaanus Kloppenburg, Type W2705 (HAW). Collected by Dr. Art Whistler 27 June 1975 on trail from Fagasa Pass & Matafas, Tutuila, Samoa. Elevation 350m. O Ac- Ro, x 2.5-3, 49. Hoya tauensis Kloppenburg & Siar. Collected by Dr. Art Whistler at 15 Jan Ta'u, Samoa. Leaves glabrous, broadly ovate elliptic, edges undulate, glabrous, apex attenuate to apiculate acute, base obtuse, nerves 5 palmate visible on both surfaces central ones extend to near the apex, anastomosing, opposite; cm. long cm. widest near the middle. Petiole long medium heavy, glabrous, cm. long ca 0.15 cm, in diameter, same color (buff) as the stem. O E Ap

134 Hoya tenggerensis Bakhauizon f Holotype 301 (U). Collecter 9 August 1907 at Tengger, Gunung, Java, Indonesia. O CAc, x 4-5.5, Hoya tomataensis Green and Kloppenburg 2006, type (BISH). Collected in dry gulley on road to Tomato to Kolonadale Central Sualwesi. Leaf 10.7 cm. long x 3.8 cm. at the widest. 33. Hoya tjadasmalangensis Bakh. f Type 448β (L). Collected 25 October 1919, Malang, Java, Indonesia. First listed as Hoya macrophylla Blume in 1919 collected by W. F. Winckel. O Ac, 18 x 11,

135 Hoya torricellensis Schlechter 1913, Type (B). Collected by Schlechter 22 September 1909 at Torricelli Mts. NE New Guinea. Elevation 800m. E Ac, 9-14 x 4-4.3, 53. Hoya treubiana Schlechter 1908, Type s. n. (B). Collected by Schlechter in the Torricelli Mts. September Elevation 800m. E Ac, 9-14 x Hoya trigonolobus Schlechter 1905, Type (B). Collected by Schlechter 5 November 1901 near Madres, New Pomerania. Elevation 50m. O E Acute, 14 x 7,

136 Hoya tsangii Burton ex Kloppenburg Isotype (L). Collected August 1912 by A. E. D. Elmer on Mt. Cabadbaran, Mindanao, Philippines. l L, AC, x Hoya tsoi Merrill Collected at Kwangtung: Sup-man-ta Shan, C. L. Tso 23455, July 29, 1933, altitude about 360 m. L Ac x º Hoya tutuilensis Kloppenburg & Siar 2008, Type 9089 (HAW). Collected by Art Whistler 5 January 1993 at Tutuila, Samoa. Elevation 180m. O L Ac, x 3-3.5, 30. Hoya uncinata Teijsmann & Binnendijk 1863, type Mt Pantjar (W); below 800; forest. Leaves fleshy 0.09 long wide, acuminate, with the base cuneate, above somewhat lacunose veined. 134

137 Hoya venusta Schlechter 1913, Type (B). Colected by Schlechter 23 November 1908 in the Bismark Mts. NE New Guinea. Elevation 2200m. L-EL Ob-Ac, 3-5 x , 49. Hoya verticillata (Vahl) G. Don 1837, type 72 (C). Native of the East Indies This leaf measures 7.5 cm. long and in the middle 3.1 cm. wide with a slight edge rolled under Hoya vitellina Blume 1849 Type s. n. (L) labeled as collected by Blume I Java, Indonesia. L E Ac, 13.5 x

138 Hoya vitellinoides Bakhuizen Brink f. 1950, type 4181 (L). Java Western part, Batavia, Mt. Tjipoetih, near Tjiampea, alt. 800m. Leaves 14 to 16 cm long, 7 to 7.5 cm wide. 50. Hoya warburgii Schlechter Unpublished, Type (B). R O Ap, x 4.9-6, 42. Hoya wariana Schlechter 1913, Type (B). Collected by Schlechter 11 March 1908 in Decheregi in the Waria Valley, NE New Guinea. Elevation 400m. l 0b-Ac, x

139 Hoya wayetii Kloppenburg 1993, type 1989 (CAHUP) Iso (UC), collected by Maximo Wayet Baguio, Benguet Prov. Luzon, Philippines Leaf Blade: cm. long x cm. wide, thick, succulent, convex to nearly flat on the upper glabrous surface, Hoya weebella Kloppenburg 2005, type (UC), base rounded (obtuse) tapering (lanceolate) to a narrowly rounded apex cm. long and widest near the base 0.09 cm. apex with small apiculation. Hoya wibergiae Kloppenburg 2001, Typus (UC). Collected by E. Wibergii at Estrella waterfall area Palawan Is., Philippines in O Ac, x

140 Hoya wightii Hooker Collected at Deccan Peninsula; from the Bombay Ghats to the Nilgherry Mts. There are 6 sheets at (US) Type Hoya, n. 27, Herb. Ind. Or. H. f. & T. (NY). 5-6 x º. Hoya whistlerii Kloppenburg 2002, Isotype 3139 (B) Holotype (BISH). O Ap, x ,